identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E5ED0AFF9ED56F2B996A19FB60FA55.text	03E5ED0AFF9ED56F2B996A19FB60FA55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megarthrus chiapas Rodríguez & Navarrete-Heredia & Arriaga-Varela & Cuccodoro 2020	<div><p>Megarthrus chiapas Cuccodoro sp nov.</p><p>(Figs 1-26, 39)</p><p>http://zoobank.org:act: 1F284006-11C5-404A-8E68-E5C7668672EA</p><p>Type material. (7♂, 5♀). Holotype ♂: labeled MEXICO: Chiapas, 8 mi [= 12.9 km] N Pueblo Nuevo Solistahuacán, 6000’ [= 1830 m], 26-27.VIII.1973, A. Newton, berlese cloud forest litter, in FMNH . Paratypes (11): labaled with the same data as holotype, 2♂ and 1♀ in FMNH, 1 ♂ &amp; 1♀ in MHNG and 1♂ in CZUG; labeled MEXICO: Chiapas, 8.9 km E Rayón, 1500m, 19.IX.1991, R. Anderson, 91-109, ex cloud forest litter, 1♂ and 2♀ in SEMC, 1 ♂ and 1♀ in MHNG .</p><p>Description. Habitus as in Figs 1–3. Combined length of head, pronotum and elytra = 1.3 - 1.4 mm; maximal pronotal width = 0.9 - 1.0 mm. Body chestnut brown, with lateral margins of pronotum and appendages slightly paler; 1, 2 or 3 yellowish apical antennomeres. Dorsal pubescence denser on head and pronotum than on elytral disc; medial frontal setae directed forward; elytral and pronotal setae almost straight, recumbent, becoming slightly denser along medial groove of pronotum and on anterior portion of elytral disc; pubescence on abdomen uniform, parallel, without macrosetae on sternites IV–VII. Frons, pronotum, humeral area of elytral and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granules about as high as their diameter, or higher; elytral disc and metaventrite coarsely granulo-fossulate.</p><p>Frons above clypeus forming sharp ridge, the latter finely carinate, evenly; mesal portion of disc moderately convex in lateral view, evenly; anterior margin in dorsal view weakly arcuate in middle and laterally oblique; Ushaped frontal impression shallow. Temples convex, granulate. Occipital ridge indistinct. Antennae (Fig. 9) 1.8-2.0 times longer than pronotum.</p><p>Pronotum (Figs 25-26) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edges, shallowly depressed along anterior and posterior margins; medial groove deep, parallel-sided, slightly arcu- ate in lateral view; hypomera granulate, with ridge extended from anterior margin to laterobasal angle (Fig. 26). Proventrite (Fig. 26) lacking medial ridge. Scutellum with anterior margin rounded; posterior margin laterally mod- erately arcuate toward strongly arcuate apex. Elytra gradually widened posteriorly (Fig. 1); disc with humeral callus strongly convex, two moderate swellings along anterior and posterior portions of suture, and moderately depressed posteriorly along lateral edge; the latter finely carinate, not denticulate, slightly arcuate in dorsal view.</p><p>Male: Frontoclypeal area not modified. Protarsomere 1 lacking ventral patch of modified adhesive setae. Mesofemora (Fig. 5) as long as metafemora. Mesotibiae (Fig. 6) shorter than metatibiae (Fig. 7), the latter straight. Metatarsomeres 1 about 1.3 times as long as combined length of metatarsomeres 2-4. Peg-like setae present only on mesotrochanters (Fig. 5; arranged in a row), on distal two thirds of mesotibiae (Fig. 6; arranged in 1-2 rows) and on distal half of metatibiae (Fig. 7; grouped in a field). Apex of abdomen as in Figs 10-11; tergite VIII as in Figs. 16-17, with apex mucronate in dorsal view; sternite VIII as in Fig. 14; segments IX-X as in Figs 12-13, with hemitergites IX as in Fig. 18 and sternite IX as in Fig 15. Aedeagus as in Figs 4 and 8, with dorsal valve triangular.</p><p>Female: Abdominal tergite VIII (Figs 23-24) forming a medioapical projection, with dorsal outline evenly convex in lateral view. Sternite VIII as in Fig. 22. Genital segments as in Figs 19-21; gonocoxal plate without me- diodorsal ridge.</p><p>Comparisons and diagnostic notes. Megarthrus chiapas Cuccodoro sp. nov. is easily distinguished from the Mexican M. altivagans and M. alatorreorum by the presence of a prohypomeral ridge (Fig. 26) and the medial frontal setae directed forward. It shares these features with M. flavosignatus Bierig (1940), M. zunilensis Sharp (1887), and M. cavianae Rodríguez, Navarrete-Heredia &amp; Arriaga-Varela sp. nov., which are also the only other members of the genus to have the posterior margin of the male abdominal sternite VIII forming two projecting pointed processes (Figs 10, 11 and 14). Within these four species, the males of M. flavosignatus can be easily distinguished by the peglike setae on the mesotibiae arranged in a row (Cuccodoro 2011: Fig. 60) instead of grouped in a field (Figs 6 and 34; Cuccodoro 2011: Fig. 174), and by the metatibiae bearing peg-like setae only in distal third (Cuccodoro 2011: Fig. 59) intead of distal half (Figs 7 and 35; Cuccodoro 2011: Fig. 173). The males of three remaining species differ from each other by the shape of the tip of the abdominal tergite VIII, which is broadly rounded to slightly concave in M. zunilensis (Cuccodoro 2011: Fig. 176), mucronate in M. chiapas (Fig. 17), and sinuate in M. cavianae (Fig. 38). The shape of the aedeagus, notably that of the apical portion of the ventral wall, is diagnostic for the four species.</p><p>The female of M. flavosignatus is still unknown. However the mediodorsal margin of the valvifers is contiguous in apical half in M. zunilensis (Cuccodoro 2011: Fig. 180), while they are not contiguous in M. cavianae and M. chiapas (Fig. 19), and the posterior apical margin of the abdominal stenite VIII is evenly narrowed to subangulate apex in M. chiapas (Fig. 22), while it is slightly sinuate to rounded at apex in M. zunilensis (Cuccodoro 2011: Fig. 180), and deeply sinuate to rounded apex in M. cavianae (Fig. 30).</p><p>Distribution and natural history. Megarthrus chiapas Cuccodoro sp nov. in known only from the northwestern part of the Sierra Madre de Chiapas (Fig. 39), where specimens were collected at elevations ranging from 1500 to 1800 m sieving cloud forest’s leaf litter.</p><p>Etymology. The specific name is a noun in apposition derived from the type locality.</p></div>	https://treatment.plazi.org/id/03E5ED0AFF9ED56F2B996A19FB60FA55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Rodríguez, William David;Navarrete-Heredia, José L.;Arriaga-Varela, Emmanuel;Cuccodoro, Giulio	Rodríguez, William David, Navarrete-Heredia, José L., Arriaga-Varela, Emmanuel, Cuccodoro, Giulio (2020): Two new species of Megarthrus (Coleoptera, Staphylinidae, Proteininae) from Mexican Cloud Forests. Zootaxa 4778 (3): 571-583, DOI: 10.11646/zootaxa.4778.3.8
03E5ED0AFF98D5622B996A19FEF9FDD1.text	03E5ED0AFF98D5622B996A19FEF9FDD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megarthrus cavianae Rodriguez, Navarrete-Heredia & Arriaga-Varela 2020	<div><p>Megarthrus cavianae Rodríguez, Navarrete-Heredia &amp; Arriaga-Varela sp. nov.</p><p>(Figs 27-39)</p><p>http://zoobank.org:act: F4814244-2EA7-493D-8086-01CC754585C0</p><p>Type material. (3♂, 2♀). Holotype ♂: labeled “ MÉXICO: Veracruz, San Andrés Tlalnelhuayocan, Piedra Parada, 19°30’58.6’’N 97°00’34.4’’W, 1522 msnm, Fragmento de Bosque Mesófilo de Montaña ripario, Cernido de Hojarasca húmeda, extraída en <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.00955&amp;materialsCitation.latitude=19.516277" title="Search Plazi for locations around (long -97.00955/lat 19.516277)">Winkler</a>, 5.V.2013, F. Alvarado, E. Arriaga, R. Chavero cols”, in CZUG . Paratypes (4): labeled “ MÉXICO: Veracruz, Xico, Matlalapa, 19°26’24.8»N 97°02’37.2»W, 1532 msnm. Fragmento de Bosque Mesófilo de Montaña ripario, Cernido de Hojarasca húmeda, extraída en embudo de <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.04366&amp;materialsCitation.latitude=19.440222" title="Search Plazi for locations around (long -97.04366/lat 19.440222)">Berlesse</a>, 20.VI.2012. F. Escobar, F. Alvarado, R. Chavero cols” , 1♀ in CZUG, 1♂ in IEXA, and 1 ♂, 1♀ in MHNG.</p><p>Description. Similar to M. chiapas from which it differs as follows: Habitus as in Fig. 29. Combined length of pronotum and elytra = 1.0 - 1.4 mm; maximal pronotal width = 0.7 - 1.1 mm. Antennae as in Fig. 36. Elytra with humeral callus strongly convex; disc with humeral callus strongly convex, two conspicuous swellings similar to humeral callus along anterior and posterior portions of suture, moderately depressed posteriorly along lateral edge. Male: Frontoclypeal area not modified. Protarsomere 1 lacking ventral patch of modified adhesive setae. Mesofemora (Fig. 33) as long as metafemora. Mesotibiae (Fig. 34) shorter than metatibiae (Fig. 35), the latter slightly compressed subapically. Peg-like setae present only on mesotrochanters (Fig. 33; arranged in a single row), on distal two thirds of mesotibiae (Fig. 34; grouped in a field) and on distal half of metatibiae (Fig. 35; grouped in a field). Apical segments of abdomen similar to those in Figs 10-15 &amp; 18; tergite VIII as in Figs 38, with apex sinuate in dorsal view. Aedeagus as in Figs 32 &amp; 37, with dorsal valve triangular.</p><p>Female: Abdominal tergite forming a medioapical projection, with dorsal outline subangulate subapically in lateral view VIII (Fig. 31). Sternite VIII as in Fig. 30. Genital segments similar to those in Figs 19-21; gonocoxal plate without mediodorsal ridge.</p><p>Comparisons and diagnostic notes. Megarthrus cavianae Rodríguez, Navarrete-Heredia &amp; Arriaga-Varela sp. nov. is similar to M. chiapas Cuccodoro sp. nov., from which it differs by its more pronounced elytral relief, as well as several other features detailed in Comparisons and diagnostic notes under M. chiapas .</p><p>Distribution and natural history. Megarthrus cavianae Rodríguez, Navarrete-Heredia &amp; Arriaga-Varela sp. nov. is apparently restricted to Mountain Cloud Forests in the south of the Transmexican Volcanic Belt (Fig. 39). The specimens were collected from sifted leaf litter in riparian remnants of cloud forests at an elevation of approximately 1500 m.</p><p>Etymology. The specific name is a composite word in honor to Carmen Moreno Jiménez, Virginia de Jesús Rodríguez, Ana Cricelia Rodríguez, and Ana María Jiménez, in recognition of their unconditional support of the first author.</p></div>	https://treatment.plazi.org/id/03E5ED0AFF98D5622B996A19FEF9FDD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Rodríguez, William David;Navarrete-Heredia, José L.;Arriaga-Varela, Emmanuel;Cuccodoro, Giulio	Rodríguez, William David, Navarrete-Heredia, José L., Arriaga-Varela, Emmanuel, Cuccodoro, Giulio (2020): Two new species of Megarthrus (Coleoptera, Staphylinidae, Proteininae) from Mexican Cloud Forests. Zootaxa 4778 (3): 571-583, DOI: 10.11646/zootaxa.4778.3.8
