taxonID	type	description	language	source
03E5E91565180530FF5BF9279B2C2CE1.taxon	description	(Fig. 1 – 6)	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565180530FF5BF9279B2C2CE1.taxon	diagnosis	The diagnosis based on workers (for males and queens, see descriptions).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565180530FF5BF9279B2C2CE1.taxon	description	External morphology Body length. 4 – 5 mm (min. to max. WL = 1.24 – 2.02, WL mean = 1.68 mm). No dimorphism or apparent polymorphism in workers. Head. 12 - segmented antennae including scape, without distinct club but with funicular segments continuously enlarged towards the tip, with distal flagellomere approximately twice as wide as the basal flagellomere (Fig. 2 h); scape curved and widened apically (Fig. 1 b, 2 h). Antennal socket relatively closer to the posterior margin of clypeus than in other Camponotini. Palp formula 6: 4, with apical segment of maxillary palp 15 – 20 % shorter than the subapical segment (Fig. 5 a). Head in frontal view oval, with HL similar to HW (min. to max. CI = 94 – 105, CI mean = 101) (Fig. 1 b). Clypeus slightly convex anteriorly and posterior clypeal margin incised in the middle by a developed suture, forming the frontal triangle; but the posterior suture of the triangle below carinae not developed, rather only outlined by bent cuticle (Fig. 1 d). Frontal carinae well developed, going from clypeal margin up to the middle of the head (Fig. 1 d) and forming a convex lifted ridge between each carina and the antennal socket apparent at lateral view (Fig. 1 a); vertical groove in the middle of frons between carinae missing (Fig. 1 d) or only outlined by a glossy smooth line (Fig. 5 d). Mandibles triangular with five teeth that decrease in size from apical to basal teeth. Eyes relatively large (min. to max. EL = 0.25 – 0.40 mm, mean EL = 0.32) and placed vertically in the middle of the head, surpassing the head margins in frontal view, with ~ 20 ommatidia in the longest axis (Fig. 1 b). Workers without ocelli. Mesosoma. Mesosoma shape resembles that of the genus Calomyrmex, with a well developed promesonotal suture. Metanotal groove limited at most of the length to the dorsum of thorax and ending laterally next to the metathoracic spiracle, with a depression and smooth area around the spiracle (Fig. 1 a). The depression next to metathoracic spiracle region is also well visible in dorsal view, with propodeum laterally constricted compared to relatively broad pronotum (Fig. 1 c). Metasoma. Petiole squamiform and triangular, with sharp apex in lateral view and of relatively broad lentil-shape in dorsal view (Fig. 1 c); a characteristic subpetiolar process present and divided from petiole node by a ridge (Fig. 1 a, c). Petiolar and propodeal spiracle with light-coloured margins. Metapleural gland vestigial, probably not functional according to Heterick (2019). Sculpture. On whole body, cuticle relatively smooth and glossy, with only fine punctation or soft lineation. No spines on thorax and petiole, and no deep wrinkles and pits present. Pilosity. Rather short, appressed pilosity; a few longer erect hairs present however at least at the scape apex, on ventral head, abdomen and clypeus (hair length ≤ EL); in contrast to Calomyrmex lacking a dense cover of long, erect hairs over thorax and legs. Colouration. Head, mesosoma and petiole dark brown to blackish colour, abdomen and appendages rather lighter (brown to light brown) and with yellowish coxae and trochanters; except frontal coxa that are darker, similar in colour to thorax (Fig. 1 a, 2 g, 4 a, 5 a).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565180530FF5BF9279B2C2CE1.taxon	discussion	Internal morphology: proventriculus in Overbeckia v. other Camponotini genera Viehmeyer (1916) provided a drawing of the proventriculus in the original genus description, in which rather short sepals were described, being slightly bent anteriorly and of a similar length to the vertical length of the valve (i. e. ‘ the stomach pump’), resembling in this respect the genus Paraparatrechina. These short sepals have been noted as a main feature to distinguish Overbeckia from other SE Asian Formicinae genera (Bolton 1994). Our dissection conforms to the original drawing of Overbeckia ’ s proventriculus by Viehmeyer (1916), showing a similar proportion of sepals compared to the valve. Furthermore, we observed that the valve has relatively thicker walls of whitish colour in Overbeckia, when compared to the other examined genera (Fig. 3 a). Our dissections of Echinopla also show sepals not much longer than the valve (Fig. 3 c), resembling the proventriculus of Overbeckia, but with softer and thinner walls (brittle structure). In contrast to Overbeckia and Echinopla, Calomyrmex has sepals approximately twice as long as the valve, similar to Opisthopsis (i. e. ‘ Formica ’ type in Eisner 1957; Fig. 3 b). Colobopsis resembles Camponotus in the long sepals, more than 2.5 × longer than the valve (i. e. ‘ Camponotus ’ type that also occurs in Polyrhachis (Eisner 1957; Fig. 3 d).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	description	(Fig. 1, 2)	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	etymology	Etymology Viehmeyer (1916) does not explain the reasoning for the species name, but this likely refers to the club-shaped anterior part of antennae, mentioned in the original description (‘ Geissel gegen Ende verdickt’: funinculus enlarged towards the tip) (Fig. 2 h).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	materials_examined	Material examined Syntypes. All examined from photos (AntWeb codes): CASENT 0101191, CASENT 0101192, workers (NHMB; F. Santschi collection); focol 0130, worker (DEIC, marked as ‘ Syntypus’); focol 2566, alate queen (marked as ‘ Typus’), focol 2567 and focol 2568, workers, focol 2564 and focol 2565, males (ZMHB, all marked as ‘ Typus’); CASENT 0178502, worker (MCZC, marked as ‘ Paratype 30741 ’); CASENT 0905180, male (MSGN) / coll. H. Overbeck, <1915 / Singapore / Nests in dry / narrow, hollow bamboo, colony in the hollow branch of a mango tree / 1 ° 17 ′ 35.0 ″ N, 103 ° 51 ′ 21.0 ″ E / det. Viehmeyer (1916). Additional material examined Indonesia: 5 workers and 1 queen; all leg. J. Drescher, canopy fogging (det. R. Nazarreta & P. Klimes): Worker (ZMHB): HJ 3.1 / 5. xi. 2013 / Harapan Rainforest / jungle rubber / dry season / 01 ° 50 ′ 57.7 ″ S, 103 ° 17 ′ 59.7 ″ E. Worker (MZB): BJ 6.2 / 1. xii. 2013 / Bukit Duabelas National Park / jungle rubber / rainy season / 02 ° 01 ′ 48.4 ″ S, 102 ° 46 ′ 14.9 ″ E. Worker (IECA): BF 3.1 / 5. x. 2013 / Bukit Duabelas National Park / jungle rubber / rainy season / 01 ° 56 ′ 34.1 ″ S, 102 ° 34 ′ 51.5 ″ E. Worker (MZB): BF 1.1 / 8. x. 2013 / Bukit Duabelas National Park / forest / dry season / 01 ° 59 ′ 41.4 ″ S, 102 ° 45 ′ 08.5 ″ E. Worker (CASC): CASENT 0872838 (AntWeb) / HymFrm 285 1. xii. 2013 / Bukit Duabelas National Park / jungle rubber / rainy season / 02 ° 00 ′ 56.2 ″ S, 102 ° 45 ′ 11.5 ″ E. Queen without wings (IECA): HF 4.1 / 12. ix. 2013 / Harapan Rainforest / forest / dry season / 02 ° 11 ′ 15.4 ″ S, 103 ° 20 ′ 34.7 ″ E. Philippines (photos): Acc. No. 3576 (MCZC) / Lot Bu. of Sci., P. I. / coll. M. Ramos (further details in General and Alpert 2012).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	diagnosis	Diagnosis Compared to the other two Overbeckia species described here, Overbeckia papuana sp. nov. and O. jambiensis sp. nov., workers and queens in O. subclavata are less hairy, with a lack of long erect hairs on dorsum of petiole, on thorax and on scape except the apical margin; head integument with fine dense punctation; no erect hairs extending past head margins in frontal view; frontal carinae approaching one another less anteriorly compared to those in O. papuana sp. nov. (FCDI <166; Fig. 1 b, 2 h). Longer erect hairs present only on clypeus and ventral abdomen. Fine straight lineation on lateral sides of thorax limited to the basal region below metanotal groove (Fig. 1 a).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	description	Description of worker Morphometrics. Non type ‘ HJ. 3.1 ’ (six syntypes and further five non-types workers, N. A. = measure not available hereafter; total measured N = 11, min. – max): HL 1.08 (0.87 – 1.25), HW 1.14 (0.88 – 1.24), IOD 0.87 (0.67 – 0.92), CL 0.16 (0.15 – 0.19), CW 0.49 (0.34 – 0.54), ML 0.45 (0.33 – 0.47), EL 0.31 (0.25 – 0.36), EW 0.22 (0.16 – 0.26), FCDp 0.38 (0.29 – 0.44), FCDa 0.28 (0.21 – 0.29), SL 0.99 (0.70 – 1.08), SWmax 0.13 (0.11 – 0.17), SWmin 0.06 (0.05 – 0.09), WL 1.62 (1.24 – 1.76), PW 0.79 (0.61 – 0.88), HFL 1.03 (0.92 – 1.08), HFT 0.24 (0.20 – 0.25), HTL N. A. (0.82 – 1.07), PetW 0.40 (0.32 – 0.47), PetL 0.22 (0.15 – 0.24), FCDI 134 (126 – 165), CI 105 (98 – 105), SI 87 (80 – 93), MI 41 (32 – 42). Head. General morphology as for the genus (see above). Frontal carinae approaching one another less anteriorly (FCDI = 134) and with relatively straight curvature, slightly concave near antennal sockets (Fig. 1 d). Mesosoma and metasoma. As for the genus (see above). Pilosity. Numerous tiny appressed hairs (approximately twice the length of 1 ommatidium or shorter) over scape, head and most of the thorax and abdomen; relatively long erect hairs (length ≥ SW) present only on clypeus, head and ventral abdomen and one pair of hairs present on frons in the middle above the eyes (Fig. 1 a); these longer hairs missing on genae and the occipital corners of the head including region above the eyes (in frontal view none of the longer hairs surpass the head margin); scape also without erect hairs except a tuft of ~ 8 setae growing from the apical tip (Fig. 1 b). No long erect hairs at the apex of petiole. Sculpture. Fine punctation on dorsal surface of the head, in particular in the area between frontal carinae and above the clypeus (Fig. 1 b, d) and on mesosoma dorsum and lateral sites of thorax above metathoracic spiracle and dorsolaterally on propodeum (Fig. 1 a); this punctuation changes to the sculpturing rather forming a cell structure than individual pits posteriorly towards the occipital region; soft lineation on lateral sites of mesosoma, below the metathoracic spiracle and on propodeum between propodeal spiracle and hind coxae; same cuticular lineation also present on frontal coxae, petiole and anteriorly on the head near clypeus, but here much softer than on thorax (Fig. 1 b, c). Colouration. Black head, thorax and petiole; dark brown scape except at the apex where light brown; brown abdomen and femurs; tarsi, funiculus and palps light brown to yellow; femurs brown in the middle but lighter near the joints (Fig. 1 a, c). Description of queen Morphometrics. Syntype ‘ focol 2566 ’: HL 1.43, HW 1.45, IOD 1.08, CL 0.30, CW 0.60, ML 0.67, EL 0.53, EW 0.35, FCDp 0.47, FCDa 0.33, SL 1.17, SWmax 0.16, SWmin 0.08, WL 2.56, PW 1.14, HFL 1.08, HFT 0.36, HTL 1.34, PetW 0.58, PetL 0.28, FCDI 142, CI 101, SI 81, MI 47. Head (Fig. 2 e). Eyes almost twice as large as in worker. Frontal carinae approaching each other anteriorly above the posterior margin of clypeus relatively less when compared to queen of O. papuana (FCDI <160). Mandibles with four teeth; apical and subapical teeth approximately twice as long as and sharper than the other three teeth. Three ocelli well developed. Mesosoma (Fig. 2 a, c). In lateral view, thorax dorsum flat at mesothorax, otherwise evenly rounded with convex prothoracic and propodeal angle. Overall morphology of thoracic segments and wings as in Camponotus queens, except apical part of the forewing being relatively broader with evenly convex margins laterally towards the tip. Metasoma. Petiole similar to worker but relatively thicker and with less sharp apex. Abdomen as in workers but larger. Pilosity. Numerous tiny appressed (≥ 1 / 2 length of ocellus) hairs over scape; erect relatively long hairs (length> 1 ocellus) present only on clypeus, ventral head, near ocelli, on mesothorax dorsum and on 3 rd to 5 th abdominal tergites (Fig. 2 a); these longer hairs missing on genae and the occipital corners of the head including region above the eyes (in frontal view no longer hairs surpass the head margins); scape also lacks erect hairs except five to six setae at the apical tip (Fig. 2 e). No long erect hairs on apex of petiole. Sculpture. Fine punctation on the dorsal surface of the head, and on mesosoma dorsum and propodeum; soft lineation on lateral sites of mesosoma, similar to workers. Colouration. Brown head, thorax, scape (except the tip that is lighter) and petiole; light brown abdomen; legs and funiculus yellow (Fig. 2 a, c). Notes The generally lighter colouring may also reflect either the old age of the museum specimen (> 100 years) or a relatively young queen collected from inside nest (alate). Description of male Morphometrics. Syntype ‘ CASENT 0905180 ’: HL 0.90, HW 1.08, IOD 0.69, CL 0.17, CW 0.40, ML 0.34, EL 0.46, EW 0.32, FCDp 0.32, FCDa 0.14, SL 0.80, SWmax 0.09, SWmin 0.07, WL 1.95, PW 1.08, HFL 1.31, HFT 0.18, HTL 1.30, PetW N. A., PetL N. A., FCDI 229, CI 120, SI 74, MI 38. Head (Fig. 2 f). Eyes largely surpass the head margins in frontal view, with EL ~ 1 / 2 of HL, placed at ~ 1 / 2 of HL posteriorly. Frontal carinae approaching relatively less one another anteriorly above the posterior margin of clypeus compared to those in male of O. papuana (FCDI = 229), although this variance is not as distinct as among workers and among queens of the two species. Mandibles blunt without teeth. Three ocelli well developed and relatively larger than in queens (diameter as SWmax). Antennae with 13 segments not broadened towards the tip of funiculus like in the workers and queen (see frontal image of another male from the type series ‘ focol 2564 ’ with intact antennae) (AntWeb, see https: // www. antweb. org). Scape long, at half of the length passing the head margin in frontal view. Mesosoma. Thorax dorsum flat at mesoscutum, otherwise evenly rounded in lateral view with convex prothoracic and propodeal angle, except the mesoscutellar disc that is placed higher and of more convex shape than the other thoracic parts in lateral view (Fig. 2 b). Suture between pronotum and mesoscutum not visible in dorsal view due to highly convex and enlarged mesoscutum compared to pronotum (Fig. 2 d). Metasoma. Petiole triangular but relatively thicker and blunter at the apex. Pilosity. Tiny appressed hairs present on scape and thorax but less numerous over the head compared to queen and workers; long erect hairs (length = SWmax) present only on clypeus, genae, ventral head, near ocelli (one long seta et each) and on 3 rd to 5 th abdominal tergites (Fig. 2 b); these longer hairs missing completely on scape, petiole and at the occipital corners of the head including region above the eyes (in frontal view, no long hairs surpass the head margin). Sculpture. Fine punctation on dorsal surface of the head, and on mesosoma dorsum and propodeum. Colouration. Light brown to yellowish body, except brown head, thorax dorsum and antennae. Mandibles and appendages pale.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156515053AFF7AFBAC9AB92A01.taxon	discussion	Remarks The specimens of O. subclavata from Indonesia that were examined have the posterior parts of the frontal carinae bent slightly inwards and therefore have slightly smaller FCDI (mean 133) than the type specimens from Singapore (mean 155) (Fig. 1 d, 2 h). However, the difference is small and this could possibly be due to measurement error from photographs. All other morphological parameters and the general appearance of the specimens are consistent between the two regions, therefore we regard this to be only a variation within the species. The specimen from the Philippines has a damaged left upper side of the head and a whitish dirt attached to the carinae. Therefore, we did not consider this for morphological measurements. However, absence of the longer hairs on the scape and the thorax, and the finely punctate head with numerous minute recumbent hairs, clearly fit the general appearance of O. subclavata. Viehmeyer (1916) reported that O. subclavata nested in a hollow bamboo stem and in a hollow branch of a mango tree, therefore there is uncertainty whether the syntype series of the species (six workers, three males and one queen, available from AntWeb) belongs to a single nest. Five vials (samples?) collected in 1913 and two collected in 1914 were reported, therefore several colonies from one or more sites in Singapore might have been pooled for the syntype series (most likely two colonies from two stems, a mango and a bamboo). There is also uncertainty as to whether all specimens collected in 1913 – 1914 are documented online at AntWeb, therefore more material might be scattered in museums worldwide. Nevertheless, all specimens collected by H. Overbeck illustrated in the AntWeb match O. subclavata. The arboricolous nesting in hollow plant structures of (probably) dead tissues suggests that the species nests in tree canopies, but is rather a generalist; that is, not using living tissues or myrmecophytic plants, as do more specialised arboreal species (Klimes and McArthur 2014). Specimens of O. subclavata were probably collected by H. Overbeck in a human-managed area but collections from Sumatra suggest that the species utilises a variety of habitats (see Distribution and Discussion for further information).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	description	(Fig. 4)	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	etymology	Etymology Named after the Jambi Province on Sumatra Island (Indonesia) where the type specimen was collected by J. Drescher in the lowland rainforest of Bukit Duabelas National Park (EFForTS project core plot BF 2, see Drescher et al. 2016).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	materials_examined	Material examined Holotype: Worker (MZB): BF 2.2 / 7. x. 2013 / wet season / canopy fogging / lowland forest / Bukit Duabelas National Park / 01 ° 58 ′ 54.2 ″ S, 102 ° 45 ′ 02.3 ″ E / leg. J. Drescher (det. R. Nazarreta, P. Klimes) Paratype: Worker (ZMHB): same data as above but marked as paratype.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	diagnosis	Diagnosis Overbeckia jambiensis workers are the hairiest of the Overbeckia species, with numerous conspicuous erect hairs present on dorsum of petiole, thorax, and also dorsally and ventrally on the head, abdomen and scape; head integument without punctation and with fine lineation; cuticle covered with sparse, short, appressed hairs that are less numerous but longer than in other species and present all over the head except in the area between the eyes and around carinae; integument darker and glossy; frontal carinae less approaching one another anteriorly compared to those of O. subclavata but curvature evenly convex when compared to those of the other species in frontal view (Fig. 4 d). Lineation present on whole lateral sides of mesosoma, including upper part of frontal coxae and the areas above metathoracic spiracle.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	description	Description of worker Morphometrics. Holotype ‘ BF 2.2. ’ (one paratype): HL 1.37 (1.29), HW 1.29 (1.30), IOD 0.94 (0.96), CL 0.24 (0.22), CW 0.53 (0.54), ML 0.55 (0.55), EL 0.40 (0.40), EW 0.30 (0.31), FCDp 0.42 (0.43), FCDa 0.31 (0.31), SL 1.30 (1.27), SWmax 0.16 (0.16), SWmin 0.08 (0.08), WL 1.94 (1.94), PW 0.96 (0.97), HFL 1.30 (N. A.), HFT 0.29 (N. A.), HTL 1.24 (N. A.), PetW 0.45 (0.44), PetL 0.24 (0.22), FCDI 137 (141), CI 94 (101), SI 101 (98), MI 40 (42). Head. General morphology as for the genus (see above). Frontal carinae less approaching one another anteriorly (FCDI = 137) and with convex curvature laterally (Fig. 4 d). Mesosoma and metasoma. As for the genus (see above). Pilosity. Tiny appressed hairs (approximately twice the length of ommatidium) present only sparsely over the head and mesosoma; numerous relatively long erect hairs (length ≥ SW) present on the clypeus and head, and all over abdomen; these longer hairs also present on genae and occipital corners of the head and on frons, surpassing the head margin in frontal view (Fig. 4 b); scape with relatively long hairs of both appressed and erect type, the erect hairs not limited to the apical tip but present along most of the scape (Fig. 4 b). Approximately eight erect hairs growing dorsally from apical margin of petiole. Sculpture. Fine shallow lines, joining each other to a cell formation over the glossy cuticle of the head (Fig. 4 b, d). These lines also present dorsolaterally on mesosoma; the lineation very soft and rather curved, except the lateral sites of mesosoma below metathoracic spiracle and towards coxae and on propodeum, where lines are somewhat deeper and straight (Fig. 4 a); soft lineation also present on frontal coxae and petiole. Colouration. Head, thorax and scape of rather glossy metallic appearance and black, except at scape apex where lighter brown; petiole, abdomen, apical parts of femurs and tibiae brown; tarsi, apex of funiculus and palps brown; femurs pale yellow near the joints; trochanters and mid and hind coxae pale yellow (Fig. 4 a, c). Description of male and queen None (no material)	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E9156511053BFCC4FE5F9D762E32.taxon	discussion	Remarks Body size is somewhat larger on average than in the other two Overbeckia species but due to the availability of only two specimens, assessing whether body size is significantly larger is difficult (the size is comparable to the largest workers measured from series of O. papuana).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	description	(Fig. 5, 6)	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	etymology	Etymology Dedicated to all Papua New Guineans, the nation of the country from which the type and nest series were collected (Papua New Guinea).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	materials_examined	Material examined Holotype. Worker (ZMHB): Morphospecies ‘ CAMP 021 ’ / Acc. N. ‘ HP 0177 ’ / Tree number code ‘ WS 4 D 0868 ’ / Secondary forest / Wanang village / Madang province / Papua New Guinea / 8. iii. 2007 / 05 ° 14 ′ S, 145 ° 11 ′ E / leg. M. Rimandai. Paratypes. Five workers (1 w ZMHB, 1 w IECA, 2 w NAIC, 1 w MZB), one male (ZMHB), one queen without wings (ZMHB): From the same nest series as holotype (the same data). Additional material examined. Papua New Guinea: Other 30 workers and 2 males, including brood, kept in one vial with ethanol in IECA, all from the same nest as the type specimens (same data). Australia (photos): 1 worker (SAMA): ‘ Camponotus ’ / Cairns NQ / on tree / Parkland / 2. viii. 1975 / leg. B. B. Lowery (frontal and lateral photo on p. 115 in McArthur (2012) and A. McArthur, pers. comm. for coll. information; previously misidentified by the original author for Camponotus janeti). 3 workers (WAM): CASENT 0872736, CASENT 0872753, CASENT 0872754 (AntWeb codes; ‘ Overbeckia wam 01 ’): Queensland / Mossman / 18. ix. 2002 / leg, R. Jordan / nest in dead twig in living tree behind sugar mill (thicket near Mossman R.) / det. B. E. Heterick, ii. 2018. 1 worker (CASC): CASENT 0887749 (AntWeb code; ‘ Overbeckia au 01 ’): Queensland / Mt Peters / 1. v. 2014 / leg, M. Collis / det. A. Andersen 30. xi. 2020. Australia (physical specimen): 1 worker (ZMHB): n # 15720 / AUST Qld / Captain Billy Landing / rainforest, ex. dead hanging twig / 11 ° 38 ′ S, 142 ° 51 ′ E / leg. P. Ward (the same collection series as CASENT 0882197 at AntWeb; ‘ Overbeckia au 01 ’).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	diagnosis	Diagnosis General appearance of worker and queen as in O. subclavata, but hairier on the head, with erect hairs on dorsum of petiole, thorax and abdomen; a few hairs also present on distal part of scape. Head integument less punctate than in O. subclavata, with many short-erect hairs extending past head margins in frontal view; tiny appressed hairs in high densities all over the head including genae; frontal carinae significantly approaching one another anteriorly (FCDI> 190) (Fig. 5 d).	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	description	Description of worker Morphometrics. Holotype, ‘ HP 0177 ’ nest series (five paratypes; total N = 6, min. – max): HL 1.08 (1.04 – 1.41), HW 1.10 (1.05 – 1.43), IOD 0.82 (0.79 – 1.10), CL 0.17 (0.14 – 0.23), CW 0.41 (0.39 – 0.55), ML 0.44 (0.36 – 0.57), EL 0.28 (0.26 – 0.34), EW 0.21 (0.19 – 0.27), FCDp 0.37 (0.35 – 0.52), FCDa 0.18 (0.15 – 0.27), SL 1.05 (1.02 – 1.24), SWmax 0.14 (0.12 – 0.18), SWmin 0.06 (0.06 – 0.09), WL 1.58 (1.52 – 2.02), PW 0.77 (0.69 – 1.03), HFL 1.09 (0.92 – 1.24), HFT 0.24 (0.24 – 0.30), HTL 0.94 (0.82 – 1.05), PetW 0.39 (0.37 – 0.47), PetL 0.17 (0.17 – 0.24), FCDI 208 (193 – 236), CI 102 (99 – 105), SI 95 (87 – 101), MI 40 (30 – 43). Head. General morphology as for the genus (see above). Frontal carinae relatively more approaching one another anteriorly above the clypeal margin compared to those of the other species (FCDI = 208), with rather straight carinae curvature all along (only slightly convex laterally adjacent to the antennal sockets at frontal view; Fig. 5 d). Mesosoma and metasoma. As for the genus (see above). Pilosity. Tiny appressed hairs (approximately twice the length of ommatidium or smaller) present in high densities over the head, mesosoma and abdomen; relatively long erect hairs (length ≥ SWmin) present on clypeus, head integument and all over abdomen; these longer hairs are also present on genae and occipital corners of the head and on frons (extending past head margins from the frontal view; Fig. 5 b); scape with relatively short, appressed hairs (as in O. subclavata), with a few erect hairs not limited to the apex but present ventrally along the anterior half of the scape. These hairs shorter and less numerous than in O. jambiensis (Fig. 5 b). Approximately six erect hairs growing dorsally from apical margin of petiole. Sculpture. Less dotted, rather covered by short lines joining each other to a cell formation with fine punctation over the glossy cuticle of the head (Fig. 5 d) and mesosoma dorsolaterally, with most of these cells like pits at smaller (<80 ×) magnification (Fig. 5 b, c). Lateral sites of thorax with soft lineation below mesothoracic spiracle towards mid and hind coxa. The lineation is also present on frontal coxa and petiole, but smoother. Colouration. Head, mesosoma and frontal coxa black; scape dark brown except at apex, where lighter brown; funiculus of antennae brown except the lighter apex; petiole dark brown; abdomen and hind tibiae brown, remaining appendages rather light brown, with mid and hind coxae, trochanters and palps pale yellow (Fig. 5 a, c). Description of queen Morphometrics. Paratype ‘ HP 0177 ’ nest series: HL 1.56, HW 1.57, IOD 1.22, CL 0.32, CW 0.59, ML 0.63, EL 0.49, EW 0.37, FCDp 0.50, FCDa 0.24, SL 1.23, SWmax 0.16, SWmin 0.09, WL 2.51, PW 1.14, HFL 1.22, HFT 0.31, HTL 1.27, PetW 0.55, PetL 0.35, FCDI 208, CI 101, SI 78, MI 40. Head, mesosoma and metasoma. Similar to O. subclavata, except frontal carinae significantly approaching one another anteriorly (FCDI = 208; Fig. 6 e). Pilosity. Tiny appressed hairs on scape surface, with three to four longer erect hairs on the lateral sites of the first half of the scape (hair length ≥ 1 ocellus); presence of many relatively long, erect hairs on the head (length ≥ 1 ocellus) on genae, above the eyes and over the occipital corners of the head (Fig. 6 e), and on thorax dorsum, apical tip of petiole, and over both ventral and dorsal sites of all five abdominal tergites. Sculpture. Fine punctation over the head, and on mesosoma dorsum and propodeum; and soft lineation on lateral sites of mesosoma (Fig. 6 a). Colouration. Black head, thorax, petiole and frontal coxae; dark brown abdomen and appendages except middle and hind coxae, trochanters, palps and apical tips of antennae and tarsi that are lighter brown (Fig. 6 a). Description of male Morphometrics. Paratype ‘ HP 0177 ’ nest series: HL 0.84, HW 1.00, IOD 0.66, CL 0.12, CW 0.35, ML 0.31, EL 0.34, EW 0.29, FCDp 0.27, FCDa 0.10, SL 0.67, SWmax 0.10, SWmin 0.06, WL 1.71, PW 0.96, HFL 1.31, HFT 0.18, HTL 0.99, PetW 0.33, PetL 0.21, FCDI 255, CI 119, SI 67, MI 37. Structures and diagnosis (Fig. 6 b, d, f). No apparent large distinction when compared to the male of O. subclavata, except for the following minor differences: The specimen is ~ 10 % larger than the measured male of O. subclavata, but morphological measures otherwise considerably overlapping between the two species’ males. Frontal carinae relatively more approaching one another anteriorly than in O. subclavata, but this difference not as distinct as between the two species’ workers (FCDI = 255 v. 229 in O. subclavata male). The hairs on clypeus more numerous than on clypeus of O. subclavata; presence of many more small hairs on frons and above ocelli (length near 1 / 2 of ocellus) over the occipital region (Fig. 6 f). These small, dense hairs are also present dorsolaterally on thorax (on mesoscutum and mesoscutellar disc; Fig. 6 b, d). Colouration as in O. subclavata male.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	distribution	Remarks The arboreal nest of O. papuana sp. nov. in Papua New Guinea was collected at 16.4 m in a dead vine growing along the tree trunk of Trichospermum pleiostigma (F. Muell.) Kosterm. (diameter at breast height = 16.3 cm). The nest consisted of less than one hundred workers, with one wingless gyne, three males, and multiple larvae and pupae enclosed in cocoons.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
03E5E91565100538FF0DFA689A022EAE.taxon	materials_examined	The material from Australia was examined and measured mainly from photographs from the following sources: (i) the book by McArthur (2012), where a worker of Overbeckia was incorrectly assigned to Camponotus janeti (minor worker fits ‘ O. papuana’, whereas major worker presumably fits C. janeti; there is uncertainty as to whether the two individuals are from different localities and pins or are on a single pin at SAMA); (ii) the photos on AntWeb of the three specimens in Heterick (2019) (see above). The Australian specimens appear to be larger in body size based on the measures from the photos (4 workers measured from photos, WL> 1.95 mm) than the Papua New Guinea (PNG) nest series (6 w measured, WL <1.74). However, we treat these as one species because the Australian single specimen sent by P. Ward for direct examination (n # 15720) matches O. papuana types in the body size (WL = 1.64; Supplementary Table S 1) and morphology.	en	Klimeš, Petr, Drescher, Jochen, Buchori, Damayanti, Hidayat, Purnama, Nazarreta, Rizky, Potocký, Pavel, Rimandai, Maling, Scheu, Stefan, Matos-Maraví, Pável (2022): Uncovering cryptic diversity in the enigmatic ant genus Overbeckia and insights into the phylogeny of Camponotini (Hymenoptera: Formicidae: Formicinae). Invertebrate Systematics 36 (6): 277-288, DOI: 10.1071/is21067, URL: http://dx.doi.org/10.1071/is21067
