taxonID	type	description	language	source
03E53859DA3D2630965F6FEFFBF89906.taxon	description	Figure 2 B	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3D2630965F6FEFFBF89906.taxon	materials_examined	Material examined. COLOMBIA • 2, 101.4 – 120.2 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10194. • 6, 121.6 – 161.8 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 075 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10346. Identification. Dentaries abruptly converging towards the mandibular symphysis. Bony tooth plates on the basihyal. Dark chevron pattern along the sides of body. A dark mid-lateral band extending along entire length of lateral line, in larger specimens this band is missing (Mattox et al. 2014).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3D2630965F686BFC589433.taxon	description	Figure 2 C	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3D2630965F686BFC589433.taxon	materials_examined	Material examined. COLOMBIA • 7, 52.9 – 121.6 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10112. • 13, 73.3 – 121.7 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 12 Dec. 2012; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10368. Identification. A broad dark mid-lateral stripe above lateral line from snout to tips of middle caudal-fin rays. Body with dark ground portion above black lateral stripe lacking projections or spots extending dorsally, base color below lateral stripe to level of axillary scale gray. Adults with a well-defined black spot from mid- dle of pectoral fin to its tip. Brown spots absent on dor- sal part of sides above lateral stripe (Londoño-Burbano et al. 2011).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6C70FDB49F9B.taxon	description	Figure 2 E	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6C70FDB49F9B.taxon	materials_examined	Material examined. COLOMBIA • 65, 23.4 – 38.8 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10150. • 44, 17.6 – 44.2 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 4 Apr. 2013; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10556. Identification. Maxillary not reaching suture between first and second infraorbital. Lateral line with 40 – 43 pored scales. Five scales between the dorsal-fin origin and the lateral line. Anal fin with 33 – 37 branched rays. A bright mid-lateral stripe extending from humeral region to caudal peduncle (Eigenmann 1913).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6EB3FEA099F0.taxon	description	Figure 2 F	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6EB3FEA099F0.taxon	materials_examined	Material examined. COLOMBIA • 12, 28.9 – 43.8 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10078. • 17, 25.9 – 44.7 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10139. Identification. Third infraorbital in contact with the preopercle. Lateral line complete, with 35 – 38 pored scales. No lamellar flaps over pores of lateral line scales. Anal fin short with 11 – 13 branched rays. Dark humeral blotch vertically elongated. Middle caudal-fin rays with a dark horizontal stripe (Harold and Vari 1994; Albornoz-Garzón et al. 2018).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6B18FC0E9D7A.taxon	description	Figure 2 G	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A263795DA6B18FC0E9D7A.taxon	materials_examined	Material examined. COLOMBIA • 10, 30.7 – 65.8 mm SL; Río Alvarado-Chucuní; 04 ° 27 ′ 56.7 ″ N, 075 ° 03 ′ 46.7 ″ W; 697 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10225. • 20, 29.3 – 56.3 mm SL; Río Alvarado-El País; 04 ° 27 ′ 37.9 ″ N, 075 ° 07 ′ 45.2 ″ W; 909 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10368. Identification. Third infraorbital not in contact with the preopercle. Lateral line complete, with 35 – 38 pored scales. No lamellar flaps over pores of lateral line scales. Anal fin short with 9 – 12 branched rays. Pelvic fins with short hooks on rays (usually shorter than the width of the branched rays). Eye diameter: 21.8 – 26.8 % HL. Body with dorsal portion with dark reticulated pattern formed by pigmentation concentrated on distal portion of scales. Dark humeral blotch vertically elongated. Middle cau- dal-fin rays without a dark horizontal stripe (Albornoz- Garzón et al. 2018).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A2637965F6C92FACD9FDF.taxon	description	Figure 2 H	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A2637965F6C92FACD9FDF.taxon	materials_examined	Material examined. COLOMBIA • 16, 53.1 – 79.4 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT- IC 10060. • 6, 55.8 – 66.6 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10372. Identification. Third infraorbital in contact with the preopercle. Lateral line complete, with 34 – 37 pored scales. No lamellar flaps over pores of lateral line scales. Anal fin short with 9 – 11 branched rays. Caudal-peduncle depth (12.6 – 15.9 % SL). Dark humeral blotch ovoid. Mid- dle caudal-fin rays without a dark horizontal stripe (Har- old and Vari 1994; Albornoz-Garzón et al. 2018).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A2637965F6EFFFA729A55.taxon	description	Figure 2 I	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA3A2637965F6EFFFA729A55.taxon	materials_examined	Material examined. COLOMBIA. • 7, 31.3 – 35.3 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 28 Jun. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10706. • 7, 36.6 – 37.5 mm SL; Quebrada La Manjarres; 04 ° 28 ′ 19.3 ″ N, 075 ° 04 ′ 26.9 ″ W; 758 m a. s. l.; 28 Jun. 2013; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10788. Identification. Outermost branched pectoral-fin ray of adult males with a fan-shaped structure distally formed. Bony hooks and a dark blotch present distally in the fanshaped structure. Caudal-fin rays 10 th and 11 th lacking of terminal lateral-line tube. Adult males with a long gill gland (7.0 – 11.4 % SL). Urogenital papillae developed in adult females. An intense dark pigmentation at the base of the five anterior dorsal-fin rays (Vanegas-Ríos 2016).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA68BBFC819CC2.taxon	description	Figure 3 A	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA68BBFC819CC2.taxon	materials_examined	Material examined. COLOMBIA • 2, 29.6 – 41.2 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10087. • 1, 32.0 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10331. Identification. Body slender and slightly short. Body deep (19.5 – 19.5 % SL). Head length: 23.1 – 27.6 % SL. Maxillary barbel not reaching pelvic-fin origin. Adi- pose fin relatively short (18.6 – 19.3 % SL) and triangleshaped. Insertion of the adipose fin equidistant between dorsal and caudal fins. Caudal fin forked with both lobes pointed. Body brown with four conspicuous pale dorsal transverse bands: one around posterior region of the head, one in front of the dorsal fin, one anterior to adipose fin, and one around the caudal peduncle (Miles 1943).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA6DEAFD7E9F0D.taxon	description	Figure 2 K	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA6DEAFD7E9F0D.taxon	materials_examined	Material examined. COLOMBIA • 7, 24.5 – 34.5 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10335. • 22, 26.2 – 32.9 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 4 Apr. 2013; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10569. Identification. Lateral line with 30 – 35 pored scales. Dorsal-fin origin equidistant from the tip of snout and the base of the caudal rays. Anal fin short with 9 – 10 branched rays. Dark humeral blotch vertically elongate. A dark silvery mid-lateral stripe extending from humeral region to caudal peduncle (Eigenmann 1913).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA392634965F6DEAFB939849.taxon	materials_examined	Material examined. COLOMBIA • 11, 47.4 – 97.9 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10197. • 3, 55.9 – 76.6 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 12 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10379. Identification. Body moderately elongate. Maxillary barbel reaching anal-fin origin, surpassing anal-fin in some specimens. Greater maximum depth of dentations on the posterior margin of the pectoral-fin spine, 1.71 – 2.90 times in the width of the spine at its base. Distal posterior margin of pectoral-fin spine without denta- tions, 32.2 – 52 % in pectoral-fin spine length. Adipose fin relatively long (22.6 – 35.8 % SL) and deep (3.5 – 6.7 % SL), forming elevated curve in lateral profile. Insertion of adi- pose fin equidistant between dorsal and caudal fins. Cau- dal fin forked with both lobes pointed, lower lobe slightly shorter than upper lobe. Body light brown. A dark brown mid-lateral stripe well defined, extending from snout to base of middle caudal-fin rays (Ardila-Rodríguez 2017 a; Conde-Saldaña et al. 2019 b).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA392634965F6961FAE69B05.taxon	description	Figure 3 C	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA392634965F6961FAE69B05.taxon	materials_examined	Material examined. COLOMBIA • 3, 149.0 – 164.7 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10351. • 1, 125.8 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 12 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10380. Identification. Body moderately elongated and relatively deep. Maxillary barbel parallel to main body axis, crossing the pelvic-fin insertion. Dorsal-fin base (14.4 – 19.4 % SL). Adipose fin long (35.9 – 43.2 % SL) and deep (4.1 – 7.5 % SL), forming elevated curve in lateral profile. Insertion of the adipose fin closer to the dorsal fin than to the caudal fin. Caudal fin forked with both lobes rounded, lower lobe slightly longer than upper lobe. Body usually brown. In some specimens a dark lateral stripe along the lateral line is present (Hernández et al. 2015).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA6E2CFEBD9937.taxon	description	Figure 2 L	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA39263495DA6E2CFEBD9937.taxon	materials_examined	Material examined. COLOMBIA • 1, 49.1 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 04 Apr. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10536. • 1, 48.8 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 28 Jun. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10680. Identification. Lateral line with 53 – 60 pored scales. Scales below lateral line 13 to 18. Anal fin with 44 – 47 branched rays. Dark humeral blotch ovoid present. Cau- dal peduncle with a conspicuous tear-drop shaped spot (Lucena 2000).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA392635965F6A56FAEA9433.taxon	description	Figure 3 D	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA392635965F6A56FAEA9433.taxon	materials_examined	Material examined. COLOMBIA • 3, 44.3 – 62.0 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10417. • 2, 40.9 – 42.4 mm SL; Río Alvarado-Chucuní; 04 ° 27 ′ 56.7 ″ N, 075 ° 03 ′ 46.7 ″ W; 697 m a. s. l.; 13 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10452. Identification. Body moderately elongated. Body deep (9.2 – 16.6 % SL). Cleithral width: 14.0 – 18.8 % SL. Head length: 18.4 – 22.1 % SL. Maxillary barbel reaching pec- toral-fin insertion. Adipose fin rounded (17.6 – 30.2 % SL, its origin anterior to anal-fin; adipose-fin origin equi- distant between dorsal and caudal fins). Adipose-fin depth (2.2 – 4.6 % SL). Caudal fin forked with both lobes rounded, upper lobe longer than lower lobe. Body brown with scattered minute melanophores along body. A conspicuous pale collar around posterior region of head and pale blotch in front of dorsal fin. Fins hyaline.	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A9658690FFC9C9A4C.taxon	description	Figure 3 I	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A9658690FFC9C9A4C.taxon	materials_examined	Material examined. COLOMBIA • 8, 34.1 – 94.7 mm SL; Río Alvarado-Chucuní; 04 ° 27 ′ 56.7 ″ N, 075 ° 03 ′ 46.7 ″ W; 697 m a. s. l.; 13 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10450. • 23, 33.1 – 101.2 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 4 Apr. 2013; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10593. Identification. Distal portion of snout without plates. Caudal peduncle with five rows of plates. Caudal fin emarginated with lower portion longer than upper portion. Black spots on head and body. Fins with scattered dark spots. Caudal fin uniformly dusky, with undulated dark stripe along distal margin (Fowler 1941; Miles 1947).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A95DA6DC8FD3F9ECA.taxon	description	Figure 3 E	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A95DA6DC8FD3F9ECA.taxon	materials_examined	Material examined. COLOMBIA • 3, 119.7 – 130.2 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10352 • 2, 117.9 – 138.1 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz- Garzón leg.; CZUT-IC 10366. Identification. Predorsal distance (34.9 – 38.9 % SL). Pre- ventral distance (42.1 – 47.0 % SL). Adipose-fin length (16.4 – 20.3 % SL). Dorsal – adipose distance (16.6 – 22.5 % SL). Interorbital width (36.2 – 42.8 % HL). Body gray without spots or stripes (Villa-Navarro et al. 2017).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A95DA6E26FDD199B5.taxon	description	Figure 3 F	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A95DA6E26FDD199B5.taxon	materials_examined	Material examined. COLOMBIA • 19, 18.2 – 37.3 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10106. • 8, 21.3 – 45.0 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10168. Identification. Relatively small body size (largest specimen 47 mm SL). Head triangular in dorsal view. Premaxilla with 27 conical teeth arranged in two or three irregular rows. Dentary with 30 conical teeth arranged in three of four irregular rows. Interopercle with 26 odontodes. Opercle with 23 to 29 odontodes. Pectoral fin with seven branched rays. Caudal fin emarginated. Body light yellow with predorsal and horizontal mid-lateral rows of large blotches (García-Melo 2009).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A96586C54FB319FA3.taxon	materials_examined	Material examined. COLOMBIA • 12, 25.7 – 75.8 mm SL; Río Alvarado-El País; 04 ° 27 ′ 37.9 ″ N, 075 ° 07 ′ 45.2 ″ W; 909 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10254. • 16, 24.8 – 54.3 mm SL; Quebrada Chumba; 04 ° 29 ′ 06.2 ″ N, 075 ° 05 ′ 48.6 ″ W; 973 m a. s. l.; 20 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10282. Identification. Relatively medium body size (largest specimen 85 mm SL). Head triangular in dorsal view. Premaxilla with 37 conical teeth arranged in three irregular rows; dentary with conical 37 teeth arranged in three irregular rows. Interopercle with 37 odontodes. Opercle with 23 odontodes. Pectoral fin with seven branched rays. Caudal fin emarginated. Body brown with dark diffuse dots on dorsum and dorsolateral region of trunk (García- Melo 2009; Ardila-Rodríguez 2017 b).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263A95DA68C5FC5D9D05.taxon	materials_examined	Material examined. COLOMBIA • 19, 18.2 – 37.3 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10106. • 8, 21.3 – 45.0 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10168. Identification. Relatively medium body size (largest specimen 75 mm SL). Head squared in dorsal view. Premaxilla with 32 incisiform teeth arranged in two regular rows. Dentary with 40 incisiform teeth arranged in two regular rows. Interopercle with 39 odontodes. Opercle with 15 odontodes. Pectoral fin with six branched rays. Caudal fin rounded. Body light brown, lighter ven- trally. Dorsal region of body with light brown blotches and dots, irregularly distributed; a light brown midlateral stripe present, extending from behind opercle to caudalfin base, becoming diffuse in larger specimens (García- Melo 2009).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263B96586B63FDA49D3A.taxon	description	Figure 3 J	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA37263B96586B63FDA49D3A.taxon	materials_examined	Material examined. COLOMBIA • 66, 23.0 – 69.2 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10121. • 14, 25.5 – 97.1 mm SL; Río Alvarado-Chucuní; 04 ° 27 ′ 56.7 ″ N, 075 ° 03 ′ 46.7 ″ W; 697 m a. s. l.; 19 Sep. 2012; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10219. Identification. Distal portion of snout without plates. Interoperculum with four to five odontodes. Caudal pedun- cle with five rows of plates. Anal fin rays I, 5. Caudal fin emarginated with lower portion longer than upper portion. Light spots usually concentrated on head. Fins with white to yellow bands. Distal margin along caudal fin pale, more evident at tips (Regan 1904).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B96586DC8FA879F79.taxon	description	Figure 4 B	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B96586DC8FA879F79.taxon	materials_examined	Material examined. COLOMBIA • 11, 25.0 – 102.7 mm SL; Quebrada Cocare; 04 ° 28 ′ 37.4 ″ N, 075 ° 08 ′ 25.4 ″ W; 1057 m a. s. l.; 13 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10500. • 6, 51.7 – 139.8 mm SL; Quebrada Cocare; 04 ° 28 ′ 37.4 ″ N, 075 ° 08 ′ 25.4 ″ W; 1057 m a. s. l.; 5 Apr. 2013; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10665. Identification. Relatively large size (largest specimen 140 mm SL). Premaxillary teeth unicuspid. Adipose fin long and deep with a short spine embedded into the posterior region of fin, posterior fin margin continuous with first caudal-fin ray. First pectoral and dorsal-fin rays ex- tended as filaments. Caudal fin truncated. Upper and lower unbranched caudal-fin rays extended as filaments. Body brownish with greenish tone, dark scattered spots and blotches on head and body (Briñez-Vásquez 2011).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B96586E91FB2F9A55.taxon	description	Figure 4 C	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B96586E91FB2F9A55.taxon	materials_examined	Material examined. COLOMBIA • 32, 20.1 – 54.4 mmSL; Quebrada Chumba; 04 ° 29 ′ 06.2 ″ N, 075 ° 05 ′ 48.6 ″ W; 973 m a. s. l.; 20 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10275. • 43, 19.2 – 59.3 mm SL; Quebrada Cocare; 04 ° 28 ′ 37.4 ″ N, 075 ° 08 ′ 25.4 ″ W; 1057 m a. s. l.; 5 Apr. 2013; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10666. Identification. Relatively medium size (largest specimen 71.4 mm SL). Premaxillary teeth bicuspid. Head and predorsal region covered with small papillae. Adipose fin short and thick, with a visible spine covered with odontodes, posteriorly connected to caudal peduncle by a membrane. First dorsal-fin ray not extended as a fila- ment. Ventral-fin origin well in advance of the dorsal-fin origin. Caudal fin with distal margin concave, upper and lower unbranched caudal-fin rays extended as filaments. Body light brown with black scattered spots and blotches on head and body, sometimes with a small yellow blotch on the adipose-spine base (Regan 1904).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B95DA6C5CFE499F62.taxon	materials_examined	Material examined. COLOMBIA • 1, 53.9 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 28 Jun. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10697. • 2, 24.2 – 49.7 mm SL; Río Alvarado-Puente; 04 ° 31 ′ 11.3 ″ N, 074 ° 59 ′ 14.0 ″ W; 521 m a. s. l.; 28 Jun. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10766. Identification. Mouth with spoon-shaped teeth. Plates in the skin between the dorsal fin and the lateral plates an- terior of the dorsal-fin spine 1 to more of 6. Pectoral-fin spine reach 2 – 3 plates beyond the pelvic fin when de- pressed ventrally to the pelvic fin. Caudal-fin spines with spots (Armbruster 2003).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B95DA6EB4FD089994.taxon	description	Figure 3 L	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B95DA6EB4FD089994.taxon	materials_examined	Material examined. COLOMBIA • 5, 29 – 74.2 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10065. • 3, 30.2 – 33.1 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 074 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10123. Identification. Whisker-like odontodes on the cheek and at the corner of the snout. Absence of plates ventrally in front of the anus. Caudal fin slightly emarginated to forked. Dark spots in the dorsal fin (Armbruster 2005).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B95DA68A6FE799433.taxon	description	Figure 4 A	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA36263B95DA68A6FE799433.taxon	materials_examined	Material examined. COLOMBIA • 13, 32.8 – 69 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10063. • 19, 24.3 – 92.3 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz- Garzón leg.; CZUT-IC 10082. Identification. Reticulations on predorsal, supraoccipital, and interorbital regions. Rostrum present. Cleithral width: 11.0 – 14.1 % SL. Coalescent plates 13 – 16. Dorsal, pectoral, pelvic, and anal fins lacking filaments. A thin dorsolateral stripe discontinuous and ill-marked present. Dorsal, pectoral, pelvic, and anal fins lacking large, dark blotches occupying almost the entire fin (Londoño-Bur- bano and Reis 2019).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA35263996586ADEFD109E14.taxon	materials_examined	Material examined. COLOMBIA • 1, 181 mm TL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10353. • 3, 249 – 314 mm TL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 12 Dec. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10367. Identification. Dorsal margin of snout profile strongly concave. Pectoral fin length: 44 – 53 % HL. Head length: 13 – 16 % LEA. Branchial opening: 20 – 28 % HL. Dark hu- meral spot absent or very diffuse (Hulen et al. 2005).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263995DA6F78FDFB98D0.taxon	description	Figure 4 F	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263995DA6F78FDFB98D0.taxon	materials_examined	Material examined. COLOMBIA • 1, 131.8 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10090. • 1, 86.2 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 28 Jun. 2013; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT- IC 10692. Identification. Mouth rictus extends to beyond the posterior margin of eyes. Body blotchy. A clear band from chin to the beginning of dorsal mid-sagittal electroreceptor organ. Two clear bands surrounding the caudal peduncle in specimens (up to 165 mm TL); bands become obscured by pigment blotching in specimens larger than 165 mm (de Santana et al. 2004).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263996586C2EFB9A9F4B.taxon	description	Figure 4 H	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263996586C2EFB9A9F4B.taxon	materials_examined	Material examined. COLOMBIA • 10, 11.7 – 24.3 mm SL; Quebrada La Manjarres; 04 ° 28 ′ 19.3 ″ N, 075 ° 04 ′ 26.9 ″ W; 758 m a. s. l.; 19 Sep. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10209. • 8, 15.9 – 23.7 mm SL; Quebrada La Manjarres; 04 ° 28 ′ 19.3 ″ N, 075 ° 04 ′ 26.9 ″ W; 758 m a. s. l.; 12 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10434. Identification. Dorsal-fin rays 6 – 7. Gonopodium with a retrorse hook on ray 5. The fleshy palp elongated, ex- tending well beyond the tip. Ventral spines on the third gonopodial ray large, giving the gonopodium a bulb-like appearance. Body silvery, with dots, stripes and blotches (only in males) (Poeser 2003).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263996586E63FA899955.taxon	description	Figure 4 I	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263996586E63FA899955.taxon	materials_examined	Material examined. COLOMBIA • 11, 10.7 – 33.9 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 075 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10126. • 7, 11.8 – 28.4 mm SL; Río Alvarado-Chucuní; 04 ° 27 ′ 56.7 ″ N, 075 ° 03 ′ 46.7 ″ W; 697 m a. s. l.; 19 Sep. 2012; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10243. Identification. Dorsal-fin rays 9 – 10. Blueish and reddish spots (males), extending from humeral region to caudal peduncle in the body sides. Dorsal fin with a yellow spot surrounded by two or three black bands (Poeser 2003).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263995DA685DFBF29CC2.taxon	description	Figure 4 G	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263995DA685DFBF29CC2.taxon	materials_examined	Material examined. COLOMBIA • 11, 16.9 – 45.7 mm SL; Quebrada La Manjarres; 04 ° 28 ′ 19.3 ″ N, 075 ° 04 ′ 26.9 ″ W; 758 m a. s. l.; 19 Sep. 2012; Juan G. Albor- noz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10208. • 13, 19.9 – 44.6 mm SL; Quebrada La Manjarres; 04 ° 28 ′ 19.3 ″ N, 075 ° 04 ′ 26.9 ″ W; 758 m a. s. l.; 12 Dec. 2012; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10439. Identification. Head scaled, except for cheek. Mouth su- perior. Longitudinal series of scales 40 – 42. Body col- oration of live males brown, sides of body bluish, from humeral region to caudal peduncle, with a lateral irregular series of red spots. Dorsal and anal fins with red spots on basal half, distal half yellow with thin dark pur- ple band on distal margin. Caudal fin dark purple along procurrent rays, distal portion yellow and white. Pectoral and pelvic fins yellowish. Body coloration of live fe- males brown, with scattered dark blotches along sides. Dorsal and anal fins with dark spots. Pelvic and pectoral fins hyaline. A dark spot on upper portion of caudal pe- duncle (Henn 1916).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263E965868DBFF1B9D88.taxon	description	Figure 4 J	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA34263E965868DBFF1B9D88.taxon	materials_examined	Material examined. COLOMBIA 2, 53.4 – 58.2 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10110. • 3, 41.6 – 45.4 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 075 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 4 Apr. 2013; Juan G. Albornoz-Garzón, Cristhian C. Conde-Saldaña leg.; CZUT-IC 10582. Identification. Body moderately deep (36.9 – 42.5 % SL). Snout slightly rounded (25.9 – 356 % HL). Mouth termi- nal. First gill arch lacking a prominent epibranchial lobe. Scales in E 1 11 – 18 and E 2 6 – 10. Dorsal fin with 14 – 16 spines and 10 – 12 soft rays, 5 th to 6 th ray longest, pro- longed as filament. Anal fin with three spines and 7 – 10 soft rays, 4 th to 5 th ray longest, prolonged as filament. Caudal fin subtruncate or slightly rounded. In sexually mature specimens, head and first half body with metallic green spots. Sides of body with nine dark vertical stripes and a dark midlateral spot. A vertically dark blotch in the mid of the caudal-fin base. Dorsal fin dark grey, with light grey bands and narrow white margin. Caudal fin dark grey with distal portion yellowish (Steindachner 1878).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA33263E95DA6CA2FEDF990B.taxon	description	Figure 4 K	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA33263E95DA6CA2FEDF990B.taxon	materials_examined	Material examined. COLOMBIA • 13, 16.7 – 44.4 mm SL; Río Alvarado-Caldas Viejo; 04 ° 36 ′ 41.2 ″ N, 074 ° 55 ′ 46.2 ″ W; 351 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde- Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10111. • 11, 13.8 – 66.7 mm SL; Quebrada La Caima; 04 ° 35 ′ 45.8 ″ N, 075 ° 56 ′ 39.6 ″ W; 374 m a. s. l.; 19 Sep. 2012; Cristhian C. Conde-Saldaña, Juan G. Albornoz-Garzón leg.; CZUT-IC 10159. Identification. Body deep (38.1 – 41.8 % SL). Snout mod- erately pointed (43.4 – 59.6 % HL). Mouth subterminal and protractile. First gill arch with prominent epibranchial lobe. Scales in E 1 19 – 20 and E 2 13. Dorsal fin with 16 spines and 13 soft rays, 4 th to 5 th ray longest, extended as filaments. Anal fin with three spines and 7 – 8 soft rays. Caudal fin with distal margin concave. Body coloration brown or light brown. Sides of body with six faint dark vertical stripes. A vertically dark blotch at middle of cau- dal-fin. Dorsal fin dark grey, with light dark bands in soft rays. Caudal fin hyaline with dark spots. Pectoral fin dark grey. Pelvic and anal fins pale grey or hyaline (Eigen- mann 1922).	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA32263F96586B44FA8E9AE1.taxon	description	17 ′ Caudal fin absent; body uniformly dark ..................... Sternopygidae: Sternopygus aequilabiatus (Fig. 4 E)	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
03E53859DA32263F95DA6F9BFA7A9A34.taxon	description	4 ′ Simple pattern of lateral line tubes in skin of snout, cheek and nape; posterior supraoccipital process not contacting nuchal plate ........................ Heptapteridae (except Pimelodella floridablancaensis): 14 5 Dark lateral stripe along body; dorsal surface of head smooth ................................................. Heptapteridae: Pimelodella floridablancaensis (Fig. 3 B) 5 ′ Dark lateral stripe along body absent; dorsal surface of head rough ..... Pimelodidae: Pimelodus yuma (Fig. 3 E) 6 Caudal fin with 12 or more rays; adipose fin present ................................................................................... 7 6 ′ Caudal fin with 12 rays; adipose fin absent ................ ......................... Sturisomatichthys leigthoni (Fig. 4 A) 7 Abdomen naked; teeth villiform ............................... 8 7 ′ Abdomen with plates; teeth spoon-shaped ................. .............. Hypostomus (Cochliodon) hondae (Fig. 3 K) 8 Anterior region of snout naked; whisker-like odon- todes on cheek absent ................................................ 9 8 ′ Anterior region of snout covered with plates; whiskerlike odontodes on cheek .............................................. ............................... Lasiancistrus caucanus (Fig. 3 L) 9 Head and body gray, usually with small dark spots; caudal fin with undulated dark stripe at distal mar- gin .................................. Chaetostoma milesi (Fig. 3 I) 9 ′ Head and body dark brown, sometimes with light spots; caudal fin without undulated dark stripe at dis- tal margin ................ Chaetostoma thomsoni (Fig. 3 J) 10 Adipose fin long and deep, with short spine embedded in posterior region; first pectoral-fin ray and first dor- sal-fin ray prolonged as filaments .............................. ................................... Astroblepus grixalvii (Fig. 4 B) 10 ′ Adipose fin short and low, with visible spine covered with odontodes; first pectoral-fin ray and first dorsalfin ray not prolonged as long filaments ................... 11 11 Premaxillary teeth bicuspid; body light brown with black scattered spots and blotches, sometimes with small yellow blotch on adipose-spine base ................. .................................. Astroblepus homodon (Fig. 4 C) 11 ′ Premaxillary teeth incisiform; body usually dark, with variable light blotch on middle of caudal peduncle, sometimes with light blotch on posterior region of dorsal-fin base .................... Astroblepus sp. (Fig. 4 D) 12 Head squared in dorsal view, dark lateral band well defined on body ........................................................... .................... Trichomycterus transandianus (Fig. 3 G) 12 ′ Head triangular in dorsal view, dark lateral band absent ....................................................................... 13 13 Body with a series of dark lateral spots, usually larger than eye; premaxilla with 27 teeth; dentary with 30 teeth ................... Trichomycterus banneaui (Fig. 3 F) 13 ′ Body with dark spots, usually smaller than eye; pre- maxilla with 37 teeth; dentary with 37 teeth .............. ......................... Trichomycterus mogotensis (Fig. 3 H) 14 Maxillary barbel not reaching pelvic-fin insertion; adipose fin short (<30 % SL), adipose-fin origin equi- distant between dorsal fin and caudal fin; body with conspicuous transverse pale bands ......................... 15 14 ′ Maxillary barbel reaching pelvic-fin insertion; adi- pose fin long (> 39 % SL), adipose-fin origin closer to dorsal fin than the caudal fin; body variably pig- mented, sometimes dark or light brown, often with a dark lateral stripe along lateral line ............................ .............................. Rhamdia guatemalensis (Fig. 3 C) 15 Adipose fin triangular; caudal-fin lobes pointed; body with conspicuous pale transverse bands ..................... ............................ Cetopsorhamdia molinae (Fig. 3 A) 15 ′ Adipose fin rounded; caudal-fin lobes rounded; body with conspicuous pale band behind head and in front of dorsal fin ...... Heptapteridae undescribed (Fig. 3 D)	en	Albornoz-Garzón, Juan G., Conde-Saldaña, Cristhian C., López-Delgado, Edwin O., García-Melo, Jorge E., Villa-Navarro, Francisco A. (2020): Fishes from the Río Alvarado drainage, Upper Río Magdalena Basin, Colombia. Check List 16 (5): 1181-1198, DOI: 10.15560/16.5.1181, URL: http://dx.doi.org/10.15560/16.5.1181
