taxonID	type	description	language	source
03E3878EFFEEFFB8FF7FFCC9FE02C7A3.taxon	type_taxon	Dichomeris Hübner, 1818. Type species: Dichomeris ligulella Hübner, 1818.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFB8FF7FFCC9FE02C7A3.taxon	synonymic_list	For synonyms, see Hodges 1986.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	description	(Figs 4, 35)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Bac Kan Prov.: Ba Be Nat. Park, 26 – 28 vii 2006, leg. Park, Chae & Cuong, gen. slide no. CIS- 6323, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	description	Description. Adult (Fig. 4). Wingspan 13.0 mm. Head dark brown dorsally, with grayish yellow erect scales laterally above compound eyes; frons grayish brown. Ocellus absent. Antenna with scape dark brown on dorsal surface, grayish white on ventral surface; flagellum alternately dark brown and yellowish brown on dorsal surface, yellowish brown on ventral surface, ciliate. Second segment of labial palpus clothed with rough scales, brown on outer surface, yellowish white on inner surface; 3 rd segment shorter than second, also clothed with rough scales brownish in basal 2 / 3, yellowish in distal 1 / 3. Thorax and tegula dark brown; mesothoracic anepisternum with yellowish-white hair pencils in male. Fore- and midleg pale yellow on inner surface; femora grayish yellow, tibiae and tarsi brown on outer surface; hind tibia pale yellow. Forewing ground color yellowish white, scattered with brown scales; a small dark-brown patch beyond 5 / 6 of costa; a longitudinal dark-brown stripe running from near 1 / 3 length of cell to termen and a slightly oblique, black fascia well-developed along fold; costa slightly arched near basal 1 / 3 and oblique beyond basal 5 / 6; apex triangularly protrudced; termen concave beyond apex; black line along termen; fringe yellowish brown. Hind wing broader than forewing, brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 35): Uncus subquadrate, bluntly rounded on posterior margin, concave in arch on anterior margin. Gnathos stout, curved at basal 2 / 5, acute at apex; culcitula semiovate, with dense spines. Valva as long as the length of tegumen plus uncus, narrow in basal 3 / 5, broader distally, with rounded apex; valvella short, about 1 / 5 the length of valva, with rounded apex, bearing long setae apically. Vinculum banded, longer than tegumen plus uncus, sparsely setose at basal 1 / 4; lateral lobes absent. Saccal region broad, slightly concave at middle anteriorly. Sicae paired lobes, long, symmetrical, fused for short distance at base, then separated in U-shape, reaching anterior margin of uncus, with rounded apex. Aedeagus extremely stout, subtriangular basally, with three sclerotized, nearly same sized processes arising from zone: left one with rounded apex, internal process gradually narrowed to apex, slightly bent downward, and right one slender, sharply pointed apically; with an additional median process arising from ventral surface, ovate, about 1 / 3 length of internal lobe. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	distribution	Distribution. Vietnam (Bac Kan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	diagnosis	Diagnosis. The new species is similar to Dichomeris moriutii Ponomarenko & Ueda, 2004 in the male genitalia, but it can be distinguished from the latter by the third segment of the labial palpus clothed with rough scales and the male genitalia with sicae reaching anterior margin of the uncus. In D. moriutii, the third segment of the labial palpus is smoothly clothed and the sicae exceeds posterior margin of the uncus.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEEFFBBFF7FFBD2FC33C5E8.taxon	etymology	Etymology. The specific name is derived from the Latin, - fascia (= banded) and lobatus (= lobe) with a Latin dimunitive suffix, - ella, referring to the banded siace in the male genitalia.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	description	(Figs 7, 36)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ninh Binh Prov.: Cuc Phuong Nat. Park, 450 m, 14 vi 2004, leg. KT Park & N Cuong, gen. slide no. CIS- 6680, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	description	Description. Adult (Fig. 7). Wingspan 11.0 mm. Head dark brown dorsally; frons orange gray. Ocellus absent. Antenna with scape deep brown on dorsal surface, pale yellow on ventral surface; flagellum yellowish brown with dark-brown annulations dorsally, yellowish brown ventrally, with fine cilia. Second segment of labial palpus dark brown on outer surface, grayish yellow on inner surface, with grayish white-tipped triangular scale tuft dorsally; 3 rd segment smoothly clothed, shorter than second, pale yellow except dark brown on ventral surface. Thorax and tegula dark brown; mesothoracic anepisternum without hair-pencils in male. Hind tibia dark brown on outer surface, grayish white on inner surface, tarsomeres grayish white apically on outer surface. Forewing elongated; ground color dark brown; a small, pale-yellow costal streak at distal 1 / 4 of costa; a pair of discal black spots, larger one placed at basal 2 / 3 of cell and smaller one at end of cell; plical spot similar to discal spot at basal 2 / 3 of fold; apex more or less obtuse; termen oblique, nearly straight, with black line along margin; fringe concolorous with ground color, with narrow yellowish-white basal line. Hind wing brownish gray; fringe conclorous with ground color. Male genitalia (Fig. 36): Uncus broadened with posterior margin bluntly rounded, sparsely setose. Gnathos heavily sclerotized, C-shaped, acute apically. Valva much longer than tegumen plus uncus, rounded apically; valvella about 1 / 6 the length of valva, gradually narrowed to sharp apex, setose. Vinculum about 3 / 5 the length of tegumen plus uncus; lateral lobes arising from distal 5 / 7 of vinculum, about 2 / 3 length, triangularly extended to sharp apex, setose apically. Saccal region broad, slightly concave on anterior margin medially. Sicae slender, paired lobes, fused basally, nearly symmetrical. Aedeagus shuttle-shaped; basal zone triangular, truncate at apex; two sclerotized, asymmetrical processes arising from left side of zone, sharply pointed apically, longer one about 3 / 5 the length of aedeagus and shorter one about 2 / 5 length. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	distribution	Distribution. Vietnam (Ninh Binh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	diagnosis	Diagnosis. The new species resembles Dichomeris parallelosa Park & Ponomarenko, 1998, but it differs from the latter by the aedeagus with two sclerotized lobes from the zone and a hook-shaped lobe absent. In D. parallelosa, the aedeagus has three sclerotized lobes from the zone, and one of them is hook-shaped.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBBFF7FFDCCFD4DC2A0.taxon	etymology	Etymology. The specific name is derived from the Latin, - flav (= yellow) and maculatus (= spotted), referring to the yellow spot at costal 3 / 4 in the forewing.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	description	(Figs 11, 37, 50)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Bac Kan Prov.: Ba Be Nat. Park, 26 – 28 vii 2006, leg. Park, Chae & Cuong, gen. slide No. CIS- 6284, in NIBR. Paratypes: VIETNAM: Vinh Phuc Prov.: 1 ♀, Tam Dao Nat. Park, 450 m, 27 iv 2006, leg. Park, Kim & Kang, gen. slide no. nklep 033. CHINA: Hainan Prov.: 3 ♀, Nankai Town (19 ° 07 ’ N 109 ° 40 ’ E), Yinggeling, 270 m, 09 – 10 xi 2016, leg. X Bai, S Qian, & W Qi, gen. slide nos. ZSN 16417, ZSN 16422, in NKU.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	description	Description. Adult (Fig. 11). Wingspan 12.0 – 13.0 mm. Head dark yellowish brown dorsally; frons orange gray. Ocellus present. Antenna with scape dark brown on dorsal surface, grayish white on ventral surface; flagellum yellowish brown with dark-brown annulations on dorsal surface, pale yellow on ventral surface. Second segment of labial palpus dark brown on outer surface, pale gray on inner surface, clothed with grayish white-tipped triangular scale tuft; 3 rd segment smoothly clothed, longer than second segment, pale yellow except dark brown on ventral surface. Thorax dark brown mesially, yellowish brown laterally; tegula dark brown; mesothorax with khaki hairpencils arising from anepisternum in male. Fore- and midleg dark brown on outer surface; femora grayish white, tibiae and tarsi pale yellow on inner surface, tarsi with each tarsomere pale yellow apically on outer surface; hindleg pale yellow on inner surface, dark brown on outer surface except tibia pale yellow in basal half, tarsomeres pale yellow at apex on outer surface. Forewing ground color yellowish brown, scattered with dark-brown scales; several oblique strigulae along costa in basal 3 / 5; ill-defined dark-brown fascias arising from 1 / 3 of costa to middle of dorsum; apex obtuse; termen slightly oblique, with black line along margin; fringe concolorous with ground color in basal half, then dark brown beyond. Hind wing dark yellowish brown; fringe concolorous with ground color. Male genitalia (Fig. 37): Uncus rather small, broadened in distal 1 / 3, with rounded posterior margin, bearing two long setae posteriorly, deeply concave in semicircle on anterior margin. Gnathos short, strongly curved at middle, sharp at apex; culcitula triangular, densely setose, rounded posteriorly. Valva longer than tegumen plus uncus complex, concave at distal 1 / 5 on dorsal margin, nearly straight on ventral margin, rounded at apex; free ventral lobe short, about 1 / 5 the length of valva, setose, pointed apically. Setose patch large sheeted. Vinculum rather broad, about equal to the length of tegumen plus uncus; lateral lobes arising from basal 1 / 3, shorter than 1 / 6 the length of vinculum, cornute, setose apically. Saccal region nearly straight on anterior margin. Sicae paired lobes, nearly symmetrical, separated at base, narrowed basally, then broadened, slightly arched on inner margin, distal half narrowed sparsely setose. Aedeagus stout, nearly parallel sided, rounded basally, bifurcated apically: left part slightly longer than right one; internal lobe sclerotized arising from disatl 1 / 3 of aedeagus, excurvate mesially, acute apically, reaching apex of aedeagus. Female genitalia (Fig. 50): Apophyses anteriores about 1 / 4 length of apophyses posteriors, triangularly inflated distally. Eighth tergite with a triangular, median protrusion on posterior margin; eighth sternite deeply concave on posterior margin medially, densely setose laterally. Antrum broadly developed in rectangular, more or less cupshaped. Ductus bursae membranous, short, narrowed between antrum and ductus bursae, with heavily sclerotized longitudinal band mesially and with various irregular shaped, heavily sclerotized plates arising from ductus bursae extending to anterior 1 / 4 of corpus bursa. Corpus bursae ovate; signum plate ovate, sclerotized, placed at the middle of corpus bursae; accessory bursae arising from middle of signum.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	distribution	Distribution. China (Hainan), Vietnam (Vinh Phuc, Bac Kan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	diagnosis	Diagnosis. This species is superficially similar to Dichomeris hodgesi Li & Zheng, 1996, but it can be distinguished from D. ferrogra by having triangular scale-tuft on the second segment of the labial palpus, khaki hairpencils on the mesothoracic anepisternum in the male; the male genitalia with uncus narrowed in basal 2 / 3, inflated in distal 1 / 3, the sicae with basal half broader, distal half narrower; the eighth tergite with median protrusion posteriorly. In D. ferrogra, the second segment of the labial palpus has a quadrate scale tuft, the mesothoracic anepisternum has gray hair- pencils in the male; the male genitalia with rectangular uncus; the eighth tergite straight on the posterior margin.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEDFFBAFF7FF884FD27C2A0.taxon	etymology	Etymology. The specific name is derived from the Latin, infundibularis (= funneled), in reference to the ductus bursae having a funneled sclerotized plate distally.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	description	(Figs 12, 38)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ninh Binh Prov.: Doi Vau Vill. (20 ° 19 ' N 105 ° 36 ' E), Cuc Phuong, 135 m, 10 – 16 ix 2013, leg. JB Heppner, gen. slide no. nklep 054, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	description	Description. Adult (Fig. 12). Wingspan 14.0 mm. Head dark yellowish brown dorsally, with erect grayish white scales laterally above compound eyes; frons grayish yellow. Ocellus present. Antenna with scape dark brown dorsally, grayish white ventrally; flagellum yellowish brown, with dark-brown annulations on dorsal surface. Second segment of labial palpus dark brown on outer surface, pale yellow on inner surface, clothed with grayish whitetipped quadrate scale tuft; 3 rd segment smoothly clothed, as long as 2 nd segment, pale yellow on dorsal surface, dark brown on ventral surface. Thorax and tegula dark yellowish brown; mesothorax with yellowish-brown hairpencil arising from anepisternum in male. Fore- and midleg dark brown on outer surface, grayish white on inner surface; hindleg yelowish white, mixed with brown on outer surface. Forewing pale yellowish brown, irregularly scattered with dark-brown scales; costal margin with dark-brown streak basally; apex acute; termen concave; fringe concolorous with groud color. Hind wing brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 38): Uncus small, elongated, inflated in distal 1 / 3, with rounded posterior margin; anterior margin gently concave. Gnathos with median process recurved in C-shape, sharply pointed at apex; culcitula triangular, densely spinose. Valva about as long as tegumen plus uncus, narrowest at base, dorsal margin slightly convex at 2 / 5, abruptly invaginate at 3 / 4; ventral margin gently concaved at middle, distal 1 / 4 slightly convex; apex rounded; valvella short, dilated in distal half, setose apically. Setose patch large sheeted. Vinculum shorter than length of tegumen plus uncus, slightly protruded in basal 2 / 5 outward, sparse with setae; lateral lobe absent. Saccal region broad, with small emargination on anterior margin medially. Sicae with two pairs of lobes, symmetrical, with each pair bearing two different length of arms, fused basally, then divided in U-shape; outer arm about 2 / 5 the length of inner arm, gently incurvate, gradually narrowed to sharp apex, inner arm nearly straight, extended to middle of tegumen. Aedeagus shuttle-shaped, bluntly rounded basally, broadest at zone; a sclerotized, bar-shaped rod arising from half of aedeagus along dorsal margin; a needle-like process, about equal to length of dorsal one, sharply pointed apically, and with a short, semiovate lobe internally. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	distribution	Distribution. Vietnam (Ninh Binh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	diagnosis	Diagnosis. This species resembles Dichomeris zonata Li & Wang, 1997, but it can be distinguished from the latter by having two pairs of sicae, wheras in D. zonata, the sicae are not furcate.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFECFFBDFF7FF884FF30C060.taxon	etymology	Etymology. The specific name is derived from the Latin, - lanceus (= lances), referring to the sicae with hastate lobes.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	description	(Figs 13, 39)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Vinh Phuc Prov.: Tam Dao. Nat. Park, 900 m, 09 vii 2003, leg. KT Park, gen. slide no. CIS- 6281, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	description	Description. Adult (Fig. 13). Wingspan 19.5 mm. Head grayish yellow dorsally, pale yellow posteriorly. Ocellus present. Antenna with scape yellowish brown dorsally, pale yellow ventrally; flagellum yellowish brown with dark-brown annulations dorsally, with fine cilia. Second segment of labial palpus dark brown on outer surface, yellowish white on inner surface, with grayish white-tipped triangular scale tuft on dorsal surface; 3 rd segment smoothly clothed, as long as 2 nd segment, dark brown excepting pale-yellow basal 5 / 6 dorsally. Thorax grayish yellow; tegula grayish yellow; mesothoracic anepisternum with pale-yellow hair-pencils in male. Fore- and midleg dark brown on outer surface, femora and tibiae yellowish white, tarsi yellow on inner surface; hindleg grayish yellow on outer surface, pale yellow on inner surface, tibia clothed with long, yellowish scales. Forewing broad; ground color grayish yellow, sparsely scattered with brown scales, with weak, pale-yellow postmedian line visible; discal and discocellular spots dark brown; plical spot dark brown, placed at middle of fold; dark brown dots along termen at end of each veins; costa slightly arched beyond 2 / 3; apex obtuse; termen slightly concave before middle, with small black dots along margin; fringe concolorous with ground color. Hind wing broader than forewing, brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 39): Uncus more or less trapezoidal, broadly expanded distally, setose sparsely on ventral surface, with two long setae near posterior margin; anterior margin concave in triangular. Gnathos slender, long, bent near basal 1 / 3, with sharply pointed apex; culcitula triangular, bluntly rounded on posterior margin, densely setose. Valva about equal to the length of tegumen plus uncus, bluntly rounded at apex; free ventral lobe about 1 / 2 length of valva, acute apically, densely setose in distal 2 / 3. Setose patch columnar, inflated distally, about 1 / 3 length of valva. Vinculum narrow band, about 2 / 3 length of tegumen plus uncus; anterior part of vinculum almost at right angles with lateral arms, excavated in saccal region; a pair of heavily sclerotized, more or less sickle-shaped, asymmetrical lateral processes arising from near middle: left one slender, about 2 / 3 length of vinculum and right one loner than left one. Saccal region concave on anterior margin medially. Sicae with a pair of slender processes, slightly asymmetrical, about 2 / 3 length of lateral arms of vinculum. Aedeagus membraneous in basal 1 / 4, narrowed basally; a heavily sclerotized flange from distal 3 / 5 along margin, exceeding apex of aedeagus, sharply acuted apically; internal lobe arising from zone, broad in basal 2 / 3, slightly narrowed to sharp apex in distal 1 / 3; two weakly sclerotized, different sized lobes arising from zone: longer one about 3 times length of shorter one. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	distribution	Distribution. Vietnam (Vinh Phuc).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	diagnosis	Diagnosis. This species is superficially similar to Dichomeris fuscusitis Li & Zheng, 1996, but it can be separated from D. fuscusitis by the forewing with fascia absent; the male genitalia with lateral lobes arising from basal 4 / 5 of the vinculum, and the aedeagus having one sclerotized internal lobe. In D. fuscusitis, the forewing has an obvious fascia distally, the lateral lobe arising from distal 2 / 3 of the vinculum, and the aedeagus has no sclerotized internal lobe.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEBFFBCFF7FFB44FECEC732.taxon	etymology	Etymology. The specific name is derived from the Latin, - longi (= long), referring to the valva with a long ventral lobe.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	description	(Figs 15, 40)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Vinh Phuc Prov.: Tam Dao Nat. Park, 950 m, 01 viii 2006, leg. Park, Chae & Cuong, gen. slide no. nklep 028, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	description	Description. Adult (Fig. 15). Wingspan 15.0 mm. Head dark brown dorsally; frons gray. Ocellus absent. Antenna with scape brown on dorsal surface, grayish white on ventral surface; flagellum yellowish brown with darkbrwon annulations dorsally, with fine cilia ventrally. Second segment of labial palpus yellowish brown on outer surface, pale gray on inner surface, clothed with rough scales; 3 rd segment thickened, nearly as long as 2 nd segment, yellowish brown. Thorax and tegula brown; mesothorax without hair pencils arising from anepisternum in male. Fore- and midleg brown on outer surface, grayish white on inner surface; hindleg pale yellow. Forewing ground color pale brownish gray, scattered with dark-brown scales, more in basal half; costa gently arched, with a short, black streak from costal 3 / 4 to near apex; a narrow, yellowish-white subterminal line well preesentyed along termen internally; apex slightly produced; termen oblique; fringe pale yellow in basal half, then grayish. Hind wing broader than forewing, pale brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 40): Uncus extremely broad, uniquely developed with triangular median process on posterior margin, densely bearing strong setae. Gnathos short, small; culcitula semicircular, densely spined. Valva narrow in basal half and broadly expanded in distal half, slightly longer than length of tegumen plus uncus; with free basal lobe bluntly rounded apically, about 1 / 5 the length of valva. Vinculum slightly shorter than the length of tegumen plus uncus; lateral lobe arising from basal 1 / 3 of vinculum, slightly incurvated, with acute apex, about 1 / 3 the length of vinculum. Saccal region convex anteriorly. Sicae arising from lateral sides of saccal region, symmetrical, curved in S-shape, acute apically. Aedeagus longer than valva, triangular in basal 1 / 6, then parallel sided, slightly narrowed distally; one sclerotized marginal lobe arising from zone slender, acute apically, reaching apex of aedeagus; another short, sclerotized lobe from zone inflated distally, about 1 / 6 the length of aedeagus. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	distribution	Distribution. Vietnam (Vinh Phuc).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	diagnosis	Diagnosis. This new species is superficially similar to Dichomeris ferrogra Li & Wang, 1997, but it can be distinguished from the latter by the third segment of the labial palpus with rough scales, whereas in D. ferrogra, the third segment is smoothly clothed. The male genitalia of this species are easy to distinguish from those of the latter by the uncus with trianguar median process on posterior margin with dense strong setae, and the valva broadly expanded in distal half.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFEAFFBFFF7FFCF5FD1BC4C8.taxon	etymology	Etymology. The specific name is derived from the Latin, - marg (= margin) with a Latin suffix - alis, referring to the uncus with dense setae on the posterior margin.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	description	(Figs 21, 42, 53)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Vinh Phuc Prov.: Tam Dao Nat. Park, 950 m, 26 iv 2006, leg. Park, Kim & Kang, gen. slide no. CIS- 6675, in NIBR. Paratypes: VIETNAM: Ninh Binh Province: 2 ♂, 1 ♀, Cuc Phuong Nat. Park, 450 m, 24 iv 2006, leg. Park, Kim & Kang, gen. slide nos. CIS- 6251 (♂), nklep 048 (♂), & nklep 061 (♀). All types in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	description	Description. Adult (Fig. 21). Wingspan 14.0 – 15.0 mm. Head yellowish brown dorsally, with yellowish scales laterally; frons grayish brown. Ocellus absent. Antenna with scape dark brown dorsally, yellowish white ventrally; flagellum yellowish brown with dark-brown annulations, ciliate ventrally. Second segment of labial palpus dark brown on outer surface, grayish yellow on inner surface, clothed with rough scales; 3 rd segment equal to length of second segment, dark brown, clothed with rough scales, yellow white apically. Thorax dark brown mesially, with yellowish scales laterally; tegula dark brown anteriorly, yellowish in distal half; mesothoracic anepisternum with pale-yellow hair-pencils in male. Fore- and midleg dark brown on outer surface, grayish on inner surface; hindleg pale yellow, mixed with brown scales on outer surface. Forewing slightly dilated distally; ground color pale yellow, irregularly scattered with dark-brown scales, especially in basal half and along fold; with an obscure longitudinal brownish streak from middle of cell to termen; costa slightly arched medially; apex obtuse; termen oblique, slightly concave before middle, scattered with dark-brown scales along termen; fringe pale yellow in basal 3 / 4, brown beyond along termen, brown around tornus. Hind wing brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 42): Uncus narrowed in basal half, inflated distally, with rounded posterior margin; anterior margin concave in triangular. Gnathos long, slender, bent from before middle, with acute apex; culcitula semicircular, spinous. Valva longer than the length of tegumen plus uncus, bent medially, parallel-sided in basal half, slightly dilated in distal half, rounded apically; valvella about 1 / 4 the length of valva, clavate, setose in distal half. Setose patch short, columnar, dilated disally. Vinculum about equal to the length of tegumen plus uncus, broad in basal 3 / 5; lateral lobes arising from distal 2 / 5 of vinculum, about 3 / 5 the length of vinculum, apex truncate. Saccal region broad, nearly straight on anterior margin. Sicae fused for short distance, asymmetrical: left lobe about 3 / 5 length of vinculum, truncate on apex with sharply produced process outwardly; right one shorter, about 2 / 5 length of vinculum, bifurcate into V-shape apically. Aedeagus shuttle-shaped, long, about equal length to saccus plus uncus, broad in basal zone; two sclerotized, asymmetrical lobes arising from zone: left one about 1 / 4 length of aedeagus, dentate on inner margin, blunt at apex; right one longer than left one, bifurcate in distal 2 / 3, dentate on inner margin, blunt at apex. Female genitalia (Fig. 53): Apophyses anteriores about 1 / 4 length of apophyses posteriores. Eighth sternite with nearly straight posterior margin and roundly concave on anterior margin. Antrum membraneous. Ductus bursae with heavily sclerotized complex of various shapes of plates, left one extended 1 / 3 to posterior of corpus bursae, excurvate near apex. Corpus bursae large, ovate, densely spinous on inner surface anteriorly; signum composed by several strong spicules placing near anterior margin medially; accessory bursa arising from middle of signum.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	distribution	Distribution. Vietnam (Vinh Phuc, Ninh Binh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	diagnosis	Diagnosis. This species is superficially similar to Dichomeris aculata Park, 2001 in the male genitalia, but it can be separated from the latter by having teeth on the ventral surface of the sicae, with different shape of sclerotized lobe in the aedeagus, whereas in D. aculata the sicae are smooth on the ventral surface. The female genitalia differ from those of the latter by having semicircular sclerotized plate in the ductus bursae, but the ductus bursae of the latter has no similar sclerotized plate.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE9FFBFFF7FFEACFD0AC36C.taxon	etymology	Etymology. The specific name is derived from the Greek, - myria (= many) and - odontos (= tooth), referring to the sicae having several teeth on the ventral surface.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	description	(Figs 17, 41)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Vinh Phuc Prov.: Tam Dao Nat. Park, 950 m, 24 v 2007, leg. KT Park, gen. slide no. CIS- 6277. Paratype: 1 ♂, same data as the holotype, 5 v 2005, gen slide no. CIS- 6690. All types in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	description	Description. Adult (Fig. 17). Wingspan 17.0 mm. Head grayish yellow dorsally; frons pale yellow. Ocellus present. Antenna with scape yellowish brown on dorsal surface, yellowish white on ventral surface; flagellum alternately yellow and yellowish brown dorsally. Second segment of labial palpus thick, with triangular scale tuft dorsally, ochreous yellow on outer surface, pale yellow on inner surface; 3 rd segment slender, smoothly clothed, shorter than 2 nd segment, pale yellow except ochreous yellow on ventral surface. Thorax and tegula yellow, mixed with yellowish-brown scales; mesothoracic anepisternum without hair-pencils in male. Fore- and midleg yellowish brown on outer surface, yellowish white on inner surface, tarsi with tarsomeres yellowish white apically on outer surface; hindleg pale yellow, tibia clothed with long yellowish scales. Forewing ground color pale yellow, scattered with yellowish-brown scales especially below costa from basal 2 / 5 and below fold; a broad, yellowish-brown fascia with arched anterior margin above fold; costa arched in basal 1 / 3 then straight; apex more or less acute; termen oblique, slightly convex medially; fringe yellowish brown with yellowish-white narrow basal line. Hind wing brownish gray; fringe concolorous with ground color, with narrow yellowish-white basal line. Male genitalia (Fig. 41): Uncus extremely large, more or less triangular distally, with a longitudinal ridge mesially; anterior margin concave in semicircular. Gnathos strongly bent in basal 1 / 3, with acute apex; culcitula triangularly rounded on posterior margin, densely spinous. Valva slightly longer than tegumen plus uncus, broadly expanded in distal half with rounded apex; free ventral lobe short, about 1 / 5 the length of valva, with rounded apex, setose. Setose patch short, irregular-shaped, inflated distally. Vinculum about 2 / 3 the length of tegumen plus uncus; lateral lobes arisig from distal 4 / 5 of vinculum, about 1 / 2 the length of vinculum. Saccal region broad, produced in triangular on anterior margin. Sica single, stout, about 2 / 3 the length of vinculum. Aedeagus shuttle-shaped, narrowed in basal 1 / 3, blunt at apex; one sclerotized internal lobe arising from zone, gradually narrowed to sharp apex, slightly exceeding apex of aedeagus; two sclerotized lobes from zone on ventral surface, about 1 / 3 length of aedeagus, symmetrical, sharp and excurvate apically. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	distribution	Distribution. Vietnam (Vinh Phuc).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	diagnosis	Diagnosis. This species is similar to Dichomeris neixiangensis Li & Wang, 1999, but it can be distinguished from the latter by the forewing with yellowish ground color; the lateral lobe of the vinculum gradually narrowed in basal 1 / 3, the aedeagus having one clerotized internal lobe. In D. neixiangensis, the forewing ground color is yellowish brown; the lateral lobe of the vinculum is gradually narrowed to apex, and the aedeagus has no clerotized internal lobe.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFBEFF7FFF1DFCEBC1F8.taxon	etymology	Etymology. The specific name is derived from the Latin prefix, - ochro (= ochreous), with a Latin suffix, – ellus, referring to the ochreous yellow fascia in the forewing.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	description	(Figs 23, 43)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ninh Binh Prov.: Cuc Phuong Nat. Park, 200 m, 23 iv 2006, leg. Park, Kim & Kang, gen. slide no. CIS- 6307. Paratypes: Ninh Binh Prov.: 1 ♂, same as holotype, 22 iv 2006, gen. slide no. CIS- 6285; 1 ♂, same locality as holotype, 450 m, 14 vi 2004, leg. KT Park & N Cuong, gen. slide no. CIS- 6678. All types in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	description	Description. Adult (Fig. 23). Wingspan 14.5 – 15.0 mm. Head dark brown dorsally; frons gray. Ocellus absent. Antenna with scape dark brown dorsally, pale yellow ventrally; flagellum dark bown dorsally, yellowish brown ventrally, ciliate. Second segment of labial palpus dark brown on outer surface, grayish on inner surface, clothed with rough scales above; 3 rd segment thickened, shorter than 2 nd segment, dark brown, with rough scales above. Thorax and tegula dark brown; mesothoracic anepisternum without hair-pencils in male. Fore- and midleg dark brown except tarsomeres grayish yellow apically on outer surface, yellowish white on inner surface; hindleg pale yellow, mixed with dark-brown scales on outer surface, tibia clothed with long, yellow scales, tarsus with each tarsomere pale yellow apically. Forewing slightly broadened distally; ground color pale yellow, scattered with dark-brown scales; costa with dark-brown streak along margin from base to 4 / 5; with dark-brown, distinct discal and discocellular spots at middle and end of cell; broad dark-brown fascia presented along dorsum, extending to 4 / 5 of dorsum; a large, quadrate, dark-brown patch well-developed along termen from beyond apex to tornus; apex acute; termen obliquely straight; fringe dark brown, with pale yellow basal line. Hind wing brownish gray; fringe concolorous with ground color. Male genitalia (Fig. 43): Uncus nearly rectangular, with rounded posterior margin, with a longitudinal ridge medially; anterior margin deeply concave in semicircular. Gnathos long, heavily sclerotized, strongly bent at basal 1 / 3, sharp at apex; culcitula semicircular, densely spinose, with a rounded process mesially on posterior margin. Valva slightly longer than tegumen plus uncus, narrowed in basal half, broadened distally, rounded at apex; valvella large, spatulate, about 2 / 5 the length of valva, with rounded apex. Setose patch small, columnar, inflated distally. Vinculum broad in anterior base, almost at right angles with lateral lobes; lateral lobes arising from middle, about 1 / 2 the length of vinculum, broadly banded; distal 1 / 5 bifurcate; apex acute. Saccal region narrow, nearly straight on anterior margin. Sicae with a pair of long, symmetrical lobes, fused for short distance, gradually narrowed to rounded apex, slightly exceeding base of vinculum. Aedeagus longer, 1.2 times as long as valva; triangularly produced at base, slightly narrowed beyond half; two short, weakly sclerotized bars placed beyond middle. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	distribution	Distribution. Vietnam (Ninh Binh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	diagnosis	Diagnosis. This species is superficially similar to Dichomeris obsepta (Meyrick, 1935), but it can be separated from the latter by having symmetrical lateral lobes in the vinculum, the symmetrical sicae, whereas in Dichomeris obsepta, lobes of the vinculum and the sicae are asymmetrical.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE8FFB1FF7FF9BDFE53C028.taxon	etymology	Etymology. The specific name is derived from the Latin, - quadrat (= quadrate), and - ellus (= small), referring to the rectangula valvella.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	description	(Figs 25, 44)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Vinh Phuc Prov.: Tam Dao Nat. Park, 950 m, 26 iv 2006, leg. Park, Kim & Kang, gen. slide no. nklep 023. Paratypes: 6 ♂, same locality as holotype, 1000 m, 04 – 05 v 2005, leg. KT Park & SR. Kim, gen. slide nos. CIS- 6305, nklep 059. All types in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	description	Description. Adult (Fig. 25). Wingspan 12.5 – 14.5 mm. Head dark brown; frons gray. Ocellus present. Antenna with scape dark brown on dorsal surface, pale yellow on ventral surface; flagellum alternately dark-brown and yellowish brown on dorsal surface. Second segment of labial palpus dark brown on outer surface, grayish brown on inner surface except yellow near base, with gray-tipped triangular scale tuft dorsally; 3 rd segment smoothly clothed, slightly shorter than 2 nd segment, dark brown ventrally. Thorax and tegula dark brown; mesothoracic anepisternum without hair pencils in male. Fore- and midleg dark brown on outer surface, femora grayish white, tibiae and tarsi pale yellow on inner surface; hindleg grayish brown on outer surface, yellowish white apically, pale yellow on inner surface, tibia clothed with long, yellowish brown scales. Forewing elongate; ground color dark brown, with ill-defined yellowish – white postmedian line from costal 3 / 4 to before tornus, strongly convex outward, followed by more darkened area; apex acute; termen oblique; fringe dark brown. Hind wing dark grayish brown; fringe concolorous with ground color. Male genitalia (Fig. 44): Uncus elongated, semiovate, densely setose on ventral surface. Gnathos strongly bent at basal 1 / 4, L-shaped, with acute apex upturned; culcitula irregular-shaped, densely spined. Valva subequal to length of tegumen plus uncus, nearly patrallel sided in basal 2 / 3, then inflated distally, with round apex; free ventral lobes small, about 1 / 5 length of valva, with round apex. Vinculum with two pairs of heavily sclerotized lateral lobes, symmetrical in both arms: posterior one, digitate, about 1 / 3 length of lateral arm, with round apex, densely setose; anterior one slightly shorter, spine-like; valvella setose, about equal to length of dorsal lobe. Saccal region narrow, slightly concave medially on anterior margin. Sica single, bifurcated apically, about equal to length of vinculum, nearly parallel sided. Aedeagus stout, as long as valva, nearly parallel-sided, bluntly rounded at base and apex; two short, sclerotized lobes from zone, at lateral side respectively, symmetrical, about 1 / 5 length of aedeagus, slightly bifurcate distally.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	distribution	Distribution. Vietnam (Vinh Phuc).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners by the dark-grayish brown forewing. It can be easily distinguished from congers by having the gnathos strongly bent into L-shape, vinculum with two pairs of strong lateral lobes, and a strong sica parallel sided, bifurcated apically.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE7FFB0FF7FFB0CFCCAC5E8.taxon	etymology	Etymology. The specific name is derived from the Latin, - sclero- (= sclerotized) and spinula (= spine), referring to the sclerotized spine on vinculum of the male genitalia.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	description	(Figs 27, 45, 54)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ninh Binh Prov.: Cuc Phuong Nat. Park, 200 m, 23 iv 2006, leg. Park, Kim & Kang, gen. slide no. nklep 003, in NIBR. Paratypes: VIETNAM: Ninh Binh Province: 2 ♂, same as holotype; 2 ♂, same locality as holotype, Bong, 450 m, 24 iv 2006, leg. Park, Kim & Kang; 4 ♂, same locality as holotype, 300 m, 30 iv – 01 v 2005, leg. KT Park & SR Kim, gen. slide no. nklep 004; Vinh Phuc Province: 1 ♀, Tam Dao Nat. Park, 450 m, 27 iv 2006, leg. Park, Kim & Kang, gen. slide no. CIS- 6287, in NIBR. CHINA: Hainan Province: 1 ♀, Diaoluoshan Nature Reserves (18.43 ° N 109.52 ° E), 922 m, 26. i. 2015, leg. P Cong, W Guan and S Hu, gen. slide no. ZSN 16242; 1 ♀, Bawangling (19.12 ° N 109.09 ° E), Changjiang County, 193 m, 11 iv 2018, leg. P Liu, X Bai & S Yu, in NKU.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	description	Description. Adult (Fig. 27). Wingspan 17.0 – 18.0 mm. Head dark brown dorsally, with yellowish erect scales laterally; frons pale gray. Ocellus absent. Antenna with scape dark brown on dorsal surface, yellowish white on ventral surface; flagellum alternately dark brown and yellowish brown on dorsal surface, pale yellow on ventral surface, ciliate. Second segment of labial palpus dark brown on outer surface, pale yellow on inner surface, with grayish white-tipped triangular scale tuft dorsally; 3 rd segment smoothly scaled, longer than 2 nd segment, pale yellow, dark brown on ventral surface. Thorax and tegula dark gray; mesothorax with golden hair pencils arising from anepisternum in male. Fore- and midleg dark brown on outer surface; hindleg grayish yellow with tibia clothed with long, grayish-yellow scales. Forewing gradually broadened distally; ground color pale yellow, scattered with brownish scales; discal and discocellular spots well-developed, dark brown, rounded and ill-defined plical spot placed at middle of fold; costa with dark-brown narrow streak from base extended to near 2 / 5 and pale-yellow narrow band along costal margin, abruptly oblique from prior to apex; ill-defined dark-brown fascia arising from 4 / 5 of costa extended to near tornus; apex blunt; termen oblique; fringe concolroue with narrow pale yellow basal line. Hind wing broader than forewing, dark grayish brown; fringe concolrous with ground color. Male genitalia (Fig. 45): Uncus subquadrate, with rounded posterior margin; anterior margin concave in semicircle. Gnathos strongly bent before middle, sharp at apex; culcitula rounded, densely spined. Valva longer than tegumen plus uncus, narrowed at base, broader toward apex; apex rounded; free ventral lobe digitate, about 1 / 4 length of valva, dentate marginally. Vinculum about equal to the length of tegumen plus uncus; lateral lobes very short, more or less triangular, with dentate margin sparsely setose, arising from basal 1 / 4 of lateral arms, shorter than 1 / 7 of arm. Saccal region broad, slightly concave at middle anteriorly. Sicae with uniquely developed lobes; left lobe hook-shaped apically, strongly bent at distal 1 / 3, reaching base of vinculum, with another shorter lobe directing to base outwardly; right lobe straight, about 2 / 5 the length of left one, fused in basal 1 / 3, then bifurcated in U shape. Aedeagus shuttle-shaped, rounded at base, sharply produced apically. broadest at zone; cornutus sickle-shaped, heavily sclerotized, about 1 / 3 the length of aedeagus. Female genitalia (Fig. 54): Apophyses anteriores about 1 / 3 the length of apophyses posteriors, inflated basally. Eighth sternite nearly straight on posterior margin. Antrum inverted trapeziform, weakly sclerotized. Ductus bursae short, broad, with complex of irregularly modified heavily sclerotized plates; left side with a short, triangular process and right side with a sword-like sclerotized plate extending to basal 2 / 5 of corpus bursae, forming an inverted U-shaped anterior margin between them; ductus seminalis arising from right side distally. Corpus bursae large, ovate; signum rounded, placed at anterior 1 / 3 ventrally, regions around signum with dense spines; accessory bursa arising from middle of signum.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	distribution	Distribution. China (Hainan), Vietnam (Ninh Binh, Vinh Phuc).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	diagnosis	Diagnosis. The new species can be distinguished from congeners by the forewing with abruptly oblique apically. The forewing pattern is more or less similar to D. longivalvata sp. nov., but it can be easily distinguished from the latter by the unique shape of apical region of the forewing. The male genitalia are similar to those of D. christophi Ponomarenko & Mey, 2002, but it can be separated from the latter by the different shape of sicae, with left lobe hook-shaped and right lobe bifurcated apically. The shape of sicae are slightly variable as illustrated in Fig. 46 (holotype) and Fig. 47 (paratype).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE6FFB3FF7FFDCCFC65C604.taxon	etymology	Etymology. The specific name is derived from the Latin, spinosus (= spinose), with a Latin dimunitive suffix, - ellus, referring to the sicae with a spinous process on inner margin.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	description	(Figs 28, 46)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ho Chi Minh City: Cat Tien Nat. Park, Thac Ngua Troi, 200 m, 09 ii 2006, leg. KT Park & N Cuong, gen. slide no. CIS- 6320, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	description	Description. Adult (Fig. 27). Wingspan 11.5 mm. Head dark yellowish brown dorsally, with yellowish scales laterally. Ocellus present. Antenna with scape dark brown on dorsal surface, grayish white on ventral surface; flagellum alternately dark brown and yellowish brown on dorsal surface, yellow on ventral surface. Second segment of labial palpus dark brown on outer surface, pale gray on inner surface, grayish white-tipped triangular scale tuft dorsally; 3 rd segment smoothly clothed, approximately equal to length of second segment, yellow except dark brown on ventral surafce. Thorax yellow, tegula dark brown in basal half, yellow in distal half; mesothorax without hair pencils arising from anepisternum in male. Fore- and midleg dark brown on outer surface except tarsomeres yellowish white apically, femora grayish white, tibiae and tarsi pale yellow on inner surface; hindleg pale yellow. Forewing slightly broadened distally; ground color pale yellow, sparesely scattered with fine brown scales; a small, dark-brown discal spot at 2 / 3 of cell; dark brown patch along termen from beyond apex before tornus; costa with blackish streak in basal 1 / 5 and ill-defined band along 4 / 5 of costal margin; apex acute; termen oblique; fringe concolorous with ground color. Hind wing broader than forewing, dark browinish gray. Male genitalia (Fig. 46). Uncus rather short, nearly quadrate, with rounded posterior margin; anterior margin deeply concaved in semicircular. Gnathos small, with median process as long as uncus. Valva about equal to the length of tegumen plus uncus, incised at distal 1 / 4 on dorsal margin, rounded at apex; valvella about 1 / 3 length of valva, inflated in distal half, setose apically. Setose patch short, irregular-shaped, inflated distally. Vinculum about 2 / 3 the length of tegumen plus uncus; lateral lobes long, about half the length of valva, arising from distal 3 / 4, rounded apically. Saccal region broad, nearly straight on anterior margin. Sica single, about 3 / 4 the length of vinculum, sparsely setose at apex. Aedeagus shuttle-shaped, rounded at base, broadest at zone, narrowed apically; dorsal margin sclerotized distally, produced; cornutus needle-shaped with round basal part, as long as 3 / 5 length of aedeagus. Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	distribution	Distribution. Vietnam (Ho Chi Minh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	diagnosis	Diagnosis. This species is very similar to Dichomeris quadratella sp. nov. in the forewing color pattern, but it can be separated from the latter by the head pale yellow, 3 rd segment of labial plapus lacking rough scales dorsally, the forewing lacking dark-brown fascia along dorsum.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE5FFB3FF7FFDE0FE4EC318.taxon	etymology	Etymology. The specific name is derived from the Latin, - uni (= one) and lobatus (= lobe), referring to the single siace in the male.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	description	(Figs 31, 47, 48)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	materials_examined	Type material. Holotype ♂, VIETNAM: Ninh Binh Prov.: Cuc Phuong Nat. Park, 300 m, 02 v 2005, leg. KT Park & SR Kim, gen. slide no. nklep 041, in NIBR.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	description	Description. Adult (Fig. 31). Wingspan 9.5 mm. Head yellowish brown. Ocellus absent. Antenna dark brown on dorsal surface, pale yellow on ventral surface. Second segment of labial palpus dark brown on dorsal surface, pale yellow in basal half, dark brown in distal half, with rough scales; 3 rd segment smoothly clothed, about equal to the length of 2 nd segment. Thorax yellowish brown; tegula dark brown; mesothorax without hair pencil arising from anepisternum in male. Fore- and midleg dark brown on outer surface, yellowish white on inner surface, tibiae with yellowish white rings at middle and end respectively; hindleg dark brown on outer surface, femur yellowish white on inner surface, tibia with basal half yellowish brown, distal half blackish bown on inner surface. Forewing dark brown; markings ill-defined, with distinct discal spots; a lager rounded black one at end of cell and smaller one near middle of cell, ill-defined; postmedian line zigzagged, slightly oblique; apex more or less acute; termen slightly convex medially; fringe dark brown. Hind wing broader than forewing; deep brown. Male genitalia (Fig. 47): Uncus extremely narrow in basal 3 / 4, with fan-shaped apical part, about 1 / 2 the length of valva. Gnathos long, longer than uncus, slightly upturned, heavily sclerotized. Valva shorter than tegumen plus uncus, narrowed at base, gradually broadened toward rounded apex. Setose patch banded. Vinculum with lateral arms heavily sclerotized along margin. Saccus triangularly produced. Aedeagus broadened in basal 3 / 5, then narrowed, elongated, bent process apically. Eighth tergite triangular, straight on anterior margin, bluntly rounded on posterior margin (Fig. 48). Female unknown.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	distribution	Distribution. Vietnam (Ninh Binh).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	diagnosis	Diagnosis. This species superficially resembles Helcystogramma flavifuscum Li & Zhen, 2011, but it differs from the latter by having gradually broadened valva, the cornute lateral lobe of vinculum, whereas in the latter, the valva is narrowest medially, inflated distally.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFE0FFA9FF7FFF1DFD96C55C.taxon	etymology	Etymology. The specific name is derived from the Latin - albus (= white) and macularis (= spot), referring to the white spot at costal 2 / 3 in the forewing.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFE50FD74C032.taxon	description	(Figs 1, 49)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFE50FD74C032.taxon	materials_examined	Material examined: VIETNAM: Bac Kan Province: 1 ♀, Ba Be Nat. Park, 26 – 28 vii 2006, leg. Park, Chae & Cuong, gen. slide no. CIS- 6283; 1 ♂, same locality, 255 m, 20 – 24 ix 2013, leg. JB Heppner, gen. slide no. nklep 057. Female genitalia (Fig. 84). Apophyses anteriores about 1 / 3 length of apophyses posteriores. Eighth tergite with anterior margin deeply concave in triangle, posterior margin convex in trapezoid; eighth sternite deeply concave in semicircular on posterior margin. Antrum ampullaceal, about same length with apophyses posteriores, gently sclerotized on anterior margin, deeply concave triangularly on posterior margin, extending onwards to middle of antrum, sclerotized marginally. Ductus bursae rectangular, about 2 / 7 length of apophyses posteriors, membranous. Corpus bursae large ovate, densely spinous on inner surface in posterior 4 / 5; two inverted triangular sclerotized lobes at entrance of corpus bursae ventrally, fused at base, left lobe extending to posterior 1 / 4 of corpus bursae, right lobe extending to posterior 1 / 5.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFE50FD74C032.taxon	distribution	Distribution. Vietnam (new record: Bac Kan), China (Hong Kong, Hubei).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFE50FD74C032.taxon	discussion	Remarks. The female is firstly described.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFBFEFC77C198.taxon	description	(Fig. 2)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFBFEFC77C198.taxon	materials_examined	Material examined. VIETNAM: Thua Thien-Hue Province: 2 ♂, Back Ma Nat. Park, 40 m, 18 vi 2004, leg. KT Park & N Cuong, gen. slide no. CIS- 6681, - 6718.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFBFEFC77C198.taxon	distribution	Distribution. Vietnam (new record: Thua Thien-Hue), Thailand.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFA5CFC07C338.taxon	description	(Fig. 3)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFA5CFC07C338.taxon	materials_examined	Material examined. VIETNAM: Ninh Binh Province: 1 ♂, 1 ♀, Cuc Phuong Nat. Park, 300 m, 30 iv – 02 v 2005, leg. KT Park & SR Kim, gen. slide no. CIS- 6279 (♀).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA9FF7FFA5CFC07C338.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh), China (Taiwan), India.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA8FF7FF8FCFAA6C55C.taxon	description	(Fig. 5)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFFFFA8FF7FF8FCFAA6C55C.taxon	distribution	Distribution. Vietnam (new record: Bac Kan, Ninh Binh), China (Yunnan), Thailand (Mae Hong Son).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFE19FA92C6BE.taxon	description	(Fig. 6)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFE19FA92C6BE.taxon	materials_examined	Material examined. VIETNAM: Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 450 m, 30 vii 2006, Park, Chae & Cuong, gen. slide no. CIS- 6289.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFE19FA92C6BE.taxon	distribution	Distribution. Vietnam (new record: Vinh Phuc), China (Hubei, Taiwan, Zhejiang), Japan, Indonesia, India.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFD7FFA8EC01B.taxon	description	(Fig. 8) Dichomeris foliforma Li & Park, 2017, In: Zhao et al.: 221. TL: China (Yunnan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFD7FFA8EC01B.taxon	materials_examined	Material examined: VIETNAM: Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 450 m, 27 iv 2006, leg. Park, Kim & Kang, gen. slide no. CIS- 6256.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFD7FFA8EC01B.taxon	distribution	Distribution. Vietnam (new record: Vinh Phuc), China (Guangxi, Hainan, Zhejiang), Cambodia (Pursat).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFBD2FC46C177.taxon	description	(Fig. 9)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFBD2FC46C177.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 1 ♂, Cuc Phuong Nat. Park, 200 m, 23. iv. 2006, leg. Park, Kim & Kang, gen. slide no. nklep 012.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFBD2FC46C177.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh), China (Sichuan),	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFA30FAA6C2ED.taxon	description	(Fig. 10) Dichomeris hainanensis Li & Park, 2017, In: Zhao et al.: 225. TL: China (Hainan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFA30FAA6C2ED.taxon	materials_examined	Material examined: VIETNAM: Vinh Phuc Province: 1 ♀, Tam Dao Nat. Park, 950 m, 24 v 2007. leg. KT Park, gen. slide no. nklep 002. Ninh Binh Province: 1 ♂, Cuc Phuong Nat. Park, 450 m, 24 iv 2006, leg. Park, Kim & Kang.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFA8FF7FFA30FAA6C2ED.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh, Vinh Phuc), China (Hainan), Cambodia (Mondul Kili).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFABFF7FF8CBFEE6C57E.taxon	description	(Fig. 14)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFABFF7FF8CBFEE6C57E.taxon	materials_examined	Material examined. VIETNAM: Thua Thien-Hue Province: 1 ♂, Back Ma Nat. Park, 400 m, 18. vi. 2004, LEG. K. T. Park & N. Cuong, gen. slide no. CIS- 6722.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFEFFABFF7FF8CBFEE6C57E.taxon	distribution	Distribution. Vietnam (new record: Thua Thien-Hue), China (Jiangxi, Yunnan, Taiwan), India, Sri Lanka, Indonesia.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFE3FFBDDC6F7.taxon	description	(Fig. 16)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFE3FFBDDC6F7.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 2 ♂, Cuc Phuong Nat. Park, 300 m, 30 iv – 02 v 2005, leg. KT Park & SR Kim, gen. slide no. nklep 013; Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 950 m, 24 v 2007, leg. KT Park.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFE3FFBDDC6F7.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh, Vinh Phuc), China (Guizhou).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFCB1FD0DC11C.taxon	description	(Fig. 18)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFCB1FD0DC11C.taxon	materials_examined	Material examined. VIETNAM: Vinh Phuc Province: 1 ♂, 1 ♀, Tam Dao Nat. Park, 450 m, 30 vii 2006, leg. Park, Chae & Cuong, gen. slide nos. nklep 031 (♂), CIS- 6290 (♀); 1 ♂, same locality, 750 m, 15 viii 2005, leg. KT Park, MY Kim & MY Chae. Ninh Binh Province: 1 ♂, Cuc Phuong Nat. Park, 200 m, 23 iv 2006, leg. Park, Kim & Kang, gen. slide no. CIS- 6253. Bac Kan Province: 1 ♂, Ba Be Nat. Park, 26 – 28 vii 2006, leg. Park, Chae & Cuong, gen. slide no. CIS- 6317. Ho Chi Minh City: 1 ♂, Cat Tien Nat. Park, Thac Ngua Troi, 200 m, 9 ii 2006, leg. KT Park & N. Cuong. Gen. slide no. CIS- 6257.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFCB1FD0DC11C.taxon	distribution	Distribution. Vietnam (new record: Bac Kan, Ho Chi Minh, Ninh Binh, Vinh Phuc), Cambodia (Sima), China (Guangdong, Hainan, Hong Kong, Taiwan, Yunnan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFAD8FC52C3F2.taxon	description	(Figs 19, 51)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFAD8FC52C3F2.taxon	materials_examined	Material examined. VIETNAM: Vinh Phuc Province: 2 ♀, Tam Dao Nat. Park, 1000 m, 04 – 05 v 2005, leg. KT Park & SR Kim, gen. slide no. CIS- 6312; 1 ♀, same locality, 750 m, 15 viii 2005, leg. KT Park, MY Kim & MY Chae, gen. slide no. nklep 060; 1 ♀, same locality, 450 m, 30 vii 2006, leg. Park, Chae & Cuong; 1 ♀, same locality, 950 m, 31 vii 2006, leg. Park, Chae & Cuong, gen. slide no. nklep 022; 1 ♂, same locality, 950 m, 24 v 2007, leg. KT Park, gen. slide no. nklep 036; 1 ♀, same locality, 950 m, 26 iv 2006, leg. Park, Kim & King, gen. slide no. nklep 035. Ninh Binh Province: 1 ♀, Cuc Phuong Nat. Park, 200 m, 23 iv 2006, leg. Park, Kim & King, gen. slide no. nklep 040. Female genitalia (Fig. 51). Apophyses anteriores inflated in basal 1 / 3, about 1 / 5 the length of apophyses posteriores. Eighth sternite deeply concave on anterior margin, slightly concave on posterior margin. Antrum membranous. Ductus bursae sclerotized along lateral margins, with two different sized, sclerotized band arising from basal 1 / 3 of ductus bursae, extending to posterior part of corpus bursae. Corpus bursae rounded, as long as ductus bursae; signum absent.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFAD8FC52C3F2.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh, Vinh Phuc), Thailand.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFDFFABFF7FFAD8FC52C3F2.taxon	discussion	Remarks. The female is described and illustrated for the first time.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFF1DFC52C6F1.taxon	description	(Figs 20, 52).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFF1DFC52C6F1.taxon	materials_examined	Material examined: VIETNAM: Bac Kan Province: 1 ♂, Ba Be Nat. Park, 26 – 28 vii 2006, leg. Park, Chae & Cuong. Vinh Phuc Province: 2 ♂, Tam Dao Nat. Park, 1000 m, 05 v 2005, leg. KT Park & SR Kim; 1 ♂, 1 ♀, same locality, 912 m, 13. viii. 2005, leg. KT Park, MY Kim & MY Chae, gen. slide nos. CIS- 6704 (♂), CIS- 6700 (♀); 1 ♀, same locality, 950 m, 31 vii 2006, leg. Park, Chae & Cuong, gen. slide no. CIS- 6701. Female genitalia (Fig. 52). Apophyses anteriores extremely short, triangularly produced. Eighth sternite weakly sclerotized, with nearly straight posterior margin. Antrum sub-rectangular, deeply concave in semicircular on anterior margin. Ductus bursae with a long, oblique digitate process in left side extending to conjunction with corpus bursae; ductus seminalis arising from basal 1 / 3 of ductus bursae on ventral surface, with a sclerotized ring at base. Corpus bursae large, rounded, densely spined on inner surface; signum ill-defined, placed at anterior 1 / 5; accessory bursa arising from middle of signum.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFF1DFC52C6F1.taxon	distribution	Distribution. Vietnam (new record: Bac Kan, Thua Thien-Hue, Vinh Phuc), China (Hong Kong).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFF1DFC52C6F1.taxon	discussion	Remarks. The female is described and illustrated for the first time.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFCB4FB0FC11E.taxon	description	(Fig. 22)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFCB4FB0FC11E.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 22 ♂, 3 ♀, Cuc Phuong Nat. Park, 300 m, 30 iv – 02 v 2005, leg. KT Park & SR Kim, gen. slide nos. nklep 016 (♂), nklep 017 (♀); 4 ♂, 1 ♀, same locality, 200 m, 22 – 23 iv 2006, leg. Park, Kim & Kang, gen. slide no. nklep 020 (♂); 1 ♀, same locality, 11 – 12. vi. 2004, leg. K. T. Park & N. Cuong, gen. slide no. CIS- 6720. Vinh Phuc Province: 4 ♂, Tam Dao Nat. Park, 950 m, 26. iv. 2006, leg. Park, Kim & Kang, gen. slide no. nklep 014; 1 ♂, same locality 1000 m, 05 v 2005, leg. KT Park & SR Kim, gen. slide no. nklep 021; 2 ♂, 1 ♀, same locality 950 m, 24 – 25 v 2007, leg. KT Park, gen. slide no. nklep 010 (♂); 1 ♂, same locality, 985 m, 01 – 07 v 2012, leg. JB Heppner.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFCB4FB0FC11E.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh, Vinh Phuc), China (Fujian, Jiangxi, Sichuan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFADEFBD7C27B.taxon	description	(Fig. 24)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFADEFBD7C27B.taxon	materials_examined	Material examined. VIETNAM: Thua Thien-Hue Province: 2 ♂, Back Ma Nat. Park, 1,100 m, 17 – 19 vi 2004, leg. KT Park & N. Cuong, gen. slide nos. CIS- 6699, CIS- 6702.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FFADEFBD7C27B.taxon	distribution	Distribution. Vietnam (new record: Thua Thien-Hue), China (Fujian, Jiangxi).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FF93DFF3AC3F1.taxon	description	(Fig. 26)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FF93DFF3AC3F1.taxon	materials_examined	Material examined: Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 450 m, 25 v 2007, leg. KT Park, gen. slide no. CIS- 6319.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFFCFFAAFF7FF93DFF3AC3F1.taxon	distribution	Distribution. Vietnam (new record: Vinh Phuc), China (Fujian, Guangdong, Guizhou, Hainna, Jiangxi, Zhejiang).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFAFFF7FFF1DFC52C6F1.taxon	description	(Figs 29, 55) Dichomeris splendiptera Li & Park, 2017, In: Zhao et al.: 217. TL: China (Yunnan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFAFFF7FFF1DFC52C6F1.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 2 ♂, Cuc Phuong Nat. Park, 200 m, 23 iv 2006, leg. Park, Kim & Kang, gen. slide nos. CIS- 6301, nklep 042. Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 950 m. 26 iv 2006, leg. Park, Kim & Kang, gen. slide no. nklep 027; 1 ♀, same locality, 1000 m, 04 v 2005, leg. KT Park & SR Kim, gen. slide no. CIS- 6677. Bac Kan Province: 1 ♂, Ba Be Nat. Park, 240 m, 21 – 26 vi 2008, leg. JB Heppner, gen. slide no. CIS- 6362. Female genitalia (Fig. 55). Apophyses anteriores about 1 / 3 length of apophyses posteriors, dilated distally. Antrum membranous. Ductus bursae not clearly defined from corpus bursa; ductus seminalis broad, arising from near conjunction with corpus bursae. Corpus bursae elongated, corpus-ductus bursae complex about 3 times length of apophyses posteriors; accessory bursae arising from anteriorly; signum absent.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFAFFF7FFF1DFC52C6F1.taxon	distribution	Distribution. Vietnam (new record: Bac Kan, Ninh Binh, Vinh Phuc), Cambodia (Mondul Kili), China (Hainan, Yunnan).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFAFFF7FFF1DFC52C6F1.taxon	discussion	Remarks. The female is described and illustrated for the first time.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFA0FF7FFCB4FBB3C5B5.taxon	description	(Fig. 30)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFA0FF7FFCB4FBB3C5B5.taxon	materials_examined	Material examined. VIETNAM: Vinh Phuc Province: 1 ♀, Tam Dao Nat. Park, 450 m, 30 vii 2006. leg. Park, Chae & Cuong, gen. slide no. CIS- 6321.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFA0FF7FFCB4FBB3C5B5.taxon	distribution	Distribution. Vietnam (new record: Vinh Phuc), China (Guizhou, Hubei, Taiwan, Yunnan), India. (Fig. 32)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFA0FF7FFCB4FBB3C5B5.taxon	materials_examined	Material examined: VIETNAM: Vinh Phuc Province: 1 ♂, Tam Dao Nat. Park, 950 m, 26 iv 2006. leg. Park, Kim & Kang, gen. slide no. nklep 010; 1 ♀, same locality, 24 v 2007. leg. KT Park, gen. slide no. nklep 011; 1 ♂, Tam Dao Nat. Park, 1000 m. 05 v 2005, leg. KT Park & S. R. Kim.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF9FFA0FF7FFCB4FBB3C5B5.taxon	distribution	Distribution. Vietnam (new record: Vinh Phuc), China (Gansu, Henan, Shaanxi).	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFE73FCC4C74F.taxon	description	(Fig. 33)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFE73FCC4C74F.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 1 ♂, 1 ♀, Cuc Phuong Nat. Park, 300 m, 02 v 2005, leg. KT Park & SR Kim, gen. slide no. CIS- 6258 (♂); 1 ♀, Cuc Phuong Nat. Park, near main gate, 200 m, 22 iv 2006, leg. Park, Kim & Kang.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFE73FCC4C74F.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh), China (Guizhou, Jiangxi, Taiwan, Zhejiang). Reamrks. This species is newly recorded for Vietnam.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFC42FC48C126.taxon	description	(Fig. 34)	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFC42FC48C126.taxon	materials_examined	Material examined: VIETNAM: Ninh Binh Province: 1 ♂, Cuc Phuong Nat. Park 300 m, 02. v. 2005, leg. KT. Park & SR. Kim, gen. slide no. nklep 025.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
03E3878EFFF6FFA0FF7FFC42FC48C126.taxon	distribution	Distribution. Vietnam (new record: Ninh Binh), China (Fujian, Gansu, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Jiangxi, Shaanxi, Sichuan, Xinjiang, Xizang), Russia, Europe.	en	Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan, Li, Houhun (2020): Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam. Zootaxa 4821 (3): 435-461, DOI: 10.11646/zootaxa.4821.3.2
