taxonID	type	description	language	source
03E37946FD51FFBDFE4CFB5FFB1088C9.taxon	description	MycoBank: MB 844536 Figs 3 – 4	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD51FFBDFE4CFB5FFB1088C9.taxon	diagnosis	Diagnosis Russula pseudopunicea sp. nov. differs from other species by the reddish brown, light brown to brownish orange pileus disc with a greyish yellow margin, small basidiospores (5.9 – 6.4 – 6.9 × 5.2 – 5.5 – 5.8 μm), short basidia (35 – 40 – 45 × 9.1 – 11 – 12 μm) and distinct position in the nrITS phylogenetic tree.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD51FFBDFE4CFB5FFB1088C9.taxon	etymology	Etymology The epithet ʻ pseudopunicea ʼ refers to its phylogenetic position relative to R. subpunicea.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD51FFBDFE4CFB5FFB1088C9.taxon	materials_examined	Material examined Holotype CHINA • Beijing, Miyun District, Sileng Mountains; 40 ° 28 ′ 23 ″ N, 117 ° 6 ′ 17 ″ E; elev. 713 m; in broadleaf forest dominated by Betula costata Trautv.; 30 Aug. 2021; C. L. Hou, G. Q. Cheng & H. Zhao; BJTC [BJTC ZH 1392]. Additional material CHINA • Beijing, Miyun District, Sileng Mountains; 40 ° 28 ′ 23 ″ N, 117 ° 6 ′ 18 ″ E; elev. 710 m; in broadleaf forest dominated by Betula costata; 30 Aug. 2021; C. L. Hou, G. Q. Cheng & H. Zhao; BJTC [BJTC ZH 1389] • Hebei Province, Chengde City, Xinglong County, Badaziling; 40 ° 18 ′ 36.6 ″ N, 117 ° 35 ′ 6.0 ″ E; elev. 879 m; in broadleaf forest dominated by Populus davidiana Dode; 22 Aug. 2020; C. L. Hou & G. Q. Cheng; BJTC [BJTC C 335] • Beijing, Huairou District, Sunzhazi Village; 40 ° 56 ′ 33.9 ″ N, 116 ° 30 ′ 25.4 ″ E; elev. 791 m; in coniferous and broad-leaved forest dominated by Populus davidiana and Pinus tabuliformis Carr.; 22 Aug. 2020; C. L. Hou & G. Q. Cheng; BJTC [BJTC C 565].	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD51FFBDFE4CFB5FFB1088C9.taxon	description	Description Basidiomata medium to large-sized. Pileus 35 – 113 mm in diam., hemispheric when young, applanate with a depressed centre at maturity; margin smooth or with indistinct striae, 3 – 12 mm, surface viscid when wet, cuticle peeling approximately to one-third of the pileus radius; reddish brown (# 9 a 603 e), light brown (# 8 e 4 d 1 e) to brownish orange (# c 5994 b), greyish brown (# a 79 e 9 a), margin greyish yellow (# ceb 98 f), pale yellow (# feeed 4) to light grey-brown (# fddaa 2). Lamellae cream (# ffffff) to pale yellow (# ffe 7 a 3), adnate to adnexed, approximately 11 – 14 pieces / cm near the pileus margin, not forked, lamellulae absent. Stipe 41 – 94 × 21 – 38 mm, white (# ffffff), grey-white (# d 3 cbba) to cream-white (# d 3 cfba), cylindrical, solid. Context 5 – 15 mm at the pileus centre, white (# ffffff), unchanging when bruised, odour not obvious, taste mild. Spore print not observed. Basidiospores (5.2 –) 5.9 – 6.4 – 6.9 (– 7.5) × (4.8 –) 5.2 – 5.5 – 5.8 (– 6.2) μm, Q = 1.16 ± 0.16, globose to ellipsoid, ornamented with small and amyloid warts of height 0.2 – 0.6 μm, which are usually connected by the lower line and sometimes form a partial reticulum; hilar appendix usually conspicuous, imamyloid. Basidia (32 –) 35 – 40 – 44 (– 50) × (7.4 –) 9.1 – 10 – 11 (– 12) μm, clavate, 2 -, 4 - spored, thin-walled, with guttate or granular contents; basidiola clavate or subcylindrical, ca 5 – 10 μm wide. Pleurocystidia dispersed to moderately numerous, (54 –) 57 – 68 – 76 (– 84) × (7.1 –) 7.4 – 8.4 – 9.4 (– 10.2) μm, cylindrical, clavate or subfusiform, apically mainly obtuse, presence of heteromorphous-crystalline, occasionally banded contents, turning grey to purplish red (# 694 f 89) in sulfovanillin. Cheilocystidia (50 –) 52 – 60 – 67 (– 76) × (5.8 –) 6.3 – 7.1 – 7.9 (– 9.2) μm, clavate, occasionally cylindrical or subfusiform, with crystalline or granulose, few banded contents. Marginal cells (15 –) 17 – 21 – 25 (– 27) × (4.0 –) 4.2 – 5.8 – 7.4 (– 11) μm, subcylindrical, often flexuous. Pileipellis orthochromatic in cresyl blue, sharply delimited from the underlying context, 210 – 410 μm thick, two-layered, suprapellis 60 – 190 μm thick, not gelatinized, composed of erect, repent or ascending and near the surface loose hyphal terminations, inflated at base and attenuated towards terminal cells; subpellis 150 – 270 μm thick, less gelatinized, composed of interwoven hyphae of width 2.5 – 5.5 μm. Hyphal terminations near the pileus margin not branched, occasionally flexuous, thin-walled; terminal cells (8.1 –) 9.2 – 22 – 36 (– 55) × (2.0 –) 2.4 – 3 – 3.6 (– 4.1) μm, subcylindrical to cylindrical, sometimes apically attenuated, subterminal cells often short or inflated, 2 – 8 μm wide, forked. Hyphal terminations near the pileus centre similar to those near the pileus margin, terminal cells (18 –) 20 – 29 – 38 (– 47) × (2.0 –) 2.4 – 2.8 – 3.2 (– 3.6) μm, mostly subcylindrical. Pileocystidia near the pileus margin (34 –) 38 – 53 – 67 (– 74) × (3.0 –) 3.4 – 4.2 – 5 (– 5.4) μm, thin-walled, cylindrical, subcylindrical or narrowly fusiform, sometimes with a 3 – 5 μm long appendage, contents heteromorphous granulose or banded, becoming purple (# 800080) in sulfovanillin; pileocystidia near the pileus centre (22 –) 25 – 39 – 53 (– 71) × (3.0 –) 3.2 – 4 – 4.8 (– 5.1) μm, thin-walled, cylindrical, subcylindrical or narrowly fusiform, apically mainly obtuse, sometimes with a 2 – 4 μm long appendage, contents heteromorphous-granulose. Cystidioid hyphae absent.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD5CFFB8FE53FBAEFD78889E.taxon	description	MycoBank: MB 844537 Figs 5 – 6	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD5CFFB8FE53FBAEFD78889E.taxon	diagnosis	Diagnosis Russula wulingshanensis sp. nov. differs from other species by the purple pinkish, pale brown or greyish white pileus disc with grey-white to grey-purple margin at maturity, small basidiospores (5.8 – 6.3 – 6.8 × 4.9 – 5.2 – 5.5), pleurocystidia turning pink in sulfovanillin and a separate phylogenetic position.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD5CFFB8FE53FBAEFD78889E.taxon	etymology	Etymology The epithet ʻ wulingshanensis ʼ refers to the locality where the type specimen was collected.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD5CFFB8FE53FBAEFD78889E.taxon	materials_examined	Material examined Holotype CHINA • Hebei Province, Xinglong County, Wulingshan Mountain National Nature Reserve; 40 ° 33 ′ 40.7 ″ N, 117 ° 28 ′ 44.9 ″ E; elev. 1248 m; in broadleaf forest dominated by Betula dahurica Pall.; 23 Aug. 2020; C. L. Hou & G. Q. Cheng; BJTC [BJTC C 403]. Additional material CHINA • Hebei Province, Xinglong County, Wulingshan Mountain National Nature Reserve; 40 ° 33 ′ 39.7 ″ N, 117 ° 28 ′ 44.7 ″ E; elev. 1248 m; in broadleaf forest dominated by Betula dahurica; 19 Aug. 2019; H. Zhao & J. Q. Li; BJTC [BJTC L 278] • ibid.; 40 ° 33 ′ 40.1 ″ N, 117 ° 28 ′ 44.7 ″ E; elev. 1256 m; in broadleaf forest dominated by B. dahurica; 23 Aug. 2020; C. L. Hou, G. Q. Cheng & R. T. Zhang; BJTC [BJTC C 399].	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
03E37946FD5CFFB8FE53FBAEFD78889E.taxon	description	Description Basidiomata small to medium-sized. Pileus 14 – 63 mm in diam., hemispheric at first, applanate with a depressed centre to infundibuliform at maturity, margin smooth when young, often fine striate at maturity, cuticle peeling often one-fifth of the pileus radius; purple pinkish (# ff 989 b), pale brown (# a 95 b 4 c) or greyish white (# ececea), margin grey-white (# e 9 e 9 e 6), grey-purple (# aaaad 5). Lamellae white (# fffff) to pale yellow (# ffffcc), adnate, fragile, up to 6 mm wide, approximately 10 – 12 pieces per cm near the pileus margin, lamellulae absent, usually forked near the stipe. Stipe 72 – 105 × 12 – 27 mm, solid, subcylindrical to slightly inflated at the base, sometimes light brown to brownish orange (# ae 6020) when bruised. Context unchanging when bruised, odour unpleasant, taste mild. Spore print not observed. Basidiospores (5.5 –) 5.8 – 6.3 – 6.8 (– 7.3) × (4.7 –) 4.9 – 5.2 – 5.5 (– 5.6) μm, Q = (1.02 –) 1.10 – 1.21 – 1.32 (– 1.45), subglobose to ellipsoid, ornamented with small and amyloid warts of height 0.2 – 0.9 μm; hilar appendix small, amyloid or weakly amyloid. Basidia (31 –) 35 – 40 – 45 (– 48) × (8.5 –) 9.4 – 11 – 12 (– 15) μm, 2 - and 4 - spored, clavate and slightly inflated in the upper half, thin-walled, with guttate or granular contents, sterigmata 4.3 – 5.8 × 0.9 – 1.2 μm; basidiola clavate or subcylindrical, 7 – 11 μm wide. Pleurocystidia dispersed, (48 –) 58 – 71 – 84 (– 95) × (8.6 –) 9.7 – 11 – 12 (– 14) μm, clavate, fusiform, subcylindrical sometimes lanceolate, apically always obtuse, few mucronate, sometimes with a 2 – 7 μm long appendage, thinwalled, contents granulose, heteromorphous-crystalline, pink in sulfovanillin. Cheilocystidia (33 –) 40 – 51 – 62 (– 67) × (6.2 –) 8.1 – 9.1 – 10 (– 11) μm, clavate and subfusiform, apically always obtuse, sometimes with 3 – 12 μm long appendage; contents granulose, heteromorphous-crystalline, often dispersed in the upper and middle layers. Marginal cells (20 –) 22 – 24 – 26 (– 29) × (4.9 –) 5.6 – 6.7 – 7.8 (– 9.4) μm, clavate or subfusiform, sometimes fusiform. Pileipellis orthochromatic in cresyl blue, sharply delimited from the underlying context, 120 – 180 μm thick, two-layered, strongly gelatinized, suprapellis 50 – 80 μm thick, composed of ascending to erect hyphal terminations; subpellis 70 – 100 μm thick, composed of horizontally oriented, interwoven hyphae. Hyphal terminations near the pileus margin intricate and sometimes branched, occasionally flexuous, thin-walled; terminal cells (11 –) 15 – 21 – 27 (– 31) × (3.0 –) 3.3 – 4.1 – 4.9 (– 5.9) μm, mostly subcylindrical, occasionally subfusiform or lageniform, apically usually obtuse, sometimes attenuated or constricted, subterminal cells frequently shorter and slightly inflated, ca 4 – 7 μm wide, rarely branched. Hyphal terminations near the pileus centre similar to those near the pileus margin; terminal cells (12 –) 18 – 26 – 35 (– 44) × (2.1 –) 3.5 – 4.4 – 5.3 (– 6.2) μm, subcylindrical, subfusiform or lageniform, apically usually obtuse, sometimes attenuated or constricted, subterminal cells frequently short, 2 – 6 μm wide, rarely branched. Pileocystidia near the pileus margin (35 –) 39 – 50 – 62 (– 68) × (5.0 –) 5.2 – 5.8 – 6.4 (– 7.5) μm, thin-walled, cylindrical, clavate or subfusiform, apically usually obtuse, sometimes with a 2 – 5 μm long appendage, contents heteromorphous-granulose, black in sulfovanillin. Pileocystidia near the pileus centre (32 –) 41 – 50 – 59 (– 71) × (4.3 –) 4.7 – 5.8 – 6.9 (– 8.9) μm, thin-walled, apically usually obtuse, sometimes with 1 – 4 μm long appendage, contents with granulose or heteromorphous-crystalline, black in sulfovanillin. Cystidioid hyphae dispersed in subpellis and context, with heteromorphous-granulose contents, oleiferous hyphae frequent in subpellis and context.	en	Zhou, Hao, Cheng, Gui-Qiang, Huang, Xiao-Bo, Hou, Cheng-Lin (2023): Two new species of Russula subgenus Heterophyllidia (Russulaceae, Russulales) from Yanshan Mountains, North China. European Journal of Taxonomy 861: 185-202, DOI: 10.5852/ejt.2023.861.2077, URL: http://dx.doi.org/10.5852/ejt.2023.861.2077
