identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E08794FFC3FFFEFF5B7E49FEDFF857.text	03E08794FFC3FFFEFF5B7E49FEDFF857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anacochylidia Brown 2019	<div><p>Anacochylidia Brown, new genus</p><p>Type species: Anacochylidia maderana Brown, new species, by monotypy.</p><p>Diagnosis. The short broad valvae and the long, slender secondary process of the phallus of Anacochylidia (Fig. 19) are most similar to those of Cochylidia Obraztsov, 1956, but other features of the male genitalia deviate from those of all known Cochylidia . In Anacochylidia the top of the tegumen is broad with a shallow, rounded concavity; the costa of the valva is short, unmodified, and upcurved, without a basal process; and the phallus has an elongate secondary process ca. 0.5 times as long as the phallus itself. In contrast, the male genitalia of Cochylidia are characterized by a dorsally attenuate tegumen, often with a narrowly bifurcate process at the top; the base of the costa of the valva has a long, erect, somewhat digitate, setose process; and the phallus has a short, curved digitate process apically. The latter is probably homologous with the extraordinarily long process of the phallus of Anacochylidia.</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna about 0.9 times flagellomere diameter in male, shorter, sparser in female; labial palpus porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus inconspicuous. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing (Fig. 1) length 4.5–4.8 mm, length ca. 2.8 times width; costa slightly and evenly arched throughout, without costal fold in male, apex rounded; termen obliquely angled toward hind margin; Sc ca. 0.45 times wing length; R 1 originating from about middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa well before apex; M 3 and CuA 1 separate; CuA 2 originating about 0.75 times length of discal cell; CuP absent; A1+2 stalked in distal 0.65. Hindwing length ca. 2.5 times width; costa weakly undulate, male with narrow costal roll in basal 0.5 of wing; apex rounded; termen slightly concave near middle; Sc+R 1 ca. 0.25 times wing length, Rs and M 1 stalked ca. 0.3 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.6 length of discal cell; frenulum with one acanthus in male, two acanthi in female. Abdomen: Unmodified. Male genitalia (Fig. 19) with tegumen short, broad, with shallow, rounded excavation at top bordered laterally by short lobelike processes; subscaphium subrectangular, semi-sclerotized; socius inconspicuous; transtilla well developed, broad, weakly bilobed with small rounded median lobe; valva short, broad, subquadrate, pointed apically, outer margin undulate, length ca. 0.4 times width; costa sclerotized, curved dorsally; sacculus represented by narrow, curved, sclerotized edge, lacking free apical process; vinculum arms weakly joined by membrane distally. Phallus elongate, ca. 4.0 times length of valva, strongly curved in basal 0.4, with slender curved spine originating dorsally just beyond curve, extending ca. 0.5 length of phallus, vesica with dense bundle of 7–8 small, non-deciduous cornuti. Female genitalia (Fig. 33) with ovipositor long, slender, telescopic; papillae anales slender, slightly pointed posteriorly; apophyses posteriores ca. 2.0 times as long as apophyses anteriores; sterigma mostly membranous, lamella postvaginalis a subquadrate plate, attenuate and distinctly emarginate posteriorly, with a pair of narrow, submedial sclerites extending anteriorly from plate; colliculum sclerotized with longitudinal wrinkles, occupying entire ductus bursae; corpus bursae rounded, differentiated from ductus bursae by absence of sclerotization, signum lacking; origin of accessory bursa uncertain.</p><p>Etymology. The name comes from the similar genus Cochylidia and the Greek “ana” (= against) and is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC3FFFEFF5B7E49FEDFF857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC2FFFFFF5B7C99FD5FFB9A.text	03E08794FFC2FFFFFF5B7C99FD5FFB9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anacochylidia maderana Brown 2019	<div><p>Anacochylidia maderana Brown, new species</p><p>Figs. 1, 19, 33</p><p>Diagnosis. Anacochylidia maderana is a small species with a narrow triangular forewing and somewhat distinctive maculation: the ground color is white, with an ill-defined, pale rust patch in the subterminal region and an ill-defined, concolorous, oblique dash from the middle of the hind margin extending to about the middle of the discal cell (Fig. 1). The species is superficially similar to Cochylis temerana (Busck, 1907) and Mimcochylis fulvotincatana (Walsingham, 1884), but the genitalia have little in common with those species. Anacochylidia maderana is easily distinguished by the male genitalia (Fig. 19), which have a broad, concavely rounded top of the tegumen; a very short, wide valva similar to species of Cochylidia; and a long, curved, slender, apically-attenuate rod from the phallus, extending ca. 0.5 the length of the phallus. The male genitalia lack the conspicuous elongate process from the base of the costa of the valva that is present in all species Cochylidia .</p><p>Description. Head: Frons and vertex buff; labial palpus light buff with scattered fuscous scales, cream on inner surface; antenna mixed fuscous and cream, sensory setae about 0.9 times flagellomere diameter in male, shorter in female. Thorax: Mesonotum and tegula buff in anterior 0.3, remainder pale fuscous; forewing length 4.5–4.8 mm (n = 4); forewing ground white, with ill-defined, pale rust, submedian dash extending from near middle of hind margin to lower edge of discal cell; small concolorous spot at costa ca. 0.6 distance from base to apex; subterminal area with broad concolorous patch extending from costa almost to tornus. Fringe white. Hindwing cream-white. Fringe concolorous with wing. Abdomen: Dorsal scaling brown, venter shiny white. Male genitalia (Fig. 19) as described for genus. Female genitalia (Fig. 33) as described for genus.</p><p>Types. Holotype ³, Madera Canyon, [Santa Cruz Co.], Arizona, 3‒5 Sep 1969, J. Powell (EME) . Paratypes (6³, 1♀). USA: Arizona: Cochise Co.: Palmerlee, no date (5³, 1♀) (USNM). Santa Cruz Co.: Madera Canyon, 3‒5 Sep 1969 (1³), J. Powell (EME) .</p><p>Etymology. The specific epithet refers to the type locality of Madera Canyon in Arizona.</p><p>Distribution and biology. This species is known from Palmerlee (Cochise Co.) (n = 7) (USNM) and Madera Canyon (Santa Cruz Co.) (n = 2) (EME) of southern Arizona. The only two specimens with capture data are from September. Nothing is known of the early stages.</p></div>	https://treatment.plazi.org/id/03E08794FFC2FFFFFF5B7C99FD5FFB9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC2FFF8FF5B78ADFB0CFC29.text	03E08794FFC2FFF8FF5B78ADFB0CFC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atroposia Brown 2019	<div><p>Atroposia Pogue, new genus</p><p>Type species: Conchylis oenotherana Riley, 1881, by monotypy.</p><p>Atroposia Pogue and Mickevich, 1990: 322 (nomen nudum).</p><p>Cochylini New Genus 1 [ oenotherana group]: Brown 2005: 207; Metzler and Brown 2014: 278.</p><p>Atroposia is proposed to accommodate the single species oenotherana . Originally described in Conchylis Sodoffsky, 1837 (an unjustified emendation of Cochylis), the species was assigned to Phalonia Hübner, [1825], 1816 by McDunnough (1939) and placed in “incertae sedis” by Powell (1983). Pogue (1986) proposed the genus Atroposia for the species in his unpublished Ph.D. dissertation; Razowski (1997) treated oenotherana under “Unplaced Cochylini ”; and Brown (2005) and Metzler &amp; Brown (2014) treated it as “ Cochylini New Genus”. According to Pogue (1986), Atroposia is most closely related to Spinipogon among North American genera. However, based on facies and genitalia, Atroposia appears to be related to Cochylis .</p><p>Diagnosis. Atroposia is easily distinguished from other genera by facies (Figs. 3, 4), which feature a somewhat two-toned forewing with a yellow-tan basal half and a pinkish red distal half with a variably developed yellowish spot. However, in some specimens both the basal and distal halves of the wing are overscaled with dark gray, reducing the two-toned pattern. Putative autapomorphies of Atroposia include the unusual shape of the valva with a distinctly pointed costa and sacculus; the distal portion of the phallus, with a pair of long, pointed, flattened, lateral sheaths and three platelike internal sclerites; and the broad sterigma composed of four fused spheroid plates. The absence of the male hindwing costal roll in A. oenotherana distinguishes it from most species of Cochylis and related genera (e.g., Cochylichroa, Cochylidia, Thyraylia Walsingham, 1897).</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with appressed scales; ocellus present; sensory setae of antenna 1.0–1.2 times flagellomere diameter in male, shorter in female; labial palpus porrect, com- bined length of all segments 1.6–1.8 times diameter of compound eye, scaling of segment II concealing segment III; maxillary palpus 1-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing in male slightly broadened distally with costa weakly convex and termen straight, moderately oblique, forewing not expanding distally in female, apex rounded in both; forewing length 3.8–5.4 mm, length 2.5–3.4 times width, slightly wider in female; without costal fold in male; Sc less than 0.5 wing length; R 1 originating at or beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating about 0.75 length of discal cell; CuP absent; A1+2 stalked at 0.5 length. Hindwing length 2.5–2.8 times width; costa straight, without costal roll; apex rounded; termen straight, oblique; Sc+R 1 ca. 0.75 wing length, Rs and M 1 stalked ca. 0.65 length of M 1; M 3 and CuA 1 separate; CuA 2 originating 0.65 length of discal cell; male frenulum with one acanthus, female with two. Abdomen: Unmodified. Male genitalia (Fig. 20) with tegumen short, broad, convexly rounded at top; uncus absent; socius small, digitate, setose; transtilla a well-developed band, with triangulate to hooklike median process, keeled ventrally; valva highly modified, short, broad basally, with well-developed costal and saccular portions; costal portion broad at base, constricted apically into a robust, curved spine; saccular half with a short, blunt dorsal process and a longer, robust, pointed basal process; inner base of sacculus with elongate, pointed, extension confluent with arms of vinculum; vinculum arms free distally; juxta large, subrectangular, with a pair of dorso-lateral lobes. Phallus large, length about equal to basal width of valva, distally with three median platelike processes covered with minute spicules, subtended laterally by a pair of flattened, distally-pointed flanges extending ventro-terminally; vesica with minute cornuti. Female genitalia (Fig. 34) with ovipositor short; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 1.2 times length of apophyses posteriores; sterigma large, composed of four fused spheroid plates, sterigma not attached to apophyses anteriores; ductus bursae short, broad, undifferentiated from corpus bursae; corpus bursae a short convoluted sac lacking a signum; accessory bursa originating from ductus bursae.</p><p>Etymology. The name comes from the Greek “atropos, meaning without turn. In Greek mythology, Atropos (or Aisa) was one of the three Moirai, the goddess of fate and destiny. The name is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC2FFF8FF5B78ADFB0CFC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC5FFF8FF5B7841FBDEF80C.text	03E08794FFC5FFF8FF5B7841FBDEF80C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atroposia oenotherana (Riley 2019) Brown 2019	<div><p>Atroposia oenotherana (Riley, 1881), new combination</p><p>Figs. 3, 4, 20, 34</p><p>Conchylis oenotherana Riley, 1881: 316 .</p><p>Phalonia oenotherana: McDunnough 1939: 60 .</p><p>[Incertae sedis] oenotherana: Powell 1983: 42; Razowski 1997: 137.</p><p>[ Cochylini New Genus 1] oenotherana: Brown 2005: 207; Metzler &amp; Brown 2014: 278.</p><p>Atroposia oenotherana is one of the more easily recognized Cochylina in the Nearctic fauna, with the basal half of the forewing yellow-tan and the distal half pinkish red with a variably developed, yellow-tan spot. The species was redescribed in detail by Razowski (1997: 137). The genitalia are highly divergent from those of other Cochylina, and on the basis of the distinct facies and unusual genitalia, oenotherana is placed in a monotypic genus.</p><p>Atroposia oenotherana is widely distributed from Nova Scotia to Manitoba, south to Florida and Texas. Adults have been collected mostly in July and August, with fewer records in May, June, and September, suggesting a single generation in the north and two generations in the southern portions of its range. Larvae are tip-tiers in the flowers of Oenothera species ( Onagraceae) (Riley 1881; Godfrey et al. 1987; many reared specimens in USNM); Oenothera biennis (L.) has been recorded as the larval host in Georgia (USNM).</p><p>The illustration of the male genitalia of A. oenotherana (from Vineland, Ontario, Canada) in Razowski (1997: figs. 123) deviates in several features (e.g., the shape of the median process of the transtilla, the shape of the juxta, the curvature of the phallus) from those of Fig. 3 (from Padre Island National Seashore, Texas, USA). Because barcodes of A. oenotherana are represented by two BINs in BOLD (Brown et al. 2019), it is possible that two species are concealed under this name, and these differences in genital morphology reflect two different species. Alternatively, the male genitalia shown in Fig. 3 and in Razowski (1997) may merely represent individual variation and/or artifacts of slide mounting.</p><p>Syntype ♀, USA, Missouri, r.f. Oenothera, M. Murtfeld (USNM). Riley (1881) referred to four specimens in his original description, but I have been able to locate only a single female syntype .</p></div>	https://treatment.plazi.org/id/03E08794FFC5FFF8FF5B7841FBDEF80C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC7FFFAFF5B7C99FAEFF9A4.text	03E08794FFC7FFFAFF5B7C99FAEFF9A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cagiva Brown 2019	<div><p>Cagiva Pogue, new genus</p><p>Type species: Phalonia cephalanthana Heinrich, 1921, by monotypy.</p><p>Cagiva Pogue and Mickevich, 1990: 322 (nomen nudum).</p><p>Cochylini New Genus 2 [ cephalanthana group]: Brown 2005: 208; Metzler and Brown 2014: 278.</p><p>Cagiva is proposed for Phalonia cephalanthana, a species that has long defied confident generic assignment. The species was described in Phalonia by Heinrich (1921) and placed in “incertae sedis” by Powell (1983). The monotypic genus Cagiva was proposed by Pogue (1986) in his unpublished Ph.D. dissertation. Brown (2005) and Metzler and Brown (2014) followed that concept, designating the manuscript name Cagiva as “ Cochylini New Genus”.</p><p>Diagnosis. Putative autapomorphies for Cagiva include the prominent, well-sclerotized socii, and the distally bifurcate phallus. The genus is further characterized by a unique combination of characters, some of which are shared with other genera: the 3-segmented maxillary palpi (shared with Thyraylia, Nycthia, and few others), the absence of cornuti in the vesica (shared with Eupinivora Brown, 2013, Honca, and Nycthia), and the connate condition of hindwing veins M 3 and CuA 1 (shared with a few other genera). As in most Cochylina, the male of Cagiva lacks a forewing costal fold and has a well-developed hindwing costal roll.</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna 0.8–1.0 times flagellomere diameter in male, much shorter in female; labial palpus porrect, combined length of all segments 1.1–1.2 times diameter of compound eye, segment III exposed; maxillary palpus 3-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing (Fig. 2) length 4.0– 4.7 mm, length 2.4–2.8 times width, slightly wider in female; costa gently arched from base to apex, without costal fold in male, apex abruptly rounded; termen straight; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, length 2.3–2.8 times width; costa straight, with costal roll extending about 0.65 length of costa in male; apex rounded; termen slightly concave below apex; Sc+R 1 ca. 0.25 wing length in male, 0.65–0.75 wing length in female; Rs and M 1 stalked ca. 0.7 length of M 1; M 3 and CuA 1 connate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female.Abdomen: Unmodified. Male genitalia (Fig. 21) with uncus absent; socius prominent, well sclerotized, setose, distally elongate and curved mesially; transtilla well developed, median process a small rounded tooth; valva length 1.4‒1.5 times width; costa sclerotized, relatively straight; sacculus well defined, confined to basal 0.33 of venter of valva, terminating in a rounded lobe; vinculum arms weakly join distally by thin membrane. Phallus large, ca. 1.3 times length of valva, straight, parallel-sided in basal 0.5, broadened in distal 0.5, bifurcate apically, asymmetric; vesica without cornuti. Female genitalia (Fig. 35) with telescopic ovipositor; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 2.0 times that of apophyses posteriores, ventral branch of apophyses anteriores reduced, apophyses with rounded swelling at anterior tip; sterigma complicated, comprised of weakly sclerotized sacs, some bearing setae, and a crescent-shaped plate posterad of ostium; sterigma not attached to apophyses anteriores; ductus bursae relatively broad, rather uniform in width, ca. 0.65 as long as corpus bursae, differentiated from corpus bursae, with caudal 0.7 weakly sclerotized; accessory bursa originating near ostium; corpus bursae oblong, covered with minute spicules, signum lacking.</p><p>Etymology. The name is an arbitrary combination of letters and is considered feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC7FFFAFF5B7C99FAEFF9A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC7FFFBFF5B7AB4FE33FE15.text	03E08794FFC7FFFBFF5B7AB4FE33FE15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cagiva cephalanthana (Heinrich 1921) Brown 2019	<div><p>Cagiva cephalanthana (Heinrich, 1921), new combination</p><p>Figs. 2, 21, 35</p><p>Phalonia cephalanthana Heinrich, 1921: 825; McDunnough 1939: 59.</p><p>[Incertae sedis] cephalanthana: Powell 1983: 42 .</p><p>[ Cochylini New Genus 2] cephalanthana: Brown 2005: 208; Metzler &amp; Brown 2014: 278.</p><p>Cagiva cephalanthana is a small (forewing length 4.0–4.7), rather dark species with a white to cream colored head, described from a series of specimens from Lake Charlotte, Texas. The forewing pattern is ill-defined, somewhat irregularly checkered with copper, pink, and reddish brown, with a darker brown to black median fascia (Fig. 2). I have examined specimens from Texas (USNM), Oklahoma (BDGU), Missouri (USNM), Indiana (MSC), Maryland (MSC), Rhode Island (MSC), and Mississippi (USNM). Specimens from Texas and Mississippi were reared from the seed pods of button-bush, Cephalanthus occidentalis L. ( Rubiaceae). The specimens from Mississippi were collected as larvae in August and emerged in September; the specimens from Texas were collected as larvae in early September and emerged in late September and October. Field-collected adults were taken in July and August. It is assumed that there are two broods annually, with adults in July‒August and September‒October.</p><p>Holotype ³, USA, Texas, Chambers Co., shores of Lake Charlotte, r.f., Cepahlanthus occidentalis, em: 16 Sep 1918, C. Heinrich (USNM).</p></div>	https://treatment.plazi.org/id/03E08794FFC7FFFBFF5B7AB4FE33FE15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC6FFFBFF5B7E25FADAF8DD.text	03E08794FFC6FFFBFF5B7E25FADAF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybilla Brown 2019	<div><p>Cybilla Pogue, new genus</p><p>Type species: Phalonia hubbardana Busck, 1907, by monotypy.</p><p>Cybilla Pogue and Mickevich, 1990: 322 (nomen nudum).</p><p>Cochylini New Genus 3 [ hubbardana group]: Brown 2005: 208; Metzler and Brown 2014: 278.</p><p>Cybilla is proposed for the single species Phalonia hubbardana, which traditionally has defied generic assignment. Described in Phalonia by Busck (1907), it was treated as such by McDunnough (1939) and was placed in “incertae sedis” by Powell (1983). Pogue (1986) proposed the name Cybilla, and Brown (2005) and Metzler and Brown (2014) treated it as “ Cochylini New Genus”. Pogue (1986) concluded that Cybilla is most closely related to Decuma (now recognized as Monoceratuncus), and that the two are the most derived Cochylina genera in North America.</p><p>Diagnosis. Cybilla shares with Monoceratuncus an unusually modified uncus, the absence of socii, and the absence of a median process of the transtilla. Cybilla can be distinguished from Monoceratuncus by the presence of small lateral plates fused to the base of the uncus, and by the 4-segmented maxillary palpi.</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna ca. 1.0 times flagellomere diameter in male, shorter, sparser in female; labial palpus upturned, porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus 4-segmented. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing (Figs. 5, 6) length 4.6–5.6 mm, length 3.0–3.1 times width, females usually slightly larger than males; costa straight, without costal fold in male, apex rounded; termen gently curved; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.75 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, costa straight, with costal roll extending about 0.4 length of costa in male; apex rounded; termen straight; Sc+R 1 ca. 0.65 wing length; Rs and M 1 stalked at 0.65–0.75 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female. Abdomen: Unmodified. Male genitalia (Fig. 22) with well-developed uncus bearing small, closely appressed, flat, lateral plates; socius absent; transtilla well developed, concave mesially, lacking median process; valva broad at base, attenuate distally, length ca. 2.1 times width; costa setose, apex produced, sacculus undulate from base to apex of valva, with patch of small spines at outer margin of first undulation from base; vinculum arms free distally. Phallus large, ca. 1.1 times length of valva, straight, parallel-sided in basal 0.33, enlarged in distal 0.5, with ventral spadelike projection; vesica with numerous cornuti. Female genitalia (Fig. 36) with papillae anales curved near middle, bulbous, slightly bilobed, setose; apophyses short, slender, length of apophyses anteriores ca. same as apophyses posteriores, ventral branch of apophyses anteriores joined to sterigma; sterigma a lateral band with prominent short-digitate, caudally-directed, subtriangular median process; ductus bursae slender, membranous throughout; corpus bursae small, rounded, with minute spicules; accessory bursa originating dorsomedially from corpus bursae.</p><p>Etymology. The generic name is from the Latin goddess of nature, Cybele, and is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC6FFFBFF5B7E25FADAF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC6FFF4FF5B7B62FEE3FD46.text	03E08794FFC6FFF4FF5B7B62FEE3FD46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybilla hubbardana , Lordsburg	<div><p>Cybilla hubbardana (Busck, 19097), new combination</p><p>Figs. 5, 6, 22, 36</p><p>Phalonia hubbardana Busck, 1907: 27; McDunnough 1939: 60.</p><p>[Incertae sedis] hubbardana: Powell 1983: 42 .</p><p>[ Cochylini New Genus 3] hubbardana: Brown 2005: 208; Metzler &amp; Brown 2014: 278.</p><p>Cybilla hubbardana is a poorly known species recorded from Arizona to western Texas. It was described from a small series of specimens from Tucson, Arizona reared from Koeberlinia spinosa (Koeberliniaceae) (Busck 1907) (USNM). In general, the forewing of C. hubbardana has a pale cream ground color faintly reticulated with gray specks and a variably defined (usually ill-defined), slightly darker gray median fascia (Fig. 5, 6). Specimens from Texas (USNM) have the basal half slightly paler buff, creating a slightly two-toned appearance between the basal and distal halves, divided by the median fascia. The male and female genitalia (Figs. 22, 36) are as described for the genus.</p><p>Reared specimens from Arizona eclosed primarily in May. Field collected specimens from Texas were collected in late May and early June.</p><p>Holotype ♀, USA, Arizona, Pima Co., Tucson, r.f. Koeberlinia spinosa, em: 12 May 1897, H. Hubbard (USNM).</p></div>	https://treatment.plazi.org/id/03E08794FFC6FFF4FF5B7B62FEE3FD46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC9FFF5FF5B7EC8FB06FDDD.text	03E08794FFC9FFF5FF5B7EC8FB06FDDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Honca Brown 2019	<div><p>Honca Pogue, new genus</p><p>Type species: Phalonia grandis Busck, 1907, by monotypy.</p><p>Honca Pogue and Mickevich, 1990: 322 (nomen nudum).</p><p>Cochylini New Genus 4 [ grandis group]: Brown 2005: 208; Metzler and Brown 2014: 278.</p><p>Honca is described to accommodate the single species Phalonia grandis, which is among the largest Cochylina in the Nearctic region. The forewing pattern (Fig. 7) somewhat resembles that of Henricus umbrabasanus, however, H. grandis is much larger (forewing length 11‒13 mm vs. 8‒10 mm) and lacks a pale brown basal patch. Honca also resembles Aphalonia Razowski, 1984, a South American genus, in size and forewing maculation, however, the genitalia of the two have little in common.</p><p>Diagnosis. The shapes of the valva and transtilla of Honca are similar to those of some species of Aethes, however, Honca lacks the sickle-shaped socii that are characteristic of most species of Aethes . The most conspicuous autapomorphy of Honca is the highly modified phallus-juxta complex in the male genitalia (Fig. 23): the juxta is rather long, slender, subrectangular, and widely bifurcate distally, and is strongly attached to the large, curved phallus. The phallus has a long, flattened process of the right side that is as long as the phallus, and a short, stout, more rounded process on the left side that is about one-fourth the length of the phallus. The latter features are reminiscent of the paired process that subtend the phallus in some species of Aethes . In the female genitalia the spiculate, clubshaped arms of the sterigma (Fig. 37) are somewhat reminiscent of some species of Eupinivora . Honca is further characterized by the absence of ocelli (usually present in Cochylina), a 3-segmented maxillary palpus (shared with Eugnosta), a reduced haustellum (shared with Eugnosta), four acanthi in the female frenulum (occurring consistently nowhere else in Cochylina), fusion of the socii with the tegumen, and the absence of cornuti in the vesica (shared with Cochylis, Thyraylia, and Cochylichroa, and few other genera).</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus absent; sensory setae of antenna about as long as flagellomere diameter; labial palpus porrect, combined length of all segments ca. 3 times horizontal diameter of compound eye, shorter in female, segment III nearly concealed by scaling of segment II; maxillary palpus 3-segmented. Thorax: Posterior crest absent. Forewing length 11.0– 12.5 mm in male, 11.5–13.0 mm in female; length 2.4–2.8 times maximum width; forewing with base of costa straight, gently curved in distal 0.33 to rounded apex, costal fold absent; termen straight, oblique; Sc less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 nearer to R 3 than to R 1; R 5 terminating at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.8 length of discal cell; CuP absent; A1+2 stalked about 0.5 of their lengths. Hindwing with costa mostly straight, gently arched in apical 0.33, male without costal roll, termen slightly concave beneath apex; Sc+R 1 ca. 0.8 wing length; Rs and M 1 stalked to ca. 0.6 length; M 3 and CuA 1 separate; CuA 2 originating ca. 0.66 length of discal cell; frenulum with one acanthus in male, four in female. Abdomen: Unmodified. Male genitalia (Fig. 23) with uncus absent; socius prominent, fused to tegumen; transtilla with large, triangular, median process, with strongly sclerotized, pointed tip; valva short, broad, with upturned, well-sclerotized costa and attenuate apex, sacculus a straight sclerotized margin along venter of valva with slightly produced distal tip; vinculum arms con- nected distally by membrane; juxta a complicated plate, narrow, subrectangular, deeply emarginate on distal 0.35 distally resulting in pair of pointed processes, basal end fused to phallus. Phallus large, ca. 0.8 times length of valva, subtended by long, flattened lateral process on right side and shorter, peglike process on left (possibly homologous with “anellus lobes” of Sabourin et al. 2002), coecum large; vesica without cornuti. Female genitalia (Fig. 37) with papillae anales weakly curved, sclerotized; apophyses short, stout, apophyses anteriores ca. 1.1 times length of apophyses posteriores; sterigma membranous, median area with a pair of mesially-directed, spiculate, slightly club-shaped processes originating from posterior base of apophyses anteriores, each with 15‒20 long setae; ostium bursae wide; colliculum absent; ductus bursae short, undifferentiated from corpus bursae; corpus bursae with vertical lines of sclerotization below ostium bursae and with large patch of minute spicules; frail accessory bursae originating anterad of middle of ductus bursae.</p><p>Etymology. The generic name is an arbitrary combination of letters and is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC9FFF5FF5B7EC8FB06FDDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC8FFF5FF5B7E6DFAEBFB39.text	03E08794FFC8FFF5FF5B7E6DFAEBFB39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Honca grandis (Busck 1907) Brown 2019	<div><p>Honca grandis (Busck, 1907), new combination</p><p>Figs. 7, 23, 37</p><p>Phalonia grandis Busck, 1907: 23; McDunnough 1939: 60.</p><p>[Incertae sedis] grandis: Powell 1983: 42 .</p><p>[ Cochylini New Genus 4] grandis: Brown 2005: 208; Metzler and Brown 2014: 278.</p><p>Honca grandis can be distinguished from nearly all other North American Cochylini by its large size (forewing length 11.0– 12.5 mm, mean = 11.8 mm, n = 5, in males; 11.5–13.0 mm, mean = 12.2 mm, n = 6, in females). The forewing pattern (Fig. 8), which is similar to that of Henricus umbrabasana, features a cream ground color with a large pale reddish-brown triangular blotch occupying the middle of the hind margin, a narrow concolorous line along the basal 0.33 of the costa, and a concolorous dot near mid-costa. The hindwing is pale cream with pale graybrown mottling. The forewing pattern and the absence of a male forewing costal fold distinguish H. grandis from species of Phtheochroa, which also are among the largest North American Cochylina.</p><p>Honca grandis is rare in collections. The early stages are unknown. Captures extend from June through July, with a single record from August. The species is apparently restricted to montane habitats in the western United States, with specimens recorded from Utah, Colorado, and Wyoming (CSUC, USNM, EME, AMNH, MSC) at elevations between 6,400 and 7,400 feet.</p><p>Holotype ♀, USA, Colorado, Jefferson Co., Golden, Chimney Gulch, 1 Jul 1904, Oslar (USNM).</p></div>	https://treatment.plazi.org/id/03E08794FFC8FFF5FF5B7E6DFAEBFB39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFC8FFF6FF5B7951FD9CFABD.text	03E08794FFC8FFF6FF5B7951FD9CFABD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nycthia Brown 2019	<div><p>Nycthia Pogue, new genus</p><p>Type species. Phalonia pimana Busck, 1907, by present designation.</p><p>Nycthia; Pogue and Mickevich, 1990: 322 (nomen nudum).</p><p>Cochylini New Genus 5 [ pimana group]; Brown, 2005: 208; Metzler and Brown 2014: 278.</p><p>Nycthia is described to accommodate two previously recognized species, N. pimana and N. yuccatana . The two were described in Phalonia and treated as such in early checklists and catalogs (e.g., McDunnough 1939). Powell (1983) transferred yuccatana to “incertae sedis” and pimana to Cochylis; Pogue (1986) proposed the name Nycthia for the two species in his unpublished dissertation. Razowski (1994: 294) placed them both in Cochylis “…until a description of the new genus is published,” referring to Pogue’s work. Brown (2005) and Metzler and Brown (2014) treated them as “ Cochylini New Genus.”</p><p>Diagnosis. Adults of Nycthia are not particularly similar to any other North American Cochylina (Figs. 9, 10), although the complete, narrow, median fascia of N. pimana is somewhat reminiscent of the forewing pattern of the eastern Eugnosta sartana (Hübner, 1823) or Cochylis caulocatax Razowski, 1984 . As in most Nearctic Cochylina, males of Nycthia lack a forewing costal fold and have a well-developed hindwing costal roll similar to species of Cochylis and related genera. The most conspicuous autapomorphy for Nycthia is the highly modified valva, with a short, free, lobelike costa; an extremely long, slender, curved, rodlike median portion; and a thumblike sacculus. The uncus is represented by a short, membranous, broadly triangular process; and the median process of the transtilla is long and slender with a curved tip, similar to that of Cochylis and related genera (e.g., Cochylichroa, Thyraylia).</p><p>Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; antennal sensory setae 1.0–1.2 diameter of flagellomere in male, shorter and sparser in female; labial palpus porrect, combined length of all segments 2.5–3.0 times horizontal diameter of compound eye, segment III nearly concealed by scaling of segment II; maxillary palpus 3-segmented. Thorax: Posterior crest present. Forewing with base of costa straight, gently curved in distal 0.33 to rounded apex; termen oblique; Sc ca. 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating at 0.65 length of discal cell; CuP absent; A1+2 stalked about 0.5 length; chorda and M-stem a trace in discal cell; costal fold absent. Hindwing length ca. 2.5 times width; costal roll present in male; Sc+R ca. 0.35 hindwing length, ending before wing margin; Rs and M 1 fused ca. 0.5 length, Rs to costa before apex, M 1 to termen; M 2, M 3, CuA 1 widely separated at base; CuP faint; frenulum with one acanthus in male, two (rarely one) in female. Abdomen: Unmodified. Male genitalia (Figs. 24, 25) with uncus represented by a small triangular lobe; socius small, rounded or broadly triangular, fused basally, sparsely setose; tegumen triangular; transtilla well developed, with elongate median process, nearly uniform in width throughout, pointed apically; valva with short costal process projecting at 90‒100 angle from vertical (tegumen), median process of valva elongate, strongly curved, extending beyond sac- culus, pointed apically, sacculus free from valva except at base, digitate, well sclerotized; vinculum narrow, arms connected by membrane distally. Phallus large, 1.5–2.0 times length of median process of valva, bent near middle, with pointed ventral termination; phallobase with a pair of small, lateral, rounded, membranous wings; vesica with patch of fine spiculae, lacking cornuti. Female genitalia (Figs. 38, 39) with ovipositor elongate, telescopic; papillae anales lightly sclerotized, slender, joined and narrowed posteriorly, sparsely setose; apophyses anteriores ca. 1.2 times length of apophyses posteriores; sterigma lightly sclerotized, lamella antevaginalis U-shaped, pointed posteriorly, not joined to apophyses anteriores; lamella postvaginalis membranous, with two lightly sclerotized, setose bands coalesced with top of colliculum; ductus bursae with elongate, strongly sclerotized colliculum, proximal ends slightly bent laterally, rounded; origin of accessory bursae just beyond anterior margin of sclerotized region; corpus bursae with short vertical convolutions from junction with ductus bursae; signum absent.</p><p>Distribution and biology. Nycthia ranges from Texas west to southern California, and south to Chiapas, Mexico; Razowski (1994) illustrated a female from the Dominican Republic. The few specimens examined from Mexico may represent one or more undescribed species that do not range as far north as U.S. border. All reported life history information indicates that larvae are specialists on the seeds of Agave and Yucca (Liliaceae) . The female genitalia have a long slender ovipositor adapted for depositing eggs in the flowers and fruit. A presumed yucca-feeder ( N. yuccatana) and a presumed agave-feeder ( N. pimana) have been recorded as sympatric in at least two localities in southern California: Pinyon Crest in Riverside County, and Blair Valley in San Diego County.</p><p>Etymology. The generic name is an anagram of Cynthia, and is named in honor of Mike Pogue’s late cousin, Cynthia L. Haines. It is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFC8FFF6FF5B7951FD9CFABD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCBFFF7FF5B79CDFC84FE85.text	03E08794FFCBFFF7FF5B79CDFC84FE85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nycthia pimana (: Powell 2019) Brown 2019	<div><p>Nycthia pimana (Busck, 1907), new combination</p><p>Figs. 9, 24, 38</p><p>Phalonia pimana Busck, 1907: 24; McDunnough 1939: 60.</p><p>Cochylis pimana: Powell 1983: 142; Razowski 1994: 294.</p><p>[ Cochylini New Genus 5] pimana: Brown, 2005: 208; Metzler &amp; Brown 2014: 278.</p><p>Diagnosis. Adults of N. pimana are easily distinguished from those of N. yuccatana by two features of the forewing: N. pimana has a complete, uninterrupted forewing median fascia, whereas the median fascia is faint and/or interrupted in the latter; and N. pimana has a paler ocherous forewing ground color, whereas that of N. yuccatana is pale grayish. In the male genitalia of N. pimana (Fig. 24) the long, rodlike, median part of the valva is rather sinuate in the distal 0.2, whereas in N. yuccatana the rod is more evenly curved. In the female genitalia, the colliculum of N. pimana is more cylindrical than that of N. yuccatana and has a distinctive thumblike process anteriorly.</p><p>Type. Holotype ♀, USA, Arizona, Pima Co., Baboquivera Mountains, August (USNM).</p><p>Nycthia pimana has been reared from the seeds of Agave species ( Liliaceae). Adult captures range from April through September, probably representing two broods. The species apparently is restricted to the xeric mountain ranges of the southwestern U.S. from Texas (Davis, Chisos, and Guadalupe mountains) to Lordsburg, New Mexico, Arizona (Baboquivari, Santa Rita, Chiricahua, and Huachuca mountains), and southern California (San Jacinto Mountains), south into Sinaloa and Baja California, Mexico, typically at elevations from about 4,000’ (in California) to 6,800’ (in New Mexico). In recent years it has been collected with some frequency in the lower elevations of the Chiricahua Mountains (near Portal) in southeastern Arizona.</p></div>	https://treatment.plazi.org/id/03E08794FFCBFFF7FF5B79CDFC84FE85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCAFFF7FF5B7D8AFC09FC09.text	03E08794FFCAFFF7FF5B7D8AFC09FC09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nycthia yuccatana (Busck 1907) Brown 2019	<div><p>Nycthia yuccatana (Busck, 1907), new combination</p><p>Figs. 10, 25, 39</p><p>Phalonia yuccatana Busck, 1907: 23; McDunnough 1939: 60.</p><p>[Incertae sedis] yuccatana: Powell 1983: 42 .</p><p>Cochylis yuccatana: Razowski 1994: 294; Powell et al. 1995: 142.</p><p>[ Cochylini New Genus 5] pimana: Brown 2005: 208; Metzler &amp; Brown 2014: 278.</p><p>Diagnosis. When compared to N. pimana (Fig. 9), N. yuccatana (Fig. 10) has a more somber chalk white to pale grayish ground color and the pattern elements reduced to a few small spots and/or an interrupted median fascia. In the male genitalia of N. yuccatana (Fig. 25), the distal tips of the long, curved, rodlike costa of the valva are less recurved than those of N. pimana, and the median process of the transtilla is slightly more dilated mesially.</p><p>Type. Holotype ♀, USA, Texas, Nueces Co., Nuecestown, r.f. Yucca baccata, 12 Apr 1897, C. Marlatt (USNM).</p><p>Nycthia yuccatana has been reared from flowers, seeds, and fruiting stalks of Yucca baccata (Liliaceae) . Adults have been collected from Texas to southern California. Specimens similar to N. yuccatana occur as far south as Chiapas, Mexico, but these may represent one or more undescribed species.</p></div>	https://treatment.plazi.org/id/03E08794FFCAFFF7FF5B7D8AFC09FC09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCAFFF0FF5B7821FEE7FBB9.text	03E08794FFCAFFF0FF5B7821FEE7FBB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pogospinia Brown 2019	<div><p>Pogospinia Brown, new genus</p><p>Type species: Pogospinia floridana Brown, new species, by present designation.</p><p>Pogospinia is proposed for three species formerly placed in Spinipogon Razowski, 1967, i.e., P. spinifera (Razowski, 1967), new combination; P. veracruzanus (Razowski and Becker 1986), new combination; and P. signata (Razowski, 1967), new combination; and one new species described herein. The last is from southern Florida and appears to be the only U.S. representative of Pogospinia. Razowski (1994) referred to “two groups of species…” in Spinipogon, which I recognize as distinct genera. Although P. spinifera, P. veracruzanus, and the new species are convincingly associated by male genitalia, P. signata is known only from the female, and its genitalia are not like those of any other Spinipogon . Females of most (but not all) species of Spinipogon and Pogospinia have an unusual and characteristic spiny, truncate-cone-shaped process emanating from the corpus bursae (e.g., Razowski and Becker 1983: figs. 111, 112), similar to that of Phalonidia and Gynnidomorpha Turner, 1916 .</p><p>The male genitalia of the species included in Pogospinia are very similar, however, they can be distinguished most easily by the shape of the median process of the transtilla. Razowski and Becker (1986) provided three illustrations of the male genitalia of S. veracruzanus, all from Estación Biologia Las Tuxtlas in Veracruz, Mexico, and recognized two different “forms” which mostly likely represent different species.</p><p>Diagnosis. Superficially, adults of Pogospinia (Fig. 8) are similar to those of Spinipogon, Lorita Busck, 1939, Cochylis, Thyraylia, and others, however, they usually are slightly smaller. The male genitalia (Fig. 26) of Pogospinia are easily distinguished by the shape of the valva: the distal 0.3–0.5 is widely bifurcate with a broad, round- ed or squarish concavity between the termination of the costa and the termination of the ventral edge (sacculus) (Fig. 26; also see Razowski &amp; Becker 1986: figs 103‒108). This shape is unique among Cochylina and distinguishes the three described species of Pogospinia for which males are known. Pogospinia appears to be most closely related to Spinipogon; the two genera share similar socii, a similar median process of the transtilla, and fused arms of the vinculum, sometimes forming a weak saccus. In Spinipogon the valva are somewhat rectangular or parallel-sided and not bifurcate distally (see Razowski and Becker 1986: figs. 109, 110; Razowski 1993: figs 33, 34; Metzler and Sabourin 2002: figs. 2a, b). The absence of cornuti in the vesica of Pogospinia is shared with a few other genera, including Spinipogon . Males of Pogospinia lack both a forewing costal fold a hindwing costal roll.</p><p>Description. Head: Frons and vertex rough scaled; ocellus present; sensory setae of antenna 0.8–1.0 times flagellomere diameter in male, shorter in female; labial palpus porrect, length of all segments combined 1.0–1.2 times diameter of compound eye, segment III weakly exposed; maxillary palpus 1-segmented. Thorax: Posterior crest weak; lateral scales of metanotum hairlike. Forewing length 3.3–4.5 mm in male, 4.4–4.5 mm in female, length 2.5–2.9 times maximum width, slightly wider in female; costa gently arched from base, apex round, male without costal fold; termen straight, oblique; Sc ca. 0.5 wing length; R 1 originating near middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating about 0.6 length of discal cell; CuP absent; A1+2 stalked at 0.4 length. Hindwing length 2.5–3.6 times width; costa straight, male without costal roll; apex rounded; termen concave between M 1 and M 2; Sc+R 1 about 0.6 wing length; Rs and M 1 stalked about 0.4 length of M 1; M 3 and CuA 1 separate; CuA 2 originating 0.65 length of discal cell; frenulum with one acanthus in male, two in female. Abdomen: Unmodified. Male genitalia (Fig. 26) with uncus absent; socius rounded, broadly attached to top of tegumen, setose; transtilla well developed, median process variable from short and blunt to bifurcate, spined or minutely dentate; valva wide at base, bifurcate in distal 0.5‒0.7 with dorsal (costal) lobe and ventral (sacculus) lobe, each with well-developed field of strong spines; vinculum arms fused distally forming V-shaped saccus. Phallus large, slender, 0.8–1.3 times length of valva, pointed apically; vesica without cornuti. Female genitalia (Fig. 40) with papillae anales weakly lobate, setose; length of apophyses anteriores ca. 0.8 times that of apophyses posteriores, apophyses anteriores connected to sclerotized sterigma; sterigma variable from complicated with dense patches of spiculation, and rounded V-shaped median process to simple with median sclerite; ductus bursae short, with colliculumlike sclerotized band, weakly differentiated from corpus bursae; corpus bursae variable, round to elongate, with minute spicules and a truncate-cone-shaped process of long spines; accessory bursa originating from anterior third of corpus bursae.</p><p>Distribution and biology. The genus ranges from Brazil ( P. spinifera) north through Mexico ( P. veracruzanus) to Florida; it likely occurs in the Caribbean. Little is known of the life history of Pogospinia, however, P. floridana was reared (n = 3 individuals) by Don Davis from Anthernanthera maritima (Amaranthaceae) at Long Key State Park in southern Florida.</p><p>Etymology. The genus name is close to, but not quite, an anagram of the related Spinipogon, and is feminine in gender.</p></div>	https://treatment.plazi.org/id/03E08794FFCAFFF0FF5B7821FEE7FBB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCDFFF1FF5B78C6FEEFFCD1.text	03E08794FFCDFFF1FF5B78C6FEEFFCD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pogospinia floridana Brown 2019	<div><p>Pogospinia floridana Brown, new species</p><p>Figs. 8, 26, 40</p><p>Diagnosis. Pogospinia floridana (Fig. 8) is a small (forewing length 4.0‒5.0 mm) cream-colored moth with brown and dark brown markings. It is distinguished from its congeners by the shape of the median process of the transtilla (Fig. 26), which is about as long as the free terminal lobes of the valva, and is emarginate distally with a tiny, short thorn on each terminal lobe. In comparison, the median process of the transtilla of P. spinifera is short and blunt. The width of the dorsal (costal) lobe of the valva of P. floridana is ca. equal to the width of ventral lobe. In comparison, the dorsal (costal) lobe of the valva of P. veracruzanus is ca. 1.5 times as wide as the width of ventral lobe.</p><p>Description. Head: Frons and vertex cream; labial palpus cream, mixed with tawny orange on outer surface, scaling only slightly expanded distally on segment II, second III slightly exposed; antenna with cream scales. Thorax: Nota mostly cream, slightly tawny; forewing length 4.0– 4.5 mm (mean = 4.3 mm, n = 5) in male, 4.5 mm (n = 2) in female; forewing ground color cream with scattered, ill-defined blotches of pale tawny; a blackish brown, triangular patch from lower edge at ca. 0.4 distance from base to apex, a small, ill-defined, brown costal blotch at ca. 0.2 distance from base to apex, a similar but larger blotch near mid-costa, and a similar colored fascia overscaled with pale tanwy at ca. 0.8 distance from base to apex, extending toward, but ending before tornus. Fringe cream. Hindwing white with pale beige overscaling. Fringe concolorous with hindwing. Abdomen: Mostly cream above and below. Male genitalia (Fig. 26) essentially as described for genus; median process of transtilla about as long as lobes of valva, somewhat chordate-emarginate distally, with a tiny spine on each terminal lobe; valva widest at base, birfurcate in distal 0.33 creating a broad, U-shaped concavity along outer margin of valva; both arms of bifurcation nearly parallel-sided, dorsal arm with field of strong, conspicuously socketed setae restricted to arm, ventral arm with field of strong spines extending length of sacculus; vinculum arms forming narrowly rounded, V-shaped saccus. Phallus large, slender, 6–7 times as long as wide, attenuate-pointed apically; vesica without cornuti. Female genitalia (Fig. 40) with papillae anales weakly lobate, setose; sterigma complicated with dense patches of spiculation, and V-shaped median region rounded at anterior margin; ductus bursae short, undifferentiated from corpus bursae; corpus bursae round, with patch of minute spicules on left side and truncate-cone-shaped process of long spines on right side.</p><p>Types. Holotype ³, USA, Florida, Monroe Co., Key Largo, 6 Mar 1977, S. Kemp (USNM) . Paratypes (23³, 9♀). USA: Florida: Collier Co.: Chokoloskee, no date, no collector (6³) (USNM) . Monroe Co.: Key Largo, 8 Oct 1964 (1♀), 26 Oct 1964 (3³), 20 Nov 1964 (1³), 25 Feb 1977 (1³), 8 Oct 1964 (1³), 19 Feb 1968 (1³), 25 Jun 1971 (1³), 15 Dec 1969 (1³), 27 Dec 1971 (1♀), 12 Nov 1967 (1³), 15 Jul 1967 (1♀), 28 Oct 1971 (1³), all S. Kemp (USNM) . Long Key State Park, 14 Jun 1998, reared from Alternanthera maritima, emerged 29 Apr‒4 May 1998 (3³), D. R. Davis (USNM) . Tavernier, 17 Sep 1955 (1♀), 21 Oct 1955 (1♀), J. Todd (USNM) . 2 mi NE Tav- ernier, 20 Jan 1973 (3♀), J. Heppner &amp; J. Powell (USNM) . Highlands Co.: Archbold Biological Station, Highlands Co., 31 Jul 1978 (1♀), Weems &amp; Frolich (USNM) . Sarasota Co.: Siesta Key, 28 Feb 1953 (1³), 8 Dec 1953 (1³), 11 Dec 1953 (1³), C. Kimball (USNM) .</p><p>Pogospinia floridana is known from the panhandle of northern Florida to the Keys at the southern tip of Florida (Escambia, Highlands, Collier, and Monroe counties). Collection records extend throughout the year, with the majority from October to March. Larvae collected on 14 April from Anthernanthera maritima (Amaranthaceae) at Long Key State Park in southern Florida produced adults that emerged on 29 April (n = 2) and 4 May (n = 1) of the same year.</p></div>	https://treatment.plazi.org/id/03E08794FFCDFFF1FF5B78C6FEEFFCD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCFFFF3FF5B78F5FB70FC0D.text	03E08794FFCFFFF3FF5B78F5FB70FC0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monoceratuncus Razowski 1992	<div><p>Monoceratuncus Razowski, 1992</p><p>Ceratuncus Razowski, 1986a: 382 . [preoccupied]</p><p>Monoceratuncus Razowski, 1992: 201 [replacement name]; Brown 2005: 265.</p><p>Type species: Ceratunus lugens Razowski, 1986, by original designation.</p><p>Although Pogue (1986) proposed the name Decuma in his unpublished Ph.D. dissertation, he subsequently recognized that it was conceptually the same as Razowski’s Ceratuncus, and the latter name was used by Pogue &amp; Mickevitch (1990) in a phylogenetic analysis of North American genera of Cochylidae . In 1992, Razowski proposed the replacement name Monoceratuncus because Ceratuncus was recognized as a junior homonym.</p><p>According to Razowski (1986a), autapomorphies for the genus include the distally expanded uncus with a hook-shaped distal process; the broad anterior portion of the tegumen; and the long basal process of the valva, extending ventro-proximally. Razowski (1986a) further commented that the presence of numerous cornuti in the vesica is a synapomorphy with Cochylis .</p><p>As currently defined, Monoceratuncus includes nine species: seven occur in Mexico, one in Peru (Razowski 1993), and one is described herein from Texas. Illustrations of the genitalia can be found in Razowski (1986a, 1993) and Razowski and Becker (1986).</p><p>Diagnosis. Adults of Monoceratuncus are among the smaller Cochylina in the New World. They can be distinguished from all other genera by the unusual, well-developed uncus with a conspicuous median swelling or lobe and a slender, hook-shaped apex. The valvae are strongly attenuate distally, usually with a longitudinal row of strong spinelike setae in the distal 0.5. The genus is further characterized by the fused distal arms of the vinculum, the absence of a median process of the transtilla, and the absence of socii. The latter three characters occur sporadically in a few other genera, but never in the same combination as in Monoceratuncus .</p><p>Redescription. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna ca. 1.0 times flagellomere diameter in male, shorter, sparser in female; labial palpus porrect, combined length of all segments 1.0‒1.5 times diameter of compound eye, segment III exposed; maxillary palpus 2-segmented. Thorax: Posterior crest small; lateral scale tufts of metanotum flat. Forewing (Fig. 12) length 3.4–3.7 mm, females slightly larger than males, forewing length 2.6–3.1 times width, slightly wider in female; costa gently arched from base to apex, without costal fold in male, apex rounded; termen straight, oblique; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 of length. Hindwing moderately broad, length 2.4–2.7 times width; costa straight, with costal roll extending ca. 0.5 length of costa in male; apex produced; termen concave below apex; Sc+R 1 ca. 0.33 wing length in male, 0.65 wing length in female; Rs and M 1 stalked ca. 0.65 length of M 1; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female. Abdomen: Unmodified. Male genitalia (Fig. 29) with anterior portion of tegumen broad; vinculum complete with small saccuslike process; uncus well developed, with expanded median lobe, produced into a curved spine apically; socius absent; transtilla well developed, broad laterally, with short median process; valva broad at base, narrowed distally, costa lightly sclerotized, apex produced, sacculus ill defined, completely fused to venter of valva. Phallus slightly shorter than valva, bent near middle, pointed apically; vesica with patch of tiny cornuti. Female genitalia (Fig. 42) with papillae anales narrow, weakly sclerotized, setose, joined posteriorly; apophyses anteriores slightly shorter than apophyses posteriores, not fused to sterigma; sterigma comprised of a few narrow trapezoidal plates parallel to each other; ductus bursae undifferentiated from corpus bursae; corpus bursae small with linear convolutions along entire length, with scattered minute spicules; accessory bursa originating from posterior end of corpus bursae.</p></div>	https://treatment.plazi.org/id/03E08794FFCFFFF3FF5B78F5FB70FC0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFCEFFECFF5B7815FED0FF3D.text	03E08794FFCEFFECFF5B7815FED0FF3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monoceratuncus lantana Brown 2019	<div><p>Monoceratuncus lantana Pogue, new species</p><p>Figs. 12, 29, 42</p><p>Diagnosis. Monoceratuncus lantana is superficially similar to most congeners. It can be distinguished by the male genitalia, which are most similar to those of M. tantulus Razowski and Becker (1986: fig. 118) and M. autolytus Razowski (1986: figs. 52‒55). In M. lantana (Fig. 29) the median lobe of the uncus is large, as in M. tantulus (larger than that of C. autolytus), and the outer margin of the valva is rather evenly curved (broad basally and abruptly narrowed in the middle in M. tantulus and angled near the middle in A. autolytus).</p><p>Description. Head: Frons and vertex buff, with cream-tipped scales giving a speckled appearance; labial palpus light buff with scattered fuscous scales, cream on inner surface; antenna mixed fuscous and cream, sensory setae about 1.0 times flagellomere diameter in male, shorter in female. Thorax: Mesonotum and tegula buff in anterior 0.3, remainder fuscous; forewing length 2.8–3.6 mm (mean = 3.3 mm, n = 4); forewing ground color buff; submedian cinnamon spot extending from immediately above hind margin to costal edge of discal cell; median costal spot cinnamon, suffused with brown; distal 0.5 of wing darker buff and brown; apical band indistinct, tawny, edged and suffused with brown. Fringe buff. Hindwing pale cream brown. Fringe concolorous with wing. Abdomen: Dorsal scaling brown, venter shining white. Male genitalia (Fig. 29) with uncus large, bearing conspicuous median lobe, apical 0.4 of uncus slender, curved; socius absent; transtilla broad, lacking median process; valva broad at base, attenuate apically, sacculus confined to basal 0.25 of valva; a patch of slender spines along ventral margin of valva from ca. 0.50‒0.75 distance from base to apex of valva. Phallus relatively slender, curved near middle, pointed apically; vesica with patch of tiny cornuti. Female genitalia (Fig. 42) with papillae slightly sclerotized along inner margin, joined at dorso-posterior end; length of apophyses anteriores ca. 0.65 times that of apophyses posteriores; sterigma with broad, shallow, U-shaped plate anterior to ostium, with smaller sclerite posterior to ostium and two narrow sclerites anterior to U-shape plate; ductus bursae undifferentiated from corpus bursae; corpus bursae small with scattered minute spicules; accessory bursa originating from posterior end of corpus bursae.</p><p>Types. Holotype ³, USA, Texas, Uvalde Co., Uvalde, [no date], reared from Lantana, L. F. Hitchcock (USNM) . Paratypes (5³, 8♀). USA: Texas: Uvalde Co.: Uvalde, Jul 1940 (1³, 2♀) (USNM) . Washington Co .: Washington, Jul 1939 (1³, 2♀), reared from Lantana, L. F. Hitchcock (USNM) . Brooks Co., Falfurrias, Mar 1939 (1³, 1♀), Jun 1939 (2³, 3♀), S. Kelley (USNM) .</p><p>Distribution and biology. The species is known only from the type locality in southeastern Texas, and all specimens were reared from Lantana sp. ( Verbenaceae) in 1939–1940, which represents the first reported host plant for the genus.</p></div>	https://treatment.plazi.org/id/03E08794FFCEFFECFF5B7815FED0FF3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD3FFEFFF5B7A5AFC67FE69.text	03E08794FFD3FFEFFF5B7A5AFC67FE69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimcochylis Razowski 1985	<div><p>Mimcochylis Razowski, 1985</p><p>Type species: Mimcochylis planola Razowski, 1985 (= Conchylis fulvotinctana Walsingham, 1884, new combination, new synonymy).</p><p>Mimcochylis Razowski, 1985 b: 61; Powell et al. 1995: 142; Brown 2005: 424.</p><p>Mimcochylis was described by Razowski (1985) along with four new species from northwestern Mexico (i.e., Baja California, Sinaloa, and Durango), with each species represented only by the holotype – one by a male and three by females. Razowski (1985) listed the following autapomorphies for the genus: the top of the tegumen with three small prominences; a median prominence on the dorsal surface of the tegumen subapically; and accessory bursa arising ventrally from the base of the ductus bursae. He also noted the following features: tegumen broad, mem- branous laterally except for base and apical one-third; vinculum arms separate distally; transtilla complete with a simple median process; juxta short, expanded laterally; valva short, broad, with differentiated costal portion; caulis anterior; ovipositor elongate, semi-telescopic; apophyses long; ductus seminalis from membranous corpus bursae, subdorsally; and anterior margin of sterigma concave or deeply incised mesially. The single known male, that of M. planola, has a distinctive, long, slender, free, upturned costal of the valva, and a long, slender, distally attenuate phallus lacking cornuti; and these features distinguish it from all other Cochylina.</p><p>Distribution and biology. Mimcochylis is known from southern Arizona (Santa Rita and Huachuca mountains), U.S.A., and Baja California, Sinaloa, and Durango, Mexico.</p></div>	https://treatment.plazi.org/id/03E08794FFD3FFEFFF5B7A5AFC67FE69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD2FFEFFF5B7941FAD1F8B7.text	03E08794FFD2FFEFFF5B7941FAD1F8B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aethes Billberg 1820	<div><p>Aethes Billberg, 1820</p><p>Type species: Pyralis smeathmanniana Fabricius, 1781 .</p><p>Aethes (with its many synonyms) is a large genus of about 150 described species (Gilligan et al. 2019), with many additional undescribed species, that is widely distributed in the Palearctic, Nearctic, and Neotropical Regions. Unlike most genera of Cochylina, nearly all North American members of Aethes can be identified to genus by facies alone. The forewing typically has a white to cream ground color with one or more narrow, brown or golden fascia (often interrupted), a small triangular patch near the middle of the hind margin, and/or a brown dash from the base of the forewing; however, there are many exceptions. Razowski (1986b) listed the Neotropical species.</p><p>Both the male and female genitalia are fairly diverse morphologically (e.g., Sabourin et al. 2002), as would be expected for a large, widely distributed genus, but males can almost always be assigned to Aethes on the basis of a pair of slender, hooklike processes from the socii. However, these characteristic processes appear to be lost secondarily in a number of species that based on other characters, including barcodes (see Brown et al. 2019), fit convincingly in Aethes . Below I transfer one species to Aethes and describe a second, the latter the undescribed type species of Pogue’s (1986) manuscript genus Morta . Both species lack the sickle-shaped processes from the socii.</p></div>	https://treatment.plazi.org/id/03E08794FFD2FFEFFF5B7941FAD1F8B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD2FFEFFF5B7E01FA84FB29.text	03E08794FFD2FFEFFF5B7E01FA84FB29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimcochylis fulvotinctana (Walsingham 1884) Brown 2019	<div><p>Mimcochylis fulvotinctana (Walsingham, 1884), new combination</p><p>Figs. 13, 28, 43</p><p>Conchylis fulvotinctana Walsingham, 1884: 132 .</p><p>Phalonia fulvotinctana: McDunnough 1939: 60 .</p><p>“Incertae sedis” [unplaced Cochylini] fulvotinctana: Razowski 1964: 384; Powell 1983: 42; Razowski 1994: 359; Brown 2005: 209; Metzler &amp; Brown 2014: 278.</p><p>Cochylis fulvotinctana: Powell et al. 1995: 142 .</p><p>Mimcochylis planola Razowski, 1985: 61, new synonym .</p><p>The generic assignment of Conchylis fulvotinctana has been elusive since its description. Walsingham (1884) assigned it to the catch-all genus Conchylis (an unjustified emendation of Cochylis); McDunnough (1939) placed it in Phalonia; Razowski (1964) treated it as “incertae sedis), and the latter assignment (unplaced) was followed by Powell (1983), Brown (2005), and Metzler and Brown (2014). Because the type (NHMUK) lacks its abdomen, the generic assignment has remained enigmatic. However, I recently discovered two series of specimens, one from Arizona (USNM) and one from Baja California (SDNHM), that are superficially indistinguishable from the type of fulvotinctana (Fig. 13). The male genitalia (Fig. 28) of the specimens match precisely those illustrated by Razowski (1985: fig. 1) for Mimcochylis planola Razowski, 1985 . Based on these observations, I propose the synonymy of fulvotinctana and planola, and transfer fulvotinctana to Mimcochylis .</p><p>According to the original description, Walsingham (1884) had two specimens from Arizona. Razowski (1994: 384) referred to the only one he examined as “ lectotype ” and indicated “without abdomen,” and I interpret this as the designation of that specimen as the lectotype; the specimen bears a holotype label (NHMUK 010921407).</p></div>	https://treatment.plazi.org/id/03E08794FFD2FFEFFF5B7E01FA84FB29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD7FFEAFF5B7C99FEE3FBC8.text	03E08794FFD7FFEAFF5B7C99FEE3FBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aethes ringsi (Metzler 2000) Brown 2019	<div><p>Aethes ringsi (Metzler, 2000), new combination</p><p>Figs. 14, 30, 44</p><p>Cochylis ringsi Metzler, 2000: 191; Brown 2005: 213; Metzler &amp; Brown 2014: 276.</p><p>Described in Cochylis by Metzler (2000), ringsi is transferred to Aethes based on facies, DNA barcodes (see Brown et al. 2019), and the similarity in the male genitalia to those of Aethes tuxtlana Razowski, 1986 . The forewing pattern of A. ringsi (Fig. 14) is somewhat reminiscent of that of a few other species of Aethes (e.g., A. fernaldana (Walsingham, 1879), A. argentilimitana (Robinson, 1869)), with a slightly shiny, buff to whitish ground color and a contrasting and characterisitic orange-brown basal patch, and a well-defined, median fascia, angled near the middle. DNA barcodes convincingly place it deeply embedded within Aethes in a maximum likelihood analyses (Brown et al. 2019).</p><p>The assignment of ringsi to Aethes is contradicted by the absence of the sickle-shaped processes from the socii that define Aethes and the absence of a median process of the transtilla, common to most Cochylini, including Aethes . However, at least two other species of Aethes (i.e., an undescribed species from the Midwestern U.S. and Aethes tuxtlana from Mexico) lack the sickle-shaped processes, and the latter species lacks a median process from the transtilla, as well (see Razowski 1986b: fig. 5). The unusual somewhat ”split” valva, with a invagintation between the sacculus and the remainder of the valva of A. ringsi (Fig. 32) is extremely similar to that of A. tuxtlana, which also has unusual socii. Hence, ringsi appears to be a derived species within Aethes, possibly the sister to A. tuxtlana . The female genitalia (Fig. 44) are fairly typical of the genus.</p><p>Aethes ringsi ranges from the southeastern U.S. (Alabama, South Carolina, North Carolina) to the mid-West (Iowa, Indiana, Missouri, Arkansas, and Oklahoma). Metzler (2000) suggests it is a species of oak barrens on sandy soils. Captures range primarily from June through August, but there are a few April and May records from the southernmost portion of its range.</p><p>Holotype ³, USA, Indiana, Newton Co., 41˚6.43’N, 87˚26.25’W, Conrad Savanna, 8 Jul 1998, E. Metzler, UV (USNM).</p></div>	https://treatment.plazi.org/id/03E08794FFD7FFEAFF5B7C99FEE3FBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD7FFEBFF5B7855FB59FABD.text	03E08794FFD7FFEBFF5B7855FB59FABD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aethes triassumenta Brown 2019	<div><p>Aethes triassumenta Pogue, new species</p><p>Figs. 15, 16, 31, 45</p><p>Diagnosis. Adults of A. triassumenta (Figs. 15, 16) are superficially extremely similar to those of A. bomonana (Kearfott, 1907) and A. obliquana (Kearfott, 1907), with a slightly shiny, pale cream forewing ground color and illdefined, contrasting brown to red-brown markings, often with numerous tiny brown specks in the apical and subapical regions. The male genitalia of A. triassumenta (Fig. 31) are distinguished from other congeners by the absence of the sickle-shaped processes of the socii and the attachment of the juxta to the valva in the male genitalia; and the extremely elongate, frail, membranous corpus bursae in the female genitalia (Fig. 45). The shape of the valva is not particular divergent from that of other congeners, with a weakly concave outer margin and small lobe or spine near the apex of the costa. The suberect, elongate-triangular socii of A. triassumenta are nearly identical to those of A. tuxtlana, and likely are homologous with the sickle-shaped socii of other Aethes . The species is assigned to Aethes on the basis of the facies and DNA barcode data (Brown et al. 2019).</p><p>Description. Head: Frons and vertex white to cream; labial palpus reddish brown on outer surface, white to cream on dorsal and inner surface. Thorax: Mesonotum and tegula white with variable cream scaling; posterior crest white; forewing length 8.9–9.1 mm (mean = 9.0 mm, n = 10); forewing ground color white; base of costa and small irregular patch above R vein reddish brown; an irregular horizontal patch extending from wing base to middle of discal cell, consisting of a posterior chestnut patch proximally contiguous with a larger anterior chestnut patch, with interior scaling shading to light cinnamon; two median patches at end of discal cell: anterior one chestnut, usually larger than posterior patch, posterior patch oblong with anterior 0.5 chestnut and posterior 0.5 buff yellow with a few tawny scales proximally; two buff-yellow patches: one from base of R vein to just before horizontal bar below Sc, second one less distinct, from base of R 1 to end of discal cell; both patches below costa; costa and hind margin with short brown striations, variable in number and intensity; subapical band from costa to hind margin white. Fringe white. Hindwing pale gray. Fringe white with basal band of ground color. Abdomen: Unmodified. White to cream with scattered pale-gray scales; genital tuft white. Male genitalia (Fig. 31) with uncus absent; socius with basal 0.25 fused to tegumen, distal part free, narrow-triangular, with pointed apex, setose; transtilla broad at base, with pointed spines laterally, median projection elongate, attenuate distally, rounded apically; valva broad, length about 1.4 times maximum width, costa weakly concave mesially, apically produced into thumblike process with variably developed thorn, sacculus undifferentiated; vinculum arms free distally; juxta shieldlike with prominent lateral arms confluent with upper edge of sacculus. Phallus large, 1.2 times length of valva, ventral projection elongate, curved, pointed; vesica with a single large cornutus. Female genitalia (Fig. 45) with papillae anales slender, lightly sclerotized; apophyses anteriores about 1.2 times length of apophyses posteriores; sterigma mostly membranous, lightly sclerotized around ositum bursae; lamella postvaginalis comprised of two semicircular plates joined mesially; colliculum sclerotized with a ventrally produced digitate process opposite of origin of accessory bursa, and second digitate process near anterior end of colliculum immediately anterad of cup-shaped concavity; ductus bursae elongate, membranous anterad of colliculum, gradually broadened, lacking distinct junction with corpus bursae; corpus bursae extremely elongate, slender, membranous, without spiculae or signum.</p><p>Types. Holotype ³, USA, Florida, Highlands Co., Archbold Biological Station, 25 Dec 1957, R. W. Pease, Jr. (USNM) . Paratypes (47³, 3♀). USA: FLORIDA: Same locality as holotype, 23 Dec 1957, 26 Jan 1958, J. R. Mal- loch (4³) (USNM) . Pinellas Co.: St. Petersburg, 31 Jan 1958, 11 Feb 1958, R. W. Pease (2³) (USNM) . St Johns Co.: Hastings, [no date], [no collector] (2³) (USNM) . Volusia Co.: Cassadaga, 19 Jan 1962 (1³), S. V. Fuller (USNM) . Cassadaga, 24 Feb 1960 (1♀), S. W. Frost (USNM) . NORTH CAROLINA: Brunswick Co.: Meyers Clemmins Tract, Green Swamp, 25 Mar 1994 (1³), Sullivan et al. (USNM) . Green Swamp, Big Island Annex, 19 Sep 1994 (3³), Sullivan et al. (USNM) . MOTSU, inter. firebreaks 32 &amp; 34, 9 May 1994 (1³), 29 Aug 1994 (1³), Sullivan et al. (USNM) . Carteret Co.: Willis Road Savannah, 18 Jul 1983, 24 Aug 1983, 27 Aug 1983, 9 Sep 1983, 20 Sep 1983, J. B. Sullivan (10³) (USNM) . Craven Co.: Rd 122-2, powerline sav., Croatan National Forest, 7 Sep 1999 (1³), J. B. Sullivan (USNM) . Croatan Road 147, 2 June 1997 (1♀), J. B. Sullivan (USNM) . New Hanover Co.: MOTSU buffer, Peter’s Point, 27 Sep 1994 (1³), 29 Aug 1994 (1³), Sullivan et al. (USNM) . Carteret Co.: Sam Hatcher Road, N34 44.719, W076 55.659, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.92765&amp;materialsCitation.latitude=34.745316" title="Search Plazi for locations around (long -76.92765/lat 34.745316)">Croatan National Forest</a>, 22 Jun 2008 (1³), J. B. Sullivan (USNM) . Camp Sam Hatcher Road, 29 Mar 2002 (5³), J. B. Sullivan (USNM) . Old landfill off Hobbs Road, 29 Mar 2002 (4³, 1♀), J. B. Sullivan (USNM) . Willis Savanna, 14 Mar 2002 (1³), J. B. Sullivan (USNM) . North of Millis Sa- vanna, Croatan National Forest, 14 Mar 2002 (3³), J. B. Sullivan (USNM) . Jct state hiway 101 &amp; 181, 8 Aug 1972 (1³), J. B. Sullivan (USNM) . Onslow Co.: Camp Lejune Marine Base, 9 Apr 1999 (1³), J. B. Sullivan (USNM) . Pender Co.: Holly shelter gamelands, 12 Apr 1995, 10 June 1995, J. B. Sullivan (3³, 1♀) (USNM) .</p><p>Distribution and biology. Aethes triassumenta is apparently restricted to the southeastern U.S., ranging from Florida to North Carolina. Three specimens from Florida were misidentified by Kearfott (1907) and included in his type series of Aethes obliquana . Aethes triassumenta is recorded sporadically throughout the year with no conspicuous peak fight period(s). The early stages are unknown.</p><p>Remarks. Pogue’s (1986) manuscript name triassumenta was proposed as the type species of his manuscript genus Morta . However, because triassumenta appears to fit rather convincingly in Aethes based on facies and genital morphology, the nomen nudum Morta Pogue and Mickevitch, 1990 is placed under Aethes .</p></div>	https://treatment.plazi.org/id/03E08794FFD7FFEBFF5B7855FB59FABD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD6FFEBFF5B79C2FD92F809.text	03E08794FFD6FFEBFF5B79C2FD92F809.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyraylia Walsingham 1897	<div><p>Thyraylia Walsingham, 1897</p><p>Type species: Conchylis bunteana Robinson, 1869</p><p>Thyraylia [with its synonym Acornutia Obraztsov, 1944; type species Cochylis nana (Haworth, 1811)] includes seven Nearctic species (including the one transferred herein) and one Holarctic species. Although Razowski (1960) treated it as a valid genus (as its junior synonym Acornutia), he later (Razowski 1970, 1997, 2011) treated Acornutia (which he considered a monotypic genus for the Holarctic species nana) as a synonym of Cochylis . However, Thyraylia has long been recognized as a valid genus in North America (e.g., Powell 1983, Pogue 1986, Brown 2005, Metzler &amp; Brown 2014), and it currently includes the following species: bana (Kearfott, 1907), bunteana (Robinson, 1869), discana (Kearfott, 1907), gunniana (Busck, 1907), hollandana (Kearfott, 1907), nana, and voxcana (Kearfott, 1907) (Metzler &amp; Brown 2014). Species of Thyraylia are characterized by an elongate, somewhat upcurved, distally attenuate valva, a sacculus with a long, free distal tip near the middle of the venter of the valva, and a slender, curved, apically pointed phallus that lacks cornuti (e.g., Razowski 1997: figs. 87‒91). Phalonia wiscana Kearfott is herein transferred to Thyraylia .</p></div>	https://treatment.plazi.org/id/03E08794FFD6FFEBFF5B79C2FD92F809	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD9FFE5FF5B78BDFD6AFE85.text	03E08794FFD9FFE5FF5B78BDFD6AFE85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyraylia wiscana (Kearfott 1907) Brown 2019	<div><p>Thyraylia wiscana (Kearfott, 1907), new combination</p><p>Figs. 17, 18, 32, 46</p><p>Phalonia wiscana Kearfott, 1907: 77; McDunnough 1939: 60; Klots 1942: 419.</p><p>Phalonia acropeda Meyrick, 1912: 35 (unnecessary replacement name).</p><p>Cochylis wiscana: Powell 1983: 42; Razowski 1977: 132.</p><p>[ Cochylini New Genus 6] wiscana: Brown, 2005: 208; Metzler &amp; Brown 2014: 278.</p><p>Described in Phalonia and treated as such by McDunnough (1939), wiscana was transferred to “incertae sedis” by Powell (1983). Pogue (1986) proposed the name Poterioparvus in his unpublished dissertation for Phalonia wiscana, and Pogue and Mickevitch (1990) used Poterioparvus in their phylogenetic analysis of the genera of North American Cochylidae . Brown (2005) treated it as “ Cochylini New Genus 6 [ wiscana group].” According to Pogue (1986), Poteriorparvus is the sister group to Thyraylia . Based on the distinctive shape of the valva with a long, free spine near the middle, which is the most convincing synapomorphy for Thyraylia, wiscana is transferred to Thyraylia .</p><p>Thyraylia wiscana is easily distinguished from other species of Thyraylia by its larger size and forewing pattern (Fig. 17, 18). Forewing length averages about 9.0 mm, and the pattern includes a pale whitish gray ground color with a pinkish hue and a poorly-defined, beige to rust median fascia that extends uninterrupted from the hind margin to the costa. The only species of Cochylina of similar size in the eastern half of the U.S. are small individuals of Phtheochroa, most of which are dark brown. The male genitalia (Fig. 32) are most easily distinguished by the somewhat trifurcate apex of the socii.</p><p>Distribution and biology. Aside from the holotype from Wisconsin (AMNH), I examined nine specimens from Kentucky (USNM), most of which were collected by malaise trap, and one from Pennsylvania (MSC). Hence, it is possible that this species is diurnal and not avidly attracted to light. Razowski (1997) reported this species from Saskatchewan (n = 2); however, the illustrations of the male genitalia that he provided (Razowski 1997: figs 100–101) belong to a species of Cochylichroa. The Kentucky specimens are all from April, the specimen from Pennsylvania is from late May‒early June, suggesting a rather early flight period.</p><p>Holotype ♀, USA, Wisconsin (AMNH).</p></div>	https://treatment.plazi.org/id/03E08794FFD9FFE5FF5B78BDFD6AFE85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD8FFE5FF5B7D8AFA62FCED.text	03E08794FFD8FFE5FF5B7D8AFA62FCED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eupinivora Brown 2013	<div><p>Eupinivora Brown, 2013</p><p>Type species: Eupinivora ponderosae Brown, 2013</p><p>Eupinivora was described for seven species ranging from the mountains of the western United States (Nevada, Utah, Wyoming, Colorado, Arizona, New Mexico, and Texas) south through Nuevo Leon and Durango to Estado de Mexico, Mexico. Eupinivora are large moths (forewing length 7.5–12.0 mm) with a rusty forewing pattern reminiscent of many species that feed on pines (e.g., Rhaycionia Hübner [1825], Eucopina Gilligan &amp; Wright, 2014, etc.). Brown (2013) commented that Eupinivora is closely related to Henricus Busck, 1943, and DNA barcodes provide additional support for this hypothesis. Four species of Eupinivora have been reared from the cones of Pinus arizonica var. cooperi Blanco (Pinaceae) and one from Pinus ponderosa P. &amp; C. Lawson, and all of the species occur in habitats dominated by conifers at elevations between ca. 1700 and 2750 m. Below one additional species is transferred to the genus.</p></div>	https://treatment.plazi.org/id/03E08794FFD8FFE5FF5B7D8AFA62FCED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
03E08794FFD8FFE5FF5B7FA1FB55FA30.text	03E08794FFD8FFE5FF5B7FA1FB55FA30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eupinivora parotanus (Razowski & Becker 2010) Brown 2019	<div><p>Eupinivora parotanus (Razowski &amp; Becker, 2010), new combination</p><p>Henricus parotanus Razowski &amp; Becker, 2010: 435 .</p><p>Henricus parotanus was described from a male and a female from Cofre de Parote, Veracruz, Mexico. Razowski and Becker (2010) commented “Facies resembling Rhyacionia Hubner [1925].” The forewing shape (long and slender) and maculation ( Rhyacionia or Retinia -like), the absence of a cornutus in the vesica, and the absence of a hair pencil from the base of the valva in the male genitalia all deviate from most species of Henricus . In facies and genitalia (male and female), Henricus parotanus (see Razowski and Becker 2010: figs. 1, 2, 23, 29, 30) is extremely similar to Eupinivora rufofascia Brown, 2013 (see Brown 2013: figs. 8, 14, 18). On the basis of these similarities, parotanus is transferred to Eupinivora . Eupinivora parotanus can be separated from E. rufofascia by the following features of the male genitalia: in E. parotanus the spines from the sacculus are shorter than those of E. rufofascia; in E. rufofascia the distal portion of the median process of the transtilla has an abruptly attenuate distal process in the apical 0.2 compared to the evenly rounded apex of E. parotanus; and in E. rufofascia the phallus has a strong internal sclerite that is absent in E. parotanus . The addition of E. parotanus to the genus increases the known geographic distribution of Eupinivora to the state of Veracruz, Mexico, and the elevational range to 3300 m.</p><p>Holotype ³, Mexico, Veracruz, Cofre de Parote 3300 m, 6 Jun 1997, V.O. Becker (VBC).</p></div>	https://treatment.plazi.org/id/03E08794FFD8FFE5FF5B7FA1FB55FA30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Brown, John W.	Brown, John W. (2019): New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae). Zootaxa 4671 (2): 195-222, DOI: 10.11646/zootaxa.4671.2.2
