taxonID	type	description	language	source
03E187D7315DFFCADBD6FDD7FC61FAC6.taxon	discussion	Werner (1990) treated the species discussed in this paper as part of the genus Zonantes Casey, 1895 (type-species: Xylophilus subfasciatus Leconte, 1855). In his monography, however, Báguena Corella (1948) considered the North American genus Zonantes a subgenus of the Australian genus Syzeton Blackburn, 1891 (type species: Syzeton laetus Blackburn, 1891, fixed by Nardi (2007 )). A superficial overview of species respectively assigned to each of these two genera shows a great degree of morphological similarities. Dr. Traci Grzymala confirmed this impression in her doctoral thesis (2016) by revisiting the taxonomic status and investigating the phylogenetic relationships of these two genera. She concluded: “ The genus Zonantes was recovered as a clade within Syzeton in the concatenated analysis, rendering Syzeton paraphyletic. No single gene analysis recovered Syzeton and Zonantes as distinct clades and there are no consistent morphological differences found between these two taxa ”. She therefore proposed Zonantes as a synonym of Syzeton. Her doctoral work not being formally published as required by the ICZN, this synonymy is not effective. I therefore propose to validate it here by confirming that Zonantes Casey, 1895 is a junior synonym of Syzeton Blackburn, 1891, syn. nov. Consequently, all North American species treated by Werner (1990) as Zonantes are now placed in Syzeton, as detailed hereafter. The genus is characterized by the following morphological features (see also Grzymala (2016) for an extensive description): elytral pubescence generally well developed, uniseriate (Figure 1 a); head transverse, bean-shaped in dorsal view, with base both truncated and concave (Figure 1 b); anterior margin of pronotum broadly convex, formed from two segments converging in an obtuse angle toward the head (Figure 1 c); antennomere 3 longer than the pedicel (Figure 1 d); ♀♀ with a setose pit on the last abdominal ventrite (Figure 1 e, f); mesothoracic tarsi with a ventral brush of long setae in both sexes (Figure 1 g); presence of a small, basal, ventral, setae-filled excavation on hind femora of both ♂♂ and ♀♀ (Figure 1 h, i); ♂ hind femora devoid of large pads of setae (contrasting with, for instance, ♂♂ of Elonus Casey, 1895). Most species exhibit distinct but moderate sexual dimorphism, such as the size of the eyes (enlarged in ♂♂) or the conformation of the antennae (shorter in ♀♀). Male aedeagus broadly cylindrical, moderately curved in lateral view, generally well sclerotized; phallobase not distinctly separated from apicale by a suture, as seen in other genera such as Mixaderus Collado & Alonso-Zarazaga, 1996 (Gompel, 2021) or Cobososia Collado & Alonso-Zarazaga, 1996 (Telnov & Gompel, 2021); apicale devoid of paired accessory lobes (Grzymala, 2016) as often found in Pseudoanidorus Pic, 1893 or Anidorus Mulsant & Rey, 1866 (N. Gompel & Barrau, 2002); ventral side of the phallobase with a large oval opening that never reaches the base of the aedeagus; a set of complex sclerotized internal plates often present in the aedeagus and visible with retrolighting. Syzeton species are broadly distributed around the world, except in Europe and North-Africa. Syzeton specimens are typically collected by sweeping grass or beating the foliage of shrubs and hardwood trees. They are also often attracted to lights or obtained from flight intercept traps and Malaise traps, in forests or grasslands. Key to US Syzeton species (modified and expanded from Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315DFFCADBD6FDD7FC61FAC6.taxon	discussion	Separation of S. hubbardi (Casey, 1895), S. subfasciatus (LeConte, 1875) and S. nubifer (LeConte, 1878) Syzeton nubifer exists in two forms, sometimes found in sympatry: a dark form (Figure 2 d), with a black pronotum, darken mid- and hind femora, and well developed elytral markings extending along the lateral margins, and a pale form, with a brown pronotum, legs entirely orange, and elytral markings reduced to a single oblique stripe on each side, often interrupted before the lateral margin. The latter can easily be mistaken for S. hubbardi (Werner, 1990) or S. subfasciatus (LeConte, 1875). In ♂♂, the curvature of the foretibiae (Figure 5 c, d, j) and the shape of the aedeagus, particularly in ventral view with the asymmetrical apicale (Figure 3 c, d, j) make the distinction unambiguous. In ♀♀, however, the separation is more subtle: ♀♀ of S. subfasciatus (LeConte, 1875) are generally stouter, with ovoid elytra. The elytral markings are variable, from a solid transverse band of black pigmentation, to blotches, and to nearly no markings at all in exceptional cases. Females of the pale form of S. nubifer (LeConte, 1878) often look similar to S. subfasciatus (LeConte, 1875) with a solid transverse band of black pigmentation, but the band always centers slightly posterior to middle of elytron in S. nubifer (LeConte, 1878) (Figure 2 o) while it is always centered near the midpoint of the elytra in S. subfasciatus (LeConte, 1875). In females of S. hubbardi (Werner, 1990), the markings form an horizontal transverse band, or, when separated in blotches, the blotches closer to the elytral suture are strongly oblique, nearly parallel to the elytral suture, forming a 20 º angle (Figure 2 n). In S. nubifer (LeConte, 1878), the inner blotches are less strongly oblique, forming a 30 – 40 º angle together (Figure 2 o). Isolated females of these three species may be difficult to name with certainty.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315FFFCCDBD6FA24FAE4FE4D.taxon	description	(Figures 1 b, d; 2 a, m; 3 a; 4 h; 5 a)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315FFFCCDBD6FA24FAE4FE4D.taxon	materials_examined	Holotype: Oklahoma: Latimer Co. (USNM) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315FFFCCDBD6FA24FAE4FE4D.taxon	description	Because the two taxa were described in the same paper, the question of priority arises. Leaning on ICZN Art. 24.2 (International Commission on Zoological Nomenclature, 1999), I decided to maintain Syzeton ouachitanus (Werner, 1990) as the senior name for two reasons. First, its description precedes that of Zonantes mississippiensis Werner, 1990 in Werner’s paper. Second. in the interest of nomenclature stability, as Zonantes mississippiensis Werner, 1990 is described from only 2 females and much less information is available for this taxon.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315FFFCCDBD6FA24FAE4FE4D.taxon	materials_examined	Material examined. United States. Florida: Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 6. V. 2007, M. C. Thomas leg. (1 ♂, NGPC); Sarasota County, Myakka River State Park, FFA / N. Plot, (10 m) [27 ° 12 ’ N 82 ° 14 ’ W], 2. XI. 2000, funnel trap / turpentine trap, J. L. Foltz leg. (1 ♀, FSCA); Suwannee County, F- 49 [30 ° 14 ’ N 82 ° 59 ’ W], 2. IX. 1976, at river, black light trap, J. R. Wiley leg. (2 ♀♀, FSCA). Georgia: Clarke County, Athens, Whitehall Forest, (210 m) [33 ° 53 ’ N 83 ° 21 ’ W], 13. IX. 1978, black light trap, R. Turnbow leg. (1 ♂, FSCA). Mississippi: Hancock County, I- 10 rest area and Welcome Center, (5 m) [30 ° 18 ’ 47 ” N 89 ° 35 ’ 59 ” W], 17. VI. 1982, E. G. Riley leg. (1 ♀ paratype of Zonantes mississippiensis Werner, 1990, ERPC). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W], UV,. VIII. 1998, K. Stephan leg. (1 sp., TAMU). Texas: Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 22. VIII. 2016, V. Belov leg. (1 ♂, NGPC); at lights, 12. IX. 2016, V. Belov leg. (1 sp., VBPC); at lights, 9. IV. 2017, V. Belov leg. (1 sp., VBPC); at lights, 16. IV. 2017, V. Belov leg. (1 ♂, NGPC); at lights, 13. V. 2017, V. Belov leg. (2 sp., VBPC); at lights, 20. VI. 2017, V. Belov leg. (2 sp., VBPC); at lights, 22. VI. 2020, V. Belov leg. (1 ♂, VBPC); at lights, 12. VII. 2020, V. Belov leg. (1 ♀, NGPC); at lights, 19. VII. 2020, V. Belov leg. (1 ♂, VBPC); at lights, 24. VIII. 2020, V. Belov leg. (1 ♂, VBPC); San Patricio County, Welder Wildlife Refuge [28 ° 07 ’ N 97 ° 19 ’ W], at light, 25. VI. 1981, J. C. Schaffner leg. (1 ♂, TAMU).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315FFFCCDBD6FA24FAE4FE4D.taxon	distribution	Distribution. United States: Texas to Florida, North to Oklahoma (extended from Werner (1990 )).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315AFFC1DBD6FF2AFA58FBD5.taxon	description	(Figures 1 e – f; 2 b; 3 b; 5 b, o)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315AFFC1DBD6FF2AFA58FBD5.taxon	materials_examined	Lectotype: Pennsylvania (MCZC) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315AFFC1DBD6FF2AFA58FBD5.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VIII. 2013, T. King leg. (1 ♂, FSCA). Arkansas: Logan County, 12 mi. NW Havana [35 ° 14 ’ N 93 ° 40 ’ W], 27. V. 1995, J. C. Schaffner leg. (1 sp., TAMU); Sharp County, Harold E. Alexander WMA [36 ° 12 ’ N 91 ° 26 ’ W], Mixed hardwood forest, Beaten from branches of hardwood tree, 22. VIII. 2010, J. P. Gruber leg. (1 ♀, WIRC). Florida: Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], 5. V. 2008, M. C. & S. Thomas leg. (1 ♀ and 2 ♂♂, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], 6. V. 2008, M. C. & S. Thomas leg. (1 ♀, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], 6. V. 2017, M. C. Thomas leg. (1 ♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 27. IV. 1991, M. C. Thomas leg. (1 ♂ and 2 ♀♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 27. IV. 1991, M. C. Thomas leg. (1 ♂ and 2 ♀♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 6. V. 2007, M. C. Thomas leg. (2 ♂♂, FSCA); Santa Rosa County, 3.5 mi. N Munson, (0 m) [30 ° 54 ’ N 86 ° 51 ’ W], miscellaneous night / light collection, 10. IV. 1999, Skelley P. leg. (1 ♂, FSCA). Georgia: Clarke County, Black Rock Mountain State Park, (200 m) [33 ° 54 ’ N 83 ° 16 ’ W], 29. V. 1992, J. Pickering leg. (1 ♂, FSCA); Clarke County, Barnett Shoals area, (220 m) [33 ° 50 ’ N 83 ° 22 ’ W], 30. IV. 1987, R. Morris leg. (1 ♀, FSCA); Dekalb County, Stone Mountain, (320 m) [33 ° 48 ’ N 84 ° 10 ’ W], 16. V. 1991, J. E. Wappes leg. (1 ♀ and 2 ♂♂, FSCA); Henry County, Nail Road, (275 m) [33 ° 25 ’ N 84 ° 11 ’ W], black light trap,. IV. 1994, R. Beshear leg. (1 ♂, FSCA); Rabun County, Black Rock Mountain State Park [34 ° 54 ’ N 83 ° 24 ’ W], 27. VI. 1982, F. N. Young leg. (1 ♂, FSCA). Indiana: Brown County, N. of Belmont [39 ° 09 ’ N 86 ° 20 ’ W], 22. VIII. 1984, C. W. Hagen Jr. leg. (1 ♂, FSCA); Brown County, Bear Wallow [39 ° 19 ’ N 86 ° 22 ’ W], 1. VI. 1977, J. E. Wappes leg. (1 ♂, FSCA); Brown County [39 ° 09 ’ N 86 ° 20 ’ W], 5. VI. 1987, M. M. Downie leg. (1 ♀, FSCA); Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], black light trap, 17. VI. 1990, F. N. Young leg. (1 ♂, FSCA); Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], 18. VI. 1992, F. N. Young leg. (1 ♂, FSCA); Orange County, Shirley Creek Horsecamp, (210 m) [38 ° 38 ’ N 86 ° 35 ’ W], mercury vapors + black light, 26. V. 2007, R. Turnbow leg. (1 ♂, FSCA). Iowa: Johnston County, Saylorville Lake [41 ° 43 ’ N 93 ° 42 ’ W], 9. VII. 1983, J. E. Wappes leg. (1 ♀, FSCA); Polk County, Brown Forest Preserve [41 ° 32 ’ N 93 ° 42 ’ W], 20. VI. 1982, J. E. Wappes leg. (1 ♀, FSCA); Polk County, Jester Park [41 ° 47 ’ N 93 ° 47 ’ W], 15. VI. 1985, J. E. Wappes leg. (1 ♀, FSCA). Maryland: Prince George’s County [39 ° 03 ’ N 76 ° 47 ’ W], 14. VI. 1993, J. Pickering leg. (1 ♂, FSCA); Prince George’s County [39 ° 03 ’ N 76 ° 47 ’ W], 21. VI. 1993, J. Pickering leg. (1 ♀, FSCA). Illinois: Jackson County, Shawnee National Forest, 7.8 km NW of Pomona [37 ° 40 ’ 50 ” N 89 ° 23 ’ 42 ” W], Upland hardwood forest, Beaten from branches of Carya sp., 23. VIII. 2010, J. P. Gruber leg. (1 ♀ and 3 ♂♂, WIRC); Chicago, Blackwelder [41 ° 41 ’ N 87 ° 40 ’ W],. VI., Wm. S. Marshall leg. (1 ♀, WIRC). Massachusetts: Middlesex County, Great Meadows National Wildlife Refuge, Concord Unit [42 ° 28 ’ N 71 ° 19 ’ W], 23. VI. 2001, E. G. Riley leg. (2 sp., TAMU). Missouri: Greene County, 10 mi. E Springfield [37 ° 12 ’ N 93 ° 17 ’ W], 15. VI. 1981, R. Haswell leg. (2 sp., TAMU), 15. VI. 1981. New Jersey: Essex County 40 ° 44 ’ N 74 ° 16 ’ W, .. 2010, V. Belov leg. (1 ♂ and 1 sp., VBPC); Maplewood [40 ° 44 ’ N 74 ° 16 ’ W], .. 2013, V. Belov leg. (1 ♂, VBPC). New York: Madison County, Georgetown, Muller Hill, Blueberry Trail, 100 Acre Evans camp [42 ° 46 ’ N 75 ° 46 ’ W], Woods, Barrier trap, 8. VII. 2002, M. H. Evans leg. (1 ♂, WIRC), Flight intercept trap, 24. VII. 2006, M. H. Evans leg. (2 ♀♀, WIRC). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W],. V. 1995, K. Stephan leg. (1 sp., TAMU), UV,. VIII. 1998, K. Stephan leg. (1 sp., TAMU). Pennsylvania: Lawrence County, 2.5 mi. S of Lebanon [40 ° 17 ’ N 76 ° 24 ’ W],. VI. 2012, B. Hays leg. (1 sp., TAMU); Lebanon County, 2.5 mi. S of Lebanon [40 ° 17 ’ N 76 ° 24 ’ W], 8. VII. 1986, J. Fetter leg. (1 ♂, WIRC). South Carolina: Chesterfield County, Sand Hills State Forest, 6.5 km WNW of Patrick [34 ° 35 ’ N 80 ° 06 ’ W], Mixed coniferous / hardwood forest, Beaten from branches of Cornus florida L., 21. VIII. 2011, J. P. Gruber leg. (1 ♂, WIRC); Fairfield County, Lake Wateree State Recreation Area, Desportes Island, 7.4 km SW of Liberty Hill [34 ° 25 ’ 58 ” N 80 ° 51 ’ 37 ” W], Beaten from forest understory vegetation, 23. VIII. 2011, J. P. Gruber leg., (1 ♀ and 1 ♂, WIRC); Newberry County, Mollys Rock Picnic Area, (110 m) [34 ° 23 ’ 16 ” N 81 ° 32 ’ 33 ” W], 12. VI. 2013, E. G. Riley leg. (1 sp., TAMU). Tennessee: Blount County, Cove Mountain, Cold Spring Gap, (760 m) [35 ° 37 ’ 24 ” N 83 ° 49 ’ 22 ” W], 28. VI. 2001, E. G. Riley leg. (1 sp., TAMU). Texas: Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W] (50 m), Lindgren funnel trap, 21. VII. 2007, E. G. Riley leg. (1 sp., TAMU), Lindgren funnel trap, 13. IV. 2008, E. G. Riley leg. (1 sp., TAMU), Lindgren funnel trap, 10. V. 2008, E. G. Riley leg. (1 sp., TAMU), Lindgren funnel trap, 7. VI. 2008, E. G. Riley leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], Ground-level flight intercept, 18. V. 2003, E. G. Riley leg. (1 sp., TAMU). Virginia: Botetourt County, Hwy. 615 near Hipes, 10.5 mi. W Jet Hwy. 220 [37 ° 39 ’ 09 ” N 79 ° 55 ’ 40 ” W], 24. V. 2006, E. G. Riley leg. (1 sp., TAMU). Wisconsin: Adams County, Quincy Bluffs - The Nature Conservancy [43 ° 51 ’ 59 ” N 89 ° 53 ’ 06 ” W], Lindgren funnel trap, 18. VII. 2003, D. K. Young leg. (1 ♂, WIRC), Lindgren funnel trap, 30. VII. 2003, D. K. Young leg. (1 ♂, WIRC), Old Populus deadfall, Malaise trap, 16. VI. 2016, D. K. Young leg. (1 ♀, WIRC), Quercus / Populus swail, Malaise trap, 27. VI. 2016, D. K. Young leg. (1 ♂, WIRC); Columbia County, Rocky Run creek State Fishery Area [43 ° 27 ’ 38 ” N 89 ° 19 ’ 31 ” W], Grassland, Swept from long grasses and other prairie-type vegetation, 23. VII. 2008, J. P. Gruber leg. (1 ♂, WIRC); Dane County, 18. VI. 1898, Wm. S. Marshall leg. (1 ♀, WIRC), 6. VII. 1899, Wm. S. Marshall leg. (1 ♀, WIRC); Dane County, Monona Lake [43 ° 04 ’ N 89 ° 21 ’ W], White oak, 13. VII. 1912, J. G. Sanders leg. (2 ♀♀ and 1 ♂, WIRC); Dane County, Madison, Eagle Heights, (300 m) [43 ° 05 ’ N 89 ° 26 ’ W], Grass sweeping, 13. VII. 2003, N. Gompel leg. (1 ♀, NGPC); Dane County, Madison [43 ° 04 ’ N 89 ° 24 ’ W], Sweeping, 8. VII. 1929, L. C. Fluke leg. (1 ♀, WIRC); Dane County, Mazomanie, (250 m) [43 ° 13 ’ N 89 ° 47 ’ W], 25. VI. 1983, N. Ravdin leg. (1 sp., WIRC); Florence County, Spread Eagle Barrens State Natural Area, (355 m) [45 ° 51 ’ 54 ” N 88 ° 08 ’ 58 ” W], Swept from vegetation in grassland / barrens pine / oak / aspen, 7. VII. 2005, J. P. Gruber leg. (1 ♀, WIRC); Grant County, Cassville Bluffs SNA [42 ° 42 ’ N 90 ° 56 ’ W], Mixed hardwood forest, Sweeping foliage near path, 18. VII. 2003, J. P. Gruber leg. (5 ♀♀ and 1 sp., WIRC); Grant County, Wyalusing State Park, (300 m) [42 ° 58 ’ N 91 ° 07 ’ W], Forest, Swept from foliage, 14. VII. 1999, J. P. Gruber leg. (1 sp., WIRC); Grant County, Witted oak, 2. VI. 1949, W. McNeel leg. (6 ♀♀ and 1 ♂, WIRC); Green County, Abraham’s wood SNA [42 ° 41 ’ N 89 ° 29 ’ W], Mesic hardwood forest, Gilled fungus baited pitfall trap, 11. VII. 2001, J. P. Gruber leg. (1 ♀, WIRC); Iowa County, Arena Pines-Sand Barrens State Natural Area, (220 m) [43 ° 10 ’ 33 ” N 89 ° 55 ’ 53 ” W], Sand barrens with jack pine, black oak, Beaten from branches of living Quercus sp., 12. VI. 2012, J. P. Gruber leg. (1 ♂ and 1 ♀, WIRC); Jackson County, Black River State Forest [44 ° 13 ’ N 90 ° 39 ’ W], Oak-pine forest, Lindgren funnel trap, 24. VI. 1999, J. P. Gruber leg. (1 ♂, WIRC; 1 sp., NGPC), 12. VII. 1999, J. P. Gruber leg. (2 ♀♀, WIRC), 1. VIII. 1999, J. P. Gruber leg. (1 ♀, WIRC); Jackson County, Brockway, (250 m) [44 ° 16 ’ N 90 ° 50 ’ W], Gypsy Moth-M. T., 4. VIII. 1975 (1 sp., NGPC); Jackson County, T 21 N R 4 W S 27, Oak-pine forest, Gypsy Moth-M. T., 15. VII. 1975 (1 ♀ and 2 ♂♂, WIRC), 28. VII. 1975 (2 ♀♀ and 5 ♂♂, WIRC); Jackson County, T 21 N R 4 W S 33, Gypsy Moth-M. T., 28. VI. 1976 (1 ♂, WIRC), 2. VIII. 1976 (1 ♀, WIRC); Juneau County, Necedah national wildlife refuge, Flight intercept trap, 1. VII. 1996, K. Pope leg. (1 ♂ and 1 ♀, WIRC), 3. VII. 1996, K. Pope leg. (1 ♂ and 1 ♀, WIRC), 9. VII. 1996, K. Pope leg. (1 ♂ and 3 ♀♀, WIRC), Pitfall trap, 23. VII. 1996, K. Pope leg., (1 ♀, WIRC); La Crosse County, Coulee Experimental State Forest [43 ° 51 ’ 32 ” N 91 ° 01 ’ 00 ” W], Grass sweeping, 25. VII. 2003, N. Gompel leg. (1 sp., NGPC); La Crosse County, Environs west of West Salem [43 ° 54 ’ N 91 ° 10 ’ W], Mixed hardwood forest, Suburban Malaise trap, 10. VII. 2010, A. M. Shorter leg. (1 ♂, WIRC); La Crosse County, West Salem [44 ° 03 ’ N 91 ° 04 ’ W], Medium-aged oak / hickory forest, Root sever; Flight intercept trap, 23. VIII. 2004, K. Werner leg. (2 ♀♀, WIRC); Lafayette County, about 3 miles north of Argyle [42 ° 44 ’ 17 ” N 89 ° 51 ’ 05 ” W], Southern mixed hardwood forest, Flight intercept trap, 24. VI. 2001, J. P. Gruber leg. (1 sp., NGPC), Unbaited Townes malaise trap, 11. VII. 2001, C. M. Brabant leg. (1 ♂, WIRC), Flight intercept trap, 11. VII. 2001, J. P. Gruber leg. (4 sp., NGPC), 5. VIII. 2001, J. P. Gruber leg. (1 sp., NGPC), 3. IX. 2001, J. P. Gruber leg. (2 sp., NGPC), 8. VII. 2004, N. Gompel leg. (1 ♂, NGPC); Lafayette County, Yellowstone Lake State Wildlife Area [42 ° 46 ’ N 89 ° 57 ’ W], Oak savanna, From 30 cm long Quercus macrocarpa log, Lightproof rearing container, 4. IX. 2003, D. Biggs leg. (1 ♂, WIRC); Marathon County, Big Eau Pleine County Park [44 ° 44 ’ N 89 ° 50 ’ W], Northern mesic forest, Beaten from branches of hardwood tree, 6. VII. 2013, J. P. Gruber leg. (1 ♀ and 2 ♂♂, WIRC), Crawling on debarked trunk of dead standing hardwood tree, With lots of borer holes, 6. VII. 2013, J. P. Gruber leg. (1 ♂, WIRC); Marquette County, Summerton Bog The Nature Conservancy, 3 mi. SE Oxford [43 ° 45 ’ N 89 ° 31 ’ W], Sweep pound / spring margin, 6. VII. 1990, D. K. Young leg. (2 ♀♀, WIRC); Monroe County, Fort McCoy, N. Post, 1 mi. W of Big Rock [43 ° 59 ’ N 90 ° 40 ’ W], Oak savanna, Flight intercept trap, 14. VII. 1997, J. A. Maxwell leg. (1 ♀, WIRC); Monroe County, Fort McCoy, N. Post, Dorn’s Cut [43 ° 59 ’ N 90 ° 40 ’ W], Mesic oak forest, Flight intercept trap, 14. VIII. 1997, J. A. Maxwell leg. (1 ♂, WIRC), 29. VIII. 1997, J. A. Maxwell leg. (1 ♀, WIRC); Monroe County, Fort McCoy Sand Quar., Sandy prairie / oak savanna, Unbaited Lindgren funnel trap, 24. VI. 1999, J. P. Gruber leg. (1 sp., NGPC); Monroe County, Fort McCoy State Natural Area [43 ° 59 ’ N 90 ° 40 ’ W], Oak savanna, Flight intercept trap, 27. VII. 2002, J. P. Gruber leg. (2 sp., NGPC); Monroe County, Fort McCoy State Natural Area [43 ° 59 ’ N 90 ° 41 ’ W], Oak savanna at margin of sandy prairie, Flight intercept trap, 17. VI. 2007, J. P. Gruber leg. (1 ♂, WIRC), 30. VI. 2007, J. P. Gruber leg. (2 ♀♀ and 1 ♂, WIRC), 14. VII. 2007, J. P. Gruber leg. (1 ♀, WIRC); Richland County, LWRSGA Lone Rock Unit [43 ° 11 ’ N 90 ° 14 ’ W], Sandy oak barrens, Leaf litter near fallen tree, 23. VIII. 2000, J. P. Gruber leg. (1 ♀, WIRC); Richland County, LWRSGA Lone Rock Unit, Station 2, (220 m) [43 ° 11 ’ N 90 ° 14 ’ W] Swept from vegetation in grassland / barrens pine / oak / aspen, 27. VI. 2001, D. K. Young leg. (3 ♂♂, WIRC), Flight intercept trap, 27. VI. 2001, D. K. Young leg. (1 ♀, WIRC), Quercus savanna, Malaise trap baited - wintergreen oil, 5. X. 2006, D. K. Young leg. (1 ♀, WIRC); Sauk County, Hemlock Draw, (330 m) [43 ° 21 ’ N 89 ° 58 ’ W], Lindgren funnel trap., 2. VII. 2003, D. K. Young leg. (1 ♂, WIRC); Sauk County, Spring Green Preserve SNA, (220 m) [43 ° 11 ’ N 90 ° 03 ’ W], Sandy oak barrens, Flight intercept trap, 1. VII. 2001, J. P. Gruber leg. (1 sp., NGPC), Dry-mesic southern forest, Swept from foliage near path, 13. VII. 2003, J. P. Gruber leg. (3 sp., NGPC), Dry-mesic southern forest, Underside of leaf on branch of Carya ovata, 12. VI. 2012, J. P. Gruber leg. (2 ♀♀ and 5 ♂♂, WIRC); Sauk County, Spring Green Preserve SNA (270 m) [43 ° 12 ’ N 90 ° 03 ’ W], Dry-mesic southern forest, Found on underside of leaf on branch of Carya ovata tree, 12. VI. 2012, J. P. Gruber leg. (1 ♀, WIRC); Sheboygan County, Kohler-Andrae State Park, Woodland Dunes Nature Trail [43 ° 39 ’ 07 ” N 87 ° 43 ’ 38 ” W], Mixed hardwood / coniferous forest, Beaten from branches of hardwood tree, 30. VII. 2014, J. P. Gruber leg. (4 ♀♀ and 7 ♂♂, WIRC); Sheboygan County, Sheboygan [43 ° 39 ’ N 87 ° 43 ’ W], 27. VI. 2001, P. Messer leg. (1 ♂, WIRC); Vernon County, Tunnelville Cliffs State Natural Area [43 ° 34 ’ N 90 ° 41 ’ W], Mixed hardwood forest, Beaten from branches of Carya sp. tree, 19. VII. 2014, J. P. Gruber leg., (3 ♂♂, WIRC); Vernon County, Romance Prairie State Natural Area [43 ° 33 ’ 11 ” N 91 ° 08 ’ 36 ” W], Junction of oldfield / grassland and mixed forest, Beaten from branches of hardwood tree, 30. VI. 2013, J. P. Gruber leg. (1 ♀, WIRC); Waukesha County [43 ° 00 ’ N 88 ° 17 ’ W], Collected from non-wilting Quercus alba, 17. VII. 1949, W. McNeel leg. (2 ♀♀, WIRC); Waukesha County, 2.8 mi W of North Prairie [42 ° 56 ’ N 88 ° 27 ’ W], 30. VI. 2001, Messer P. leg. (1 ♂, WIRC); Waupaca County [44 ° 21 ’ N 89 ° 05 ’ W], 20. VI. 1920, L. G. Gentner leg. (1 ♀ and 1 ♂, WIRC); Wood County, Wood County SWA [44 ° 18 ’ N 90 ° 07 ’ W], Dry-mesic northern hardwood forest, Flight intercept trap, 6. VII. 2000, J. P. Gruber leg. (1 sp., NGPC), 21. VII. 2000, J. P. Gruber leg. (1 sp., NGPC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7315AFFC1DBD6FF2AFA58FBD5.taxon	distribution	Distribution. United States: Florida to Texas, north to Quebec and Michigan (extended from Werner (1990 )).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73154FFC0DBD6FBDEFAE3F8ED.taxon	description	(Figures 2 c, n; 3 c; 5 c)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73154FFC0DBD6FBDEFAE3F8ED.taxon	materials_examined	Holotype: “ Washington DC ” (USNM) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73154FFC0DBD6FBDEFAE3F8ED.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VIII. 2012, T. King leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 20. IV. 2017, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 22. VI. 2017, R. Turnbow leg. (1 ♀, FSCA); Alachua County, (200 m) [29 ° 36 ’ N 82 ° 22 ’ W], 21. III. 1992, J. Pickering leg. (1 ♂, FSCA); Alachua County, (200 m) [29 ° 36 ’ N 82 ° 22 ’ W], 11. IV. 1992, J. Pickering leg. (1 ♂, FSCA; 1 ♂, NGPC); Alachua County, (200 m) [29 ° 36 ’ N 82 ° 22 ’ W], 28. VIII. 1992, J. Pickering leg. (1 ♂, FSCA); Alachua County, 6 mi. SW Gainesville [29 ° 36 ’ N 82 ° 24 ’ W], 28. IV. 1975, L. R. Davis Jr. leg. (1 ♂, FSCA); Alachua County, E side of Newnan’s Lake, (40 m) [29 ° 38 ’ N 82 ° 11 ’ W], 10. VI. 1984, M. C. Thomas leg. (1 ♀, FSCA); Alachua County, Austin Cary Forest, (40 m) [29 ° 43 ’ N 82 ° 13 ’ W], 7. V. 1983, M. C. Thomas leg. (1 ♀ and 1 ♂, FSCA); Alachua County, SE of Gainesville, Kincaid Road, 0.5 mi. S Paynes Prairie [29 ° 38 ’ N 82 ° 17 ’ W], old field near Prunus umbellata, Malaise trap, 5. V. 1996, B. D. Sutton leg. (1 ♀, FSCA); Alachua County, 2 mi. N Cross Creek, (20 m) [29 ° 30 ’ N 82 ° 10 ’ W], 1. IX. 1991, M. C. Thomas leg. (3 ♂♂, FSCA); Alachua County, Archer [29 ° 32 ’ N 82 ° 31 ’ W],. V. 1984, M. J. Plagens leg. (1 ♂, FSCA); Alachua County, Gainesville, San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 17. IV. 1982, M. C. Thomas leg. (1 ♀, FSCA); Alachua County, Gainesville, San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 14. III. 1990, M. C. Thomas leg. (1 ♂, FSCA); Alachua County, Orange Heights, (50 m) [29 ° 43 ’ N 82 ° 08 ’ W], cornfield and adjacent area, sticky wire trap,. VI. 1983, M. J. Plagens leg. (3 ♀♀ and 1 ♂, FSCA),. VI. 1983; Columbia County, O’Leno State Park, (20 m) [29 ° 55 ’ N 82 ° 36 ’ W], 28. IV. 1991, R. Turnbow leg. (1 ♂, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 27. IV. 1983, M. C. Thomas & L. Parker leg. (1 ♀, NGPC; 1 ♂, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 5. V. 1983, M. C. Thomas & L. Parker leg. (2 ♂♂, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 15. VI. 1983, M. C. Thomas & L. Parker leg. (2 ♂, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 6. IX. 1983, M. C. Thomas & L. Parker leg. (1 ♀, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap,. V. 1991, J. Gleason leg. (1 ♂, NGPC); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m) [28 ° 38 ’ N 82 ° 20 ’ W], 27. IV. 1991, R. Turnbow leg. (1 ♀, NGPC; 1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m) [28 ° 38 ’ N 82 ° 20 ’ W], 28. IV. 1991, R. Turnbow leg. (2 ♀♀, FSCA); Walker County, Jasper [33 ° 49 ’ N 87 ° 16 ’ W], 10. XI. 1984, T. King leg. (1 ♀, UNHC). Arkansas: Logan County, Mt. Magazine Electronics site [35 ° 10 ’ N 93 ° 38 ’ W], 26. VI. 1993, E. G. Riley leg. (1 sp., TAMU). Florida: Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W],. IX. 1995, SPB study, funnel trap / turpentine trap, J. L. Foltz leg. (1 ♀, FSCA); Clay County, Gold Head Branch State Park, (70 m) [29 ° 50 ’ N 81 ° 57 ’ W], Screen sweep, 28. V. 1991, J. B. Woolley leg. (1 sp., TAMU) Highlands County, Archbold Biological Station, 7 mi. S. Lake Placid, (40) [27 ° 10 ’ N 81 ° 21 ’ W], 26. V. 1997, P. E. Skelley leg. (1 ♂, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], black light trap, 5. V. 2008, M. C. & S. Thomas leg. (1 ♂, FSCA); Levy County, Rt 24, near relay fower, 6 mi. NE Bronson, (30 m) [29 ° 30 ’ N 82 ° 34 ’ W], sweeping rosemary, 29. IV. 1994, F. W. Head leg. (1 ♀, FSCA); Levy County, 3.5 mi. SW Archer [29 ° 29 ’ N 82 ° 35 ’ W], black light trap, 25. V. 2018, R. Turnbow leg. (1 ♂, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], black light trap, 3. V. 2003, M. C. Thomas leg. (1 ♀ and 1 ♂, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], black light trap, 6. V. 2007, M. C. Thomas leg. (2 ♂♂, FSCA); Marion County, Silver River State Park, (20 m) [29 ° 12 ’ N 82 ° 03 ’ W], beating burned schrub, 4. VIII. 1999, C. W. Mills leg. (1 ♀ and 1 ♂, FSCA); Marion County, Ocala National Forest, 2 mi. W Grassy Pond, (30 m) [29 ° 22 ’ N 81 ° 52 ’ W], mercury vapor light, 3. VI. 1994, R. Woodruff & P. Landolt leg. (1 ♂, FSCA); Marion County, Citra, NW 24 th Avenue, (20 m) [29 ° 24 ’ N 82 ° 10 ’ W], mercury vapor and ultraviolet light, 26. III. 1993, F. W. Skillman Jr. leg. (1 ♂, FSCA); Seminole County, Econ River Wilderness Area, (20 m) [28 ° 36 ’ N 81 ° 10 ’ W], scrub oak / saw palmetto (burned), Malaise trap, 11. III. 2000, T. Smith leg. (1 ♀, FSCA); Seminole County, Econ River Wilderness Area, (20 m) [28 ° 36 ’ N 81 ° 10 ’ W], scrub oak / saw palmetto (burned), Malaise trap, 15. IV. 2000, T. Smith leg. (3 ♀♀, FSCA). Georgia: Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 5. XII. 1992, J. Pickering leg. (1 ♂, FSCA). Indiana: Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], black light trap, 28. VIII. 1988, F. N. Young leg. (1 ♂, FSCA); Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], black light trap, 11. VII. 1990, F. N. Young leg. (1 ♂, FSCA); Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], black light trap, 26. VII. 1990, F. N. Young leg. (1 ♂, FSCA); Monroe County, Bloomington [39 ° 09 ’ N 86 ° 31 ’ W], black light trap, 7. VII. 1991, F. N. Young leg. (1 ♂, FSCA). Maryland: Dorchester County, Castle Haven, (0 m) [38 ° 37 ’ N 76 ° 10 ’ W], 4. VII. 1961, H. L. Dozier leg. (1 ♂, FSCA). Missouri: Christian County, Finley River, 10 mi. S Rogersville [36 ° 57 ’ N 93 ° 03 ’ W], 8. VII. 1983, R. Haswell leg. (1 sp., TAMU). New Jersey: Essex County [40 ° 44 ’ N 74 ° 16 ’ W], 2010, V. Belov leg. (1 ♂). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W], UV,. VIII. 1998, K. Stephan leg. (2 sp., TAMU; 1 ♂, NGPC). Texas: Brazos County, College Station, Lick Creek Park, (70 m) [30 ° 33 ’ N 96 ° 13 ’ W], Night sweep bottomland trails, 14. V. 1997, E. G. Riley leg. (1 sp., TAMU); Sabine County, Mill Creek Cove, Beech Bottom, 8.8 miles NE Hemphill [31 ° 23 ’ 06 ” N 93 ° 42 ’ 32 ” W], UV light, 16. VI. 2007, Heffern D. J. leg. (1 sp., TAMU); Sabine County, Sabine National Forest, 9 mi. E Hemphill, Beech Bottom [31 ° 19 ’ N 93 ° 42 ’ W], 4. IV. 1989, Beech-Magnolia forest, Malaise trap, R. R. Anderson & E. Morris leg. (1 sp., TAMU); 6. VIII. 1989 R. R. Anderson & E. Morris leg. (1 ♀, TAMU); 7. X. 1989, R. R. Anderson & E. Morris leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], UV light trap, 10. V. 2008, Anderson & Morris leg. (1 ♂, NGPC); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], 18. V. 2003, E. G. Riley leg. (1 ♂, ERPC); 8. VI. 2003, E. G. Riley leg. (1 sp., ERPC); Walker County: Ellis Prison Unit, (60 m) [30 ° 52 ’ N 95 ° 27 ’ W], Suction trap, 3. V. 1980, D. A. Dean leg. (1 sp., TAMU).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73154FFC0DBD6FBDEFAE3F8ED.taxon	distribution	Distribution. United States: Florida to eastern Texas and Oklahoma, north to New Jersey and Missouri (Werner, 1990). The record from New Jersey is a single male, unambiguously assigned to S. hubbardi (Casey, 1895). This locality is very surprising and the stable presence of the species in this region will need to be confirmed.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73156FFC5DBD6FF2AFA5DFABC.taxon	description	(Figures 1 a, d; 2 d, o; 3 d; 5 d) Xylophilus nubifer LeConte, 1878: 425;	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73156FFC5DBD6FF2AFA5DFABC.taxon	materials_examined	Holotype: Florida, Volusia County, Enterprise (MCZC) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73156FFC5DBD6FF2AFA5DFABC.taxon	description	Zonantes nubifer (LeConte, 1878): Werner (1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73156FFC5DBD6FF2AFA5DFABC.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VII. 2013, T. King leg. (2 ♂♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 28. IV. 2011, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 31. V. 2014, R. Turnbow leg. (2 ♀♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 3. VII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 6. VII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 11. VII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 30. VII. 2014, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 10. VIII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 19. VIII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 6. IV. 2015, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 6. IV. 2015, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 11. VI. 2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 30. VI. 2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 11. VII. 2015, R. Turnbow leg. (6 ♀♀ and 2 ♂♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 16. VII. 2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 24. VII. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 3. VIII. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 11. VIII. 2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 10. VI. 2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 3. VIII. 2016, R. Turnbow leg. (1 ♀, FSCA). Jefferson County, Birmingham [33 ° 31 ’ N 86 ° 48 ’ W], black light trap, 28. IX. 1981, F. N. Young leg. (1 ♂, FSCA). Florida: Alachua County, (200 m) [29 ° 36 ’ N 82 ° 22 ’ W], 11. IV. 1992, J. Pickering leg. (2 ♀♀, FSCA); Alachua County, Highway 236, 2 km W Highway 241, 19. III. 1999, C. W. Mills leg. (1 ♀, FSCA); Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W], black light trap, 5. IV. 1983, N. Hostettler leg. (2 ♂♂, FSCA); Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W], black light trap,. IX. 2000, electric light fixture G. Buckingham leg. (1 ♀, FSCA); A Alachua County, Gainesville, San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 15. IX. 1988, M. C. Thomas leg. (1 ♀, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], 20. I. 2000, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], 4. III. 2000, P. E. Skelley leg. (2 ♂♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], live oak, black light, 10. II. 2008, P. E. Skelley leg. (1 ♀, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], black light, 24. III. 2008, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], flight intercept trap, 27. III. 2011, P. E. Skelley leg. (2 ♀♀, FSCA); Alachua County, Gainesville, 3517 NW 10 th Avenue, (30 m) [29 ° 39 ’ N 82 ° 22 ’ W], black light trap, 24. XI. 1972, R. E. Woodruff leg. (2 ♀, FSCA); Alachua County, Gainesville, 3517 NW 10 th Avenue, (30 m) [29 ° 39 ’ N 82 ° 22 ’ W], at light, 31. VIII. 2008, R. E. Woodruff leg. (1 ♀, FSCA); Alachua County, Gainesville, 610 NW 54 th Terrace, (30 m) [29 ° 39 ’ N 82 ° 24 ’ W], 25. VIII. 1994, L. Stange leg. (1 ♀, FSCA); Dade County, Camp Mahachee, South Miami, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], 10. IV. 1993, M. C. Thomas leg. (2 ♀♀ and 1 ♂, FSCA); Escambia County, Cantonment, 2666 Monica Lane, (40 m) [30 ° 33 ’ N 87 ° 18 ’ W], black light trap, 10. IV. 2009, T. Dobbs leg. (1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), 10. IV. 1991, R. Turnbow leg. (1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), 27. IV. 1991, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], black light trap, 5. V. 2008, M. C. & S. Thomas leg. (2 ♀♀ and 1 ♂, FSCA); Leon County, Tall Timbers Research Station, Woodyard Hammok, (60) [30 ° 39 ’ N 84 ° 12 ’ W], Berlese funnel, 24. XI. 1972, C. W. O’Brien leg. (1 ♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], black light trap, 3. V. 2003, M. C. Thomas leg. (3 ♀♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 6. V. 2007, M. C. Thomas leg. (2 ♀♀, FSCA); Liberty County, Aspalaga Landing, (20 m) [30 ° 37 ’ N 84 ° 54 ’ W], Malaise trap, 31. X. 2002, DJP leg. (2 ♀♀, FSCA); Pasco County, Green Swamp West, McNeil Trailhead [28 ° 18 ’ N 82 ° 03 ’ W], Hardwood / coniferous forest, Beaten from branches of hardwood tree, 15. I. 2014, J. P. Gruber leg. (1 ♂, WIRC); Hardwood / coniferous forest, Beaten from branches of hardwood tree, 16. I. 2014, J. P. Gruber leg. (2 ♀♀, WIRC); 17. I. 2014, J. P. Gruber leg. (1 ♀, WIRC); Pasco County, Green Swamp West, Main Entrance, (30 m) [28 ° 22 ’ 02 ” N 82 ° 07 ’ 04 ” W], Mixed hardwood / coniferous forest, Beaten from fan-like leaf of palm, 12. I. 2014, J. P. Gruber leg. (1 ♀, WIRC); Pasco County, Green Swamp West, Main Entrance, (30 m) [28 ° 22 ’ 02 ” N 82 ° 07 ’ 04 ” W], Mixed hardwood / coniferous forest, Beaten from branches of various hardwood trees, 12. I. 2014, J. P. Gruber leg. (5 ♀♀ and 1 ♂, WIRC); Pasco County, Upper Hillsborough Wildlife Management Area [28 ° 15 ’ 32 ” N 82 ° 06 ’ 50 ” W], Hardwood / coniferous forest, Beaten from branches of hardwood tree, 17. I. 2014, J. P. Gruber leg. (1 ♂, WIRC). Georgia: Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 14. IX. 1991, J. Pickering leg. (1 ♀, FSCA), 14. IX. 1991. Louisiana: Allen Parish, Whisky Chitto Creek, (25 m) [30 ° 41 ’ N 92 ° 52 ’ W], Wet-mesic hardwood forest, Ex. foliage of mixed hardwoods, along streamlet; bordering Pinus cutover, 7. XI. 2014, J. P. Gruber leg. (1 sp., NGPC); East Baton Rouge Parish, Baton Rouge, Place Du Plantier Apartments (5 m) [30 ° 23 ’ 55 ” N 91 ° 09 ’ 13 ” W], Berlese forest litter, 30. XII. 1993, E. G. Riley leg. (1 sp., TAMU); Rapides Parish, Indian Creek Reservoir, (25 m) [31 ° 08 ’ N 92 ° 28 ’ W], Dry-mesic open Pinus palustris forest, Ex. foliage of mixed hardwoods, 7. XI. 2014, J. P. Gruber leg. (2 sp., NGPC). Mississippi: Oktibbeha County, Osborn Prairies, (100 m) [33 ° 30 ’ N 88 ° 44 ’ W], 27. V. 2017, R. Turnbow leg. (1 ♀, FSCA). North Carolina: Haywood County, I- 40, N of Waynesville, (840 m) [35 ° 34 ’ N 82 ° 58 ’ W], 2. VII. 1985, H. V. Weems Jr. leg. (1 ♀, FSCA). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W], UV,. VIII. 1998, K. Stephan leg. (1 sp., TAMU). South Carolina: Berkeley County, Francis Marion National Forest, Guilliard Lake Recreational Arena, (10 m) [33 ° 17 ’ 03 ” N 79 ° 37 ’ 07 ” W], 27. V. 2006, E. G. Riley leg. (1 sp., TAMU). Texas: Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], 16. VII. 2016, at lights, V. Belov leg. (1 ♂, VBPC), 30. VII. 2016, at lights, V. Belov leg. (1 ♂, VBPC), 23. VIII. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 6. IX. 2016, at lights, V. Belov leg. (1 ♀, VBPC and 1 ♀, NGPC), 7. IX. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 8. IX. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 12. IX. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 17. IX. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 26. IX. 2016, at lights, V. Belov leg. (1 ♂, VBPC), 27. IX. 2016, at lights, V. Belov leg. (2 ♂♂, VBPC), 5. X. 2016, at lights, V. Belov leg. (1 ♀, 2 ♂♂ and 1 sp., VBPC), 6. X. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 14. X. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 16. X. 2016, at lights, V. Belov leg. (1 ♀, VBPC), 17. X. 2016, at lights, V. Belov leg. (2 ♂♂, VBPC), 21. X. 2016, at lights, V. Belov leg. (1 ♂, VBPC), 24. X. 2016, at lights, V. Belov leg. (1 ♀ and 3 ♂♂, VBPC), 31. X. 2016, at lights, V. Belov leg. (2 ♀♀ and 3 ♂♂, VBPC), 1. XI. 2016, at lights, V. Belov leg. (3 ♂♂, VBPC), 2. XI. 2016, at lights, V. Belov leg. (1 ♀ and 2 ♂♂, VBPC), 4. XI. 2016, at lights, V. Belov leg. (3 ♂♂, VBPC), 18. III. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 27. III. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 29. III. 2017, at lights, V. Belov leg. (1 ♂, VBPC), 13. IV. 2017, at lights, V. Belov leg. (2 ♂♂, VBPC), 14. IV. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 15. IV. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 16. IV. 2017, at lights, V. Belov leg. (2 ♀♀ and 1 ♂, VBPC), 18. IV. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 19. IV. 2017, at lights, V. Belov leg. (7 ♂♂, VBPC), 21. IV. 2017, at lights, V. Belov leg. (3 ♀♀ and 2 ♂♂, VBPC), 22. IV. 2017, backyard, vegetation, V. Belov leg. (1 ♀ and 1 sp., VBPC), 25. IV. 2017, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 1. V. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 13. V. 2017, at lights, V. Belov leg. (2 ♀♀ and 2 ♂♂, VBPC), 15. V. 2017, at lights, V. Belov leg. (1 ♂, VBPC), 19. V. 2017, at lights, V. Belov leg. (1 ♀, VBPC), 4. VI. 2017, at lights, V. Belov leg. (1 ♂, VBPC), 8. VI. 2017, at lights, V. Belov leg. (1 ♂, VBPC), 11. VI. 2017, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 20. VI. 2017, at lights, V. Belov leg. (2 ♀♀ and 1 ♂, VBPC), 26. VI. 2017, at lights, V. Belov leg. (1 ♂ and 2 sp., VBPC), 14. VII. 2017, at lights, V. Belov leg. (1 ♂, VBPC), 8. X. 2018, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 19. V. 2019, at lights, V. Belov leg. (1 ♀, VBPC), 26. V. 2019, at lights, V. Belov leg. (1 ♀, VBPC), 7. IV. 2020, at lights, V. Belov leg. (1 ♀, VBPC), 17. IV. 2020, at lights, V. Belov leg. (1 ♀, VBPC), 19. IV. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 20. IV. 2020 at lights, V. Belov leg. (1 ♂, VBPC), 24. IV. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 19. V. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 28. V. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 31. V. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 21. VI. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 26. VI. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 3. VII. 2020, at lights, V. Belov leg. (1 ♂, VBPC), 28. VII. 2020, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 17. IX. 2020, at lights, V. Belov leg. (1 ♀, VBPC); Brazos County, College Station, Riley Estate, (80 m) [30 ° 35 ’ N 96 ° 15 ’ W], 27. VII. 2003, E. G. Riley leg. (1 sp., TAMU), 19. IV. 2005, E. G. Riley leg. (1 ♀, TAMU); Brazos County, College Station, Lick Creek Park, (70 m) [30 ° 33 ’ 38 ” N 96 ° 13 ’ 03 ” W], 14. IV. 1996, E. G. Riley leg. (1 ♀, TAMU), 5. V. 2002, E. G. Riley leg. (1 ♀, TAMU); Brazos County, Bryan [30 ° 40 ’ N 96 ° 22 ’ W], taken at light, 11. X. 1965, J. C. Schaffner leg. (2 ♂♂, TAMU); Brazos County, Bryan [30 ° 40 ’ N 96 ° 22 ’ W], 13. VI. 1988, E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Ref., Site 8, (3 m) [25 ° 51 ’ 06 ” N 97 ° 25 ’ 13 ” W], Palm forest margin, Resaca bank, 20. IX. 2008, E. G. Riley leg. (2 ♀♀, ERPC); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25 ° 51 ’ 03 ” N 97 ° 25 ’ 04 ” W], 5. IV. 1987, E. G. Riley & Rider D. leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25 ° 51 ’ 03 ” N 97 ° 25 ’ 04 ” W], 20. X. 1990, T. Carlow & E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25 ° 51 ’ 03 ” N 97 ° 25 ’ 04 ” W], 23. IV. 1994, E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25 ° 51 ’ 03 ” N 97 ° 25 ’ 04 ” W], 4. XI. 2013, E. G. Riley & M. A. Quinn leg. (1 sp., TAMU); Fort Bend County, Brazos Bend State Park, Whiteoak Trail, (15 m) [29 ° 22 ’ 32 ” N 95 ° 34 ’ 38 ” W], Buckeye-sycamore forest, Beating / sweeping, 25. III. 2000, E. G. Riley leg. (1 sp., TAMU); Fort Bend County, Brazos Bend State Park, Whiteoak Trail, (15 m) [29 ° 22 ’ 32 ” N 95 ° 34 ’ 38 ” W], Hardwood forest, Beaten from vegetation, 18. IV. 2014, J. P. Gruber leg. (5 ♀♀ and 2 ♂♂, WIRC; 2 ♀♀ and 2 ♂♂, NGPC); Houston County, Crocket National Forest, Big Slough Wilderness Area, FR 511 & Four C’s trail, (70 m) [31 ° 28 ’ N 95 ° 08 ’ W], 15. III. 1996, E. G. Riley leg. (1 sp., TAMU); Montgomery County, The Woodlands [30 ° 10 ’ N 95 ° 30 ’ W], 25. V. 1978, J. S. Wappes leg. (3 ♂♂, FSCA); Sabine County, Sabine National Forest, 9 mi. E Hemphill, Beech Bottom [31 ° 19 ’ N 93 ° 42 ’ W], Beech-Magnolia forest, Malaise trap, 16. IV. 1989, R. Anderson & E. Morris leg. (1 sp.); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], Malaise trap, 15. III. 2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], UV light trap, 30. III. 2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], Lindgren funnel, 13. IV. 2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], UV light trap, 10. V. 2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31 ° 23 ’ 10 ” N 93 ° 42 ’ 29 ” W], UV light trap, 15. VI. 2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31 ° 16 ’ 42 ” N 93 ° 44 ’ 10 ” W], Mixed hardwood / coniferous forest, Beaten from branches of Quercus sp., 13. III. 2013, J. P. Gruber leg. (2 ♂♂ and 1 sp., NGPC); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31 ° 16 ’ 42 ” N 93 ° 44 ’ 10 ” W], Mixed hardwood / coniferous forest, Beaten from branches of Quercus sp., 14. III. 2013, J. P. Gruber leg. (4 ♀♀ and 2 ♂♂, WIRC); San Jacinto County, Sam Houston National Forest, Big Creek Scenic Area [30 ° 30 ’ 22 ” N 95 ° 05 ’ 20 ” W], Mixed hardwood / coniferous forest, Beaten from branches of hardwood tree (Quercus and Carya), 11. III. 2013, J. P. Gruber leg. (1 ♀ and 4 ♂♂, WIRC; 8 sp., NGPC); Tyler County, 5 mi. E Spurger, (20 m) [30 ° 41 ’ N 94 ° 05 ’ W], Flight intercept trap under pines, 23. III. 1997, Gillogly A. R. leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], Elevated flight intercept, 27. IV. 2003, E. G. Riley leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], Ground-level flight intercept trap, 27. IV. 2003, E. G. Riley leg. (3 sp., TAMU); Walker County, 4 mi. N Huntsville [30 ° 43 ’ N 95 ° 33 ’ W], at light, 30. IV. 1974, R. Phelps leg. (1 ♀, TAMU); Waller County, 15 mi. ENE of Hempstead, (80 m) [30 ° 10 ’ N 95 ° 50 ’ W], Beating, 22. IV. 1997, Chamberlain W. E. leg. (1 sp., TAMU).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73156FFC5DBD6FF2AFA5DFABC.taxon	distribution	Distribution. United States: Florida to eastern Oklahoma and Texas, north to North Carolina (Werner, 1990).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73150FFC4DBD6FA76FB50FE4C.taxon	description	(Figures 2 e; 3 e; 4 d; 5 e)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73150FFC4DBD6FA76FB50FE4C.taxon	materials_examined	Holotype: Texas, Belfrage (MCZC) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73150FFC4DBD6FA76FB50FE4C.taxon	materials_examined	Material examined. United States. Arkansas: Polk County, Mena, (350 m) [34 ° 35 ’ N 94 ° 14 ’ W], UV light traps, 19. VII. 2007, B. Baldwin leg. (1 ♂, UNHC); UV light traps, 6. IX. 2010, B. Baldwin leg. (1 ♀, UNHC). Missouri: Jackson County, Blue Springs [39 ° 01 ’ N 94 ° 16 ’ W], on Celtis occidentalis L., 5. VII. 1999, G. H. Nelson leg. (1 ♀, FSCA); Jackson County, Raytown [39 ° 00 ’ N 94 ° 27 ’ W], 21. VI. 1978, G. H. Nelson leg. (1 ♂, NGPC). Ohio: Cuyahoga County, Fairview Park, MacBeth Avenue, (220 m) [41 ° 25 ’ N 81 ° 52 ’ W], UV light, 22. VIII. 2007, H. J. Lee leg. (1 ♀, UNHC); Franklin County, Columbus [39 ° 57 ’ N 82 ° 59 ’ W] 28. VI. 1991, B. & B. Valentine leg. (1 ♀, FSCA); Franklin County, Columbus [39 ° 57 ’ N 82 ° 59 ’ W] 5. VII. 1991, B. & B. Valentine leg. (1 ♀, FSCA); Franklin County, Columbus [39 ° 57 ’ N 82 ° 59 ’ W] 8. VII. 1991, B. & B. Valentine leg. (1 ♀, NGPC); Franklin County, Columbus [39 ° 57 ’ N 82 ° 59 ’ W] 29. VIII. 2000, B. & B. Valentine leg. (1 ♀, FSCA). Texas: Comal County, Bulverde [29 ° 47 ’ N 98 ° 25 ’ W], mercury vapors + black light, 11. VI. 1998, R. Turnbow leg. (1 ♂, FSCA). Oklahoma: Grady County, Tuttle, Silver Ridge Drive, (390 m) [35 ° 19 ’ N 97 ° 44 ’ W] Blacklight trap, 20. V. 2005, B. Baldwin leg. (1 ♂, UNHC); UV light traps, 19. V. 2006, B. Baldwin leg. (3 ♂♂, UNHC). Texas: Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], At lights, 31. X. 2016, V. Belov leg. (1 ♂, VBPC); At lights, 4. IX. 2020, V. Belov leg. (1 ♂, VBPC); Wise County, Black Creek Reservoir, (300 m) [33 ° 20 ’ N 97 ° 35 ’ W], Dry oak savannah, UV sheet, Woodland with scattered red cedar, 15. IX. 2009, J. P. Gruber leg. (1 ♂, WIRC); Dry oak savannah-woodland, Attracted by UV light, 15. IX. 2009, J. P. Gruber leg. (1 ♀, NGPC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73150FFC4DBD6FA76FB50FE4C.taxon	distribution	Distribution. United States: Louisiana and Texas to Ohio and Illinois (Werner, 1990).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	description	(Figures 2 f; 3 f; 5 f)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	etymology	Etymology. This taxon is named after my good friend Jeffrey Gruber, who first collected it, in appreciation of his long-term support of my work and his generosity with sharing material.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	materials_examined	Type locality. United States: Arizona, Cochise County, Coronado NF: Huachuca Mountains, Upper Sonoran zone, Carr Canyon [31 ° 26 ’ 25 ” N 110 ° 17 ’ 07 ” W].	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	distribution	Distribution. United States: Arizona.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	description	Description. Body length. ♂: 1.93 ± 0.08 mm (n = 4) ♀: 2.21 ± 0.08 mm (n = 4). Body brown to orange, elytra progressively darker toward apex, pronotum and head darker, appendages entirely orange. Head. Transverse, dark-brown to black, slightly wider than pronotum. Frons convex. Palpi orange. Punctures sparse (spaces between punctures much wider than the puncture diameters), fine and superficial. Pubescence uniseriate, relatively long, spaced, hirsute, golden. Eyes convex, bulging, separated from the posterior edge of the head in side view by a distance the width of two rows of ommatidia. Eyes emarginate, with a deep anterior notch where the antennae are inserted, separated from each other by less than one eye diameter in front view in both sexes. Antennae. Thick, covered with thin and stout setae, 80 % the length of elytra in ♂♂, ~ 70 % in ♀♀, uniformely orange. Antennomeres 2 – 7 bead-shaped, longer than wide, antenommeres 8 – 10 weakly conical, somewhat elongated in ♂♂, subquadrate in ♀♀, regularly thicker toward apex. Last antennomere twice as long as penultimate, rounded. Pronotum. Dark orange to nearly black. Subquadrate in ♂♂, transverse in ♀♀, regularly convex, pentagonal, with a straight base flanked by rounded posterior angles, sides straight, hardly diverging from base to apex, and apex composed of two segments converging toward the head insertion in an obtuse rounded angle. Surface densely and coarsely punctate, the space between punctures equal to, or smaller than the puncture diameters, punctures large, deep. Pubescence uniseriate, raised, hirsute, golden. Elytra. Length. ♂: 1.39 ± 0.05 mm (n = 3) ♀: 1.60 ± 0.08 mm (n = 4). Elongate (width / length ♂: 0.52 ± 0.04 (n = 3) ♀: 0.56 ± 0.01 (n = 4), sides subparallel in both sexes, humeral angles well marked. Orange to pale brown, progressively infuscated in posterior half. Punctures dense, strong and deep, coarser than on the pronotum, space between punctures smaller than the puncture diameters. Disc weakly but regularly convex. Pubescence uniseriate, moderately long, raised, golden. Legs. Simple, inconspicuous, orange. Foretibia straight and simple in both sexes, with a barely perceptible kink at apex in ♂♂ (Figure 5 f). Aedeagus (Figure 3 f). Elongate, straight, sides subparallel in ventral view in the basal two thirds, then regularly tapered to the apex. Nearly straight along its dorsal side in lateral view. Apex terminated in a button in lateral view. Sexual dimorphism. Weak sexual differentiation. Eyes more separated in ♀♀ than in ♂♂. Antennae shorter in females.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73151FFC4DBD6FE66FC5DF87D.taxon	materials_examined	Type material. United States. Arizona: Cochise County, Coronado NF: Huachuca Mountains, Upper Sonoran zone, Carr Canyon (1670 m) [31 ° 26 ’ 25 ” N 110 ° 17 ’ 07 ” W], Evergreen oak-grassland habitat, Beaten from branches of hardwood tree, 9. VIII. 2013, J. P. Gruber leg. (1 ♂ holotype and 3 ♂♂ paratypes, NGPC); Evergreen oak-grassland habitat, at side of stream, Beaten from branches of Quercus sp. or other hardwood tree, 10. VIII. 2013, J. P. Gruber leg. (2 ♀♀ and 1 ♂ paratypes, WIRC; 1 ♀ paratype, NGPC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	description	(Figures 2 g; 3 g; 4 a; 5 g, q)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	etymology	Etymology. This taxon is named after its geographic origin.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	materials_examined	Type locality. United States: Arizona. Cochise County, Coronado National Forest, Chiricahua Mountains, 1 mi. N Rustler Park [31 ° 54 ’ 53 ” N 109 ° 16 ’ 07 ” W].	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	distribution	Distribution. United States: Arizona.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	description	Description. Body length. ♂ holotype: 1.50 mm. Body and appendages entirely orange-yellow. Head (Figure 4 a). Transverse, orange-yellow, slightly wider than pronotum. Frons relatively depressed (compared to S. belovi n. sp.). Palpi orange-yellow. Punctures sparse (spaces between punctures much wider than the puncture diameters), fine and shallow. Pubescence uniseriate, short, appressed, yellow, reclinate. Eyes small, convex, bulging, separated from the posterior edge of the head in side view by a distance half the eye diameter. Eyes nearly round, their outline hardly modified near the antennal insertions, separated from each other by three eye diameters in front view (Figure 4 a, double arrow). Antennae. Filiform, thinner than in S. belovi n. sp., covered with thin and stout setae, 75 % the length of elytra in the ♂ holotype, orange-yellow. Antennomeres 3 – 10 weakly conical, subquadrate, regularly thicker toward apex. Last antennomere twice as long as penultimate, rounded. Pronotum. Transverse, oval-shaped, posterior and anterior angles hard to discern, sides convex and rounded. Surface densely and coarsely punctate, the space between punctures much smaller than the puncture diameters, punctures large, deep. Pubescence uniseriate, short, appressed, yellow. Elytra. Length. ♂ holotype: 1.05 mm. Elongate (width / length ♂ holotype: 0.52), sides rounded, humeral angles well marked. Punctures dense, strong and deep, coarser than on the pronotum, space between punctures much smaller than the puncture diameters. Disc weakly but regularly convex. Pubescence uniseriate, short, appressed, yellow (Figure 5 q). Legs. Simple, inconspicuous, yellow. Foretibia of the ♂ holotype straight with an apical inward kink and an inner tooth. These are male-specific characters. Aedeagus (Figure 3 g). Elongate, straight, regularly thinner from base to apex. Apex briefly tapered, with a ventral sinuosity in lateral view. Dorsal side with two sinuosities in lateral view, and an elongated process protruding ventrally (contrary to S. belovi n. sp., Figure 3 h).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFC6DBD6FF2AFAFAFADC.taxon	materials_examined	Type material. United States. Arizona: Cochise County, Coronado National Forest, Chiricahua Mountains, 1 mile N Rustler Park, 21. VIII. 2000, B. Rodriguez Velez leg. (1 ♂ holotype, TAMU); Cochise County, Sunnyside, (1780 m) [31 ° 26 ’ N 110 ° 24 ’ W], 20. VIII. 1972, K. Stephan leg. (1 ♀ paratype, FSCA); Cochise County, Sunnyside, (1780 m) [31 ° 26 ’ N 110 ° 24 ’ W], 19. IX. 1972, (1 ♀ paratype, FSCA; 1 ♂ paratype, NGPC) K. Stephan leg.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	description	(Figures 2 h; 3 h; 4 b; 5 h, m, p)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	etymology	Etymology. This taxon is named after my colleague Vassili Belov, who provided most of the type series, in appreciation of his systematic efforts to collect Aderidae.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	materials_examined	Type locality. United States: Texas. Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ 08 ” N 96 ° 20 ’ 10 ” W].	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	distribution	Distribution. United States: Oklahoma, Texas, Florida.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	description	Description. Body length. ♂: 1.78 ± 0.11 mm (n = 3) ♀: 1.83 ± 0.09 mm (n = 5). Body and appendages entirely orange, head somewhat darker. Head (Figure 4 b). Transverse, dark-orange, slightly wider than pronotum. Frons convex. Palpi orange. Punctures sparse (spaces between punctures much wider than the puncture diameters), fine and deep. Pubescence uniseriate, long, very dense, white-golden, reclinate and converging toward the midline between the eyes. Eyes convex, bulging, separated from the posterior edge of the head in side view by a distance the width of three rows of ommatidia. Eyes emarginate, with a deep anterior notch where the antennae are inserted, separated from each other by nearly two eye diameters in front view (Figure 4 b, double arrow). Antennae. Thick, covered with thin and stout setae, 80 % the length of elytra in ♂♂, 60 % in ♀♀, orange. Antennomeres 3 – 10 weakly conical, subquadrate in ♂♂, transverse in ♀♀, regularly thicker toward apex. Last antennomere twice as long as penultimate, rounded. Pronotum. Transverse, regularly convex, pentagonal, with a straight base flanked by broadly rounded posterior angles, weakly diverging straight sides, and apex composed of two segments converging toward the head insertion in an obtuse rounded angle. Surface densely and coarsely punctate, the space between punctures equal to, or smaller than the puncture diameters, punctures large, deep. Pubescence uniseriate, long, raised, dense, golden. Elytra. Length. ♂: 1.21 ± 0.06 mm (n = 3) ♀: 1.32 ± 0.07 mm (n = 5). Elongate (width / length ♂: 0.55 ± 0.01 (n = 3) ♀: 0.52 ± 0.03 (n = 5), sides subparallel, humeral angles well marked. Punctures dense, strong and deep, coarser than on the pronotum, space between punctures smaller than the puncture diameters. Disc weakly but regularly convex. Pubescence uniseriate, long, dense, silky, raised, golden (Figure 5 p). Legs. Simple, inconspicuous, orange. Foretibia straight and simple in both sexes. Aedeagus (Figure 3 h). Elongate, straight, regularly thinner from base to apex. Apex abruptly truncated in ventral view. Somewhat bent along its dorsal side in lateral view, with a slight sinuosity before the base, and no distinct process protruding ventrally (contrary to S. arizonae n. sp., Figure 3 g). Sexual dimorphism. Eyes more separated in ♀♀ than in ♂♂. Antennae shorter with antennomeres 3 – 10 transverse in females (note the difference with S. pallidus (Werner, 1990), Figure 5 m, n and insets).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73153FFD9DBD6FAD4FA82FA59.taxon	materials_examined	Type material. United States. Florida: Alachua County, Gainesville, 3517 NW 10 th Avenue, (30 m) [29 ° 39 ’ N 82 ° 22 ’ W], at light, 6. V. 2007, R. E. Woodruff leg. (1 ♀ paratype, FSCA); Alachua County, Orange Heights, (50 m) [29 ° 43 ’ N 82 ° 08 ’ W], sticky wire trap,. XII. 1983, M. J. Plagens leg. (1 ♂ paratype, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 6. V. 2007, M. C. Thomas leg. (1 ♂ paratype, NGPC); Palm Beach County, Palm Beach, Loxahatchee, (10 m) [26 ° 46 ’ N 80 ° 14 ’ W], suction trap, 4. IV. 2006, S. Halbert leg. (1 ♀ paratype, FSCA). Oklahoma: Latimer County, 4 mi. W Red Oak, (180 m) [34 ° 56 ’ N 95 ° 09 ’ W], 4. VII. 1987, D. S. Chandler leg. (1 ♀ paratype, UNHC). Texas: Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 19. X. 2016, V. Belov leg. (1 ♀ paratype, NGPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 24. X. 2016, V. Belov leg. (2 ♀♀ paratypes, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 18. III. 2017, V. Belov leg. (2 ♀♀ paratypes, VBPC; 1 ♂, NGPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 19. IV. 2017, V. Belov leg. (1 ♀ paratype, NGPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 22. V. 2017, V. Belov leg. (1 ♀ paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 26. V. 2017, V. Belov leg. (1 ♀ paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 31. V. 2017, V. Belov leg. (1 ♀ paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 11. VI. 2017, V. Belov leg. (1 ♀ paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 20. VI. 2017, V. Belov leg. (1 ♂, paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 26. VI. 2017, V. Belov leg. (1 ♀ paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], found dead,. V. 2019, V. Belov leg. (1 ♂, paratype, VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 6. VI. 2019, V. Belov leg. (1 ♀ paratype, VBPC); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], Ground-level flight intercept trap, 27. IV. 2003, E. G. Riley leg. (1 ♀ paratype, ERPC); Walker County, Ellis Prison Unit, (60 m) [30 ° 52 ’ N 95 ° 27 ’ W], Suction trap, 24. V. 1980, D. A. Dean leg. (1 ♂ paratype, TAMU).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314CFFDBDBD6FA52FF24FF10.taxon	description	(Figures 1 g – i, 2 i; 3 i; 4 e; 5 i, n)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314CFFDBDBD6FA52FF24FF10.taxon	materials_examined	Holotype: Oklahoma: Latimer Co., 5 mi W Red Oak (USNM) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314CFFDBDBD6FA52FF24FF10.taxon	materials_examined	Material examined. United States. Florida: Alachua County, Orange Heights, (50 m) [29 ° 43 ’ N 82 ° 08 ’ W],. VII. 1983, sticky wire trap, M. J. Plagens leg. (1 ♂, FSCA). Georgia: Fulton County, College Park [33 ° 39 ’ N 84 ° 26 ’ W], 1. X. 1977, A. R. Gillogly leg. (1 ♀, TAMU). Missouri: Benton County, Warsaw [38 ° 14 ’ N 93 ° 22 ’ W], 5. VIII. 1971, ultraviolet light, G. H. Nelson leg. (1 ♀, FSCA); Greene County, 10 mi. E Springfield [37 ° 12 ’ N 93 ° 17 ’ W], 23. VIII. 1984, R. Haswell leg. (1 sp., TAMU). New Hampshire: Strafford County, Durham, 1 mi. SW Durham, (20 m) [43 ° 08 ’ N 70 ° 57 ’ 00 ” W], 26. V. 1987, Flight intercept trap, D. S. Chandler leg. (3 ♀♀, UNHC); Strafford County, Durham, 1 mi. SW Durham, (20 m) [43 ° 08 ’ N 70 ° 57 ’ 00 ” W], 10. VI. 1987, Flight intercept trap, D. S. Chandler leg. (2 ♀♀, UNHC). New York State: Madison County, Georgetown, Muller Hill, 100 Acre Evans camp, (610 m) [42 ° 46 ’ N 75 ° 46 ’ W], Barrier trap at top of Vista Meadow, 10. VI. 2002, M. H. Evans leg. (1 ♀, WIRC). Oklahoma: Grady County, Tuttle [35 ° 18 ’ N 97 ° 45 ’ W], Blacklight trap, 29. IV. 2004, B. Baldwin leg. (1 ♂, UNHC), 20. V. 2005, B. Baldwin leg. (1 ♀, UNHC), 11. VII. 2005, B. Baldwin leg. (1 ♂, UNHC); Grady County, Tuttle [35 ° 18 ’ N 97 ° 45 ’ W], UV light traps, 11. VII. 2005, B. Baldwin leg. (1 ♂, UNHC), 13. VII. 2005, B. Baldwin leg. (1 ♂, UNHC), 14. VII. 2005, B. Baldwin leg. (1 ♂, UNHC), 15. VII. 2005, B. Baldwin leg. (1 ♀, UNHC), 2. VIII. 2005, B. Baldwin leg. (1 ♀ and 1 ♂, UNHC), 6. VIII. 2005, B. Baldwin leg. (1 ♀, UNHC), 8. VIII. 2005, B. Baldwin leg. (1 ♀ and 1 ♂, UNHC), 9. VIII. 2005, B. Baldwin leg. (3 ♂♂, UNHC), 9. IX. 2005, B. Baldwin leg. (1 ♀, UNHC), 17. IX. 2005, B. Baldwin leg. (1 ♀, UNHC), 21. IX. 2005, B. Baldwin leg. (1 ♂, UNHC), 22. V. 2006, B. Baldwin leg. (1 ♂, UNHC), 29. VI. 2006, B. Baldwin leg. (3 ♂♂, UNHC), 30. VI. 2006, B. Baldwin leg. (1 ♂, UNHC), 1. VII. 2006, B. Baldwin leg. (3 ♀♀ and 1 ♂, UNHC), 2. VII. 2006, B. Baldwin leg. (1 ♂, UNHC), 3. VII. 2006, B. Baldwin leg. (1 ♀, UNHC); Grady County, Tuttle [35 ° 18 ’ N 97 ° 45 ’ W], Malaise trap, 9. V. 2006, B. Baldwin leg. (1 ♂, UNHC); Grady County, Tuttle [35 ° 18 ’ N 97 ° 45 ’ W], Lindgren traps, 2. V. 2007, B. Baldwin leg. (1 ♂, UNHC). Grady County, Tuttle, Silver Ridge Drive, (390 m) [35 ° 19 ’ N 97 ° 44 ’ W], Blacklight trap, 19. V. 2005, B. Baldwin leg. (2 ♀♀ and 3 ♂♂, UNHC), 20. V. 2005, B. Baldwin leg. (1 ♂, UNHC), 7. VI. 2005, B. Baldwin leg. (1 ♂, UNHC); Latimer County [34 ° 50 ’ N 95 ° 18 ’ W], UV,. VIII. 1998, K. Stephan leg. (2 sp., TAMU). Texas: Bastrop County, Bastrop State Park, (150 m) [30 ° 06 ’ N 97 ° 17 ’ W], 2. IV. 1990, E. G. Riley leg. (1 ♀ and 1 ♂, ERPC); Bastrop County, Bastrop State Park, (150 m) [30 ° 06 ’ N 97 ° 17 ’ W], UV light, 7. IV. 2000, E. G. Riley leg. (1 sp., TAMU); Bell County, 1 km W Youngsport, Bowmer Rch., Lampasas River, (200 m) [30 ° 57 ’ 43 ” N 97 ° 43 ’ 45 ” W], Flight intercept trap, 21. IV. 2010, Robbins T. leg. (1 sp., TAMU); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], at lights, 6. VIII. 2016, V. Belov leg. (1 ♂, VBPC), 17. IX. 2016, V. Belov leg. (1 sp., VBPC), 5. X. 2016, V. Belov leg. (1 sp., VBPC), 28. X. 2016, V. Belov leg. (1 sp., VBPC), 26. V. 2017, V. Belov leg. (1 sp., VBPC), 6. IV. 2020, V. Belov leg. (1 sp., VBPC), 7. IV. 2020, V. Belov leg. (1 sp., VBPC), 2. X. 2020, V. Belov leg. (1 sp., VBPC); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], found dead,. IX. 2016, V. Belov leg. (1 ♂, VBPC); Brazos County, Bryan [30 ° 40 ’ N 96 ° 22 ’ W], taken at light, 10. VI. 1970, J. C. Schaffner leg. (1 sp., TAMU); Brazos County, College Station, Riley Estate, (80 m) [30 ° 35 ’ N 96 ° 15 ’ W], UV, 14. IX. 2006, E. G. Riley leg. (1 sp., TAMU); Burnet County, Ca. 8 mi. WNW Burnet, Reveille Peak Ranch, (300 m) [30 ° 47 ’ 22 ” N 98 ° 21 ’ 18 ” W], UV light, 17. VII. 2003, J. D. Oswald leg. (1 sp., TAMU); Edwards County, 24 mi. S Junction Hwy. 377, (600 m) [30 ° 15 ’ N 99 ° 57 ’ W], UV light, 10. IV. 2002, E. G. Riley & M. J. Yoder leg. (1 sp., TAMU); Hays County, 6 mi. NW Dripping Springs, (400 m) [30 ° 13 ’ N 98 ° 11 ’ W], Juniperus unmanaged plot, Flight intercept trap, 27. IV. 2006, E. G. Riley leg. (1 sp., TAMU), 2. VI. 2006, E. G. Riley leg. (1 sp., TAMU), 1. IX. 2006, E. G. Riley leg. (1 sp., TAMU); Jeff Davis County, Davis Mounts Resort, Marqua Residence, (1770 m) [30 ° 37 ’ 42 ” N 104 ° 05 ’ 01 ” W], UV, 5. VII. 2009, E. G. Riley leg. (1 ♂, ERPC); Jeff Davis County, Davis Mounts State Park [30 ° 37 ’ N 104 ° 05 ’ W], UV, 24. VI. 1994, E. G. Riley leg. (1 ♂, ERPC); Kerr County, 6.5 mi. SW Hunt, (580 m) [29 ° 59 ’ N 99 ° 23 ’ W], Upland deciduous forest, Lindgren funnel trap, 12. X. 2005, E. G. Riley leg. (1 ♀, TAMU); Kerr County, 6.5 mi. SW Hunt, (580 m) [29 ° 59 ’ N 99 ° 23 ’ W], Upland deciduous forest, Pitfall trap, 27. IV. 2006, E. G. Riley leg. (1 ♀, TAMU); Kerr County, 6.5 mi. SW Hunt, (580 m) [29 ° 59 ’ N 99 ° 23 ’ W], Upland deciduous forest, Lindgren funnel trap, 30. IV. 2006, E. G. Riley leg. (1 ♀, TAMU), 27. VII. 2006, E. G. Riley leg. (1 ♀ and 1 ♂, TAMU); Lamar County, Lake Gibbons, Prairie area [33 ° 42 ’ N 95 ° 39 ’ W], Sweep native prairie, 9. IV. 2015, E. G. Riley leg. (1 ♀, ERPC); San Patricio County, Welder Wildlife Refuge, Headquarters, (10 m) [28 ° 06 ’ 45 ” N 97 ° 25 ’ 02 ” W], UV light, 1. XI. 2013, E. G. Riley & M. A. Quinn leg. (1 sp., TAMU); Travis County, vic. Long Hollow Creek, (280 m) [30 ° 27 ’ N 97 ° 52 ’ W], On Quercus virginiana, 2. IV. 1990, Quinn M. leg. (1 ♂, ERPC); Wise County, Black Creek Reservoir, (300 m) [33 ° 20 ’ N 97 ° 35 ’ W], Dry oak savannah, UV sheet, Woodland with scattered red cedar, 15. IX. 2009, J. P. Gruber leg. (7 ♀♀ and 3 ♂♂, WIRC); Wise County, Black Creek Reservoir, (300 m) [33 ° 20 ’ N 97 ° 35 ’ W], Dry oak savannah-woodland, Beaten from branches, morning, 16. IX. 2009, J. P. Gruber leg. (2 ♀♀ and 3 ♂♂, NGPC). Wisconsin: Dane County [43 ° 08 ’ N 89 ° 30 ’ W], 10. V. 1900, W. S. Marshall leg. (1 ♀, WIRC); Iron County, Frog Lake and Pines State Natural Area [46 ° 04 ’ 26 ” N 90 ° 00 ’ 16 ” W], Northern hardwoods, By open field, Malaise trap, 14. VIII. 2012, J. Janicki leg. (1 ♀, WIRC); Manitowoc County, Point Beach State Forest, Red Pine Trail [44 ° 12 ’ 52 ” N 87 ° 30 ’ 56 ” W], Mixed hardwood / coniferous forest, Beaten from branches of Quercus, Acer and Corylus, 2. IX. 2012, J. P. Gruber leg. (2 ♀♀ and 1 ♂, WIRC); Manitowoc County, Woodland Dunes State Natural Area [44 ° 08 ’ 19 ” N 87 ° 36 ’ 11 ” W], Mixed hardwood / coniferous forest, Beaten from interspersed branches of Acer sp. and Carya sp., 1. IX. 2012, J. P. Gruber leg. (1 ♀ and 1 ♂, WIRC); Oconto County, Hwy W, 12.2 km WSW of Mountain [45 ° 07 ’ N 88 ° 36 ’ W], Edge of mixed hardwood / coniferous forest, Beaten from branches of hardwood tree, 14. VIII. 2012, J. P. Gruber leg. (5 ♀♀ and 4 ♂♂, WIRC; 3 ♀♀ and 4 ♂♂, NGPC); Sheboygan County, Kohler-Andrae State Park, Woodland Dunes Nature Trail [43 ° 39 ’ 07 ” N 87 ° 43 ’ 38 ” W], Mixed hardwood / coniferous forest, Beaten from branches of hardwood tree, 13. VIII. 2012, J. P. Gruber leg. (23 ♀♀ and 9 ♂♂, WIRC; 1 ♂ and 8 sp., NGPC), 21. VIII. 2013, J. P. Gruber leg. (1 ♀, WIRC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314CFFDBDBD6FA52FF24FF10.taxon	distribution	Distribution. United States: Eastern Oklahoma and Texas to Florida, north to Ontario and Quebec (Werner, 1990).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314EFFDDDBD6FE9BFE5CFB01.taxon	description	(Figures 2 j; 3 j; 4 g; 5 j)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314EFFDDDBD6FE9BFE5CFB01.taxon	materials_examined	Lectotype: “ Middle States ” (MCZC) (Werner, 1990) Zonantes subfasciatus (LeConte, 1875): Werner (1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314EFFDDDBD6FE9BFE5CFB01.taxon	description	Note. The MCZC hosts a male of Syzeton subfasciatus (LeConte, 1875) labeled “ ♂ / Type 4886 / X. quercicola Schwarz / NOT = quercicola F Werner ‘ 90 ” and misidentified as Xylophilus quercicola Schwarz, 1878 (photos from this specimen are available from MCZBase: The Database of the Zoological Collections (https: // mczbase. mcz. harvard. edu )). The male lectotype of Xylophilus quercicola Schwarz, 1878, now Ariotus quercicola (Schwarz, 1878) is kept at the USMN (Werner, 1990). Schwarz ‘ s original description (Schwarz, 1878) leaves no doubt that the MCZC specimen is not Ariotus quercicola (Schwarz, 1878) and was mistakenly assigned to this taxon. It should not be considered part of the type material of Xylophilus quercicola Schwarz, 1878.	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314EFFDDDBD6FE9BFE5CFB01.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VIII. 2012, T. King leg. (1 ♀ and 2 ♂♂, FSCA); Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VIII. 2013, T. King leg. (2 ♀ and 4 ♂♂, FSCA); Covington County, Solon Dixon Forestry Education Center, (80 m) [31 ° 08 ’ N 86 ° 41 ’ W], 5. II. 2003, C. W. O’Brien leg. (1 ♂, FSCA); Dale County, Fort Rucker Military Resort, (110 m) [31 ° 19 ’ N 85 ° 43 ’ W], leaf litter, 31. III. 1993, R. Turnbow leg. (1 ♀, FSCA); Dale County, Fort Rucker Military Resort, (110 m) [31 ° 19 ’ N 85 ° 43 ’ W], Berlese beech litter, 27. X. 2001, O’Brien C. leg. (1 ♀, FSCA); Dale County, Fort Rucker Military Resort, (110 m) [31 ° 19 ’ N 85 ° 43 ’ W], leaf litter, 8. V. 2016, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], 30. VII. 2014, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 19. VIII. 2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 19. VIII. 2014, R. Turnbow leg. (3 ♂♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 8. IV. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 13. IV. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 30. VI. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], 3. VIII. 2015, R. Turnbow leg. (4 ♀♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], 10. VIII. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], 17. III. 2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], flight intercept trap, 22. IV. 2016, R. Turnbow leg. (2 ♂♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], 18. IV. 2018, R. Turnbow leg. (1 ♀, FSCA). Arkansas: Prairie County, Wattensaw Wildlife Management Area, 9.6 km NNE of Hazen [34 ° 51 ’ 39 ” N 91 ° 32 ’ 35 ” W], Forest / oldfield, Beaten from branches of hardwood tree, 22. VIII. 2010, J. P. Gruber leg. (2 ♂♂, WIRC); Sharp County, Harold E. Alexander WMA [36 ° 12 ’ 28 ” N 91 ° 26 ’ 34 ” W], Mixed hardwood forest, Beaten from branches of hardwood tree, 22. VIII. 2010, J. P. Gruber leg. (2 ♀♀, WIRC). Florida: Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W], sticky wire trap,. VII. 1984, M. J. Plagens leg. (1 ♂, FSCA); San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 13. III. 1985, M. C. Thomas leg. (2 ♂♂, FSCA); San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 3. IV. 1991, Mills C. W. & M. C. Thomas leg. (2 ♀♀ and 1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], flight intercept trap, 27. III. 2011, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, Hogtown Creek, (50 m) [29 ° 39 ’ N 82 ° 21 ’ W], light trap, 19. VI. 1988, P. E. Skelley leg. (1 ♀, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], black light, 1. IX. 1997, J. Cicero leg. (1 ♀, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30 ° 48 ’ N 85 ° 13 ’ W], 5. V. 2008, M. C. & S. Thomas leg. (2 ♀♀ and 2 ♂♂, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 31. III. 1985, M. C. Thomas leg. (1 ♂, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 4. VIII. 1996, black light, J. Cicero leg. (1 ♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], 31. III. 1985, M. C. Thomas leg. (3 ♀♀ and 1 ♂, FSCA); Walton County, Eglin AFB, Alaqua-Blount Creek confluence [30 ° 37 ’ N 86 ° 09 ’ W], beech magnolia leaf litter, 21. XI. 1998, P. W. Kovarik leg. (1 ♀, FSCA); Walton County, Eglin AFB, 5.1 mi. W Highway 331 on Range Road 205 [30 ° 30 ’ N 86 ° 14 ’ W], black light trap, 17. VI. 1995, P. E. Skelley leg. (1 ♀, FSCA). Georgia: Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 15. IV. 1992, J. Pickering leg. (1 ♂, FSCA); Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 22. IV. 1992, J. Pickering leg. (1 ♀, FSCA); Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 3. V. 1992, J. Pickering leg. (1 ♂, FSCA); Clarke County [33 ° 54 ’ N 83 ° 16 ’ W], 12. VIII. 1992, J. Pickering leg. (1 ♂, FSCA). Illinois: Jackson County, Shawnee National Forest, 7.8 km NW of Pomona [37 ° 40 ’ 50 ” N 89 ° 23 ’ 42 ” W], Upland hardwood forest, Beaten from branches of hardwood tree, 23. VIII. 2010, J. P. Gruber leg. (3 ♀♀ and 2 ♂♂, WIRC); Jackson County, Shawnee National Forest, 7.8 km NW of Pomona [37 ° 40 ’ 50 ” N 89 ° 23 ’ 42 ” W], Upland hardwood forest, Beaten from Carya sp. sapling, 16. VII. 2011, J. P. Gruber leg. (2 ♀♀ and 2 ♂♂, WIRC). Indiana: Montgomery County, Shades State Park [39 ° 55 ’ N 87 ° 04 ’ W], Mixed hardwood forest, Beaten from branches of hardwood tree, 24. V. 2013, J. P. Gruber leg. (5 ♀♀ and 1 ♂, WIRC); Morgan County, Catholic Cemetery Road, 5 mi. south of Martinsville, (220 m) [39 ° 21 ’ 31 ” N 86 ° 24 ’ 57 ” W], Medium-aged oak / hickory forest, Beaten from branches of hardwood tree, 26. V. 2013, J. P. Gruber leg. (2 ♀ and 4 ♂♂, WIRC). Louisiana: West Feliciana Parish, Nature Conservancy Easement, 15 mi. E of St. Francisville [30 ° 47 ’ 40 ” N 91 ° 15 ’ 15 ” W], Woods near pond, UV, 30. III. 2015, M. H. Evans leg. (2 ♀♀, WIRC). Maryland: Prince George’s County [39 ° 03 ’ N 76 ° 47 ’ W], 27. IV. 1992, J. Pickering leg. (1 ♀, FSCA); Prince George’s County [39 ° 03 ’ N 76 ° 47 ’ W], 4. V. 1992, J. Pickering leg. (1 ♂, FSCA); Prince George’s County [39 ° 03 ’ N 76 ° 47 ’ W], 11. V. 1992, J. Pickering leg. (1 ♂, FSCA). Mississippi: Alcorn County, Corinth [34 ° 56 ’ N 88 ° 31 ’ W], 25. V. 2008, R. Turnbow leg. (1 ♀, FSCA); Clay County, West Point [33 ° 36 ’ N 88 ° 39 ’ W], black light trap, 11. VI. 2016, R. Turnbow leg. (1 ♂, FSCA); Lowndes County, Highway 182 at Catalpa Creek, (60 m) [33 ° 28 ’ N 88 ° 37 ’ W], 11. VI. 2016, M. C. Thomas leg. (1 ♀ and 1 ♂, FSCA). Missouri: Phelps County, Mark Twain National Forest, Mill Creek Recreational Area, 16.4 km WSW of Rolla [37 ° 52 ’ 23 ” N 91 ° 55 ’ 46 ” W], Hardwood forest, near creek, at blacklight, night, 20. VIII. 2010, K. Katovich & J. P. Gruber leg. (1 ♀, WIRC); Shannon County, Mark Twain National Forest, 8.4 km S of Birch Tree, (220 m) [36 ° 55 ’ 01 ” N 91 ° 30 ’ 33 ” W], Pine-oak woodland, Beaten from branches of hardwood tree, 23. VIII. 2010, J. P. Gruber leg. (1 ♀ and 2 ♂♂, WIRC); Taney County, Branson, 5751 State Highway, (290 m) [36 ° 35 ’ N 95 ° 18 ’ W], on a tree, foliage, 27. V. 2018, N. Gompel leg. (1 ♂, NGPC). New Jersey: Essex County [40 ° 44 ’ N 74 ° 16 ’ W], .. 2010, V. Belov leg. (1 sp., VBPC); Essex County, Maplewood [40 ° 44 ’ N 74 ° 16 ’ W], Yard lights, .. 2013, V. Belov leg. (1 sp., VBPC). Ohio: Licking County, Blackhand Gorge State Nature Preserve, (230 m) [40 ° 03 ’ 21 ” N 82 ° 13 ’ 05 ” W], Under Liriodendron bark, 4. VI. 1998, P. W. Kovarik leg. (1 ♂, TAMU). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W],. X. 2002, K. Stephan leg. (2 sp., TAMU),. XI. 2002, K. Stephan leg. (2 sp., TAMU). North Carolina: Henderson County, Fletcher [35 ° 25 ’ N 82 ° 30 ’ W], 17. VI. 1970, L. L. Lampert leg. (1 ♀, FSCA). South Carolina: Chesterfield County, Sand Hills State Forest, 6.5 km WNW of of Patrick [34 ° 35 ’ N 80 ° 06 ’ W], Mixed coniferous / hardwood forest, Beaten from branches of Cornus florida L., 22. VIII. 2011, J. P. Gruber leg. (1 ♂, NGPC); Chesterfield County, Sand Hills State Forest, 6.5 km WNW of of Patrick [34 ° 35 ’ N 80 ° 06 ’ W], Mixed coniferous / hardwood forest, Beaten from branches of Carya sp., 25. VIII. 2011, J. P. Gruber leg. (3 ♀♀ and 3 ♂♂, WIRC). Tennessee: Blount County, Great Smoky Mountains National Park, Cades Cove area, (560 m) [35 ° 36 ’ 00 ” N 83 ° 50 ’ 42 ” W], UV, 28. VI. 2001, E. G. Riley leg. (1 sp., TAMU). Texas: Brazos County, San Bernard National Wildlife Refuge, Palm Unit, 5 mi. S Sweeny, (5 m) [28 ° 58 ’ 26 ” N 95 ° 40 ’ 45 ” W], 30. III. 2007, E. G. Riley leg. (1 sp., TAMU); Brazos County, 5 km SW Wellborn, (80 m) [30 ° 30 ’ N 96 ° 20 ’ W], 18. III. 2017, at lights, V. Belov leg. (1 sp., VBPC); Brazos County, College Station, Lick Creek Park, (70 m) [30 ° 33 ’ 38 ” N 96 ° 13 ’ 03 ” W], 21. V. 2004, E. G. Riley leg. (1 sp., TAMU); Brazos County, College Station, Lick Creek Park (site 1), (70 m) [30 ° 33 ’ 30 ” N 96 ° 12 ’ 31 ” W], Bottomland forest, Berlese litter, 20. III. 202, E. G. Riley leg. (1 ♀ and 2 ♂♂, NGPC); Cameron County, Sabal Palm Grove [25 ° 50 ’ N 97 ° 25 ’ W], 24. X. 1992, E. G. Riley leg. (1 ♀, ERPC); Cameron County, Sabal Palm Grove Ref., Site 1, (8 m) [25 ° 50 ’ 53 ” N 97 ° 25 ’ 08 ” W], Palm forest, 16. IX. 2008, E. G. Riley leg. (1 sp., ERPC); Cameron County, Sabal Palm Grove Ref., Site 8, (3 m) [25 ° 51 ’ 06 ” N 97 ° 25 ’ 13 ” W], Palm forest margin, Resaca bank, 18. X. 2008, E. G. Riley leg. (1 sp., ERPC); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25 ° 51 ’ 03 ” N 97 ° 25 ’ 04 ” W], 10. X. 1994, E. G. Riley leg. (1 ♂, ERPC); Fort Bend County, Brazos Bend State Park, Whiteoak Trail, (15 m) [29 ° 22 ’ 32 ” N 95 ° 34 ’ 38 ” W], Hardwood forest, Beaten from vegetation, 18. IV. 2014, J. P. Gruber leg. (7 ♂♂, WIRC; 2 ♂♂ and 2 ♀♀, NGPC); Sabine County, Sabine National Forest, 9 mi. E Hemphill, Beech Bottom [31 ° 19 ’ N 93 ° 42 ’ W], Flight intercept trap, 16. IV. 1989, R. Anderson & E. Morris leg. (2 sp, TAMU); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31 ° 16 ’ 42 ” N 93 ° 44 ’ 10 ” W], Mixed hardwood / coniferous forest, Beaten from branches of hardwood tree, 14. III. 2013, J. P. Gruber leg. (1 ♂, NGPC); San Jacinto County, Sam Houston National Forest, Double Lake [30 ° 32 ’ 56 ” N 95 ° 07 ’ 53 ” W], at light, 29. VII. 1991, W. F. Chamberlain leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30 ° 34 ’ N 94 ° 25 ’ W], Ground-level flight intercept trap, 27. IV. 2003, E. G. Riley leg. (1 ♂, ERPC), 8. VI. 2003, E. G. Riley leg. (1 ♀, ERPC). Virginia: Montgomery County, Caldwell Fields, Craig Creek Road, (520 m) [37 ° 20 ’ 14 ” N 80 ° 19 ’ 35 ” W], taken at light, 23. V. 2006, E. G. Riley leg. (1 sp., TAMU); Montgomery County, Caldwell Fields, Craig Creek Road, (520 m) [37 ° 20 ’ N 80 ° 19 ’ W], 24. V. 2006, M. C. Thomas leg. (1 ♀, FSCA). Wisconsin: Eau Claire County, Eau Claire County Forest [44 ° 41 ’ N 90 ° 56 ’ W], Dry-mesic northern hardwood forest, Flight intercept trap, 3. IX. 2001, J. P. Gruber leg. (1 ♂, WIRC); Grant County, Cassville Bluffs SNA [42 ° 42 ’ N 90 ° 56 ’ W], Mixed hardwood forest, Swept from foliage near path, 18. VII. 2003, J. P. Gruber leg. (1 sp., NGPC); Grant County, Blue River Sand Barrens State Natural Area [43 ° 11 ’ 32 ” N 90 ° 31 ’ 49 ” W], Sandy oak barrens, Flight intercept trap, 9. VI. 2000, J. P. Gruber leg. (1 ♀, WIRC); Grant County, Blue River Sand Barrens State Natural Area [43 ° 11 ’ 32 ” N 90 ° 31 ’ 49 ” W], Sandy oak barrens, Beaten from branches of Quercus sp. tree, 23. VI. 2012, J. P. Gruber leg. (1 ♀, WIRC); Green County, Gasner Hollow Prairie SNA, (290 m) [42 ° 55 ’ N 91 ° 07 ’ W], Prairie / savanna, Swept from foliage near path, 29. VI. 2003, J. P. Gruber leg. (1 sp., NGPC); Iowa County, Arena Pines-Sand Barrens State Natural Area, (220 m) [43 ° 10 ’ 33 ” N 89 ° 55 ’ 53 ” W], Sand barrens vegetated with jack pine, Beaten from branches of living Quercus sp. tree, 12. VI. 2012, J. P. Gruber leg. (1 ♀, WIRC); Lafayette County, about 3 miles north of Argyle, Wedel property [42 ° 44 ’ 17 ” N 89 ° 51 ’ 05 ” W], Southern mixed hardwood forest, Flight intercept trap, 27. V. 2001, J. P. Gruber leg. (1 ♀ and 2 ♂♂, WIRC; 6 sp., NGPC), 10. VI. 2001, J. P. Gruber leg. (3 sp., NGPC), 24. VI. 2001, J. P. Gruber leg. (1 sp., NGPC), 11. VI. 2001, J. P. Gruber leg. (2 ♀♀ and 2 ♂♂, WIRC), 3. IX. 2001, J. P. Gruber leg. (1 ♀ and 1 ♂, WIRC), 9. XII. 2001, J. P. Gruber leg. (1 ♂, WIRC); Lafayette County, about 3 miles north of Argyle, Wedel property [42 ° 44 ’ 17 ” N 89 ° 51 ’ 05 ” W], Southern mixed deciduous hardwood forest, Unbaited Townes malaise trap, 5. VIII. 2001, C. M. Brabant leg. (2 ♀♀, WIRC); Lafayette County, about 3 miles north of Argyle, Wedel property [42 ° 44 ’ 17 ” N 89 ° 51 ’ 05 ” W], Beating a dead oak, 1. IX. 2003, N. Gompel leg. (4 sp., NGPC); Sauk County, Spring Green Preserve SNA, (220 m) [43 ° 11 ’ N 90 ° 03 ’ W], Dry-mesic southern forest, Beaten from branches of Quercus sp. tree, 1. X. 2010, J. P. Gruber leg. (1 ♀, WIRC); Sauk County, Spring Green Preserve SNA, (220 m) [43 ° 11 ’ N 90 ° 03 ’ W], Dry-mesic southern forest, Found on underside of leaf on branch of Carya ovata tree, 12. VI. 2012, J. P. Gruber leg. (2 ♂♂, WIRC); Sauk County, Spring Green Preserve SNA, (220 m) [43 ° 11 ’ N 90 ° 03 ’ W], Dry-mesic southern forest, Beaten from branches of Cornus sp. tree, 22. VII. 2012, J. P. Gruber leg. (1 ♀ and 1 ♂, WIRC); Shawano County, Jung Hemlock-Beech Forest State Natural Area [44 ° 48 ’ 39 ” N 88 ° 46 ’ 14 ” W], Old-growth northern mesic forest, Beaten from branches of Acer sp. tree, 17. VIII. 2012, J. P. Gruber leg. (1 ♀, WIRC); Vernon County, Tunnelville Cliffs State Natural Area [43 ° 34 ’ 17 ” N 90 ° 41 ’ 33 ” W], Mixed coniferous / hardwood forest, Beaten from branches of Carya sp., 19. VII. 2014, J. P. Gruber leg. (1 ♀, WIRC); Waukesha County, Kettle Moraine State Forest, Southern Unit [42 ° 52 ’ 21 ” N 88 ° 30 ’ 50 ” W], Sandy oak barrens, Unbaited Townes malaise trap, 15. VII. 2001, C. M. Brabant leg. (1 ♂, WIRC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314EFFDDDBD6FE9BFE5CFB01.taxon	distribution	Distribution. United States: From Florida to eastern Texas and Oklahoma, north to New Hampshire and Wisconsin (Werner, 1990).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73148FFDCDBD6FAAAFC46FB9D.taxon	description	(Figures 1 c; 2 k; 3 k; 4 c; 5 k)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73148FFDCDBD6FAAAFC46FB9D.taxon	materials_examined	Holotype: Florida, Dade County, Camp Mahachee, nr. Matheson Hammock (FSCA) (Werner, 1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73148FFDCDBD6FAAAFC46FB9D.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 21. VII. 2013, T. King leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 17. III. 2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], flight intercept trap, 22. IV. 2016, R. Turnbow leg. (1 ♂, FSCA). Florida: Alachua County, (200 m) [29 ° 36 ’ N 82 ° 22 ’ W], 28. VIII. 1992, J. Pickering leg. (1 ♀, FSCA); Alachua County, West of Gainesville, Pierce’s homestead [29 ° 39 ’ N 82 ° 19 ’ W], Malaise, 4. I. 1974, W. H. Pierce leg. (1 ♀, FSCA); Alachua County, SE of Gainesville, Kincaid Road, 1 mi. N Paynes [29 ° 38 ’ N 82 ° 17 ’ W], old field / dry oak hammock, Malaise, 10. VII. 1999, B. D. Sutton leg. (1 ♂, FSCA); Alachua County, Gainesville, San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 15. IX. 1988, M. C. Thomas leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], 26. II. 2000, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29 ° 37 ’ N 82 ° 20 ’ W], 20. III. 2008, P. E. Skelley leg. (1 ♀, FSCA); Collier County, Immokalee, Southwest Florida Research and Education, (10 m.) [26 ° 27 ’ N 82 ° 26 ’ W], suction trap, 20. IX. 2007, S. Halbert leg. (1 ♂, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 5. V. 1983, M. C. Thomas & L. Parker leg. (1 ♀, FSCA); Dade County, Camp Mahachee, Near Matheson Hammock, (5 m) [25 ° 40 ’ N 80 ° 16 ’ W], black light trap, 23. VIII. 1983, M. C. Thomas & L. Parker leg. (1 ♀, FSCA); Escambia County, Cantonment, 2666 Monica Lane, (40 m) [30 ° 33 ’ N 87 ° 18 ’ W], black light trap, 10. IV. 2009, T. Dobbs leg. (1 ♂, FSCA); Franklin County, 2 mi. NW Ochlockonee Bay, (2 m) [30 ° 00 ’ N 84 ° 25 ’ W], 2. XII. 2004, C. W. O’Brien & M. Haseeb leg. (1 ♀, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), 27. IV. 1991, R. Turnbow leg. (4 ♀♀ and 1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), beating vegetation, 15. IV. 2005, R. Turnbow leg. (1 ♂, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], black light trap, 3. V. 2003, M. C. Thomas leg. (1 ♀, FSCA); Liberty County, Torreya State Park, (40 m) [30 ° 33 ’ N 84 ° 56 ’ W], black light trap, 6. V. 2007, M. C. Thomas leg. (1 ♀, FSCA); Marion County, 2.5 mi. SSE Micanopy, (40 m) [29 ° 29 ’ N 82 ° 14 ’ W], Malaise trap, 2. IV. 1989, D. Wahl D. B. leg. (1 ♀, ERPC); Monroe County, Upper Key Largo, (40 m) [25 ° 05 ’ N 80 ° 26 ’ W], 30. V. 1997, M. C. Thomas leg. (1 ♂, FSCA); Monroe County, Key Largo, John Pennekamp Coral Reef State Park [25 ° 07 ’ N 80 ° 24 ’ W], black light trap, 29. V. 1997, P. E. Skelley leg. (6 ♂♂, FSCA); Pasco County, Green Swamp West, McNeil Trailhead [28 ° 18 ’ N 82 ° 03 ’ W], Grassland / savanna with oaks and palms, Beaten from branches of Quercus sp., 12. I. 2014, J. P. Gruber leg. (4 ♀♀ and 3 ♂♂, NGPC); Hardwood / coniferous forest, Beaten from branches of hardwood tree, 15. I. 2014, J. P. Gruber leg. (4 ♀♀ and 5 ♂♂, WIRC; 7 sp. and 1 ♂, NGPC); 17. I. 2014, J. P. Gruber leg. (2 ♀♀ and 4 ♂♂, WIRC); County, Green Swamp West, Cumpressco Entrance, (30 m), Mixed hardwood / coniferous forest, Beaten from branches of Quercus sp. tree, 13. I. 2014, J. P. Gruber leg. (2 ♀♀ and 1 6, WIRC); Pasco County, Green Swamp West, Main Entrance, (30 m) [28 ° 22 ’ 02 ” N 82 ° 07 ’ 04 ” W], Mixed hardwood / coniferous forest, Beaten from fan-like leaf of palm, 12. I. 2014, J. P. Gruber leg. (16 ♀♀ and 16 ♂♂, WIRC); Polk County, Winter Haven [28 ° 01 ’ N 81 ° 43 ’ W], suction trap, 25. II. 2005 (1 ♂, FSCA). Georgia: Thomas County, Wade Forest Tract, 10 km S Thomasville, (75 m) [30 ° 45 ’ N 84 ° 00 ’ W], open pine forest, flight interception trap, 27. IV. 2010, S. Peck leg. (1 ♀, FSCA); Thomas County, Wade Forest Tract, 10 km S Thomasville, (75 m) [30 ° 45 ’ N 84 ° 00 ’ W], open pine forest, Malaise trap, 31. IV. 2010, S. Peck leg. (1 ♀, FSCA). Louisiana: Allen Parish, Whisky Chitto Creek, (25 m) [30 ° 41 ’ N 92 ° 52 ’ W], Wet-mesic hardwood forest, Ex. foliage of mixed hardwoods, Along streamlet; bordering Pinus cutover, 7. XI. 2014, J. P. Gruber leg. (1 ♀, NGPC). Texas: Sabine County, Sabine National Forest, 9 mi. E Hemphill, Beech Bottom [31 ° 19 ’ N 93 ° 42 ’ W], Flight intercept trap, 2. IV. 1989, Anderson leg. (1 ♀, ERPC). Mississippi: Lee County, Highway 45 at Brewer road, (80 m) [34 ° 08 ’ N 88 ° 42 ’ W], 15. V. 2008, R. Turnbow leg. (2 ♀, FSCA).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D73148FFDCDBD6FAAAFC46FB9D.taxon	distribution	Distribution. United States: Texas to Florida (Werner, 1990).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314BFFDEDBD6FF2AFF37FA57.taxon	description	(Figures 2 l; 3 l; 4 f; 5 l) Euglenes signatus Haldeman, 1848: 97;	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314BFFDEDBD6FF2AFF37FA57.taxon	materials_examined	Lectotype: “ New York, Carolina ” (MCZC) (Werner, 1990) Zonantes signatus (Haldeman, 1848): Werner (1990)	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314BFFDEDBD6FF2AFF37FA57.taxon	materials_examined	Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33 ° 53 ’ N 86 ° 25 ’ W], light trap, 29. VII. 2012, T. King leg. (3 ♀♀, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 6. IV. 2015, R. Turnbow leg. (2 ♂♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 24. VI. 2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 17. III. 2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 13. V. 2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31 ° 17 ’ N 85 ° 46 ’ W], black light trap, 17. VII. 2016, R. Turnbow leg. (1 ♀, FSCA). Florida: Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W], black light trap, 3. XII. 1978, M. C. Thomas leg. (1 ♂, FSCA); Alachua County, Gainesville [29 ° 39 ’ N 82 ° 19 ’ W], black light trap,. VII. 1984, M. J. Plagens leg. (1 ♀, FSCA); Alachua County, Gainesville, San Felasco Hammock Preserve State [29 ° 42 ’ N 82 ° 27 ’ W], 15. IX. 1988, M. C. Thomas leg. (1 ♂, FSCA); Columbia County, O’Leno State Park, (20 m) [29 ° 55 ’ N 82 ° 36 ’ W], 28. IV. 1991, R. Turnbow leg. (1 ♂, FSCA); Leon County, Woodville, Natural Wells Road [30 ° 17 ’ N 84 ° 14 ’ W], 26. III. 2015, M. Haseeb & G. B. Marshall leg. (1 ♂, FSCA). Louisiana: West Feliciana Parish, Nature Conservancy Easement, 15 mi. E of St. Francisville [30 ° 47 ’ 40 ” N 91 ° 15 ’ 15 ” W], Woods near pond, UV, 30. III. 2015, M. H. Evans leg. (2 ♀♀ and 1 ♂, WIRC). Missouri: Shannon County, Mark Twain National Forest, 8.4 km S of Birch Tree, (220 m) [36 ° 55 ’ 01 ” N 91 ° 30 ’ 33 ” W], Pine-oak woodland, Beaten from branches of hardwood tree, 23. VIII. 2010, J. P. Gruber leg. (1 ♂, WIRC). Oklahoma: Latimer County [34 ° 50 ’ N 95 ° 18 ’ W],. X. 2002, K. Stephan leg. (1 sp., TAMU). South Carolina: Chesterfield County, Sand Hills State Forest, 6.5 km WNW of Patrick, (100 m) [34 ° 35 ’ 12 ” N 80 ° 06 ’ 56 ” W], Mixed coniferous / hardwood forest, Beaten from branches of Cornus florida, 22. VIII. 2011, J. P. Gruber leg. (2 ♂♂, WIRC); Chesterfield County, Sand Hills State Forest, 9.6 km East of Patrick, Parking area of Middendorf Pond [34 ° 32 ’ 39 ” N 80 ° 08 ’ 37 ” W], Forest edge, Beaten from branches of hardwood tree, 22. VIII. 2011, J. P. Gruber leg. (1 sp., NGPC). Texas: Brazos County, College Station, Lick Creek Park, (70 m) [30 ° 33 ’ 38 ” N 96 ° 13 ’ 03 ” W], 21. V. 2004, E. G. Riley leg. (6 sp., TAMU); Hardin County, Roy E. Larsen Preserve, (20 m) [30 ° 20 ’ N 94 ° 14 ’ W], beating burned schrub, 19. IV. 1980, J. E. Wappes leg. (1 ♀, FSCA); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31 ° 16 ’ 42 ” N 93 ° 44 ’ 10 ” W], Mixed hardwood / coniferous forest, Beaten from branches of Quercus sp., 13. III. 2013, J. P. Gruber leg. (1 ♂, WIRC; 2 sp., NGPC), 14. III. 2013, J. P. Gruber leg. (1 ♀ and 3 ♂♂, WIRC); San Jacinto County, Sam Houston National Forest, Big Creek Scenic Area [30 ° 30 ’ 22 ” N 95 ° 05 ’ 20 ” W], Mixed hardwood / coniferous forest, Beaten from branches of hardwood tree, 11. III. 2013, J. P. Gruber leg. (2 and 10 ♂♂, WIRC; 8 sp., NGPC).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
03E187D7314BFFDEDBD6FF2AFF37FA57.taxon	distribution	Distribution. United States: Florida to Texas, north to North Carolina and Missouri (extended from Werner (1990 )).	en	Gompel, Nicolas (2022): The genus Syzeton Blackburn, 1891 (Coleoptera, Aderidae) in the United States of America. Zootaxa 5162 (3): 243-267, DOI: 10.11646/zootaxa.5162.3.3
