identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E187FC3616494BFF5FFCECFBBAFE93.text	03E187FC3616494BFF5FFCECFBBAFE93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asthenodipsas stuebingi Quah & Grismer & Lim & Anuar & Imbun 2019	<div><p>Asthenodipsas stuebingi sp. nov.</p><p>Sabah Mountain Smooth Slug Snake</p><p>Figs. 2 &amp; 3.</p><p>urn:lsid:zoobank.org:act: 99B2763A-B16E-4320-8176-FC7E311305BA</p><p>Amblycephalus laevis (in part): Boulenger 1896: 441 &amp; 442; de Rooij 1917: 276.</p><p>Amblycephalus vertebralis (in part): Loveridge 1938: 43.</p><p>Asthenodipsas laevis (in part): Das 2006: 007, 2010: 344; Grossmann &amp; Tillack 2003: 188; Stuebing et al. 2014: 85.</p><p>Internatus laevis: Malkmus et al. 2002: 342 &amp; 343, Figs, 337 &amp; 338.</p><p>Pareas laevis: Malkmus 1996: 293; Manthey &amp; Grossmann 1997: 377, Abb. 284; Stuebing 1991: 330 &amp; 331 (in part); Stuebing &amp; Inger 1999: 87 (in part); Haile 1958: 766 (in part).</p><p>Holotype. Adult female, SP 04679 collected by Paul Yambun Imbun, Fred Tuh Yit Yu and Safrie on 25 March 2009 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.34724&amp;materialsCitation.latitude=5.82324" title="Search Plazi for locations around (long 116.34724/lat 5.82324)">Minduk Sirung</a> (Alab–Mahua trail), Crocker Range Park, Sabah, East Malaysia (estimated: N 5.823240, E 116.347238, 1859 m a.s.l.).</p><p>Paratypes. Adult male (SP 04806) collected by Siti Azizah Kusop, Junaidi Ais and R., Adzmie on 26 October 2014 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.57016&amp;materialsCitation.latitude=5.988087" title="Search Plazi for locations around (long 116.57016/lat 5.988087)">Kampung Desa Aman</a>, Kundasang (estimated: N5.988087, E 116.570162, 1341 m a.s.l.) . Adult female (ZMB 65429) collected by Rudolf Malkmus and Andreas N ӧllert in August 1994 on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.5473&amp;materialsCitation.latitude=6.0292" title="Search Plazi for locations around (long 116.5473/lat 6.0292)">Kamborangoh Road</a>, Mount Kinabalu, Sabah, West Malaysia (estimated: N6.0292, E116.5473, 1854 m a.s.l.) . Adult male (MCZ R43591) collected by John A. Griswold Jr. in 1937 from Kadmayan R, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.48973&amp;materialsCitation.latitude=6.029788" title="Search Plazi for locations around (long 116.48973/lat 6.029788)">Kiau</a>, Sabah, West Malaysia (estimated: N6.029788, E 116.489732, 971 m a.s.l.) .</p><p>Diagnosis. Morphological analyses of the four specimens from the highlands of Sabah confirms the placement of this population in the genus Asthenodipsas based on the combination of the following characters: smooth dorsal scales; absence of preoculars and suboculars; one or more supralabials in contact with the eye; a single anterior inframaxillary, followed by two or three pairs of inframaxillaries which are wider than long (Grossmann &amp; Tillack 2003). Asthenodipsas stuebingi sp. nov. can be differentiated from its congeners by the following combination of characters: a maximum SVL of 557 mm; 15/15/15 dorsal scale rows; 165–175 ventrals; 35–47 subcaudal scales; one or two postoculars; 2(+0–1)+2 temporals; six supralabials, 3 rd &amp; 4 th touching the eye; 4–6 infralabials, 2 nd or 3 rd pair in contact; a sharp vertebral keel; dorsum of adults beige to brown and overlain with 30–42 irregularly-shaped, rhomboidal dark brown bands beginning posterior to dark patch on the neck that and extends the length of body and tail and onto lateral edges of the ventral scales to form spots, but not encircling body; a narrow, light-coloured vertebral stripe; throat and ventrals beige to yellow and mottled with small, dark spots; head whitish to light-brown with dark speckling on the snout and crown; and iris and pupils black (Table 4, 5 &amp; 6; Fig. 2 &amp; 3).</p><p>Description of holotype. Adult female SVL 557 mm and Tal 62 mm; rostral as wide as high; head somewhat bulbous and elongate in shape, longer than wide; nasals undivided; internasals shorter than prefrontals; posterior margin of prefrontals contact eye; frontal hexagonal, slightly wider than long; loreals present, longer than wide; supraoculars subpentagonal, approximately half the length and half the width of frontal; preoculars absent; upper and lower postoculars present on right side of head, lower postocular extending to below orbit; single postocular on left side of head, lower margin extending to below orbit; suboculars absent; supralabials 6/6 with 3rd and 4th contacting orbit and 6th elongate; temporals 2(+1)+2/2+2, i.e. on left side of head there are two anterior temporals and two posterior temporals but between lower anterior and lower posterior temporal is an additional scale resulting in two upper temporals and three lower temporals; mental triangular wider than long; anterior inframaxillary pentagonal, in contact with infralabials 1–3; two pairs of posterior inframaxillaries following anterior inframaxillary, first pair of posterior inframaxillaries slightly oval, rhomboid-shaped, second pair more heaxagonal, elongated; infralabials 6/6 with 3 rd pair in medial contact. Body long, bulky, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsal scales smooth, in 15/15/15 rows, vertebrals slightly enlarged; 170 ventrals; 36 divided subcaudals; cloacal scute entire; tail tapering to a point.</p><p>Colouration in preservation (Fig 2A &amp; B). The ground colour of the head, body and tail is beige with dark speckling. The speckling is heavier on the dorsum nearer to the vertebral scale rows and lightens up on the flanks. The top of the head is also heavily speckled and the rostral, nasal, loreal, postoculars, supralabials, mental, inframaxillaries and infralabials are mottled with dark spots. On the neck is a dark patch starting from the nape, behind the posterior margin of the jaw, extending to approximately the position of the 11 th ventral. The dark patch starts as a narrow stripe on the 4 th &amp; 5 th dorsal scale rows, gradually widens to dorsal scale rows 1–5, and does not cross the vertebral scales or meet on the ventrals. Starting behind the dark neck patch are 42 dark, rhomboidal bands ranging from 1–2 dorsal scales in length along the body and tail. The venter is yellow and bears, dark lateral blotches from the extensions of the dorsal bands that meet the ventral scales as well as scattered dark spots between them. The thin vertebral stripe is light-yellow.</p><p>laevis group. Abbreviations are in the Materials and methods. A. jamilinaisi A. stuebingi A. laevis A. laevis Thai-Ma- A. laevis A. laevis</p><p>sp. nov. sp. nov. Borneo lay Peninsula Java Sumatra Variation (Table 4; Fig 2 &amp; 3). The paratypes closely resemble the holotype in overall external morphology, colouration, and pattern. All paratypes have a slightly darker ground colour than the holotype and a larger dark neck patch. In SP 04806 the markings from the dark neck patch extend further onto the edges of the ventrals, leaving a narrow, light-coloured median stripe on the underside of the neck and the tail is more heavily mottled. Paratypes ZMB 65429 and MCZ R43591 also have fewer infralabials than the holotype (4–5 vs. 6) and the second instead of the third pair are in contact. Variation in scale counts and size measurements within the type series are shown in Table 4.</p><p>Comparison. Asthenodipsas stuebingi sp. nov. can be differentiated from Aplopeltura boa by its higher number of mid-dorsal scale rows (15 vs. 13) and divided subcaudals (de Rooij 1917; Grossmann &amp; Tillack 2003; Stuebing et al. 2014). Asthenodipsas stuebingi sp. nov. can be differentiated from members of the genus Pareas by their possession of preocular and subocular scales (absent vs. present), supralabials in contact with orbit (3 rd &amp; 4 th contact orbit vs. no supralabials in contact with orbit) and anterior single inframaxillary (present vs. absent) (Grossmann &amp; Tillack 2003). Asthenodipsas lasgalenensis, A. tropidonotus and A. vertebralis can be differentiated from the new species by their higher number subcaudals (54–77 vs. 35–47), pairs of infralabials in contact (1 st vs. 3 rd) and more pairs of posterior inframaxillaries (three vs. two). In addition, A. vertebralis and A. tropidonotus can be differentiated from A. stuebingi sp. nov. by their higher number of ventrals (195–215 vs. 165–175). Adult A. lasgalensis can also be differentiated from A. stuebingi sp. nov. by their dorsal colour pattern that is solid dark brown to black and their whitish labials versus a beige to brown dorsum with dark-bands and a dark patch on the neck in the latter (Loredo et al. 2013). From Bornean populations of A. malaccana to which it is most similar in colour pattern, A. stuebingi sp. nov. can be differentiated by a lower number of supralabials (6 vs. 7–8) (Chan-ard et al. 2015; Das 2010; de Rooij 1917; Stuebing et al. 2014). From A. laevis to which it was previously confused, A. stuebingi sp. nov. can be differentiated by its larger adult length, (Max SVL 557mm vs. 373mm), dorsal scales rows (15/15/15 vs. 15/15/13) and sharp vertebral keel (present vs. absent) (Figs. 3F &amp; 4F) (Tables 2–6). A. stuenbingi sp. nov. can be differentiated from A. jamilinaisi sp. nov. by its lower number of subcaudals (35–47 vs. 52–53), lower number of ventrals in males (165 vs. 173–175), size of vertebral scales (slightly enlarged vs. greatly enlarged), colour pattern (light-coloured head and dorsum with a dark neck patch and distinct bands vs. dark overall colouration of dorsum with muted banding) and body form (robust and stout vs. gracile and laterally compressed) (Tables 4–6). A key to the family Pareidae of Borneo is presented below.</p><p>Distribution (Fig. 8). Asthenodipsas stuebingi sp. nov. is only known from the highlands of Sabah, East Malaysia at Mount Kinabalu, Kundasang and the Crocker Range. It is likely to range further throughout the island and found in similar upland habitats in Sarawak, Malaysia and Kalimantan, Indonesian Borneo.</p><p>Etymology. The specific epithet stuebingi is a patronym in honour of Robert B. Stuebing for his contributions to the field of herpetology in Borneo, especially on the study of snakes.</p><p>Natural History. Asthenodipsas stuebingi sp. nov. is a montane species that so far has only been found on the highlands of Sabah from 900–2000 m a.s.l.. This is in contrast to its congener A. laevis that is widely distributed in Borneo and recorded from many lowland localities up to 1150 m in elevation (Stuebing et al. 2014). At some locations such as the Crocker Range, both species occur (SP 04476, SP 04570 &amp; SP 04679; see Table 5). Similar to other species of slug snakes, A. stuebingi sp. nov. is nocturnal. It is reportedly both terrestrial and arboreal (Malkmus et al. 2002). Malkmus et al. (2002) reported finding a specimen crossing the road at night in rainy weather and another specimen in a bush approximately 1m above the ground. The paratype ZMB 65429 was also collected at night fol- lowing light rains at approximately 22:00 h. A specimen was observed at 09:00 h close the power station of the Kinabalu park headquarters along Kamborangoh road (Andreas N ӧllert in litt. 2018). When threatened, the snakes roll themselves up into tight coils and remain motionless (Fig. 3D &amp; E) — a behaviour observed in Aplopeltura boa (Jablonski &amp; Hegner 2016) . Asthenodipsas stuebingi sp. nov. presumably feeds on snails or slugs like other mem- bers of the family and nothing is known about the reproductive biology of the species.</p></div>	https://treatment.plazi.org/id/03E187FC3616494BFF5FFCECFBBAFE93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Quah, Evan S. H.;Grismer, L. Lee;Lim, Kelvin K. P.;Anuar, M. S. Shahrul;Imbun, Paul Y.	Quah, Evan S. H., Grismer, L. Lee, Lim, Kelvin K. P., Anuar, M. S. Shahrul, Imbun, Paul Y. (2019): A taxonomic reappraisal of the Smooth Slug Snake Asthenodipsas laevis (Boie, 1827) (Squamata: Pareidae) in Borneo with the description of two new species. Zootaxa 4646 (3): 501-526, DOI: 10.11646/zootaxa.4646.3.4
03E187FC36124943FF5FF8AFFDEAFBAF.text	03E187FC36124943FF5FF8AFFDEAFBAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asthenodipsas jamilinaisi Quah & Grismer & Lim & Anuar & Imbun 2019	<div><p>Asthenodipsas jamilinaisi sp. nov.</p><p>Sabah Mountain Slender Slug Snake</p><p>Fig. 4 &amp; 5.</p><p>urn:lsid:zoobank.org:act: FE0ABCEE-6DD3-449E-A828-057CFB16639A</p><p>Amblycephalus laevis (in part): Boulenger 1896: 441 &amp; 442; de Rooij 1917: 276.</p><p>Asthenodipsas laevis (in part): Das 2010: 344; Stuebing et al. 2014: 82.</p><p>Asthenodipsas vertebralis (in part): Stuebing et al. 2014: 82.</p><p>Internatus laevis (in part): Malkmus et al. 2002: 343.</p><p>Pareas laevis: Stuebing 1991: 330 &amp; 331 (in part); Stuebing &amp; Inger 1999: 87 (in part); Haile 1958: 766 (in part).</p><p>Pareas vertebralis (in part): Stuebing &amp; Inger 1999: 87.</p><p>Holotype. Adult male, SP 04076 collected by Paul Yambun Imbun and other unspecified members of Sabah Parks on 14 May 1991 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.51667&amp;materialsCitation.latitude=5.552776" title="Search Plazi for locations around (long 116.51667/lat 5.552776)">Mount Trusmadi</a>, Tambunan, Sabah, East Malaysia (estimated: N 5.552776, E 116.516667, 2612 m a.s.l.).</p><p>Paratypes. Juvenile male (ZRC 2.2742) collected by F.N. Chasen and H.M. Pendlebury in May 1929 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.51902&amp;materialsCitation.latitude=6.080644" title="Search Plazi for locations around (long 116.51902/lat 6.080644)">Marei Parei</a>, Mount Kinabalu, Sabah, East Malaysia (estimated: N 6.080644, E 116.519022, 1668 m a.s.l.) .</p><p>Diagnosis. Morphological examination of the type series determined their placement in the genus Asthenodipsas owing to their possession of smooth dorsal scales; absence of preoculars and suboculars; one or more supralabials in contact with the eye; a single anterior inframaxillary, followed by two or three pairs of inframaxillaries which are wider than long (Grossmann &amp; Tillack 2003)—character states that diagnose Asthenodipsas from other genera of the family Pareidae . Asthenodipsas jamilinaisi sp. nov. can be differentiated from its congeners by the following combination of characters: a maximum SVL of 378 mm; 15/15/15 dorsal scale rows; 173–175 ventrals (males); 52–53 subcaudal scales (male); postoculars; 1–2(+0–1)+2 temporals; 6–7 supralabials, 3 rd &amp; 4 th touching the eye; 5–6 infralabials, 3 rd pair in contact; a sharp vertebral keel; dorsum of adults dark-brown with 44–48 indistinct dark bands that are more prominent on the lower dorsal rows along the flanks and extend onto the corners of the ventrals; chin &amp; throat dark-brown; ventrals cream to light-yellow; and iris and pupils black (Tables 4–6; Fig. 4 &amp; 5).</p><p>Description of holotype. Adult male SVL 378 mm and Tal 78 mm; rostral wider than tall; head narrow, elongate, somewhat bulbous, longer than wide; nasals undivided; internasals shorter than prefrontals; posterior margin of prefrontals contact eye; frontal hexagonal, slightly wider than long; loreals present, longer than tall; supraoculars subpentagonal, approximately half the length and a third the width of frontal; preoculars absent; two postoculars present on both left and right side of head, lower postocular extending to below orbit; suboculars absent; supralabials 6/6 with 3rd and 4th contacting orbit and 6th elongate; temporals 1+2/2(+1)+2, i.e. on left side of head, upper anterior temporal is fused with parietal resulting in a single lower anterior temporal and two posterior temporals, while on right side of head there are two anterior temporals and two posterior temporals but between lower anterior and lower posterior termporal is an additional scale resulting in two upper temporals and three lower temporals; mental triangular wider than long; anterior inframaxillary hexagonal, in contact with infralabials 1–3; two pairs of posterior inframaxillaries following anterior inframaxillary, first pair of posterior inframaxillaries slightly squarish, rhomboid-shaped, second pair more heaxagonal, elongated; infralabials 6/6 with 3 rd pair in medial contact. Body long, slender, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsal scales smooth, in 15/15/15 rows, vertebrals greatly enlarged; 173 ventrals; 53 divided subcaudals; cloacal scute entire; tail tapering to a point.</p><p>Colouration in preservation (Figs. 4A &amp; B). The ground colour of the head, body and tail is dark-brown. The colour is darker on the back along the vertebral scale rows and lightens up to light-brown on the lower flanks. Along the body and tail are 44 dark, rhomboidal bands ranging from 1–3 dorsal scales in length starting from neck. These bands are indistinct on the dorsum and are only clearly visible along the lower flanks and extend onto the corners of the ventral scales. The chin and throat are dark-brown while the iris and pupils are black. The venter is cream to light-yellow and bears dark, lateral blotches from the extensions of the dorsal bands that meet corners of the ventral scales.</p><p>Variation (Figs. 4 &amp; 5). The juvenile paratype ZRC 2.2742 approaches the holotype in pholidosis but has seven supralabials on the right side of the head, five pairs of infralabials, 48 body bands that are more prominent than those on the adult holotype and a lighter vertebral region (Table 4; Fig. 4C &amp; D). Based on photographs of other specimens from Mount Kinabalu, Sabah that have been tentatively identified as this species, the ground colour of some are dark-grey with nearly obscure banding (Fig. 5A), while in others there is a poorly defined, broken, whitish, vertebral stripe (Fig. 5D) and the ventral colouration is white (Fig. 5B). The juveniles are more contrastingly patterned than the adults with the lower dorsal scale rows white, forming a poorly-defined white stripe along the lower flanks; more prominent banding on the body and tail; and a very prominent, broken, white, vertebral stripe that appear as a series of white spots along the back (Fig. 5C).</p><p>A. jamilinaisi sp. nov. A. jamilinaisi A. stuebingi sp. nov.</p><p>sp. nov.</p><p>SP 04076 ZRC 2.2742 SP 04679 SP 04806 ZMB 65429 MCZ R43591</p><p>(holotype) (paratype) (holotype) (paratype) (paratype) (paratype)</p><p>Mount Trusmadi, Tam- Marei Parei, Mt. Minduk Sirung, Crocker Kg. Desa, Kundasang, Mount Kinabalu, Kadmayan Rd., Kiau,</p><p>bunan, Sabah Kinabalu, Sabah Range Park, Sabah Sabah Sabah Mount Kinabalu, Sabah</p><p>......continued on the next page</p><p>A. jamilinaisi sp. nov. A. jamilinaisi A. stuebingi sp. nov.</p><p>sp. nov.</p><p>SP 04076 ZRC 2.2742 SP 04679 SP 04806 ZMB 65429 MCZ R43591</p><p>(holotype) (paratype) (holotype) (paratype) (paratype) (paratype)</p><p>Mount Trusmadi, Tam- Marei Parei, Mt. Minduk Sirung, Crocker Kg. Desa, Kundasang, Mount Kinabalu, Kadmayan Rd., Kiau,</p><p>bunan, Sabah Kinabalu, Sabah Range Park, Sabah Sabah Sabah Mount Kinabalu, Sabah</p><p>Comparison. Asthenodipsas jamilinaisi sp. nov. can be differentiated from Aplopeltura boa by its higher number of mid-dorsal scale rows (15 vs. 13) and divided subcaudals (de Rooij 1917; Grossmann &amp; Tillack 2003; Stuebing et al. 2014). Asthenodipsas jamilinaisi sp. nov. can be differentiated from members of the genus Pareas by its possession of preocular and subocular scales (absent vs. present), supralabials in contact with orbit (3 rd &amp; 4 th contact orbit vs. no supralabials in contact with orbit) and anterior single inframaxillary (present vs. absent) (Grossmann &amp; Tillack 2003). Asthenodipsas lasgalenensis, A. tropidonotus and A. vertebralis can be differentiated from A. jamilinaisi sp. nov. by their possession of more pairs of posterior inframaxillaries (three vs. two) and pairs of infralabials in contact (1 st vs. 3 rd). In addition, A. vertebralis and A. tropidonotus can be differentiated A. jamilinaisi sp. nov. by their higher number of ventrals (195–215 vs. 173–175) (Loredo et al. 2013). A. jamilinaisi sp. nov. can be differentiated from A. malaccana by its lower number of supralabials (6 vs. 7–8) and colour pattern (dark dorsum with indistinct banding vs. light-grey to brown body with distinct banding or blotches and a prominent dark patch on the neck) (Chan-ard et al. 2015; Das 2010; de Rooij 1917; Stuebing et al. 2014). Asthenodipsas jamilinaisi sp. nov. can be distinguished from A. laevis by its larger adult length, (max SVL 378 mm vs. 373 mm), higher number of ventral scales (173–175 vs. 148–173), dorsal scales rows (15/15/15 vs. 15/15/13) and sharp vertebral keel (present vs. absent) (Figs. 4E &amp; 4F) (Tables 3–6). A. jamilinaisi sp. nov. can be differentiated from A. stuebingi sp. nov. by its higher number of subcaudals (52–53 vs. 35–47), higher number of ventrals in males (173–175 vs. 165), size of vertebral scales (greatly enlarged vs. slightly enlarged), colour pattern (dark overall colouration of dorsum with indistinct banding vs. light-coloured head and dorsum with a dark neck patch and distinct bands) and body form (gracile and laterally compressed vs. robust and stout) (Tables 3–6). A key to the family Pareidae of Borneo is presented below.</p><p>Distribution (Fig. 8). Asthenodipsas jamilinaisi sp. nov. is only known from the highlands of Sabah, East Malaysia at Mount Trusmadi and Mount Kinabalu. It may possibly be found at other montane areas in Borneo.</p><p>Etymology. The specific epithet jamilinaisi is a patronym in honour of Dr. Jamili Nais, the Director of Sabah Parks for his contributions to the research and conservation of biodiversity in the state and the first Malaysian to be appointed as a member of the World Heritage panel of the International Union for Conservation of Nature (IUCN).</p><p>Natural History. Little is known about the natural history of A. jamilinaisi sp. nov. except that it is a montane species that has only been found from 1500 to 2500 m a.s.l. in Sabah. Similar to other slug snakes it is nocturnal and has been found crawling in the vegetation. The paratype and another juvenile (Fig. 5C) were found during May which indicates that breeding takes place in the earlier half of the year. Presumably it also feeds on snails and slugs like other members of the family.</p></div>	https://treatment.plazi.org/id/03E187FC36124943FF5FF8AFFDEAFBAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Quah, Evan S. H.;Grismer, L. Lee;Lim, Kelvin K. P.;Anuar, M. S. Shahrul;Imbun, Paul Y.	Quah, Evan S. H., Grismer, L. Lee, Lim, Kelvin K. P., Anuar, M. S. Shahrul, Imbun, Paul Y. (2019): A taxonomic reappraisal of the Smooth Slug Snake Asthenodipsas laevis (Boie, 1827) (Squamata: Pareidae) in Borneo with the description of two new species. Zootaxa 4646 (3): 501-526, DOI: 10.11646/zootaxa.4646.3.4
