identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E1981EB533FFE4FF59F96F044DEFE6.text	03E1981EB533FFE4FF59F96F044DEFE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala aglaos Filippini, Galante	<div><p>Anomala aglaos Filippini, Galante, &amp; Micó, new species</p><p>Fig. 1</p><p>Material examined. Holotype: ♂ "La Montura, Prov. San José, Costa Rica 07/11/2007 Leg. M. Moraga / CEUA 00106184" deposited in INBIO .</p><p>Paratypes (6): 1 ♂ " Isla Bonita, Prov. Alajuela, Costa Rica. 11/06/2007 Leg. M. Moraga / CEUA 00106183"; 1 ♂ " COSTA RICA Prov. San José. P.N. Braulio Carrillo. Sitio La Montura. 1100m. 13 JUN 2007. A. García, M. Moraga, M.A. Zumbado. Tp. Luz 1. L_N_232500_539125 #92217 / CEUA 00106185"; 1 ♀ " COSTA RICA. Prov. Alajuela. Guatuso. P.N. Volcán Tenorio. Punto 1: Catarata Río Buenavista. 800m. 5 JUN 2008. J. A. Azofeifa. Tp de Luz 2. L_N_298380_427850 #93849 / CEUA 00106186"; 1 ♀ " COSTA RICA Prov. San José. P.N. Braulio Carrillo. Sitio La Montura. 1100m. 12 JUL 2007. A. García, M. Moraga. Tp. Luz 2. L_N_232500_539125 #92245 / CEUA 00106187"; 1 ♀ "La Montura Prov. San José Costa Rica 13/06/2007 Det. D. Briceño" INBIOCRI; 1 ♀ " Isla Bonita, Prov. Alajuela, Costa Rica. 6/11/2007 Leg. A. García, M. Moraga / CEUA 00106189".</p><p>Holotype male description. Body shape elongate. Length 17.5 mm. Width 9.7 mm.</p><p>Color. Head, pronotum, and scutellum metallic green. Elytra metallic green with branched ochre lines along apical portion of costae and coppery luster. Pygidium bronze. Legs metallic green with coppery luster. Venter dark metallic green, abdominal sternites copper.</p><p>Clypeus trapezoidal, surface densely punctate, convex. Clypeus anterior margin slightly sinuate, anteriorly thick. Clypeus ratio width/length 2.5. Frons moderately punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.0. Antenna: ratio funiculus/club 0.7.</p><p>Pronotum subtrapezoidal, ratio width/length 1.9. Lateral margins sinuate. Anterior angles acute, sharp; posterior angles obtuse, blunt. Basal margin sinuate, subapical bead obliterated in the middle. Surface with fine punctures.</p><p>Figures 7–12. Shape of protibia in 7: Anomala aglaos (male: holotype; female: paratype, Isla Bonita, Costa Rica). 8: A. estrella (male: holotype; female: paratype, Los Almendros, Costa Rica). 9: A. inbio (male: holotype; female: paratype, Reserva Biológica Hitoy Cerere, Costa Rica). 10: A. pincelada (male: paratype, Parque Nacional Santa Rosa, Costa Rica; female: paratype, Tierras Morenas, Costa Rica). 11: Callistethus ruteloides (male: holotype; female: paratype, El Copal, Cartago). 12: Shape of protibia of Callistethus yalizo (male: paratype, Esperanza del Guarco, Cartago; female: paratype, Quebrada Segunda, Cartago). Scale = 1 mm.</p><p>Scutellum. Shape subtriangular, with rounded sides and blunt apex. Ratio width/length 1.4.</p><p>Elytra with costae defined by irregular rows of sparse punctures. Interstices with sparse, shallow punctures; surface slightly irregular.</p><p>Pygidium sparsely punctate with long setae at apical margin. Ratio width/length 1.5.</p><p>Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1–2 rows of setae (2–3 rows on 2nd sternite) and 6–8 rows of punctures per sternite. Last sternite punctate, with subapical bead thin and sinuate, apical membrane narrow.</p><p>Protibia (Fig. 7) bidentate; apical tooth long and curved; basal tooth at same level as internal apex of protibia, triangular in shape, acute. Metatibia slender, fusiform. Ratio length/width 3.2. First external carina obliterated. Surface punctate above second external carina and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower one, inferior margin sinuate.</p><p>Aedeagus (Fig. 13): parameres large and stout in a lateral view, but narrow dorsally. Ventral plate with curved apex in a lateral view. Endophallus (Fig. 19) with 2 long sacculi; 1 ending with a pair of thick, long, sclerotized projections with a long diverticle in between, and with a patch of sclerotized spines medially; the other sacculi with swollen base, where ejaculatory sclerite and duct opening are located, and a ridged plate of thick spines at apex. Median lobes asymmetric, 1 with a triangular fold near apex.</p><p>Variation. Ochre lines on elytra of variable length; elytra with green or copper luster. Female. Similar to male, protibia (Fig. 7) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw narrower with curved inferior margin; last sternite with curved apex.</p><p>Body length 17.5–18.8 mm, body width 9.7–10.2 mm. Clypeus w/l: 2.3–2.7. Interocular ratio (interocular width/width of eye): 3.0–3.4. Antenna: ratio funiculus/club 0.7–0.9. Pronotum w/l: 1.8–1.9. Scutellum w/l: 1.4– 1.6. Pygidium w/l: 1.5–1.6. Metatibia l/w: 2.9–3.5.</p><p>Diagnosis. Size large, shape elongate, metallic green color with branched ochre lines on apical portion of elytra, elytral surface irregular, parameres large and stout, endophallus with a pair of long, sclerotized projections.</p><p>This species is similar to A. obovata Ohaus, 1933, with the slightly bigger size, oblong shape, and dark green color; but A. obovata lacks the ochre lines on elytra, its elytral surface is strongly irregular with large and deep coalescing punctures, and it has slender parameres (Fig. 27), endophallus with a sacculus with a large patch of dense setae on basal portion, instead of sclerotized projections (Fig. 28).</p><p>Etymology. From the Greek adjective aglaós, splendid, beautiful, treated as a noun in apposition, for the nice and elegant aspect of this species.</p><p>Distribution. Guanacaste and Central mountain ranges, Costa Rica, from 800–1100 m (Fig. 34).</p></div>	https://treatment.plazi.org/id/03E1981EB533FFE4FF59F96F044DEFE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
03E1981EB537FFEBFF59F895044EECA5.text	03E1981EB537FFEBFF59F895044EECA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala estrella Filippini, Galante	<div><p>Anomala estrella Filippini, Galante, &amp; Micó, new species</p><p>Fig. 2</p><p>Material examined. Holotype: ♂ "Est. La Casona, 1520m, Res. Biol. Monteverde, Prov. Puntarenas, Costa Rica, N. Obando, Mar 1992 L-N 253250,449700 / INBIOCRI000898153" deposited in INBIO .</p><p>Paratypes (8): 1♀ "Estac. Maritza, 600 m. W side Volcan Orosi Guanac. Pr. COSTA RICA. 17 May 1988. Janzen &amp; Hallwachs W85 29 37 ", N10 57 39 " / INBIOCRI002517167"; 1♂ "San Jose. Costa Rica Hda. Tiquires. 1500m Rio Tiquires 28 marzo 1988 Col.A.Solis / CEUA 00106164"; 2♀ "Est. La Casona, 1520m, Res. Biol. Monteverde, Prov. Puntarenas, Costa Rica, N. Obando, Mar 1992 L-N 253250,449700" / INBIOCRI000898149 and INBIOCRI000898152; 1♂ "Est. Las Pailas, 800 m, P.N. Rincon de la Vieja, Prov. Guan., COSTA RICA. 6 ene a 13 abr 1993, J. Sihezar, G. Rodriguez, L- N 306300_388600 / INBIOCRI001331532"; 1♀ "Est. Las Pailas, 800 m, P.N. Rincon de la Vieja, Prov. Guan., COSTA RICA. 10 a 27 Mar 1993, K. Taylor, L- N 306300_388600 / INBIOCRI001355780"; 1♂ "Est. Maritza, 600 m, lado O Vol. Orosi, Prov. Guan. COSTA RICA. R. Blanco, Abr 1990, L- N 326900_373000 / INBIOCRI000238024"; 1♀ "Los Almendros, P. N. Guanacaste, Prov. Guanacaste, Costa Rica, 28 mar a 24 abr 1992, M. Reyes, L- N 334800_369800 / CEUA 00105806".</p><p>Holotype male description. Body shape elongate. Length 14.9 mm. Width 8.4 mm.</p><p>Color. Head dark brown. Pronotum orange ochre with 1 large irregular dark brown macula on disc, not reaching basal margin, crossed by a lighter weak longitudinal line; small maculae on lateral foveae; surface with green luster. Scutellum light brown with lighter center. Elytra orange ochre. Pygidium ochre. Legs reddish brown; protibiae with darker margins, metafemura ochre, metatibiae and metatarsi dark brown with green luster. Venter, sternum reddish brown with darker margins, abdominal sternites ochre.</p><p>Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.3. Frons densely punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 2.9.</p><p>Pronotum subtrapezoidal. Ratio width/length 1.7. Lateral margins angulated at 1/3 of pronotum length. Apical angles acute, sharp; basal angles obtuse, blunt. Basal margin sinuate, subapical bead complete. Surface densely punctate.</p><p>Scutellum. Shape subtriangular, with rounded sides, apex acute. Ratio width/length 1.5. Surface densely punctate.</p><p>Elytra with costae defined by regular rows of punctures. Subsutural interstice with 3–5 rows of punctures, 2nd– 4th interstices with 1–2 irregular rows of punctures. Surface covered with secondary punctures.</p><p>Pygidium finely granulate with long setae at apical margin. Ratio width/length 1.4.</p><p>Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of sparse setae and 5–6 rows of punctures per sternite. Terminal sternite punctate, with subapical bead narrow and sinuate, apical membrane narrow.</p><p>Protibia (Fig. 8) bidentate; apical tooth long and curved; basal tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, slightly narrower subapically. Ratio length/width 2.6. First external carina obliterated; surface densely punctate above second external carina, rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower branch; inferior margin sinuate.</p><p>Aedeagus (Fig. 14): parameres with sinuate dorsal and ventral margins, apex acute in a lateral view. Endophallus (Fig. 20) with 1 long sacculus with wide apex, covered with setae medially and with a strip of long and short, thick spines at apex. Ejaculatory duct opening placed on a ventral inflation.</p><p>Variation. Macula on pronotum with variable extension either fusing with maculae on foveae, forming a starshaped macula or reaching the posterior margin; elytra ochre to red. Female. Similar to male, protibia (Fig. 8) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw with curved inferior margin.</p><p>Body length 14.4–16.1 mm, body width 7.9–9.6 mm. Clypeus w/l: 2.2–2.3. Interocular ratio (interocular width/width of eye): 2.5–3.0. Antenna: ratio funiculus/club 0.6–0.7. Scutellum w/l: 1.2–1.5. Pygidium w/l: 1.4–1.6. Metatibia l/w: 2.5–2.8.</p><p>Diagnosis. Size large; macula on pronotum irregular, dark; elytra ochre; parameres with sinuate dorsal and ventral margins in lateral view; endophallus with a strip of long and short, thick spines at apex.</p><p>Anomala jansoni Ohaus, 1897 is a similar species of similar size but with two small maculae on pronotum and parameres with long, narrow, curved apex (Fig. 29) (Monte Rotondo, Costa Rica; MNHUB). Anomala antica Ohaus, 1897, A. luciae Blanchard, 1851, and A. quiche Ohaus, 1897 are similar but smaller species, with one small and pentagonal macula on pronotum ( A. antica), pronotum with sinuate lateral margins ( A. quiche), maculae on elytra ( A. luciae and A. quiche), and differences in the parameres: A. antica has a long thin apex (Fig. 30), A. luciae has lobes on anterior portion of inferior margins (Fig. 31), and A. quiche has wide rounded parameres, and the ventral plate curved at apex in a lateral view (Fig. 32).</p><p>Etymology. Latinized from the Spanish noun estrella, star, for the star-shaped macula on pronotum. Treated here as a noun in apposition,.</p><p>Distribution: Guanacaste and Tilarán mountain ranges, Costa Rica, from 280–1520 m (Fig. 34).</p></div>	https://treatment.plazi.org/id/03E1981EB537FFEBFF59F895044EECA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
03E1981EB539FFE8FF59FE9F022BEDF0.text	03E1981EB539FFE8FF59FE9F022BEDF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala inbio Filippini, Galante	<div><p>Anomala inbio Filippini, Galante, &amp; Micó, new species</p><p>Fig. 3</p><p>Material examined. Holotype: ♂ " COSTA RICA, Prov. Limon, Est. Hitoy Cerere, 100m. 21 MAR-7 ABR 1998. E. Rojas. Tp. Luz L_N_184600_643400 #49943 / INBIOCRI002415599" deposited in INBIO .</p><p>Paratypes (10): 1♂ " COSTA RICA. Prov. Limón, R.B. Hitoy Cerere, Estación Hitoy Cerere, 100m, 21 MAR 2003, B. Gamboa, Tp. de Luz, L_N_184120_643471 #73276 / INB0003703490" (INBIOCRI); 1♂ " COSTA RICA. Prov. Limón. R.B. Hitoy Cerere. Send. Tepezcuintle. 0- 100m. 25 MAR 2008. A. Solis, C. Hernández. Libre. L_N_184300_643300 #93807 / CEUA 00106172"; 2♂ " COSTA RICA. Prov. Heredia. Sarapiquí. Est. Biol. La Tirimbina. Mirador. 167m. 23-24 ABR 2009. C. Hernández, D. Díaz. Tp. Luz. L_N_266268_523359 #96599" / INB0004211777 and INB0004211778 (INBIOCRI); 1♂ "Est. Hitoy Cerere, 100 m, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limon, COSTA RICA. Abr 1993. G. Carballo. L- N 184200_643300 / INBIOCRI000975461"; 3♂, 2♀ "R.B. Hitoy Cerere, Valle La Estrella, Prov. Limon, COSTA RICA. 100- 200m. 4-20 Abr 1994, G. Carballo, L N 643400_184600 # 2855" / INBIOCRI001802267, INBIOCRI001802285, INBIOCRI001802287, INBIOCRI001802301, and CEUA 00106173.</p><p>Holotype male description. Body shape elongate. Length 16.8 mm. Width 8.6 mm.</p><p>Color. Head, pronotum, scutellum, pygidium, legs, and venter black. Elytra ochre with irregular black maculae: 3 pairs along basal margin, 1 median, 1 on sides towards apex; the latter with small flecks near edges.</p><p>Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.2. Frons densely punctate, flat. Ocular canthum long, narrow, with rounded apex. Interocular ratio (interocular width/width of eye): 2.7. Antenna: ratio funiculus/club 0.7.</p><p>Pronotum subtrapezoidal. Ratio width/length 1.8. Lateral margins regularly convex. Anterior angles right, blunt; posterior angles obtuse, blunt. Basal margin sinuate, subapical bead complete. Surface densely punctate.</p><p>Scutellum. Shape subpentagonal, with rounded sides and acute apex. Ratio width/length 1.3. Surface sparsely punctate.</p><p>Elytra with costae defined by regular rows of punctures. Subsutural interstice with 2–3 rows of punctures, 2nd– 4th interstices with 1–2 irregular secondary striae.</p><p>Pygidium finely strigate, with long setae at apical margin. Ratio width/length 1.8.</p><p>Space between the mesocoxae narrow, slightly convex. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of setae (2–3 rows on 2nd sternite) and 3–5 rows of punctures per sternite. Terminal sternite strigate, with subapical bead thick and sinuate, apical membrane wide.</p><p>Protibia (Fig. 9) bidentate; apical tooth long and curved; basal tooth at the same level as internal apex of protibia, triangular in shape, sharp, acute. Metatibia stout, fusiform. Ratio length/width 3.1. First external carina constituted by a row of sparse setae, surface punctate above second external carina, and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower branch, inferior margin sinuate.</p><p>Aedeagus (Fig. 15): parameres stout; with protruding, narrow apex ending sharply. Ventral angle nearly quadrate, blunt. Endophallus (Fig. 21) with 1 long sacculus with a small patch of sclerotized spines near apex; ejaculatory duct opening ventrally at base; median lobes heavily sclerotized.</p><p>Variation. Extension of dark maculae on elytra variable. Female similar to male, protibia (Fig. 9) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw narrower with curved inferior margin; terminal sternite with curved apex.</p><p>Body length 15.7–17.6 mm, body width 8.6–9.3 mm. Clypeus w/l: 2.00–2.2. Interocular ratio (interocular width/width of eye): 2.5–2.7. Antenna: ratio funiculus/club 0.7–0.8. Pronotum w/l: 1.7–1.8. Scutellum w/l: 1.2– 1.6. Pygidium w/l: 1.7–1.9. Metatibia l/w: 2.8–3.1.</p><p>Diagnosis. Large size; black pronotum and irregular maculae on elytra; parameres stout, with protruding narrow, acute apex.</p><p>The most similar species are A. inconstans and A. pincelada, but they are smaller, have ochre margins on pronotum, have a different pattern of maculae on elytra, and have different shaped aedeagus: A. pincelada has a bifurcate apex of parameres (Fig. 16), A. inconstans has parameres with a frontal lobe-like expansion on anterior part of ventral margin (Fig. 33).</p><p>Etymology. This species is dedicated to INBio, Instituto Nacional de Biodiversidad in Costa Rica, for its efforts in increasing the knowledge of Costa Rican biodiversity. This name should be treated as a noun in apposition.</p><p>Distribution: Caribbean lowlands, from 0–167 m (Fig. 35).</p></div>	https://treatment.plazi.org/id/03E1981EB539FFE8FF59FE9F022BEDF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
03E1981EB53AFFE9FF59FE2C0295EF65.text	03E1981EB53AFFE9FF59FE2C0295EF65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala pincelada Filippini, Galante	<div><p>Anomala pincelada Filippini, Galante, &amp; Micó, new species</p><p>Fig. 4</p><p>Material examined. Holotype: ♂"Fca. Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 18-25 Abr 1993, E. Araya, L- N 316200_364400 / INBIOCRI001167801" deposited in INBIO .</p><p>Paratypes (9): 1♂ "3 Km este de Cuajiniquil, 300m, Prov. Guanacaste, Costa Rica, 25 jun 1992, III curso Parataxon. L-N 325600_355200 / INBIOCRI000863924"; 1♂ "Santa Rosa National Park Guanacaste Prov. COSTA RICA. 9-11 May 1980 DH Janzen &amp; W Hallwachs / INBIOCRI001116980"; 1♂, 1♀ "Fca. Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 9-14 May 1993. E. Araya, L-N 316200_364400" INBIOCRI001183402 and INBIOCRI001183321; 1♀ "Tierras Morenas, Rio San Lorenzo, Tenorio, Prov. Guana, COSTA RICA, 1050m. May 1993. G. Rodriguez, L S 283950_424500 # 2118 / INBIOCRI001180936"; 1♂ "Sector Las Pailas, P. N. Guanacaste, A. C. Guanacaste, Prov. Guana, COSTA RICA. 800 m. 6-26 Jun 1994, K. Taylor, L N 309500_389500 # 3063 / INBIOCRI001909257"; 2♀ "Finca Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 18-25 Abr 1993, E. Araya, L- N 316200_364400" INBIOCRI001167798 and CEUA 00106161; 1♂ "Est. Murcielago, 8 km S. O. de Cuajiniquil, Prov. Guana, COSTA RICA. 100m. 16 Jun- 4 Jul 1993. F. Quesada, L N 320300_347200 # 2177 / CEUA 00106160".</p><p>Holotype male description. Body shape elongate. Length 13.2 mm. Width 7.9 mm.</p><p>Color. Head reddish brown. Pronotum ochre with an irregular median brown macula and small maculae on lateral foveae. Scutellum dark brown. Elytra ochre with a dark brown macula at each side of scutellum and dark stripes on apical third of first interstice, from basal to apical calli and near lateral margins. Pygidium ochre. Legs ochre with base and apex of tibiae dark brown; protarsi and mesotarsi reddish brown, metatarsi dark brown. Venter ochre.</p><p>Clypeus trapezoidal, with anterior angles widely curved; surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.1. Frons densely punctate, flat. Ocular canthum long, thin, with acute apex. Interocular ratio (interocular width/width of eye): 2.6. Antenna: ratio funiculus/club 0.8.</p><p>Pronotum subtrapezoidal, ratio width/length 1.7, lateral margins regularly convex. Anterior angles quadrate and blunt. Posterior angles obtuse and blunt. Basal margin sinuate, subapical bead complete. Surface with fine, dense punctures.</p><p>Scutellum. Shape subtriangular, with rounded sides and blunt apex. Ratio width/length 1.4. Surface densely punctate.</p><p>Elytra with costae defined by regular rows of punctures, subsutural interstices with 2–3 rows of punctures; maximum width towards apex.</p><p>Pygidium with coalescing punctures and long setae at apical margin. Ratio width/length 1.6.</p><p>Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of setae and 2–3 rows of sparse, fine punctures per sternite. Terminal sternite punctate with subapical bead narrow and sinuate, apical membrane wide.</p><p>Protibia (Fig. 10) tridentate; apical tooth long and curved; middle tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, fusiform, ratio length/width 3.0. First external carina consists of a row of sparse setae, surface punctate above second external carina and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and half the width of the lower branch. Inferior margin with a sharp angle near base.</p><p>Aedeagus (Fig. 16): parameres deeply and widely sinuate anteriorly in a lateral view, becoming nearly bifurcate. Endophallus (Fig. 22) with 1 long lateral sacculus tapering at apex; ejaculatory duct opening on the other side, at base; 1 small diverticle with long spines ventrally at base.</p><p>Variation. Macula on pronotal disc of different size, from a subpentagonal macula not reaching posterior margin to covering most of the surface with ochre sides; a median narrow, faint, lighter band may be present; maculae on elytra of variable length; metatibia from ochre to red with darker apex. Female similar to male, protibia (Fig. 10) with longer and wider apical tooth; internal protarsal claw with curved inferior margin.</p><p>Body length 13.2–16.1 mm, body width 7.9–9.2 mm. Clypeus w/l: 2.0–2.3. Interocular ratio (interocular width/width of eye): 2.5–2.9. Antenna: ratio funiculus/club 0.7–0.8. Pronotum w/l: 1.7–1.8. Scutellum w/l: 1.4– 1.6. Pygidium w/l: 1.5–1.6. Metatibia l/w: 2.5–3.0.</p><p>Diagnosis. Externally nearly identical to A. inconstans Burmeister, 1844 (Colombia, MLUH), it is separated from this species by the shape of male genitalia, which has bifurcated apex of parameres, whereas A. inconstans has parameres with thin and curved apex, blunt and wide ventral angle, and a frontal lobe-like expansion on anterior part of ventral margin (Fig. 33). Specimens from Mexico, Paraguay, Guatemala, and Venezuela from MNHUB collection, also similar to A. inconstans, all show different aedeagi, so these species are part of a large and widely distributed species complex.</p><p>Etymology. Latinized from Spanish noun pincelada, brush stroke, for the elytral pattern. This name should be treated as a noun in apposition.</p><p>Distribution. Guanacaste mountain range, from 200–600 m (Fig. 35).</p></div>	https://treatment.plazi.org/id/03E1981EB53AFFE9FF59FE2C0295EF65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
03E1981EB53BFFEFFF59FCAD00E4EC88.text	03E1981EB53BFFEFFF59FCAD00E4EC88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus ruteloides Filippini, Galante	<div><p>Callistethus ruteloides Filippini, Galante, &amp; Micó, new species</p><p>Fig. 5</p><p>Type material. Holotype: ♂ "Quebrada Segunda, P. N. Tapanti, 1250 m, Prov. Cartago, Costa Rica, G. Mora, May 1992 L-N 194000_560000 / INBIOCRI000959028" deposited in INBIO .</p><p>Paratypes (6): 1♂ "Est. Biol. Las Alturas, 1500 m, Coto Brus, Prov. Puntarenas, Costa Rica. M. Ramirez. Mar 1992 L-S 322500, 591300 / INBIOCRI000987456"; 1♀ "Tapantí (Costa Rica) 1300 m 29-9-06 T. luz 17:00-23:00 Leg.: Micó, García, Galante / CEUA 00019649"; 1♀ "El Copal, Prov. Cartago, Costa Rica. 17/06/2007 Leg. M. Moraga / CEUA 00106211"; 1♀ " COSTA RICA. Prov. Alajuela. Upala. P.N. Volcán Tenorio. Alb. Heliconias. Send. a Laguna Danta. 900 m. 16 JUN 2007. J. D. Gutiérrez. Tp. Luz 2. L_N_299100_424000 #92208 / CEUA 00106212"; 1♀ "Q. Segunda, P.N. Tapantí, Prov. Carta, COSTA RICA. 1300 m. Set 1993. G. Mora, L N 194000_559800 #2326 / INBIOCRI001669692"; 1♀ " PANAMA - CHIRIQUI Santa Clara env. 1546 m 08°51'42,2"N; 082°44'36,5" W 17.6.- 4.7.06; V. Malý lgt. P 7 / coll. Vladislv Malý Česká Rep. Praha" Vladislv Malý (Česká, Czech Republic), personal collection.</p><p>Holotype male description. Body shape oval. Length 17.5 mm. Width 9.8 mm.</p><p>Color. Head pale ochre with a sinuate brown macula on frons. Pronotum pale ochre with 2 lateral brown bands. Scutellum pale ochre with red margins. Elytra pale ochre with 2 transversal sinuate beige bands, 1 median and 1 apical. Pygidium and legs ochre. Venter orange-ochre.</p><p>Head large, almost 2/3 of the width of pronotum. Clypeus rectangular, with anterior angles widely curved. Clypeus ratio width/length 2.1; surface densely punctate-reticulate, convex; anterior margin straight, nearly as long as frons. Frons moderately punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.0. Antenna: ratio funiculus/club 0.6. Mandibles with apex curved ventrally, maxillae with 2nd and 3rd tooth fused together. Labium and labrum quadrilobulate.</p><p>Pronotum trapezoidal. Width 1.9 times length. Lateral margins curved. Anterior angles quadrate with blunt vertex; basal angles obtuse, blunt. Basal margin narrow, sinuate, and obliterated in the middle. Surface with fine and dense punctures.</p><p>Scutellum. Shape subtriangular, surface sparsely punctate; ratio width/length 1.6.</p><p>Elytra with striae defined by regular rows of punctures. Marginal membrane very narrow.</p><p>Pygidium strigate. In lateral view slightly convex. Triangular in shape; with short setae on disc, long setae at apical margin.</p><p>Mesosternal process long, with tapering apex in ventral view; width at base: 1.0 mm (Fig. 25). Mesometasternal suture at about half the length of the process.</p><p>Abdominal sternites with 1–3 rows of setae, denser at sides, and 4–6 rows of punctures per sternite. Terminal sternite strigate, with apical margin thick and sinuate; apical membrane wide.</p><p>Legs. Protibia bidentate (Fig. 11); apical tooth long and straight; basal tooth below internal apex of protibia triangular in shape, obtuse. Metatibia slender, slightly narrower subapically. Ratio length/width 4.1. First external carina well developed. Surface rugose. Protarsomere 1 equal in length to protarsomeres 2–4, tarsomeres stout. Protarsal claws: external claw curved, internal claw bifurcate, upper branch equal in length and 2/3 the width of the lower branch, inferior margin sinuate.</p><p>Aedeagus (Fig. 17): parameres with a narrow and acute apex in a lateral view, pointing downwards, flattened dorsoventrally; apical portion of ventral plate membranous. Endophallus (Fig. 23) short, with patches of short and long setae in different positions. Ejaculatory duct opening frontal.</p><p>Variation. Background color from pale ochre to orange; transversal bands on elytra more or less evident. Female similar to male, antennal club shorter than in male; protibia with longer and wider apical tooth (Fig. 11); internal protarsal claw narrower with curved inferior margin; terminal sternite with slightly sinuate apex.</p><p>Body length 17.5–20.1 mm, body width 9.8–11.4 mm. Clypeus w/l: 1.9–2.1. Interocular ratio (interocular width/width of eye): 3.2–4.2. Pronotum w/l: 1.8–1.9. Scutellum w/l: 1.5–1.6. Pygidium w/l: 1.5–1.6. Metatibia w/l: 3.8–4.1. Width of mesosternal process at base: 1.0– 1.2 mm.</p><p>Diagnosis. Size large, color pale ochre with darker bands on pronotum and elytra, clypeus long, head large, pronotum short, mesosternal process long, parameres flattened dorsoventrally.</p><p>Etymology. Latinized from the name of genus Rutela and suffix -oides, for its general resemblance with species of the tribe Rutelini .</p><p>Distribution. Guanacaste and Talamanca mountain ranges, Costa Rica and northern Panama from 900–1600 m, (Fig. 36).</p><p>Note. This species is tentatively placed in the genus Callistethus, as it has some distinct features not found in other Callistethus species. It could be placed in a separate genus, but the lack of known similar species would make a proper description and diagnosis of the new genus difficult. Singular characters are: long clypeus (nearly as long as frons); head large (2/3 of pronotum width); mandibles with apex curved ventrally; maxillae with second and third tooth fused together; labrum and labium quadrilobulated; pronotum short; tarsomeres short and stout; mesometasternal suture high, nearly at half the length of the mesosternal process; aedeagus with parameres flattened dorsoventrally.</p></div>	https://treatment.plazi.org/id/03E1981EB53BFFEFFF59FCAD00E4EC88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
03E1981EB53DFFEDFF59FEF405B6EF01.text	03E1981EB53DFFEDFF59FEF405B6EF01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus yalizo Filippini, Galante	<div><p>Callistethus yalizo Filippini, Galante, &amp; Micó, new species</p><p>Fig. 6</p><p>Type material. Holotype: ♂ "P.N. Tapanti, La Represa, A. C. Amistad, Prov. Carta, COSTA RICA. 1650 m. 23 Mar 1994, G. Mora, A. Solis, E. Ulate, L N 185900_563360 # 2783 / INBIOCRI001964129" deposited in INBIO . Paratypes (8): 1♂ "R. Grande de Orosi, desde Puente R. Dos Amigos hasta la Represa, Prov. Carta, COSTA RICA. 1400-1800m. MAR 1995. R. Delgado. L_N_186600_562000 #4418 / INBIOCRI002253282"; 1♂ " Costa Rica, Cartago Prov., Tapanti N. Pk. Dam area, vi-4-1997 RW Hamilton / RWHC " INBIO; 1♂ "Cerro Chompipe, Res. Biol. Chompipe, Prov. Here, COSTA RICA, J. F. Corrales, 7 Abr 1991, L- N 230000_528000 / INBIOCRI000183743"; 1♀ "Quebrada Segunda, P. N. Tapanti, 1250M, Prov. Cartago, Costa Rica, Mar 1992, R. Vargas, L-N 194000,560000 / INBIOCRI000894947"; 1♂ "Esperanza del Guarco, Cartago, Costa Rica. 2300 m 5/ 6/2008 Leg. A. García, M.Zumbado / CEUA 00106167"; 1♀ " COSTA RICA. Prov. Cartago. P.N. Tapanti. La Esperanza del Guarco. 2334m. 5 MAY 2008. A. García, M.A. Zumbado. Tp. Luz 2. L_N_188418_552219 #93739 / CEUA 00106168"; 1♂ "V. Platanar P.N. Juan Castro Blanco (Costa Rica) 1800 m 29-1-2006. T. luz (18:30-21:00). Leg.: Micó, García, Galante / CEUA 00003201"; 1♂ " MUSEO DE INSECTOS UNIVERSIDAD DE COSTA RICA. COSTA RICA, PROV. Cartago. Río Macho, Orosí. 4 may. 1973. E. Monge." MUCR.</p><p>Holotype male description. Body shape oval. Length 15.1 mm; width 8.6 mm.</p><p>Color. Head, pronotum, and scutellum metallic green. Margins of pronotum yellow, with green/bronze luster. Elytra and pygidium metallic green with brown luster. Metasternum metallic green, legs (including coxae) and lateral sternites yellow, with green base and apex; protibiae and tarsi metallic green, claws red. Abdominal sternites reddish brown with green/bronze luster. Antennae red with club dark brown.</p><p>Clypeus nearly rectangular, ratio width/length 2.3; surface granulate; anterior margin slightly sinuate. Frons with dense, coarse punctures; rest of head with sparse, fine punctures. Ocular canthus long, tapering towards apex; apex acute. Eyes small, interocular ratio (interocular width/width of eye) 3.0. Antenna: ratio funiculus/club 0.6.</p><p>Pronotum width 1.8 times length; in dorsal view broadest at base. Lateral margins rounded. Anterior angles quadrate, with rounded vertex; posterior angles obtuse, with rounded vertex. Basal margin without subapical sulcus, sinuate. Entire surface with small, shallow, sparse punctures.</p><p>Scutellum. Shape subpentagonal, with a median sulcus and sparse, deep punctures on the entire surface; ratio width/length 1.5.</p><p>Elytra. Surface with costae defined by rows of shallows punctures. Interstices with 3–4 (1st interstice) or 1–2 (2nd–5th interstices) rows of sparse shallow punctures.</p><p>Pygidium granulate with a shallow median sulcus, setae short on disc and longer on margins. Ratio width/ length 1.6.</p><p>Mesosternal process short, not going beyond anterior margin of mesocoxae, uniformly wide (half as wide as metafemura) with rounded apex; width at base: 0.6 mm (Fig. 26). Mesometasternal suture well defined. Abdominal sternites with sparse, shallow punctures; laterally forming 6–7 rows of punctures. Short, blonde setae arranged in 1 row per sternite, denser on sides (in 2–3 rows). Terminal sternite with sparse, shallow punctures; apical margin height 2/3 of basal portion, surface smooth; apical membrane narrow. Abdominal spiracle 6th convex.</p><p>Legs. Protibia bidentate (Fig. 12); apical tooth long, oblique in relation to the vertical axis of protibia, nearly straight. Basal tooth obtuse, blunt, positioned at same level with internal apex of protibia. Metatibia very thin, slightly narrower subapically. Ratio length/width 5.1. Both external carinae well defined. Surface punctate. Protarsal claws: external claw curved; internal claw bifurcate, with branches strongly diverging; upper branch of equal length and 2/3 the width of the lower branch, inferior margin sinuate.</p><p>Aedeagus (Fig. 18). Parameres nearly bilobed in a lateral view, ventral plate partially fused with parameres. Endophallus (Fig. 24): with 1 long, sinuate sacculus, with a row of small, globoid diverticles in basal half, partially covered with setae; ejaculatory duct opening ventrally at base, laterally with a small diverticle covered in long setae.</p><p>Variation. Elytra from red with green luster to metallic green. Abdominal sternites from ochre to dark brown with green or bronze luster. Female similar to male. Antennal club slightly shorter than in male. Eyes smaller than in male. Protibia (Fig. 12): apical tooth longer and wider, basal tooth above the internal apex. Protarsal claw: internal claw narrower than in male, bifurcation narrower, inferior margin curved. Margin of terminal sternite narrower than in male.</p><p>Body length 15.1–16.2 mm; width 8.6–9.5 mm. Clypeus w/l: 2.2–2.3. Interocular ratio: 3.0–3.1 (male), 3.5–4.2 (female). Antenna: ratio funiculus/club: 0.6–0.8. Pronotum w/l: 1.8–1.9. Scutellum w/l: 1.3–1.5. Pygidium w/l: 1.5–1.7. Width of mesosternal process at base: 0.4–0. 6 mm. Metatibia w/l: 4.6–5.4.</p><p>Diagnosis. Metallic green color; elytra with shallow punctures that give an irregular appearance; mesosternal process short; parameres nearly bilobed in a lateral view, ventral plate partially fused with parameres.</p><p>Etymology. Latinized from the Greek verb yalízo, to be green like glass, treated as a noun in apposition, for the green shiny color of this species.</p><p>Distribution. Central and Talamanca mountain ranges, Costa Rica, from 200–2300 m (Fig. 37).</p></div>	https://treatment.plazi.org/id/03E1981EB53DFFEDFF59FEF405B6EF01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Filippini, Valentina;Galante, Eduardo;Micó, Estefanía	Filippini, Valentina, Galante, Eduardo, Micó, Estefanía (2015): Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1), DOI: 10.11646/zootaxa.3948.1.2
