taxonID	type	description	language	source
03E1D765D54F510EFF72FB5509BD0BF4.taxon	description	Figs 1 A, 3 A, 12 A	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54F510EFF72FB5509BD0BF4.taxon	materials_examined	Material studied. Holotype male. CANADA, Ontario, 7 mi S of Griffith, 10. VII. 1991, B. E. Cooper (in MZH, http: // id. luomus. fi / GE. 1808). Paratype. CANADA, Ontario, Sudbury Co., Chapleau, Northeast Superior Forest, 47.573 ° N, 82.859 ° W, mature (99 years old) stand composed of 90 % Pinus banksiana and 10 % Picea mariana with understory shrub layer of Vaccinium, very wet with considerable bryophyte coverage, Pinus banksiana log, photoeclector, 22. VII. 2013, R. Deady, 1 male (in SDEI).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54F510EFF72FB5509BD0BF4.taxon	description	Description. Male. Head. Face and antenna concolorous pale brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 11 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 setae, with large dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.8 x as long as wide, neck slightly longer than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 2 setae. Proepisternum with 5 setae. Scutellum with 2 long, 2 moderately long and some short and fine setae. Wing. Hyaline. Length 2.1 – 2.2 mm. Width / length 0.45. Anal lobe small. Veins distinct, except for stM. c / w 0.60. R 1 / R 0.60. stM longer than fork of M. bM shorter than r-m, stCuA shortest. bM and r-m non-setose. Halters of the studied specimen missing. Legs. Pale brown; setae pale. Fore tibial organ forming small patch of some setae in demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale, short and fine. Hypopygium (Fig. 1 A). Brown, like abdomen. Intergonocoxal area moderately long, with 2 indistinct lobes each with 4 long setae. Gonocoxa broad, longer than gonostylus, with sparse and fine setosity. Gonostylus (Fig. 3 A) wide, roundish laterally, convex medially, slightly narrowed apically; with short and dense setosity apically, with straight and conical apical tooth, with 1 megaseta on apical, 1 on dorsal, 1 on ventral side of apical tooth, with 3 subapical megasetae very near apical tooth; slightly differentiated whip-lash seta on apical quarter of gonostylus. Tegmen (Fig. 12 A) slightly shorter than wide, roundish and membraneous apically, straight and strongly sclerotized laterally, with long apodemes, and a large area of aedeagal teeth. Aedeagal apodeme rather short. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54F510EFF72FB5509BD0BF4.taxon	discussion	Discussion. Lycoriella ampla sp. n. has long antennae, a narrow eye bridge with one or two rows of facets, an extremely short maxillary palpus with a strongly deepened sensory pit (2 nd and 3 rd palpal segments are short and oval and together as long as 1 st segment), basally narrowed wings, a very short wing vein R 1, and a small demarcated fore tibial organ. The intergonocoxal area has apically two small groups of only four or five closely placed setae, the medial side of the gonostylus is straight, with a relatively short whip-lash seta on its apical half, and a slender apical megaseta. These characters are shared with Lycoriella micria Mohrig & Menzel, 1990. Lycoriella ampla differs from L. micria in having longer antennal flagellomeres, a strongly widened gonocoxa and gonostylus, apically roundish tegmen with straight sclerotized lateral sides, a nearly straight apical tooth, and a higher number of apical-subapical gonostylar megasetae (see the description). Lycoriella micria has a narrower gonocoxa and gonostylus with more widely placed and stronger subapical megasetae, a wider and straighter apex of the tegmen, more indistinct groups of setae in the intergonocoxal area, and a more strongly curved apical tooth of the gonostylus with only one apical and three to four subapical megasetae. The extremely voluminous gonostylus of L. ampla, with its extremely widened lateral side, resembles that of Bradysiopsis vittigera (Zetterstedt, 1851) and the unusually sclerotized lateral sides of its tegmen, unknown in any other known species of Lycoriella, resemble those of Hemineurina riparia (Holmgren, 1883) [see Figs 159 and 374 in Menzel & Mohrig (2000)].	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54F510EFF72FB5509BD0BF4.taxon	etymology	Etymology. The name is Latin, ampla, widened, referring to the widened gonostylus of the male hypopygium.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54C5108FF72FC910D6B0B1C.taxon	description	Figs 1 B, 3 B	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54C5108FF72FC910D6B0B1C.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 12 – 22. VII. 2010, A. Barkalov (in ISEA). Paratypes. RUSSIA, same data as holotype, 2 males (1 in MZH, http: // id. luomus. fi / GE. 1810; 1 in SDEI); same data but 9 – 20. VII. 2010, 2 males (1 in MZH, http: // id. luomus. fi / GE. 1809; 1 in SDEI); Krasnodarsk region, Taimyr Peninsula, River Zakharova Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 1 male (in ISEA).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54C5108FF72FC910D6B0B1C.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 3 facets wide. Face with 20 – 26 setae. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 5 – 7 setae, with small dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.40 – 2.95 x as long as wide, neck as long as wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 4 – 7 setae. Proepisternum with 3 – 10 setae. Scutellum with 2 long and some short and fine setae. Wing. Hyaline. Length 1.6 – 1.8 mm. Width / length 0.40. Anal lobe small. Veins distinct, except for stM. c / w 0.65 – 0.75. R 1 / R 0.70 – 0.85. stM longer than fork of M. bM longer than r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming large patch of many setae in demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately long. Hypopygium (Fig. 1 B). Brown, like abdomen. Intergonocoxal area short, without lobe or seta group. Gonocoxa narrow, longer than gonostylus, with dense and moderately long setosity, at medial margin short. Gonostylus (Fig. 3 B) very narrow, widest basally, distinctly narrowed in basal third, impressed medially at the narrowed part; apex densely setose, with strong apical tooth; with 3 – 4 medial megasetae in apical third, megasetae narrow, slightly curved, slightly shorter than apical tooth, orientation variable; with well-differentiated long whip-lash seta in middle of gonostylus. Tegmen wider than long, truncate apically, curved basolaterally, weakly sclerotized, with small area of aedeagal teeth. Aedeagal apodeme rather short. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54C5108FF72FC910D6B0B1C.taxon	discussion	Discussion. Lycoriella barkalovi sp. n. is very similar to L. lundstromi (Frey, 1948): Both species have the eye bridge with two or three rows of facets, a large, basally bow-formed, distinctly demarcated fore tibial organ, lack the intergonocoxal lobe and have a very narrow gonostylus with a few megasetae (spines) latero-medially. In Lycoriella barkalovi the wing is much narrower due to the reduced anal lobe, bM is longer than r-m, coxae and legs are darker brown, the 2 nd and 3 rd palpal segments are subequal in length and the antennal flagellomeres have only some short sensilla. The medial margin and the membraneous area of the gonocoxa have a strikingly short and dense setosity. Furthermore, L. barkalovi has only three or four short and strong megasetae (spines) in the apical half of the gonostylus, the megasetae are all evenly directed basad, the apex of gonostylus is richly setose and the apical part of the gonostylus more impressed and appearing much narrower than the basal part. By comparison, in L. lundstromi the wing is broad with a well-developed anal lobe, bM and r-m are subequal in length, coxae and legs are yellowish (at least coxae yellow), the medial margin and the membraneous area of gonocoxa have only a few weak setae, antennal flagellomeres have long, strongly curved and brownish setae (the vestiture of flagellomeres appears distinctly uneven; the 2 nd segment of maxillary palpus short and the 3 rd segment distinctly longer; the gonostylus is evenly impressed and narrowed towards apex; the gonostylar apex is less setose with weaker setae; the medial side of the gonostylus has four to six megasetae, one to three of which are shorter and located near the base of the apical tooth and three to five are longer, in the apical half of the gonostylus (often two megasetae on the middle of the gonostylus are directed towards each other and may cross).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54C5108FF72FC910D6B0B1C.taxon	etymology	Etymology. The species is named after the Russian dipterist A. V. Barkalov (Novosibirsk), who collected the type specimens.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54A510AFF72FDB90FA50E27.taxon	description	Figs 2 A, 3 C	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54A510AFF72FDB90FA50E27.taxon	materials_examined	Material studied. Holotype male. CANADA, British Columbia, Lake Kinbasket, Canoe Reach of Kinbasket Reservoir, Malaise trap, 21. VI. 2010, Cooper Beauchesne and Associates Ltd. (in RBCM).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54A510AFF72FDB90FA50E27.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 3 facets wide. Face with 12 long and short setae. Clypeus with 1 seta. Maxillary palpus with 3 segments; segment lengths not measurable in the specimen studied; 1 st segment with 3 setae and dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.75 x as long as wide, neck shorter than wide, longest setae slightly shorter than width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 4 setae. Proepisternum with 2 setae. Scutellum with 2 long and some short setae. Wing. Hyaline. Length 1.5 mm. Width / length 0.50. Anal lobe small. Veins distinct, except for stM. c / w 0.55. R 1 / R 0.65. bM shorter than r-m, stCuA shortest. stM subequal to fork of M. bM and r-m non-setose. Halter yellow. Legs. Pale brown, coxae yellow; coxal setae pale. Fore tibial organ forming very small patch of few setae in demarcated depression. Fore tibial spur longer than apical width of tibia. Abdomen. Pale brown; setae pale and long. Hypopygium (Fig. 2 A). Pale brown, like abdomen. Intergonocoxal area with distinct broad lobe with apical setae. Gonocoxa longer than gonostylus, with rather short and sparse vestiture. Gonostylus (Fig. 3 C) wide, narrowed apically, impressed medially, with dense apical setosity, with long and narrow apical tooth, with 4 long and straight medial megasetae in apical fourth; with indistinct whip-lash seta in middle of gonostylus. Tegmen slightly sclerotized, wider than long, with small area of aedeagal teeth. Aedeagal apodeme rather strong. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54A510AFF72FDB90FA50E27.taxon	discussion	Discussion. Lycoriella canningsi sp. n. belongs to the group of Lycoriella which share short and broad wings, a small fore tibial organ, a large Trichocoelina - like and medially impressed gonostylus, four conical basomedially directed and slightly curved megasetae as well as a large, broad and roundish intergonocoxal lobe of with strong setae. Among these species, Lycoriella canningsi most resembles the Palaearctic L. latilobata Menzel & Mohrig, 2000 because of its pale body setosity, the eye bridge with two to three rows of facets, relatively densely setose antennal flagellomeres with only few sensilla, a similarly long 4 th flagellomere and a fairly long maxillary palpus. Lycoriella canningsi differs from L. latilobata in its wider gonostylus, a shorter whip-lash seta basally on the medial margin of the gonostylus, a longer apical tooth placed at the very apex of the gonostylus, the broad and short tegmen, stronger gonostylar megasetae, and a longer intergonocoxal lobe with weaker setae (all setae are curved and not spine-like). By comparison, L. latilobata has a distinctly narrower gonostylus, a longer whip-lash seta, the apical tooth placed subapically on the medial side of the gonostylus, the tegmen much longer and conical, shorter and narrower gonostylar megasetae and a shorter, a bow-shaped intergonocoxal lobe with apical margin with dense, straight, spine-like setae.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D54A510AFF72FDB90FA50E27.taxon	etymology	Etymology. The species is named after the Canadian entomologist Robert A. Cannings, formerly the Curator of Entomology in the Royal British Columbia Museum, Victoria, BC.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5485104FF72F8A70E120C34.taxon	description	Figs 4 B, 6 C, 13 A	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5485104FF72F8A70E120C34.taxon	materials_examined	Material studied. Holotype male. CANADA, Quebec, Old Chelsey, 3. VIII. 1958, J. R. Vockeroth (in CNC).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5485104FF72F8A70E120C34.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 3 facets wide. Face with 14 setae. Clypeus with 4 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 5 setae, with small dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.2 x as long as wide, neck slightly shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 5 setae. Proepisternum with 8 setae. Scutellum with 2 long and some short and fine setae. Wing. Fumose. Length 1.7 mm. Width / length 0.45. Anal lobe small. Veins distinct, except for stM. c / w 0.75. R 1 / R not detectable in the specimen studied. stM shorter than fork of M. bM and r-m subequal in length, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming large patch of many setae in demarcated depression. Fore tibial spur slightly shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately long. Hypopygium (Fig. 4 B). Brown, like abdomen. Intergonocoxal area rather long, with short setose lobe. Gonocoxa longer than gonostylus, slightly roundish laterally, with rather dense and fine setosity. Gonostylus (Fig. 6 C) widest subbasally, narrowed towards apex, strongly impressed medially, densely setose apically, with strong and curved apical tooth; with 5 medial megasetae in apical half, megasetae slender, long and straight, oblique, the apicalmost (with sockets) shorter than the others, as long as apical tooth; with well-differentiated long whip-lash seta in basal third of gonostylus. Tegmen (Fig. 13 A) much wider than long, smoothly curved and membraneous apically, straight laterally, sclerotized basolaterally, with small area bearing a few aedeagal teeth. Aedeagal apodeme moderately long. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5485104FF72F8A70E120C34.taxon	discussion	Discussion. Lycoriella eurystylata sp. n. resembles L. taimyrensis sp. n. and L. tundrae sp. n. in having the tegmen smoothly curved apicolaterally, whereas L. acutostylia Mohrig & Menzel, 1990 [described in Menzel et al. (1990)], L. jakovlevi sp. n. and L. kinbasketi sp. n. have the tegmen distinctly angled apicolaterally (Fig. 13). L. eurystylata differs from L. taimyrensis and L. tundrae in having the tegmen very short, strongly sclerotized basolaterally, semicircular apically and without tooth-like structures, whereas the other two have their tegmen only slightly sclerotized laterally, less curved apically and with tooth-like structures. Furthermore, L. eurystylata has five, L. tundrae four and L. taimyrensis eight to nine gonostylar megasetae and the fore tibial organ of L. eurystylata is more distinctly demarcated than that of L. taimyrensis and L. tundrae. L. eurystylata is similar to L. acutostylia, L. jakovlevi and L. kinbasketi in having rather long antennal flagellomeres (4 th flagellomere 2.1 – 2.3 times as long as wide), whereas L. taimyrensis and L. tundrae have shorter flagellomeres (4 th flagellomere 1.4 – 1.6 times as long as wide). To distinguish L. taimyrensis and L. tundrae, see under the former, and to distinguish L. acutostylia, L. jakovlevi and L. kinbasketi, see under L. jakovlevi sp. n.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5485104FF72F8A70E120C34.taxon	etymology	Etymology. The name is derived from the Greek words eurys, broad, and stylos, style, referring to the thick gonostylus of the male hypopygium.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5465105FF72FA510D380BF4.taxon	description	Figs 2 B, 3 D, 12 B	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5465105FF72FA510D380BF4.taxon	materials_examined	Material studied. Holotype male. USA, Colorado, Laramer Co., Pinewood Springs, 40.2677 ° N, 105.3678 ° W, Malaise trap, 26. VII – 7. IX. 2014, S. & B. Fitzgerald (in MZH, http: // id. luomus. fi / GE. 1811).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5465105FF72FA510D380BF4.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 1 – 2 facets wide. Face with 27 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 setae, with deep dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.8 x as long as wide, neck slightly longer than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 9 setae. Proepisternum with 10 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.9 mm. Width / length 0.40. Anal lobe small. Veins distinct, except for stM. c / w 0.65. R 1 / R 0.75. stM longer than fork of M. bM as long as r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Pale brown; setae pale. Fore tibial organ forming large patch of setae in demarcated depression. Fore tibial spur longer than apical width of tibia. Abdomen. Pale brown; setae pale, short and weak. Hypopygium (Fig. 2 B). Brown, like abdomen. Intergonocoxal area moderately long, with semicircular setose lobe. Gonocoxa narrow and long, longer than gonostylus, with sparse, rather short setosity. Gonostylus (Fig. 3 D) widest subapically, slightly narrowed towards apex, deeply excavated dorsally; densely setose apically, with long and narrow apical tooth, with 1 subapical and 7 – 8 medial megasetae on ventral margin of apical half of gonostylus; with long whip-lash seta in middle. Tegmen (Fig. 12 B) longer than wide, slightly notched apically, weakly sclerotized laterally, with area of minute teeth apicoventrally, with area of small aedeagal teeth. Aedeagal apodeme moderately long. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5465105FF72FA510D380BF4.taxon	discussion	Discussion. Lycoriella excavata sp. n. with its subapically widened, lobe-shaped gonostyli resembles Lycoriella piristylata Vilkamaa, Hippa & Heller, 2013, which has (in contrast to the original description) also a palpal sensory pit and therefore belongs to Lycoriella Frey. In contrast to Lycoriella excavata sp. n., in L. piristylata a long whiplash seta is missing, the intercoxal lobe is small (or absent in some specimens), the apical tooth is significantly shorter or completely reduced and the tegmen is short and trapezoidal (Vilkamaa et al. 2013: fig. 3 A – C). Lycoriella excavata sp. n. (Fig. 2 B) also differs from this species in its ventrally enlarged lobe-shaped gonostylus with a very deep dorsal excavation, all medial megasetae of the gonostylus on its ventral margin, and its tegmen with apicolateral teeth. Apicolateral teeth of the tegmen are currently known only in Lycoriella taimyrensis described here (Fig. 13 E), in some species of angustostylata species group in Bradysia Winnertz, 1867 (Menzel & Heller 2005; Mohrig 2016), as well in Odontosciara Rübsaamen, 1908 (Mohrig 2003).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5465105FF72FA510D380BF4.taxon	etymology	Etymology. The name is a Latin adjective, excavata, meaning hollowed, and refers to the strongly hollowed gonostylus.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5475100FF72FC910FD409D0.taxon	description	Figs 5 A, 6 A, 13 C, 14 B	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5475100FF72FC910FD409D0.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Karelia, Kivach, poplar stand, pitfall trap, 11 – 13. VI. 1986, J. Jakovlev, (in MZH, http: // id. luomus. fi / GE. 1812). Paratypes. RUSSIA, same locality as holotype but pine-lichen habitat, Malaise trap, 5 – 7. VIII. 1986, J. Jakovlev, 1 male (in MZH, http: // id. luomus. fi / GE. 1813); same data as previous but window trap, 8 – 10. VII. 1985, 1 male (in SDEI).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5475100FF72FC910FD409D0.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 26 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer or as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 4 setae, with moderate dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.10 – 2.25 x as long as wide, neck much shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 4 setae. Proepisternum with 4 – 7 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.4 mm. Width / length 0.40 – 0.45. Anal lobe small. Veins distinct, except for stM. c / w 0.70 – 0.75. R 1 / R 0.70 – 0.90. stM as long as fork of M. bM longer than r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming moderate patch of many setae in demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately long. Hypopygium (Fig. 5 A). Brown, like abdomen. Intergonocoxal area (Fig. 14 B) moderately long, with semicircular setose lobe. Gonocoxa longer than gonostylus, rather straight laterally, with dense and moderately long setosity, shorter on medial margin. Gonostylus (Fig. 6 A) widest subbasally, gradually tapered towards apex, strongly impressed medially, densely setose apically, with a strong slightly curved apical tooth, 2 pairs of medial megasetae on apical third, megasetae slender, shorter than apical tooth, slightly procurved; with well-differentiated long whip-lash seta in middle of gonostylus. Tegmen (Fig. 13 C) as long as wide, with distinct and sharp apicolateral angles; surrounded by hyaline membrane laterally (and ventrally?), membraneous apically, sclerotized laterally, with indistinct sclerotizations dorsally, with small area of aedeagal teeth. Aedeagal apodeme rather short. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5475100FF72FC910FD409D0.taxon	discussion	Discussion. This species is characterised by its apically gradually strongly narrowed gonostylus with only four to five medial megasetae and its lack of any apical megasetae. Lycoriella jakovlevi sp. n. resembles L. acutostylia Menzel & Mohrig, 1990, L. kinbasketi sp. n. and L. tundrae sp. n. Lycoriella jakovlevi is closer to L. acutostylia and L. kinbasketi and differs from L. tundrae in having the tegmen distinctly angled apicolaterally. The tegmen of Lycoriella jakovlevi is about as long as wide, whereas the tegmen of L. acutostylia and L. kinbasketi is shorter than wide. Furthermore, the intergonocoxal lobe L. acutostylia is very short, and hardly produced from the level of the basomedial margin of the gonocoxae, whereas L. jakovlevi has a longer lobe, nearly semi-circular, and L. kinbasketi ’ s lobe is long but narrowed apically (Fig. 14). Unlike Lycoriella jakovlevi and L. kinbasketi, the two basalmost medial megasetae of the gonostylus of L. acutostylia are not parallel, but diverge, the whip-lash setae are positioned more basally, and the basal rim of the hypopygium turned caudad medially (Figs 4 A, 6 B, 6 C). The gonostyli of Lycoriella jakovlevi and that of L. kinbasketi are almost identical, both having four gonostylar megasetae on the apical quarter of the gonostylus but L. jakovlevi has a longer apical tooth of gonostylus. The differences to Lycoriella eurystylata sp. n. and L. taimyrensis sp. n. are discussed elsewhere.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5475100FF72FC910FD409D0.taxon	etymology	Etymology. The species is named after the Finnish mycetophilid specialist Jevgeni Jakovlev (Finnish Environment Institute, Helsinki, Finland), who collected the type specimens.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425100FF72FEF509BD0DEC.taxon	description	Figs 5 B, 6 B, 13 D, 14 C	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425100FF72FEF509BD0DEC.taxon	materials_examined	Material studied. Holotype male. CANADA, British Columbia, Canoe Reach of Kinbasket Reservoir near Valemount, Malaise trap, 21. VI. 2010, V. Prigmore, V. Smith., N. Shaw & H. Schindler (in MZH, http: // id. luomus. fi / GE. 1814). Paratype. CANADA, same data as holotype, 1 male (in SDEI).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425100FF72FEF509BD0DEC.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 25 setae. Clypeus with 3 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 4 – 5 setae, with a moderate dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.10 – 2.30 x as long as wide, the longest setae longer than the width of the flagellomere, neck shorter than wide. Thorax. Brown; setae pale. Anterior pronotum with 5 – 8 setae. Proepisternum with 8 – 9 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.4 mm. Width / length 0.40. Anal lobe small. Veins distinct, except for stM. c / w 0.70 – 0.75. R 1 / R 0.45. stM as long as fork of M. bM longer than r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Brown, coxae yellow; setae pale. Fore tibial organ forming moderate patch of many setae in demarcated depression. Fore tibial spur shorter than tibial width. Abdomen. Pale brown; setae pale and moderately long. Hypopygium (Fig. 5 B). Brown, like abdomen. Intergonocoxal area (Fig. 14 C) moderately long, with a triangular setose lobe. Gonocoxa longer than gonostylus, roundish laterally, with dense and short setosity. Gonostylus (Fig. 6 B) widest subbasally, strongly narrowed towards apex, strongly impressed dorsally, densely setose apically, with a rather short and strong apical tooth; with 4 medial megasetae at apical fourth, 2 basalmost ones close to each other, megasetae slender, nearly straight; with well-differentiated long whip-lash seta near the middle of the gonostylus. Tegmen (Fig. 13 D) surrounded by hyaline membrane laterally (and ventrally?); shorter than wide wide, membraneous apically, sclerotized laterally, with indistinct sclerotizations dorsomedially, with a large area of aedeagal teeth. Aedeagal apodeme moderately long. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425100FF72FEF509BD0DEC.taxon	discussion	Discussion. See under Lycoriella eurystylata sp. n. and L. jakovlevi sp. n.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425100FF72FEF509BD0DEC.taxon	etymology	Etymology. The species is named after the 19 th- century Chief Kinbasket of the Columbia River Shushap people.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425102FF72FAE90DD70B1C.taxon	description	Figs 7 A, 9 A	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425102FF72FAE90DD70B1C.taxon	materials_examined	Material studied. Holotype male. USA, Colorado, Laramer Co., Pinewood Springs, Malaise trap, 12. IX – 1. XII. 2014, S. & B. Fitzgerald (in MZH, http: // id. luomus. fi / GE. 1815). Paratypes. USA, Colorado, Pinewood Springs, 40.259476 ° N, 105.360503 ° W, wood along small creek, Malaise trap, 7. IX – 12. X. 2014, S. & J. Fitzgerald, 5 males (3 in MZH, http: // id. luomus. fi / GE. 1816, http: // id. luomus. fi / GE. 1817 and http: // id. luomus. fi / GE. 1818; 2 in SDEI).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425102FF72FAE90DD70B1C.taxon	description	Description. Male. Head. Face paler brown than antenna, maxillary palpus pale brown. Eye bridge 2 – 3 facets wide. Face with 15 – 30 setae. Clypeus with 2 – 5 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 5 setae, with small dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.35 – 2.50 x as long as wide, neck shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 4 – 6 setae. Proepisternum with 6 – 11 setae. Scutellum with 2 moderate and some short and fine setae. Wing. Hyaline. Length 1.4 mm. Width / length 0.45. Anal lobe rather small. Veins distinct, except stM. c / w 0.55 – 0.70. R 1 / R 0.65 – 85. stM shorter than fork of M. bM shorter, longer or as long as r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Yellow; setae pale. Fore tibial organ forming large patch of many setae in demarcated depression. Fore tibial spur as long as apical width of tibia. Abdomen. Pale brown; setae pale, short and fine. Hypopygium (Fig. 7 A). Brown, like abdomen. Intergonocoxal area moderate, with short setose lobe. Gonocoxa narrow, much longer than gonostylus, with dense and short setosity. Gonostylus (Fig. 9 A) very narrow, evenly narrowed towards apex, strongly impressed medially; densely setose apically, with long apical tooth; with 2 subapical and 3 medial megasetae in medial impression, 2 of medial megasetae close to subapical megasetae, 1 longer and stronger one near middle of gonostylus, megasetae straight or very slightly curved, all shorter than apical tooth; with well-differentiated long whip-lash seta in middle of gonostylus. Tegmen broadly conical, membraneous apically, weakly sclerotized laterally, with small area of aedeagal teeth. Aedeagal apodeme rather long. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425102FF72FAE90DD70B1C.taxon	discussion	Discussion. With its slender gonostylus, Lycoriella longa sp. n. possesses the typical gonostylar structure of the genus, but differs in having its gonostylus distinctly impressed dorsomedially and in having a single conspicuous medial megaseta in the middle of the gonostylus, well separated from the other megasetae.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5425102FF72FAE90DD70B1C.taxon	etymology	Etymology. The name is a Latin adjective, longa (long), referring to the long gonostylus, the long medial megaseta and the long antenna.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D540511DFF72FDB90DB50A3C.taxon	description	Figs 8 A, 9 B Literature. Lycoriella (Lycoriella) sp. n. — Wirta et al. (2016): supporting information, table S 1, fig. S 1.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D540511DFF72FDB90DB50A3C.taxon	materials_examined	Material studied. Holotype male. CANADA, Quebec, Great Whale River, 4 – 9. VIII. 1949, J. R. Vockeroth, 1 male (in CNC). Paratypes. CANADA, same data as holotype, 1 male (in MZH, http: // id. luomus. fi / GE. 1899); Nunavut, Ellesmere Island, ‘ Fosheim Pns. ’ [= Fosheim Peninsula], Hot Weather Creek, 79 ° 58 ’ N, 84 ° 28 ’ W, 22. VII. 1990, F. Brodo (in MZH, http: // id. luomus. fi / GE. 1821); Nunavut, Ellesmere Island, Alert, 82.30.096 ° N, 62.20.902 ° W, 31. VII. 1963, J. R. Vockeroth, 1 male (in CNC); Quebec, Payne Bay, 19. VII. 1958, J. R. Vockeroth, 1 male (in CNC); N. E. GREENLAND, Kap Köbenhavn, 68 ° 30 ’ N, 22 ° 30 ’ W, VII. 1986, J. Böcher (in ZMUC); same data but 68 ° 30 ’ N, 22 ° 34 ’ W 13. VII. 1986, 1 male (in ZMUC); Zackenberg Valley, Northeast Greenland National Park, 74 ° 28 ’ N, 20 ° 34 ’ W (UTM 8265758: 05147529), 14 m, Malaise trap, 2 – 11. VII. 2011, T. Roslin & G. Varkonyi, 1 male (hypopygium only) (in MZH, http: // id. luomus. fi / GE. 1819, BOLD sample ID GRPV 15, sequence ID GRAFW 2354 - 13); same data but UTM 82654758: 0514752, 18 – 24. VI. 2011, 1 male (in MZH, http: // id. luomus. fi / GE. 1825); same data but UTM 8264450: 0512722, 35 m, 12 – 20. VII. 2011, 1 male (in SDEI); same locality as previous but without date, J. Böcher, 1 male (in ZMUC); N. GREENLAND, ‘ Nedre Midsommer Sö’ [= Nedre Midsommer Sø], 82.0980 ° N, 35.9498 ° W, 10. VII. 1966, Canadian Peary Land Expedition, 1 male (in MZH, http: // id. luomus. fi / GE. 1820); same data but 14. VII. 1966, 1 male (in SDEI); E. GREENLAND, Jameson Land, Muslingeelvdal, 71 ° 23 ’ N, 24 ° 38 ’ W, 1994, J. Böcher, 1 male (in ZMUC); GREENLAND, ‘ Sydl. Zool. Station’ [= Sydlig Zoologisk Station], ‘ 597 ’, 28. VI. 1974, collector unknown, 1 male (in ZMUC); RUSSIA, Arkhangelsk oblast, Yamalo-Nenets Autonomous Okrug, near Seyakh, 70.7788 ° N, 72.0750 ° E, nival meadow, Malaise trap, 13 – 29. VII. 2014, N. Zubryi, 1 male (in MZH, http: // id. luomus. fi / GE. 1900).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D540511DFF72FDB90DB50A3C.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 – 3 facets wide. Face with 32 – 45 setae. Clypeus with 1 – 2 setae or non-setose. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 4 – 6 setae, with large dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.3 – 2.7 x as long as wide, neck shorter than wide, longest setae as long as width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 4 – 7 setae. Proepisternum with 6 – 13 setae. Scutellum with 2 moderate and some short and fine setae. Wing. Hyaline. Length 2.1 – 2.5 mm. Width / length 0.40. Anal lobe small. Veins distinct, except for stM. c / w 0.60 – 0.75. R 1 / R 0.6 – 1.0. stM longer than fork of M. bM and r-m subequal in length, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Yellow. Fore tibial organ forming large distinct patch of fine setae in demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale and short. Hypopygium (Fig. 8 A). Brown, like abdomen. Intergonocoxal area moderately long, with wide setose lobe. Gonocoxa wide, longer than gonostylus, with sparse setosity, shortest in apical quarter. Gonostylus (Fig. 9 B) long and narrow, narrowed apically, impressed medially; with short setosity, densely setose apically, with strong apical tooth and 10 – 12 medial megasetae, megasetae straight, shorter than apical tooth; with well-differentiated whip-lash seta on basal third of gonostylus. Tegmen longer than wide, apically roundish, straight laterally, weakly sclerotized, with area of small aedeagal teeth. Aedeagal apodeme long and strong. BIN. BOLD: AAL 7874.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D540511DFF72FDB90DB50A3C.taxon	discussion	Remarks. The barcoded male of Lycoriella nivicola sp. n. from Greenland with the specimen ID GRPV 15 (Wirta et al. 2016) was identified by us and is designated as a paratype (see above). However, 225 further barcoded specimens with the BIN BOLD: AAL 7874 are found on BOLD, collected in N. E. Greenland (56), the United States (1) [Alaska] and Canada (168) [Alberta, British Columbia, Manitoba, Newfoundland and Labrador, Northwest Territories, Nunavut Territory, Quebec, Saskatchewan, Yukon Territory]. Discussion. Lycoriella nivicola resembles L. suboptica Mohrig & Mamaev, 1990, described from one specimen from the European part of Russia, in having a similar structure of the eye bridge, maxillary palpus and antenna (narrow eye bridge with one or two rows of facets, a thickened 1 st palpal segment with a large, deep sensory pit, 2 nd and 3 rd palpal segments short, and rather long antennal flagellomeres). Lycoriella nivicola has a large, distinct intergonocoxal lobe with about 20 setae, whereas L. suboptica has the lobe short with 10 – 12 setae. In L. nivicola, the medial margin of the gonostylus is less impressed than that in L. suboptica (therefore in the former the gonostylus is broader, especially in its apical half). Lycoriella nivicola has seven to ten hyaline gonostylar megasetae (only four-five mentioned for the holotype of L. suboptica), those close to the apical tooth much much shorter than the others. Furthermore, the new species has a very variable C-w index (0.48 – 0.66; about 0.66 in L. suboptica), the halter is pale (brown in L. suboptica), and the colour of setae on mesonotum, scutellum and abdomen is pale (dark in L. suboptica).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D540511DFF72FDB90DB50A3C.taxon	etymology	Etymology. The name is derived from the Latin words nivis, snow, and - cola, inhabitant, referring to the cold habitats of the species.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55F511EFF72FC590D9C0818.taxon	description	Figs 8 B, 9 D	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55F511EFF72FC590D9C0818.taxon	materials_examined	Material studied. Holotype male. N. GREENLAND, Nedre Midsommer Sö, 82.0980 ° N, 35.9498 ° W, 14. VII. 1966, Canadian Peary Land Expedition (in MZH, http: // id. luomus. fi / GE. 1822).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55F511EFF72FC590D9C0818.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with about 20 setae. Clypeus with 2 setae. Maxillary palpus with 2 segments (2 nd and 3 rd segments partly fused); 1 st segment with 4 setae, with large dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.3 x as long as wide, neck shorter than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 4 setae. Proepisternum with 7 setae. Scutellum with 2 long and some short and fine setae. Wing (in poor condition in the specimen studied). Hyaline. Length 1.9 mm. Width / length and anal lobe not detectable in the specimen studied. Veins distinct, except for stM. c / w 0.65. Halter yellow. Legs. Brown; setae pale. Fore tibial organ forming large patch of setae in demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale and moderately sized. Hypopygium (Fig. 8 B). Pale brown, like abdomen. Intergonocoxal area moderately long, with wide setose lobe. Gonocoxa wide, longer than gonostylus, with sparse setosity, shortest in apical fourth. Gonostylus (Fig. 9 D) long and narrow, apically narrowed, weakly impressed; with short setosity, densely setose apically, with strong apical tooth, 9 – 10 megasetae medially, megasetae straight, shorter than apical tooth; with well-differentiated whip-lash seta on basal third of gonostylus. Tegmen longer than wide, roundish apically, straight laterally, weakly sclerotized, with small area of tiny aedeagal teeth. Aedeagal apodeme long and strong. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55F511EFF72FC590D9C0818.taxon	discussion	Discussion. In having the maxillary palpus 2 - segmented, Lycoriella pearyi sp. n. resembles L. felix (Schmitz, 1919) described from the Netherlands, L. deserticola Mohrig & Mamaev, 1983, described from Uzbekistan, and L. stylata Mohrig & Mamaev, 1985, described from the European part of Russia, all of which may have the palpus either three- or two-segmented. Lycoriella pearyi is similar to L. felix, and differs from L. deserticola and L. stylata in having a distinct intergonocoxal lobe and five megasetae on the apical half of its gonostylus, whereas the other species have at most a row of setae on the intergonocoxal area and two or three subapical gonostylar megasetae. Lycoriella pearyi differs from L. felix in having the eye bridge continuous, with two to three rows of facets, a normal, not a very narrow wing, and an undivided intergonocoxal lobe, whereas L. felix has the eye bridge broken medially, with a discontinuous single row of facets, a strongly reduced wing and distinctly two intergonocoxal lobes. The hypopygium of Lycoriella pearyi is very similar to that of L. nivicola sp. n. but the latter can be distinguished by its two-segmented palpus, narrower gonostylus which is more evenly narrowed towards apex, in having the gonostylar megasetae relatively longer, and in being smaller (wing length 1.9 versus 2.1 – 2.5 mm).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55F511EFF72FC590D9C0818.taxon	etymology	Etymology. The species is named after the polar explorer Robert Edwin Peary (1856 – 1920), one of the pioneers in the study of Greenland.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C511EFF72FEBD0D510F0C.taxon	description	Figs 10 A, 11 A, 13 E	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C511EFF72FEBD0D510F0C.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov (in ISEA). Paratypes. RUSSIA, Krasnoyarsk region, Taimyr Peninsula, on River Zakharova Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 3 males (2 in MZH, http: // id. luomus. fi / GE. 1793 and http: // id. luomus. fi / GE. 1794; 1 in SDEI).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C511EFF72FEBD0D510F0C.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide, medially narrowed and without facets. Face with 16 setae. Clypeus with 2 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 6 setae, with large dorsal pit with setae; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.45 – 1.55 x as long as wide, neck shorter than wide, longest setae shorter than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 5 setae. Proepisternum with 5 setae. Scutellum with 2 long and some short and fine setae. Wing. Fumose. Length 1.4 – 1.5 mm. Width / length 0.45. Anal lobe small. Veins distinct, except for stM. c / w 0.65 – 0.70. R 1 / R 0.60 – 0.70. stM slightly longer than fork of M. bM and r-m subequal in length, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Yellow. Fore tibial organ forming moderate indistinct patch of sensilla in poorly demarcated depression. Fore tibial spur slightly shorter than apical width of tibia. Abdomen. Pale brown; setae pale and short. Hypopygium (Fig. 10 A). Brown, like abdomen. Intergonocoxal area moderately long, with wide setose lobe. Gonocoxa wide, longer than gonostylus, with sparse, rather long and fine setosity. Gonostylus (Fig. 11 A) widest basally, strongly narrowed towards apex, slightly impressed medially; densely setose apically, with strong apical tooth, with 4 – 5 short hyaline megasetae medially and dorsally very near apical tooth, 8 – 9 long and narrow megasetae on high basal bodies in a dense group more basally near whip-lash seta. Tegmen (Fig. 13 E) wider than long, smoothly curved apically, weakly sclerotized, with small teeth apicolaterally; straight laterally, with strongly sclerotized apodemes, with small area of minute aedeagal teeth. Aedeagal apodeme short and strong. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C511EFF72FEBD0D510F0C.taxon	discussion	Discussion. Unlike any other known Lycoriella species, Lycoriella taimyrensis sp. n. and L. tundrae sp. n. have some short but strong and well-sclerotized teeth-like structures laterally on their tegmen, the former in an apicolateral position (Fig. 13 E), the latter in a basolateral position (Fig. 13 F). Lycoriella taimyrensis has its tegmen relatively broader, with straighter lateral sides and with stronger apodemes, its gonostylus and gonocoxa wider, its gonostylus with more megasetae, including apical ones (lacking in L. tundrae) and the setae of its intergonocoxal lobe weaker and more numerous. See also under Lycoriella eurystylata sp. n. and L. jakovlevi sp. n.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C511EFF72FEBD0D510F0C.taxon	etymology	Etymology. The species is named after the area where the collection localities are situated, the Taimyr Peninsula in northern Siberia.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C5119FF72F989091E09FC.taxon	description	Figs 10 B, 11 B, 13 F, 14 D	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C5119FF72F989091E09FC.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Peninsula, on River Zakharova Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov (in ISEA). Paratype. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov (in MZH, http: // id. luomus. fi / GE. 1823).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C5119FF72F989091E09FC.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 – 3 facets wide. Face with 28 – 31 setae. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st and 3 rd segments subequal in length, 2 nd segment shortest; 1 st segment with 2 – 5 setae, with deep dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.5 – 1.6 x as long as wide, neck shorter than wide, longest setae shorter than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 5 – 7 setae. Proepisternum with 5 – 6 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.4 – 1.5 mm. Width / length 0.45. Anal lobe small. Veins distinct, except for stM. c / w 0.65 – 0.70. R 1 / R 0.75 – 0.90. stM shorter than fork of M. bM longer than r-m. stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Pale brown; setae pale. Fore tibial organ forming small indistinct patch of setae in poorly demarcated depression. Fore tibial spur shorter than apical width of tibia. Abdomen. Pale brown; setae pale and short. Hypopygium (Fig. 10 B). Brown, like abdomen. Intergonocoxal area (Fig. 14 D) moderately long, with wide setose lobe. Gonocoxa wide, longer than gonostylus, with sparse, rather long and fine setosity. Gonostylus (Fig. 11 B) widest basally, strongly tapered towards apex, slightly impressed medially; densely setose apically, with strong apical tooth, with 4 – 5 long and narrow medial megasetae on basal bodies on apical third of gonostylus; with long whip-lash seta in the middle. Tegmen (Fig. 13 F) about as long as wide, smoothly curved apically and laterally, weakly sclerotized, with some small teeth basolaterally, apodemes strongly sclerotized, with area of small aedeagal teeth. Aedeagal apodeme short and strong. BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C5119FF72F989091E09FC.taxon	discussion	Discussion. See under Lycoriella eurystylata sp. n., L. jakovlevi sp. n. and L. taimyrensis sp. n.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D55C5119FF72F989091E09FC.taxon	etymology	Etymology. The species is named after the habitat, arctic tundra, where the specimens were collected.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D558511BFF72FF0C0E3D09FC.taxon	description	Figs 7 B, 9 C Synonyms: = curvispina Tuomikoski, 1960; = difficilis var. obscuratipes (Frey, 1948). Literature. Bradysia (Chaetosciara) difficilis var. obscuratipes Frey — Frey (1948): 59, 82. Lycoriella (Lycoriella) curvispina Tuomikoski — Tuomikoski (1960): 79, 85; figs 17 e, 18 j and 19 d. Lycoriella curvispina Tuomikoski — Gerbachevskaja (1969): 339; figs 208.5 and 208.9; Krivosheina & Mohrig (1986): 157; Gerbachevskaja (1988): 514, figs 208.5 and 208.9; Jakovlev (1994): 76; Rudzinski (1994): 286. Sciara parva Holmgren — Holmgren (1869): 16, 52; Jacobson (1898): 204; Lengersdorf (1930): 56; Edwards (1935): 535, fig. 1 d; Bertram & Lack (1938): 51. Lycoria (Neosciara) parva (Holmgren) — Lengersdorf (1928 – 30): 65. Bradysia (Bradysia) parva (Holmgren) — Frey (1948): 67, 85; Stone & Laffoon (1965): 234. Bradysia parva (Holmgren) — McAlpine (1964): 128. Lycoriella (Lycoriella) parva (Holmgren) — Tuomikoski (1967): 49; Gerbachevskaja-Pavluchenko (1986): 33; Menzel & Mohrig (2000): 385, 398, 611; Coulson & Refseth (2004): 113; Menzel et al. (2006): 114; Coulson (2008): 162; Coulson (2013): 154; Mohrig et al. (2013): 271. Lycoriella parva (Holmgren) — Heller et al. (2009): 42; Salmela et al. (2015): 87; Vilkamaa & Menzel (2019): 52; Menzel et al. (2020): 66.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D558511BFF72FF0C0E3D09FC.taxon	materials_examined	Material studied. CANADA, Quebec, Kuujjuarapik, 55 ° 17 ’ N, 77 ° 48 ’ W, 150 m, hilltop, lichen- Sphagnum, 16. VII – 3. VIII. 1990, S. Koponen, 3 males (in MZH, http: // id. luomus. fi / GE. 1778, http: // id. luomus. fi / GE. 1779 and http: // id. luomus. fi / GE. 1780); same locality but treeline forest, window trap, 27. VII – 10. VIII. 1990, S. Koponen, 1 male (in MZH, http: // id. luomus. fi / GE. 1781); Quebec, Mt. Logan, 48 ° 52 ’ N, 66 ° 38 ’ W, 1100 m, alpine zone, S. Koponen, 3 males (in MZH, http: // id. luomus. fi / GE. 1782, http: // id. luomus. fi / GE. 1783 and http: // id. luomus. fi / GE. 1784); RUSSIA, Murmansk Province, Lapland Nature Reserve, 5 km SE of Chunozero settlement (67.64169 ° N, 32.68118 ° E), Malaise trap, 30. V – 23. VI. 2014, A. Polevoi, 1 male (in MZH, http: // id. luomus. fi / GE. 1777); USA, Vermont, Mt. Mansfield, 44.5436 ° N, 66.6347 ° W, 4000 ft., alpine zone, 18. VI – 18. VIII. 1991, S. Koponen, 10 males (8 in MZH, http: // id. luomus. fi / GE. 1785 – 1792; 2 in SDEI); New Hampshire, Mt. Washington, 44.2705 ° N, 71.3032 ° W, 5700 ft., alpine zone, 15. VI – 17. VIII. 1991, S. Koponen, 3 males (in MZH, http: // id. luomus. fi / GE. 1795, http: // id. luomus. fi / GE. 1796 and http: // id. luomus. fi / GE. 1797).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D558511BFF72FF0C0E3D09FC.taxon	description	Redescription. Male. Head. Face slightly darker brown than antenna, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 11 – 24 setae. Clypeus with 1 – 3 setae. Maxillary palpus with 3 segments, 1 st segment slightly longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 2 – 5 (rarely 6) setae, with large dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.85 – 2.35 x as long as wide, neck slightly shorter than wide, longest setae as long as or slightly longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 2 – 4 setae. Proepisternum with 4 – 7 setae. Scutellum with 2 long and some short and fine setae. Wing. Fumose. Length 1.2 – 1.4 mm. Width / length 0.35 – 0.40. Anal lobe small. Veins indistinct. c / w 0.55 – 0.70. R 1 / R 0.60 – 0.85. stM longer than fork of M. bM longer than r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Fore coxa yellow, mid- and hind coxae pale brown, other segments yellow; setae pale. Fore tibial organ forming small patch of setae in demarcated depression. Fore tibial spur longer than apical width of tibia. Abdomen. Yellow; setae pale, short and fine. Hypopygium (Fig. 7 B). Yellow, like abdomen. Intergonocoxal area rather long, with large conical setose lobe. Gonocoxa narrow and long, longer than gonostylus, with rather sparse and short setosity, on medial margin very short. Gonostylus (Fig. 9 C) very narrow, widest basally, narrowed in apical third, impressed on apical half, evenly narrowed towards apex, densely setose apically, with narrow and curved apical tooth; with 2 – 3 apical megasetae, 4 (rarely 5) medial megasetae in apical third, megasetae rather strong, slightly recurved, about as long as apical tooth; without well-differentiated whip-lash seta. Tegmen wider than long, membraneous and smoothly curved or truncate apically, slightly sclerotized and roundish laterally, with small area of aedeagal teeth. Aedeagal apodeme rather short. BIN. BOLD: ABA 5291.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D558511BFF72FF0C0E3D09FC.taxon	discussion	Discussion. There is some variation in the structure of the gonostylus between specimens from different localities, the northernmost specimens studied by us (Kuujjuarapik, Quebec) having a narrower gonostylus than the specimens from the more southern Mt. Logan, Mt. Mansfield and Mt. Washington (Figs 7 B, 9 C). However, as even the European and North American specimens have the same DNA barcode — e. g., specimens with the IDs 24536 - E 11 from Norway (Vestvold) and BIOUG 11301 - E 11 from Canada (Newfoundland and Labrador) — they most probably belong to one and the same species. With the character combination of the gonostylus with strong apical megasetae, few curved medial megasetae, an indistinct whip-lash seta and a strong intergonocoxal lobe, Lycoriella parva is unique among Lycoriella. The species has been recorded from Austria, Canada, Finland, Norway (mainland and Svalbard), Russia, Sweden and United Kingdom (Wirta et al. 2016; Heller & Menzel 2017; Menzel et al. 2020; BOLD Systems 2021).	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D559511BFF72FE410DF60A90.taxon	description	Figs 4 A, 13 B, 14 A Literature. Lycoriella (Lycoriella) acutostylia Mohrig & Menzel — Menzel et al. (1990): 340, fig. 7 a – c; Menzel & Mohrig (2000): 385, 611. Lycoriella acutostylia Mohrig & Menzel — Menzel et al. (2000): 77; Mukkala et al. (2005): 7, 16, 32; Heller et al. (2009): 41; Vilkamaa & Menzel (2019): 51.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D559511BFF72FE410DF60A90.taxon	materials_examined	Material studied. CZECH REPUBLIC, Bílina, Vétrak, 50 ° 33 ’ 29 ” N, 13 ° 45 ’ 57 ” E, dump restoration SW, 200 m, 8 – 10. VI. 1997, M. Barták, 1 male (in SDEI); same data but 9 – 23. IV. 1997, 2 males (in SDEI). FINLAND, Lkor (Lapponia kemensis orientalis), Pelkosenniemi, Kemihaara, Sadinvaara (Grid 7445310: 3537269), Malaise trap, 31. VII – 29. IX. 2015, J. Salmela, 3 males (in LMM; specimen IDs JS-COI- 2016 - 0011, JS-COI- 2016 - 0012 and JS-COI- 2016 - 0242); Ks (Regio kuusamoensis), Kuusamo, Matinjärvi (Grid 7367909: 3615372), Malaise trap, 1. VII – 3. VIII. 2005, J. Salmela, 1 male (in MZH, http: // id. luomus. fi / GE. 1824); Ab (Regio Aboensis), Karkkila, Iilammi, rich fen, Malaise trap, 6. VI. 2004, J. Salmela, 1 male (in LMM). BIN. BOLD: ACV 2466.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D559511BFF72FC350F7F0EC4.taxon	materials_examined	Material studied. RUSSIA, Chukotka, bank of river Anadyr, 64.72 ° N, 175.21 ° E, 25. VI – 19. VII. 2014, A. Barkalov, 6 males (3 in MZH, http: // id. luomus. fi / GE. 1798, http: // id. luomus. fi / GE. 1799 and http: // id. luomus. fi / GE. 1807; 1 in ISEA; 2 in SDEI); Arkhangelsk Oblast, Yamalo-Nenets Autonomous District, near Seyakh, 70.7788 ° N, 72.0750 ° E, nival meadow, Malaise trap, 13 – 29. VII. 2014, N. Zubryi, 1 male (in MZH, http: // id. luomus. fi / GE. 1800). BIN. BOLD: ACC 1679, BOLD: ACO 1394, BOLD: ACP 4705, [? BOLD: AAM 9236].	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D559511BFF72FC350F7F0EC4.taxon	discussion	Remarks. Lycoriella brevipila Tuomikoski, 1960 was often confused with Lycoriella ingenua (Dufour, 1839) in the past (e. g., in Menzel & Mohrig 2000) because the male genital structures are very similar. The specimens identified as ‘ Lycoriella brevipila ’ so far with pale setose flagellomeres, thorax, abdomen and male genital are almost certainly still a mixture of species whose morphological differences have not yet been sufficiently studied. The genetic diversity indicates that extensive taxonomic work remains to be done in this complex of very similar species. Thus, the Canadian specimens with BIN BOLD: AAM 9236 could possibly correspond to Lycoriella pearyi sp. n. (Figs 8 A, 9 D). There are currently 58 specimens on BOLD that have been assigned to Lycoriella brevipila. Their COI sequences form 4 clusters with the BIN IDs BOLD: AAM 9236 (27 specimens from Canada), BOLD: ACC 1679 (5 from Belarus, Finland, Norway), BOLD: ACO 1394 (16 from Belarus, Norway, Sweden), and BOLD: ACP 4705 (10 from Belarus and Norway). The time-consuming verification of the discussed BINs by morphological studies on the corresponding specimens is postponed to a later date. There are almost certainly other species in this complex that do not belong to Lycoriella brevipila Tuomikoski.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5565115FF72F8CF0E6B0B33.taxon	materials_examined	Material studied. USA, Alaska, 11 mi S Anderson Jct, Rte 3, mi 270, forest (alder, poplar, spruce), Malaise trap, 23. VI – 11. VIII. 1984, S. & J. Peck, 1 male (in MZH, http: // id. luomus. fi / GE. 1803). BIN. BOLD: ACC 1227.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5565115FF72F8CF0E6B0B33.taxon	discussion	Remarks. New for the Nearctic region. On BOLD, there are currently registered 50 specimens with four public BINs: 5 with the ID code BOLD: ACC 1227 (Germany, Finland), 35 specimens with BOLD: ACG 8695 (Canada, Finland), 9 with BOLD: ACG 4621 (Belarus, Germany, Norway), and one single female in poor condition from Bulgaria with BOLD: ACP 4988 (not verifiable without a male). Certainly, there may be more species in their morphology very similar to Lycoriella lundstromi (Frey), lacking the intergonocoxal lobe or seta groups.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5575115FF72FD540ED00F75.taxon	materials_examined	Material studied. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov, 1 male (in MZH, http: // id. luomus. fi / GE. 1801); Krasnoyarsk region, Taimyr Peninsula, on River Zakharova Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 2 males (in ISEA and SDEI); Russia, Chukotka, bank of river Anadyr, 64.72 ° N, 175.21 ° E, 25. VI – 19. VII. 2014, A. Barkalov, 1 male (in MZH, http: // id. luomus. fi / GE. 1802). BIN. BOLD: ABA 1215.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5575115FF72FD540ED00F75.taxon	discussion	Remarks. In the BOLD database were found 1039 COI-sequenced specimens with the BIN ID BOLD: ABA 1215. These were collected from the following regions and countries: PALAEARCTIC: Austria, Belgium, Belarus, Bulgaria, China, Germany, Norway, Russia (Primorskiy Kray), South Korea; NEARCTIC: Canada, United States; AFROTROPICAL: South Africa; SUBANTARCTIC: French Southern and Antarctic Lands (Kerguelen Island); AUSTRALASIA: Australia, New Zealand. Thirty-seven other specimens named Lycoriella sativae are recorded with BIN IDs BOLD: ACD 3588 (13 specimens from Canada, Norway, Sweden), BOLD: ACM 4797 (21 from Canada, Iran, Pakistan), as well BOLD: ACR 0675 (3 from Norway and Russia: Primorskiy Kray). Representatives of these clusters must be subjected to a morphological review.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
03E1D765D5575116FF72F9110D5D0844.taxon	materials_examined	Material studied. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov, 1 male (in SDEI); RUSSIA, Krasnoyarsk region, Taimyr Peninsula, on River Zakharova Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 1 male (in ISEA), RUSSIA, Krasnoyarsk region, Taimyr Peninsula, 12.5 km S of Dixon, 73 ° 24 ’ N, 80 ° 39 ’ E, on the River Lemberova, pan trap, 7 – 10. VII. 2012, A. Barkalov, 2 males (1 in MZH http: // id. luomus. fi / GE. 1804; 1 in SDEI); RUSSIA, Yamalo-Nenets Autonomous Okrug, 13 – 29. VII. 2014, N. Zubryi, 1 male (in MZH, http: // id. luomus. fi / GE. 1805); N. Greenland, Nedre Midsommer Sö, 62.0980 ° N, 35.9498 ° W, 14. VII. 1966, Canadian Peary Land Expedition, 1 male (in MZH, http: // id. luomus. fi / GE. 1806). BIN. Unknown.	en	Menzel, Frank, Vilkamaa, Pekka (2021): New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region. Zootaxa 5072 (6): 501-530, DOI: 10.11646/zootaxa.5072.6.1
