identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EE8791F25BD8170897D17CA6698864.text	03EE8791F25BD8170897D17CA6698864.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclogethes tibialis Liu, Huang & Audisio 2024	<div><p>Cyclogethes tibialis Liu, Huang &amp; Audisio, sp. nov.</p><p>Figs 1a–d; 2a–i</p><p>Diagnosis</p><p>Narrowly elongate, scarcely transversely convex, densely, and rather deeply punctuated, uniformly testaceous-orange; vaguely recalling the Oriental Cyclogethes abnormis Kirejtshuk, 1979 (from NE India, N Indochina, and SW China), but easily differentiated by the peculiarly shaped (narrow and markedly sinuate) hind tibiae in both sexes, the more elongate body shape, the male pygidium markedly pointed distad, and the different male and female genitalia.</p><p>Etymology. The specific epithet is derived from the unusually narrow shape of the metatibiae in both sexes, markedly sinuate along their inner edge even in females.</p><p>Material examined</p><p>Holotype. CHINA ♂; Yunnan: Yuxi, Xinping YizuDaizu Autonomous County, Gasa Town, Ailao Mountain Nature Reserve, ca. 1965 m a.s.l., ca. 24.003372°N, 101.547364°E, 09.iv.2017, Chen &amp; He lgt, beating yellow and incompletely flowered corymbs of Pseudognaphalium affine (D. Don) Anderberg ( Asteraceae, subfamily Gnaphalieae; Fig. 3) (NWAU).</p><p>Paratypes. CHINA 3 ♀♀; Yunnan: same data as holotype (NWAU, CAR-MZUR).</p><p>Description (male holotype)</p><p>Measurements. Body length 2.5 mm, width 1.3 mm.</p><p>Colour. Body uniformly testaceous-orange (Fig. 1a), without distinctly paler or darker areas, including peripheral margins of pronotum, only with vaguely infuscate discal portions of elytra; legs and antennae uniformly yellowishorange, testaceous, including antennal club.</p><p>Body shape. Body moderately elongate, scarcely transversely convex (Figs 1a, 2a).</p><p>Dorsal punctation. Surface moderately densely, coarsely, and deeply punctate (spaces between pronotal and elytral punctures ca. 1–1.5× their diameter), with barely shagreened and rather shining interspaces; elytra without traces of transverse strigose sculpturing.</p><p>Prothorax. Pronotum with trapezoidal shape, widely arcuated lateral sides, maximum width nearly at its posterior angles (Figs 1a, 2a). Pubescence sparse, golden-whitish, rather long, and distinct, each individual seta nearly as long as 5 th antennomere, slightly longer only along posterior base. Notosternal sutures distinct, slightly darker than the remaining part of prosternum, only very slightly raised. Median flat portion of the prosternal process narrowly rounded distad, rather parallel-sided, its maximum width nearly in the middle of its whole length, here nearly as wide as antennal club (Fig. 2c).</p><p>Elytra. Elytra rather parallel-sided, ratio LELY/WELY ≈ 1.02. Pubescence sparse, golden-whitish, rather long and distinct, each individual seta nearly as long as 5 th antennomere.</p><p>Metathorax. Metaventrite with a rather shallow, nearly pentagonal impression, occupying its posterior half, deeper in its longitudinal middle. Last abdominal ventrite bearing a couple of rather large proximal semicircular impressions, their diameter nearly as 1.3× the maximum diameter of each eye.</p><p>Pygidium. Proximal base of pygidium with normal, “ V ” shaped carina in the middle, directed backwards (Fig. 2h). Pygidium distinctly pointed distad, with small, acute, and evident apical projection directed backwards (Figs 2a, h).</p><p>Legs. Front tibiae rather wide, triangular, ratio PTLE/PTWI ≈ 0.3, with a series of very small and short tegumental teeth (Fig. 2d); front tarsi nearly as wide as front tibiae, and nearly as wide as the length of the 1 st antennomere (Fig. 2a); middle tibiae rather narrow, moderately sinuated (Fig. 2e), their outer edge with a normal and distinct series of small dense spinules. Hind tibiae narrow (ratio MTLE/MTWI ≈ 4.5), peculiarly shaped, along their inner side strongly modified and narrowed in their posterior half, and markedly sinuate (Figs 1a, 2a, f), their outer edge without a series of spinules (only with a short series of very small hairs distad), without any similitude with any other hitherto known member of its genus.</p><p>Antennae. Antennal club elongate, symmetrical, without evident sexual differentiation (Fig. 2b).</p><p>Male genitalia. Distinctly shaped, rather small, with elongate and subparallel-sided tegmen (Fig. 1b), and roundly pointed apex of each paramere; ratio DTIN/LETE ≈ 0.42–0.43, the excision’s inner margins with a small pre-distal gibbosity; ratio LETE/WITE ≈ 1.70. Aedeagal median lobe peculiarly shaped, slightly narrowed in its distal third, with maximum width near its proximal third; subtruncate but weakly rounded distad (Fig. 1c); ratio LEAE/WIAE ≈ 2.6.</p><p>Female. Front tibiae moderately wide, slender, distinctly narrower than in males, front tarsi slightly narrower than front tibiae (ratio WFTA/LFTA ≈ 0.25). Middle tibiae nearly as in males. Hind tibiae slightly narrower than in males (MTLE/MTWI ≈ 4.7), but along their inner side markedly sinuate, nearly as in males (Fig. 2g). Pygidium regularly rounded distad, without any pointed projection (Fig. 2i). Metaventrite in females almost flat, without distinct impression, in the middle only with a barely impressed longitudinal line. Ovipositor rather small and scarcely sclerotized, not darkened towards the moderately pointed and distinctly divaricated distal apex, exhibiting moderately long styli, inserted rather far from the apex (Fig. 1d). Ratio STLE/DSIA ≈ 0.42; ratio STLE/CGOW ≈ 0.23; ratio GONL/CGOW ≈ 3.1. Ratio OVPL/GONL ≈ 2.15.</p><p>Variation. Body size 2.2–2.5 mm (length) and 1.2–1.3 mm (width). Antennae without evident differences among sexes (Fig. 2b).</p><p>Distribution</p><p>SW China (Yunnan) (Fig. 4).</p><p>Phenology</p><p>The thus far available specimens of the new species were collected in early-middle April, which likely indicates adult local activity in Spring (maybe from early April to late June or early July).</p><p>Hostplants</p><p>As above reported, the four known adult specimens of the new species were all collected in Yunnan on the yellow buds (inflorescence corymbs in prefloration) of a member of the family Asteraceae, subfamily Gnaphalieae, Pseudognaphalium affine (D. Don) Anderberg (Fig. 3). This observation alone does not allow to demonstrate any possible larval host-plant relationship, although the high females/males sex ratio (3:1) could make a true larval trophic relationship slightly more likely. As discussed in the Introduction, the larval hostplants of all members of the genus Cyclogethes are unknown. However, Audisio et al. (2009b, 2015b) have hypothesized a probably close phylogenetic relationship between Oriental Cyclogethes and the southern and eastern African genus Tarchonanthogethes Audisio &amp; Cline, 2009 . Being all known members of the latter genus associated as larvae with male inflorescences of small trees and shrubs of the family Asteraceae (subfam. Carduoideae, tribe Tarchonantheae: Panero &amp; Funk, 2002; Funk et al. 2009; Audisio et al. 2009b, 2015b), we cannot exclude that similar insect-hostplant relationships could link members of Cyclogethes with inflorescences of some Oriental Asteraceae . In this scenario, the best candidates could perhaps be represented by members of some Asian genera that include shrublets and small trees, such as, e.g., Gymnanthemum Cass., Monosis DC., or Vernonia Schreber, but also Pseudognaphalium Kirp. (in the tribe Gnaphalieae, not far from Tarchonantheae: Funk et al. 2009; Mandel et al. 2019) could certainly be taken in account. New insights and further contributions also on phylogenetic relationships between African and Oriental Asteraceae (Mandel et al., 2019) could maybe help in identification of the true hostplants. However, further field research in SW China has been programmed in the next few years, to discover with certainty (i.e., finding the larval stages) the hostplants of Cyclogethes species.</p><p>Habitat</p><p>Locality data indicates that the new species could prefer sparsely forested and bushy areas or thickets, in rocky and rather steep habitats, at the edge of subtropical mountain forest (ca. 2000 m a.s.l.). The other known species of Cyclogethes have been all collected in tropical and subtropical forest habitats, thickets, and forest clearings (Table 1), at altitudes comprised between ca. 400 and ca. 2800 m a.s.l. (Kirejtshuk 1979, 1980; Kirejtshuk &amp; Kabakov 1997; Jelínek 2000b; Audisio et al. 2009b; Liu 2019).</p><p>Taxonomic remarks</p><p>As reported above, this new species certainly belongs to the Cyclogethes abnormis species-group, being vaguely related to Cyclogethes abnormis from NE India, N Indochina, and SW China, and to C. aldridgei from N India and Nepal (Table 1). The new species is otherwise unmistakable due to the very peculiar shape of the male and female hind tibiae, the larger and more elongated body shape, the deeper and coarser dorsal punctation, the dorsal elytral surface without any trace of transversal strigosity, the acutely pointed apex of pygidium in males, as well as by its distinctly shaped male genitalia and ovipositor. However, the peculiar shape of the ovipositor of the new species, markedly bifid distad, and with minute pre-distal styli (Fig. 1d), probably suggests a rather isolated position for the new species, the more related taxa of the Cyclogethes abnormis species-group sharing, in fact, a simply rounded apex of gonostyloids, with larger distal styli (Fig. 43 in Kirejtshuk 1979, Fig. 6k herein; Fig. 84 in Kirejtshuk 1980; Fig. 6h herein).</p></div>	https://treatment.plazi.org/id/03EE8791F25BD8170897D17CA6698864	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liu, Meike;Wang, Xinyue;Yang, Xingke;Huang, Min;Audisio, Paolo;Gardini, Pietro;Sabatelli, Simone	Liu, Meike, Wang, Xinyue, Yang, Xingke, Huang, Min, Audisio, Paolo, Gardini, Pietro, Sabatelli, Simone (2024): A new Chinese Cyclogethes pollen beetle, with an updated key to species of the genus and notes on its phylogenetic positioning (Coleoptera: Nitidulidae: Meligethinae). Zootaxa 5406 (2): 359-372, DOI: 10.11646/zootaxa.5406.2.8, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.8
03EE8791F251D8170897D4C6A27B8F11.text	03EE8791F251D8170897D4C6A27B8F11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclogethes abnormis Kirejtshuk 1979	<div><p>C. abnormis:</p><p>CHINA: Yunnan, Weibaoshan (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.24&amp;materialsCitation.latitude=25.11" title="Search Plazi for locations around (long 100.24/lat 25.11)">Weibao Mts</a>), 25.11°N, 100.24°E, W Slope, 2000–2800 m, 25–28.vi.1992, V. Kubáň lgt., 1♂, 2♀♀ (NMPC, CAR-MZUR) ; CHINA: Yunnan, Cangshan (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.08&amp;materialsCitation.latitude=25.42" title="Search Plazi for locations around (long 100.08/lat 25.42)">Cang Mts</a>), 25.42°N, 100.08°E, E Slope, 2000–2500 m, 21.vi.1992 V. Kubáň lgt., 5♂♂, 5♀♀ (NMPC, CAR-MZUR); ibidem, 2500–3000, 24.vii.1992, Vít Kubáň lgt., 3♀♀ ; CHINA: Yunnan, Dali, 31.v.–1.vi.1993, R. Cervenka lgt., 1♂ ; CHINA: Yunnan, Qujing, Shizong, Junzishan ( Junzi Mts) 08.vi.2016, M. Liu lgt., 1 ex. (used for molecular analysis purposes: CAR-MZUR) ;</p><p>CHINA: Guizhou, Bijie City, Dafang co., Shiniujiao village, E Slope, 2000–2800 m, 09.vii.2018, Y. Chen lgt., 1♂, 1♀ (NWAU, CAR-MZUR) .</p><p>NEPAL: Kathmandu valley, Godavari, 1500 m, 21– 27.05.1989, V . C. Holzschuh lgt. 1♀ (CAR-MZUR) .</p></div>	https://treatment.plazi.org/id/03EE8791F251D8170897D4C6A27B8F11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liu, Meike;Wang, Xinyue;Yang, Xingke;Huang, Min;Audisio, Paolo;Gardini, Pietro;Sabatelli, Simone	Liu, Meike, Wang, Xinyue, Yang, Xingke, Huang, Min, Audisio, Paolo, Gardini, Pietro, Sabatelli, Simone (2024): A new Chinese Cyclogethes pollen beetle, with an updated key to species of the genus and notes on its phylogenetic positioning (Coleoptera: Nitidulidae: Meligethinae). Zootaxa 5406 (2): 359-372, DOI: 10.11646/zootaxa.5406.2.8, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.8
03EE8791F251D8170897D2A0A56D8C26.text	03EE8791F251D8170897D2A0A56D8C26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclogethes aldridgei Kirejtshuk 1980	<div><p>C. aldridgei:</p><p>NEPAL: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.40111&amp;materialsCitation.latitude=27.584723" title="Search Plazi for locations around (long 85.40111/lat 27.584723)">Kandbari</a>, 1200–1500 m, 29.ix.1978, B. Bhakta lgt., 1♀ (CAR-MZUR); near Kathmandu, Phalchoki mount, 2300–2730 m., 27.584722°N, 85.401111°E, 14.v.2000, sweeping in forest, Kostantinov, Lingfelter &amp; Volkovitsh lgt., 1♂, 1♀ (CAR-MZUR) .</p><p>INDIA: Uttarakhand, Sunderdhunga Valley, 8–12.000 fts, W. Almora lgt., on flowering shrubs, 3♀♀ (no additional data; NHM, CAR-MZUR) .</p></div>	https://treatment.plazi.org/id/03EE8791F251D8170897D2A0A56D8C26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liu, Meike;Wang, Xinyue;Yang, Xingke;Huang, Min;Audisio, Paolo;Gardini, Pietro;Sabatelli, Simone	Liu, Meike, Wang, Xinyue, Yang, Xingke, Huang, Min, Audisio, Paolo, Gardini, Pietro, Sabatelli, Simone (2024): A new Chinese Cyclogethes pollen beetle, with an updated key to species of the genus and notes on its phylogenetic positioning (Coleoptera: Nitidulidae: Meligethinae). Zootaxa 5406 (2): 359-372, DOI: 10.11646/zootaxa.5406.2.8, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.8
03EE8791F251D8170897D46FA3D989FC.text	03EE8791F251D8170897D46FA3D989FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclogethes spathulatus Kirejtshuk 1979	<div><p>C. spathulatus:</p><p>CHINA: Yunnan, Cheli, 500 m, 8.IV.1955, unknown collector, 6♂♂, 3♀♀ (NWAU, CAR-MZUR) .</p></div>	https://treatment.plazi.org/id/03EE8791F251D8170897D46FA3D989FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liu, Meike;Wang, Xinyue;Yang, Xingke;Huang, Min;Audisio, Paolo;Gardini, Pietro;Sabatelli, Simone	Liu, Meike, Wang, Xinyue, Yang, Xingke, Huang, Min, Audisio, Paolo, Gardini, Pietro, Sabatelli, Simone (2024): A new Chinese Cyclogethes pollen beetle, with an updated key to species of the genus and notes on its phylogenetic positioning (Coleoptera: Nitidulidae: Meligethinae). Zootaxa 5406 (2): 359-372, DOI: 10.11646/zootaxa.5406.2.8, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.8
03EE8791F251D8140897D1EFA2B18ECB.text	03EE8791F251D8140897D1EFA2B18ECB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclogethes Kirejtshuk 1979	<div><p>Key to identification of species of the genus Cyclogethes</p><p>[from Jelínek (2000b), updated and modified; Figs 5, 6 herein re-inked and slightly modified from Jelínek (2000b) and Kirejtshuk (1979, 1980)].</p><p>1 (2) Metatibiae peculiarly narrow (ratio MTLE/MTWI= 4.5–4.7), in both sexes markedly sinuated along their inner edge (Figs 2f–g), their outer edge without a series of spinules (only with a short series of very small hairs distad). Elytra without any trace of transversal strigosity, distinctly longer than their combined width, ratio ELLE/ELWI≈ 1.15 (Figs 1a, 2a). The whole body yellowish-brown, at most with barely darker, infuscate elytral discal portion. Male genitalia and ovipositor as figured (Figs 1b–d). Body length: 2.2–2.5 mm. SW China (Yunnan) (Fig. 4)................................... C. tibialis sp. nov.</p><p>2 (1) Metatibiae in both sexes wider (ratio MTLE/MTWI= 2.7–3.6), simply arcuated along their inner edge and not sinuated (Figs 11, 22–23 in Jelínek 2000b; Figs 6e–g herein), their outer edge with a series of normal, dense and moderately long spinules. Elytra with more or less distinct traces of transversal strigosity, shorter than or at most as long as their combined width (ratio ELLE/ELWI≈ 0.8–1.0)................................................................................. 3</p><p>3 (4) Elytra nearly as long as their combined width, ratio ELLE/ELWI ≈ or &lt;1.0. Meso- and metatibiae markedly narrower than in the following species, ratio MTLE/MTWI= 3.5–3.6 (Fig. 22 in Jelínek 2000b; Fig. 6f herein). Prosternal process narrowed apically (Fig. 5 in Jelínek 2000b; Fig. 6a herein). Body yellowish-brown, elytra, metaventrite and abdomen blackish-brown to blackish, tips of elytra and apical margin of hypopygium yellowish. Male genitalia as figured (Figs 16–17 in Jelínek 2000b; Figs 5e–f herein). Ovipositor: Fig. 84 in Kirejtshuk 1980; Fig. 6h herein. Body length: 2.1–2.3 mm. N India (Uttar Pradesh, Uttarakhand), Nepal (Fig. 4)..................................................... C. aldridgei Kirejtshuk, 1980</p><p>4 (3) Elytra shorter than their combined width, ratio ELLE/ELWI &lt;1.0. Meso- and metatibiae markedly wider than in the preceding species, ratio MTLE/MTWI= 2.8–3.0 (Figs 11, 23 in Jelínek 2000b; Figs 6e, g herein). Prosternal process not narrowed apically (Figs 6 –7, 9 in Jelínek 2000b; Figs 6b–d herein)............................................................. 5</p><p>5 (6) Prosternal process narrow, parallel-sided (Fig. 6 in Jelínek 2000b; Fig. 6b herein). Male: tegmen with deep V-shaped excision and smooth ventrolateral margins, median lobe of aedeagus broadly arcuately emarginate apically (Figs 18–19 in Jelínek 2000b; Figs 5g –h herein). Ovipositor: Fig. 43 in Kirejtshuk 1979 and in Fig. 6k herein. Length 1.9–2.1 mm. Vietnam, N Thailand, NE India (Darjeeling), SW China (Yunnan, Guizhou) (Fig. 4).............................. C. abnormis Kirejtshuk, 1979</p><p>6 (5) Prosternal process wider, and with more or less arcuate lateral margins (Figs 7, 9 in Jelínek 2000b; Figs 6c–d herein). Male: tegmen more elongate, its ventrolateral margins with series of incisions (Figs 12–13, 20 in Jelínek 2000b; Figs 5a–b, 5i herein).............................................................................................. 7</p><p>7 (10) Elytra transversely strigose with distinct, sometimes oblong, punctures.......................................... 8</p><p>8 (9) Male: genitalia (Figs 34–36 in Kirejtshuk 1979; Figs 5k–m herein) similar to those of C. orientalis, tegmen with shallow apical emargination reaching about one fifth of its length, incisions of ventrolateral margins situated more proximally than the bottom of the emargination; median lobe of the aedeagus parallel-sided, slender, with narrow terminal protuberance gently curved in lateral view. Ovipositor: Fig. 37 in Kirejtshuk 1979; Fig. 6j herein. Body length 1.9–2.1 mm. Vietnam, SW China (Yunnan) (Fig. 4).................................................................... C. spathulatus Kirejtshuk, 1979</p><p>9 (8) Male: tegmen (Figs 12–13 in Jelínek 2000b; Figs 5a–b herein) with apical emargination reaching about two fifths of its length, ventrolateral margins with series of incisions situated immediately at the tips of lateral lobes; median lobe of aedeagus without terminal protuberance (Figs 14–15 in Jelínek 2000b; Figs 5c–d herein). Ovipositor unknown. Body length 1.7–1.8 mm. NW Thailand (Fig. 4)................................................................ C. fuscipennis Jelínek, 2000</p><p>10 (7) Elytra transversely strigose without distinct punctures. Male: genitalia resembling those of C. spathulatus, tegmen with apical emargination reaching about one third of its length and series of incisions on ventrolateral margins situated besides the bottom of the incision; median lobe of the aedeagus slender, parallel-sided with narrow terminal protuberance strongly hooked in lateral view (Figs 20–21 in Jelínek 2000b; Figs 5i–j herein). Ovipositor: Fig. 30 in Kirejtshuk 1979; Fig. 6i herein. Body length 1.9–2.1 mm. Vietnam, N and S Thailand (Fig. 4)..................................... C. orientalis Kirejtshuk, 1979</p></div>	https://treatment.plazi.org/id/03EE8791F251D8140897D1EFA2B18ECB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liu, Meike;Wang, Xinyue;Yang, Xingke;Huang, Min;Audisio, Paolo;Gardini, Pietro;Sabatelli, Simone	Liu, Meike, Wang, Xinyue, Yang, Xingke, Huang, Min, Audisio, Paolo, Gardini, Pietro, Sabatelli, Simone (2024): A new Chinese Cyclogethes pollen beetle, with an updated key to species of the genus and notes on its phylogenetic positioning (Coleoptera: Nitidulidae: Meligethinae). Zootaxa 5406 (2): 359-372, DOI: 10.11646/zootaxa.5406.2.8, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.8
