taxonID	type	description	language	source
03EE87FEFFF1FFFC91A6F887FF18A1CA.taxon	materials_examined	Type species: Laccophilus pumilio LeConte, 1878, here designated.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF1FFFC91A6F887FF18A1CA.taxon	description	Description. Habitus (Figs 1; 3 – 4). Small Laccophilini (TL: 1.8 – 2.5 mm; MW: 0.9 – 1.3 mm), oval or oblong-oval, teardrop-shaped, outline between head, pronotum and elytra continuously curved. Maximum width at about proximal 1 / 5 of elytra. Elytral tip pointed, gradually to abruptly narrowed. Colouration testaceous or reddishyellow, darker on elytra, which often bear pale markings; in one species elytra marmorate. Body surface smooth, without strongly impressed punctures. Elytra with characteristic iridescence, also occurring (although sometimes less evident) on pronotum, ventral surface, and metatarsi. Head. Broad, smooth, shiny, never iridescent, with fine and poorly impressed microreticulation, sometimes with few, scattered and shallow punctures. Pronotum. Short, with angled projection at middle of base (Figs 5 – 6), lateral margins not bordered, hind angles always rounded (Fig. 7). Surface smooth, with fine microreticulation; often with a subtle iridescence. Punctures mostly along lateral sides, hind and fore margins, arranged in more or less regular series. Elytra. Elongate, apically attenuate and often strongly narrowed. Sutural line between elytron and epipleuron not visible in dorsal view, therefore sides of elytra apparently not bordered. Surface smooth, iridescent, with fine microreticulation of elongate cells (Figs 14 – 17). Small and superficial punctures visible in most species, sparse and / or arranged in irregular, longitudinal series. Epipleuron broad to metacoxae / first abdominal ventrite, then strongly narrowed, distinctly attenuate just before elytral tip (Figs 10 – 13). Underside. (Figs 8 – 13) Smooth, more or less iridescent, with microreticulation as on elytra (Figs 18; 19), usually more impressed, especially in females. Lateral expansions of metaventrite (“ metasternal wings ”) thin and arched. Prosternum and head separated by a more or less pronounced step, (Figs 20 – 31). Prosternal process more or less blunt (Figs 20 – 31), in lateral view keeled to saddle shaped, depending on species; in ventral view broadly lanceolate to cordiform, bordered, ending in short, pointed to broadly rounded tip, reaching or barely reaching metaventrite. Metacoxal lines sinuate, strongly convergent anteriorly (Figs 32 – 41); fore laminae of metacoxal processes divided by deep V-shaped incision, either broadly rounded to angulate or ending in sharp spine. Abdominal ventrites lacking curved scratchlike sculpture. Last ventrite as long as rest of abdomen, or almost so (Fig. 42), with large spiculiferous punctures at about distal 2 / 3. Legs. Pro- and mesothoracic legs short, tarsomeres with width greater than length except for fifth almost the length of third and fourth together. Pro- and mesotarsal claws as long as or slightly longer than fifth tarsomere. Spurs of mesotibiae very long, almost as long as entire mesotarsus (Figs 52; 53). Swimming setae on mesotibiae present only apically. Posterior surface of mesofemora always with a series of 3 – 4 long, stiff setae (Figs 8; 12; 52). Spurs of metatibiae simple, not bifid (Figs 54 – 55). Metatarsi with iridescence as on elytra; segments 1 – 4 with short posterolateral lobes and small combs of flat spines. Last metatarsomere with inner claw smaller, outer claw more developed, spur shaped. Males. Last abdominal ventrite tectiform, distally slightly sinuate to almost straight at sides, ending in more or less pointed medial tip (Fig. 42). Pro- and mesotarsomeres 1 – 3 weakly but visibly dilated, each one bearing a pair of stalked suction palettes (Figs 44 – 51). In all but one species, outer claw of forelegs modified, scimitar-shaped (Figs 49; 70 – 76), in some cases with few small denticles on ventral margin. Aedeagus (Figs 84 – 115) asymmetrical as in other genera of Laccophilini; median lobe elongate, generally divided into three parts, variable in shape and proportions depending on species: i) with a thicker base, ii) with distal portion gradually narrowed to apex, or iii) ending in apical expansion (Fig. 91). Left paramere long and thin, arched, either sinuate or straight, reaching half or more than half length of median lobe; right paramere smaller, subtriangular or oval. Both parameres with single long seta at tip. Females. Often duller than males due to more impressed reticulation; iridescence on elytra, pronotum and metacoxae more intense than in males in most cases. Last abdominal ventrite triangular, with almost pointed to almost truncate apex (Fig. 43). Female genitalia (Figs 116 – 120): distal segments of gonocoxae blade-shaped, sharply acute at tip, with small median tooth-like process; an additional subapical denticle present in two species. Ramus short, with more or less densely dentate “ saw ”, depending on species, and lateroproximal processes rudimentary. In general, the species of Laccomimus are hard to identify without examination of male sexual characters. External features are often homogeneous or very variable and difficult to evaluate.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF1FFFC91A6F887FF18A1CA.taxon	distribution	Distribution and ecology. A mainly Neotropical genus, widespread in Central and South America. Three species reach Argentina, and another one reaches Florida in the US. Little ecological information is known for this genus. A large portion of the studied material was collected at light. Available field data (see Figs 130 – 138) suggest a preference for lentic or stagnant waters that are rich in debris and vegetation. Apparently, a few species live in running waters or have a wider ecological range. Young (1954) gave a quite detailed description of the habitat of L.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF1FFFC91A6F887FF18A1CA.taxon	etymology	Etymology. Laccomimus is derived from the words lacco, meaning “ lake ”, and mimus, meaning “ actor ”. Maybe “ water actor ” for their animated and aquatic habits. Name given in litteris by Frank N. Young and Paul J. Spangler. The gender of the name is masculine. Phylogenetic relationships of Laccomimus. The phylogenetic analysis included the following characters: Character 0. Prosternum + head: (0) divided by a shallow step; (1) divided by a deep step; (2) prosternum protruding anteriorly. In Laccomimus the anterior side of the prosternum, just posterior to the head, is quite thick. Pronotum and head are therefore separated by a kind of step (character 0.1; Figs 20 – 22; 29 – 31), which is particularly visible in those species with a protruding prosternum (character 0.2; Figs 24 – 27). In other genera such as Laccodytes, Neptosternus or Laccophilus, the anterior rim of the prosternum is thin (character 0.0). Character 1. Base of pronotum: (0) with median angular projection; (1) straight or almost so. In most genera of Laccophilini, including Laccomimus and the L. phalacroides - group of Laccodytes, a median, posterior lobe of the hind margin of the pronotum projects between the elytra covering the scutellum (character 1.0; Figs 5; 6). In some genera (e. g. Neptosternus, the L. apalodes- group of Laccodytes) the hind margin of the pronotum is straight, lacking a visible projection, although the scutellum remains concealed (character 1.1). Character 2. Posterior angles of pronotum: (0) rounded or angulate; (1) ending in a spiniform process. In Neptosternus and in the L. apalodes - group of Laccodytes (Toledo et al. 2010), the posterior angles of the pronotum end in a spiniform process, projected backward (character 2.1; see fig. 26 in Toledo et al. 2010). In the remaining Laccophilini genera (including Laccomimus) and in the L. phalacroides - group of Laccodytes, the posterior angles of the pronotum are rounded or slightly angulate (character 2.0; Fig. 7 and see fig. 27 in Toledo et al. 2010). Character 3. Prosternal process (ventral view): (0) elongate, reaching mesocoxae but not extending beyond their hind margins; (1) very long, extending beyond hind margins of mesocoxae; (2) broad, not or barely reaching mesocoxae. In Laccophilus, the phalacroides - group of Laccodytes (Toledo et al. 2010), and some other Laccophilini genera, the prosternal process is elongate, narrow, with its apex extending between the mesocoxae (at most extending a little over the hind margins of the mesocoxae as in some species of Laccophilus) (character 3.0; see figs 6, 8 in Toledo et al. 2010 and figs 101 – 103 in Brancucci 1983 b). In the L. apalodes - group of Laccodytes and in Neptosternus, the prosternal process ends in a long, almost needle-like apex, visibly going beyond the hind margins of the mesocoxae (character 3.1; see figs 4, 7 in Toledo et al. 2010). Finally, in Laccosternus and Laccomimus the prosternal process is short and broad, with its apex not or barely reaching the mesocoxae (character 3.2; Figs 23 – 31); the only exception occurs in Laccomimus distinctus, in which the prosternal process is almost similar to that of Laccophilus (Figs 20 – 22). Character 4. Prosternal process (ventral view): (0) simple; (1) trifid. Members of the genus Neptosternus bear a peculiar three pointed prosternal process, resembling a fork (character 4.1). In the remaining Laccophilini genera the prosternal process is single pointed (character 4.0). Character 5. Prosternal process (lateral view): (0) elevated or carinate, visibly reaching and positioned on same plane as metaventrite; (1) elevated or carinate, barely reaching and positioned on same plane as metaventrite; (2) saddle-shaped, barely reaching and positioned below metaventrite. In most genera of Laccophilini the prosternal process in lateral view is elevated or even carinate, and its apex clearly reaches the metaventrite between the mesocoxae. It is also positioned on the same frontal plane as the mesocoxae (character 5.0). In Laccosternus and in most species of Laccomimus the situation is similar but the apex barely reaches the metaventrite (character 5.1; Figs 23; 28; 29 – 31), with the exception of Laccomimus distinctus in which the prosternal process in lateral view looks more like that described for character 5.0 (Figs 20 – 22). In a few species of Laccomimus (L. amazonas, L. spinosus, L. variegatus), the prosternal process can be thought as saddle-shaped, i. e. the outline of the process is strongly sinuate in lateral view and the apex is bent upward, not or barely reaching the metaventrite (character 5.2; Fig. 27). Character 6. Strong and sparse punctures on elytra: (0) absent; (1) present. Both species of Laccosternus (more evident in L. grouvellei (Régimbart, 1895 )) share a strong punctation of sparsely distributed punctures on the whole elytral surface (character 6.1). A strong and diffuse punctation on the elytra was not observed in other Laccophilini, with the exception of the longitudinal series of punctures (more or less clearly impressed, and usually less regular and somewhat sparse in posterior half) commonly present in all species (character 6.0). Character 7. Epipleural carina: (0) not visible dorsally; (1) visible dorsally. In Neptosternus, Laccodytes and Napodytes the carina between the elytron and epipleuron is visible in dorsal view. This gives the appearance of a border running along the lateral side of each elytron (character 7.1). In all other genera of Laccophilini, including Laccomimus, this carina is not visible and the elytra do not appear laterally bordered in dorsal view (character 7.0). Character 8. Silky sheen, mainly on elytra and metatarsi: (0) absent; (1) present. Members of Laccomimus are unique within the Laccophilini genera studied in the presence of a characteristic silky sheen which is more evident on the elytra and metatarsi (character 8.1). This sheen is absent in other genera of Laccophilini (character 8.0). Character 9. Metacoxal lines: (0) sinuate, converging anteriorly; (1) straight, subparallel. In some genera of Laccophilini, including Neptosternus and Laccodytes, the metacoxal lines are straight and parallel, or almost so (character 9.1; see figs 9 – 11 in Toledo et al. 2010). In other genera the metacoxal lines are sinuate and converge anteriorly at the level of the suture separating the metacoxae and the metaventrite (character 9.0; Figs 8 – 13; 32 – 41). Character 10. Lobes of metacoxal processes: (0) separated by a deep notch; (1) notch almost absent. In most genera of Laccophilini, including Laccomimus, the lobes of the metacoxal processes are separated by a more or less deep notch (character 10.0; Figs 32 – 41). In Laccophilus and Philodytes these lobes are very close to each other (character 10.1; see figs 105, 107, 108 in Brancucci 1983 b). Character 11. Lobes of metacoxal processes: (0) rounded to slightly angulate; (1) strongly angulate to acuminate; (2) truncate. Within the studied genera of Laccophilini, Laccosternus and most species of Laccomimus share rounded or slightly angulate lobes of the metacoxal processes (character 11.0; Figs 37; 39 – 41). In some species of Laccomimus the lobes end in a sharp spine in both sexes or in a sort of spine in males and in a more or less marked angulation in females (character 11.1; Figs 32; 33 – 36; 38). In Laccophilus and Philodytes the lobes are distally truncate (character 11.2; see figs 105, 107, 108 in Brancucci 1983 b). Character 12. Larger mesotibial spur: (0) not longer than first two tarsomeres; (1) almost as long as whole mesotarsus. In Laccomimus the mesotibial spurs are very long, the larger one is almost as long as the whole mesotarsus, excluding the claws (character 12.1; Figs 52; 53). In all other genera of Laccophilini studied, including Laccosternus, the mesotibial spurs are shorter, reaching at most the second mesotarsomere (character 12.0). Character 13. Apex of metatibial spurs: (0) simple; (1) bifid. The apically bifid condition of the metatibial spurs (character 13.1) is typical of all known species of Laccophilus except one (see key to genera of Laccophilini, below). In all other genera of Laccophilini, including Philodytes, the apices of the metatibial spurs are acute (character 13.0). Character 14. Claws of male protarsus: (0) equal in shape, unmodified; (1) different in shape, outer claw scimitarshaped. In all species of Laccomimus, except L. distinctus, the outer protarsal claws of males are modified, typically in the shape of a scimitar, different from the inner claw which has a normal sabre shape (character 14.1; Figs 47; 49 – 50). In all other genera of Laccophilini, both male anterior claws are unmodified (character 14.0). Character 15. Median lobe of aedeagus: (0) without a lateral membranous expansion; (1) with a lateral membranous expansion. A lateral membranous expansion is present on the median lobe of the aedeagus of species of the L. apalodes - group of Laccodytes (character 15.1; see figs 44, 45 in Toledo et al. 2010). No other Laccophilini studied exhibits this feature (character 15.0). Character 16. Left paramere: (0) short and broad, shorter than half of length of median lobe; (1) elongate, at least half length of median lobe. In Laccomimus the left paramere is elongate and very long, at least as long as half the length of the median lobe, usually longer (character 16.1; Figs 84 – 98; 101 – 105). The right paramere is, instead, very small. In all other genera of Laccophilini, the left paramere is at most as long as half of the length of the median lobe, usually shorter although generally broad; the right paramere is normally shorter then the left paramere, but the difference is not marked (character 16.0). Character 17. Gonocoxal blades: (0) with two denticles; (1) smooth or with one denticle. In Laccosternus and Laccomimus (with the exception of L. distinctus and L. malkini), the gonocoxal blades are smooth or at most exhibit a single small rounded tooth at about half the length on the dorsal side (character 17.1; Figs 118 – 119); In all other genera of Laccophilini studied and in two species of Laccomimus, the gonocoxal blades bear a subapical, sharp tooth on dorsal side, together with the median tooth described before (character 17.0; Figs 116 – 117). Character 18. Lateral proximal processes of ramus: (0) well developed; (1) short, almost rudimentary. In females of Laccosternus and Laccomimus the lateral proximal process of the female ramus is very short, almost rudimentary (character 18.1; Figs 116 – 120). In Laccophilus and in all other studied genera, the lateral proximal processes of the ramus are long and well developed (character 18.0). The final data matrix includes 20 taxa and 19 characters (15 binary and four multistate) (Table 1). The analysis with TNT yielded a single most parsimonious tree of 26 steps (CI = 0.92; RI = 0.97), which supports the monophyly of Laccomimus and the paraphyly of Laccodytes as presently defined. It also demonstrated weak support for the internal nodes of Laccomimus, with the exception of a robustly supported clade formed by L. variegatus, L. spinosus and L. amazonas (Fig. 121). Among the New World genera of Laccophilini, Laccomimus has been confounded with Laccodytes. The confusion may have arisen from the small size of members of both genera and the scarce knowledge of Laccodytes until recently. In spite of this, Laccodytes and Laccomimus are very different morphologically. They have different body shapes, very flat with short to truncate elytral tips and lateral sides of the elytra finely bordered in dorsal view in Laccodytes (character 7.1, Fig. 2), less flat, with strongly attenuate elytral tip and elytra not bordered in dorsal view in Laccomimus (character 7.0). In Laccodytes the metacoxal lines are straight and subparallel (character 9.1), whereas in Laccomimus they are sinuate and strongly converging anteriorly (character 9.0). In Laccodytes the pro- and mesothoracic legs are long and slender, whereas in Laccomimus they are short and robust. The distant relationship of Laccodytes and Laccomimus suggested by our analysis (Fig. 121) is supported by previous results of and analysis by Ribera et al. (2008). In fact, the specimens identified as " Laccodytes sp 1 " and “ Laccophilini ” in fig. 3 of that paper actually belong to Laccomimus, whereas the specimen identified as " Laccodytes sp 2 " is actually Laccodytes (Ignacio Ribera, 2015, personal communication). Laccomimus is therefore related to Africophilus Guignot, 1948 based on results by Ribera et al. (2008), with a rather isolated position within Laccophilinae, and more distantly related to Laccodytes. Apomorphies of Laccomimus are: i) the prosternum and head divided by a deep step (character 0) (this is particularly marked in species with a protruding prosternum); ii) prosternal process short and blunt, cordiform or almost so, not or barely reaching mesocoxae (character 3); iii) aedeagus with very long left paramere, normally longer than half the length of the median lobe (character 16); iv) larger mesotibial spur almost as long as mesotarsus (character 12); v) surface of body, especially of elytra and metatarsi, with a characteristic silky sheen (character 8); vi) male outer claw of protarsi modified (except in one species) (character 14); vii) gonocoxae with one tooth (with two teeth in two species) (character 17); viii) lateral proximal process of ramus short, almost rudimentary (character 18). None of these character states occurs in Laccodytes in its current concept (Toledo et al. 2010). On the other hand, all species of Laccodytes are stream dwellers as may be indicated by the variegate dorsal colour patterns, a colouration generally exhibited by reophilic Laccophilini (Balke et al. 2000). Laccomimus, instead, apparently inhabits mainly stagnant or slowly flowing waters, although two or three species appear to be found mainly in running waters. The third genus of American Laccophilini, Napodytes, is morphologically very close to Laccodytes, and apparently also similar in ecology, differing only in few, very specialised characters. Laccomimus is a quite homogeneous genus, although some species groups are distinctive. Species Character Laccodytes is presently divided into two very distinct groups: the L. apalodes - group, including two species that appear similar to Neptosternus, and the L. phalacroides - group, including the remaining species (Toledo et al. 2010). Characters 1, 2, 3 and 15 define these two groups, with all the derived conditions present in the L. apalodes - group, and the primitive states present in the L. phalacroides - group (Table 1). The strong differentiation of these two groups, supported by our phylogenetic results (Fig. 121), suggests that Laccodytes in its current form is not monophyletic. Since this study is focused on Laccomimus, however, we prefer not to make changes to Laccodytes at this stage.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	materials_examined	Type locality. Panamá, Canal Zone, Albrook Forest. Type material. Holotype male (USNM): dissected for genitalia probably by Young; aedeagus and last two ventrites glued on pointed card pinned under the holotype “ Panama, C. Z. [Canal Zone], Albrook Forest site, ground, XI. 16 - 17.1967, BLT R. S. Hutton ” [printed label with date and “ ground ” handwritten], “ HOLOTYPE Laccomimus distinctus Toledo & Michat ” [red, rectangular, printed label]. Paratypes (244 exs all bearing red printed label “ PARATYPE Laccomimus distinctus Toledo & Michat ”): Panamá: same data as holotype, except for: “ 9 – 16. X. 1967 ” (1 ex. USNM); “ 16 – 17. XI. 1967 ” (1 ex. USNM); “ 14 – 16. XII. 1967 ” (1 ex. USNM); “ 21 – 22. XII. 1967 ” (1 ex. USNM); “ 28 – 29. XII. 1967 ” (2 exs USNM); “ Canal Zone, 5 mi SSW Gatun Dam, 30. XII. 1987, MacDonald & Schiefer ”, “ William H. Cross Expedition ” (1 ex. MEM); “ Darien, 5 km W Meteti, 9. I. 1988, MacDonald & Schiefer ”, “ William H. Cross Expedition ” (13 exs MEM); “ Tocumen, 6 – 10. VII. 1970, D. Navas BLT ” [hw] (1 ex. USNM); idem “ 26 – 30. VII. 1970 ” [hw] (5 exs USNM – all but one unlabelled); idem “ 31. VIII – 4. IX. 1970 ” [hw] (1 ex. USNM); idem “ 7 – 11. IX. 1970 ” [hw] (2 exs USNM – one unlabelled); idem “ 9 – 16. IX. 1970 ” [printed] (14 exs USNM). Argentina: Prov. Corrientes, Mbrucuyá N. P., 15 – 17. I. 2008, light trap., leg. M. C. Michat ” (14 exs MCMC); “ Prov. Santiago del Estero, Salada Lagoon, Los Juríes, 3. IV. 2002, leg. M. C. Michat ” (2 exs MCMC); “ Prov. Santiago del Estero, Copo N. P., 27. III. 2001, light trap, leg. M. C. Michat ” (1 ex. MCMC); “ Prov. Formosa, El Bagual Reserve, 7. X. 2002, leg. M. C. Michat ” (1 ex. MCMC); “ Prov. Chaco, El Cachapé, 4. XII. 2008, ditch, leg. M. C. Michat ” (7 exs MCMC); “ Prov. Chaco, 97 km NW of Resistencia, 10 km W of Capitán Solari, 26 ° 48 ' 32 " S 59 ° 36 ' 29 " W, 3 – 7. II. 2004. [collector not given] ” (10 exs NHMP – two of them bearing additional “ Coll. Jiří Hájek National Museum Prague, Czech Republic ”); “ Chaco, San Bernardo, III. 1994, leg. O. Di Iorio ” (1 ex. NMW). Bolivia: “ Guayaramerin, Beni, river bank, Soil Zoological Exp., Balogh, Mahunka, Zicsi ”, “ No. 428, beaten from vegetation, 2. XII. 1966 ” (4 exs FF HNMB); “ Guayaramerin, Beni, outskirts of town, Soil Zoological Exp., Balogh, Mahunka, Zicsi ”, “ No. 424, lamping., 1. XII. 1966 ” (1 ex. HNMB); “ Santa Cruz, Buena Vista, 400 m, m. v. light, 20. X. 1994, R. Ward. leg. ” (1 ex. LHB); “ Santa Cruz, Sara, 3 km S Santa Rosa, 22. II. 1969, BLT A Martinez & RS Woodruff ” (14 exs USNM); idem “ 2 km NW Santa Rosa, 21. II. 1969 ” (2 exs USNM). Brazil: “ Mato Grosso, Cuiabà [Cuyabà], BLT 30. III. 1972, WH Whitcomb ” (2 exs USNM); idem “ 4. IV. 1972 ” (1 ex. USNM); idem “ 5. IV. 1972 ” (1 ex. USNM); idem “ 7. IV. 1972 ” (2 exs USNM); idem “ 9. IV. 1972 ” (1 ex. USNM); idem “ 21. IV. 1972 ” (1 ex. USNM); idem “ 22. IV. 1972 ” (3 exs USNM); idem “ 24. IV. 1972 ” (2 exs USNM); “ Sao Paulo, Piracicaba, 10. II. 1966, Black light, C. A. Triplehorn ” (17 exs USNM); “ Minas Gerais, Aguas Vermelhas, XII. 1998, leg. A. Bello & F. Z. Vaz-de-Mello, coll. L. Hendrich ” (11 exs ZSM); “ Minas Gerais Prov., Cordisburgo Faz., Pontinha, XII. 1997, leg. F. Z. Vaz-de-Mello, coll. L. Hendrich ” (3 exs ZSM). Colombia: “ Rio Frio, Magdalena, VI. 1916, Darlington ” (17. exs USNM); “ Aracataca, Magdalena., Darlington ” (2 exs USNM). Costa Rica: “ Prov. Puntarenas (CR 4), Qu. Negra, ca. 8 km NW Golfito, nr. La Gamba (vill.), at Field Station La Gamba, 90 m a. s. l., 8 ° 42 ' 06 " N, 83 ° 12 ' 07 " W, 1 – 3. III. 2009, leg. M. Brojer ” (1 ex. NMW); same data except for “ 1 – 7. III. 2009 ” (1 ex. NMW). French Guyana: “ FG / 09, Piste N 2 Auberge ‘ Elevage de la Chaumiere’, 3. X. 2009, lg. A. Dostal ” (1 ex. NMW). Paraguay: “ Paraguay, Vezényi ”, “ Asuncion, 1904 ”, “ Laccodytes perparvulus Rég., F. Guignot det., 1955 ” (1 ex. HNMB); “ Dept. S. Pedro, Nueva Germania, 12 – 22. I. 1991, leg. U. Drechsel ” (2 exs NMW); “ Dep. Central, Asunciòn, 30. IX. 1991, U. Drechsel leg. ” (1 ex. ZSM); idem “ 2. X. 1992 ” (1 ex. NMW); idem “ Capiata, 3. X. 1994 ” (1 ex. NMW); idem “ Capiata, Sabana de Agrocomia Totai, 2. X. 1994 ” (1 ex. NMW); “ Dep. Cordillera, Arroyos y Esteros, 11. IX. 1991, Drechsel leg. ” (3 exs LHB); “ Dep. Guairá, Garsy [or Garay?], 20. VIII. 1992, Ulf Drechsel leg. ” (17 exs LHB); “ Dep. Paraguari, Sapucal, 4. X. 1992, Dreschsel leg. ” (1 ex. LHB); “ Dep. Alto Paraguay, Defensores del Chaco N. P., Fortín Madrejón, 23. XI. 2002, light trap, leg. M. C. Michat ” (1 ex. MCMC); “ Dep. Presidente Hayes, Transchaco rd., 5 km SE Pozo Colorado, 22. XI. 2002, light trap, leg. M. C. Michat ” (3 exs MCMC). Venezuela: “ Guárico State, 9 ° 46.320 ' N, 67 ° 21.177 ' W, 280 m, rio San Antonio, N. Dos Caminos, side stream, 8. I. 2009, leg. Miller & Short, VZ 09 - 0108 - 02 B ” (1 ex. VAIP); idem except “ mic. habitats, leg. K. B. Miller, VZ 09 - 0108 - 02 C ” (1 ex. VAIP); “ Guárico State, 9 ° 6.794 ' N, 66 ° 59.595 ' W, 152 m, stream at road crossing N. of Palenque, stream, 8. I. 2009, leg. Short, Garcia, Miller, Camacho & Joly, VZ 09 - 0108 - 03 X ” (3 exs VAIP); “ Guárico State, 8 ° 316.7050 ' N, 66 ° 22.602 ' W, 145 m, ca. 65 km S. Las Mercedes, large lagoon with vegetation, 9. I. 2009, leg. Short, Miller, Garcia & Camacho, VZ 09 - 0109 - 01 X ” (6 exs VAIP); “ Apure State, 7 ° 37.298 ' N, 69 ° 3.679 ' W, 83 m, side road ca. 10 km E Mantecal, marshy area and pool by road, 18. I. 2009, leg. Short, Garcia & Camacho, VZ 09 - 0118 - 02 X ” (19 exs VAIP; one ex. bearing additional label “ Laccodytes det. A. E. Z. Short 2009 ”); “ Apure State, 7 ° 38.660 ' N, 69 ° 18.004 ' W, 90 m, between ‘ La Ye’ [La Yeguera?] and Bruzual, lagoon, 18. I. 2009, leg. Short, Garcia & Camacho, VZ 09 - 0118 - 03 X ” (2 exs VAIP); “ Apure State, 8 ° 2.534 ' N, 69 ° 20.530 ' W, 83 m, edge of Bruzual, large marsh, 18. I. 2009, leg. Short, Garcia & Camacho, VZ 09 - 0118 - 04 X ” (1 ex. VAIP); “ Portuguesa State, 8 ° 25.773 ' N, 69 ° 35.202 ' W, 185 m, Rio Guanare N. of Guanare, 19. I. 2009, leg. Garcia, VZ 09 - 0119 - 02 B ” (1 ex. VAIP); “ Zulia State, 9 ° 51.883 ' N, 72 ° 43.285 ' W, 96 m, between Machiques and Tukuko, road side marsh, 29. I. 2009, leg. Short, Garcia, Camacho, VZ 09 - 0129 - 03 X ” (1 ex. VAIP); “ Barinas State, 8 ° 28.477 ' N, 70 ° 32.194 ' W, 205 m, E. Los Pasitos, marshy pond on hacienda, 14. VII. 2009, leg. Short et al, VZ 09 - 0714 - 05 A ” (1 ex. VAIP); “ Amazonas State, 2 ° 10.591 ' N, 63 ° 08.295 ' W, 45 m, S. of Maturin, vegetated river ”, “ morichal margin, 2. II. 2010, leg. Short & Garcia, VZ 10 - 0202 - 01 B ” (2 exs VAIP); “ Sucre State, 10 ° 31.419 ' N, 63 ° 07.070 ' W, 2 m, ca. 5 km SE El Pilar, forested swamp by road, 28. I. 2010, leg Short & Garcia, VZ 10 - 0129 - 04 A ” (5 exs VAIP); “ Delta Amacuro, 9 ° 012 ' 00 " N, 61 ° 53 ' 21.3 " W, 3 m, transect # 3, temporary rain pond, 28. VIII. 2009, leg. L. Cordero, VZ 09 - 0828 - 36 A ” (1 ex. VAIP); “ Delta Amacuro, 9 ° 10.504 ' N, 61 ° 54.610 ' W, 8 m, between Tucupita & Los Güires, marsh by road, 3. II. 2010, leg Short & Garcia, VZ 10 - 0203 - 01 A ” (1 ex. VAIP); each specimen in VAIP bearing additional label (KUNHM-ENT) with its own barcode and catalogue number.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	diagnosis	Diagnosis. One of the most distinctive species of Laccomimus based on the following combination of characters: elytra typically with sutural band darker than rest of surface, which is normally pale brown, gradually faded subbasally and subapically. Male: lobes of metacoxal processes ending in spine, foreclaws unmodified and median lobe characteristically curved. Female: lobes of metacoxal processes in most cases angulated, with gonocoxal blades with two teeth.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	description	Description. Habitus (Fig. 59). TL: 2.05 – 2.4 mm; MW: 1.0 – 1.2 mm (holotype: TL: 2.2 mm; MW: 1.1 mm); A-TL: 2.2 mm; A-MW: 1.1 mm (NUM: 91 exs); drop-shaped, regularly narrowed posteriorly. Smooth; strongly iridescent on elytra and, often, pronotum. Colour. Pronotum testaceous; head and elytra darker. Head gradually paler on front and clypeus; elytra normally faded on apical fourth and with confuse and often fragmented yellow band at base; sutural band normally visibly darker than rest of elytra. Underside mainly testaceous, often darkened on metaventrite and metacoxal plates. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker. Structures and sculpture. Head smooth, shiny, with microreticulation fine and hard to recognise. Pronotum smooth, with few shallow and fine punctures scattered on disc, more numerous along sides. File of punctures along hind margin poorly impressed. Elytra shiny, strongly iridescent, covered by fine but quite impressed microreticulation, together with shallow punctures arranged in irregular, longitudinal series and light roughness in some parts. Prosternal process elongate, clearly carinate, acuminate at tip, reaching mesocoxae (Figs 20 – 22). Lobes of metacoxal processes ending in tooth or simply acute tip, depending on sex (see below). Male. Lobes of metacoxal processes ending in acute tooth (Fig. 32). Last abdominal ventrite tectiform, with antero-lateral sides from rectilinear to visibly sinuate. Outer claw of forelegs unmodified, similar to inner claw (Figs 44 – 45; 73). Median lobe of aedeagus, in left lateral view (Fig. 84 a) straight, then typically curved on apical third, which is flattened like a leaf, ending in small apical expansion. In dorsal view (Fig. 107) somewhat wide, with almost parallel sides, abruptly narrowed on apical fourth; dorsal lamina narrow and straight; ejaculator groove very wide up to narrowed apical fourth. Left paramere (Fig. 84 b) poorly but regularly curved, bent down apically. Female. Lobes of metacoxal processes ending in acute angle; rarely with small apical tooth, but less developed than in males, otherwise almost rounded (Figs 33 – 36). Last abdominal ventrite triangular, almost pointed at apex. Lateral tergites wide and sclerotized; gonocoxal blade (Fig. 116 a) with second subapical denticle at about ¾ of its length. Ramus (Fig. 116 b) with moderately large but dense teeth; normally first tooth, following apical hook, less developed.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	distribution	Distribution. Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guyana, Panamá, Paraguay, Venezuela (Fig. 122). Habitat. As for most species of this genus, the principal source of material studied was collected at lights. Nevertheless, recent field data suggest a wide ecological range for this species, in both stagnant and running waters (Figs 131 – 132; 138). In El Cachapé refuge (Argentina) some specimens were collected in the vegetated margins of a small semipermanent pond about 15 m long, 5 m wide and 70 cm in depth, with clear water and exposed to insolation (Figs 131 – 132).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	etymology	Etymology. The specific epithet refers to the distinctive features of both sexes of this species. It is an adjective in the nominative singular. Notes on variability. Paler patterns on the elytra vary from more or less extended and more or less defined to obscure. In females, the lobes of the metacoxal processes often vary in shape within the same population. The aedeagus and female sexual structures seem to be quite homogeneous.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFAFFFF91A6FD90FD52A306.taxon	discussion	Remarks. This is a very distinct species, characterised by the long, almost Laccophilus - like prosternal process, unmodified male protarsal claws, and angulated or even toothed lobes of the metacoxal processes in females. These features are not found in any other species of Laccomimus, whereas two-toothed gonocoxal blades in females occur also in L. malkini (see below).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	materials_examined	Type locality. Colombia, Amazonas, Leticia. Type material. Holotype male (USNM): with genitalia dissected probably by Young; median lobe of aedeagus and parameres included in DMHF on plastic label pinned under the holotype “ Colombia, Amaz. [= Amazonas], Leticia, III. 12 – 15.1969, P. & P. Spangler ” [printed], “ HOLOTYPE Laccomimus malkini Toledo & Michat ” [red, rectangular, printed label]. Paratypes (68 exs, all bearing red, printed label “ PARATYPE Laccomimus malkini Toledo & Michat ”): Colombia: same data as holotype (28 exs USNM, MTP). Bolivia: “ Beni, 40 km E San Borja, Estacion Biologica Beni, Estancia El Polvenir, 6 – 8. IX. 1987, W. E. Steiner ”, “ At black light: open grass savanna and marsh ” (2 exs USNM). Brazil: “ Sao Paulo, Piracicaba, blacklight, 12. II. 1966, C. A. Triplehorn ” (4 exs USNM); “ Para, Belem, 10. VII. 1969, P. & P. Spangler ” (1 ex. USNM). Ecuador: “ Napo, Lago Agrio (2 km N), 26. IX. 1975, Andrea Langley ” (3 exs USNM); idem except for “ 16. VIII. 1975, blacklight ” (1 ex. USNM). Paraguay: “ Jejuí-Mí, light trap, 19. XII. 2003, leg. M. C. Michat ” (5 exs MCMC); idem “ 20. XII. 2003 ” (2 exs MCMC). Perú: “ Madre de Dios, Rio Tambopata Res, 30 air km SW Pto-Maldonado, 290 m, 2 – 5. XI. 1979, J. B. Heppner, subtropical moist forest ” (8 exs USNM). Suriname: “ Langaman, Kondre-Marowijne distr., 25. X. 1963, Borys Malkin ” (5 exs USNM); “ Zanderijsavanne, 1 st tributary of Colakreek, 55 ° 14 ' W, 05 ° 27 ' N, 4. IX. 1969, leg N. Nieser (SN 114) ” (1 ex. NMW); “ Paramaribo, Guest House of Museum, at light, 20. VIII – 23. IX. 1969, leg N. Nieser ” (2 exs NMW). Venezuela: “ Guarico, 12 km S Calabozo, II. 6 – 12.1969, P. & P. Spangler ” (3 exs USNM); idem “ 15 km S Calabozo, II. 9 – 13.1969 ” (1 ex. USNM); “ Bolivar State, 04 ° 41.878 ' N, 61 ° 04.246 ' W, 815 m, Gran Sabana, N Santa Elena, marshy area long Rt. 10, 17. VII. 2010, leg. Short, Tellez & Camacho, VZ 10 - 0717 - 03 A ” (4 exs VAIP – each specimen with additional label KUNHM-ENT, with its own barcode and catalogue number).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	diagnosis	Diagnosis. Easily recognised by the combination of (usually) uniformly brown elytra and bright pale pronotum, along with male (scimitar-shaped outer foreclaw, characteristic bisinuate left paramere) and female (two-toothed gonocoxal blades) characters.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	description	Description. Habitus (Fig. 60). TL: 2.0 – 2.3 mm; MW: 1.0 – 1.2 mm (holotype: TL: 2.05 mm; MW: 1.1 mm); A-TL: 2.1 mm; A-MW: 1.2 mm (NUM: 36 exs); almost regularly oval, gradually narrowed backward; elytral tip almost truncate. Glossy to almost dull; iridescence on elytra more or less visible. Colour. Head chestnut brown, gradually paler on front and clypeus; pronotum testaceous, with darker band along hind margin. Elytra uniformly chestnut brown, rarely with more or less defined paler markings on subbasal and preapical position. Underside yellowish, at most slightly darkened on sutures. Antennae and mouthparts testaceous. Legs testaceous, metatarsi slightly darker. Structures and sculpture. Head smooth, shiny, with quite impressed microreticulation of rounded cells. Pronotum shiny, often with slight iridescence. Few scattered punctures only along sides. Elytra with no or hardly visible subapical sinuation; shiny to almost dull, with iridescence of variable intensity; microreticulation quite impressed, accompanied by subtle scratches and scattered, poorly visible punctures. Prosternal process (Fig. 23) rounded at tip, slightly elevated. Lobes of metacoxal processes regularly rounded in both sexes (Fig. 37). Male. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped (Fig. 71). Median lobe of aedeagus, in left lateral view (Fig. 85 a), slender; base and distal portion forming an angle of about 160 °; distal portion straight, gradually narrowed to apex, ending in small button-like expansion, quite variable in shape. In dorsal view (Fig. 108) slender and acuminate, gradually tapering from base to tip; dorsal lamina strongly sinuate close to base; ejaculator groove narrow. Left paramere (Fig. 85 c) straight, visibly bi-sinuate on ventral side. Female. Elytra and pronotum often slightly duller than in males. Last abdominal ventrite triangular, narrowly rounded at apex. Lateral tergites similar to L. distinctus: wide and sclerotized. Gonocoxal blades (Fig. 117 a) with second denticle at about distal 4 / 5. Ramus also distinctive (Fig. 117 b), with numerous densely distributed small teeth of uniform size.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	distribution	Distribution. Bolivia, Brazil, Colombia, Ecuador, Paraguay, Perú, Suriname, Venezuela (Fig. 123). Habitat. In Santa Elena (Venezuela) some specimens were collected in a marshy area exposed to direct sunlight (Fig. 135).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	etymology	Etymology. Name given in litteris by Frank N. Young and Paul J. Spangler, in honour of the collector Borys Malkin (cf. Young 1974: 2, 3). The specific epithet is a noun in the genitive case. Notes on variability. Specimens from most localities have uniformly brown elytra. Paler markings, variably developed but never sharply defined, have been observed in most specimens from Paraguay. The few exemplars seen from Brazil (Piracicaba) share peculiar characters: larger size (TL: 2.4 – 2.5 mm; MW: 1.2 – 1.25 mm), more elongate body shape (with apical sinuation of elytra missing or almost so), pale patterns on the elytra well developed and clear (Fig. 60 b), and a button-like apical expansion of the median lobe projecting posteriorly into a small, acute process (Fig. 85 e). Such specimens might represent a distinct species, but at present this is difficult to determine due to the few available specimens.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFF8FFF891A6FC31FC91A56B.taxon	discussion	Remarks. This species is close to L. pumilio and related species (see below), sharing with them a series of characters: scimitar-shaped male outer foreclaws, the lobes of the metacoxal processes rounded in both sexes, and the prosternal process broad and short, barely reaching the metaventrite. However, it is one of only two species, along with L. distinctus, that exhibits female gonocoxal blades with two teeth. The peculiar ramus, bearing dense, numerous and uniformly small teeth, is unique within the genus.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	materials_examined	Type locality. Perú, Colonia Calleria, Río Calleria, 15 km E Ecayali. Type material. Holotype male (USNM): dissected by us; median lobe of aedeagus and parameres included in DMHF on plastic label pinned under the holotype “ Peru, Colonia Calleria, 15 [km?] E Ecayali, muddy R. Shore, ix. 10 – 25.61, B. Malkin # 14 ” [printed white label; number “ 14 ” hw], “ HOLOTYPE Laccomimus variegatus Toledo & Michat ” [red, rectangular, printed label]. Paratypes (24 exs, all bearing red, printed label “ PARATYPE Laccomimus variegatus Toledo & Michat ”): Perú: same data as holotype (9 exs USNM); idem, exception made for “ ix. 10 – 30.61 ” plus “ # 1 ” (1 ex. USNM); idem, exception made for “ # 16 ” (1 ex. USNM); “ Loreto, 80 km NE Iquitos, Explorama Lodge, 1 km up Rio Yanamono from Amazon River, 1 – 5. IX. 1992, P. Skelley, at light / coll. Hendrich Berlin " (1 ex. ZSM). Bolivia: “ Guayaramerin, Beni, river bank, Balogh, Mahunka, Zicsi ”, “ No. 428, beaten from vegetation, 2. Xii. 1966 ” (1 ex. HNMB); “ Beni, 40 km E San Borja, Estacion Biologica Beni, Estancia El Porvenir, 6 – 8. IX. 1987, W. E. Steiner ”, “ At black light: open grass savanna and marsh ” (8 exs USNM); “ Beni, 40 km E San Borja, Estacion Biologica Beni, Palm Camp at Rio Curitaba, 13. IX. 1987 / At black light, mud flat in small open area of inondation forest, W- E. Steiner " (3 exs – two of which unlabelled – USNM).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	diagnosis	Diagnosis. Unmistakable species based on the peculiar elytral colouration, with vermiculation of irregular obscure markings and pale spots (Fig. 61), unique within Laccomimus. Lobes of metacoxal processes ending in long spine in both sexes. Aedeagal features distinctive, median lobe expanded on apical fourth (Fig. 86 a) and left paramere strongly narrowed on distal half (Fig. 86 b).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	description	Description. Habitus (Fig. 61). TL: 2.2 – 2.4 mm; MW: 1.15 – 1.25 mm; (holotype: TL: 2.3 mm; MW: 1.2 mm); A-TL: 2.3 mm; A-MW: 1.1 mm (NUM: 16 exs); elongate, drop-shaped, strongly narrowed posteriorly. Smooth; dorsal surface with faint silky sheen. Colour. Head testaceous, slightly darker between eyes in some specimens; pronotum testaceous with darker band along fore margin and obscure darker area close to hind margin. Elytra testaceous to reddish-brown with characteristic vermiculation of darker, obscure and irregular markings together with tiny, rounded, pale spots; these markings are concentrated mostly on disc and along lateral sides of elytra, giving illusion of very irregular, longitudinal patterns. Underside reddish-yellow to brown, paler on prosternum. Antennae and mouthparts testaceous. Legs testaceous. Structures and sculpture. Head smooth, shiny, with fine and hard to recognise microreticulation. Pronotum smooth, with few shallow and fine punctures scattered on disc and, more numerous along inner and fore margins; microreticulation as on head; iridescence absent. Elytra elongate with pronounced subapical sinuation; surface shiny, faintly iridescent; microreticulation fine, poorly impressed; small, shallow, irregular grooves visible together with fine scattered punctures. Underside shiny, with hardly visible silky sheen. Prosternal process broad, slightly elevated, obtuse at tip. Lobes of metacoxal processes ending in long sharp spine in both sexes. Male. Last abdominal ventrite tectiform, lateral sides almost rectilinear. Outer claw of forelegs scimitar-shaped (Fig. 76). Median lobe of aedeagus, in left lateral view (Fig. 86 a), widely arched, slender but robust, expanded on apical fourth, then narrowed into subtruncate tip. In dorsal view (Fig. 109), sinuate on both sides, with visible constriction on distal third of left side; apex with narrow tip turned to left; ejaculator groove almost uniformly wide up to apex. Left paramere (Fig. 86 b) regularly rounded along ventral side, strongly narrowed beyond basal half on dorsal side. Female. Not duller than male. Last abdominal ventrite elongate, with fore margin broadly rounded. Lateral tergites thin, partly membranous. Gonocoxal blade (Fig. 119 a) with single medial denticle; ramus (Fig. 119 b) with large and spaced dentation; first tooth, following apical hook, as developed as second tooth.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	distribution	Distribution. Bolivia, Perú (Fig. 124).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	etymology	Etymology. The specific epithet is an adjective in the nominative singular and refers to the distinctive elytral colouration, unique within this genus. Notes on variability. All but one specimens from Bolivia (all immature) bear obscure markings on the elytra, very difficult to see, colouration appearing uniform. This might also depend on the way the beetles have been killed and preserved.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFFFFF991A6FA27FB09A636.taxon	discussion	Remarks. This species is very close to L. amazonas and L. spinosus, but readily recognisable by the peculiar elytral colouration. These three species form a distinct group within Laccomimus, characterised by the following combination of peculiar characters: prosternum protruding anteriorly, almost bulbous (Figs 24 – 26); prosternal process very broad and short, saddle shaped in lateral view, barely reaching metaventrite and positioned below its level (Figs 24 – 26); lobes of metacoxal processes ending in long sharp spine in both sexes (Fig. 38).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	materials_examined	Type locality. Brazil, Matto Grosso, Jacaré, Xingu National Park. Type material. Holotype male (USNM), dissected by us; genitalia mounted in DMHF on plastic label pinned under the holotype “ Brazil, Matto Grosso, Jacaré, P. N. Xingu, XI. 1961, Alvarenga e [sic!] Werner leg. ” [printed], “ HOLOTYPE Laccomimus spinosus Toledo & Michat ” [red, rectangular, printed label].	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	diagnosis	Diagnosis. Easily recognised by the following combination of characters: lobes of metacoxal processes ending in long sharp spine in both sexes; male with outer claw of forelegs scimitar-shaped. Aedeagal features distinctive, median lobe lacking apical expansion but with faint constriction at about mid length (Fig. 87 a).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	description	Description. (based on holotype). TL: 2.3 mm; MW: 1.2 mm; similar to L. variegatus except body somewhat broader and shorter, strongly but regularly narrowed posteriorly, lacking elytral patterns. Elytral surface with visible silky sheen. Colour (it is an old specimen and colouration might be altered). Head uniformly testaceous; pronotum testaceous with darker band along posterior margin. Elytra almost uniformly reddish-brown, with darker parasutural line. Underside uniformly reddish-brown. Antennae and mouthparts testaceous. Legs testaceous. Structures and sculpture. Head smooth, shiny, with hardly visible microreticulation and scattered punctures of different size. Pronotum with lateral sides widely rounded toward anterior angles; surface as on head, with a faint silky sheen and hardly visible microreticulation. Elytra shorter than in preceding species; subapical sinuation visible although poorly developed; elytral apex narrow, almost acute; surface with faint silky sheen; irregular shallow grooves and scattered punctures visible as in L. variegatus. Underside with intense iridescence. Prosternal process as in L. variegatus but centrally quite convex, rounded at tip. Lobes of metacoxal processes ending in long sharp spine. Male. Last abdominal ventrite tectiform with lateral sides almost straight. Outer claw of forelegs scimitarshaped. Median lobe of aedeagus, in left lateral view (Fig. 87 a), regularly curved, narrowed to apex, lacking expansions but with faint constriction dividing basal and apical halves. Left paramere (Fig. 87 b) similar to that of L. variegatus but apical half thicker and less elongate. Female. Unknown.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	distribution	Distribution. Brazil. Known only from the type locality (Fig. 124).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	etymology	Etymology. The specific epithet is an adjective in the nominative singular and alludes to the acuminate spine on each lobe of metacoxal processes, characteristic of this species and those closely related.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFEFFFA91A6F9E1FDABA341.taxon	discussion	Remarks. See under L. variegatus.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	materials_examined	Type locality. Venezuela, Amazonas State, Communidad caño Gato on Río Sipapo, 4 ° 58.838 ' N, 67 ° 44.341 ' W, 95 m. Type material. Holotype male (MIZA), dissected by us; genitalia mounted in DMHF on plastic label pinned under the holotype “ Venezuela, Amazonas State, 4 ° 58.838 ' N, 67 ° 44.341 ' W, 95 m, Comunidad caño Gato on Rio Sipapo, 16. I. 2009, leg. Short, Miller, Camacho, Joly & García, VZ 09 - 0116 - 01 X, along stream ” [printed], “ SM 0843379 KUNHM-ENT ” [printed, together with barcode], “ HOLOTYPE Laccomimus amazonas Toledo & Michat ” [red, rectangular, printed label]. Paratypes (33 exs, all bearing red, printed label “ PARATYPE Laccomimus amazonas Toledo & Michat ”): Venezuela: same data as holotype (30 exs VAIP, MTP, MCMC, NMW); “ Amazonas State, 4 ° 55.849 ' N, 67 ° 44.645 ' W, 87 m, stream along Rio Sipapo, 16. I. 2009, leg Short, Garcia, Camacho, Miller & Joly, VZ 09 - 0116 - 02 X, stream habitats ” (2 exs VAIP); “ Amazonas State, 5 ° 20.497 ' N, 67 ° 45.358 ' W, 72 m, ca. 12 km N Samariapo, small stream at rd. x-ing, 6. I. 2006, AS- 06 - 013, leg. A. E. Z. Short ” (1 ex. VAIP). Each specimen bearing second VAIP label with its own barcode and catalogue number.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	diagnosis	Diagnosis. Easily recognised by the following combination of characters: lobes of metacoxal processes ending in long sharp spine in both sexes; elytra dark with distinct pale markings; male with very large stalked suction palettes and outer claw of forelegs scimitar-shaped. Aedeagal features distinctive, median lobe expanded on apical fourth (Fig. 88 a) and left paramere triangular, gradually tapering apically (Fig. 88 b).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	description	Description. Habitus (Fig. 62). TL: 2.2 – 2.4 mm; MW: 1.15 – 1.2 mm; (holotype: TL: 2.3 mm; MW: 1.15 mm); A-TL: 2.4 mm; A-MW: 1.2 mm (NUM: 16 exs); drop-shaped, strongly but regularly narrowed posteriorly. Smooth; elytra with faint silky sheen. Colour. Head testaceous, often darkened behind eyes. Pronotum testaceous, with darker transverse marking along posterior margin, normally divided into two parts. Elytra chestnut-brown; each elytron with three distinct yellow markings: transverse, subbasal band reaching lateral side; submedial faded spot close to lateral side; subapical marking not reaching apex. Underside reddish-yellow. Antennae and mouthparts testaceous. Legs reddish-yellow. Structures and sculpture. Head shiny, with faint microreticulation and scattered fine punctures on disc. Pronotum smooth, sculpture as on head, with faint silky sheen. Elytra elongate, with subapical sinuation poorly developed and elytral apex narrow, almost acute; surface smooth, with faint silky sheen; fine and scattered punctures are visible in most specimens but no grooves occur, in contrast to preceding species. Underside shiny, lacking silky sheen. Prosternal process broad, slightly elevated especially close to rounded tip (Fig. 24 – 27). Lobes of metacoxal processes ending in long sharp spine in both sexes (Fig. 38). Male. Last abdominal ventrite tectiform with lateral sides almost rectilinear. Stalked suction palettes on pro- and mesotarsi very broad, broader than in any other Laccomimus species (Figs 47 – 48; compare with Fig. 44 and Figs 49; 51). Outer claw of forelegs scimitar-shaped (Fig. 75). Median lobe of aedeagus, in left lateral view (Fig. 88 a), resembling that of L. variegatus, with leaf-like expanded apical fourth but visibly narrower, especially before apical expansion. In dorsal view (Fig. 110) slightly turning to right, gradually tapering from base to apex, with nipple-like tip; dorsal lamina sinuate and ejaculator groove narrower on basal half and wider near apex. Left paramere distinctive (Fig. 88 b): triangular, shorter than in any other Laccomimus species, and more gradually tapering apically. Female. Last abdominal ventrite elongate, with fore margin broadly rounded. Lateral tergites thin and partly membranous. Gonocoxal blade with single medial denticle; ramus with large and spaced dentation; first tooth, following apical hook, very small with respect to second tooth.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	distribution	Distribution. Venezuela, Amazonas State (Fig. 124). Habitat. Collected in running waters (Fig. 137).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	etymology	Etymology. The specific epithet relates to type locality. It is treated as a noun in the nominative singular.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFDFFFB91A6FC69FDABA362.taxon	discussion	Remarks. See under L. variegatus.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFCFFE491A6FC1FFE48A683.taxon	materials_examined	Type locality. USA, Florida, Enterprise. Type material. Holotype (MCZ) not studied. Material examined. Belize: British Honduras (Belize), Corozal Town, 4 – 14. V. 1967, George & Ruth Lacy (3 exs USNM); Toledo Dist., Blue Creek Village, 17. VI. 1981, W. E. Steiner, taken at blacklight in rainforest valley near creek, Earthwatch Belize expedition 1981, D. H. Messersmith, W. E. Steiner et al. (1 ex. USNM); Cayo District, San Ignacio (14 km S), colln 17, 23. V. 1987, Paul J. Spangler & Robin A. Faitoute / Collected at blacklight at Chaa River Cottages (16 exs USNM). Cuba: Cayamas, E. A. Schwarz collector (different dates hw: 7. V; 8. V; 9. V; 11. V. Year not signed. Two specimens with “ Coll’n JD Sherman Jr. 1928 ”), most specimens bearing also an additional label “ Laccophilis pumilo LeC. Det. F. N. Young ” [handwritten by Young] (16 exs USNM); Laguna, Base J. A. Mella, 24. III. 1973, V. Decu (1 ex. USNM). Guatemala: Izabal, Rio Dulce Fy., 2. VII. 1974, W. E. Steiner (1 ex. USNM). USA: Florida, Dade Co., Homestead, 5. I. 1977, W. E. Steiner (6 exs USNM); Florida, Alachua County, San Felasco, Hammock, T- 9 - 6 R- 19 - E, Sec. 19, 478, 28. X. 1947, F. N. Young (3 exs USNM; one bearing also “ P. J. Spangler coll. 1958, 221697 ”); Florida, Alachua County, 28. X. 1947, 478, F. Young / FIG [small, green printed label] (1 ex. USNM); Florida, Alachua County, San Felasco, Hammock, 1931, 24. VIII. 1961, F. N. Young (1 ex. USNM); Florida, Sanford, 28. X. 1960, BLT Gindess … [illegible] (1 ex. USNM); Florida, Gainesville, Doyle Cónner Building, 1 – 7. IX. 1972, H. R. Dodge, light trap (1 ex. USNM); Florida, Collier Co., Marco, 1. I. 1977, W. E. Steiner (1 ex. USNM); Florida, Collier Co., S of Indian Riserve, at light, 30. V. 1993, T. A. Lott & D. Mattews leg. (3 exs MTP).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFCFFE491A6FC1FFE48A683.taxon	diagnosis	Diagnosis. Recognisable from most species of Laccomimus by aedeagal characters only, and from L. alvarengi (see below) by aedeagal characters and, usually, by a more uniform elytral colouration. Median lobe differentiated into basal and distal portion, divided by an angle or a curve (Figs 91 – 93), with apex ending in small button-like expansion. The acuminate tip of the prosternal process is the most striking character, although hard to appreciate at low magnifications.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFCFFE491A6FC1FFE48A683.taxon	description	Description. Habitus (Fig. 63). TL: 2.1 – 2.35 mm; MW: 1.05 – 1.2 mm; A-TL: 2.2 mm; A-MW: 1.1 mm (NUM: 34 exs); regularly drop-shaped, gradually narrowed backward; subapical sinuation not strongly pronounced; elytral tip rounded to almost truncate. Glossy; iridescence clearly visible on elytra and pronotum. Colour. Head uniformly testaceous or slightly darkened on front, between eyes; pronotum testaceous, normally with wide, transverse brown band on basal half, where two darker, elongate spots are often visible close to posterior margin. Elytra chestnut brown, often with paler, hardly defined areas in subbasal, subapical and, normally, lateral position; otherwise uniformly brown or almost so. Underside testaceous, at most slightly darkened on sutures. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker. Structures and sculpture. Head smooth, shiny, with poorly impressed microreticulation of rounded cells and small scattered punctures. Pronotum shiny, with clear iridescence. Regular row of small punctures visible along both anterior and posterior margin, this latter interrupted at disc. Elytra gradually narrowed at tip, with subapical sinuation poorly marked; shiny, with clear iridescence; microreticulation quite impressed, accompanied by shallow and scattered punctures; a more or less regular, longitudinal row of small, shallow punctures is often (hardly) visible. Prosternal process (Fig. 67) slightly carinate, acuminate at tip. Lobes of metacoxal processes regularly rounded in both sexes. Male. Last abdominal ventrite tectiform, with antero-lateral sides visibly but not strongly sinuate. Outer claw of forelegs scimitar-shaped, sinuate, with ventral margin often slightly dentate (Figs 82 – 83). Median lobe of aedeagus, in left lateral view (Figs 91 a; 92 a; 93 a), with base and distal portion forming distinct angle; distal portion elongate and broader at middle length, then gradually narrowed to tip, visibly sinuate along both dorsal and, especially, ventral side, ending in small button-like expansion. In dorsal view same as in L. alvarengi (see Fig. 111); broadly lanceolate, gradually tapering apically and slightly turning right; dorsal lamina sinuate along its whole length; ejaculator groove strongly narrowed apically. Left paramere (Figs 91 c; 92 c; 93 c) curved, apically thin and very poorly sinuate. Female. Occasionally elytra and pronotum duller than in males. Last abdominal ventrite triangular, narrowly rounded at apex. Lateral tergites thin and partly membranous. Gonocoxal blade with single medial denticle. Ramus with large and spaced teeth; first tooth contiguous to apical hook, less developed with respect to second tooth, sometimes very reduced.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFCFFE491A6FC1FFE48A683.taxon	distribution	Distribution. So far known with certainty from USA (Florida), Belize, Guatemala, and Cuba (Fig. 128). Young (1954) reported to have seen specimens from Jamaica and Haiti with male genitalia very similar to specimens from Florida and Cuba. All the specimens we have seen in USNM from Jamaica and Haiti are L. bordoni (see below). Mexico remains a large gap in the known distribution of this species (at present, Laccomimus is known to be represented in this country only by L. spangleri, see below) in spite of numerous field expeditions. Nevertheless, the occurrence of this beetle in Mexico is likely. Notes on variability. At present this species is known to us from three populations: Florida, Belize, and Cuba, plus a single male from Guatemala. No substantial differences have been observed in the specimens from Belize and Guatemala in comparison with those from Florida. The specimens from Cuba exhibit some differences, mainly in the shape of the median lobe of aedeagus, where the separation between the base and the distal portion is arched instead of angulated (Fig. 93 a), in the prosternal process which is less elevated and with less acuminate tip (Fig. 68), and in having more developed pale patterns on elytra. These populations are somewhat similar to L. alvarengi (see below), and based on available information, we are not sure about the systematic position of the specimens. We hesitate to describe it as a new species at this time. Future collecting activities and / or molecular studies may clarify its status.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFFCFFE491A6FC1FFE48A683.taxon	discussion	Remarks. The combination of a short and broad prosternal process, which is in lateral view straight and on the same level as the metaventrite, the outer protarsal claw of the male scimitar-shaped, the rounded or at most obtusely angulated metacoxal lobes, and the female gonocoxal blades with one tooth, distinguishes L. pumilio and related species (see below) within the genus. Among these, L. pumilio and L. alvarengi can be separated by aedeagal characters only.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	materials_examined	Type locality. Brazil, Mato Grosso, Jacaré, Xingu National Park. Type material. Holotype male (USNM): dissected for genitalia probably by Young; genitalia and last two abdominal ventrites glued on separate label under the holotype “ Brazil, Matto [sic] Grosso, Jacaré, P. N. Xingu, XI. 1961, Alvarenga & Werner leg ” [printed], “ FIC ” [small, pink, printed label] ”, “ Laccomimus alvarengi Young ” [white label, handwritten by Young], “ HOLOTYPE Laccomimus alvarengi Toledo & Michat ” [red, rectangular, printed label]. Paratypes (200 exs all bearing red, printed label “ PARATYPE Laccomimus alvarengi Toledo & Michat ”): Brazil: same data as holotype (10 exs USNM); same data as holotype except for “ XI. 1965 ” (65 exs USNM – most of them without label); “ Mato Grosso, Cuiabá, 30. III. 1972, W. H. Withcomb ” (8 exs USNM); idem but “ 9. IV. 1972 ” (6 exs USNM); idem but “ 11. IV. 1972 ” (3 exs USNM); idem but “ 21. IV. 1972 ” (1 ex. USNM); idem but “ 22. IV. 1972 ” (7 exs USNM – four of which without label); “ Mato Grosso, Rio Bento Gomes (Pantanal), 1993 – 1994, E. Stuhr leg. ”, “ Flussabschnitt bei [= section of river at] Periquito, 15 ° 54 ' S, 56 ° 33 ' W ” (1 ex. NMW); “ Rondôna, 62 km SW Ariquemes, nr. fz. Rancho Grande, 8 – 20. IX. 1994, JE Eger & C. W. O’Brien leg ” (3 exs ZSM); “ Minas Gerais, Prov. Cordisburgo, Faz. Pontinha, XII. 1997, leg. F. Z. Vaz-de-Mello, coll. L. Hendrich ” (16 exs ZSM); “ Par. Curitiba, 28. VI. 1969, leg. P. & P. Spangler ” (1 ex. USNM). Argentina: “ Prov. Corrientes, Mbrucuyá N. P., 15 – 17. I. 2004, light trap, leg. M. C. Michat ” (24 exs MCMC). Bolivia: “ Beni, 40 km E San Borja, Estacion Biologica Beni, Estancia El Porvenir, 6 – 8. IX. 1987, M. E. Steiner ”, “ At black light, open grass savanna and marsh ” (7 exs USNM); “ Guayaramerin, Beni, river bank, Soil Zoological Exp., Balogh, Mahunka, Zicsi ”, “ No. 428, beaten from vegetation, 2. XII. 1966 ” (2 exs HNMB); “ El Beni, Beni Station, NE of San Borja, 31. VII. 1988 ”, “ Robert W. Brooks ex., at lights, BIOLAT-SI / MAB ” (2 exs VAIP); “ El Beni, El Porvenir Station, NE of San Borja, 6 – 9. VIII. 1988 ”, “ Robert W. Brooks ex., at lights, BIOLAT-SI / MAB ” (1 ex. VAIP); “ Santa Crus, Sara, 2 km NW Santa Rosa, 21. II. 1969, BLT A. Martinez & R. E. Woodruff ” (5 exs USNM); idem “ 3 km S Santa Rosa, 22. II. 1969 ” (7 exs USNM). Ecuador: “ Esmeraldas, 9 km S San Lorenzo, 25. V. 1979, ultraviolet light, Jos. J. Anderson ”, “ Ecuador Peace Corps, Smithsonian Institution Aquatic Insects Survey ” (3 exs USNM – one ex. without label); “ Los Rios, Babahoyo, 21. VI. 1975, Cohen, Langley & Monnig ”, “ Ecuador Peace Corps, Smithsonian Institution Aquatic Insects Survey ” (2 exs USNM). Panamá: “ C. Z. [= Canal Zone], Albrook Forest site, 21 – 22. IX. 1967, BLT RS Hutton ” (1 ex. USNM). Perú: “ Prov. Loreto, Rio Yarapa, Puerto Miguel ”, “ 200 m, 16 – 23. XII. 1994, T. Hàcz & G. Holzinger ” (3 exs HNMB); “ Loreto, Yacumama Lodge, 73.6 ° W, 4.8 ° S, nr. junction Rio Maranon - Rio Ucayali, 6 – 20. VIII. 1994, Skelly leg., small light in wood, coll. L. Hendrich ” (8 exs ZSM); “ Colonia Calleria, Rio Calleria, 15 km E Ecayali, muddy river shore, 12 – 25. X. 1961, B. Malkin ” (2 exs USNM); “ Madre de Dios, Rio Tambopata Res., 30 air km SW Pto. Maldonado, 290 m, 2 – 5. XI. 1979, J. B. Heppner, subtropical moist forest ” (3 exs USNM). Paraguay: “ Jejuí-Mí, light trap, 18. XII. 2003, leg. M. C. Michat ” (2 exs MCMC); “ Dep. Alto Paraguay, Campo Grande Farm, light trap, 7. XII. 2002, leg. M. C. Michat ” (2 exs MCMC); “ Dep. Alto Paraguay, rd. to Fortín Patria, 3. XII. 2002, light trap, leg. M. C. Michat ” (4 exs MCMC); “ Dep. Alto Paraguay, Defensores del Chaco N. P., Fortín Madrejón, 23. XI. 2002, light trap, leg. M. C. Michat ” (2 exs MCMC); “ Dep. Cordillera, Arroyos y Esteros, 8. IX. 1991, Ulf Drechsel leg., Coll. Hendrich ” (1 ex. ZSM); “ Dep. Central, Asuncion, Rio Paraguay, 2. XI. 1990, Ulf Drechsel leg., Coll. Hendrich ” (1 ex. ZSM). Suriname: “ Paramaribo, Guest House of Museum, at light, 19 – 20. VII. 1969, leg N. Nieser ” (1 ex. NMW); idem except for “ 21 – 25. VIII. 1969 ” (1 ex. NMW).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	diagnosis	Diagnosis. Recognisable from most species of Laccomimus by aedeagal characters only, and from the very close L. pumilio (see above) by aedeagal characters and, frequently, by the more distinct pattern on elytra. Median lobe differentiated into basal and distal portion, divided by an angle or a curve (Figs 94 – 102), with apex ending in small button-like expansion. The rounded tip of the prosternal process, which is very different from the acuminate tip of L. pumilio, is constant in this species. The average size is smaller with respect to L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	description	Description. Habitus (Fig. 64). TL: 1.9 – 2.25 mm; MW: 0.95 – 1.2 mm (holotype: TL: 2.2 mm; MW: 1.1 mm); A-TL: 2.1 mm; A-MW: 1.1 mm (NUM: 47 exs); very similar to L. pumilio except for aedeagal characters, shape of prosternal process and, frequently, elytral colouration. Colour. Variable. Pale patterns of elytra normally (but not always) more sharply defined on darker background and often more extended with respect to L. pumilio. Most commonly pale subbasal band of elytra fragmented in semicircular spots, although strong variability occurs (see below). Pronotum normally with narrower dark transversal band, more evidently bicolour; head, antennae, mouthparts, legs, and underside as in L. pumilio. Structures and sculpture. Mostly as in L. pumilio, but prosternal process rounded at tip and elevated, not carinate (Figs 28; 69); iridescence on pronotum and mainly on elytra often more intense. Male. Last abdominal ventrite as in L. pumilio. Outer claw of forelegs scimitar-shaped, variable in shape, often with denticles on ventral margin (Figs 77 – 81). Median lobe of aedeagus in dorsal view as in Fig. 111; same as in L. pumilio (see description above). In left lateral view variable (Figs 94 a; 95 a, 96 a; 97 a; 98 a; 99 a; 100 a; 101 a; 102 a): in holotype (Fig. 94 a) base and distal portion forming a curve instead of an angle, distal portion less elongate with respect to L. pumilio and more uniformly wide along its length, although visibly sinuate on apical half. Left paramere (Figs 94 c; 95 c; 96 c; 97 c; 98 c; 101 c; 102 c) strongly sinuate apically, expanded and thickened. Female. Often duller than males, with more intense iridescence. Genital pieces (Fig. 118) as in L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	distribution	Distribution. Laccomimus alvarengi is one of the most widespread and variable member of Laccomimus: Argentina, Bolivia, Brazil, Ecuador, Panamá, Paraguay, Perú, Suriname (Figs 125; 128). It has not been recorded from Venezuela, however, in spite of VAIP’s intensive collections during the last decade.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	etymology	Etymology. Frank N. Young and Paul J. Spangler named, in litteris, this taxon “ Laccomimus alvarengi n. sp. ”, in honour of the collector Moacir Alvarenga (cf. Young 1974: 2, 3). The specific epithet is a noun in the genitive case. Notes on variability. L. alvarengi is more variable than L. pumilio. Except for the striking differences in the prosternal process, morphological separation of the two species is often difficult. The visibly sinuate and expanded apex of the left paramere and the less broadened distal portion of the median lobe seem to be constant diagnostic characters of L. alvarengi with respect to L. pumilio. In other features the median lobe is very variable, and this variability has been observed in different populations. Intermediate forms between the holotype of L. alvarengi and Florida specimens of L. pumilio have been observed frequently, even in exemplars from the same locality (see for example Fig. 97). In most cases, however, the aedeagus is distinctive with respect to L. pumilio, particularly in the angulated separation between the base and distal portion and the somewhat widened distal portion. An aberrant median lobe was found dissected from a specimen collected at the type locality (Fig. 95). Colouration is also very variable. Normally the elytral patterns in L. alvarengi are more defined than in L. pumilio, and some populations exhibit very bright elytral patterns, including large and complete pale subbasal bands (Fig. 64 a, b). We have observed, however, specimens (especially from Brazil) with obscure or reduced pale markings, similar to L. pumilio. Uniformly dark-chestnut elytra occur in few specimens from Brazil and Paraguay. Since the shape of the prosternal process unambiguously separates L. alvarengi from L. pumilio, and a visibly sinuate and expanded apex of the left paramere and a less broadened distal portion of the median lobe are also apparently constant differences, we decided to describe L. alvarengi as a distinct species and not as a subspecies within L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE3FFE691A6F8B2FDCAA469.taxon	discussion	Remarks. See under L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	materials_examined	Type locality. Venezuela, Guarico State, 8 ° 31.705 ' N, 66 ° 22.602 ' W, 145 m, ca. 65 km S Las Mercedes, large lagoon with vegetation. Type material. Holotype male (MIZA): dissected by us; last two metaventrites and genitaia included in DMHF on plastic label, pinned under the holotype “ Guarico State, 8 ° 31.705 ' N, 66 ° 22.602 ' W, 145 m, ca. 65 km S Las Mercedes, large lagoon with vegetation, 9. I. 2009, leg. Short, Miller, Garcia & Camacho, VZ 09 - 0109 - 01 X ” [printed], “ KUNHM-ENT SM 0845078 ” [printed, together with barcode], “ HOLOTYPE Laccomimus improvidus Toledo & Michat ” [red, rectangular, printed label]. Paratypes (26 exs, all bearing red, printed label “ PARATYPE Laccomimus improvidus Toledo & Michat ”): Venezuela: same data of holotype (1 male VAIP); “ Monagas State, 9 ° 36.591 ' N, 63 ° 8.295 ' W, 45 m, S. of Marturin, vegetated river ”, “ morichal margin, 2. II. 2010, leg. Short & Garcia, VZ 10 - 0202 - 01 B ” (1 male VAIP); “ Monagas State, 8 ° 46.439 ' N, 62 ° 14.306 ' W, 19 m, between Tucupita and Temblador, pond margin, 3. II. 2010, leg. Short, Garcia & Joly, VZ 10 - 0203 - 02 A ” (1 male VAIP); each one of these two latter specimens bearing second label (KUNHM-ENT) with its own barcode and catalogue number; “ Km 88 El Dorado, Santa Elena BO, 160 m, 14. XI. 1966, J. & B. Bechyne & E. Osuna ” (1 male, 3 females MIZA); “ Bolivar, Guri, Rio Caroni, 100 m, 16. XI. 1966, J. & B. Bechyne & E. Osuna ” (1 female MIZA); “ Bolivar, Ana [illegible letter] oco, 60 m, 6 ° 5 ', N 61 ° 8 ' W, 10 – 30. VIII. 1979 ” (1 male MIZA); “ Aragua, El Limon, 450 m, 13. VII. 1977, leg. B. Bechyne ” (1 female MIZA); “ Coje, Galeras del Pao, 28. VI. 1963, leg. C. J. Rosales & A. Perez ” (2 females MIZA); all these specimens in MIZA, bearing an additional printed green label “ MIZA ”. Suriname: “ Paramaribo, Guest House of Museum, at light, 20. VIII – 23. IX. 1969, leg. N. Nieser ” (1 male, 2 females NMW); idem except for “ 21 – 25. VIII. 1969 ” (1 female NMW); idem except for “ 26. VII – 7. VIII. 1969 ” (1 female NMW); “ 55 ° 06 ' W, 05 ° 45 ' N, Oost-West Verbinding, 7 – 8 km E Meerzorg, 2. X. 1969, leg N. Nieser (SN 157) ” (1 male NMW); “ 55 ° 11 ' W, 05 ° 31 ' N, N banks of Coropinakreek, 25. IX. 1969, leg N. Nieser (SN 156 A) ” (1 female NMW). Guyana: “ Karanambo, 3 ° 45.1 ' N, 59 ° 18.6 ' W, blacklight on bank of Rupununi, 31. III. 1994, P. J. Spangler, colln # 2 ” (1 female USNM); idem except for “ Buffalo Pond, 24. IV. 1994, colln # 11 ” (1 male USNM); “ Pirara Ranch, 3 ° 32.1 ' N, 59 ° 40.5 ' W, Pirara River, 24. IV. 1995, P. J. Spangler & A. Perry, colln # 18 ” (1 female USNM).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	diagnosis	Diagnosis. Externally indistinguishable from other species of Laccomimus, in particular from dark specimens of L. alvarengi and L. malkini with marked elytral pattern. Examination of female genitalia is also not diagnostic, except from L. malkini; male genitalia are distinctive, however, with median lobe more straight and robust than in L. pumilio, with base and distal portion forming a more obtuse angle and apex ending in a wider expansion (Fig. 89 a, b); left paramere similar to that of L. alvarengi, although not or very poorly thickened apically (Fig. 89 c).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	description	Description. TL: 2.0 – 2.25 mm; MW: 1.0 – 1.2 mm (holotype: TL: 2.2 mm; MW: 1.1 mm); A-TL: 2.2 mm; A- MW: 1.1 mm (NUM: 19 exs); drop-shaped, regularly tapering posteriorly; elytral subapical sinuation missing or hardly visible. Iridescence well visible. Colour. Head and pronotum markedly paler than elytra. Head uniformly reddish-yellow, at most slightly darkened behind eyes; pronotum of same colour as head, with two brown, rectangular markings along both sides of hind margin. Elytra chestnut-brown, with vague yellow subbasal markings normally fragmented in spots; rarely missing. Apical fourth more or less gradually paler; a poorly defined paler area is often visible on lateral side. Underside uniformly testaceous to reddish-yellow. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker. Structures and sculpture. Head smooth, shiny, with fine microreticulation of rounded cells. Pronotum with microreticulation as on head, shiny, with faint iridescence. Smooth, lacking punctures except along hind and fore margins. Elytra apically not markedly pointed and subapical sinuation poorly or not pronounced; surface smooth, with punctures very scarce or lacking and often with few, very shallow, gnarled scratches; reticulation poorly impressed; iridescence well visible. Prosternal process and lobes of metacoxal processes as in L. alvarengi. Male. Last abdominal ventrite tectiform, with antero-lateral sides markedly sinuate. Outer claw of forelegs scimitar-shaped. Median lobe of aedeagus very similar to that of L. pumilio except as follows: in left lateral view (Fig. 89 a, b) more straight and robust, with base and distal portion forming a more obtuse angle; apex ending in a wider expansion, laterally flattened; in dorsal view (Fig. 112) more gently attenuate at tip, almost straight, and dorsal lamina more strongly sinuate. Left paramere (Fig. 89 c) very similar to that of L. alvarengi, although not or very poorly thickened apically. Female. Elytra duller than in males, with more evident iridescence. Last abdominal ventrite triangular, very narrowly rounded to subtruncate at apex. Genital pieces as in L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	distribution	Distribution. Venezuela, Guyana, Suriname (Fig. 129). Habitat. In Guarico State (Venezuela) some specimens were collected in a large lagoon with vegetation, exposed to direct sunlight (Fig. 133).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	etymology	Etymology. Latin adjective for “ surprising ”, “ unexpected ”; quite late, in fact, we recognised it as a separate species.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE1FFE791A6FB1CFE6EA683.taxon	discussion	Remarks. This species is close to L. pumilio and L. alvarengi. At first, after dissecting the first male specimen (Suriname, Paramaribo), we were convinced that it was a dark specimen of L. alvarengi with an aberrant median lobe. We also considered that it could represent a possible case of hybridisation between L. alvarengi and L. bolivari since the median lobe appears as a broader tip added to an L. alvarengi median lobe. Both species occur at that locality. Additional material found in USNM and NMW, however, made it clear that the specimens belong to an undescribed species, very uniform in elytral colouration and in aedeagal features. Apparently it is a rare species, at present known only from few specimens, often found together with other species. The females are impossible to identify with certainty.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	materials_examined	Type locality. Venezuela, Higuerote. Type material. Holotype male (USNM): dissected probably by Young; genitalia and last two abdominal ventrites glued on separate label under the holotype “ Venezuela ”, “ Higuerote m. 5, (Barlovento - Edo. Miranda) ”, “ 4 febr. 1962, leg. Bordon ” [three rectangular, white labels, printed], “ Holotype Laccomimus bordoni Young 69 ” [red, handwritten by Young], “ HOLOTYPE Laccomimus bordoni Toledo & Michat ” [red, rectangular, printed label]. Paratypes (218 exs, all bearing red, printed label “ PARATYPE Laccomimus bordoni Toledo & Michat ”): Venezuela: same data as holotype, except for white label, instead of red, handwritten by Young “ Laccomimus bordoni Young ' 69 ” (12 exs USNM); “ Bolivar, El Dorado (65 km S), 1. XII. 1982, John L. Hellman ” (4 exs USNM). Argentina: “ Prov. Misiones, Yacutinga res., 4 – 3. II. 2006, light trap, leg. M. C. Michat ” (1 ex. MCMC); “ Prov. Jujuy, Calilegua N. P., lagoon, 16. XII. 2003, leg. M. C. Michat ” (1 ex. MCMC); idem “ 18. IX. 2005 ” (2 exs MCMC); “ Prov. Chaco, 97 km NW of Resistencia, 10 km W of Capitán Solari, 26 ° 48 ' 32 " S, 59 ° 36 ' 29 " W, 3 – 7. II. 2004 [collector not given] ” (1 ex. NHMP). Bolivia: “ Guayaramerin, Beni, river bank, Soil Zoological Exp., Balogh, Mahunka, Zicsi ”, “ No. 428, beaten from vegetation, 2. XII. 1966 ” (5 exs HNMB); “ Guayaramerin, Beni, outskirts of town, Soil Zoological Exp., Balogh, Mahunka, Zicsi ”, “ No. 391, lamping, 22. XI. 1966 ” (2 exs HNMB); idem “ No. 424, lamping., 1. XII. 1966 ” (2 exs HNMB); “ Santa Cruz, San Antonio, Rio de Parapeti, Rio Parapeti, 15 – 19. VII. 1964, B. M. Malkin ” (1 ex. USNM); “ Santa Cruz, Sara, 2 km NW Santa Rosa, 21. II. 1969, BLT A. Martinez & R. E. Woodruff ” (3 exs USNM); idem, “ 3 km S Santa Rosa, 22. II. 1969 ” (10 exs USNM); “ Santa Cruz, 60 mi. N Santa Cruz, Saavedra Expl. Sta., 27 – 30. XII. 1959, Robert Cumming, collected at black light ” (1 ex. USNM). Brazil: “ Mato Grosso, Cuiabà (Cuyabà), 30. III. 1972, BLT W. H. Whitcomb ” (1 ex. USNM); idem “ 11. IV. 1972 ” (1 ex. USNM). Dominican Republic: “ Sanchez, VII. 1938, Darlington ” (2 exs USNM); “ Dajabon Province, 13 km S. Loma de Cabrera, ca. 400 m, 20 – 22. V. 1973, Don & Mignon Davis ” (1 ex. USNM); “ Juan Dolio 10 – 18. XII. 2005, R. Fencl leg. ” (3 exs NHMP). Jamaica: “ Porus, 23. II. 1937 ”, “ Sta 414, Chapin and Blackwelder ”, “ Flying at dusk ” (2 exs USNM); “ Try., Good Hope, VIII. II. 1966, A. T. Howden ” (2 exs USNM); “ Parrish, St. Catherine, Worthy Park Est. ”, “ R. E. Woodroof, 17. V. 1969, blacklight trap ” (3 exs USNM). Haiti: “ Miragoane, 30. X – 2. XI. 1934, Darlington ” (1 ex. USNM); “ Etang Lac haux, SW peninsula, under 1000 ft, 26 – 27. X. 1934, Darlington ” (1 ex. USNM). Panamá: “ Canal Zone, Albrook Forest site, 28 – 29. V. 1968, BLT R. S. Hutton ” (2 exs USNM); “ Canal Zone, 5 mi SSW Gatun Dam, 30. XII. 1987, MacDonald & Schiefer ”, “ William H. Cross Expedition ” (2 exs MEM); “ Darien, 5 km W Meteti, 9. I. 1988, MacDonald & Schiefer ”, “ William H. Cross Expedition ” (1 ex. MEM). Paraguay: “ Dep. Guaira, Garay, 20. VIII. 1992, U. M. Drechsel leg ” (1 ex. LHB); “ Dep. Boqueron, X. 1990, Col. New Land ” (1 ex. NMW); “ Dep. Central, Capiata, 3. X. 1994 ” (1 ex. NMW); “ Jejuí-Mí, 18 – 20. XII. 2003, light trap, leg. M. C. Michat ” (22 exs MCMC); “ Dep. Alto Paraguay, rd. to Fortín Patria, 3. XII. 2002, light trap, leg. M. C. Michat ” (12 exs MCMC); “ Dep. Alto Paraguay, Defensores del Chaco N. P., Fortín Madrejón, 23. XI. 2002, light trap, leg. M. C. Michat ” (7 exs MCMC); “ Dep. Presidente Hayes, Transchaco rd., 5 km SE Pozo Colorado, 22. XI. 2002, light trap, leg. M. C. Michat ” (1 ex. MCMC); “ Dep. Concepcion, Est. San Luis, 8. IX. 1997, Bach im Cerrado-Wald, Uferschotter [= brook in Cerrado-Forest, shore gravel], leg. Drechsel ” (1 ex. NMW). Virgin Islands: “ St. Croix, Bethlehem, grassy pond, 11 – 15. II. 1966, Paul J. Spangler ” (107 exs USNM, MTP). Additional material studied. Jamaica: Moneague, 26. VIII. 1934, Darlington (2 females USNM – most likely belonging to this species).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	diagnosis	Diagnosis. Dorsally very similar to other species of the genus, except for (frequently) very elongate body and elytra markedly sinuate subapically. Underside dark, at least in part, darker than in any other Laccomimus species. Median lobe of aedeagus distinctive, apex ending in characteristic small, bird head-like expansion (Fig. 103 a, b).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	description	Description. Habitus (Fig. 65). TL: 2.1 – 2.5 mm; MW: 1.0 – 1.3 mm (holotype: TL: 2.3 mm; MW: 1.2 mm); A- TL: 2.3 mm; A-MW: 1.2 mm (NUM: 67 exs); body elongate, drop-shaped, elytra often markedly sinuate subapically. Colour. Head and pronotum paler than elytra. Head testaceous or reddish-brown, frequently darker between eyes; pronotum testaceous to reddish, often with darker, elongate marking along hind margin. Elytra brown, with quite sharply defined pale subbasal band, often fragmented in spots; subapical lateral yellow band extending to apex; a third, lateral, less defined yellow marking is frequently visible. Underside brown or dark brown, normally head, prosternum, epipleura, metacoxal processes, and last abdominal ventrite paler. Dark areas always present, at least in part, although very variable in extension. Antennae and mouthparts testaceous. Legs uniformly testaceous except for dark metatarsi. Structures and sculpture. Head smooth, shiny, with fine microreticulation of rounded cells. Pronotum with microreticulation as on head, shiny, with faint iridescence. Few scattered punctures mostly along fore and hind margins. Elytra with subapical sinuation quite pronounced; smooth with quite impressed reticulation and intense iridescence. Prosternal process (Figs 29 – 31) slightly elevated, rounded at tip. Lobes of metacoxal processes rounded in both sexes (Fig. 40). Male. Last abdominal ventrite tectiform, with antero-lateral sides markedly sinuate (Fig. 42). Outer claw of forelegs scimitar-shaped (Figs 50; 70). Median lobe of aedeagus, in left lateral view (Fig. 103 a, b), slender but robust, base and distal portion forming a very wide angle, then straight, regularly narrowed to apex, ending in characteristic small, bird head-like expansion. In dorsal view (Fig. 113) similar to that of L. pumilio. except for a visible constriction on left side, close to base, and apical fourth strongly narrowed with hooked tip; dorsal lamina straight along its whole length; ejaculator groove wide, gradually narrowed to apex. Left paramere (Fig. 103 c) curved, gradually tapering to apex. Female. Last abdominal ventrite triangular, very narrowly rounded to subtruncate at apex (Fig. 43). Lateral tergites thin, partly membranous. Gonocoxal blade as in preceding species; ramus (Fig. 120) similar to that of L. pumilio. except for teeth a bit larger.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	distribution	Distribution. Argentina, Bolivia, Brazil (Mato Grosso), Dominican Republic, Haiti, Jamaica, Panamá, Paraguay, Venezuela, Virgin Islands (Figs 126; 128). Ecology. Field data suggest a wide ecological range with collections from both stagnant and running waters. In Young’s letter to Spangler, the following field observations are given for this species in the Virgin Islands: “ adults and larvae of Laccomimus were collected from shallow water at the edge of the pond in an open field on St. Croix (…). They were found beneath a mat of wiry grass growing over the margin of the pond. By bending the mat of vegetation back onto the bank and anchoring it, the shallow margins of the pond were exposed and could be easily worked with a dip net. Vegetation accumulated in working the pond margin with the net was dumped onto a ground cloth. Adult Laccomimus were surprisingly agile. They jumped rapidly much like fleas as soon as they were dumped onto the cloth and they often jumped completely off the cloth and escaped. Most of the adults would have escaped without the rapid use of a suction bulb aspirator. The slow-moving larvae could be easily hand picked from the cloth. ” In Calilegua National Park (Argentina) some specimens were collected from the margins of a large vegetated pond at 750 m a. s. l., partially exposed to direct sunlight, with turbid water rich in organic matter, sparse marginal vegetation and absence of floating vegetation (Fig. 130).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	etymology	Etymology. The specific name is a noun in the genitive case which was given in litteris by Frank N. Young and Paul J. Spangler, in honour of the collector Carlos Bordon (cf. Young 1974: 2, 3). Notes on variability. In spite of its wide distribution, no relevant morphological variation was observed between populations. An exception is the partially dark underside, which is always present, but varies in extent (often this is the only diagnostic feature to distinguish females of this species). The shape of the median lobe is very constant among the studied specimens.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE0FFE191A6F8B2FDA8A551.taxon	discussion	Remarks. This species is similar to L. pumilio in the modified male anterior claws, the shape of the prosternal and metacoxal processes, and the female genitalia. However, it is highly characteristic in the dark underside and in the aedeagal features mentioned above.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	materials_examined	Type locality. Mexico, Sinaloa, Mazatlan. Type material. Holotype male (USNM): dissected by us; median lobe of aedeagus and parameres included in DMHF on plastic label, pinned under the holotype “ Mexico, Sinaloa, Mazatlan, VII. 17 – 23.1963, P. & P. Spangler ” [rectangular, white label, printed], “ HOLOTYPE Laccomimus spangleri Toledo & Michat ” [red, rectangular, printed label]. Paratypes (6 exs, all bearing red, printed label “ PARATYPE Laccomimus spangleri Toledo & Michat ”): Mexico: same data as holotype; two specimens bearing the following additional label “ Collected in blk litetrap ” [smaller white, rectangular, printed label] (3 exs USNM). Costa Rica: “ 16 Mi. S La Cruz, VII. 13.1965, Paul J. Spangler ”, “ Laccomimus det. PJ Spangler ” (2 exs USNM).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	diagnosis	Diagnosis. Externally very similar to most of the species of the genus, but visibly smaller. Median lobe of aedeagus distinctive, distal portion straight, almost parallel-sided up to abruptly narrowed apex, ending in a small button-like expansion (Fig. 104 a).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	description	Description. Habitus (Fig. 66). TL: 1.8 – 2.1 mm; MW: 0.9 – 1.0 mm (holotype: TL: 1.9 mm; MW: 1.0 mm); A- TL: 1.9 mm; A-MW: 1.1 mm (NUM: 6 exs); smaller than any other Laccomimus species; body quite broad, regularly tapering to elytral apex. Colour. Head and pronotum paler than elytra. Head testaceous or reddish-brown, often darker between eyes; pronotum testaceous, with more or less defined dark, elongate marking along hind margin. Elytra chestnut-brown, with quite defined pale subbasal band, fragmented in irregular spots; subapical lateral yellow area reduced to a narrow lateral band or completely obsolete. Whole underside testaceous or pale-brown. Antennae and mouthparts testaceous. Legs testaceous. Structures and sculpture. Head smooth, shiny, with fine microreticulation of rounded cells. Pronotum with microreticulation as on head, shiny, with faint iridescence. Surface with few scattered punctures. Elytra gradually tapering, with subapical sinuation obsolete; smooth, with poorly impressed reticulation but visible and quite intense iridescence. Prosternal process slightly elevated, almost acuminate at tip. Lobes of metacoxal processes regularly rounded in both sexes. Male. Last abdominal ventrite tectiform, pointed, with anterolateral sides rectilinear. Outer claw of forelegs scimitar-shaped (Fig. 72). Median lobe of aedeagus, in left lateral view (Fig. 104 a), elongate, slender, base and distal portion forming an angle of about 100 °; distal portion straight, almost parallel-sided up to abruptly narrowed apex, ending in a small button-like expansion. In dorsal view (Fig. 114) straight and slender, with almost parallel sides, then gradually narrowed and turned to right; apex quite dilated; dorsal lamina covering distal half of ejaculator groove. Left paramere (Fig. 104 b) sabre-shaped, gradually tapering to apex. Female. Not duller than male. Last abdominal ventrite subtruncate, with small medial notch at apex. Genital pieces as in L. pumilio.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	distribution	Distribution. Mexico, Costa Rica (Fig. 128).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	etymology	Etymology. Dedicated to the late Paul J. Spangler, who collected this species from both known localities. The specific epithet is a noun in the genitive case. Notes on variability. The only specimen seen from Costa Rica (the second one is known only from a single elytron) is the smallest known specimen of Laccomimus (TL: 1.8 mm; MW: 0.9 mm).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE6FFE291A6FA08FCB8A49E.taxon	discussion	Remarks. Laccomimus spangleri is close to L. pumilio and similar species, sharing with them external male characters, similar prosternal and metacoxal processes, and similar female genitalia. It is, however, well characterised by aedeagal features and by its very small size.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	materials_examined	Type locality. Venezuela, Bolivar State, Los Pijiguaos, 6 ° 35.617 ' N, 66 ° 49.238 ' W, 80 m (Fig. 134). Type material. Holotype male (MIZA) dissected by us; genitalia glued on the same rectangular card as holotype “ Venezuela, Bolivar State, 6 ° 35.617 ' N, 66 ° 49.238 ' W, 80 m, Los Pijiguaos, outcrop / morichal, 12. I. 2009, leg. Short, Garcia, Camacho, Miller & Joly, VZ 09 - 0112 - 01 A, morichal margin ” [printed], “ KUNHM-ENT SMC 0856789 ” [printed, together with barcode], “ HOLOTYPE Laccomimus bolivari Toledo & Michat ” [red, rectangular, printed label]. Paratypes (87 exs, all bearing red, printed label “ PARATYPE Laccomimus bolivari Toledo & Michat ”): Venezuela: same data as holotype (4 exs VAIP); “ Bolivar State, 5 ° 50 ' 49.2 " N, 61 ° 28 ' 2.4 " W, 1340 m, Rio Aponowao at Hwy 10, 31. VII. 2008, leg. A. Short, M. Garcia, AS- 08 - 060 a, small vegetated pool ” (1 ex. VAIP); “ Amazonas State, 5 ° 20.514 " N, 67 °. 45,315 " W, 87 m, S- Communidad Porvenir, 15. I. 2009, leg. Miller & Short, VZ 09 - 0115 - 03 B: small streamlet ” (19 exs VAIP, MTP); “ Amazonas State, 5 ° 20.479 ' N, 67 ° 45.358 ' W, 72 m, ca. 12 km N Samariapo, small stream at rd x-ing, 6. I. 2006, AS- 06 - 013, leg. A. E. Z. Short ” (2 exs VAIP); “ Amazonas State, 5 ° 16.639 ' N, 67 °. 48.044 ' W, 60 m, ca. 4 km N Samariapo, small stream at rd x-ing, 6. I. 2006, AS- 06 - 012, leg. A. E. Z. Short ” (2 exs VAIP); “ Amazonas State, 5 ° 48.414 ' N, 67 ° 26.313 ' W, 80 m, nr. Iboruwa: " Tobogancito ", 13. I. 2009, leg. Short et al., VZ 09 - 0113 - 02 X ” (1 ex. VAIP); “ Guarico State, 8 ° 31.705 ' N, 66 ° 22.602 ' W, 145 m, ca. 65 km S Las Mercedes, 9. I. 2009, leg Short, Miller, Garcia & Camarcho, large lagoon w / veg., VZ 09 - 0109 - 01 X ” (2 exs VAIP); “ Guarico State, 9 ° 46.320 ' N, 67 ° 21.177 ' W, 280 m, Rio San Antonio, N Dos Caminos, 8. I. 2009, leg K. B. Miller, mic. Habitats, VZ 09 - 0108 - 02 C ” (1 ex. VAIP) [each specimen in VAIP bearing second label (KUNHM- ENT) with its own barcode and catalogue number]; “ Guarico, 12 km S Calabozo, 11 – 6. XII. 1969, P. & P. Spangler ”, “ Est. Biologica Los Llanos, black light ” (3 exs USNM). Suriname: “ Brokopondo, 55 ° 10 ' W, 04 ° 59 ' N, rd. Brownsberg – Brownsweg, 24. X. 1969, leg. N. Nieser (SN 201 A) ” (1 ex. NMW). Trinidad: “ Cumuto, Trinidad, 1929, Darlington ”; all printed label, except for one specimen, handwritten, presumably by Young; two specimens with additional label “ Laccomimus trinidadensis sp. nov. det. F. N. Young ” handwritten by Young (51 exs NMHS).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	diagnosis	Diagnosis. Externally hard to tell apart from other members of Laccomimus. Aedeagal characters distinctive, median lobe with angle between base and distal portion almost indistinct, apex ending in wide and flattened expansion (Figs 105 a; 106 a); left paramere straight (Fig. 105 c). Females impossible to identify without association with males.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	description	Description. Habitus (Fig. 1). TL: 2.0 – 2.4 mm; MW: 1.0 – 1.2 mm (holotype: TL: 2.3 mm; MW: 1.2 mm); A- TL: 2.1 mm; A-MW: 1.1 mm (NUM: 35 exs); very similar to L. alvarengi in body shape and colouration; elytra markedly sinuate subapically; ventral surface uniformly testaceous. Colour. Head, pronotum, and elytra as in L. alvarengi. Pale markings on elytra always well defined, subbasal band in most cases complete, occasionally fragmented in spots; lateral markings missing or very reduced. Subapical band extending to lateral margin, not reaching elytral apex. Underside testaceous, slightly darker along sutures and apex of abdominal ventrites 3, 4 and 5. Antennae and mouthparts testaceous. Legs testaceous; metatarsi slightly darker. Structures and sculpture. Head smooth, shiny, with hardly visible microreticulation of rounded cells. Pronotum with quite impressed microreticulation, with iridescence visible as on elytra. Few scattered punctures mostly along fore and hind margins. Elytra lacking subapical sinuation; less narrowed at tip than preceding species; smooth with quite impressed reticulation and intense iridescence. Underside with faint iridescence. Prosternal process quite elongate and visibly carinate, almost acute at tip; lobes of metacoxal processes ending in obtuse angulation (Fig. 41). Male. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped (Fig. 74). Median lobe of aedeagus, in left lateral view (Figs 105 a; 106 a), elongate, almost straight, with angle between base and distal portion almost indistinct; gradually narrowed to apex, ending in characteristic, wide and flattened expansion. In dorsal view (Fig. 115), slender, gently but visibly turned to right; ejaculator groove gradually narrowed to apex, closed before apical fifth of median lobe. Left paramere (Fig. 105 c) straight, gradually tapering to apex. Female. Iridescence as in male or slightly more intense. Last abdominal ventrite triangular, very narrowly rounded at apex, almost pointed. Lateral tergites thin and partly membranous. Gonocoxal blade with single denticle; ramus as in preceding species, with large and spaced teeth.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	distribution	Distribution. Venezuela, Trinidad, Suriname (Fig. 127). Ecology. VAIP’s field data suggest a wide ecological range, with collections from a morichal, a lagoon and a small streamlet (Figs 133 – 134; 136).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	etymology	Etymology. Young and Spangler named in litteris this species Laccomimus trinidadensis, based on material in USNM, in great part collected from Trinidad. However, it is clear that this species has a much wider distribution. We decided to name this taxon bolivari in reference to the type locality and in honour of the Venezuelan patriot Simón José Antonio de la Santísima Trinidad Bolivar, “ El Libertador ”. The specific epithet is a noun in the genitive case. Notes on variability. The only specimen seen from Suriname has the median lobe in lateral view thicker in distal portion and more abruptly narrowing to apex in distal fourth; the apical expansion is slightly smaller (Fig. 106 a).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE5FFE391A6FB54FB71A6DE.taxon	discussion	Remarks. This species is close to L. pumilio and related species, sharing with them external male characters, similar female genitalia, and a similar shape of the prosternal and metacoxal processes. It is, however, well characterised by aedeagal features, unusual compared to the other species of Laccomimus.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	materials_examined	Type locality. Brazil, Mato Grosso, Jacaré, Parque Nacional Xingu. Type material. Holotype male (USNM): dissected by us; median lobe of aedeagus and parameres included in DMHF on plastic label pinned under the holotype “ Brazil, MG [= Mato Grosso], Jacare, PNX [= Parque Nacional Xingu], XI. 1965, Alvarenga ” [rectangular, white, handwritten], “ HOLOTYPE Laccomimus youngi Toledo & Michat ” [red, rectangular, printed label]. Paratypes (7 exs, all bearing red, printed label “ PARATYPE Laccomimus youngi Toledo & Michat ”): Brazil: same data as holotype (7 exs USNM: most of them unlabelled: printed label added by the present authors).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	diagnosis	Diagnosis. Very close to L. bolivari but a bit shorter and with less clearly defined markings on elytra. Aedeagus similar to that of L. bolivari but longer, with apex ending in distinctive expansion, resembling an axe (Fig. 90 a).	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	description	Description. TL: 2.0 – 2.3 mm; MW: 1.0 – 1.2 mm (holotype: TL: 2.3 mm; MW: 1.15 mm); A-TL: 2.0 mm; A- MW: 1.1 mm (NUM: 7 exs); body drop-shaped, short, regularly tapering to apex, pale markings poorly defined. Colour (based on old specimens: colouration might be altered). Head and elytra reddish-brown; pronotum paler, with more or less defined dark elongate marking along hind margin. Elytra with poorly defined pale subbasal band – normally not fragmented – and subapical lateral area. Whole underside testaceous or pale-brown. Antennae and mouthparts testaceous. Legs testaceous except for slightly darkened metatarsi. Structures and sculpture. Head and pronotum as in preceding species: iridescence lacking or faint on pronotum. Elytra with hardly visible subapical sinuation, gently narrowed at tip; smooth with quite impressed reticulation and shallow grooves on basal half; iridescence faint. Iridescence on underside missing or almost so. Prosternal and metacoxal processes as in preceding species. Male. Last abdominal ventrite tectiform, with antero-lateral sides visibly sinuate. Outer claw of forelegs scimitar-shaped. Median lobe of aedeagus, in left lateral view (Fig. 90 a), straight as in L. bolivari but longer, with more elongate, narrow, almost stem-like apex, ending in distinctive expansion, widely rounded dorsally, short and acute ventrally, resembling an axe. In dorsal view as in L. bolivari but more slender. Left paramere straight as in L. bolivari but blunt, less acute apically, very short in other dissected specimens (Fig. 90 d, e). Female. Last abdominal ventrite triangular, very narrowly rounded at apex, almost pointed. Genital pieces as in preceding species.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	distribution	Distribution. Brazil, Mato Grosso (Fig. 127). Known only from the type locality.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	etymology	Etymology. Species dedicated to Prof. Frank N. Young. The specific epithet is a noun in the genitive case.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
03EE87FEFFE4FFEC91A6F889FEE2A4FB.taxon	discussion	Remarks. L. youngi is very close to L. bolivari, from which it can be distinguished only by minor aedeagal differences.	en	Toledo, Mario, Michat, Mariano C. (2015): Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990 (3): 301-354, DOI: 10.11646/zootaxa.3990.3.1
