taxonID	type	description	language	source
03EEDE4BFFE1FFB9EF03FD83B2CAF9EB.taxon	description	(Fig. 2 A)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE1FFB9EF03FD83B2CAF9EB.taxon	materials_examined	LECTOTYPE (by inference of holotype by Türkay 1974 a: 969). — MNHN-IU- 2000 - 3735 (= MNHN-B 3735), ♂ 72.8 × 88.1 mm, Samoa, ‘ Astrolabe’ Exp., presumed Type, D. Guinot 1972. — Original label: “ Cardisoma urvillei Edw., ♂, C. obesum Dana?, Samoa, Exp. de l’ ‘ Astrolabe’ ”; on the right chela: “ C. urvillei, Samoa ”. — Additional label: “ Type présumé, D. Guinot 1972 ”. PRESERVATION. — Dry. Carapace with a crack on right gastric / cardiac region; RP 2, RP 3 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE1FFB9EF03FD83B2CAF9EB.taxon	discussion	REMARKS Türkay (1974 a: 969, 973) considered the specimen MNHN-IU- 2000 - 3735 (= MNHN-B 3735) the only type specimen and regarded it as the holotype. Henri Milne Edwards (1853) mentioned that the chelae changed according to the age, a sentence suggesting that several individuals had been seen and the description not based on a unique specimen. Since the original description was based on more than one specimen, a holotype could not be assumed. By inference that the specimen was a holotype, Türkay (1974 a) can be deemed to have designated that specimen as the lectotype (Code, Art. 74.6). The Astrolabe Expedition (1826 - 1829) directed by Dumont d’Urville visited Samoa: in the CAA, 1829 is the entry date for material collected during this exploration by “ Quoy & Gaimard ” (see Appendix). The holotype of Cardisoma obesum Dana, 1851, another junior subjective synonym of C. carnifex, was recently discovered in the USNM (USNM 2356). This dried holotype now consists of only the carapace and a part of the cephalothorax only, so its sex cannot be determined (Ng 2017: 5, figs 4, 5).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE1FFB7EFECF927B50DFCB5.taxon	description	(Fig. 2 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE1FFB7EFECF927B50DFCB5.taxon	materials_examined	PARATYPES (by original designation). — MNHN-IU- 2014 - 11214 (= MNHN-B 26950), 1 ♀ 22.1 × 25.7 mm, station 85 - 062, Tuala Cave, Panglao Island, Bohol, 0 - 1 m depth, mud bottom, in dark section, salinity 4 ppt., with dip net, coll. T. M. Iliffe, 3. IV. 1985. — MNHN-IU- 2014 - 11215 (= MNHN-B 26951), 1 ♂ 15.5 × 17.7 mm, station 85 - 067, Calingoob Cave, Panglao Island, Bohol, 1 m depth, bottom rocks, with dip net, coll. T. M. Iliffe, 6. IV. 1985. — MNHN-IU- 2014 - 11216 (= MNHN-B 26952), 1 ♂ 15.8 × 18.0 mm, station 85 - 077, Underpass Cave, Panglao Island, Bohol, 1 m depth, salinity 8 ppt, coll. T. M. Iliffe, 8. IV. 1985. — MNHN-IU- 2014 - 11217 (= MNHN-B 27770), 1 ♂ 26.3 × 29.6 mm, 1 ♀ 19.9 × 22.7 mm, Tawala Cave, in pools near entrance, Tawala, Panglao Island, Bohol, Philippines, coll. L. Liao et al., 17. XII. 2000. — MNHN-IU- 2014 - 11218 (= MNHN-B 27771), 2 ♂ 30.9 × 35.6 mm, 40.6 × 46.9 mm, Virata Cave, Libaong, Panglao Island, Bohol, Philippines, coll. local villagers, 18. XII. 2000. PRESERVATION. — In alcohol.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE1FFB7EFECF927B50DFCB5.taxon	discussion	REMARKS The holotype, a male 45.9 × 53.1 mm, Virata Cave, Libaong, Panglao Island, Bohol, Philippines, coll. local villagers, 18. XII. 2000, is deposited at USC, Philippines. Other paratypes are deposited at NSM, USC and ZRC.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB7ECA7FCE2B303FE57.taxon	description	(Fig. 2 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB7ECA7FCE2B303FE57.taxon	materials_examined	LECTOTYPE (by inference of holotype by Türkay 1974 c: 259, as Cardisoma longipes; Ng & Guinot 2001: 317). — MNHN-IU- 2000 - 3763 (= MNHN-B 3763), ♂ 54.0 × 55.0 mm, New Caledonia; presumed type, D. Guinot 1972; Discoplax longipes A. Edw., B 3763, det. Guinot & Ng 2001. — Original label: “ Cardisoma longipes, A. M. Edw. (= Discoplax longipes A. Edw.), Nouvelle-Calédonie ”. Additional labels: “ Cardisoma longipes A. M. Edw. (= Discoplax longipes A. M. Edw.). Nelle Calédonie ”. “ Type présumé, D. Guinot 1972 ”. “ Cardisoma longipes (A. Milne-Edwards 1873), Révision M. Türkay VI. 1972 ”; “ Discoplax longipes A. Edw., B 3763, det. Guinot et Ng 2001 ”. PRESERVATION. — Dry. LP 3, RP 3 dactyli missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB7ECA7FCE2B303FE57.taxon	discussion	REMARKS This specimen, the only one under this name in the MNHN collections, was regarded as the holotype by Türkay (1974 c: 259, as Cardisoma longipes, with measurements 54.0 × 49 mm); on account of the measurements similar to those noted by A. Milne-Edwards (1867), it was considered the holotype by monotypy by Ng & Guinot (2001: 317, with measurements 54.0 by 55.0 mm) and also by Ng & Shih (2015: 383, but with measurements 54.6 × 48.5 mm). In the absence of mention of several specimens in the original description, it is considered the lectotype. Ng & Shih (2015: figs 1 A-D, 11 C, D) reproduced the figures of A. Milne-Edwards (1873 a: pl. 15) and figured the gonopods of this specimen. Alphonse Milne-Edwards (1873 a: 294) regarded Discoplax longipes as “ very rare ” in New Caledonia. The specimen was simply labelled as from “ New Caledonia ”, without any other indication as is often the case in old descriptions and records, but the presence of the species on the main island of New Caledonia is doubtful. It seems that, since A. Milne-Edwards (1867), no specimens of D. longipes have been collected from the main Island where the species is not known locally (Ng & Guinot 2001). The specimen on which A. Milne-Edwards (1867) based his description, i. e., the present lectotype, may have actually been collected from the nearby Loyalty Islands where D. longipes is not uncommon (see Ng & Shih 2014).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB4EFC7FD8CB6B0FDB6.taxon	description	(Fig. 2 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB4EFC7FD8CB6B0FDB6.taxon	materials_examined	HOLOTYPE (by monotypy, see Rathbun 1899). — MNHN-IU- 2000 - 10951 (= MNHN-B 10951), ♂ 46.3 × 69.0 mm, Baja California, coll. M. Diguet, E. - L. Bouvier det. — Original label: “ Gecarcinus planatus Stimpson, Basse-Californie, M. Diguet, (1898), E. - L. Bouvier det. ”. PRESERVATION. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEFFFB4EFC7FD8CB6B0FDB6.taxon	discussion	REMARKS The species is known based on a short description by Bouvier (1895: 8) for a unique specimen brought by Diguet from Baja California in Mexico in 1894 (“ Basse-Californie ” in Bouvier 1895: 6). This specimen MNHN-IU- 2000 - 10951 (= MNHN-B 10951), a male specimen, dry, 67.0 mm cw according to Bouvier (1898: 372), can be considered the holotype by monotypy of Gecarcinus digueti. Rathbun (1899: 73), who wrote “ The type and only specimen hitherto collected is from Lower California and is in the Paris Museum ”, provided several measurements of body parts of this specimen, notably for the carapace 46.3 × 69.0 mm (measurements presently verified). The species was redescribed by Bouvier (1898: 372) with more detail (but not figured) based on another specimen collected in 1897 by Diguet in the same region, a male MNHN-IU- 2016 - 8892 (= MNHN-B 16029), now preserved in alcohol. This second specimen was not taken into account by Rathbun (1899: 73; 1918: 359, under Gecarcinus planatus Stimpson, 1860). Perger et al. (2011: 58, 59), based on information from the MNHN, agreed that the male MNHN-IU- 2000 - 10951 (= MNHN-B 10951) was most likely the holotype of Gecarcinus digueti, being “ the only specimen in the MNHN concordant with the type specimen of G. digueti described by Bouvier (1895) and subsequently reported by Rathbun (1899) […]. Rathbun (1899) recognised G. digueti as a distinct species, recording it from María Cleofa Island [i. e., Cleofas Island] in the Gulf of California, and apparently examined the type in the Paris Museum which she stated was a male measuring 69.0 by 46.3 mm […]. On the basis of the available data, this MNHN specimen is almost certainly the holotype of G. digueti Bouvier, 1895. ” Diguet is known for his trips in Mexico (see Appendix); “ Basse-Californie ” of Bouvier (1895; 1898) and on the MNHN label corresponds to Baja California, Mexico (not Southern California, United States).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB4ECB8FDE3B380FCF5.taxon	description	(Fig. 2 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB4ECB8FDE3B380FCF5.taxon	materials_examined	LECTOTYPE (designated by Türkay 1973: 96). — MNHN-IU- 2000 - 3750 (= MNHN-B 3750), ♂ 63.7 × 82.4 mm (measurements by Türkay 63 × 81 mm),? Australia (inaccurate locality; in fact, Brazil, see Remarks), coll. Quoy and Gaimard, M. Türkay revision VI. 1972. — Original label: “ Gecarcinus lagostoma M. Edwards,? Australie, MM. Quoy et Gaimard ”. Additional label: “ Lectotype! Gecarcinus (Johngarthia) lagostoma H. Milne-Edw., 1837, ♂, Révision M. Türkay VI. 1972 ”. PRESERVATION OF THE LECTOTYPE. — Dry. Carapace with a fissure radiating from the cardiac region to hepatic region. PARALECTOTYPES (designated by Türkay 1973: 96). — MNHN-IU- 2000 - 3748 (= MNHN-B 3748), 1 ♀ 73.9 × 95.5 mm, same data as lectotype. — MNHN-IU- 2000 - 3749 (= MNHN-B 3749), 1 ♀ 73.4 × 97.2 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB4ECB8FDE3B380FCF5.taxon	discussion	REMARKS The binomen Gecarcinus lagostoma introduced in a footnote by H. Milne Edwards in Fréminville (1835: 218) is a nomen nudum since it is not accompanied by a description or indication of the taxon it represents. Later, H. Milne Edwards (1837: 27) recorded that the specimen of Gecarcinus lagostoma was collected by Quoy & Gaimard in “ Australasia ” (and not in Australia), and he (H. Milne Edwards 1837: 25) described in the same paper Gecarcoidea lalandii from Brazil. Ortmann (1894: 739, as G. lalandei) considered inexact the origin of Gecarcoidea lalandii, whereas Ortmann (1897: 338) and then Bouvier (1907: 498) considered inexact the type locality of Gercarcinus lagostoma (see also Tesch 1918; Tweedie 1947; Tavares 1989; Lai et al. 2017). As effectively Johngarthia lagostoma and G. lalandii were currently known from islands of the southern Atlantic Ocean and from the Indo-West Pacific regions, respectively, Türkay (1970: 346) concluded that H. Milne Edwards (1837) switched the collecting records of Gecarcinus lagostoma and Gecarcoidea lalandii, or, perhaps, the labels were exchanged during the same expedition. The diagnosis of G. lagostoma conforms, however, with the specimens of the type series (see also Rathbun 1918: 361, under Gecarcinus lagostoma; 1918: 365, under? Gecarcoidea lalandii; Türkay 1973: 96). Thus, the locality “ Australasia ” of the lectotype and paralectotypes must be recognised as inaccurate, and Brazil must be considered the type locality of Gecarcinus lagostoma, whereas “ Australasia ” is the original type locality of Gecarcoidea lalandii. Lai et al. (2017: 410, 416, figs 5 E, 9 D) have recently selected a fresh male specimen 64.2 × 88.8 mm from Quang Nam, central Vietnam collected in 2009, as the neotype of G. lalandii (ZRC 2010.0321). Johngarthia lagostoma is only known from Ascension Island in the South Atlantic and from three islands off Brazil (Trinidad, Fernando de Noronha, Atol das Rocas), western Atlantic (Melo 1996; Hartnoll et al. 2006; Hartnoll et al. 2010: fig. 1). The differences between the measurements by Türkay (63.0 × 81.0 mm) and ours (63.7 × 82.4 mm) are due to the restoration of the lectotype in the 1990 s.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB5EF19FC22B688F869.taxon	description	(Fig. 2 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB5EF19FC22B688F869.taxon	materials_examined	NEOTYPE (by present designation). — MNHN-IU- 2000 - 3758 (= MNHN-B 3758), ♂ 36.7 × 47.2 mm, Guadeloupe, coll. Beaupertuis. — Original label: “ Gecarcinus lateralis, Fréminv., Guadeloupe, Mr Beaupertuis ”. — Additional labels: “ Lectotype. J. F. [J. Forest] 1.3.69 ”; “ Gecarcinus lateralis Fréminville, Guadeloupe, M. Beaupertuis, Lectotype ”; “ Gecarcinus (Gecarcinus) lateralis lateralis (Fréminville, 1835), Révision M. Türkay, VI. 1972 ”. PRESERVATION. — Dry. Specimen complete, mounted on cardboard.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFECFFB5EF19FC22B688F869.taxon	discussion	REMARKS Two assertions previously published in the literature, related to the authorship and the type material of Gecarcinus lateralis need to be corrected. The authorship of G. lateralis is traditionally attributed to Fréminville (1835) (e. g. H. Milne Edwards 1837; Chace & Hobbs 1969; Türkay 1970, 1974 a; Bliss et al. 1978; Hartnoll 1988; Ng & Guinot 2001; Ng et al. 2008; Perger & Wall 2014). Low et al. (2013) stated that Guérin (1832 in Guérin 1829 - 1837) must take precedence. Fréminville (1835) described Ocypoda lateralis from La Désirade, Guadeloupe, Marie-Galante, Martinique, Les Saintes in the West Indies, based on specimens that he has probably himself collected (see Appendix). In the introduction of this paper (1835: 213, footnote), the editor of the volume, actually H. Milne Edwards, indicates about Fréminville’s “ tourlourou ”: “ son Ocypoda lateralis est une espèce de Gécarcin bien distincte et dont j’ai donné une figure dans mon Histoire naturelle des Crustacés. M. Guérin l’a également figuré dans son Iconographie du règne animal ” [“ his Ocypoda lateralis is a clearly distinct species of ‘ Gécarcin’ and to which I have given a figure in my Histoire naturelle des Crustacés. M. Guérin has also figured it in his Iconographie du règne animal ”]. Guérin (1832, in Guérin 1829 - 1837: 7) had previously used the name “ Gecarcinus lateralis Fréminville ” and illustrated what is clearly the same species. It is very likely that Guérin used the same material as Fréminville and that he was aware of the future species by Fréminville. According to the official dates, Guérin’s plate has been published before the description by Fréminville in 1835. According to Low et al. (2013) the authorship of G. lateralis must be credited to Guérin (1832). But, as Fréminville is solely responsible for the species name in a way satisfying the criteria of availability (Code, Art. 50.1.1), the authorship of G. lateralis should be credited to Fréminville in Guérin (1832). Türkay (1970), based on erroneous indications given by J. Forest, the curator of Crustacea Collection in the MNHN at the time, regarded as the type series of G. lateralis material collected by two French naturalists, Beaupertuis and Bélanger, and deposited in this institution. Türkay (1970: 337) selected as lectotype of Gecarcinus (Gecarcinus) lateralis a specimen from Guadeloupe collected by Beaupertuis. Türkay (1974 a: 974, 975) then designated as paralectotypes one male and one female from Guadeloupe MNHN-IU- 2000 - 3757 (= MNHN-B 3757), and two males and one female collected by Bélanger from Martinique (MNHN-IU- 2000 - 3755 [= MNHN-B 3755], MNHN-IU- 2000 - 3756 [= MNHN-B 3756]). A colour figure of the lectotype was published by Perger & Wall (2014: fig. 6 A-C). However, Beaupertuis and Bélanger’s specimens are likely to not have been collected during this period (i. e., before 1832), these two naturalists having traveled in the Antilles later on as shown by the entry dates of their material in the CAA (Beaupertuis: 1839 from Guadeloupe; Bélanger: 1859, 1860, 1864 from Martinique) (see Appendix), and cannot constitute the original material seen by Guérin (1832). According to the Code (Art. 74.2), “ the name-bearing type of any nominal taxon, once fixed in conformity with the provisions of the Code, is not subject to change ” except if the lectotype is “ found not to have been a syntype ”. Thus, “ if it is demonstrated that a specimen designated as a lectotype was not a syntype, it loses its status of lectotype ”. A neotype of G. lateralis must be fixed. Having been previously referred to, although unsupported, as the lectotype for the species by Türkay (1970), then by Perger & Wall (2014), and considering its locality, the specimen from Guadeloupe registered MNHN-IU- 2000 - 3758 (= MNHN-B 3758) is eligible for neotype designation: it is hereby selected as the neotype of Gecarcinus lateralis Guérin, 1832.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB5EFE9FF4DB35FFAF4.taxon	description	(Fig. 2 G)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB5EFE9FF4DB35FFAF4.taxon	materials_examined	PARATYPE. — MNHN-IU- 2014 - 11211 (= MNHN-B 12314), 1 ♀ 36.6 × 44.3 mm Ecuador, St. Elena, Festa coll. — Original label: “ Gecarcinus Festae Nob. (cotype), St Helena, Festa, Ecuador, Nobili 1901, Muséum Paris ”. PRESERVATION. — In alcohol.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB5EFE9FF4DB35FFAF4.taxon	discussion	REMARKS Actually Nobili never published a description of Gecarcinus festae. In 1901 he described Sesarma festae and Uca festae and in the same paper referred to two specimens of Gecarcinus collected in Ecuador as Gecarcinus ruricola, a species restricted to western Atlantic islands (Türkay 1970; Perger & Wall 2014). The abovementioned specimen, with the name of Nobili (1901) on the label, was likely collected by Enrico Festa during his 1895 - 1898 trip to Ecuador (Viaggio del Dr. Enrico Festa nella Republica dell’Ecuador et regioni vicine), then studied by Nobili who probably first considered it as new and dedicated to Festa as “ Gecarcinus festae ” without finally establishing it. The unpublished binomen Gecarcinus festae is a nomen nudum. The MNHN specimen examined by Nobili and selected as paratype of the new species Gecarcinus nobilii dedicated to Nobili by Perger & Wall (2014) is figured in their fig. 3 D-F. The holotype is preserved in LACM. Furthermore, Pacific Gecarcinus (Gecarcinus) lateralis quadratus sensu Türkay (1970: 338; see Prahl & Manjarrés 1984: 155) and Gecarcinus lateralis sensu Türkay 1987 (: 147, fig. 7) have been referred to as Gecarcinus nobilii Perger & Wall, 2014. Note that Gecarcinus quadratus Saussure, 1853 was recently recognised as a valid species (Toledano Carrasco & Villalobos Hiriart 2018).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB2EF06FA20B608FED7.taxon	description	(Fig. 2 H)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB2EF06FA20B608FED7.taxon	materials_examined	HOLOTYPE (by monotypy, see Türkay 1974 a: 973). — MNHN-IU- 2000 - 3751 (= MNHN-B 3751), ♂ 54.0 × 75.0 mm, Loyalty Islands, coll. M. Caillot, Type. — Original labels: “ Pelocarcinus cailloti ”; “ Pelocarcinus cailloti A. M. Edw. (Type), I. Loyauté, Caillot, 2523 - 86 ”; with “ 2523 86 ” on a small label stuck onto the carapace of the specimen. Additional label: “ Type présumé (Holotype) de Pelocarcinus cailloti A. Milne Edwards 1890; Gecarcoidea lalandii H. Milne Edwards 1837, Révision M. Türkay VI. 1972 ”. PRESERVATION. — Dry. Carapace cracked; LP 1, LP 5 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEDFFB2EF06FA20B608FED7.taxon	discussion	REMARKS Alphonse Milne-Edwards (1890: 175) clearly indicated that the species in his hands was represented by a unique specimen, so it is a holotype by monotypy, which corroborates Türkay’s (1974 a: 972) inference of the holotype. The mention 2523 - 86 on the label corresponds to the information recorded in the CAA.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECB7FE0CB693FAEB.taxon	description	(Fig. 3 A)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECB7FE0CB693FAEB.taxon	materials_examined	LECTOTYPE (designation by Türkay 1974 a: 973). — MNHN-IU- 2000 - 3752 (= MNHN-B 3752), ♂ 51.5 × 70.9 mm (measurements by Türkay: 54.0 × 70.0 mm), Dos Hermanas Islands, Philippines, coll. Alfred Marche. — Original labels: “ Pelocarcinus Marchei ”, “ Pelocarcinus Marchei A. M. Edw., no 942, 5 crabes, île des Deux Soeurs, Iles Philippines, Alf. Marche ”. Additional labels: “ Pelocarcinus Marchei A. M. Edw. (d’après A. M. Edwards), Gecarcoidea Lalandei Edw. (d’après Ortman), Iles Philippines, Ile des Deux Soeurs, Marche, 2725 - 1882 ”; “ Gecarcoidea lalandii H. M. Edwards 1837, Révision M. Türkay; VI- 1972. Lectotype de Pelocarcinus marchei A. Milne-Edw. 1890 ”. PRESERVATION OF THE LECTOTYPE. — Dry. RP 1, LP 1, LP 4 missing. PARALECTOTYPES (designation by Türkay 1974 a: 974). — MNHN-IU- 2000 - 3753 (= MNHN-B 3753), 1 ♂ 57.0 × 79.1 mm, 3 ♀ 47.6 × 63.3 mm, 48.1 × 61.4 mm, ca 57.5 × c. 73.7 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry. 1 ♂ and 1 ♀ in good condition but pereopods detached; 2 ♀ in bad condition.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECB7FE0CB693FAEB.taxon	discussion	REMARKS Türkay (1974 a: 973) gave the carapace measurements as 54.0 × 70.0 mm, but presently the specimen measures 51.5 × 70.9 mm. The mention 2725 - 1882 on the label corresponds to the information recorded in the CAA.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECA8FA20B324F86A.taxon	description	(Fig. 3 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECA8FA20B324F86A.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3745 (= MNHN-B 3745), ♂ 50.0 × 63.0 mm, Oceania, M. Guérin. — Original label: “ Cardisoma rotundum (Quoy & Gaimard 1824), Océanie, M. Guérin ”. Additional labels: “ Cardisoma rotundum (Quoy & Gaimard 1824), Type présumé de C. frontalis H. Milne-Edw. (1853) (Holotype), Rev. M. Türkay VI. 1972 ”. PRESERVATION. — Dry. Specimen stuck onto corkboard; carapace cracked on posterior region; two pereopods missing; pleon detached; G 1 rehydrated, in alcohol.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEAFFB2ECA8FA20B324F86A.taxon	discussion	REMARKS The species, described by Quoy & Gaimard (1824: 527, pl. 77, fig. 1) and originating from the banks of rivers in Guam, Mariana Islands, was collected during the Expedition of L. de Freycinet on the Uranie (1817 - 1820) (see Appendix). Quoy & Gaimard (1824: 527) did not explicitly mention if the taxon was based on a single individual, but their description provides information on the behaviour of several crabs: “ ces animaux extrêmement défiants ” [“ these extremely challenging animals ”], an indication that numerous specimens were probably collected in Guam. “ Océanie ” is a large area that can include Guam, thus the MNHN specimen such labelled may constitute a part of the type material. Türkay (1974 a: 971, 972; pers comm., Jan. 2013) did not considered any specimen as type of Thelphusa rotunda, assuming the species was a synonym of Cardisoma frontalis H. Milne Edwards, 1853, and regarded the specimen of Quoy & Gaimard (1824) from Guam as the holotype of C. frontalis, instead of T. rotunda; Türkay (1974 b: 236) confirmed this choice (see Opinion 1205). In the description of C. frontalis, H. Milne Edwards (1853: 170 [204]) stated that the locality of C. frontalis was not known. Although Davie (2002: 185) mentions under “ Type data ” of Cardisoma frontalis that “ syntypes ” are most probably deposited at the MNHN, the type material could not be traced in the MNHN collections and may be considered no longer extant. Since the description and illustration of Thelphusa rotunda Quoy & Gaimard, 1824 (1824: pl. 77, fig. 1) from Guam in the Mariana Islands (being considered good enough to identify the species) correctly corresponded with Cardisoma frontalis and the identity of the two species was not doubted, Türkay (1974 a) judged the designation of a neotype for C. frontalis superfluous. He selected the specimen MNHN-IU- 2000 - 3745 (= MNHN-B 3745) labelled “ Cardisoma rotundum (Quoy & Gaimard 1824) ” as the holotype of Cardisoma frontalis. By that action Türkay (1974 b: 234 - 236, fig. 14) considered Guam the type locality of C. rotundum. But such an act by Türkay (1974 a) is unwarranted (see Code, Art. 73.1.3), and the label in question actually concerns the type of Thelphusa rotunda Quoy & Gaimard, 1824. The statements that the type of Thelphusa rotunda was ‘ in all likelihood’ lost and that Cardisoma frontalis was the junior synonym of C. rotunda are unjustified, with unsatisfactory arguments. Ng & Guinot (2001: 333), regarding Discoplax A. Milne-Edwards, 1867 as a distinct genus (see Guinot 1988, 1994), treated Cardisoma frontalis as the junior synonym of Discoplax rotunda. Even though the two species are synonymous, the specimen of Quoy & Gaimard (1824) from Guam is not the type of C. frontalis but, instead, that of Tuerkayana rotundum (Quoy & Gaimard, 1824). The MNHN specimen labelled “ Océanie ” and regarded as the holotype of C. frontalis by Türkay (1974 a, b) is eligible to be designated as the lectotype of Tuerkayana rotundum (Quoy & Gaimard, 1824), see Guinot et al. (2018).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEBFFB3EC81FE8DB241F9AA.taxon	materials_examined	TYPE MATERIAL. — None extant.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEBFFB3EC81FE8DB241F9AA.taxon	discussion	REMARKS The binomen “ Grapse albo-lineatus ” (“ grapse rayé de blanc ” or white-striped grapse) has been introduced by Latreille, in Milbert (1812: 275) but Grapsus albolineatus is usually credited to Lamarck (1818: 249, as Grapsus albo-lineatus, “ grapse raies-blanches ”). As is often the case, Latreille in Milbert (1812) and Lamarck (1818) were based on the same material, namely a crab collected by Mathieu from Mauritius (= Île de France; see also Latreille 1825: 148), and referred to the same species. Latreille in Milbert (1812: 275) wrote “ grapse ” (“ Ce sont les grapses de M. Lamarck ” [These are the grapses of M. Lamarck]). Although “ G. albo-lineatus ” is a binominal combination with latinised species names, the absence of any diagnosis makes it not available (see the discussion below). The genus name Grapsus belongs to Lamarck (1801: 150), with Cancer grapsus Linnaeus, 1758 as type species by tautonomy (see also designation by Latreille 1810: 422). Grapsid species cited in Latreille in Milbert (1812: 275), i. e., Grapsus albolineatus, G. erytrhocheles, G. tuberculatus and G. tessellatus, correspond to two different nomenclatural situations. Grapsus tessellatus “ grapse damier ” (“ checkered grapse ”) is described in a few lines: this constitutes a diagnosis making the name available: consequently, it can be credited to Latreille in Milbert (1812) as Lybia tessellata (see Guinot 1976). The case is different for the three other “ grapses ”: G. albo-lineatus “ grapse rayé de blanc ” (white-striped grapse); G. erytrhocheles “ grapse à pinces rouges ” (grapse with red claws), G. tuberculatus “ grapse tuberculé ” (tuberculated grapse). In most French and German scientific works at this time, the Latin name was accompanied by its translation in French (e. g. Latreille 1806; Lamarck 1818; H. Milne Edwards 1834, 1837), or in German, (e. g. Herbst 1782 - 1804). These specific spellings are only the latinisation of the names depicting morphological (shape, colour, ornamentation), behavioural, geographical and other features, by simple juxtaposition of both names. This simple translation of the names is not enough to constitute a diagnosis and to make the taxa available, and does not correspond to a brief description. These names cannot satisfy the requirement of Article 12.1 of the Code stipulating that (for the names published before 1931) to be available a taxon must be accompanied by a description (even if only the colour) or a definition of the taxon that it denotes, or by an indication. According to Ng et al. (2008: 22) these three species were “ defined by their colour or shape ”, but this is not the case and these names are not available. Moreover, Ng et al. (2008: 22) have treated the three above-mentioned grapses in the same manner as the case of Grapsus tessellatus, and credited Grapsus albolineatus, Grapsus erytrhocheles and Grapsus tuberculatus, like G. tessellatus, to Latreille in Milbert (1812). It is an erroneous generalisation: G. tessellatus is an available name with the authorship of Latreille in Milbert (1812), whereas the other three names are clearly nomina nuda. Grapsus albolineatus belongs to Lamarck, 1818, G. erytrhocheles is not available (or at least corresponds to a species inquirenda); G. tuberculatus must be credited to Lamarck (1818: 247, as Plagusia tuberculata) (Schubart & Ng 2000: 327, 334, fig. 3 A). To avoid ambiguity, a point that requires discussion is the meaning of “ vernacular ”. According to the English glossary of the Code (ICZN 1999: 110) a vernacular name is “ A name of an animal or animals in a language used for general purposes as opposed to a name proposed only for zoological nomenclature ”, whereas a zoological name is “ The scientific name of an animal taxon in binominal nomenclature ”. Thus in French scientific works at the period of Henri Milne Edwards (1834, 1837), the new taxa are proposed in a scientific context and are not vernacular names, conversely to the views of most carcinologists (including Ng et al. 2008: 20). The type material from Mauritius collected by Mathieu was not found in the MNHN collection. A neotype should be fixed for Grapsus albolineatus, preferably from the type locality, Mauritius or its proximity (Code, Art. 75.3.6), as it is considered the senior synonym of numerous species (see Banerjee 1960: 147, 154; Holthuis 1977: 145, 147), such as G. strigosus (Herbst, 1799), from the Indian Ocean; G. (Goniopsis) flavipes MacLeay, 1838, from Cape of Good Hope; G. peroni H. Milne Edwards, 1853, from Australia (see below); and G. longipes Stimpson, 1858, from Kikaishima, southern Japan, and Hong Kong (Davie 2002). The type locality “ les mers de l’Ile-de-France ” and the occurrence of this species on the east coast of Africa was questioned by Crosnier (1965: 17) but according to Holthuis (1977: 147) it is one on the most common and conspicuous crabs of the Red Sea. A neotype designation is unfortunately not possible at this time because the MNHN collection does not possess any material from Mauritius or a nearby locality (Code, Art. 75.3.6). Grapsus granulosus H. Milne Edwards, 1853, from the Red Sea, identified as G. albolineatus by some authors (e. g. Banerjee 1960; Davie 2002), is regarded as distinct by others (e. g. Crosnier 1965; Holthuis 1977; Vannini & Valmori 1981; Zaouali et al. 2007; Ng et al. 2008); see below.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEBFFB1EFF6F9E0B5E9FE17.taxon	description	(Fig. 3 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEBFFB1EFF6F9E0B5E9FE17.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 10978 (= MNHN-B 10978), ♂ 28.5 × 36.0 mm, patria ignota. — Original label: “ Graspsus brevipes M. Edw. ”. PRESERVATION. — Dry. Specimen stuck to cardboard; a crack on the left branchial region of the carapace.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFEBFFB1EFF6F9E0B5E9FE17.taxon	discussion	REMARKS The description did not mention how many specimens were examined to diagnose the species. Only one specimen was found in the MNHN collection but, as there is no assumption it is a holotype (i. e., fixed by monotypy), it is designated here as lectotype (Code, Recommendation 73 F).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFB1ED75FE4CB5EDF869.taxon	description	(Fig. 3 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFB1ED75FE4CB5EDF869.taxon	materials_examined	NEOTYPE (by present designation). — MNHN-IU- 2000 - 3407 (= MNHN-B 3407), ♂ 40.9 × 49.8 mm, Martinique, coll. M. Plée. — Original label: “ Grapsus cruentatus Latreille, Martinique, M. Plée ”; “ Plée, Martinique, 1826, n ° 28 ” is written with Chinese ink directly on the left cheliped. PRESERVATION. — Dry. LP 4, RP 5 dactylus, right eye missing, LP 2 detached.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFB1ED75FE4CB5EDF869.taxon	discussion	REMARKS Latreille (1803: 70) described this species from the islands of South America (“ îles de l’Amérique méridionale ”), imprecise information (as it was common in the past) indicating the geographical origin. Henri Milne Edwards (1837: 85) has added two localities, Brazil and Antilles, based on specimens present at that time in the collection of the MNHN (as “ C. M. ”). Today, the MNHN collection contains several samples of dry specimens labelled Grapsus or Goniopsis cruentatus, from Brazil, Mexico, French Guiana and Martinique. They were all collected too late to belong to the type series; moreover, some of the localities cannot be considered “ islands of South America ”. For example: (1) the sample with one specimen MNHN-IU- 2000 - 3406 (= MNHN-B 3406) collected from Brazil by Delalande in 1816 (see Appendix), a date that is too late to have enabled Latreille (1803) to examine the material; (2) the sample with one specimen MNHN-IU- 2000 - 3403 (= MNHN-B 3403) labelled “ Brésil, (Destero), M. Müller ” (with a note by “ Monod 1953 ”, indicating this specimen as being a new species of Metopograpsus). Concerning the sample from Martinique in the MNHN collection, which could correspond to the type locality, consisting of two males collected by M. Plée, it cannot belong to the type series: the material of Plée was collected after 1820 (Bauchot et al. 1990: 117; see Appendix), and this is confirmed by the entry date “ 1826 ” of Plée’s material in the CAA. Since no MNHN material conforms to a presumed syntype and no name-bearing type is believed to be extant, a neotype should be designated. The more complete male of the MNHN-B 3406 sample has been selected as neotype, MNHN-IU- 2000 - 3407 (= MNHN-B 3407). The slightly smaller male specimen has been separated and is registered as MNHN-IU- 2000 - 1109 (= MNHN-B 3407).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFAEEFD6FF4DB64AFDD6.taxon	description	(Fig. 3 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFAEEFD6FF4DB64AFDD6.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2008 - 10635 (= MNHN-B 11803), ♂ 28.0 × 31.7 mm, Nosy Bé, Madagascar, coll. and det. A. Crosnier. PRESERVATION OF THE LECTOTYPE. — In alcohol. RP 1 missing. PARALECTOTYPES. — MNHN-IU- 2014 - 11212 (= MNHN-B 11531), 3 ♂ 22.6 × 25.2 – 24.2 × 27.8 mm, 1 ♀ 21.2 × 24.4 mm, 2 ovig. ♀ 24.1 × 28.6 – 29.5 × 33.4 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — In alcohol.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFE9FFAEEFD6FF4DB64AFDD6.taxon	discussion	REMARKS Crosnier (1965: 14, table p. 16) has based the species on several specimens from Nosy Bé, Madagascar, but he did not expressly mention any type specimens in his publication. He mentions in the text a large specimen (length: 34.1; width: 37.7 mm) and provides measurements of five males and two females in a table (Crosnier 1965: 16). Two lots of specimens deposited at the Muséum, MNHN-IU- 2008 - 10635 (= MNHN-B 11803) and MNHN-IU- 2014 - 11212 (= MNHN-B 11531) are labelled “ Syntypes ”. Our measurements of most of the specimens do not correspond to the dimensions provided by Crosnier (1965: 16), except for one male MNHN-IU- 2008 - 10635 (= MNHN-B 11803), 28.0 × 31.7 mm. This discrepancy can be easily accounted for by how the specimens were measured. The largest specimen (length: 34.1; width: 37.7 mm) could not be located. The next largest male specimen MNHN-IU- 2008 - 10635 (= MNHN-B 11803) is here designated as lectotype of G. fourmanoiri. The other specimens from the type series become paralectotypes. It is very likely that Crosnier (1965) has collected and examined more than eight specimens belonging to his type series. Actually, one specimen was donated to Leiden Museum by Crosnier in 1966 (RMNH D 22683, male 23.7 × 27.0 mm, Nosy Bé, Madagascar, coll. and det. A. Crosnier): it is indicated as syntype in the Catalogue of the Types of the Nationaal Natuurhistorisch Museum, Leiden, by Fransen et al. (1997: 123) and shows the same ‘ type and form’ of label as the labels of the MNHN jars. This RMNH sample also becomes paralectotype. Fransen et al. (1997: 123, footnote) regarded two additional samples as syntypes of G. fourmanoiri, but this mention corresponds to an erroneous statement. Crosnier (1965: 17, footnote) wrote: “ At Leiden, we have reexamined specimens from Banerjee (1960: 149) from Durban and Reunion Rocks, initially identified as Grapsus albolineatus ”. According to Fransen (pers. comm., December 2012), two samples (RMNH D 3210, one female 32.5 × 35.0 mm, South Africa, Reunion Rocks, 28. X. 1938, leg. L. D. Brongersma; RMNH D 3250, 2 males 10.8 - 16.9 × 13.0 - 19.6 mm, South Africa, Durban, 10 - 11. XI. 1938, leg. H. Engel) have been deemed by Crosnier as suitable syntypes of his G. fourmanoiri. However, despite the Code ’ s Recommendation (Art. 74 E) about the verification of locality (“ When selecting a lectotype, the author should, if possible, verify the accuracy of the locality ascribed to it ”), it is obvious that these two samples, RMNH D 3250 from Durban and RMNH D 3210 from Reunion Rocks, have not been collected from Nosy Bé, Madagascar by A. Crosnier. It is likely that they were only subsequently identified by him as G. fourmanoiri and, therefore, they cannot be considered syntypes, even if they correspond to G. fourmanoiri.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEECA7FD03B501F869.taxon	description	(Fig. 3 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEECA7FD03B501F869.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3411 (= MNHN-B 3411), ♂ 24.6 × 27.2 mm, East Indies. — Original label: “ Grapsus grapsus Linné = Grapsus gracilipes Edw., Indes Orientales ”. PRESERVATION OF THE LECTOTYPE. — Dry. LP 3, RP 3, RP 5 dactyli, RP 4, LP 2 meri and dactyli missing. PARALECTOTYPES. — MNHN-IU- 2000 - 3408 (= MNHN-B 3408), 1 ♀ 27.1 × 30.1 mm, same data as lectotype. — MNHN-IU- 2000 - 10767 (= MNHN-B 10767), 1 ♂ 23.3 × 27.1 mm, same data as lectotype. — MNHN-IU- 2000 - 10769 (= MNHN-B 10769), 1 ♀ 29.0 × 32.2 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEECA7FD03B501F869.taxon	discussion	REMARKS In the published description by H. Milne Edwards (1853) a locality is mentioned, i. e., “ Mers de Chine, Taoranne ”. A quick check with Dr Tran Anh Duc and M. Nguyen Thanh Son (Vietnam National University-University of Science) regarding “ Taoranne ” has revealed that the locality should be Tourane, the French name of Da Nang, a port city in Vietnam. The term “ Indes Orientalis ”, or East Indies, was used in colonial times to designate the territories of South and Southeast Asia, including countries like Iran, Pakistan, India, Nepal, Bhutan, Bangladesh, Myanmar, Sri Lanka, Maldives, Thailand, Malaysia, Vietnam, Cambodia, Laos, the Philippines, Brunei, Singapore, East Timor, and Indonesia. It is thus possible to consider that “ Indes Orientales ” may correspond to “ Mers de Chine ”, and “ Taoranne ” the type locality. The type status was suspected unknown according to Davie (2002: 214). Four samples of dry Grapsus gracilipes are actually present in the MNHN collection and hereby designated as lectotype and paralectotypes, respectively.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEF04FF4DB399FB54.taxon	description	(Fig. 3 G)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEF04FF4DB399FB54.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3417 (= MNHN-B 3417), ♂ 64.2 × 71.9 mm, Red Sea, coll. M. Roux. — Original label: “ Grapsus granulosus, Mer Rouge, M. Roux ”. PRESERVATION OF THE LECTOTYPE. — Dry. Carapace with a vertical crack on the right, a small chip off on the right posterior margin. PARALECTOTYPES. — MNHN-IU- 2000 - 3455 (= MNHN-B 3455), 1 ♂ 61.8 × 67.0 mm, same data as lectotype. — MNHN-IU- 2000 - 10937 (= MNHN-B 10937), 1 ♂ 60.4 × 68.4 mm, 1 ♀ 54.1 × 59.2 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEF04FF4DB399FB54.taxon	discussion	REMARKS The type locality is the Red Sea, and the collector “ M. Roux ” that is indicated on the label is likely J. L. F. P. Roux who collected material in the Mediterranean Sea and in the Near East (Bauchot et al. 1990, 1997; see Appendix). Material brought from the Red Sea by Roux has 1835 as entry date in the CAA. Thus, these samples unambiguously belong to the type series, and a lectotype can be designated. According to Ng et al. (2008: 216, 218), Grapsus granulosus was often regarded as a junior synonym of Grapsus albolineatus (e. g. Banerjee 1960: 147, 153 - 154; Davie 2002: 213), but Crosnier (1965: 15, pl. 2, fig. 1) and Holthuis (1977: 147, 148) commented that it is a distinct species (see also Vannini & Valmori 1981; Zaouali et al. 2007).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEFE6FA81B2D5F869.taxon	description	(Fig. 3 H)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEFE6FA81B2D5F869.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3421 (= MNHN-B 3421), ♀ 34.1 × 42.1 mm, Australia. — Original label: “ Grapsus grayi Edw., Australie, Museum Britannique! ”. PRESERVATION. — Dry. Carapace cracked on the gastric, cardiac regions; RP 2 dactylus, RP 3, RP 4 meri and dactyli, LP 2 - LP 4 carpi, meri and dactyli missing; pleonal somites 5, 6 damaged; telson missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF6FFAEEFE6FA81B2D5F869.taxon	discussion	REMARKS Davie (2002: 213) indicated that the type status was uncertain. There is no doubt, however, that the above-mentioned female specimen was identified by H. Milne Edwards and corresponds to the type material. The mention on the label “ Museum Britannique! ” punctuated by an exclamation mark corresponds to the indication in the CAA of an exchange with the British Museum in 1849. According to Recommendation 73 F of the Code, this specimen is hereby designated as lectotype.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEC99FF4DB606FB54.taxon	description	(Fig. 4 A)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEC99FF4DB606FB54.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3426 (= MNHN-B 3426), ♂ 16.4 × 19.1 mm, Antilles, Martinique, coll. M. Plée. — Original label: “ Grapsus lividus Edw. Martinique, M. Plée ”. PRESERVATION OF THE LECTOTYPE. — Dry. RP 2 missing. PARALECTOTYPES. — MNHN-IU- 2000 - 1100 (= MNHN-B 3426), 1 ♀ 13.7 × 15.8 mm, same data as lectotype. — MNHN-IU- 2000 - 3427 (= MNHN-B 3427), 1 ovig. ♀ 14.7 × 15.8 mm, 1 ♀ 13.0 × 16.9 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEC99FF4DB606FB54.taxon	discussion	REMARKS Henri Milne Edwards (1837) indicated that the species was from the Antilles. His description mentioning at least two colour morphs (yellow with red marks; and almost entirely red) reveals that he has examined several specimens. The material collected by Plée in Martinique, his country of birth, most likely belongs to the type series according to the entry date “ 1826 ” of Plée’s material mentioned in the CAA (see also Bauchot et al. 1990: 117; Appendix). The specimen of “ 14 lignes ” (1 ligne = 0.2255 cm), thus measuring 31.5 mm carapace length, cited by H. Milne Edwards (1837: 85) is no longer present in the MNHN collection, and all the MNHN specimens are smaller. The only male in the Plée material was selected as lectotype.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFECAAFA81B36EFEB7.taxon	description	(Fig. 4 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFECAAFA81B36EFEB7.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3414 (= MNHN-B 3414), ♂ 36.5 × 40.0 mm, Haiti, coll. M. Ricord. — Original label: “ Grapsus grapsus (= Grapsus pictus, Grapsus maculatus), M. Ricord, Haiti ”. PRESERVATION OF THE LECTOTYPE. — Dry. LP 23, RP 12, RP 5 missing. PARALECTOTYPE. — MNHN-IU- 2000 - 1102 (= MNHN-B 3414), 1 ♂ 29.2 × 32.7 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPE. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFECAAFA81B36EFEB7.taxon	discussion	REMARKS The origin of the species is the Antilles. The original sample MNHN-B 3414 with two dry specimens (2 males) from Haiti is presumed to belong to the type series, the collector Ricord being known to have collected material in this region and sent to the MNHN. In the CAA, Ricord (see Appendix) is listed as a collector of material from Cuba with two entries in 1833.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEFAEFEECB25FFA0B.taxon	description	(Fig. 4 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEFAEFEECB25FFA0B.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3523 (= MNHN-B 3523), ♂ 17.0 × 19.5 mm, Oran, northwestern coast of Algeria, M. Deshayes, M. Lucas. — Original label: “ Leptograpsus maurus Edw., Oran, M. Lucas ”. PRESERVATION. — Dry. Specimen attached onto cardboard by a stub; RP 5 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFAFEFAEFEECB25FFA0B.taxon	discussion	REMARKS The species described by Lucas (1846: 20) was based on two male specimens, one measuring 17.0 × 19.5 mm, collected by Deshayes at Oran, located on the northwestern coast of Algeria. Lucas (1846) noted: “ This species, of which I have only two male individuals, was found during the summer, in Oran harbour, by M. Deshayes ” (“ Cette espèce, dont je ne possède que deux individus mâles, a été rencontrée pendant l’été, dans la rade d’Oran, par M. Deshayes ”). Rathbun (1918: 244) indicated that the ‘ type’ was in the “ Paris Museum ”, but Poupin et al. (2005: 28) noted that “ the first author was unable to locate it ” in the MNHN. A sample MNHN-IU- 2000 - 3523 (= MNHN-B 3523), with only one male specimen labelled “ Oran, M. Lucas ”, thus from the type locality and obviously collected and identified by Lucas, fully corresponds to the male specimen described and measured by Lucas (1846). It is hereby designated as lectotype. The second specimen of the type series is missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFADEF15FA40B6DFFDB6.taxon	description	(Fig. 4 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFADEF15FA40B6DFFDB6.taxon	materials_examined	LECTOTYPE (designated by Castro 2011: 115). — MNHN-IU- 2000 - 10992 (= MNHN-B 10992), ♂ 32.5 × 38.1 mm, Poulo Han (= Hon Island, Papua New Guinea), coll. Hombron & Jacquinot. — Original label: “ Metopograpsus oceanicus Hombr. & Jacq., MM. Hombron & Jacquinot, Poulo Han ”. PRESERVATION OF THE LECTOTYPE. — Dry. PARALECTOTYPE. — MNHN-IU- 2000 - 1104 (= MNHN-B 10992), 1 ♂ 23.0 × 26.9 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPE. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF7FFADEF15FA40B6DFFDB6.taxon	discussion	REMARKS For authorities as cited on the plate captions or in the text of the Crustacea collected during the Voyage au Pôle Sud (1842 - 1854) [Voyage to the South Pole], see Clark & Crosnier 2000: table 4; Holthuis 2002: 422.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADEC9CFDE3B63EFA2B.taxon	description	(Fig. 4 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADEC9CFDE3B63EFA2B.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3415 (= MNHN-B 3415), ♂ 64.1 × 69.0 mm, Chile, coll. M. Fontaine. — Original label: “ Grapsus grapsus Linné (= Grapsus ornatus M. Edw.), M. Fontaine, Chili ”. PRESERVATION OF THE LECTOTYPE. — Dry. PARALECTOTYPES. — MNHN-IU- 2000 - 3429 (= MNHN-B 3429), 1 ♂ 65.5 × 73.5 mm, Chile, coll. M. Fontaine. — MNHN-IU- 2000 - 3431 (= MNHN-B 3431), 1 ♂ 69.9 × 78.0 mm, Chile. PRESERVATION OF THE PARALECTOTYPES. — Dry. All specimens are incomplete.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADEC9CFDE3B63EFA2B.taxon	discussion	REMARKS The two data, the type locality and the name of Fontaine, who is known as a collector of crustaceans in South America (see Guinot & Cleva 2002; Appendix), ensure recognition of type series material and justify such a lectotype designation. The entry of material from Chile collected by Fontaine in the CAA is dated 1834.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADECA5FA60B27DFBD4.taxon	description	(Fig. 4 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADECA5FA60B27DFBD4.taxon	materials_examined	HOLOTYPE (by monotypy, present paper). — MNHN-IU- 2000 - 3450 (= MNHN-B 3450), ♀ 45.8 × 49.9 mm, Stephen Island, Torres Strait. — Original label: “ Grapsus strigosus Herbst (= Grapsus pelagicus M. Edwards), Ile Stephen, Détroit de Torrès ”. PRESERVATION. — Dry. Only RP 2, RP 5 attached, LP 3 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADECA5FA60B27DFBD4.taxon	discussion	REMARKS Henri Milne Edwards (1853) described Grapsus pelagicus as a new species from the Torres Strait. This species has two senior secondary homonyms: Grapsus pelagicus Roux, 1830 from the Mediterranean Sea (see under this name) and Grapsus pelagicus Say, 1818 (Say 1818: 442) from the Gulf Stream, both synonymised under Planes minutus (Linnaeus, 1758) (see Chace 1951: 81; Manning & Holthuis 1981: 236). Although the species-group name Grapsus pelagicus H. Milne Edwards, 1853 is a junior secondary homonym, it was no longer considered congeneric with its two homonyms and has never been replaced, the junior name not having to be rejected since these species are placed in a different genera (Code, Art. 59.2). The only specimen of H. Milne Edwards’ (1853) species MNHN-IU- 2000 - 3450 (= MNHN-B 3450) deposited at the MNHN labelled “ Grapsus strigosus (= Grapsus pelagicus), Ile Stephen, Détroit de Torrès ” is certainly the type specimen of Grapsus pelagicus H. Milne Edwards, 1853. Our examination confirms that the specimen is indeed Grapsus albolineatus. Alcock (1900: 394) has synonymised Grapsus granulosus, G. pelagicus and G. peroni under Grapsus strigosus (Herbst) without providing any treatment. The specific nomen strigosus Herbst, 1799, as published in the combination Cancer strigosus, is not available because it is a junior homonym of Cancer strigosus, which is actually Galathea strigosa (Linnaeus, 1761), but many authors continued to record Grapsus albolineatus under the name Grapsus strigosus (see Banerjee 1960: 153). Grapsus strigosus has been determined to be the junior synonym of Grapsus albolineatus, e. g. by Banerjee (1960: 147, 154), Sakai (1976: 630), and Ng et al. (2008: 323).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADEFEFFB02B37BF86A.taxon	description	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 10934 (= MNHN-B 10934), ♂ 44.8 × 48.9 mm, Australia, coll. Péron & Lesueur. — Original label: “ Grapsus strigosus Herbst (= Grapsus Peroni Edw.), Nouvelle-Hollande, MM. Péron & Lesueur ”. PRESERVATION. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF5FFADEFEFFB02B37BF86A.taxon	discussion	REMARKS Grapsus peroni was dedicated to François Péron, who attended the Baudin Expedition to the “ Terres Australes ” (1800 - 1804) (Baudin 1974), i. e., the southern coast of Australia, at that time “ Nouvelle-Hollande [New Holland] ”. The other name on the original label refers to Charles-Alexandre Lesueur, also a member of the Baudin Expedition (see Appendix). Crosnier (1965: 17) could not find the type of this species in the MNHN collection. Davie (2002: 213), however, presumed the presence of syntypes, but a single specimen was found in the MNHN.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF2FFABECA8F887B6BFFDB6.taxon	description	(Fig. 5 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF2FFABECA8F887B6BFFDB6.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3439 (= MNHN-B 3439), ♂ 55.0 × 57.4 mm, Red Sea, coll. M. Roux. — Original label: “ Grapsus pharaonis Edw., Mer Rouge, M. Roux ”. PRESERVATION OF THE LECTOTYPE. — Dry. PARALECTOTYPES. — MNHN-IU- 2000 - 3438 (= MNHN-B 3438), 2 ♀ 58.3 × 59.5 mm, 60.2 × 60.9 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF2FFABECA8F887B6BFFDB6.taxon	discussion	REMARKS “ M. Roux ” on the label is J. L. F. P. Roux who collected material in the Mediterranean Sea, Red Sea and in the Near East (Bauchot et al. 1990; Appendix) and published the volume Crustacés de la Méditerranée et de son littoral, décrits et lithographiés (Roux 1828 - 1830). Grapsus pharaonis was synonimised with Grapsus grapsus (Linnaeus, 1758) by A. Milne-Edwards & Bouvier (1900: 111) and, subsequently, with Grapsus tenuicrustatus (Herbst, 1783) by Banerjee (1960), see Ng et al. 2008: 217. In the CAA, material collected by Roux in the Red Sea has 1835 as entry date.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFABED5BFDE3B2D5FBB4.taxon	description	(Fig. 5 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFABED5BFDE3B2D5FBB4.taxon	materials_examined	NEOTYPE (by present designation). — MNHN-IU- 2000 - 3409 (= MNHN-B 3409), mature ♀ 31.6 × 33.3 mm, Grapsus grapsus Linné (= Grapsus pictus Latr.), attacked by bopyrids, Martinique, coll. M. Rivière. — Original label: “ Grapsus grapsus Linné (= Grapsus pictus Latr.), attaqué par les Bopyres, Martinique, M. Rivière ”. PRESERVATION. — Dry. All pereopods detached.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFABED5BFDE3B2D5FBB4.taxon	discussion	REMARKS The specific name pictus was introduced by Lamarck (1801: 150) when he established the new generic name Grapsus (its etymology is related to the ancient Greek grapsaios that means “ crab ”) in indicating “ Grapsus pictus. n. Cancer grapsus. Lin. ”, i. e., Cancer grapsus of Linnaeus (1758). As a specific description is lacking and there is no mention of any material or locality, the specific name pictus of Lamarck (1801) is a nomen nudum. Later, Lamarck (1818: 248) provided a description and a geographical origin “ mers de l’Amérique méridionale ” (seas of South America), still considering it a nomen for Cancer grapsus, now Grapsus grapsus (Linnaeus, 1758). Meanwhile, based on the same material as Lamarck (1818), Latreille (1803: 69, pl. 47, fig. 2) used the name Grapsus pictus in giving a short diagnosis and a figure of a specimen from the “ îles de l’Amérique méridionale ” (islands of South America) that he likewise referred to as the Cancer grapsus of Linnaeus. Therefore, the authorship of G. pictus is not Lamarck (1801) but Latreille (1803). Note that in his list of genera with the indication of their type species, called ‘ genotypes’ (“ Tableau des genres avec l’indication de l’espèce qui leur sert de type ”), Latreille (1810: 422) quoted Cancer grapsus as the type (onomatophore) of the genus Grapsus, and not G. pictus (see also H. Milne Edwards 1836 - 1844: pl. 22). G. grapsus (Linnaeus, 1758) is the type species of Grapsus by tautonomy (Manning & Holthuis 1981; Ng et al. 2008).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFABED5BFDE3B2D5FBB4.taxon	discussion	The MNHN collection contains a sample with two specimens: a preadult female 26.2 × 32.4 mm, MNHN-IU- 2000 - 1101 (= MNHN-B 3409); and a mature female 31.6 × 33.3 mm, MNHN-IU- 2000 - 3409 (= MNHN-B 3409), dry, labelled “ Grapsus grapsus Linné (= Grapsus pictus Latr.), attaqué par les Bopyres, Martinique, M. Rivière ” [this collector is unknown according to the available information and thus is not included in the Appendix]. These individuals are parasitised by bopyrids, the preadult female on both sides and the mature female on the right side. In the handwritten Catalogues of Crustacea by Latreille (LC 1807, LC 1814), two specimens, a male and a female, are cited without locality, date or collector: there is no mention of bopyrids (Fig. 1 B). The figure of Grapsus pictus in Latreille (1803: pl. 47, fig. 2) does not show any deformation of the carapace by a parasite but such an unnatural shape was perhaps not taken into account by the drawer. Even if Latreille (1803) could have confused the preadult female with a male, we do not have enough objective proof to consider that these specimens belong to the type series of Grapsus pictus. Consequently, for clarification purposes the designation of a neotype is justified.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFA8EFF4FBE2B575FC95.taxon	description	(Fig. 5 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFA8EFF4FBE2B575FC95.taxon	materials_examined	LECTOTYPE (designated by Poupin et al. 2005: 37, 38). — MNHN-IU- 2000 - 3585 (= MNHN-B 3585), ♀ 15.0 × 19.0 mm, Sandwich Islands [Hawaiian Is.], coll. Louis Claude de Saulces de Freycinet, 1817 - 1820. — Original labels: “ Pachygrapsus plicatus (Stimp.), Grapsus plicatus Edw., Type, Iles Sandwich, M. Freycinet ” and on the back of the cardboard box “ Type de Grapsus plicatus (M. Edw.), c’est un Metopograpsus ” (“ Type of Grapsus plicatus (M. Edw.), it is a Metopograpsus ”). PRESERVATION OF THE LECTOTYPE. — Dry. Carapace with a crack on the left cardiac / branchial region; RP 1, LP 3 missing; pleon damaged. PARALECTOTYPE (designated by Poupin et al. 2005: 37, 38). — MNHN-IU- 2000 - 1103 (= MNHN-B 3585), 1 ♀ 12.0 × 16.5 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPE. — Dry. Left side of carapace cracked, intestinal region with a gap; LP 1, LP 3, LP 2 dactlyus missing; only coxa LP 4; pleon intact.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF3FFA8EFF4FBE2B575FC95.taxon	discussion	REMARKS Based on the examination of the two MNHN female syntypes from Hawaiian Islands, the lectotype and paralectotype designation by Poupin (2005) (see also Castro 2011: 117) for the lot MNHN-IU- 2000 - 1103 (= MNHN-B 3585) is valid. Previously, Fransen et al. (1997: 126) recorded the RMHN D 42256 specimen from Indonesia as a type of P. plicatus, which actually refers to a nomen nudum of Herklots (1861). Similarly, the Indonesian specimens RMHN D 42234, D 42235, D 45029, and D 45030 that are indicated as “ types of Grapsus plicatus (de Haan, MS) Herklots, 1861, nomen nudum ” by Fransen et al. (1997: 125, under Metopograpsus frontalis Miers, 1880) are not in any way part of the type material of P. plicatus; this was confirmed by C. Fransen (pers comm., December 2012). These erroneous data from Fransen et al. (1997) were confusing for Davie (2002: 218) and Castro (2011: 117) who presumed that a part of the type material of P. plicatus could be in the RMHN. We are unsure as to what Castro (2011: 117) is referring when he mentions in the synonymy of Pachygrapsus plicatus “ Poupin et al. 2005: 37, figs 11, 14 e, 15 e [type material, O’ahu] ”, the mention of the island O’ahu not being cited anywhere in the original publication.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EC8EFCC3B257FD97.taxon	description	(Fig. 5 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EC8EFCC3B257FD97.taxon	materials_examined	HOLOTYPE (by monotypy, see Castro 2011: 114). — MNHN-IU- 2000 - 10875 (= MNHN-B 10875), ♂ 47.5 × 49.8 mm, Sandwich Islands [Hawaiian Is.], coll. Quoy & Gaimard. — Original label: “ Grapsus rudis Edw., Iles Sandwich, MM. Quoy & Gaimard ”. PRESERVATION. — Dry. Carapace stuck with elderberry stub onto cardboard; hairline cracks on the intestinal / branchial regions; RP 5, LP 3 missing,	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EC8EFCC3B257FD97.taxon	discussion	REMARKS Henri Milne Edwards (1837: 86, 87) briefly described a new species for a single specimen that was “ brought from the Sandwich Islands [the Hawaiian Islands] by Quoy & Gaimard ”, in writing “ GRAPSE RUDE ” (i. e., written in small capital letters), not followed in this case by a Latin nomen. Later, H. Milne Edwards (1853: 134 [168]) used the Latin nomen Grapsus rudis. The French name GRAPSE RUDE, not “ used for general purposes ”, is not a vernacular name (see Kottelat 2001). Gibbes (1850: 17) had latinised the name for the first time but, since several carcinologists (Rathbun 1906: 838; Banerjee 1960: 134; Paul’son 1875: 68 [reprinted 1961: 74]; Ng et al. 2008: 217) including Gibbes himself (1850) have attributed the taxon to H. Milne Edwards, it is clear that the authorship of H. Milne Edwards with the date 1837 can be retained for Grapsus rudis (Code, Art. 50.1.1). The MNHN collection contains one specimen, a male labelled “ Grapsus rudis, Sandwich Islands, Quoy & Gaimard ” with the mention “ Type ” on the back of the cardboard box (MNHN-B 10875). Castro (2011: 114) in his catalogue of Brachyura from the Hawaiian Islands regarded it as the holotype of G. rudis, actually a holotype by monotypy. Presently, G. rudis is regarded as a junior synonym of G. tenuicrustatus (Herbst, 1783) (Rathbun 1906: 838; Banerjee 1960: 134; Griffin 1973: 423; Ng et al. 2008: 217; Castro 2011: 114). Griffin (1973: 423) stated that the Grapsus pictus of Quoy & Gaimard (1824: 523, pl. 76, fig. 2) from Guam was synonym of Grapsus rudis, itself a synonym of Grapsus tenuicrustatus, as previously stated by several authors, notably Banerjee (1960: 143) (see under Grapsus pictus).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EFB5FDCCB3D5F9AB.taxon	description	(Fig. 7 H)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EFB5FDCCB3D5F9AB.taxon	materials_examined	NEOTYPE (by present designation). — MNHN-IU- 2000 - 3526 (= MNHN-B 3526), ♂ 24.0 × 27.1 mm, Mediterranean Sea. — Original label: “ Leptograpsus marmoratus Fabr., Méditerranée ”. PRESERVATION. — Dry. RP 5 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA8EFB5FDCCB3D5F9AB.taxon	discussion	REMARKS Latreille (1803: 67, 68, and footnote) stated the material was given to him by a young naturalist, Marcel Serres, who had collected it near Montpellier, on the coast of the Mediterranean Sea; and the species closely resembled Cancer marmoratus (Fabricius, 1787). Henri Milne Edwards (1837: 88) examined the material deposited in the MNHN collection, noting this species was very common along the rocky areas of Bretagne (Brittany), France, Italy, etc. Alphonse Milne-Edwards & Bouvier (1900: 109) synonymised Grapsus varius with Pachygrapsus marmoratus Fabricius, 1787, the type locality of which is unknown (Fabricius 1787: 319). From the original sample MNHN-B 3526, with one male and one female, from the Mediterranean Sea, the male is hereby selected as the neotype of Grapsus varius Latreille, 1803, MNHN-IU- 2000 - 3526 (= MNHN-B 3526). The female is registered as MNHN-IU- 2000 - 1108 (= MNHN-B 3526).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA9EFEDF9E0B593FC55.taxon	description	(Fig. 5 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA9EFEDF9E0B593FC55.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3457 (= MNHN-B 3457), ♂ 28.4 × 31.0 mm, Canary Islands, coll. M. Berthelot. Grapsus webbi H. Milne Edwards, 1853, syntype, revid. M. Türkay, IV- 1977. — Original label: “ Grapsus Webbi Edw. maculatus Catesby, Iles Canaries, M. Berthelot ”. Additional label: “ Grapsus webbi H. Milne Edwards, 1853, syntype, revid. M. Türkay, IV- 1977 ”. PRESERVATION OF THE LECTOTYPE. — Dry. RP 1, RP 5 missing. PARALECTOTYPES. — MNHN-IU- 2000 - 10979 (= MNHN-B 10979), 1 ♂ 54.6 × 62.2 mm, 1 ♀ 32.6 × 38.5 mm, Canary Islands, coll. M. Berthelot. Grapsus webbi H. Milne Edwards, 1853. — Original label: “ Grapsus grapsus Linné (Grapsus Webbi Edw.), Iles Canaries, M. Berthelot ”. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF0FFA9EFEDF9E0B593FC55.taxon	discussion	REMARKS Henri Milne Edwards (1853) described this species based on the material collected by Webb & Berthelot who are well known for their exploration of the Canary Islands where they have gathered a considerable amount of material of all kinds (see Appendix). In their masterpiece “ Histoire naturelle des îles Canaries ”, the part Crustacea has been authored by G. A. Brullé, and this crab was identified as Grapsus strigosus by Brullé (1839: 15), before it was recognised as new by H. Milne Edwards, (1853: 133 [167]) and dedicated to Webb, then synonymised to Grapsus adscensionis (Osbeck, 1765).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9ECB0FB82B572F869.taxon	description	(Fig. 6 A)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9ECB0FB82B572F869.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 4023 (= MNHN-B 4023), ♂ 19.8 × 23.9 mm, Juan Fernández Islands, Chile. — Original label: “ Leptograpsus Ansoni H. M. Edwards, syntypes, Juan Fernandez ”. PRESERVATION OF THE LECTOTYPE. — Dry. Carapace with a small crack on left cardiac region; LP 3, RP 4 missing. PARALECTOTYPES. — MNHN-IU- 2000 - 1105 (= MNHN-B 4023), 1 juv. 10.3 × 12.1 mm, same data as lectotype. — MNHN-IU- 2000 - 3519 (= MNHN-B 3519), 1 ♂ 19.6 × 23.2 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Incomplete specimens.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9ECB0FB82B572F869.taxon	discussion	REMARKS The original samples MNHN-B 4023 and MNHN-B 3519 have an additional label by D. Guinot in 1972 with the indication “ presumed syntypes ”. This was cited by Griffin (1973: 418), who mentioned “ three syntypes ” in the MNHN when he recognised the species as a junior synonym of Leptograpsus variegatus. The specific name ansoni refers to the Commodore George Anson, famous for his Voyage around the world to disrupt or capture Spain’s Pacific possessions, reaching the Juan Fernández Archipelago in 1741.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9EF18FF4DB4B6F94A.taxon	description	(Fig. 6 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9EF18FF4DB4B6F94A.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 10944 (= MNHN-B 10944), ♀ 38.0 × 43.2 mm, Canary Islands, coll. MM. Webb & Berthelot. — Original label: “ Leptograpsus Bertheloti M. Edw., MM. Webb et Berthelot, Iles Canaries ”. PRESERVATION. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA9EF18FF4DB4B6F94A.taxon	discussion	REMARKS Alphonse Milne-Edwards & Bouvier (1894: 48) wrote under Leptograpsus marmoratus: “ Ce Crabe, que l’on trouve communément sur les côtes océaniques de la France, semble avoir une large dissémination géographique, il se trouve à Madère, aux îles Canaries, et nous avons lieu de penser qu’il ne diffère pas spécifiquement des formes décrites sous les noms de Leptograpsus variegatus Fabricius et de L. Bertheloti H. Milne Edwards. ” [“ This crab, commonly found on the oceanic coasts of France, seems to have a widespread geographical distribution, it is found in Madeira, the Canary Islands, and there may be reason to believe that it does not differ specifically from the forms described under the names Leptograpsus variegatus Fabricius and L. Bertheloti H. Milne Edwards ”]. Kingsley (1880: 196) placed many species in the synonymy of L. variegatus including L. bertheloti, but Griffin (1973: 423), who also gave a long list of synonyms, did not recognise L. bertheloti as a synonym of L. variegatus [considering instead a possible synonymy with Pachygrapsus marmoratus (Fabricius)], both without giving an explanation of their statements. Rathbun (1918: 250) and Ng et al. (2008: 217) placed Leptograpsus bertheloti, with reservation, in the synonymy of Pachygrapsus marmoratus, a species found in the Mediterranean Sea and on the Atlantic coast of Europe, from Britanny to Morocco, including Madeira, the Azores, and the Canary Islands (Manning & Holthuis 1981: 225; d’Udekem d’Acoz 1989: 255). Our examination of the specimen MNHN-IU- 2000 - 10944 (= MNHN-B 10944) has confirmed that Leptograpsus bertheloti is indeed Pachygrapsus marmoratus, and not Leptograpsus variegatus (Fabricius, 1793) as stated by Kingsley (1980). Recent publication on the phylogeography of Pachygrapsus marmoratus from the African Mediterranean coast (Deli et al. 2016) has revealed a genetic homogeneity across the Siculo-Tunisian Strait.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA7EFFAF880B5D3FE56.taxon	description	(Fig. 6 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA7EFFAF880B5D3FE56.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3531 (= MNHN-B 3531), ♂ 22.6 × 25.6 mm, Valparaiso, Chile, coll. C. Gay, 1828 - 1832. — Original labels: “ Leptograpsus planifrons Dana, Valparaiso, M. Gay, 1828 - 1832 ”; “ L. planifrons (= Leptograpsus gayi, Edw. Type) ”. PRESERVATION. — Dry. LP 3 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFF1FFA7EFFAF880B5D3FE56.taxon	discussion	REMARKS Henri Milne Edwards (1853) dedicated this species to Claude Gay, who collected material in Chile (see Appendix). Previously, Nicolet in Gay (1849: 167) in the Historia de Chile identified their material from Chile as Grapsus variegatus (Fabricius, 1793). Griffin (1973: 418, 423), based on the inaccurate information provided by D. Guinot, stated that the type of L. gayi was not extant, and he synonymised this species with L. variegatus. According to Davie (2002: 216) the holotype of L. gayi was deposited at the MNHN, and indeed, one specimen, MNHN-IU- 2000 - 3531 (= MNHN-B 3531), from Valparaiso, Chile, with the name of Gay and labelled “ Type ” has been found in the MNHN dry collection. It is designated here as the lectotype of Leptograpsus gayi.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1FD8CB54DF9EA.taxon	description	(Fig. 6 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1FD8CB54DF9EA.taxon	materials_examined	LECTOTYPE (present designation). — MNHN-IU- 2000 - 3534 (= MNHN-B 3534), ♂ 12.9 × 16.1 mm, Brazil, coll. Gaudichaud. — Original label: “ Leptograpsus rugulosus Lamk., Brésil, M. Gaudichaud ”. PRESERVATION. — Dry. RP 2 - 3, LP 1, RP 5 propodus and dactylus missing; LP 4 - 5 carpus, merus and dactylus missing; ventral surface with elder stub.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1FD8CB54DF9EA.taxon	discussion	REMARKS Rathbun (1918: 245) mentioned that the type was in “ Paris Museum ” but without listing any material. Although Leptograpsus rugulosus was recognised by Kingsley (1880), it is the junior synonym of Pachygrapsus transversus (Gibbes, 1850) according to Poupin et al. (2005: 44). The MNHN collection contains two samples that could have been seen by H. Milne Edwards in 1853: a complete and dry female specimen MNHN-IU- 2000 - 3535 (= MNHN-B 3535), collected from Rio de Janeiro by Peter Claussen (see Appendix), and a specimen MNHN-IU- 2000 - 3534 (= MNHN-B 3534) from Brasil collected by Charles Gaudichaud-Beaupré, who was a well-known, extremely active collector at that time in the MNHN (see Appendix). The first entry in the MNHN of South American material collected by Gaudichaud is from 1832, as indicated in the CAA. Henri Milne Edwards was used to mention the localities in accordance with the information provided by the collector, thus the type locality is “ Brésil ” and not “ Rio de Janeiro ”. The specimen MNHN-IU- 2000 - 3534 (= MNHN-B 3534) of Gaudichaud is selected as lectotype.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1F920B3A2FC56.taxon	description	(Fig. 6 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1F920B3A2FC56.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 4024 (= MNHN-B 4024), ♂ 34.9 × 39.6 mm, Australia, coll. J. P. Verreaux, 1842. Presumed Syntype, D. Guinot 1972. — Original label: “ Leptograpsus Verreauxi H. M. Edwards, Syntype, Australie, M. Verreaux ”. Additional label: “ Syntype présumé, D. Guinot 1972 ”. PRESERVATION OF THE LECTOTYPE. — Dry. Pleon with a small crack; telson folded inwards. PARALECTOTYPE. — MNHN-IU- 2000 - 3546 (= MNHN-B 3546), 1 ♂ 30.7 × 34.1 mm, same data as lectotype. — Additional label: “ Presumed Syntype, D. Guinot 1972 ”. PRESERVATION OF THE PARALECTOTYPE. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7ECC1F920B3A2FC56.taxon	discussion	REMARKS In 1972 Guinot wrote a new label “ presumed syntypes ” for the samples MNHN-IU- 2000 - 4024 (= MNHN-B 4024) and MNHN-IU- 2000 - 3546 (= MNHN-B 3546) collected by Verreaux in Australia. Based on this information, Griffin (1973: 418, 423) mentioned that two syntypes were deposited at the MNHN when he recognised the species as a junior synonym of Leptograpsus variegatus. The larger male from the type series is selected here as lectotype. The entry date in the CAA of a considerable material of the Verreaux collection from Tasmania, Australia is 1847, which corroborates the hypothesis that H. Milne Edwards has established his new species after this Verreaux material (see Appendix).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7EF18FB82B354F869.taxon	description	(Fig. 6 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7EF18FB82B354F869.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3547 (= MNHN-B 3547), ♂ 16.6 × 20.1 mm, Sandwich Islands [Hawaiian Is.], Type, coll. J. T. Eydoux & L. F. A. Souleyet. — Original label: “ Metopograpsus eydouxi Edw., Type. Iles Sandwich, MM. Eydoux et Souleyet ”. PRESERVATION. — Dry. RP 1 - 3, 5, LP 4 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFFFFA7EF18FB82B354F869.taxon	discussion	REMARKS Davie (2002: 217) wrote “ type status unknown MNHP * ”, thus only a probable MNHN depository. The sample MNHN-IU- 2000 - 3547 (= MNHN-B 3547) was collected from the type locality, the Sandwich Islands, i. e., the Hawaiian Is., by the well-known scientists Eydoux and Souleyet who visited these islands, notably during the Expedition of the Bonite (1836 - 1837) (see Appendix). Their names are inscribed on the original label, so this material is here selected as lectotype. In contrast, the dry sample MNHN-IU- 2000 - 3551 (= MNHN-B 3551) from the Sandwich Islands, with two specimens, 1 ♂ and 1 ♀, was collected later as shown by the label “ 788 - 66 ” corresponding to the entry in 1866 listed in the CAA as material purchased from “ M. Robson, London ”.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4EC89FF4DB5B0F98B.taxon	description	(Fig. 7 A)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4EC89FF4DB5B0F98B.taxon	materials_examined	LECTOTYPE (by inference from holotype designation by Türkay 1974 c). — MHNG, ♀ 16.0 × 21.0 mm, St Thomas, Virgin Is. PARALECTOTYPE. — MNHN-IU- 2000 - 10991 (= MNHN-B 10991), 1 ♀ 8.4 × 11.1 mm, Saint Thomas, coll. M. de Saussure. — Original label: “ Pachygrapsus gracilis Saussure, ♀, St Thomas, M. de Saussure ”; on the back of the cardboard “ Metopograpsus gracilis Sauss., ♀, St Thomas = Pachygrapsus, M. de Saussure, 270.58 ”. PRESERVATION. — Carapace cracked on right side at cardiac region, RP 4 missing.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4EC89FF4DB5B0F98B.taxon	discussion	REMARKS Saussure (1858 a: 443, pl. 2, fig. 15, 15 a-c) established this species from a specimen measuring 16.0 × 21.0 mm collected from St Thomas, Virgin Islands (see Appendix), but his description did not explicitly mention how many specimens were examined. A female, 16.0 × 21.0 mm, in the MHNG was regarded as holotype by Türkay (1974 c: 144, pl. 1, figs 5, 6) in his paper on the Grapsidae deposited at the MHNG: it is a dry specimen (P. Schwendinger, pers. comm., Dec. 2012). In assuming a holotype rather than designating a lectotype, Türkay (1974 b: 145) proceeded as though syntypes did not exist. But there is a specimen, a dry female, collected by Saussure from the same locality, St Thomas, deposited in the MNHN collection, MNHN-IU- 2000 - 10991 (= MNHN-B 10991), as confirmed by the mention in the CAA of an exchange between Saussure and the MNHN in 1858, entry 270, corresponding to “ 270.58 ” on the original label. We may consider that this specimen has been collected at the same time as the holotype and could belong to the syntype series. Therefore, Türkay (1974 c), who has published the assumption that the species-group taxon was based upon a single type specimen, is deemed to have designated that specimen as the lectotype (Code, Art. 74.6: Fixation of lectotype by inference of “ holotype ”). The female MNHN-IU- 2000 - 10991 (= MNHN-B 10991) may be considered paralectotype.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4ECD9F9C0B2CDFA0B.taxon	description	(Fig. 7 B)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4ECD9F9C0B2CDFA0B.taxon	materials_examined	LECTOTYPE (designated by Tweedie 1954: 128). — MNHN-IU- 2014 - 11210 (= MNHN-B 12888), ♂ 18.0 × 23.2 mm, Type, patria ignota. “ Metopograpsus thukuhar (Owen) after M. Tweedie 1948 who has examined the pleopods drawn by Miss I. Gordon ” [“ Metopograpsus thukuhar (Owen, 1839) selon M. Tweedie 1948 qui a examiné les pléopodes dessinés par Miss I. Gordon ”]. — Original label: “ Metopograpsus intermedius Edw. (type) ”. PRESERVATION. — Alcohol. Six pereopods detached from the body.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA4ECD9F9C0B2CDFA0B.taxon	discussion	REMARKS De Man (1888: 359) then Tweedie (1949: 470) suspected a problem with the identity of Metopograpsus intermediu s. Upon Tweedie’s request, Isabella Gordon examined the type material at the MNHN. According to Tweedie (1949: 470), Gordon found two specimens in the MNHN labelled as “ cotypes ”, patria ignota, but her sketches of the G 1 of each of these two “ cotypes ” revealed that they belonged to two different species. Tweedie (1954: 127, 128, under M. gracilipes) stated the name M. intermedius could not be used because it was ‘ composite’, and his first opinion was that the bigger specimen MNHN-IU- 2014 - 11880 (= MNHN-B 12856) could be selected as the lectotype of M. intermedius, a species that he recognised to be a senior synonym of M. gracilipes De Man, 1891. Nevertheless, in order to preserve stability of nomenclature Tweedie (1954: 127) has accepted the recommendation of I. Gordon and L. B. Holthuis in writing: “ The two authorities whom I consulted [Gordon and Holthuis] have suggested that gracilipes would be established as a valid name if the smaller of Milne Edwards’ cotypes were designated as lectotype of his species M. intermedius ”. Tweedie (1954) finally selected the smaller specimen MNHN-IU- 2014 - 11210 (= MNHN-B 12888) as lectotype of M. intermedius, on which M. thukuhar (Owen, 1839) has priority (see Banerjee 1960: 186; Ng et al. 2008: 217). The bigger specimen MNHN-IU- 2014 - 11880 (= MNHN-B 12856) belonging to the syntype material of Metopograpsus intermedius (see Code, Art. 72.4.4) is M. gracilipes De Man, 1891, presently a synonym of M. frontalis Miers, 1880 (Banerjee 1960: 182; Davie 2002: 216; Ng et al. 2008: 217). It is a male, 22.1 × 30.2 mm, in alcohol, with ten detached pereopods, pleon absent. — Original label: “ Metopograpsus intermedius Edw. (Type) ”. Additional labels: “ Metopograpsus gracilipes de Man emend. according to M. Tweedie 1948 who has examined the pleopods drawn by Miss I. Gordon [Metopograpsus gracilipes de Man emend., d’après M. Tweedie 1948 qui a examiné les pléopodes dessinés par Mlle I. Gordon] ”.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA5EF26FA41B46DFED0.taxon	description	(Fig. 7 C)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA5EF26FA41B46DFED0.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3554 (= MNHN-B 3554), ♂ 31.2 × 35.3 mm, Batavia [Jakarta], coll. M. Meder. — Original label: “ Metopograpsus maculatus Edw., Type, Batavia, M. Meder ”. PRESERVATION OF THE LECTOTYPE. — Dry. Right region of pleonal somite 6 and telson missing. PARALECTOTYPES. — MNHN-IU- 2000 - 3553 (= MNHN-B 3553), 1 ♂ 24.8 × 27.7 mm, 1 ♀ 27.5 × 30.7 mm, same data as lectotype. — MNHN-IU- 2000 - 1106 (= MNHN-B 3554), 2 ♂ 28.9 × 32.0 mm, 30.3 × 33.2 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — Dry.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFCFFA5EF26FA41B46DFED0.taxon	discussion	REMARKS Meder is a well-known collector in this region (see Appendix).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EC90FE0DB205FE57.taxon	description	(Fig. 7 D)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EC90FE0DB205FE57.taxon	materials_examined	LECTOTYPE (by inference of holotype by Türkay 1974 c). — MHNG, ♂, St Thomas, Virgin Is. PRESERVATION OF THE LECTOTYPE. — Carapace with sides distended by a bopyrid (see below); RP 2 - 3, 5 and LP 2 - 4 missing. PARALECTOTYPE. — MNHN-IU- 2000 - 10994 (= MNHN-B 10994), 1 ♂ 10.4 × 14.2 mm, Saint Thomas, coll. M. de Saussure. — Original label: “ Metopograpsus miniatus Saussure, St Thomas, M. de Saussure ”. PRESERVATION OF THE PARALECTOTYPE. — Dry. Left side distended by a parasite. RP 1 - 2, RP 5 missing, RP 3, 4 detached.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EC90FE0DB205FE57.taxon	discussion	REMARKS Saussure (1858 a: 444, pl. 2, fig. 17, 17 a-c) described a crab collected from St Thomas, Virgin Is., measuring 12.0 × 16.0 mm as Metopograpsus miniatus, but did not explicitly mention how many specimens he had examined. Rathbun (1918: 244, 245), who synonymised M. miniatus with Pachygrapsus transversus, wrote that the type of M. miniatus was deposited at the MHNG and “ its sides [carapace] are distended by a parasite, one side more than the other ”. She probably received this information from Zehntner, the Swiss scientist who took care of the MHNG collections at the end of the 19 th century (see Appendix). In his paper on the Grapsidae deposited in the MHNG, Türkay (1974 c: 142, figs 1, 2, pl. 1, figs 1, 2) regarded this specimen, a male, as holotype of M. miniatus; he figured the G 1 and provided a note written on a label by Zehntner signaling the presence of a parasite on the branchial regions. The holotype photographed by Türkay (1974 c: 143, pl. 1, figs 1, 2) shows a slightly swollen left branchial region, while neither the description nor the figure of Saussure (1858 a: 444, pl. 2, fig. 17) indicates the presence of a parasite deforming the carapace. In assuming a holotype rather than selecting a lectotype, Türkay (1974 c: 142, 143) proceeded as though other syntypes did not existed. But we have found supplementary material, a male MNHN-IU- 2000 - 10994 (= MNHN-B 10994), collected by Saussure from St Thomas, presumably at the same time, as confirmed by the mention in the CAA of an exchange between Saussure and the MNHN in 1858, corresponding to the entry “ 262.58 ”. This specimen likewise shows a deformation of the carapace. The fact that both specimens are similarly parasitised may be an indication that they should be from the same lot and, presumably, they belong to the syntype series, despite there being no indication of swelling on the carapace in Saussure’s paper. It is reasonable to expect that the original description was based on more than one specimen. Therefore, Türkay (1974 c), who published the assumption that the species-group taxon was based upon a single type specimen in recognising a holotype, is deemed to have designated that specimen as the lectotype (Code, Art 74.6). The male MNHN-IU- 2000 - 10994 (= MNHN-B 10994) may be considered a paralectotype. It is also the case with Metopograpsus gracilis Saussure, 1858, also from Saint Thomas (see above).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EF0CFD8CB3ECF869.taxon	description	(Fig. 7 E)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EF0CFD8CB3ECF869.taxon	materials_examined	LECTOTYPE (by inference of holotype by Davie (2002: 217). — MZUT, Cr. 1026, New Caledonia. PARALECTOTYPES. — MNHN-IU- 2000 - 3573 (= MNHN-B 3573), 1 ♂ 28.2 × 31.8 mm, New Caledonia. — Original label: “ Metopograpsus pictus A. M. Edw., cat. du Muséum 225 - 63, Nelle Calédonie ”. — MNHN-IU- 2000 - 10893 (= MNHN-B 10893), 1 ♂ 24.2 × 28.1 mm, 1 ♀ 24.4 × 24.9 mm, Nlle Calédonie, M. Beaudouin; “ 369 - 65 ” is written with Chinese ink directly on the carapaces of both specimens. — MNHN-IU- 2000 - 3572 (= MNHN-B 3572), 1 ♂ 17.2 × 20.6 mm, New Caledonia, coll. Beaudouin. PRESERVATION. — Dry. LP 5 missing, RP 4 - 5 dactyli missing for male MNHN-IU- 2000 - 3573; LP 2 missing for female of MNHN-IU- 2000 - 10893; most of pereiopods missing for male MNHN-IU- 2000 - 3572.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFDFFA5EF0CFD8CB3ECF869.taxon	discussion	REMARKS Davie (2002: 217) stated that the holotype of Metopograpsus pictus as “ MZUT Cr. 1026 ” was deposited at the Museo Regionale di Scienze Naturali, Torino, Italy (MRSN). The previous curator of this institution has confirmed that a “ holotype, MZUT Cr. 1026 ” was present in their collection (Alberto Chiarle, pers. comm., Dec. 2012). It must be treated as the lectotype. According to the CAA, the entry 225 - 63 designates the sample MNHN-IU- 2000 - 3573 (= MNHN-B 3573) that was collected by M. De Planche (although not mentioned on the original label) in New Caledonia and given to the Muséum by M. Aubry-Lecomte, director of the Musée des Colonies. The entry 369 - 65 designates the samples MNHN-IU- 2000 - 10893 (= MNHN-B 10893) and MNHN-IU- 2000 - 3572 (= MNHN-B 3572) collected in New Caledonia and given to the Muséum by M. Beaudouin (Navy Captain, see Appendix). They could be considered paralectotypes (Code, Art. 72.4). A dry specimen labelled Metopograpsus pictus deposited at the Peabody Museum of Natural History in Yale University (YPM IZ 001463) does not belong to the type series: it was collected from Dominica, West Indies, by A. E. Verrill’s sons, George and Alpheus, and sent over by A. Milne-Edwards in 1873, according to the information that we received (E. A. Lazo-Wasem, pers comm. Dec 2012).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECBAFF4CB6C7FCD6.taxon	description	(Fig. 7 F)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECBAFF4CB6C7FCD6.taxon	materials_examined	LECTOTYPE (designated by Poupin et al. 2005: 31). — MNHN-IU- 2000 - 3582 (= MNHN-B 3582), ♂ 7.6 × 10.4 mm, coll. M. Balansa, New Caledonia. — Original label: “ Pachygrapsus minutus A. Edw., Nelle Calédonie, M. Balansa ”. PRESERVATION OF THE LECTOTYPE. — Dry. PARALECTOTYPES. — MNHN-IU- 2000 - 1107 (= MNHN- B 3582), 4 ♂ 5.5 × 7.7 – 7.3 × 10.0 mm, same data as lectotype. PRESERVATION OF THE PARALECTOTYPES. — All are dry specimens.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECBAFF4CB6C7FCD6.taxon	discussion	REMARK Balansa collected much carcinological material from New Caledonia (see Appendix).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECB7FC02B383FB34.taxon	description	(Fig. 7 G)	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECB7FC02B383FB34.taxon	materials_examined	LECTOTYPE (by present designation). — MNHN-IU- 2000 - 3586 (= MNHN-B 3586), ♂ 13.5 × 16.0 mm, Sandwich Islands [Hawaiian Is.], coll. M. Ballieu. — Original label: “ Pachygrapsus striatus A. M. Edw., M. Ballieu, îles Sandwich; P. striatus = Grapsus striatus Edw. ”. PRESERVATION OF THE LECTOTYPE. — Dry. PARALECTOTYPES. — MNHN-IU- 2000 - 1110 (= MNHN-B 3586), 1 ♂ 12.1 × 15.1 mm and 1 ♂ with carapace cl 12.3 mm, same data as lectotype. MNHN-IU- 2000 - 3583 (= MNHN- 3583), 1 ♀ female 13.5 × 10.4 mm, Upolu — Other label: “ Pachygrapsus plicatus A. M. Edwards, Upolu (Samoa), 73 - 70 ”. PRESERVATION OF THE PARALECTOTYPES. — MNHN-IU- 2000 - 1110 (= MNHN-B 3586), both dry, the larger one with damaged carapace, LP 1, RP 5 detached. MNHN-IU- 2000 - 3583 (= MNHN- 3583), dry, RP 2 missing, LP 1 detached.	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
03EEDE4BFFFBFFA3ECB7FC02B383FB34.taxon	discussion	REMARKS In a paper dealing with the crabs from the Museum of M. C. Godeffroy, a private museum in Hamburg, Germany, which existed from 1861 to 1885 (see Appendix at Godeffroy Museum), A. Milne-Edwards (1873 b) described as Pachygrapsus striatus material collected in the Sandwich Islands and in Samoa at Upolu: he quotes the number 5806 and cites a specimen measuring 14.0 × 11.0 mm. The species was synonymised to P. plicatus by Kingsley (1880), Ortmann (1894) and Tesch (1918: 77). This synonymy was recognised by Poupin et al. (2005: 38, 42) but not mentioned by Castro (2011: 114) in his Catalogue of Brachyura from the Hawaiian Islands. The original sample MNHN-B 3586, with three males, from Sandwich Islands, collected by M. Ballieu (poorly handwritten on the label, thus erroneously cited as “ Baillien ” by Poupin 2005: 38) (see Appendix) presumably belongs to the type series, as stated by Poupin et al. (2005: 42). Even though the original sample MNHN-B 3583 is labelled as P. plicatus, in the CAA entry data 73 - 70 (thus corresponding to 1870), it is registered as P. striatus. Thus it could have been examined by A. Milne Edwards and belong to the type series. Ortmann (1894: 708, see footnote n ° 3) recorded a male and a female of P. plicatus from Upolu, as “ Mus. Godeffroy (vend.) ” and with a footnote that mentions the same number 5806 [“ Vend. ” could indicated that the material was sold by the Godeffroy trading company]. Thus the P. plicatus of Ortmann (1894) could correspond to the P. striatus material of A. Milne Edwards (1873 b). The MZS handles one sample (MZS Cru 0742) labelled as “ Pachygrapsus plicatus Upolu, Godeffroy Mus., 1888 ”, with one male (with all pereiopods detached) and one female (with LP 1 - P 5, RP 1, RP 4 - 5 detached) in the alcool collection (M. - D. Wandhammer, Chief Curator of MZS, personal communication, September 2017). Although there is no mention on the label of the number 5806 quoted by A. Milne-Edwards (1873 b) and Ortmann (1894), this sample could belong to the type series, and the two specimens can be considered paralectotypes. Poupin (2005: 39) recorded a dry sample of P. plicatus A. M. Edwards, Upolu, God. 88 (MZS Cru 1210), but this catalogue number deals with Orithyia sinica (Linnaeus, 1771) in the catalogue of the Strasbourg Museum (M. - D. Wandhammer, pers. comm., December 2017).	en	Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle, Mollaret, Noémy (2019): Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema 41 (7): 91-130, DOI: 10.5252/zoosystema2019v41a7
