identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EF87AA6527FFCECA0DE0BEFA4BF802.text	03EF87AA6527FFCECA0DE0BEFA4BF802.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afroriccardia comosa (Steph.) Reeb & Gradst.	<div><p>Afroriccardia comosa (Steph.) Reeb &amp; Gradst.</p> <p>Aneura comosa Steph., Botanical Gazette 15: 281 (1890). (Riccardia comosa [Steph.] E.W.Jones nom. inval.). — Type: Réunion, 1889, Rodriguez s.n. (holo-, G[G00045027]!; iso-, PC[PC0103522]!).</p> <p>FURTHER SPECIMENS EXAMINED. — Madagascar. Angavokely Forest, 18°55’16”S 47°44’30”E, 1600 m, 2.II.2011 Reeb CR11188* (PC0763877, TAN); Zahamena National Park, 17°38’19’’S, 48°36’46”E, 1156 m, 28.XII.2013, Reeb &amp; Andriamanantena 13Z28* (PC0763872, TAN), 13Z32* (PC0763873, TAN). Zahamena National Park, on seeping rocks, highest part of the river</p></div> 	https://treatment.plazi.org/id/03EF87AA6527FFCECA0DE0BEFA4BF802	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6527FFCECAD6E39FFBE8F9A6.text	03EF87AA6527FFCECAD6E39FFBE8F9A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afroriccardia Reeb & Gradst.	<div><p>Genus Afroriccardia Reeb &amp; Gradst.</p> <p>REMARKS</p> <p>A monospecific African genus. For characters of Afroriccardia see under the species.</p> </div>	https://treatment.plazi.org/id/03EF87AA6527FFCECAD6E39FFBE8F9A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6527FFCECA80E278FA55FA87.text	03EF87AA6527FFCECA80E278FA55FA87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneuraceae	<div><p>Family ANEURACEAE</p> <p>A family of five genera worldwide (Preussing et al. 2010; Rabeau et al. 2017), including three in Africa: Riccardia Gray, Aneura Dumort. and Afroriccardia Reeb &amp; Gradst. The principal characteristics of Aneuraceae are: 1) thallus pinnate or unbranched; 2) midrib usually indistinct; 3) gametoecia at thallus margins or on short lateral branches; 4) sporophyte in a fleshy calyptra; and 5) elaters attached to the tips of the valves.</p> </div>	https://treatment.plazi.org/id/03EF87AA6527FFCECA80E278FA55FA87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6524FFCDCA91E31FFA4AF9E7.text	03EF87AA6524FFCDCA91E31FFA4AF9E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneura Dumort.	<div><p>Genus Aneura Dumort.</p> <p>REMARKS</p> <p>About 20 species worldwide (Söderström et al. 2016); two species in Africa. Aneura is recognized by the rather broad and fleshy, glossy green, scarcely branched, dioicous thallus with entire to somewhat crisped margins and without midrib.</p> </div>	https://treatment.plazi.org/id/03EF87AA6524FFCDCA91E31FFA4AF9E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6524FFCCCA95E07EFD42FD1A.text	03EF87AA6524FFCCCA95E07EFD42FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneura latissima Spruce	<div><p>Aneura latissima Spruce</p> <p>Riccardia latissima (Spruce) Schiffn., Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 67: 177 (1898) — Type: Brazil, Amazonas, Rio Negro, on rotten logs, Spruce s.n. (syn-, MANCH!).</p> <p>Aneura pseudopinguis (Herzog) Pócs, Acta Botanica Hungarica 29: 133 (1983) (Riccardia pinguis Herzog). — Type: Brazil, St. Catarina, Jaraguá, 3.X.1937, Carl s.n. (holo-, JE n.v.), syn. fide Gradstein (2013).</p> <p>SPECIMENS EXAMINED. — Madagascar. Alaotra-Mangoro, Vohimana Reserve 18°55’09.9”S, 49°30’49.4”E, 827 m, 15.IV.2010, Reeb, Andriamanantena &amp; Bidault CRAE142, (PC0724341A, TAN); Maromizaha Reserve 18°58’34.4”S, 48°27’54.5”E, 1095 m, 05.VII.2012, Reeb MTM1280 (PC, TAN); Sava, Makirovana, 14°10’29.3”S, 49°57’07.7”E, 695 m, 06.V.2010, Reeb CR282 (PC0763839).</p> <p>DISTRIBUTION. — Tropical South America and tropical Africa (Gradstein et al. 1983; Meenks &amp; Pócs 1985). Widespread in tropical Africa, recorded from Angola, Bioko, Congo, Ethiopia, Gabon, Ghana, Ivory Coast, Mauritius, Madagascar, Malawi, Nigeria, Reunion, Rwanda, São Tomé and Principe, Seychelles, Sierra Leone, Tanzania, Uganda and Zimbabwe (Wigginton 2018). Records of A. latissima from South Africa are referred to A. pinguis (Perold 2001a).</p> <p>HABITAT. — On rotten wood and bark, rock and muddy banks in humid submontane and montane forests (Meenks &amp; Pócs 1985).</p> <p>DESCRIPTION</p> <p>Meenks &amp; Pócs (1985), Wigginton (2004). Mycorrhiza-like fungi may be observed in thallus cross-section, but they do not seem to be constantly present in African Aneura.</p> </div>	https://treatment.plazi.org/id/03EF87AA6524FFCCCA95E07EFD42FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6525FFCCC858E3FFFED6F926.text	03EF87AA6525FFCCC858E3FFFED6F926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneura pinguis	<div><p>Aneura pinguis (L.) Dumort.</p> <p>Jungermannia pinguis L., Species Plantarum 1: 1136 (1753). — Riccardia pinguis (L.) Gray, A Natural Arrangement of British Plants 1: 684 (1821). — Type: Europa (Grolle 1976).</p> <p>DESCRIPTION</p> <p>Perold (2001a).</p></div> 	https://treatment.plazi.org/id/03EF87AA6525FFCCC858E3FFFED6F926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6525FFCCCADEE25FFA54F802.text	03EF87AA6525FFCCCADEE25FFA54F802.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia angusticosta (Steph.) Grolle	<div><p>Riccardia angusticosta (Steph.) Grolle</p> <p>(Fig. 2 A-E)</p> <p>Aneura angusticosta Steph., Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 88: 724 (1913). — Type: Tanzania, Usambara, 1909, Brunnthaler s.n. (G[G0067547]!).</p> <p>Aneura congoana Steph., Species Hepaticarum 6: 23 (1917). — Type: Zaire (Reeb &amp; Bardat 2014).</p> <p>Aneura stephanii Besch. ex Steph., Revue Bryologique 20: 60 (1893). (Riccardia stephanii [Besch. ex Steph.] E.W.Jones nom. illeg.). — Type: Congo (Jones 1956 as R. stephanii).</p> <p>Aneura travisiana Pearson, Memoirs and Proceedings of the Manchester Literary and Philosophical Society 65: 1 (1921). — Type: Cameroon (Jones 1956 as R. stephanii).</p> <p>Riccardia amazonica auct., (e.g. Meenks &amp; Pócs 1985; Wigginton 2004; Reeb &amp; Bardat 2014), non typus.</p> <p>Riccardia capensis S.W.Arnell, see Index Hepaticarum — Type: South Africa (Reeb &amp; Bardat 2014).</p> <p>FURTHER SPECIMENS EXAMINED. — Gabon. Ougoué-Invido, réserve intégrale d’Ipassa, 00°30’N, 12°48’E, 510 m, 18.IV.2006, Vanderpoorten GAB1695* (LG).</p> <p>Ghana. West Region, Aiyinasi, 13.II.1971, Jones 1364 C (E 00018737), 1361 C (G 00264220).</p> <p>Madagascar. Alaotra-Mangoro, Vohibola reserve, 18°34’53.4”S, 49°14’32.2”E, 5 m, 18.IV.2010, Reeb, Andriamanantena &amp; Bidault CRAE 176 (PC 0764349 A); Sava, Makirovana, 14°10’29.3”S, 49°57’07.7”E, 695 m, 06. V.2010, Reeb CR 275 (PC 0763840 A),</p></div> 	https://treatment.plazi.org/id/03EF87AA6525FFCCCADEE25FFA54F802	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6525FFCCCA86E7FAFBE0FBC5.text	03EF87AA6525FFCCCA86E7FAFBE0FBC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia Gray	<div><p>Genus Riccardia Gray</p> <p>REMARKS</p> <p>About 100 species worldwide (Söderström et al. 2016). Based on the results of species delimitation analysis (Reeb et al. 2018), 12 species are recognized in Africa. Eleven of these occur in eastern Africa (Fig. 1) and three are new to science: R. gasparii sp. nov., R. martinii sp. nov. and R. vohimanensis sp. nov. In addition, Riccardia multifida, a species known from South Africa, may be expected to occur in eastern Africa and is therefore included in this treatment. The genus Riccardia is characterized by the 1-3-pinnately or palmately branched thalli with a narrow, 0.3-1.5(-2) mm wide axis and short sexual branches. The archegonia and antheridia are developed in two straight rows on separate branches. The genus is closely related to Aneura and Afroriccardia but the latter two genera have broader, less branched axes (more than 2 mm wide). In early literature, Riccardia was part of Aneura (e.g. Stephani 1898-1924).</p> </div>	https://treatment.plazi.org/id/03EF87AA6525FFCCCA86E7FAFBE0FBC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652AFFC3C88CE6F4FA99FEA8.text	03EF87AA652AFFC3C88CE6F4FA99FEA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia	<div><p>KEY TO THE SPECIES OF RIccARdIA OF EASTERN AFRICA</p> <p>1. Thallus margins with numerous small, simple or branched papillae. Thallus very small, 2-10 mm long...................................................................................................................... R. inconspicua (Steph.) Reeb &amp; Bardat</p></div> 	https://treatment.plazi.org/id/03EF87AA652AFFC3C88CE6F4FA99FEA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652BFFC0CAE7E3DEFC14FB04.text	03EF87AA652BFFC0CAE7E3DEFC14FB04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia chamedryfolia (With.) Grolle	<div><p>Riccardia chamedryfolia (With.) Grolle</p> <p>(Fig. 2 F-I)</p> <p>Jungermannia chamedryfolia With., A Botanical Arrangement of the Vegetables of Great Britain 2: 699 (1776). — Type: England (Grolle 1976).</p> <p>SPECIMENS EXAMINED. — Comores. Grandes Comores, Karthala, 1600 m, 01.V.2008, Bardat KarP1Q2Hu1* (PC0763845).</p> <p>Ethiopia. Bonga, Araba Yel forest, 37°80’77.54”N, 18°73’34”E, 1798 m, Hylander KH 5490* (ETH, PC 0763857); Boka forest, 02.XI.2006, Hylander KH 5497 * (ETH, PC 0763862); Bale Mts., Harenna forest, 06°42’58.3”N, 39°43’32.1”E, 2380 m, 10.VIII.2011, Reeb, Queinnec &amp; Wégé CR 11418 * (ETH, PC 0763898), ibid., road to Rira, 06°43’09.4”N, 39°43’11.3”E, 2392 m, 10.VIII.2011, Reeb, Queinnec &amp; Wégé CR 11425 *(ETH, PC 0763893), CR 11428 * (PC 0763896, ETH).</p> <p>Madagascar. Analamanga, Station d’Angavokely, 18°55’44.8”S, 47°45’04.8”E, 1507 m, Reeb &amp; Andriamiarisoa CR 11156* (PC 0763897, TAN). Malawi. Mulanje Mts., 15°53’32.1”S, 35°29’24”E, 2180 m, 22. VI.1991 O’Shea M 7327A-1* (E 00430482), Longton M 8299 A * (E 00430472). France. Réunion, Forêt de Belouve, Plateau Citrons, 21°31.5” S, 55°33.5” E, 1500 m, 18. VI.1996, Pócs 9647 A (EGR), 1150 m, 09.IX.2013, Bardat REU 1365* (PC 0146834). Uganda. Kabala, Bwindi National Park, 10°40’S, 02°E, 2050 m, 30. I.1996, Wiggington U 5137 C * (E 00430554), 2070 m, 26. I.1996, Porley U 97 A * (E 00430540). Zimbabwe. Melsetter District, Kasipiti, 1200 m, 28.XII.1964, Loveridge 1289 (EGR).</p> <p>DISTRIBUTION. — Widespread in temperate regions of the Northern Hemisphere, also in tropical America (e.g. Gradstein &amp; Costa 2003; Schäfer-Verwimp et al. 2013; Rabeau et al. 2017). Newly reported from Africa where the species has been found in St. Helena (Rabeau et al. 2017), Comores, Ethiopia, Madagascar, Malawi, Reunion and Uganda.</p> <p>HABITAT. — On dead wood and tree trunks in in sheltered environments in humid forests, also on humid rocks in streams and at the entrance of caves, between 900 and 2400 m.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous or paroicous).</p> <p>Thallus</p> <p>Green to brownish when fresh, yellow to bright brown in herbarium, flaccid, dull to glossy, to 20(-40) mm long, loosely to closely attached to the substrate with creeping to erect and sometimes recurved branches, mostly 2-3-bipinnate, sometimes palmate, branches alternate to subopposite, occasionally flagelliform, stolons rare.</p> <p>Main axis</p> <p>Not well-defined, variable in width, in cross section plano-convex to concave-convex, 5-6(-7) cells thick, margins acute, with narrow, 1-2 cells wide wings, epidermis cells 3-4× smaller than inner cells.</p> <p>Primary branches</p> <p>To 3 mm wide, 5-6(-7) cells thick, rarely overlapping, axis narrowed towards basis.</p> <p>Ultimate branches</p> <p>Tongue-shaped, oblong and dense, often fan-like arranged, straight and somewhat widened to the apex, to 3.5 mm long and 1.2 mm wide, in cross section concave-convex to plano-convex, winged, wings 1-4 cells wide, narrower than the midrib, apex enlarged, broad.</p> <p>Mucilage papillae</p> <p>Apical and in two ventral rows, persistent.</p> <p>Oil bodies</p> <p>Present in nearly all cells, grayish to light brown, 1-2 and rounded in epidermis cells, 1-4(-5) and fusiform in inner cells.</p> <p>Sexual branches</p> <p>Solitary or sometimes grouped, sessile, occurring along all the axis, laterally inserted.</p> <p>Male branches</p> <p>Bordered by convex cells.</p> <p>Female branches</p> <p>Straight, not recurved, margin with more than two cells long scales and cilia, somewhat crisped, fimbriate.</p> <p>Calyptra</p> <p>With scattered thick-walled cells, umbo lacking.</p> <p>Gemmae</p> <p>Occasionally present, 2-celled.</p> <p>COMMENTS</p> <p>Riccardia chamedryfolia has not previously been recognized in Africa, probably due to confusion with the polymorphic R. longispica. A well-delimited cluster of R. chamedryfolia was recovered in the molecular analysis with new records from Guadeloupe, St. Helena and the mainland of tropical Africa, indicating a more widespread distribution of the species than was previously known (Rabeau et al. 2017). The species is mainly characterized by the winged, plano-convex to concave-convex axis and branches, the broad branch apices, the epidermis cells much smaller than inner thallus cells, and monoicy. Sterile plants may be confused with robust phenotypes of R. longispica and R. saccatiflora but R. longispica is normally dioicous whereas R. saccatiflora lacks an axial wing and is usually palmate, and has dark brown oil bodies.</p> </div>	https://treatment.plazi.org/id/03EF87AA652BFFC0CAE7E3DEFC14FB04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6529FFC7CA39E31FFE36F926.text	03EF87AA6529FFC7CA39E31FFE36F926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia corbieri (Reeb & Gradstein 2020) Reeb & Gradstein 2020	<div><p>Riccardia corbieri (Steph.) Reeb &amp; Gradst. comb. nov. Fig. 2 J-N</p> <p>Aneura corbieri Steph., Species Hepaticarum 6: 23, 1917. — Type:</p> <p>Madagascar, 1898, Leloutre 126a (G[G00045028]!).</p> <p>Aneura nudiflora Steph., Hedwigia 31: 199 (1892). — Riccardia nudiflora (Steph.) Grolle, Bryophytorum Bibliotheca 48: 130 (1995). — Type: Mauritius, Robillard s.n. (lecto- fide Reeb &amp; Bardat [2014], G[G0037569]!) syn. nov.</p> <p>FURTHER SPECIMENS EXAMINED. — Madagascar. Sava, Makirovana, 14°10’17.4”S, 49°57’07.5”E, 508 m, 07.V.2010, Reeb &amp; Pierrot CR295* (PC0763837A, TAN), CR297* (PC0763838A, TAN).</p> <p>Seychelles. Mahé, Pointe au sel, 139 m, 20.IX.2012, Senterre &amp; Labiche-Barreau 6310* (PC 0763853, SEY), Mare aux cochons, 562 m, Senterre &amp; Labiche-Barreau 6258* (PC 0763852, SEY), 18.II.2012, Beaver B 304* (PC 0763848, SEY).</p> <p>DISTRIBUTION. — Madagascar, Mauritius, Seychelles.</p> <p>HABITAT. — On rock or dead wood in humid evergreen forests, from 100 to 560 m.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous).</p> <p>Thallus</p> <p>Green to olive green when fresh, yellow green to bright brown in herbarium, fleshy but not leathery, usually dull (rarely glossy), mostly flat, to 20 mm long, main axis creeping and attached to the substrate, branches ascending to erect, mostly 1-2-pinnate, not palmate, branches alternate to subopposite, flagelliform branches absent, stolons often present on the main axis, especially near the base in young plants, ultimate ascending branches wider than creeping ones.</p> <p>Main axis</p> <p>Very irregularly defined, in cross section ellipsoid to biconvex, 5-6(-9) cells thick, margins rounded, not winged (exceptionally margin cells bulging outwards), epidermis cells 2-4× smaller than inner cells, all cell walls thickened in mature thalli.</p> <p>Primary branches</p> <p>Erect, to 15 mm long, rarely overlapping.</p> <p>Ultimate branches</p> <p>Tongue-shaped to triangular, often sinuose, 3-5 mm long, 0.8- 1.2 mm wide, cells usually in straight rows, rarely in diverging rows, in cross section plano-convex to concave-convex, with 4-5 cells wide wings, the wings narrower to as wide as the midrib, epidermis cells 2-3× smaller than inner cells, branch apex rounded to truncate to emarginate.</p> <p>Mucilage papillae</p> <p>In two ventral rows, persistent.</p> <p>Oil bodies</p> <p>Not seen.</p> <p>Male branches</p> <p>Along main axis and primary branches, solitary, sessile, with a narrow, one cell wide wing.</p> <p>Female branches</p> <p>Straight, laterally to ventro-laterally inserted, margins with up to eight cells long cilia and with broad wings as wide as or wider than the axis, the wings sometimes recurved and branch becoming cup-shaped.</p> <p>Calyptra</p> <p>Smooth, with a triangular umbo of aggregate cells.</p> <p>Gemmae</p> <p>Occasionally present, 2-celled.</p></div> 	https://treatment.plazi.org/id/03EF87AA6529FFC7CA39E31FFE36F926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652EFFC6CAD0E43AFE94F801.text	03EF87AA652EFFC6CAD0E43AFE94F801.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia fastigiata (Lehm.) Trevis.	<div><p>Riccardia fastigiata (Lehm.) Trevis.</p> <p>(Fig. 2 M-R)</p> <p>Jungermannia fastigiata Lehm., Linnaea 4: 370 (1829). — Aneura fastigiata (Lehm.) Lehm. &amp; Lindenb., Synopsis Hepaticarum 500 (1846). — Type: Cape Town, „Montis Teufelsberg et in vertice montis Tafelberg“, Ecklon s.n. (lecto-, S-B104928!; isolecto-, G[G0067556]!).</p> <p>? Aneura caespitans Steph., Bulletin de la Société Royal de Botanique de Belgique 30: 194 (1891) — Type: Réunion, Rodriguez s.n., ex hb. Renauld (holo-, BM[BM001167936]!). syn. nov.</p> <p>FURTHER SPECIMENS EXAMINED. — Réunion. Cirque de Salazie, plaine des Merles, 21°02’50”S, 55°28’08”E, 1800 m, 24.VIII.1994, Vojko 9422AP*, 9422V* (EGR); Forêt de Belouve, plateau des Cochons, 21°02’00”S, 55°33’00”E, 1430 m, 02.VII.1996, Orban 9615CB* (EGR, PC0146833); Piton de la Fournaise. 21°18’00”S 55°42’07”E. 1390-2100 m. 30.VI.1996. Pócs 9613CD* (EGR).</p> <p>South AFrica. Pietermaritzburg, 914 m, XII.1914, Sim 7536, 7538 (PRE); Knysna, 01.IV.1929, Bottomley 3671 (PRE); eastern slope of Table Mountain, 300 m, 19.III.1995, Arts 3318CD (PRE).</p> <p>DISTRIBUTION. — South Africa, Réunion.</p> <p>HABITAT. — Among Sphagnum and on rocks in streams, between 300 and 2100 m.</p> <p>DESCRIPTION</p> <p>Monoicous (Dioicous).</p> <p>Thallus</p> <p>Gray green, olive to deep green when fresh, becoming darker colored in herbarium, flaccid, strap-shaped, loosely or not attached to the substrate, to 25 mm long, 1-3-pinnate, not palmate, branches alternate, flagelliform branches sometimes present, stolons not seen.</p> <p>Main axis</p> <p>Well-defined, to 1 mm wide, in cross section plano-convex to concave-convex, usually with mycorrhiza-like fungi in the ventral part, 4-6 cells thick, margin rounded to acute, with a translucent band of protruding cells but not clearly winged, epidermis cells c. 2-3× smaller than inner cells to equal in size, regularly quadrate, radial walls of the epidermis cells often thickened (in material from Réunion).</p> <p>Ultimate branches</p> <p>Tongue-shaped to oblong, mostly 1-2(-4) mm long and 200- 400(-500) µm wide, at an angle of 60-80° with the axis, cross section similar to axis, margins with protruding cells but not clearly winged, branch apex often broadened, rounded or cucullate, not dissected.</p> <p>Mucilage papillae</p> <p>In two rows, clavate.</p> <p>Oil bodies</p> <p>Not seen.</p> <p>Male branches</p> <p>Short, straight or reflexed, often shortly stalked, up to 900 µm, max. of 7-8 antheridial pairs, regularly bordered by a plane or undulate, 2-3 cells wide wing.</p> <p>Female branches</p> <p>Mostly solitary, laterally inserted, present along the main axis and along primary branches, sessile or shortly stalked, recurved when immature, 300-800 µm long, with wing-like scales, the scales 2-4 cells wide; archegonia in up to five pairs.</p> <p>Calyptra</p> <p>With an umbo of aggregated cells and pachydermal scattered cells.</p> <p>Gemmae</p> <p>Occasionally present, 2-celled.</p> <p>COMMENTS</p> <p>Riccardia fastigiata resembles R. chamedryfolia but the latter species is autoicous and has clearly winged branches. A noteworthy feature of the specimens of R. fastigiata from Réunion are the thickened radial walls of the epidermis cells. In the material from South Africa, including the type, epidermal thickenings are usually lacking. As the plants from the two regions are identical in all other respects, we treat them as conspecific. Fresh collections are needed to verify this (DNA of the South African plants could not be amplified).</p> <p>Aneura caespitans Steph. is probably a synonym of Riccardia fastigiata. The type material (Réunion, leg. Rodriguez) is sterile and poorly developed. Reeb &amp; Bardat (2014) did not see the type (it is lacking in G) and treated A. caespitans as a dubious name. They noted that the description of the species in Species Hepaticarum (Stephani, 1898-1924) is heterogeneous and includes characters of two different species, including the type and a specimen from Cameroon (Dusén 251) not mentioned in the protologue. The confusion was worsened by Stephani’s illustration of A. caespitans in Icones Ineditae (Stephani 1985), which was based on the specimen from Cameroon and not on the type. We have now been able to examine both specimens (kept in BM) and found that the type belongs probably to R. fastigiata, as mentioned, whereas the plant from Cameroon is R. saccatiflora. Already E.W. Jones (in sched.) noted that the type of A. caespitans and the specimen from Cameroon are different taxa. Since no duplicates of the type are present in G or other herbaria, the collection in BM may be considered the holotype.</p> </div>	https://treatment.plazi.org/id/03EF87AA652EFFC6CAD0E43AFE94F801	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652FFFC4CAF9E6FBFE36FE1A.text	03EF87AA652FFFC4CAF9E6FBFE36FE1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia gasparii Reeb & Gradstein 2020	<div><p>Riccardia gasparii Reeb &amp; Gradst. sp. nov.</p> <p>(Fig. 3 A-E)</p> <p>DIAGNOSIS. — Dioicous. Thallus very fleshy, unwinged. Axis circular to biconvex, branches with rounded margins. Male branches stalked.</p> <p>TYPE. — Madagascar. Alaotra-Mangoro, Vohimana Reserve, Grande Cascade, 18°55’43.6”S 49°29’56.6”E, 849 m, 30. VI.2012, Reeb MTV1246* (holo-, PC [PC 0771050]; iso-, PC [PC 0763876], TAN).</p> <p>ETYMOLOGY. — Dedicated to Gaspard Reeb-Leurent, born 21/07/2015.</p> <p>FURTHER SPECIMENS EXAMINED. — Madagascar. Alaotra-Mangoro, Vohimana Reserve, Grande Cascade, 18°55’43.6”S, 49°29’56.6”E, 849 m, 30.VI.2012, Reeb MTV1257 (PC, TAN); Maromizaha Reserve, 18°58’53.02”S, 48°27’50.1”E, 1002 m, 08.VII.2012, Reeb MTM12124 * (PC, TAN), 18°58’53.10”S, 48°27’51”E, 921 m, 08.VII.2012, Reeb MTM12127A *, MTM12133 *(PC0763885), MTM12134 * (PC0763884, TAN); Analanjirofo, Mananara Nord, E slope of Mahavoho hills, 16°27’S, 49°46.9’47.5”E, 220-300 m, 14.VIII.1998, Pócs &amp; Szabo 9878/W (EGR).</p> <p>DISTRIBUTION. — Alaotra-Mangoro and Analanjirofo regions of Madagascar.</p> <p>HABITAT. — On rocks or boulders sometimes covered with a humus layer, in humid evergreen lowland and montane forests, between about 200 and 1000 m.</p> <p>DESCRIPTION</p> <p>Dioicous.</p> <p>Thallus</p> <p>Dull olive green to green when fresh, green to yellow brown in herbarium, very fleshy, rigid but not leathery (easily breaking when manipulated), to 2 cm long, 1-3-pinnate, not palmate, branches alternate to subopposite, reiterations sometimes present, stolons common, branches mostly wider than the main axis.</p> <p>Main axis</p> <p>Irregularly defined to well-defined in mature thalli, width variable, in cross section circular to ellipsoid at the base, circular to biconvex in the middle, 7-9(-16) cells thick, margins obtuse to rounded, not winged, epidermis cells 3-4× smaller than inner cells.</p> <p>Primary branches</p> <p>To 7 mm long, often overlapping.</p> <p>Ultimate branches</p> <p>Tongue-shaped to oblong, often sinuose (horizontally curved), to 6 mm long and 1.8 mm wide, in cross section plano-convex, 5-6(-7) cells thick, margins obtuse, not winged, epidermis cells 2-3× smaller than inner cells, branch apex rounded.</p> <p>Mucilage papillae</p> <p>Apical, not persistent.</p> <p>Oil bodies</p> <p>Not seen.</p> <p>Male branches</p> <p>On the main axis, solitary, stalked, with a scale-like wing narrower than the axis, the wing crenulate by larger cells.</p> <p>Female branches</p> <p>Laterally inserted, straight, bordered by a ciliate, scale-like wing.</p> <p>Gemmae</p> <p>Occasionally present, 2-celled.</p></div> 	https://treatment.plazi.org/id/03EF87AA652FFFC4CAF9E6FBFE36FE1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652DFFC4C9B3E578FAB0FB24.text	03EF87AA652DFFC4C9B3E578FAB0FB24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia inconspicua (Steph.) Reeb & Bardat	<div><p>Riccardia inconspicua (Steph.) Reeb &amp; Bardat</p> <p>(Fig. 3 F-K)</p> <p>Aneura inconspicua Steph., Hedwigia 32: 23 (1893). — Type: Cameroon, Junger 5 (lecto-, here designated, G[G00067564]!; isolecto-, S-B20523!).</p> <p>Riccardia tenuicostata Schiffn., Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 67: 166 (1898). — Aneura tenuicostata (Schiffn.) Steph., Species Hepaticarum 1: 245 (1899). — Type: Singapore (Furuki 1994).</p> <p>Aneura exigua Steph., Species Hepaticarum 6: 25 (1917). — Type: Seychelles (Reeb &amp; Bardat 2014).</p> <p>FURTHER SPECIMENS EXAMINED. — Cameroon. Ekundu N’dene, 16.III.1892, Dusén 841 (G00264075, PC0101721). Cameroon: Kumba, Banga, S. Bakundu Forest Reserve, 5°5’50”N, 9°18’35”E, 15.III.1948, Brenan &amp; Jones 269 (E).</p> <p>Madagascar. Alaotra-Mangoro, Maromizaha Reserve, 18°58’45”S, 48°27’51.8”E, 1011 m, 06.VII.2012, Reeb MTM1295* (PC 0763882, TAN).</p> <p>Seychelles. Mahé, Congo Rouge, 46°27’40.68’’S 55°26’12.768’’E, 820 m, 16.IV.2012 Senterre &amp; Lariche SEY 6265* (PC, SEY).</p> <p>Tanzania. Morogoro, Uluguru Mts., Kilangala, SE of Bundunki, 1750-1950 m, 21.IX.1971, Pócs &amp; Mwanjabe 6464 BP (EGR).</p> <p>DISTRIBUTION. — Paleotropical: tropical Africa (Cameroon, Madagascar, Seychelles, Tanzania), tropical Asia, Australia (Queensland).</p> <p>HABITAT. — On rotten wood in humid lowland and montane forests, between 120 and 1950 m.</p> <p>DESCRIPTION</p> <p>Dioicous.</p> <p>Thallus</p> <p>Green to pale-green when fresh, yellow to clear brown in herbarium, flaccid and fragile, glossy, mostly flat, very small, to 10 mm long, main axis creeping and attached to the substrate, branches erect, 1-2-pinnate, not palmate, branches alternate to subopposite, stolons rare, thallus margins with mucilage papillae.</p> <p>Main axis</p> <p>Irregularly to clearly defined, 150-300(-350) µm wide, in cross section plano-convex, (3-)4(-5) cells thick, margins acute, with a 1-2 cells wide wing.</p> <p>Ultimate branches</p> <p>Tongue-shaped to oblong, up to 1.5 mm long, in cross section 3-4 cells thick, lunate to concavo-convex, with 3-4 cells wide wings, the wings as wide as or wider than the midrib.</p> <p>Mucilage papillae</p> <p>On all branch margins (including sexual branches), especially on ultimate branches, simple to 2-3-furcate, to 100 µm long.</p> <p>Oil bodies</p> <p>Not seen.</p> <p>Male branches</p> <p>Solitary, straight to recurved, sessile or stalked, to 1.8 mm long, with up to 20 antheridial pairs, the margin with a scale-like wing equal to sometimes wider than the axis.</p> <p>Female branches</p> <p>Inserted laterally, straight, solitary or grouped, with up to 22 archegonial pairs, bordered by numerous, 4-5 cells long cilia, the cilia to 250 µm long, exceeding the width of the axis.</p> <p>Calyptra</p> <p>625-800 µm long, with an umbo of aggregate cells.</p> <p>Gemmae</p> <p>Abundant, present in all collections seen, 2-celled.</p></div> 	https://treatment.plazi.org/id/03EF87AA652DFFC4C9B3E578FAB0FB24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA652DFFDACAD9E09DFC0CFEFB.text	03EF87AA652DFFDACAD9E09DFC0CFEFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia longispica (Steph.) Pearson	<div><p>Riccardia longispica (Steph.) Pearson</p> <p>(Fig. 3 L-W)</p> <p>Aneura longispica Steph., Botanical Gazette 15: 281 (1890). —</p> <p>Type: Réunion, Boivin s.n. (G [G 00067568]!).</p> <p>Aneura congoensis Steph., Species Hepaticarum 6: 23 (1917). — Type: Congo (Reeb &amp; Bardat 2014).</p> <p>Aneura grosselimbata Steph., Species Hepaticarum 6: 29 (1917). — Type: Tanzania (Reeb &amp; Bardat 2014 as R. limbata).</p> <p>Aneura holstii Steph., Species Hepaticarum 6: 31 (1917). — Riccardia holstii (Steph.) E.W.Jones, Transactions of the British Bryological Society 3: 78 (1956). — Type: Tanzania (Reeb &amp; Bardat 2014 as R. limbata).</p> <p>Aneura lepervanchei Steph., Species Hepaticarum 1: 255 (1899). — Type: Réunion (Reeb &amp; Bardat 2014).</p> <p>Aneura limbat a Steph., Hedwigia 30: 204 (1891). — Riccardia limbata (Steph.) E.W.Jones, Transactions of the British Bryological Society 3: 79 (1956). — Type: Cameroon, Lower Bomania, 1890, Dusén 33 (lecto-, here designated, G[G00067566]!) syn. nov.</p> <p>Aneura piliflora Steph., Species Hepaticarum 6: 38 (1917). — Type: Madagascar (Jones 1956).</p> <p>Aneura reticulata Steph., Hedwigia 30: 204 (1891). — Riccardia reticulata (Steph.) S.W. Arnell, Botaniska Notiser 1952: 152 (1952). — Type: Cameroon (Jones 1956 as R. limbata).</p> <p>Aneura erosa Steph., Hedwigia 30: 269 (1891). — Riccardia erosa (Steph.) E.W.Jones, Transactions of the British Bryological Society 3: 83 (1956). — Type: Sao Tomé, 1989, Quintas 7 (G[G0067552]!) syn. nov.</p> <p>Aneura tenera Steph., Species Hepaticarum 6: 43 (1917). — Type: Tanzania (Reeb &amp; Bardat 2014) syn. nov.</p> <p>? Riccardia kilimanjarica S.W.Arnell., Svensk Botanisk Tidskrift 53: 543 (1959). — Type: Tanzania (Reeb &amp; Bardat 2014).</p> <p>SELECTED FURTHER SPECIMENS EXAMINED. — Central AFrica Republic. Plateau de Kagadouba, 480 m, Assel 729 (E00018738, EGR).</p> <p>Comores. Karthala, 2010, Bardat Karp1q2li2 * (PC 0763850).</p> <p>Congo. Kimpese, road Lombo Fuese, 5°23’52”S, 14°28’12”E, 640 m, 13. I.2013, Vanderpoorten 13111*.</p> <p>Equatorial Guinea. Evinayong, Mt. Alen, 950 m. 25.VII.1994, Heras VIT 398/94*.</p> <p>Ethiopia. Bonga, Gera Beki forest, 37°80’82.07”N, 19°96’66”E, 1881 m, 12.XII.2006, Hylander KH 5488*, KH 5491* (ETH, PC 0763858).</p> <p>Gabon. Ogoué-Ivindo, Réserve intégrale d’Ipassa, 00°30’N, 12°48’00”E, 510 m, 01.IV.06, Vanderpoorten GAB 1396*, 03°04’8”N, 12°46’12”E, 11.IV.06, Vanderpoorten GAB 1398*.</p> <p>Ghana. Central region, Pra-Suhien Forest Reserve, 11.II.1971, Jones 1339 A (E 00018734, G 00264219).</p> <p>Madagascar. Alaotra-Mangoro, Vohimana Reserve, 18°55’24.2”S, 48°30’21.1”E, 860 m, 14.IV.2010, Reeb, Andriamanantena, &amp; Bidault CRAE 74 * (PC 0724433 B), 18°55’43.6”S, 48°29’56.6”E, 849 m, 15.IV.2010, Reeb, Andriamanantena, &amp; Bidault CRAE 84 (PC 0724365), CRAE 88 * (PC 0763869), 18°55’13.1”S, 48°30’57.1”E, 789 m, 29. VI.2012, Reeb MTV1225 * (PC 0763889), MTV1226 (PC), 18°55’45.7”S, 48°29’59.3”E, 918 m, 30. VI.2012, Reeb MTV1234 (PC 0763887), MTV1237 *(PC 0763888), MTV1238 (PC 0763892); Alaotra-Mangoro, Maromizaha Réserve, 18°59’S, 48°28’E, 1084- 1095 m, 07.VII.2012, Reeb MTM12103 * (PC 0763854), MTM12106 * (PC 0763855); Antsiranana, Manongarivo Special Reserve, 13°59’S, 48°26’E, 1300-1350 m, Geissler 19708/5 (G); Sava, Parc du Marojejy, 29.XI.2009, Bardat JBMAD 104 * (PC 0763861), Makirovana, 14°10’29.3”S, 49°57’07”E, 695 m, 06. V.2010, Reeb CR 276 (PC 0763841), CR 288 (PC 0763836 A), 14°17’10.04”S, 49°57’07.5”E, 508 m, 07. V.2010, Reeb CR 304 * (PC 0763833).</p> <p>Malawi. Mulanje Mt., 15°53’21”S, 35°39’24”E. 2180 m, 25. VI.91, Longton M 8302 A * (E), 15°55’01”S, 35°36’04”E, 2000 m, 24. VI.1991, Kungu M 3199 A * (E-00430533).</p> <p>Mauritius. Mt. Le Pouce, 20°13’5”S, 57°31’20”E, 750-811 m, 20.III.1995, Pócs 9537 H * (EGR).</p> <p>Réunion. Cirque de Salazie, 21°2’50”S, 55°28’8”E, 1899 m, 24.VIII.1994, Szabo 9422 J * (EGR); Plaine des Palmistes, 21°6’33”S, 55°39’E, 900 m, 30.VIII.1994, Vojko 9435 L * (EGR); Forêt de Belouve, 21°2’S 55°33.5’E, 1440 m, 16.VII.1996, Konya 9644/D * (G 00264189, EGR); Piton de la Fournaise, 21°18’08”S, 55°44’00”E, 1200 m, Szabo 9609/ CP * (EGR), 21°18’05”S, 55°42’05”E, 1000- 1390 m, 30. VI.1996, Pócs 9612/CD (EGR). Plaine des fougères, 1325 m, 19.IV2011, Bardat PF.2 (PC 0763860).</p> <p>South AFrica. Port Shepstone, Iberzana, 10 m, 23.VIII.1918, Eyles 1407 (EGR).</p> <p>Tanzania. Trail Kitoto to Meru crater, 2500 m, 11.VII.1968, Sharp et al. P 507 (E 00018742, EGR); Mt. Kilimanjaro, trail Mweka to Kibo Peak, 2400-2800 m, 29.VII.1968, Sharp et al. 7632 (E).</p> <p>Uganda. Rukungiri, 1°00’00”S, 29°37’00”E, 1650 m, 07.II.1997, Wiggington U 8318 C *, U 8321 A, U 8324 A *.</p> <p>PHENOTYPE EROSA. — Gabon. Province Ogououé-Ivindo, chutes de Kongou sur l’Ivindo 0°17’24”N, 12°25’18”E, 09.IV.2006, Vanderpoorten GAB 1435*.</p> <p>Madagascar. Alaotra-Mangoro, Vohibola Reserve, 18°35’16.1”S, 49°14’28.8”E, 20.IV.2010, Reeb, Andriamanantena &amp; Bidault CRAE 193 (PC 0763859); Atsimo-Andrefana, massif du Makay, 21°34’05.9”S, 45°04’33.7”E, 303 m, 14. I.2011, Reeb CR 1126 * (PC), 21°34’09.8”S, 45°04’35.4”E, 290 m, 16. I.2011, Reeb CR 1169 * (PC 0763891), 21°38’53”S, 44°59’52”E, 189 m, 23. I.2011, Reeb CR 11126 * (PC 0763894), CR 11127 * (PC 0763895).</p> <p>Tanzania. Usambara Mts., Amanani, 950 m, 17. VI.1970, Rasmussen &amp; Esbessen B 27 (EGR).</p> <p>Uganda. Nyakafunjo Nat. Reserve, 24. I.1997, 1075 m, Porley U 416 A (E), 1060 m, Hodgetts U 4346 A (E); Kabale Bwindi Nat. Park, 01°23’7”S, 29°46’50”E, 2250 m, 25. I.1997, Hodgetts U 5009 A * (E 00430552), 2415 m, 25. I.1996, Porley U 35 B * (E 00430508).</p> <p>DISTRIBUTION. — Riccardia longispica is one of the most common African Riccardia species, recorded from almost all West and East African countries and Indian Ocean Islands (Wigginton 2018).</p> <p>HABITAT. — The large phenotype of R. longispica is mainly found on dead wood, sometimes on humid rock along streams or near waterfalls, or on dense bryophyte mats (in Réunion), between about 500 and 2500 m. The small erosa phenotype seems to have the same geographical distribution as the large phenotype but is mainly found on shaded lateritic soil, occasionally on rotten wood, and occurs at slightly lower elevation (200 m) on Madagascar.</p> <p>DESCRIPTION</p> <p>Dioicous, occasionally monoicous (R. limbata, R. holstii).</p> <p>Thallus</p> <p>Green to olive green when fresh, yellow to dark brown or green in herbarium, soft to fleshy, (10-) 20-30 mm long, very variable when dry, crisped (in phenotype erosa) or flat, ultimate branches usually slightly recurved, main and secondary axes closely attached to the substrate from which arise the ultimate branches, branching 1-2(-3)-pinnate, sometimes palmate on young thalli, branches alternate or subopposite, reiterations sometimes present, stolons mainly seen in young thalli.</p> <p>Main axis</p> <p>Irregularly defined in young plants, well-defined in mature thalli, (600-)800-1200(-1400) µm wide, in cross section ellipsoid to biconvex at the base, plano-convex in the middle, 5-7(-8) cells high, margins acute, with narrow, 1-2 cells wide wings, epidermis cells at least 4-5× smaller than inner cells.</p> <p>Primary branches</p> <p>To 700 µm wide, occasionally overlapping.</p> <p>Ultimate branches</p> <p>Tongue-shaped to oblong, straight in well-developed thalli (stocky and fan-shaped in young thalli), 2.5-4.2 mm long, (0.4-)0.5-0.8(-1.0) mm wide, in cross section plano-convex to slightly concave-convex, 3-4(-5) cells thick, wings 2-4 cells wide, narrower than the midrib, epidermis cells usually in straight rows, rarely in diverging rows (obliquely to the axis), branch apex rounded to truncate with sometimes overlapping lateral edges.</p> <p>Mucilage papillae</p> <p>Ventral (in two rows) and apical, persistent.</p> <p>Oil bodies</p> <p>Present in all cells, spherical to ellipsoid to bean-shaped, finely granular, 1-2 per cell.</p> <p>Male branches</p> <p>Solitary, rarely grouped by two, along axis and branches, straight except when young, to 2 mm long, wing equal to narrower than the axis, scale-like, antheria in 6-12(-20) pairs.</p> <p>Female branches</p> <p>Ventro-laterally inserted, bordered by a scale-like, 1-2 cells wide wing, wing narrower than the axis, archegonia in 5-12 pairs.</p> <p>Calyptra</p> <p>To 2 mm long, terminating in a crown-like umbo and very scattered thick-walled cells armed with striations.</p> <p>Gemmae</p> <p>Occasionally present, especially at branch junctions, produced on deformed thallus margins, 2-celled.</p> <p>eROSA PHENOTYPE (Fig. 2 R-W)</p> <p>Plants</p> <p>Usually growing on lateritic soil.</p> <p>Thallus</p> <p>Very small, to 5-6 mm long only, 1-2-pinnate without reiterations, crisped when dry.</p> <p>Main axis</p> <p>180-320 µm wide, in cross section circular ellipsoid at the base to plano-convex in middle, 3-4 cells high, with 1-2 cells wide wings, epidermis cells 2-3× smaller than inner cells.</p> <p>Ultimate branches</p> <p>280-2000 µm long, 70-330 µm wide, linear to oblong, sometimes tongue-shaped, in cross section plano-convex, 3-4 cells thick, wings 3-4 cells wide, as wide as or wider than the midrib, wing cells in straight rows.</p> <p>Sexual branches</p> <p>Equal or longer than ultimate branches.</p> <p>Male branches sessile</p> <p>Or shortly stalked.</p></div> 	https://treatment.plazi.org/id/03EF87AA652DFFDACAD9E09DFC0CFEFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6533FFD8CAFCE0DDFBE8FD1A.text	03EF87AA6533FFD8CAFCE0DDFBE8FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia martinii Reeb & Gradstein 2020	<div><p>Riccardia martinii Reeb &amp; Gradst. sp. nov.</p> <p>(Fig. 4 A-I)</p> <p>DIAGNOSIS. — Dry thallus with white margins and with an orange-colored network of thick-walled dorsal epidermis cells.</p> <p>TYPE. — Madagascar. Alaotra-Mangoro, Zahamena Nat. Park, 17°38’22”S, 48°38’47”E 1247 m, 26.XII.2013, Reeb &amp; Andriamanantena CR 13 Z 9* (holo-, PC [PC 0763851]).</p> <p>FURTHER SPECIMENS EXAMINED. — Réunion. St Denis, 1500 m, 11.X.1962. Een R 039 (S-B12531); Piton de la Fournaise, 21°19’00”S, 55°42’00”E, 800-870 m, 23.II.1995, Pócs 9501 H * (G 00264150, EGR), 21°18.5’S, 55°42’E, 800-1080 m, Pócs 9602/ CU * (G 00264153, EGR), 21°18’0S, 55°42.7’E, 1800 m, 30. VI.1996, Pócs 9612/ CC, 9 613/ CC *, 9613/CW (EGR); Plaine des Caffres, 21°09’S, 55°33’E, 1850 m, 22.VII.1996, Vojko 9660 / CS (G 00264121, EGR); Forêt de Bélouve, 21°4’S, 55°31.6’E, 1610-2000 m, Pócs 9618/ CU (EGR).</p> <p>Madagascar. Alaotra-Mangoro, Zahamena Nat. Park, 17°38’22”S, 48°38’47”E 1247 m, 26.XII.2013, Reeb &amp; Andriamanantena CR 13 Z 9 * (PC 0763856), CR 13 Z 25 * (PC 0763863); Vohimana Reserve, 18°55’43.6”S, 48°29’56.6”E, 849 m, 15.IV.2010, Reeb, Andriamanantena &amp; Bidault; CRAE 106 (PC); Sava, Marojejy Nat. Park, X.2010, Bardat JBMAD 78* (PC 0763890).</p> <p>Tanzania. Uluguru Mts., Kinole Village, 1480-1530 m, 05.II.1973, Pócs &amp; Lubgwecha 6877/ K (G 00264154, EGR), Kibale Forest, 1730 m, 19.X.1971, Pócs 6477/ M (G 00264161, EGR)., Bondwa, 2125 m, 13.XII.1972, Pócs &amp; Crosby 6947/ K (EGR), ridge and Lupanga peak, 2100-2400 m, 21.XI.1969, Pócs &amp; Csontos 6072/ BA (EGR).</p> <p>DISTRIBUTION. — Tanzania, Madagascar and Réunion.</p> <p>HABITAT. — On dead wood, bark on peaty soils covered a dense bryophyte layer, occurring between 400 and 4000 m but mainly found at mid elevations (800-1500 m).</p> <p>ETYMOLOGY. — Dedicated to Martin Reeb-Leurent, born 25/05/2011.</p> <p>DESCRIPTION.</p> <p>Monoicous (autoicous) or dioicous.</p> <p>Thallus</p> <p>Fleshy to leathery,rigid, green, glossy,green to dark brown or black in herbarium, to 22 mm long, when dry crisped or with recurved branches and with white margins, main axis closely attached to the substrate or to other thalli, 1-3-pinnate, sometimes palmate in young thalli, reiterations and stolons frequently present.</p> <p>Main axis</p> <p>Irregularly defined, 400-500 µm wide, with mycorrhiza-like fungi on the ventral side (cross section), in cross section ellipsoid to biconvex, rarely plano-convex, 7-10(-11) cells thick, with ventral subepidermis, margin acute with one bulging cell, not winged, epidermis cells more than 4-5× smaller than inner cells, radial walls of the dorsal epidermis cells usually thickened and orange-colored in dry material.</p> <p>Primary branches</p> <p>4-12 mm long.</p> <p>Ultimate branches</p> <p>Oblong, stocky to tongue-shaped, margins white or clearer, (0.2-) 1.5-3 mm long, 0.3-0.4-0.7(-1.0) mm wide, in cross section lunate to concavo-convex, 4-7 cells thick, margins acute or with a 1-2 cells wide wing, epidermis cells equal or 2-4× smaller than inner cells, branch apex emarginate, dissected or retuse, without overlapping edges, young thalli with numerous multi-apical areas.</p> <p>Mucilage papillae</p> <p>Present, apical and in two ventral rows, 70(-100) µm long, exceeding the apex.</p> <p>Male branches</p> <p>Solitary or often grouped, sessile to or stalked, sometimes prolonged by vegetative growth, curved and reflexed with age, to 1 mm long, narrowly winged. the wing narrower than the axis, margins crenate by larger cells, antheridia in up to 20 pairs.</p> <p>Female branches</p> <p>Very short when young (seemingly immersed in the vegetative tissue), ventrally to ventro-laterally inserted, narrowly winged, the wing ornamented by short, straight cilia.</p> <p>Calyptra</p> <p>Rugose by scattered thick-walled, bulging cells.</p> <p>Gemmae</p> <p>Occasionally present.</p></div> 	https://treatment.plazi.org/id/03EF87AA6533FFD8CAFCE0DDFBE8FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6531FFDFCAB2E3BEFF17F9A7.text	03EF87AA6531FFDFCAB2E3BEFF17F9A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia multifida	<div><p>Riccardia multifida (L.) Gray (Illustration: Perold 2001b)</p> <p>Jungermannia multifida L., Species Plantarum 1: 1136 (1753). — Aneura multifida (L.) Dumort., Commentationes Botanicae 112 (1822). — Type: England (Grolle 1976).</p> <p>SPECIMEN EXAMINED. — South AFrica. Diepwalle Forest Reserve, 07.X.2000, Perold 4449 (PRE-P581351.0).</p> <p>DISTRIBUTION. — Riccardia multifida is widespread in temperate regions of the Northern Hemisphere; in addition, the species has been recorded from Brazil (Rio de Janeiro; Gradstein &amp; Costa 2003) and from South Africa where it is known from several localities (Perold 2001b). The occurrence of R. multifida in South African was confirmed by molecular analysis (Reeb et al. 2018). The species is not yet known from East Africa and the Indian Ocean Islands, but may be expected there and is therefore included in this treatment.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous).</p> <p>Thallus</p> <p>Glossy green when fresh, green to brown in herbarium, fleshy, mostly flat, in compact layers, to 30 mm long, 1-3(-4)-pinnate, with regulary pinnate secondary branches, branches alternate to subopposite, reiterations and flagelliform branches present, stolons absent.</p> <p>Main axis</p> <p>Well-defined, in cross section ellipsoid, biconvex(to plano-convex) in the middle, 4-5(-8) cells thick, with a 2-3 cells wide wing, epidermis cells 2-4 × smaller than inner cells.</p> <p>Primary branches</p> <p>Up to 5 mm, regularly branched, branching at an angle of c. 45⁰ with the axis, rarely overlapping.</p> <p>Ultimate branches</p> <p>Usually narrowly linear, occasionally tongue-shaped to oblong, straight, to 1.6 mm long, 0.5-0.6 mm wide, in cross section plano-convex to biconvex, 3-4 cells thick, winged, wings 3-4 cells wide, narrower than the midrib, branch apex rounded to truncate, margins crenulate by bulging cells.</p> <p>Mucilage papillae</p> <p>Apical and in two ventral rows, persistent.</p> <p>Oil bodies</p> <p>Absent or rare in epidermis cells, present in inner cells, 1(-2) per cell, spherical to ellipsoidal, light brown, finely granular.</p> <p>Male branches</p> <p>Along the axis, solitary to grouped, 600-750 µm long, margins crenate by larger, bulging cells.</p> <p>Female branches</p> <p>Laterally inserted, to 450 µm long, margins with mostly straight cilia or scales.</p> <p>Calyptra</p> <p>Up to 3 mm long, ornamented by loosely aggregated umbo cells and scattered thick-walled cells.</p> <p>Gemmae</p> <p>Not seen.</p></div> 	https://treatment.plazi.org/id/03EF87AA6531FFDFCAB2E3BEFF17F9A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6536FFDEC987E0BEFF17FDDB.text	03EF87AA6536FFDEC987E0BEFF17FDDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia ramosissima (Steph.) Grolle	<div><p>Riccardia ramosissima (Steph.) Grolle</p> <p>(Fig. 4 J-O)</p> <p>Aneura ramosissima Steph., Bulletin de la Societé Royal de Botanique de Belgique 30: 196 (1891) — Type: Réunion, Rodriguez s.n. (G-00045033).</p> <p>Aneura compacta Steph., Hedwigia 32: 19 (1893). — Riccardia compacta (Steph.) S.W.Arnell, Botaniska Notiser 1952: 141 (1952). — Type: South Africa, Cape province, Jelinek s.n. (lecto-, here designated, G[G00067549]!; isolecto-, PC[PC0101714]!, S[SB20511]!) syn. nov. FURTHER SPECIMEN EXAMINED. — Ethiopia. Bale Montane, above Rira W of Aduka, 06°43’N, 39°42’E, 3320 m, 13.II.1990, Miehe 2541 (EGR).</p> <p>Madagascar. Alaotra-Mangoro, Zahamena Reserve, highest part of the river above base camp, 17°38’22.7”S, 48°38’45.3”E, 1294 m, 01. I.2014, Reeb CR 13 Z 51* (PC 0763864).</p> <p>South AFrica. Mpulalanga province, Mriepskop Mt., 1550 m, 18. VI.1969, Vorster 828 B (E-00430437); Cape province, Apollo Peak, 1219 m, 31.III.1956, Esterhuysen &amp; Arnell 4361 (PRE-0501890).</p> <p>Tanzania. Mt. Kilimanjaro, 1972, Pócs 6718, 6789/ B (EGR, G 00264060), ibid., 1984, Pócs 6981/ A (EGR, G 00264135, G 00264136).</p> <p>DISTRIBUTION. — Ethiopia, Tanzania, Uganda, South Africa, Madagascar, Réunion.</p> <p>HABITAT. — On rotten wood, moist rock and humic soil in forests and along streams, between 1200 and 3800 m.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous) or dioicous.</p> <p>Thallus</p> <p>Dull or glossy yellow-green to green when fresh, dark brown to black in herbarium, leathery, rigid, growing in numerous layers, to 15 mm long, 1-2(-3)-pinnate, corymbose, not palmate, branches alternate to subopposite, reiterations frequently present in mature thalli.</p> <p>Main axis usually</p> <p>Ill-defined due to the densely corymbose ramification, in cross section rounded to ellipsoid at the base, 5-10 cells thick, 300-400 µm wide, in the middle ellipsoid to biconvex (rarely plano-convex), 300-500 µm wide, margins obtuse by bulging cells, not winged, epidermis cells c. 2-4 × smaller than inner cells at least on the ventral side of the axis, occasionally equal is size.</p> <p>Primary branches</p> <p>And reiterations to 7 mm long.</p> <p>Ultimate branches</p> <p>Linear to triangular, sometimes sinuous (horizontally curved), 0.5-3(-5) mm long, 0.2-0.3 mm wide, in cross section plano-convex to biconvex, 3-6 cells thick, margins with 1-3 cells wide wings, outer margins often crenate by bulging cells, epidermis cells 2-4 × smaller than inner cells, branch apex rounded to acute, emarginate to weakly dissected.</p> <p>Mucilage papillae</p> <p>Apical and in two ventral rows.</p> <p>Oil bodies</p> <p>Ellipsoid, finely granular, 2-4 per cell.</p> <p>Male branches</p> <p>Solitary, along the axis, shortly stalked, sometimes prolonged by vegetative growth, straight, 400-450 µm long, with a narrow, 1 cell wide wing.</p> <p>Female branches</p> <p>Along the axis and primary branches, laterally inserted, to 500 µm long, margin scaly and with hair-like, straight cilia, length of cilia and scales exceeding the width of the axis.</p> <p>Calyptra</p> <p>To 2.5 mm long, terminating in a conical umbo of densely aggregate cells.</p> <p>Gemmae</p> <p>Not seen.</p></div> 	https://treatment.plazi.org/id/03EF87AA6536FFDEC987E0BEFF17FDDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6537FFDDC99BE25FFF17FCFA.text	03EF87AA6537FFDDC99BE25FFF17FCFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia saccatiflora (Steph.) S. W. Arnell	<div><p>Riccardia saccatiflora (Steph.) S.W.Arnell</p> <p>(Fig. 4 P-T)</p> <p>Aneura saccatiflora Steph., Botanical Gazette 15: 282 (1890). — Type: Réunion, Rodriguez 186 (lecto-, fide Reeb &amp; Bardat [2014], PC[PC102136]!; isolecto-, G[G00067576]!).</p> <p>Aneura dismieri Steph., Species Hepaticarum 6: 24 (1917). — Type: Madagascar (Reeb &amp; Bardat 2014).</p> <p>Aneura papulosa Steph., Species Hepaticarum 6: 36 (1917). nom. illeg. — Type: Seychelles (Reeb &amp; Bardat 2014 as synonym of R. obtusa).</p> <p>Riccardia obtusa S.W.Arnell, Botaniska Notiser 105: 142 (1952). — Type: Cape Province, Knysna, 28.XI.1951, Arnell 1602 (holotype, UPS; isotypes, BOL!, S-B20192!*) syn. nov.</p> <p>Riccardia rhodesiae S.W.Arnell., Botaniska Notiser 105: 150 (1952). — Type: Zimbabwe (Reeb &amp; Bardat 2014).</p> <p>FURTHER SPECIMENS EXAMINED. — Cameroon. Buea, 13.VII.1891, Dusén 251, as Aneura caespitans (BM-00167935, G00067548).</p> <p>Comores. Karthala, 1600 m, 01. V.2008, Bardat KarP1 Q 2Li1* (PC 0763849).</p> <p>Madagascar. Alaotra-Mangoro, Vohimana Reserve, 18°55’15.86”S, 48°30’55.76”E, 782 m, 28. VI.2012, Reeb MTV1211* (PC 0763881 A, TAN), 18°55’13.1”S, 48°30’57.1”E, 789 m, 28. VI.2012, Reeb MTV1220 (PC 0763875), MTV1221* (PC 0763880, TAN), 18°55’43.6”S, 48°29’56.6”E, 849 m, 30. VI.2012, Reeb MTV1235* (PC 0763886, TAN); Alaotra-Mangoro, Zahamena Reserve, 17°38’19”S, 48°36’46.3E, 1156 m, 28.XII.2013, Reeb CR 13 Z 26 * (PC 0763870), CR 13 Z 27 * (PC 0763871), 17°38’25”S 48°38’21E, 1100 m, 29.XII.2013, Reeb CR 13 Z 40 * (PC 0763867), 17°38’22.7”S 48°38’45.3, 1294 m, 30.XII.2013, Reeb CR 13 Z 57 * (PC 0763866); Analamanga, Angavokely Reserve, 18°55’12.8”S 47°44’29.3E, 1470 m, 31. I.2011, Reeb CR 11142 * (PC 0763883).</p> <p>Malawi. Mulanje Mt., 15°56’09”S, 35°31’02”E, 980 m, 30. VI.1991, Wigginton M 1758 B * (E 00430531), 15°52’50”S, 35°36’36”E, 1800 m, 23. VI.1991, O’Shea M 7409 A * (E 00430479).</p> <p>Mauritius. Black River Gorges Nat. Park, 20°24.5’S, 57°27.5’E, 540 m, 07.VIII.1996, Pócs 9667/ F * (EGR); Mt. Cocotte, 700 m, 08.X.1962, Een M 064 (S-B7132).</p> <p>Réunion. 13 km W of St. Anne, 021°4’35”S, 55°37’10”E, 520- 800 m, 30.VIII.1994, Orban 9436/ T * (G 00264122, EGR); Cirque de Salazie, 21°03’00”S, 55°32’00”E, 1550-1800 m, 01.IX.1994, Vojko 9438 / CF* (EGR); Langevin Valley, 21°17’05”S, 55°38’05”E, 550-850 m, 20.VIII.1996, Pócs 9686/X* (G 00264118, EGR); Forêt de Tévelave, 1470 m, 27.IV.2011, Bardat TEV.1* (PC 0763865).</p> <p>Tanzania. Uluguru Mts., Bondwa Peak, 1900-2000 m, VIII-X /1969 Pócs 6011/ S, 6011 T; 6011/ I, 6051/AG (EGR), 1400 m, 13.XII.1970, Jones 1210 (EGR).</p> <p>Uganda. Ruwenzori Nat. Park, 2100 m, 01.II.1997, Porley U 554 C * (E 00430545), 22°00” S, 29°59’00”E, 2080 m, 02.II.1997, Wigginton U 8053 B * (E).</p> <p>DISTRIBUTION. — Cameroon, East Africa, South Africa and Indian Ocean Islands.</p> <p>HABITAT. — Often on rock, sometimes on bark or, at highest elevations, on soil, always in humid evergreen forests and usually in or near river beds, from 540 to over 2000 m.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous or paroicous) or dioicous (R. obtusa).</p> <p>Thallus</p> <p>Fleshy, green to black green, convex with incurved branches when dry, dull to glossy, to 20 mm long, main axis closely attached to the substrate, branches arising upwards, branching 1-2-pinnate in the lower half of the thallus and palmate in the upper half, branches alternate to subopposite, reiterations present, flagelliform branches occasional, stolons frequent.</p> <p>Main axis</p> <p>Well-defined, 100-420 µm wide, in cross section biconvex to plano-convex at the base, 5-8 cells high, biconvex to plano-convex or concave-convex in the middle, 5-7 cells high, margins acute, of 1-2 rows of bulging cells, not clearly winged, often asymmetrical, epidermis cells more than 5× smaller than inner cells.</p> <p>Primary branches</p> <p>To 7 mm long, crowded and often overlapping, angles with the axis variable.</p> <p>Ultimate branches</p> <p>Crowded, oblong, sometimes tongue-shaped, 600-1500 µm long, 250-400(-650) µm wide, in cross-section plano-convex to concave-convex, 4-5(-7) cells thick, with 2-3(-4) cells wide wings, epidermis cells 2-4 × smaller than inner cells, margins irregularly undulate, apex widened, rounded to truncate to emarginate to dissected, without overlapping edges, with numerous multi-apical areas. Mucilage papillae apical and ventral. Oil bodies (Meenks &amp; Pócs 1985) dark brown, present in epidermis cells.</p> <p>Sexual branches</p> <p>Usually grouped, lateral to ventro-laterally inserted.</p> <p>Male branches</p> <p>Mainly sessile, 300-500(-900 µm) long, with 2-3 cells wide scale-like wings, the wings narrowed than the axis, antheridia in 2-10 pairs.</p> <p>Female branches</p> <p>500-750 µm long, ventrally inserted, bordered by straight, hair-like cilia, archegonia in 2-6 pairs.</p> <p>Calyptra</p> <p>1.5-3 mm long, terminating in corona, surface with numerous thick-walled cells.</p> <p>Gemmae</p> <p>Not seen.</p></div> 	https://treatment.plazi.org/id/03EF87AA6537FFDDC99BE25FFF17FCFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
03EF87AA6534FFDDC9BEE2FFFC04F987.text	03EF87AA6534FFDDC9BEE2FFFC04F987.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riccardia vohimanensis Reeb & Gradstein 2020	<div><p>Riccardia vohimanensis Reeb &amp; Gradst. sp. nov.</p> <p>(Fig. 4 U-X)</p> <p>DIAGNOSIS. — Monoicous. Thallus rigid, unwinged, with short branches. Axis plano-convex to biconvex, 8-12 cells thick, ventral side with mycorrhiza-like fungi. Immature calyptra strawberry-like.</p> <p>TYPE. — Madagascar. Vohimana Reserve, 18°55’43.6S 48°29’53.6”E 849 m, 15.IV.2010, Reeb, Andriamantena &amp; Bidault CRAE 96 * (holo-, PC [PC 0771053]).</p> <p>FURTHER SPECIMENS EXAMINED. — Madagascar. Alaotra-Mangoro, Zahamena Reserve, 17°38’22”S, 48°38’47”E, 1247 m, 27.XII.2013, Reeb CR 13 Z 23* (PC 0763868); Maromizaha Reserve, 18°58’33.9”S 48°27’54.3”E, 1021 m, 05.VII.2012, Reeb MTM1278* (PC 0763879), 18°58’45”S, 48°27’51.8”E, 1011 m, 06.VII.2012, Reeb MTM1290* (PC 0763878); Haute Matsiatra, Andringitra Nat. Park, 22°08’4061” S, 46°53’47.4”E, 1590 m, 21. V.2016, Reeb CR 16 M 52 (PC).</p> <p>Malawi. Mulanje Mt., 15°53’32.1”S, 35°29’24”E, 2180 m, 22. VI.1991, O’Shea M 7327A-2* (E 00430480).</p> <p>Mauritius. Macabé, 550 m, 03.X.1962, Een M 028* (S-B7097).</p> <p>DISTRIBUTION. — Malawi, Madagascar, Mauritius.</p> <p>HABITAT. — On dead logs (bare, smooth and very hard wood) and buttresses, rarely on humid rock, from 550 m to 2200 m.</p> <p>DESCRIPTION</p> <p>Monoicous (autoicous or paroicous).</p> <p>Thallus</p> <p>Leathery and rigid, green to blackish green when fresh and forming rosettes on bare substrates, dull brown to black in herbarium, closely attached to the substrate, to 22 mm long, 1-2-pinnate, not palmate, branches alternate, not flagelliform, stolons lacking.</p> <p>Main axis</p> <p>Well-defined, 700-1000(-1200) µm wide, in cross section plano-convex to concave-convex, with mycorrhizal fungi in the ventral part, 8-12 cells thick, margins obtuse, not winged, epidermis cells 2-4× smaller than inner cells.</p> <p>Ultimate branches</p> <p>Oblong and stocky, to 2 mm long and 1 mm wide, in cross section biconvex to plano-convex, 7-9 cells thick, margins acute, not winged, branch apex rounded and often widened, emarginate to weakly dissected, without overlapping edges.</p> <p>Mucilage papillae</p> <p>Apical and lateral.</p> <p>Oil bodies</p> <p>Not seen.</p> <p>Sexual branches</p> <p>Numerous all along axis, often close to each other.</p> <p>Male branches</p> <p>Mainly grouped, sessile, 300-400 µm long, not winged, with 4-8 antheridial pairs.</p> <p>Female branches</p> <p>Grouped, laterally inserted, 300-500 µm long, base often elongate, female branch bordered by scales or long hair-like, straight cilia, the cilia longer than the width of the axis, archegonia in 4-6 pairs.</p> <p>Calyptra</p> <p>Less than 1 mm long, strawberry-like when immature, densely covered by rounded, thick-walled cells.</p> <p>Gemmae</p> <p>Sometimes present.</p></div> 	https://treatment.plazi.org/id/03EF87AA6534FFDDC9BEE2FFFC04F987	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Reeb, Catherine;Gradstein, Robbert	Reeb, Catherine, Gradstein, Robbert (2020): A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key. Cryptogamie, Bryologie 20 (2): 11-34, DOI: 10.5252/cryptogamie-bryologie2020v41a2, URL: http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2
