taxonID	type	description	language	source
03EFA657FFC1CC3DFF4BFA68FDA4FD28.taxon	materials_examined	Type species: Tillicera javana Spinola, 1844 Spinola 1841: 73; Spinola 1844: 157; Lacordaire 1857: 440; Desmarest, in Chenu 1860: 241; Schenkling 1903: 26, 37. Body: Stout, compact, length 6 – 16 mm.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC1CC3DFF4BFA68FDA4FD28.taxon	description	Head: Head including eyes not conspicuously broader than pronotum; eyes margined, protruding laterally, finely facetted, separated by more than one eye width, emarginate proximal to antennal base; labrum bilobed; terminal maxillary palpomeres digitiform; terminal labial palpomeres securiform; head from posterior margin of eyes towards lower part with fine longitudinal rugosity; antennae 11 - segmented, basal funicular antennomeres filiform, antennomeres from A 4 to A 7 onwards (though most often from A 5) triangularly dilated, A 2 shorter than A 3, A 10 mostly broader than long or nearly as wide as long. Gular sutures apically divergent, gular process narrow (not bilobed). Thorax: Procoxal cavities open behind; pro-intercoxal process narrow, linear, slightly dilated distally; pronotum almost always longer than broad (except T. callosa n. sp., T. michaeli n. sp.), broadest in middle or just behind middle; anterior surface of mesepisternum transversely wrinkled, postero-laterally punctate or coarsely wrinkled, smooth towards middle; anterior mesosternal process present, forming a triangular bulge; posterior meso-intercoxal process stout, depressed or more or less conspicuously erect; scutellum circular. Elytra: Elongate, subparallel, almost two times longer than broad; each elytron with ten rows of coarse punctures; elytral base not margined; scutellum more or less circular. Legs: Short; tibiae with (each on the inner and outer faces) or without longitudinal carina; tarsi pentamerous; tarsal pulvillar formula 4 - 4 - 2; tibial spur formula 1 - 2 - 2; pretarsal claws unipartite, with basal denticles. Aedeagus: With the exception of Tillicera tonkinensis, all investigated species show tooth-like or hookshaped structures at parameres. Important differential character for the separation of Tillicera from similar genera is the tarsal pulvillar formula: Clerus 4 - 3 - 2, Thanasimus 4 - 4 - 3, Omadius 4 - 1 - 1 and Hemitrachys 4 - 2 - 2. A revision of the ‘ Tillicera group’ of genera is in preparation.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC3AFF4BFF05FE31FD5A.taxon	description	(Fig. 18)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC3AFF4BFF05FE31FD5A.taxon	description	Length: 6.5 mm Head: Black, glossy; frons with dense, long, white hairs; vertex with a few scattered black bristles. Antennae: Yellow, clubbed. Pronotum: Red, glossy; with stiff, scattered, black bristles, a “ bunch ” of white bristles projecting from the lateral margin just above the base. Elytra: Black, with two reddish-yellow transverse fasciae vested with dense white pubescence, white posterior transverse fascia narrower; apex black, with few white hairs. Ventral surface: Black; thorax clothed with long thick white hairs. Legs: Black, basal part of femora yellow, with long yellow and white hairs.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC3AFF4BFF05FE31FD5A.taxon	distribution	Distribution: India.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC36FF4BFCC7FCE8FBA9.taxon	description	(Figs. 1, 19 – 20)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC36FF4BFCC7FCE8FBA9.taxon	description	Paratype: Pseudoclerops auratofasciatus Pic, 1927, ZMAN type COLE. 1801.1, Cotype: Pseudoclerops auratofasciatus 1927, M. Pic, Tonkin, Chapa, 1932, ex coll. M. Pic (ZMAN). Others: Laos, Phongsaly Prov., Phongsaly env., Phu Fa, h: 1450 – 1600 m, 27.7.2006, leg. M. Geiser, Bergregenwald (2 ex., MGCB); Thailand, Chiangmai, Doi Pui, 14.5.85 (1 ex., RGCM); Thailand, Chiangmai, Doi Pui, 26.5.1985 (2 ex., RGCM); Doi Pui, 1400 – 1500 m Chiang Mai, N. Thailand, 19 - VI- 1983, T. Shimomura leg. (1 ex., RGCM); Thailand, Soppong Pai 1. - 8.5.1993, Pacholatko & Dembicky leg. (1 ex., RGCM); N. Thailand, Meo Village, near Chiang Mai, 5.1988 (1 ex., RGCM); Mt. Tam Dao, Vinh Phu, N. Vietnam, 5.1998 (1 ex., RGCM); Tam Dao, N. Vietnam, 8.1999 (1 ex., RGCM); 2. – 8.5.1994, Tam Dao, (N. Vietnam), Local-coll (1 ex., RGCM); Xia-Cha-Yu, 1900 – 2600 m, Cha Yu County, Tibet, China, 1. – 28.7.2004, Li Jingke (1 ex., RGCM; 1 ex., JSBC); Carin Cheba, 900 – 1100 m, L. Fea V. XII- 88 (2 ex., MCSN); Chapa, Coll. M. Pic (1 ex., MNHN); Tam Dao, Tonkin, H. Perrot, Coll. M. Pic (1 ex., MNHN); Hoa-Binh, Tonkin 1912, Tonkin, V. Laboissiere, 1913 (1 ex., MNHN); Carin Cheba, 900 – 1100 m, L. Fea V. XII- 88, ex Coll. R. Oberthur (1 ex., MNHN); Carin Cheba, 900 – 1100 m, L. Fea V. XII- 88, Coll. Gorham Oliver, 1914 (1 ex., MNHN); H te Birmanie, Mines des Rubis, 1200 m – 1300 m, Doherty 1890, ex Coll. R. Oberthur (2 ex., MNHN); Laos, Phongsaly prov., 21 ° 41 - 2 ’ N, 102 ° 06 - 8 ’ E, Phongsaly env., 28.5. – 20.6.2003, ~ 1500 m, M. Brancucci leg. (5 ex., NHMB); N E. Burma, Kambaiti, 4. – 8.6.1934, R. Malaise (1 ex., NHRS); Tonkin, Chapa, 7.1916, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN); Laos, Ban Pan, 2.5.1918, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN); Laos, Vien Pou Kha, 9.5.1918, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN); Laos, Xieng Khouang, 13. V. 1919, Laos, X. Khouang, le 13 - V 1919, R. Vitalis de Salvaza, Homotype, Tillicera auratofasciata Pic, Ekis 1974 (1 ex., ZMAN). Length: 9 – 16 mm (34 specimens). Collecting period: May – August, December. Head: Black or brown; anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black to brown (or paler); distributed with dense, fine punctation; cranium vested with golden and dark pubescence; head including eyes only slightly broader than anterior width of pronotum. Antennae: A 1 to A 4 light brown with erect yellow setae, from A 5 onwards dark brown to black, vested with depressed and erect black setae; antennomeres triangularly dilated from A 5 or A 6 onwards, A 10 broader than long or breadth and width subequal. Pronotum: Unicoloured red-brown to brown, surface anterior to transverse depression sometimes black or with a semicircular black spot anterior to middle; surface punctation anterior to transverse depression fine, densely and irregularly distributed, surface behind transverse depression more coarsely punctate; vested with anteriorly directed erect black and golden setae; length: width ratio 1.14: 1 (broadest in middle). Scutellum: Brown, finely punctate, vested with a few golden-yellow setae. Elytra: Black, basal third red-brown to brown (side margins bordered black, in two specimens this redbrown part is reduced to a macula on each elytron), margined posteriorly by two yellowish pigmented transverse maculae (spanning from the lateral margin to mid-way towards suture) with dense, golden-yellow, depressed setae, between these maculae with a semicircular sickle-like pattern of pale setae (sometimes with a pale pigmented spot in the middle), apical third with a pale pigmented transverse fascia (not always pigmented towards suture), with dense, golden-yellow, depressed setae; disc regularly and deeply punctate in rows, diameter of punctures larger than interstices, interstices partly with tubercles; the black middle part without the first two or three rows of punctation, apex densely and very finely punctate; basal third with erect golden and dark setae, the black part with posteriorly directed and semi-decumbent, predominantly dark and isolated yellow (mainly at apex) setae; length: width ratio 2.21: 1. Legs: Brown to black, often paler at base; with more or less dense, golden-yellow, erect setae; tibiae with longitudinal carinae. Ventral surface: Abdomen black to brown, pro- and mesothorax brown, metathorax black or brown with golden-yellow, depressed pubescence. Aedeagus: Conspicuously asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC7CC36FF4BFCC7FCE8FBA9.taxon	distribution	Distribution: China, Myanmar, Thailand, Laos, Vietnam.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCBCC35FF4BFBF0FE31FBAA.taxon	description	(Figs. 2, 21 – 22)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCBCC35FF4BFBF0FE31FBAA.taxon	description	Others: India, Mysore, Coorg, Mercara, 5.73 1200 m, leg. Th. R. S. Nathan (1 ex., RGCM); Maissour, Manganali, 1897 (1 ex., RGCM); Bombay, Fontanier 301 - 39 (1 ex., MNHN); India, Bombay, Ex coll. Janson, ex Coll. R. Oberthur (1 ex., MNHN); India, Ex Museo James Thomson, ex Coll. R. Oberthur (1 ex., MNHN); Inde Occidentale, Règion de Coorg, Chasseurs Indigènes, 2 e semestre 1905, Ex Coll. M. Maindros, Coll. G. Babault 1930 (1 ex., MNHN); Wallardi (Travancore), Ex Coll. M. Maindros, Coll. G. Babault 1930 (1 ex., MNHN); Nilgiri Hills, don. H. E. Andrewes (1 ex., ZMAN); Anamalai hills, 2400, Madras, J. C. M. Gardner, 10.5.1930 (1 ex., ZMAN). Length: 11 – 15 mm (10 specimens). Collecting period: May. Head: Red-brown to brown, clypeus, labrum and palpi brown; distributed with fine, dense punctation; cranium vested with short, erect, predominantly golden pubescence; head including eyes only slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 5 paler, vested with depressed golden setae; triangularly dilated from A 5 or A 6 onwards, antennomeres generally narrower than in all other Tillicera species; A 10 as long as wide in females, longer than broad in males; A 11 conspicuously extended. Pronotum: Red-brown to brown; surface finely and irregularly punctate; with anteriorly directed erect dark and short golden setae; length: width ratio 1.07: 1 (broadest in middle). Scutellum: Brown to black, finely punctate, partially vested with very dense golden setae. Elytra: Black, humeral and apical regions predominantly with long, erect, golden setae, posterior black region with loose depressed dark posteriorly directed setae; with two broad, pale pigmented transverse fasciae with dense, depressed, golden setae, the posterior fascia almost reaching the sutural angle; a pale pigmented macula is dilated around the scutellum; disc deeply punctate in regular rows from base towards posterior fascia, diameter of punctures broader than interstices; apex densely and finely punctate, sutural angle slightly emarginated; length: width ratio 2.11: 1. Legs: Brown, with erect golden setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, pro-and mesothorax brown, metathorax dark brown to black with dense, depressed golden setae; penultimate abdominal sternite conspicuously emarginated in males. Aedeagus: With tooth-like structures; parameres at lateral view strongly broadened.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCBCC35FF4BFBF0FE31FBAA.taxon	distribution	Distribution: India.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC8CC32FF4BFBF7FB82F846.taxon	description	(Figs. 3, 23 – 24)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC8CC32FF4BFBF7FB82F846.taxon	description	Paralectotype: ♀. Syntypus, Tillicera, bibalteata, Gorham, 1892, Carin Chebá, 900 – 1100 m, L. Fea V XII- 88, Museo Civico di Genova (MCSN). Others: Coll. I. R. Sc. N. B., Cambodia Siem Reap, 28.4. – 04.5.2003 Yellow, tray under neon light, Leg. Steph. De Greef (1 ex., RGCM); Laos, Umg. Vientiane, 3. – 6.1963 (1 ex., RGCM); Thailand, 19.5.1993, Lom Sak – Dan Sai, leg. Pacholatko & Dembicky (3 ex., RGCM); Thailand, Maemo, Lampang, 1.8.1991 (1 ex., RGCM); Thailand, Soppong Pai, 1. – 8.5., 1993 Pacholatko & Dembicky leg. (2 ex., RGCM); Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo (bio. station), 05. – 12.5.2001, Secondary forest, Leg. Constant & Grootaert (1 ex., RGCM); Coll. I. R. Sc. N. B., Thailand: Loei, Na Haeo, 1. – 8.4.2001, Malaise trap, Leg. Constant & Grootaert (1 ex., RGCM); Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo 22.5.2003, Malaise trap 1, Leg. P. Grootaert (1 ex., RGCM); N. Thailand, Meo Village, near Chiang Mai, 5.1988 (1 ex., RGCM); Carin Cheba, 900 – 1100 m, L. Fea V XII- 88 (2 ex., MCSN); Bengale, Collection Léon Fairmaire, 1906 (1 ex., MNHN); Phedong, R. P. Desgodins, Coll. M. Pic (1 ex., MNHN); Inde Anglaise, Pedong, District de Darjeeling, Chasseurs indigènes, 1932, ex Coll. R. Oberthur (1 ex., MNHN); Annam, Quinhone, R. P. Guerlach, 1908 (1 ex., MNHN); Annam, Keng Trap?, 27.5.1917, R. Vitalis de Salvaza, Coll. M. Pic (1 ex., MNHN); Tonkin, Coll A Sicaro 1930 (1 ex., MNHN); Tonkin, Collection Léon Fairmaire, 1906 (1 ex., MNHN); Tonkin, Env. S. Hanoi, (Demange), D. Grouvelle? 1909 (1 ex., MNHN); Cochinchine, Bien – Hoa, Coll. de Marseul (1 ex., MNHN); Laos on, Cambodge, Bonotte 1911 (1 ex., MNHN); Thailand, Nakhon rat- chasima, 19.5.1961 (1 ex., MNHN); Laos, Vieng Xai, 10.5.1918, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN). Length: 8 – 13.5 mm (28 specimens). Collecting period: April, May, August, December. Head: Red-brown to brown, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus and sometimes frons black; distributed with fine and diffuse punctation; anterior part of cranium vested with pale, erect, pubescence, posteriorly also with dark setae; head including eyes only slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 3 slightly paler, vested with depressed, short pale and dark setae; antennomeres triangularly dilated from A 5 onwards, A 10 broader than long. Pronotum: Red-brown to brown; surface finely and irregularly punctate; vested with anteriorly directed erect, short white and longer black setae; length: width ratio 1.02: 1 (broadest in middle). Scutellum: Brown, finely punctate, vested with pale setae. Elytra: Black, with a brown macula around scutellum and two broad, pale pigmented transverse fasciae in middle and apical third; disc with regular, deep, rasp-like punctation, diameter of punctures larger than interstices, only rows 4 to 10 extending towards apical fascia, rows 3 to 9 terminating just behind the middle fascia, apex densely and finely punctate; setae at basal third along the suture and between the punctation short and pale, remainder with posteriorly directed, depressed, short and some erect, long, dark setae; length: width ratio 2.05: 1. Legs: Dark brown to black, with dense, pale, erect setae; tibiae mostly without longitudinal carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with dense, depressed, pale setae; penultimate ventrite of males deeply emarginated. Aedeagus: Tip of phallus slightly asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFC8CC32FF4BFBF7FB82F846.taxon	distribution	Distribution: Bhutan, Myanmar, Thailand, Laos, Vietnam, Cambodia.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCECC30FF4BF977FD4EFC64.taxon	description	(Figs. 4, 25)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCECC30FF4BF977FD4EFC64.taxon	materials_examined	Holotype: ♀. India, West Bengalen, Distr. Darjeeling, Kurseong, Rapesing, 1400 m, 13.8.1996, leg. N. Dangal (RGCM). Length: 9 mm (1 specimen). Collecting period: August. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; distributed with fine and diffuse punctation; cranium vested with erect, pale and dark setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, vested with depressed, short, dark setae; antennomeres triangularly dilated from A 5 onwards, A 10 approximately as long as wide. Pronotum: Red-brown to brown; surface finely and densely punctate; vested with anteriorly directed erect, predominantly dark and shorter, depressed, pale setae; length: width ratio 0.96: 1 (broadest in middle). Scutellum: Brown, finely punctate, vested with pale setae. Elytra: Black, with two broad, pale pigmented transverse fasciae in middle and apical third, with dense, depressed, pale setae; humeral callous present; basal third regularly and deeply punctate in rows towards posterior fascia, diameter of punctures broader than interstices, apex densely and finely punctate; basal region with long, erect, dark setae, central region with semi-decumbent, posteriorly directed, short, dark setae, apex with longer, pale setae; length: width ratio 2.03: 1. Legs: Dark brown to black, meso- and metafemora basally paler, with dense, pale, erect setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with dense, depressed, pale setae.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCECC30FF4BF977FD4EFC64.taxon	distribution	Distribution: India.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCECC30FF4BF977FD4EFC64.taxon	etymology	Etymology: The trivial name callosa is a Latin adjective; we refer to the characteristic elytral humeral callosity.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCECC30FF4BF977FD4EFC64.taxon	diagnosis	Differential diagnosis to T. michaeli: The new species is similar to T. michaeli n. sp., but the vestiture of T. michaeli is more strongly golden and glossy than in T. callosa n. sp.; the transverse fasciae are pigmented pale in T. callosa n. sp., not in T. michaeli n. sp.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCDCC2EFF4BFB80FBB9FE87.taxon	description	(Figs. 5 – 6, 26 – 27)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCDCC2EFF4BFB80FBB9FE87.taxon	description	Specimens examined: Holotype: ♀. Typus, Syntypus, Tillicera cleroides, Gorham, 1892, cleroides, Gorh., Carin Chebá, 900 – 1100 m, L. Fea V XII- 88, Museo Civico di Genova (MCSN). Lectotype: ♀. Type, Tonkin, Thanasimus mimomutillus n. sp., Museum Paris, Coll. M. Pic, (MNHN) Tillicera mimomutilla n. syn. Paralectotypes: ♀, same data as Lectotypus (pinned together) (1 ex., MNHN); Thanasimus, mimomutillus, Pic, 1926, ZMAN type COLE. 1796.1, Cotype: Thanasimus mimomutillus Pic 1926, Tonkin, Lac Thò, Thanasimus mimomutillus Pic (1 ex., ZMAN); (on one pin): Paratype, Thanasimus mimomutillus Pic, Cotypes, Hoa Binh, Tonkin, Brit. Mus., 1951 - 293 (2 ex., BMNH).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCDCC2EFF4BFB80FBB9FE87.taxon	materials_examined	Holotype: ♂. Type, v. Rouyeri, Pic, Mt. Semerou, Volcans Java, (Rouyer), Museum Paris, Coll. M. Pic (MNHN), Dasyceroclerus rouyeri n. comb, n. syn. Holotype: ♂. Type, Chapa, Tonkin, v. nov. bicoloricollis, Museum Paris, Coll. M. Pic (MNHN) Paratype: Thanasimus mimomutillus bicoloricollis, Pic, 1931, ZMAN type COLE. 1797.1, Cotype: Thanasimus mimomutillus Pic, v. bicoloricollis Pic, Tonkin, Hanoi, 1932, ex Coll., M. Pic (ZMAN). 2 Types ab. bipartita: ♂. v. bipartita, Pic, Hoa Binh, Museum Paris, Coll. M. Pic (MNHN) Tillicera mimomutilla ab. bipartita n. syn. Others: Laos, Vientiane Prov., Vang Vieng env., Umgebung der Ortschaft, 20.7.2006, Bananenplantagen und Reisfelder, an Strauch (1 ex., RGCM); China: S-Yunnan, (Xishuangbanna), 23 km NW Jinghong, vic. Na Ban, N 22 09.49, E 100 39.92, 05.6.2008 leg. A. Weigel, 730 m second. forest KF (1 ex., RGCM); Chiang Dao, Chiang Mai Prov., N. Thailand, 24.6.1979, W. Suzuki leg. (1 ex., RGCM); Nord, Vietnam Tonkin, Hoa-Binh, ex. Coll. Oberthur, I. G. 18293 (4 ex., RGCM); Tonkin: Thai-Nien. Banks, of Fleuve Rouge, 1924, H. Stevens (1 ex., RGCM); Hanoi (Vauloger), (Coll. Ph Francois), Coll. L. Bedel 1922 (2 ex., MNHN); D. Nodier, Hanoi, Coll. M. Pic (4 ex., MNHN); Hanoi, 7.02, Coll. M. Pic (1 ex., MNHN); Tonkin, Coll. M. Pic (1 ex., MNHN); Museum Paris, Tonkin, Dap Can, H. Donckier 1907, 14. au 18.5.06 (1 ex., MNHN); Museum Paris, Tonkin, Dap Can, H. Donckier 1906, 4. au 8.5.06 (1 ex., MNHN); Dong Jang, Tonkin, 4.1903, Vauloger, (Coll. Ph Francois), Coll. L. Bedel 1922 (4 ex., MNHN); D. Nodier, Dong-Dang, Coll. M. Pic (1 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa Binh, A. de Cooman 1987 (1 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa Binh, A. de Cooman, ex Coll. R. Oberthur (3 ex., MNHN); Tonkin occ., Env. de Hoa-Binh, R. P. A. de Cooman, 1919 (1 ex., MNHN); Tonkin occ., Rég. de Hoa-Binh, 1919, ex Coll. R. Oberthur (1 ex., MNHN); Tonkin, Région de, Hoa-Binh, Museum Paris, 1934, A. de Cooman (1 ex., MNHN); Tonkin occ., Rég. de Hoa-Binh, 1919, ex Coll. R. Oberthur (1 ex., MNHN); Hoa-Binh, Coll. M. Pic (1 ex., MNHN); Hoa Binh, Coll. M. Pic (3 ex., MNHN); Hoa Binh, Tonkin, Coll. M. Pic (5 ex., MNHN); Museum Paris, Tonkin, Rég de Hoa-Binh, A. de Cooman 1926 (1 ex., MNHN); Lactho, Tonkin, de lineCooman, Coll. M. Pic (1 ex., MNHN); Env. de Lam, (Tonkin), L. Blaise, (Coll. Ph Francois), Coll. L. Bedel 1922 (4 ex., MNHN); Rég. de Luc-Nam, (Tonkin), L. Blaise, (Coll. Ph Francois), Coll. L. Bedel 1922 (5 ex., MNHN); Museum Paris, Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901 (1 ex., MNHN); Sept-Pagodes, (Tonkin), L. Blaise, (Coll. Ph Francois), Coll. L. Bedel 1922 (3 ex., MNHN); Museum Paris, Lakhon, Marmand 1878 (1 ex., MNHN); Phuyen Binh, 11.1907, Coll. M. Pic (2 ex., MNHN); Carin Chebà, 900 – 1100 m, L. Fea V XII. 88 (1 ex., MNHN); Samberbajem, Res. Kediri, E. Java, ex Louwerens, Coll. M. Pic (1 ex., MNHN); Coll. M. Pic (2 ex., MNHN); Hainan Is., NamChaChuen, S. W. of Nodoa, July 8, Lingnan, University, 5 th. Hainan Is., Exped. 1929 (1 ex., ZMAN); Depok, 4.1948, C. v. Nidek, Collectie, C. v. Nidek, Acq. 1969 (1 ex., ZMAN); C. J. Louwerens O. - Java, Res. Kediri, Sambong (2 ex., ZMAN); C. J. Louwerens, O. - Java, Res. Kediri, Glodok, Metatype: Thanasimus mimomutillus 1937 M. Pic (1 ex., ZMAN); 1 Ex. C. J. Louwerens, O. - Java, Res. Kediri, Plengong Qentir-, Qentiran?, Metatype: Thanasimus mimomutillus, 1937 M. Pic (1 ex., ZMAN); C. J. Louwerens, O. - Java, Res. Kediri, Zuidergebergte, Popoh, Thanasimus mimomutillus, Pic, Metatype: Thanasimus mimomutillus, 1937 M. Pic (1 ex., ZMAN); C. J. Louwerens, Java 84 m, Toeloengagoeng (1 ex., ZMAN); Babakan. Z. K. -, Java Drescher, 7.11.1937 (1 ex., ZMAN); Dammerman, O. Soemba, 450 M., Mao Marroe, 5.1925 (1 ex., ZMAN). Length: 7 – 10 mm (83 specimens). Collecting period: April – July, November, December. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus brown to black; distributed with fine and diffuse punctation; cranium vested with erect, pale setae; head including eyes as broad as anterior width of pronotum. Antennae: A 1 to A 4 light brown, from A 5 onwards dark brown to black, with brown and golden depressed setae; antennomeres triangularly dilated from A 5 or A 6 (sometimes from A 7) onwards, A 10 broader than long. Pronotum: Black, glossy, anteriorly and posteriorly narrowly brown to red-brown (at sides sometimes posteriorly dilated towards apex); surface finely and diffusely punctate; vested with predominantly anteriorly directed erect, dark setae, towards apex and base also with golden setae; length: width ratio 1.14: 1 (broadest in middle). Scutellum: Brown, finely punctate, vested with pale setae. Elytra: Basal half red-brown to brown, remainder black, behind middle with a yellowish pigmented fascia which bears dense golden setae (spanning from the lateral margin to mid-way towards suture), and a posterior pale pigmented fascia, bearing dense golden setae, and not reaching the lateral margin; disc deeply punctate in rows, diameter of punctures larger than interstices, the basal red-brown part with tubercles, apex densely and finely punctate, emarginated; basal half with erect, dark and golden setae, black part with posteriroly directed, semi-decumbent, and isolated erect dark setae, apex with golden setae gradually more erect approaching apices; length: width ratio 2.19: 1. Legs: Dark brown to black, pro- and mesofemora paler, mostly with a dark marking dorsally, metafemora paler only basally, metatibiae often with a pale marking, legs with erect, pale setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, with few pale setae, thorax brown, with dense, pale setae; penultimate ventrite of males deeply emarginated. Aedeagus: Conspicuously asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFCDCC2EFF4BFB80FBB9FE87.taxon	distribution	Distribution: China, Myanmar, Thailand, Laos, Vietnam, Indonesia (Java).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD2CC2CFF4BFED8FBDEFCD2.taxon	description	(Figs. 7 – 8, 28 – 29)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD2CC2CFF4BFED8FBDEFCD2.taxon	description	Others: W - Sumatra 9. – 29.10.1991, Payakumbuth, Harau-Vall., 1000 m, leg. A. Riedel, coll. R. Gerstmeier (1 ex., RGCM); Harau, W. Sumatra, Indonesia, 7.1996 (1 ex., RGCM); Lac Thô, Tonkin (1 ex., RGCM); Phu yen binh, 1907 (1 ex., RGCM); Sud Borneo, Goenong Pandjang, Tanggarang, M. E. Walsh, 6.1937 (1 ex., MNHN); Est Borneo, Babi djoelon, M. M. E. Walsh, 1937 (1 ex., MNHN); Tonkin occ., Règ. de Hoa Binh, 1919, ex Coll. R. Oberthur (1 ex., MNHN); Hoa Binh, Coll. M. Pic (1 ex., MNHN); Pho. Vi, 6.1907, Coll. M. Pic (1 ex., MNHN); Sumatra, Palembang, H. Donckier 1898 (1 ex., MNHN); Sandakan, Borneo, Baker (1 ex., ZMAN); Sarawak, Kuching, 17.6.1899 (1 ex., ZMAN). Length: 7 – 14 mm (13 specimens). Collecting period: June, July, October. Head: Black or red-brown to red, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus dark brown to black; distributed with dense, fine punctation; cranium vested with erect pale and dark setae; head including eyes scarcely broader than anterior width of pronotum. Antennae: Dark brown, A 1 to A 4 or A 5 light brown, vested with depressed, short pale and dark setae; antennomeres triangularly dilated from A 6 onwards, A 10 broader than long. Pronotum: Red-brown to brown; surface finely and irregularly punctate, punctation anterior to transverse depression denser; vested with anteriorly directed erect, short white and longer black setae; length: width ratio 1.09: 1 (broadest in middle). Scutellum: Brown, finely punctate, vested with pale setae. Elytra: Black, with two yellowish pigmented transverse fasciae (spanning from the lateral margin to midway towards suture), with dense, depressed, pale setae, and a brown macula surrounding the scutellum; humeral callous present; basal third deeply punctate in rows, appearing irregular, near humeral callous only rows 4 to 8 extending almost to posterior fascia, rows 2, 3 and 9 only obscure or terminating shortly behind the anterior fascia, diameter of punctures larger than interstices; apex densely and finely punctate; basal third with erect golden and dark setae, remainder with predominantly posteriorly directed, semi-decumbent, long, dark setae, apex with long, erect, golden setae; length: width ratio 2.01: 1. Legs: Brown to red-brown or black, base frequently paler, with dense, pale, erect and isolated dark setae; hind tibiae with or without carinae. Ventral surface: Abdomen dark brown to black, thorax brown to red-brown, with dense, depressed, pale setae. Aedeagus: Tip slightly asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD2CC2CFF4BFED8FBDEFCD2.taxon	distribution	Distribution: Vietnam, Indonesia (Sumatra), Malaysia (Borneo, Sabah, Sarawak).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD2CC2CFF4BFED8FBDEFCD2.taxon	diagnosis	Differential diagnosis to T. bibalteata: The antennomeres of T. hirsuta are dilated from A 6 onwards, in T. bibalteata from A 5 onwards; the head of T. hirsuta is unicoloured black, T. bibalteata has at least the posterior part red-brown; the apex of elytra is emarginated in T. hirsuta, not in T. bibalteata.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD1CC2AFF4BF947FDBAFD47.taxon	description	(Figs. 9, 30 – 31)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD1CC2AFF4BF947FDBAFD47.taxon	description	Others: Java, J. D. Pasteur 1899 (2 ex., MNHN); F. C. Drescher, G. Tangkoeban Prahoe, 4000 – 5000 Voet, Preanger, Java, 11.1937 (1 ex., ZMAN); same data: 11.1934 (1 ex., RGCM). Length: 6 – 7 mm (5 specimens). Collecting period: November. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus brown to black; distributed with fine and dense punctation; cranium vested with erect, pale setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 3 or A 4 light brown, vested with depressed, brown setae; antennomeres triangularly dilated from A 5 onwards, A 10 broader than long. Pronotum: Surface anterior to transverse depression black, remainder brown, glossy; surface finely, diffusely and irregularly punctate; vested with anteriorly directed erect, pale and dark, setae; length: width ratio 1.11: 1 (broadest in middle). Scutellum: Brown, irregularly punctate, vested with a few pale setae. Elytra: Basal third red-brown to brown, remainder black, with two yellowish pigmented transverse fasciae (spanning from the lateral margin to mid-way towards suture), with dense white setae; basal third with tubercles and deeply punctate in rows, only rows 3 to 7 extending towards the black part, becoming obsolete towards apex, diameter of punctures larger than interstices, apex densely and finely punctate; basal third with erect, pale and isolated dark setae, black part with posteriorly directed, semi-decumbent, predominantly dark setae; length: width ratio 2.24: 1. Legs: Dark brown to black, base paler, with dense, erect, pale setae; meso- and metatibiae with carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with few pale setae. Aedeagus: Very small, tooth-like structures only visible with higher microscopic magnification.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD1CC2AFF4BF947FDBAFD47.taxon	distribution	Distribution: Indonesia (Java).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD7CC28FF4BFCA7FD33FD87.taxon	description	(Figs. 10 – 11, 32 – 33)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD7CC28FF4BFCA7FD33FD87.taxon	materials_examined	Holotype: Tillicera javana Luchti, Corporaal, 1949, ZMAN type COLE. 1799.1, J. B. Corporaal: Type: Holotype: Tillicera javana Spin., ab. Luchti, 1948, Corp., J. B. Corporaal det. 1921: Tillicera javana Spin., Boekit Gabah, Z. W. K. Sumatra, leg. H. Lucht, 2.1919, ex coll. F. C. Drescher (ZMAN). Paratypes: same data as Holotype (2 ex., ZMAN). Others: Indonesien, Ost-Java, Mt Argopura, Nov. 1997 (2 ex., RGCM); F. C. Drescher, G. Tangkoeban Prahoe, 4000 – 5000 Voet, Preanger, Java, 3.1937 (1 ex., HNHM), same label data: 7.11.1928 (1 ex., ZMAN), 27.10.1929 (1 ex., ZMAN), 3.12.1929 (1 ex., ZMAN), 18. – 28.12.1932 (1 ex., ZMAN), 3.1934 (1 ex., RGCM), 9.1934 (1 ex., ZMAN), 10.1934 (2 ex., ZMAN), 1.1936 (2 ex., ZMAN), 9.1936 (1 ex., RGCM), 11.1936 (1 ex., ZMAN), 12.1936 (1 ex., RGCM), 2.1937 (1 ex., ZMAN), 3.1937 (2 ex., ZMAN), 4.1937 (1 ex., ZMAN), 11.1937 (2 ex., ZMAN), 12.1937 (1 ex., MNHN), 1.1938 (1 ex., ZMAN), 8.1938 (1 ex., ZMAN), 1.1940, C. v. Nldek, Acq. 1969 (1 ex., ZMAN); Sumatra, Si-Rambe, 12.90 – 3.91, E. Modigliani (5 ex., MCSN, 3 ex., MNHN, 1 ex. RGCM); Sumatra, Modigliani 1890 (1 ex., MCSN); Java, J. D. Pasteur 1899 (5 ex., MNHN); Java, J. D. Pasteur 268 - 94 (1 ex., MNHN); Java orient., Montes Tengger, 4000 1890, H. Fruhstorfer, ex Coll. R. Oberthur (1 ex., MNHN); Java Occ., Toegoe, ex Coll. R. Oberthur (1 ex., MNHN); Java merid., 1500 1891, H. Fruhstorfer, ex Coll. R. Oberthur (1 ex., MNHN); Kina Balu, Waterstradt?, ex Coll. R. Oberthur (1 ex., MNHN); Sumatra, Si-Rambe, 12.90 – 3.91, E. Modigliani, H. Donckier 1910 (3 ex., MNHN); Palembang, Sumatra, Coll. M. Pic (1 ex., MNHN); Sumatra (W.), G. Merapi S, of Bukittinggi, 1050 – 1800 m, 17.2.1991, Bocák & Bocáková leg. (1 ex., NHMB); Megamendoeng, 11.1934, Handschin (1 ex., NHMB); F. C. Drescher, G. Tangkoeban Prahoe, 4000 – 5000 Voet, Preanger, Java, Sitoe Lembang, 10.1936 (1 ex., ZMAN); Preanger, Java, Slopes M. Gede, 1000 m 1.1935, Native collector (1 ex., ZMAN); Coll. F. C. Drescher, Java, Preanger, N. O. I., Mt. Boerangrang, 1600 m, 11.1936 (2 ex., ZMAN); J. B. Corporaal, Java, Preanger, Djampang (1 ex., ZMAN). G. nePapandajan, Java, Drescher, 2.1915, ex coll. F. C. Drescher (2 ex., ZMAN), same data 2.1916 (1 ex., ZMAN); S. Leefmans, Java, 10.1922, Goen. Tjisalak, G. Tjisalak october, S. Leefmans (1 ex., ZMAN); Pasir Kopo 4.1932, Ned. Indie, W. Java, De Vos tot, Nederveen Cappea (1 ex., ZMAN); ex Mus. Lugd. Bat., J. D. Pasteur, Java occ., Toegoe (1 ex., ZMAN); Homotype, Ekis, 1974 Tillicera, javana, Spinola, Batoerraden, G. Slamet. Java, F. C. Drescher, II. 1926 (1 ex., ZMAN); Batoerraden, G. Slamet. Java, F. C. Drescher, 2.1926 (1 ex., ZMAN), same data 1.1926 (1 ex., ZMAN), 24.4.1932 (1 ex., ZMAN); Tjibodas, 400 M., 1923, Karny 102, 1925 ex coll. Deutsches Entomolog. Institut Berl. - Dahlem (1 ex., ZMAN); Sumatra, Si-Rambe, 12.90 - 3.91, E. Modigliani, 1946 ex Mus. Kopenhagen (1 ex., ZMAN); H. Lucht leg., Sumatra 2. W. K., Boekit Gabah, 4.1919, ex coll. F. C. Drescher (1 ex., ZMAN); Boekit Gabah, Z. W. K. Sumatra, Leg. H. Lucht, 3.1919, ex coll. F. C. Drescher (1 ex., ZMAN); H. Lucht leg., Sumatra 2. W. K., Boekit Gabah, 3.1919, ex coll. F. C. Drescher (1 ex., ZMAN); Collectie, P. F. Sijthoff, Ind. Austr. Arch., L. Kalshoven, don. 1930 (2 ex., ZMAN). Length: 8 – 12.5 mm (78 specimens). Collecting period: August – April. Head: Black, anterior part of clypeus and labrum brown, posterior part of clypeus black, palpi brown to black; distributed with dense punctation; cranium vested with predominantly dark, and isolated golden, setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 3 light brown, vested with depressed, short, golden and dark setae; antennomeres triangularly dilated from A 4 or A 5 onwards, A 10 broader than long. Pronotum: Red-brown to brown with a semicircular black or dark brown spot at apex; surface anterior to transverse depression finely, densely and irregularly punctate or rugose, surface behind transverse depression with more coarse punctation; vested with anteriorly directed erect, black and golden setae; length: width ratio 1.15: 1 (broadest in middle). Scutellum: Brown, finely punctate, almost without setae. Elytra: Basal third red-brown to brown, remainder black, margined posteriorly by two yellowish pigmented, slightly curved maculae (spanning from the lateral margin to mid-way towards suture), which bears dense, light golden, depressed, setae; between these maculae a semicircular sickle-like pattern of light golden setae; apical third with a pale pigmented transverse fascia with light golden setae; basal third regularly and deeply punctate in rows, (only rows 7 and 8 developed within black region), diameter of punctures larger than interstices, interstices with tubercles, apex densely and very finely punctate; basal third with erect golden and dark setae, the black part with posteriorly directed and semi-decumbent, predominantly dark and isolated golden setae; length: width ratio 2.2: 1. Legs: Dark brown to black, with dense, erect, golden setae; tibiae with longitudinal carinae. Ventral surface: Abdomen dark brown to black, pro- and mesothorax brown, metathorax black or brown, with dense, depressed, golden setae. Aedeagus: Asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD7CC28FF4BFCA7FD33FD87.taxon	distribution	Distribution: Indonesia (Sumatra, Java), Malaysia (Borneo, Sabah).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD7CC28FF4BFCA7FD33FD87.taxon	diagnosis	Differential diagnosis to T. auratofasciata: In T. auratofasciata the posterior transverse fascia is mostly pigmented, whereas in T. javana this fascia consists only of pattern-forming setae; distribution, ultimate abdominal ventrites and aedeagi are different.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD5CC26FF4BF9D2FEA6FD32.taxon	description	(Figs. 12, 34 – 35) Specimens examined: Holotype: ♂. Laos, Phongsaly Prov., Phongsaly env., Phu Fa, h: 1450 – 1600 m, 27.7.2006, leg. M. Geiser, Bergregenwald, Umgestürzter Baum (RGCM). Paratypes: Same data as Holotype: 27.7.2006 (2 ex., RGCM, 2 ex., MGCB), 28.7.2006 (1 ex., RGCM). Length: 9 – 10 mm (6 specimens). Collecting period: July. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; relatively densely distributed with fine punctation; anterior part of cranium vested with erect golden setae, posterior part with dark setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, vested with depressed, dark setae; antennomeres triangularly dilated from A 4 or A 5 onwards, A 10 as long as wide. Pronotum: Red-brown to brown, apex sometimes slightly darker; surface finely and densely punctate; vested with anteriorly directed long, pale and dark, erect, setae, and shorter, pale, depressed, setae; length: width ratio 0.93: 1 (broadest just behind middle). Scutellum: Brown, finely punctate, vested with dense, pale setae. Elytra: Black, with two transverse fasciae, the anterior (black) fascia narrowly lined with yellowish depressed setae, the posterior, narrowly yellowish pigmented fascia very broadly lined with yellowish depressed setae; humeral callous present; disc deeply punctate in rows, extending towards posterior transverse fascia, diameter of punctures larger than interstices; apex densely and finely punctate; basal third and apex with erect, long, predominantly golden setae, along suture and at side margins also with shorter depressed, golden setae, middle part with posteriorly directed, short, semi-decumbent, setae, and isolated, long, erect, dark setae; length: width ratio 2.01: 1. Legs: Brown, pro- and mesofemora and base of metafemora paler; with dense, pale, erect setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with dense, depressed, pale setae. Aedeagus: Tip asymmetrical, tooth-like structures only indistinct at base.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD5CC26FF4BF9D2FEA6FD32.taxon	distribution	Distribution: Laos.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD5CC26FF4BF9D2FEA6FD32.taxon	etymology	Etymology: This species is dedicated to Michael Geiser, a young Cleroidea colleague and the collector of this species.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDBCC27FF4BFD7FFE93FC89.taxon	description	(Figs. 13, 36) Specimens examined: Holotype: ♀. Malaysia-W, Pahang, 30 km E of Ipoh, 1500 m, Cameron Highlands, Tanah Rata, 20.2. - 3.3.1998, P. Chechovsky leg., coll. R. Gerstmeier, Munich (RGCM). Paratype: ♀. Malaysia-W, Perak, 25 km E of Ipoh, 2100 m, Banjaran Titi Wangsa mts., Korbu mt., 4. - 13.3.1998, P. Chechovsky leg. (RGCM). Length: 12 – 13 mm (2 specimens). Collecting period: March. Head: Black, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; distributed with fine and diffuse punctation; cranium vested with erect predominantly pale setae; head including eyes only slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 4 or A 5 slightly paler, vested with depressed pale and dark setae; antennomeres triangularly dilated from A 5 or A 6 onwards, A 10 as long as wide. Pronotum: Red-brown to brown, anterior to transverse depression darker; surface finely and diffusely punctate; predominatly vested with anteriorly directed erect, dark and pale, setae; length: width ratio 1.14: 1 (broadest in middle). Scutellum: Brown, with dark margin, finely punctate, scarcely vested with setae. Elytra: Black, basal third with a brown macula, beginning just behind base and reaching posteriorly towards the anterior, brown pigmented, curved, transverse fascia, just behind scutellum with a small macula of pattern-forming pale setae, and a posterior, brown pigmented transverse fascia; basal third regularly and deeply punctate in rows, rows 3 to 10 reaching the black part in the middle, then becoming obsolete, diameter of punctures larger than interstices, apex densely, finely and irregularly punctate, emarginated; basal third with erect pale and dark setae, middle part with posteriorly directed and semi-decumbent, dark setae, apex with predominantly longer, pale setae, the transverse fascia with dense, depressed, pale setae; length: width ratio 2.21: 1. Legs: Brown, femora at least at base slightly paler, with dense erect setae on tibiae and depressed, pale setae on femora; hindtibiae with carinae. Ventral surface: Abdomen black to brown, thorax brown, with dense, pale, depressed setae.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDBCC27FF4BFD7FFE93FC89.taxon	distribution	Distribution: Malaysia.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDBCC27FF4BFD7FFE93FC89.taxon	etymology	Etymology: The trivial name obscura is a Latin adjective, we refer to the comparatively dark colour of the beetle.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDACC27FF4BFC10FE31F86A.taxon	description	(Figs. 14, 37 – 38)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDACC27FF4BFC10FE31F86A.taxon	description	Others: S. India: Nilgiri, Hills: Cherangode, 3500 ft 11.1950, P. S. Nathan (RGCM). Length: 6 – 7 mm (2 specimens). Collecting period: November. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; distributed with fine and diffuse punctation; cranium vested with erect predominantly pale setae; head including eyes as broad as anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 5 paler, vested with depressed, brown and golden setae; antennomeres triangularly dilated from A 5 or A 6 onwards, A 10 broader than long. Pronotum: Red-brown to brown, anterior to transverse depression slightly darker; surface finely and densely punctate; vested with anteriorly directed, erect, dense, golden setae, surface posterior of transverse depression also with long, dark, erect setae; length: width ratio 1.1: 1 (broadest at middle). Scutellum: Brown, finely punctate, vested with few golden setae. Elytra: Basal third red-brown, posteriorly margined with a yellowish pigmented transverse fascia (which is vested with dense, depressed, golden setae), middle part black, apical third brown, with a transverse fascia of dense, pattern-forming, depressed, golden setae. Basal third regularly and deeply punctate in rows, rows 3 to 5, or 3 to 6, obsolete in the black part, with small tubercles, diameter of punctures larger than interstices, apex finely and diffusely punctate; basal third with long, erect golden and dark setae, the black middle part with posteriorly directed and semi-decumbent, dark setae, apical third with depressed and erect golden setae, apex only with erect, golden setae; length: width ratio 2.11: 1. Legs: Brown, with erect pale setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with dense, pale setae. Aedeagus: Asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDACC27FF4BFC10FE31F86A.taxon	distribution	Distribution: India.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD9CC22FF4BF97FFB83FE79.taxon	description	(Figs. 15, 39 – 40) Specimens examined: Holotype: 19 m. or Tana Rata, Cameron Highlands, Malaysia, 4.1987, Wong Tet Fatt leg., (RGCM). Paratypes: same data as Holotype (8 ex., RGCM); J. B. Corporaal, Sumatra`s O. K., Medan 5.12.1918, 20 M. (1 ex., HNHM), same label data: 20.9.19 (2 ex., MNHN), 6.1.21 (1 ex., MNHN), 4.10.17 (1 ex., ZMAN), 19.10.17 (1 ex., ZMAN), 30.10.17 (1 ex., ZMAN), 9.11.17 (1 ex., ZMAN), 8.12.17 (1 ex., ZMAN), 28.1.1918 (1 ex., ZMAN), 20.9.1919 (1 ex., ZMAN), 21.2.1920 (1 ex., ZMAN), 25.2.1920 (1 ex., ZMAN), 26.2.1920 (1 ex., ZMAN), 20.8.1920 (1 ex., ZMAN), 10.1.21 (1 ex., ZMAN), 11.1.21 (1 ex., ZMAN), 1.2.21 (1 ex., ZMAN), 16.4.1921 (1 ex., ZMAN), 9.8.1921 (1 ex., ZMAN); Museum Paris, Java, Soekaboemi, E. L. Moult. 1925 (1 ex., MNHN); J. B. Corporaal, Sumatra`s O. K., Pagar Marbau 26.12.17, 24 M. (1 ex., ZMAN), same label data: 20.4.19 (1 ex., ZMAN), 4.4.20 (1 ex., ZMAN), 9.4.20 (1 ex., ZMAN); J. B. Corporaal, Sumatra`s O. K., Lau Rakit 2.1918, 300 M. (2 ex., ZMAN); J. B. Corporaal, Sumatra`s O. K., Balimbingan 15.4.20, 370 M. (1 ex., ZMAN); J. B. Corporaal, Medan, 25.2.1920 (1 ex., ZMAN); Java, Collection Le Moult, 1923 (1 ex., ZMAN). Length: 6.5 – 9 mm (39 specimens). Collecting period: August – February, April. Head: Black, glossy, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus brown to black; distributed with fine and diffuse punctation; cranium vested with erect pale setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 4 paler, vested with predominantly brown, depressed setae; antennomeres triangularly dilated from A 5 or A 6 onwards, A 10 broader than long. Pronotum: Red-brown to brown, anterior to transverse depression dark brown to black, apex with a narrow brown to red-brown stripe; surface finely and diffusely punctate; anterior to transverse depression vested with predominantly anteriorly directed, erect, pale and dark setae, apex with dense pale setae, posterior third also with longer, posteriorly directed dark setae; length: width ratio 1.14: 1 (broadest at middle). Scutellum: Brown, finely punctate, vested with few pale setae. Elytra: Basal part red-brown to brown, remainder black, anterior to middle with a yellowish pigmented transverse fascia, which is vested with pale setae, posterior third with a transverse fascia of pattern-forming dense, pale setae, sometimes reduced to a circular macula; basal half regularly and deeply punctate in rows, with small tubercles, rows 1 and 3 to 6 reduced to fine ridges within the black part, diameter of punctures larger than interstices, interstices with tubercles, apex finely and diffusely punctate; basal part with erect dark and pale setae, the black part with posteriorly directed and semi-decumbent, isolated, erect, dark setae, apex with predominantly longer, pale setae; length: width ratio 2.18: 1. Legs: Dark brown to black, femora at least basally paler, sometimes also tibiae with a pale stripe, with dense, erect, pale setae; tibiae without carinae. Ventral surface: Abdomen dark brown to black, thorax brown, with few, pale setae. Aedeagus: Distinct asymmetrical, with tooth-like structures.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD9CC22FF4BF97FFB83FE79.taxon	distribution	Distribution: Malaysia, Indonesia (Sumatra, Java).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFD9CC22FF4BF97FFB83FE79.taxon	etymology	Etymology: We refer to the similarity of this species with T. cleroides.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDFCC20FF4BFAD9FD1FFB01.taxon	description	(Figs. 16, 41 – 42)	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDFCC20FF4BFAD9FD1FFB01.taxon	description	Paratypes: ♀. Tillicera soror, Schenkling, 1902, ZMAN type COLE. 1800.1, Type: Tillicera soror Schklg., Cotype, British Bootang, Maria Basti, L. Durel, ex Mus., Lugd. Bat. (1 ex., ZMAN); ♀. Tillicera soror, Schenkling, 1902, ZMAN type COLE. 1800.2, Type: Tillicera soror Schklg., coll. R. Oberthür, British Bootang, Maria Basti, 1899, ex Mus., Lugd. Bat. (1 ex., ZMAN). Others: British Bootang, Padong Himalaya, 1914 rec.: L. Durel, ex coll. R. Oberthur (2 ex., RGCM); Inde, Ex Museo, Mniszech, Coll. M. Pic (1 ex., MNHN); Assam, India, Ex Museo, E. Hintz, ex Coll. R. Oberthur (1 ex., MNHN); British Bootang, Maria Basti, 1900, ex Coll. R. Oberthur (2 ex., MNHN); British Bootang, Maria Basti, L. Durel, ex Coll. R. Oberthur (8 ex., MNHN); British Bootang, Maria Basti, 1899, ex Coll. R. Oberthur (18 ex., MNHN); British Bootang, Maria Basti, L. Durel, Coll. M. Pic (1 ex., MNHN); Maria Basti, British Bootang, ex Coll. R. Oberthur (3 ex., MNHN); British Bootan, Padong, L. Durel, 1914, ex Coll. R. Oberthur (6 ex., MNHN); British Bootan, Padong, L. Durel 1914, Coll. A. Kuwert, Coll. R. Oberthur (3 ex., MNHN); Pedong, A. Desgodins, ex Coll. R. Oberthur (1 ex., MNHN); Pedong, A. Desgodins, Coll. M. Pic (1 ex., MNHN); British Bootang, L. Durel, 1898, ex Coll. R. Oberthur (20 ex., MNHN); British Bootang, Sakiou, L. Durel, ex Coll. R. Oberthur (2 ex., MNHN); Bhoutan, Anglais, R. Oberthur 1900 (1 ex., MNHN); Rongli, 26. – 27.7.1900, ex Coll. R. Oberthur (4 ex., MNHN); Khasis, 7.1894, Nat. Coll, ex Coll. R. Oberthur (1 ex., MNHN); Khasis, 7.1894, Nat. Coll, Coll. M. Pic (1 ex., MNHN); Gnatong, 1. – 2.7.1900, ex Coll. R. Oberthur (1 ex., MNHN); Ex Museo, Parry, ex Coll. R. Oberthur (3 ex., MNHN); Nepal: Kosi - # 13, Depitar 27 ° 27 ’ N, 87 ° 17 ’ E to Barabishe, 27 ° 26 ’ N, 87 ° 18 ’ E, 1250 – 560 m, 12.6.01, NHMB Basel, expedition to Nepal, 2001 (2 ex., BMNH); Nepal: Kosi - # 9 a, Chichilla, 1900 – 2000 m, 27 ° 28 ’ N, 87 ° 14 ’ E, 3. – 5.6.2001, NHMB Basel, expedition to Nepal, May / June 2001 (1 ex., NHMB); India, W Bengalen, 1000 m, Darjeeling Distr., Kurseong, Malloutar Umg. 6. – 14.7., leg. N. Dangal 1989 (1 ex., NMEG); Nepal 469 Myagdi Distr., Myagdi Khola S Boghara, 1400 m, 27. – 28.5.1995, Martens & Schawaller (1 ex., SMNS); 413 Sankhua Sabha Distr., Arun Valley betw. Hurure and Chichila, tree, rich cultural land, 2000 m, 17.6.1988 Martens & Schawaller leg., Nepal-Expedition, Jochen Martens (1 ex., SMNS); British Bootang, Maria Basti, 1899, ex Mus. R. Oberthur 1922 (1 ex., ZMAN); British Bootang, Maria Basti, L. Durel 1899, ex Mus. R. Oberthur 1922 (2 ex., ZMAN); British Bootang, Maria Basti, L. Durel, ex Mus. R. Oberthur 1922 (1 ex., ZMAN); Gopaldhara, Darjeeling, 3440 – 4720 ft., H. Stevens, from Foliage, 1925 from, Brit. Mus. (1 ex., ZMAN); Gopaldhara, Nepal-Sikhim Frontier, H. Stevens 16.9.18, 1925 from, Brit. Mus. (1 ex., ZMAN); Gopaldhara, Nepal-Sikhim Frontier, H. Stevens 1920, 1925 from Brit. Mus. (1 ex., ZMAN); Gopaldhara, Bw., Darjiling, 4720 ft. - 19., H. Stevens, 1925 from Brit. Mus. (1 ex., ZMAN); Ex Chinde, Deprepani 6000, Darjeeling, Bengal, J. C. M. Gardner, 19.4.1930, R. R. D. 22, B. C. R. 483, Cage 192 (1 ex., ZMAN); Zool. Surv. Ind., Sureil, 5000 ft., Darjiling dist. E. Himalayas, 11. – 31.10.17, N. A. & F. G., 1937, Retained from Indian Museum Calcutta (1 ex., ZMAN); Zool. Surv. Ind., Above Tura, Garo hills, Assam, 3500 – 3900 ft., 15.7. – 30.8.17, S. Kemp., on fallen tree, 1937, Retained from Indian Museum Calcutta (1 ex., ZMAN); Tonkin, Tugun, Quang, 5.1915, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN). Length: 9 – 13 mm (101 specimens) Collecting period: April – October, December Head: Black, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; distributed with fine punctation; cranium vested with dense, predominantly pale, setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark-brown to black, A 1 to A 4 sometimes slightly paler, with depressed pale and dark setae; antennomeres triangularly dilated from A 6 or A 7 onwards, A 10 broader than long, or as long as wide. Pronotum: Red-brown to brown, surface anterior to transverse depression mostly black; surface anterior to transverse depression finely and diffusely punctate, posterior of transverse depression more coarsely and irregularly punctate; vested with anteriorly directed and erect dark and pale setae; length: width ratio 1.14: 1 (broadest at middle). Scutellum: Brown, with pale setae, finely punctate. Elytra: Black; basal third with a red-brown macula (side margins bordered black), margined posteriorly by two yellowish pigmented transverse maculae (spanning from the lateral margin to mid-way towards suture) with dense, pale, depressed setae, between these maculae with a semicircular sickle-like pattern of pale setae, apical third with a pale yellowish pigmented transverse fascia (not always pigmented towards suture), with dense, pale, depressed setae; basal third regularly and deeply punctate with 6 to 7 deep rows, only sometimes extending towards posterior transverse fascia, the black part with fine, dense, punctation, diameter of punctures larger than interstices, apex densely and finely punctate; basal third with longer, erect, pale and dark setae, middle part predominantly with dark, posteriorly directed and semi-decumbent, setae, apex with dark setae gradually more erect approaching apices; length: width ratio 2.24: 1. Legs: Brown to black, base sometimes paler, with dense, erect, pale setae; tibiae with longitudinal carina. Ventral surface: Abdomen black to brown, thorax brown, with pale, depressed setae; 5 th ventrite of males deeply emarginated. Aedeagus: Asymmetrical, tooth-like structures distinct, large parameres.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDFCC20FF4BFAD9FD1FFB01.taxon	distribution	Distribution: Nepal, Bhutan, India.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDFCC20FF4BFAD9FD1FFB01.taxon	diagnosis	Differential diagnosis to T. auratofasciata and T. javana: The antennae of T. soror are triangularly (less stronger) dilated from A 6 onwards; besides differences in punctation, also the coloration is different; the brown basal macula of T. soror is smaller, and less precisely bordered than that of T. auratofasciata and T. javana; parameres and pygidium also differ.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDCCC1EFF4BFF05FAADFE79.taxon	description	(Figs. 17, 43 – 44) Specimens examined: Holotype: ♂. N-Vietnam, Tonkin, Cuc-Phuong NP 2. - 12.5.1991, leg. E. Jendek, coll. R. Gerstmeier (RGCM). Paratypes: Tonkin, (Coll. Ph Francois), Coll. L. Bedel 1922 (1 ex., MNHN); Vietnam, Tam dao, 27.5. – 2.6.1986, Vinh phu prov., lgt. J. Rybnícek (1 ex., RGCM). Length: 10 mm (3 specimens). Collecting period: May. Head: Black, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus brown to black; distributed with fine and diffuse punctation; cranium vested with short, erect, golden setae; head including eyes slightly broader than anterior width of pronotum. Antennae: Dark brown to black, A 1 to A 2 or A 3 paler, predominantly vested with brown, depressed setae; antennomeres triangularly dilated from A 4 onwards, A 10 broader than long or as long as wide. Pronotum: Unicoloured red-brown to brown or black anterior to transverse depression; surface finely and diffusely punctate, coarser behind transverse depression; vested with anteriorly directed, short, erect, golden and dark setae; length: width ratio 1.07: 1 (broadest at middle). Scutellum: Brown, finely punctate, vested with golden setae. Elytra: Black, base red-brown to brown, basal third with a yellowish pigmented transverse fascia, vested with dense, depressed, golden setae, apical third with a pale pigmented transverse fascia, vested with dense, depressed, golden setae, not reaching the lateral margin; basal half regularly and deeply punctate, punctation extending towards posterior transverse fascia, punctations bearing tubercles, diameter of punctures larger than interstices, apex densely and finely punctate; basal red-brown part with erect, golden and dark setae and shorter, depressed, golden setae, middle part with posteriorly directed and semi-decumbent, dark setae, apex longer golden and dark setae; length: width ratio 2.04: 1. Legs: Brown, femora paler, but sometimes with a dark stripe, with erect, golden setae; tibiae with longitudinal carina. Ventral surface: Abdomen black to brown, thorax brown, with pale, depressed setae. Aedeagus: Very long and narrow, without tooth-like structures distinct (even when viewed at 500 x magnification).	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDCCC1EFF4BFF05FAADFE79.taxon	distribution	Distribution: Vietnam.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
03EFA657FFDCCC1EFF4BFF05FAADFE79.taxon	etymology	Etymology: Named after its region of occurrence; Tonkin is the former term for North Vietnam.	en	Gerstmeier, Roland, Bernhard, Nadine (2010): Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1-34, DOI: 10.11646/zootaxa.2359.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2359.1.1
