identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EFF252F3007F6714BFA356FB7EFE18.text	03EFF252F3007F6714BFA356FB7EFE18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachinus Weber 1801	<div><p>Brachinus Weber, 1801</p><p>There are a total of 4 species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). Three species belong to sg. Metabrachinus Jeannel, 1949 and one as incertae sedis.</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6714BFA356FB7EFE18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA38FFF2CFCE7.text	03EFF252F3007F6714BFA38FFF2CFCE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachinus Weber 1801	<div><p>Brachinus (sg. incertae) apicalis Erichson, 1843</p><p>Distribution in Angola (Provinces): 1) Benguela.</p><p>Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227/8) (BENGUELA), 3.IV.1998, 1♀, PF, A. Neves leg., ASC.</p><p>Remarks. A species found in D. R. of the Congo, Zimbabwe and Angola (Liebke 1934). The singleton female adult was collected by pitfall trapping in open bush habitat with sparse grassy vegetation on sandy-clayey yellowreddish soil in the outskirts of Lobito together with some other ground beetles [ Calosoma (Calosoma) senegalense Dejean, 1831, Parophonus (Hyparpalus) tomentosus (Dejean, 1829), Anthia (s. str.) cinctipennis actaeon Erichson, 1843].</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6714BFA38FFF2CFCE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA442FAE0F90A.text	03EFF252F3007F6714BFA442FAE0F90A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calosoma Weber 1801	<div><p>Calosoma Weber, 1801</p><p>This genus was not recorded in the catalogue by Ferreira (1965), but there are four species reported to Angola (Insectoid.info 2017). The nomenclature here adopted follows Häckel (2013) and Häckel et al. (2016).</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6714BFA442FAE0F90A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA1DFFE72FB90.text	03EFF252F3007F6714BFA1DFFE72FB90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Styphlomerus Chaudoir in Putzeys 1875	<div><p>Styphlomerus Chaudoir in Putzeys, 1875</p><p>There is only one polytypic species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6714BFA1DFFE72FB90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA6F0FEF1FA28.text	03EFF252F3007F6714BFA6F0FEF1FA28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Styphlomerus ludricus (Erichson 1843)	<div><p>Styphlomerus (s. str.) ludricus ludricus (Erichson, 1843)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.</p><p>Material examined. Cachingues-Chipica (13 ° 10´15.74´´ S, 16 ° 45´48.17´´ E, 1649 m alt., 281) (BIÉ), 2.IV.2014, 1♂, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A polymorphic species widely distributed throughout western and central Africa (see Serrano 2008, 2009 and references herein). Adults were sampled in a secondary open forest. It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6714BFA6F0FEF1FA28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6614BFA57BFE91FEFF.text	03EFF252F3007F6614BFA57BFE91FEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calosoma (Calosoma) senegalense Dejean 1831	<div><p>Calosoma (Calosoma) senegalense Dejean, 1831</p><p>Distribution in Angola (Provinces): 1) Unknown; 2) Benguela. Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227/8) (BENGUELA), 3.IV.1998, 1♀, PF, A. Neves leg., ASC.</p><p>Remarks. A species widespread throughout Africa south of Sahara, Cape Vert Islands and Madagascar (Häckel et al. 2016). It was already cited to Angola (Häckel et al. 2016, Insectoid.info 2017), but without exact data. The single female adult was collected by pitfall trapping in an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Lobito together with some other ground beetles (see B. apicalis remarks).</p></div>	https://treatment.plazi.org/id/03EFF252F3007F6614BFA57BFE91FEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA3E7FC8FFC88.text	03EFF252F3017F6614BFA3E7FC8FFC88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calosoma (Calosoma) planicolle Chaudoir 1869	<div><p>Calosoma (Calosoma) planicolle Chaudoir, 1869</p><p>Distribution in Angola (Provinces): 1) Unknown; 2) Luanda.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.25&amp;materialsCitation.latitude=-8.833333" title="Search Plazi for locations around (long 13.25/lat -8.833333)">Material</a> examined. Luanda (8° 50´S, 13° 15´E, 70 m alt., 89) (LUANDA), 27.IV.1974, 1♀, DO , A. Serrano leg., ASC; Luanda near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.083333&amp;materialsCitation.latitude=-8.983334" title="Search Plazi for locations around (long 13.083333/lat -8.983334)">Motor</a> racing circuit (8° 59´S, 13° 05´E, 50 m alt., 89) (LUANDA), 12.VI.2016, 1♀, DO, R. Capela leg., ASC .</p><p>Remarks. A species widespread thoughout central, southern and eastern Africa (Angola, Botswana, D. R. of the Congo, Zambia, R. of South Africa, Lesotho, Swaziland, Zimbabwe, Malawi, Uganda, Mozambique, Tanzania, Kenya, Ethiopia, Somalia) and Madagascar (Häckel et al. 2016). Like the precedent species it was already known to Angola (Häckel et al. 2016, Insectoid.info 2017), but without any recorded locality. The first female adult was found in an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Luanda and the second on ground during night near a public light.</p></div>	https://treatment.plazi.org/id/03EFF252F3017F6614BFA3E7FC8FFC88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA1A2FF13FBEB.text	03EFF252F3017F6614BFA1A2FF13FBEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Distichus Motschulsky 1858	<div><p>Distichus Motschulsky, 1858</p><p>There is only one species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3017F6614BFA1A2FF13FBEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA6DBFB51FA04.text	03EFF252F3017F6614BFA6DBFB51FA04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Distichus picicornis (Dejean 1831)	<div><p>Distichus (s. str.) picicornis (Dejean, 1831)</p><p>Distribution in Angola (Provinces): 1) Unknown; 2) Luanda.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.166269&amp;materialsCitation.latitude=-9.315394" title="Search Plazi for locations around (long 13.166269/lat -9.315394)">Barra do Cuanza</a> (09° 18´55.42´´ S, 13° 09´58.57´´ E, 11 m alt., 107) (LUANDA), 28.IV.2014, 4♂, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widely spread throughout all Africa south of Sahara (e.g. Ferreira 1965). This author only refers to the presence of the species in Angola, without specifying localities. Adults were sampled close to River Cuanza by light trapping within a riparian forest. It is a new record for the Luanda Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3017F6614BFA6DBFB51FA04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA47DFDB5F965.text	03EFF252F3017F6614BFA47DFDB5F965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites Fabricius 1775	<div><p>Scarites Fabricius, 1775</p><p>There are seven species in this genus, two of them polytypic [ Scarites (Orientolobus) lucidus (Chaudoir, 1880) and Scarites (Scarites) tenebricosus Dejean, 1831] known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). Four species belong to Scarites (s. str.), two to sg. Orientolobus Dostall, 1996 and one to sg. Parallelomorphus Motschulsky, 1849 .</p></div>	https://treatment.plazi.org/id/03EFF252F3017F6614BFA47DFDB5F965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6514BFA55EFC7FFCAA.text	03EFF252F3017F6514BFA55EFC7FFCAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites (Orientolobus) lucidus subsp. strigiceps Quendenfeldt 1883	<div><p>Scarites (Orientolobus) lucidus strigiceps Quendenfeldt, 1883</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Malanje; 2) Huambo, Bié.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.450277&amp;materialsCitation.latitude=-12.281388" title="Search Plazi for locations around (long 15.450277/lat -12.281388)">Bembua-Camenhe</a> (12° 16´53´´ S, 15° 27´01´´ E, 1686 m alt., 231) (HUAMBO), 1. XI.2015, 3 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.789444&amp;materialsCitation.latitude=-13.556389" title="Search Plazi for locations around (long 16.789444/lat -13.556389)">Chitembo</a> (6 km SE) (13° 33´23´´ S, 16° 47´22´´ E, 1586 m alt., 302) (BIÉ), 9. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.983055&amp;materialsCitation.latitude=-13.59611" title="Search Plazi for locations around (long 16.983055/lat -13.59611)">Chingueia-Mandongue</a> (13° 35´46´´ S, 16° 58´59´´ E, 1648 m alt., 302) (BIÉ), 9. XI.2015, 2 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Mumbué (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.32389&amp;materialsCitation.latitude=-13.828055" title="Search Plazi for locations around (long 17.32389/lat -13.828055)">Cuanza</a> riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 8‒17. XI.2015, 1 ♂, 1♀, PF, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.4&amp;materialsCitation.latitude=-14.010278" title="Search Plazi for locations around (long 17.4/lat -14.010278)">Catota</a> (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8‒ 17. XI.2015, 1 ♂, PF, 12. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species spread throughout Democratic Republic of the Congo, Zimbabwe, Angola, Tanzania and Mozambique (Basilewsky, 1955). Adults were found within secondary open forest on reddish soil together with some adults of tiger and ground beetles [Bembua-Camenhe: Prothyma (s. str.) erythropyga erythropyga (Putzeys, 1880), Dromica (Foveodromica) spp., Prothymidia angusticollis angusticollis (Boheman, 1848), Cylindera agualusai Serrano &amp; Capela, 2015, Graphipterus albomarginatus Quedenfeldt, 1883] or on sandy whitish soil [Chitembo: Trichotaenia nzingae Oesterle, Serrano &amp; Capela, 2015, Lophyra (Stenolophyra) uncivittata (Quedenfeldt, 1883); Chingueia-Mandongue: Dromica fredericoi Serrano &amp; Capela, 2015, T. nzingae, Elliptica muata f. parallelestriata (W. Horn, 1923), Graphipterus congoensis lundanus Basilewsky, 1977 and Cypholoba rohani (Bénard, 1921); Mumbué: Dromica pentheri W. Horn, E. muata f. parallelestriata, Graphipterus andersoni Chaudoir, 1870, Anthia (Odontanthia) convexipennis Putzeys, 1880; Catota: D. fredericoi, Trichotaenia pepetela Serrano &amp; Capela, 2015, Chlaenius (Pachydinodes) bipustulatus Boheman, 1848, Abacetus pseudoangolensis, Siopelus (Neosiopelus) wrasei Facchini, 2012, Graphipterus andersoni and Atractonotus puncticollis Schle &amp; Heinz, 2013]. It is a new record for the Huambo and Bié Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3017F6514BFA55EFC7FFCAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA11BFD94F9CC.text	03EFF252F3027F6514BFA11BFD94F9CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites (Orientolobus) vilhenai Basilewsky 1955	<div><p>Scarites (Orientolobus) vilhenai Basilewsky, 1955</p><p>Distribution in Angola (Provinces): 1) Huambo; 2) Malanje.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.356112&amp;materialsCitation.latitude=-13.433056" title="Search Plazi for locations around (long 15.356112/lat -13.433056)">Catata-Nova Monção</a> (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6. XI.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.9783325&amp;materialsCitation.latitude=-9.064167" title="Search Plazi for locations around (long 15.9783325/lat -9.064167)">Kalandula</a> (1,5 km N) (09° 03´51´´ S, 15° 58´42´´ E, 1126 m alt., 112) (MALANJE), 18. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Cambamba (Catoio- <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.009167&amp;materialsCitation.latitude=-9.245" title="Search Plazi for locations around (long 17.009167/lat -9.245)">Quela</a>) (09° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Mutubanzamba (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.527777&amp;materialsCitation.latitude=-9.795834" title="Search Plazi for locations around (long 16.527777/lat -9.795834)">Cangandala</a>) (09° 47´45´´ S, 16° 31´40´´ E, 1136 m alt., 132) (MALANJE), 21. XI.2015, 2 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Carima (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.700277&amp;materialsCitation.latitude=-9.583333" title="Search Plazi for locations around (long 15.700277/lat -9.583333)">Cacuso-Pungandongo</a>) (09° 35´00´´ S, 15° 42´01´´ E, 1117 m alt., 130) (MALANJE), 23. XI.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An Angolan scaritid beetle endemic species whose description was based on one single female from Monte Moco (Huambo province). Our specimens conform well to the Basilewsky (1955) description. Adult specimens were found whitin secondary open forests on sandy whitish soil (first locality) or on reddish soil (Malanje localities) together with some adults of tiger and ground beetles [Catata-Nova Monção: Euryarthron reticostatum (Wellman &amp; W. Horn, 1908), Dromica (Foveodromica) sp., P. angusticollis, Trichodela nubifera (Quedenfeldt, 1883), Lophyra (Stenolophyra) saraliensis saraliensis (Guérin-Méneville, 1849), Lophyra (Bothryolophyra) wellmani (W. Horn, 1907), Chlaenius ovalipennis, Euleptus intermedius Péringuey, 1896, Orthotrichus patroboides (Murray, 1859), Orthogonius sp. 1, Netrodera vethi (Bates, 1889), Cypholoba graphipteroides monardi Burgeon, 1935 and Anthia (Odontanthia) kleinfeldi Puchner, 2013; Malanje localities: Dromica (Foveodromica) spp., P. angusticollis, T. nubifera, Epitrichodes villosus (Putzeys, 1880), Lophyra (Stenolophyra) infuscatula (W. Horn, 1915), L. saraliensis, L. wellmani, Graphipterus ellipticus Burgeon, 1928]. It is a new record for the Malanje Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3027F6514BFA11BFD94F9CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA4B5FA9AF865.text	03EFF252F3027F6514BFA4B5FA9AF865.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites tenebricosus subsp. fatalis Harold 1879	<div><p>Scarites (s. str.) tenebricosus fatalis Harold, 1879</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 31.X.2014, 1♂, 2.XI.2014, 3♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A subspecies spread throughout Angola and D. R. Congo (Katanga) (Bäninger 1938). Adults were found in activity on sandy whitish soil within a secondary open forest during night together with other ground beetles (e.g. Systolocranius tibialis Lecordier, 1972, C. bipustulatus, Termophilum massilicata Guérin-Méneville, 1845).</p></div>	https://treatment.plazi.org/id/03EFF252F3027F6514BFA4B5FA9AF865	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA31DFCBEFC86.text	03EFF252F3037F6414BFA31DFCBEFC86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clivina capensis Kult 1959	<div><p>* Clivina capensis Kult, 1959</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Satchijamba (10 km E) (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 283) (BIÉ), 7.IV.2014, 1♀, VM, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species only reported to South Africa (Cape Province) (Kult 1959). The specimen was identified using Kult`s (1959) key, taking into account that our specimen is dark ferruginous, shining (elytra without microsculture), with frons impunctate, proepisternae finely punctate, protibiae sulcate with 3rd spine almost indistinct, prothorax with transverse wrinkles and group of punctures on each side and mesotibiae without denticle. The unique feature that does not agree with C. capensis description is the antennal antenomers 4 to10: moniliform in C. capensis, but subelongate in our specimen. For the present, we consider that this specimen belongs to this species with some reserves. The single female specimen was found in open secondary forest and collected by means of light trapping. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6414BFA31DFCBEFC86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA2EBFC5CFEA4.text	03EFF252F3037F6414BFA2EBFC5CFEA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clivina Latreille 1802	<div><p>Clivina Latreille, 1802</p><p>There are nine species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017), belonging seven to Clivina (s.str.) and two to sg. Eoclivina Kult, 1959 .</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6414BFA2EBFC5CFEA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA6F6FAB7F9BA.text	03EFF252F3037F6414BFA6F6FAB7F9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Morion guineensis Imhoff 1843	<div><p>Morion guineensis Imhoff, 1843</p><p>Distribution in Angola (Provinces): 1) Cunene; 2) Luanda, Bié.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.166269&amp;materialsCitation.latitude=-9.315394" title="Search Plazi for locations around (long 13.166269/lat -9.315394)">Barra do Cuanza</a> (09° 18´55.42´´ S, 13° 09´58.57´´ E, 11 m alt., 107) (LUANDA), 28. IV.2014, 1 ♀, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.18396&amp;materialsCitation.latitude=-13.74625" title="Search Plazi for locations around (long 17.18396/lat -13.74625)">Satchijamba-Somakwanza</a> (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 283) (BIÉ), 2. XI.2014, 1 ♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout almost all Africa south of Sahara, including S. Tomé Island (e.g. Straneo 1959, Ferreira 1965, Serrano 1995). Adults were collected by light trapping within a riparian forest (Barra do Cuanza) and an open secondary forest (Satchijamba-Somakwanza). In the later locality the adults were collected together with ant nest beetles [ Paussus (Shuckardipaussus) vanrooni Wasmann, 1922, Paussus (Katapaussus) chappuisi Reichensperger, 1938] (see Serrano 2015a). It is a new record for the Luanda and Bié Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6414BFA6F6FAB7F9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA1A0FA71FB8D.text	03EFF252F3037F6414BFA1A0FA71FB8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Morion Latreille 1810	<div><p>Morion Latreille, 1810</p><p>There is one species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6414BFA1A0FA71FB8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA42BFBECF964.text	03EFF252F3037F6414BFA42BFBECF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stereostoma Murray 1857	<div><p>Stereostoma Murray, 1857</p><p>There are five species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017), belonging one to Stereostoma (s.str.) and four to sg. Stereodema Chaudoir, 1872 .</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6414BFA42BFBECF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6B14BFA55CFD06FEDA.text	03EFF252F3037F6B14BFA55CFD06FEDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stereostoma (Stereodema) angolense Muller 1940	<div><p>Stereostoma (Stereodema) angolense Müller, 1940</p><p>Distribution in Angola (Provinces): 1) Huambo; 2) Bi.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described on one specimen from Bailundo (Huambo province) (Müller 1940) and known also from D.R. of the Congo (Straneo 1952, 1991). The single adult was sampled within an open secondary forest by light trapping together with other ground beetles (e.g. Stereostoma guineense Müller, 1940, Brachyodes pseudoguineensis Lecordier, 1990, Parophonus escheri (Dejean, 1831), Siopelus harpaloides (Guérin-Méneville, 1847), Siopelus natalicus Péringuey, 1896, Axinotoma latipalpis Basilewsky, 1968 and Meladroma informicollis Liebke, 1928). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3037F6B14BFA55CFD06FEDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA682FEFEF962.text	03EFF252F30C7F6B14BFA682FEFEF962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caelostomus (Drimostomellus) intermedius (Chaudoir 1878)	<div><p>Caelostomus (Drimostomellus) intermedius (Chaudoir, 1878)</p><p>Distribution in Angola (Provinces): 1) Huíla, Huambo; 2) Malanje, Kwanza Sul.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X‒ 2. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Mutubanzamba (Cangandala) (09° 47´45´´ S, 16° 31´40´´ E, 1136 m alt., 132) (MALANJE), 21. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Alto Ventura (Colé- Calulo) (09° 59´42´´ S, 14° 50´32´´ E, 1116 m alt., 128) (KWANZA SUL), 4. XII.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC;</p><p>Remarks. A species with a distribution throughout Angola, D. R. of the Congo, Zimbabwe, Tanzania (Zanzibar) and Mozambique (Straneo 1942, 1952). In the first locality the adult specimen was found in a wall during night near a public light. In the second locality, the adult was collected within the litter which covered the soil and, in the third locality, the adults were found under bark of trees together with adults of other Coleoptera families (e.g. Endomychidae, Trogossitidae, Tenebrionidae). It is a new record for the Malanje and Kwanza Sul Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F30C7F6B14BFA682FEFEF962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA668FB27FB21.text	03EFF252F30C7F6B14BFA668FB27FB21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caelostomus MacLeay 1825	<div><p>Caelostomus MacLeay, 1825</p><p>There are four species in this genus known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017), belonging two to the Caelostomus (s.str.) and the other two to sg. Drimostomellus Jeannel, 1948 .</p></div>	https://treatment.plazi.org/id/03EFF252F30C7F6B14BFA668FB27FB21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA3CBFAAFFC78.text	03EFF252F30C7F6B14BFA3CBFAAFFC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stereostoma (Stereodema) guineense Muller 1940	<div><p>Stereostoma (Stereodema) guineense Müller, 1940</p><p>Distribution in Angola (Provinces): 1) Huambo, Lunda Norte; 2) Bié, Kwanza Sul.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.75565&amp;materialsCitation.latitude=-13.515339" title="Search Plazi for locations around (long 16.75565/lat -13.515339)">Chitembo</a> (13° 30´55.22´´ S, 16° 45´20.34´´ E, 1628 m alt., 302) (BIÉ), 8. IV.2014, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.900122&amp;materialsCitation.latitude=-13.60487" title="Search Plazi for locations around (long 16.900122/lat -13.60487)">Chitembo</a> (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30. X.2014, 1 ♂, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.549167&amp;materialsCitation.latitude=-12.222221" title="Search Plazi for locations around (long 15.549167/lat -12.222221)">Alto Hama</a> (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31. X.2015, 3 ♂, 3♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.551945&amp;materialsCitation.latitude=-12.853056" title="Search Plazi for locations around (long 15.551945/lat -12.853056)">Caála</a> (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.549167&amp;materialsCitation.latitude=-12.222221" title="Search Plazi for locations around (long 15.549167/lat -12.222221)">Calulo</a> (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (KWANZA SUL), 27.XI‒5. XII.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described on one specimen from Bailundo also (Huambo Province) (Müller 1940) and widespread throughout western and central Africa (Straneo 1952, 1991). The single male adult collected in the second locality was sampled by light trapping within an open secondary forest together with some other ground beetles (e.g. see S. angolense remarks). The remaining adults were found in activity on ground in the road edges during night, probably attracted to the public lights. It is a new record for the Bié and Kwanza Sul Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F30C7F6B14BFA3CBFAAFFC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA543FF13F80C.text	03EFF252F30C7F6B14BFA543FF13F80C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Strigomerus Chaudoir 1872	<div><p>Strigomerus Chaudoir, 1872</p><p>There is one species until now known from Angola in this genus (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F30C7F6B14BFA543FF13F80C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA051FEC9FCAE.text	03EFF252F30D7F6A14BFA051FEC9FCAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus Dejean 1828	<div><p>Abacetus Dejean, 1828</p><p>A very speciose genus with thirty two species known from Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). Within the genus there are two species belonging to sg. Abacetillus Straneo, 1942, six to the nominal genus, seven to sg. Astigis Rambur, 1838, one to sg. Bisulcillus Straneo, 1942, nine to sg. Caricus Motschulsky, 1866, four to sg. Distrigodes Motschulsky, 1864, one to sg. Triaenabacetus Straneo, 1942 and two as Incertae sedis.</p></div>	https://treatment.plazi.org/id/03EFF252F30D7F6A14BFA051FEC9FCAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA116FE11FAEF.text	03EFF252F30D7F6A14BFA116FE11FAEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus distinctus Chaudoir 1878	<div><p>Abacetus (s. str.) distinctus Chaudoir, 1878</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Calulo (Faz. Klein) (10° 02´14´´ S, 14° 54´38´´ E, 1059 m alt., 147) (KWANZA SUL), 3. XII.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Calulo-Cabuta (9° 53´59´´ S, 14° 54´26´´ E, 831 m alt., 128) (KWANZA SUL), 29.XI‒6. XII.2015, 6 ♂, 2♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout Angola, D. R. of the Congo, Zimbabwe and South Africa (Straneo 1958, 1959). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e.g. Dromica (Foveodromica) sp., Hipparidium interruptum (Fabricius, 1775), Abacetus optimus Péringuey, 1904, Disphericus tarsalis Bates, 1886, Craspedophorus magnicollis discrepans Basilewsky, 1987, Galerita procera capelai ssp. n.] and collected also by pitfall trapping in the same habitats. It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30D7F6A14BFA116FE11FAEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA7D7FAE3F908.text	03EFF252F30D7F6A14BFA7D7FAE3F908.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus natalensis Chaudoir 1869	<div><p>Abacetus (s. str.) natalensis Chaudoir, 1869</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Bié.</p><p>Material examined. Cachingues-Chipica (13° 10´15.14´´ S, 16° 45´33.52´´ E, 1668 m alt., 282) (BIÉ), 2.IV.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widely spread in Africa (e.g. Cape Vert, Ivory Coast, Ethiopia, Mozambique, Nigeria, Angola, Congo, D. R. of the Congo, Zimbabwe and Rwanda) (Straneo 1956, 1958, 1959, 1968). The single male was collected by light trapping within a secondary open forest. It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30D7F6A14BFA7D7FAE3F908	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA2EBFAA4FE70.text	03EFF252F30D7F6A14BFA2EBFAA4FE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Strigomerus sulcipennis (Dejean 1831)	<div><p>Strigomerus sulcipennis (Dejean, 1831)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Bié.</p><p>Material examined. Satchijamba (10 km E) (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 283) (BIÉ), 7.IV.2014, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a distribution throughout Angola, D. R. of the Congo, Zimbabwe and South Africa (Straneo 1942, 1958, 1959). The singleton adult was collected under the bark of a decomposing trunk together with Zygentoma, Formicidae and Tenebrionidae adults and the ant beetle Paussus (Klugipaussus) pseudoklugi Luna de Carvalho, 1963 within an open secondary forest (see Serrano 2015a). It is a new record for the Bié province.</p></div>	https://treatment.plazi.org/id/03EFF252F30D7F6A14BFA2EBFAA4FE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6914BFA578FCE7FE37.text	03EFF252F30D7F6914BFA578FCE7FE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus (Caricus) pseudangulanus Straneo 1952	<div><p>Abacetus (Caricus) pseudangulanus Straneo, 1952</p><p>Distribution in Angola (Provinces): 1) Cunene; 2) Bié.</p><p>Material examined. Mumbué (Cuanza riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 8‒ 17.XI.2015, 1♂, PF, A. Serrano &amp; R. Capela leg., ASC; Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8‒17.XI.2015, 2♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic ground beetle species of Angola from which is known so far the male and female types (Straneo 1952). Both adults were sampled through pitfall trapping. The Mumbué specimen was collected in sandy whitish soil within a mixed open herbaceous/bush habitat close to the riverhead together with adults of the tiger beetle E. muata f. parallelestriata and the ground beetles S. lucidus strigiceps, E. intermedius, Systolocranius tibialis and Anthia convexipennis Putzeys, 1880 . The Catota specimens were collected through pitfall trapping in sandy clayish reddish soil within a secondary open forest together with adults of some ground beetles (see S. lucidus remarks). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30D7F6914BFA578FCE7FE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA3AFFD95FC1C.text	03EFF252F30E7F6914BFA3AFFD95FC1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus (Abacetillus) optimus Peringuey 1904	<div><p>Abacetus (Abacetillus) optimus Péringuey, 1904</p><p>Distribution in Angola (Provinces): 1) Moxico; 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI.2015, 1 ♂, 2♀, 5. XII.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Mussende (Calulo) (9° 57´00´´ S, 14° 47´36´´ E, 905 m alt., 128) (KWANZA SUL), 28.XI‒5. XII.2015, 3 ♀, PF, A. Serrano &amp; R. Capela leg., ASC; Calulo-Cabuta (9° 53´59´´ S, 14° 54´26´´ E, 831 m alt., 128) (KWANZA SUL), 29.XI‒ 6. XII.2015, 8 ♂, 9♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout Angola, Zimbabwe, Mozambique and South Africa (Straneo 1958, 1959, 1991). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e.g. Dromica (Foveodromica) sp., A. distinctus, Disphericus tarsalis, C. magnicollis discrepans, G. procera capelai ssp. n.] and collected also by pitfall trapping in the same habitats. It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30E7F6914BFA3AFFD95FC1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA185FB2FFA91.text	03EFF252F30E7F6914BFA185FB2FFA91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abacetus (Distrigodes) monardianus Straneo 1952	<div><p>Abacetus (Distrigodes) monardianus Straneo, 1952</p><p>Distribution in Angola (Provinces): 1) Benguela, Moxico; 2) Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 2.XI.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic ground beetle species of Angola from which is known so far the male type (Straneo 1952) and a second specimen (unknown sex) from Calundo Lake (Straneo 1959). The specimen fits very well with the original description and was captured by light trapping within a secondary open forest together with ground and ant nest beetles ( M. guineensis, P. vanrooni, P. chappuisi). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30E7F6914BFA185FB2FFA91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA447FD78F830.text	03EFF252F30E7F6914BFA447FD78F830.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aristopus elisabethanus (Burgeon 1935)	<div><p>* Aristopus elisabethanus (Burgeon, 1935)</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 2.XI.2014, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species only reported to D. R. of the Congo (Straneo 1983). The specimen was identified using Straneo`s (1983) key and the specimen fits well the original description (Burgeon 1935). For the present, until the acquisition of new data, we consider that this specimen belongs to this species with some reserves. The single adult was collected under the bark of a decomposing trunk together with Zygentoma, Formicidae and Tenebrionidae adults. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F30E7F6914BFA447FD78F830	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA7F2FA66FA5F.text	03EFF252F30E7F6914BFA7F2FA66FA5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aristopus LaFerte-Senectere 1853	<div><p>Aristopus LaFerté-Sénectère, 1853</p><p>There are five species known from Angola in this genus (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F30E7F6914BFA7F2FA66FA5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA1DFFD9FFB90.text	03EFF252F30F7F6814BFA1DFFD9FFB90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Craspedophorus Hope 1838	<div><p>Craspedophorus Hope, 1838</p><p>There are ten species in this genus known from Angola (Alves 1963, Ferreira 1965, Häckel &amp; Farkač 2012, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F30F7F6814BFA1DFFD9FFB90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA6F0FCC9F9F5.text	03EFF252F30F7F6814BFA6F0FCC9F9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Craspedophorus magnicollis subsp. discrepans Basilewsky 1987	<div><p>Craspedophorus magnicollis discrepans Basilewsky, 1987</p><p>Distribution in Angola (Provinces): 1) Malanje, Huíla; 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 2.XII.2015, 1♂), 4.XII.2015, 1♂, 5.XII.2015, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. The nominal species was described from Angola (Malanje) (Quedenfeldt 1883) and has a distribution throughout Angola and D. R. of the Congo (Häckel &amp; Farkač 2012). Basilewsky (1987) described two subspecies, C. magnicollis discrepans from the Lulua region (D. R. of the Congo) and Zambia and C. magnicollis inquilinus from Mayidi (D. R. of the Congo), giving the drawings of aedeagus apex (dorsal view) of the three taxa. Interestingly our Calulo specimens present the aedeagus apex similar to the C. magnicollis discrepans (Basilewsky 1987, Fig. 4c), enlarging its distribution to Angola. Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e.g. Dromica (Foveodromica) sp., A. distinctus, A. optimus]. It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30F7F6814BFA6F0FCC9F9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6F14BFA4EEFC68FF47.text	03EFF252F30F7F6F14BFA4EEFC68FF47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Craspedophorus rikatlensis (Peringuey 1896)	<div><p>Craspedophorus rikatlensis (Péringuey, 1896)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.‒ 2. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout Angola, D. R. of the Congo, Burundi, Zambia and Mozambique (Basilewsky 1955, Häckel &amp; Farkač 2012, 2013a, Insectoid.info 2017), presenting some variability on the elytra maculae. Adults were found during night near public lights together with other ground beetles [e.g. S. guineense, Chlaenius cupreocinctus Reiche, 1847, Ch. tenuicollis (Fabricius, 1801), Progonochaetus planicollis (Putzeys, 1880), Agonidium alacre (Boheman, 1848), Agonidium natalense (Boheman, 1848), E. intermedius, Euleptus peringueyi Csiki, 1931]. It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30F7F6F14BFA4EEFC68FF47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA3C6FDC3FCE7.text	03EFF252F30F7F6814BFA3C6FDC3FCE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Disphericus tarsalis Bates 1886	<div><p>Disphericus tarsalis Bates, 1886</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Kwanza Sul.</p><p>Material examined. Mussende (Calulo) (9° 57´00´´ S, 14° 47´36´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI.2015, 1 ♂, DO, 28.XI‒5. XII.2015, 2 ♀, PF, A. Serrano &amp; R. Capela leg., ASC; Alto Ventura (Faz. Monte Café) (Calulo) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI.2015, 1 ♂, 5. XII.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic ground beetle species of Angola from which is known so far the type (Bates 1886) and a second specimen (unknown sex) from Dundo (Basilewsky 1955). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles (e.g. Dromica (Foveodromica) sp., A. distinctus, A. optimus, C. magnicollis discrepans) and also felt in pitfall traps in the same habitats. It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30F7F6814BFA3C6FDC3FCE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA2ADFD9FFEDE.text	03EFF252F30F7F6814BFA2ADFD9FFEDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Disphericus Waterhouse 1842	<div><p>Disphericus Waterhouse, 1842</p><p>There are two species in this genus known from Angola (Ferreira 1965, Häckel &amp; Farkač 2012, 2013a, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F30F7F6814BFA2ADFD9FFEDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA3E1FC3DFE4D.text	03EFF252F3087F6F14BFA3E1FC3DFE4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachyodes Jeannel 1949	<div><p>* Brachyodes Jeannel, 1949</p><p>A speciose genus in Africa (Lorenz 2005) but never recorded for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3087F6F14BFA3E1FC3DFE4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA036FB5AFCC2.text	03EFF252F3087F6F14BFA036FB5AFCC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachyodes pseudoguineensis Lecordier 1990	<div><p>* Brachyodes pseudoguineensis Lecordier, 1990</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a distribution throughout D. R. Congo and Burundi (Lecordier 1990). It can be segregated from B. guineensis (Chaudoir, 1882), to which is very close, by the smaller antennae and the unarmed mesosternum groove. The adult was sampled by light trapping within a secondary open forest together with some other ground beetles (see S. angolense remarks). It is a new genus/species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3087F6F14BFA036FB5AFCC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA426FC81F86A.text	03EFF252F3087F6F14BFA426FC81F86A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius Bonelli 1810	<div><p>Chlaenius Bonelli, 1810</p><p>A very speciose genus with fifty two species/subspecies known from Angola (Alves 1963, Ferreira 1965, Kirschenhofer 2007, Anichtchenko 2016, Insectoid.info 2017). Within this assemblage there are one species/ subspecies belonging to sg. Ocybatus LaFerté-Sénectère, 1851, four to sg. Lissauchenius MacLeay, 1825, three to sg. Pachydinodes Kuntzen, 1919, one to sg. Agostenus Waldheim, 1829, two to sg. Epomis Bonelli, 1810, two to sg. Rhysotrachelus Boheman, 1848, two to sg. Tomochilus LaFerté-Sénectère, 1851, one to sg. Pseudochlaeniellus Jeannel, 1949, eight to sg. Chlaeniostenus Bates, 1892, four to sg. Amblygenius LaFerté-Sénectère, 1851, six to the nominal genus, two to sg. Leptorembus Kolbe, 1889, one to sg. Macrochlaenites Kuntzen, 1919, seven to sg. Homalolachnus LaFerté-Sénectère, 1851, one to sg. Pleroticus Péringuey, 1896, two to sg. Paracallistoides Basilewsky, 1965 and five to sg. Callistoides Motschulsky, 1865 .</p></div>	https://treatment.plazi.org/id/03EFF252F3087F6F14BFA426FC81F86A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA1A3FCF5FBEC.text	03EFF252F3087F6F14BFA1A3FCF5FBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Systolocranius Chaudoir 1857	<div><p>Systolocranius Chaudoir, 1857</p><p>There are two species within this genus known for Angola (Ferreira 1965, Lecordier 1972, Insectoid.info 2017), of which one was sampled during our entomological trips.</p></div>	https://treatment.plazi.org/id/03EFF252F3087F6F14BFA1A3FCF5FBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA6D5FE3CF9BD.text	03EFF252F3087F6F14BFA6D5FE3CF9BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Systolocranius tibialis Lecordier 1972	<div><p>Systolocranius tibialis Lecordier, 1972</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 2. XI.2014, 3 ♂, 3♀, DO, A. Serrano &amp; R. Capela leg., ASC; Mumbué (Cuanza riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 8‒17. XI.2015, 1 ♂, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species spread throughout Angola and D. R. of the Congo (Katanga) (Lecordier 1972). Adults in the first locality were found in activity on sandy whitish soil within a secondary open forest during night together with other ground beetles [e.g. S. tenebrosus fatalis, C. (Pachydinodes) bipustulatus, T. massilicata] It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3087F6F14BFA6D5FE3CF9BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA38DFD88FCE5.text	03EFF252F3097F6E14BFA38DFD88FCE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Lissauchenius) assecla (Laferte-Senectere 1851)	<div><p>* Chlaenius (Lissauchenius) assecla (Laferté-Sénectère, 1851)</p><p>Distribution in Angola (Provinces): 2) Luanda.</p><p>Material examined. Luanda near Motor racing circuit (08° 59´S, 13° 05´E, 50 m alt., 89) (LUANDA), 12.VI.2016, 1♂, 1♀, DO, R. Capela leg., ASC.</p><p>Remarks. A species with a scattered distribution throughout Africa south of Sahara (Mauritania, Senegal, Ivory Coast, Togo, Mali, Chad, Democratic Republic of the Congo, Eritrea, Ethiopia, Kenya, Tanzania, Zimbabwe, South Africa), but known from West Asia (Oman) and some Atlantic Islands (Cape Verde, Saint Helena and São Tomé and Príncipe) also (Insectoid.info 2017). Adult specimens were found on a wall during night near a public light. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3097F6E14BFA38DFD88FCE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA1DEFB93F9BA.text	03EFF252F3097F6E14BFA1DEFB93F9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Lissauchenius) goryi Gory 1833	<div><p>Chlaenius (Lissauchenius) goryi Gory, 1833</p><p>Distribution in Angola (Provinces): 1) Lunda Norte?; 2) Malanje, Kwanza Sul.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.064444&amp;materialsCitation.latitude=-8.9375" title="Search Plazi for locations around (long 16.064444/lat -8.9375)">Kibundi</a> (15 km N Nambua) (8° 56´15´´ S, 16° 03´52´´ E, 1164 m alt., 95) (MALANJE), 19.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Alto Ventura (Faz. Monte Café) (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.836666&amp;materialsCitation.latitude=-9.9875" title="Search Plazi for locations around (long 14.836666/lat -9.9875)">Calulo</a>) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 30.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout Africa south of Sahara (Senegal, Gambia, Guinea, Ivory Coast, Cameroon, Burkina Faso, Chad, Democratic Republic of the Congo, Burundi, Rwanda, Zambia, Zimbabwe, Uganda, Kenya, Tanzania, Ethiopia, Eritrea, Yemen) (Insectoid.info 2017), but not recorded to Mozambique, South Africa and Namibia. It was reported for Angola by Alves (1963). However, the data given in this work (locality, date) for the species seems similar to the specimen which was identified as C. deyrollei (see the precedent species) by Basilewsky (1955). The female adult specimen (first locality) was found in the edges of a countryside road within an open secondary forest while the male adult specimen (second locality) was collected within the litter of a secondary rainforest. Both adults were found together with adults of some tiger and ground beetles [first locality: e.g. Dromica (Foveodromica) sp., A. optimus, A. distinctus, D. tarsalis, C. magnicollis discrepans, G. procera capelai n. ssp.; second locality: Dromica (Foveodromica) sp., L. wellmani]. These new data confirm this species’ presence in Angola and extends its distribution to Malanje and Kwanza Sul Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3097F6E14BFA1DEFB93F9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA2EBFE49FE14.text	03EFF252F3097F6E14BFA2EBFE49FE14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Ocybatus) deyrollei (LaFerte-Senectere 1851)	<div><p>Chlaenius (Ocybatus) deyrollei (LaFerté-Sénectère, 1851)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 5.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widespread throughout western and central Africa since Senegal to Angola (Basilewsky 1955, Insectoid.info 2017). The singleton adult male was found during night near a public light. It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3097F6E14BFA2EBFE49FE14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6D14BFA42BFBE7FF6A.text	03EFF252F3097F6D14BFA42BFBE7FF6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Pachydinodes) bipustulatus Boheman 1848	<div><p>Chlaenius (Pachydinodes) bipustulatus Boheman, 1848</p><p>Distribution in Angola (Provinces): 1) Huíla, Huambo; 2) Bié.</p><p>Material examined. Cruzeiro (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.902636&amp;materialsCitation.latitude=-12.778094" title="Search Plazi for locations around (long 15.902636/lat -12.778094)">Pedra do Alemão</a>) (12° 46´41.14´´ S, 15° 54´09.49´´ E, 1743 m alt., 232) (HUAMBO), 7. XI.2015, 1 ♂, 1♀, DO, A Serrano &amp; R. Capela leg., ASC ; Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 2. XI.2014, 3 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8‒17. XI.2015, 2 ♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A polymorphic ground beetle species with a distribution throughout Africa south of Sahara. The nominal subspecies seems restricted to the southern countries of Africa (Angola, Namibia, Zimbabwe, South Africa and Mozambique) (Insectoid.info 2017). Adults were found beneath litter (first locality) or in activity on sandy whitish soil within a secondary open forest (second locality) during night together with other ground beetles (e.g. S. tenebrosus fatalis, S. tibialis, T. massilicata). In the third locality the specimens were collected by pitfall trapping within a secondary open forest. It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3097F6D14BFA42BFBE7FF6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA682FBADF997.text	03EFF252F30A7F6D14BFA682FBADF997.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Chlaeniostenus) tenuicollis (Fabricius 1801)	<div><p>* Chlaenius (Chlaeniostenus) tenuicollis (Fabricius, 1801)</p><p>Distribution in Angola (Provinces): 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X‒ 2. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a known scattered distribution throughout Africa: Ethiopia, D. R. of the Congo and South Africa (Anichtchenko 2016, Insectoid.info 2017). Adults were found also on ground during night near public lights (see comments on C. rikatlensis remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F30A7F6D14BFA682FBADF997	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA40FFBB1F868.text	03EFF252F30A7F6D14BFA40FFBB1F868.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Chlaenius) obtusus subsp. tazieffi Basilewsky 1949	<div><p>* Chlaenius (Chlaenius) obtusus tazieffi Basilewsky, 1949</p><p>Distribution in Angola (Provinces): 2) Huambo, Bié.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Cachingues (13° 04´54´´ S, 16° 46´08´´ E, 1647 m alt., 282) (BIÉ), 11. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A subspecies known exclusively from D. R. of the Congo (Basilewsky 1949, Insectoid.info 2017). Our specimens fit very well the Basilewsky`description. Adults were found on ground during night near public lights (first locality) (see also C. rikatlensis remarks) and beneath litter on sandy whitish soil within an open secondary forest (second locality). It is a new species/subspecies record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F30A7F6D14BFA40FFBB1F868	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA35BFA66FCAC.text	03EFF252F30A7F6D14BFA35BFA66FCAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Pachydinodes) ovalipennis Quedenfeldt 1883	<div><p>Chlaenius (Pachydinodes) ovalipennis Quedenfeldt, 1883</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Malanje, Bié, Huambo.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 2. XI.2014, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Mutubanzamba (9° 47´45´´ S, 16° 31´40´´ E, 1136 m alt., 132) (MALANJE), 21. XI.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described from Quango based on one male of collections by Mechow (Quedenfeldt 1883). Our specimens fit very well the original description, including the figure 10 of the plate 3. There is a report of this species to Lunda by Monard et al. (1956) (sub Chlaenius ovalipennis Gaf. ?). Surprisingly the species is not reported for Angola in the Ferreira (1965)`s catalogue or even for the neighbouring countries (e.g. D. R. of the Congo, Zambia, Namibia) in Anichtchenko (2016) and Insectoid.info (2017) sites. Adults were found in all localities beneath litter within secondary open forests together with some tiger and ground beetles (Catata-Nova Monção and Mutubanzamba: see S. vilhenai remarks). It is a new record for Malanje, Huambo and Bié Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F30A7F6D14BFA35BFA66FCAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA115FBA9FB21.text	03EFF252F30A7F6D14BFA115FBA9FB21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Tomochilus) cupreocinctus Reiche 1847	<div><p>Chlaenius (Tomochilus) cupreocinctus Reiche, 1847</p><p>Distribution in Angola (Provinces): 1) Huambo, Lunda Norte.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♂, 1♀, 4.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered distribution in Africa: Angola, Zimbabwe, Ethiopia and Eritrea (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). Adults were found on ground during night near public lights together with other ground beetles (e.g. see C. rikatlensis and P. incrassatus remarks).</p></div>	https://treatment.plazi.org/id/03EFF252F30A7F6D14BFA115FBA9FB21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA5FDFBFDF851.text	03EFF252F30B7F6C14BFA5FDFBFDF851.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistomimus Chaudoir 1872	<div><p>* Callistomimus Chaudoir, 1872</p><p>A genus very speciose in Africa (Lorenz 2005), but never recorded for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA5FDFBFDF851	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA3A9FAE1FCAD.text	03EFF252F30B7F6C14BFA3A9FAE1FCAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Callistoides) maculiceps subsp. maculiceps Boheman 1848	<div><p>Chlaenius (Callistoides) maculiceps maculiceps Boheman, 1848</p><p>Distribution in Angola (Provinces): 1) Huambo; 2) Kwanza Sul.</p><p>Material examined. Calulo (Faz. Klein) (10° 02´14´´ S, 14° 54´38´´ E, 1059 m alt., 147) (KWANZA SUL), 1.XII.2015, 1♂, 3.XII.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. The species was reported for Angola by Basilewsky (1955) under the name C. hessei Basylewsky, 1949 . The nominal species is widespread throughout Angola, D. R. of the Congo, Zambia, Zimbabwe, South Africa and Tanzania (Basilewsky 1955, Anichtchenko 2016, Insectoid.info 2017). Adults were found in the edges of small rain pounds, running quickly when disturbed. It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA3A9FAE1FCAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA116FD77FBC7.text	03EFF252F30B7F6C14BFA116FD77FBC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Callistoides) pulchellus Boheman 1848	<div><p>Chlaenius (Callistoides) pulchellus Boheman, 1848</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♀, 5.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout all Africa south of Sahara (Basilewsky 1955, Anichtchenko 2016, Insectoid.info 2017). Adults were found on ground during night near public lights (see also C. rikatlensis remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA116FD77FBC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA6BFFAA6F9BC.text	03EFF252F30B7F6C14BFA6BFFAA6F9BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Callistoides) pumilio Kolbe 1889	<div><p>* Chlaenius (Callistoides) pumilio Kolbe, 1889</p><p>Distribution in Angola (Provinces): 2) Malanje.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.064444&amp;materialsCitation.latitude=-8.9375" title="Search Plazi for locations around (long 16.064444/lat -8.9375)">Kibundi</a> (15 km N Nambua) (8° 56´15´´ S, 16° 03´52´´ E, 1164 m alt., 95) (MALANJE), 19.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Cambamba (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.009167&amp;materialsCitation.latitude=-9.245" title="Search Plazi for locations around (long 17.009167/lat -9.245)">Catoio-Quela</a>) (9° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20.XI.2015, 3♂, 3♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered known distribution in Africa (D.R. of the Congo and Sudan) (Anichtchenko 2016, Insectoid.info 2017). Adults were found in the edges of small rain pounds together with adults of the tiger beetle C. agualusai, running quickly when disturbed. It is a new species record to Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA6BFFAA6F9BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA2EBFD61FE38.text	03EFF252F30B7F6C14BFA2EBFD61FE38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Homalolachnus) lineatus Putzeys 1880	<div><p>Chlaenius (Homalolachnus) lineatus Putzeys, 1880</p><p>Distribution in Angola (Provinces): 1) Huíla, Malanje.</p><p>Material examined. Bicuar, Parque Nacional (unknown) (HUÍLA), 21.II.2014, 2♂, PF, R. Capela leg., ASC. Remarks. A species restricted to Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). It seems a common species in the Huíla Province by the reported localities given (Ferreira 1965). Adults were collected by pitfall trapping within a corn field plantation.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA2EBFD61FE38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA425FCC1F8F9.text	03EFF252F30B7F6C14BFA425FCC1F8F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Callistoides) Bonelli 1810	<div><p>Chlaenius (Callistoides) sp.</p><p>Distribution in Angola (Provinces): Malanje.</p><p>Material examined. Carima (Cacuso-Pungandongo) (9° 35´00´´ S, 15° 42´01´´ E, 1117 m alt., 130) (MALANJE), 23.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. It was not possible to ascribe the unique female specimen to any species. The adult was found in the edges of a small rain pound within a secondary open forest.</p></div>	https://treatment.plazi.org/id/03EFF252F30B7F6C14BFA425FCC1F8F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA2EBFB57FE70.text	03EFF252F3147F7314BFA2EBFB57FE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistomimus convexicollis Britton 1937	<div><p>* Callistomimus convexicollis Britton, 1937</p><p>Distribution in Angola (Provinces): 2) Malanje.</p><p>Material examined. Cambamba (Catoio-Quela) (9° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species with a known distribution throughout D. R. of the Congo, Zambia and Mozambique (Alves 1963, Anichtchenko 2016, Insectoid.info 2017). The singleton adult was collected beneath litter in a countryside road together with adults of tiger beetles [ Dromica (Foveodromica) sp., L. wellmani) and of ground beetles ( Lasiocera schuelei sp. n., G. ellipticus]. It is a new genus/species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3147F7314BFA2EBFB57FE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA16FFEA9FAB8.text	03EFF252F3147F7314BFA16FFEA9FAB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Notiobia (Diatypus) picina (Chaudoir 1878)	<div><p>Notiobia (Diatypus) picina (Chaudoir, 1878)</p><p>Distribution in Angola (Provinces): 1) Huambo, Cunene.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.‒ 2. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Cachimbamba (12° 28´11´´ S, 15° 40´56´´ E, 1560 m alt., 232) (HUAMBO), 3. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A common species widespread throughout Africa south of Sahara (Ferreira 1965, Anichtchenko 2016), but also present in South America (Argentina, Brazil, Uruguay) (Insectoid.info 2017) where was introduced probably by means of the nautical slave trade held between Africa (e.g. Luanda) and Brazil (e.g. Salvador) during the XVI‒XVII centuries. Adults were found on ground during night near public lights (first locality) together with some other ground beetles [e.g. S. guineense, C. intermedius, C. rikatlensis, C. cupreocinctus, C.. tenuicollis, Progonochaetus planicollis (Putzeys, 1880), Platymetopus pseudocrenulatus Facchini, 2016, Agonidium natalense (Boheman, 1848), Agonidium alacre (Boheman, 1848), E. peringuey, E. intermedius] or on soil beneath litter (second locality).</p></div>	https://treatment.plazi.org/id/03EFF252F3147F7314BFA16FFEA9FAB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA01AFCA1FD77.text	03EFF252F3147F7314BFA01AFCA1FD77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Notiobia Perty 1830	<div><p>Notiobia Perty, 1830</p><p>There is only one species reported for Angola within this genus (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017) which was sampled during our entomological trips also.</p></div>	https://treatment.plazi.org/id/03EFF252F3147F7314BFA01AFCA1FD77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA728FE31FA61.text	03EFF252F3147F7314BFA728FE31FA61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progonochaetus G. Muller 1938	<div><p>Progonochaetus G. Müller, 1938</p><p>A genus with five species known to Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017), of which were sampled two species.</p></div>	https://treatment.plazi.org/id/03EFF252F3147F7314BFA728FE31FA61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA455FEB8F852.text	03EFF252F3147F7314BFA455FEB8F852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progonochaetus incrassatus (Boheman 1848)	<div><p>Progonochaetus incrassatus (Boheman, 1848)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Benguela, Huambo, Huíla, Cunene, Cuando Cubango.</p><p>Material examined. Bembua-Camenhe (12° 16´53´´ S, 15° 27´01´´ E, 1686 m alt., 231) (HUAMBO), 1.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 2♂, 4.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout central and southern Africa (Democratic Republic of the Congo, Angola, Namibia, Zambia, Zimbabwe, Burundi, Rwanda, Malawi, Mozambique and Tanzania) (Basilewsky 1950a, Anichtchenko 2016, Insectoid.info 2017). In Angola the species presents a wide distribution throughout the country. Adults were found on soil beneath litter (first locality, see S. lucidus strigiceps remarks) or on ground and walls during night near public lights (second locality) together with several other ground beetles [e.g. S. guineense, C. rikatlensis, C. cupreocinctus, C. tenuicollis, P. planicollis (Putzeys, 1880), A. natalense, A. alacre, E. peringuey, E. intermedius].</p></div>	https://treatment.plazi.org/id/03EFF252F3147F7314BFA455FEB8F852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA2EBFC64FDE0.text	03EFF252F3157F7214BFA2EBFC64FDE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Progonochaetus planicollis (Putzeys 1880)	<div><p>Progonochaetus planicollis (Putzeys, 1880)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Malanje; 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.‒ 2. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3. XI.2015, 1 ♂, 1♀, 4. XI.2015, 2 ♂, 6♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species originally described from Angola (Putzeys 1880), but with a wide distribution throughout Africa south of Sahara (Senegal, Guinea-Bissau, Guinea, Mali, Burkina Faso, Chad, Ivory Coast, Cameroon, Ghana, Central African Republic, Nigeria, Equatorial Guinea, Gabon, Congo, D. R. of the Congo, Angola, Namibia, Zambia, Zimbabwe, Rwanda, Botswana, Saudi Arabia, Yemen, Djibouti, Eritrea, Ethiopia, Somalia, Kenia, Tanzania, Mozambique) (Serrano 1999b, Anichtchenko 2016, Insectoid.info 2017). Adults were collected in activity during night on ground and walls near public lights together with other ground beetles (see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3157F7214BFA2EBFC64FDE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA1DEFDD1FB7F.text	03EFF252F3157F7214BFA1DEFDD1FB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenolophus (Egadroma) barbarae Facchini 2012	<div><p>* Stenolophus (Egadroma) barbarae Facchini, 2012</p><p>Distribution in Angola (Provinces): 2) Malanje.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.064444&amp;materialsCitation.latitude=-8.9375" title="Search Plazi for locations around (long 16.064444/lat -8.9375)">Kibundi</a> (15 km N Nambua) (8° 56´15´´ S, 16° 03´52´´ E, 1164 m alt., 95) (MALANJE), 19.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species known from Central African Republic, Cameroon, Burundi, Zambia and Malawi (Facchini 2012b). The single male fits well the original description, including the aedeagus features. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3157F7214BFA1DEFDD1FB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA767FB88F9D0.text	03EFF252F3157F7214BFA767FB88F9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenolophus (Egadroma) humeralis (Dejean 1831)	<div><p>* Stenolophus (Egadroma) humeralis (Dejean, 1831)</p><p>Distribution in Angola (Provinces): 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 5.XII.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widespread throughout western Africa (Senegal, Gambia, Guinea-Bissau, Guinea, Sierra Leone, Gabon, Mali, D. R. of the Congo) (Serrano 1999b, Anichtchenko 2016, Insectoid.info 2017). The single adult fits well the description and aedeagus of this species and was collected beneath litter together with adults of some tiger and ground beetles [e.g. Dromica (Foveodromica) sp., A. distinctus, A. optimus, D. tarsalis, C. magnicollis discrepans, G. procera capelai ssp.n.]. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3157F7214BFA767FB88F9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA4B0FA7FF80D.text	03EFF252F3157F7214BFA4B0FA7FF80D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenolophus (Egadroma) Dejean 1821	<div><p>Stenolophus (Egadroma) sp.</p><p>Distribution in Angola (Provinces): Bié.</p><p>Material examined. Satchijamba (10 km E) (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 303) (BIÉ), 7.IV.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species close to S. barbarae Facchini, 2012 as well as to S. basirufus (Basilewsky, 1948) . However, a different shape of the median lobe of the aedeagus did not allow its inclusion on one of these species.</p></div>	https://treatment.plazi.org/id/03EFF252F3157F7214BFA4B0FA7FF80D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA0C1FCADFCE5.text	03EFF252F3157F7214BFA0C1FCADFCE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenolophus Dejean 1821	<div><p>Stenolophus Dejean, 1821</p><p>A speciose genus with eleven species and two subspecies known to Angola, belonging to sg. Egadroma Motschulsky, 1855 (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017, Facchini, 2016a), of which were sampled three species, two of them new records to this country.</p></div>	https://treatment.plazi.org/id/03EFF252F3157F7214BFA0C1FCADFCE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA2EBFAD9FF48.text	03EFF252F3167F7114BFA2EBFAD9FF48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Harpalus Latreille 1802	<div><p>Harpalus Latreille, 1802</p><p>A genus with two species recorded for Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA2EBFAD9FF48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA339FA9EFD85.text	03EFF252F3167F7114BFA339FA9EFD85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Harpalus Latreille 1802	<div><p>Harpalus sp.</p><p>Distribution in Angola (Provinces): Benguela.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.839158&amp;materialsCitation.latitude=-12.339533" title="Search Plazi for locations around (long 13.839158/lat -12.339533)">Baixo Pundo</a> (12° 20´22.32´´ S, 13° 50´20.97´´ E, 275 m alt., 227/8) (BENGUELA), 25‒27.X.2013, 1♀, PF, R. Capela leg., ASC.</p><p>Remarks. The single female adult specimen was collected by pitfall trapping within an open bush habitat with sparse grassy vegetation and sandy soil, together with the ground beetle Cypholoba brevivittis Chaudoir, 1866 .</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA339FA9EFD85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA4B2FABDF86A.text	03EFF252F3167F7114BFA4B2FABDF86A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus (Hyparpalus) escheri (Dejean 1831)	<div><p>Parophonus (Hyparpalus) escheri (Dejean, 1831)</p><p>Distribution in Angola (Provinces): 1) Huíla; 2) Bié.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout Africa south of Sahara (Serrano 1999c, Anichtchenko 2016, Insectoid.info 2017). The single adult was collected by means of light trapping within a secondary open forest together with some other ground beetles (see S. angolense remarks). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA4B2FABDF86A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA7B5FE34F9D1.text	03EFF252F3167F7114BFA7B5FE34F9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus Ganglbauer 1891	<div><p>Parophonus Ganglbauer, 1891</p><p>A genus with ten species known for Angola, belonging one to sg. Heterohyparpalus Basilewsky, 1946 and the remaining species to sg. Hyparpalus Alluaud, 1930, respectively (Ferreira 1965, Lecordier 1988, Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA7B5FE34F9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA0FEFD8EFCAF.text	03EFF252F3167F7114BFA0FEFD8EFCAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platymetopus Dejean 1829	<div><p>Platymetopus Dejean, 1829</p><p>A genus with nine species known for Angola (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017, Facchini 2016b), of which was sampled one species.</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA0FEFD8EFCAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA117FB69FACD.text	03EFF252F3167F7114BFA117FB69FACD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platymetopus pseudocrenulatus Facchini 2016	<div><p>Platymetopus pseudocrenulatus Facchini, 2016</p><p>Distribution in Angola (Provinces): 1) Huambo, Huíla, Bié.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.900122&amp;materialsCitation.latitude=-13.60487" title="Search Plazi for locations around (long 16.900122/lat -13.60487)">Chitembo</a> (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 1. IV.2014, 1 ♂, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.789444&amp;materialsCitation.latitude=-13.556389" title="Search Plazi for locations around (long 16.789444/lat -13.556389)">Chitembo</a> (6 km SE) (13° 33´23´´ S, 16° 47´22´´ E, 1586 m alt., 302) (BIÉ), 9. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.549167&amp;materialsCitation.latitude=-12.222221" title="Search Plazi for locations around (long 15.549167/lat -12.222221)">Alto Hama</a> (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.‒2. XII.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Mutubanzamba (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.527777&amp;materialsCitation.latitude=-9.795834" title="Search Plazi for locations around (long 16.527777/lat -9.795834)">Cangandala</a>) (9° 47´45´´ S, 16° 31´40´´ E, 1136 m alt., 132) (MALANJE), 21. XI.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described recently by Facchini (2016b) with a wide distribution throughout central and southern Africa (Angola, Zambia, Zimbabwe, Rwanda, Burundi, Malawi, Tanzania, Mozambique and South Africa). Adults were collected by means of light trapping within an open secondary forest (Chitembo), on walls near public lights (Alto Hama) and beneath litter (Mutubanzamba) together with other ground beetles (Chitembo: S. guineense; Alto Hama: C. intermedius, C. rikatlensis, C. cupreocinctus, C. tenuicollis, P. planicollis) and tiger beetles [Mutubanzamba: T. nubifera, E. villosus, L. saraliensis, L. uncivittata, L. wellmani].</p></div>	https://treatment.plazi.org/id/03EFF252F3167F7114BFA117FB69FACD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA612FBBCF9F2.text	03EFF252F3177F7014BFA612FBBCF9F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus (Heterohyparpalus) integer (Peringuey 1896)	<div><p>Parophonus (Heterohyparpalus) integer (Péringuey, 1896)</p><p>Distribution in Angola (Provinces): 1) Huíla, Cuando Cubango, Moxico; 2) Huambo, Bié, Malanje.</p><p>Material examined. Mumbué (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.32389&amp;materialsCitation.latitude=-13.828055" title="Search Plazi for locations around (long 17.32389/lat -13.828055)">Cuanza</a> riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 9. IV.2014, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.551945&amp;materialsCitation.latitude=-12.853056" title="Search Plazi for locations around (long 15.551945/lat -12.853056)">Caála</a> (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 5. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.759167&amp;materialsCitation.latitude=-12.713889" title="Search Plazi for locations around (long 16.759167/lat -12.713889)">Kakande</a> (12° 42´50´´ S, 16° 45´33´´ E, 1716 m alt., 258) (BIÉ), 7. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Mutubanzamba (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.527777&amp;materialsCitation.latitude=-9.795834" title="Search Plazi for locations around (long 16.527777/lat -9.795834)">Cangandala</a>) (09° 47´45´´ S, 16° 31´40´´ E, 1136 m alt., 132) (MALANJE), 21. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. Another species widespread throughout almost all Africa south of Sahara (Senegal, Guinea-Bissau, Mali, Ivory Coast, Chad, D. R. of the Congo, Angola, Zimbabwe, Rwanda, Eritrea, Sudan, Kenya, Tanzania, Mozambique, South Africa) (Serrano 1999c, Anichtchenko 2016, Insectoid.info 2017). Adults were found in countryside roads within savanah (Mumbué) or open secondary forest (Kakande and Mutubanzamba) with withish sandy soil beneath the marginal litter, and on a wall near public lights (Caála) with other ground beetles (see P. holosericus remarks). It is a new record for the Huambo, Bié and Malanje Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3177F7014BFA612FBBCF9F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA2EBFAD9FDA8.text	03EFF252F3177F7014BFA2EBFAD9FDA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus (Hyparpalus) holosericus (Dejean 1829)	<div><p>Parophonus (Hyparpalus) holosericus (Dejean, 1829)</p><p>Distribution in Angola (Provinces): 1) Huíla, Cuando-Cubango, Lunda Norte, Lunda Sul; 2) Huambo, Bié.</p><p>Material examined. Cachingues-Chipica (13° 10´15.74´´ S, 16° 45´48.17´´ E, 1649 m alt., 281) (BIÉ), 2.IV.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 5.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A very common species widespread throughout all Africa south of Sahara, including Madagascar, Comoros and Seychelles islands (Serrano 1999c, Anichtchenko 2016, Insectoid.info 2017). Adults were collected by means of light trapping within an open secondary forest (Cachingues-Chipica) and on a wall near public lights (Caála) together with other ground beetles [Cachingues-Chipica: S. ludricus, A. natalensis; Caála: C. deyrollei, C. pulchelus, Parophonus integer (Péringuey, 1896)]. It is a new record for the Huambo and Bié Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3177F7014BFA2EBFAD9FDA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA019FEF1FBB1.text	03EFF252F3177F7014BFA019FEF1FBB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus (Hyparpalus) tomentosus (Dejean 1829)	<div><p>Parophonus (Hyparpalus) tomentosus (Dejean, 1829)</p><p>Distribution in Angola (Provinces): 1) Cunene; 2) Benguela.</p><p>Material examined. Lobito (12° 21´S, 13° 33´E, 10 m alt., 227/8) (BENGUELA), 3.IV.1998, 1♀, PF, A. Neves leg., ASC.</p><p>Remarks. A very common species widespread throughout all Africa south of Sahara (Serrano 1999c, Anichtchenko 2016, Insectoid.info 2017). It was firstly recorded to Angola under two names (Putzeys 1880): Hypolithus tomentosus Dejean, 1829 and Hypolithus prolixus Putzeys, 1880 . Later Basilewsky (1960) confirmed the synonymy of the latter species with the former one. The single female adult was collected by pitfall trapping within an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Lobito together with some other ground beetles (see B. apicalis remarks). It is a new record for the Benguela Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3177F7014BFA019FEF1FBB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA4D3FE70F89C.text	03EFF252F3177F7014BFA4D3FE70F89C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohyparpalus Basilewsky 1946	<div><p>Pseudohyparpalus Basilewsky, 1946</p><p>A genus with two species known for Angola (Ferreira 1965, Clarke 1981, Insectoid.info 2017), of which was sampled one species.</p></div>	https://treatment.plazi.org/id/03EFF252F3177F7014BFA4D3FE70F89C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7714BFA505FACEFEDB.text	03EFF252F3177F7714BFA505FACEFEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohyparpalus elegans Clarke 1981	<div><p>Pseudohyparpalus elegans Clarke, 1981</p><p>Distribution in Angola (Provinces): 1) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. An Angolan ground beetle endemic species (Anichtchenko 2016, Insectoid.info 2017). The species was known until now by two type specimens from Luimbale, Mt. Moco at 1800-1900 m (Clarke 1981). The singleton male fits very well the original description, including the aedeagus. The adult was collected on a wall near public lights with other ground beetles ( C. cupreocinctus, C. pulchelus, P. incrassatus, P. planicollis).</p></div>	https://treatment.plazi.org/id/03EFF252F3177F7714BFA505FACEFEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA6BEFC49F8BF.text	03EFF252F3107F7714BFA6BEFC49F8BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Africobatus) harpaloides (Guerin-Meneville 1847)	<div><p>Siopelus (Africobatus) harpaloides (Guérin-Méneville, 1847)</p><p>Distribution in Angola (Provinces): 1) Benguela, Huíla, Cuando Cubango, Moxico; 2) Huambo, Bié.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.76338&amp;materialsCitation.latitude=-13.17104" title="Search Plazi for locations around (long 16.76338/lat -13.17104)">Cachingues-Chipica</a> (13° 10´15.74´´ S, 16° 45´48.17´´ E, 1649 m alt., 281) (BIÉ), 2.IV.2014, 1♂, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.20493&amp;materialsCitation.latitude=-13.727433" title="Search Plazi for locations around (long 17.20493/lat -13.727433)">Satchijamba-Somakwanza</a> (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 303) (BIÉ), 7.IV.2014, 1♂, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.900122&amp;materialsCitation.latitude=-13.60487" title="Search Plazi for locations around (long 16.900122/lat -13.60487)">Chitembo</a> (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.551945&amp;materialsCitation.latitude=-12.853056" title="Search Plazi for locations around (long 15.551945/lat -12.853056)">Caála</a> (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♂, 1♀, 5.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A common species known almost from all Africa south of Sahara (Senegal, Guinea, Togo, Ivory Coast, Cameroon, Central African Republic, Chad, Gabon, D.R. of the Congo, Burundi, Rwanda, Angola, Zambia, Zimbabwe, Eritrea, Somalia, Ethiopia, Kenya, Tanzania, Mozambique, Uganda, Malawi, South Africa) (Basilewsky 1950a, Ferreira 1965, Insectoid.info 2017). In the first three localities the adults were collected by light trapping within open secondary forests together with some other ground beetles (e.g. S. ludricus, S. guineense, A. natalensis). Adults in Caála were collected on a wall near public lights with other ground beetles also (see P. elegans remarks). It is a new record for the Huambo and Bié Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3107F7714BFA6BEFC49F8BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA16DFEF1FBC5.text	03EFF252F3107F7714BFA16DFEF1FBC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Laparhetes) gracilis (Harold 1879)	<div><p>Siopelus (Laparhetes) gracilis (Harold, 1879)</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Kwanza Sul, Malanje, Moxico; 2) Huambo. Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species originally described from Angola (Harold 1879), but with a wide distribution throughout Africa (Guinea, Central African Republic, Congo, D. R. of the Congo, Angola, Zimbabwe, Burundi, Malawi, Mozambique, Sudan, Tanzania) (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017). The adult was collected on a wall near public lights with other ground beetles (see P. elegans remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3107F7714BFA16DFEF1FBC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA3CBFCB5FD75.text	03EFF252F3107F7714BFA3CBFCB5FD75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus Murray 1859	<div><p>Siopelus Murray, 1859</p><p>A genus with ten species known for Angola (Ferreira 1965, Insectoid.info 2017, Facchini 2017b), distributed by six subgenera [ Africobatus Basilewsky, 1946, Aulacoryssus Alluaud, 1916, Laparhetes Jeannel, 1946 and Siopelus (s. str.) with one species each one, Neosiopelus Basilewsky, 1946 and Pseudosiopelus Alluaud, 1916 with two and four species, respectively] (Ferreira 1965, Anichtchenko 2016, Insectoid.info 2017, Facchini 2017b). A total of eigth species were sampled within Africobatus, Aulacoryssus, Laparhetes (each one with one species), Neosiopelus (two species) and the remaining three within Pseudosiopelus .</p></div>	https://treatment.plazi.org/id/03EFF252F3107F7714BFA3CBFCB5FD75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7614BFA535FC8FFE37.text	03EFF252F3107F7614BFA535FC8FFE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Neosiopelus) punctatellus subsp. punctatellus (Reiche 1847) Reiche 1847	<div><p>Siopelus (Neosiopelus) punctatellus punctatellus (Reiche, 1847)</p><p>Distribution in Angola (Provinces): 1) Benguela, Huíla, Lunda Norte, Malanje; 2) Huambo. Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species present throughout Arabian Peninsula and eastern and central-southern Africa (Yemen, Eritrea, Somalia, Ethiopia, Kenya, Tanzania, Mozambique, Malawi, Rwanda, Burundi, D. R. of the Congo, Angola, Namibia, Zambia, Zimbabwe, Niger) (Basilewsky 1950a, Ferreira 1965, Insectoid.info 2017). It is with some reserve that we ascribe to this species the sampled female singleton. It should be necessary a male to confirm this identification because there are some species belonging to this sg. with similar habitus (see Facchini 2012c). The adult was found on ground during night near public lights together with several other ground beetles (see P. incrassatus remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3107F7614BFA535FC8FFE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA3AFFD7AFC88.text	03EFF252F3117F7614BFA3AFFD7AFC88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Neosiopelus) wrasei Facchini 2012	<div><p>* Siopelus (Neosiopelus) wrasei Facchini, 2012</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8‒17.XI.2015, 1♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described from Zimbabwe and Botswana (Facchini 2012). The single female adult fits well the original description, but without a male specimen it is with some reserve that we ascribe it to this species. The singleton adult was collected by pitfall trapping within an open secondary forest together with other ground beetles ( S. lucidus strigiceps, C. bipustulatus, A. pseudoangolensis and Atractonotus puncticollis Schle &amp; Heinz, 2013). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3117F7614BFA3AFFD7AFC88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA74AFC8FF92A.text	03EFF252F3117F7614BFA74AFC8FF92A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Pseudosiopelus) barbarae Facchini 2017	<div><p>Siopelus (Pseudosiopelus) barbarae Facchini, 2017</p><p>Distribution in Angola (Provinces): Benguela, Bié, Huíla; 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X.‒ 2. XI.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3. XI.2015, 1 ♂, 4. XI.2015, 1 ♂, 5. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described recently with a scattered distribution in central, southern and eastern Africa (Nigeria, D. R. of the Congo, Angola, Zambia, Zimbabwe, Uganda) (Facchini 2017b). Adults were collected on walls during night near public lights together with several other ground beetles (e.g. see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3117F7614BFA74AFC8FF92A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA1F9FC3CFB5A.text	03EFF252F3117F7614BFA1F9FC3CFB5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Neosiopelus) Murray 1859	<div><p>Siopelus (Neosiopelus) sp.</p><p>Distribution in Angola (Provinces): Malanje.</p><p>Material examined. Carima (Cacuso-Pungandongo) (9° 35´00´´ S, 15° 42´01´´ E, 1117 m alt., 130) (MALANJE), 23.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. The single female adult is quite similar to S. punctatellus, but with a shape of pronotum slightly different [anterior angles not protruding, anterior margin almost straight between angles and basal margin not sinuate at each side of the middle region). The adult was found in a set aside country road beneath litter within an open secondary forest together with adults of tiger and ground beetles [e.g. Dromica (Foveodromica) auropunctata Quedenfeldt, 1883, P. angusticollis, T. nubifera, E. villosus, G. ellipticus].</p></div>	https://treatment.plazi.org/id/03EFF252F3117F7614BFA1F9FC3CFB5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7514BFA49BFAFEFEFF.text	03EFF252F3117F7514BFA49BFAFEFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Pseudosiopelus) natalicus Peringuey 1896	<div><p>Siopelus (Pseudosiopelus) natalicus Péringuey, 1896</p><p>Distribution in Angola (Provinces): 1) Huambo, Huíla, Lunda Norte; 2) Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 303) (BIÉ), 7.IV.2014, 2♀, LT, A. Serrano &amp; R. Capela leg., ASC; Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♂, 2♀, LT, A. Serrano &amp; R. Capela leg., ASC; Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4.XI.2015, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered distribution throughout eastern, central and southern Africa (Kenya, Tanzania, Rwanda, D. R. of the Congo, Angola, Zimbabwe, South Africa) (Basilewsky 1950a, Ferreira 1965, Insectoid.info 2017). Adults were sampled by light trapping in Bié province within open secondary forests together with other ground beetles (see S. angolense remarks) and on walls during night (Caála) near public lights together with several other ground beetles (see P. incrassatus remarks). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3117F7514BFA49BFAFEFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA766FF13FA27.text	03EFF252F3127F7514BFA766FF13FA27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Axinotoma Dejean 1829	<div><p>* Axinotoma Dejean, 1829</p><p>A genus apparently without any species known to Angola (Facchini 2011a, 2017a, however see Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3127F7514BFA766FF13FA27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA3E7FD06FD50.text	03EFF252F3127F7514BFA3E7FD06FD50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Pseudosiopelus) pallidior (Burgeon 1936)	<div><p>Siopelus (Pseudosiopelus) pallidior (Burgeon, 1936)</p><p>Distribution in Angola (Provinces): 1) Huíla, Cuando Cubango; 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♂, 5.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known on western, eastern and central Africa (Guinea, Ivory Coast, Central African Republic, Equatorial Guinea, Gabon, D. R. of the Congo, Angola, Burundi, Rwanda, Zimbabwe, Ethiopia, Kenya, Tanzania, Mozambique) (Basilewsky 1950a, Ferreira 1965, Insectoid.info 2017). Adults were collected on walls during night near public lights together with several other ground beetles (e.g. see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3127F7514BFA3E7FD06FD50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA131FCFFFB7D.text	03EFF252F3127F7514BFA131FCFFFB7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Pseudosiopelus) pulchelus (Dejean 1829)	<div><p>* Siopelus (Pseudosiopelus) pulchelus (Dejean, 1829)</p><p>Distribution in Angola (Provinces): 2) Cabinda.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.218611&amp;materialsCitation.latitude=-5.7280555" title="Search Plazi for locations around (long 12.218611/lat -5.7280555)">Material</a> examined. Chipita (5° 43´41´´ S, 12° 13´07´´E, 7 m alt., 7) (CABINDA), 11.II.2011, 1♂, DO, R. Capela leg., ASC.</p><p>Remarks. A species known on western, eastern and central Africa, including atlantic and indic islands (Mauritania, Senegal, Gambia, Guinea-Bissau, Guinea, Ivory Coast, Benin, Sierra Leone, Ghana, Togo, Burkina Faso, Mali, Chad, Cameroon, Central African Republic, Equatorial Guinea, Gabon, Niger, Nigeria, Democratic Republic of the Congo, Burundi, Rwanda, Tanzania, Mozambique, São Tomé and Príncipe, Comoros, Seychelles and Madagascar) (Basilewsky 1950a, Insectoid.info 2017). The single adult was collected under a stone. Surprisingly the species was never recorded for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3127F7514BFA131FCFFFB7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7414BFA79FFBDCFEA2.text	03EFF252F3127F7414BFA79FFBDCFEA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Axinotoma latipalpis Basilewsky 1968	<div><p>* Axinotoma latipalpis Basilewsky, 1968</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC; Chitembo (13° 22´40.78´´ S, 16° 41´58.07´´ E, 1666 m alt., 281) (BIÉ), 1.XI.2014, 1♀, LT, A. Serrano leg., ASC .</p><p>Remarks. A species known only to Central African R. and Uganda (Basilewsky 1968, Facchini 2011a). Interestingly our male specimen fits very well the median lobe shape of aedeagus in lateral view of A. latipalpis Basilewsky, 1968 (Basilewsky 1968, Fig. 3, Facchini 2003, Fig. 18a) with the apex slightly bent up. However, in dorsal view the median lobe apex is similar to the drawing given in Facchini (2011a, Fig. 12b) for A. dilatipalpis Facchini, 2011 . Despite this asymmetry we ascribe our identification to the Basilewsky`s species by some morphological features (e.g. last labial palpomere only dilated, elytra striae crenulated, third interval with a discal setigerous puncture in the last third). However, our specimens do not present any small pubescence in apical margin, as was point out by Basilewsky (1968) for its species, and the labial palpi are clearly bicolored.</p><p>Unfortunately, Facchini (2011a) do not give in the affinities between both species any comment on the elytra apical pubescence. Probably in a near future the status of these two species will be reviewed. Adult specimens were collected by light trapping within open secondary forests together with some other ground beetles (e.g. see S. angolense and S. guineense remarks). It is a new genus/species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3127F7414BFA79FFBDCFEA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA0A6FEFEFB7F.text	03EFF252F3137F7414BFA0A6FEFEFB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthogonius MacLeay 1825	<div><p>Orthogonius sp. 1</p><p>Distribution in Angola (Provinces): Huambo, Bié.</p><p>Material examined. Lumbanje (Morro Lubiri) (12° 07´55´´ S, 15° 30´59´´ E, 1424 m alt., 232) (HUAMBO), 30.X.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC; Chitembo (6 km SE) (13° 33´23´´ S, 16° 47´22´´ E, 1586 m alt., 302) (BIÉ), 9.XI.2015, 1♂, DO, A. Serrano leg., ASC .</p><p>Remarks. Adult specimens were collected always within litter in open secondary forests together with a common tiger beetle ( P. angusticollis). Further adult specimens of tiger and ground beetles were collected in the same habitats namely L. uncivittata, L. saraliensis and L. wellmani in Lumbanje, Euryarthron reticostatum (Wellman &amp; W. Horn, 1908), Dromica (Foveodromica) sp., T. nubifera, L. saraliensis, L. wellmani, S. vilhenai, C. ovalipennis, E. intermedius, O. patroboides, N. vethi, C. graphipteroides monardi and A. (Odontanthia) kleinfeldi in Catata-Nova Monção and T. nzingae, Ophryodera rufomarginata poggei (Harold, 1878) and L. uncivittata in Chitembo.</p></div>	https://treatment.plazi.org/id/03EFF252F3137F7414BFA0A6FEFEFB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA3C5FAFCFD3D.text	03EFF252F3137F7414BFA3C5FAFCFD3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthogonius MacLeay 1825	<div><p>Orthogonius MacLeay, 1825</p><p>A genus with four species known for Angola (Ferreira 1965, Insectoid.info 2017). Nevertheless we had collected the same number of species during our entomological trips in Angola, we were not well succeeding to assign the specimens obtained to any known one ( Orthogonius alutaceus Quedenfeldt, 1883, Orthogonius brevilabris H. Kolbe, 1889, Orthogonius clarkei Murray, 1858 and Orthogonius impunctipennis Quedenfeldt, 1883).</p></div>	https://treatment.plazi.org/id/03EFF252F3137F7414BFA3C5FAFCFD3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA767FCBDF9F5.text	03EFF252F3137F7414BFA767FCBDF9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthogonius MacLeay 1825	<div><p>Orthogonius sp. 2</p><p>Distribution in Angola (Provinces): Bié.</p><p>Material examined. Bembua-Camenhe (12° 16´53´´ S, 15° 27´01´´ E, 1686 m alt., 302) (BIÉ), 2.XI.2015, 3♂, DO, A. Serrano leg., ASC .</p><p>Remarks. Adult specimens were collected within litter in open secondary forests together with some tiger and ground beetles ( Prothyma erythropyga erythropyga (Putzeys, 1880), Dromica (Foveodromica) soror (W. Horn, 1935), P. angusticollis, T. nubifera, O. rufomarginata distanti (Heath, 1905), L. uncivittata, L. saraliensis, L. wellmani, C. agualusai, S. lucidus strigiceps and A. kleinfeldi .</p></div>	https://treatment.plazi.org/id/03EFF252F3137F7414BFA767FCBDF9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA4E2FDEEF853.text	03EFF252F3137F7414BFA4E2FDEEF853.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthogonius MacLeay 1825	<div><p>Orthogonius sp. 3</p><p>Distribution in Angola (Provinces): Bié.</p><p>Material examined. Bembua-Camenhe (12° 16´53´´ S, 15° 27´01´´ E, 1686 m alt., 302) (BIÉ), 2.XI.2015, 2♂, DO, A. Serrano leg., ASC; Chingueia-Mandongue (13° 35´46´´ S, 16° 58´59´´ E, 1648 m alt., 302) (BIÉ), 9.XI.2015, 2♂, DO, A. Serrano leg., ASC .</p><p>Remarks. Like the precedent Orthogonius, these adult specimens were collected within litter in open secondary forests together with some tiger and ground beetles (for the first locality see Orthogonius sp. 2 remarks). Adult specimens in Chingueia-Mandongue were collected together with some different tiger and ground beetles (see S. lucidus strigiceps remarks).</p></div>	https://treatment.plazi.org/id/03EFF252F3137F7414BFA4E2FDEEF853	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA7B7FA6EF944.text	03EFF252F31C7F7B14BFA7B7FA6EF944.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agonidium alacre (Boheman 1848)	<div><p>* Agonidium alacre ((Boheman, 1848)</p><p>Distribution in Angola (Provinces): 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3.XI.2015, 1♀, 4.XI.2015, 1♂, 5.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a distribution in southern and eastern Africa (Botswana, Namibia, South Africa, Zimbabwe and Mozambique), but recorded also to Yemen (Insectoid.info 2017). Adult specimens fit well the descrition of this species, including the aedeagus conformation (Basilewsky 1950b). Adults were found on ground during night near public lights (e.g. see comments on C. rikatlensis remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7B14BFA7B7FA6EF944	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA762FBDBFACF.text	03EFF252F31C7F7B14BFA762FBDBFACF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agonidium Jeannel 1948	<div><p>Agonidium Jeannel, 1948</p><p>A genus with one species known for Angola (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7B14BFA762FBDBFACF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA117FD3EFBC8.text	03EFF252F31C7F7B14BFA117FD3EFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexagonia angustula Peringuey 1904	<div><p>Hexagonia angustula Péringuey, 1904</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Malanje.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.064444&amp;materialsCitation.latitude=-8.9375" title="Search Plazi for locations around (long 16.064444/lat -8.9375)">Kibundi</a> (15 km N Nambua) (8° 56´15´´ S, 16° 03´52´´ E, 1164 m alt., 95) (MALANJE), 19.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species known from Angola, D. R. of the Congo and Zimbabwe (Basilewsky 1948, Ferreira 1965, Insectoid.info 2017). The single adult was collected on the ground in a countryside road within an open secondary forest. It is a new record for the Malanje Province.</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7B14BFA117FD3EFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA01AFD3EFCAF.text	03EFF252F31C7F7B14BFA01AFD3EFCAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexagonia Kirby 1825	<div><p>Hexagonia Kirby, 1825</p><p>A genus with two species known for Angola (Ferreira 1965, Insectoid.info 2017). This genus is surely poorly known in Angola when compared with the neighbouring country D. R. of the Congo with a total of nine recorded species (Anichtchenko 2016, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7B14BFA01AFD3EFCAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA293FDB1FE70.text	03EFF252F31C7F7B14BFA293FDB1FE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthogonius MacLeay 1825	<div><p>Orthogonius sp. 4</p><p>Distribution in Angola (Provinces): Bié.</p><p>Material examined. Chitembo (13° 30´55´´ S, 16° 45´20´´ E, 1628 m alt., 302) (BIÉ), 9.XI.2015, 2♀, DO, A. Serrano leg., ASC .</p><p>Remarks. Adult specimens were found in activity within the village on soil in the road edges during night, probably attracted to the public lights.</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7B14BFA293FDB1FE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7A14BFA53DFA72FEA2.text	03EFF252F31C7F7A14BFA53DFA72FEA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agonidium natalense (Boheman 1848)	<div><p>Agonidium natalense ((Boheman, 1848)</p><p>Distribution in Angola (Provinces): 1) Unknown; 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species known already to Angola without known localities (Putzeys 1880, under the name Platynus urens) and with a distribution in southern and part of eastern Africa (Angola, Botswana, South Africa, Zimbabwe and Mozambique) (Basilewsky 1950b, Insectoid.info 2017). The singleton adult was found on ground during night near public lights (e.g. see C. rikatlensis remarks). It seems a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F31C7F7A14BFA53DFA72FEA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA051FE4CFC55.text	03EFF252F31D7F7A14BFA051FE4CFC55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euleptus intermedius Peringuey 1896	<div><p>* Euleptus intermedius Péringuey, 1896</p><p>Distribution in Angola (Provinces): 2) Huambo, Bié.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 4. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catota (2,5 km S) (14° 00° 37´´ S, 17° 24° 00´´ E, 1532 m alt., 323) (BIÉ), 8‒17. XI.2015, 1 ♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known from South Africa R. and Zimbabwe (Basilewsky 1950b, Insectoid.info 2017). Adult specimens fit well the original description (Péringuey 1896). Adults of the first locality were found on ground during night near public lights (e.g. see C. rikatlensis remarks) and in the second locality beneath litter within an open secondary forest (e.g. see comments on S. vilhenai remarks), respectively. The Catota adult specimen was collected by pitfall trapping together with other ground beetles (see S. wrasei remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F31D7F7A14BFA051FE4CFC55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA303FBCDFE70.text	03EFF252F31D7F7A14BFA303FBCDFE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euleptus Klug 1833	<div><p>Euleptus Klug, 1833</p><p>A genus with one species recorded for Angola (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F31D7F7A14BFA303FBCDFE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA64EFE4CFA02.text	03EFF252F31D7F7A14BFA64EFE4CFA02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euleptus peringueyi Csiky 1931	<div><p>* Euleptus peringueyi Csiky, 1931</p><p>Distribution in Angola (Provinces): 2) Huambo, Kwanza Sul.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 3. XI.2015, 1 ♂, 6. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Calulo (9° 59´55´´ S, 14° 53´35´´ E, 999 m alt., 128) (BIÉ), 27.XI‒5. XII.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered known distribution (Cameroon, Ivory Coast, D. R. of the Congo, South Africa and Mozambique) (Basilewsky 1950b, Insectoid.info 2017). Adult specimens fit very well the original description (as albicornis Péringuey 1896), including the aedeagus (see Basilewsky 1950b, Fig. 6). Adults were found on ground during night near public lights (e.g. for the first locality see C. rikatlensis remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F31D7F7A14BFA64EFE4CFA02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7914BFA4B0FDFBFF6A.text	03EFF252F31D7F7914BFA4B0FDFBFF6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptagonum elegans (Peringuey 1904)	<div><p>* Leptagonum elegans (Péringuey, 1904)</p><p>Distribution in Angola (Provinces): 2) Huambo.</p><p>Material examined. Bembua-Camenhe (12° 16´53´´ S, 15° 27´01´´ E, 1686 m alt., 231) (HUAMBO), 31.X.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Londuimbali-Tinguita (12° 18´10´´ S, 15° 21´15´´ E, 1633 m alt., 231) (HUAMBO), 2.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known only from Zimbabwe and Mozambique (Basilewsky 1950b, Insectoid.info 2017). Adult specimens fit very well its description (Péringuey 1904, Basilewsky 1950b) and were found within litter in open secondary forests together with some tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new genus/species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F31D7F7914BFA4B0FDFBFF6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA463FC21F9D0.text	03EFF252F31D7F7A14BFA463FC21F9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptagonum Kolbe 1898	<div><p>* Leptagonum Kolbe, 1898</p><p>A genus not yet recorded for Angola (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F31D7F7A14BFA463FC21F9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA051FE11FB59.text	03EFF252F31E7F7914BFA051FE11FB59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthotrichus gilvipes (Boheman 1848)	<div><p>Orthotrichus gilvipes (Boheman, 1848)</p><p>Distribution in Angola (Provinces): 1) Unknown, 2) Kwanza Sul.</p><p>Material examined. Cateculo (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.7920475&amp;materialsCitation.latitude=-9.955486" title="Search Plazi for locations around (long 14.7920475/lat -9.955486)">Calulo</a>) (9° 57´19.75´´ S, 14° 47´31.37´´ E, 872 m alt., 128) (KWANZA SUL), 5. XI.2014, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Mussende ( Calulo) (9° 57´00´´ S, 14° 47´´ 36´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Alto Ventura (Faz. Monte Café) (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.836666&amp;materialsCitation.latitude=-9.9875" title="Search Plazi for locations around (long 14.836666/lat -9.9875)">Calulo</a>) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI.2015, 4 ♂, 5♀, 2. XII.2015, 1 ♂, 5♀, 5. XII.2015, 1 ♂, 2♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Calulo (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.910555&amp;materialsCitation.latitude=-10.037223" title="Search Plazi for locations around (long 14.910555/lat -10.037223)">Faz. Klein</a>) (10° 02´14´´ S, 14° 54´38´´ E, 1059 m alt., 147) (KWANZA SUL), 1. XII.2015, 1 ♂, 1♀, 3. XII.2015, 2 ♂, 2♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout eastern, central and southern Africa (Ethiopia, Mozambique, Zimbabwe, Burundi, Rwanda, D. R. of the Congo, Angola, Botswana, South Africa R.) (Putzeys 1880, Insectoid.info 2017). The species was recorded for Angola by Putzeys (1880) without information on localities. Basilewsky (1960a) studying the 24 types described by this author did not pay attention to the other 117 species of carabids recorded for Angola in that work, referring that most of them were misinterpreted. Unfortunately we can not confirm the identification of the Putzeys study because all the material deposited in MLZA (Museu Bocage) was destroyed during a catastrophic fire in 1978. However, as the species is so well characterized, it seems to us that Putzeys identification was almost surely correct. Adult specimens were found within litter in secondary rainforests together with other tiger and ground beetles (see comments on A. distinctus remarks). It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F31E7F7914BFA051FE11FB59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA74AFC4CF82F.text	03EFF252F31E7F7914BFA74AFC4CF82F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthotrichus patroboides (Murray 1859)	<div><p>Orthotrichus patroboides (Murray, 1859)</p><p>Distribution in Angola (Provinces): 2) Kwanza Sul, Huambo.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.356112&amp;materialsCitation.latitude=-13.433056" title="Search Plazi for locations around (long 15.356112/lat -13.433056)">Catata-Nova Monção</a> (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6. XI.2015, 2 ♂, 1♀,DO, A. Serrano &amp; R. Capela leg., ASC ; Cruzeiro (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.902636&amp;materialsCitation.latitude=-12.778094" title="Search Plazi for locations around (long 15.902636/lat -12.778094)">Pedra do Alemão</a>) (12° 46´41.14´´ S, 15° 54´09.49´´ E, 1743 m alt., 232) (HUAMBO), 7. XI.2015, 2 ♀, DO, A Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.907222&amp;materialsCitation.latitude=-9.899722" title="Search Plazi for locations around (long 14.907222/lat -9.899722)">Calulo-Cabuta</a> (9° 53´59´´ S, 14° 54´26´´ E, 831 m alt., 128) (KWANZA SUL), 29. XI.2015, 2 ♂, 4♀, 29.XI‒6. XII.2015, 26 ♂, 20♀, PF, A. Serrano &amp; R. Capela leg., ASC ; Mussende (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.793334&amp;materialsCitation.latitude=-9.95" title="Search Plazi for locations around (long 14.793334/lat -9.95)">Calulo</a>) (9° 57´00´´ S, 14° 47´36´´ E, 905 m alt., 128) (KWANZA SUL), 28.XI‒5. XII.2015, 14 ♂, 9♀, PF, A. Serrano &amp; R. Capela leg., ASC ; Alto Ventura (Faz. Monte Café) (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.836666&amp;materialsCitation.latitude=-9.9875" title="Search Plazi for locations around (long 14.836666/lat -9.9875)">Calulo</a>) (9° 59´15´´ S, 14° 50´12´´ E, 1086 m alt., 128) (KWANZA SUL), 5. XII.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout western, central and eastern Africa (Senegal, Guinea, Ivory Coast, Nigeria, D. R. of the Congo, Angola, Zambia, Burundi, Rwanda, Uganda) (Putzeys 1880, Insectoid.info 2017). The species was described by Putzeys (1880) from Angola under the name Platynus calefactus without information on localities. The species is very abundant in Calulo region and the adult specimens were found within litter or collected by pitfall trapping in secondary rainforests together with other tiger and ground beetles (see A. distinctus remarks). It is a new record for the Kwanza Sul and Huambo Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F31E7F7914BFA74AFC4CF82F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA35BFE7EFE70.text	03EFF252F31E7F7914BFA35BFE7EFE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthotrichus Peyron 1856	<div><p>Orthotrichus Peyron, 1856</p><p>A genus with seven species recorded for Angola (Ferreira 1965, Insectoid.info 2017). Its representatives are all within Metagonum Jeannel, 1948 in Lorenz (2005) . However, this genus was synonimized with Orthotrichus (Anichtchenko 2016) .</p></div>	https://treatment.plazi.org/id/03EFF252F31E7F7914BFA35BFE7EFE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7814BFA2EBFACFFDA8.text	03EFF252F31F7F7814BFA2EBFACFFDA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthotrichus umtalianum (Peringuey 1904)	<div><p>* Orthotrichus umtalianum (Péringuey, 1904)</p><p>Distribution in Angola (Provinces): 2) Kwanza Sul, Malanje.</p><p>Material examined. Cambamba (Catoio-Quela) (09° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20.XI.2015, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Calulo (Faz. Klein) (10° 02´14´´ S, 14° 54´38´´ E, 1059 m alt., 147) (KWANZA SUL), 1.XII.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known from central, southern and part of eastern Africa (D. R. of the Congo, Souh Africa R., Burundi, Zimbabwe) (Basilewsky 1950b, Insectoid.info 2017). Adults fit very well its description, including the aedeagus features (Péringuey 1904, Basilewsky 1950b). Adult specimens were found within litter in an open secondary forest and in a secondary rainforest together with some other tiger and ground beetles (see S. vilhenai and O. gilvipes remarks for first and second localities, respectively). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F31F7F7814BFA2EBFACFFDA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7814BFA0C2FBBDFC8A.text	03EFF252F31F7F7814BFA0C2FBBDFC8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perigona Laporte de Castelnau 1835	<div><p>Perigona Laporte de Castelnau, 1835</p><p>A genus with five species recorded to Angola (Ferreira 1965, Insectoid.info 2017), distributed by two subgenera ( Euripogena Basilewsky, 1989: one species; Trechicus LeConte, 1853: four species).</p></div>	https://treatment.plazi.org/id/03EFF252F31F7F7814BFA0C2FBBDFC8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7C14BFA1FBFB06FE12.text	03EFF252F31F7F7C14BFA1FBFB06FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perigona liboloensis Serrano & Capela & Santos 2017	<div><p>Perigona (s. str.) liboloensis Serrano &amp; Capela sp. n.</p><p>(Figs 1a, 2, 3)</p><p>Type series. Holotype, ♂; Angola (Kwanza Sul), Alto Ventura-Col (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.842222&amp;materialsCitation.latitude=-9.995" title="Search Plazi for locations around (long 14.842222/lat -9.995)">Calulo</a>) (9° 59´42´´ S, 14° 50´32´´ E, 1116 m alt., 128), 4.XII.2015, DO, A. Serrano &amp; R. Capela leg., ASC . Allotype, 1♀, same locality and date as holotype, ASC. Derivatio nominis. This species is named after Libolo (noun in apposition), a municipality of the Kwanza Sul Province where the specimens were found and home of one of the most remarkable Angolan Sports Club: the Sports Club Recreativo do Libolo .</p><p>Diagnosis. Body elongate, sub-parallel, slightly convex and reddish-brown (Fig. 1a); integument with microreticulation, shiny; surface with sparse micropunctures visible at high magnification. Elytral color as Perigona pallida Castelnau, 1835 . Internal sac of median lobe of aedeagus (Fig. 2) with a conspicuous twisted sclerite closer to the apex and a posterior bone shaped cut half sclerite (lateral view).</p><p>Description. Length of Holotype: 3.36 mm. Length of allotype: 3.16 mm.</p><p>Head. Slightly wider than long [length: 0.59 mm (holotype), 0.64 mm (allotype), width: 0.72 mm (holotype), 0.75 mm (allotype)], darker (black-brownish) than pronotum and elytra, clypeus, labrum, mouth parts and antennae piceous; surface with sparce fine punctuation, microreticulation distinct, isodiametric, rather glossy; eyes large, moderately protruding, labrum moderately notched; clypeal-frontal furrows distinct, oblique, extended to level of anterior supraorbital seta; frons and occiput slightly convex; antenna short just attaining base of pronotum, scape two times longer than wide, 3rd to 9th antennomeres as long as wide, slightly globulose, last antennomere 1.6‒1.8 times longer than wide. Cephalic chaetotaxy (large setae): Labrum with three pairs of setae gradually shorter from exterior to inner sides, one pair on sides of clypeus and two pairs of supraocular setae present over each eye.</p><p>Thorax. Pronotum (Fig. 1a) 1.27‒1.29 times wider than long [length: 0.66 mm (holotype), 0.74 mm (allotype), width: 0.85 mm (holotype), 0.94 mm (allotype)], reddish-brown dark, wider than head, barely subcordiform, widest at first third; surface sparcely micropunctate, with microreticulation distinct, consisting of isodiametric meshes in diskal area and transverse meshes and lines laterally, shiny; apical margin not margined, lateral margins with complete and distinct margination continuing in the two thirds of the basal margin after the posterior angle; apical margin arcuate, apical angles roundly acute, protruded; lateral margin slightly curved at first two thirds, followed by a slight sinuosity and ending straight obliquely before the roundly obtuse basal angles, posterior margin distinctly emarginated; median line distinct, not reaching anterior or posterior margins, anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous; anterior marginal setae situated close to the widest width, posterior marginal seta situated at basal angle, none short setae at disk or sides.</p><p>Elytra (Fig. 1a) 1.61‒1.62 times longer than wide [length: 1.90 mm (Holotype), 1.97 mm (allotype), width: 1.17 mm (holotype), 1.22 mm (allotype)], reddish-brown lighter than pronotum, sutural stripe slightly darker; surface glossy, with sparse very fine punctuation and superficial microreticulation of fine transverse meshes; narrow and elongate, sub-parallel, dorsally moderately convex, very slightly depressed on disk, widest at middle; humeral angles distinct but broadly rounded; mediam marginal punctures arranged in a straight line; marginal channel towards apex widened, depressed, first half scarsely pilose, second half densely pilose; only inner three striae recognizable at two thirds by inconspicuous sulci not marked by punctures, anterior setiferous puncture situated at 3rd striae and at basal quarter, median setiferous puncture situated in 3rd interval close to 2nd striae slightly behind middle, apical setiferous puncture (long setae) situated at 3rd striae just above sub-apical border.</p><p>Abdominal segments brown-yellowish, covered with dense dressed short hairs; last segment with 3 pairs (male) or 5 pairs (female) of setae near the posterior margin; terminal abdominal sternite in both sexes with fringe of short hairs. Legs brown-testaceous.</p><p>Aedeagus (Figs 2 a, b, c) with median lobe short and stout, ventral margin slightly curved, apex downwards (lateral view); internal sac with a conspicuous twisted sclerite and a bone shaped cut in half sclerite at right and left sides, respectively; genital ring (Fig. 2 d) rather asymmetric, with wide, obliquely rounded apex, rather parallel, base evenly rounded.</p><p>Female genitalia (Fig. 3) with the ordinary pattern shape of the genus. Stylomere 1 with two ventral large setae on subapical margin. Stylomere 2 with two medium sized ensiform ventro-lateral setae, one quite large ensiform dorso-median seta located about in middle of stylomere, and a single nematiform ventro-seta originating from a groove in apical third of stylomere. Some sensillae (probably chemoreceptors) are dispersed over ventral lateroapical margins of stylomere 1, and other sensillae over basal, medial and sub-apical areas of ventral, lateral and dorsal sides of stylomere 2.</p><p>Remarks. The new species belongs to the subgenus Perigona s.str. based on the arrangement of the three lateral marginal elytral pores in a straight line and by the presence of distinct clypeo-orbital sulci. Until now there are six species known for continental Africa belonging to this subgenus (Basilewsky 1989, Baehr 2004 and Serrano 2008). Perigona liboloensis sp. n. is easily segregate from P. parallela Chaudoir, 1878, P. nigrociliata Basiewsky, 1953, P. mediornata Basilewsky, 1989, P. wachteli Baehr, 2004 and P. principensis Serrano, 2008, among other characters, by the quite dissimilar aedeagi of all these species. Certainly the new species is next related to P. pallida Laporte de Castelnau, 1835 with which it shares the general color, the elongate, almost parallel body shape and male median lobe general shape, though differs in its general body length (3.16‒3.36 mm vs. 2.30‒2.70 mm), pronotum conformation (slightly sinuouse before the posterior angles vs. not sinuouse before the posterior angles), elytral index (1.61‒1.62 vs. 1.47‒1.55), and in the shape of the sclerotized plates in the internal sac of the median lobe of aedeagus (cf. Figs 2a, b vs. Fig. 5b in Basilewsky 1989). Moreover, P. pallida presents the pronotal widest situated in the middle while in the new species is in the first third. In the former species the elytral striae are conspicuous with punctuation well marked while in the latter the elytral striae are almost indistinct and without marked punctuation. By the Perigona pallida previous records (e.g. Basilewsky 1989) and the new species known locality, the two taxa seem to be sister species with vicariant (allopatric) distribution, a fact referred already for tiger beetle and ant nest beetle species of Angola also (Serrano et al. 2015, Serrano &amp; Capela 2015a).</p><p>Ecological notes. A species known until now only from Angola. Adults were found under the bark of a fallen tree within an abandoned coffee plantation (Fig. 9a) with Catascopus beauvoisi Laporte de Castelnau, 1835 and Coptodera (Coptoderina) congolensis Burgeon, 1937 . Further adults of Dermaptera [ Apachyus depressus (Palisot de Beauvois, 1805 and Echinosoma afrum (Palisot de Beauvois, 1805)] and Coleoptera Endomychidae, Trogossitidae, Tenebrionidae (several species) were found together (syntopic) with the new species.</p></div>	https://treatment.plazi.org/id/03EFF252F31F7F7C14BFA1FBFB06FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F7C14BFA443FC21F930.text	03EFF252F31B7F7C14BFA443FC21F930.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lasiocera Dejean 1831	<div><p>* Lasiocera Dejean, 1831</p><p>A genus not recorded for Angola yet (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F31B7F7C14BFA443FC21F930	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F7C14BFA073FA6FFA5F.text	03EFF252F31B7F7C14BFA073FA6FFA5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perigona Serrano & Capela & Santos 2017	<div><p>Key to the African Perigona (s.str.) species</p><p>For identification, a new key of the nominate subgenus Perigona Laporte de Castelnau s. str. is provided that is modified from the keys to the African species by Baehr (2004) and Serrano (2008).</p><p>1. Elytra narrow and elongate,&gt;1.4x as long as wide, subparallel; pronotum narrow, not cordiform, widest diameter less close to apex; eyes large and prominent; macropterous, with elongate metepisternum (c. 3x as long as wide), aedeagus with shorter apex............................................................................................... 2</p><p>- Elytra wide and short, &lt;1.3x as long as wide, laterally convex; pronotum fairly wide, subcordiform, widest diameter close to apex; eyes smaller and little prominent; brachypterous, with short metepisternum (c. 1.5x as long as wide); aedeagus with elongate apex (Fig. 5d in Basilewsky 1989). Central Africa..................................... nigrociliata Basilewsky</p><p>2. Smaller species, length &lt;3.9 mm; elytra either uniformly dark yellowish to light brown, or not either uniformly of the same color; terminal abdominal sternite in both sexes with fringe of short hairs; aedeagus short and stout, with rather thin apex of variable length (Fig. 1 in Baehr 2004).................................................................... 3</p><p>- Larger species, length&gt; 4.6 mm; elytra black with light sutural stripe that is enlarged behind middle to a wide spot; terminal abdominal sternite only in female with fringe of short hairs; aedeagus elongate and more depressed, with short, thick apex (Fig. 5c in Basilewsky 1989). Eastern Central Africa............................................ mediornata Basilewsky</p><p>3. Elytra and pronotum uniformly dark yellowish to light brown.................................................. 4</p><p>- Elytra not uniformly of the same color..................................................................... 5</p><p>4. Smaller species, length &lt;2.7 mm; pronotum narrower, little narrowed towards base, lateral margin not sinuate near base, anterior angles not produced; aedeagus with fairly elongate apex (Fig. 5b in Basilewsky 1989). West Africa.... pallida Castelnau</p><p>- Larger species, length&gt; 3.25 mm; pronotum wider, with narrower base, lateral margin gently sinuate near base, anterior angles slightly produced; aedeagus with very short apex (Fig. 5a in Basilewsky 1989). Tropical Africa from Senegal to Mozambique ..................................................................................... parallela Chaudoir</p><p>5. Elytra and pronotum reddish-brown dark or lighter.......................................................... 6</p><p>- Elytra black with narrow reddish sutural stripe; pronotum black, sometimes with indistinct dark reddish discal spot; anterior sclerite in internal sac large, characteristically dentate at upper margin (Fig. 1 in Baehr 2004,). Zambia, southern Africa. ............................................................................................. wachteli Baehr</p><p>6. Elytra reddish-brown dark with light sutural stripe, enlarged behind middle to apex; pronotum reddish-brown dark homogeneous; Elytrae less long (1.50–1.60 times longer than wide); Aedeagus with short apex (Fig. 3 in Serrano 2008). Príncipe Island, Golf of Guinea ................................................................... principensis Serrano</p><p>- Elytra reddish-brown lighter than pronotum, sutural stripe slightly darker; Elitrae slightly longer (1.61–1.62 times longer than wide). Aedeagus with fairly elongate apex (Figs 2a, c, this work). Angola............................ liboloensis sp. n.</p></div>	https://treatment.plazi.org/id/03EFF252F31B7F7C14BFA073FA6FFA5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.text	03EFF252F31B7F4114BFA490FE18FB8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lasiocera schuelei Serrano & Capela & Santos 2017	<div><p>Lasiocera schuelei Serrano sp. n.</p><p>(Figs 1 b, 4, 5)</p><p>Type series. Holotype, ♂; Angola (KWANZA SUL), Munenga 35 km W Calulo (unknown, 147), 3.XII.2012, DO, P. Schüle leg., PSC . Allotype 1♀: same data as Holotype, P. Schüle leg., ASC . Paratypes: Angola (HUÍLA) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.47111&amp;materialsCitation.latitude=-14.148055" title="Search Plazi for locations around (long 14.47111/lat -14.148055)">3.5 km SW Negola</a> (14° 08´53´´ S, 14° 28´16´´ E, 1615 m, 317), 8.XII.2012, 2♀, DO, P. Schüle leg., PSC, ZSM ; Cambamba (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.009167&amp;materialsCitation.latitude=-9.245" title="Search Plazi for locations around (long 17.009167/lat -9.245)">Catoio-Quela</a>) (09° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20.XI.2015, 1♀, DO, A. Serrano leg., ASC .</p><p>Derivatio nominis. This species is cordially dedicated to Peter Schüle, from Herrenberg, a specialist on tiger and ground beetles of Africa which has deeply collaborated with us and kindly provided most of the specimens of this new species.</p><p>Diagnosis. Body elongate, parallel, slightly convex and greenish-bronze (Fig. 1 b); integument densely and deeply punctate, shagrened; surface among punctures with micropunctures visible at high magnification. Pronotum subrectangular or subtrapezoidal shaped (Fig. 4). Four reduced elytral spots. Internal sac of median lobe of aedeagus (Fig. 5) with tangled membranes, the dorsal twisted one closer to the apex (lateral view), this slightly bent to the left (dorsal view) (Fig. 5).</p><p>Description. Length of Holotype: 4.82 mm. Length of paratypes (all females): 4.95‒5.74 mm.</p><p>Head. Triangular (Fig. 1 b), 1.2‒1.3 times wider than long [length: 0.77 mm (holotype), 0.83‒0.88 mm (female allotype and paratypes), width: 0.99 mm (holotype), 1.02‒1.14 mm (female allotype and paratypes)], greenish bronze, shagreened, closely and deeply punctate, surface intervals distinctly micropunctate; mandibles and labrum brownish dark; eyes large, proeminent; palpi light yellowish, apical half of the last articles darker, last article of both palpi fusiform, sharp at the tip; labrum with anterior margin triangularly emarginated; frontal forrows distinct, subparallel, extended to level of anterior supraorbital seta, joining to the inner ocular suture; frons slightly convex or depressed, occiput slightly concave; antennae flavescent, brown dark in inner side of 1st to 10th antennomers, surpassing the posterior margin of pronotum by the last two-three antennomers, 3rd to 10th with 4‒5 long bristles just bellow the joints, scape 2.5 times (holotype) and 2.3‒2.7 times (female allotype and paratypes) longer than wide [length: 0.32 mm (holotype), 0.31‒0.38 mm (female allotype and paratypes), width: 0.13 mm (holotype), 0.13‒0.14 mm (female allotype and paratypes)], pedicele 1.3 times (holotype) and 1.1‒1.4 times (female allotype and paratypes) longer than wide [length: 0.10 mm (holotype), 0.10‒0.11 mm (female allotype and paratypes), width: 0.08 mm (holotype), 0.08‒0.10 mm (female allotype and paratypes)], 3rd to 4th antennomers 2.1 times (holotype) and 2.1‒2.5 times (female allotype and paratypes) longer than wide [length: 0.21 mm (holotype), 0.19‒ 0.21 mm (female allotype and paratypes), width: 0.10 mm (holotype and female allotype and paratypes)], 5th to 10th 2.3‒2.6 times (holotype) and 2.2‒3.2 times (female allotype and paratypes) longer than wide [length: 0.21‒0.24 mm (holotype), 0.22‒0.32 mm (female allotype and paratypes), width: 0.09‒0.10 mm (holotype), 0.09‒0.10 mm (female allotype and paratypes)], and 11th 2.7 times (holoptype) and 3‒3.4 times (female allotype and paratypes) longer than wide [length: 0.27 mm (holotype), 0.27‒0.31 mm (female allotype and paratypes), width: 0.10 mm (holotype), 0.09 mm (female allotype and paratypes)]. Cephalic chaetotaxy (large setae): Labrum with three pairs of setae gradually shorter from external to inner sides, one pair on sides of clypeus and two pairs of supraocular setae present over each eye.</p><p>Thorax. Pronotum (Fig. 4) 1.36 times (holotype) and 1.23‒1.35 times (female allotype and paratypes) longer than wide [length: 0.99 mm (holotype), 1.01‒1.18 mm (female allotype and paratypes), width: 0.73 mm (holotype), 0.75‒0.91 mm (female allotype and paratypes)], greenish bronze, shiny, closely and deeply punctate, surface intervals distinctly micropunctate; wider than head, slightly subrectangular or inverted subtrapezoidal shaped, concussively widest at first third or close to the anterior angles; apical and basal margins not margined, lateral margins with complete and distinct margination in the two first thirds since the anterior angles, reaching the posterior angles, proepisterna well visible from above in the two first thirds; apical margin slightly emarginated, apical angles largely rounded, not protruded; lateral margins slightly subparallel or oblique at first two thirds, followed by a slight sinuosity before the roundly right basal angles, posterior margin slightly but distinctly arcuate; median line absent or very superflous, not reaching anterior or posterior margins, anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous; one marginal setae situated in the end of first third, the posterior marginal seta absent.</p><p>Elytra (Fig. 1 b) 1.76 times (holotype) and 1.71‒1.82 times (female allotype and paratypes) longer than wide [length: 2.78 mm (holotype), 2.88‒3.36 mm (female allotype and paratypes), width: 1.58 mm (holotype), 1.68‒ 1.87 mm (female allotype and paratypes)], greenish bronze, shiny and with punctual metallic reflections; wider than pronotum, humeral angles broadly rounded; elongate and parallel, dorsally moderately convex, truncated obliquely behind, posterior obtuse angle sometimes slightly produced in a short spine, apical margin with 7 to 10 small denticles since the posterior angle to the sutural angle; striae deeply punctate, intervals narrow and convexes, transversely strongly rugose, surface distinctly micropunctate; yellowish patches: one sub-basal and other submedian on 6th interval, more or less rounded or in a small longitudinal line, and two joint small longitudinal subapicals, respectively on 5th and 6th intervals; three setiferous punctures situated at 3rd interval, one at basal quarter, other at median region and the last one at beginning of the last quarter.</p><p>Ventral surface. Blachish-bronze, shiny; genae sparsely finely punctate, glabrous; proepisterna closely and deeply alveolo-punctate in the upper sides, becoming less densely punctate towards the ventral region; sternum glabrous; elytral epipleura smooth with some sparse and deeply punctures at first half; abdominal segments sparcely micropunctate, the first two covered lateraly with sparce dressed short hairs; last segment with 1 pair (male) or 2 pairs (female) of setae near the posterior margin;</p><p>Legs. Slender, yellowish, almost transparent; knees, apical region of tibiae and tarsi brownish dark; trochanters black.</p><p>Aedeagus (Fig. 5). Slender, median lobe slightly curved and apex slightly upwards (lateral view); internal sac with tangled membranes very weekly sclerotized twisted in a pre-dorso-apical position; apex slightly bent to left (dorsal view); parameres like figure 5a.</p><p>Intraspecific variation. The range of variability observed in L. schuelei sp. n. (5 specimens) affects the pronotum shape (see description), the elytral yellow pattern and the number of teeth in apical elytrae margin (see description).</p><p>Concerning the variability within the elytral yellow patches, sometimes the sub-basal patch is absent in one elytron or on both, other times both the sub-basal and the median ones are also absent, remaining solely the subapicals. Asymmetries in the length of left and right elytron are common too.</p><p>Remarks. The species of this genus are characterized, among other features (see Péringuey 1896), by the convex pronotum inflated lateraly in the median region and, consequently, presenting an oblong-ovate or fusiform shape more or less developed. The new species by the singular pronotum shape (subrectangular or even subtrapezoidal), presenting the lateral margins parallel or slightly oblique convergently backwards, is easily segregate from the remaining species. It seems closer to L. egregia Péringuey, 1896 by the reduced elytral patches, the elongate body, parallel elytra and the posterior elytral angle slightly denticled.</p><p>Ecological notes. Adult specimens from Munenga (Calulo) were found during day on open, moist patches with sparse vegetation within a meadow with a few single trees. Adult specimens from Negola were found also during day, walking on a sunshine loamy path after rain, within an open secondary forest. The single female specimen from Cambamba (Catoio-Quela) was found beneath litter in the edges of a countryside road with sandy soil whitin a secondary open forest (Fig. 9b) together with the tiger beetle L. wellmani and the ground beetles C. convexicollis and G. ellipticus .</p></div>	https://treatment.plazi.org/id/03EFF252F31B7F4114BFA490FE18FB8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3267F4114BFA6FBFA6FF87B.text	03EFF252F3267F4114BFA6FBFA6FF87B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lasiocera Dejean 1831	<div><p>Key to the Southern African species of Lasiocera</p><p>An identification key for the southern African species, including the new species, is provided.</p><p>1. Elytra punctato-striate, intervals more or less plain, not transversely rugose....................................... 2</p><p>-. Elytra punctato-striate, intervals transversely rugose.......................................................... 3</p><p>2. Pronotum elongate-ovatum; elytrae depressed, intervals larger, an elongated sub-basal yellow patch on the 5th interval, a small dot below it on the 4th, three adjoining one another in the median region on 5th, 6th and 7th intervals respectively, two in the posterior region on the 3rd and 4th, two more a little lower down on the 5th and 6th and a supra-apical one close to the suture; legs yellowish, knees and tarsi brownish (Botswana, Namibia and South Africa)........................ peringueyi Kuntzen</p><p>-. Pronotum more transverse-ovatum; elytrae sub-convex, intervals narrower, three yellowish, arched transverse patches composed of elongated spots, the first in the basal region on the 3rd, 4th, 5th and 6th intervals respectively, the second middle stripe on the 2nd, 3rd, and 4th intervals, and the last from the 3rd to the 7th interval forming an arch, the last spot of which is somewhat separate and no longer elongated but round and a single standing small spot on the 7th interval close to the tip of the elytra; legs with femora, tibia tip and tarsi brown-blackish, tibia yellowish (Kenya, Tanzania, Mozambique, Zimbabwe and South Africa).......................................................................................... tesselata Klug</p><p>3. Pronotum subrectangular or trapezoidal inverted; elytrae yellowish patches: one sub-basal, other sub-median, both on 6th interval, more or less rounded or in a small longitudinal line, and two joint small longitudinal sub-apicals, respectively on 5th and 6th intervals (Fig. 1 b) (Angola).................................................................. schuelei sp. n.</p><p>-. Pronotum oblong-ovate or fusiform....................................................................... 4</p><p>4. Pronotum oblong-ovate; elytrae depressed, parallel, deeply and closely punctato-striate, intervals narrow, sub-convex, an elongated longitudinal pale-yellow patch slightly notched in the middle on both sides situated in the first half near the lateral margin; another post-median, moderately large, transverse, and uneven one of the same color (South Africa R.)..................................................................................................... gracilis Boheman</p><p>-. Pronotum fusiform; elytrae a little convex, sub-parallel, deeply and broadly punctato-striate, intervals very narrow, convex; four round, rather indistinct, yellowish patches: a sub-basal median and a post-median one in a longitudinal line, a nearly juxtasutural round dot close to the post-median patch a little lower down (Mozambique, Zimbabwe).......... egregia Peringuey</p></div>	https://treatment.plazi.org/id/03EFF252F3267F4114BFA6FBFA6FF87B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA47FFEC3F86C.text	03EFF252F3277F4014BFA47FFEC3F86C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus andersoni Chaudoir 1870	<div><p>Graphipterus andersoni Chaudoir, 1870</p><p>Distribution in Angola (Provinces): 1) Huíla, Cunene; 2) Bié.</p><p>Material examined. Mumbué (Cuanza riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 3. IV.2014, 1 ♀, 12. XI.2015, 4 ♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8. XI.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. Basilewsky (1977) refered to this species as a synonymous of G. amabilis Boheman, 1860 . Lorenz (2005) restablished G. andersoni as a good species and considers the subspecies ovipennis Péringuey, 1892 as well as subovipennis Basilewsky, 1977 as synonymous, both with the status of subspecies of G. amabilis in Basilewsky (1977) . Here we follow Lorenz (2005) criterium. The species is known from Angola, Botswana, Namibia and Zimbabwe (Basilewsky 1977, Insectoid.info 2017). Adults were found on sandy whitish soil or on reddish soil together with some adults of tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3277F4014BFA47FFEC3F86C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA1BDFAF6FACD.text	03EFF252F3277F4014BFA1BDFAF6FACD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus congoensis subsp. lundanus Basilewsky 1977	<div><p>Graphipterus congoensis lundanus Basilewsky, 1977</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.</p><p>Material examined. Chingueia-Mandongue (13° 35´46´´ S, 16° 58´59´´ E, 1648 m alt., 302) (BIÉ), 9.XI.2015, 2♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A subspecies known only from Angola and described on the basis of Caluango specimens (Basilewsky 1977). Our specimens fit very well this subspecies. Adult specimens were found within an open secondary forest together with the tiger beetle E. muata parallelestriata, among other tiger and ground beetles (see S. lucidus strigiceps remarks). Interestingly, both species present a similar pattern of elytral bands (Figs 6a, 6b) suggesting a possible mimicry. Moreover, both species ressemble in the running behaviour also, thus being difficult to segregate them in situ without its capture. Similar observations were already reported to other Graphipterini / Cicindelini associations in Africa (Cassola &amp; Taglianti 1988). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3277F4014BFA1BDFAF6FACD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA0FEFDE9FC8B.text	03EFF252F3277F4014BFA0FEFDE9FC8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus Latreille 1802	<div><p>Graphipterus Latreille, 1802</p><p>A very speciose genus with nineteen species and ten subspecies known for Angola (Ferreira 1965, Basilewsky 1977, Insectoid.info 2017). The genus was split in two super-groups (similitarsi and diversitarsi), each one with several groups (Basilewsky 1977).</p></div>	https://treatment.plazi.org/id/03EFF252F3277F4014BFA0FEFDE9FC8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA2EBFB25FF48.text	03EFF252F3277F4014BFA2EBFB25FF48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pentagonica Schmidt-Goebel 1846	<div><p>Pentagonica Schmidt-Goebel, 1846</p><p>A very speciose genus with three species known for Angola (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3277F4014BFA2EBFB25FF48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA339FB4AFD85.text	03EFF252F3277F4014BFA339FB4AFD85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pentagonica Schmidt-Goebel 1846	<div><p>Pentagonica sp.</p><p>Distribution in Angola (Provinces): Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 6.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A singleton female specimen was found on a wall near public lights (Caála) with other ground beetles (see E. peringueyi remarks). It was not possible to ascribe the specimen to any species.</p></div>	https://treatment.plazi.org/id/03EFF252F3277F4014BFA339FB4AFD85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4714BFA2ADFC9FFDE1.text	03EFF252F3207F4714BFA2ADFC9FFDE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus erychssoni subsp. erychssoni Peringuey 1892	<div><p>Graphipterus erychssoni erychssoni Péringuey, 1892</p><p>Distribution in Angola (Provinces): 1) Cunene; 2) Bié.</p><p>Material examined. Chitembo (6 km SE) (13° 33´23´´ S, 16° 47´22´´ E, 1586 m alt., 302) (BIÉ), 9.XI.2015, 2♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known from Angola and Namibia (Basilewsky 1977, Insectoid.info 2017) with two subspecies (jambanus Basilewsky, 1977 and ornatissimus Basilewsky, 1977), both restricted to the southern territory of Angola (Basilewsky 1977, 1986a). Adults were found within an open secondary forest on sandy whitish soil together with some adults of tiger and ground beetles (see S. lucidus strigiceps remarks). The comments given to G. congoensis lundanus on mimicry associations between Cicindelini and Graphipterini can be reported also to this species (Figs 6a, 6c). It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3207F4714BFA2ADFC9FFDE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4614BFA791FC36FE37.text	03EFF252F3207F4614BFA791FC36FE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus albomarginatus subsp. albomarginatus Quedenfeldt 1883	<div><p>Graphipterus albomarginatus albomarginatus Quedenfeldt, 1883</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Malanje, Benguela, Huambo, Huíla, Bié; 2) Kwanza Sul.</p><p>Material examined. Cruzeiro (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.902636&amp;materialsCitation.latitude=-12.778094" title="Search Plazi for locations around (long 15.902636/lat -12.778094)">Pedra do Alemão</a>) (12° 46´41.14´´ S, 15° 54´09.49´´ E, 1743 m alt., 232) (HUAMBO), 4. XI.2014, 2 ♀, 7. XI.2015, 1 ♀, DO, A Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.914744&amp;materialsCitation.latitude=-9.921345" title="Search Plazi for locations around (long 14.914744/lat -9.921345)">Calulo-Cabuta</a> (9° 55´16.84´´ S, 14° 54´53.08´´ E, 826 m alt., 128) (KWANZA SUL), 5. XI.2014, 3 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.930278&amp;materialsCitation.latitude=-10.038334" title="Search Plazi for locations around (long 14.930278/lat -10.038334)">Calulo-Cabeba</a> (10° 02´18´´ S, 14° 55´49´´ E, 1061 m alt., 147) (KWANZA SUL), 5. XI.2014, 3 ♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.354167&amp;materialsCitation.latitude=-12.302778" title="Search Plazi for locations around (long 15.354167/lat -12.302778)">Londuimbali-Tinguita</a> (12° 18´10´´ S, 15° 21´15´´ E, 1633 m alt., 231) (HUAMBO), 2. XI.2015, 5 ♂, 21♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.221666&amp;materialsCitation.latitude=-13.020833" title="Search Plazi for locations around (long 15.221666/lat -13.020833)">Longonjo-Catabola</a> (13° 01´15´´ S, 15° 13´18´´ E, 1528 m alt., 279) (HUAMBO), 4. XI.2015, 3 ♂, 5♀, DO, A. Serrano &amp; R. Capela leg., ASC ; Cuima- <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.572778&amp;materialsCitation.latitude=-13.345833" title="Search Plazi for locations around (long 15.572778/lat -13.345833)">Cusse</a> (13km) (13° 20´45´´ S, 15° 34´22´´ E, 1633 m alt., 280) (HUAMBO), 5. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Ceilunga (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=17.005556&amp;materialsCitation.latitude=-12.2277775" title="Search Plazi for locations around (long 17.005556/lat -12.2277775)">Kuito</a>) (12° 13´40´´ S, 17° 00´20´´ E, 1683 m alt., 235) (BIÉ), 10. XI.2015, 1 ♂,DO, A. Serrano &amp; R. Capela leg., ASC ; Kinglês (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=7.7666664&amp;materialsCitation.latitude=-9.383333" title="Search Plazi for locations around (long 7.7666664/lat -9.383333)">Lombe-Kalandula</a>) (9° 23´00´´ S, 16° 0´7 46´´ E, 1121 m alt., 113) (MALANJE), 18. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Santa Maria (= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.9783325&amp;materialsCitation.latitude=-8.920556" title="Search Plazi for locations around (long 15.9783325/lat -8.920556)">Quiluanje</a>) (8° 55´14´´ S, 15° 58´42´´ E, 1157 m alt., 94) (MALANJE), 19. XI.2015, 2 ♂, DO, A. Serrano &amp; R. Capela leg., ASC ; Calulo (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.910555&amp;materialsCitation.latitude=-10.037223" title="Search Plazi for locations around (long 14.910555/lat -10.037223)">Faz. Klein</a>) (10° 02´14´´ S, 14° 54´38´´ E, 1059 m alt., 147) (KWANZA SUL), 3. XII.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.943055&amp;materialsCitation.latitude=-9.944722" title="Search Plazi for locations around (long 14.943055/lat -9.944722)">Calulo</a> (N. Faz. Belo Horizonte) (9° 56´41´´ S, 14° 56´35´´ E, 882 m alt., 128) (KWANZA SUL), 3. XII.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. The nominal species is known only from Angola (Basilewsky 1977, 1986b). Probably is the most common and widespread Graphipterus species in Angola. Adults were found in footpaths and country roads within open secondary forests running quickly when disturbed to hide beneath the litter. They are syntopic with some tiger and ground beetles, changing the species in accordance with the locality.</p></div>	https://treatment.plazi.org/id/03EFF252F3207F4614BFA791FC36FE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA683FC07F908.text	03EFF252F3217F4614BFA683FC07F908.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus alternatus Burgeon 1928	<div><p>Graphipterus alternatus Burgeon, 1928</p><p>Distribution in Angola (Provinces): 1) Huambo, Huíla, Cuando Cubango; 2) Bié.</p><p>Material examined. Cachingues-Chipica (13° 10´15.74´´ S, 16° 45´48.17´´ E, 1649 m alt., 281) (BIÉ), 1. XI.2014, 6 ♂, 4♀, DO, A. Serrano &amp; R. Capela leg., ASC; Cachingues (2 km E) (13° 04´54´´ S, 16° 46´08´´ E, 1647 m alt., 282) (BIÉ), 11. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Cuima-Cusse (13km) (13° 20´45´´ S, 15° 34´22´´ E, 1633 m alt., 280) (HUAMBO), 5. XI.2015, 1 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic species of Angola (Basilewsky 1977). In Bié Province the adult specimens were found within secondary meadows on sandy whitish soil together with some tiger and ground beetles [Cachingues- Chipica: P. angusticollis, Dromica similis Cassola, 1980, T. nubifera, L. infuscatula; Cachingues: P. angusticollis, D. similis, D. ( Foveodromica) sp., L. infuscatula, L. saraliensis, Lophyra sumlini Cassola, 1976, Trianogenius ferox (Erichson, 1843), A. convexipennis]. In the Huambo locality the adult specimen was collected together with L. saraliensis and G. albomarginatus . It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3217F4614BFA683FC07F908	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA051FE8AFBE9.text	03EFF252F3217F4614BFA051FE8AFBE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus limbatus Laporte de Castelnau 1840	<div><p>Graphipterus limbatus Laporte de Castelnau, 1840</p><p>Distribution in Angola (Provinces): 1) Cunene; 2) Bié, Cuando Cubango.</p><p>Material examined. Chitembo (6 km SE) (13° 33´23´´ S, 16° 47´22´´ E, 1586 m alt., 302) (BIÉ), 9. XI.2015, 2 ♂, DO, A. Serrano &amp; R. Capela leg., ASC; Munquengue (14° 42´01´´ S, 17° 24´14´´ E, 1440 m alt., 343) (CUANDO CUBANGO), 14. XI.2015, 1 ♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout southern Africa (Namibia, South Africa, Bostwana, Zambia, Zimbabwe) (Basilewsky 1977, 1986b). Putzeys (1880) recorded already the species for Angola (Humbe, Cunene Province), but Basilewsky (1977) considered erroneous this report. However, our specimens fit very well the species description, confirming the species presence in Angola. Moreover, their presence in the southeastern territory of Angola is very plausible, according to the known distribution in the northern territory of Namibia close to the Cunene River. Adult specimens were found on whitish sandy soil near Chitembo within an open secondary forest together with some tiger and ground beetles ( T. nzingae, O. rufomarginata poggei, L. uncivittata, G. erikssoni) and near Munquengue within shrub savannah together with some tiger beetles ( Dromica pentheri W. Horn, 1899 and Ophryodera rufomarginata bradshawi Péringuey, 1888). It is a new record for the Bié and Cuando Cubango Provinces.</p></div>	https://treatment.plazi.org/id/03EFF252F3217F4614BFA051FE8AFBE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4514BFA55BFB54FE37.text	03EFF252F3217F4514BFA55BFB54FE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Graphipterus ellipticus subsp. ellipticus Burgeon 1928	<div><p>* Graphipterus ellipticus ellipticus Burgeon, 1928</p><p>Distribution in Angola (Provinces): 2) Malanje.</p><p>Material examined. Cambamba (Catoio-Quela) (09° 14´42´´ S, 17° 00´33´´ E, 1178 m alt., 115) (MALANJE), 20.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC; Santa Maria (=Quiluanje) (8° 55´14´´ S, 15° 58´42´´ E, 1157 m alt., 94) (MALANJE), 19.XI.2015, 1♂, 2♀, DO, A. Serrano &amp; R. Capela leg., ASC; Carima (Cacuso-Pungandongo) (09° 35´00´´ S, 15° 42´01´´ E, 1117 m alt., 130) (MALANJE), 23.XI.2015, 3♂, 12♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. The nominal species is known from D. R. of the Congo and Zambia (Basilewsky 1977, 1986b) and the subspecies G. ellipticus upembanus Basilewsky, 1977 from the same countries plus Tanzania (Basilewsky op. cit.). Adult specimens collected in Malanje province fit well the nominal species description. Adults were found within open secondary forests together with tiger and ground beetles [Cambamba (Catoio-Quela) and Carima: see S. vilhenai and C. convexicollis remarks, respectively]. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3217F4514BFA55BFB54FE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA4B0FD19F8F9.text	03EFF252F3227F4514BFA4B0FD19F8F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catascopus Kirby 1825	<div><p>Catascopus Kirby, 1825</p><p>A speciose genus with three species known for Angola (Alves 1963, Ferreira 1965, Insectoid.info 2017). One species was collected during our entomological trips.</p></div>	https://treatment.plazi.org/id/03EFF252F3227F4514BFA4B0FD19F8F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA3AFFE03FDE0.text	03EFF252F3227F4514BFA3AFFE03FDE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Somoplatus Dejean 1829	<div><p>Somoplatus Dejean, 1829</p><p>A genus with three species known from Angola (Basilewsky 1986a, Insectoid.info 2017), of which was collected one in our entomological trips.</p></div>	https://treatment.plazi.org/id/03EFF252F3227F4514BFA3AFFE03FDE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA0C1FB8BFBE9.text	03EFF252F3227F4514BFA0C1FB8BFBE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Somoplatus fulvus Mulsant & Godart 1867	<div><p>Somoplatus fulvus Mulsant &amp; Godart, 1867</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.</p><p>Material examined. Satchijamba (10 km E) (13° 43´38.76´´ S, 17° 12´17.74´´ E, 1601 m alt., 283) (BIÉ), 7.IV.2014, 1♀, VM, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout western, central, southern and eastern Africa (Senegal, Gambia, Guinea, Burkina Faso, Ivory Coast, Nigeria, Cameroon, D. R. of the Congo, Angola, South Africa R., Malawi, Kenya) (Basilewsky 1986a, Schüle 2009, Insectoid.info 2017). It was imported to France also (Basilewsky 1986a). The only record for Angola was reported by Basilewsky (1986a) apparently to Lunda Norte province, but without any exact locality. The species seems associated with vacated termite nests of Macrotermes sp. ( Isoptera) (Basilewsky 1986a). The single adult female was collected by light trapping within an open secondary forest with other ground beetles (see S. harpaloides remarks). It is a new reord for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3227F4514BFA0C1FB8BFBE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA683FEF1F92C.text	03EFF252F3227F4514BFA683FEF1F92C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetragonoderus quadrimaculatus Gory 1833	<div><p>Tetragonoderus quadrimaculatus Gory, 1833</p><p>Distribution in Angola (Provinces): 1) Lunda Norte?; 2) Bié.</p><p>Material examined. Chitembo (10 km N) (13° 22´40.78´´ S, 16° 41´58.07´´ E, 1666 m alt., 282) (BIÉ), 1.XI.2014, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout western, central and southern Africa (Mauritania, Senegal, Gambia, Guinea-Bissau, Guinea, Chad, Ivory Coast, Mali, Angola, D. R. of the Congo, South Africa R.) (Serrano 2009, Insectoid.info 2017). It is with some reserve that we ascribe the singleton male adult specimen to this species. The apex of median lobe of aedeagus is slightly different of T. quadrimaculatus and more similar to the same organ of T. quadrum (Fabricius, 1792) . However, this species is distributed on northern territories of Africa. The adult specimen was found in a country road within an open secondary forest. It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3227F4514BFA683FEF1F92C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4414BFA506FDC3FE12.text	03EFF252F3227F4414BFA506FDC3FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catascopus beauvoisi Laporte de Castelnau 1835	<div><p>Catascopus beauvoisi Laporte de Castelnau, 1835</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Colé-Calulo) (09° 59´42´´ S, 14° 50´32´´ E, 1116 m alt., 128) (KWANZA SUL), 4.XII.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widespread throughout western, central and eastern Africa (Guinea, Ivory Coast, Togo, Liberia, Benin, Cameroon, Angola, D. R. of the Congo, Malawi, Tanzania) (Alves 1963, Insectoid.info 2017). The species was recorded for Angola by Alves (1963) (Dundo, Lunda Norte Province), but it was not given in Ferreira`s catalogue (1965). The single male collected near Calulo confirms its presence in Angola. The adult specimen was found under tree bark together with C. congolensis (see also C. intermedius remarks). It is a new record for the Kwanza Sul Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3227F4414BFA506FDC3FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA767FE4CF9D0.text	03EFF252F3237F4414BFA767FE4CF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arsinoe laevigata Basilewsky 1970	<div><p>* Arsinoe laevigata Basilewsky, 1970</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Cachingues-Chipica (13° 10´15.74´´ S, 16° 45´48.17´´ E, 1649 m alt., 281) (BIÉ), 2.IV.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known to western and central Africa (Guinea-Bissau, Guinea, Ivory Coast, Central African Republic, D. R. of the Congo) (Basilewsky 1970, Serrano 2009, Insectoid.info 2017). The single female specimen fits well with the original description and with specimens from Guinea-Bissau. The adult specimen was collected by light trapping together with other ground beetles (see P. holosericus remarks). It is a new genus/ species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4414BFA767FE4CF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA612FB26FB7F.text	03EFF252F3237F4414BFA612FB26FB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arsinoe Laporte de Castelnau 1835	<div><p>* Arsinoe Laporte de Castelnau, 1835</p><p>A genus not recorded until now for Angola (Ferreira 1965, Facchini 2011b, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4414BFA612FB26FB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA0A5FD58FBB1.text	03EFF252F3237F4414BFA0A5FD58FBB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coptodera (Coptoderina) congolensis Burgeon 1937	<div><p>* Coptodera (Coptoderina) congolensis Burgeon, 1937</p><p>Distribution in Angola (Provinces): 2) Kwanza Sul.</p><p>Material examined. Alto Ventura (Colé-Calulo) (09° 59´42´´ S, 14° 50´32´´ E, 1116 m alt., 128) (KWANZA SUL), 4.XII.2015, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widespread throughout western, central and eastern Africa (Senegal, Togo, Cameroon, Equatorial Guinea, Gabon, D. R. of the Congo, Rwanda, Uganda) (Hansen 1968, Insectoid.info 2017). Adult specimens were found under tree bark together with P. liboloensis sp. n. and C. beauvoisi (see also C. intermedius remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4414BFA0A5FD58FBB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA073FAA3FD3C.text	03EFF252F3237F4414BFA073FAA3FD3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coptodera Dejean 1825	<div><p>Coptodera Dejean, 1825</p><p>A genus with four species known for Angola (Ferreira 1965, Hansen 1968, Insectoid.info 2017) distributed by three subgenera ( Coptoderina Jeannel, 1949, Coptoderinella Hansen, 1968 and Haplocrepis Jeannel, 1949).</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4414BFA073FAA3FD3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA4B0FCFFF89D.text	03EFF252F3237F4414BFA4B0FCFFF89D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dontolobus Basilewsky 1970	<div><p>Dontolobus Basilewsky, 1970</p><p>A genus recently recorded for Angola (Facchini 2012a).</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4414BFA4B0FCFFF89D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4B14BFA506FE04FE37.text	03EFF252F3237F4B14BFA506FE04FE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dontolobus aemiliae Facchini 2012	<div><p>Dontolobus aemiliae Facchini, 2012</p><p>Distribution in Angola (Provinces): 1) Lunda Norte, Huambo, Bié.</p><p>Material examined. Satchijamba-Somakwanza (13° 44° 46.50°´S, 17° 11° 2.26´´ E, 1611 m alt., 303) (BIÉ), 2.XI.2014, 2♂, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered known distribution in Africa (Angola, Zambia, Tanzania) (Facchini 2012a). It was described recently by Facchini (2012a) and our adult specimens fit very well its description, including the aedeagus features. Adult specimens were collected by light trapping within an open secondary forest together with some other ground and ant nest beetles (e.g. P. vanrooni, P. chappuisi, M. guineensis, A. monardianus, Stenolophus sp.).</p></div>	https://treatment.plazi.org/id/03EFF252F3237F4B14BFA506FE04FE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA612FBECFB7F.text	03EFF252F32C7F4B14BFA612FBECFB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymindoidea Laporte de Castelnau 1833	<div><p>Cymindoidea Laporte de Castelnau, 1833</p><p>A genus with one species reported for Angola (Basilewsky 1961, Ferreira 1965).</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4B14BFA612FBECFB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA767FDFAF9D0.text	03EFF252F32C7F4B14BFA767FDFAF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymindoidea virgulifera Chaudoir 1875	<div><p>Cymindoidea virgulifera Chaudoir, 1875</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X‒ 2.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species with a large distribution in Africa south of Sahara (Senegal, Guinea-Bissau, Ivory Coast, Mali, Chad, D. R. of the Congo, Angola, Zambia, Zimbabwe, Uganda, Mozambique, Tanzania, Kenya, Ethiopia, Sudan) (Basilewsky 1961, Ferreira 1965, Serrano 2009, Insectoid.info 2017). The single female specimen was collected in a wall near a public light together with other ground beetles (see C. rikatlensis remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4B14BFA767FDFAF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA4B0FBE9F89D.text	03EFF252F32C7F4B14BFA4B0FBE9F89D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyaulacus Chaudoir 1878	<div><p>* Polyaulacus Chaudoir, 1878</p><p>A genus not recorded until now for Angola (Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4B14BFA4B0FBE9F89D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA3AFFBF1FDE0.text	03EFF252F32C7F4B14BFA3AFFBF1FDE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyreopterus Dejean 1831	<div><p>Thyreopterus Dejean, 1831</p><p>A genus with tree species known for Angola (Ferreira 1965, Insectoid.info 2017) distributed by two subgenera [ Thyreopterus (two species) and Thyreopterinus Alluaud, 1932 (one species)].</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4B14BFA3AFFBF1FDE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA0C1FB16FBB1.text	03EFF252F32C7F4B14BFA0C1FB16FBB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyreopterus flavosignatus Dejean 1831	<div><p>Thyreopterus flavosignatus Dejean, 1831</p><p>Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.</p><p>Material examined. Satchijamba (10 km E) (13° 43° 38.76´´ S, 17° 12° 17.74´´ E, 1601 m alt., 283) (BIÉ), 7.IV.2014, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout western, central, southern and eastern Africa (Senegal, Gambia, Guinea-Bissau, Cameroon, Central African Republic, D. R. of the Congo, Angola, Namibia, South Africa R., Kenya) (Basilewsky 1955, Ferreira 1965, Serrano 2009, Insectoid.info 2017). Adult specimens were collected under the bark of decomposing trunks together with Zygentoma, Formicidae and Tenebrionidae adults and the ant nest beetle P. pseudoklugi within an open secondary forest. It is a new record for the Bié Province.</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4B14BFA0C1FB16FBB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4A14BFA506FE06FE6F.text	03EFF252F32C7F4A14BFA506FE06FE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyaulacus brunneus Chaudoir 1878	<div><p>* Polyaulacus brunneus Chaudoir, 1878</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Chitembo (10 km N) (13° 22´40.78´´ S, 16° 41´58.07´´ E, 1666 m alt., 282) (BIÉ), 1.XI.2014, 1♀, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species widespread throughout western, central, southern and eastern Africa (Guinea-Bissau, Guinea, Ivory Coast, D. R. of the Congo, South Africa R., Rwanda, Zambia, Zimbabwe, Mozambique, Tanzania) (Serrano 2009, Insectoid.info 2017). The singleton female specimen fits well the original description and is similar to Guinea-Bissau specimens (see Serrano 2009). The adult specimen was collected by light trapping within an open secondary forest together with a tiger beetle ( L. sumlini) and ant nest beetles [ Heteropaussus trapezicollis (Wasmann,1922), Paussus (Spinicoxipaussus) cridae Gestro, 1915] (see Serrano &amp; Capela 2015a, 2015b). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F32C7F4A14BFA506FE06FE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA057FC67FD3C.text	03EFF252F32D7F4A14BFA057FC67FD3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cylindrocranius Chaudoir 1878	<div><p>Cylindrocranius Chaudoir, 1878</p><p>A genus with one species reported for Angola (Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F32D7F4A14BFA057FC67FD3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA0C1FE4CFB8D.text	03EFF252F32D7F4A14BFA0C1FE4CFB8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cylindrocranius errans Peringuey 1896	<div><p>* Cylindrocranius errans Péringuey, 1896</p><p>Distribution in Angola (Provinces): 2) Luanda.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.166269&amp;materialsCitation.latitude=-9.315394" title="Search Plazi for locations around (long 13.166269/lat -9.315394)">Barra do Cuanza</a> (09° 18´55.42´´ S, 13° 09´58.57´´ E, 11 m alt., 107) (LUANDA), 28.IV.2014, 2♀, LT, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species known until now from Botswana and Namibia (Insectoid.info 2017). Adult specimens fit very well the original description and are similar to a specimen from Namibia [South of Walvis Bay, "Musee Royal de l’Afrique Centrale (Belgium, Tervuren)”] and determined by P. Basilewsky (in Anichtchenko 2016). Adults were collected by light trapping close to River Cuanza within a riparian forest together with one ant nest beetle [ Carabidomemnus (Carabidodoxus) sp.] and two ground beetles ( D. picicornis and M. guineensis). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F32D7F4A14BFA0C1FE4CFB8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA7D0FA67F909.text	03EFF252F32D7F4A14BFA7D0FA67F909.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drypta (Deserida) disctincta (Rossi 1792)	<div><p>* Drypta (Deserida) disctincta (Rossi, 1792)</p><p>Distribution in Angola (Provinces): 2) Huambo.</p><p>Material examined. Caála (12° 51´11´´ S, 15° 33´07´´ E, 1758 m alt., 256) (HUAMBO), 6.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species widespread throughout the Mediterranean Region, West Asia and Africa (Basilewsky 1960b, Insectoid.info 2017). Curiously, it was never recorded for Angola. A singleton female specimen was found on a wall near public lights with other ground beetles (see E. peringueyi remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F32D7F4A14BFA7D0FA67F909	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA6BFFDD0FAF0.text	03EFF252F32D7F4A14BFA6BFFDD0FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drypta Latreille 1796	<div><p>Drypta Latreille, 1796</p><p>There are two species in this genus known from Angola (Ferreira 1965, Insectoid.info 2017). We collected two species not recorded to Angola yet.</p></div>	https://treatment.plazi.org/id/03EFF252F32D7F4A14BFA6BFFDD0FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4914BFA493FE22FEDC.text	03EFF252F32D7F4914BFA493FE22FEDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drypta (Drypta) thoracica Boheman 1848	<div><p>* Drypta (Drypta) thoracica Boheman, 1848</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 1.IV.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species with a scattered distribution throughout western, central, southern and eastern Africa (Guinea, Ivory Coast, Cameroon, D. R. of the Congo, South Africa R., Rwanda, Tanzania, Kenya) (Basilewsky 1960b, Insectoid.info 2017). The single male was collected by light trapping within an open secondary forest together with P. pseudocrenulatus . It is a new species record for Angola. Dendrocellus Schmidt-Göbel, 1846</p><p>A genus considered a synonymous of Desera Dejean 1825 by Lorenz (2005). However, it was reestablished before as a valid genus by Liang et al. (2004). There are three species known from Angola ( Liang &amp; Kavanaugh 2007, Serrano 2008‒2009).</p></div>	https://treatment.plazi.org/id/03EFF252F32D7F4914BFA493FE22FEDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4914BFA3DAFCF7FC97.text	03EFF252F32E7F4914BFA3DAFCF7FC97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dendrocellus micropectinatus Liang & Kavanaugh 2007	<div><p>Dendrocellus micropectinatus Liang &amp; Kavanaugh, 2007</p><p>Distribution in Angola (Provinces): 1) Malanje; 2) Huambo.</p><p>Material examined. Alto Hama (12° 13´20´´ S, 15° 32´57´´ E, 1512 m alt., 232) (HUAMBO), 31.X‒ 2.XI.2015, 1♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC.</p><p>Remarks. A species known for Angola, Zambia and D. R. of the Congo (Liang &amp; Kavanaugh 2007, Serrano 2008‒2009). It was recorded for the first time to Angola based on one male specimen collected in “Parque Nacional da Cangandala, a protected area for the emblematic giant sable antelope Hippotragus niger variani Thomas, 1916 . The new adult specimens collected in Huambo confirm its presence in Angola. Adult specimens were found at a wall during night near a public light together with other ground beetles (see C. rikatlensis remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F32E7F4914BFA3DAFCF7FC97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4914BFA105FE74FC45.text	03EFF252F32E7F4914BFA105FE74FC45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Galerita Fabricius 1871	<div><p>Galerita Fabricius, 1871</p><p>A speciose genus with six species and one subspecies known from Angola (Basilewsky 1963, Ferreira 1965, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F32E7F4914BFA105FE74FC45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.text	03EFF252F32E7F4E14BFA63BFD41F857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Galerita procera subsp. capelai Serrano & Capela & Santos 2017	<div><p>Galerita procera capelai Serrano ssp. n.</p><p>(Figs 7a, 7c, 8a, 8b, 8c)</p><p>Type series. Holotype, ♂; Angola (Kwanza Sul), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.907222&amp;materialsCitation.latitude=-9.899722" title="Search Plazi for locations around (long 14.907222/lat -9.899722)">Calulo-Cabuta</a> (9° 53´59´´ S, 14° 54´26´´ E, 831 m alt., 128) (KWANZA SUL), 29.XI‒6.XII.2015, PF , A. Serrano &amp; R. Capela leg., ASC . Allotype 1♀: Mussende (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.793334&amp;materialsCitation.latitude=-9.95" title="Search Plazi for locations around (long 14.793334/lat -9.95)">Calulo</a>) (9° 57´00´´ S, 14° 47´36´´ E, 905 m alt., 128) (KWANZA SUL), 28.XI‒5.XII.2015, PF , A. Serrano &amp; R. Capela leg., ASC; Paratypes, 2♂, same locality and date as Holotype, A. Serrano &amp; R. Capela leg., ASC, PSC; 1♀, same locality and date as Allotype, A. Serrano &amp; R. Capela leg., ASC .</p><p>Derivatio nominis. This subspecies is named in honour of Rúben Capela, a specialist on Diptera Culicidae and Ceratopogonidae, which has collaborated with the first author (AS) in the Angola entomological trips.</p><p>Diagnosis. Similar in form and color to G. procera Gerstaecker, 1867; five pairs of impar elytral carinae well developed, the 2nd par carinae (4th carinae) only slightly distinct in the first quarter, 3rd par carinae (6th carinae) distinct in the first and last thirds of elytrae, remaining par carinae almost indistinct, scutellar carina almost indistinct joining the first carina; median lobe of aedeagus with a hooked apex ventraly backwards (Figs 8a, 8b).</p><p>Description. Length of Holotype: 25.5 mm. Length of paratypes (without labrum): 25.32‒25.33 mm (males), 25.89‒26.30 mm (females).</p><p>Head (Fig, 7a) 1.2‒1.3 times longer than wide [length: 4.62 mm (holotype and male paratypes), 4.62‒4.82 mm (female allotype and paratype), width: 3.68 mm (holotype), 3.62‒3.68 mm (male paratypes), 3.71‒3.81 mm (female allotype and paratype)], black, eyes small, slightly wider than temples; front longitudinaly protruded in the median region, almost smooth, two short deeply furrows lateraly; occiput surface largely rugose-punctate, covered with very sparce blackish semierected pubescence, surface intervals smooth; mandibles and labrum blackish; palpi blackish, apical region of the last articles brownish, last article of both palpi strongly securiform; labrum with anterior margin straight or slightly arcuate; antennae proeminent, densely pubescent, decreasing in thickness towards apex, first four antennomers blackish, gradually brown dark to lighter brown since the 5th to the 11th, reaching or slightly surpassing the middle of elytra; Cephalic chaetotaxy (large setae, all brownyellowish): Labrum with three submarginal pairs of long setae, two pairs of long setae in a transversal middle line throughout the first half of clypeus disk, one pair on sides behind this anterior series and two pairs of very long supraocular setae present over each eye; some other not so long setae dispersed over the sides of clypeus and frons.</p><p>Thorax (Fig. 7a) Pronotum 1.13‒1.21 times longer than wide [length: 5.68 mm (holotype), 5.71‒5.74 mm (male paratypes), 5.68‒5.87 mm (female allotype and paratype), width: 5.02 mm (holotype), 4.82‒4.95 mm (male paratypes), 4.69‒4.88 mm (female allotype and paratype)], black, wider than head, barely subcordiform, widest near the middle; lateral margins slightly raised, narrow, regularly round in the first two thirds, slightly subparallel or sinuate before the rectangular basal angles, round at tip; apical angles roundly acute, slightly protruded; apex and base finely margined, the former arcuate, the latter almost straight; disk gently convex; median line in middle slightly more impressed than anteriorly and posteriorly, neither reaching apex nor base; anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous in males, absent in females; basal grooves barely indicated; lateral margin with two elongate setae, one situated slightly before the middle, the other before the basal angle, two‒three short setae near the anterior angles, a fringe of brown-reddish hairs at anterior and posterior margins (shorter in the former); surface strongly transversely punctate-rugose in sides and base, slighter in the disk, covered with sparce blackish semierected pubescence, microreticulation distinct, extremely fine, consisting of transverse meshes, shiny.</p><p>Elytra (Fig. 7a) 1.66‒1.79 times longer than wide [length:, 14.36 mm (holotype), 14.23‒14.36 mm (male paratypes), 14.90‒15.30 mm (female allotype and paratype), width: 8.65 mm (holotype), 8.05‒8.11 mm (male paratypes), 8.31‒8.91 mm (female allotype and paratype)], blackish, shiny, sometimes with violet reflections; wider than pronotum, broadly elongate-ovate, humeral angles not distinct, widest point behind the middle; dorsally very slightly convex, truncated obliquely at apex; each elytron with five distinct impair carinae, the 2nd par carinae (4th carinae) only slightly distinct in the first quarter, 3rd par carinae (6th carinae) distinct in the first and last thirds of elytrae, remaining par carinae almost indistinct, scutellar carina almost indistinct joining the first carina; intervals between two consecutive impair carinae with two longitudinal deep punctate rows, each flanked lateraly by a row of brownish setae from base to apex, totalizing three rows; 4th interval narrower than others, by this with only one longitudinal punctate row flanked lateraly by a row of brownish setae from base to apex; apical truncature with blackish pubescence; surface intervals covered with distinct, dense, transversely punctate-granulate meshes (Fig. 7c); hind wings absent.</p><p>Ventral surface. Blach to brown dark, shiny in the head and thorax, dull in abdomen; genae sparsely punctate, sides covered with sparse black semierected setae; proepisterna smooth, not punctate in the upper sides, becoming sparsely punctate towards the ventral region; metaepisterna, pro, meso and metasterna densely punctate and pubescent (brownish color); elytral epipleura moderately punctate; abdominal segments densely punctate and pubescent like thoracic sterna; posterior margin of last segment deeply (males) or slightly (females) truncate in the middle region, one pair (males and females) of setae near the posterior margin.</p><p>Legs are conformed to the genus morphological pattern. Male forelegs with the three basal tarsomeres of tarsi more dilated in the inner region than in the outer (asymmetrical dilated), the 4th lesser asymmetrical, bilobated, presenting two rows of elongate, sensorial phaneres beneath.</p><p>Aedeagus (Figs 8a, 8b, 8c). Median lobe short, robust, apex hooked ventraly backwards (Fig. 8b). Endophalous with a large postero-latero-dorsal scaly plate. Left paramere large, round at apex, right atrophied.</p><p>Intraspecific variation. The range of variability observed in G. procera capelai ssp. n. (5 specimens) affects the median region of frons, more or less longitudinaly protruded and the pronotum shape (more or less subcordiform). Asymmetries in the length of left and right elytron are common too. The pronotum is partially pubescent or almost glabrous, accordingly with the presence/absence of decumbent setae.</p><p>Remarks. The subspecies of G. procera, including the nominal one, are easily segregated by the apical conformation of the median lobe of aedeagus than by external morphological characters (see Basilewsky1963). The representants of this complex (eight subspecies including the nominal one) are distributed from Angola to Kenya and Tanzania, throughtout D. R. of the Congo, Zambia, Burundi, Rwanda and Uganda. Galerita procera angolana Basilewsky, 1963 was the unique subspecies known until now from Angola. By the recorded localities given (Basilewsky 1963, Ferreira 1965) it seems distributed in the central and eastern territories of Angola (Huambo, Lunda Norte and Moxico Provinces). Interestingly, G. procera capelai ssp. n. by the external morphological characters and shape of apical median lobe of aedeagus seems closer to the nominal subspecies than the already known Angolan subspecies. The nominal subspecies is proper of eastern Africa (Kenya and Tanzania), very faraway of the new subspecies locality. The other seven subspecies are distributed between the two taxa. In southern Africa this singular relationship in which species closer morphologicaly are strongly allopatric (western vs eastern) seems more common than expected (e.g. Serrano 1995). The new subspecies by the singular apical shape of median lobe (ventraly hooked backwards) (Fig. 8b) is easily segregate from the remaining subspecies (see Fig. 20, in Basilewsky 1963). Galerita procera capelai ssp. n. can be separated also from G. procera angolana, the subspecies closer territorialy of the new subspecies, by a very different shape of the median lobe apex (cf Figs 8b vs 8e) and further by the elytral carinae less elevated, the intervals slightly larger and less excavated, the rows between impair carinae shallower punctate and surface intervals covered with more distinct and dense transversely punctate-granulate meshes.</p><p>Ecological notes. Adult specimens were collected by means of pitfall trapping within secondary rainforest patches with moist soils, covered with dense and high litter (Fig. 9c), together with adults of some tiger and ground beetles [ Dromica (Foveodromica) sp., A. optimus, A. distinctus, D. tarsalis, C. magnicollis discrepans, O. gilvipes and O. patroboides]. As happens with some carabid species living mainly at moist habitats (e.g. Rossi &amp; Santamaria 2001) the adult specimens of the new species were more or less infected with ectoparasitic fungi of the order Laboulbeniales on pronotum as well on elytra. Some study cases were already reported to other African carabid species (e.g. Santamaria &amp; Faille 2009).</p></div>	https://treatment.plazi.org/id/03EFF252F32E7F4E14BFA63BFD41F857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32A7F4D14BFA496FCCAF8FB.text	03EFF252F32A7F4D14BFA496FCCAF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meladroma Motschulsky 1855	<div><p>Meladroma Motschulsky, 1855</p><p>There is one species recorded to Angola (Basilewsky 1962).</p></div>	https://treatment.plazi.org/id/03EFF252F32A7F4D14BFA496FCCAF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32A7F4C14BFA5FEFBACFE37.text	03EFF252F32A7F4C14BFA5FEFBACFE37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meladroma informicollis Liebke 1928	<div><p>* Meladroma informicollis Liebke, 1928</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Chitembo (18 km SE) (13° 36´17.53´´ S, 16° 54´00.44´´ E, 1538 m alt., 302) (BIÉ), 30.X.2014, 1♂, LT, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species known from Cameroon, Ivory Coast and D. R. of the Congo (Basilewsky 1962, Insectoid.info 2017). Curioursly M. katangensis Burgeon, 1937, the species reported to Angola by Basilewsky (1962), is referred neither in Ferreira catalogue (1965) nor in Insectoid.info (2017). The singleton male adult specimen of M. informicollis was collected by light trapping within an open secondary forest together with some other ground beetles (see S. angolense remarks). It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F32A7F4C14BFA5FEFBACFE37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA408FB44F965.text	03EFF252F32B7F4C14BFA408FB44F965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractonotus Perroud 1847	<div><p>Atractonotus Perroud, 1847</p><p>There is one species reported to Angola (Kleinfeld &amp; Puchner 2012, Häckel &amp; Farkač 2013b).</p></div>	https://treatment.plazi.org/id/03EFF252F32B7F4C14BFA408FB44F965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA64EFA7FFB22.text	03EFF252F32B7F4C14BFA64EFA7FFB22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Netrodera Chaudoir 1850	<div><p>Netrodera Chaudoir, 1850</p><p>There are three species recorded for Angola (Ferreira 1965, Kleinfeld &amp; Puchner 2012, Häckel &amp; Farkač 2013b).</p></div>	https://treatment.plazi.org/id/03EFF252F32B7F4C14BFA64EFA7FFB22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA683FBBCF998.text	03EFF252F32B7F4C14BFA683FBBCF998.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Netrodera vethi (Bates 1889)	<div><p>Netrodera vethi (Bates, 1889)</p><p>Distribution in Angola (Provinces): 1) Benguela, Huíla; 2) Huambo.</p><p>Material examined. Cuima-Cusse (13° 22´52´´ S, 15° 30´15´´ E, 1583 m alt., 280) (HUAMBO), 5.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic species of Angola described originaly from Benguela (Kleinfeld &amp; Puchner 2012). Adult specimens were found beneath litter within open secondary forests together with some tiger and other ground beetles (see Orthogonius sp. 1 remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F32B7F4C14BFA683FBBCF998	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA0FDFF02FC55.text	03EFF252F32B7F4C14BFA0FDFF02FC55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trianogenius ferox (Erichson 1843)	<div><p>Trianogenius ferox (Erichson, 1843)</p><p>Distribution in Angola (Provinces): 1) Benguela, Namibe, Huambo; 2) Bié.</p><p>Material examined. Cachingues (2 km E) (13° 04´54´´ S, 16° 46´08´´ E, 1647 m alt., 282) (BIÉ), 11.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species spread in Angola and Namibia (Basilewsky 1959, Insectoid.info 2017). Basilewsky (1959) reported that the species was distributed in the western and southwestern Angolan areas. The singleton male adult was found on sandy whitish soil within an open secondary forest together with some tiger beetles (see G. alternatus remarks). It is a new record for the Bié Province enlarging its distribution to the eastern territory of Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F32B7F4C14BFA0FDFF02FC55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F5314BFA55EFAB5FEDA.text	03EFF252F32B7F5314BFA55EFAB5FEDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractonotus puncticollis Schule & Heinz 2013	<div><p>* Atractonotus puncticollis Schüle &amp; Heinz, 2013</p><p>Distribution in Angola (Provinces): 2) Bié.</p><p>Material examined. Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BIÉ), 8‒17.XI.2015, 1♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species described and known only from Zambia (Schüle &amp; Heins 2013). The single female specimen was checked with one female paratype of this species. There is another species within this genus reported to Angola ( A. mulsanti mulsanti Perroud, 1847) which presents a larger distribution throughout central, southern and eastern Africa) (Háva &amp; Hovorka 2007, Kleinfeld &amp; Puchner 2012). The adult was collected through pitfall trapping within an open secondary forest and is syntopic with some tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new species record for Angola, enlarging its distribution to southwestern Africa.</p></div>	https://treatment.plazi.org/id/03EFF252F32B7F5314BFA55EFAB5FEDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA0FDFD0BFC1E.text	03EFF252F3347F5314BFA0FDFD0BFC1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypholoba brevivittis Chaudoir 1866	<div><p>Cypholoba brevivittis Chaudoir, 1866</p><p>Distribution in Angola (Provinces): 1) Benguela, Namibe.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.839158&amp;materialsCitation.latitude=-12.339533" title="Search Plazi for locations around (long 13.839158/lat -12.339533)">Baixo Pundo</a> (12° 20´22.32´´ S, 13° 50´20.97´´ E, 275 m alt., 227/8) (BENGUELA), 25‒27.X.2013, 1♂, 2♀, PF, R. Capela leg., ASC.</p><p>Remarks. An endemic species of Angola originaly described from Benguela (Kleinfeld &amp; Puchner 2012). Adult specimens were collected within bush habitat with sparse grassy vegetation and sandy soil by pitfall trapping, together with the ground beetle Harpalus sp.</p></div>	https://treatment.plazi.org/id/03EFF252F3347F5314BFA0FDFD0BFC1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA3CBFAE9FD84.text	03EFF252F3347F5314BFA3CBFAE9FD84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypholoba Chaudoir 1850	<div><p>Cypholoba Chaudoir, 1850</p><p>A genus very well represented in Angola with fifteen species and two subspecies known within its territory (Ferreira 1965, Kleinfeld &amp; Puchner 2012, Häckel &amp; Farkač 2013b, Schüle 2015, Insectoid.info 2017).</p></div>	https://treatment.plazi.org/id/03EFF252F3347F5314BFA3CBFAE9FD84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA186FDC9FA27.text	03EFF252F3347F5314BFA186FDC9FA27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypholoba gracilis (Dejean 1831)	<div><p>* Cypholoba gracilis (Dejean, 1831)</p><p>Distribution in Angola (Provinces): 2) Benguela.</p><p>Material examined. Chamume (12° 41´38´´ S, 13° 08´51.9´´m E, 6 m alt., 251) (BENGUELA), 28.X.2013, 1♀, PF, R. Capela leg., ASC.</p><p>Remarks. The species is polymorphic with four recognized subspecies distributed mainly throughout southern and eastern Africa (Kleinfeld &amp; Puchner 2012, Insectoid.info 2017). The nominal subspecies occurs in several South Africa Provinces, Botswana and Namibia (Kleinfeld &amp; Puchner 2012). We do not ascribe our singleton female to any known subspecies, but she looks like the nominal form with the gray-whitish haired longitudinal median pronotal sulcus and the elytra totally glabrous. It could be ascribed to Cypholoba edax Péringuey, 1892 also, but our specimen does not present any withish pubescence at the elytral base and the elytrae are more domed. It is a new species record for Angola.</p></div>	https://treatment.plazi.org/id/03EFF252F3347F5314BFA186FDC9FA27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA79FFC08F80C.text	03EFF252F3347F5314BFA79FFC08F80C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypholoba graphipteroides subsp. monardi Burgeon 1935	<div><p>Cypholoba graphipteroides monardi Burgeon, 1935</p><p>Distribution in Angola (Provinces): 1) Huíla, Cunene; 2) Huambo.</p><p>Material examined. Cuima-Cusse (13° 22´52´´ S, 15° 30´15´´ E, 1583 m alt., 280) (HUAMBO), 5.XI.2015, 7♂, 3♀, 6.XI.2015, 1♂, 2♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13° 25´59´´ S, 15° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6.XI.2015, 2♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. The species is very polymorphic (differences in the setal patterns of the elytra), with fifteen recognized subspecies distributed mainly throughout central and eastern Africa (Lorenz 2005, Kleinfeld &amp; Puchner 2012, Insectoid.info 2017). As referred by Mawdsley et al. (2012) “Many of these taxa are doubtfully distinct from the nominate form and the whole group is in need of a careful revision”, which is not the scope of this work. Cypholoba graphipteroides monardi seems proper to the south of Angola (Ferreira 1965). Adults were found in diurnal activity beneath litter and patches within open secondary forests together with other tiger and ground beetles (see S. vilhenai remarks). It is a new record for the Huambo Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3347F5314BFA79FFC08F80C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA45BFDECF815.text	03EFF252F3357F5214BFA45BFDECF815.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthia cinctipennis	<div><p>Anthia (s.str.) cinctipennis actaeon Erichson, 1843</p><p>Distribution in Angola (Provinces): 1) Namibe; 2) Benguela.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.55&amp;materialsCitation.latitude=-12.35" title="Search Plazi for locations around (long 13.55/lat -12.35)">Lobito</a> (12 ° 21´S, 13 ° 33´E, 10 m alt., 227/8) (BENGUELA), 20.III.1998, 2♀, 10.V.1998, 1♂, 1♀, PF, A. Neves leg., ASC; Biópio ‒ Benguela (12 ° 27´38.17´´ S, 13 ° 44´02.92´´ E, 148 m alt., 228) (BENGUELA), 27.X.2013, 1♂, PF, R. Capela leg., ASC .</p><p>Remarks. The nominal species has a wide distribution throughout central, southern and eastern Africa (Angola, Namibia, South Africa, Zambia, Bostwana, Zimbabwe, Mozambique), but this subspecies is proper to South Angola and North Namibia (Kleinfeld &amp; Puchner 2012). Adult specimens were collected by pitfall trapping in the outskirts of Lobito (see also B. apicalis remarks) and near Biópio within open xeric bush in sandy soil. It is a new record for Benguela Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3357F5214BFA45BFDECF815	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA7F3FACAFA60.text	03EFF252F3357F5214BFA7F3FACAFA60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthia Weber 1801	<div><p>Anthia Weber, 1801</p><p>There are three species and one subspecies known for Angola (Ferreira 1965, Kleinfeld &amp; Puchner 2012).</p></div>	https://treatment.plazi.org/id/03EFF252F3357F5214BFA7F3FACAFA60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA2EBFB0BFDA8.text	03EFF252F3357F5214BFA2EBFB0BFDA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cypholoba rohani (Benard 1921)	<div><p>Cypholoba rohani (Bénard, 1921)</p><p>Distribution in Angola (Provinces): 1) Cuando Cubango; 2) Bié.</p><p>Material examined. Chingueia-Mandongue (13° 35´46´´ S, 16° 58´59´´ E, 1648 m alt., 302) (BI), 9. XI.2015, 1 ♂, 2♀, DO, A. Serrano &amp; R. Capela leg., ASC; Munquengue (14° 42´01´´ S, 17° 24´14´´ E, 1440 m alt., 343) (CUANDO CUBANGO), 14. XI.2015, 2 ♀, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic species of Angola described on the basis of one specimen from the Menongue region (not Huíla District, but Cuando Cubango Province) (Bénard 1921 in Kleinfeld &amp; Puchner 2012). Ferreira (1965) reported “distrito do Moxico, Menong (=Menongue) (MP)”, which is not correct also. Adults were found in activity in open secondary forest (first locality) and bush savannah (second locality) on sandy withish soil together with some tiger and ground beetles (see S. lucidus strigiceps and G. limbatus remarks, respectively).</p></div>	https://treatment.plazi.org/id/03EFF252F3357F5214BFA2EBFB0BFDA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA019FEE6FCAD.text	03EFF252F3357F5214BFA019FEE6FCAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Termophilum Basilewsky 1950	<div><p>Termophilum Basilewsky, 1950</p><p>There are six species and three subspecies recorded for Angola (Ferreira 1965, Kleinfeld &amp; Puchner 2012). Lorenz (2005) considers this genus as a subgenus of Anthia Weber, 1801 . Here we follow Kleinfeld &amp; Puchner (2012) systematic.</p></div>	https://treatment.plazi.org/id/03EFF252F3357F5214BFA019FEE6FCAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA116FBA6FA92.text	03EFF252F3357F5214BFA116FBA6FA92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Termophilum massilicatum Guerin-Meneville 1845	<div><p>Termophilum massilicatum Guérin-Méneville, 1845</p><p>Distribution in Angola (Provinces): 1) Unknown; 2) BIÉ.</p><p>Material examined. Satchijamba-Somakwanza (13° 44´46.50´´ S, 17° 11´2.26´´ E, 1611 m alt., 303) (BIÉ), 31.X.2014, 1♂, 1♀, 2.XI.2014, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catota (2,5 km S) (14° 00´37´´ S, 17° 24´00´´ E, 1532 m alt., 323) (BI), 2.XI.2014, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. A species spread throughout southern, central and part of eastern Africa (Angola, Namibia, South Africa, Zambia, Bostwana, Zimbabwe, Mozambique) (Ferreira 1965, Kleinfeld &amp; Puchner 2012). The species was recorded for Angola without any reference to a known locality (Ferreira 1965). Adult specimens were found in nocturnal activity (first locality) and diurnal activity (second locality) within open secondary forests together with other ground beetles (first locality: see S. tenebricosus fatalis remarks) and the tiger beetles P. angusticollis, D. fredericoi, D. pentheri, Dromica serietuberculata W. Horn, 1929, T. pepetela, L. uncivittata and C. agualusai (second locality) (see also Serrano et al. 2015). It is a new record for the Bi Province.</p></div>	https://treatment.plazi.org/id/03EFF252F3357F5214BFA116FBA6FA92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA2EBFEFFFDDF.text	03EFF252F3367F5114BFA2EBFEFFFDDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthia (Odontanthia) convexipennis Putzeys 1880	<div><p>Anthia (Odontanthia) convexipennis Putzeys, 1880</p><p>Distribution in Angola (Provinces): 1) Huíla; 2) Bié.</p><p>Material examined. Cachingues (2 km E) (13 ° 04´54´´ S, 16 ° 46´08´´ E, 1647 m alt., 282) (BIÉ), 11. XI.2015, 1 ♂, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Mumbué (Cuanza riverhead) (13° 49´41´´ S, 17° 19´26´´ E, 1511 m alt., 303) (BIÉ), 8‒17. XI.2015, 1 ♀, PF, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. An endemic species of Angola (Puchner 2013). It was synonimized with Anthia calida Harold, 1878 by Basilewsky (1960), but re-established as a good species by Puchner (2013). Our specimens fit very well this species, presenting the typical subhumeral band (Puchner 2013, figs 9, 10). Adult specimens were found within secondary meadows on sandy whitish soil together with some tiger and ground beetles (see G. alternatus and S. lucidus strigiceps remarks for the first and second localities, respectively). The species was known only for the south of Angola (Huíla Province), but the new localities enlarges its distribution for the eastern territory (Bié Province).</p></div>	https://treatment.plazi.org/id/03EFF252F3367F5114BFA2EBFEFFFDDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA0C7FE04FB89.text	03EFF252F3367F5114BFA0C7FE04FB89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthia (Odontanthia) kleinfeldi Puchner 2013	<div><p>Anthia (Odontanthia) kleinfeldi Puchner, 2013</p><p>Distribution in Angola (Provinces): 1) Huambo, Huíla.</p><p>Material examined. Bembua-Camenhe (12 ° 16´53´´ S, 15 ° 27´01´´ E, 1686 m alt., 231) (HUAMBO), 1.XI.2015, 1♀, DO, A. Serrano &amp; R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59´´ S, 15 ° 21´22´´ E, 1607 m alt., 279) (HUAMBO), 6.XI.2015, 1♂, DO, A. Serrano &amp; R. Capela leg., ASC .</p><p>Remarks. Another endemic species of Angola localized in the Huambo and Huíla Provinces (Puchner 2013). Adult specimens fit very well the Puchner (2013) description, including the aedeagus. Adults were found beneath litter within an open secondary forest (first locality) and in an open secondary bush savannah (second locality) together with some other tiger and ground beetles (see S. lucidus strigiceps and S. vilhenai remarks for the first and second localities, respectively).</p></div>	https://treatment.plazi.org/id/03EFF252F3367F5114BFA0C7FE04FB89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Serrano, Artur R. M.;Capela, Rúben A.;Santos, Carmen Van-Dúnem Neto	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
