taxonID	type	description	language	source
03EFF252F3007F6714BFA38FFF2CFCE7.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA38FFF2CFCE7.taxon	materials_examined	Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227 / 8) (BENGUELA), 3. IV. 1998, 1 ♀, PF, A. Neves leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA38FFF2CFCE7.taxon	discussion	Remarks. A species found in D. R. of the Congo, Zimbabwe and Angola (Liebke 1934). The singleton female adult was collected by pitfall trapping in open bush habitat with sparse grassy vegetation on sandy-clayey yellowreddish soil in the outskirts of Lobito together with some other ground beetles [Calosoma (Calosoma) senegalense Dejean, 1831, Parophonus (Hyparpalus) tomentosus (Dejean, 1829), Anthia (s. str.) cinctipennis actaeon Erichson, 1843].	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA6F0FEF1FA28.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA6F0FEF1FA28.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.74 ´´ S, 16 ° 45´48.17 ´´ E, 1649 m alt., 281) (BIÉ), 2. IV. 2014, 1 ♂, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6714BFA6F0FEF1FA28.taxon	discussion	Remarks. A polymorphic species widely distributed throughout western and central Africa (see Serrano 2008, 2009 and references herein). Adults were sampled in a secondary open forest. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6614BFA57BFE91FEFF.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown; 2) Benguela. Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227 / 8) (BENGUELA), 3. IV. 1998, 1 ♀, PF, A. Neves leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3007F6614BFA57BFE91FEFF.taxon	discussion	Remarks. A species widespread throughout Africa south of Sahara, Cape Vert Islands and Madagascar (Häckel et al. 2016). It was already cited to Angola (Häckel et al. 2016, Insectoid. info 2017), but without exact data. The single female adult was collected by pitfall trapping in an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Lobito together with some other ground beetles (see B. apicalis remarks).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA3E7FC8FFC88.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown; 2) Luanda.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA3E7FC8FFC88.taxon	materials_examined	Material examined. Luanda (8 ° 50´S, 13 ° 15´E, 70 m alt., 89) (LUANDA), 27. IV. 1974, 1 ♀, DO, A. Serrano leg., ASC; Luanda near Motor racing circuit (8 ° 59´S, 13 ° 05´E, 50 m alt., 89) (LUANDA), 12. VI. 2016, 1 ♀, DO, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA3E7FC8FFC88.taxon	discussion	Remarks. A species widespread thoughout central, southern and eastern Africa (Angola, Botswana, D. R. of the Congo, Zambia, R. of South Africa, Lesotho, Swaziland, Zimbabwe, Malawi, Uganda, Mozambique, Tanzania, Kenya, Ethiopia, Somalia) and Madagascar (Häckel et al. 2016). Like the precedent species it was already known to Angola (Häckel et al. 2016, Insectoid. info 2017), but without any recorded locality. The first female adult was found in an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Luanda and the second on ground during night near a public light.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA6DBFB51FA04.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown; 2) Luanda.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA6DBFB51FA04.taxon	materials_examined	Material examined. Barra do Cuanza (09 ° 18´55.42 ´´ S, 13 ° 09´58.57 ´´ E, 11 m alt., 107) (LUANDA), 28. IV. 2014, 4 ♂, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6614BFA6DBFB51FA04.taxon	discussion	Remarks. A species widely spread throughout all Africa south of Sahara (e. g. Ferreira 1965). This author only refers to the presence of the species in Angola, without specifying localities. Adults were sampled close to River Cuanza by light trapping within a riparian forest. It is a new record for the Luanda Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6514BFA55EFC7FFCAA.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Malanje; 2) Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6514BFA55EFC7FFCAA.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 231) (HUAMBO), 1. XI. 2015, 3 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Chitembo (6 km SE) (13 ° 33´23 ´´ S, 16 ° 47´22 ´´ E, 1586 m alt., 302) (BIÉ), 9. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Chingueia-Mandongue (13 ° 35´46 ´´ S, 16 ° 58´59 ´´ E, 1648 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♂, 1 ♀, PF, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♂, PF, 12. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3017F6514BFA55EFC7FFCAA.taxon	discussion	Remarks. A species spread throughout Democratic Republic of the Congo, Zimbabwe, Angola, Tanzania and Mozambique (Basilewsky, 1955). Adults were found within secondary open forest on reddish soil together with some adults of tiger and ground beetles [Bembua-Camenhe: Prothyma (s. str.) erythropyga erythropyga (Putzeys, 1880), Dromica (Foveodromica) spp., Prothymidia angusticollis angusticollis (Boheman, 1848), Cylindera agualusai Serrano & Capela, 2015, Graphipterus albomarginatus Quedenfeldt, 1883] or on sandy whitish soil [Chitembo: Trichotaenia nzingae Oesterle, Serrano & Capela, 2015, Lophyra (Stenolophyra) uncivittata (Quedenfeldt, 1883); Chingueia-Mandongue: Dromica fredericoi Serrano & Capela, 2015, T. nzingae, Elliptica muata f. parallelestriata (W. Horn, 1923), Graphipterus congoensis lundanus Basilewsky, 1977 and Cypholoba rohani (Bénard, 1921); Mumbué: Dromica pentheri W. Horn, E. muata f. parallelestriata, Graphipterus andersoni Chaudoir, 1870, Anthia (Odontanthia) convexipennis Putzeys, 1880; Catota: D. fredericoi, Trichotaenia pepetela Serrano & Capela, 2015, Chlaenius (Pachydinodes) bipustulatus Boheman, 1848, Abacetus pseudoangolensis, Siopelus (Neosiopelus) wrasei Facchini, 2012, Graphipterus andersoni and Atractonotus puncticollis Schle & Heinz, 2013]. It is a new record for the Huambo and Bié Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA11BFD94F9CC.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo; 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA11BFD94F9CC.taxon	materials_examined	Material examined. Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Kalandula (1,5 km N) (09 ° 03´51 ´´ S, 15 ° 58´42 ´´ E, 1126 m alt., 112) (MALANJE), 18. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Cambamba (Catoio- Quela) (09 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Mutubanzamba (Cangandala) (09 ° 47´45 ´´ S, 16 ° 31´40 ´´ E, 1136 m alt., 132) (MALANJE), 21. XI. 2015, 2 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Carima (Cacuso-Pungandongo) (09 ° 35´00 ´´ S, 15 ° 42´01 ´´ E, 1117 m alt., 130) (MALANJE), 23. XI. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA11BFD94F9CC.taxon	discussion	Remarks. An Angolan scaritid beetle endemic species whose description was based on one single female from Monte Moco (Huambo province). Our specimens conform well to the Basilewsky (1955) description. Adult specimens were found whitin secondary open forests on sandy whitish soil (first locality) or on reddish soil (Malanje localities) together with some adults of tiger and ground beetles [Catata-Nova Monção: Euryarthron reticostatum (Wellman & W. Horn, 1908), Dromica (Foveodromica) sp., P. angusticollis, Trichodela nubifera (Quedenfeldt, 1883), Lophyra (Stenolophyra) saraliensis saraliensis (Guérin-Méneville, 1849), Lophyra (Bothryolophyra) wellmani (W. Horn, 1907), Chlaenius ovalipennis, Euleptus intermedius Péringuey, 1896, Orthotrichus patroboides (Murray, 1859), Orthogonius sp. 1, Netrodera vethi (Bates, 1889), Cypholoba graphipteroides monardi Burgeon, 1935 and Anthia (Odontanthia) kleinfeldi Puchner, 2013; Malanje localities: Dromica (Foveodromica) spp., P. angusticollis, T. nubifera, Epitrichodes villosus (Putzeys, 1880), Lophyra (Stenolophyra) infuscatula (W. Horn, 1915), L. saraliensis, L. wellmani, Graphipterus ellipticus Burgeon, 1928]. It is a new record for the Malanje Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA4B5FA9AF865.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA4B5FA9AF865.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 31. X. 2014, 1 ♂, 2. XI. 2014, 3 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3027F6514BFA4B5FA9AF865.taxon	discussion	Remarks. A subspecies spread throughout Angola and D. R. Congo (Katanga) (Bäninger 1938). Adults were found in activity on sandy whitish soil within a secondary open forest during night together with other ground beetles (e. g. Systolocranius tibialis Lecordier, 1972, C. bipustulatus, Termophilum massilicata Guérin-Méneville, 1845).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA31DFCBEFC86.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA31DFCBEFC86.taxon	materials_examined	Material examined. Satchijamba (10 km E) (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 283) (BIÉ), 7. IV. 2014, 1 ♀, VM, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA31DFCBEFC86.taxon	discussion	Remarks. A species only reported to South Africa (Cape Province) (Kult 1959). The specimen was identified using Kult`s (1959) key, taking into account that our specimen is dark ferruginous, shining (elytra without microsculture), with frons impunctate, proepisternae finely punctate, protibiae sulcate with 3 rd spine almost indistinct, prothorax with transverse wrinkles and group of punctures on each side and mesotibiae without denticle. The unique feature that does not agree with C. capensis description is the antennal antenomers 4 to 10: moniliform in C. capensis, but subelongate in our specimen. For the present, we consider that this specimen belongs to this species with some reserves. The single female specimen was found in open secondary forest and collected by means of light trapping. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA6F6FAB7F9BA.taxon	distribution	Distribution in Angola (Provinces): 1) Cunene; 2) Luanda, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA6F6FAB7F9BA.taxon	materials_examined	Material examined. Barra do Cuanza (09 ° 18´55.42 ´´ S, 13 ° 09´58.57 ´´ E, 11 m alt., 107) (LUANDA), 28. IV. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC; Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 283) (BIÉ), 2. XI. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6414BFA6F6FAB7F9BA.taxon	discussion	Remarks. A species widespread throughout almost all Africa south of Sahara, including S. Tomé Island (e. g. Straneo 1959, Ferreira 1965, Serrano 1995). Adults were collected by light trapping within a riparian forest (Barra do Cuanza) and an open secondary forest (Satchijamba-Somakwanza). In the later locality the adults were collected together with ant nest beetles [Paussus (Shuckardipaussus) vanrooni Wasmann, 1922, Paussus (Katapaussus) chappuisi Reichensperger, 1938] (see Serrano 2015 a). It is a new record for the Luanda and Bié Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6B14BFA55CFD06FEDA.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo; 2) Bi.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6B14BFA55CFD06FEDA.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3037F6B14BFA55CFD06FEDA.taxon	discussion	Remarks. A species described on one specimen from Bailundo (Huambo province) (Müller 1940) and known also from D. R. of the Congo (Straneo 1952, 1991). The single adult was sampled within an open secondary forest by light trapping together with other ground beetles (e. g. Stereostoma guineense Müller, 1940, Brachyodes pseudoguineensis Lecordier, 1990, Parophonus escheri (Dejean, 1831), Siopelus harpaloides (Guérin-Méneville, 1847), Siopelus natalicus Péringuey, 1896, Axinotoma latipalpis Basilewsky, 1968 and Meladroma informicollis Liebke, 1928). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA682FEFEF962.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Huambo; 2) Malanje, Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA682FEFEF962.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X ‒ 2. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Mutubanzamba (Cangandala) (09 ° 47´45 ´´ S, 16 ° 31´40 ´´ E, 1136 m alt., 132) (MALANJE), 21. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Alto Ventura (Colé- Calulo) (09 ° 59´42 ´´ S, 14 ° 50´32 ´´ E, 1116 m alt., 128) (KWANZA SUL), 4. XII. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC;	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA682FEFEF962.taxon	discussion	Remarks. A species with a distribution throughout Angola, D. R. of the Congo, Zimbabwe, Tanzania (Zanzibar) and Mozambique (Straneo 1942, 1952). In the first locality the adult specimen was found in a wall during night near a public light. In the second locality, the adult was collected within the litter which covered the soil and, in the third locality, the adults were found under bark of trees together with adults of other Coleoptera families (e. g. Endomychidae, Trogossitidae, Tenebrionidae). It is a new record for the Malanje and Kwanza Sul Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA3CBFAAFFC78.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Lunda Norte; 2) Bié, Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA3CBFAAFFC78.taxon	materials_examined	Material examined. Chitembo (13 ° 30´55.22 ´´ S, 16 ° 45´20.34 ´´ E, 1628 m alt., 302) (BIÉ), 8. IV. 2014, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC; Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. 2015, 3 ♂, 3 ♀, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (KWANZA SUL), 27. XI ‒ 5. XII. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30C7F6B14BFA3CBFAAFFC78.taxon	discussion	Remarks. A species described on one specimen from Bailundo also (Huambo Province) (Müller 1940) and widespread throughout western and central Africa (Straneo 1952, 1991). The single male adult collected in the second locality was sampled by light trapping within an open secondary forest together with some other ground beetles (e. g. see S. angolense remarks). The remaining adults were found in activity on ground in the road edges during night, probably attracted to the public lights. It is a new record for the Bié and Kwanza Sul Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA116FE11FAEF.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA116FE11FAEF.taxon	materials_examined	Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo (Faz. Klein) (10 ° 02´14 ´´ S, 14 ° 54´38 ´´ E, 1059 m alt., 147) (KWANZA SUL), 3. XII. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo-Cabuta (9 ° 53´59 ´´ S, 14 ° 54´26 ´´ E, 831 m alt., 128) (KWANZA SUL), 29. XI ‒ 6. XII. 2015, 6 ♂, 2 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA116FE11FAEF.taxon	discussion	Remarks. A species widespread throughout Angola, D. R. of the Congo, Zimbabwe and South Africa (Straneo 1958, 1959). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e. g. Dromica (Foveodromica) sp., Hipparidium interruptum (Fabricius, 1775), Abacetus optimus Péringuey, 1904, Disphericus tarsalis Bates, 1886, Craspedophorus magnicollis discrepans Basilewsky, 1987, Galerita procera capelai ssp. n.] and collected also by pitfall trapping in the same habitats. It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA7D7FAE3F908.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA7D7FAE3F908.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.14 ´´ S, 16 ° 45´33.52 ´´ E, 1668 m alt., 282) (BIÉ), 2. IV. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA7D7FAE3F908.taxon	discussion	Remarks. A species widely spread in Africa (e. g. Cape Vert, Ivory Coast, Ethiopia, Mozambique, Nigeria, Angola, Congo, D. R. of the Congo, Zimbabwe and Rwanda) (Straneo 1956, 1958, 1959, 1968). The single male was collected by light trapping within a secondary open forest. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA2EBFAA4FE70.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Moxico; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA2EBFAA4FE70.taxon	materials_examined	Material examined. Satchijamba (10 km E) (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 283) (BIÉ), 7. IV. 2014, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6A14BFA2EBFAA4FE70.taxon	discussion	Remarks. A species with a distribution throughout Angola, D. R. of the Congo, Zimbabwe and South Africa (Straneo 1942, 1958, 1959). The singleton adult was collected under the bark of a decomposing trunk together with Zygentoma, Formicidae and Tenebrionidae adults and the ant beetle Paussus (Klugipaussus) pseudoklugi Luna de Carvalho, 1963 within an open secondary forest (see Serrano 2015 a). It is a new record for the Bié province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6914BFA578FCE7FE37.taxon	distribution	Distribution in Angola (Provinces): 1) Cunene; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6914BFA578FCE7FE37.taxon	materials_examined	Material examined. Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♂, PF, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 2 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30D7F6914BFA578FCE7FE37.taxon	discussion	Remarks. An endemic ground beetle species of Angola from which is known so far the male and female types (Straneo 1952). Both adults were sampled through pitfall trapping. The Mumbué specimen was collected in sandy whitish soil within a mixed open herbaceous / bush habitat close to the riverhead together with adults of the tiger beetle E. muata f. parallelestriata and the ground beetles S. lucidus strigiceps, E. intermedius, Systolocranius tibialis and Anthia convexipennis Putzeys, 1880. The Catota specimens were collected through pitfall trapping in sandy clayish reddish soil within a secondary open forest together with adults of some ground beetles (see S. lucidus remarks). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA3AFFD95FC1C.taxon	distribution	Distribution in Angola (Provinces): 1) Moxico; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA3AFFD95FC1C.taxon	materials_examined	Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI. 2015, 1 ♂, 2 ♀, 5. XII. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Mussende (Calulo) (9 ° 57´00 ´´ S, 14 ° 47´36 ´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI ‒ 5. XII. 2015, 3 ♀, PF, A. Serrano & R. Capela leg., ASC; Calulo-Cabuta (9 ° 53´59 ´´ S, 14 ° 54´26 ´´ E, 831 m alt., 128) (KWANZA SUL), 29. XI ‒ 6. XII. 2015, 8 ♂, 9 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA3AFFD95FC1C.taxon	discussion	Remarks. A species widespread throughout Angola, Zimbabwe, Mozambique and South Africa (Straneo 1958, 1959, 1991). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e. g. Dromica (Foveodromica) sp., A. distinctus, Disphericus tarsalis, C. magnicollis discrepans, G. procera capelai ssp. n.] and collected also by pitfall trapping in the same habitats. It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA185FB2FFA91.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Moxico; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA185FB2FFA91.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA185FB2FFA91.taxon	discussion	Remarks. An endemic ground beetle species of Angola from which is known so far the male type (Straneo 1952) and a second specimen (unknown sex) from Calundo Lake (Straneo 1959). The specimen fits very well with the original description and was captured by light trapping within a secondary open forest together with ground and ant nest beetles (M. guineensis, P. vanrooni, P. chappuisi). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA447FD78F830.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA447FD78F830.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30E7F6914BFA447FD78F830.taxon	discussion	Remarks. A species only reported to D. R. of the Congo (Straneo 1983). The specimen was identified using Straneo`s (1983) key and the specimen fits well the original description (Burgeon 1935). For the present, until the acquisition of new data, we consider that this specimen belongs to this species with some reserves. The single adult was collected under the bark of a decomposing trunk together with Zygentoma, Formicidae and Tenebrionidae adults. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA6F0FCC9F9F5.taxon	distribution	Distribution in Angola (Provinces): 1) Malanje, Huíla; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA6F0FCC9F9F5.taxon	materials_examined	Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 2. XII. 2015, 1 ♂), 4. XII. 2015, 1 ♂, 5. XII. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA6F0FCC9F9F5.taxon	discussion	Remarks. The nominal species was described from Angola (Malanje) (Quedenfeldt 1883) and has a distribution throughout Angola and D. R. of the Congo (Häckel & Farkač 2012). Basilewsky (1987) described two subspecies, C. magnicollis discrepans from the Lulua region (D. R. of the Congo) and Zambia and C. magnicollis inquilinus from Mayidi (D. R. of the Congo), giving the drawings of aedeagus apex (dorsal view) of the three taxa. Interestingly our Calulo specimens present the aedeagus apex similar to the C. magnicollis discrepans (Basilewsky 1987, Fig. 4 c), enlarging its distribution to Angola. Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles [e. g. Dromica (Foveodromica) sp., A. distinctus, A. optimus]. It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6F14BFA4EEFC68FF47.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6F14BFA4EEFC68FF47.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. ‒ 2. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6F14BFA4EEFC68FF47.taxon	discussion	Remarks. A species widespread throughout Angola, D. R. of the Congo, Burundi, Zambia and Mozambique (Basilewsky 1955, Häckel & Farkač 2012, 2013 a, Insectoid. info 2017), presenting some variability on the elytra maculae. Adults were found during night near public lights together with other ground beetles [e. g. S. guineense, Chlaenius cupreocinctus Reiche, 1847, Ch. tenuicollis (Fabricius, 1801), Progonochaetus planicollis (Putzeys, 1880), Agonidium alacre (Boheman, 1848), Agonidium natalense (Boheman, 1848), E. intermedius, Euleptus peringueyi Csiki, 1931]. It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA3C6FDC3FCE7.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA3C6FDC3FCE7.taxon	materials_examined	Material examined. Mussende (Calulo) (9 ° 57´00 ´´ S, 14 ° 47´36 ´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI. 2015, 1 ♂, DO, 28. XI ‒ 5. XII. 2015, 2 ♀, PF, A. Serrano & R. Capela leg., ASC; Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI. 2015, 1 ♂, 5. XII. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30F7F6814BFA3C6FDC3FCE7.taxon	discussion	Remarks. An endemic ground beetle species of Angola from which is known so far the type (Bates 1886) and a second specimen (unknown sex) from Dundo (Basilewsky 1955). Adults were found in activity within the litter of secondary rainforests together with adults of some tiger and ground beetles (e. g. Dromica (Foveodromica) sp., A. distinctus, A. optimus, C. magnicollis discrepans) and also felt in pitfall traps in the same habitats. It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA036FB5AFCC2.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA036FB5AFCC2.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA036FB5AFCC2.taxon	discussion	Remarks. A species with a distribution throughout D. R. Congo and Burundi (Lecordier 1990). It can be segregated from B. guineensis (Chaudoir, 1882), to which is very close, by the smaller antennae and the unarmed mesosternum groove. The adult was sampled by light trapping within a secondary open forest together with some other ground beetles (see S. angolense remarks). It is a new genus / species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA6D5FE3CF9BD.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA6D5FE3CF9BD.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 3 ♂, 3 ♀, DO, A. Serrano & R. Capela leg., ASC; Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♂, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3087F6F14BFA6D5FE3CF9BD.taxon	discussion	Remarks. A species spread throughout Angola and D. R. of the Congo (Katanga) (Lecordier 1972). Adults in the first locality were found in activity on sandy whitish soil within a secondary open forest during night together with other ground beetles [e. g. S. tenebrosus fatalis, C. (Pachydinodes) bipustulatus, T. massilicata] It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA38DFD88FCE5.taxon	distribution	Distribution in Angola (Provinces): 2) Luanda.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA38DFD88FCE5.taxon	materials_examined	Material examined. Luanda near Motor racing circuit (08 ° 59´S, 13 ° 05´E, 50 m alt., 89) (LUANDA), 12. VI. 2016, 1 ♂, 1 ♀, DO, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA38DFD88FCE5.taxon	discussion	Remarks. A species with a scattered distribution throughout Africa south of Sahara (Mauritania, Senegal, Ivory Coast, Togo, Mali, Chad, Democratic Republic of the Congo, Eritrea, Ethiopia, Kenya, Tanzania, Zimbabwe, South Africa), but known from West Asia (Oman) and some Atlantic Islands (Cape Verde, Saint Helena and São Tomé and Príncipe) also (Insectoid. info 2017). Adult specimens were found on a wall during night near a public light. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA1DEFB93F9BA.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte?; 2) Malanje, Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA1DEFB93F9BA.taxon	materials_examined	Material examined. Kibundi (15 km N Nambua) (8 ° 56´15 ´´ S, 16 ° 03´52 ´´ E, 1164 m alt., 95) (MALANJE), 19. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA1DEFB93F9BA.taxon	discussion	Remarks. A species widespread throughout Africa south of Sahara (Senegal, Gambia, Guinea, Ivory Coast, Cameroon, Burkina Faso, Chad, Democratic Republic of the Congo, Burundi, Rwanda, Zambia, Zimbabwe, Uganda, Kenya, Tanzania, Ethiopia, Eritrea, Yemen) (Insectoid. info 2017), but not recorded to Mozambique, South Africa and Namibia. It was reported for Angola by Alves (1963). However, the data given in this work (locality, date) for the species seems similar to the specimen which was identified as C. deyrollei (see the precedent species) by Basilewsky (1955). The female adult specimen (first locality) was found in the edges of a countryside road within an open secondary forest while the male adult specimen (second locality) was collected within the litter of a secondary rainforest. Both adults were found together with adults of some tiger and ground beetles [first locality: e. g. Dromica (Foveodromica) sp., A. optimus, A. distinctus, D. tarsalis, C. magnicollis discrepans, G. procera capelai n. ssp.; second locality: Dromica (Foveodromica) sp., L. wellmani]. These new data confirm this species’ presence in Angola and extends its distribution to Malanje and Kwanza Sul Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA2EBFE49FE14.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA2EBFE49FE14.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 5. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6E14BFA2EBFE49FE14.taxon	discussion	Remarks. A species widespread throughout western and central Africa since Senegal to Angola (Basilewsky 1955, Insectoid. info 2017). The singleton adult male was found during night near a public light. It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6D14BFA42BFBE7FF6A.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Huambo; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6D14BFA42BFBE7FF6A.taxon	materials_examined	Material examined. Cruzeiro (Pedra do Alemão) (12 ° 46´41.14 ´´ S, 15 ° 54´09.49 ´´ E, 1743 m alt., 232) (HUAMBO), 7. XI. 2015, 1 ♂, 1 ♀, DO, A Serrano & R. Capela leg., ASC; Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 3 ♀, DO, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 2 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3097F6D14BFA42BFBE7FF6A.taxon	discussion	Remarks. A polymorphic ground beetle species with a distribution throughout Africa south of Sahara. The nominal subspecies seems restricted to the southern countries of Africa (Angola, Namibia, Zimbabwe, South Africa and Mozambique) (Insectoid. info 2017). Adults were found beneath litter (first locality) or in activity on sandy whitish soil within a secondary open forest (second locality) during night together with other ground beetles (e. g. S. tenebrosus fatalis, S. tibialis, T. massilicata). In the third locality the specimens were collected by pitfall trapping within a secondary open forest. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA682FBADF997.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA682FBADF997.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X ‒ 2. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA682FBADF997.taxon	discussion	Remarks. A species with a known scattered distribution throughout Africa: Ethiopia, D. R. of the Congo and South Africa (Anichtchenko 2016, Insectoid. info 2017). Adults were found also on ground during night near public lights (see comments on C. rikatlensis remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA40FFBB1F868.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA40FFBB1F868.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Cachingues (13 ° 04´54 ´´ S, 16 ° 46´08 ´´ E, 1647 m alt., 282) (BIÉ), 11. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA40FFBB1F868.taxon	discussion	Remarks. A subspecies known exclusively from D. R. of the Congo (Basilewsky 1949, Insectoid. info 2017). Our specimens fit very well the Basilewsky`description. Adults were found on ground during night near public lights (first locality) (see also C. rikatlensis remarks) and beneath litter on sandy whitish soil within an open secondary forest (second locality). It is a new species / subspecies record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA35BFA66FCAC.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Malanje, Bié, Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA35BFA66FCAC.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Mutubanzamba (9 ° 47´45 ´´ S, 16 ° 31´40 ´´ E, 1136 m alt., 132) (MALANJE), 21. XI. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA35BFA66FCAC.taxon	discussion	Remarks. A species described from Quango based on one male of collections by Mechow (Quedenfeldt 1883). Our specimens fit very well the original description, including the figure 10 of the plate 3. There is a report of this species to Lunda by Monard et al. (1956) (sub Chlaenius ovalipennis Gaf.?). Surprisingly the species is not reported for Angola in the Ferreira (1965)`s catalogue or even for the neighbouring countries (e. g. D. R. of the Congo, Zambia, Namibia) in Anichtchenko (2016) and Insectoid. info (2017) sites. Adults were found in all localities beneath litter within secondary open forests together with some tiger and ground beetles (Catata-Nova Monção and Mutubanzamba: see S. vilhenai remarks). It is a new record for Malanje, Huambo and Bié Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA115FBA9FB21.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Lunda Norte.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA115FBA9FB21.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 1 ♀, 4. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30A7F6D14BFA115FBA9FB21.taxon	discussion	Remarks. A species with a scattered distribution in Africa: Angola, Zimbabwe, Ethiopia and Eritrea (Ferreira 1965, Anichtchenko 2016, Insectoid. info 2017). Adults were found on ground during night near public lights together with other ground beetles (e. g. see C. rikatlensis and P. incrassatus remarks).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA3A9FAE1FCAD.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA3A9FAE1FCAD.taxon	materials_examined	Material examined. Calulo (Faz. Klein) (10 ° 02´14 ´´ S, 14 ° 54´38 ´´ E, 1059 m alt., 147) (KWANZA SUL), 1. XII. 2015, 1 ♂, 3. XII. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA3A9FAE1FCAD.taxon	discussion	Remarks. The species was reported for Angola by Basilewsky (1955) under the name C. hessei Basylewsky, 1949. The nominal species is widespread throughout Angola, D. R. of the Congo, Zambia, Zimbabwe, South Africa and Tanzania (Basilewsky 1955, Anichtchenko 2016, Insectoid. info 2017). Adults were found in the edges of small rain pounds, running quickly when disturbed. It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA116FD77FBC7.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA116FD77FBC7.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♀, 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA116FD77FBC7.taxon	discussion	Remarks. A species widespread throughout all Africa south of Sahara (Basilewsky 1955, Anichtchenko 2016, Insectoid. info 2017). Adults were found on ground during night near public lights (see also C. rikatlensis remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA6BFFAA6F9BC.taxon	distribution	Distribution in Angola (Provinces): 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA6BFFAA6F9BC.taxon	materials_examined	Material examined. Kibundi (15 km N Nambua) (8 ° 56´15 ´´ S, 16 ° 03´52 ´´ E, 1164 m alt., 95) (MALANJE), 19. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Cambamba (Catoio-Quela) (9 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 3 ♂, 3 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA6BFFAA6F9BC.taxon	discussion	Remarks. A species with a scattered known distribution in Africa (D. R. of the Congo and Sudan) (Anichtchenko 2016, Insectoid. info 2017). Adults were found in the edges of small rain pounds together with adults of the tiger beetle C. agualusai, running quickly when disturbed. It is a new species record to Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA2EBFD61FE38.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA2EBFD61FE38.taxon	materials_examined	Material examined. Bicuar, Parque Nacional (unknown) (HUÍLA), 21. II. 2014, 2 ♂, PF, R. Capela leg., ASC. Remarks. A species restricted to Angola (Ferreira 1965, Anichtchenko 2016, Insectoid. info 2017). It seems a common species in the Huíla Province by the reported localities given (Ferreira 1965). Adults were collected by pitfall trapping within a corn field plantation.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA425FCC1F8F9.taxon	distribution	Distribution in Angola (Provinces): Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA425FCC1F8F9.taxon	materials_examined	Material examined. Carima (Cacuso-Pungandongo) (9 ° 35´00 ´´ S, 15 ° 42´01 ´´ E, 1117 m alt., 130) (MALANJE), 23. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F30B7F6C14BFA425FCC1F8F9.taxon	discussion	Remarks. It was not possible to ascribe the unique female specimen to any species. The adult was found in the edges of a small rain pound within a secondary open forest.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA2EBFB57FE70.taxon	distribution	Distribution in Angola (Provinces): 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA2EBFB57FE70.taxon	materials_examined	Material examined. Cambamba (Catoio-Quela) (9 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA2EBFB57FE70.taxon	discussion	Remarks. A species with a known distribution throughout D. R. of the Congo, Zambia and Mozambique (Alves 1963, Anichtchenko 2016, Insectoid. info 2017). The singleton adult was collected beneath litter in a countryside road together with adults of tiger beetles [Dromica (Foveodromica) sp., L. wellmani) and of ground beetles (Lasiocera schuelei sp. n., G. ellipticus]. It is a new genus / species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA16FFEA9FAB8.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Cunene.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA16FFEA9FAB8.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. ‒ 2. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Cachimbamba (12 ° 28´11 ´´ S, 15 ° 40´56 ´´ E, 1560 m alt., 232) (HUAMBO), 3. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA16FFEA9FAB8.taxon	discussion	Remarks. A common species widespread throughout Africa south of Sahara (Ferreira 1965, Anichtchenko 2016), but also present in South America (Argentina, Brazil, Uruguay) (Insectoid. info 2017) where was introduced probably by means of the nautical slave trade held between Africa (e. g. Luanda) and Brazil (e. g. Salvador) during the XVI ‒ XVII centuries. Adults were found on ground during night near public lights (first locality) together with some other ground beetles [e. g. S. guineense, C. intermedius, C. rikatlensis, C. cupreocinctus, C .. tenuicollis, Progonochaetus planicollis (Putzeys, 1880), Platymetopus pseudocrenulatus Facchini, 2016, Agonidium natalense (Boheman, 1848), Agonidium alacre (Boheman, 1848), E. peringuey, E. intermedius] or on soil beneath litter (second locality).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA455FEB8F852.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Benguela, Huambo, Huíla, Cunene, Cuando Cubango.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA455FEB8F852.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 231) (HUAMBO), 1. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 2 ♂, 4. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3147F7314BFA455FEB8F852.taxon	discussion	Remarks. A species widespread throughout central and southern Africa (Democratic Republic of the Congo, Angola, Namibia, Zambia, Zimbabwe, Burundi, Rwanda, Malawi, Mozambique and Tanzania) (Basilewsky 1950 a, Anichtchenko 2016, Insectoid. info 2017). In Angola the species presents a wide distribution throughout the country. Adults were found on soil beneath litter (first locality, see S. lucidus strigiceps remarks) or on ground and walls during night near public lights (second locality) together with several other ground beetles [e. g. S. guineense, C. rikatlensis, C. cupreocinctus, C. tenuicollis, P. planicollis (Putzeys, 1880), A. natalense, A. alacre, E. peringuey, E. intermedius].	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA2EBFC64FDE0.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Malanje; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA2EBFC64FDE0.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. ‒ 2. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 1 ♀, 4. XI. 2015, 2 ♂, 6 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA2EBFC64FDE0.taxon	discussion	Remarks. A species originally described from Angola (Putzeys 1880), but with a wide distribution throughout Africa south of Sahara (Senegal, Guinea-Bissau, Guinea, Mali, Burkina Faso, Chad, Ivory Coast, Cameroon, Ghana, Central African Republic, Nigeria, Equatorial Guinea, Gabon, Congo, D. R. of the Congo, Angola, Namibia, Zambia, Zimbabwe, Rwanda, Botswana, Saudi Arabia, Yemen, Djibouti, Eritrea, Ethiopia, Somalia, Kenia, Tanzania, Mozambique) (Serrano 1999 b, Anichtchenko 2016, Insectoid. info 2017). Adults were collected in activity during night on ground and walls near public lights together with other ground beetles (see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA1DEFDD1FB7F.taxon	distribution	Distribution in Angola (Provinces): 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA1DEFDD1FB7F.taxon	materials_examined	Material examined. Kibundi (15 km N Nambua) (8 ° 56´15 ´´ S, 16 ° 03´52 ´´ E, 1164 m alt., 95) (MALANJE), 19. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA1DEFDD1FB7F.taxon	discussion	Remarks. A species known from Central African Republic, Cameroon, Burundi, Zambia and Malawi (Facchini 2012 b). The single male fits well the original description, including the aedeagus features. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA767FB88F9D0.taxon	distribution	Distribution in Angola (Provinces): 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA767FB88F9D0.taxon	materials_examined	Material examined. Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 5. XII. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA767FB88F9D0.taxon	discussion	Remarks. A species widespread throughout western Africa (Senegal, Gambia, Guinea-Bissau, Guinea, Sierra Leone, Gabon, Mali, D. R. of the Congo) (Serrano 1999 b, Anichtchenko 2016, Insectoid. info 2017). The single adult fits well the description and aedeagus of this species and was collected beneath litter together with adults of some tiger and ground beetles [e. g. Dromica (Foveodromica) sp., A. distinctus, A. optimus, D. tarsalis, C. magnicollis discrepans, G. procera capelai ssp. n.]. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA4B0FA7FF80D.taxon	distribution	Distribution in Angola (Provinces): Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA4B0FA7FF80D.taxon	materials_examined	Material examined. Satchijamba (10 km E) (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 303) (BIÉ), 7. IV. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3157F7214BFA4B0FA7FF80D.taxon	discussion	Remarks. A species close to S. barbarae Facchini, 2012 as well as to S. basirufus (Basilewsky, 1948). However, a different shape of the median lobe of the aedeagus did not allow its inclusion on one of these species.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA339FA9EFD85.taxon	distribution	Distribution in Angola (Provinces): Benguela.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA339FA9EFD85.taxon	materials_examined	Material examined. Baixo Pundo (12 ° 20´22.32 ´´ S, 13 ° 50´20.97 ´´ E, 275 m alt., 227 / 8) (BENGUELA), 25 ‒ 27. X. 2013, 1 ♀, PF, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA339FA9EFD85.taxon	discussion	Remarks. The single female adult specimen was collected by pitfall trapping within an open bush habitat with sparse grassy vegetation and sandy soil, together with the ground beetle Cypholoba brevivittis Chaudoir, 1866.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA4B2FABDF86A.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA4B2FABDF86A.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA4B2FABDF86A.taxon	discussion	Remarks. A species widespread throughout Africa south of Sahara (Serrano 1999 c, Anichtchenko 2016, Insectoid. info 2017). The single adult was collected by means of light trapping within a secondary open forest together with some other ground beetles (see S. angolense remarks). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA117FB69FACD.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Huíla, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA117FB69FACD.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 1. IV. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC; Chitembo (6 km SE) (13 ° 33´23 ´´ S, 16 ° 47´22 ´´ E, 1586 m alt., 302) (BIÉ), 9. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. ‒ 2. XII. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Mutubanzamba (Cangandala) (9 ° 47´45 ´´ S, 16 ° 31´40 ´´ E, 1136 m alt., 132) (MALANJE), 21. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3167F7114BFA117FB69FACD.taxon	discussion	Remarks. A species described recently by Facchini (2016 b) with a wide distribution throughout central and southern Africa (Angola, Zambia, Zimbabwe, Rwanda, Burundi, Malawi, Tanzania, Mozambique and South Africa). Adults were collected by means of light trapping within an open secondary forest (Chitembo), on walls near public lights (Alto Hama) and beneath litter (Mutubanzamba) together with other ground beetles (Chitembo: S. guineense; Alto Hama: C. intermedius, C. rikatlensis, C. cupreocinctus, C. tenuicollis, P. planicollis) and tiger beetles [Mutubanzamba: T. nubifera, E. villosus, L. saraliensis, L. uncivittata, L. wellmani].	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA612FBBCF9F2.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Cuando Cubango, Moxico; 2) Huambo, Bié, Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA612FBBCF9F2.taxon	materials_examined	Material examined. Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 9. IV. 2014, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Kakande (12 ° 42´50 ´´ S, 16 ° 45´33 ´´ E, 1716 m alt., 258) (BIÉ), 7. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Mutubanzamba (Cangandala) (09 ° 47´45 ´´ S, 16 ° 31´40 ´´ E, 1136 m alt., 132) (MALANJE), 21. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA612FBBCF9F2.taxon	discussion	Remarks. Another species widespread throughout almost all Africa south of Sahara (Senegal, Guinea-Bissau, Mali, Ivory Coast, Chad, D. R. of the Congo, Angola, Zimbabwe, Rwanda, Eritrea, Sudan, Kenya, Tanzania, Mozambique, South Africa) (Serrano 1999 c, Anichtchenko 2016, Insectoid. info 2017). Adults were found in countryside roads within savanah (Mumbué) or open secondary forest (Kakande and Mutubanzamba) with withish sandy soil beneath the marginal litter, and on a wall near public lights (Caála) with other ground beetles (see P. holosericus remarks). It is a new record for the Huambo, Bié and Malanje Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA2EBFAD9FDA8.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Cuando-Cubango, Lunda Norte, Lunda Sul; 2) Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA2EBFAD9FDA8.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.74 ´´ S, 16 ° 45´48.17 ´´ E, 1649 m alt., 281) (BIÉ), 2. IV. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 5. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA2EBFAD9FDA8.taxon	discussion	Remarks. A very common species widespread throughout all Africa south of Sahara, including Madagascar, Comoros and Seychelles islands (Serrano 1999 c, Anichtchenko 2016, Insectoid. info 2017). Adults were collected by means of light trapping within an open secondary forest (Cachingues-Chipica) and on a wall near public lights (Caála) together with other ground beetles [Cachingues-Chipica: S. ludricus, A. natalensis; Caála: C. deyrollei, C. pulchelus, Parophonus integer (Péringuey, 1896)]. It is a new record for the Huambo and Bié Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA019FEF1FBB1.taxon	distribution	Distribution in Angola (Provinces): 1) Cunene; 2) Benguela.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA019FEF1FBB1.taxon	materials_examined	Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227 / 8) (BENGUELA), 3. IV. 1998, 1 ♀, PF, A. Neves leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7014BFA019FEF1FBB1.taxon	discussion	Remarks. A very common species widespread throughout all Africa south of Sahara (Serrano 1999 c, Anichtchenko 2016, Insectoid. info 2017). It was firstly recorded to Angola under two names (Putzeys 1880): Hypolithus tomentosus Dejean, 1829 and Hypolithus prolixus Putzeys, 1880. Later Basilewsky (1960) confirmed the synonymy of the latter species with the former one. The single female adult was collected by pitfall trapping within an open bush habitat with sparse grassy vegetation on sandy-clayey yellow-reddish soil in the outskirts of Lobito together with some other ground beetles (see B. apicalis remarks). It is a new record for the Benguela Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7714BFA505FACEFEDB.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7714BFA505FACEFEDB.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3177F7714BFA505FACEFEDB.taxon	discussion	Remarks. An Angolan ground beetle endemic species (Anichtchenko 2016, Insectoid. info 2017). The species was known until now by two type specimens from Luimbale, Mt. Moco at 1800 - 1900 m (Clarke 1981). The singleton male fits very well the original description, including the aedeagus. The adult was collected on a wall near public lights with other ground beetles (C. cupreocinctus, C. pulchelus, P. incrassatus, P. planicollis).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA6BEFC49F8BF.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Huíla, Cuando Cubango, Moxico; 2) Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA6BEFC49F8BF.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.74 ´´ S, 16 ° 45´48.17 ´´ E, 1649 m alt., 281) (BIÉ), 2. IV. 2014, 1 ♂, 1 ♀, LT, A. Serrano & R. Capela leg., ASC; Satchijamba-Somakwanza (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 303) (BIÉ), 7. IV. 2014, 1 ♂, 1 ♀, LT, A. Serrano & R. Capela leg., ASC; Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 1 ♀, 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA6BEFC49F8BF.taxon	discussion	Remarks. A common species known almost from all Africa south of Sahara (Senegal, Guinea, Togo, Ivory Coast, Cameroon, Central African Republic, Chad, Gabon, D. R. of the Congo, Burundi, Rwanda, Angola, Zambia, Zimbabwe, Eritrea, Somalia, Ethiopia, Kenya, Tanzania, Mozambique, Uganda, Malawi, South Africa) (Basilewsky 1950 a, Ferreira 1965, Insectoid. info 2017). In the first three localities the adults were collected by light trapping within open secondary forests together with some other ground beetles (e. g. S. ludricus, S. guineense, A. natalensis). Adults in Caála were collected on a wall near public lights with other ground beetles also (see P. elegans remarks). It is a new record for the Huambo and Bié Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA16DFEF1FBC5.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Kwanza Sul, Malanje, Moxico; 2) Huambo. Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7714BFA16DFEF1FBC5.taxon	discussion	Remarks. A species originally described from Angola (Harold 1879), but with a wide distribution throughout Africa (Guinea, Central African Republic, Congo, D. R. of the Congo, Angola, Zimbabwe, Burundi, Malawi, Mozambique, Sudan, Tanzania) (Ferreira 1965, Anichtchenko 2016, Insectoid. info 2017). The adult was collected on a wall near public lights with other ground beetles (see P. elegans remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7614BFA535FC8FFE37.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Huíla, Lunda Norte, Malanje; 2) Huambo. Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3107F7614BFA535FC8FFE37.taxon	discussion	Remarks. A species present throughout Arabian Peninsula and eastern and central-southern Africa (Yemen, Eritrea, Somalia, Ethiopia, Kenya, Tanzania, Mozambique, Malawi, Rwanda, Burundi, D. R. of the Congo, Angola, Namibia, Zambia, Zimbabwe, Niger) (Basilewsky 1950 a, Ferreira 1965, Insectoid. info 2017). It is with some reserve that we ascribe to this species the sampled female singleton. It should be necessary a male to confirm this identification because there are some species belonging to this sg. with similar habitus (see Facchini 2012 c). The adult was found on ground during night near public lights together with several other ground beetles (see P. incrassatus remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA3AFFD7AFC88.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA3AFFD7AFC88.taxon	materials_examined	Material examined. Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA3AFFD7AFC88.taxon	discussion	Remarks. A species described from Zimbabwe and Botswana (Facchini 2012). The single female adult fits well the original description, but without a male specimen it is with some reserve that we ascribe it to this species. The singleton adult was collected by pitfall trapping within an open secondary forest together with other ground beetles (S. lucidus strigiceps, C. bipustulatus, A. pseudoangolensis and Atractonotus puncticollis Schle & Heinz, 2013). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA74AFC8FF92A.taxon	distribution	Distribution in Angola (Provinces): Benguela, Bié, Huíla; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA74AFC8FF92A.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X. ‒ 2. XI. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 4. XI. 2015, 1 ♂, 5. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA74AFC8FF92A.taxon	discussion	Remarks. A species described recently with a scattered distribution in central, southern and eastern Africa (Nigeria, D. R. of the Congo, Angola, Zambia, Zimbabwe, Uganda) (Facchini 2017 b). Adults were collected on walls during night near public lights together with several other ground beetles (e. g. see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA1F9FC3CFB5A.taxon	distribution	Distribution in Angola (Provinces): Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA1F9FC3CFB5A.taxon	materials_examined	Material examined. Carima (Cacuso-Pungandongo) (9 ° 35´00 ´´ S, 15 ° 42´01 ´´ E, 1117 m alt., 130) (MALANJE), 23. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7614BFA1F9FC3CFB5A.taxon	discussion	Remarks. The single female adult is quite similar to S. punctatellus, but with a shape of pronotum slightly different [anterior angles not protruding, anterior margin almost straight between angles and basal margin not sinuate at each side of the middle region). The adult was found in a set aside country road beneath litter within an open secondary forest together with adults of tiger and ground beetles [e. g. Dromica (Foveodromica) auropunctata Quedenfeldt, 1883, P. angusticollis, T. nubifera, E. villosus, G. ellipticus].	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7514BFA49BFAFEFEFF.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Huíla, Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7514BFA49BFAFEFEFF.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 303) (BIÉ), 7. IV. 2014, 2 ♀, LT, A. Serrano & R. Capela leg., ASC; Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♂, 2 ♀, LT, A. Serrano & R. Capela leg., ASC; Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3117F7514BFA49BFAFEFEFF.taxon	discussion	Remarks. A species with a scattered distribution throughout eastern, central and southern Africa (Kenya, Tanzania, Rwanda, D. R. of the Congo, Angola, Zimbabwe, South Africa) (Basilewsky 1950 a, Ferreira 1965, Insectoid. info 2017). Adults were sampled by light trapping in Bié province within open secondary forests together with other ground beetles (see S. angolense remarks) and on walls during night (Caála) near public lights together with several other ground beetles (see P. incrassatus remarks). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA3E7FD06FD50.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Cuando Cubango; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA3E7FD06FD50.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA3E7FD06FD50.taxon	discussion	Remarks. A species known on western, eastern and central Africa (Guinea, Ivory Coast, Central African Republic, Equatorial Guinea, Gabon, D. R. of the Congo, Angola, Burundi, Rwanda, Zimbabwe, Ethiopia, Kenya, Tanzania, Mozambique) (Basilewsky 1950 a, Ferreira 1965, Insectoid. info 2017). Adults were collected on walls during night near public lights together with several other ground beetles (e. g. see N. picina and P. incrassatus remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA131FCFFFB7D.taxon	distribution	Distribution in Angola (Provinces): 2) Cabinda.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA131FCFFFB7D.taxon	materials_examined	Material examined. Chipita (5 ° 43´41 ´´ S, 12 ° 13´07 ´´ E, 7 m alt., 7) (CABINDA), 11. II. 2011, 1 ♂, DO, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7514BFA131FCFFFB7D.taxon	discussion	Remarks. A species known on western, eastern and central Africa, including atlantic and indic islands (Mauritania, Senegal, Gambia, Guinea-Bissau, Guinea, Ivory Coast, Benin, Sierra Leone, Ghana, Togo, Burkina Faso, Mali, Chad, Cameroon, Central African Republic, Equatorial Guinea, Gabon, Niger, Nigeria, Democratic Republic of the Congo, Burundi, Rwanda, Tanzania, Mozambique, São Tomé and Príncipe, Comoros, Seychelles and Madagascar) (Basilewsky 1950 a, Insectoid. info 2017). The single adult was collected under a stone. Surprisingly the species was never recorded for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7414BFA79FFBDCFEA2.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7414BFA79FFBDCFEA2.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC; Chitembo (13 ° 22´40.78 ´´ S, 16 ° 41´58.07 ´´ E, 1666 m alt., 281) (BIÉ), 1. XI. 2014, 1 ♀, LT, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3127F7414BFA79FFBDCFEA2.taxon	discussion	Remarks. A species known only to Central African R. and Uganda (Basilewsky 1968, Facchini 2011 a). Interestingly our male specimen fits very well the median lobe shape of aedeagus in lateral view of A. latipalpis Basilewsky, 1968 (Basilewsky 1968, Fig. 3, Facchini 2003, Fig. 18 a) with the apex slightly bent up. However, in dorsal view the median lobe apex is similar to the drawing given in Facchini (2011 a, Fig. 12 b) for A. dilatipalpis Facchini, 2011. Despite this asymmetry we ascribe our identification to the Basilewsky`s species by some morphological features (e. g. last labial palpomere only dilated, elytra striae crenulated, third interval with a discal setigerous puncture in the last third). However, our specimens do not present any small pubescence in apical margin, as was point out by Basilewsky (1968) for its species, and the labial palpi are clearly bicolored. Unfortunately, Facchini (2011 a) do not give in the affinities between both species any comment on the elytra apical pubescence. Probably in a near future the status of these two species will be reviewed. Adult specimens were collected by light trapping within open secondary forests together with some other ground beetles (e. g. see S. angolense and S. guineense remarks). It is a new genus / species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA0A6FEFEFB7F.taxon	distribution	Distribution in Angola (Provinces): Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA0A6FEFEFB7F.taxon	materials_examined	Material examined. Lumbanje (Morro Lubiri) (12 ° 07´55 ´´ S, 15 ° 30´59 ´´ E, 1424 m alt., 232) (HUAMBO), 30. X. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Chitembo (6 km SE) (13 ° 33´23 ´´ S, 16 ° 47´22 ´´ E, 1586 m alt., 302) (BIÉ), 9. XI. 2015, 1 ♂, DO, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA0A6FEFEFB7F.taxon	discussion	Remarks. Adult specimens were collected always within litter in open secondary forests together with a common tiger beetle (P. angusticollis). Further adult specimens of tiger and ground beetles were collected in the same habitats namely L. uncivittata, L. saraliensis and L. wellmani in Lumbanje, Euryarthron reticostatum (Wellman & W. Horn, 1908), Dromica (Foveodromica) sp., T. nubifera, L. saraliensis, L. wellmani, S. vilhenai, C. ovalipennis, E. intermedius, O. patroboides, N. vethi, C. graphipteroides monardi and A. (Odontanthia) kleinfeldi in Catata-Nova Monção and T. nzingae, Ophryodera rufomarginata poggei (Harold, 1878) and L. uncivittata in Chitembo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA767FCBDF9F5.taxon	distribution	Distribution in Angola (Provinces): Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA767FCBDF9F5.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 302) (BIÉ), 2. XI. 2015, 3 ♂, DO, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA767FCBDF9F5.taxon	discussion	Remarks. Adult specimens were collected within litter in open secondary forests together with some tiger and ground beetles (Prothyma erythropyga erythropyga (Putzeys, 1880), Dromica (Foveodromica) soror (W. Horn, 1935), P. angusticollis, T. nubifera, O. rufomarginata distanti (Heath, 1905), L. uncivittata, L. saraliensis, L. wellmani, C. agualusai, S. lucidus strigiceps and A. kleinfeldi.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA4E2FDEEF853.taxon	distribution	Distribution in Angola (Provinces): Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA4E2FDEEF853.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 302) (BIÉ), 2. XI. 2015, 2 ♂, DO, A. Serrano leg., ASC; Chingueia-Mandongue (13 ° 35´46 ´´ S, 16 ° 58´59 ´´ E, 1648 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♂, DO, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3137F7414BFA4E2FDEEF853.taxon	discussion	Remarks. Like the precedent Orthogonius, these adult specimens were collected within litter in open secondary forests together with some tiger and ground beetles (for the first locality see Orthogonius sp. 2 remarks). Adult specimens in Chingueia-Mandongue were collected together with some different tiger and ground beetles (see S. lucidus strigiceps remarks).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA7B7FA6EF944.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA7B7FA6EF944.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♀, 4. XI. 2015, 1 ♂, 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA7B7FA6EF944.taxon	discussion	Remarks. A species with a distribution in southern and eastern Africa (Botswana, Namibia, South Africa, Zimbabwe and Mozambique), but recorded also to Yemen (Insectoid. info 2017). Adult specimens fit well the descrition of this species, including the aedeagus conformation (Basilewsky 1950 b). Adults were found on ground during night near public lights (e. g. see comments on C. rikatlensis remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA117FD3EFBC8.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA117FD3EFBC8.taxon	materials_examined	Material examined. Kibundi (15 km N Nambua) (8 ° 56´15 ´´ S, 16 ° 03´52 ´´ E, 1164 m alt., 95) (MALANJE), 19. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA117FD3EFBC8.taxon	discussion	Remarks. A species known from Angola, D. R. of the Congo and Zimbabwe (Basilewsky 1948, Ferreira 1965, Insectoid. info 2017). The single adult was collected on the ground in a countryside road within an open secondary forest. It is a new record for the Malanje Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA293FDB1FE70.taxon	distribution	Distribution in Angola (Provinces): Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA293FDB1FE70.taxon	materials_examined	Material examined. Chitembo (13 ° 30´55 ´´ S, 16 ° 45´20 ´´ E, 1628 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♀, DO, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7B14BFA293FDB1FE70.taxon	discussion	Remarks. Adult specimens were found in activity within the village on soil in the road edges during night, probably attracted to the public lights.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7A14BFA53DFA72FEA2.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7A14BFA53DFA72FEA2.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31C7F7A14BFA53DFA72FEA2.taxon	discussion	Remarks. A species known already to Angola without known localities (Putzeys 1880, under the name Platynus urens) and with a distribution in southern and part of eastern Africa (Angola, Botswana, South Africa, Zimbabwe and Mozambique) (Basilewsky 1950 b, Insectoid. info 2017). The singleton adult was found on ground during night near public lights (e. g. see C. rikatlensis remarks). It seems a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA051FE4CFC55.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA051FE4CFC55.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 4. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00 ° 37 ´´ S, 17 ° 24 ° 00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA051FE4CFC55.taxon	discussion	Remarks. A species known from South Africa R. and Zimbabwe (Basilewsky 1950 b, Insectoid. info 2017). Adult specimens fit well the original description (Péringuey 1896). Adults of the first locality were found on ground during night near public lights (e. g. see C. rikatlensis remarks) and in the second locality beneath litter within an open secondary forest (e. g. see comments on S. vilhenai remarks), respectively. The Catota adult specimen was collected by pitfall trapping together with other ground beetles (see S. wrasei remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA64EFE4CFA02.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo, Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA64EFE4CFA02.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 3. XI. 2015, 1 ♂, 6. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Calulo (9 ° 59´55 ´´ S, 14 ° 53´35 ´´ E, 999 m alt., 128) (BIÉ), 27. XI ‒ 5. XII. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7A14BFA64EFE4CFA02.taxon	discussion	Remarks. A species with a scattered known distribution (Cameroon, Ivory Coast, D. R. of the Congo, South Africa and Mozambique) (Basilewsky 1950 b, Insectoid. info 2017). Adult specimens fit very well the original description (as albicornis Péringuey 1896), including the aedeagus (see Basilewsky 1950 b, Fig. 6). Adults were found on ground during night near public lights (e. g. for the first locality see C. rikatlensis remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7914BFA4B0FDFBFF6A.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7914BFA4B0FDFBFF6A.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 231) (HUAMBO), 31. X. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Londuimbali-Tinguita (12 ° 18´10 ´´ S, 15 ° 21´15 ´´ E, 1633 m alt., 231) (HUAMBO), 2. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31D7F7914BFA4B0FDFBFF6A.taxon	discussion	Remarks. A species known only from Zimbabwe and Mozambique (Basilewsky 1950 b, Insectoid. info 2017). Adult specimens fit very well its description (Péringuey 1904, Basilewsky 1950 b) and were found within litter in open secondary forests together with some tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new genus / species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA051FE11FB59.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown, 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA051FE11FB59.taxon	materials_examined	Material examined. Cateculo (Calulo) (9 ° 57´19.75 ´´ S, 14 ° 47´31.37 ´´ E, 872 m alt., 128) (KWANZA SUL), 5. XI. 2014, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Mussende (Calulo) (9 ° 57´00 ´´ S, 14 ° 47 ´´ 36 ´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 30. XI. 2015, 4 ♂, 5 ♀, 2. XII. 2015, 1 ♂, 5 ♀, 5. XII. 2015, 1 ♂, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo (Faz. Klein) (10 ° 02´14 ´´ S, 14 ° 54´38 ´´ E, 1059 m alt., 147) (KWANZA SUL), 1. XII. 2015, 1 ♂, 1 ♀, 3. XII. 2015, 2 ♂, 2 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA051FE11FB59.taxon	discussion	Remarks. A species widespread throughout eastern, central and southern Africa (Ethiopia, Mozambique, Zimbabwe, Burundi, Rwanda, D. R. of the Congo, Angola, Botswana, South Africa R.) (Putzeys 1880, Insectoid. info 2017). The species was recorded for Angola by Putzeys (1880) without information on localities. Basilewsky (1960 a) studying the 24 types described by this author did not pay attention to the other 117 species of carabids recorded for Angola in that work, referring that most of them were misinterpreted. Unfortunately we can not confirm the identification of the Putzeys study because all the material deposited in MLZA (Museu Bocage) was destroyed during a catastrophic fire in 1978. However, as the species is so well characterized, it seems to us that Putzeys identification was almost surely correct. Adult specimens were found within litter in secondary rainforests together with other tiger and ground beetles (see comments on A. distinctus remarks). It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA74AFC4CF82F.taxon	distribution	Distribution in Angola (Provinces): 2) Kwanza Sul, Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA74AFC4CF82F.taxon	materials_examined	Material examined. Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 2 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Cruzeiro (Pedra do Alemão) (12 ° 46´41.14 ´´ S, 15 ° 54´09.49 ´´ E, 1743 m alt., 232) (HUAMBO), 7. XI. 2015, 2 ♀, DO, A Serrano & R. Capela leg., ASC; Calulo-Cabuta (9 ° 53´59 ´´ S, 14 ° 54´26 ´´ E, 831 m alt., 128) (KWANZA SUL), 29. XI. 2015, 2 ♂, 4 ♀, 29. XI ‒ 6. XII. 2015, 26 ♂, 20 ♀, PF, A. Serrano & R. Capela leg., ASC; Mussende (Calulo) (9 ° 57´00 ´´ S, 14 ° 47´36 ´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI ‒ 5. XII. 2015, 14 ♂, 9 ♀, PF, A. Serrano & R. Capela leg., ASC; Alto Ventura (Faz. Monte Café) (Calulo) (9 ° 59´15 ´´ S, 14 ° 50´12 ´´ E, 1086 m alt., 128) (KWANZA SUL), 5. XII. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31E7F7914BFA74AFC4CF82F.taxon	discussion	Remarks. A species widespread throughout western, central and eastern Africa (Senegal, Guinea, Ivory Coast, Nigeria, D. R. of the Congo, Angola, Zambia, Burundi, Rwanda, Uganda) (Putzeys 1880, Insectoid. info 2017). The species was described by Putzeys (1880) from Angola under the name Platynus calefactus without information on localities. The species is very abundant in Calulo region and the adult specimens were found within litter or collected by pitfall trapping in secondary rainforests together with other tiger and ground beetles (see A. distinctus remarks). It is a new record for the Kwanza Sul and Huambo Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7814BFA2EBFACFFDA8.taxon	distribution	Distribution in Angola (Provinces): 2) Kwanza Sul, Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7814BFA2EBFACFFDA8.taxon	materials_examined	Material examined. Cambamba (Catoio-Quela) (09 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo (Faz. Klein) (10 ° 02´14 ´´ S, 14 ° 54´38 ´´ E, 1059 m alt., 147) (KWANZA SUL), 1. XII. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7814BFA2EBFACFFDA8.taxon	discussion	Remarks. A species known from central, southern and part of eastern Africa (D. R. of the Congo, Souh Africa R., Burundi, Zimbabwe) (Basilewsky 1950 b, Insectoid. info 2017). Adults fit very well its description, including the aedeagus features (Péringuey 1904, Basilewsky 1950 b). Adult specimens were found within litter in an open secondary forest and in a secondary rainforest together with some other tiger and ground beetles (see S. vilhenai and O. gilvipes remarks for first and second localities, respectively). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7C14BFA1FBFB06FE12.taxon	materials_examined	Type series. Holotype, ♂; Angola (Kwanza Sul), Alto Ventura-Col (Calulo) (9 ° 59´42 ´´ S, 14 ° 50´32 ´´ E, 1116 m alt., 128), 4. XII. 2015, DO, A. Serrano & R. Capela leg., ASC. Allotype, 1 ♀, same locality and date as holotype, ASC. Derivatio nominis. This species is named after Libolo (noun in apposition), a municipality of the Kwanza Sul Province where the specimens were found and home of one of the most remarkable Angolan Sports Club: the Sports Club Recreativo do Libolo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7C14BFA1FBFB06FE12.taxon	diagnosis	Diagnosis. Body elongate, sub-parallel, slightly convex and reddish-brown (Fig. 1 a); integument with microreticulation, shiny; surface with sparse micropunctures visible at high magnification. Elytral color as Perigona pallida Castelnau, 1835. Internal sac of median lobe of aedeagus (Fig. 2) with a conspicuous twisted sclerite closer to the apex and a posterior bone shaped cut half sclerite (lateral view).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7C14BFA1FBFB06FE12.taxon	description	Description. Length of Holotype: 3.36 mm. Length of allotype: 3.16 mm. Head. Slightly wider than long [length: 0.59 mm (holotype), 0.64 mm (allotype), width: 0.72 mm (holotype), 0.75 mm (allotype)], darker (black-brownish) than pronotum and elytra, clypeus, labrum, mouth parts and antennae piceous; surface with sparce fine punctuation, microreticulation distinct, isodiametric, rather glossy; eyes large, moderately protruding, labrum moderately notched; clypeal-frontal furrows distinct, oblique, extended to level of anterior supraorbital seta; frons and occiput slightly convex; antenna short just attaining base of pronotum, scape two times longer than wide, 3 rd to 9 th antennomeres as long as wide, slightly globulose, last antennomere 1.6 ‒ 1.8 times longer than wide. Cephalic chaetotaxy (large setae): Labrum with three pairs of setae gradually shorter from exterior to inner sides, one pair on sides of clypeus and two pairs of supraocular setae present over each eye. Thorax. Pronotum (Fig. 1 a) 1.27 ‒ 1.29 times wider than long [length: 0.66 mm (holotype), 0.74 mm (allotype), width: 0.85 mm (holotype), 0.94 mm (allotype)], reddish-brown dark, wider than head, barely subcordiform, widest at first third; surface sparcely micropunctate, with microreticulation distinct, consisting of isodiametric meshes in diskal area and transverse meshes and lines laterally, shiny; apical margin not margined, lateral margins with complete and distinct margination continuing in the two thirds of the basal margin after the posterior angle; apical margin arcuate, apical angles roundly acute, protruded; lateral margin slightly curved at first two thirds, followed by a slight sinuosity and ending straight obliquely before the roundly obtuse basal angles, posterior margin distinctly emarginated; median line distinct, not reaching anterior or posterior margins, anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous; anterior marginal setae situated close to the widest width, posterior marginal seta situated at basal angle, none short setae at disk or sides. Elytra (Fig. 1 a) 1.61 ‒ 1.62 times longer than wide [length: 1.90 mm (Holotype), 1.97 mm (allotype), width: 1.17 mm (holotype), 1.22 mm (allotype)], reddish-brown lighter than pronotum, sutural stripe slightly darker; surface glossy, with sparse very fine punctuation and superficial microreticulation of fine transverse meshes; narrow and elongate, sub-parallel, dorsally moderately convex, very slightly depressed on disk, widest at middle; humeral angles distinct but broadly rounded; mediam marginal punctures arranged in a straight line; marginal channel towards apex widened, depressed, first half scarsely pilose, second half densely pilose; only inner three striae recognizable at two thirds by inconspicuous sulci not marked by punctures, anterior setiferous puncture situated at 3 rd striae and at basal quarter, median setiferous puncture situated in 3 rd interval close to 2 nd striae slightly behind middle, apical setiferous puncture (long setae) situated at 3 rd striae just above sub-apical border. Abdominal segments brown-yellowish, covered with dense dressed short hairs; last segment with 3 pairs (male) or 5 pairs (female) of setae near the posterior margin; terminal abdominal sternite in both sexes with fringe of short hairs. Legs brown-testaceous. Aedeagus (Figs 2 a, b, c) with median lobe short and stout, ventral margin slightly curved, apex downwards (lateral view); internal sac with a conspicuous twisted sclerite and a bone shaped cut in half sclerite at right and left sides, respectively; genital ring (Fig. 2 d) rather asymmetric, with wide, obliquely rounded apex, rather parallel, base evenly rounded. Female genitalia (Fig. 3) with the ordinary pattern shape of the genus. Stylomere 1 with two ventral large setae on subapical margin. Stylomere 2 with two medium sized ensiform ventro-lateral setae, one quite large ensiform dorso-median seta located about in middle of stylomere, and a single nematiform ventro-seta originating from a groove in apical third of stylomere. Some sensillae (probably chemoreceptors) are dispersed over ventral lateroapical margins of stylomere 1, and other sensillae over basal, medial and sub-apical areas of ventral, lateral and dorsal sides of stylomere 2.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31F7F7C14BFA1FBFB06FE12.taxon	discussion	Remarks. The new species belongs to the subgenus Perigona s. str. based on the arrangement of the three lateral marginal elytral pores in a straight line and by the presence of distinct clypeo-orbital sulci. Until now there are six species known for continental Africa belonging to this subgenus (Basilewsky 1989, Baehr 2004 and Serrano 2008). Perigona liboloensis sp. n. is easily segregate from P. parallela Chaudoir, 1878, P. nigrociliata Basiewsky, 1953, P. mediornata Basilewsky, 1989, P. wachteli Baehr, 2004 and P. principensis Serrano, 2008, among other characters, by the quite dissimilar aedeagi of all these species. Certainly the new species is next related to P. pallida Laporte de Castelnau, 1835 with which it shares the general color, the elongate, almost parallel body shape and male median lobe general shape, though differs in its general body length (3.16 ‒ 3.36 mm vs. 2.30 ‒ 2.70 mm), pronotum conformation (slightly sinuouse before the posterior angles vs. not sinuouse before the posterior angles), elytral index (1.61 ‒ 1.62 vs. 1.47 ‒ 1.55), and in the shape of the sclerotized plates in the internal sac of the median lobe of aedeagus (cf. Figs 2 a, b vs. Fig. 5 b in Basilewsky 1989). Moreover, P. pallida presents the pronotal widest situated in the middle while in the new species is in the first third. In the former species the elytral striae are conspicuous with punctuation well marked while in the latter the elytral striae are almost indistinct and without marked punctuation. By the Perigona pallida previous records (e. g. Basilewsky 1989) and the new species known locality, the two taxa seem to be sister species with vicariant (allopatric) distribution, a fact referred already for tiger beetle and ant nest beetle species of Angola also (Serrano et al. 2015, Serrano & Capela 2015 a). Ecological notes. A species known until now only from Angola. Adults were found under the bark of a fallen tree within an abandoned coffee plantation (Fig. 9 a) with Catascopus beauvoisi Laporte de Castelnau, 1835 and Coptodera (Coptoderina) congolensis Burgeon, 1937. Further adults of Dermaptera [Apachyus depressus (Palisot de Beauvois, 1805 and Echinosoma afrum (Palisot de Beauvois, 1805)] and Coleoptera Endomychidae, Trogossitidae, Tenebrionidae (several species) were found together (syntopic) with the new species.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.taxon	materials_examined	Type series. Holotype, ♂; Angola (KWANZA SUL), Munenga 35 km W Calulo (unknown, 147), 3. XII. 2012, DO, P. Schüle leg., PSC. Allotype 1 ♀: same data as Holotype, P. Schüle leg., ASC. Paratypes: Angola (HUÍLA) 3.5 km SW Negola (14 ° 08´53 ´´ S, 14 ° 28´16 ´´ E, 1615 m, 317), 8. XII. 2012, 2 ♀, DO, P. Schüle leg., PSC, ZSM; Cambamba (Catoio-Quela) (09 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 1 ♀, DO, A. Serrano leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.taxon	etymology	Derivatio nominis. This species is cordially dedicated to Peter Schüle, from Herrenberg, a specialist on tiger and ground beetles of Africa which has deeply collaborated with us and kindly provided most of the specimens of this new species.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.taxon	diagnosis	Diagnosis. Body elongate, parallel, slightly convex and greenish-bronze (Fig. 1 b); integument densely and deeply punctate, shagrened; surface among punctures with micropunctures visible at high magnification. Pronotum subrectangular or subtrapezoidal shaped (Fig. 4). Four reduced elytral spots. Internal sac of median lobe of aedeagus (Fig. 5) with tangled membranes, the dorsal twisted one closer to the apex (lateral view), this slightly bent to the left (dorsal view) (Fig. 5).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.taxon	description	Description. Length of Holotype: 4.82 mm. Length of paratypes (all females): 4.95 ‒ 5.74 mm. Head. Triangular (Fig. 1 b), 1.2 ‒ 1.3 times wider than long [length: 0.77 mm (holotype), 0.83 ‒ 0.88 mm (female allotype and paratypes), width: 0.99 mm (holotype), 1.02 ‒ 1.14 mm (female allotype and paratypes)], greenish bronze, shagreened, closely and deeply punctate, surface intervals distinctly micropunctate; mandibles and labrum brownish dark; eyes large, proeminent; palpi light yellowish, apical half of the last articles darker, last article of both palpi fusiform, sharp at the tip; labrum with anterior margin triangularly emarginated; frontal forrows distinct, subparallel, extended to level of anterior supraorbital seta, joining to the inner ocular suture; frons slightly convex or depressed, occiput slightly concave; antennae flavescent, brown dark in inner side of 1 st to 10 th antennomers, surpassing the posterior margin of pronotum by the last two-three antennomers, 3 rd to 10 th with 4 ‒ 5 long bristles just bellow the joints, scape 2.5 times (holotype) and 2.3 ‒ 2.7 times (female allotype and paratypes) longer than wide [length: 0.32 mm (holotype), 0.31 ‒ 0.38 mm (female allotype and paratypes), width: 0.13 mm (holotype), 0.13 ‒ 0.14 mm (female allotype and paratypes)], pedicele 1.3 times (holotype) and 1.1 ‒ 1.4 times (female allotype and paratypes) longer than wide [length: 0.10 mm (holotype), 0.10 ‒ 0.11 mm (female allotype and paratypes), width: 0.08 mm (holotype), 0.08 ‒ 0.10 mm (female allotype and paratypes)], 3 rd to 4 th antennomers 2.1 times (holotype) and 2.1 ‒ 2.5 times (female allotype and paratypes) longer than wide [length: 0.21 mm (holotype), 0.19 ‒ 0.21 mm (female allotype and paratypes), width: 0.10 mm (holotype and female allotype and paratypes)], 5 th to 10 th 2.3 ‒ 2.6 times (holotype) and 2.2 ‒ 3.2 times (female allotype and paratypes) longer than wide [length: 0.21 ‒ 0.24 mm (holotype), 0.22 ‒ 0.32 mm (female allotype and paratypes), width: 0.09 ‒ 0.10 mm (holotype), 0.09 ‒ 0.10 mm (female allotype and paratypes)], and 11 th 2.7 times (holoptype) and 3 ‒ 3.4 times (female allotype and paratypes) longer than wide [length: 0.27 mm (holotype), 0.27 ‒ 0.31 mm (female allotype and paratypes), width: 0.10 mm (holotype), 0.09 mm (female allotype and paratypes)]. Cephalic chaetotaxy (large setae): Labrum with three pairs of setae gradually shorter from external to inner sides, one pair on sides of clypeus and two pairs of supraocular setae present over each eye. Thorax. Pronotum (Fig. 4) 1.36 times (holotype) and 1.23 ‒ 1.35 times (female allotype and paratypes) longer than wide [length: 0.99 mm (holotype), 1.01 ‒ 1.18 mm (female allotype and paratypes), width: 0.73 mm (holotype), 0.75 ‒ 0.91 mm (female allotype and paratypes)], greenish bronze, shiny, closely and deeply punctate, surface intervals distinctly micropunctate; wider than head, slightly subrectangular or inverted subtrapezoidal shaped, concussively widest at first third or close to the anterior angles; apical and basal margins not margined, lateral margins with complete and distinct margination in the two first thirds since the anterior angles, reaching the posterior angles, proepisterna well visible from above in the two first thirds; apical margin slightly emarginated, apical angles largely rounded, not protruded; lateral margins slightly subparallel or oblique at first two thirds, followed by a slight sinuosity before the roundly right basal angles, posterior margin slightly but distinctly arcuate; median line absent or very superflous, not reaching anterior or posterior margins, anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous; one marginal setae situated in the end of first third, the posterior marginal seta absent. Elytra (Fig. 1 b) 1.76 times (holotype) and 1.71 ‒ 1.82 times (female allotype and paratypes) longer than wide [length: 2.78 mm (holotype), 2.88 ‒ 3.36 mm (female allotype and paratypes), width: 1.58 mm (holotype), 1.68 ‒ 1.87 mm (female allotype and paratypes)], greenish bronze, shiny and with punctual metallic reflections; wider than pronotum, humeral angles broadly rounded; elongate and parallel, dorsally moderately convex, truncated obliquely behind, posterior obtuse angle sometimes slightly produced in a short spine, apical margin with 7 to 10 small denticles since the posterior angle to the sutural angle; striae deeply punctate, intervals narrow and convexes, transversely strongly rugose, surface distinctly micropunctate; yellowish patches: one sub-basal and other submedian on 6 th interval, more or less rounded or in a small longitudinal line, and two joint small longitudinal subapicals, respectively on 5 th and 6 th intervals; three setiferous punctures situated at 3 rd interval, one at basal quarter, other at median region and the last one at beginning of the last quarter. Ventral surface. Blachish-bronze, shiny; genae sparsely finely punctate, glabrous; proepisterna closely and deeply alveolo-punctate in the upper sides, becoming less densely punctate towards the ventral region; sternum glabrous; elytral epipleura smooth with some sparse and deeply punctures at first half; abdominal segments sparcely micropunctate, the first two covered lateraly with sparce dressed short hairs; last segment with 1 pair (male) or 2 pairs (female) of setae near the posterior margin; Legs. Slender, yellowish, almost transparent; knees, apical region of tibiae and tarsi brownish dark; trochanters black. Aedeagus (Fig. 5). Slender, median lobe slightly curved and apex slightly upwards (lateral view); internal sac with tangled membranes very weekly sclerotized twisted in a pre-dorso-apical position; apex slightly bent to left (dorsal view); parameres like figure 5 a. Intraspecific variation. The range of variability observed in L. schuelei sp. n. (5 specimens) affects the pronotum shape (see description), the elytral yellow pattern and the number of teeth in apical elytrae margin (see description). Concerning the variability within the elytral yellow patches, sometimes the sub-basal patch is absent in one elytron or on both, other times both the sub-basal and the median ones are also absent, remaining solely the subapicals. Asymmetries in the length of left and right elytron are common too.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F31B7F4114BFA490FE18FB8B.taxon	discussion	Remarks. The species of this genus are characterized, among other features (see Péringuey 1896), by the convex pronotum inflated lateraly in the median region and, consequently, presenting an oblong-ovate or fusiform shape more or less developed. The new species by the singular pronotum shape (subrectangular or even subtrapezoidal), presenting the lateral margins parallel or slightly oblique convergently backwards, is easily segregate from the remaining species. It seems closer to L. egregia Péringuey, 1896 by the reduced elytral patches, the elongate body, parallel elytra and the posterior elytral angle slightly denticled. Ecological notes. Adult specimens from Munenga (Calulo) were found during day on open, moist patches with sparse vegetation within a meadow with a few single trees. Adult specimens from Negola were found also during day, walking on a sunshine loamy path after rain, within an open secondary forest. The single female specimen from Cambamba (Catoio-Quela) was found beneath litter in the edges of a countryside road with sandy soil whitin a secondary open forest (Fig. 9 b) together with the tiger beetle L. wellmani and the ground beetles C. convexicollis and G. ellipticus.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA47FFEC3F86C.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Cunene; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA47FFEC3F86C.taxon	materials_examined	Material examined. Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 3. IV. 2014, 1 ♀, 12. XI. 2015, 4 ♀, DO, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA47FFEC3F86C.taxon	discussion	Remarks. Basilewsky (1977) refered to this species as a synonymous of G. amabilis Boheman, 1860. Lorenz (2005) restablished G. andersoni as a good species and considers the subspecies ovipennis Péringuey, 1892 as well as subovipennis Basilewsky, 1977 as synonymous, both with the status of subspecies of G. amabilis in Basilewsky (1977). Here we follow Lorenz (2005) criterium. The species is known from Angola, Botswana, Namibia and Zimbabwe (Basilewsky 1977, Insectoid. info 2017). Adults were found on sandy whitish soil or on reddish soil together with some adults of tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA1BDFAF6FACD.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA1BDFAF6FACD.taxon	materials_examined	Material examined. Chingueia-Mandongue (13 ° 35´46 ´´ S, 16 ° 58´59 ´´ E, 1648 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA1BDFAF6FACD.taxon	discussion	Remarks. A subspecies known only from Angola and described on the basis of Caluango specimens (Basilewsky 1977). Our specimens fit very well this subspecies. Adult specimens were found within an open secondary forest together with the tiger beetle E. muata parallelestriata, among other tiger and ground beetles (see S. lucidus strigiceps remarks). Interestingly, both species present a similar pattern of elytral bands (Figs 6 a, 6 b) suggesting a possible mimicry. Moreover, both species ressemble in the running behaviour also, thus being difficult to segregate them in situ without its capture. Similar observations were already reported to other Graphipterini / Cicindelini associations in Africa (Cassola & Taglianti 1988). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA339FB4AFD85.taxon	distribution	Distribution in Angola (Provinces): Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA339FB4AFD85.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 6. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3277F4014BFA339FB4AFD85.taxon	discussion	Remarks. A singleton female specimen was found on a wall near public lights (Caála) with other ground beetles (see E. peringueyi remarks). It was not possible to ascribe the specimen to any species.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4714BFA2ADFC9FFDE1.taxon	distribution	Distribution in Angola (Provinces): 1) Cunene; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4714BFA2ADFC9FFDE1.taxon	materials_examined	Material examined. Chitembo (6 km SE) (13 ° 33´23 ´´ S, 16 ° 47´22 ´´ E, 1586 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4714BFA2ADFC9FFDE1.taxon	discussion	Remarks. A species known from Angola and Namibia (Basilewsky 1977, Insectoid. info 2017) with two subspecies (jambanus Basilewsky, 1977 and ornatissimus Basilewsky, 1977), both restricted to the southern territory of Angola (Basilewsky 1977, 1986 a). Adults were found within an open secondary forest on sandy whitish soil together with some adults of tiger and ground beetles (see S. lucidus strigiceps remarks). The comments given to G. congoensis lundanus on mimicry associations between Cicindelini and Graphipterini can be reported also to this species (Figs 6 a, 6 c). It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4614BFA791FC36FE37.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Malanje, Benguela, Huambo, Huíla, Bié; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4614BFA791FC36FE37.taxon	materials_examined	Material examined. Cruzeiro (Pedra do Alemão) (12 ° 46´41.14 ´´ S, 15 ° 54´09.49 ´´ E, 1743 m alt., 232) (HUAMBO), 4. XI. 2014, 2 ♀, 7. XI. 2015, 1 ♀, DO, A Serrano & R. Capela leg., ASC; Calulo-Cabuta (9 ° 55´16.84 ´´ S, 14 ° 54´53.08 ´´ E, 826 m alt., 128) (KWANZA SUL), 5. XI. 2014, 3 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo-Cabeba (10 ° 02´18 ´´ S, 14 ° 55´49 ´´ E, 1061 m alt., 147) (KWANZA SUL), 5. XI. 2014, 3 ♀, DO, A. Serrano & R. Capela leg., ASC; Londuimbali-Tinguita (12 ° 18´10 ´´ S, 15 ° 21´15 ´´ E, 1633 m alt., 231) (HUAMBO), 2. XI. 2015, 5 ♂, 21 ♀, DO, A. Serrano & R. Capela leg., ASC; Longonjo-Catabola (13 ° 01´15 ´´ S, 15 ° 13´18 ´´ E, 1528 m alt., 279) (HUAMBO), 4. XI. 2015, 3 ♂, 5 ♀, DO, A. Serrano & R. Capela leg., ASC; Cuima- Cusse (13 km) (13 ° 20´45 ´´ S, 15 ° 34´22 ´´ E, 1633 m alt., 280) (HUAMBO), 5. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Ceilunga (Kuito) (12 ° 13´40 ´´ S, 17 ° 00´20 ´´ E, 1683 m alt., 235) (BIÉ), 10. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Kinglês (Lombe-Kalandula) (9 ° 23´00 ´´ S, 16 ° 0´7 46 ´´ E, 1121 m alt., 113) (MALANJE), 18. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Santa Maria (= Quiluanje) (8 ° 55´14 ´´ S, 15 ° 58´42 ´´ E, 1157 m alt., 94) (MALANJE), 19. XI. 2015, 2 ♂, DO, A. Serrano & R. Capela leg., ASC; Calulo (Faz. Klein) (10 ° 02´14 ´´ S, 14 ° 54´38 ´´ E, 1059 m alt., 147) (KWANZA SUL), 3. XII. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Calulo (N. Faz. Belo Horizonte) (9 ° 56´41 ´´ S, 14 ° 56´35 ´´ E, 882 m alt., 128) (KWANZA SUL), 3. XII. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3207F4614BFA791FC36FE37.taxon	discussion	Remarks. The nominal species is known only from Angola (Basilewsky 1977, 1986 b). Probably is the most common and widespread Graphipterus species in Angola. Adults were found in footpaths and country roads within open secondary forests running quickly when disturbed to hide beneath the litter. They are syntopic with some tiger and ground beetles, changing the species in accordance with the locality.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA683FC07F908.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Huíla, Cuando Cubango; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA683FC07F908.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.74 ´´ S, 16 ° 45´48.17 ´´ E, 1649 m alt., 281) (BIÉ), 1. XI. 2014, 6 ♂, 4 ♀, DO, A. Serrano & R. Capela leg., ASC; Cachingues (2 km E) (13 ° 04´54 ´´ S, 16 ° 46´08 ´´ E, 1647 m alt., 282) (BIÉ), 11. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Cuima-Cusse (13 km) (13 ° 20´45 ´´ S, 15 ° 34´22 ´´ E, 1633 m alt., 280) (HUAMBO), 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA683FC07F908.taxon	discussion	Remarks. An endemic species of Angola (Basilewsky 1977). In Bié Province the adult specimens were found within secondary meadows on sandy whitish soil together with some tiger and ground beetles [Cachingues- Chipica: P. angusticollis, Dromica similis Cassola, 1980, T. nubifera, L. infuscatula; Cachingues: P. angusticollis, D. similis, D. (Foveodromica) sp., L. infuscatula, L. saraliensis, Lophyra sumlini Cassola, 1976, Trianogenius ferox (Erichson, 1843), A. convexipennis]. In the Huambo locality the adult specimen was collected together with L. saraliensis and G. albomarginatus. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA051FE8AFBE9.taxon	distribution	Distribution in Angola (Provinces): 1) Cunene; 2) Bié, Cuando Cubango.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA051FE8AFBE9.taxon	materials_examined	Material examined. Chitembo (6 km SE) (13 ° 33´23 ´´ S, 16 ° 47´22 ´´ E, 1586 m alt., 302) (BIÉ), 9. XI. 2015, 2 ♂, DO, A. Serrano & R. Capela leg., ASC; Munquengue (14 ° 42´01 ´´ S, 17 ° 24´14 ´´ E, 1440 m alt., 343) (CUANDO CUBANGO), 14. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4614BFA051FE8AFBE9.taxon	discussion	Remarks. A species widespread throughout southern Africa (Namibia, South Africa, Bostwana, Zambia, Zimbabwe) (Basilewsky 1977, 1986 b). Putzeys (1880) recorded already the species for Angola (Humbe, Cunene Province), but Basilewsky (1977) considered erroneous this report. However, our specimens fit very well the species description, confirming the species presence in Angola. Moreover, their presence in the southeastern territory of Angola is very plausible, according to the known distribution in the northern territory of Namibia close to the Cunene River. Adult specimens were found on whitish sandy soil near Chitembo within an open secondary forest together with some tiger and ground beetles (T. nzingae, O. rufomarginata poggei, L. uncivittata, G. erikssoni) and near Munquengue within shrub savannah together with some tiger beetles (Dromica pentheri W. Horn, 1899 and Ophryodera rufomarginata bradshawi Péringuey, 1888). It is a new record for the Bié and Cuando Cubango Provinces.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4514BFA55BFB54FE37.taxon	distribution	Distribution in Angola (Provinces): 2) Malanje.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4514BFA55BFB54FE37.taxon	materials_examined	Material examined. Cambamba (Catoio-Quela) (09 ° 14´42 ´´ S, 17 ° 00´33 ´´ E, 1178 m alt., 115) (MALANJE), 20. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC; Santa Maria (= Quiluanje) (8 ° 55´14 ´´ S, 15 ° 58´42 ´´ E, 1157 m alt., 94) (MALANJE), 19. XI. 2015, 1 ♂, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Carima (Cacuso-Pungandongo) (09 ° 35´00 ´´ S, 15 ° 42´01 ´´ E, 1117 m alt., 130) (MALANJE), 23. XI. 2015, 3 ♂, 12 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3217F4514BFA55BFB54FE37.taxon	discussion	Remarks. The nominal species is known from D. R. of the Congo and Zambia (Basilewsky 1977, 1986 b) and the subspecies G. ellipticus upembanus Basilewsky, 1977 from the same countries plus Tanzania (Basilewsky op. cit.). Adult specimens collected in Malanje province fit well the nominal species description. Adults were found within open secondary forests together with tiger and ground beetles [Cambamba (Catoio-Quela) and Carima: see S. vilhenai and C. convexicollis remarks, respectively]. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA0C1FB8BFBE9.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA0C1FB8BFBE9.taxon	materials_examined	Material examined. Satchijamba (10 km E) (13 ° 43´38.76 ´´ S, 17 ° 12´17.74 ´´ E, 1601 m alt., 283) (BIÉ), 7. IV. 2014, 1 ♀, VM, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA0C1FB8BFBE9.taxon	discussion	Remarks. A species widespread throughout western, central, southern and eastern Africa (Senegal, Gambia, Guinea, Burkina Faso, Ivory Coast, Nigeria, Cameroon, D. R. of the Congo, Angola, South Africa R., Malawi, Kenya) (Basilewsky 1986 a, Schüle 2009, Insectoid. info 2017). It was imported to France also (Basilewsky 1986 a). The only record for Angola was reported by Basilewsky (1986 a) apparently to Lunda Norte province, but without any exact locality. The species seems associated with vacated termite nests of Macrotermes sp. (Isoptera) (Basilewsky 1986 a). The single adult female was collected by light trapping within an open secondary forest with other ground beetles (see S. harpaloides remarks). It is a new reord for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA683FEF1F92C.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte?; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA683FEF1F92C.taxon	materials_examined	Material examined. Chitembo (10 km N) (13 ° 22´40.78 ´´ S, 16 ° 41´58.07 ´´ E, 1666 m alt., 282) (BIÉ), 1. XI. 2014, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4514BFA683FEF1F92C.taxon	discussion	Remarks. A species widespread throughout western, central and southern Africa (Mauritania, Senegal, Gambia, Guinea-Bissau, Guinea, Chad, Ivory Coast, Mali, Angola, D. R. of the Congo, South Africa R.) (Serrano 2009, Insectoid. info 2017). It is with some reserve that we ascribe the singleton male adult specimen to this species. The apex of median lobe of aedeagus is slightly different of T. quadrimaculatus and more similar to the same organ of T. quadrum (Fabricius, 1792). However, this species is distributed on northern territories of Africa. The adult specimen was found in a country road within an open secondary forest. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4414BFA506FDC3FE12.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4414BFA506FDC3FE12.taxon	materials_examined	Material examined. Alto Ventura (Colé-Calulo) (09 ° 59´42 ´´ S, 14 ° 50´32 ´´ E, 1116 m alt., 128) (KWANZA SUL), 4. XII. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3227F4414BFA506FDC3FE12.taxon	discussion	Remarks. A species widespread throughout western, central and eastern Africa (Guinea, Ivory Coast, Togo, Liberia, Benin, Cameroon, Angola, D. R. of the Congo, Malawi, Tanzania) (Alves 1963, Insectoid. info 2017). The species was recorded for Angola by Alves (1963) (Dundo, Lunda Norte Province), but it was not given in Ferreira`s catalogue (1965). The single male collected near Calulo confirms its presence in Angola. The adult specimen was found under tree bark together with C. congolensis (see also C. intermedius remarks). It is a new record for the Kwanza Sul Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA767FE4CF9D0.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA767FE4CF9D0.taxon	materials_examined	Material examined. Cachingues-Chipica (13 ° 10´15.74 ´´ S, 16 ° 45´48.17 ´´ E, 1649 m alt., 281) (BIÉ), 2. IV. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA767FE4CF9D0.taxon	discussion	Remarks. A species known to western and central Africa (Guinea-Bissau, Guinea, Ivory Coast, Central African Republic, D. R. of the Congo) (Basilewsky 1970, Serrano 2009, Insectoid. info 2017). The single female specimen fits well with the original description and with specimens from Guinea-Bissau. The adult specimen was collected by light trapping together with other ground beetles (see P. holosericus remarks). It is a new genus / species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA0A5FD58FBB1.taxon	distribution	Distribution in Angola (Provinces): 2) Kwanza Sul.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA0A5FD58FBB1.taxon	materials_examined	Material examined. Alto Ventura (Colé-Calulo) (09 ° 59´42 ´´ S, 14 ° 50´32 ´´ E, 1116 m alt., 128) (KWANZA SUL), 4. XII. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4414BFA0A5FD58FBB1.taxon	discussion	Remarks. A species widespread throughout western, central and eastern Africa (Senegal, Togo, Cameroon, Equatorial Guinea, Gabon, D. R. of the Congo, Rwanda, Uganda) (Hansen 1968, Insectoid. info 2017). Adult specimens were found under tree bark together with P. liboloensis sp. n. and C. beauvoisi (see also C. intermedius remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4B14BFA506FE04FE37.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte, Huambo, Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4B14BFA506FE04FE37.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44 ° 46.50 °´S, 17 ° 11 ° 2.26 ´´ E, 1611 m alt., 303) (BIÉ), 2. XI. 2014, 2 ♂, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3237F4B14BFA506FE04FE37.taxon	discussion	Remarks. A species with a scattered known distribution in Africa (Angola, Zambia, Tanzania) (Facchini 2012 a). It was described recently by Facchini (2012 a) and our adult specimens fit very well its description, including the aedeagus features. Adult specimens were collected by light trapping within an open secondary forest together with some other ground and ant nest beetles (e. g. P. vanrooni, P. chappuisi, M. guineensis, A. monardianus, Stenolophus sp.).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA767FDFAF9D0.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA767FDFAF9D0.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X ‒ 2. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA767FDFAF9D0.taxon	discussion	Remarks. A species with a large distribution in Africa south of Sahara (Senegal, Guinea-Bissau, Ivory Coast, Mali, Chad, D. R. of the Congo, Angola, Zambia, Zimbabwe, Uganda, Mozambique, Tanzania, Kenya, Ethiopia, Sudan) (Basilewsky 1961, Ferreira 1965, Serrano 2009, Insectoid. info 2017). The single female specimen was collected in a wall near a public light together with other ground beetles (see C. rikatlensis remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA0C1FB16FBB1.taxon	distribution	Distribution in Angola (Provinces): 1) Lunda Norte; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA0C1FB16FBB1.taxon	materials_examined	Material examined. Satchijamba (10 km E) (13 ° 43 ° 38.76 ´´ S, 17 ° 12 ° 17.74 ´´ E, 1601 m alt., 283) (BIÉ), 7. IV. 2014, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4B14BFA0C1FB16FBB1.taxon	discussion	Remarks. A species widespread throughout western, central, southern and eastern Africa (Senegal, Gambia, Guinea-Bissau, Cameroon, Central African Republic, D. R. of the Congo, Angola, Namibia, South Africa R., Kenya) (Basilewsky 1955, Ferreira 1965, Serrano 2009, Insectoid. info 2017). Adult specimens were collected under the bark of decomposing trunks together with Zygentoma, Formicidae and Tenebrionidae adults and the ant nest beetle P. pseudoklugi within an open secondary forest. It is a new record for the Bié Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4A14BFA506FE06FE6F.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4A14BFA506FE06FE6F.taxon	materials_examined	Material examined. Chitembo (10 km N) (13 ° 22´40.78 ´´ S, 16 ° 41´58.07 ´´ E, 1666 m alt., 282) (BIÉ), 1. XI. 2014, 1 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32C7F4A14BFA506FE06FE6F.taxon	discussion	Remarks. A species widespread throughout western, central, southern and eastern Africa (Guinea-Bissau, Guinea, Ivory Coast, D. R. of the Congo, South Africa R., Rwanda, Zambia, Zimbabwe, Mozambique, Tanzania) (Serrano 2009, Insectoid. info 2017). The singleton female specimen fits well the original description and is similar to Guinea-Bissau specimens (see Serrano 2009). The adult specimen was collected by light trapping within an open secondary forest together with a tiger beetle (L. sumlini) and ant nest beetles [Heteropaussus trapezicollis (Wasmann, 1922), Paussus (Spinicoxipaussus) cridae Gestro, 1915] (see Serrano & Capela 2015 a, 2015 b). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA0C1FE4CFB8D.taxon	distribution	Distribution in Angola (Provinces): 2) Luanda.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA0C1FE4CFB8D.taxon	materials_examined	Material examined. Barra do Cuanza (09 ° 18´55.42 ´´ S, 13 ° 09´58.57 ´´ E, 11 m alt., 107) (LUANDA), 28. IV. 2014, 2 ♀, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA0C1FE4CFB8D.taxon	discussion	Remarks. A species known until now from Botswana and Namibia (Insectoid. info 2017). Adult specimens fit very well the original description and are similar to a specimen from Namibia [South of Walvis Bay, " Musee Royal de l’Afrique Centrale (Belgium, Tervuren) ”] and determined by P. Basilewsky (in Anichtchenko 2016). Adults were collected by light trapping close to River Cuanza within a riparian forest together with one ant nest beetle [Carabidomemnus (Carabidodoxus) sp.] and two ground beetles (D. picicornis and M. guineensis). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA7D0FA67F909.taxon	distribution	Distribution in Angola (Provinces): 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA7D0FA67F909.taxon	materials_examined	Material examined. Caála (12 ° 51´11 ´´ S, 15 ° 33´07 ´´ E, 1758 m alt., 256) (HUAMBO), 6. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4A14BFA7D0FA67F909.taxon	discussion	Remarks. A species widespread throughout the Mediterranean Region, West Asia and Africa (Basilewsky 1960 b, Insectoid. info 2017). Curiously, it was never recorded for Angola. A singleton female specimen was found on a wall near public lights with other ground beetles (see E. peringueyi remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4914BFA493FE22FEDC.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4914BFA493FE22FEDC.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 1. IV. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32D7F4914BFA493FE22FEDC.taxon	discussion	Remarks. A species with a scattered distribution throughout western, central, southern and eastern Africa (Guinea, Ivory Coast, Cameroon, D. R. of the Congo, South Africa R., Rwanda, Tanzania, Kenya) (Basilewsky 1960 b, Insectoid. info 2017). The single male was collected by light trapping within an open secondary forest together with P. pseudocrenulatus. It is a new species record for Angola. Dendrocellus Schmidt-Göbel, 1846 A genus considered a synonymous of Desera Dejean 1825 by Lorenz (2005). However, it was reestablished before as a valid genus by Liang et al. (2004). There are three species known from Angola (Liang & Kavanaugh 2007, Serrano 2008 ‒ 2009).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4914BFA3DAFCF7FC97.taxon	distribution	Distribution in Angola (Provinces): 1) Malanje; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4914BFA3DAFCF7FC97.taxon	materials_examined	Material examined. Alto Hama (12 ° 13´20 ´´ S, 15 ° 32´57 ´´ E, 1512 m alt., 232) (HUAMBO), 31. X ‒ 2. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4914BFA3DAFCF7FC97.taxon	discussion	Remarks. A species known for Angola, Zambia and D. R. of the Congo (Liang & Kavanaugh 2007, Serrano 2008 ‒ 2009). It was recorded for the first time to Angola based on one male specimen collected in “ Parque Nacional da Cangandala, a protected area for the emblematic giant sable antelope Hippotragus niger variani Thomas, 1916. The new adult specimens collected in Huambo confirm its presence in Angola. Adult specimens were found at a wall during night near a public light together with other ground beetles (see C. rikatlensis remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.taxon	materials_examined	Type series. Holotype, ♂; Angola (Kwanza Sul), Calulo-Cabuta (9 ° 53´59 ´´ S, 14 ° 54´26 ´´ E, 831 m alt., 128) (KWANZA SUL), 29. XI ‒ 6. XII. 2015, PF, A. Serrano & R. Capela leg., ASC. Allotype 1 ♀: Mussende (Calulo) (9 ° 57´00 ´´ S, 14 ° 47´36 ´´ E, 905 m alt., 128) (KWANZA SUL), 28. XI ‒ 5. XII. 2015, PF, A. Serrano & R. Capela leg., ASC; Paratypes, 2 ♂, same locality and date as Holotype, A. Serrano & R. Capela leg., ASC, PSC; 1 ♀, same locality and date as Allotype, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.taxon	etymology	Derivatio nominis. This subspecies is named in honour of Rúben Capela, a specialist on Diptera Culicidae and Ceratopogonidae, which has collaborated with the first author (AS) in the Angola entomological trips.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.taxon	diagnosis	Diagnosis. Similar in form and color to G. procera Gerstaecker, 1867; five pairs of impar elytral carinae well developed, the 2 nd par carinae (4 th carinae) only slightly distinct in the first quarter, 3 rd par carinae (6 th carinae) distinct in the first and last thirds of elytrae, remaining par carinae almost indistinct, scutellar carina almost indistinct joining the first carina; median lobe of aedeagus with a hooked apex ventraly backwards (Figs 8 a, 8 b).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.taxon	description	Description. Length of Holotype: 25.5 mm. Length of paratypes (without labrum): 25.32 ‒ 25.33 mm (males), 25.89 ‒ 26.30 mm (females). Head (Fig, 7 a) 1.2 ‒ 1.3 times longer than wide [length: 4.62 mm (holotype and male paratypes), 4.62 ‒ 4.82 mm (female allotype and paratype), width: 3.68 mm (holotype), 3.62 ‒ 3.68 mm (male paratypes), 3.71 ‒ 3.81 mm (female allotype and paratype)], black, eyes small, slightly wider than temples; front longitudinaly protruded in the median region, almost smooth, two short deeply furrows lateraly; occiput surface largely rugose-punctate, covered with very sparce blackish semierected pubescence, surface intervals smooth; mandibles and labrum blackish; palpi blackish, apical region of the last articles brownish, last article of both palpi strongly securiform; labrum with anterior margin straight or slightly arcuate; antennae proeminent, densely pubescent, decreasing in thickness towards apex, first four antennomers blackish, gradually brown dark to lighter brown since the 5 th to the 11 th, reaching or slightly surpassing the middle of elytra; Cephalic chaetotaxy (large setae, all brownyellowish): Labrum with three submarginal pairs of long setae, two pairs of long setae in a transversal middle line throughout the first half of clypeus disk, one pair on sides behind this anterior series and two pairs of very long supraocular setae present over each eye; some other not so long setae dispersed over the sides of clypeus and frons. Thorax (Fig. 7 a) Pronotum 1.13 ‒ 1.21 times longer than wide [length: 5.68 mm (holotype), 5.71 ‒ 5.74 mm (male paratypes), 5.68 ‒ 5.87 mm (female allotype and paratype), width: 5.02 mm (holotype), 4.82 ‒ 4.95 mm (male paratypes), 4.69 ‒ 4.88 mm (female allotype and paratype)], black, wider than head, barely subcordiform, widest near the middle; lateral margins slightly raised, narrow, regularly round in the first two thirds, slightly subparallel or sinuate before the rectangular basal angles, round at tip; apical angles roundly acute, slightly protruded; apex and base finely margined, the former arcuate, the latter almost straight; disk gently convex; median line in middle slightly more impressed than anteriorly and posteriorly, neither reaching apex nor base; anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous in males, absent in females; basal grooves barely indicated; lateral margin with two elongate setae, one situated slightly before the middle, the other before the basal angle, two ‒ three short setae near the anterior angles, a fringe of brown-reddish hairs at anterior and posterior margins (shorter in the former); surface strongly transversely punctate-rugose in sides and base, slighter in the disk, covered with sparce blackish semierected pubescence, microreticulation distinct, extremely fine, consisting of transverse meshes, shiny. Elytra (Fig. 7 a) 1.66 ‒ 1.79 times longer than wide [length:, 14.36 mm (holotype), 14.23 ‒ 14.36 mm (male paratypes), 14.90 ‒ 15.30 mm (female allotype and paratype), width: 8.65 mm (holotype), 8.05 ‒ 8.11 mm (male paratypes), 8.31 ‒ 8.91 mm (female allotype and paratype)], blackish, shiny, sometimes with violet reflections; wider than pronotum, broadly elongate-ovate, humeral angles not distinct, widest point behind the middle; dorsally very slightly convex, truncated obliquely at apex; each elytron with five distinct impair carinae, the 2 nd par carinae (4 th carinae) only slightly distinct in the first quarter, 3 rd par carinae (6 th carinae) distinct in the first and last thirds of elytrae, remaining par carinae almost indistinct, scutellar carina almost indistinct joining the first carina; intervals between two consecutive impair carinae with two longitudinal deep punctate rows, each flanked lateraly by a row of brownish setae from base to apex, totalizing three rows; 4 th interval narrower than others, by this with only one longitudinal punctate row flanked lateraly by a row of brownish setae from base to apex; apical truncature with blackish pubescence; surface intervals covered with distinct, dense, transversely punctate-granulate meshes (Fig. 7 c); hind wings absent. Ventral surface. Blach to brown dark, shiny in the head and thorax, dull in abdomen; genae sparsely punctate, sides covered with sparse black semierected setae; proepisterna smooth, not punctate in the upper sides, becoming sparsely punctate towards the ventral region; metaepisterna, pro, meso and metasterna densely punctate and pubescent (brownish color); elytral epipleura moderately punctate; abdominal segments densely punctate and pubescent like thoracic sterna; posterior margin of last segment deeply (males) or slightly (females) truncate in the middle region, one pair (males and females) of setae near the posterior margin. Legs are conformed to the genus morphological pattern. Male forelegs with the three basal tarsomeres of tarsi more dilated in the inner region than in the outer (asymmetrical dilated), the 4 th lesser asymmetrical, bilobated, presenting two rows of elongate, sensorial phaneres beneath. Aedeagus (Figs 8 a, 8 b, 8 c). Median lobe short, robust, apex hooked ventraly backwards (Fig. 8 b). Endophalous with a large postero-latero-dorsal scaly plate. Left paramere large, round at apex, right atrophied. Intraspecific variation. The range of variability observed in G. procera capelai ssp. n. (5 specimens) affects the median region of frons, more or less longitudinaly protruded and the pronotum shape (more or less subcordiform). Asymmetries in the length of left and right elytron are common too. The pronotum is partially pubescent or almost glabrous, accordingly with the presence / absence of decumbent setae.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32E7F4E14BFA63BFD41F857.taxon	discussion	Remarks. The subspecies of G. procera, including the nominal one, are easily segregated by the apical conformation of the median lobe of aedeagus than by external morphological characters (see Basilewsky 1963). The representants of this complex (eight subspecies including the nominal one) are distributed from Angola to Kenya and Tanzania, throughtout D. R. of the Congo, Zambia, Burundi, Rwanda and Uganda. Galerita procera angolana Basilewsky, 1963 was the unique subspecies known until now from Angola. By the recorded localities given (Basilewsky 1963, Ferreira 1965) it seems distributed in the central and eastern territories of Angola (Huambo, Lunda Norte and Moxico Provinces). Interestingly, G. procera capelai ssp. n. by the external morphological characters and shape of apical median lobe of aedeagus seems closer to the nominal subspecies than the already known Angolan subspecies. The nominal subspecies is proper of eastern Africa (Kenya and Tanzania), very faraway of the new subspecies locality. The other seven subspecies are distributed between the two taxa. In southern Africa this singular relationship in which species closer morphologicaly are strongly allopatric (western vs eastern) seems more common than expected (e. g. Serrano 1995). The new subspecies by the singular apical shape of median lobe (ventraly hooked backwards) (Fig. 8 b) is easily segregate from the remaining subspecies (see Fig. 20, in Basilewsky 1963). Galerita procera capelai ssp. n. can be separated also from G. procera angolana, the subspecies closer territorialy of the new subspecies, by a very different shape of the median lobe apex (cf Figs 8 b vs 8 e) and further by the elytral carinae less elevated, the intervals slightly larger and less excavated, the rows between impair carinae shallower punctate and surface intervals covered with more distinct and dense transversely punctate-granulate meshes. Ecological notes. Adult specimens were collected by means of pitfall trapping within secondary rainforest patches with moist soils, covered with dense and high litter (Fig. 9 c), together with adults of some tiger and ground beetles [Dromica (Foveodromica) sp., A. optimus, A. distinctus, D. tarsalis, C. magnicollis discrepans, O. gilvipes and O. patroboides]. As happens with some carabid species living mainly at moist habitats (e. g. Rossi & Santamaria 2001) the adult specimens of the new species were more or less infected with ectoparasitic fungi of the order Laboulbeniales on pronotum as well on elytra. Some study cases were already reported to other African carabid species (e. g. Santamaria & Faille 2009).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32A7F4C14BFA5FEFBACFE37.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32A7F4C14BFA5FEFBACFE37.taxon	materials_examined	Material examined. Chitembo (18 km SE) (13 ° 36´17.53 ´´ S, 16 ° 54´00.44 ´´ E, 1538 m alt., 302) (BIÉ), 30. X. 2014, 1 ♂, LT, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32A7F4C14BFA5FEFBACFE37.taxon	discussion	Remarks. A species known from Cameroon, Ivory Coast and D. R. of the Congo (Basilewsky 1962, Insectoid. info 2017). Curioursly M. katangensis Burgeon, 1937, the species reported to Angola by Basilewsky (1962), is referred neither in Ferreira catalogue (1965) nor in Insectoid. info (2017). The singleton male adult specimen of M. informicollis was collected by light trapping within an open secondary forest together with some other ground beetles (see S. angolense remarks). It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA683FBBCF998.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Huíla; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA683FBBCF998.taxon	materials_examined	Material examined. Cuima-Cusse (13 ° 22´52 ´´ S, 15 ° 30´15 ´´ E, 1583 m alt., 280) (HUAMBO), 5. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA683FBBCF998.taxon	discussion	Remarks. An endemic species of Angola described originaly from Benguela (Kleinfeld & Puchner 2012). Adult specimens were found beneath litter within open secondary forests together with some tiger and other ground beetles (see Orthogonius sp. 1 remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA0FDFF02FC55.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Namibe, Huambo; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA0FDFF02FC55.taxon	materials_examined	Material examined. Cachingues (2 km E) (13 ° 04´54 ´´ S, 16 ° 46´08 ´´ E, 1647 m alt., 282) (BIÉ), 11. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F4C14BFA0FDFF02FC55.taxon	discussion	Remarks. A species spread in Angola and Namibia (Basilewsky 1959, Insectoid. info 2017). Basilewsky (1959) reported that the species was distributed in the western and southwestern Angolan areas. The singleton male adult was found on sandy whitish soil within an open secondary forest together with some tiger beetles (see G. alternatus remarks). It is a new record for the Bié Province enlarging its distribution to the eastern territory of Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F5314BFA55EFAB5FEDA.taxon	distribution	Distribution in Angola (Provinces): 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F5314BFA55EFAB5FEDA.taxon	materials_examined	Material examined. Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F32B7F5314BFA55EFAB5FEDA.taxon	discussion	Remarks. A species described and known only from Zambia (Schüle & Heins 2013). The single female specimen was checked with one female paratype of this species. There is another species within this genus reported to Angola (A. mulsanti mulsanti Perroud, 1847) which presents a larger distribution throughout central, southern and eastern Africa) (Háva & Hovorka 2007, Kleinfeld & Puchner 2012). The adult was collected through pitfall trapping within an open secondary forest and is syntopic with some tiger and ground beetles (see S. lucidus strigiceps remarks). It is a new species record for Angola, enlarging its distribution to southwestern Africa.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA0FDFD0BFC1E.taxon	distribution	Distribution in Angola (Provinces): 1) Benguela, Namibe.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA0FDFD0BFC1E.taxon	materials_examined	Material examined. Baixo Pundo (12 ° 20´22.32 ´´ S, 13 ° 50´20.97 ´´ E, 275 m alt., 227 / 8) (BENGUELA), 25 ‒ 27. X. 2013, 1 ♂, 2 ♀, PF, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA0FDFD0BFC1E.taxon	discussion	Remarks. An endemic species of Angola originaly described from Benguela (Kleinfeld & Puchner 2012). Adult specimens were collected within bush habitat with sparse grassy vegetation and sandy soil by pitfall trapping, together with the ground beetle Harpalus sp.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA186FDC9FA27.taxon	distribution	Distribution in Angola (Provinces): 2) Benguela.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA186FDC9FA27.taxon	materials_examined	Material examined. Chamume (12 ° 41´38 ´´ S, 13 ° 08´51.9 ´´ m E, 6 m alt., 251) (BENGUELA), 28. X. 2013, 1 ♀, PF, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA186FDC9FA27.taxon	discussion	Remarks. The species is polymorphic with four recognized subspecies distributed mainly throughout southern and eastern Africa (Kleinfeld & Puchner 2012, Insectoid. info 2017). The nominal subspecies occurs in several South Africa Provinces, Botswana and Namibia (Kleinfeld & Puchner 2012). We do not ascribe our singleton female to any known subspecies, but she looks like the nominal form with the gray-whitish haired longitudinal median pronotal sulcus and the elytra totally glabrous. It could be ascribed to Cypholoba edax Péringuey, 1892 also, but our specimen does not present any withish pubescence at the elytral base and the elytrae are more domed. It is a new species record for Angola.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA79FFC08F80C.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla, Cunene; 2) Huambo.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA79FFC08F80C.taxon	materials_examined	Material examined. Cuima-Cusse (13 ° 22´52 ´´ S, 15 ° 30´15 ´´ E, 1583 m alt., 280) (HUAMBO), 5. XI. 2015, 7 ♂, 3 ♀, 6. XI. 2015, 1 ♂, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 2 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3347F5314BFA79FFC08F80C.taxon	discussion	Remarks. The species is very polymorphic (differences in the setal patterns of the elytra), with fifteen recognized subspecies distributed mainly throughout central and eastern Africa (Lorenz 2005, Kleinfeld & Puchner 2012, Insectoid. info 2017). As referred by Mawdsley et al. (2012) “ Many of these taxa are doubtfully distinct from the nominate form and the whole group is in need of a careful revision ”, which is not the scope of this work. Cypholoba graphipteroides monardi seems proper to the south of Angola (Ferreira 1965). Adults were found in diurnal activity beneath litter and patches within open secondary forests together with other tiger and ground beetles (see S. vilhenai remarks). It is a new record for the Huambo Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA45BFDECF815.taxon	distribution	Distribution in Angola (Provinces): 1) Namibe; 2) Benguela.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA45BFDECF815.taxon	materials_examined	Material examined. Lobito (12 ° 21´S, 13 ° 33´E, 10 m alt., 227 / 8) (BENGUELA), 20. III. 1998, 2 ♀, 10. V. 1998, 1 ♂, 1 ♀, PF, A. Neves leg., ASC; Biópio ‒ Benguela (12 ° 27´38.17 ´´ S, 13 ° 44´02.92 ´´ E, 148 m alt., 228) (BENGUELA), 27. X. 2013, 1 ♂, PF, R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA45BFDECF815.taxon	discussion	Remarks. The nominal species has a wide distribution throughout central, southern and eastern Africa (Angola, Namibia, South Africa, Zambia, Bostwana, Zimbabwe, Mozambique), but this subspecies is proper to South Angola and North Namibia (Kleinfeld & Puchner 2012). Adult specimens were collected by pitfall trapping in the outskirts of Lobito (see also B. apicalis remarks) and near Biópio within open xeric bush in sandy soil. It is a new record for Benguela Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA2EBFB0BFDA8.taxon	distribution	Distribution in Angola (Provinces): 1) Cuando Cubango; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA2EBFB0BFDA8.taxon	materials_examined	Material examined. Chingueia-Mandongue (13 ° 35´46 ´´ S, 16 ° 58´59 ´´ E, 1648 m alt., 302) (BI), 9. XI. 2015, 1 ♂, 2 ♀, DO, A. Serrano & R. Capela leg., ASC; Munquengue (14 ° 42´01 ´´ S, 17 ° 24´14 ´´ E, 1440 m alt., 343) (CUANDO CUBANGO), 14. XI. 2015, 2 ♀, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA2EBFB0BFDA8.taxon	discussion	Remarks. An endemic species of Angola described on the basis of one specimen from the Menongue region (not Huíla District, but Cuando Cubango Province) (Bénard 1921 in Kleinfeld & Puchner 2012). Ferreira (1965) reported “ distrito do Moxico, Menong (= Menongue) (MP) ”, which is not correct also. Adults were found in activity in open secondary forest (first locality) and bush savannah (second locality) on sandy withish soil together with some tiger and ground beetles (see S. lucidus strigiceps and G. limbatus remarks, respectively).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA116FBA6FA92.taxon	distribution	Distribution in Angola (Provinces): 1) Unknown; 2) BIÉ.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA116FBA6FA92.taxon	materials_examined	Material examined. Satchijamba-Somakwanza (13 ° 44´46.50 ´´ S, 17 ° 11´2.26 ´´ E, 1611 m alt., 303) (BIÉ), 31. X. 2014, 1 ♂, 1 ♀, 2. XI. 2014, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catota (2,5 km S) (14 ° 00´37 ´´ S, 17 ° 24´00 ´´ E, 1532 m alt., 323) (BI), 2. XI. 2014, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3357F5214BFA116FBA6FA92.taxon	discussion	Remarks. A species spread throughout southern, central and part of eastern Africa (Angola, Namibia, South Africa, Zambia, Bostwana, Zimbabwe, Mozambique) (Ferreira 1965, Kleinfeld & Puchner 2012). The species was recorded for Angola without any reference to a known locality (Ferreira 1965). Adult specimens were found in nocturnal activity (first locality) and diurnal activity (second locality) within open secondary forests together with other ground beetles (first locality: see S. tenebricosus fatalis remarks) and the tiger beetles P. angusticollis, D. fredericoi, D. pentheri, Dromica serietuberculata W. Horn, 1929, T. pepetela, L. uncivittata and C. agualusai (second locality) (see also Serrano et al. 2015). It is a new record for the Bi Province.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA2EBFEFFFDDF.taxon	distribution	Distribution in Angola (Provinces): 1) Huíla; 2) Bié.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA2EBFEFFFDDF.taxon	materials_examined	Material examined. Cachingues (2 km E) (13 ° 04´54 ´´ S, 16 ° 46´08 ´´ E, 1647 m alt., 282) (BIÉ), 11. XI. 2015, 1 ♂, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Mumbué (Cuanza riverhead) (13 ° 49´41 ´´ S, 17 ° 19´26 ´´ E, 1511 m alt., 303) (BIÉ), 8 ‒ 17. XI. 2015, 1 ♀, PF, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA2EBFEFFFDDF.taxon	discussion	Remarks. An endemic species of Angola (Puchner 2013). It was synonimized with Anthia calida Harold, 1878 by Basilewsky (1960), but re-established as a good species by Puchner (2013). Our specimens fit very well this species, presenting the typical subhumeral band (Puchner 2013, figs 9, 10). Adult specimens were found within secondary meadows on sandy whitish soil together with some tiger and ground beetles (see G. alternatus and S. lucidus strigiceps remarks for the first and second localities, respectively). The species was known only for the south of Angola (Huíla Province), but the new localities enlarges its distribution for the eastern territory (Bié Province).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA0C7FE04FB89.taxon	distribution	Distribution in Angola (Provinces): 1) Huambo, Huíla.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA0C7FE04FB89.taxon	materials_examined	Material examined. Bembua-Camenhe (12 ° 16´53 ´´ S, 15 ° 27´01 ´´ E, 1686 m alt., 231) (HUAMBO), 1. XI. 2015, 1 ♀, DO, A. Serrano & R. Capela leg., ASC; Catata-Nova Monção (13 ° 25´59 ´´ S, 15 ° 21´22 ´´ E, 1607 m alt., 279) (HUAMBO), 6. XI. 2015, 1 ♂, DO, A. Serrano & R. Capela leg., ASC.	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
03EFF252F3367F5114BFA0C7FE04FB89.taxon	discussion	Remarks. Another endemic species of Angola localized in the Huambo and Huíla Provinces (Puchner 2013). Adult specimens fit very well the Puchner (2013) description, including the aedeagus. Adults were found beneath litter within an open secondary forest (first locality) and in an open secondary bush savannah (second locality) together with some other tiger and ground beetles (see S. lucidus strigiceps and S. vilhenai remarks for the first and second localities, respectively).	en	Serrano, Artur R. M., Capela, Rúben A., Santos, Carmen Van-Dúnem Neto (2017): Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae). Zootaxa 4353 (2): 201-256, DOI: 10.11646/zootaxa.4353.2.1
