taxonID	type	description	language	source
03EC87C09F4BF30A20D8FF6DFC44FE50.taxon	description	(Fig. 1 – 3, 7, 10 – 12)	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
03EC87C09F4BF30A20D8FF6DFC44FE50.taxon	materials_examined	Material studied: 1 ♁, Madagascar, Ranomafana National Park, 21.26 S, 47.43 E, 900 – 1000 m, 12.04.2019, sifted from leaf litter in secondary forest, leg. P. Jałoszyński (MNHW). Emended diagnosis. Head with arcuate, anteriorly convex frontal groove accompanied at each side by elongate, oblique impression and with a pair of large shallow impressions each near mesal margin of supraantennal region; pronotum with six pairs of longitudinal sulci, of which only the lateral pair reaches anterior pronotal margin, each of four median sulci strongly broadened behind middle, pronotum with transverse, oval antebasal impression between the two lateralmost sulci; all antennomeres less than 3 times as long as broad; ‘ cavities’ of abdominal segment VIII in male conspicuously large, each about as broad as 1 / 4 width of segment; posterior margin of sternite IX in male with broad median area devoid of denticles; aedeagus slender, in dorsal view parallel-sided between base and distal 1 / 4, in lateral view weakly sinuate. Supplementary description. Head in male with paired impressions, these internally with smooth surface, not filled with setae (Fig. 1 – 2); antennomeres 3 – 9 elongate but each less than 3 times as long as broad (Fig. 3); male abdominal segment VIII (Fig. 7) with a pair of large internal ‘ cavities’, each nearly as broad as 1 / 4 of width of segment; posterior margin of sternite IX in male (Fig. 10) with 4 – 5 sharp denticles on each side, median area arcuate, lacking denticles. Aedeagus (Fig. 11 – 12) slender and conspicuously large, ~ 1 / 3 as long as body, in dorsal view (Fig. 11) parallelsided from base to distal 1 / 4, in lateral view (Fig. 12) weakly sinuate; endophallic structures indistinct, with numerous transverse folds of internal sac barely discernible under light microscope (better so with phase contrast).	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
03EC87C09F4BF30A20D8FF6DFC44FE50.taxon	discussion	Remarks. Details of the aedeagus and abdominal structures were not described previously for this species. As a part of diagnosis, “ large, shallow, setae bearing pits on the head ” were mentioned by Darby (2013). His SEM micrographs of the pronotum and head (Darby 2013: Fig. 20) agree with the characters seen in the specimen examined during the present study. The antenna (Darby 2013: Fig. 28) generally agrees with that illustrated here (Fig. 3), although with slight differences in the shape of some flagellomeres. Similar differences between the antennae of D. bisulcatus figured by Darby (2013: Fig. 27) and that in Fig. 6 in the present paper can be seen, and they can be attributed to different study methods (light microscopy of a specimen in liquid medium and hand drawing, vs. SEM of dried specimens). A close-up of the head of D. vasilii in Darby (2013: Fig. 23 – 24) shows paired impressions, similar to those found during the present study, except that there is another, small median impression in front of the transverse arcuate sulcus. The impressions do not seem to bear any setae, but appear as filled with dirt. The same paired impressions in the specimen examined and illustrated in the present study (Fig. 2) are smooth and empty, and the median impression is missing. The presence and absence of the latter may reflect variability within species, or sexual dimorphism (the specimen illustrated by Darby (2013) is of unknown sex). Discheramocephalus vasilii, as noted by Darby (2013), shares the same number of pronotal sulci with D. elisa- bethae Grebennikov, 2008, and D. mikaeli Grebennikov, 2008. Discheramocephalus elisabethae [Cameroon] lacks the arcuate transverse frontal groove, its postocular transverse groove bears submedian longitudinal carinulae flanking a narrow smooth area, and the four median pronotal sulci are indistinctly broadening posterad. Each of these characters is different in D. vasilii, which has a transverse frontal groove on the head, the smooth median region of the postocular groove not demarcated by longitudinal carinulae, and the pronotal sulci strongly, almost abruptly broadening posterad. Discheramocephalus mikaeli [Tanzania] differs from D. vasilii in the same cephalic characters as D. elisabethae, and additionally in the median pair of pronotal sulci much broader and reaching anterior pronotal margin, vs. narrow and not reaching the anterior margin in D. vasilii.	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
03EC87C09F4AF30D20D8F8AAFC0FFC49.taxon	description	(Fig. 1 – 3, 7, 10 – 12)	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
03EC87C09F4AF30D20D8F8AAFC0FFC49.taxon	materials_examined	Material studied: 1 ♁, 2 ♀♀, Madagascar, Ranomafana National Park, 21.26 S, 47.43 E, 900 – 1000 m, 12.04.2019, sifted from leaf litter in secondary forest, leg. P. Jałoszyński (MNHW). Emended diagnosis. Head with biarcuate (broadly M-shaped) frontal groove, lacking well-defined paired impressions; frons in front of groove strongly microsculptured; pronotum with two pairs of longitudinal sulci reaching anterior pronotal margin, inner pair gradually broadening posterad, pronotum also with two pairs of oval antebasal impressions between sulci; proximal flagellomeres more than 3 times as long as broad; ‘ cavities’ of abdominal segment VIII in male and female narrower than 1 / 4 width of segment; posterior margin of sternite IX in male and female similar, with broad median area devoid of denticles; aedeagus stout, in dorsal view with rounded sides, in lateral view abruptly bent ventrad at obtuse angle. Supplementary description. Male (Fig. 4 – 5) not differing externally from female; antennae (Fig. 6) with conspicuously elongate antennomeres 3 – 9, each 3 times or more longer than broad; abdominal segment VIII in both sexes with pair of small lateral internal ‘ cavities’, each narrower than 1 / 4 of width of segment, in male (Fig. 8) ‘ cavities’ appear slightly smaller than in female (Fig. 9); posterior margin of sternite IX in both sexes (Fig. 13 – 14) of similar shape, with lateral groups of 2 – 3 large denticles and arcuate median region lacking denticles. Aedeagus (Fig. 15 – 17) relatively stout and short, less than 0.2 times as long as body, in dorsal view (Fig. 15) with evenly rounded sides, apex (Fig. 16) subtrapezoidal, in lateral view distinctly bent ventrad by an obtuse angle (Fig. 17). Endophallus poorly visible (also with phase contrast). Spermatheca (Fig. 18) simple, lacking loops and coils, with elongate funnel.	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
03EC87C09F4AF30D20D8F8AAFC0FFC49.taxon	discussion	Remarks. Males of this species were not known so far; they are externally indistinguishable from females. The latter were adequately described by Darby (2013), except for the details of abdominal structures, which are added in the present study. Discheramocephalus bisulcatus clearly differs from its sympatric Malagasy relative D. vasilii in cephalic, pronotal and genital characters while both species have a similar shape of the posterior margin of the sternite IX. This latter structure, i. e. the median portion of the posterior margin lacking denticles and flanked by groups of large teeth, differentiates the Malagasy species from those known from other regions.	en	Jałoszyński, Paweł (2020): Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae). Zootaxa 4885 (4): 591-595, DOI: 10.11646/zootaxa.4885.4.8
