taxonID	type	description	language	source
03EC132F702F8D0AE2AD94CDDF58AF4C.taxon	materials_examined	Type species: Neophyllobius lombardinii Summers & Schlinger, 1955, 550.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702F8D0AE2AD94CDDF58AF4C.taxon	diagnosis	Diagnosis. According to Bolland (1986) the genus can readily be recognized by the following combination of characters: Idiosoma with 14 or 15 pairs of dorsal setae, five or six pairs of dorsomedian pdx, c 1, d 1, e 1, f 1 and h 1; four pairs of propodosomal vi, ve, sci, sce and five pairs of hystero-lateral setae c 2, d 2, e 2, f 2 and h 2; seta pdx present or absent (if present then with one or two pdx); palp tarsus with or without solenidion, with one or two eupathidia, and with one or two simple setae. Two pairs of eyes present. Peritreme with 1 – 3 loops or complex; coxa I with 1 or 2 setae; femur I with 3 or 4 setae; genua I – IV with one seta; tibia I – II with 9 (1 or 2 φ) - 8 or 9 (1 φ) setae; tibia I – IV with 9 (1 φ) - 8 (1 φ) - 8 (1 φ) - 7 (1 φ) setae; tarsi I – IV 9 (1 ω) or 10 (1 ω) - 10 (1 ω) or 8 (1 ω) – 7 or 7 (1 ω) – 7 or 7 (1 ω) and each of tarsi I – II with one sub-basal solenidion, and two variously spaced setae; tarsi III – IV with one medially placed seta.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702C8D0EE2AD94CDDA3CAA7E.taxon	diagnosis	Diagnosis. Dorsal idiosoma without pdx seta; palp tarsus with two simple setae; peritreme with one loop; dorsal seta c 1 190 (183 – 208) [193]; the ratio of dorsal setae c 1: d 1: e 1: f 1 as follows: 1.9 (2.6 – 3.05) [2.68]: 1.7 (2.04 – 2.42) [2.25]: 0.93 (1.19 – 1.42) [1.26]: 1.0 (1.0) [1.0]; tarsus I – II with 9 (ω) – 8 (ω); femora I – IV with 4 – 3 – 3 – 2.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702C8D0EE2AD94CDDA3CAA7E.taxon	description	Female (n = 4). Length of body (excluding gnathosoma) 320 (260 – 330) [281], width 220 (170 – 250) [203]. Gnathosoma (Figs. 6, 7). Gnathosoma 90 (90 – 96) [92] long (from base of subcapitulum to tip of palp) and 70 (65 – 67) [66] wide. Peritreme with one loop (Fig. 6). Subcapitulum with setae m 28 (25 – 30) [27] and two pairs of adoral setae (or 1 – 2), or 1 7 (7) [7], or 2 8 (7 – 10) [8]; m-m 29 (25 – 30) [28] (Fig. 7). Palpi five segmented with following setal distribution (Fig. 7): tarsus with one eupathidium, two simple setae, one small solenidion; tibia with three smooth setae and one long claw 17 (18 – 21) [20]; genu with one long and slender seta 40 (34 – 42) [39]; femur with two serrated setae 18 (15 – 20) [18], 33 (33 – 38) [37]; chelicerae fused 27 (27 – 30) [29] long. Palp coxa with one supra-coxal seta 3 (3) [3] long (Fig. 7). Dorsum (Figs. 1 – 3). Dorsal idiosoma region with 14 pairs of long and thick serrated setae set on tubercles (Figs. 1 – 3). Length of dorsal setae as follows: vi 88 (83 – 90) [86], ve 51 (45 – 59) [54], sci 41 (58 – 62) [61], sce 37 (35 – 41) [38], c 1 190 (183 – 208) [193], c 2 71 (50 – 80) [64], d 1 170 (145 – 170) [162], d 2 53 (51 – 60) [54], e 1 93 (85 – 99) [91], e 2 50 (48 – 59) [52], f 1 100 (60 – 83) [72], f 2 45 (41 – 50) [46], h 1 50 (44 – 48) [47], h 2 40 (38 – 42) [40]. Distances between setae: vi-vi 38 (30 – 40) [35], ve – ve 108 (95 – 110) [102], vi – ve 41 (20 – 40) [33], ve-sci 19 (15 – 20) [18], scisci 135 (120 – 160) [140], sce – sce 185 (155 – 200) [182], sci – sce 58 (46 – 61) [53], c 1 - c 1 15 (20 – 28) [24], c 1 - c 2 95 (75 – 110) [95], c 2 – c 2 215 (160 – 215) [194], c 1 – d 1 70 (40 – 70) [57], c 2 - d 2 50 (45 – 56) [52], d 1 – d 1 25 (18 – 20) [19], d 1 – d 2 85 (60 – 100) [86], d 2 – d 2 195 (150 – 192) [174], d 1 – e 1 38 (42 – 52) [48], d 2 – e 2 60 (50 – 58) [54], e 1 - e 1 20 (25 – 30) [28], e 1 – e 2 80 (50 – 90) [73], e 2 – e 2 145 (110 – 160) [137], e 1 – f 1 62 (40 – 65) [55], e 2 – f 2 45 (18 – 53) [27], f 1 – f 1 20 (20 – 21) [20], f 1 – f 2 50 (30 – 60) [47], f 2 – f 2 120 (85 – 115) [104], f 1 – h 1 50 (30 – 60) [44], f 2 – h 2 30 (20 – 40) [30], h 1 - h 1 17 (15 – 26) [21], h 1 – h 2 19 (15 – 20) [18], h 2 – h 2 60 (52 – 63) [58]. Seta c 1 the longest and seta sce the smallest. The ratio of dorsal setae vi / c 1 0.46 (0.43 – 0.45) [0.45]; c 1 / d 1 1.12 (1.22 – 1.26) [1.2]; d 1 / e 1 1.83 (1.70 – 1.71) [1.78]; e 1 / f 1 0.93 (1.19 – 1.42) [1.26]; f 1 / h 1 2.0 (1.36 – 1.73) [1.53]; vi / ve 1.72 (1.52 – 1.84) [1.59]; sci / sce 1.1 (1.51 – 1.66) [1.60]; c 1 / c 2 2.67 (2.6 – 3.66) [3.01]; d 1 / d 2 3.2 (2.83 – 2.84) [3.0]; e 1 / e 2 1.86 (1.68 – 1.77) [1.75]; f 1 / f 2 2.22 (1.46 – 1.66) [1.57]; h 1 / h 2 1.25 (1.14 – 1.16) [1.18]; c 1: d 1: e 1: f 1 as follows: 1.9 (2.6 – 3.05) [2.68]: 1.7 (2.04 – 2.42) [2.25]: 0.93 (1.19 – 1.42) [1.26]: 1.0 (1.0) [1.0]. Two pairs of eyes above seta sce, 8 (7 – 8) [8] and 14 (13 – 14) [13] diameter (Fig. 1). Venter (Figs. 4, 5, 8). Ventral surface striated, coxae I – IV with smooth striation, coxal setae stout and serrate, ventral setae 1 a, 3 a and 4 a slender and slightly serrate (1 a set on coxa I). Endopodal shields absent. Anogenital area with one pair of aggenital setae (ag), one pair of genital seta (g) and three pairs of small and smooth anal setae (ps 1 – 3). Length of ventral setae as follows: 1 a 30 (25 – 33) [29], 1 b 30 (20 – 33) [28], 1 c 55 (44 – 65) [52], 2 b 41 (35 – 48) [40], 3 a 30 (33 – 35) [34], 3 b 50 (45 – 56) [50], 3 c 40 (31 – 46) [37], 4 a 29 (27 – 31) [29], 4 b 18 (16 – 20) [19], 4 c 28 (24 – 27) [25], ag 19 (17 – 18) [18], g 15 (11) [11], ps 1 15 (14 – 16) [15], ps 2 11 (12 – 17) [15], ps 3 9 (10 – 15) [12] (Figs. 4, 5, 8). Legs (Figs. 9 – 16). Measurements of leg I 675 (615 – 643) [633], leg II 525 (535 – 580) [553], leg III 635 (585 – 643) [616], leg IV 755 (675 – 740) [712]. Chaetotaxy of leg segments as follows (solenidia in parentheses and not included in setal counts): coxae 3 – 1 – 2 – 2, trochanters 1 – 1 – 1 – 1, femora 4 – 3 – 3 – 2, genua 1 (κ) – 1 (κ) – 1 – 1, tibiae 9 (φ) – 8 (φ) – 8 (φ) – 7 (φ), tarsi 9 (ω) – 8 (ω) – 7 – 7 (Figs. 9 – 16). Genua I – IV with one long, serrated setae, genual setae III – IV longer than I – II (Figs. 9 – 12). Number of tenent hairs in tarsi I – IV as follows: 7; 8; 4 – 8; 2 – 6. Genual setae I – IV length as follows: 63 (58 – 76) [65] – 133 (132 – 160) [148] – 163 (130 – 173) [183] – 190 (170 – 195) [183]. Solenidia of legs I – IV length as follows: I κ 3 (3) [3], II κ 4 (3) [3], I φ 16 (13 – 15) [14], II φ 12 (10 – 12) [11], III φ 9 (10) [10], IV φ 8 (8 – 12) [10], I ω 8 (9 – 10) [9], II ω 5 (5 – 6) [5]. Coxa I with one supra-coxal seta 4 (4) [4] long (Fig. 8). Male. Unknown.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702C8D0EE2AD94CDDA3CAA7E.taxon	discussion	Remarks. The new species, T. iranensis sp. nov., closely resembles T. hawaiiensis, T. ueckermanni sp. nov. and T. farsiensis sp. nov. in lacking seta pdx and having the same chaetotaxy of genua I – IV and tibia II – IV, however, it differs from T. hawaiiensis in: 1) coxa I with 3 setae and tarsus II 8 (1) in new species, instead of 1 and 9 (1) in T. hawaiiensis; 2) peritreme with one loop in the former while complex in the latter; 3) the smallest dorsal seta sce vs. h 2 in T. hawaiiensis; 4) length of dorsal setae c 1 183 – 208, d 1 145 – 170 and h 2 38 – 42 vs. c 1 130 – 155, d 1 121 – 148 and h 2 23 – 28; 5) the ratio of dorsal setae c 1: d 1: e 1: f 1: 1.9 – 3.05: 1.7 – 2.42: 0.93 – 1.42: 1.0 instead of 2.03 – 2.28: 1.95 – 2.12: 0.96 – 1.04: 1.0. It differs from T. ueckermanni in: 1) the smallest dorsal setae is sce in T. iranensis but h 2 in T. ueckermanni; 2) femur III with 3 setae and tarsi I – II with 9 (ω) – 8 (ω) setae in the former whereas femur III with 2 setae and tarsi I – II with 10 (ω) – 9 (ω) setae in the latter; 3) palp tarsus with two simple setae in the former oppose to one simple seta in the latter; 4) ratio of dorsal setae c 1: d 1: e 1: f 1: 1.9 – 3.05: 1.7 – 2.42: 0.93 – 1.42: 1.0; sci / sce 1.1 – 1.66 in T. iranensis instead of c 1: d 1: e 1: f 1: 2.17 – 2.7: 2.12: 1.1 – 1.19: 1.0; sci / sce 1.92 – 2.0 in T. ueckermanni; 5) ventral surface of coxae I – IV with striation in the former whereas ventral surface of coxae I – IV with smooth reticulations plus striations in the latter. It also differs from T. farsiensis sp. nov. in: 1) the longest and smallest dorsal setae is c 1 and sce respectively in T. iranensis but d 1 and c 1 in T. farsiensis; 2) tarsi I – II with 9 (ω) – 8 (ω) setae in the former whereas 10 (ω) – 9 (ω) in T. farsiensis; 3) seta e 1 extending to over level of e 2 in new species opposed to not reaching e 2 in the second species; 4) ratio of dorsal setae c 1: d 1: e 1: f 1: 1.9 – 3.05: 1.7 – 2.42: 0.93 – 1.42: 1.0; vi / c 1 0.43 – 0.45; c 1 / d 1 1.12 – 1.26; sci / sce 1.1 – 1.66; 2.6 – 3.66; c 1 / c 2 2.6 – 3.66 instead of 0.39 – 0.45: 2.29 – 2.53: 0.97 – 1.14: 1.0; vi / c 1 2.90 – 3.16; c 1 / d 1 0.16 – 0.17; sci / sce 2.14 – 2.43; c 1 / c 2 0.40 – 0.55.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702C8D0EE2AD94CDDA3CAA7E.taxon	etymology	Etymology. This species is named after country of its origin, Iran.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F702C8D0EE2AD94CDDA3CAA7E.taxon	materials_examined	Type materials. Holotype female from soil and rotten leaves under oak tree (Quercus brantii Lindl, Fagaceae), Koohmare Sorkhi region, Fars province, Iran (29 ° 32 ' 05 '' N, 52 ° 12 ' 56 '' E, 1916 m a. s. l.), 0 1 ix 2010. Three paratypes female from the same host and data collection (P 1 – 3) respectively: P 1, Koohmare Sorkhi region, Fars province, Iran (29 ° 28 ' 09 '' N, 52 ° 10 ' 11 '' E, 1769 m a. s. l.), 0 7 - 03 - 2010. P 2, Koohmare Sorkhi region, Fars province, Iran (29 ° 31 ' 40 '' N, 52 ° 12 ' 17 '' E, 1827 m a. s. l.), 0 1 iii 2010. P 3 from soil and rotten leaves under oak tree, Koohmare Sorkhi region, Fars province, Iran (29 ° 28 ' 52 '' N, 52 ° 10 ' 27 '' E, 1652 m a. s. l.), 0 3 i 2010, S. Yazdanpanah. The type materials are preserved as slide mounted specimens and the holotype female and two paratypes are deposited in the Acari Collection of the Department of Plant Protection, Faculty of Agriculture, University of Bu-Ali Sina, Hamedan, Iran and one paratype female will be deposited in the Mites collection of Arachnida, Biosystematic Division, ARC-Plant Protection Research Institute, Pretoria, South Africa.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70278D07E2AD94CDDC2EAA96.taxon	diagnosis	Diagnosis. Dorsal idiosoma without pdx seta; palp tarsus with one simple seta; peritreme with one loop; dorsal seta c 1 190 (180 – 185 [183]; femoral formula I – IV 4 - 3 - 2 - 2. The ratio dorsal setae c 1: d 1: e 1: f 1 as follows: 2.3 (2.17 – 2.7) [2.4]: 2.12 (2.12 – 2.4) [2.28]: 1.1 (1.18 – 1.19) [1.18]: 1.0 (1.0) [1.0].; tarsus I – II with 10 (ω) – 9 (ω).	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70278D07E2AD94CDDC2EAA96.taxon	description	Female (n = 4). Length of body (excluding gnathosoma) 342 (290 – 320) [303], width 280 (230 – 245) [237]. Gnathosoma (Figs. 20 – 22). Gnathosoma 101 (75 – 95) [86] long (from base of subcapitulum to tip of palp) and 79 (68 – 95) [78] wide. Peritreme with one loop (Fig. 20). Chelicerae fused 36 (27 – 28) [28] long (Fig. 20). Palpi five segmented with following setal pattern (Fig. 21): tarsus with one eupathidium, one simple seta, and one small solenidion; tibia with three smooth setae and one long claw 25 (21 – 25) [23] long; genu with one long and slender seta 36 (34 – 35) [35]; femur with two serrated setae 16 (17 – 19) [18], 38 (35 – 38) [37]. Subcapitulum with setae m 25 (24) [24] and two pairs of adoral setae (or 1 – 2), or 1 8 (6 – 8) [7], or 2 10 (8) [8]; m-m 25 (26 – 28) [27] (Fig. 22). Palp coxa with one supra-coxal seta 4 (4) [4] long (Fig. 22). Dorsum (Figs 17 – 19). Dorsal idiosoma region with 14 pairs of long and thick serrated setae set on tubercles (Fig. 17 – 19). Length of dorsal setae as follows: vi 83 (73 – 85) [81], ve 62 (47 – 51) [49], sci 69 (48 – 66) [58], sce 35 (25 – 33) [29], c 1 190 (180 – 185 [183], c 2 64 (50 – 65) [56], d 1 174 (160 – 180) [173], d 2 55 (40 – 44) [42], e 1 90 (80 – 100) [90], e 2 58 (40 – 55) [47], f 1 82 (67 – 85) [76], f 2 42 (43 – 45) [44], h 1 45 (39 – 41) [40], h 2 31 (35 – 39) [37]. Distances between setae: vi-vi 36 (35 – 38) [36], ve – ve 86 (85 – 90) [87], vi – ve 30 (30) [30], ve-sci 43 (35 – 40) [38], sci-sci 145 (130 – 140) [133], sce – sce 210 (180 – 195) [188], sci – sce 60 (51 – 60) [55], c 1 - c 1 20 (20 – 25) [25], c 1 - c 2 105 (104 – 120) [115], c 2 – c 2 240 (200 – 220) [207], c 1 – d 1 41 (50 – 60) [55], c 2 – d 2 71 (61 – 70) [65], d 1 – d 1 33 (27 – 33) [28], d 1 – d 2 98 (77 – 100) [90], d 2 – d 2 200 (151 – 190) [167], d 2 – e 2 65 (55 – 60) [58], d 1 – e 1 47 (40 – 60) [50], e 1 - e 1 27 (20 – 22) [21], e 1 – e 2 80 (80 – 85) [83], e 2 – e 2 150 (130 – 150) [140], e 1 – f 1 50 (55 – 58) [57], e 2 – f 2 42 (26 – 50) [37], f 1 – f 1 15 (15 – 18) [17], f 1 – f 2 52 (45 – 58) [51], f 2 – f 2 110 (94 – 95) [95], f 1 – h 1 50 (44 – 45) [45], f 2 – h 2 30 (25 – 26) [26], h 1 - h 1 10 (11 – 15) [13], h 1 – h 2 16 (16 – 17) [17], h 2 - h 2 50 (50) [50]. Seta c 1 the longest while seta h 2 the smallest. The ratio of dorsal setae as follows: vi / c 1 0.44 (0.41 – 0.46) [0.44]; c 1 / d 1 1.09 (1.03 – 1.13) [1.06]; d 1 / e 1 1.93 (1.80 – 2.0) [1.92]; e 1 / f 1 1.1 (1.17 – 1.19) [1.18]; f 1 / h 1 1.82 (1.72 – 2.07) [1.90]; vi / ve 1.34 (1.55 – 1.67) [1.65]; sci / sce 1.97 (1.92 – 2.0) [2.0]; c 1 / c 2 2.97 (2.84 – 3.6) [3.27]; d 1 / d 2 3.16 (4.0 – 4.01) [4.12]; e 1 / e 2 1.55 (1.81 – 2.0) [1.91]; f 1 / f 2 1.95 (1.56 – 1.89) [1.73]; h 1 / h 2 1.45 (1.05 – 1.11) [1.08]; c 1: d 1: e 1: f 1: 2.3 (2.17 – 2.7) [2.4]: 2.12 (2.12 – 2.4) [2.28]: 1.1 (1.18 – 1.19) [1.18]: 1.0 (1.0) [1.0]. Two pairs of eyes above the seta sce present, 8 (6 – 8) [7] and 15 (11 – 14) [13] diameter (Fig. 17). Venter (Figs. 23 – 25). Ve n tra l sur fac e striated coxae with soft reticulation, coxal setae stout and serrate, ventral setae 1 a, 3 a and 4 a slender and slightly serrate (1 a set on coxa I). Endopodal shields absent. Anogenital area with one pair of aggenital setae (ag), genital valve with one pair of genital seta (g 1) and three pairs of small and smooth anal setae (ps 1 – 3). Length of ventral setae: 1 a 32 (31) [31], 1 b 32 (24 – 29) [27], 1 c 63 (61 – 61) [61], 2 b 45 (39) [39], 3 a 31 (30 – 33) [32], 3 b 44 (34 – 35) [35], 3 c 34 (40) [40], 4 a 25 (23 – 28) [26], 4 b 16 (15 – 18) [17], 4 c 26 (23 – 25) [24], ag 19 (15 – 16) [16], g 15 (15) [15], ps 1 15 (15 – 17) [16], ps 2 15 (15) [15], ps 3 16 (14 – 16) [15]. (Figs. 23 – 25). Legs (Figs. 26 – 33). Measurements of leg I 658 (635 – 645) [640], leg II 570 (513 – 537) [525], leg III 628 (580 – 583) [582], leg IV 719 (665 – 667) [666]. Chaetotaxy of leg segments as follows (solenidia in parentheses and not included in setal counts): coxae 3 – 1 – 2 – 2, trochanters 1 – 1 – 1 – 1, femora 4 – 3 – 2 – 2, genua 1 (κ) – 1 (κ) – 1 – 1, tibiae 9 (φ) – 8 (φ) – 7 (φ) – 7 (φ), tarsi 10 (ω) – 9 (ω) – 7 – 7 (Figs. 26 – 33). Number of tenent hairs in tarsi I – IV as follows: 6 – 8; 9; 6 – 8; 2 – 6 (Figs. 30 – 33). Genua I – IV with one long, serrated seta, genual setae III – IV longer than I – II. Genual setae length as follows: I – IV 74 (60 – 65) [62] – 133 (115 – 125) [119] – 150 (128 – 155) [142] – 163 (150 – 158) [154] (Figs. 26 – 29). Solenidia of legs I – IV length as follows: I κ 3 (3) [3], II κ 3 (3 – 4) [4], I φ 20 (18 – 19) [19], II φ 14 (10 – 12) [11], III φ 8 (11 – 12) [12], IV φ 9 (10 – 12) [11], I ω 11 (8 – 11) [10], II ω 7 (5 – 7) [6]. Coxa I with one supra-coxal seta 4 (3 – 4) [4] long (Fig. 25). Male. Unknown.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70278D07E2AD94CDDC2EAA96.taxon	discussion	Remarks. Tycherobius ueckermanni sp. nov. is similar to T. superbus (Canestrini, 1889) in having two setae on femur III however differs from the latter by: 1) femur I with four setae in T. ueckermanni whereas three in T. superbus; 2) length of dorsal setae c 1 180 – 190 and d 1 160 – 180 in the former vs. 140 and 115 in the latter. The new species also resembles T. farsiensis sp. nov. in lacking dorsal setae pdx and the same tibial setae and tarsal chaetotaxy but can be distinguished from the latter by: 1) femur III with two setae in T. ueckermanni sp. nov. instead of three in T. farsiensis; 2) palp tarsus with one simple seta (plus one eupathidion, one ω) in the former whereas two simple setae (plus one eupathidion, one ω) in the latter; 3) the longest and smallest dorsal setae c 1 and h 2 respectively in T. ueckermanni instead of d 1 and c 1 in T. farsiensis; 4) length of dorsal seta c 1 180 – 190 whereas 27 – 31 in the latter; 5) ventral surface of coxae I – IV with soft reticulation plus striation in the new species opposed to without reticulation pattern in T. farsiensis; 6) the ratio dorsal setae as follows: c 1: d 1: e 1: f 1 2.17 – 2.7: 2.12 – 2.4: 1.10 – 1.19: 1.0; vi / c 1 0.41 – 0.46; c 1 / c 2 2.84 – 3.6 versus 0.39 – 0.45: 2.29 – 2.53: 0.97 – 1.14: 1.0; vi / c 1 2.90 – 3.16; c 1 / c 2 0.40 – 0.55.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70278D07E2AD94CDDC2EAA96.taxon	etymology	Etymology. This species is named in honor of Prof. Edward A. Ueckermann, who kindly assisted the senior author through excellent collaborations and has helped other Iranian Acarologists in last two decades.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70278D07E2AD94CDDC2EAA96.taxon	materials_examined	Type materials. Holotype from soil and rotten leaves under oak tree (Quercus brantii Lindl, Fagaceae), Koohmare Sorkhi region, Fars province, Iran (29 ° 28 ' 41 '' N, 52 ° 10 ' 08 '' E, 1688 m a. s. l.), 31 xii 2010. Three paratypes (P 1 – 3) from same host respectively: P 1, Koohmare Sorkhi region, Fars province, Iran (29 ° 32 ' 05 '' N, 52 ° 12 ' 56 '' E, 1916 m a. s. l.), 0 1 ix 2010. P 2 from soil & rotten leaves under oak tree, Koohmare Sorkhi region, Fars province, Iran (29 ° 31 ' 41 '' N, 52 ° 12 ' 24 '' E, 1843 m a. s. l.), 0 4 x 2010. P 3, Koohmare Sorkhi region, Fars province, Iran (29 ° 31 ' 40 '' N, 52 ° 12 ' 17 '' E, 1827 m a. s. l.), 0 1 iii 2010, S. Yazdanpanah. The type materials are preserved as slide mounted specimens and the holotype female and two paratypes are deposited in the Acari Collection of the Department of Plant Protection, Faculty of Agriculture, University of Bu-Ali Sina, Hamedan, Iran and one paratype female will be deposited in the Mite Collection of Arachnida, Biosystematic Division, ARC-Plant Protection Research Institute, Pretoria, South Africa.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70238D1BE2AD94CDDC2EA896.taxon	diagnosis	Diagnosis. Dorsal idiosoma without pdx seta; palp tarsus with two simple seta; peritreme with one loop; dorsal seta c 1 31 (27 – 31) [29]; The ratio dorsal setae c 1: d 1: e 1: f 1 as follows: 0.45 (0.39) [0.39]: 2.53 (2.29 – 2.4) [2.36]: 0.97 (1.06 – 1.14) [1.11]: 1.0 (1.0) [1.0]; tarsus I – II with 10 (ω) – 9 (ω); femora I – IV with 4 – 3 – 3 – 2.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70238D1BE2AD94CDDC2EA896.taxon	description	Female (n = 4). Length of body (excluding gnathosoma) 350 (300 – 360) [330], width 282 (220 – 290) [258]. Gnathosoma (Figs. 37 – 38). Gnathosoma 98 (100) [100] long (from base of infracapitulum to tip of palp) and 78 (86) [86] wide. Peritreme with one loop (Fig. 37); chelicerae length 42 (32 – 42) [32]. Infracapitulum with setae m 25 (25) [25] and two pairs of adoral setae (or 1 – 2), or 1 7 (7) [7], or 2 7 (6 – 7) [6]; m-m 28 (25 – 28) [25] (Fig. 38). Palpi five segmented with following setal pattern (Fig. 38): tarsus with one eupathidium, two simple setae, one small solenidion; tibia with three smooth setae and one long claw 22 (20 – 25) [23]; genu with one long, slender seta 40 (38 – 41) [40]; femur with two serrated setae 20 (20 – 25) [22], 40 (33 – 42) [38]. Palp coxa with one supra – coxal seta 4 (4) [4] long (Fig. 38). Dorsum (Figs. 34 – 36). Dorsal idiosoma region with 14 pairs of long and thick serrated setae set on tubercles (Figs 34 – 36). Length of dorsal setae as follows: vi 90 (83 – 98) [92], ve 42 (49 – 50) [49], sci 63 (73 – 75) [74], sce 28 (30 – 35) [32], c 1 31 (27 – 31) [29], c 2 56 (59 – 77) [70], d 1 175 (168 – 183) [177], d 2 48 (46 – 50) [49], e 1 67 (80 – 85) [83], e 2 45 (50) [50], f 1 69 (70 – 80) [75], f 2 40 (40 – 48) [45], h 1 41 (37 – 45) [41], h 2 38 (38 – 42) [40]. Distances between setae: vi-vi 35 (40 – 65) [48], ve – ve 95 (85 – 95) [89], vi – ve 37 (30 – 39) [33], ve-sci 49 (44 – 53) [48], sci-sci 167 (130 – 167) [140], sce – sce 167 (130 – 167) [140], sci – sce 59 (53 – 60) [56], c 1 - c 1 20 (16 – 20) [17], c 1 - c 2 120 (119 – 120) [120], c 2 – c 2 235 (220 – 245) [235], c 1 – d 1 45 (50 – 60) [57], c 2 - d 2 70 (63 – 75) [71], d 1 – d 1 30 (20 – 30) [26], d 1 – d 2 109 (95 – 113) [103], d 2 – d 2 181 (187 – 200) [193], d 1 – e 1 70 (65 – 70) [67], d 2 – e 2 54 (57 – 62) [60], e 1 - e 1 27 (18 – 27) [22], e 1 – e 2 81 (85 – 95) [90], e 2 – e 2 132 (135 – 155) [142], e 1 – f 1 44 (38 – 55) [47], e 2 – f 2 34 (28 – 44) [36], f 1 – f 1 20 (20 – 22) [21], f 1 – f 2 65 (65 – 67) [66], f 2 – f 2 98 (90 – 115) [104], f 1 – h 1 53 (55 – 65) [60], f 2 – h 2 23 (22 – 31) [26], h 1 - h 1 12 (15 – 20) [17], h 1 – h 2 16 (16 – 20) [17], h 2 – h 2 52 (57 – 60) [58]. Seta d 1 the longest while seta c 1 the smallest. The ratio of dorsal setae as follows: vi / c 1 2.90 (3.07 – 3.16) [3.17]; c 1 / d 1 0.17 (0.16 – 0.17) [0.16]; d 1 / e 1 2.16 (2.1 – 2.2) [2.13]; e 1 / f 1 0.97 (1.14 – 1.06) [1.11]; f 1 / h 1 1.68 (1.78 – 1.9) [1.83]; vi / ve 2.14 (1.7 – 1.96) [1.88]; sci / sce 2.25 (2.14 – 2.43) [2.31]; c 1 / c 2 0.55 (0.40 – 0.46) [0.41]; d 1 / d 2 3.65 (3.5 – 3.66) [3.61]; e 1 / e 2 1.49 (1.6 – 1.7) [1.67]; f 1 / f 2 1.73 (1.67 – 1.75) [1.67]; h 1 / h 2 1.08 (0.97 – 1.07) [1.03]; c 1: d 1: e 1: f 1 0.45 (0.39) [0.39]: 2.53 (2.29 – 2.4) [2.36]: 0.97 (1.06 – 1.14) [1.11]: 1.0 (1.0) [1.0]. Two pairs of eyes above seta sce, 8 (8 – 9) [9] and 13 (12 – 13) [12] diameter (Fig. 34). Venter (Figs. 39 – 41). Ventral surface striated, coxae with smooth striations, coxal setae stout and serrate, ventral setae 1 a, 3 a and 4 a slender and slightly serrate (1 a set on coxa I). Endopodal shields absent. Anogenital area with one pair of aggenital setae (ag), with one pair of genital setae (g) and three pairs of small and smooth anal setae (ps 1 – 3). Length of ventral setae: 1 a 28 (29 – 33) [31], 1 b 33 (43) [43], 1 c 78 (77 – 78) [77], 2 b 50 (49 – 50) [49], 3 a 25 (30) [30], 3 b 55 (45 – 55) [46], 3 c 45 (59) [59], 4 a 22 (22) [22], 4 b 24 (20 – 24) [21], 4 c 24 (25) [25], ag 17 (18 – 20) [19], g 15 (15) [15], ps 1 15 (15 – 20) [18], ps 2 18 (15 – 18) [16], ps 3 17 (15 – 17) [16] (Figs. 39 – 41). Legs (Figs. 42 – 49). Measurements of leg I 759 (770 – 775) [773], leg II 620 (620 – 658) [639], leg III 670 (680 – 723) [702], leg IV 820 (805 – 848) [827]. Chaetotaxy of leg segments as follows (solenidia in parentheses and not included in setal counts): coxae 3 – 1 – 2 – 2, trochanters 1 – 1 – 1 – 1, femora 4 – 3 – 3 – 2, genua 1 (κ) – 1 (κ) – 1 – 1, tibiae 9 (φ) – 8 (φ) – 7 (φ) – 7 (φ), tarsi 10 (ω) – 9 (ω) – 7 – 7 (Figs. 42 – 49). Number of tenent hairs in tarsi I – IV as follows: 8; 7; 8; 7 (Figs. 46 – 49). Genua I – IV with one long, serrated setae, genual setae III – IV longer than I – II (Figs. 42 – 45). Genual setae I – IV length as follows: 69 (74 – 92) [83] – 113 (122 – 125) [124] – 135 (144 – 146) [145] – 175 (160 – 195) [176]. Solenidia of legs I – IV length as follows: I κ 4 (3 – 4) [3], II κ 3 (3 – 4) [3], I φ 20 (13 – 20) [15], II φ 10 (8 – 11) [9], III φ 10 (10) [10], IV φ 10 (8 – 10) [9], I ω 11 (11 – 12) [12], II ω 8 (7 – 8) [7]. Coxa I with one supra-coxal seta 4 (3 – 4) [4] long (Fig. 41). Male. Unknown.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70238D1BE2AD94CDDC2EA896.taxon	discussion	Remarks. The new species T. farsiensis closely resembles T. polonicus Bolland, 1986 in having the same chaetotaxy of legs I – IV (except tarsus II) and lacking dorsal setae pdx. However, it differs from the latter by: 1) tarsus II with 9 (ω) in the former but 10 (ω) in the latter; 2) peritreme with one loop in T. farsiensis whereas two loops in T. polonicus; 3) dorsal setae: c 1 27 – 31, d 1 168 – 183 and d 2 46 – 50 versus 235, 205 and 95; 4) the ratio of dorsal setae c 1: d 1: e 1: f 1 0.39 – 0.45: 2.29 – 2.53: 0.97 – 1.14: 1.0 versus 2.8: 2.4: 0.71: 1.0; 5) the longest and smallest dorsal setae d 1 and c 1 respectively in T. farsiensis opposed to c 1 and sce, f 2, h 2 in T. polonicus.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70238D1BE2AD94CDDC2EA896.taxon	etymology	Etymology. This species is named locality name, Fars province.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
03EC132F70238D1BE2AD94CDDC2EA896.taxon	materials_examined	Type material. Holotype from soil & rotten leaves under oak tree (Quercus brantii Lindl, Fagaceae), Koohmare Sorkhi region, Fars province, Iran (29 ° 30 ' 09 '' N, 52 ° 10 ' 20 '' E, 1717 m a. s. l.), 31 xi 2010. Three paratypes (P 1 – 3) from same host, respectively: P 1 from soil & rotten leaves under oak tree, Koohmare Sorkhi region, Fars province, Iran (29 ° 29 ' 26 '' N, 52 ° 11 ' 01 '' E, 1636 m a. s. l.), 16 xii 2010. P 2, Koohmare Sorkhi region, Fars province, Iran (29 ° 28 ' 09 '' N, 52 ° 10 ' 11 '' E, 1769 m a. s. l.), 0 7 iii 2010. P 3, Koohmare Sorkhi region, Fars province, Iran (29 ° 28 ' 52 '' N, 52 ° 10 ' 27 '' E, 1652 m a. s. l.), 0 3 xi 2010, S. Yazdanpanah. The type materials are preserved as slide mounted specimens and the holotype female and two paratypes are deposited in the Acari Collection of the Department of Plant Protection, Faculty of Agriculture, University of Bu-Ali Sina, Hamedan, Iran and one paratype female will be deposited in the mite Collection of Arachnida, Biosystematic Division, ARC-Plant Protection Research Institute, Pretoria, South Africa.	en	Khanjani, Mohammad, Yazyanpanah, Shima, Ostovan, Hadi, Fayaz, Bahman Asali (2012): Three new species of the genus Tycherobius Bolland (Acari: Camerobiidae) from Iran. Zootaxa 3266: 23-40, DOI: 10.5281/zenodo.280753
