taxonID	type	description	language	source
03ED87B7CB771B2DF967FEB7FD93FC6B.taxon	description	(Figs. 2 – 15)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB771B2DF967FEB7FD93FC6B.taxon	diagnosis	Diagnosis. Australian species of the tribe Macrotomini can be separated from all other Australian Prioninae by the following characters: eyes shallowly or not emarginate near antennal insertions (Fig. 13 M); mesocoxal cavities open to mesanepisternum and mesepimeron (Fig. 4 D); mesotrochantin obviously exposed (Fig. 4 D); metanepisternum broad; metepimeron narrow; pronotum with lateral carina dentate.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB771B2AF969FC4AFE4CF952.taxon	description	(Figs. 2, 14 A, 15 D, 16 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB771B2AF969FC4AFE4CF952.taxon	diagnosis	Diagnostic combination (male). Large beetles, 30 – 70 mm long. Dorsum bicoloured, head and pronotum darker than elytra. Sexually dimorphic dense punctures (Fig. 2 B) present on most of the pronotum, all thoracic sterna and at least on anterior parts of abdominal ventrites. Head distinctly narrower than prothorax; frontoclypeal suture angulate; median groove complete. Antennal tubercles prominent, round- ed apically. Eyes very large, weakly emarginate near antennal insertion, narrowly separated dorsally. Mandibles less than half of head length, not sexually dimorphic, strongly bent and unidentate apically. Antenna almost reaching elytral apex in male. Scape longer than half of head length, posteriorly extending to anterior margin of pronotum, expanding apically, flat with sharp external edge; antennomere 3 about as long as scape and almost twice as long as 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps weakly expanded and apically truncate. Prothorax transverse with lateral carina regularly dentate; anterior and posterior margins with complete beads. Pronotal disc uneven with weakly elevated smooth and coarsely punctate callosities on densely punctate, feebly shiny background. Prosternal process subparallel, extending beyond procoxae, rounded apically. Elytral surfaces weakly coriaceous; inner apical angle with short spine. Legs relatively long; femora and tibiae with rows of sharp spines. Protibia with sharp apical spine and pair of subequal spurs; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB771B2AF969FC4AFE4CF952.taxon	description	Description. Male. Length 30 – 70 mm. Head and pronotum reddish or dark brown, elytra lighter than pronotum often yellowish and semitransparent. Head approximately as long as wide, narrower than pronotum. Mandibles shorter than half of head length, moderate to strongly curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base; unidentate at apex, with two extra teeth on incisor edge. Labrum weakly constricted at base, anterior margin angulate or pointed medially with very dense setae; labrum separated from clypeus by a deep groove; dorsal surface of labrum and clypeus covered with long golden setae. Frontoclypeal suture angulate; median groove distinct and complete. Antennal tubercles prominent, apex rounded. Antenna 11 - segmented, filiform, extending almost to elytral apices; scape long and flat, reaching beyond anterior margin of pronotum; antennomere 3 as long as or slightly shorter than scape, almost twice as long as antennomere 4; antennomeres 5 – 11 with blunt posterior angles. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; rather narrowly separated dorsally. Submentum weakly curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps weakly expanded and apically truncate. Pronotum transverse, trapezoid with posteri- or margin broader than anterior margin; anterior margin emarginate medially with complete bead; posterior margin with complete bead; lateral margin with rather regular small sharp teeth. Disc surface very densely punctate and matt except for shiny, coarsely and deeply punctate islands consisting of two triangular median areas, two smaller lateral areas and a transverse area in front of the posterior margin. Prosternum covered with fine and deep punctures; hypomeron well-defined, as broad as prosternal process; prosternal process projecting towards mesoventrite, rounded apically. Mesoventral process emarginate at apex. Ventral side of thorax and coxae covered with golden hairs. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface very finely punctate. Elytral surfaces dull covered with irregular coriaceous sculpture; two weak traces of venation often present on basal half of elytra; elytral apices rounded with short sutural spine; epipleuron almost complete, narrowing apically. Legs strong and relatively long; femur sub-parallel, with two rows of sharp spines on ventral side; tibia slender, with spines along outer margin, bearing sensory setae on ventral side apically; each tibia with distinct apical spine and a pair of spurs; tarsi strong; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1 – 4 densely punctate basally with smooth and apparently less sclerotised apical part; ventrite 5 smooth; ventral surface with very sparse setae, but bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 A). Tegmen longer than sternite VII; parameres moderately long, 0.16 times length of tegmen, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex narrowly rounded. Female. Antennae usually shorter than in male, only extending to middle of elytra. Pronotal disc shiny, surfaces irregular with very deep and coarse punctures; fine and dense punctures on thorax and abdomen absent. Abdominal ventrite 5 truncate or slightly emarginate at apex. Ovipositor (Fig. 15 D) long, apical sclerotised part ñ length of baculus; distal gonocoxites relatively short and stout; stylus moderately long, inserted laterally and very close to gonocoxite apex, gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB771B2AF969FC4AFE4CF952.taxon	discussion	Remarks. The genus Agrianome can be distinguished from the remaining genera of the Australian Macrotomini by the large body, pale yellow and slightly transparent elytra with apicosutural angle bearing a long spine, and the antennal scape as long as antennomere 3, and almost twice the length of antennomere 4. Four species have been recognised in the genus Agrianome. The type species, A. fairmairei (Montrouzier, 1861) was described from New Caledonia, and was later recorded from other islands nearby. A. loriae Gestro, 1893 was described from Papua New Guinea, and Gressitt (1951) reported two specimens from northern Queensland, Australia. A. spinicollis (Macleay, 1826) was described from intertropical and western coast of Australia, but it was subsequently record- ed from Queensland and New South Wales. A. howei (Olliff, 1889) stat. nov. originally described from Lord Howe Island was regarded as a subspecies of A. spinicollis since Lameere (1912) and our study confirmed its species level status based on both molecular and morphological evidence.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB701B2BFB39FDA6FEEEFD12.taxon	description	(Figs. 2 A – D, 16 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB701B2BFB39FDA6FEEEFD12.taxon	diagnosis	Diagnosis. A. howei can be separated from other Agrianome species by the antennomere 3 as long as or slightly shorter than antennomeres 4 and 5 combined; pronotum with strongly emarginate anterior margin; protibia without lateral spines and being geographically restricted to the Lord Howe Island. Notes. When reviewing the insect fauna of the Lord Howe Island, Olliff (1889) found a large Prioninae species he assumed to be Arimaspes howei Thomson (currently in Hermerius) described from that island. Based on the locality alone Olliff (1889) catalogued this species as Cnemoplites howei (Thomson), and commented the specimens he had at hand were apparently similar to Agrianome gemella Pascoe. After examination of the type specimens of Arimaspes howei Thomson, Agrianome gemella Pascoe and available specimens from Lord Howe Island, we found the following: (a) all Lord Howe Island Macrotomini belong to a single species of Agrianome; (b) Arimaspes howei Thomson belongs to Hermerius and its type locality of Lord Howe Island is most probably wrong; (c) Olliff (1889) had specimens of Agrianome from Lord Howe Island that he misidentified as Arimaspes howei Thomson, and his comments on his specimens were diagnostic enough to treat them as species description. However, the specimens Olliff examined at that time (obtained by Mr. Etheridge’s party) were probably lost (D. Smith, pers. comm.) and a neotype of Cnemoplites howei Olliff is therefore designated here to stabilise the taxonomic status of the species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB701B2BFB39FDA6FEEEFD12.taxon	materials_examined	Type. Cnemoplites howei Olliff, neotype here designated: a male specimen deposited in ANIC with the following labels: “ Lord Howe Island, 1957, I. Liepa | ANIC Database 25 - 039504 ” (genitalia were dissected and stored separately, and the right hind coxa was used for DNA extraction). Material examined (Fig. 16 A). Lord Howe Island: AM (17 ♂♂; 13 ♀♀): 9: Dec. 2002, Ushijima and Chikakura; 5: A. M. Lea; 3: Research Centre, Dec. 2000, C. Reid; 2: Research Centre, Nov. 2000, C. Reid; Apr. 1979, T. Kingston; Blackburn Island near Lord Howe Island, Feb. 2001, S. Fellenberg and M. Humphrey; Coral Coult, Jan. 1979, T. Kingston; Feb. 1921, A. R. McCulloch; Feb. 1922, A. R. McCulloch; Jan. 1972, Lagoon Rd., Mar. 1979, T. Kingston; Paddock behind Sea Breege, Jan. 1979, T. Kingston; Rabbit Island, Jan. 1979, T. Kingston; Research Centre, Feb. 2017, Stahon; Sea Breege, Feb. 1979, T. Kingston; ANIC (5 ♂♂; 5 ♀♀): 2: Jan. 1985, G. Bornemissza; 2: Salmon Beach, Dec. 1972, Z. Liepa; A. M. Lea; Dec. 1955, S. J. Paramonov; Dec. 1981, P. Macnicol; Jan. - Dec. 1960, L. Moore; Nov. 1956, Z. Liepa; Stevens Reserve, nr. Signal Point, Dec. 1988, D. C. F. Rentz; MAMU (4 ♂♂; 2 ♀♀): 6: [no data]; MV (3 ♂♂; 1 ♀): 4: [no data]. Molecular data. BioSample ID: SAMN 12650064 (Neotype).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B2BF956FCB7FEB3F976.taxon	description	(Figs. 2 E – H)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B2BF956FCB7FEB3F976.taxon	diagnosis	Diagnosis. A. loriae can be separated from other Agrianome species by antennomeres 3 - 7 having acute posterior angles; antennomere 3 being extremely long, longer than antennomeres 4 and 5 combined, and extending beyond the humeral angle of the elytra and with the ventral surface bearing dense spines (Fig. 2 G). Notes. Gressitt (1951) recorded two specimens of A. loriae from North Queensland. We were not able to examine his specimens to assess the correctness of Gressitt’s identification and we consider this record as being dubious, as we found no specimens to be A. loriae in all the Australian Agrianome specimens we have examined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B2BF956FCB7FEB3F976.taxon	materials_examined	Material examined (Fig. 16 A). Papua New Guinea: ANIC (8 ♂♂; 1 ♀): 2: Wau, Nov. 1978, [H. Ohlmus Coll.]; Mandang, Oct. 1972, [H. Ohlmus Coll.]; Mt. Sinewit, Jun. – Sep. 1963, W. W. Brandt; Wau, Aug. 1970, [H. Ohlmus Coll.]; Wau, Dec. 1969, [H. Ohlmus Coll.]; Wau, Dec. 1970, [H. Ohlmus Coll.]; Wau, Feb. 1970, [H. Ohlmus Coll.]; Wau, May 1971, [H. Ohlmus Coll.]; BNHM (1 ♂): Wau Ecology Institute, Jul. 1990, G. W. Becealoni; QM (2 ♂♂): Bulolo Unitech Morobe Province, Jul. 1999, P. Bouchard; Tikling, nr Buso Riv- er, 20 km E of Lae, Jul. 1999, D. Yeates, P. Bouchard and S. Winterton.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B29F8A0F948FDE9FBC8.taxon	description	(Figs. 2 I – K, 14 A, 15 D, 16 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B29F8A0F948FDE9FBC8.taxon	diagnosis	Diagnosis. A. spinicollis can be separated from A. loriae by antennomeres 3 - 7 having blunt posterior angles; antennomere 3 being shorter than antennomeres 4 and 5 combined, and only extending to the posterior margin of the pronotum and mostly with a smooth ventral surface. It can be separated from A. howei by the anterior margin of the pronotum being weakly emarginate, and the protibia has dense lateral spines.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB711B29F8A0F948FDE9FBC8.taxon	materials_examined	Types. Prionus spinicollis Macleay Lectotype designated here: female specimen deposited in MAMU, with the following original labels “ Prionus spinicollis, New Holland, Capt. King | MAMU EN. 30675 ”. Macrotoma gemella Pascoe Holotype male deposited in BNHM (BNHM- 010799857), with the following original labels “ Macrotoma gemella Pasc. type | Pascoe Coll. 93 - 60 | Agrianome gemella Pasc ”. Material examined (Fig. 16 A). New South Wales: ANIC (63 ♂♂; 49 ♀♀): 13: Lower South Portland, Dec. 1976, D. P. Carne; 9: Wingham (Lots 11 and 12 Combined St), {from Dec. 1990 to Jan. 1992}, J. Stockard; 5: Cambridge Plateau, Jan. 1996, J. Bugeja; 4: Pennefathers Rd, Cherry Tree State Forest via Casino, {from Dec. 1996 to Jan. 1997}, S. Watkins; 3: 7 mi. W of Wisemans Ferry, Dec. 1974, D. P. Carne; 3: Binnyguy via Moree [Biniguy], Oct. 1987, R. Delves; 3: Murrurundi, {from Nov. to Dec. 1947}, B. L. Middleton; 2: Armidale, Jan. 1966, E. T. Giles; 2: Batemans Bay, Apr. 1976, R. S. McInnes; 2: Cambridge Plateau, Richmond Range SF, WNW Casino, Jan. - Feb. 1997, S. Watkins; 2: Mitchell Lab., Trangie, Apr. 1953, R. E. Cameron; 2: Pennefathers Rd., CherryTree SF, SE Mallanganee, Dec. 1997, Watkins; 2: Roote Hill [Rooty Hill], J. L. C. Lawson; 2: Sydney, Apr. 1905, W. W. Froggatt; 2: Wauchope, c. 72 km W on Oxley Highway, Jan. 1970, Britton, Holloway and Misko; 2: Woodburn, Jan. 1986, G. O’Reilly; 3 km W Araluen, Jan. 1974, J. R. Hosking; 4 k N Batlow, Dec. 1977, R. Sutherland; Allyn R. F. Pk. [Allyn River Forest Park], Jan. 1986, R. B. Halliday; Armidale, Feb. 1964, F. Herbert; Avoca Beach, N Gosford, Dec. 1972, A. Morton; Balmain, Dec. 1993, J. M. Disney; Batemans Bay, Aug. 1962, D. P. Carne; Bonville, Dec. 1975, P. B. Carne; Bonville, Jan. 1970, A. T. Dunn; Border Ranges Nat. Pk., Forest Tops camp, Feb. 1999, M. Neave, E. D. Edwards, M. Yee and B. Edmonds; Cabbage Tree Island, Jan. 1969, J. McKean; Caparra (Lots 72, 73, 148), Dec. 1994, S. Watkins; Caparra (Lots 72, 73, 148), Jan. 1993; Caparra (Lots 72, 73, 148), Jan. 1995; Casino, Dec. 1971, Key and Balderson; Coolatai Station, Dec. 1966, T. G. Campbell; Crosslands via Hornsby, Dec. 1976, D. P. Carne; Dorrigo N. P. via Dorrigo, Mar. 1955; Exp. Station, Trangie, Nov. 1946, L. J. Chinnick; Findon Ck., 27 km E of Woodenbong, Nov. 1990, D. C. F. Rentz and K. McCarron; Forbes, Nov. 1979, L. P. Kelsey; Georges River, Pleasure Point, Jan. 1995, A. Sundholm and R. de Keyzer; Gocup, Tumut, Feb. 1977; Gosford, Jan. 1911, Jenkins; Grafton, Dec. 1955, F. S. P.; Homebush, Apr. 1905; Ingleside, Katandra Bushland Sanct., Oct. 1998, G. Hangay; Kangaroo Valley, Feb. 1992, F. Prichard; Leeton, Dec. 1970, P. Atkinson; Leets Vale, Dec. 1975, D. P. Carne; Lismore, W. W. Froggatt; Maimuru, 6 km SSE of Young, Dec. 1993, J. Worner; Moruya River, Jan. 1930; Mt. Keira Base, nr. Wollongong, Jan. 1985, J. E. Feehan; Mt. Keira scout camp, Mar. 1981, J. F. Lawrence and A. Calder; Mt. Keira, Mar. 1967, E. B. Britton and S. Misko; Murwillumbah, Dec. 1973, G. May; Narrabeen, Jan. 2003, G. and K. Hangay; Narrabeen, Nov. 2000, G. Hangay, I. Rozner, A. Podlussany and K. Hangay; Park College, Sydney, Apr. 1905, W. W. Froggatt; Port Macquarie, Jan. 1970, K. H. L. Key; Pumpenbil, Nov. 1977, G. May; Pumpenbil, Upper Tweed R. [Pumpenbil], Mar. 1978, G. May; Singleton, Dec. 1945, P. B. Carne and E. B. Britton; Singleton, Dec. 1948, K. H. I. and B. Key; Temora, Feb. 1923; Trangie, Dec. 1947, L. J. Chinnick; Trangie, Dec. 1951, B. Cameron; Tyalgum, Mar. 1976, G. May; W Wyalong, Dec. 1973, J. C. Le Souef; Warrah, Feb. 1924; Wilberforce, Oct. 1973, P. Zborowski; Wingham Brush, Dec. 1984, G. Hoye; Wingham Scrub, Jan. 1970, E. B. Britton and S. Misko; MAMU (3 ♂♂; 2 ♀♀): 2: [no data]; 2: Bobbin Head, Jan. 1970, A. B. Rose; Urunga, Dec. 1982, B. Lewis; MV (6 ♂♂; 9 ♀♀): 4: [no data]; 4: Muswellbrook, E. Smith; Barmedman; Batemans Bay, Dec. 1975, M. S. and B. J. Moulds; Boambee, Jan. 1955; Canowindra, Dec. 1954; Muswellbrook; Sth. Boambee, Dec. 1952, B. Bamford; Sydney, Mar. 1940; OAI (1 ♂): Kempsey, Jan. 1985, E. E. Taylor. Queensland: ANIC (97 ♂♂; 74 ♀♀): 74: Kuranda, 19 Bulter Dr., {from Apr. 2003 to Apr. 2017}, D. C. F. Rentz; 10: Shiptons Flat, Oct. 1980, T. A. Weir; 9: Mt. Hypipamee NP., Nov. 1998, A. A. Calder; 6: Binna Burra, Lamington Nat. Pk., {from Dec. 1972 to Dec. 1973}, R. J. Kohout; 3: Bucasia, {from Nov. 2008 to Sep. 2002}, K. J. Sandery; 3: Mission Beach, {from Oct. 1991 to Jul. 1992}, G. O’Reilly; 2: 13.6 m up Whitefield Ra., Timber Rd., Cairns, Nov. 1970, J. G. Brooks; 2: Bunya Mts, Dec. 2004, 2: Clump Point, Jan. 1948, J. G. Campbell; 2: Gympie, Mar. 1949, J. King; 2: Mt. Webb Nat. Pk, Sep. 1980, T. A. Weir; 2: Tamborine, O. King; 2: The Crater, Mt. Hypipamee N. P., Feb. 1998, A. A. Calder; 12 km SE of Daintree, Nov. 1981, J. Balderson; Airlie Beach, Feb. 1978, T. Draper; Atherton Tablelands, Forty Mile Scrub Nat. Pk., Feb. 1998, S. Grossee, E. D. Edwards, R. Oberprieler and J. Cardale; Bellenden Ker., Jan. 1973, A. P. Richardson and J. G. Brooks; Black Mtn Rd., 5 mi. N of Kuranda, Dec. 1968, Britton, Brooks and Misko; Brisbane, Dec. 1983, J. F. R. Kerr; Broken R. Rd. rest area, 2 mi S of Eungella, Jan. 1967, J. G. Brooks; Broken Riv- er Rd. Rest Area 2 m. S of Eungella, Jan. 1967, J. G. and J. A. Brooks; Byfield (Yeppoon), Oct. 1924, H. J. Carter; Byfield, Nov. 1960, C. V.; Cairns, Apr. 1980, [Carnaby Coll.]; Cape Tribulation, Mar. 1989, H. Spencer; Cardstone, Jan. 1972, P. Zborowski; Credition nr. Broken River, Eungella NP, Dec. 1992, A. Colville and P. Greenslade; Danbulla F. R. 13 km NE by N of Yungaburra [Danbulla Forestry Reserve], Nov. 1981, J. Balderson; Danbulla Rd. at Lake Tinaroo, Mar. 2016, D. C. F. Rentz and B. Richardson; Eungella Nat. Pk., I. B. F. Common and M. S. Upton; Eungella; Jooro, Nov. 1933, D. V. Line; Jubilee Rd., 4.5 mi. NE Innisfail, Nov. 1966, E. B. Britton; Julatten, Jan. 2000, B. P. Moore; Kuranda; Kuranda, Feb. 1982, W. N. B. Quick; Kuranda, May 1905, Harris; Kuranda, Nov. 1967, J. A. G. Brooks; Lake Barrine, Atherton Tableland, Dec. 1985, J. Balderson; Lake Eacham NP., Nov. 1998, A. A. Calder; Lolworth [Lolworth Homestead], Mar. 1900; Mission Beach nr. Tully, Oct. 1967, R. Dobson; Mt. Baldy Loop Rd., nr Atherton, Nov. 2001, D. C. F. Rentz and M. S. Moulds; Mt. Cook Nat. Pk, May 1981, A. A. Calder; Mt. Isa, Dec. 1967, J. L. McRean; Mt. Lewis, Feb. 1969, J. G. Brooks; Mt. Lewis, Jan. 1977, B. P. Moore; Mungkan Kandju Nat. Pk., Coen Fiver camp, Jul. 1998, Grosse, Pullen, Calder, et al.; nr. Butchers Creek (village), Atherton Tablelands, Mar. 1988, D. C. F. Rentz; Paddington, Brisbane, Dec. 1970, R. J. Kohout; Palm Park, 4 ml. SE by E of Byfield, Dec. 1969, R. Jealous; Paluma, 5 km W, Jan. 1970, E. B. Britton and S. Misko; Paluma, Jan. 1980, D. W. Frith; Paluma, Mar. 1970, J. MacQueen; Rockhampton, Dec. 1961, C. V.; R’ton [Rockhampton], Dec. 1963, V. C.; St. Bernards, Mt. Tamborine, Jan. 1981, C. W. Frazier; St. Lucia, Bris. [Brisbane], Nov. 1977, T. Jordan; St. Lucia, Dec. 1989, D. Sinclair; Tamborine Mt., Davidson; Toowoomba, Dec. 1977, J. M.; Townsville, May 1905, C. Breuer; Wongabel St. For. Pavey Rd., Apr. 2013, D. C. F. Rentz; Yeppoon, Mar. 1935; Yeppoon, Nov. 1965, J. C. Le Souef; MAMU (2 ♂): 2: [no data]; MV (10 ♂♂; 6 ♀♀): 3: [no data]; 2: Goodna; 2: Mount Tamborine, {from Dec. 1950 to Dec. 1951}; 2: Stony crk., {from Aug. to Nov, 1938}, A. L. Brown; Blackall Range, Jan. - Dec. 1919; Cairns; Clump Point, Dec. 1999, J. O. Campbell; Clump Point, Jan. 1948; Cooktown, Feb. 1908, J. O. Campbell; Endeavour River; Gordonvale, Nov. 1925; QDPC (22 ♂♂; 20 ♀♀): 9: Toowong, Brisbane, Dec. 1977, R. H. Broadley; 5: Brisbane, {from Nov. 1958 to Nov. 1974}, R. A. Yule; 5: Newell Street, Atherton, {from Oct. 1970 to Nov. 1974}, J. H. Barrett; 4: Upper Barron, Nov. 1960, G. W. S.; 2: Brisbane, Dec. 1979, R. Hughes; 2: Long Pocket, Indooroopilly, Brisbane, {from Dec. 1976 to Dec. 1978}, J. F. Donaldson; 2: Mt. Fort William, 740 m via Kalpowar, Apr. 2000, M. de Baar; 2: Winder L. A. via Mt. Glorious, 50 km NW Brisbane, {from Dec. 1979 to Dec. 1980}, M. Hockey and M. de Baar; Atherton, Dec. 1963, G. W. Saunders; Atherton, Dec. 1968, J. H. Barrett; Atherton, Sep. 1968, B. Nelson; Bongeen, Dec. 1987, P. Twine; Brisbane, Nov. 1970, R. H. Broadley; Brisbane, Nov. 1985, L. Johnson; Bundamba, Ipswich, Oct. 2029; Corinda, Brisbane, Dec. 1995, B. Kennedy; Crater Lakes National Park, Jan. 1967, R. J. Elder; Garradunga, Innisfail, Dec. 1989; Imbil, A. C. Arvier; Imbil, Jan. – Dec. 1965, M. de Baar; Indooroopilly, Brisbane, Dec. 1987, M. Pilleris; Indooroopilly, Brisbane, Mar. 1978, C. C. Barrett; Kingaroy, Apr. 1978, H. Brier; Mount Tamborine, H. Hacker; Mt. Glorious SF., 40 km WNW Brisbane, Dec. 1979, D. Hockey; Nambour, Jan. 1957, T. H. K.; National Park, Dec. 1933, H. Hacker; National Park, H. Hacker; Taringa, Brisbane, Jan. 1930; Theodore; Toowong, Brisbane, Dec. 1964, C. J. Warson; Toowong, Brisbane, Nov. 1947, J. Jarvis; QM (8 ♂♂; 6 ♀): 4: Windsor Tableland NW of Mossman (site 1), Jan. 1994, G. and A. Daniels, R. Eastwood; 3: Mission Beach, Oct. 1978, K. L. Walker; „ Kahmoo ”, 20 km W of Cunnamulla, Jan. 1993 - Dec. 1994, K. Campbell; Garradunga nr Innisfail, Dec. 1993, G. and A. Daniels, R. Eastwood; Hurdle Gully, 13 km WSW Monto, Dec. 1997, Monteith, Cook, Thompson; Ipswich, Nov. 1985, G. Daniels; Massey Ck., Eungella, Dec. 1993, J. Pease; Mt Macartney Cathu SF, Jan. 1996, G. and A. Daniels; Mt Misery, SW of Cooktown, Jan. 1994, G. and A. Daniels, R. Eastwood. Victoria: MV (1 ♂; 1 ♀): Ballarat District (Ballarat); French Island. Western Australia: ANIC (1 ♀): Port Hedland, Sep. 1965, P. Carne. Molecular data. BioSample ID: SAMN 12650019, SAMN 12650065, SAMN 11080862	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB731B26F958FBEAFDA7F9DE.taxon	description	(Figs. 3, 14 B, 15 E, 16 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB731B26F958FBEAFDA7F9DE.taxon	diagnosis	Diagnostic combination (male). Moderately large beetles, 25 – 40 mm long. Dorsum unicoloured, brown or dark brown. Sexually dimorphic dense punctures present on lateral or antero-lateral parts of pronotum and ventral part of prothorax. Head slightly narrower than prothorax; frontoclypeal suture straight or arcuate; median groove complete. Antennal tubercles relatively flat, directed laterally, angulate at apex. Eyes moderately large, weakly emarginate near antennal insertion, very broadly separated dorsally. Mandibles less than half of head length, not sexually dimorphic except for more apparent and denser setation in male, broad at base, strongly bent and unidentate apically. Antenna not reaching mid length of elytra in male. Scape about 1 / 4 of head length, posteriorly not extending to posteri- or margin of eye, globular; antennomere 3 about as long as scape and about as long as 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps ovoid and apically rounded. Prothorax transverse with lateral carina almost smooth; anterior and posterior margins with complete beads. Pronotal disc mostly shiny with incomplete median longitudinal groove and coarsely punctate, slightly depressed lateral irregular areas. Prosternal process distinctly expanded beyond procoxae, extending to mesoventrite; rounded apically. Elytral surfaces weakly coriaceous; inner apical angle without spine. Legs strong; femora and tibiae without rows of spines. Protibia smooth externally with sharp external spine and pair of subequal spurs; lobes on tarsomere 3 narrow and broadly separated; tarsomere 5 as long as tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB731B26F958FBEAFDA7F9DE.taxon	description	Description. Male. Length 25 - 40 mm. Dorsum usually uniformly brown or dark brown, elytra sometimes light brown. Head approximately as long as wide, slightly narrower than pronotum. Mandibles without sexual dimorphism, less than half of head length; apex moderately curved inwards, surface bearing dense setae along incisor edge; each mandible wedge-shaped, thick and strong on lateral side and near the base, unidentate at apex, with one extra tooth on incisor edge. Labrum triangular, weakly elongate near the base, dorsal surface covered with long golden setae. Frontoclypeal suture straight or broadly rounded. Antennal tubercles relatively flat, very distant to each other, apex bluntly angulate. Antenna 11 - segmented, filiform or slightly moniliform, extending to humeral angles of elytra; scape short and stout, as long as eye length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; broadly separated dorsally. Submentum weakly curved at apex; mentum well exposed. Terminal palpomere of maxillary and labial palps ovoid and apically rounded. Pronotum transverse, anterior margin emarginate medially with complete bead; posterior margin with complete bead; lateral margin with rather small and short teeth. Disc shiny and sparsely punctate medially, with median longitudinal groove; dense fine punctures near anterior angles and rather coarse punctures near posterior angles. Prosternum covered with fine and deep punctures; hypomeron well-defined, slightly broader than prosternal process; prosternal process projecting towards mesoventrite, laterally expanded near apex and pointed apically. Mesoventral process emarginate at apex. Ventral side of pterothorax and hind coxae covered with golden hairs. Metanepisternum slightly narrowing posteriorly. Scutellum narrowly rounded apically, surface shiny and smooth. Elytral surface shiny with irregular coriaceous sculpture; two weak traces of venation often present on apical 2 / 3 of elytra; elytral apices rounded with distinct suture angles, but without spines; epipleuron almost complete, narrowing apically. Legs strong and relatively short; femur broad and relatively flattened, weakly restricted at both ends, surface smooth, ventral side bearing few short spines apically; each tibia with distinct apical spine and a pair of strong spurs; tarsi slender; tarsomere 3 with relatively narrow lobes; tarsomere 5 as long as tarsomere 1 and 2 combined. Abdominal ventrites only with sparse long hairs or few fine punctures, but bearing dense long setae along the edges, especially on the apex of ventrite 5. Male genitalia (Fig. 14 B). Tegmen longer than sternite VII; parameres relatively short, truncate apically with triangular projections at the base; penis longer than tegmen, dorsal apex strongly emarginate while ventral apex narrowly pointed. Female. Mandible surface glabrous. Pronotal disc with coarser punctures near anterior angles; fine and dense punctures on prosternum absent. Ovipositor (Fig. 15 E) short, apical sclerotised part as long as baculus; distal gonocoxites elongate; stylus inserted laterally and relatively distant to gonocoxite apex, very weakly expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB731B26F958FBEAFDA7F9DE.taxon	discussion	Remarks. Analophus can be distinguished from the other genera of the Australian Macrotomini by having short antennae that only extend to the elytral humeral angle, the prothorax is transverse with the lateral carina almost smooth, and the pronotal disc is mostly shiny, coarsely punctate with an incomplete median longitudinal groove. Four species are currently recognised in Analophus worldwide, including the one new species described here, and possibly one species has been incorrectly assigned to this genus. The type species Analophus parallelus Waterhouse was described from Queensland, Australia. Gahan added the second species A. niger Gahan from the Arfak Mountains, Indonesia. About 100 years later, the third species A. vicksoni Nýlander and Komiya was described from Papua New Guinea. However, based on the images and description from Nýlander and Komiya (2015), A. vicksoni is not likely to be a species of Analophus because of the general body shape and the pattern of sexual dimorphism. Without examining the specimens, we cannot attempt to place it into an appropriate genus. On the other hand, we have discovered a new Australian species close to A. parallelus, based on both morphological characters and molecular sequences. The new species is restricted to Northern Queensland, while A. parallelus is mainly distributed in south-eastern Queensland and New South Wales.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B26FB39FF12FC00F958.taxon	description	(Figs. 3 A – C, 3 G, 3 I, 14 B, 15 E, 16 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B26FB39FF12FC00F958.taxon	diagnosis	Diagnosis. Male of A. parallelus can be separated from the male of A. septentrionalis sp. nov. by its sexually dimorphic punctures occupying most of the antero-lateral side of pronotum (Fig. 3 I), and by very short parameres, about 0.11 times length of entire tegmen. The female is distinguished from the new species in having the clypeus densely setose.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B26FB39FF12FC00F958.taxon	materials_examined	Types. Analophus parallelus Waterhouse, Holotype male deposited in BNHM, with the following original labels “ Analophus parallelus (Type) C. Waterh. | Queensland, 77.17 ”. Material examined (Fig. 16 B). New South Wales: AM (1 ♂): 50 km N of Warren, Dec. 1988, M. S. and B. J. Moulds; ANIC (1 ♂) Trangie, K. H. L. Key; MV (1 ♂): [no data]. Queensland: ANIC (2 ♂♂; 1 ♀): 2: Emerald, Dec. 1935, [S. R. E. Brock Coll.]; Edungalba, Jan. 1940, [S. R. E. Brock Coll.]; QDPC (2 ♂♂; 3 ♀♀): Clermont; Jandowae, Dec. 1936, J. Turner; Research Station, Biloela, Nov. 1967, F. D. Page; Toowoomba, Nov. 1973, R. Lloyd; [no data]; QM (8 ♂♂; 18 ♀♀): 2: Expedition Ra. NP, Amphitheatre camp, Dec. 1997, Monteith, Cook and Thompson; 2: Hurdle Gully, 13 km WSW Monto, Dec. 1997, Monteith, Cook and Thompson; 2: Wonga Hills site, Dec. 2001, Monteith, Cook, and Wright; “ Allinga ” Chinchilla, Lithgow, G.; Blackbutt Ra. W of Moore, Jan. 2001, L. Popple, J. Moss and M. Moulds; Boggomoss No. 19 via Taroom, Nov. 1996, G. Monteith; Boggomoss No. 21 via Taroom, Nov. 1996, G. B. Monteith; Braigalow Res. Stn, site 10, Dec. 2000, A. Eward; C. Deane; Cameron’s Scrub, knoll top, Nov. 1998 - Jan. 1999, Monteith, Cook, Thompson; Chinchilla (town), Lithgow, G.; Cobbs Hill, Site 2, Mar. 1993, S. Hamlet; Conway Range N. P., Nov. 1992, Monteith, Thompson, Cook and Janetzki; Drummond Ra. Summit, Oct. 2002, Cook, Wright and Vanderduys; Expedition Ra. NP, Amphitheatre camp, Dec. 1977, Monteith, Cook and Thompson; Nipping Gully, site 6, Dec. 1998, G. Monteith, C. Gough and G. Maywald; Southwood NP site 5, Dec. 2005, G. Monteith; St Lawrence, 5 km E, Oct. 2000, Cook, Wright and Vanderduys; Taroom District, Jan. 1997, Burwell, Cook and Janetzki; Taroom, Nov. 1992, G. Daniels; Taroom, Nov. 1999, G. and A. Daniels; Theodore Brigalow Reserve, Dec. 1999; Yarramulla, Feb. 2003, G. Monteith. Molecular data. BioSample ID: SAMN 12650056, SAMN 12650111.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	description	(Figs. 3 D – F, 3 H, 3 J, 16 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	diagnosis	Diagnosis. Male of A. septentrionalis can be separated from A. parallelus in having sexually dimorphic punctures limited only to the antero-lateral corner of the pronotum (Fig. 3 J), and relatively long parameres, about 0.17 times length of the entire tegmen. The female of A. septentrionalis has the clypeus sparsely setose or glabrous. This species is restricted to northern Queensland.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	description	Description. Male. Body length 30 - 35 mm. Body sub-parallel, slightly convex on dorsum; head length about 0.7 × of width; mandibles shorter than head, weakly curved towards ventral side; pronotum shorter than head, about 0.7 × of head length; elytra 3.5 × longer than pronotum. Antennal scape 1.2 × of antennomere 3 length; interorbital width over 3 × of eye length. Prosternal process 0.75 × as diameter of procoxal cavity, twice as hypomeron width; mesoventral process narrower than prosternal process. Abdominal ventrite 1 moderately longer than the others, ventrite 5 rounded at apex. Metatarsomeres 1 – 3 length ratio as 1: 0.7: 0.8, tarsomere 5 shorter than 1 – 3 combined. Female. Body length 30 - 45 mm; pronotum as long as head and slightly broader than head; elytra about 4 times of pronotum length.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	etymology	Etymology. The species name is a Latin word, meaning “ northern ” in English, indicating its Northern Queensland distribution.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	biology_ecology	Biology information. Some specimens were caught boring in the candlenut tree (Aleurites moluccanus.)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7C1B27FBDDF97AFC58FE83.taxon	materials_examined	Types. Holotype. ♂: “ 16 ° 48 ’ S, 145 ° 38 ’ E (GPS), QLD: Kuranda (335 m elev.) (Top of the Range) 19 Butler Dr., 15 - 30 December 2003, D. C. F. Rentz | ANIC 25 - 067075 ” (ANIC). Paratypes (7). 4 ♂♂: “ Paluma N. Q., Jan 1968, at light | ANIC 25 - 073238 ” (ANIC); “ Crystal Ck., Q. 23 mi. SSE of Ingham, 18.58 Sx 146.16 E, 9. xii. 1968, at light, Britton and Misko | ANIC 25 - 073245 ” (ANIC); “ Black light, Kareeya Power Station, Tully, N. E. QLD, 27 / 12 / 2000, S. Fearn ” (QVMAG); “ Black light, Kareeya Power Station, Tully, N. E. QLD, 28 / 12 / 2000, S. Fearn ” (QVMAG); 3 ♀♀: “ 16 ° 48 ’ S, 145 ° 38 ’ E (GPS), QLD: Kuranda (335 m elev.) (Top of the Range) 19 Butler Dr., 1 - 15 January 2004, D. C. F. Rentz | ANIC 25 - 073260 ” (ANIC); “ 16 ° 48 ’ S, 145 ° 38 ’ E (GPS), QLD: Kuranda (335 m elev.) (Top of the Range) 19 Butler Dr., 15 - 29 February 2004, D. C. F. Rentz | ANIC 25 - 073260 ” (ANIC); “ Black light, Kareeya Power Station, Tully, N. E. QLD, 28 / 12 / 2000, S. Fearn ” (QVMAG). Other material examined (Fig. 16 B). Queensland: ANIC (2 ♂♂; 9 ♀♀): 2: Atherton, Jan. 1930, R. H. Doggrell; Cardstone, Feb. 1962, K. H.; Cardstone, Feb. 1963, G. B.; Green Forest Rd., Kuranda, Dec. 2013, J. Hasenpusch; Kirrama Range, Nov. 1968, W. and A. E. Y. Muir; Lake Placid, Cairns, Jan. 1968, J. G. Brooks; Mount Aunt, 10 km SSW of Mareeba, Mar. 1988, D. C. F. Rentz; Polly Ck., Garradunga, Dec. 2014, J. Hasenpusch; Polly Ck., Garradunga, Feb. 2014, J. Hasenpusch; Smithfield, Cairns, Dec. 2013, J. Hasenpusch; MV (1 ♂): [no data]; OAI (1 ♀): Atherton, Feb. 1975, G. R. Brown; QVMAG (2 ♂♂; 2 ♀♀): 3: Kareeya Power Station, Tully, Dec. 2000, S. Fearn; Kareeya Power Station, Tully, Nev. 2001, S. Fearn. Molecular data. BioSample ID: SAMN 11080863 (Holotype).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7D1B24FBC5FE25FB04FB80.taxon	description	(Figs. 4 A-G, 14 C, 15 F, 16 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7D1B24FBC5FE25FB04FB80.taxon	diagnosis	Diagnostic combination (male). Large beetles, 25 – 90 mm long. Dorsum usually uniformly dark brown, sometimes with light or reddish-brown elytra. Sexually dimorphic dense punctures present on most of the pronotum and to various extent on ventral side. Head narrower than prothorax; frontoclypeal suture angulate; median groove complete. Antennal tubercles prominent, rounded at apex. Eyes large, weakly emarginate near antennal insertion, narrowly separated dorsally. Mandibles about 1 / 3 of head capsule length, not sexually dimorphic, broad at base, not bent towards ventral side and unidentate apically. Antenna extends to or slightly beyond elytral apex. Scape about half of head capsule length, posteriorly extending beyond posterior margin of eye, gradually expanded; antennomere 3 three times of scape length, much longer and thicker than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps oval and apically weakly truncate. Prothorax transverse with posterior angles blunt but projecting posteriorly; lateral carina very weak, regularly dentate. Anterior and posterior margins with very fine beads not extending to anterior or posterior angles. Pronotal disc with almost smooth and shiny callosities on densely punctate background. Prosternal process relatively broad, laminate and arcuate, weakly narrowing towards apex, extending beyond procoxae and rounded apically. Elytral surfaces coriaceous; elytral apices with sharp spine. Legs relatively long; rows of spines present on femora and tibiae. Protibial surface rough but glabrous, with pair of unequal spurs apically; tarsomere 3 moderately lobed; tarsomere 5 as long as or slightly shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7D1B24FBC5FE25FB04FB80.taxon	description	Description. Male. Length 25 – 90 mm. Body usually uniformly brown or dark brown, in some species with reddish- or yellowish-brown elytra. Head approximately as long as wide, narrower than pronotum. Mandibles about 1 / 3 of head length, weakly curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two or three teeth on incisor edge. Labrum very short, anterior margin rounded with very dense setae; labrum separated from clypeus by a deep groove; dorsal surface of labrum with longer and denser setae than on clypeus. Frontoclypeal suture angulate; median groove relatively broad in anterior third, complete. Antennal tubercles prominent with apex rounded. Antenna 11 - segmented, filiform, extending to or beyond elytral apices; scape moderately long and gradually expanded, reaching beyond posterior margin of eye; antennomere 3 at least three times longer than scape or antennomere 4, and thicker than all following segments, surface very rough, bearing tiny spines on ventral side; antennomeres 5 – 11 with rather blunt posterior angles. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; narrowly separated dorsally. Submentum curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps oval and apically truncate. Pronotum transverse, trapezoid with posterior margin broader than anterior margin; beads present on both anterior and posterior margin not extending laterally to anterior or posterior angles; lateral carina very fine with regular sharp teeth. Disc very densely punctate and matt except for two shiny, sparsely punctate islands in admedian area. Prosternum covered with fine and deep punctures; hypomeron well-defined, narrower than prosternal process. Prosternal process relatively broad, laminate and bent towards mesoventrite, rounded apically. Mesoventral process sub-parallel and emarginate apically, without median impression or groove to receive prosternal process. Metaventrite with dense pubescent triangular area medially and densely punctate area elsewhere. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface with dense fine hairs. Elytral surfaces dull covered with irregular coriaceous sculpture; two or three weak traces of venation especially on basal half; elytral apices rounded with sharp sutural spine; epipleuron almost complete, narrowing apically. Legs strong and long; femur sub-parallel, rows of spines present on ventral side; tibia slender and relatively flat, with spines along outer margin, surface very rough but glabrous; each tibia with distinct apical spine and a pair of unequal spurs; tarsi strong; tarsomere 3 moderately lobed; tarsomere 5 as long as or slightly shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1 – 5 smooth, only with fine punctures near antero-lateral angles surfaces almost glabrous, only bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 C). Tegmen longer than sternite VII; parameres moderately long, 0.14 times length of entire tegmen, rounded apically without triangular projections at the base; penis longer than tegmen, dorsal apex truncate or emarginate while ventral apex narrowly pointed. Female. Antenna usually shorter than in male, extending beyond middle of elytra but not reaching the apex; antennomere 3 smooth, much longer than scape or antennomere 4, but not thicker than antennomeres 4 – 11. Pronotal disc relatively shiny, with sparse and fine punctures near centre and coarse dense punctures laterally. Fine and dense punctures on thorax but absent from abdomen. Legs much slender than males. Abdominal ventrite 5 truncate or weakly emarginate medially. Ovipositor (Fig. 15 F) long, apical sclerosed part consists of about half length of baculus; distal gonocoxites relatively short and small, rounded near apex; stylus large, inserted laterally and very close to gonocoxite apex, gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7D1B24FBC5FE25FB04FB80.taxon	discussion	Remarks. Anomophysis can be distinguished from the remaining genera of the Australian Macrotomini by the extremely elongate antennomere 3, which is at least 3 times longer than the scape or antennomere 4, and very narrowly separated dorsal eye lobes. Male with sexual fine punctures on pronotum and ventral side of thorax, antennomere 3 thick with surfaces very rough and bearing tiny spines on ventral side. Twenty-one species are currently recognised in Anomophysis worldwide, mostly distributed in the Indo-Malaysian region. Only two species have been recorded in Australia, including A. coxalis restricted to the Christmas Island. The only species from mainland Australia, A. australis was removed from Pseudoplites by Queentin and Villiers (1981), but its distribution is uncertain as very limited number of specimens have been found for this species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7E1B25FB15FA1EFD0BFE83.taxon	description	(Figs. 4 A – C, 16 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7E1B25FB15FA1EFD0BFE83.taxon	diagnosis	Diagnosis. A. australis can be separated from A. coxalis by its distribution and the elytra brown or reddish brown, with weak venations. Male can also be recognised by its acute posterior angles of pronotum.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7E1B25FB15FA1EFD0BFE83.taxon	materials_examined	Types. Anomophysis australis Quentin et Villiers, Holotype male deposited in MNHN, male genitalia dissected and stored separately, with the following labels “ northern Lemty [partially unintelligible writing] | Ex-Musaeo Mniszech | Muséum Paris 1952, Coll. R. Oberthur | Anomophysis australis nobis Holotype ♂, Quentin and Villiers Det. 1979 ”. Material examined (Fig. 16 C). Australia: ANIC (2 ♀♀): NT, Berrimah, Sep. 1976; [no data]. Molecular data. BioSample ID: SAMN 11080891.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B25F8B1FE22FE5CFACF.taxon	description	(Figs. 4 D – G, 14 C, 15 F, 16 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B25F8B1FE22FE5CFACF.taxon	diagnosis	Diagnosis. A. coxalis can be separated from A. australis by its geographic distribution and the elytra yellowish brown with strong venation, especially on the basal half. Male can also be recognised by posteri- or angles of pronotum narrowly rounded, and the anterior angles distinct.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B25F8B1FE22FE5CFACF.taxon	materials_examined	Types. Quentin and Villiers (1981) designated two lectotype specimens for this species, here we specify the lectotype as the male specimen (BNHM- 010799864) with the following original labels “ Flying Fish Cove, Oct 1897 | Christmas I., C. W. Andrews, 1898, 98 – 20 | Prinobius coxalis, Gahan, ♂, Type ”, the female specimen (BNHM- 010799827) is here designated as paralectotype, with the following original labels “ Christmas I., C. W. Andrews, Dec. 1897, 98 - 20 | Prinobius coxalis Gahan, Type ♀ ”. Material examined (Fig. 16 C). Christmas Island: ANIC (5 ♂♂; 7 ♀♀): 3: nr. Central area, Apr. 1989, J. F. Lawrence; 2: 1 km ENE Aldrich Hill, Oct. 1983, L. Hill; 2: 2 km WSW Waddle Hill, Oct. 1983, L. Hill; 2: Jungle Bean Hill, Oct. 1964, T. G. Campbell; Drumsite; settlement, Oct. 1964, T. G. Campbell; vic. Of Gtants Well, Apr. 1989, J. F. Lawrence.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B22F96CFAEEFE04F9BB.taxon	description	(Figs. 4 H – K, 14 D, 15 G, 16 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B22F96CFAEEFE04F9BB.taxon	diagnosis	Diagnostic combination (Male). Moderately large beetles, 26 – 40 mm long. Dorsum bicoloured, head and pronotum darker than elytra. Sexually dimorphic, dense fine punctures present on lateral sides of pronotum. Head as broad as prothorax; frontoclypeal suture broadly rounded; median groove complete. Antennal tubercles almost flat, weakly angulate apically. Eyes transverse, weakly emarginate near antennal insertion, broadly separated dorsally. Mandibles almost same length as head capsule, sexually dimorphic in the length and thickness, weakly curved and unidentate apically. Antenna reaching mid elytra. Scape about 1 / 4 of head length, posteriorly not extending to posterior margin of eye, globular; antennomere 3 about as long as scape and about as long as 4. Mentum fused to submentum (Fig. 4 K). Terminal palpomere of maxillary and labial palps ovoid and apically rounded. Prothorax transverse, as long as or slightly shorter than head capsule; lateral carina almost smooth; anterior margin with bead incomplete medially, posterior margin with complete bead. Pronotal disc mostly shiny and smooth, with fine and deeply punctate, slightly depressed lateral irregular areas. Prosternal process distinctly expanded beyond procoxae, extending to mesoventrite, narrowly pointed apically. Elytral surface semitransparent; inner apical angle angulate or with weak spine. Legs strong, femora and tibiae without rows of spines. Protibia smooth externally with sharp external spine and pair of subequal spurs; lobes on tarsomere 3 narrow and broadly separated, tarsomere 5 much longer than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B22F96CFAEEFE04F9BB.taxon	description	Description. Male. Length 26 – 40 mm. Head and pronotum reddish or dark brown, elytra lighter than pronotum often yellowish and semitransparent. Head transverse, as broad as pronotum. Mandibles show sexual dimorphism on length and thickness which are much longer and stronger in males, dorsum bearing dense hairs, apex weakly curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with several teeth on incisor edge. Labrum weakly constrict- ed at base, anterior margin pointed medially with very dense setae; labrum separated from clypeus by a deep groove; dorsal surface of labrum and clypeus covered with long golden setae. Frontoclypeal suture almost straight; median groove distinct and complete. Antennal tubercles relatively flat, very distant to each other, apex blunt angulate. Antennae 11 - segmented, filiform or slightly moniliform, extending to mid elytra; scapes short and stout, as long as eye length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; broadly separated dorsally. Mentum fused to submentum, submentum emarginate medially and weakly depressed on apical central area, covering the base of maxilla and labium. Terminal palpomere of maxillary and labial palps ovoid and apically rounded. Pronotum transverse, almost sub-squared and with posterior margin slightly narrower than anterior margin; with incomplete bead on anterior and complete bead on posterior margins; lateral carina rather smooth. Disc surface shiny, glabrous and smooth medially, with deep and fine punctures on lateral area. Prosternum with deep fine punctate area along procoxal cavities, and a smooth shiny T-area from anterior prosternum to the apex of prosternal process; hypomeron well-defined, as broad as prosternal process; prosternal process tridentate at apex, median lobe projecting towards mesoventrite, narrowly rounded apically. Mesoventral process broad, apex truncate and emarginate medially. Ventral side of pterothorax and coxae covered with golden hairs. Metanepisternum weakly constricted at base. Scutellum narrowly round- ed at apex, surface smooth. Elytra surfaces relatively shiny with weak irregular coriaceous sculpture, two to three traces of venation present; elytral apices angulate without sutural spines; epipleuron almost complete, very narrowing apically. Legs short and strong, femur broad and relatively flat, weakly restricted at both ends, surface smooth, ventral side without spines; each tibia with distinct apical spine and a pair of strong spurs; tarsi slender; tarsomere 3 with relatively narrow lobes; tarsomere 5 much longer than tarsomere 1 and 2 combined. Abdominal ventrites surface smooth, only bearing dense long setae along the edge, especially on the apex of ventrite 5. Male genitalia (Fig. 14 D). Tegmen longer than sternite VII; parameres relatively short, only 0.1 times length of entire tegmen, rounded apically with triangular projections at the base; penis longer than tegmen, dorsal apex rounded while ventral apex narrowly rounded. Female. Mandibles much shorter than in males, surface with less setae; pronotum constricted near anterior margin, with coarsely punctate, irregular lateral area; prosternum and hypomeron with sparse fine punctures and loose hairs. Ovipositor (Fig. 15 G) long, apical sclerosed part 1 / 3 of barculus length; distal gonocoxites short and stout; stylus inserted laterally and close to gonocoxite apex, relatively long and gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7F1B22F96CFAEEFE04F9BB.taxon	discussion	Remarks. Archetypus can be distinguished from the remaining genera of the Australian Macrotomini by its medium body size, the pale yellow elytra, the tibia without lateral spines, and the tarsomere 5 much longer than tarsomeres 1 and 2 combined. Males are recognised by modified elongate and strong mandibles, bearing dense golden hairs on dorsal surface. Only two species are recognised in Archetypus. The type species, A. fulvipennis was described from Dorey, Papua New Guinea, and later recorded from Aru, Lifu, Solomon Islands, and northern Queensland. However, according to our molecular results, the specimens from the northern Queensland are genetically different from the Papuan populations and are described here as a new species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB781B22F94CF91AFC4BF8AB.taxon	description	(Figs. 4 H – K, 14 D, 15 G, 16 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB781B22F94CF91AFC4BF8AB.taxon	diagnosis	Diagnosis. Male of A. marginatus sp. nov. can usually be separated from the Papuan A. fulvipennis by the median carina deeply impressed anteriorly, forming shallow groove while joining the frontoclypeal depression (median groove is indistinct anteriorly in A. fulvipennis). Otherwise this species differs from the Papuan congener by its reddish-brown head and pronotum, the elytra always pale yellow with dark brown margin and the inner apical angle angulate (with weak spine in A. fulvipennis). It is restricted to northern Australia.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB781B22F94CF91AFC4BF8AB.taxon	description	Description. Male. Body length 29 – 37 mm; Body sub-parallel, head length about 0.7 times width; mandibles as long as head capsule; pronotum as long as head and slightly broader than head; elytra 3 times of head length. Antennal scape 1.5 times of antennomere 3 length; interobital width 4 times of eye length. Prosternal process moderately broad, as wide as the length of procoxal cavity, and 2 times of hypomeron width; mesoventral process broader than prosternal process. Abdominal ventrite 1 being the longest, about twice length of ventrite 2; ventrite 5 slightly shorter than 4. Metatarsomeres 1 – 3 length ratio as 2: 1: 1; tarsomere 5 slightly longer than 1 – 3 combined. Female. Body length 26 – 34 mm; mandibles shorter than head capsule; pronotum slightly shorter and broader than head; elytra over 5 times of pronotum length.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB781B22F94CF91AFC4BF8AB.taxon	etymology	Etymology. The species name is derived from the Latin word “ marginatus ”, which indicates the dark margin on elytra of this new species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB781B22F94CF91AFC4BF8AB.taxon	materials_examined	Types. Holotype. ♂: “ Iron Range, Cape York Pen., N. QLD, Jun 30 - July 4 1977, G. B. Monteith | ANIC 25 - 068370 ” (QM). Paratypes (5). 2 ♂♂: “ Iron Range, Cape York Pen., N. QLD, Jun 30 - July 4 1977, G. B. Monteith ” (QM); “ 12.44 S 143.14 E, 3 km ENE of Mt. Tozer, QLD, 28 Jun. – 4 Jul. 1986, T. Weir and A. Calder | collected at light ” (ANIC); 3 ♀♀: “ Iron Range, Cape York Pen., N. QLD, Jun 30 – July 4 1977, G. B. Monteith ” (QM); “ 12 ° 42 ’ 37 S 143 ° 17 ’ 33 E, Iron Range Nat. Pk., Cooks Hut, QLD, 15 – 20 July 1998, Grosse, Pullen, Calder, Oberprieler and Edwards | ANIC 25 - 072962 ” (ANIC); “ 12.43 S 143.18 E GPS, Iron Range N. P., QLD, 1 km NE Lamond Hill, 16 - 19 Jul. 1998, A. A. Calder, at light ” (ANIC). Other material examined (Fig. 16 C). Queensland: ANIC (1 ♂; 2 ♀♀): 15 km WNW of Bald Hill, McIhwralth Range, Jun. 1989, T. A. Weir; Iron Range Nat. Pk., Cooks Hut, July 1998, M. Neave, T. Weir, S. Pratt and L. Mckenzie; Captain Billy’s Landing, May 2015, A. Zwick; Iron Range Nat. Pk., Cooks Hut, July 1998, M. Neave, T. Weir, S. Pratt and L. Mckenzie; QM (2 ♂♂; 2 ♀♀): 2: Iron Range, Cape York Pen., Jun. 1977, G. B. Monteith; Cape York Pen., Nov. 1969, B. Cantrell; Iron Range, Cape York Pen., Jun. 1975, M. Walford- Huggins. Molecular data. BioSample ID: SAMN 12650014 (Holotype), SAMN 12650110 (Paratype), SAMN 12650122.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB791B20F976FF12FE62FDD9.taxon	description	(Figs. 5 A – I, 14 E, 15 H, 16 D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB791B20F976FF12FE62FDD9.taxon	diagnosis	Diagnostic combination (Male). Moderately large beetles, 20 – 42 mm long. Dorsum uniformly brown to dark brown, some individuals may have slightly yellowish-brown elytra. Sexually dimorphic dense punctures apparent only in some males and present on dorsal and ventral side of prothorax. Head narrower than prothorax; frontoclypeal suture angulate; median carina deeply grooved, complete. Antennal tubercles prominent, rounded apically. Eyes moderately large, weakly emarginate near antennal insertion, relatively distant dorsally. Mandibles short, about half of head capsule length, strongly angulate and unidentate apically. Antenna reaching the middle of elytra. Scape about 1 / 2 – 2 / 3 of head capsule length, posteriorly extending to or beyond posterior margin of eye, not or very weakly expanded apically, flattened with blunt external edge; antennomere 3 about as long as or slightly shorter than scape and subequal to antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps weakly expanded and apically truncate. Prothorax transverse with lateral carina weakly or regularly dentate; anterior and posterior margins with complete beads. Pronotal disc uneven with two admedian weakly elevated smooth and less punctate callosities on densely and coarsely punctate background. Prosternal process expanded beyond procoxae, extending beyond procoxae, rounded apically. Elytral surfaces strongly coriaceous; inner apical angle with sharp spine. Legs strong; femora and tibiae with rows of sharp spines. Protibia with two sharp apical projections and a pair of subequal spurs; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB791B20F976FF12FE62FDD9.taxon	description	Description. Male. Length 20 – 42 mm. Dorsum usually uniformly brown to dark brown, sometimes with yellowish elytra. Head approximately as long as wide, narrower than pronotum. Mandibles about half of head length, moderately curved inwards, sexually dimorphic, longer and stronger in males; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two extra teeth on incisor edge. Labrum transverse, anterior weakly pointed medially with very dense setae; labrum separated from clypeus by a deep arcuate groove; dorsal surface of labrum and clypeus almost glabrous or covered with fine hairs. Frontoclypeal suture angulate; median groove distinct and complete. Antennal tubercles prominent, apex rounded. Antenna 11 - segmented, filiform, extending to or beyond middle of elytra; scape relatively short and stout, flat and bearing blunt external edge, reaching to or beyond posterior margin of eye; antennomere 3 as long as or slightly shorter than scape or antennomere 4; antennomeres 5 – 11 with blunt posterior angles. Eyes moderately large, transverse, coarsely facetted, slightly emarginate near antennal foramen; relatively distant dorsally. Submentum weakly curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps weakly expanded and apically truncate. Pronotum transverse with rounded posterior angles; anterior margin weakly emarginate medially with complete bead; posterior margin with complete bead; lateral margins moderately to strongly dentate or with regular small sharp teeth. Disc surface very densely and coarsely punctate and matt except for two elevated shiny and less punctate triangular admedian islands. Prosternum covered with coarse coriaceous punctures; hypomeron well-defined but very narrowed anteriorly, narrower than prosternal process; prosternal process moderately expanded beyond procoxae, projecting towards mesoventrite, abruptly narrowed and rounded apically. Mesoventral process emarginate at apex. Ventral side of thorax and coxae covered with long dense golden hairs. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface very finely punctate or smooth. Elytral surfaces dull covered with irregular coriaceous sculpture, without traces of venation; elytral apices rounded with short sutural spine; epipleuron almost complete, narrowing apically. Legs strong and relatively long; femur flat and weakly emarginate at both ends, with two rows of sharp spines on ventral side; tibia moderately strong, with regular spines along outer margin; each tibia with two distinct apical projections and a pair of subequal spurs; tarsi slender; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdominal ventrites glabrous, smooth and shiny medially, lateral area covered with fine and dense punctures; adpressed hairs present along the edges, sparse near posterior margin, relatively dense close to lateral margin; very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 E). Tegmen longer than sternite VII; parameres very long, 0.23 times length of entire tegmen, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex narrowly pointed. Female. Entire body usually much slender than male, with longer elytra. Eyes larger than in male. Antennae usually shorter than in male; antennal scape thicker than the following segments, but not as distinctly expanded as in male. Abdominal ventrites bearing relatively dense long hairs. Ovipositor (Fig. 15 H) long, apical sclerosed part about half length of baculus, distal gonocoxites relatively short and sharp; stylus long, inserted laterally and very close to gonocoxite apex, gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB791B20F976FF12FE62FDD9.taxon	discussion	Remarks. Brephilydia can be distinguished from the remaining genera of the Australian Macrotomini by having the frontoclypeal suture angulate and the external margin of protibia with very sharp spines. Males are recognised by the stout antennal scape, more or less in a shape of triangular prism. The type species B. jejuna is widely distributed in New South Wales and south-eastern Queensland, while a new species, B. fearni sp. nov., was found in the northern Queensland.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B20F96CFBE9FB1BFB21.taxon	description	(Figs. 5 A – D, 16 D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B20F96CFBE9FB1BFB21.taxon	diagnosis	Diagnosis. The northern Queensland species B. fearni is not sympatric with more southern and widely distributed vicariant. It can be recognised from B. jejuna by having frontoclypeal suture approximately forming a right angle (Fig. 5 D), the mandibular surface with very short and sparse fine hairs and the longer elytra (about 5 times of pronotum length in male, 6 times length of pronotum in female).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B20F96CFBE9FB1BFB21.taxon	description	Description. Male. Body length 26 – 42 mm; body sub-parallel and slender, head capsule length about 0.7 × of width; mandibles strong, as long as frons; pronotum slightly longer and broader than head; elytra over 4 × of head capsule length. Antennal scape longer than antennomere 3, and almost twice thicker than antennomere 3; interorbital width 1.4 × of eye length. Prosternal process narrow medially, gradually expand- ed and pointed at apex, narrowest point about 0.3 × length of procoxal cavity, and same as hypomeron width; mesoventral process broader than prosternal process, width about half of mesocoxal cavity width. Elytra about 5 × length of pronotum. Abdominal ventrite 1 slightly longer than 2 or 3, ventrite 5 truncate or very weakly emarginate at apex. Metatarsomere 1 – 3 length ratio as 2: 1: 1, tarsomere 5 shorter than 1 and 2 combined. Female. Body length 25 – 45 mm; antennal scape about 1.5 × thicker than antennomere 3; elytra about 6 x length of pronotum.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B20F96CFBE9FB1BFB21.taxon	etymology	Etymology. The species name is dedicated to Mr. Simon Fearn from QVMAG, who collected the most type specimens and generously loaned us many prionine specimens for this project.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B20F96CFBE9FB1BFB21.taxon	materials_examined	Types. Holotype. ♂, “ Black light, Kareeya Power Station, Tully Gorge, N. E. Qld, 8 / 2 / 2002, S. Fearn ” (QVMAG). Paratypes (5). 2 ♂♂, “ D. Perkins, Cardstone, Qld, Oct. – Dec. 1972 | ANIC 25 068137 ” (ANIC), “ Black light, Kareeya Power Station, Tully Gorge, N. E. Qld, 23 / 11 / 2001, S. Fearn ” (QVMAG); 3 ♀♀: “ 17 ° 19 ’ 06.63 ’’ S, 145 ° 30 ’ 43.58 ’’ E (GE), Wongabel St. For. Pavey Rd., QLD, 746 m, 17 APR 2013, DCF Rentz, Stop 13 - 14 | ANIC 25 066032 ” (ANIC), “ Black light, Kareeya Power Station, Tully Gorge, N. E. Qld, 8 / 2 / 2002, S. Fearn ” (QVMAG), “ Black light, Kareeya Power Station, Tully Gorge, N. E. Qld, 23 / 11 / 2001, S. Fearn ” (QVMAG). Other material examined (Fig. 16 D). Queensland: ANIC (7 ♂♂; 6 ♀♀): 2: 15.4 m up whitfield, Cairns, Oct. 1971, J. G. Brooks; 2: Boar PKT, Dec. 1969, J. G. Brooks; 2: Cardstone, May 1972, D. Perkins; 2: Cardstone, Oct. – Dec. 1972, D. Perkins; 2: Lake Barrine, Atherton, Tableland, Dec. 1985, J. Balderson; 8 km WbyN of Bald Hill, McIlwraith Range, Jun. – Jul. 1989, T. Weir and A. Calder; Cardstone, Feb. 1973, D. Perkins; Townsville, Dec. 1958, S Brock; QM (1 ♂): Windsor Tableland, NW of Mossman (site 1), Jan. 1994, G. and A. Daniels R. Eastwood; QVMAG (1 ♀): Kareeya Power Station, Tully Gorge, Mar. 2001, S. Fearn. Molecular data. BioSample ID: SAMN 12650119 (Holotype), SAMN 12650118 (Paratype).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B21FBE5FA83FC00FA43.taxon	description	(Figs. 5 E – I, 14 E, 15 H, 16 D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B21FBE5FA83FC00FA43.taxon	diagnosis	Diagnosis. This species is highly variable but can be separated from B. fearni by having obtuse angled frontoclypeal suture (Fig. 5 E), the mandibular surface with relatively long and dense hairs and relatively shorter elytra (about 4 times pronotum length in male, about 5 times length of pronotum in female). It is widely distributed in New South Wales and south-eastern Queensland.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B21FBE5FA83FC00FA43.taxon	materials_examined	Types. Pascoe (1864) had several specimens when describing Mallodon jejunum Pascoe, but only single specimen remains in the BNHM. Herein we designate this male specimen as the lectotype, with the following original labels “ Richmond R. | Pascoe Coll. 93 - 60. | Mallodon jejunum Pasc., Type ”. Macrotoma papyria Pascoe, holotype female deposited in BNHM, with the following original labels “ Macrotoma papyria Pasc. | Australia | Pascoe Coll. 93 - 60 ”. Material examined (Fig. 16 D). New South Wales: AM (6 ♂♂; 2 ♀♀): 4: Mt. Nardi, Dec. 1976, F. T. Fricke; 2: Dorrigo, Feb. 1981, D. A. Doolan; Alstonville, Dec. 1976, F. T. Fricke; Wilson R. Bridge nr. Bluff Point Picnic Area, Jan. 1999, B. J. Day; ANIC (14 ♂♂; 18 ♀♀): 8: Cambridge Plateau, {from Nov. 1996 to Feb. 1997}, S. Watkins; 6: Dorrigo NP., {from Feb. 1962 to Feb. 1967}, C. W. Frazier; 4: Cambridge Plateau, Jan. 1996, J. Bugeja; 2: Dorrigo NP., Jan. 1970, S. Misko; 2: Grafton, Nov. 1926; 2: Little Run Ck., at Potoroo Rd, Jan. 1996, [S. G. Watkins Coll.]; “ Big Scrub ” Whian Whian St. Forest NE Lismore, Jan. 1981, J. Powell; Brown’s lookout, Dingo SF, Dec. 1990, [S. G. Watkins Coll.]; Cherry Tree West SF., WSW Cassino, Nov. 1996, S. Watkins; Dingo Tops Forest Pk., Dingo SF, NW Wingham, Jan. 1993, [S. G. Watkins Coll.]; Dorrigo NP., Nov. 1971, R. Hardie; Pennefathers Rd., Cherry Tree SF, via Casino, Dec. 1996, S. Watkins; Richmond Rage S. F., Feb. 1983, T. Weir and A. Calder; Tooloom, Oct. 1961, I. B. F. Common and M. S. Upton; ASC (1 ♀): Yabbra SF, 394 Nth Yabbra Rd. and Castle Spur Road, Dec. 1999, Allen Sundholm; MV (1 ♀): [no data]; OAI (1 ♂; 5 ♀♀): 3: Grafton, 1926; Dorrigo, Dec. 1975, A. B. Rose; Grafton, W. W. Froggatt; Urbenville, Nov. 1945, K. L. Taylor; QDPC (1 ♀): Tooloom, H. Hacker; QM (5 ♂♂; 2 ♀♀): 3: Tooloom Plateau via Urbenville, Dec. 1972, I. Naumann; Kyogle, Nov. 1970, G. Monteith; Tooloom Plateau via Urbenville, Feb. 1983, Monteith and Thompson; Tooloom scrub via Urbenville, Oct. 1972, G. B. Monteith; Woodenbong, Dec. 1966, G. Monteith; RDKC (2 ♂♂; 2 ♀♀): 2: The Glade Picnic Area, Dorrigo NP, Dec. 2009, R. de Keyzer; 2: The Glade Picnic Area, Dorrigo NP, Jan. 2006, R. de Keyzer. Queensland: ANIC (12 ♂♂; 11 ♀♀): 4: St. Bernard’s Mt. Tamborine, Jan. 1961, C. W. Frazier; 3: Binna Burra, Lamington Nat. Pk., Dec. 1972, R. J. Kohout; 3: Mt. Glorious, Dec. 1968, R. Hardie; 2: Lamington NP., Feb. 1984, L. Lowman; 2: National Park, Jan. 1928, H. J. Carter; Lamington NP, IBISCA 700 A, Dec. 2008, G. Monteith; Lamington NP, IBISCA 700 c, Dec. 2008, G. Monteith; Lamington NP, O’Reily’s, Nov. 2011, H. Escalona; Lamington NP., Feb. 1983, H. Lowman; Lamington NP., Jan. 2009, G. Monteith; Lamington NP., May 1981, L. Lowman; Lamington NP., Nov. 2015, A. Zwick; Mt. Glorious, Jan. 1973, J. Kerr; National Park, Macpherson Rge., Jan. 1928, H. J. Carter; MV (2 ♂♂; 1 ♀): 2: National Park QLD, Dec. 1919; Mount Tamborine, Feb. 1943; QDPC (5 ♂♂; 2 ♀♀): 4: Yarraman, 1947, A. R. Brimblecombe; 2: Mount Tamborine, 2015, H. Hacker; Imbil, Apr. 1937, A. R. Brimblecombe; Yarraman, May 1947, A. R. Brimblecombe; Yarraman, Nov. 1947, A. R. Brimblecombe; QM (32 ♂♂; 17 ♀♀): 4: Lam [ington] NP, Feb. 1964, G. Monteith and H. A. Rose; 3: [Lamington] Nat. Pk, R. Illidge; 3: Davidson’s property hmstd, Tamborine Mtn, {from Nov. to Dec. 1992}, K. J. Lambkin; 3: Mount Glorious, Feb. 1969, B. Cantrell; 3: Mount Glorious, Jan. 1972, G. B. and S. R. Monteith; 3: Mount Glorious, Oct. 1971, B. Cantrell; 3: Mt. Glorious, Nov. 1992, R. Sheridan; 2: Lamington NP, R. Illidge; 2: Mount Glorious, Dec. 1966, G. Monteith; 2: Mt. Gannon summit via West Bunleigh, Jan. 1987, G. Monteith and D. Cook; [Lamington] Nat. Pk; Barkers Hills, Bunya Mtns, Jan. 1993, K. J. Lambkin; Bunya [Mtns], Jan. 1926; Bunya Mtns, Feb. 1964, B. Cantrell; Dayboro, Jan. 1971, G. B. Monteith; Lam [ington] Nat. Pk, Feb. 1963, G. Monteith; Lam [ington] NP, Feb. 1963, G. Monteith; Lamington NP., Feb. – Mar. 2009, Monteith; Mount Glorious, Feb. 1963; Mount Glorious, Mar. 1975, K. J. Lambkin; Mount Glorious, Oct. 1971, G. B. Monteith; Mt Glorious, Oct. 1971, B. Cantrell; Mt Mee St. For., Apr. 1972, G. B. Monteith; Mt Tenison-Woods via Mt Glorious, Mar. 1975, K. J. Lambkin; Mt. Glorious, Dec. 1970, A. Hiller; National Park, Dec. 1919, H. Hacker; National Park, Nov. 1920, H. Hacker; Tamborine Mountain, Nov. 1925, H. Hacker; Tamborine; Upper Laceys Ck, via Dayboro, Jan. 1971, G. B. Monteith.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B21FBE5FA83FC00FA43.taxon	discussion	Remarks. There are two distinctive morphological male forms in this species (Figs. 5 F, 5 G); one of them appearing to be a stronger masculine form (Fig. 5 F) with the pronotum surface bearing sexually dimorphic punctures, the lateral margin of pronotum bearing very small teeth, the antenna short and only extending to the middle of elytra but the antennal scape shorter and thicker. Both types of males have been collected in the same area, and even reared from the same log (personal observation, R. de Keyzer). We have found no significant differences in mitochondrial genome sequences and have kept them under the same species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7A1B21FBE5FA83FC00FA43.taxon	materials_examined	Molecular data. BioSample ID: SAMN 12650016, SAMN 12650115, SAMN 12650116, SAMN 12650117, SAMN 11080866.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7B1B3FFB83FA62FF15FDBE.taxon	description	(Figs. 6, 7, 14 F, 15 I, 16 E – F, 17 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7B1B3FFB83FA62FF15FDBE.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 25 – 60 mm long. Dorsum usually unicoloured, black to dark brown, sometimes with elytra slightly yellowish or reddish brown. Sexually dimorphic dense punctures on prothorax and scutellum present in some species only. Head narrower than prothorax; frontoclypeal suture arcuate or weakly angulate; median groove complete. Antennal tubercles moderately prominent, rounded apically. Eyes relatively large, weakly emarginate near antennal insertion, narrowly or moderately separated dorsally. Mandibles short, at most about half of head length, not sexually dimorphic, weakly bent towards ventral side in few species, unidentate apically. Antennal length variable. Scape at least half of head length, posteriorly extending beyond posterior margin of eye or even reaching anterior margin of pronotum, expanding apically, flat with blunt external edge; antennomere 3 longer than scape and antennomere 4 in most species, except in C. cephalotes. Mentum not fused to submentum. Terminal palpomere of maxillary and labial palps weakly expanded and apically slightly truncate. Prothorax transverse with lateral carina regularly dentate; anterior and posterior margins with complete beads. Pronotal disc usually glabrous, sometimes with very sparse and fine hairs, surface with dense coarse punctures. Prosternal process subparallel, extending beyond procoxae, rounded apically. Elytral surfaces moderately to strongly coriaceous, traces of wing venation present on few species; inner apical angle with short spine. Legs relatively long; femora with rows of sharp spines on ventral side. Protibia usually with rows of spines except in C. cephalotes, two or three sharp external projections apically and a pair of subequal spurs; tarsomere 3 moderately lobed; tarsomere 5 length varies in different species.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7B1B3FFB83FA62FF15FDBE.taxon	description	Description. Male. Length 25 – 60 mm. Dorsum usually uniformly brown or dark brown, sometimes with slightly reddish or yellowish elytra. Head approximately as long as wide, narrower than pronotum. Mandibles usually about 1 / 3 of head length, in C. cephalotes and C. gahani longer than half length of head; mandible weakly to moderately curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with one or two extra teeth on incisor edge. Labrum transverse, anterior margin pointed medially with very dense setae; labrum separated from clypeus by arcuate groove; dorsal surface of labrum and clypeus with sparse fine hairs. Frontoclypeal suture angulate or weakly arcuate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11 - segmented, filiform, usually extending to the middle of elytra, in C. cephalotes relatively short, only extending beyond elytral humeral angle, in C. cnemoplitoides extremely long, almost extending beyond elytra apex. Scape long and weakly flat, at least reaching beyond posteri- or margin of eyes, in C. cnemoplitoides reaching the anterior margin of pronotum; antennomere 3 as long as or slightly longer than scape, in C. cephalotes distinctly shorter than scape; and always longer than antennomere 4. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; narrowly to moderately separate dorsally. Submentum weakly emarginate at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate oval to weakly expanded, apex slightly truncate. Pronotum transverse, more or less trapezoid with posterior margin broader than anterior margin, sub rectangular in C. cephalotes; complete bead present on both anterior and posterior margin. Lateral carina with rather regular teeth, usually with prominent posterior angles except in C. cephalotes. Disc surface almost glabrous, sparse and fine hairs present near lateral margin in C. edulis; surface almost uniformly densely and coarsely punctate, with very weakly elevated and shiny area medially. Prosternum usually coriaceous punctate, in C. cephalotes only with few fine punctures; surface usually with sparse fine hairs, densely setose in C. edulis; hypomeron well-defined, as broad as or slightly narrower than prosternal process; prosternal process sub-parallel, strongly projecting towards mesoventrite, rounded apically. Mesoventral process relatively narrow, emarginate at apex. Ventral side of pterothorax and coxae covered with golden hairs. Metanepisternum weakly constricted near posterior margin. Scutellum rounded or weakly emarginate apically; surface variable. Elytral surfaces covered with irregular coriaceous sculpture, glabrous and usually shiny, relatively dull in C. cnemoplitoides; without or with very weak traces of venation presenting on basal elytra; elytral apices rounded usually with short sutural spine, spines absent in C. edulis; epipleuron almost complete, narrowing apically. Legs strong; femur subparallel, with rows of sharp spines on ventral side; tibia weakly expanded towards apex, surface rough and usually with rows of spines along outer margin, spines absent in C. cephalotes, ventral surface bearing sensory setae apically; each tibia with one or two apical projections and a pair of subequal spurs; tarsi moderately strong; tarsomere 3 moderately to strongly lobed; tarsomere 5 usually as long as or shorter than tarsomeres 1 and 2 combined, longer than tarsomeres 1 and 2 combined in C. cephalotes. Abdominal ventrites 1 – 4 with sexual setal patches except in C. australis and C. cnemoplitoides, the sizes of setal patches and length of setae varies in different species; setae present along the edge of ventrites 1 – 5, especially on the apex of ventrite 5. Male genitalia (Fig. 14 F). Tegmen longer than sternite VII; parameres moderately long, rounded apically with triangular projections at the base; penis usually as long as tegmen, longer in C. argodi and C. cephalotes; dorsal apex broadly rounded while ventral apex narrowly rounded. Female. Body usually more elongate than in male, sexual setal patches absent on abdominal ventrites. Ovipositor long; apical sclerosed part usually less than half length of baculus, in C. edulis about 2 / 3 length of baculus (Fig. 15 I); distal gonocoxites very short and small, moderately to strongly truncate apically, stylus large, inserted laterally and close to gonocoxite apex, expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB7B1B3FFB83FA62FF15FDBE.taxon	discussion	Remarks. Based on the molecular results (Jin et al. 2020), we excluded some species from the Cnemoplites, and placed them in two separated genera: Hermerius and Hagrides. We have subsequently found morphological evidence to support these molecular clades (discussed below). The monophyly of the clade with the remaining species of Cnemoplites was not strongly supported on the molecular tree, and we have not been able to find morphological characters to support that clade, and its monophyly requires a further research.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3FF8BCF94BFB41FAAC.taxon	description	(Figs. 6 A – D, 16 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3FF8BCF94BFB41FAAC.taxon	diagnosis	Diagnosis. Cnemoplites. argodi can be separated from other Cnemoplites species by the relatively smaller body length, the elytra dark brown to almost black with very dense irregular sculpture.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3FF8BCF94BFB41FAAC.taxon	description	Description. Length 30 – 35 mm. Male. Antennae extending to or slightly beyond apices of elytra. Antennal scape and antennomere 3 thicker than the following segments; antennomere 3 slightly longer than scape and distinctly longer than antennomere 4. Prothorax shiny with irregular coarse sculpture; scutellum shiny, punctate. Elytra with very coarse and irregular sculpture, in lateral areas with additional small granules. Abdominal sexual setae long and erect, the setal area transverse, relatively narrow and short. Female. Antennae extending beyond middle length of elytra; sexual setae on abdominal ventrites absent.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3FF8BCF94BFB41FAAC.taxon	materials_examined	Types. Lectotype male (BNHM- 010799749) deposit- ed in BNHM is designated here, with the following labels “ Moreton Bay | Cnemoplites argodi, Type, Lameere ”. Paralectotype male (BNHM- 010799859) deposited in BNHM, with the following labels “ Moreton Bay | Cnemoplites argodi, Cotype. Lameere ”; and paralectotype male deposited in MNHN, with the following labels “ Salt Lakes, S. Rustral [Austral] | Cnemoplites argodi, ♂, Type, Lameere ” (specimen without abdomen). Material examined (Fig. 16 E). New South Wales: AM (1 ♂): Ropes Creek; ANIC (1 ♂; 2 ♀♀): Ogilvie Trail, Banyabba NR, SSW Whiporie, Nov. 2000, S. Watkins; Olives Swamp Myrtle SF., Dec. 1996; Sydney, 1899, W. W. F.; OAI (1 ♂): Graman, Dec. 1960, Bourke, T. V. Queensland: QM (4 ♂♂; 2 ♀♀): 2: “ Allinga ” Chinchilla, Lithgow, G.; 2: Blackdown Tableland, Stoney Ck. Via Dingo, Dec. 1985, S. Hamlet; Blackdown Tableland, Via Dingo, Dec. 1978, Maywald and Czechura; Durikai, 2 km NE, Dec. 2001, L. Popple and J. Moss; RDKC (1 ♂): Dunmore S. F., Dec. 1987, A. Hiller. Molecular data. BioSample ID: SAMN 12650053, SAMN 12650106, SAMN 12650107.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3CFBC4FA0FFB41F968.taxon	description	(Figs. 7 E – H, 16 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3CFBC4FA0FFB41F968.taxon	diagnosis	Diagnosis. Cnemoplites. australis can be separated from other Cnemoplites species by the antennomere 3 very long, about twice as long as antennomere 4 and the elytra surface with irregular sculptures near scutellum, and very fine dense nodules on remaining areas. It is similar to C. cnemoplitoides (Thomson), but can be easily recognised by the following characters: protibia short, male elytra about 3 × of protibia length, female over 3 × of protibia length, distributed from Tasmania to south-eastern Queensland.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3CFBC4FA0FFB41F968.taxon	description	Description. Length 25 – 45 mm. Male. Antennae extending beyond the middle of elytra. Antennomere 3 about 1.5 × length of scape, both stronger than the remaining segments; prothorax dorsal and ventral surfaces with the combination of coarse punctures and very fine punctures; scutellum densely punctate; abdominal ventrites relatively smooth and glabrous, only bearing fine punctures and sparse short adpressed setae. Female. antennal scape slightly longer and stronger than antennomere 3; pronotum with dense coarse punctures, each bearing short hair; scutellum shiny without distinct punctures; femur surface with less spines and nodules; abdominal ventrite 5 emarginate at apex.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB651B3CFBC4FA0FFB41F968.taxon	materials_examined	Type. Macrotoma servilis, Holotype female deposited in BNHM (BNHM- 010799847), with the following labels “ Macrotoma servilis Type ♀, Pasc., Melbourne | Pascoe Coll. 93 - 60 ”. Material examined (Fig. 16 F). Tasmania: ANIC (88 ♂♂; 33 ♀♀): 52: Kelton, Longford, {from Jan. 1982 to Feb. 1985}, S. Fearn; 42: Greens Beach, {from Feb. 1984 to Feb. 1985}, S. Fearn; 20: Beechford, {from Jan. 1977 to Jan. 1982}, S. Fearn; 2: Park Beach, near Sorrel, Apr. 1987, S. Fearn; Beechford, Jan. 1979; Freycinet Nat. Pk., Feb. 1963, I. B. F. Common and M. S. Upton; Kelso; Kelton, Longford, Feb. 1982, A. Walker; Whitemark, Flinders Island, Mar. 1952, J. H. Calaby; MV (1 ♂; 2 ♀♀): 3: Flinders Island. Victoria: ANIC (4 ♂♂; 10 ♀♀): 3: Lakes Entrance, Feb. 1955, M. and G. Coulson; 3: Rosebud, Jan. 1962, Le Souef; Blairgowrie, Feb. 1967, J. C. Le Souef; Manns Beach, Feb. 1963, M. J. Coulson; Maryvale, Mar. 1969, G. May; Red Hill, Jan. 1972, D. R. Holmes; Rosebud, Jan. 1964, Le Souef; Seaford, Feb. 1952, D. Smith; Waratah Bay, Mar. 1969, G. May; Warragul, Jan. 1972, C. Elton; MV (4 ♂♂ 1 ♀): Point Lonsdale, Feb. 1941; Inverloch, Jan. 1936; Inverloch, Jan. 1937; North Melbourne; nr. Melb; QM (1 ♀): Cann Riv., Jan. 1967, G. Monteith. South Australia: ANIC (1 ♂) Mt. Barker, Dec. 1980, D. P. Carne. New South Wales: AM (2 ♂♂; 1 ♀): 2: Barrington Tops NP, Jan. 1987, R. de Keyzer; Church Point, Mar. 1971, D. A. Doolan; ANIC (20 ♂♂; 22 ♀♀): 4: Bowley Point, {from Dec. 1998 to Jan. 1999}, D. C. F. Rentz; 3: Pennefathers Rd, Cherry Tree State Forest via Casino, Jan. 1997, Watkins; 3: Wingham (Lots 11 and 12 Combined St), {from Dec. 1991 to Jul. 1992}, J. Stockard; 2: Bermagui, Jan. 2015, B. P. Moore; 2: Congo, Dec. 1980, Upton, M. S.; 2: Green cape, Jan. 1968, L. Hall; 2: Park College, Sydney, Apr. 1905, W. W. Froggatt; Bawley Point, Jan. 1997, D. C. F. Rentz and K. McCarron; Benandra SF, Jan. 1959, K. G. Campbell; Botanical Gardens, Sydney, Dec. 1904; Broulee, Dec. 1975, D. Ferguson; Broulee, Dec. 2003, B. Boyed; Broulee, Jan. 1974, T. E. Bellas; Burrawarra Pt. [Burrewarra Pt.], Jan. 1978, M. J. Coulson; Cambridge Plateau, Jan. 1996, J. Bugeja; Congo, Jan. 1982, H. S. Upton; Durras North, Dec. 1978, H. Cameron; Durras, Jan. 1977, D. J. Ferguson; Jan. 1974, M. Dendy; La Perouse, Apr. 1986, E. E. Taylor; Minnamurra, Dec. 1937, A. H. W.; Mollymook, nr. Ulladulla, Jan. 1971, E. B. Britton; Moruya Heads, Jan. 1937, F. G. Holdaway; Mossy Pt., Jan. 1993, D. F. Waterhouse; Mount Keira Clive Bissell Drive, near Mount Ousley Rd., Illawarra Escarpment State Conservation area, Jul. 2004, R. de Keyzer; Myall Lakes, Dec. 1987, M. Dixon; Pambula, Jan. 1964, M. Key; Pennefathers Rd., CherryTree SF, SE Mallanganee, Nov. 1997, Watkins; Rosedale, Jan. 1999, D. F. Waterhouse; Wallaga Lake, Jan. 1981, H. P. Black; Wingham Brush, Jan. 1984, G. Williams; MV (2 ♂♂; 1 ♀): Otford, Mar. 1902; Sydney; [no data]; OAI (8 ♂♂; 12 ♀♀): 3: Lisarow, Jun. 1956, K. M. Moore; 2: Mt Keira, near Wollongong, D. A. Walsh; 2: Narara, near Gosford, Dec. 1949, I. Mosse-Robinson; Benandarah State Forest, Jan. 1959, K. G. Campbell; La Perouse, Feb. 1982, E. E. Taylor; Narara, Dec. 1949, K. D. Fairey; Nowra, Sep. 1986, G. R. Brown and H. M. Holmes; Palm Beach, Dec. 1934, Mr Bilby; Park Beach, Coffs Harbour, Dec. 1998, J. A. Macdonald; Pearl Beach, near Woy Woy, Feb. 1995, A. E. Westcott and F. Swindley; Roseville, near Sydney, Feb. 1971, C. E. Chadwick; Royal Botanic Gardens, Sydney, Jan. 1904; Uralla, Feb. 1937, K. L. Taylor; Uralla, Jan. 1938, K. L. Taylor; Vaucluse, Jan. – Dec. 1927, A. Eady; Vaucluse, near Sydney, Apr. 1957; QM (2 ♂♂; 3 ♀♀): 2: Woollongong, Feb. 1967, G. Monteith; Glen Innes, Jun. 1913, C. Deane; Bulahdelah SF, Bulahdelah, Jan. 1966, B. Cantrell; Mt Keira via Wollongong, Feb. 1967, G. Monteith. Queensland: ANIC (3 ♂♂; 7 ♀♀): 5: Binna Burra, Lamington Nat. Pk., Dec. 1972, R. J. Kohout; 3: Binna Burra, Lamington Nat. Pk., Dec. 1973, R. J. Kohout; Bongan [Bongeen], Oct. 1932, Fuller; Eukey, Apr. 1944; MV (2 ♂♂; 1 ♀): Eukey, Jan. 1934; Mount Tamborine, Jan. 1912; Stanthorpe; QDPC (2 ♂♂): Baldy Mountain Road, near Atherton, Jan. 1978, R. I. Storey; Crater Lakes National Park, Jan. 1967, R. J. Elder; QM (2 ♂♂; 2 ♀♀): Birthday Ck via Paluma, Jan. 1973, B. Cantrell; Davidson’s property hmstd, Tamborine Mtn, Dec. 1992, K. J. Lambkin; Glen Aplin, Feb. 1964, P. Kerridge; Landsborough, Nov. 1995, C. Cowell; QVMAG (1 ♂): The Gap, Brisbane, Feb. 1994, S. Fearn. Molecular data. BioSample ID: SAMN 12650017, SAMN 12650018, SAMN 12650057, SAMN 12650058, SAMN 12650059, SAMN 12650061, SAMN 12650062, SAMN 12650063, SAMN 11080874.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB661B3DFBC5F94BFA6AFE83.taxon	description	(Figs. 6 E – H, 17 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB661B3DFBC5F94BFA6AFE83.taxon	diagnosis	Diagnosis. C. cephalotes can be separated to oth- er Cnemoplites species by the following characters: antennal scape much longer than other antennal segments; pronotum more or less squared, dorsum coarsely punctate without sexual punctures; legs strong and broad. Males can be recognised by the dense and long sexual setae on abdominal ventrites, setal area covering the whole ventrite (Fig. 6 H); several small spines on ventral side of protibia.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB661B3DFBC5F94BFA6AFE83.taxon	description	Description. Length 30 – 60 mm. Male. Antennae short, extending just beyond the elytral humeral angles. Antennal scape longer and thicker than antennomere 3. Pronotum completely covered with dense and coarse punctures; scutellum shiny and smooth. Elytra with coarse and irregular sculptures, without additional small granules. Abdominal sexual setae long and almost adpressed, the setal area almost covering the whole ventrite. Tarsomere 5 elongate, almost as long as tarsomeres 1 – 3 combined. Female. Mandibles less thick than in male. Pronotum with more or less smooth area medially; sexual setae on abdominal ventrites absent.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB661B3DFBC5F94BFA6AFE83.taxon	materials_examined	Type. Mallodon cephalotes Pascoe, Holotype male deposited in BNHM (BNHM- 010799747), with following labels “ Mallodon cephalotes | Queensland | Pascoe Coll. 93 - 30 | Cnemoplites cephalotes Pasc ”. Material examined (Fig. 17 A). New South Wales: AM (3 ♂♂; 2 ♀♀): 2: Wheeny Ck., N of Kurmond, Feb. 1992, M. S. and B. J. Moulds; Lansdowne, Feb. 1981, G. Williams; Carningford, Jan. 1963, W. Newham; Gosford, Nov. 1975, F. T. Fricke; ANIC (1 ♂): Moonon Brook, Feb. 2015, B. Lessard, A. Landford, D. Ferguson and D. Yeates; MV (2 ♀♀): Coraki; Inverell; OAI (2 ♀♀): Erina Heights, Feb. 1953, J. G. Gellatley; Lisarow, Feb. 1952, R. Moore; QM (1 ♀): Barrington House via Salisbury, Feb. 1965, G. Monteith. Queensland: ANIC (2 ♂♂; 10 ♀♀): 3: Kuranda, 19 Butler Dr., Feb. 2016, D. C. F. Rentz; Bluff Rg., Foothills, via Biggenden, Aug. 1975, H. Frauca; Garradunga, Nov. 1996, J. Hasenpusch; Innisfail, Garradunga, Dec. 1995, J. Hasenpusch; Kuranda, 19 Butler Dr., Feb. 2010, D. C. F. Rentz; Mission Bay, Mar. 1993, G. O’Reilly; Mossman Gorge, Dec. 1972, P. Zborowski; Polly Ck., Garradunga, Jan. 1999, J. Hasenpusch; Polly Ck., Garradunga, Nov. 1996, J. Hasenpusch; Tully Falls, Jan. 1972, P. Zborowski; QDPC (1 ♂; 2 ♀♀): Thulimbah, R. A. McLachlan; Burleigh, H. Jarvis; Oxley, Brisbane, Feb. 1980, K. Houston; QM (7 ♂♂ 2 ♀♀): 4: Mt. Goonaneman, via Childers, Feb. 1981, G. B. Monteith; Blackdown Tableland via Dingo, Feb. 1981, G. B. Monteith; Garradunga, Innisfail, Feb. 1991, J. Hasenpusch; Lake Boronto (= Wincheura) Newcastle Bay, Jan. – Feb. 1975, Monteith; Paluma Range, Feb. 1981, S. J. Johnson; Yuleba SF, site 1, Mar. – Feb. 2002, Monteith and Wright; QVMAG (4 ♂♂; 1 ♀♀): 4: The Gap, Brisbane, Mar. 1994, S. Fearn.; Kareeya Power Station, Tully, Jan. 2001, S. Fearn; RDKC (5 ♂♂ 2 ♀♀): 4: Polly Ck., Garradunga, {from Jan. 2011 to Feb. 2012}, J. Hasenpusch; J. Transverse Track, Conondale NP, 1 km from Bellthorpe, Jan. 2014, A. Scott; Cathu Lookout, Cathu S. F., Feb. 2016, A. Sundholm and J. Bugeja; Diwan, near Cow Bay, Jan. 2008, S. Lamond. Molecular data. BioSample ID: SAMN 12650098.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB671B3DFB1DFE25FB41F8AB.taxon	description	(Figs. 7 I – L, 16 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB671B3DFB1DFE25FB41F8AB.taxon	diagnosis	Diagnosis. This species is very similar to C. australis, as they both with tiny nodules present on elytra and long antennomere 3, and males without sexual patch on abdominal ventrites, but can be recognised by the following characters: protibia longer, elytra only 2 – 2.5 times of protibia length in male, elytra less than 3 times of protibia length in female.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB671B3DFB1DFE25FB41F8AB.taxon	description	Description. Length 30 – 55 mm. Male. Antennae extending to almost apices of elytra. Antennal scape and antennomere 3 thicker than the following segments. Prothorax almost rectangular, matt with dense punctures in addition to regular deep sculpture; scutellum densely punctate. Abdominal ventrites smooth and shiny. Female. Antenna slightly shorter. Prothorax trapezoidal shiny with regular deep sculpture; scutellum shiny and smooth.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB671B3DFB1DFE25FB41F8AB.taxon	materials_examined	Type. Macrotoma cnemoplitoides Thomson, Holotype male deposited in MNHN, with the following labels “ Moret. Bay | Th. TYPE | 143 ”. Material examined (Fig. 16 F). Queensland: AM (1 ♂): Innisfail, Dec. 1975, F. T. Fricke; ANIC (24 ♂♂; 19 ♀♀): 27: Kuranda, 19 Butler Dr., {from 2003 to 2016}, D. C. F. Rentz; 2: Cardstone, Oct. – Dec. 1972, D. Perkins; 2: Mission Beach, Nov. 1976, G. O’Reilly; 2: Mt. Hypipamee NP, Nov. 1998, A. A. Calder; 10 km SSW of Cape York, Oct. 1992, T. A. Weir and P. Zborowski; Black Mtn Rd., 5 mi. N of Kuranda, Dec. 1968, Britton, Brooks and Misko; Cairns, Dec. 1971, A. Walford-Higgins; Cairns, F. H. Taylor; Julatten, Dec. 1997, B. P. Moore; Julatten, Jan. 2008, B. P. Moore; Kuranda, J. Hasenpusch; Paluma, Jan. 1970, E. B. Britton; Polly Ck., Garradunga, Dec. 1996, J. Hasenpusch; Polly Ck., Garradunga, Dec. 2010, A. Zwick; QDPC (3 ♂♂): 2: Windsor Tablelands, via Mount Carbine, Jan. 1978, R. I. Storey; Kirrama Range, via Kennedy, Dec. 1977, R. I. Storey; QM (3 ♂♂): Graham Range, Monteith, Thompson and Cook; Nr. Plane Crash, 11 km NW Mossman, Dec. 1989, [ANZSES Expedition]; West Claudie R., Iron Range, Dec. 1985, G. Monteith and D. Cook; QVMA G (4 ♂♂; 8 ♀♀): 11: Kareeya Power Station, Tully Gorge, Dec. 2000, S. Fearn; Paluma, Dec. 2000, S. Fearn. Molecular data. BioSample ID: SAMN 12650060, SAMN 12650120, SAMN 12650121.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB601B3BF8B9FF12FE45FCF7.taxon	description	(Figs. 7 A – D, 14 F, 15 I, 16 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB601B3BF8B9FF12FE45FCF7.taxon	diagnosis	Diagnosis. C. edulis can be separated from other Cnemoplites species by the elytral apicosutural angle rounded, the hypomeron relatively narrow, about half the width of prosternal process and the tarsomere 3 with very broad lobe, about twice the width of tarsomere 2.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB601B3BF8B9FF12FE45FCF7.taxon	description	Description. Length 25 – 50 mm. Male. Antennae extending beyond the middle of elytra, antennal scape and antennomere 3 much thicker than the following segments; pro- and mesotibia strongly spinose on lateral margins, metatibia only with few spines; abdominal ventrites 1 – 4 with extensive setal patches. Female. Only antennal scape slightly thicker than the following segments; pro- and mesotibia with few spines, metatibia glabrous along external margin; sexual setae on abdominal ventrites absent.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB601B3BF8B9FF12FE45FCF7.taxon	materials_examined	Types. Newman (1842) described this species from a series of specimens purchased from Mr. Edmund Higgins from Port Phillip. Since no specimen from the original series has been found in the BMNH, the neotype of C. edulis is designated here to stabilise taxonomic status of this species. Neotype male deposited in ANIC, with the following labels “ Queanbeyan, NSW, 30 Jan. 1992, A. M. E. Roach, at light | ANIC 25 - 066984 ”. The mitochondrial genome was generated for this type, NCBI Access ID as MK 614556, and BioSample ID as SAMN 11080884. Material examined (Fig. 16 E). Victoria: ANIC (19 ♂♂; 12 ♀♀): 5: Red Hill, Jan. 1974, D. R. Holmes; 3: Mizpah Rd., Jan. 1977, C. Elton; 2: Hazelwood, Feb. 1949, J. H. Courtenay; Gould, Tyers River, Feb. 1960, G. T. Coulson; Hazelwood, Feb. 1948, J. H. Courtenay; Hazelwood, Feb. 1950, J. H. Courtenay; Malvern, Ferr; Maryvale, Feb. 1969, G. May; Moe, Feb. 1948, C. G. L. Gooding; Moe, Feb. 1949, C. G. L. Gooding; Mt. Beauty, Dec. 1960, C. E. Wilkinson; Mt. Cannibal, Feb. 1987, W. N. B. Quick; Newboraugh, Jan. 1956, G. T. Coulson; Orrvale, Mar. 1969, H. Carins; Red Hill, Mar. 1958, B. P. Moore; Warragul, Feb. 1959, L. Thar; Warragul, Feb. 1961, C. G. L. Gooding; Warragul, Feb. 1963, P. J; Warragul, Feb. 1963, R. P. W; Warragul, Feb. 1963, R. R. W.; Warragul, Feb. 1965, C. G. L. Gooding; Warragul, Feb. 1969, F. Zinner; Warragul, Jan. 1968, Woods and Reeves; Warragul, Jan. 1972, J. G. Brooks; MV (10 ♂♂; 14 ♀♀): 2: Mitcham, Jan. 1946; 2: North Melbourne; 2: [no data]; Cranbourne, Dec. 1947, E. Smith; Croydon; J. E. Dixon; Macedon, Jan. 1903; Meredith, Feb. 1959; Montmorency, Feb. 1969, L. J. Cookson; Montmorency, Mar. 1969, L. J. Cookson; Mordialloc, Jan. 1895; Mordialloc; nr. Melb, Feb. 1937; nr. Melb, Jan. 1923; Porepunkah, Jan. 1960; The basin, Feb. 1958; Tyabb, Jan. 1919; Warrandyte, Feb. 1996, L. J. Cookson; Warrandyte, Feb. 1998, L. J. Cookson; Warrandyte, Feb. 2002, L. J. Cookson; Woori Yallock, Dec. 1939; QM (4 ♂♂; 4 ♀♀): 2: Ferntree Gully; 2: Maffra, Jan. 1966, B. Cantrell; Ferntree Gully, Mar. 1932, A. H. Westley; river x-ing, 10 miles N of Valencia Ck via Maffra, Jan. 1966, T. Weir; Winchelsea; Yallourn; RDK C (1 ♂): Airey’s Inlet, Jan. 1998, P. J. Krake. Australian Capital Territory: ANIC (19 ♂♂; 3 ♀♀): 14: Black Mt., {from 1965 to 1974}, {E. Britton; B. P. Moore; K. R. Pullen; I. F. B. Common; E. Nielsen; M. Neave and M. Yee}; Acton, Feb. 1949, W. Price; Canberra, Dec. 1988, J. Cotterill; Canberra, Jan. 1958, W. J. M. Vestjens; Lyneham, Feb. 1974, B. P. Moore; Mt Majura, Feb. 1998, A. Polak; No. 9 glasshouse, CSIRO, Black Mt., Feb. 1969, J. A. Bull; Orroral Vally, Feb. 1969, M. Carins; Red Hill, Feb. 1974, H. Rozdarz; OAI (1 ♂): Canberra, Mar. 1992, A. E. Westcott. New South Wales: AM (4 ♂♂; 1 ♀♀): 2: Barren grounds, Jan. 1966; 7.4 km W of Tomerong, Jan. 1979, G. Daniels; Bald Rock Nat. Park, Jan. 1978, G. Daniels; Hanging Rock, Jan. 1966, C. N. Smithers; ANIC (22 ♂♂; 31 ♀♀): 10: Bawley Point, {from 1995 to 1999}, D. C. F. Rentz and K. McCarron; 3: Cumberland NF, {from 1988 to 1990}, {E. E. Taylor; R. H. Eldridge}; 2: “ Warragun ” nr Braidwood, Jan. 1984, M. Whitten; 2: Armidale, Feb. 1961, A. Glover; 2: Burra Burra Pnt., Dec. 1972, Coulson; 2: Burrewarra Pt., Jan. 1978, M. J. Coulson; 2: Durras Nth, Nov. 1964, I. Cameron; 2: Fairfield, Dec. 1923, M. Fuller; 2: Ogilvie Trall, Barryabba SF, N of Grafton, Dec. 1997, Watkins; 3.3 km SSWest on North Belmore Rd., Busby’s Flat, Dec. 2000, Watkins; Armidale, Jan. 1964, J. Frazier; Barrewarra Pt., Jan. 1978, M. J. Coulson; Beecroft, Jan. 1966, M. R. Allen; Broulee, Jan. 1969, W. J. M. Vestjens; Cambridge Plateau, Jan. 1996, J. Bugeja; Carlingford NE Parramatta, Dec. 1973, J. Walk- er; Chiswick Expt Stn, Dec. 1965, R. J. Roberts; Congo, Dec. 1981, Upton, M. S.; Fortis Creek SF, N Graton, Dec. 1996, S. Watkins; Hornsby Heights, Feb. 1972, J. King; Lilyvale, Dec. 1971, D. A. Doolan; Lisarow, Jan. 1958, K. M. Moore; London Foundation, Kioloa, Jan. 1981, J. Conran; Mt. Wilson, Jun. 1967, R. A. Faragher; Powerful Owl Resv., Girand SF, West of Drake, Dec. 1998, Watkins; MV (1 ♂; 2 ♀♀): Dorrigo; Sydney; Sydney, Dec. 1902; OAI (10 ♂♂; 9 ♀♀): 6: Barren Ground Nature Reserve, Jan. 1966, C. E. Chadwick; 2: Kosciusko, Waste Point, Jan. 1966, C. E. Chadwick; Arding, 10 - 15 km S / SW Armidale, Jan. 1987, C. R. Haywood; Armidale, Dec. 1980, C. R. Haywood; Borenore, Feb. 1998, P. Thornberry; Cabramatta, near Sydney, Jan. 1958, M. I. Nikitin; Meryla, Jan. 1950, J. E. Wells; Normanhurst, near Sydney, Oct. 1958, F. Bagshaw; Orange Agricultural Institute grounds, Orange, Dec. 1998, P. Slattery; Orange Agricultural Institute grounds, Orange, Feb. 1992, L. Penrose; Orange, Mar. 1982, W. G. Thwaite; Patonga, near Woy Woy, Mar. 1966, N. Douglas; Swallowfield, [14.5 km E by N of] Armidale, Dec. 1980, C. R. Haywood; QM (3 ♂♂): Glencoe, Jan. 1990, M. Goombs; Rivertree, E. Sutton; Woodford, Jan. 1909, G. A. Waterhouse. Queensland: ANIC (1 ♂; 2 ♀♀): 3: Binna Burra, Lamington Nat. Pk., {from 1972 to 1973}, R. J. Kohout; QM (1 ♂; 1 ♀): 2: Brisbane; QVMAG (2 ♀♀): The Gap, Brisbane, {from 1994 to 1995}; S. Fearn. Molecular data. BioSample ID: SAMN 11080884 (Neotype), SAMN 12650094.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB601B3BF8B9FF12FE45FCF7.taxon	discussion	Remarks. This species is very common in southern part of Australia (VIC, ACT, NSW), usually found under the bark of Eucalyptus and attracted to MV light. During the sample examinations and molecular data analyses, we found out that some specimens collected in Queensland (i. e. Cnemoplites sp 6 in Jin et al. 2020) are morphologically similar but can be separated from C. edulis by sp 6 with smaller lobe on tarsomere 3 and less spinose tibiae. Molecular data also supported sp 6 being distant to C. edulis, but more specimens and molecular data are needed to resolve the status of the Queensland populations.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B3BF8B3FCC9FADFFE64.taxon	description	(Figs. 6 I – L, 17 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B3BF8B3FCC9FADFFE64.taxon	diagnosis	Diagnosis. Cnemoplites gahani can be separated from other Cnemoplites species by the following characters: pronotum with at least some smooth area medially in both sexes; antennal scape as long as or slightly longer than antennomere 3; antennomere 3 about 1.5 times as long as antennomere 4; antennomere 11 elongate, almost twice of antennomere 10 length.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B3BF8B3FCC9FADFFE64.taxon	description	Description. Male. Length 50 – 60 mm. Antennae extending to the middle of elytra; antennal scape extending beyond the posterior margin of eyes; antennomeres 3 – 8 weakly expanded apically; tibia bearing very dense hairs especially on ventral side; abdominal ventrites 1 – 4 covered with very dense and extensive setal patches. Female. Identification uncertain.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B3BF8B3FCC9FADFFE64.taxon	materials_examined	Types. Lectotype male deposited in BNHM designated here (BNHM- 010799796, Figs. 6 I – J), with label “ Hermannsburg, Central Australia, H. J. Hillier, 1911 - 311 ”. Other Material examined (Fig. 17 A). South Australia: AM (1 ♂): S. Australia, A. H. C. Seitz. Queensland: ANIC (1 ♂): Edungalba, 1957, E. E. A.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B3BF8B3FCC9FADFFE64.taxon	discussion	Remarks. Although the female syntype (BNHM- 010799817, Fig. 6 K) bears the identical collecting labels as the male lectotype, we are uncertain if it belongs to the same species and have excluded it from the type series. This female differs from the male lectotype by having shorter antennal scape, shorter antennomeres 3 and 11, broader hypomeron; almost smooth elytra, and elongate tarsomere 1. This specimen is morphologically identical to the female of Eurynassa tuberculicollis sp. nov. (Fig. 8 K), occurring in the same area. Taxonomic status of this species is uncertain as we have no molecular samples and very limited material for disposal. In addition to the lectotype we have found only two males that largely agree with the type but no females were collected associated with them.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B38FB37FE46FBF0FD1F.taxon	description	(Figs. 5 J – M, 14 G, 15 J, 17 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B38FB37FE46FBF0FD1F.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 25 – 50 mm long. Dorsum uniformly reddish brown to dark brown. Sexually dimorphic dense fine punctures present near lateral margins of pronotum, ventral side of prothorax and mesoventrite. Head as broad as prothorax; frontoclypeal suture arcuate or broadly angulate; median groove complete. Antennal tubercles almost flat, rounded apically. Eyes transverse, weakly emarginate near antennal insertion, relatively distant dorsally. Mandibles shorter than head, sexually dimorphic longer and thicker in male than in female, weakly arcuate and unidentate apically. Antenna reaching mid elytra in males. Scape about half of head length, posteriorly not extending to posterior margin of eye, gradually expanded apically; antennomere 3 shorter than scape and about as long as 4. Mentum not fused to submentum (Fig. 5 M). Terminal palpomere of maxillary and labial palps weakly expanded and apically round- ed. Prothorax transverse with lateral carina weakly dentate; anterior margin without bead, posterior margin with complete bead. Pronotal disc shiny and almost smooth medially surrounded by large and deep, sometimes merged punctures laterally and posteriorly separating disc from irregular lateral densely punctate areas. Prosternal process weakly expanded beyond procoxae, extending to mesoventrite, narrowly pointed apically. Elytra surface almost glabrous and shiny; inner apices with acute sutural angles. Legs strong, rows of tiny spines present on femora while absent on tibia. Protibia smooth externally with two sharp apical projections and pair of subequal spurs; lobes on tarsomere 3 narrow and short, tarsomere 5 much longer than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B38FB37FE46FBF0FD1F.taxon	description	Description. Male. Length 25 – 50 mm. Dorsum globrous and shiny, uniformly reddish brown to dark brown. Head oval, slightly narrower than pronotum. Mandibles about as long as head capsule, dorsally bearing sparse hairs, apex curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with one extra tooth on incisor edge. Labrum transverse, anterior margin pointed medially with relatively dense setae; labrum separated from clypeus by a deep groove; dorsal surface of clypeus with sparse long setae. Frontoclypeal suture almost arcuate or broadly angulate; median groove distinct and complete. Antennal tubercles relatively flat, very distant to each other, apex rounded. Antennae 11 - segmented, filiform, extending to mid elytra; scapes short, as long as eye length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; relatively distant dorsally. Submentum weakly curved at apex, mentum well exposed; gular area triangular and coarsely punctate. Terminal palpomere of maxillary and labial palps weakly expanded and apically rounded. Pronotum transverse with anterior margin emarginate medially; anterior margin without bead, posterior margin with complete bead; lateral carina weakly dentate. Disc surface shiny, glabrous and smooth medially, with deep and fine punctures on lateral area and two small patches of coarsely punctate area posteriorly. Prosternum bearing uniformly deep fine punctures; hypomeron well-defined, broader than prosternal process; prosternal process moderately expand beyond procoxae, with median lobe projecting towards mesoventrite, narrowly rounded apically. Mesoventral process relatively broad, apex emarginate medially. Ventral side of pterothorax and coxae covered with golden hairs. Metanepisternum constricted at base. Scutellum narrowly rounded at apex, surface smooth and shiny. Elytra shiny with very fine and weak irregular coriaceous sculpture, three traces of venation present on apical half of elytra; elytral apices rounded with small but sharp sutural angulation; epipleuron almost complete, very narrowing apically. Legs strong, femur broad and relatively flattened, weakly constricted at both ends, ventral side with rows of tiny spines; each tibia with two distinct apical spines and a pair of strong spurs. Tarsi slender; tarsomere 3 with narrow and short lobes; tarsomere 5 much longer than tarsomere 1 and 2 combined. Abdominal ventrites smooth, only bearing dense long setae along the edges, especially on the apex of ventrite 5. Male genitalia (Fig. 14 G). Tegmen longer than sternite VII; parameres short, less than 0.1 times length of entire tegmen, truncate apically with small triangular projections at the base; penis longer than tegmen, dorsal apex broadly rounded while ventral apex narrowly pointed. Female. Mandibles much shorter and narrower than in male; pronotum weakly constricted near anterior margin, with coarsely punctate, irregular lateral area devoid of very sense punctation; prosternum and hypomeron with sparse fine punctures and sparse hairs, relatively shiny. Ovipositor (Fig. 15 J) long, apical sclerosed part half of baculus length; distal gonocoxites short and small; stylus inserted laterally and close to gonocoxite apex, relatively long and gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB611B38FB37FE46FBF0FD1F.taxon	discussion	Remarks. Cryptipus can be distinguished from the remaining genera of the Australian Macrotomini by having the uniformly reddish-brown body, relatively glabrous and smooth elytra, surface only with very fine and shallow punctures usually very hardly seen, the elongate tarsomere 5, which is longer than tarsomeres 1 – 3 combined, and the tibia without any teeth or spines along external margins. Males are recognised by longer and stronger mandibles. Cryptipus is currently regarded as a monotypic genus to include single Australian species described as Teispes frenchi by Blackburn (1892) and subsequently transferred to Archetypus by Lameere (1903 a). Our molecular results (Jin et al. 2020) revealed a relatively isolated placement for this species, hence it was placed in a separate genus.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB621B39FBD1FCBFFD0BF970.taxon	description	(Figs. 5 J – M, 14 G, 15 J, 17 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB621B39FBD1FCBFFD0BF970.taxon	diagnosis	Diagnosis. Cryptipus frenchi is morphologically similar to Archetypus, Utra and Teispes. It can be separated from Archetypus by its uniformly brown body and mentum not fused to submentum; it can be separated from Utra by relatively smaller body size, dorsum with sparse and fine punctures, and small but distinctly lobed tarsomere 3; it can be separated from Teispes by pro- and mesotibia without lateral spines, much longer tarsomere 5, and anterior margin of pronotum less emarginate.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB621B39FBD1FCBFFD0BF970.taxon	materials_examined	Types. Teispes frenchi Blackburn lectotype designated here: the male specimen deposited in MV (T- 11007) with the following original labels “ Teispes frenchi, Blackb., Queensland | Nat. Mus. Victoria, C. French’s Coll. 5.11.08. ”; paralectotype designated here: the male specimen deposited in BNHM (BNHM- 010799949) with the following labels “ 4475, N. Qu, T. | Australia. Blackburn Coll. B. M. 1910 - 236. | Teispes Frenchi, Blackb. ”. Material examined (Fig. 17 B). Queensland: ANIC (66 ♂♂; 45 ♀♀): 56: Kuranda, 19 Bulter Dr., {from Nov. 2003 to Jan. 2017}, D. C. F. Rentz; 5: Mt. Hypipamee NP, Nov. 1998, A. A. Calder; 4: Cardstone, {from Jan. 1962 to Jan. 1963}, J. G. Brooks; 4: Kuranda, Black Mtn Rd., 10 miles N., {from Oct. 1970 to Nov. 1971}, A. and M. Walford-Huggins; 3: Clump Point, {from Jan. 1948 to Nov. 1949}, J. O. Campbell; 3: Kirrama Range, Dec. 1968, A. and K. Ey.; 3: Lync-Haven, Daintree area, Dec. 2012, [QM / ANIC]; 3: Mt. Carbine, Dec. 2005; 3: Whitefiled Range, W of Cairns, {from Nov. to Dec. 1970}, R. Hardie; 2: Daintree, Jan. 2000; 2: Davies ck., 18 km E by S of Mareeba, Nov. 1981, J. Balderson; 2: Little Mulgrave Valley, 5 M. N. - W Gillies Highway, Oct. 1969, J. G. Brooks; 2: Millaa Millaa, F. H. Taylor; 2: Palmerston Nat. P., Dec. 1966, E. Britton; 2: Whitefiled, Cairns, Nov. 1973, A. and M. Walford-Huggins; 2: Yandina; Cairns, [E. W. Ferguson Coll.]; Cathedral Fig., Danbulla Forest Reserve, 13 km NE by N of Yungaburra, Feb. 1988, D. C. F. Rentz; Kuranda, Jan. 1977, Dhane; Kuranda, Jan. 1978, A. E. D. Morton; Longlands Gap, Jan. 1977, B. P. Moore; Moses Ck., 4 km NbyE of Mt. Finnigan, Oct. 1980, T. Weir; Paluma Range, 1 / 2 M. below Crest, Jan. 1967, J. G. Brooks; Paluma, Dec. 1970, J. G. Brooks; Paluma, Jan. 2000; Paluma, Mt. Spec., Jan. 1959, C. Vallis; Tully Falls Rd., Jan. 1962, J. G. Brooks; Wallaman Falls, Jan. 1970, J. G. Brooks; Whitefiled, Cairns, Nov. 1970; MV (5 ♂♂; 2 ♀♀): 2: Cooktown; Cairns; Cardwell; Clump Point, Nov. 1949; Clump Point, Oct. 1951; Kuranda, Jan. 1950; OAI (1 ♂): Mulgrave Riv- er, North QLD, Dec. 1969, N. C. Coleman; QDPC (1 ♀): Imbil, Feb. 1936; QM (22 ♂♂; 5 ♀♀): 2: “ Chowchilla ”, {from Nov. to Dec. 1991}, C. T. L. and W. T. C.; „ Chowchilla ”, Feb. 2000, W. T. C.; 70 km S Ravenshoe, Jan. 2002, D. Kitchin and T. Jack; Atherton, Dec. 1972, B. Cantrell; Babinda, Jan. 1964, W. Panting; Cardwell Range, upper Broadwater Ck. valley, Dec. 1986, Monteith, Thompson and Hamlet; Cardwell, Dec. 1966, B. Cantrell; Chujeba Peak Summit, 7 km SW Redlynch, Dec. 1989, G. Monteith and G. Thompson; Cooktown, Jan. 1994, G. and A. Daniels and R. Eastwood; Daintree, Aug. 1998, G. E. Teakle; Garradunga, Jan. 2006, J. Hasenpusch; Gordonvale, Dec. 1967, G. Monteith; Julatten, Feb. 1999, B. P. Moore; Karnak-Devil’s Thumb, 8 - 12 km NW Mossman, Dec. 1989 - Jan. 1990; Kuranda, Black Mtn. Rd., 10 miles N., Feb. 1972, A. and M. Walford-Huggins; Lake Eacham, Dec. 1964, G. Monteith; Mantaka, Feb. 1972, A. and M. Walford-Huggins; Mt. Finnigan, Dec. 1990 - Jan. 1991, [QLD. Mus and ANZS- ES]; Mt. Misery, via Shiptons Flat, Dec. 1990, Monteith, Thompson, Cook, et al.; Mt. Spec, Jan. 1958, E. C. V.; Paluma Dam, Dec. 1963, G. Monteith; Paluma, Dec. 1965, G. Monteith; Paluma, Jan. 1973, B. Cantrell; Shiptons Flat, Dec. 1990, Monteith, Thompson, Cook, et al.; Stone Ck., Mar. – Nov. 2000, J. Hasenpusch; Whitfield Ra. Rd., 6 miles along, Feb. 1972, Mul.; QVMAG (3 ♂♂; 3 ♀♀): 3 Kareeya Power Station, Tully, Dec. 2000, S. Fearn; 3: Kareeya Power Station, Tully, Feb. 2002, S. Fearn. Molecular data. BioSample ID: SAMN 11080864.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB631B36F96BF97BFAA9FDDA.taxon	description	(Figs. 8, 14 H, 15 K, 17 C – D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB631B36F96BF97BFAA9FDDA.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 30 – 60 mm long. Dorsum usually with dark brown to black head and prothorax, and yellowish- or reddishbrown elytra. Sexually dimorphic dense punctures present on most of the pronotum, scutellum, all thoracic sterna and at least part of abdominal ventrites. Head narrower than prothorax; frontoclypeal suture arcuate to broadly angulate; median groove complete. Antennal tubercles prominent, rounded at apex. Eyes moderately large, weakly emarginate near antennal insertion, relatively distant dorsally. Mandibles about half of head capsule length, not sexually dimorphic, broad at base, not bent towards ventral side, and unidentate apically. Antenna extend to the middle of elytra. Scape about 1 / 3 of head capsule length, posteriorly usually extending to posterior margin of eye, gradually expanded; antennomere 3 shorter than scape, as long as or longer than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps elongate oval or weakly expanded, apex rounded. Prothorax transverse with lateral carina distinct, regularly dentate; anterior and posterior margins with complete beads, posterior angles relatively blunt. Pronotal disc uneven with weakly elevated smooth and coarsely punctate callosities on densely punctate, relatively dull background. Prosternal process subparallel, extending beyond procoxae, rounded apically. Elytral surfaces coriaceous or finely punctate; inner apical angle with sharp spine. Legs relatively long and strong; rows of or at least few spines present on femora and tibiae. Protibia surface rough and densely setose apically, with pair of subequal spurs; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB631B36F96BF97BFAA9FDDA.taxon	description	Description. Male. Length 30 – 60 mm. Head and prothorax usually dark brown to black, elytra often reddish or yellowish brown. Head approximately as long as wide, narrower than pronotum. Mandibles about half of head capsule length, weakly curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two or three teeth on incisor edge; dorsal surface with distinct carinae in E. servillei and E. tuberculicollis. Labrum weakly concave near apex, anterior margin weakly pointed with very dense setae; labrum separated from clypeus by a less sclerotised deep groove; dorsal surface of clypeus almost glabrous. Frontoclypeal suture arcuate to broadly angulate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11 - segmented, filiform, extending to the middle of elytra; scape moderately long and gradually expanded, reaching to or slightly beyond posterior margin of eye; antennomere 3 shorter than scape, as long as or slightly longer than antennomere 4; antennomeres 5 – 11 with rather blunt posterior angles. Eyes moderately large, transverse, coarsely facetted, slightly emarginate near antennal foramen; relatively distant dorsally. Submentum curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate oval to weakly expanded, apex rounded. Pronotum transverse, usually sub rectangular, weakly trapezoid in E. stigmosa, with posterior margin slightly broader than anterior margin; complete bead present on both anteri- or and posterior margins; lateral margin distinct with regular sharp teeth, with weak teeth in E. stigmosa. Disc surface very densely punctate and matt except for shiny, relatively smooth islands consisting of two triangular admedian areas, two smaller lateral areas and transverse area in front of the posterior margin. Prosternum covered with fine and deep punctures; hypomeron well-defined, slightly broader than prosternal process. Prosternal process parallel sided, extending beyond procoxae towards mesoventrite, rounded apically. Mesoventral process sub-parallel and emarginate apically. Ventral side of thorax and coxae with dense golden hairs. Metaventrite with shiny arrowshaped area medially, finely and densely setose elsewhere. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface with very fine, dense punctures. Elytral surfaces relatively shiny, covered with irregular coriaceous sculpture or very fine punctures; without weak traces of venation; elytral apices rounded with sharp sutural spine; epipleuron complete, narrowing apically. Legs very strong; femur sub-parallel or weakly constricted near both ends, surface rough, rows of spines present on ventral side; tibia expanded towards apex, with spines along outer margin, bearing sensory setae on ventral side apically; each tibia with distinct apical projection and a pair of subequal spurs; tarsi strong; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1 – 5 densely punctate basally with smooth and appearing less sclerotised apical part; ventral surface almost glabrous, only bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 H). Tegmen longer than sternite VII; parameres relatively short, about 0.13 times length of entire tegmen, round- ed apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex narrowly rounded. Female. Body usually more elongate than male, antennae usually extending to elytra humeral angle or just reach the middle of elytra. Pronotal disc entirely shiny, surfaces irregular with smooth central area and coarsely punctate lateral area; fine and dense punctures on thorax and abdomen absent. Legs much slenderer than in male. Ovipositor (Fig. 15 K) long, apical sclerosed part about 2 / 3 length of baculus, distal gonocoxites usually slender, relatively short and stout in E. stigmosa, apex rounded in most species, but weakly truncate in E. stigmosa; stylus small, inserted laterally and moderately distant to gonocoxite apex, weakly expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB631B36F96BF97BFAA9FDDA.taxon	discussion	Remarks. Eurynassa can be distinguished from the remaining genera of the Australian Macrotomini by having a short antennal scape, usually posteriorly extending to the middle of eye, antennomere 3 shorter than the scape; lateral margin of the pronotum moderately serrate; femur and tibia at least with some spines along external margins. Males always with distinguishable fine sexual punctures on ventral surface.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6C1B37FBCDFA21FEA1FC2B.taxon	description	(Figs. 8 A – B, 8 I, 17 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6C1B37FBCDFA21FEA1FC2B.taxon	diagnosis	Diagnosis. Male E. australis can be separated from other species by the following characters: body broad, elytra length usually less than twice of elytra width; maxillary and labial palps short; legs strong, surface very rough, lateral margin of each tibia with continuous rows of spines; large lobe on tarsomere 3; abdominal ventrites 1 – 5 fully covered with sexual punctures, except a pair of oval smooth area near lateral margins.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6C1B37FBCDFA21FEA1FC2B.taxon	materials_examined	Types. The type series of Mallodon australis has been lost. To establish identity of the well-known Boisduval’s species and to confirm established synonymy, the holotype of Mallodon figuratum Pascoe is here designated as also the neotype of Mallodon australis, which is a male specimen deposited at the BMNH with the following labels “ Pascoe Coll. 93 - 60. | Mallodon fignatum Pasc. Type | Eurynassa figurata Pasc. ” and ID as BNHM- 010799746. Material examined (Fig. 17 C). New South Wales: ANIC (7 ♂♂): Burrewarra Point, Apr. 2004, M. F. Braby; Burrier, Dec. 1933, Rodway; Hawkesbury R., Dec. 1975, D. P. Carne; Lameere; N. Repton, J. G. Brooks; nr. Gosford, Dec. 1965, J. Overell; Pk. Avenue, N. Repton, Dec. 1934, J. G. Brooks; MV (2 ♂♂): Como, Jan. 1956; Sydney; OAI (12 ♂♂): 2: Lisarow, Oct. 1956, K. M. Moore; Avoca, Feb. 1953, J. G. Gellatley; Castlecrag, Feb. 1969, R. D. Pares; Gladesville, nr Sydney, Jan. 1950; Lane Cove, near Sydney, Jan. 1979, G. R. Brown; Lisarow, Feb. 1956, R. Moore; Middle Harbour, Sydney, Oct. 1914, E. D. Bolton; Oatley, near Sydney, Jan. 1958; Penrith, near Sydney, Feb. 1965; Roseville, near Sydney, Feb. 1956; Rydalmere, near Sydney, Jul. 1965, E. Jacobs. Queensland: QM (3 ♂♂): 2: Brisbane, C. Deane; nr Fresh Water Lake, Cooloola, Mar. 1970, E. Dahms. Molecular data. BioSample ID: SAMN 12650006, SAMN 12650089.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6D1B34F886FB8AFEA1FC96.taxon	description	(Figs. 8 C – D, 8 J, 14 H, 15 K, 17 D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6D1B34F886FB8AFEA1FC96.taxon	diagnosis	Diagnosis. Male of E. servillei can be separated from E. australis by its slender body shape, smaller lobes on tarsomere 3; mandibular dorsal surface deeply concave (Fig. 8 J); longer maxillary ad labial palps; abdominal ventrite 5 not fully covered with sexual punctures; it differs to E. stigmosa by having larg- er and bowknot-shaped shiny area on pronotum, lateral margin of pronotum strongly dentate, abdominal ventrites 1 – 4 with smooth oval area near lateral edge; it can be discriminated to E. tuberculicollis sp. nov. by the sexual puncture area without tiny nodules.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6D1B34F886FB8AFEA1FC96.taxon	materials_examined	Types. Eurynassa servillei Thomson, lectotype here designated: male syntype in the MNHN with the following labels “ Eurynassa Thoms., n. pr. Sc. 303 | Serveillei Thoms. Type 304, Austr. Bor. | Th. TYPE | Museum Paris Coll. J. Thomson 1952 ”. The female specimen in the same collection is here designated as paralectotype with the following labels “ Museum Paris Coll. J. Thomson 1952 ”. Mallodon odewahnii Pascoe, Holotype male deposited in BNHM (BHNM- 010799838), with the following labels “ Mallodon odewahnii | Eurynassa odewahnii Pasc. | Pascoe Coll. 93 - 60 ”. Rhesus caesariensis Pic, Lectotype male deposit- ed in MNHN, with the following labels “ Caesarea | Type | Rehsus caesarensis Pic | Lectotipo | Eurynassa australis (Boisduval, 1835) ♂, G. Tavakilian det. 2007 ”. Material examined (Fig. 17 L). Queensland: ANIC (29 ♂♂): 3: 36 km SW of Mt. Garnet, Jan. 1977, B. P. Moore; 3: Edungalba, {from Dec. 1958 to Jan. 1964}, J. G. Brooks; 2: Biggenden, Jan. 1972, H. Frauca; 2: Silver Plains Homestead, Cape York Pens., {from Nov. 1961 to Dec. 1964}, J. L. Wassell; 2 mi NE of Noccundra, Jan. 1965, L. J. Chinnick; 20 km WSW Eungella, Nov. 1981, A. Gillison; Biggenden, May 1972, S. Allen; Coongara Rock, E of Sandy Ck., via Coalstoun Lakes, Jan. 1975, H. Frauca; Coongara Rock, E of Sandy Creek., via Coalstoun Lakes, Jan. 1975, H. Frauca; Davies Ck. Nat. Pk., Mar. 1993; Karumba, Jan. 1971, H. Heatwole; Lamberts Beach via Mackay, Jan. 1991, K. J. Sandery; Lolworth, Mar. 1900; Millmerran, [J. G. Brooks Bequest]; Mt. Garnet, Dec. 1960; Mt. Maria, via Rosedale, Mar. 1975, H. Frauca; Mt. Molloy, Dec. 1957, [J. G. Brooks Bequest]; Pilgrim Sands, Feb. 1998, E. D. Edwards and R. Oberprieler; Rockhampton, [C. G. L. Goods Coll.]; Roma, Jan. 1991, S. Watkins; Townsville, Jan. – Dec. 1964, C. Brweur; Toowoomba, Jan. 1974, J. MacQueen; Wandowan [Wandoan], Jan. 1957, J. Frazier; ASC (1 ♂): Barkly Highway, 7.14 km ESE of Camooweal, Feb. 2016, A. Sundholm and J. Bugeja; MV (21 ♂♂): 6: Coen; 4: Cooktown; 4: Endeavour River; 2: Townsville; Bowen, Feb. 1933; Claudie River, Jan. 1914; Edungalba, E. E. Adams; Mutchilba; [no data]; OAI (2 ♂♂): 2: Prairie, Jul. 1901; QM (25 ♂♂): 3: Rochford Scrub, Feb. 2007, Monteith; 2: Lake Broadwater via Dalby, Jan. 1985, G. Monteith and G. Thompson; “ Allinga ” Chinchilla, Lithgow, G.; “ Milroy ”, 70 km N of Qullpie, Nov. 1998, A. Eward; “ Wonga Hills ”, site 7, Mar. 2002, Monteith and Eright; Brisbane, Jan. – Dec. 1962, A. Gillison; Cooktown, Jan. 2003, A. Eward; Drillham, Jan. 2002, C. Daring; Drummond Ra, Dec. 2000, G. B. Monteith and D. J. Cook; Gayndah, Jan. 1962, H. A. Rose; Gundale, Brisbane, Dec. 2001, S. J. Johnson; Jandowae; Kogan via Chinchilla, Lithgow, G.; Mareeba, Dec. 1965, I. C. C.; Nipping Gully, Jan. – Jun. 1999, Monteith and Thompson; Nipping Gully, Jan. 1999, Monteith, Gough and Thompson; Oak Wells, old homestead, Dec. 2001, Monteith, Wright and Cook; Richmond, Jan. 1957, P. J. Coleman; Rochford Scrub, Dec. 2006, Monteith and Wright; Rockhampton; Taroom, Feb. 1991, G. and A. Daniels and C. J. Burwell; Thangool, Feb. 1991, G. and A. Daniels and C. J. Burwell; QVMAG (3 ♂♂): 2: Mt. Cooth’a [Mt. Coot- Tha], Brisbane, Oct. 1993 – Mar. 1994, S. Fearn; the Gap, Brisbane, Jan. 1994, S. Fearn. Northern Territory: ANIC (12 ♂♂): 8: Heffernan Rd., Alice Springs, Dec. 1989, D. P. Carne; 2: 39 km SW of Barrow Creek, Aug. 1980, M. S. Harvey; Alice Springs, Sep. 1992, R. Panersonm; Rimbija Island, Wessel Island, Feb. 1977, T. A. Weir; MV (5 ♂♂): Alice Springs (NT POPL); Alice Springs; Alice Springs, Nov. 1954; Bathurst Island, Oct. 1916; [no data]; QM (3 ♂♂): 2: Pellew Group, Feb. 1968, B. Cantrell; South West Is., Pellew Group, Feb. 1968, B. Cantrell. Western Australia: MV (2 ♂♂): Geraldton; [no data]. New South Wales: MV (1 ♂): [no data]. South Australia: QM (1 ♂): [no data].	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6D1B34F886FB8AFEA1FC96.taxon	description	Molecular data. BioSample ID: SAMN 12650007, SAMN 12650037, SAMN 12650045, SAMN 11080871, SAMN 12650090, SAMN 12650092, SAMN 12650039, SAMN 12650084, SAMN 12650046, SAMN 12650003, SAMN 12650004, SAMN 12650036, SAMN 12650042, SAMN 12650043, SAMN 12650093, SAMN 12650040, SAMN 12650041, SAMN 12650008, SAMN 12650044, SAMN 12650091.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B34F8F2FC25FDEEF909.taxon	description	(Figs. 8 E – F, 17 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B34F8F2FC25FDEEF909.taxon	discussion	Diagosis. Male E. stigmosa can be separated from other species of Eurynassa by the following characters: pronotum with a pair of small shiny triangular area medially, lateral margin weakly crenulate; tibia and femur without or with very few spines along external margins; abdominal ventrites 1 – 4 with a pair of pits near lateral edges.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B34F8F2FC25FDEEF909.taxon	materials_examined	Type. Mallodon stigmosum Newman, Holotype male deposited in BNHM (BNHM- 010799852), with the following original labels “ (Mallodon) stigmosum, Newm. Type | 49 4 - 1 454 ”. Material examined (Fig. 17 C). Western Australia: ANIC (2 ♂♂): Goldfields, Salmon Gum, Feb. 1981, W. P. Humphreys; Yellowdine, Feb. 1972, [Carnaby Coll.]; MV (4 ♂♂): 2: Dedari; Geraldton; Middalya. Northern Territory: ANIC (7 ♂♂): 7: Heffernan Rd., Alice Springs, Dec. 1989, D. P. Carne. New South Wales: AM (1 ♂): 4 km N of „ Talyeelye ” Hsd. Molecular data. BioSample ID: SAMN 12650039, SAMN 12650084, SAMN 12650046.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B35F8B9F8A4FD24FECE.taxon	description	(Figs. 8 G – H, 8 K – L, 17 D)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B35F8B9F8A4FD24FECE.taxon	diagnosis	Diagnosis. Male of E. tuberculicollis sp. nov. can be separated from other species of Eurynassa by having very small granules distributed between sexually dimorphic punctures (Fig. 17 D), especially visible on pronotum and ventral side of thorax, sometimes extending on the abdominal ventrites.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B35F8B9F8A4FD24FECE.taxon	description	Description. Length 30 – 50 mm. Male. Body sub-parallel, head as long as wide; mandibles half of head length; pronotum 1.2 × of head length and 1.8 x of head width; elytra over 5 × of head length. Antennal scape slightly longer than antennomere 3 length; interorbital width 1.2 × of eye length. Prosternal process narrow, less than 0.5 × width of procoxal cavity, and about half of hypomeron width; mesoventral process broader than prosternal process. Abdominal ventrites 1 – 5 about similar length, ventrite 5 emarginate at apex. Metatarsomeres 1 – 3 length ratio as 2: 1: 1, tarsomere 5 as long as tarsomere 1. Female. Pronotum weakly broader near posterior margin, dorsal surface only with sparse or dense regular punctures; elytra about 1.4 × width of pronotum.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B35F8B9F8A4FD24FECE.taxon	etymology	Etymology. The species name referring to the characteristic tiny granules present on areas with sexually dimorphic punctures.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6E1B35F8B9F8A4FD24FECE.taxon	materials_examined	Types. Holotype. ♂ “ WA: 10 km W. Menzies, 13 Jan. 2015, 400 W MV Light, P. M. Hutchinson | ANIC 25 - 066040 ” (ANIC). Paratypes (2): ♂: “ WA: 10 km W. Menzies, 13 Jan. 2015, 400 W MV Light, P. M. Hutchinson | ANIC 25 - 066007 ” (ANIC); ♀: “ WA: 10 km W. Menzies, 13 Jan. 2015, 400 W MV Light, P. M. Hutchinson | ANIC 25 - 066027 ” (ANIC); Other material examined (Fig. 17 D). Western Australia: AM (1 ♂): Southern Cross, H. W. Brown; ANIC (11 ♂♂): 2: 30 mi ENE of Eucla Motel, Oct. 1968, Key, Upton and Balderson; 2: Marloo Stn, Wurarga, Jan. 1931 – Dec. 1941, A. Goerling; 10 N Rawlinna, Jan. 1969, J. Bywater; 17 mi W by S of Cocklebiddy, Oct. 1968, Britton, Upton, Balderson; Kalgoorlie, Jan. 1946, Prof. Lameere; Kalgoorlie, Mar. 1900; Munclrabilla Station, Jan. 1990, R. Parterson; Port Hedland, Dec. 1965, P. Carne; Rawlinna, Jan. 1969, J. Bywater; MV (12 ♂♂): 3: Baandee, Mar. 1918; Carnarvon; Geraldton; Kellerberrin, Feb. 1907; Kellerberrin, Mar. 1907; Norseman, Mar. 1907; 4: [no data]. South Australia: ANIC (1 ♂): Arkaroola, Oct. 1981, P. Zborowski; MV (8 ♂♂): Emu Camp; 7: [no data]; QM (1 ♂): [no data]. Queensland: ANIC (1 ♂): 7 km N of Cunnamulla, Jan. 1971, R. C. Lewis; MV (4 ♂♂): Jandowae, Dec. 1920; May 1909; 2: [no data]; QM (4 ♂♂): Brisbane; Jandowae; Southwood NP. Camp, Dec. 2005, Monteith and Wright; Southwood NP., general, Dec. 2005, R. Hobson. New South Wales: ANIC (5 ♂♂): 2: Coolamon, F. H. Taylor; Sydney, Oct. 1924, W. W. Froggatt; Wagga Wagga, Apr. 1904, W. W. Froggatt; White Cliffs, Jan. 1976, R. S. Hogan; MV (1 ♂): Hen ty; OAI (3 ♂♂): 2: Brewarrina, Jan. – Dec. 1914, W. W. Froggatt; Whitton, Jan. – Dec. 1891; QM (3 ♂♂): 2: Broken Hill, C. Deane; Olive Downs, Jan. 1999, P. Bouchard. Molecular data. BioSample ID: SAMN 12650088 (Holotype); SAMN 12650087 (Paratype), SAMN 1265 0086 (Paratype); SAMN 12650038, SAMN 12650081.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6F1B32F8EFFEE0FF24FDDD.taxon	description	(Figs. 11 H – J, 14 J, 15 L, 17 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6F1B32F8EFFEE0FF24FDDD.taxon	diagnosis	Diagnostic combination (Male). Moderately large beetles, 30 – 40 mm long. Dorsum unicoloured, brown or dark brown. Sexually dimorphic dense punctures present on entire pronotum, scutellum and most of the ventral side including abdominal ventrites. Head slightly narrower than prothorax; frontoclypeal suture angulate, weakly impressed; median groove complete. Antennal tubercles prominent, rounded at apex. Eyes moderately large, weakly emarginate near antennal insertion, relatively narrowly separated dorsally. Mandibles less than 1 / 3 of head capsule length, not sexually dimorphic, broad at base, not bent towards ventral side and unidentate apically. Antenna extend beyond mid elytra in male. Scape about 0.7 times of head capsule length, posteriorly extending slightly beyond posterior margin of eye, gradually expanded; antennomere 3 distinctly longer than scape or antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps weakly expanded and apically rounded. Prothorax transverse with lateral carina weak and partly invisible, feebly dentate; anterior and posterior margins with complete beads. Pronotal disc with smooth and coarsely punctate callosities on densely punctate, feebly shiny background. Prosternal process subparallel, only slightly extending beyond procoxae, rounded apically. Elytral surfaces moderately coriaceous; inner apical angle with sharp spine. Legs relatively long; femora and tibiae with rows of spines. Protibia external surface granulose, ventral surface rough with sparse spines and dense yellow setae apically; apex with pair of subequal spurs; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6F1B32F8EFFEE0FF24FDDD.taxon	discussion	Description. Male. Length 30 – 40 mm. Body uniformly brown or dark brown. Head approximately as long as wide, narrower than pronotum. Mandibles shorter than 1 / 3 of head capsule length, weakly curved inwards, without sexual dimorphism; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with one extra tooth on incisor edge. Labrum strongly concave near apex, anterior margin pointed medially with very dense setae; dorsal surface with longer and denser setae than on clypeus; labrum separated from clypeus by a deep arcuate groove. Frontoclypeal suture angulate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11 - segment- ed, filiform, extending beyond mid elytra and sometimes almost to elytral apices; scape moderately long and gradually expanded, reaching beyond posterior margin of eye; antennomere 3 distinctly longer than scape, almost twice as long as antennomere 4; antennomeres 5 – 11 with blunt posterior angles. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; narrowly separated dorsally. Submentum straight at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps weakly expanded and apically truncate. Pronotum transverse, subrectangular with weak anterior angles and prominent posterior angles; anterior margin slightly emarginate medially with complete bead; posterior margin with complete bead; lateral carina weak and sometimes partially invisible with rather regular small sharp teeth. Disc surface very densely punctate and matt except for two triangular shiny, coarsely and deeply punctate admedian islands, two smaller lateral areas and transverse area in front of the posterior margin. Prosternum covered with fine and deep punctures; hypomeron well-defined, as broad as prosternal process; prosternal process parallel sided, only slightly extending beyond procoxae, rounded apically. Mesoventral process narrowing towards apex and emarginate apically. Ventral side of thorax and coxae with sparse golden hairs; metanepisternum densely setose. Metaventrite with shiny arrow-shaped area medially, finely and deeply punctate elsewhere. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface very densely and finely punctate. Elytral surfaces dull covered with irregular coriaceous sculpture; two or three weak traces of venation often present on apical 2 / 3 of elytra; elytral apices rounded with sharp sutural spine; epipleuron complete, narrowing apically. Legs strong and relatively long; femur sub-parallel, surface rough but without obvious spines on ventral side; tibia slender and relatively flat, without spines along outer margin, bearing sensory setae on ventral side apically; each tibia with distinct apical spine and a pair of subequal of spurs. Protibia external surface granulose, ventral surface rough with sparse spines and dense yellow setae apically; tarsi strong; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1 – 5 densely punctate basally with smooth and appearing less sclerotised apical part; ventral surface almost glabrous, only bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 J). Tegmen longer than sternite VII; parameres moderately long, 0.16 times length of entire tegmen, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex acute angulate. Female. Antennae usually shorter than in male, only extending to middle of elytra; antennomere 3 as long as or slightly longer than scape. Pronotal disc shiny, surfaces irregular with very deep and coarse punctures; fine and dense punctures on thorax and abdomen absent. Legs much slender than in male. Abdominal ventrite 5 truncate. Ovipositor (Fig. 15 L) relatively short, apical sclerosed part 3 / 4 length of baculus, distal gonocoxites very short and stout, with tiny lateral projection; stylus large, inserted laterally and very close to gonocoxite apex, gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B32F8F8FDFFFDBBF976.taxon	description	(Figs. 11 H – J, 14 J, 15 L, 17 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B32F8F8FDFFFDBBF976.taxon	diagnosis	Diagnosis. This species is morphologically similar to Pseudoplites and Eurynassa. Male can be separat- ed from P. hamali by its antennomere 3 not thicker than the following segments, the tibiae without lateral spines, and protarsi without long hairs. Female of G. queenslanda can be separated from P. hamali by its dense and coarse punctures on pronotum. Both sexes can be distinguished from Eurynassa species by having elongate antennomere 3 and prominent posteri- or angles on pronotum.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B32F8F8FDFFFDBBF976.taxon	materials_examined	Type. Geoffmonteithia queenslanda Jin, de Keyzer et Ślipiński, Holotype male deposited in ANIC, with the following labels “ QLD: Townsville, 29 September 1995, Woodger T. | ANIC 25 - 067089 ”. Material examined (Fig. 17 E). Queensland: AM (1 ♂): 5 km N Leyburn, Dec. 1987, A. and G. Daniels; ANIC (3 ♂♂): Eidsvold, Dec. 1965, J. Bancraft; Townsville, Sep. 1995, T. Woodger; Yeppoon, Nov. 1965, Le Souef; QDPC (1 ♂): Long Pocket, Indooroopilly, Brisbane, Nov. 1980, J. F. Donaldson; QM (2 ♂♂; 2 ♀♀): 2: Gympie; Boggimoss No. 21 via Taroom, Nov. 1996, G. B. Monteith; Dipperu, Nov. 1971, B. Baldwin; QVMAG (4 ♂♂; 2 ♀♀): 6: The Gap, Brisbane, {from Jan. 1994 to Dec. 1995}, S. Fearn. New South Wales: ANIC (1 ♂; 1 ♀): Stumpys Rd., Braemar S. F., Nov. 2016, R. de Keyz- er, A. Scott and Sundholm, A.; south boundary Fortis Creek NP., north of Grafton, Dec. 1998, Watkins; MV (1 ♂): Canowindra, Jan. 1956, F. E. Wilson. Molecular data. BioSample ID: SAMN 11080875 (Holotype), SAMN 11080889 (Paratype).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B33F893F94BFB87FD10.taxon	description	(Figs. 9, 14 I, 15 M, 17 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B33F893F94BFB87FD10.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 30 – 70 mm long. Dorsum usually unicoloured, black or dark brown, sometimes with elytra reddish brown. Sexually dimorphic dense punctures absent. Head distinctly narrower than prothorax; frontoclypeal suture arcuate or broadly angulate; median groove complete. Antennal tubercles moderately prominent, rounded apically. Eyes large, weakly emarginate near antennal insertion, moderately separated dorsally. Mandibles about half of head capsule length, not sexually dimorphic, not bent towards ventral side, and unidentate apically. Antenna in male extending to or beyond the middle of elytra. Scape about 1 / 3 – 1 / 2 of head length, extending beyond posterior margin of eye, expanded apically, flat with blunt external edge; antennomere 3 slightly longer than scape and distinctly longer than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps elongate oval and apically slightly truncate. Prothorax transverse with lateral carina regularly spinose; anterior and posterior margins with complete beads. Pronotal disc usually glabrous, sometimes with very sparse and fine hairs, surface with dense coarse punctures. Prosternal process subparallel, extending far beyond procoxae, narrowly rounded or slightly pointed apically. Elytral surfaces strongly coriaceous, with at least two distinct costae, without adpressed hairs; inner apical angle with short spine. Abdominal ventrites 1 – 4 covered by lighter tomentose pile separated from glabrous posterior patch on each ventrite by rows of long erected setae. Legs relatively long and strong; femora with rows of sharp spines ventrally. Protibia with rows of spines, two or three sharp external projections apically and a pair of subequal spurs; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B33F893F94BFB87FD10.taxon	description	Description. Male. Length 30 – 70 mm. Dorsum usually uniformly brown or dark brown, sometimes with slightly reddish elytra. Head capsule approximately as long as wide, narrower than pronotum, and with very sharp genal projection. Mandibles about as long or slightly longer than half of head capsule, moderately curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two extra teeth on incisor edge. Labrum transverse, anterior margin pointed medially with very dense setae; labrum separated from clypeus by a deep arcuate groove; dorsal surface of labrum and clypeus almost glabrous. Frontoclypeal suture angulate or broadly arcuate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11 - segmented, filiform, extending to or beyond the middle of elytra; scape long and weakly flat, reaching beyond posterior margin of eye; antennomere 3 slightly longer than scape and distinctly longer than 4. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; moderately separated dorsally. Submentum weakly emarginate at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate oval and apex slightly truncate. Pronotum sub rectangular, transverse, usually with slightly curved anterior margin; complete bead present on both anterior and posterior margins; lateral carina distinct with regular sharp teeth. Disc almost glabrous, rarely with very sparse and fine hairs, almost uniformly densely and coarsely punctate with very weakly elevated and shiny area medially. Prosternum coriaceous; hypomeron welldefined, broader than prosternal process. Prosternal process sub-parallel, strongly projecting beyond procoxae, narrowly pointed or rounded apically. Mesoventral process relatively narrow, emarginate at apex. Ventral side of thorax and coxae covered with golden hairs, ventral side of trochanter with erect golden setae. Metanepisternum weakly constricted at both ends. Scutellum rounded apically; surface with or without dense punctures. Elytral surfaces covered with irregular coriaceous sculpture, glabrous and usually shiny, with at least two weak traces of venation presenting on each elytron; elytral apices rounded with short sutural spine; epipleuron almost complete, narrowing apically. Legs strong; meso- and metatrochanter with patches of dense yellowish setae; femora sub-parallel, with two rows of sharp spines on ventral side; tibiae weakly expanded towards apex, surface rough and with rows of spines along outer margin, bearing sensory setae on ventral side apically; each tibia with two or three apical projections and a pair of subequal spurs; tarsi moderately strong; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdominal ventrites 1 – 4 covered by lighter tomentose pile separated from glabrous posterior patch on each ventrite by rows of long erected setae; lateral margin with complete carinae; setae present along the edge of ventrites 1 – 5, especially on the apex of ventrite 5. Male genitalia (Fig. 14 I). Tegmen longer than sternite VII; parameres length varies in different species, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex emarginate while ventral apex narrowly rounded. Female. Body usually larger than male, length up to 80 mm. Antennae usually shorter than in male, filiform, only extending beyond elytral humeral angle; antennal scape about as long as antennomere 3 and distinctly longer than antennomere 4. Pronotal disc trapezoid with posterior margin broader than anterior margin; sexual setal patches absent on abdominal ventrites. Ovipositor (Fig. 15 M) long; apical sclerosed part longer than half length of baculus, distal gonocoxites length and shape varies in different species, stylus moderately short and small, inserted laterally and relatively distant to gonocoxite apex, very weakly expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB681B33F893F94BFB87FD10.taxon	discussion	Remarks. Two species, H. princeps and H. blackburni, were originally included in Cnemoplites because of sexual patches on male abdominal ventrites. Our molecular results (Jin et al. 2020) revealed a long distance between them and the type species of Cnemoplites, and strongly indicated that the sexual abdominal patches evolved multiple times in the Prioninae history. Actually, these two species can be easily distinguished from Cnemoplites by the following characters: antennal scape shorter, only extending to the posterior margin of eyes; genal projections very sharp; pronotum more or less square with hypomeron relatively broader at anterior angle; female ovipositor with very small styli. One new species H. mandibularis sp. nov. (indicated as Hagrides sp 2 in Jin et al. 2020) is accordingly described here.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB691B30FBCFFA5CFD0BF9C7.taxon	description	(Figs. 9 A – E, 17 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB691B30FBCFFA5CFD0BF9C7.taxon	diagnosis	Diagnosis. H. blackburni can be separated from H. princeps by its completely punctate pronotum with the median smooth shiny area completely absent. Male distinctiveness is also supported by very lightly pubescent tarsi, the abdominal setal patch smaller and the penis with longer apical sclerotised part. Female is distinguished from H. princeps by the ovipositor with rounded distal gonocoxal apices. It can be separated from H. mandibularis by having shorter mandibles, the male abdominal ventrite 5 with transverse carinae (Fig. 9 D) and the tegmen with longer parameres while the female is distinguished by the longer distal gonocoxites.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB691B30FBCFFA5CFD0BF9C7.taxon	materials_examined	Type. Lectotype designated here: the female specimen (BNHM- 010799923) deposited in BNHM with the following labels “ Albany, 8658 | Cnemoplites blackburni Lameere ♀ cotype ”. Material examined (Fig. 17 F). Western Australia: ANIC (8 ♀♀): 3: Yellowdine, Feb. 1972, [Carnaby Coll.]; Eucla, Jan. 1986, C. Reid; Floreat Park, Feb. 1981, G. P. Hall; Kalgoorlie, Jan. 1898 – Dec. 1989; Madura Hotel, Madura, Nov. 1968, A. M. Richards; Ravensthorpe, Feb. 1974; ASC (1 ♂; 1 ♀): Balladonia- Zanthus track, 2.78 km N of Balladonia Roadhouse, Jan. 2017, A. M. Sundholm and J. Bugeja; Cundalee- Queen Victoria Springs Road, 0.6 km NNW of Cundalee, Jan. 2017, A. M. Sundholm and J. Bugeja; MV (1 ♀): Perth; QM (1 ♂): Cocklebiddy, Jan. 1964, R. C. Shepherd. South Australia: AM (1 ♂): [no data]; ANIC (1 ♂; 4 ♀♀): Brookfield, Conserv. Park, Feb. 1992, A. M. E. Roach; Hale Nat. Park, Williamstown, Jul. 1976, C. A. Kiakby; Murray river, H. S. Cope; Renmark, Feb. 1950, A. S.; Murray river, H. S. Cope; ASC (2 ♂♂): Cumberland Park, Mar. 1963; Winkie, Jan. 1963, R. T. Wheaton; MV (6 ♀♀): 4: Renmark, Jan. 1937; Kychering Soak, 24 mls West; Renmark, F. E. Wilson; QM (1 ♂; 9 ♀♀): Ooldea, A. M. Lea; [no data]. Victoria: ANIC (5 ♀♀): 2: Rainbow, Jan. 1966, R. Whyte; Mildura, Dec. 1978, M. Clements; Moroondah, Feb. 1921, L. J. Perry; Parkville, Nov. 1922, A. Wade; MV (7 ♂♂; 9 ♀♀): 2: Birchip, Nov. 1920; 2: Linga, Apr. 1905; 2: Red Cliffs, Jan. 1946; Dimboola, Dec. 1913; Hattah; Mallee District {Ouyen lat / long}; Mildura; Natya; Ouyen; Panitya, Apr. 1905; Red Cliffs; Redcliffs, Mar. 1925; Wimmera; QM (1 ♂): Speed, J. E. Dixon. New South Wales: AM (1 ♂): Round Hill Res., Jan. 1949, I. V. Paters; ANIC (1 ♂; 3 ♀♀): 2: Yenda, Jan. 1949, R. Sheridan; Cronulla, Jun. 1955; Murray R.; MV (2 ♂♂): Apr. 1911; Mar. 1909; OAI (2 ♂♂): Peak Hill, Apr. 1931; Royal Botanic Gardens, Sydney, Jan. 1904. Molecular data. BioSample ID: SAMN 12650009.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B30F94EF999FB1BFA1D.taxon	description	(Figs. 9 F – I, 17 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B30F94EF999FB1BFA1D.taxon	diagnosis	Diagnosis. H. mandibularis is similar to H. blackburni but can be recognised by having longer mandibles (Fig. 9 I), the male abdominal ventrite 5 without transverse elevations at base, the tegmen with shorter parameres. A female is diagnosed by having the ovipositor with shorter distal gonocoxites. It can be separated from H. princeps by having completely punctate pronotal disc, smaller sexual patches on abdominal ventrites, and the tegmen with very short parameres and basal triangular projection. A female can be separated by the rounded distal gonocoxal apices.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B30F94EF999FB1BFA1D.taxon	description	Description. Male. Body length 30 – 52 mm. Body sub-parallel, slightly convex; head weakly transverse; mandibles longer than eye length; pronotum as long as head capsule and pronotal width 1.5 × of head width; elytra over 4 × longer than pronotum. Antennal scape about 1.5 × of antennomere 3 length; interorbital width over 1.2 × of eye length. Prosternal process width about the same as hypomeron width and 0.7 × length of procoxal cavity; mesoventral process as broad as prosternal process. Abdominal ventrites 1 and 5 longer than the others; ventrite 5 emarginate apically. Metatarsomeres 1 – 3 equal in length, tarsomere 5 shorter than 1 – 3 combined. Female. Body length 50 – 80 mm.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B30F94EF999FB1BFA1D.taxon	etymology	Etymology. The species name emphasises its elongate mandibles, which is a distinctive character to separate it from the morphologically similar species. Types. Holotype. ♂ “ WA: 10 km W. Menzies, 13 Jan. 2015, 4000 w MV Light, P. Hutchinson | ANIC 25 - 066034 ” (ANIC). Paratypes (5). 2 ♂♂: “ WA: 2 Feb. 2017, Lake Barlee-Youanmi Road., 5.43 km NNW of Diemels Homestead. 29 ° 37 ’ 17.67 ’’ S 119 ° 17 ’ 11.35 ’’ E, Elv. 426 m, attracted to 250 w MV Light, Allen M. Sundholm, Joe Bugeja ” (ASC); “ WA: Lake Douglas, 12 km S of Kalgoorlie, 13 Jan. 1989, M. S. and B. J. Moulds | AM K 503085 ” (AM); 3 ♀♀: “ WA: 10 km W. Menzies, 13 Jan. 2015, 4000 w MV Light, P. Hutchinson | ANIC 25 - 066004 ” (ANIC); “ 10 N. Rawlinna, WA, Jan 1969, J. Bywater | ANIC 25 - 072811 ” (ANIC); “ WA: 1 Feb. 2017, Lake Barlee-Youanmi Road., 5.43 km NNW of Diemels Homestead. 29 ° 37 ’ 17.67 ’’ S 119 ° 17 ’ 11.35 ’’ E, Elv. 426 m, attracted to 250 w MV Light, Allen M. Sundholm, Joe Bugeja ” (ASC); Other material examined (Fig. 17 F). Australia: MV (1 ♀): [no data]. Western Australia: ANIC (1 ♂; 1 ♀): 2: 10 km W. Menzies, 13 Jan. 2015, Hutchinson, P.; MV (2 ♀♀): Marvel Loch; Rottnest Island, Douglas, A. Molecular data. BioSample ID: SAMN 12650095 (Holotype); SAMN 12650096 (Paratype).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B31FBE0F9BFFDEEFA44.taxon	description	(Figs. 9 J – M, 14 I, 15 M, 17 F)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B31FBE0F9BFFDEEFA44.taxon	diagnosis	Diagnosis. H. princeps can be separated from oth- er Hagrides species by the meso- and metatrochanter bearing dense setae along ventral surfaces, the prosternal process with narrowed pointed apex and the pronotum with some smooth shiny area medially. This species is mostly distributed in eastern Australia. Males can also be recognised by having large setal patch (Fig. 9 M), and the tegmen with longer basal triangular projection. The female can be diagnosed by the ovipositor with truncate distal gonocoxites apices.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6A1B31FBE0F9BFFDEEFA44.taxon	materials_examined	Type. Holotype male deposited in BNHM (BNHM- 010799836), with the following labels “ Cnemoplites princeps Gahan, ♂, Type | Queensland, 81.14 ”. Material examined (Fig. 17 F). Victoria: MV (4 ♀♀): Lake Boga, R. C. Sloane; Lake Hattah; underbool, Mar. 1928; [no data]. New South Wales: AM (1 ♀): Cobar, Mar. 1987, R. H. Mulder; ANIC (2 ♂♂; 2 ♀♀): 2: Steam Plains, Booroorban, Jan. – Feb. 1937, H. L. Jenniy; Cooinbil, Carrathool, Jan. 1938, T. G. Campbell; Toganmain, Jan. 1966, J. Rawley; MV (2 ♀♀): Broken Hill, Feb. 1917; Forbes; OAI (1 ♂): Haythorpe Station, Broken Hill, May 1992. Queensland: ANIC (4 ♂♂; 7 ♀♀): 2: Mareeba, Dec. 1952, [J. G. Brooks Bequest]; 2: Quilpie, Jan. 1966; Edungalba, Dec. 1969, E. Hoar; Eulo, Aug. 1963, J. C. Le Souef; Longreach, Dec. 1962, S. Williamsen; Mt. Garnet, Dec. 1960, [Uni of New England Coll.]; Mt. Isa, Dec. 1967, J. L. Mckeown; Muttaburra, Nov. 1971, R. A. H. Davies; Slatey Creek, Edungalba, Jan. 1953, Brigalow; ASC (1 ♂; 1 ♀): 2: 6.3 km N of Tambo, Jan. 2015, A. Sundholm and J. Budeja; MV (2 ♂♂; 1 ♀): 2: Mount Tamborine, Dec. 1951; Townsville; QDPC (1 ♀): Barcaldine, Jan. 1959; QM (5 ♂♂; 5 ♀♀): 2: Yarramulla, Dec. 2002, Monteith; “ Kahmoo ”, 15 km W of Cunnamulla, Jan. – Mar. 1993, K. Campbell; Akaray Station, Augathella, May 1991, M. Wade; Colimboola, Lithgow, G.; Jundah, “ Byudes ” Hmstd, Nov. 1992, A. Emmott; Mazeppa NP, N end, Dec. 2000 – Mar. 2001, Cook and Monteith; Sth of Starke R., Nth of Cooktown, Nov. 1992, P. Davie; Tara, Feb. 2002, C. Wayd; Wowan, Jan. 1979, W. Thorlow. Northern Territory: ANIC (1 ♂): Heffernan Rd., Alice Springs, Dec. 1989, D. P. Carne; MV (1 ♂): Alice Springs, Mar. 1955. Molecular data. BioSample ID: SAMN 12650097, SAMN 12650049, SAMN 12650050.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6B1B0EF974FA67FC48FD9D.taxon	description	(Figs. 10, 11 A – G, 14 K, 15 N, 18 A – B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6B1B0EF974FA67FC48FD9D.taxon	diagnosis	Diagnostic combination (Male). Moderately large beetles, 25 – 40 mm long. Dorsum usually unicoloured, black to dark brown or slightly reddish brown, sometimes with elytra yellowish. Sexually dimorphic, dense punctures absent. Head distinctly narrower than prothorax; frontoclypeal suture arcuate or angulate; medi- an groove complete. Antennal tubercles prominent, rounded or slightly pointed apically. Eyes large, weakly emarginate near antennal insertion, narrowly to moderately separated dorsally. Mandibles 1 / 3 – 1 / 2 of head capsule length, not sexually dimorphic, not bent towards ventral side and unidentate apically. Antenna in male extending beyond middle of elytra, sometimes reaching elytral apex. Scape about as long or slightly shorter than half of head capsule length, posteriorly extending beyond posterior margin of eye, moderately expanding apically, flat with blunt external edge; antennomere 3 longer than or about as long as scape and distinctly longer than 4. Mentum not fused to submentum. Terminal palpomere of maxillary and labial palps elongate oval and apically rounded. Prothorax transverse with lateral carina regularly dentate; anterior and posterior margins with complete beads. Pronotal disc at least bearing some hairs, surface usually with smaller and sparse punctures near central and coarser and denser punctures laterally. Prosternal process subparallel, extending beyond procoxae, rounded apically. Elytral surfaces coriaceous, with or without adpressed hairs; inner apical angle with short spine. Legs relatively long; femora with rows of sharp spines on ventral side. Protibia without rows of spines, with two sharp external projections apically and a pair of subequal spurs; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6B1B0EF974FA67FC48FD9D.taxon	description	Description. Male. Length 25 - 40 mm. Dorsum usually unicolored, black, dark brown or reddish brown, sometimes with yellowish elytra. Head approximately as long as wide, narrower than pronotum. Mandibles not sexually dimorphic, about as long or shorter than half of head capsule length, moderately curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two extra teeth on incisor edge. Labrum transverse, anterior margin weakly emarginate medially with very dense setae; labrum separated from clypeus by deep arcuate groove, weakly scelerotised; dorsal surface of labrum and clypeus covered with long golden setae. Frontoclypeal suture angulate or arcuate; median groove distinct and complete. Antennal tubercles prominent, apex rounded. Antenna 11 - segmented, mostly filiform, in H. prionoides antennae moderately serrate, mostly extending beyond the middle of elytra, reaching to elytral apices in H. prionoides. Scape long and weakly flat, reaching beyond posterior margin of eyes; antennomere 3 as long as or slightly longer than scape, distinctly longer than antennomere 4. Eyes large, transverse, coarsely facetted, slightly emarginate near antennal foramen; narrowly or moderately separated dorsally. Submentum rather straight at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate oval and apically truncate. Pronotum transverse, usually trapezoid with posterior margin broader than anterior margin; anterior emarginate medially with complete bead; posterior margin with complete bead; lateral carina with rather regular small sharp teeth. Disc surface covered with dense long hairs or at least with long adpressed setae, usually with less and smaller punctures in the central, and much coarser and dense punctures laterally. Prosternum moderately punctate; hypomeron well-defined, broader than prosternal process; prosternal process projecting towards mesoventrite, narrowly rounded apically. Mesoventral process narrow, emarginate at apex. Ventral side of thorax and coxae covered with golden hairs. Metanepisternum almost sub-parallel, only weakly constricted at base. Scutellum rounded apically; surface moderately to densely setose. Elytral surfaces dull covered with irregular coriaceous sculpture, mostly glabrous except in H. howei, only H. prionoides with two weak traces of venation presenting on basal half of elytra; elytral apices rounded with short sutural spine; epipleuron almost complete, narrowing apically. Legs strong; femur sub-parallel, with two rows of sharp spines on ventral side; tibia weakly expanded towards apex, surface rough but without spines along outer margin, bearing sensory setae on ventral side apically; each tibia with two or three apical projections and a pair of subequal spurs; tarsi strong; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdominal ventrites 1 – 4, each with sexual setal patch in all species, except of H. prionoides, surfaces smooth and bearing few adpressed hairs on lateral areas. Long setae present along the edge of ventrites 1 – 5, especially on the apex of ventrite 5. Male genitalia (Fig. 14 K). Tegmen longer than sternite VII; parameres moderately long, 0.15 – 0.3 times length of tegmen, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex emarginate while ventral apex narrowly pointed. Female. Antennae usually shorter than in male, filiform, only extending to middle of elytra. Pronotal disc usually shiny and smooth in the central, with fine punctures in H. impar; the whole body less setose than in males, sexual setal patches absent on abdominal ventrites. Ovipositor moderately long in H. impar (Fig. 15 N) and H. prionoides, relatively short in H. howei; apical sclerosed part at least longer than 2 / 3 length of baculus, distal gonocoxites long and expanded; stylus rather short, inserted laterally and relatively distant to apex, expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB6B1B0EF974FA67FC48FD9D.taxon	discussion	Remarks. The monophylly and internal classification of Hermerius were supported by the previous molecular analyses (Jin et al. 2020). With further mor- phological research, we revealed the following morphological characters which can be used to support this newly defined taxon: (1) pronotum completely covered with golden hairs in males, with the length and density of hairs variable in different species; (2) the dorsal piece of penis emarginate apically; (3) ovipositor with very small styli and distal gonocoxites moderately emarginate near stylus insertion. Of the five species currently recognised in Hermerius, four were formerly classified in different subgenera of Cnemoplites, and one is here newly described from Western and South Australia. Unfortunately, females of H. fairmairei and H. occidentalis sp. nov. are still unknown.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB541B0FFBC0FAE4FD0BFE79.taxon	description	(Fig. 10 A – B, 11 D, 18 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB541B0FFBC0FAE4FD0BFE79.taxon	diagnosis	Diagnosis. Male H. fairmairei can be separated from remaining species of Hermerius species in having the pronotum covered with long and fine hairs, the abdominal ventrites 1 – 4 with long and semi-erect sexual setae medially and the filiform antennae.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB541B0FFBC0FAE4FD0BFE79.taxon	materials_examined	Type. Holotype male deposited in MNHN, with the following labels “ MUSEUM PARIS Collection Léon Fairmaire, 1906 | 4067 | Cnemoplites fairmairei, A. Lameere vid. 1914 | Cnemoplites 9 | TYPE ”. Material examined (Fig. 18 A). Queensland: ANIC (2 ♂♂): Millmerran, Jan. 1951, J. M. C.; Mourange, Edungalba, Dec. 1968, E. E. Adams; MV (2 ♂♂): Jandowae, Oct. 1900; [no data]; QM (5 ♂♂): 2: Blackdown Tableland, Stoney ck via Dingo, Dec. 1985, S. Hamlet; 2: Virgin Brigalow forest, ~ 10.5 km NNW., R. Wicks Res. Stn., Bringalily S. F., via Inglewood, Dec. 2006, A. E.; Brigalow Re. Stn. Site 4, Oct. – Dec. 2000, Cook and Monteith; RDKC (2 ♂♂): Blackdown Tableland, Expedition Rg., Jan. 1987, M. S. and B. J. Moulds; Blackdown Tbland, Jan. 1973, M. S. Moulds; Molecular data. BioSample ID: SAMN 12650100.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0FF8B9FE58FC43F9BD.taxon	description	(Figs. 10 E – G, 11 G, 18 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0FF8B9FE58FC43F9BD.taxon	diagnosis	Diagnosis. Males of H. howei can be separated from other Hermerius species by the entire dorsum, including head, pronotum and elytra completely covered with relatively long golden hairs. Female of this species can be separated to other Hermerius species by the pronotum with moderately smooth area medially, the posterior angles blunt and the prosternum relatively glabrous.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0FF8B9FE58FC43F9BD.taxon	materials_examined	Types. Arimaspes howei Thomson, Holotype male deposited in MNHN, with the following labels “ Arimaspes Thoms., s. c. 301, n. pr. | howei Thoms. Type 302, Lord Howe’s Is. | Th. TYPE ”. Cnemoplites (Hermerius) intermedia Wilson, Holotype male in MV, with the following labels “ Moorilla, Grenfell, NSW, T. G. Sloane | Cnemoplites intermedia Wilson, Type | F. E. Wilson Collection | Holotype T- 2909 ♂, Cnemoplites intermedia Wilson ”. Material examined (Fig. 18 B). Queensland: MV (2 ♂♂; 1 ♀): Millmerran, Jan. 1946; Mount Tamborine, Dec. 1951; [no data]; QDPC (1 ♀): North Pine; QM (13 ♀♀): 3: „ Glen Witheren ”, main scrub, Dec. 1991 – May 1992, G. Monteith; Aramar SF., 3.8 km W of Maryborough, Dec. 1992, J. Ferguson and T. Cameron; Blackdown Tableland via Dingo, Jan. – Dec. 1982, S. Pearson; Brigalow Res. Stn., site 2, Dec. 2000 – Mar. 2001, Cook and Monteith; Brisbane; Brisbane, Jan. 1965, J. C. Cardale; Curtis Farm, Canungra, Jan. 1990, G. B. Monteith; Glen Witheren, Canungra Ck., Jan. 1987, G. Monteith; Gurgeena, Oct. – Dec. 2012, G. Monteith; Gympie; Taroom, 1.5 km N Hwy, Dec. 1997 – Apr. 1998, G. Monteith; QVMAG (2 ♂♂; 12 ♀♀): 7: 176 Mons School Road, Buderim, Dec. 2014, S. Fearn; 7: the Gap, Brisbane, {from Jan. to Feb. 1994}, S. Fearn; RDKC (2 ♂♂): 41 km W of Boonah, White Swamp – Boonah Rd., Jan. 2014, A. Scott; Mt. Nebo, Feb. 1987, O. Thomas. New South Wales: AM (4 ♂♂; 3 ♀♀): Avoca Beach, Feb. 1986, S. Hunter; Bruxner Park, near Coffs Harbour, Jan. 1971, D. K. McAlpine; Lansdowne, Dec. 1980, C. Cross; Lansdowne, Dec. 1980, G. and T. Williams; Lansdowne, Jan. 1981, G. and T. Williams; Lansdowne, Jan. 1981, G. Williams; Otford, Jan. 1966, R. H. Mulder; ANIC (8 ♂♂; 22 ♀♀): 5: Powerful Owl Resv., Girard SF., west of Drake, Dec. 1998, S. Watkins; 4: Western boundary Yarratt State Forest, NE Wingham, Dec. 1994, S. Watkins; 2: south boundary Fortis Creek NP., north of Grafton, Dec. 1998, S. Watkins; Bonville, Jan. 1976, P. B. Carne; Caparra, Jan. 1992, S. Watkins; Durras North, nr. Batemans Bay, Jan. – Feb. 1987, D. C. F. Rentz; Durras, Jan. 1939, G. F. H.; Fortis Creek SF, N. Grafton, Dec. 1996, S. Watkins; Mackellar Ra. Rd., Bungabee SF, 15 km SE Kyogle, Dec. 1997, S. Watkins; N. Durras, Mar. 1969, R. J. Bartell; Ogilvie Trail, Banyabba SF, SSW Whiporie, Dec. 1998, S. Watkins; Powerful Owl Resv., Girand SF, West of Drake, Jan. 1999, Watkins; Powerful Owl Resv., Girard SF, west of Drake, Jan. 1999, S. Watkins; south boundary Fortis Creek NP, northern of Grafton, Dec. 1998, S. Watkins; Starr’s Creek, Lansdowne SF, Dec. 1994, S. Watkins; Stumpys Rd., Braemar S. F., Nov. 2016, R. de Keyzer, A. Scott and Sundholm, A.; Sydney; West Pennant Hills, Cumberland N. F., Jan. 1974, R. Trueman; Western boundary Yarratt State Forest NE Wingham, Dec. 1994, S. Watkins; Wingham, Yarratt SF., Jan. 1982, G. and T. Williams; MV (2 ♂♂; 2 ♀♀): Acacia Creek, Dec. 1910; Sydney; Gosford; Moorilla (HMSD), Grenfell; OAI (4 ♀♀): 2: Graman, Feb. 1961, P. G. Regan; Gosford, Dec. 1908; Wauchope, Jan. 1930; QM (3 ♀♀): 2: c. 3 km SE Mt. Pikapene, Mt. Pikapene SF, NW Grafton, Jan. 1992, J. Stanisic and J. Chaseling; Geary’s Flat Rd., Kippara SF., NW Wauchope, Jan. 1992, J. Stanisic and J. Chaseling; RDKC (1 ♂): Wauchope, Feb. 2003, R. de Keyzer. Victoria: MV (3 ♀♀): 2: Mallee District (Ouyen); Mildura. Molecular data. BioSample ID: SAMN 12650011, SAMN 12650099, SAMN 11080888.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0FF8B9FE58FC43F9BD.taxon	discussion	Remarks. As discussed above under the Agrianome we believe that the type locality of Arimaspes howei Thomson (1864) was wrongly recorded and the specimens were collected in the mainland Australia. Based our material and continuous observations conducted by Australian Museum (C. Reid, pers. comm.) on Lord Howe Island Agrianome howei and Howea angulata were only two Prioninae species taken from the Island.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0CFBEDF91FFEA1FCDE.taxon	description	(Figs. 10 H – J, 11 F, 14 K, 15 N, 18 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0CFBEDF91FFEA1FCDE.taxon	diagnosis	Diagnosis. Male of H. impar can be separated to other Hermerius species by the following characters: pronotum and ventral side of the whole thorax bearing relatively long and dense hairs; elytra glabrous; sexual setal patch present on abdominal ventrites 1 – 5; antennae filiform, with antennomere 3 longer than scape or antennomere 4. Females are densely setose on ventral side of thorax; pronotum bears few hairs laterally; legs are glabrous and smooth, protibia is without teeth along external margin.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB551B0CFBEDF91FFEA1FCDE.taxon	materials_examined	Types. Lectotype designated here: male specimen (BNHM- 010799794) deposited in BNHM, with the following labels “ Ent. Club. 44 - 12 | Hermerius impar ♂ Newm., Kangaroo Island, 4 / 5 ”. Paralectotype female deposited in BNHM, ID as BNHM- 010799855. Material examined (Fig. 18 A). Australia: MV (1 ♂): [no data]. South Australia: ANIC (1 ♀): Vivonne Bay, Hall, G. P., Jan. 1976; SAM (1 ♂): Kangaroo Island. Molecular data. BioSample ID: SAMN 12650013, SAMN 12650048.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0CF94CFCF1FEE9F8AB.taxon	description	(Figs. 10 C – D, 11 E, 18 A)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0CF94CFCF1FEE9F8AB.taxon	diagnosis	Diagnosis. Male of H. occidentalis is very similar to H. fairmairei but can be recognised by its large setal patches placed on the oval concave areas on the abdominal ventrites 1 – 4 (Fig. 11 E). The species is restricted to Western and South Australia.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0CF94CFCF1FEE9F8AB.taxon	description	Description. Male. Length 28 – 36 mm. Body subparallel, slightly convex dorsally; head capsule length about 0.7 times width; mandibles shorter than head capsule, weakly curved towards ventral side; pronotum 0.7 times of head capsule length; elytra 3.5 times longer than pronotum. Antennal scape 1.2 times longer than antennomere 3; interorbital width over 3 times of eye length. Prosternal process width 0.75 times length of procoxal cavity, twice hypomeron width; mesoventral process narrower than prosternal process. Abdominal ventrite 1 moderately longer than each of remaining segments; ventrite 5 rounded at apex. Metatarsomeres 1 – 3 length ratio as 1: 0.7: 0.8; tarsomere 5 shorter than 1 – 3 combined. Female. Unknown.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0CF94CFCF1FEE9F8AB.taxon	etymology	Etymology. The species name is derived from the Latin word “ occidentalis ”, meaning “ western ”, indicating its type locality in Western Australia.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0CF94CFCF1FEE9F8AB.taxon	materials_examined	Types. Holotype. ♂ “ WA: Kalgarin, 30 Jan. 1976, Ex. Carnaby Coll. | ANIC 25 - 072899 ” (ANIC). Paratype ♂ “ WA: Kalgarin, 30 Jan. 1976, Ex. Carnaby Coll. | ANIC 25 - 072908 ” (ANIC); Other material examined (Fig. 18 A). Western Australia: ANIC (1 ♂): Kalgarin, Jan. 1976. South Australia: RDKC (1 ♂): Yalata, Feb. 1987, Sundholm, A., Bugeja, J.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0DFBC5FF13FEA1FDBB.taxon	synonymic_list	(Figs. 11 A – C, 18 B)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0DFBC5FF13FEA1FDBB.taxon	diagnosis	Diagnosis. Male of H. prionoides can be separat- ed from the remaining Hermerius species by the abdominal ventrites without sexual patches, the long weakly serrate antennae extending to elytral apex, and the pronotum covered with dense long hairs. The female of that species can be recognised by smooth pronotum surface bearing only sparse fine punctures, the lower eye lobes narrowly separated on ventral side of head, the short antennae, not extending to mid elytra, and the protibial surfaces glabrous with teeth along external margin.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB561B0DFBC5FF13FEA1FDBB.taxon	materials_examined	Types. Thomson (1864) described this species from a single specimen which he believed was a female, however Lameere (1885) found it to be a male, and he described and discussed the female characters of this species. Results of our specimen examinations and molecular sequence comparison support Lameere’s (1885) conclusion. Currently, there are two specimens (one male and one female) labelled as “ Lectotype ” in the MNHN. Here we specify the male specimen as the holotype of Dioclides prionoides Thomson, 1864, which bears the following labels “ Prionoides Thoms. Type 303, Adel. Austra. | ♂ Australia, King George Lam. | Th. TYPE ”. The female specimen cannot be recognised as a part of the type series as it was probably the female specimen discussed by Lameere (1885). Material examined (Fig. 27 J). Western Australia: AM (2 ♂♂): Salt Lake; John Forrest NP, Darling Ranges, Jan. 1971, G. A. Holloway and H. Hughes; ANIC (47 ♂♂; 6 ♀♀): 8: 11 km N of Geraldton, Dec. 1972, N. McFarland; 6: Horrocks Beach, Dec. 1985, K. and E. Carnaby; 3: Cliff Heads, Feb. 1985, K. and E. Carnaby; 3: Red Bluff, Caravan Park, Dec. 1971, N. McFarland; 3: Woodridge, Feb. 1985, K. and E. Carnaby; 2: 6 miles N of Mandurah, Jan. 1967, Upton, M. S.; 2: Bunbury, Whilock; 2: Claremont, Jan. 1955, M. M. W. Wallace; 2: Dongara, Feb. 1979, K. and E. Carnaby; 1 km N of Bowelling, Jan. 1979, P. Ollerinshaw; Albany Hwy, 25 km S Armadale, Jan. 2014, P. Kay; Albany, Feb. 1977, K. and E. Carnaby; Balcatta, Jan. 1971, K. D. Wright; Bullsbrook, Feb. 1979, K. and E. Carnaby; Collie, Jan. 1973, [Carnaby Coll.]; Coolimba Lake Hill via Leeman, Dec. 2014, D. G. Knowles; Deepdene, Karridale, Jan. 1967, Upton, M. S.; Esperance R., 13 km W Norseman, Jan. 1985, K. and E. Carnaby; Fremantle, Dec. 1963, K. R. Norris; Garden Island, Feb. 1935, K. R. Norris; Garden Island, Feb. 1980; Garden Island, Mar. 1951, A. B.; Guilderton; Jan. 1970, T. B. Ellas; Mt. Yokine, Jan. 1957; Nedlands, Dec. 1959, M. M. W. Wallace; Perry Lake, Perth, Feb. 1968, A. J. Mahon; Perry Lake, Perth, Jan. 1972; Pinjarra, Feb. 1986, K. and E. Carnaby; White Hills Rd., S Mandurah, Jan. 2015, P. Kay; [no data]; MV (13 ♂♂; 2 ♀♀): 3: Bunbury; 2: Geraldton; 2: Rottnest Island, E. T. Smith; Champion Bay; Dongara; Harvey, A. Douglas; Mandurah; Mundaring, Feb. 1958; Perth, Jan. 1955, A. Douglas; S. Perth, A. Douglas; South Perth, Jan. 1907; QM (1 ♂; 1 ♀): 2: [no data]. South Australia: MV (1 ♀): Adelaide, Schlapes Gully. New South Wales: AM (1 ♂): 50 km N of Warren, Dec. 1988, M. S. and B. J. Moulds. Queensland: MV (2 ♂♂): Edungalba, Dec. 1968, E. E. Adams; Mount Tamborine, Dec. 1957. Molecular data. BioSample ID: SAMN 11080868, SAMN 12650101.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB571B0AF973FD1DFF01FB8E.taxon	description	(Fig. 12)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB571B0AF973FD1DFF01FB8E.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 30 – 85 mm long. Dorsum usually uniformly dark brown, sometimes with yellowish or reddish elytra. Sexually dimorphic dense punctures present on pronotum and ventral side of thorax, except two elevated smooth islands in the centre of pronotal disc. Head narrower than prothorax; frontoclypeal suture straight or broadly angulate; median groove complete. Antennal tubercles weakly prominent, directed laterally, rounded at apex. Eyes medium size, weakly emarginate near antennal insertion, moderately separated dorsally. Mandibles 0.3 – 0.5 times of head capsule length, broad at base, weakly bent and unidentate apically; sexual dimorphism expressed by longitudinal carina on dorsal surface. Antenna extending beyond middle of elytra in Australian species but can reach elytral apex in some other species. Scape slightly shorter than head capsule length, posteriorly extending to anterior margin of pronotum; cylindrical or slightly flattened, gradually expanded; antennomere 3 distinctly shorter than scape and slightly longer than 4. Mentum not fused to submentum. Terminal palpomere of maxillary and labial palps weakly expanded and rounded apically. Prothorax transverse with lateral carina weakly dentate; anterior and posterior margins with complete beads. Pronotal disc uneven with weakly elevated smooth shiny callosities on densely punctate background. Prosternal process sub-parallel, extending to mesoventrite, rounded or narrowly pointed apically. Elytral surfaces punctate, punctures variable in different species; inner apical angle with sharp spine. Legs strong; femo- ra and tibiae surface with dense tiny nodules. Protibia without lateral spines, with one sharp projection and a pair of subequal spurs apically; lobes on tarsomere 3 moderate or extremely large, tarsomere 5 as long as or longer than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB571B0AF973FD1DFF01FB8E.taxon	description	Description. Male. Length 32 – 82 mm. Dorsum usually uniformly dark brown, elytra sometimes yellowish or reddish brown. Head slightly transverse, narrower than pronotum. Mandible with dorsal surface bearing longitudinal carina and dense setae along incisor edge; length of mandibles variable in different species, but at least longer than half of head capsule length, apex moderately curved inwards; each mandible wedgeshaped, thick and strong on lateral side and near the base, unidentate at apex, with one or two extra teeth on incisor edge. Labrum narrow, dorsal surface covered with long golden setae. Frontoclypeal suture straight or broadly angulate. Antennal tubercles weakly prominent, distant to each other, apex rounded. Antenna 11 - segmented, filiform, extending at least beyond middle of elytra, in some species reaching to elytral apices; scapes long, longer than 2 / 3 of head capsule length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; moderately separate dorsally. Submentum straight at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps weakly expanded and rounded apically. Pronotum sub rectangular and transverse, usually with prominent anterior angles; complete bead present on both anterior and posterior margins; lateral margin sharp with rather small and short teeth. Disc uneven with shiny elevated triangular admedian islands among dense punctures elsewhere. Prosternum covered with fine and deep punctures; hypomeron welldefined, distinctly broader than prosternal process. Prosternal process projecting towards mesoventrite, sub-parallel with narrowly pointed apex. Mesoventral process emarginate at apex; surface flat. Ventral side of pterothorax and hind coxae covered with golden hairs. Metanepisternum slightly narrowing posteriorly. Scutellum narrowly rounded apically, surface rather smooth and shiny. Elytra surface with variable punctures and pubescence in different species: in O. tyrannus tyrannus elytra with fine sparse punctures entirely and recumbent fine yellowish white sparse pubescence; in O. tyrannus salomonum elytra with fine sparse punctures on base, and only sparse hairs along lateral margin; in O. laevipennis elytra smooth and completely glabrous; in O. scabripennis elytra with dense coarse punctures entirely. Elytra without traces of wing venation; elytral apices rounded with distinct and sharp sutural spine; epipleuron broad, complete, narrowing apically. Legs strong and moderately long; femur sub-parallel and relatively flat, surface very rough, covered with dense tiny nodules especially on front legs, ventral side without short spines apically; each tibia with distinct apical projection and a pair of strong spurs; tarsi moderately strong; tarsomere 3 with median or large lobes; tarsomere 5 as long as or longer than tarsomere 1 and 2 combined. Abdominal ventrites with sparse long hairs or few fine punctures, but bearing dense long setae along the edges, especially on the apex of ventrite 5. Male genitalia. Tegmen longer than sternite VII; parameres moderately long, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex about truncate while ventral apex narrowly pointed. Female. Mandibular surface without longitudinal carina. Pronotal disc with coarser punctures on sexual punctate area; fine and dense punctures on prosternum absent.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB571B0AF973FD1DFF01FB8E.taxon	discussion	Remarks. The genus Olethrius was discussed by Vitali (2008) who clarified taxonomic statuses of many species in the Australo-Pacific Region. According to this publication potentially two or three species may occur in Australia, however all the Australian records are based on very old and doubtfully labelled specimens, including the type locality of O. lavipennis Vitali said to be “ Australia ”. We have examined Prioninae collections from all the major world and Australian museums and found only a single specimen of Olethrius tyrannus salomonum labelled “ Richmond River ”, the locality located in the northern New South Wales, which is unlikely locality for this species. Mitochondrial genome was generated for this particular sample.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBE7FF12FC70FD4E.taxon	diagnosis	Diagnosis. Olethrius laevipennis can be recognised by the following characters: elytra are strongly convergent and longitudinally ridged apically, elytral surface entirely glabrous and completely unpunctuat- ed, the antennal scape is relatively shorter, flattened and sparsely punctured on the dorsal surface.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBE7FF12FC70FD4E.taxon	materials_examined	Type. Holotype, male, “ Australie, ex. Coll. de Moffarts | Olethrius tyrannus var. laevipennis Lam. Type | Holotype Olethrius laevipennis ♂, Vitali 2008 ” (IRSN). Notes. So far, this species is only known from the holotype.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBC6FD16FC8BF960.taxon	description	(Figs. 12 A – B, G, J)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBC6FD16FC8BF960.taxon	diagnosis	Diagnosis. Olethrius scabripennis can be recognised by the following combination of characters: the pronotal smooth sexual lateral area subquadrate, the medial area short and broad; the lateral carina of pronotum with numerous teeth and acutely produced; the elytra with dense coarse punctures and recumbent yellow pubescence. Male mandible without distinct longitudinal carina.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBC6FD16FC8BF960.taxon	materials_examined	Type. Olethrius scabripennis Thomson. Lectotype male in MNHN, with the following labels “ Scabripennis Thoms. Type, s. c. Ap., Ils Fegee | Museum Paris Coll. J. Thomson 1952 | Th. TYPE ”. Material examined. Fiji: AM (1 ♂): Fulton College, N. of Suva, N. Poultet, Jan. 1975; ANIC (1 ♂): Viti Levu, Rakiraki, B. P. Moore, Nov. 1974. New Caledonia: ANIC (1 ♀): Moindou, B. P. Moore, Jan. 1966.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0AFBC6FD16FC8BF960.taxon	discussion	Remarks. According to Vitali (2008) O. scabripennis is widespread from the southern Solomon Islands to Tonga, through Vanuatu, Fiji and New Caledonia, and it was also recorded from “ Australia ”, but no new material is available to confirm this.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0BFB24F94BFE45FC23.taxon	description	(Figs. 12 C – D, H, K)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0BFB24F94BFE45FC23.taxon	diagnosis	Diagnosis. O. tyrannus tyrannus can be recognised by the elytra surface with fine sparse punctures entirely, bearing short recumbent fine yellowish white pubescence.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0BFB24F94BFE45FC23.taxon	materials_examined	Type. Olethrius tyrannus Thomson, Lectotype male in MNHN, with the following labels “ Olethrius Thoms. E. 316 | tyrannus Thoms. Type, N. iles Hebrides | Museum Paris Coll. J. Thomson 1952 | Th. TYPE ”. Material examined. Vanuatu: ANIC (1 ♂): S. Bule, Efate, Jul. 2004; OAI (1 ♂): Banks Island, 1914, Allen. Locality?: OAI (1 ♀): “ Port Sunlight, Lever Bros, 4.5.09 / ex W. W. Froggatt Collection ”.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB501B0BFB24F94BFE45FC23.taxon	discussion	Remarks. The specimen from OAI from the W. Froggatt collection bears labels similar to the specimens of O. t. salomonum collected from the Lever Bros plantations in the Solomon Islands but that locality does not agree with its known distribution and is here treated as doubtful.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B0BF883FB85FD0BF8AA.taxon	description	(Figs. 12 E – F, I, L)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B0BF883FB85FD0BF8AA.taxon	diagnosis	Diagnosis. O. tyrannus salomonum can be recognised by the antennal scape cylindrical (Fig. 12 I) and covered with very dense fine punctures, the pronotum finely punctate on the elevated shiny area, nearly smooth on the sides, and the elytra with fine sparse punctures near base, smooth apically, only bearing sparse hairs along lateral margin.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B0BF883FB85FD0BF8AA.taxon	materials_examined	Type. Holotype male in IRSN, with the following labels “ Buin, Ile Bougainville, N. ile Guinee, Ex. 7, Coll. Muller | Holotypus ♂, Olethrius tyrannus salomonum, mihi, det. F. Vitali 2008 ”. Material examined. Australia?: AM (1 ♂): [New South Wales] Richmond River. Solomon Islands: ANIC (9 ♂♂; 2 ♀♀): Near Lever Bros Pacific Plantation Co. Lever Brothers, Jan. 1907; 5: W. W. Froggatt, Jul. - Aug. 1909; 5: [no data]; OAI (15 ♂♂; 1 ♀): “ longicorn beetle reputed to be a Coconut pest. Solomon Is ”: 3: Port Sunlight, Lever Brothers, May. 1909; 5: W. W. Froggatt Collection, Jul. - Aug. 1909; 4: Sep. 1910; 4: Hickey, Nov. 1911. Molecular data. BioSample ID: SAMN 11080872.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B08FB81FF12FB2BFE65.taxon	description	(Figs. 13 A – C, 14 L, 15 O, 17 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B08FB81FF12FB2BFE65.taxon	diagnosis	Diagnostic combination (Male). Moderately large beetles, 25 – 45 mm long. Dorsum unicoloured, brown or dark brown. Sexually dimorphic dense punctures present on most of the pronotum, scutellum and most of the ventral side of prothorax and abdomen. Head narrow- er than prothorax; frontoclypeal suture arcuate to broadly angulate; median groove complete. Antennal tubercles prominent, rounded at apex. Eyes large, weakly emarginate near antennal insertion, narrowly separated dorsally. Mandibles about 1 / 2 of head capsule length, not sexually dimorphic, broad at base, not bent towards ventral side and unidentate apically. Antenna extend to apical elytra in male. Scape distinctly longer than half of head capsule length, posteriorly extending beyond posterior margin of eye, gradually expanded; antennomere 3 longer than scape, twice length of antennomere 4 and thicker than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps oval and apically rounded. Prothorax transverse with lateral carina weak, sometimes incomplete but regularly dentate; anterior and posterior margins with complete bead. Pronotal disc with smooth and coarsely punctate callosities on densely punctate, feebly shiny background. Prosternal process subparallel, extending well beyond procoxae, rounded apically. Elytra surfaces coriaceous; inner apical with sharp spine. Legs relatively long; rows of spines present on femora and tibiae except for hind tibiae. Protibia with ventral surface bearing several short spines, densely setose apically, with pair of unequal spurs; protarsi bearing both long hairs and short setae; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB511B08FB81FF12FB2BFE65.taxon	description	Description. Male. Length 25 – 45 mm. Body uniformly brown or dark brown. Head approximately as long as wide, narrower than pronotum. Mandibles about 1 / 2 of head capsule length, weakly curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two or three teeth along incisor edge. Labrum weakly concave near apex, anterior margin truncate with very dense setae; labrum separated from clypeus by deep groove; dorsal surface of labrum with longer and denser setae than on clypeus. Frontoclypeal suture arcuate to broadly angulate; median groove distinct and complete. Antennal tubercles relatively prominent, apex rounded. Antenna 11 - segmented, filiform, extending almost to elytral apices; scape moderately long and gradually expanded, reaching beyond posterior margin of eye; antennomere 3 longer than scape, over twice length of antennomere 4, and apically thicker than all following segments; antennomeres 5 – 11 with rather blunt posterior angles. Eyes large, convex, coarsely facetted, slightly emarginate near antennal foramen; narrowly separated dorsally. Submentum curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps oval and apically truncate. Pronotum transverse, trapezoid with posterior margin broader than anterior margin; complete bead present on both anterior and posterior margins; lateral carina weak sometimes incomplete with regular sharp teeth. Disc very densely punctate and matt except for shiny, coarsely and deeply punctate islands consisting of two admedian triangular areas, two smaller lateral areas and transverse area in front of the posterior margin. Prosternum covered with fine and deep punctures; hypomeron well-defined, slightly narrower than prosternal process. Prosternal process extending beyond coxae, projecting towards mesoventrite, rounded apically. Mesoventral process sub-parallel and emarginate apically. Ventral side of thorax and coxae with dense golden hairs. Metaventrite with shiny arrow-shaped area medially, finely and densely punctate elsewhere. Metanepisternum weakly constricted at base. Scutellum rounded apically; surface with very fine punctures. Elytral surfaces dull covered with irregular coriaceous sculpture; without traces of venation; elytral apices rounded with sharp sutural spine; epipleuron complete, narrowing apically. Legs strong and relatively long; femora sub-parallel, surface rough, rows of spines present on ventral side; tibia slender and relatively flat, with spines along outer margin on front- and mid-tibia, bearing sensory setae on ventral side apically; each tibia with distinct apical spine and a pair of unequal spurs. Protibia with several spines along external edge, ventral surface with short spines, densely setose apically. Tarsi strong; tarsomere 3 moderately lobed, front tarsi covered with dense long hairs and short setal patch; tarsomere 5 shorter than tarsomeres 1 and 2 combined. Abdomen with ventrites 1 – 5 densely punctate basally with smooth and appearing less sclerotised apical part; ventral surface almost glabrous, only bearing long setae along the edge, and very densely setose on the apex of ventrite 5. Male genitalia (Fig. 14 L). Tegmen longer than sternite VII; parameres moderately long, 0.17 times length of entire tegmen, rounded apically with triangular projections at the base; penis as long as tegmen, dorsal apex truncate while ventral apex narrowly rounded. Female. Antennae usually shorter than in male, only extending to middle of elytra; antennomere 3 longer than scape, but not thicker than antennomeres 4 – 11. Pronotal disc shiny, surfaces irregular with smooth central area and coarsely punctate lateral area; lateral carina complete and with longer spines. Fine and dense punctures on thorax and abdomen absent. Legs much slender than on males. Abdominal ventrite 5 truncate or weakly emarginate medially. Ovipositor (Fig. 15 O) long, apical sclerosed part less than half length of baculus, distal gonocoxites very short and stout, abruptly narrowed near apex; stylus large, inserted laterally and very close to gonocoxite apex, gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB521B09FB0FFE44FD0FFE04.taxon	description	(Figs. 13 A – C, 14 L, 15 O, 17 E)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB521B09FB0FFE44FD0FFE04.taxon	diagnosis	Diagnosis. Pseudoplites hamali is morphologically similar to species of Geoffmonteithia, Eurynassa and some species of Cnemoplites. It can be separated from G. queenslanda by having distinct lateral spines on protibia and distinctly longer prosternal process extending well beyond procoxae. It is distinguished from all Eurynassa species by having elongate antennomere 3. Pseudoplites hamali can be separated from Cnemoplites species by having shiny, smooth area on the centre of pronotum.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB521B09FB0FFE44FD0FFE04.taxon	materials_examined	Types. Cnemoplites (Cnemoplites) hamali Lameere, Holotype female deposited in IRSN, with the following labels “ Cbenoplites hamali Lamee., Type ♀ | hamali Lameere ex. coll. de Moffarts ”. Macrotoma (Pseudoplites) inexpectata Lameere, Lectotype male deposited in MNHN, with the following labels “ Macrotoma inexoectata, A. Lameere vid. 1916 | TYPE | Victoria | Pseudoplites inexpectatus Lam. Lectotype ♂, Quentin and Villiers det. 1979 ”. Material examined (Fig. 17 E). Queensland: AM (3 ♂♂; 3 ♀♀): Byfield, Dec. 1975, F. T. Fricke; Cairns; Kuranda, May 1971, R. A. Faragher; Kuranda, Nov. 1969, R. A. Faragher; L. M. P.; [no data]; ANIC (7 ♂♂): 2: Smithfield, Cairns, Feb. 2014, J. Hasenpusch; 2: 14 km ENE Heathlands, Nov. 1993, P. Zborowski and M. Horak; Bucasia, Nov. 2005, K. J. Sandery; Crystal Falls, Paluma, Jan. 1968, R. Dorson; Smithfield, Cairns, Jan. 2014, N. Laird; MV (1 ♂; 1 ♀): Cooktown; [no data]; QM (2 ♂♂; 1 ♀): Cape Hillsborough, Jan. 1979; 2: Fraser Island, Dec. 1979, K. J. Lambkin; QVMAG (2 ♂♂; 3 ♀♀): 2: Maroochy River Resort, Maroochydore, Dec. 2014, S. Fearn; 2: Rowes Bay, Townsville, Nov. 2000, S. Fearn; School Road, Buderim, Dec. 2014, S. Fearn; RDKC (3 ♂♂ 4 ♀♀): 3: Cairns, {from Jan. 2015 to Jan. 2016}, N. Laird; 2: Smithfield, Cairns, Jan. 2014, N. Laird; Bald Hills Stn. Nr. Cookton, Dec. 1986, M. S. Moulds; Bloomfield, Dec. 2015, S. Lamond. Molecular data. BioSample ID: SAMN 12650112, SAMN 12650113, SAMN 12650114, SAMN 11080877. Notes. Our examination of both female types found them being morphologically identical and distinguished from Cnemoplites females by the following characters: antennomere 3 long, about 1.5 times of antennal scape and twice as long as antennomere 4; tarsi surface very rough, covered with spines and dense nodules; the whole body more or less flattened dorsally.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB531B06F905FDA6FDE7FF2F.taxon	description	(Figs. 13 D – F, 15 A, 15 P, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB531B06F905FDA6FDE7FF2F.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 35 – 55 mm long. Head and pronotum black, elytra brown or light brown. Sexually dimorphic dense fine punctures present near lateral margin of pronotum and most of ventral side of prothorax. Head slightly narrower than prothorax; frontoclypeal suture straight to broadly angulate; median groove complete. Antennal tubercles almost flat, rounded apically. Eyes transverse, weakly emarginate near antennal insertion, broadly separated dorsally. Mandibles 2 / 3 of head capsule length, sexually dimorphic, not bending towards ventral side and unidentate apically. Antenna reaching to middle of elytra. Scape about 1 / 4 of head length, posteriorly not reaching posterior margin of eye, gradually expanded apically; antennomere 3 as long as scape or antennomere 4. Mentum not fused to sub-mentum, constricted underneath sub-mentum and not visible ventrally. Terminal palpomere of maxillary and labial palps oval and apically weakly truncate. Prothorax transverse with lateral carina moderately strongly dentate; anterior and posterior margins with complete bead. Pronotal disc mostly shiny, with fine and dense punctate, slightly depressed lateral irregular areas. Prosternal process sub-parallel, extending to mesoventrite, rounded apically. Elytral surface shiny with irregular coriaceous sculptures; inner apical angle with short spine. Legs moderately strong, rows of small spines present on all femora and protibia. Protibia with two sharp apical projections and pair of subequal spurs; tarsomere 3 moderately lobed, tarsomere 5 as long as tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB531B06F905FDA6FDE7FF2F.taxon	description	Description. Male. Length 35 – 55 mm. Dorsum glabrous and shiny; head and pronotum black, elytra uniformly reddish brown to dark brown. Head oval, slightly narrower than pronotum. Mandibles much longer and stronger in males; dorsum bearing short setae, apex curved inwards; each mandible wedgeshaped, very thick and strong on lateral side and near the base, unidentate at apex, with two extra teeth on incisor edge. Labrum transverse, anterior margin pointed medially with relatively dense setae; labrum separated from clypeus by a deep groove; dorsal surface of clypeus with sparse short setae. Frontoclypeal suture almost broadly angulate; median groove distinct and complete. Antennal tubercles relatively flat, very distant to each other, apex rounded. Antennaa 11 - segmented, filiform, extending to middle of elytra; scapes short, slightly longer than eye. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; broadly separated dorsally. Submentum weakly curved at apex, mentum restricted and not visible ventrally. Terminal palpomere of maxillary and labial palps oval and apically rounded. Pronotum transverse, width 2.5 times of length, with anterior margin emarginate medially; complete bead present on both anterior and posterior margins; lateral carina distinct and dentate. Disc shiny, glabrous and with sparse punctures medially; lateral and anterolateral area with deep and fine punctures. Prosternum bearing uniformly deep fine punctures; hypomeron well-defined, slightly broader than prosternal process; prosternal process sub-parallel, projecting towards mesoventrite, narrowly rounded apically. Mesoventral process relatively broad, apex emarginate medially. Ventral side of pterothorax and coxae covered with golden hairs, shiny and glabrous arrow-shapped area present in central part of metaventrite. Metanepisternum slightly broader posteriorly, but sharply constricted at base. Scutellum broadly rounded at apex, surface smooth and shiny. Elytral surfaces shiny with very fine and weak irregular coriaceous sculpture, two or three traces of venation present; elytral apices rounded with small sutural spine; epipleuron complete. Legs strong, femur broad and relatively flat, constricted at both ends, surface smooth, ventral side with several tiny spines; each tibia with distinct apical projections and a pair of subequal spurs; tarsi relatively strong; tarsomere 3 moderately lobed; tarsomere 5 longer than tarsomere 1 and 2 combined. Abdominal ventrites smooth, only bearing dense long setae along the edge, especially on the apex of ventrite 5. Male genitalia (Fig. 15 A). Tegmen longer than sternite VII; parameres moderately long, 0.15 times length of entire tegmen, truncate apically with small triangular projections at the base; penis about as long as than tegmen, dorsal apex broadly rounded while ventral apex narrowly pointed. Female. Mandibles much shorter than in males. Pronotum weakly emarginate at anterior margin; anterior angles prominent; lateral carina with longer lateral spines. Pronotal surface coarsely punctate, punctures coarser and irregular in lateral area; prosternum and hypomeron with sparse fine punctures and loose hairs, relatively shiny. Ovipositor (Fig. 15 P) moderately long, apical sclerosed part 2 / 3 of baculus length; distal gonocoxites short and small; stylus inserted laterally and close to gonocoxite apex, relatively long and gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B06F951FE8EFEA1FADF.taxon	description	(Figs. 13 D – F, 15 A, 15 P, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B06F951FE8EFEA1FADF.taxon	diagnosis	Diagnosis. This species is morphologically similar to Cryptipus and some species of Eurynassa. It can be separated from C. frenchi by having pronotum length only about 0.4 times of its width, the elytral surface with irregular coriaceous punctures and the tarsomere 3 moderately lobed. Male T. insularis can be separated from Eurynassa species by having abdomen smooth and shiny, without fine deep punctures; while the female can be separated from Eurynassa species by having strongly restricted mentum, not visible ventrally and the ovipositor with longer and larger styli.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B06F951FE8EFEA1FADF.taxon	materials_examined	Type. Teispes dorsalis Thomson. Holotype male in MNHN, with the following labels “ Teispes Thoms., s. c. 304, (Nom pr.) | dorsalis thoms. Type. 305, Austr. bor. | Museum Paris Coll. J. Thomson, 1952 | Th. TYPE ”. Material examined (Fig. 18 C). Northern Territory: AM (4 ♂♂; 2 ♀♀): 5: Yirrkala, W. S. Chaseling; E. Alligator Riv., Oenpelli, Sep. 1948, J. E. Bray; ANIC (1 ♀): Baroalba Creek Springs, 19 km NE by E of Mt. Cahill, Oct. 1972, M. S. Upton. Western Australia: RDKC (1 ♂): Weelarrana, Oct. 1996, D. Knowles. Molecular data. BioSample ID: SAMN 11080880, SAMN 12650055.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B07F925FAFEFF3CFB22.taxon	description	(Figs. 13 G – I, 15 B, 15 Q, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B07F925FAFEFF3CFB22.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 45 – 65 mm long. Dorsum brown to dark brown, elytra often weakly yellowish brown near apex. Sexually dimorphic dense fine punctures present near lateral margin of pronotum and ventral side of prothorax. Head as broad as or slightly narrower than prothorax; frontoclypeal suture straight or broadly arcuate; medi- an groove complete. Antennal tubercles almost flat, pointed apically. Eyes transverse, weakly emarginate near antennal insertion, broadly separated dorsally. Mandibles shorter than head capsule, sexually dimorphic, weakly arcuate and bidentate apically. Antenna extending beyond elytral humeral angle in males. Scape very short, less than 1 / 4 length of head capsule, posteriorly extending only to middle of eye, moderately expanded apically; antennomere 3 longer than scape and slightly longer than antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps elongate and apically rounded. Prothorax transverse with lateral carina weakly dentate; anterior margin without bead; posterior margin with complete bead. Pronotum distinctly shiny and smooth, with fine and shallow punctures laterally. Prosternal process sub-parallel with lateral carina, extending to mesoventrite, narrowly rounded apically. Elytra surface glabrous and shiny; inner apical angle bluntly angulate. Legs strong, spines absent on femora and tibia. Protibia smooth, apices with two sharp projections and pair of subequal spurs; lobes on tarsomere 3 very narrow and short, tarsomere 5 much longer than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B07F925FAFEFF3CFB22.taxon	description	Description. Male. Length 45 – 60 mm. Dorsum glabrous and shiny, brown to dark brown, with elytra apical area usually lighter in colour. Head slightly transverse, narrower than pronotum. Mandibles narrow and elongate strongly arcuate apically, bidentate at apex, without extra teeth on incisor edge; dorsum bearing dense short hairs near base. Labrum transverse, anterior margin strongly concave and emarginate medially, covered with dense setae; labrum separated from clypeus by a deep arcuate groove; dorsal surface of labrum and clypeus with dense erect long setae. Frontoclypeal suture almost straight or broadly arcuate; median groove distinct and complete. Antennal tubercles relatively flat, very distant to each other, apex moderately pointed. Antenna 11 - segmented, filiform or weakly monoliform, extending to or shortly beyond elytral humeral angle; scape short, shorter than eye length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; broadly separated dorsally. Submentum weakly curved at apex, mentum well exposed; terminal palpomere of maxillary and labial palps elongated oval and apically rounded. Pronotum transverse, trapezoid with posteri- or margin slightly narrower than anterior margin, anterior margin emarginate medially and without bead; posterior margin with complete bead; lateral carina dentate. Disc shiny, glabrous and smooth, with shallow and fine punctures on lateral area. Prosternum bearing uniformly shallow fine punctures; hypomeron well-defined, same width as prosternal process. Prosternal process sub-parallel with complete lateral carinae, projecting towards mesoventrite and narrowly rounded apically. Mesoventral process as broad as prosternal process, apex emarginate medially. Ventral side of pterothorax and coxae covered with golden hairs, except for a shiny triangular central area on metaventrite. Metanepisternum weakly constricted at base. Scutellum rounded at apex, surface smooth and shiny. Elytra smooth, glabrous and shiny, without any traces of venation; elytral apices rounded or bluntly angulate without sutural spines; epipleuron almost complete, very strongly narrowing apically. Legs strong, femur broad and relatively flat, weakly constricted at both ends, surface smooth and shiny, ventral side without rows of spines; each tibia with two distinct apical spines and a pair of long spurs; tarsi slen- der; tarsomere 3 with very narrow and short lobes; tarsomere 5 much longer than tarsomere 1 and 2 combined. Abdominal ventrites moderately smooth and shiny; each ventrite with small finely punctate area near anterior angle; ventrites bear sparse fine hairs and dense long setae along the edges, especially on the apex of ventrite 5. Male genitalia (Fig. 15 B). Tegmen longer than sternite VII; parameres long, about 0.17 times length of entire tegmen, slightly truncate apically with small triangular projections at the base; penis about as long as tegmen, dorsal apex broadly truncate while ventral apex narrowly rounded. Female. Mandibles shorter than in male, wedgeshaped, thick and strong with broad base, strongly arcuate apically, bearing single apical tooth and one subapical tooth on inner edge. Pronotum more rectangular; surface completely smooth and glabrous. Prosternum and hypomeron with sparse hairs, relatively smooth and shiny. Ovipositor (Fig. 15 Q) short and strong, apical sclerosed part less than half of baculus length; distal gonocoxites moderately long and very sharp; stylus inserted laterally and distant to gonocoxite apex, relatively short and gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5C1B07F925FAFEFF3CFB22.taxon	discussion	Remarks. Monotypic genus distributed only in Australia.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B07F97DFA84FA6AFD7C.taxon	description	(Figs. 13 G – I, 15 B, 15 Q, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B07F97DFA84FA6AFD7C.taxon	diagnosis	Diagnosis. Male Utra can be separated from Cryptipus by having strongly elongate and narrow mandibles and very fine shallow punctures on pronotum. The female of that species is distinguished from C. frenchi by having completely smooth pronotum and very narrow and short tarsomere 3.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B07F97DFA84FA6AFD7C.taxon	materials_examined	Material examined (Fig. 18 C). Queensland: AM (1 ♀): Atherton, Jan. 1977, F. T. Fricke; ANIC (1 ♂; 11 ♀♀): Black MT Rd., 2 m N of Kuranda, Dec. 1967, J. G. Brooks; Cairns, Dec. 1947, J. G. Brooks; Cairns, Nov. 1970, [Carnaby Coll.]; Cardstone, Oct. – Dec. 1972, D. Perkins; Crystal Cascades, Cairns, Jan. 1955, J. G. Brooks; Kuranda, 19 Bulter Dr., Dec. 2003, D. C. F. Rentz; Kuranda, 19 Bulter Dr., Dec. 2009, D. C. F. Rentz; Kuranda, Jan. 1978, D. Larie; Mossman Gorge, Dec. 1967, R. Dobson; Polly Ck., Garradunga, Jan. 2014, J. Hasenpusch; The Boulders Babinda, Dec. 1967, R. Dobson; Yungaburra, Jan. 1946, K. R. Norris; MV (2 ♂♂; 1 ♀): Mareeba, Jan. 1950; Cairns; [no data]; OAI (1 ♂): Kuranda State Forest, Jan. 1969, N. C. Coleman; QM (1 ♂; 3 ♀♀): 2: “ Chowchilla ”, Jan. 1988, W. E. and W. T. Cooper; Little Mulgrave, Dec. 1949; Cairns, Dec. 1969.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B07F97DFA84FA6AFD7C.taxon	discussion	Remarks. This species has been recorded from the Northern Territory, Queensland and New South Wales (McKeown, 1947), but so far, all the specimens we have examined were collected from the Northern Queensland. In spite of our efforts we have not found the type specimen of Utra nitida at the MNHN. Molecular data. BioSample ID: SAMN 11080882.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B04FB97FD5FFA7AFD99.taxon	description	(Figs. 13 J – M, 15 C, 15 R, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B04FB97FD5FFA7AFD99.taxon	diagnosis	Diagnostic combination (Male). Large beetles, 45 – 70 mm long. Head and pronotum dark brown while elytra reddish brown. Pronotum without sexually dimorphic dense fine punctures. Head narrower than prothorax; frontoclypeal suture broadly angulate; median groove complete. Antennal tubercles moderately prominent, rounded apically. Eyes transverse, weakly emarginate near antennal insertion, closely separat- ed dorsally. Mandibles about 2 / 3 length of head, sexually dimorphic on the length, not bending towards ventral side and unidentate apically. Antenna reaching elytra apex. Scape longer than half of head length, posteriorly extending beyond the posterior margin of eye, gradually expanded apically, surface covered with small spines; antennomere 3 longer than scape and antennomere 4. Mentum not fused to sub-mentum. Terminal palpomere of maxillary and labial palps elongate and weakly expanded, apices rounded. Prothorax transverse with lateral carina regularly dentate; anterior margin with incomplete bead while posterior margin with complete bead. Pronotal disc bearing dense adpressed hairs and sparse tiny nodules. Prosternal process sub-parallel, extending to mesoventrite, round- ed apically. Elytra surface covered with adpressed golden hairs; inner apical angle with sharp spine. Legs long, sharp spines present on femora and tibia. Protibia surface very rough with two sharp apical projections and single spur; tarsomere 3 moderately lobed, tarsomere 5 shorter than tarsomeres 1 and 2 combined.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B04FB97FD5FFA7AFD99.taxon	description	Description. Male. Length 45 – 70 mm. Dorsum covered with dense adpressed short hairs, head and pronotum dark brown while elytra reddish brown. Head weakly elongate, narrower than pronotum. Mandibles sexually dimorphic, much longer in males, dorsum bearing sparse hairs, apex curved inwards; each mandible wedge-shaped, very thick and strong on lateral side and near the base, unidentate at apex, with two extra teeth on incisor edge. Labrum transversely oval, anterior margin weakly emarginate medially; labrum separated from clypeus by a membrane-like groove; dorsal surface of clypeus with dense adpressed hairs as the remaining head. Frontoclypeal suture angulate; median groove distinct and complete. Antennal tubercles moderately prominent, relatively close to each other, apex rounded. Antennae 11 - segmented, filiform, extending towards apex of elytra; scapes long, much longer than eye length. Eyes transverse, coarsely facetted, very weakly emarginate near antennal foramen; closely separated dorsally. Submentum not curved at apex, mentum well exposed. Terminal palpomere of maxillary and labial palps elongate and weakly expanded, apices rounded. Pronotum transverse, almost semi-circle; anterior margin with incomplete bead while posterior margin with complete bead; lateral carina regularly dentate, posterior margin weakly dentate near humeral angle. Disc surface uniformly setose, with sparse tiny nodules. Prosternum with dense irregular punctures; hypomeron welldefined, about same width as prosternal process; prosternal process sub-parallel, projecting towards mesoventrite and rounded apically. Mesoventral process relatively broad, apex emarginate medially. Ventral side of pterothorax and coxae covered with golden hairs. Metanepisternum weakly constricted at both ends. Scutellum rounded or slightly emarginate at apex, surface densely setose. Elytra surfaces covered with golden adpressed hairs, venation present on apical 1 / 3 of elytra; elytral apices rounded with sharp sutural spine; epipleuron complete, narrowing towards apex. Legs long and strong, femur sub-parallel and relatively flat, surface rough especially on front femur, ventral side with rows of tiny spines; each tibia with two distinct apical projections and single strong spur; tarsi slender; tarsomere 3 moderately lobed; tarsomere 5 shorter than tarsomere 1 and 2 combined. Abdominal ventrites surface completely pubescent, and with dense long setae along the edge, especially on the apex of ventrite 5. Male genitalia (Fig. 15 C). Tegmen longer than sternite VII; parameres long, about 0.2 times length of entire tegmen, rounded apically without small triangular projections at the base; penis as long as tegmen, dorsal apex broadly rounded while ventral apex narrowly pointed. Female. Mandibles much shorter than in male, less than half of head capsule length; antennae shorter, only extending to middle of elytra; pronotum trapezoid with relatively distinct anterior angles; surface of antennal scape and front femur rough, but not as strongly dentate as in males; prosternum and hypomeron without dense irregular punctures. Ovipositor (Fig. 15 R) long, apical sclerosed part less than half of baculus length; distal gonocoxites short and small; stylus inserted laterally and close to gonocoxite apex, relatively long and gradually expanded apically.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5D1B04FB97FD5FFA7AFD99.taxon	discussion	Remarks. There are currently 26 species or subspecies recognised in the genus Xixuthrus widely distributed in Indo-Australian region, with single species- X. domingoensis Fisher, from the Dominican Republic (Titan database, Tavakilian and Chevillotte 2019).	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5E1B05FBD6FD38FD3DFB4D.taxon	description	(Figs. 13 J – M, 15 C, 15 R, 18 C)	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5E1B05FBD6FD38FD3DFB4D.taxon	diagnosis	Diagnosis. The Australian species can be easily separated from other Australian Prioninae in having entire body setose, the protibia with single spur apically and with random spines not arranged in rows.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5E1B05FBD6FD38FD3DFB4D.taxon	materials_examined	Types. Macrotoma microcera White, Holotype male in BNHM (BNHM- 010799863), with the following labels “ M. microcera, n. s., Java | Macrotoma microcera White Type | 49 - 5 ”. Xixuthrus nycticorax Thomson, Holotype male in MNHN, with the following labels “ nycticorax, Type, An. Fr. 1877, Bull. 226, Thoms., Austral. | Th. TYPE ”. Material examined (Fig. 18 C). Queensland: ANIC (19 ♂♂; 18 ♀♀): 11: Kuranda, 19 Butler Dr., {from Dec. 2001 to Mar. 2018}, D. C. F. Rentz; 2: Cape York, May 2015, A. Zwick; 2: Cardstone, Jan. 1972, P. Zborowski; 2: Gordonvale, Nov. 1990, R. Hardie; 2: Mission Beach, Nov. 1976, G. O’Reilly; 2: Polly Ck., Garradunga, Oct. 2013, J. Hasenpusch; 2: Shiptons Flat, Oct. 1980, T. Weir; Cairns, Dec. 1950, G. S.; Clump Point, Dec. 1949, J. O. Campbell; Cooktown, Jun. 1970, Le Souef; Edge Hill, Cairns, Jan. 1973, P. Winfield and J. G. Brooks; Gillies Rd., 6 mi. From Gordonvale, Dec. 1968, S. Misko; Julatten, Dec. 1999, B. P. Moore; Julatten, Jan. 1993, B. P. Moore; Little mulgrave valley, Nov. 1969, R. Parrot; Lync-Haven, Daintree area, Dec. 2012, [QM / ANIC]; Mission Beach, May 1993, G. O’Reilly; Palmerston Nat. Pk., Dec. 1967, R. Dobson; Pilgrim Sands, Feb. 1998, E. D. Edwards and R. Oberprieler; Polly Ck., Garradunga, Nov. 2010, J. Hasenpusch; Townsville, Jan. - Dec. 1970, C. Breuer; MV (5 ♂♂; 4 ♀♀): 3: Innisfail, Nov. 2009, L. J. Cookson; Atherton, Dec. 1910; 5: north Queensland; QM (8 ♂♂; 3 ♀♀): 2: Bloomfield Riv., Nov. 1965, G. Monteith; 2: Iron Range, Cape York Pen., Nov. 1965, G. Monteith; 4 km SW of road junct., West Claudie Riv., Nov. 1986, G. Daniels and M. A. Schneider; Cairns, Dec. 1966, B. Cantrell; Iron Range, Cape York Pen., Dec. 1995, S. J. Johnson; Iron Range, Cape York Pen., Dec. 1996, S. J. Johnson; Iron Range, Cape York Pen., May 1968, G. Monteith; Mossman, Dec. 1966, B. Cantrell; Tron Range, Oct. 1991, J. Hasenpusch. Molecular data. BioSample ID: SAMN 11080883.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
03ED87B7CB5E1B05FBD6FD38FD3DFB4D.taxon	discussion	Remarks. X. microcerus was originally described from Java but is widely distributed in the Australo- Pacific region and the taxonomic statuses of many isular populations have always been problematic. While treating the region Lameere (1912, 1919) recognised 5 allopatric sub-species X. microcerus, and further three separate species as X. heros (Fiji), X. ganglbaueri (Fiji) and X. costatus (Papua New Guinea). Marazzi et al. (2006) and Titarenko and Zubov (2018) reinstated many subspecies and local populations as valid species, but these actions cannot be assessed without molecular data that is currently not available. However, examination of the types revealed no major morphological differences between Macrotoma microcera White, 1853 and Xixuthrus nycticorax Thomson, 1877, and following Lameere (1912, 1919) we recognise only single species, Xixuthrus microcerus in the Australian fauna. We generated a complete mitochondrial genome sequences for the Australian specimen, but without sequences from other taxa / populations to compare with, no further conclusion can be made here.	en	Jin, Mengjie, Keyzer, Roger De, Hutchinson, Paul, Pang, Hong, Ślipiński, Adam (2020): A Review Of The Australian Macrotomini (Coleoptera: Cerambycidae: Prioninae). Annales Zoologici 70 (1): 33-96, DOI: 10.3161/00034541ANZ2020.70.1.003
