taxonID	type	description	language	source
03EA8794A131965DFDB7FB4DFE889B98.taxon	description	(Figures 1 – 7) Type locality Naksan sandy beach, east coast of South Korea (38 ◦ 06 ′ 20 ′′ N, 128 ◦ 38 ′ 46 ′′ E); washings of sandy sediments from a brackish water system (salinity 4 – 7 psu) near a small unnamed stream. Material examined Holotype female (NIBRV 0000238524) dissected on six slides, and paratype male (NIBRV 0000238525) dissected on four slides. Additional paratypes represented by 14 females (NIBRV 0000238526) and three males (NIBRV 0000238527) in ethanol. All specimens were deposited in the NIBR, Korea; seven females and four males were used for scanning electron microscopy while 12 females were used for DNA extraction. All specimens were collected from the type locality by J. Back on 23 April 2010. DNA-barcode (mtCOI) sequences and traces were submitted to GenBank. Description of female Body fusiform, slightly depressed dorsoventrally (Figure 1 A, B), with sensilla as illustrated. Total body length, 553 µm (n = 15, mean = 541 µm); largest width (153 µm) measured at posterior margin of cephalic shield. Body somites with well-developed arthrodial membranes. Cephalothorax and somites bearing P 2 – P 4 with crenulate posterior hyaline frill. Cephalothorax bell-shaped, with several sensilla; pleural areas weakly developed, posterolateral angles rounded. Urosomites (Figure 1 A – C), except for anal somite, with striations on hyaline frills dorsally and laterally. Fifth urosomite and anal somite without sensilla; the latter small, without anal operculum, but with well-developed flimsy serrate pseudoperculum arising from penultimate somite. Genital somite (second urosomite) and third urosomite completely fused forming genital double-somite; genital field located in posterior half of second urosomite (proximal half of genital double-somite), with copulatory pore positioned medially, and two small pores located on either side of copulatory pore. Caudal rami (Figures 1 C, 2 C, 7 H) rectangular, about 1.6 times as long as wide; with three transverse spinular rows dorsally; with spinules at bases of setae III (ventrally) and VII (dorsally), and around distal margin of rami (ventrally); secretory pores located dorsally near base of seta III and ventrally near base of seta IV; with seven setae, setae I – II located halfway down the outer margin; setae III – VII located in distal third of ramus; seta II short and pinnate; seta I bare; seta III plumose; seta IV well developed and plumose, seta V longest and pinnate in distal half; seta VI pinnate; dorsal seta VII bi-articulate at base, naked. Rostrum (Figure 1 A) well developed, with rounded tip, defined at base; with two small lateral sensilla. Antennule (Figure 2 A) slender, eight-segmented; segment 1 with short row of long spinules along anterior margin and one pinnate seta; segments 1 and 2 similar in length; segment 4 with sub-cylindrical process bearing one bare seta fused basally to aesthetasc; armature formula: 1 - [1 pinnate], 2 - [7 pinnate + 1 spinulose + 3 bare], 3 - [5 pinnate + 3 bare], 4 - [3 pinnate + (1 + ae)], 5 - [1 pinnate + 1 bare], 6 - [3 pinnate + 1 spinulose + 1 bare], 7 - [1 pinnate + 1 spinulose seta + bare], 8 - [5 bare + (2 + ae)]. Acrothek consisting of apical aesthetasc and two basally fused bare setae. Antenna (Figures 2 B, 7 A): coxa and basis without surface ornamentation. Exopod two-segmented; exp- 1 shorter than exp- 2, the former with one pinnate seta distally; exp- 2 with row of strong spinules apically and with four setae; armature formula: 1 - [1 pinnate], 2 - [3 pinnate + 1 bare]. Endopod two-segmented; enp- 1 with lateral seta, without surface ornamentation; lateral armature of enp- 2 consisting of two bare setae, one small spinulose element, and one pinnate seta; distal armature of enp- 2 consisting of four geniculate and two bare setae (one small bare seta laterally and one long bare seta fused at base to largest geniculate seta). Mandible (Figures 3 A, 7 B): coxa well developed, with process; gnathobase with six small blunt teeth and one pinnate seta at dorsal corner. Palp biramous; basis elongate, with four pinnate setae; exopod one-segmented, with three inner and two distal pinnate setae, and ornamented as figured; endopod two-segmented; enp- 1 as long as enp- 2, with two bare distal setae; enp- 2 with seven setae (two bare setae, three distal setae fused at base, and one apical long seta and one slender bare seta confluent at base). Maxillule (Figure 3 B): praecoxa subquadrate, without ornamentation, arthrite well developed, with five strong spines and one seta apically, three bare lateral elements, and two surface setae. Coxa with five pinnate setae. Basis with five setae. Exopod onesegmented, with four slender bare setae of different lengths and with lateral row of spinules. Endopod one-segmented, longer than exopod, with three apical, and two bare lateral setae. Maxilla (Figure 3 C): syncoxa with four endites. First endite small, with one pinnate and one bipinnate setae; second endite with two pinnate setae; third endite with two pinnate and one bare setae; distal endite close to third endite, with one pinnate lateral seta, two pinnate distal elements, and with one small process proximally (arrowed in Figure 3 C – inset 4). Allobasis with one bare seta, one stout claw-like element, and one bare accompanying seta. Endopod two-segmented; enp- 1 rectangular, with five slender setae; enp- 2 with one pinnate claw, two apical, and one naked lateral setae. Maxilliped (Figures 3 D, 7 C) three-segmented. Syncoxa with two rows of spinules. Basis elongate, with rows of spinules halfway down inner and outer margin. Endopod indistinctly segmented, with one bare seta proximally, one stout claw medially, two stout claw-like bi-articulate setae, and one small seta distally. P 1 (Figure 4 A): coxa and basis with spinules as figured; the latter with one outer bare and one inner pinnate seta. Exopod three-segmented, shorter than endopod; exp- 1 longest, with some spinules along outer margin and one pinnate outer spine; exp- 2 somewhat swollen distally, with one pinnate outer spine and one spinulose inner seta; exp- 3 with six spines / setae. Endopod prehensile; enp- 1 elongate and approximately as long as exopod, with row of spinules along outer margin and one long plumose inner seta; enp- 2 small, slightly longer than wide, apically with two strong claw-like setae and one small pinnate element. P 2 – P 4 (Figure 4 B – D): coxae with row of long (P 2) or slender (P 3 – P 4) spinules on outer distal corner. Basis with long inner spinules and with rows of spinules on anterior surface. Exopod three-segmented; exp- 1 with one strong pinnate outer spine; exp- 2 shortest, with one strong pinnate outer spine and one bare inner seta; exp- 3 with two pinnate outer spines, two plumose apical setae, and one bare (P 2 – P 3) or plumose (P 4) inner seta. Endopod two-segmented; enp- 1 with one plumose inner seta; enp- 2 as long as enp- 1 (P 2) or about 1.6 times as long as enp- 1 (P 3 – P 4), with one bare (P 2) or pinnate (P 3 – P 4) outer spine and two pinnate apical setae; with one pore close to inner distal corner, except for P 4. Armature formula as follows: exopod: P 2, 0.1.122; P 3, 0.1.122; P 4, 0.1.122; endopod: P 2, 1.021; P 3, 1.021 (1.020 in male); P 4, 1.021. P 5 (Figures 2 D, 7 E) with medially fused baseoendopods and discrete exopods. Baseoendopod with short pinnate outer basal seta; endopodal lobes shorter than exopods, each lobe with two apical pinnate setae of almost equal length, with long spinules along inner margin. Exopod well developed, with rows of spinules along inner and outer margins, with two pinnate apical setae and two spinulose short outer elements. P 6 (Figures 2 E, 7 D) represented by narrow transverse plate, armed with one bare inner seta and two pinnate outer elements of different lengths, the former longest. Description of male Slightly smaller and more slender than female, body length 498 µm (n = 4, mean = 488 µm) (Figure 5 A). Largest width (92 µm) measured at posterior margin of cephalic shield. Cephalothorax (Figure 5 A) slightly more angular and with more sensilla than in female. General body shape and ornamentation as in female except for separation of genital somite; additional sexual dimorphism in antennule, P 2, P 3, P 5 and P 6. Antennule (Figures 5 B, 7 F): eight-segmented, subchirocer; segment 6 swollen, largest. Aesthetascs on sixth and eighth segments. Sixth segment with surface suture dorsoanteriorly (arrowed in Figure 7 F). Armature formula: 1 - [1 pinnate], 2 - [4 bare + 1 pinnate], 3 - [5 bare + 3 pinnate], 4 - [3 bare + 3 pinnate], 5 - [2 bare + 1 pinnate], 6 - [5 bare + 6 pinnate + (1 + ae)], 7 - [4 bare], 8 - [7 bare + (2 + ae)]. P 2 (Figure 6 A) Basis with surface spinules and small outer seta. Exopod as in female. Enp- 1 as in female; enp- 2 armed with two long and one small bare setae apically, the latter smaller than homologous element in female. General shape, segmentation, and armature formula as in female. P 3 (Figure 6 B). Coxa, basis, and exopod as in female. Endopod two-segmented; enp- 1 with rows of long spinules along outer margin and with one inner plumose seta. Enp- 2 twice as long as enp- 1, with two apical setae and well developed outer apophysis. P 5 (Figure 6 C): baseoendopods confluent, forming large transverse plate; unarmed except for outer slender bare seta on either side. Exopod ovate, with row of spinules along inner margin; with two outer spinulose setae, one long apical bare seta and one small naked inner seta. P 6 (Figure 6 D): pair of P 6 symmetrical, fused medially. Each P 6 with one outer naked lateral seta proximally and one pinnate seta close to distal outer corner. Etymology The specific name refers to the type locality, the Naksan sandy beach area on the east coast of Korea.	en	Back, Jinwook, Lee, Wonchoel, Huys, Rony (2011): A new species of Remanea Klie, 1929 (Copepoda: Harpacticoida: Paramesochridae) with a redescription of the type species. Journal of Natural History 45 (47 - 48): 2939-2964, DOI: 10.1080/00222933.2011.622057, URL: http://dx.doi.org/10.1080/00222933.2011.622057
03EA8794A13E9644FE73FB3FFC899FA0.taxon	description	(Figures 8, 9) Original description. Klie (1929): 364 – 367; figs 44 – 56. Type locality. Germany, Kiel Bay, off Bülk lighthouse (vicinity of “ Nebenfahrwassertonne A ”); clean coarse sand, 10 m depth. Material examined Zoologisches Museum der Universität Kiel. Walter Klie collection. Two females (Cop 586, 589) and two males (Cop 584, 588) dissected on individual slides. Unfortunately the slides were partly dried out and not in a sufficiently good condition to make a complete detailed redescription. Description of female General shape of appendages similar to those given in Klie’s original description (Klie 1929). Caudal ramus (Figure 8 C) rectangular, about 1.5 times as long as wide; with three inner rows of spinules dorsally; with seven setae; setae I – II arising from anterior outer margin, setae III – VI around posterior margin, and seta VII in distal third of ramus; seta I short and bipinnate; seta II spinulose in distal half; seta III plumose; seta IV well developed and plumose, seta V longest and pinnate in distal half; seta VI spinulose; seta VII bi-articulate at base and naked. Ventral spinules present along posterior margin and at base of seta III. Antenna (Figure 8 A): basis with row of spinules distally. Exopod arising from distal margin of basis, two-segmented; exp- 1 as long as exp- 2; exp- 1 with one long pinnate seta; exp- 2 with two lateral and two pinnate apical setae. Enp- 1 with pinnate seta, without surface ornamentation; enp- 2 with two bare setae of different lengths and one spinulose seta laterally, with five geniculate setae and one bare seta fused with largest geniculate seta at base. P 1 (Figure 8 D): coxa well developed, with rows of spinules as figured. Basis with one inner and one outer pinnate seta. Exopod much shorter than endopod; exp- 1 longest, with outer spinules and one pinnate outer spine; exp- 2 somewhat swollen distally, with one pinnate outer spine and one long spinulose inner seta; exp- 3 with six spines / setae. Endopod prehensile; enp- 1 elongate, longer than exopod, with outer row of spinules, with one long plumose inner seta in proximal third; enp- 2 small, quadrangular, with two strong setae and one small pinnate apical element. P 5 (Figure 9 C) with medially fused baseoendopods and discrete exopods. Baseoendopod with pinnate outer basal seta; endopodal lobes narrow, shorter than exopod, each with one short outer seta and one long slender plumose inner seta, with row of spinules along inner margin. Exopod well developed, with inner and outer rows of spinules; with two short spinulose outer setae and two plumose apical elements (subequal in length). P 6 (Figure 8 B) forming small narrow transverse plate with one long outer and two bare inner setae; middle seta shortest. Description of male Sexual dimorphism expressed in antennule, P 3 endopod, P 5, P 6 and urosomal segmentation. P 3 (Figure 9 A). Coxa, basis, and exopod virtually similar to those of female (see Klie 1929). Endopod two-segmented; enp- 1 with outer rows of spinules and one inner plumose seta; enp- 2 longer than enp- 1, with two apical pinnate setae and with well developed outer apophysis (see inset in Figure 9 A). P 5 (Figure 9 B). Baseoendopods confluent, each with one outer bare basal seta. Exopod ovate, discrete, with inner row of spinules, and with two spinulose outer setae, one pinnate apical element, and one pinnate inner seta.	en	Back, Jinwook, Lee, Wonchoel, Huys, Rony (2011): A new species of Remanea Klie, 1929 (Copepoda: Harpacticoida: Paramesochridae) with a redescription of the type species. Journal of Natural History 45 (47 - 48): 2939-2964, DOI: 10.1080/00222933.2011.622057, URL: http://dx.doi.org/10.1080/00222933.2011.622057
