taxonID	type	description	language	source
03EA87FCFFC2FFE8FF19FE54034EFE30.taxon	description	(Figs 3, 25, 26, 31, 32 – 40, 63, 64, 68) lsid: zoobank. org: act: 13111 F 27 - B 26 F- 472 E-B 3 EC-A 23954 B 7 A 6 E 1	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC2FFE8FF19FE54034EFE30.taxon	materials_examined	Type material. Holotype: ♁, ECUADOR: SW of Quito, western slopes of the equatorial Andes, Via Aloag to Sto. Domingo, 0 ° 26 ’ 46 ” S, 78 ° 37 ’ 39 ” W, elevation ca. 3100 m, mining larva on Minthostachys mollis (Kunth) Griseb. (Lamiaceae), 24. ii. 2000, ex pupa iii. 2000, leg. R. Puplesis, genitalia slide no. AD 697 (USNM). Paratypes: 2 ♁, 2 ♀, same label data as holotype, genitalia slides nos AD 688 ♁, AD 1033 ♁, AD 1026 ♀ (USNM).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC2FFE8FF19FE54034EFE30.taxon	diagnosis	Diagnosis. The new species belongs to the Stigmella singularia species group, designated and characterized by Stonis et al. (2017 a). Externally, males of S. mentholica sp. nov. differ from the most similar S. aromatica sp. nov. in the slender, subapical fascia of the forewing and indistinctive patch of silvery or golden shiny scales on the apex. In the male genitalia, S. mentholica differs from all other known Neotropical Stigmella, including other species of the S. singularia group, by the combination of a distinctly truncate uncus and very long caudal processes of the gnathos (Fig. 34). Only two other species, S. aromatica and S. odora sp. nov, feed on the same host plant Minthostachys mollis, but there are significant differences in the morphology of the leaf mine: in S. mentholica, the leaf mine is a wide, relatively short gallery (Fig. 3), while the leaf mine of S. aromatica is a slender and long gallery, almost fully filled with frass (Fig. 10), and the leaf mine of S. odora is a combination of a very slender gallery and an irregular blotch (Fig. 6). Male (Figs 25, 26, 31). Forewing length 2.6 – 3.0 mm; wingspan 5.8 – 6.7 mm (n = 3). Head: palpi cream; frontal tuft dark orange; scape golden cream; collar golden cream but at certain angle may look pale grey, metallic glossy; antenna distinctly longer than half the length of forewing; flagellum with 36 segments, dark grey to brown-grey or pale grey, with golden gloss and some purple iridescence. Thorax and tegula golden brown. Forewing pale golden brown, strongly shining, at base with intense purple iridescence; fascia subapical, slender, comprised of silvery or golden shiny scales; apex of forewing brown-purple, with indistinctive patch of silvery or golden shiny scales; fringe grey; underside of forewing black-brown, without spots or androconia, except for slender scaleless spot at base. Hindwing pale grey to dark brown depending from angle of view, without androconia; fringe grey. Legs glossy grey cream; on upper side, covered with grey to brown-black scales with purple iridescence. Abdomen blackish grey to grey-brown on upper side and underside; genital segments pale brown; anal tufts short but distinctive, dark grey to fuscous. Female. Smaller than male; forewing length 2.3 – 2.4 mm; wingspan 5.2 – 5.4 mm (n = 2). Palpi and scape snow white. Frontal tuft pale yellow. Fascia of forewing post median-subapical, glossy white, distinctive. Abdomen fuscous to grey-brown on upper side, ochreous grey to grey-brown on underside; genital segments cream-ochre on underside, ovipositor pointed. Otherwise as in male. Male genitalia (Figs 32 – 40). Capsule longer (215 – 220 μm) than wide (195 – 200 μm). Vinculum with short, pointed lateral lobes. Uncus truncate caudally (Figs 33, 34). Gnathos with two very long, close-set caudal processes and a slender plate (Fig. 34). Valva (Fig. 38) 210 – 215 μm long, 75 – 80 μm wide, with slender apical process; transtilla with sublateral processes (Figs 35, 36). Juxta membranous, indistinctive. Phallus (Figs 37, 39, 40) 270 – 335 μm long, 85 – 90 μm wide; vesica with about 15 – 16 large spine-like cornuti and some minuscular cornuti (Fig. 37). Female genitalia (Figs 63, 64). Total length 1070 – 1080 μm. Anterior apophyses gradually narrowing distally; posterior apophyses very slender, longer than anterior ones, distally bent (Fig. 63). Vestibulum without sclerites. Corpus bursae with a strongly folded distal part and oval-shaped basal part with many indistinctive pectinations. Accessory sac very small, indistinctive; ductus spermathecae without coils, but with a small, tube-like vesicle. Bionomics (Figs 1 – 3). Host plant is Minthostachys mollis (Benth.) Griseb., Lamiaceae: Mentheae (Figs 1, 2). Larva pale green with brown-green intestine and pale brown head; feeds in February and probably in late January (note that in late February some leaf mines are already vacant). Prefer open, sunny areas. Leaf mine (Fig. 3) is a wide, sinuous, gradually widening gallery (easy to spot because the generally whitish appearance); at the beginning, with an interrupted line of brown-black frass, further on, with a wide margin, unfilled with the frass. Cocoon brown. Adults probably fly in March (indoors, emerged in March).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC2FFE8FF19FE54034EFE30.taxon	distribution	Distribution (Fig. 68). This species is known from a single locality in Ecuador, on the western slopes of the equatorial Andes, at elevation of ca. 3100 m, but the host plant has a much wider distribution in the northern and central Andes (see Discussion).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC2FFE8FF19FE54034EFE30.taxon	etymology	Etymology. The species name is derived from menthol, one of the principal components of essential oil of its host plant, Minthostachys mollis, in reference to the strong mint smell of its leaf mines.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC3FFE6FF19FDC8046EF899.taxon	description	(Figs 9 – 11, 22 – 24, 28 – 30, 41 – 47, 65, 66, 68) lsid: zoobank. org: act: 42 F 003 AD-AAE 4 - 4 A 52 - AD 3 B-E 2 DE 745 B 8 E 19	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC3FFE6FF19FDC8046EF899.taxon	materials_examined	Type material. Holotype: ♁, ECUADOR: SW of Quito, western slopes of the equatorial Andes, Via Aloag to Sto. Domingo, 0 ° 26 ’ 46 ” S, 78 ° 37 ’ 39 ” W, elevation ca. 3100 m, mining larva on Minthostachys mollis (Benth.) Griseb. (Lamiaceae), 24. ii. 2000, ex pupa iii. 2000, leg. R. Puplesis, genitalia slide no. AD 1022 (USNM). Paratypes: 9 ♁, 6 ♀, same label data as holotype, genitalia slides nos AD 815 ♁, AD 819 ♁ (from adult in pupal skin, pinned adult unavailable), AD 1021 ♁, AD 1032 ♁, AD 1027 ♀, AD 1031 ♀ (USNM).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC3FFE6FF19FDC8046EF899.taxon	diagnosis	Diagnosis. The new species belongs to the Stigmella singularia species group, designated and characterized by Stonis et al. (2017 a). Externally, males of S. aromatica sp. nov. differ from the most similar S. mentholica sp. nov. in the wide, median fascia of the forewing and distinctive apical fascia. In the male genitalia, S. aromatica differs from all other known Neotropical Stigmella, including species of the S. singularia group, by the combination of an uncus with two unique caudal lobes (Fig. 41), a large gnathos with two very close-set caudal processes (Fig. 43), a bifid juxta (Fig. 44), and a set of large, spine like cornuti (Figs 45 – 47). The host plant, Minthostachys mollis, also make this species distinctive among other Stigmella species, except for S. mentholica and S. odora sp. nov. However, S. aromatica differs from both species in the morphology of the leaf mine: the leaf mine of S. aromatica is a long, slender, sinuous gallery almost entirely filled with frass (Fig. 10), while the leaf mine of S. mentholica is a relatively short and wide gallery (Fig. 3), and the leaf mine of S. odora is combined of a very slender gallery and irregular blotch (Fig. 6). Male (Figs 22 – 24, 30). Forewing length 2.0 – 2.5 mm; wingspan 4.5 – 5.6 mm (n = 5). Head: palpi golden cream; frontal tuft large, dark orange; scape golden cream; collar golden cream but at certain angle may look metallic grey; antenna about 2 / 3 of the length of forewing; flagellum with 33 – 37 segments, brown-grey, golden glossy. Thorax and tegula concolorous with the forewing base. Forewing pale golden brown, strongly shining, with some purple iridescence along costa; postmedian fascia wide, comprised of silvery or golden shiny scales; apex of forewing brown with strong purple iridescence, and with a distinctive apical fascia of silvery or golden shiny scales (occasionally apical fascia is indistinctive, see Figs 28, 29); fringe grey, glossy, distally whitish; underside of forewing dark greybrown, without spots or androconia, except for a slender, scaleless spot at base. Hindwing grey to dark grey, without androconia; fringe grey. Legs glossy cream to grey; on upper side, covered with dark grey-brown scales. Abdomen blackish grey with purple iridescence on upper side, brown-grey on underside; genital segments pale brown; anal tufts short but distinctive, grey. Female. Smaller than male; forewing length 2.1 – 2.3 mm; wingspan 4.7 – 5.2 mm (n = 5). Antenna shorter than in male, only slightly longer than half the length of forewing. Abdomen grey-brown on upper side and underside; genital segments pale brown; anal tufts short but distinctive, blackish grey to grey; ovipositor pointed. Otherwise as in male. Male genitalia (Figs 41 – 47). Capsule longer (290 – 300 μm) than wide (175 – 210 μm). Vinculum with short, pointed lateral lobes. Uncus with unique caudal lobes (Fig. 41). Gnathos large, with two very close-set caudal processes (Fig. 43). Valva (Fig. 44) 200 μm long, with pointed, partially divided apical process (Fig. 44); transtilla without sublateral processes (Fig. 42). Juxta triangular, distally split (Fig. 44). Phallus (Figs 45 – 47) 210 – 330 μm long, 90 – 115 μm wide; vesica with about numerous large spine-like cornuti and a lateral set of small cornuti (Fig. 45). Female genitalia (Figs 65, 66). Total length 995 – 1000 μm. Anterior apophyses gradually narrowing and bent distally; posterior apophyses slender, approximately as long as anterior ones (Fig. 66). Vestibulum without sclerites. Corpus bursae with a strongly folded distal part and round or oval-shaped basal part with many distinctive pectinations. Accessory sac small; ductus spermathecae without coils, but with a small, tube-like vesicle. Bionomics (Figs 7 – 11). Host plant is Minthostachys mollis (Kunth) Griseb., Lamiaceae: Mentheae (Figs 7, 8). Larva yellow with indistinctive, yellowish brown intestine and pale brown head; feeds in February and probably in late January (note that in late February most of the leaf mines are already vacant). Prefer to occur in shady places. Leaf mine (Figs 9 – 11) is a long, slender, sinuous gallery almost entirely filled with black frass (Fig. 10); in old, vacant leaf mines, frass may turn brown or black (Fig. 9). Cocoon brown. Adults probably fly in March (indoors, emerged in March).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC3FFE6FF19FDC8046EF899.taxon	distribution	Distribution (Fig. 68). This species is known from a single locality in Ecuador, on the western slopes of the equatorial Andes, at elevation of ca. 3100 m, but the host plant has a much wider distribution in the northern and central Andes (see Discussion).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFC3FFE6FF19FDC8046EF899.taxon	etymology	Etymology. The species name is derived from Latin aromaticus (aromatic), due to the minty aroma of essential oil of the host plant, Minthostachys mollis, and its leaf mines.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	description	(Figs 4 – 6, 48 – 53, 68) lsid: zoobank. org: act: 5 BEE 514 E-FA 26 - 4 B 23 - A 414 - BC 1 F 8 FDB 12 F	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	materials_examined	Type material. Holotype: ♁, ECUADOR: SE of Ambato, Baños de Agua Santa, on the slopes of the Tungurahua volcano, 1 ° 24 ꞌ 13 ꞌꞌS, 78 ° 25 ꞌ 43 ꞌꞌW, elevation ca. 2080 m, mining larva on Minthostachys mollis (Benth.) Griseb. (Lamiaceae), 10. ii. 2000, leg. R. Puplesis, genitalia slide no. AD 821 ♁ (pinned adult is lost) (USNM). Paratypes: 2 ♁, same label data as holotype, genitalia slides nos AD 685 ♁ (pinned adult is lost), AD 806 ♁ (from adult in pupal skin, pinned adult unavailable) (USNM).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	diagnosis	Diagnosis. The new species belongs to the Stigmella singularia species group, designated and characterized by Stonis et al. (2017 a). In the male genitalia, S. odora sp. nov. differs from the similar S. mentholica sp. nov. and S. aromatica sp. nov. by the unique uncus (see Fig. 48), very slender plate of the gnathos, presence of numerous small cornuti (Fig. 50), and the wide inner lobe and large apical process of the valva (Figs 51, 52). The host plant, Minthostachys mollis, makes this species distinctive among Stigmella species, except for S. mentholica and S. aromatica. However, S. odora differs from both species in the morphology of the leaf mine: the leaf mine of S. odora is a combination of a very slender gallery and an irregular blotch (Fig. 6), while the leaf mine of S. aromatica is a slender gallery almost entirely filled with frass (Fig. 10), and the leaf mine of S. mentholica is a relatively short and wide gallery (Fig. 3). Male. External characters are unknown (see Remarks). Female. Unknown. Male genitalia (Figs 48 – 53). Capsule longer (260 – 280 μm) than wide (175 – 180 μm). Vinculum with short or moderately long, triangular lateral lobes. Uncus gradually narrowing towards apex, with characteristic caudal lobes (Fig. 48). Median plant of gnathos very slender, with two slender caudal processes (Fig. 48). Valva (Figs 51, 52) 170 – 190 μm long, with a large apical process and wide inner lobe (Fig. 52); transtilla with short sublateral processes (Fig. 49). Juxta membranous, indistinctive. Phallus (Figs 50, 53) 275 – 280 μm long, 80 – 100 μm wide; vesica with a set of numerous large spine-like cornuti and many small, triangular cornuti (Fig. 53). Bionomics (Figs 1, 2, 4 – 6). Host plant is Minthostachys mollis (Benth.) Griseb., Lamiaceae: Mentheae (Figs 1, 2). Larva pale, grey yellowish green with indistinctive, ochre-brown intestine and dark brown head; feeds in February. Leaf mine (Figs 4 – 6) is combined of a long, very slender gallery and a large, irregular blotch; the slender gallery is heavily contorted in the initial part and entirely or almost entirely filled with black-brown frass (Fig. 6); the blotch-like part of the leaf mine is with medially scattered black-brown frass (Figs 4, 5). Adults fly in February and March.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	distribution	Distribution (Fig. 68). This species is known from a single locality in equatorial Andes, on the slopes of the Tungurahua volcano (Ecuador: Baños) at the elevation of ca. 2100 m, but the host plant has a much wider distribution in the northern and central Andes (see Discussion).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	etymology	Etymology. The species name is derived from Latin odorus (odorous, scented), in reference to the mintyscented host plant and leaf mines of Stigmella odora.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFE4FF19FF19042BF9B0.taxon	discussion	Remarks. The new species is described based on the genitalia of three male specimens and the leaf mines only. The pinned adult of the holotype was lost, and the two paratypes are from adults in pupal skins, and therefore pinned specimens are unavailable. Although external characters of the species remain unknown, the new species possesses unique male genitalia and a particularly unique morphology of the leaf mine. We describe this species for the record and hope that the adults will be discovered in the future.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	description	(Figs 12 – 17, 27, 54 – 59, 69) lsid: zoobank. org: act: D 19 FFC 7 B- 9 E 35 - 4997 - 88 CA- 65 AD 65907308	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	materials_examined	Type material. Holotype: ♁, ECUADOR: SE of Ambato, Baños de Agua Santa, on the slopes of the Tungurahua volcano, 1 ° 24 ꞌ 5 ꞌꞌS, 78 ° 25 ꞌ 45 ꞌꞌW, elevation 1980 m, mining larva on Clinopodium tomentosum (Kunth) Govaerts (Lamiaceae), 9. ii. 2000, leg. R. Puplesis, genitalia slide no. AD 796 (adult documented but lost) (USNM).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	diagnosis	Diagnosis. The new species belongs to the Stigmella singularia species group, designated and characterized by Stonis et al. (2017 a). In the male genitalia, S. tomentosella sp. nov. differs from the similar S. odora sp. nov. and other species of the group by the unique, long and very slender cornutus, as well as the combination of a slender valva, large median element of the gnathos, and a transtilla with short sublateral processes. The host plant Clinopodium tomentosum makes this species distinctive among Stigmella species, except for the Bolivian S. clinopodiella Diškus & Stonis, 2016 that feeds on the related host plant, Clinopodium axillare (Rusby) Harley (Fig. 18) (See Remarks below). However, S. tomentosella differs in the morphology of the leaf mine: the leaf mine of the new species is slender, almost entirely filled with brown-black frass (Fig. 15); the leaf mine of S. clinopodiella is a combination of a long, very slender gallery and an elongated blotch with wide margin unfilled with frass (Figs 19 – 21). Male (Fig. 27). Forewing length 2.1 mm; wingspan 4.6 mm (n = 1). Head: palpi brownish cream; frontal tuft dark orange; scape and collar brownish cream; antenna (flagellum) brownish grey. Thorax and tegula concolorous with the forewing base. Forewing densely speckled with brown and dark brown scales; fascia ill-defined, subapical, comprised of brownish white scales; fringe pale brown; underside of forewing brown, without spots or androconia. Hindwing brownish grey, without androconia; fringe grey. Legs brownish cream; on upper side, covered with blackgrey scales. Abdomen blackish grey on upper side, brownish cream on underside; genital segments cream; anal tufts distinctive, cream, half the width of terminal segment of the abdomen. Female. Unknown. Male genitalia (Figs 54 – 59). Capsule longer (290 μm) than wide (170 μm). Vinculum with distinctive, thickened, slightly angular lateral lobes (Fig. 57). Uncus gradually narrowing towards apex, with characteristic caudal lobes (Fig. 55). Gnathos with a moderately large median plate and two slender caudal processes (Fig. 54). Valva (Fig. 56) 180 μm long, slender, gradually narrowing to an apical process (Fig. 56); transtilla with short sublateral processes (Fig. 57). Juxta membranous, indistinctive. Phallus (Figs 58, 59) about 295 μm long, 55 μm wide in the middle and 90 μm wide at apical part; vesica with a set of about ten large spine-like cornuti; one of them is very slender and long (Fig. 59). Bionomics (Figs 12 – 17, 69). Host plant is Clinopodium tomentosum (Kunth) Govaerts, Lamiaceae: Mentheae (Figs 12, 13). Larva pale, brownish yellow with an indistinctive intestine and pale brown head; feeds in February and in January (note that in early February most of leaf mines are already vacant). Leaf mine (Figs 14 – 17) is a gradually widening gallery, almost entirely filled with black-brown frass (Fig. 15). Cocoon beige to ochre-beige, 1.9 – 2.2 mm long, 1.1 – 1.4 mm wide (n = 4). Adults fly in February and probably in early March.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	distribution	Distribution (Fig. 69). This species is known from a single locality in equatorial Andes, on the slopes of the Tungurahua volcano (Ecuador: Baños) at elevation of ca. 2000 m, but the host plant has a wider distribution in the equatorial Andes (see Discussion).	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	etymology	Etymology. The species is named after the host plant, Clinopodium tomentosum.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
03EA87FCFFCFFFFBFF19F9490636F9CC.taxon	discussion	Remarks. Stigmella clinopodiella Diškus & Stonis, discovered feeding on a congeneric host plant to this species, was described from Bolivia (Fig. 70), and illustrated by Stonis et al. 2016: 79 – 82. The species also belongs to the Stigmella singularia species group. In the male genitalia (Figs 60 – 62), S. clinopodiella differs from all other species of the group in the combination of a large gnathos with two very close-set caudal processes, truncate uncus, large apical process of the valva, triangular juxta and long cornuti of the phallus. In the female genitalia, S. clinopodiella is characterized by the angular anterior apophyses (Fig. 67) and the rounded basal part of the bursa copulatrix.	en	Stonis, Jonas R., Diškus, Arūnas, Fernández-Alonso, José L., Remeikis, Andrius, Solis, M. Alma (2021): New leaf-miningNepticulidae (Lepidoptera): potential pests of aromatic Lamiaceae plants from equatorial Andes. Zootaxa 4926 (3): 363-383, DOI: 10.11646/zootaxa.4926.3.3
