identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EA8A53492AFFD9AFD0FEDEFAF8C57D.text	03EA8A53492AFFD9AFD0FEDEFAF8C57D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica Peringuey 1904	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> KEY TO SPECIES OF THE  HYBOSERICA COMPLEX (♂♂) </p>
            <p> 1 Epipleural carina reduced after anterior third of elytra. Base of pronotum with fine marginal line.................................................................. 2 (  Leoserica gen. nov. ) </p>
            <p> 1’ Epipleural carina complete. Base of pronotum without marginal line............... 5 (  Hyboserica ) </p>
            <p> 2 Protibia bidentate. Metacoxae densely setose. Mesotarsomere 1 very long, more than 1.5 times longer than the subsequent tarsomere............................................  Leoserica jamesi</p>
            <p>2’ Protibia tridentate. Metacoxae glabrous. Mesotarsomere 1 not much longer than the subsequent tarsomere............................................................................3</p>
            <p> 3 Hind wings reduced (Fig. 7C). Marginal line of anterior margin of pronotum narrowly interrupted medially..............................................................  Leoserica capensis</p>
            <p>3’ Hind wings complete (Fig. 8I). Marginal line of anterior margin of pronotum complete............ 4</p>
            <p> 4 Parameres basally convexly widened (lateral view; Fig. 7A)...................  Leoserica knysnana</p>
            <p> 4’ Parameres nearly flat (lateral view), basally not widened (Fig. 7B)...............  Leoserica hotovyi</p>
            <p> 5 Aedeagus, including parameres, strongly asymmetrical (Fig. 6F).............  Hyboserica muellerae</p>
            <p>5’ Aedeagus symmetric or only slightly asymmetrical (e.g. Fig. 6C–E)............................ 6</p>
            <p>6 Meso- and metatarsomeres punctate..................................................... 7</p>
            <p>6’ Meso- and metatarsomeres not punctate................................................. 15</p>
            <p>7 Antennal club longer than the remaining antennomeres combined............................ 8</p>
            <p>7’ Antennal club not longer than the remaining antennomeres combined......................... 9</p>
            <p>8 Parameres slightly asymmetrical (e.g. Fig. 4H). Anterior margin of labroclypeus weakly sinuate... 10</p>
            <p>8’ Parameres symmetrical (e.g. Fig. 5A). Anterior margin of labroclypeus straight................. 11</p>
            <p>9 Antennal club shorter than the remaining antennomeres combined.......................... 13</p>
            <p>9’ Antennal club as long as the remaining antennomeres combined............................. 14</p>
            <p> 10 Parameres slender. Distal teeth of parameres small (Fig. 5B)...............  Hyboserica uitsoekensis</p>
            <p> 10’ Parameres more robust. Distal teeth of parameres large (Fig. 4H)..........  Hyboserica asymmetrica</p>
            <p> 11 Distal dorsomedian lamella of phallobase incised medially at apex. Basal lobe of parameres displaced beyond the middle. Sclerotization of median lamella of phallobase continuous and in one plain with the rest of phallobase (Fig. 5A ..............................................  Hyboserica madiba</p>
            <p>11’ Sclerotization of median lamella of phallobase not continuous to the rest of phallobase, widely membranous between lamella and principal phallobase and concavely excavated mesopreapically (e.g. Fig. 4F)............................................................................ 12</p>
            <p> 12 Distal dorsomedian lamina of phallobase wide and semi-circular and weakly curved (lateral view) (Fig. 4F)........................................................  Hyboserica soutpansbergensis</p>
            <p> 12’ Distal dorsomedian lamina of phallobase narrower and ligula-like, and strongly curved (lateral view) (Fig. 4G)........................................................  Hyboserica entabeniensis</p>
            <p> 13 Parameres in lateral view oval (Fig. 6B).................................  Hyboserica shulmani</p>
            <p> 13’ Parameres in lateral view elongate (Fig. 5C)........................  Hyboserica shilouvaneensis</p>
            <p> 14 Distal dorsomedian lamina of phallobase wide and semi-circular. Distal teeth of parameres narrow, sharply pointed and directed distally. Basal lobes of parameres of moderate size and sharply pointed (Fig. 4E)........................................................  Hyboserica nelspruitensis</p>
            <p> 14’ Distal dorsomedian lamina of phallobase narrow, longitudinally concave and ligula-like. Distal teeth of parameres very small, nearly invisible. Basal lobes of parameres very small (Fig. 5G)................................................................  Hyboserica schoenhoferi</p>
            <p> 15 Shape of parameres (in lateral view) transverse (Fig. 6E)......................  Hyboserica uhligi</p>
            <p>15’ Parameres (in lateral view) at least as long as wide (e.g. Fig. 6C)............................. 16</p>
            <p> 16 Shape of parameres (in lateral view) spherical (Fig. 6C)......................  Hyboserica haladai</p>
            <p>16’ Shape of parameres (in lateral view) elongate (e.g. Fig. 6D)................................. 17</p>
            <p> 17 Antennal club longer than the remaining antennomeres combined...............  Hyboserica kochi</p>
            <p>17’ Antennal club not longer than the remaining antennomeres combined.........................18</p>
            <p>18 Antennal club as long as the remaining antennomeres combined..............................19</p>
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	https://treatment.plazi.org/id/03EA8A53492AFFD9AFD0FEDEFAF8C57D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534926FFD7AF29F931FBEBC745.text	03EA8A534926FFD7AF29F931FBEBC745.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica Peringuey 1904	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA PÉRINGUEY, 1904</p>
            <p> Hyboserica Péringuey, 1904: 27 . Type species (designated by Ahrens, 2007):  Hyboserica caffra (Fåhreus, 1857) ; Dalla Torre (1912: 65); Machatschke (1959: 744). </p>
            <p>Diagnosis: Body of medium size (7.5–11.0 mm), elongately oval, reddish to yellowish or dark brown, dorsal surface with iridescent shine; dorsal surface nearly glabrous. With a few exceptions, the labroclypeus wide and trapezoidal, with fine and dense punctures; anterior margin of labroclypeus weakly sinuate, rarely distinctly sinuate or straight.Mentum anteriorly slightly elevated. Antenna composed of nine antennomeres, club short, in both sexes with three antennomeres. Pronotum subtrapezoidal and moderately wide, lateral margins sparsely setose, anterior angles well-produced and moderately sharp, anterior margin convexly produced medially; basal margin without marginal line. Hypomeron simple, not carinate. Elytra without reduced epipleural margin, apical margin chitinous, without membranous rim of microtrichomes, intervals flat, evenly finely punctate, glabrous, odd intervals with punctures concentrated along the weakly impressed striae, apical setae on sutural interval absent. Ventral surface sparsely covered with short setae. Metatibia dorsally with two groups of spines and longitudinally convex; at apex not truncate or sinuate near tarsal articulation. Protibia tridentate, basal tooth small.</p>
            <p> Remarks: Two species were originally included in the genus,  Hyboserica caffra (Fåhreus, 1857) and  H. globuliformis (Brenske, 1901) . Ahrens (2007) designated the former as the type species. The latter is an Indian species, which was never found in South Africa (Ahrens, 2007) and which was either temporarily introduced from India, or, more likely, erroneously labelled. However, apart from the only so far examined female type specimen (see Ahrens, 2007) we recently came across another male specimen that obviously belonged to the same type series, labelled ‘Cap b. spec. Coll. Felsche/ No. 261/ der carbona ähnlig [sic]/ Coll. Brenske’ (ZMHB); its aedeagus was virtually identical in shape with that of  Maladera fumosa (Brenske, 1898) . Thus,  Serica globuliformis Brenske, 1901 has to be considered a junior synonym of  Maladera fumosa (Brenske, 1898) , a species extensively revised in Ahrens (2004) and Ahrens &amp; Fabrizi (2016). </p>
            <p> The Species 2000 &amp; ITIS catalogue of life (Schoolmeesters, 2017) and in consequence other online databases (e.g. Wikispecies, Encyclopedia of Life, Global Biodiversity Information Facility, Insectoid.info, Bionames.org, Global Species) list an additional nine  Hyboserica taxa, which is because of a typing error (‘  Hyboserica ’ instead of correct ‘  Hyposerica ’):  Hyposerica brunneipennis Moser, 1915 ,  H. brunneipennis Moser, 1918 ,  H. brunneipes Moser, 1915 ,  H. fenerivensis (sic, not feneriventris) Moser, 1915,  H. luridipennis Moser, 1915 ,  H. piceonigra Moser, 1915 ,  H. sericeomicans Moser, 1915 ,  H. silvicola Moser, 1915 and  H. truncatipennis Moser, 1915 . These Madagascan species, described in the genus  Hyposerica Brenske, 1897 , were never associated with the southern African  Hyboserica Péringuey, 1904 . </p>
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	https://treatment.plazi.org/id/03EA8A534926FFD7AF29F931FBEBC745	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534924FFD0AD42FAAEFB9EC53E.text	03EA8A534924FFD0AD42FAAEFB9EC53E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica alexandriana Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA ALEXANDRIANA SP. NOV.</p>
            <p>(FIGS 3H, 8, 9)</p>
            <p>urn:lsid:zoobank.org:act: 92C0E3ED-A27A-4B43-A474- 64AC67C3B246</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., SE. Cape Prov.  
                <a title="Search Plazi for locations around (long 26.23/lat -33.43)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.23&amp;materialsCitation.latitude=-33.43">Alexandria For. St.</a>
                 , 33.43S, 26.23E, 4.12.1987, E-Y:2552 UV light collection, leg. Endrödy-Younga’ (TMSA)  .  Paratypes: 15 ♂♂, 14 ♀♀ same data (TMSA, ZFMK) ,   1 ♂ ‘ S.Afr.,  
                <a title="Search Plazi for locations around (long 29.44/lat -31.27)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.44&amp;materialsCitation.latitude=-31.27">Transkei Ntsubane forest</a>
                 , 31.27 S, 29.44 E, 26.11.1988, E-Y: 2582 forest floor litter, leg. Endrödy-Younga’ (TMSA)  . 
            </p>
            <p>Diagnostic description: Body length: 8.0 mm, elytral length: 5.8 mm, maximum width: 4.6 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.42. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.37. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Metatibia, ratio of width/length: 1/3.89. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3H.</p>
            <p>Etymology: The new species is named after the type locality, Alexandria Forest.</p>
            <p>Variation: Body length: 7.9–9.1 mm, elytral length: 5.8–6.5 mm, maximum width: 4.6–5.1. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A534924FFD0AD42FAAEFB9EC53E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534922FFD1AED3FF14FDFAC402.text	03EA8A534922FFD1AED3FF14FDFAC402.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica asymmetrica Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA ASYMMETRICA SP. NOV.</p>
            <p>(FIGS 4H, 9)</p>
            <p>urn:lsid:zoobank.org:act: D004A6F6-C8A6-40E5- A08E-F5CCC3F02DEA</p>
            <p>
                 Type material examined:  Holotype: ♂ ‘ S. Afr., Tvl. Nelshoogte Knuckles rocks for, 25.47S, 30.50E, 8.12.1986, E-Y: 2357 UV light collection, leg. Endrödy-Younga’ (TMSA) .   Paratypes: 1 ♂ ‘ S. Afr.,  
                <a title="Search Plazi for locations around (long 30.5/lat -25.5)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.5&amp;materialsCitation.latitude=-25.5">Tvl. Nelshoogte Forest station</a>
                 , 25.50S, 30.50E,/ 2.12.1986, E-Y: 2346 UV  light collection, leg. Endrödy-Younga’ (ZFMK) . 
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            <p>Diagnostic description: Body length: 9.6 mm, elytral length: 6.1 mm, maximum width: 5.1 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.59. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.36. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.86. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4H. Female unknown.</p>
            <p>Etymology: The new species is named in reference to its asymmetric dorsomedian lobe of the male phallobase.</p>
            <p>Variation: Body length: 9.0– 9.6 mm, elytral length: 6.1–6.2 mm, maximum width: 4.6–5.1 mm.</p>
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	https://treatment.plazi.org/id/03EA8A534922FFD1AED3FF14FDFAC402	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534922FFD1AEC0F913FBA7C68C.text	03EA8A534922FFD1AEC0F913FBA7C68C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica bednariki Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA BEDNARIKI SP. NOV.</p>
            <p>(FIGS 6A, 9)</p>
            <p>urn:lsid:zoobank.org:act: 973005D1-ADE6-4647-833C- A162BF815309</p>
            <p> Type material examined:   Holotype: ♂ ‘ South Africa, Kwazulu-Natal, 20 km S of  Richmond , 500 m, in Mkomazi riv. valley, 4.–6.i.2007, M. Bednařik leg./, Coll. P. Pacholatko Brno’ (CPPB). </p>
            <p>Diagnostic description: Body length: 9.58 mm, elytral length: 6.81 mm, maximum width: 5.77 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.4. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly.Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.54. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.81. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 6A. Female unknown.</p>
            <p>Etymology: The new species is named after its collector, M. Bednařik.</p>
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	https://treatment.plazi.org/id/03EA8A534922FFD1AEC0F913FBA7C68C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534922FFCCADBEFAFDFE6BC32A.text	03EA8A534922FFCCADBEFAFDFE6BC32A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica caffra (Fahreus 1857)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA CAFFRA (FÅHREUS, 1857)</p>
            <p>(FIGS 3A, 9)</p>
            <p> Triodonta caffra Fåhreus, 1857: 139 ; Brenske, 1900: 213. </p>
            <p> Hyboserica caffra : Péringuey, 1904: 27, 28. </p>
            <p> Type material examined:   Lectotype (here designated): ♀ ‘ Caffraria / J. Wahlb. / Type / 3935 E91+/ Typus / ♀ /  caffra Fåhrs. /  Hyboserica Pér. ’ (NHRS). </p>
            <p> Additional material examined: 1 ♂ ‘Natal, D’urban/  Calloserica caffra Fhs. /  Triodonta caffra Fhs. comp. cum type / SAM-Col A035/35’ (SAMC), 1 ♂, 1 ♀ ‘Natal/ 20 Dec./ Trägardh/  Hyboserica /  caffra Fahr ?’ (ZMHB). </p>
            <p>Diagnostic description: Body length: 9.23 mm, elytral length: 6.12 mm, maximum width: 5.31 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.5. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.6. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Metatibia, ratio of width/length: 1/3.95. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3A.</p>
            <p> Remarks: One additional specimen under this name in the SAMC, Cape Town. The only available syntype of the species is a female making the species difficult to interpret. The type specimen is most likely (also from viewpoint of nearly identical external morphology) corresponding to the specimens collected at Port Natal (where J. Wahlberg quite likely collected his material) and which were assigned already by Péringuey and Brenske to  H. caffra . Therefore, we retain their interpretation of the species. </p>
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	https://treatment.plazi.org/id/03EA8A534922FFCCADBEFAFDFE6BC32A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493FFFCCAF32FE42FDDBC4EF.text	03EA8A53493FFFCCAF32FE42FDDBC4EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica davisorum Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA DAVISORUM SP. NOV.</p>
            <p>(FIGS 3G, 9)</p>
            <p>urn:lsid:zoobank.org:act: F227B870-DD3A-43F6-8C20- EA5837EEDB56</p>
            <p> Type material examined:   Holotype: ♂ ‘ South Africa, Cape Prov., Thomas Baines Nature res., 13 km SW Grahamstown,  Palmiet R [iver], c. 350 m, 6 Mar' [19]78, D. &amp; M. Davis, B. Akersberge’ (USNM). </p>
            <p>Diagnostic description: Body length: 9.12 mm, elytral length: 6.81 mm, maximum width: 5.08 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.45. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.62. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.68. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3G. Female unknown.</p>
            <p>Etymology: The new species is dedicated to D. &amp; M. Davis, collectors of the new species.</p>
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	https://treatment.plazi.org/id/03EA8A53493FFFCCAF32FE42FDDBC4EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493FFFCCAF30F900FAA0C79A.text	03EA8A53493FFFCCAF30F900FAA0C79A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica dukuduku Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA DUKUDUKU SP. NOV.</p>
            <p>(FIGS 5H, 9)</p>
            <p>urn:lsid:zoobank.org:act: 4BBAE215-1C7D-40A0-BC43- 3FB2D4985158</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘  Dukuduku Forest , 20–21.xi.1956, G. van Son’ (TMSA)  .  Paratypes: 1 ♂ ‘ SOUTH AFRICA, NATAL ,   
                <a title="Search Plazi for locations around (long 31.46/lat -28.58)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.46&amp;materialsCitation.latitude=-28.58">Mtunzini</a>
                 , 28.58S, 31.46E, 19.iv.1990, P. REAVELL/ NATIONAL COLL  .   OF INSECTS,  Pretoria , S. Afr. ’ (SANC)  ,   1 ♂ ‘ S. Afr.;,  
                <a title="Search Plazi for locations around (long 32.35/lat -28.22)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.35&amp;materialsCitation.latitude=-28.22">Zululand</a>
                 , St. Lucia, Mission Rock, 28.22 S, 32.35 E,/ 8.12.1975, E-Y: 960 at black light, leg. Endrödy-Younga’ (TMSA)  . 
            </p>
            <p>Diagnostic description: Body length: 9.0 mm, elytral length: 6.7 mm, maximum width: 5.4 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.44. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.52. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.42. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5H. Female unknown.</p>
            <p> Etymology: The new species is named after the type locality  Dukuduku Forest.</p>
            <p>Variation: Body length: 8.0– 9.6 mm, elytral length: 6.0– 6.7 mm, maximum width: 4.6–5.4 mm.</p>
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	https://treatment.plazi.org/id/03EA8A53493FFFCCAF30F900FAA0C79A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493FFFCDADB3F9F1FDFAC68C.text	03EA8A53493FFFCDADB3F9F1FDFAC68C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica entabeniensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA ENTABENIENSIS SP. NOV.</p>
            <p>(FIGS 4G, 9)</p>
            <p>urn:lsid:zoobank.org:act: 7E2046B0-CE80-4CCC-8017- 1FCD2813A29B</p>
            <p> Type material examined:  Holotype: ♂ ‘ Entabeni, Z [out]p[ans]b[er]g., Nov., 1931, G. van Son’ (TMSA) .  Paratypes: 1 ♂ ‘ South Africa, N. Prov., Lajuma Farm, Soutpansberg, 23.02S, 29.26E, 22–23.xi.1997 R . Stals,/ collecting technique: light trap’ (SANC). </p>
            <p>Diagnostic description: Body length: 8.54 mm, elytral length: 6.58 mm, maximum width: 4.96 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.53. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.41. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.59. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4G. Female unknown.</p>
            <p>Etymology: The new species is named after the type locality Entabeni.</p>
            <p>Variation: Body length: 8.5–8.6 mm, elytral length: 5.9–6.6 mm, maximum width: 4.6–5.0 mm.</p>
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	https://treatment.plazi.org/id/03EA8A53493FFFCDADB3F9F1FDFAC68C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493EFFCDAEF2FAFDFA92C6AD.text	03EA8A53493EFFCDAEF2FAFDFA92C6AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica grootkloofensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA GROOTKLOOFENSIS SP. NOV.</p>
            <p>(FIGS 3E, 9)</p>
            <p>urn:lsid:zoobank.org:act: 11D5A8AC-F76F-4A76-97C2- 5A44E18E5602</p>
            <p>
                 Type material examined:  Holotype: ♂ ‘ S. Afr., Tvl. Uitsoek Grootkloof ind[igenous] for[est], 25.15S, 30.33E,/ 25.10.1986, E-Y: 2318 ground trap, 50 days, leg. Endrödy-Younga /, ground trap with faeces bait’ (TMSA) .   Paratypes: 1 ♂ ‘S.  
                <a title="Search Plazi for locations around (long 30.33/lat -25.15)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.33&amp;materialsCitation.latitude=-25.15">Afr.</a>
                 , Tvl. Uitsoek Grootkloof, ind. for., 25.15S, 30.33E,/ 16.12.1986, E-Y: 2395 for litter, bef. rain, leg. Endrödy-Younga’ (ZFMK)  ,   1 ♂ ‘ S. Afr.,  
                <a title="Search Plazi for locations around (long 30.33/lat -25.16)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.33&amp;materialsCitation.latitude=-25.16">Tvl. Uitsoek Waterfall area</a>
                 , 25.16S, 30.33E,/ 13.12.1986, E-Y: 2385, UV  light collection, leg. Endrödy-Younga’ (TMSA),   1 ♂ ‘ S. Afr.,  
                <a title="Search Plazi for locations around (long 30.33/lat -25.15)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.33&amp;materialsCitation.latitude=-25.15">Tvl</a>
                 , Uitsoek Grootkloof, ind. for. 25.15S, 30.33E /, 25.10.1986;, E-Y: 2319 intercept trap, 50 d., leg. Endrödy-Younga’ (TMSA)  . 
            </p>
            <p>Diagnostic description: Body length: 10.3 mm, elytral length: 7.5 mm, maximum width: 5.8 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.47. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.4. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.7. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3E. Female unknown.</p>
            <p>Etymology: The new species is named after the type locality Grootkloof.</p>
            <p>Variation: Body length: 10.3–10.4 mm, elytral length: 7.4–7.5 mm, maximum width: 5.7–5.8 mm.</p>
        </div>
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	https://treatment.plazi.org/id/03EA8A53493EFFCDAEF2FAFDFA92C6AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493EFFCEAD9BFADEFEA9C13E.text	03EA8A53493EFFCEAD9BFADEFEA9C13E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica haladai Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA HALADAI SP. NOV.</p>
            <p>(FIGS 6C, 9)</p>
            <p>urn:lsid:zoobank.org:act: FD6CFE7A-1FBB-4F26- A6E4-8266AAD06F6D</p>
            <p> Type material examined:   Holotype: ♂ ‘Mozambik-S., 25 km N  Massinga , 36 m, S23°07’, E35°17’, J. Halada, 22.12.2005,/ Coll. P. Pacholatko Brno’ (CPPB). </p>
            <p>Diagnostic description: Body length: 9.23 mm, elytral length: 6.35 mm, maximum width: 4.96 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.47. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.67. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.48. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Metatarsomeres dorsally impunctate. Mesotarsomere missing. Aedeagus and habitus: Fig. 6C. Female unknown.</p>
            <p>Etymology: The new species is named for J. Halada, its collector.</p>
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	https://treatment.plazi.org/id/03EA8A53493EFFCEAD9BFADEFEA9C13E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493DFFCEAF13FCAEFAA0C0F6.text	03EA8A53493DFFCEAF13FCAEFAA0C0F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica kochi Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA KOCHI SP. NOV.</p>
            <p>(FIGS 6D, 9)</p>
            <p>urn:lsid:zoobank.org:act: D3993ACA-BCD9-4524-94DA-41650D4C0FA8</p>
            <p>
                 Type material examined:  Holotype: ♂ ‘ Südafrika, Transvaal Blyderiverspoort Nat. Res., 24°39’S, 30°50’E 13.vii.1995, F. Koch’ (ZMHB) .   Paratype: 1 ♂ ‘X-DA3708/ X-DA3708 South Africa, Mpumalanga, Blyderiverspoort,  
                <a title="Search Plazi for locations around (long 30.780445/lat -24.573112)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.780445&amp;materialsCitation.latitude=-24.573112">Forever Resort</a>
                 , 1194 m, at light, 24°34’23.2’’S, 30°46’49.6’’E, 29.I.2013, leg. Ahrens &amp; Fabrizi  Hyboserica sp 20’ (ZFMK)  . 
            </p>
            <p>Diagnostic description: Body length: 8.0 mm, elytral length: 5.4 mm, maximum width: 4.3 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.48. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.49. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.72. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 6D. Female unknown.</p>
            <p>Etymology: The new species is dedicated to F. Koch, one of the collectors of the new species.</p>
            <p>Variation: Body length: 8.0– 8.8 mm, elytral length: 5.4–6.6 mm, maximum width: 4.3–4.6 mm.</p>
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	https://treatment.plazi.org/id/03EA8A53493DFFCEAF13FCAEFAA0C0F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493DFFCFAD94FD18FD28C38E.text	03EA8A53493DFFCFAD94FD18FD28C38E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica kosiensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA KOSIENSIS SP. NOV.</p>
            <p>(FIGS 5F, 9)</p>
            <p>urn:lsid:zoobank.org:act: 90E67199-62DE-4B4C-98EB- FB9AB2494DA6</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., KwaZulu Natal,  
                <a title="Search Plazi for locations around (long 32.5/lat -26.58)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.5&amp;materialsCitation.latitude=-26.58">Kosi Bay</a>
                 camp, 26.58S, 32.50E,/ 15.11.2002, E-Y: 3540 light trap Burger, Harrison, Müller’ (TMSA)  .   Paratypes: 2 ♂, 1 ♀ ‘ RSA, KwaZulu-Natal,  Kosi Bay Nature Reserve , 11.–17.XI.2002, leg. W. Schawaller’ (SMNS, ZFMK)  ,   1 ♀ ‘X-JE076 South Africa, KwaZulu Natal, Lala Nek, iSmangaliso  World Heritage Site , 4.iv.2011, leg. A. Armstrong’ (TMSA)  . 
            </p>
            <p>Diagnostic description: Body length: 9.5 mm, elytral length: 6.4 mm, maximum width: 5.4 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Antennal club missing in holotype. Eyes, ratio diameter/interocular width: 0.44.</p>
            <p>Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted.Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.67. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.68. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5F.</p>
            <p>Etymology: The new species is named after the type locality Kosi Bay.</p>
            <p>Variation: Body length: 9.5–9.8 mm, elytral length: 6.4–7.0 mm, maximum width: 5.4–5.6 mm. Antennal club 0.7 times as long as remaining antennomeres combined. Without evident sexual dimorphism.</p>
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</html>
	https://treatment.plazi.org/id/03EA8A53493DFFCFAD94FD18FD28C38E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493CFFCFAF2DFDFEFE42C532.text	03EA8A53493CFFCFAF2DFDFEFE42C532.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica madiba Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA MADIBA SP. NOV.</p>
            <p>(FIGS 5A, 9)</p>
            <p>urn:lsid:zoobank.org:act: E9F1279A-C55B-44AA-85C5- 46F2840C240C</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘DA0403  Hyboserica sp , South Africa,  
                <a title="Search Plazi for locations around (long 29.952696/lat -23.895195)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.952696&amp;materialsCitation.latitude=-23.895195">Cheerio Farm</a>
                 , Haenertsburg, ca. 20 km W of Tzaneen, 1492 m, 023°53’42.7’’S, 029°57’09.7’’E, 21.XI.2006, leg. D. Ahrens &amp; S. Fabrizi’ (TMSA). 
            </p>
            <p>Diagnostic description: Body length: 8.08 mm, elytral length: 6 mm, maximum width: 4.27 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.58. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.33. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.89. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5A. Female unknown.</p>
            <p>Etymology: The new species is dedicated to President Nelson Mandela (Madiba).</p>
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	https://treatment.plazi.org/id/03EA8A53493CFFCFAF2DFDFEFE42C532	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493CFFCFAD52FF14FBE4C47F.text	03EA8A53493CFFCFAD52FF14FBE4C47F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica maphelaneensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> HYBOSERICA MAPHELANEENSIS SP. NOV.</p>
            <p>(FIGS 3D, 9)</p>
            <p>urn:lsid:zoobank.org:act: 04246D1A-3A48-4450-9AA5- E578D5187C61</p>
            <p> Type material examined:   Holotype ♂ ‘X-DA4402 South Africa, KZN,  Maphelane , 28.xi.2008, leg. R. Müller’ (TMSA)  .  Paratypes: 1 ♀ ‘X-DA4403 South Africa, KZN ,   Maphelane , 28.xi.2008, leg. R  .   Müller’ (ZFMK)  . </p>
            <p>Diagnostic description: Body length: 8.9 mm, elytral length: 8.1 mm, maximum width: 5.0 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct.Antennal club shorter than remaining antennomeres combined.Eyes, ratio diameter/interocular width: 0.43. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.62. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.65. Basal group of dorsal spines of metatibia at first-quarter of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation shallowly and narrowly sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3D.</p>
            <p>Etymology: The new species is named after its type locality, Maphelane.</p>
            <p>Variation: Body length: 8.8–8.9 mm, elytral length: 6.9–8.1 mm, maximum width: 5.0– 5.3 mm. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A53493CFFCFAD52FF14FBE4C47F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493CFFC8AD4EF990FE62C41E.text	03EA8A53493CFFC8AD4EF990FE62C41E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica marieskopensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> HYBOSERICA MARIESKOPENSIS SP. NOV.</p>
            <p>(FIGS 4D, 9)</p>
            <p>urn:lsid:zoobank.org:act: 21913568-C65C-46E4-B04C- 7FC0191981E8</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., Mpumalanga  
                <a title="Search Plazi for locations around (long 30.51/lat -24.35)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.51&amp;materialsCitation.latitude=-24.35">Marieskop forest</a>
                 , 24.35S, 30.51E,/ 25.11.2008, E-Y: 3805 light trap; 990 m, leg. Ruth Müller’ (TMSA)  .   Paratypes: 1 ♀ ‘ S. Afr., Mpumalanga  
                <a title="Search Plazi for locations around (long 30.51/lat -24.35)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.51&amp;materialsCitation.latitude=-24.35">Marieskop forest</a>
                 , 24.35S, 30.51E /, 25.11.2008, E-Y: 3805 light trap, 990 m, leg. Ruth Müller’ (ZFMK)  ,   1 ♂ ‘ Südafrika,  
                <a title="Search Plazi for locations around (long 30.833334/lat -24.65)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.833334&amp;materialsCitation.latitude=-24.65">Transvaal Blyderiverspoort Nat. Res.</a>
                 , 24°39’S, 30°50’E, 13.xii.1995, leg. F. Koch’ (ZMHB)  ,   1 ♂ ‘X-DA3682/ X-DA3682 South Africa, Mpumalanga: Blyderiverspoort,  
                <a title="Search Plazi for locations around (long 30.777945/lat -24.577168)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.777945&amp;materialsCitation.latitude=-24.577168">Forever Resort</a>
                 , 1148 m, at light, 24°34’37.8’’S, 30°46’40.6’’E, 16.I.-20.I.2013, leg. Ahrens &amp; Fabrizi  Hyboserica sp 20’ (ZFMK)  ,   7 ♂♂ ‘ South Africa, Mpumalanga, Blyderiverspoort,  
                <a title="Search Plazi for locations around (long 30.777945/lat -24.577168)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.777945&amp;materialsCitation.latitude=-24.577168">Forever Resort</a>
                 , 1148 m, at light, 24°34’37.8’’S, 30°46’40.6’’E, 16.I.-20.I.2013, leg. Ahrens &amp; Fabrizi’ (ZFMK)  . 
            </p>
            <p>Diagnostic description: Body length: 9.8 mm, elytral length: 7.0 mm, maximum width: 5.7 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.44. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.51. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.39. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4D.</p>
            <p>Etymology: The new species is named after its type locality, Marieskop Forest.</p>
            <p>Variation: Body length: 9.6–9.8 mm, elytral length: 7.0– 7.1 mm, maximum width: 5.6–5.7 mm. Without evident sexual dimorphism.</p>
        </div>
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	https://treatment.plazi.org/id/03EA8A53493CFFC8AD4EF990FE62C41E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493BFFC8AEC4F94EFBDAC41E.text	03EA8A53493BFFC8AEC4F94EFBDAC41E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica makutsiensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> HYBOSERICA MAKUTSIENSIS SP. NOV.</p>
            <p>(FIGS 5D, 9)</p>
            <p>urn:lsid:zoobank.org:act: A9EB6F9F-C2F6-4BAB- 9B7F-EED839AD8665</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘840739 South Africa, Limpopo, Legkamalatse Nat. Res.,  
                <a title="Search Plazi for locations around (long 30.337862/lat -24.199888)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.337862&amp;materialsCitation.latitude=-24.199888">Makutsi Bush Camp</a>
                 , 26.11.2009, 24°11’59.6’’S, 30°20’16.3’’E, 816 m, D.Ahrens &amp; S. Fabrizi  Hyboserica sp. ’ (TMSA)  .   Paratypes: 1 ♂, 1 ♀ ‘ S. Afr., Limpopo Prov.  
                <a title="Search Plazi for locations around (long 30.2/lat -24.11)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.2&amp;materialsCitation.latitude=-24.11">Lekgalameetse</a>
                 , 850 m, 24.11S, 30.20E /, 28.11.2008, E-Y: 3811 light trap, leg. Ruth Müller’ (ZFMK, TMSA)  ,   1 ♂ ‘840740 South Africa, Limpopo, Legkamalatse Nat. Res.,  
                <a title="Search Plazi for locations around (long 30.337862/lat -24.199888)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.337862&amp;materialsCitation.latitude=-24.199888">Makutsi Bush Camp</a>
                 , 26.11.2009, 24°11’59.6’’S, 30°20’16.3’’E, 816 m, D. Ahrens &amp; S. Fabrizi  Hyboserica sp. ’ (ZFMK)  ,   1 ♂ ‘ South Africa, N Prov Lekgalameetse Nat. Res.,  Makhutsi Camp , 24.12.1S, 30.20.4E, 14–16.i.1997, R  .   Stals’ (SANC)  . 
            </p>
            <p>Diagnostic description: Body length: 7.5 mm, elytral length: 5.7 mm, maximum width: 5.0 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.44. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.59. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.5. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5D.</p>
            <p>Etymology: The new species is named for its occurrence close to Makutsi Bush Camp.</p>
            <p>Variation: Body length: 7.5–8.8 mm, elytral length: 5.7–6.0 mm, maximum width: 4.8–5.0 mm. Body coloration from yellowish brown to blackish brown. Female: antennal club half as long as remaining antennomeres combined.</p>
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	https://treatment.plazi.org/id/03EA8A53493BFFC8AEC4F94EFBDAC41E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493BFFC9AD9CF94EFDDAC752.text	03EA8A53493BFFC9AD9CF94EFDDAC752.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica muellerae Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA MUELLERAE SP. NOV.</p>
            <p>(FIGS 6F, 9)</p>
            <p>urn:lsid:zoobank.org:act: 62CDC786-7C39-4007-B60B- D5CA41484E6B</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr.; Mpumalanga  
                <a title="Search Plazi for locations around (long 30.51/lat -24.35)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.51&amp;materialsCitation.latitude=-24.35">Marieskop forest</a>
                 , 24.35S, 30.51E,/ 25.11.2008, E-Y: 3805 light trap; 990 m, leg. R. Müller’ (TMSA). 
            </p>
            <p>Diagnostic description: Body length: 9.35 mm, elytral length: 6.23 mm, maximum width: 4.96 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially distinctly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.56. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.4. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.57. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 6F. Female unknown.</p>
            <p>Etymology: The new species is dedicated to Ruth Müller, collector of the new species, who dedicated her life to the study of the beetle fauna of South Africa and who immensely supported our research.</p>
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	https://treatment.plazi.org/id/03EA8A53493BFFC9AD9CF94EFDDAC752	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493AFFC9AEE4FABAFA92C62A.text	03EA8A53493AFFC9AEE4FABAFA92C62A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica nelspruitensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> HYBOSERICA NELSPRUITENSIS SP. NOV.</p>
            <p>(FIGS 4E, 9)</p>
            <p>urn:lsid:zoobank.org:act: 7CE19E10-FF51-463E-8BC2- 95A826BF89F6</p>
            <p> Type material examined:  Holotype: ♂ ‘ S. Afr., Tvl. Nelspruit Nat. Res., Koppee, 25.29S, 30.55E,/ 18.12.1986, E-Y: 2401 ground traps, 53 days, leg. Endrödy-Younga’ (TMSA) .  Paratype: 1 ♂ ‘ S. Afr.; Tvl. Nelspruit Nat. Res., dry valley, 25.29S, 30.55E,/ 8.2.1987, E-Y: 2432 UV light, top valley leg. Endrödy-Younga’ (ZFMK) . </p>
            <p>Diagnostic description: Body length: 8.3 mm, elytral length: 5.3 mm, maximum width: 4.6 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.47. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.56. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.5.Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4E. Female unknown.</p>
            <p>Etymology: The new species is named after its occurrence in the Nelspruit Nature Reserve.</p>
            <p>Variation: Body length: 8.3–8.5 mm, elytral length: 5.3–5.6 mm, maximum width: 4.6–4.7 mm.</p>
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</html>
	https://treatment.plazi.org/id/03EA8A53493AFFC9AEE4FABAFA92C62A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A53493AFFCAADBAFB44FE66C1FC.text	03EA8A53493AFFCAADBAFB44FE66C1FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica ongoyeensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA ONGOYEENSIS SP. NOV.</p>
            <p>(FIGS 3C, 9)</p>
            <p>urn:lsid:zoobank.org:act: 496D5157-D58E-4B82-B21D- 7AE5F1638A30</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., KWZ  
                <a title="Search Plazi for locations around (long 31.44/lat -28.5)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.44&amp;materialsCitation.latitude=-28.5">Natal</a>
                 , Ongoye Forest, 294 m, 28.50S, 31.44E, 4– 5.12.2010, E-Y: 3890 sifting forest litter, leg. Ruth Müller’ (TMSA)  .   Paratype: 1 ♂ ‘ SOUTH AFRICA:  Natal , Nkandla Forest, 1100 m, 12 km S. of Nkandla, 2–3 March 1978, D. &amp; M. Davis, B. Akerbergs’ (USNM)  . 
            </p>
            <p>Diagnostic description: Body length: 9.3 mm, elytral length: 6.6 mm, maximum width: 5.3 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.5. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.56. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.64. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3C. Female unknown.</p>
            <p>Etymology: The new species is named after its type locality, Ongoye Forest.</p>
            <p>Variation: Body length: 9.3–9.6 mm, elytral length: 6.6–7.6 mm, maximum width: 5.3–5.5 mm. Colour from dark to reddish brown.</p>
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	https://treatment.plazi.org/id/03EA8A53493AFFCAADBAFB44FE66C1FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534939FFCAAECAFC6DFA9FC38C.text	03EA8A534939FFCAAECAFC6DFA9FC38C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica schoenhoferi Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SCHOENHOFERI SP. NOV.</p>
            <p>(FIGS 5G, 9)</p>
            <p>urn:lsid:zoobank.org:act: F61E1801-C2D9-4BD6-95A1- E98944000DE5</p>
            <p> Type material examined:   Holotype: ♂ ‘ RSA:  Northern Province , W Soutpansberg, Lajuma, X / XI.2002, trap 7, 1300 m, leg. A. Schönhofer’ (SMNS). </p>
            <p>Diagnostic description: Body length: 8.19 mm, elytral length: 5.54 mm, maximum width: 4.15 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.28. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.4. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.83. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5G. Female unknown.</p>
            <p>E t y m o l o g y: T h e n e w s p e c i e s i s d e d i c a t e d t o A. Schönhofer, collector of the new species.</p>
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	https://treatment.plazi.org/id/03EA8A534939FFCAAECAFC6DFA9FC38C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534939FFCBAD9AFDFDFDFAC3C8.text	03EA8A534939FFCBAD9AFDFDFDFAC3C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica sebastiani Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SEBASTIANI SP. NOV.</p>
            <p>(FIGS 1, 4B, 9)</p>
            <p>urn:lsid:zoobank.org:act: 2688BA11-0E32-4E0E-AA86- E9C277E5E28A</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., Cape  
                <a title="Search Plazi for locations around (long 23.07/lat -34.0)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.07&amp;materialsCitation.latitude=-34.0">Keurboomstrand</a>
                 , 34.00S, 23.07E /, 11.12.1976, E-Y: 1308 light collection, leg. Endrödy-Younga’ (TMSA)  .   Paratypes: 1 ♂ ‘X-DA4029 South Africa, Western Cape, Goukamma Nature Reserve, Dunes near  
                <a title="Search Plazi for locations around (long 22.947443/lat -34.06747)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.947443&amp;materialsCitation.latitude=-34.06747">Rondavell Chalet</a>
                 , 10 m, 34°04’02.9’’S; 22°56’50.8’’E, 24.–26.xi.2013, lg. Ahrens, Eberle, Fabrizi’ (ZFMK)  ,   6 ♂♂ ‘ South Africa Western Cape, Goukamma Nature Reserve, Dunes near  
                <a title="Search Plazi for locations around (long 22.947443/lat -34.06747)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.947443&amp;materialsCitation.latitude=-34.06747">Rondavell Chalet</a>
                 , 10 m, 34°04’02.9’’S; 22°56’50.8’’E, 24.–26.xi.2013, lg. Ahrens, Eberle, Fabrizi’ (ZFMK)  . 
            </p>
            <p>Diagnostic description: Body length: 8.7 mm, elytral length: 6.0 mm, maximum width: 4.8 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.43. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.48. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.84. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4B. Female unknown.</p>
            <p>Etymology: The new species is named for the collector of the type specimen, Sebastian Endrödy-Younga.</p>
            <p>Variation: Body length: 7.2–8.7 mm, elytral length: 5.8–6.5 mm, maximum width: 4.2–4.9 mm.</p>
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	https://treatment.plazi.org/id/03EA8A534939FFCBAD9AFDFDFDFAC3C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534938FFCBAF31FE21FDA9C4EA.text	03EA8A534938FFCBAF31FE21FDA9C4EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica sergeji Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SERGEJI SP. NOV.</p>
            <p>(FIGS 4A, 9)</p>
            <p>urn:lsid:zoobank.org:act: ECFC931A-6AD1-4CBD-890C-6811B98D427F</p>
            <p> Type material examined:   Holotype: ♂ ‘ S. Africa Eastern Cape pr., Near  Port St. John’s, 55 m, 10.01.2002, S. Murzin leg.’ (ZFMK). </p>
            <p>Diagnostic description: Body length: 10.73 mm, elytral length: 7.38 mm, maximum width: 5.83 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.48. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially narrowly interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.45. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/ length: 1/3.91. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spinesattwo-thirdsofmetatibiallength.Apexofmetatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate.Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4A. Female unknown.</p>
            <p>Etymology: The new species is dedicated to Sergej Murzin, collector of the new species.</p>
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	https://treatment.plazi.org/id/03EA8A534938FFCBAF31FE21FDA9C4EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534938FFCBAEECF902FA92C731.text	03EA8A534938FFCBAEECF902FA92C731.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica shilouvaneensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SHILOUVANEENSIS SP. NOV.</p>
            <p>(FIGS 5C, 9)</p>
            <p>urn:lsid:zoobank.org:act: 84831D63-0F78-4B1C-A740- 7BA711E351B7</p>
            <p> Type material examined:  Holotype: ♂ ‘Shilouvane, 2.1906, Rev. Junod’ (TMSA) .  Paratype: 1 ♂ ‘ SOUTH AFRICA: Trsvl. Mooketsi, 14–18-II-1968, Krombein &amp; Spangler’ (USNM) . </p>
            <p>Diagnostic description: Body length: 6.8 mm, elytral length: 5.0 mm, maximum width: 4.3 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.42. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.5. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.76. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5C. Female unknown.</p>
            <p>Etymology: The new species is named after the type locality Shilouvane.</p>
            <p>Variation: Body length: 6.8–8.5 mm, elytral length: 5.0– 6.1 mm, maximum width: 4.3–4.5 mm.</p>
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	https://treatment.plazi.org/id/03EA8A534938FFCBAEECF902FA92C731	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534938FFC4ADA5FA5AFE62C458.text	03EA8A534938FFC4ADA5FA5AFE62C458.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica shulmani Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> HYBOSERICA SHULMANI SP. NOV.</p>
            <p>(FIGS 6B, 9)</p>
            <p>urn:lsid:zoobank.org:act: 94AEDC3B-8064-40C6-81A4- 99CAA8FA7958</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr.;  KwaZulu-Natal , Malta forest, 10.2.1974, B. Shulman’ (TMSA)  .   Paratypes: 1 ♂ ‘ S. Afr.,  
                <a title="Search Plazi for locations around (long 31.35/lat -24.59)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.35&amp;materialsCitation.latitude=-24.59">Kruger Nat. Park</a>
                 , Skukuza, 24.59S, 31.35E,/ 17.1.1996, E-Y: 3202 UV  light, leg. Endrödy-Younga’ (ZFMK),   1 ♂ ‘RSA-Mpumalanga, 20 km SE  Nelspruit , 5.-6.2004, J. Halada leg.’ (CPPB)  ,  1 ♂, 1 ♀ ‘ SOUTH AFRICA, E.Tvl., Barbeton, 25.48S, 31.03E, Dec. 1978, G.L. Prinsloo &amp; C.D. Eardley, At light / NATIONAL COLL .   OF INSECTS,  Pretoria , S.Afr. ’ (SANC)  ,   1 ♂ ‘ SOUTH AFRICA,  
                <a title="Search Plazi for locations around (long 30.54/lat -24.32)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.54&amp;materialsCitation.latitude=-24.32">Tvl.</a>
                 , Swadini, Blydepoort Nat.Res., 24.32S, 30.54E, 26–29.i.1987, B. Grobbelaar / Collected at light / NATIONAL COLL  .   OF INSECTS,  Pretoria , S.Afr. ’ (SANC)  ,   1 ♂ ‘Südafrika- [M]  Pumalanga Prov. 2.5 km w Barberton, 650 m, Buschsavanna TF (SA03), S25°44,496’, E030°59,532’, 14.01.2007, leg. J.-P. Rudloff’ (ZFMK)  ,   1 ♂ ‘Südafrika- [M]  Pumalanga Prov. 2.5 km W Barberton, 650 m (SA04), Inner Krater Westhang NF Subarabes Busch und Gartenland, S 25°45,035’, E030°59,532’, 15.01.2007, leg. J.-P. Rudloff’ (ZFMK)  ,   1 ♂ ‘  Nelspruit nr. Barberton Transvaal/SAM-COL A070352’ (SAMC)  . 
            </p>
            <p>Diagnostic description: Body length: 9 mm, elytral length: 6.5 mm, maximum width: 5.4 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.48. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.56. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.48. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 6B.</p>
            <p>Etymology: The new species is named for B. Shulman, collector of the type specimen.</p>
            <p>Variation: Body length: 8.4–9.8 mm, elytral length: 5.9–7.1 mm, maximum width: 4.5–5.8 mm. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A534938FFC4ADA5FA5AFE62C458	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534937FFC4AF14F9B9FB9EC52F.text	03EA8A534937FFC4AF14F9B9FB9EC52F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica sipeki Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SIPEKI SP. NOV.</p>
            <p>(FIGS 5E, 9)</p>
            <p>urn:lsid:zoobank.org:act: 6A0754F1-7712-459E-8D85- B1DCDF0062CF</p>
            <p> Type material examined:  Holotype: ♂ ‘ RSA, Eastern C a p e, C a c a d u d i s t r i c t, C a m d e b o o, S n e e b e r g, S31.8447, N[sic!; E]24.5286, 29.–31.XII.2014, P. Sipek &amp; P. Malec leg.’ (SANC) .   Paratypes: 2 ♂♂ ‘ RSA, Eastern Cape, Cacadu district,  Camdeboo , Sneeberg, S 31.8447, N[sic!; E]24.5286, 29.–31. XII  .2014, P. Šípek &amp; P. Malec leg.’ (ZFMK),   1 ♂ ‘X-DA4491 South Africa, Eastern Cape, Cacadu district,  Cambeboo local municipality Sneeuberge; Nieu Bethseda env. Near road to Compassberg 29.–31. XII  .2014, S31.844745, E24.5286, P. Šípek &amp; P. Malec lgt’ (ZFMK) ,   1 ♀ ‘X-DA4495 South Africa, Eastern Cape, Cacadu district,  Cambeboo local municipality; Unnamed road between N9 and Nieu Bethseda (Bethesdaweg) 29.–31. XII  .2014, between S31.906327, E24.752422 and S31.836980°, E24.577667°, P. Šípek &amp; P. Malec lgt’ (ZFMK) ,   2 ♂♂ ‘ RSA / Eastern Cape, Sneeuberge,  Nieu Bethseda between S31.844745°, E24.5286°, 29.–31. XII  .2014, black light, P. Malec &amp; P. Šípek lgt.’ (ZFMK) . </p>
            <p>Diagnostic description: Body length: 9.8 mm, elytral length: 7.5 mm, maximum width: 5.5 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Antennal club shorter than remaining antennomeres combined. Eyes, ratio diameter/interocular width: 0.47. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.41. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.48. Basal group of dorsal spines of metatibia at first-quarter of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation shallowly and narrowly sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5E.</p>
            <p>Etymology: The new species is named after Petr Šípek (Prague), one of the collectors of the new species.</p>
            <p>Variation: Body length: 9.8–11.6 mm, elytral length: 7.5–8.5 mm, maximum width: 5.5–6.0 mm. Colour varies from reddish to dark brown. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A534937FFC4AF14F9B9FB9EC52F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534936FFC5AED3FF14FB48C6CF.text	03EA8A534936FFC5AED3FF14FB48C6CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica sodwanensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SODWANENSIS SP. NOV.</p>
            <p>(FIGS 4C, 9)</p>
            <p>urn:lsid:zoobank.org:act: 57D6A780-DA41-43E1-94EA-20840FA860B3</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr., KWZ  Natal , Sodwana Bay, December 2010, at light leg. Martin Krüger’ (TMSA)  .   Paratypes: 1 ♀ ‘ S. Afr., KWZ Natal,  Sodwana Bay , December 2010, at light leg. Martin Krüger’ (TMSA)  ,   1 ♂ ‘ RSA:  
                <a title="Search Plazi for locations around (long 32.23/lat -27.59)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.23&amp;materialsCitation.latitude=-27.59">Natal False Bay</a>
                 Park cmpgrd [= camp ground], 27.59S, 32.23E, Evans, d’Hotman, Nel’ (TMSA)  ,   1 ♂ ‘835154 X-DA0781 South Africa, Kwazulu-Natal: Ithala Game Reserve, Ntshodwe camp (  Louwsburg env.), 1036 m, at light, 027°32’’36’’S, 031°17’00.3’’E, leg. D. Ahrens &amp; S. Fabrizi  Hyboserica spSA  2 ♂ ’ (ZFMK) ,   1 ♂ ‘842092 X-DA0784 South Africa, Kwazulu-Natal: Ithala Game Reserve, Ntshodwe camp (  Louwsburg env.), 1036 m, at light, 027°32’’36’’S, 031°17’00.3’’E, 27.XI.2006, leg. D. Ahrens &amp; S. Fabrizi  Hyboserica spSA  2 ♂ ’ (ZFMK) ,   1 ♂ ‘842091 X-DA0784 South Africa, Kwazulu-Natal: Ithala Game Reserve, Ntshodwe camp (  Louwsburg env.), 1036 m, at light, 027°32’’36’’S, 031°17’00.3’’E, 27.XI.2006, leg. D. Ahrens &amp; S. Fabrizi  Hyboserica spSA  2 ♂ ’ (ZFMK) ,   1 ♂ ‘842090 X-DA0783 South Africa, Kwazulu-Natal: Ithala Game Reserve, Ntshodwe camp (  Louwsburg env.), 1036 m, at light, 027°32’’36’’S, 031°17’00.3’’E, 27.XI.2006, leg. D. Ahrens &amp; S. Fabrizi  Hyboserica spSA  2 ♂ ’ (ZFMK) ,   1 ♂ ‘DA0782  Hyboserica spSA2 South Africa, Kwazulu-Natal: Ithala Game Reserve, Ntshodwe camp (  Louwsburg env.), 1036 m, at light, 027°32’’36’’S, 031°17’00.3’’E, 27.XI.2006, leg. D. Ahrens &amp; S. Fabrizi’ (ZFMK)  ,   1 ♂ ‘ Zoulouland [Zululand] Dr. Martin, Slg. R.  Oberthür (Coll. C. Martin) Eing. Nr. 4, 1956’ (ZFMK)  ,  1 ♂, 1 ♀ ‘ R. S.A., 10.– 23.12.1992, Natal: Itala Game Res. Louwsburg, leg. F. Koch’ (ZMHB) ,   2 ♂♂ ‘ Südafrika: 5.xii.1995, KwaZulu Natal,  
                <a title="Search Plazi for locations around (long 32.083332/lat -28.033333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.083332&amp;materialsCitation.latitude=-28.033333">Hluhluwe Game Res.</a>
                 28°02’S / 32°05’E leg. F. Koch’ (ZMHB)  ,   1 ♂ ‘ Südafrika: 10.ii.1995, KwaYulu Natal,  
                <a title="Search Plazi for locations around (long 32.083332/lat -28.033333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.083332&amp;materialsCitation.latitude=-28.033333">Hluhluwe Game Res.</a>
                 28°02’S, 32°05’E, leg. F. Koch’ (ZMHB)  ,   1 ♂, 1 ♀ ‘ RSA Transvaal,  Pullen-Farm , 30 km SE Nelspruit, 24./ 26. xi.2004, leg. W. Mey’ (ZMHB)  ,   1 ♂ ‘ Südafrika: 8.xii.1995, KwaZulu Natal:  
                <a title="Search Plazi for locations around (long 31.333334/lat -27.5)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.333334&amp;materialsCitation.latitude=-27.5">Itala Game Res.</a>
                 27°30’S, 31°20’E, leg. F. Koch’ (ZMHB)  ,   1 ♀ ‘X-DA4088 SA10 -ITH-L01 South Africa,  Ithala Research Camp , Oct. 2010, light trap, S. von Noort’ (SAMC)  ,   1 ♂ ‘X-DA4094 South Africa,  Ithala Research Camp , 9 Oct. 2010, light trap, S. von Noort’ (SAMC)  ,   1 ♂ ‘X-DA4095 South Africa,  Ithala Research Camp , 9 Oct. 2010, light trap, S. von Noort’ (SAMC)  . 
            </p>
            <p>Diagnostic description: Body length: 9.2 mm, elytral length: 6.9 mm, maximum width: 5.6 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.41. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.56. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.43. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4C.</p>
            <p>Etymology: The new species is named after its type locality, Sodwana Bay.</p>
            <p>Variation: Body length: 8.5–9.2 mm, elytral length: 6.5–6.9 mm, maximum width: 4.8–5.6 mm. Coloration from dark brown to reddish or yellowish brown. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A534936FFC5AED3FF14FB48C6CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534936FFC6AD66FB3DFE8FC752.text	03EA8A534936FFC6AD66FB3DFE8FC752.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica soutpansbergensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA SOUTPANSBERGENSIS SP. NOV.</p>
            <p>(FIGS 4F, 9)</p>
            <p>urn:lsid:zoobank.org:act: 673302BE-F54B-49B5-BCD5- BEA21AF4E35F</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘840800 South Africa, Limpopo,  
                <a title="Search Plazi for locations around (long 29.869862/lat -22.97089)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.869862&amp;materialsCitation.latitude=-22.97089">Soutpansberg</a>
                 , 10 km NW of Makhado, 24.– 25.11.2009, 22°58’15.2’’S, 29°52’11.5’’E, 1338 m, D. Ahrens &amp; S. Fabrizi,  Hyboserica sp. ’ (TMSA)  .   Paratypes: 1 ♀ ‘842660 South Africa, Limpopo,  
                <a title="Search Plazi for locations around (long 29.869862/lat -22.97089)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.869862&amp;materialsCitation.latitude=-22.97089">Soutpansberg</a>
                 , 10 km NW of Makhado 24.– 25.11.2009, 22°58’15.2’’S, 29°52’11.5’’E, 1338 m, D. Ahrens &amp; S. Fabrizi,  Hyboserica sp. ’ (CA)  ,   1 ♀ ‘842659 South Africa, Limpopo,  
                <a title="Search Plazi for locations around (long 29.869862/lat -22.97089)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.869862&amp;materialsCitation.latitude=-22.97089">Soutpansberg</a>
                 , 10 km NW of Makhado, 24.– 25.11.2009, 22°58’15.2’’S, 29°52’11.5’’E, 1338 m, D. Ahrens &amp; S. Fabrizi  Hyboserica sp. ’ (CA)  ,   1 ♂ ‘ RSA: Limpopo Prov.,  Soutpansberg , 15 km N Makhado (Louis Trichardt), 1400–1500 m, 2.–5. XII  .2008, leg. W. Schawaller’ (SMNS). 
            </p>
            <p>Diagnostic description: Body length: 8.19 mm, elytral length: 6.23 mm, maximum width: 4.62 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.55. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.41. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.5. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 4F.</p>
            <p>Etymology: The new species is named after the type locality Soutpansberg.</p>
            <p>Variation: Body length: 8.2–9.2 mm, elytral length: 6.2–6.4 mm, maximum width: 4.6–4.7 mm. Female: Antennal club 0.8 times as long as remaining antennomeres combined, otherwise no differences to the male.</p>
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	https://treatment.plazi.org/id/03EA8A534936FFC6AD66FB3DFE8FC752	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534935FFC6AF17FABAFBF6C1C0.text	03EA8A534935FFC6AF17FABAFBF6C1C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica uhligi Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA UHLIGI SP. NOV.</p>
            <p>(FIGS 6E, 9)</p>
            <p>urn:lsid:zoobank.org:act: 5705128B-9CD1-4748-81D2- 34487FC1BBBD</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ Zimbabwe, 7.xii.1993, 18°27’S / 32°47’E, Nyanga NP: Pungwe- Gorge,  
                <a title="Search Plazi for locations around (long 32.783333/lat -18.45)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.783333&amp;materialsCitation.latitude=-18.45">Pungwe River</a>
                 banks + wood litter sievings, leg. M. Uhlig’ (ZMHB). 
            </p>
            <p>Diagnostic description: Body length: 9 mm, elytral length: 6.35 mm, maximum width: 4.62 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.52. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.35. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.75. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 6E. Female unknown.</p>
            <p>Etymology: The new species is dedicated to M. Uhlig, collector of the new species.</p>
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	https://treatment.plazi.org/id/03EA8A534935FFC6AF17FABAFBF6C1C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534935FFC7ADABFC29FD03C1FC.text	03EA8A534935FFC7ADABFC29FD03C1FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica uitsoekensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA UITSOEKENSIS SP. NOV.</p>
            <p>(FIGS 5B, 9)</p>
            <p>urn:lsid:zoobank.org:act: A2B048E9-BF3A-4716-B9E3- C5DE41B17A13</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr.;  
                <a title="Search Plazi for locations around (long 30.33/lat -25.16)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.33&amp;materialsCitation.latitude=-25.16">Tvl. Uitsoek Waterfall area</a>
                 , 25.16S – 30.33E / 13.12.1986; E-Y: 2385 UV light collection, leg. Endrödy-Younga’ (TMSA)  .  Paratypes: 1 ♂ ‘ S. Afr.; Tvl. Uitsoek Grootkloof, ind. for. 25.15S – 30.33E / 25.10.1986; E-Y: 2319 intercept trap, 50d., leg. Endrödy-Younga’ (TMSA) ,  1 ♂ ‘Marieps Mnt., Dec. 1925, G. van Son’ (TMSA) ,   1 ♂ ‘840794 South Africa Mpumalanga  
                <a title="Search Plazi for locations around (long 30.808332/lat -24.99311)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.808332&amp;materialsCitation.latitude=-24.99311">Mac forest</a>
                 retreat, c. 8km N of Sabie, 1.– 2.12.2009, 24°59’35,2’’S, 30°48’30,0’’E, 1278 m, D. Ahrens &amp; S. Fabrizi  Hyboserica sp. ’ (ZFMK)  . 
            </p>
            <p>Diagnostic description: Body length: 8.0 mm, elytral length: 5.9 mm, maximum width: 4.6 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/ interocular width: 0.68. Antennal club slightly longer than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.24. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/4.12. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 5B. Female unknown.</p>
            <p>Etymology: The new species is named after the type locality Uitsoek.</p>
            <p>Variation: Body length: 8.0–10.0 mm, elytral length: 5.9–6.6 mm, maximum width: 4.6–5.1 mm. Body colour varies from dark brown to yellowish brown.</p>
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	https://treatment.plazi.org/id/03EA8A534935FFC7ADABFC29FD03C1FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534934FFC7AED1FC6DFC67C073.text	03EA8A534934FFC7AED1FC6DFC67C073.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica umgeniensis Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA UMGENIENSIS SP. NOV.</p>
            <p>(FIGS 3F, 9)</p>
            <p>urn:lsid:zoobank.org:act: 3F29D375-47FB-42B1-B3C7- 68EB6B4A122D</p>
            <p> Type material examined:   Holotype: ♂ ‘  Umgeni River Valley Bothas Hill distr. , Ntl. Dec. 196., R. F. Lawrence’ (TMSA). </p>
            <p>Diagnostic description: Body length: 9.46 mm, elytral length: 6.69 mm, maximum width: 5.77 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.44. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially narrowly interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.42. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.28. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3F. Female unknown.</p>
            <p>Etymology: The new species is named after the type locality Umgeni.</p>
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	https://treatment.plazi.org/id/03EA8A534934FFC7AED1FC6DFC67C073	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534934FFC0AD8BFD8EFE62C088.text	03EA8A534934FFC0AD8BFD8EFE62C088.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyboserica vogti Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> HYBOSERICA VOGTI SP. NOV.</p>
            <p>(FIGS 3B, 9)</p>
            <p>urn:lsid:zoobank.org:act: 7C98CB52-B2E6-4FD5-967C-407AA3964816</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ SOUTH AFRICA, KZN, near  
                <a title="Search Plazi for locations around (long 32.16/lat -28.38)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.16&amp;materialsCitation.latitude=-28.38">Lake Nhalabane</a>
                 , 25 km NE of Richards Bay, 28.38S, 32.16E, 20.xi.-12.xii.1992, M. Vogt / Univ. of Pretoria Project, Richards Bay Minerals / NATIONAL COLL. OF INSECTS, Pretoria, S. Afr. dune forest rehabilitation survey’ (SANC)  .  Paratypes: 2 ♂♂, 2 ♀♀ ‘ SOUTH AFRICA, KZN ,   near  
                <a title="Search Plazi for locations around (long 32.16/lat -28.38)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.16&amp;materialsCitation.latitude=-28.38">Lake Nhalabane</a>
                 , 25 km NE of Richards Bay, 28.38S, 32.16E, 20.xi.-12.xii.1992, M. Vogt / Univ. of Pretoria Project, Richards Bay Minerals / NATIONAL COLL  .   OF INSECTS,  Pretoria , S.Afr. dune forest rehabilitation survey’ (SANC, ZFMK)  ,  1 ♂ ‘ South Africa: KZN  
                <a title="Search Plazi for locations around (long 31.25/lat -27.49)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.25&amp;materialsCitation.latitude=-27.49">Ngome State Forest</a>
                 , 27.49S, 31.25E, 1100 m, 5./ 8.ii.1996, R  .   Stals / Collected at light trap in indigenous forest’ (SANC)  ,  1 ♂ ‘ SOUTH AFRICA, NATAL ,   nr.  
                <a title="Search Plazi for locations around (long 32.15/lat -28.25)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.15&amp;materialsCitation.latitude=-28.25">Richards Bay</a>
                 , 28.25S, 32.15E, XII  .1981, P. Reavell / NATIONAL COLL .   OF INSECTS,  Pretoria , S. Afr. ’ (SANC)  . 
            </p>
            <p>Diagnostic description: Body length: 9.2 mm, elytral length: 7.1 mm, maximum width: 5.1 mm. Labroclypeus narrow and long; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially straight. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.43. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum widely absent. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge complete over entire elytral length. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/ metacoxa: 1/1.66. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metatibia, ratio of width/length: 1/3.6. Basal group of dorsal spines of metatibia at first-third of metatibial length; apical group of external spines at two-thirds of metatibial length. Apex of metatibia interiorly near tarsal articulation not sinuate or truncate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally impunctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 3B.</p>
            <p>Etymology: The new species is dedicated to Mark Vogt, one of the collectors of the new species.</p>
            <p>Variation: Body length: 8.4–9.2 mm, elytral length: 6.6–7.1 mm, maximum width: 4.5–5.1 mm. Without evident sexual dimorphism.</p>
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	https://treatment.plazi.org/id/03EA8A534934FFC0AD8BFD8EFE62C088	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534933FFC0AD7BFF14FB30C37F.text	03EA8A534933FFC0AD7BFF14FB30C37F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leoserica capensis (BRENSKE 1902) Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LEOSERICA CAPENSIS (BRENSKE, 1902) COMB. NOV.</p>
            <p>(FIGS 7C, 9)</p>
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	https://treatment.plazi.org/id/03EA8A534933FFC0AD7BFF14FB30C37F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534933FFC0AF6BFCE1FECFC50D.text	03EA8A534933FFC0AF6BFCE1FECFC50D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leoserica Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LEOSERICA GEN. NOV.</p>
            <p>urn:lsid:zoobank.org:act: 59B4AE95-CB98-4FE4-9A0B-75395E303CE1</p>
            <p>Ty p e s p e c i e s: L e o s e r i c a k n y s n a n a (b y p r e s e n t designation).</p>
            <p>Diagnosis: Body of medium size (8–11 mm), elongately oval, reddish to yellowish or dark brown, dorsal surface with iridescent shine; dorsal surface nearly glabrous. Labroclypeus moderately wide or wide, trapezoidal, with fine and dense punctures; anterior margin of labroclypeus weakly sinuate, rarely distinctly sinuate. Mentum anteriorly slightly elevated.Antenna composed of nine antennomeres, club short, in both sexes with three antennomeres. Pronotum subtrapezoidal and moderately wide, lateral margins sparsely setose, anterior angles well produced and moderately sharp, anterior margin convexly produced medially; basal margin with complete marginal line. Hypomeron simple, not carinate. Elytra with reduced epipleural margin, apical margin chitinous, without membranous rim of microtrichomes, intervals flat, evenly finely punctate, glabrous, odd intervals with punctures concentrated along the weakly impressed striae, apical setae on sutural interval absent. Ventral surface sparsely covered with short setae. Metatibia dorsally with two groups of spines and longitudinally convex; at apex not truncate or sinuate near tarsal articulation. Protibia bi- or tridentate, basal tooth small.</p>
            <p> Etymology: The name of the new genus is a combination of the traditional genus name  Serica and the prefix ‘Leo-’, lion, but which refers to Silvia and Dirk’s cat, Leo. </p>
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	https://treatment.plazi.org/id/03EA8A534933FFC0AF6BFCE1FECFC50D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534933FFC0AD04FEC3FB7AC4EE.text	03EA8A534933FFC0AD04FEC3FB7AC4EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Serica capensis Brenske 1902	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Serica capensis Brenske, 1902: 26 . </p>
            <p> Type material examined:   Lectotype (here designated). ♂ ‘ Cap b. sp. Coll Chevr. / Coll. Brenske /  Serica capensis Typ. Brsk. ’ (ZMHB)  .   Paralectotype: 1 ♂ ‘CT [Cape  Town ] 1885/ v. Peringuey/ Caloserica capicola co-type P[éringue]y’ (ZMHB)  . </p>
            <p> Additional material examined:  1 ♂ ‘Kimberley Bro. Power 1912/ SAM COL A070308’ (SAMC, record doubtful) . </p>
            <p>Diagnostic description: Body length: 8.77 mm, elytral length: 6.35 mm, maximum width: 4.62 mm. Labroclypeus narrow and long; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.37. Antennal club shorter than remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum present. Marginal line of anterior margin of pronotum medially narrowly interrupted. Hind wings reduced in length. Epipleural edge reduced behind anterior half of elytra. Metasternum and metepisernites in length reduced. Ratio of length of metepisternum/metacoxa:1/2.Protibia, externally tridentate; external margin between apical two teeth moderately sinuated. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Metatibia, ratio of width/length: 1/3.5. Basal group of dorsal spines of metatibia before basal quarter of metatibial length; apical group of external spines about at half of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 7C.</p>
            <p>Remark: Paralectotype doubtful; it does not bear Brenske’s labels. The name on its labels Caloserica capiloa is a nomen nudum. The additional specimen from Kimberly probably bears an erroneous locality label. The species is so far only strictly known from the Cape, but no recent records are known.</p>
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	https://treatment.plazi.org/id/03EA8A534933FFC0AD04FEC3FB7AC4EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534933FFC1AD80F907FDEBC6AE.text	03EA8A534933FFC1AD80F907FDEBC6AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leoserica hotovyi Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LEOSERICA HOTOVYI SP. NOV.</p>
            <p>(FIGS 7B, 9)</p>
            <p>urn:lsid:zoobank.org:act: 2F1DDB79-7C13-4238-B20C- 4E9675337B0B</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ RSA (Western Cape), 4.ii.2005,  
                <a title="Search Plazi for locations around (long 20.4005/lat -34.452835)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.4005&amp;materialsCitation.latitude=-34.452835">De Hoop Nature Reserve</a>
                 34°27.17’S; 20°24.03’E; 24 m, J. Hotovy &amp; J. Mateju leg.’ (ZFMK). 
            </p>
            <p>Diagnostic description: Body length: 7.96 mm, elytral length: 5.19 mm, maximum width: 4.04 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus distinct. Eyes, ratio diameter/interocular width: 0.4. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum present. Marginal line of anterior margin of pronotum complete. Hind wings fully developed. Epipleural edge reduced behind anterior half of elytra. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.61. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth deeply sinuated. Metatibia, ratio of width/length: 1/4.69. Basal group of dorsal spines of metatibia before basal quarter of metatibial length; apical group of external spines about at half of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 7B. Female unknown.</p>
            <p>Etymology: The new species is named for J. Hotovy, one of the collectors of the type specimen.</p>
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	https://treatment.plazi.org/id/03EA8A534933FFC1AD80F907FDEBC6AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534932FFC1AF21FAF1FB6EC1E2.text	03EA8A534932FFC1AF21FAF1FB6EC1E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leoserica jamesi Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LEOSERICA JAMESI SP. NOV.</p>
            <p>(FIGS 7D, 9)</p>
            <p>urn:lsid:zoobank.org:act: 2B8BDC33-7007-4F59-8CD6- D3F528642D67</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr. W coast  
                <a title="Search Plazi for locations around (long 18.02/lat -31.24)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.02&amp;materialsCitation.latitude=-31.24">Vredendal distr.</a>
                 Elsie Erasmus Kloof 158/ 31.24S – 18.02E, 7.–8.XII.1996, leg. J. du G. Harrison’ (TMSA). 
            </p>
            <p>Diagnostic description: Body length: 10.85 mm, elytral length: 7.15 mm, maximum width: 5.19 mm. Labroclypeus narrow and long; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially weakly sinuate. Angle between lateral margin of labroclypeus and ocular canthus indistinct. Eyes, ratio diameter/interocular width: 0.48. Antennal club as long as remaining antennomeres combined. Mentum flat and longitudinally concave anteriorly. Basal marginal line of pronotum present. Marginal line of anterior margin of pronotum medially widely interrupted. Hind wings fully developed. Epipleural edge reduced behind anterior half of elytra. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.51. Metacoxa densely setose; its anterior marginal line complete. Protibia externally bidentate; external margin between apical two teeth deeply sinuated. Metatibia, ratio of width/length: 1/3.5. Basal group of dorsal spines of metatibia before basal quarter of metatibial length; apical group of external spines about at half of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face with a convex elevated carina dorsal of tarsal articulation. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1, 1.5 times as long as subsequent tarsomere. Aedeagus and habitus: Fig. 7D. Female unknown.</p>
            <p>Etymology: The new species is dedicated to James du G. Harrison, collector of the new species, who greatly supported our research in South Africa.</p>
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	https://treatment.plazi.org/id/03EA8A534932FFC1AF21FAF1FB6EC1E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
03EA8A534932FFC2AD9DFC38FD08C1FC.text	03EA8A534932FFC2AD9DFC38FD08C1FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leoserica knysnana Fabrizi & Eberle & Ahrens 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LEOSERICA KNYSNANA SP. NOV.</p>
            <p>(FIGS 7A, 9)</p>
            <p>urn:lsid:zoobank.org:act: F2461379-62CA-4B55-A871- A1CC70AAB73C</p>
            <p>
                 Type material examined:   Holotype: ♂ ‘ S. Afr. W Cape 20 km W  Knysna 15– 20.2.1978 ’ (TMSA)  .   Paratypes: 2 ♂♂ ‘ S. Afr. W Cape 20 km W  Knysna 15– 20.2.1978 ’ (TMSA, ZFMK)  ,   1 ♂ ‘X-DA4364 South Africa Western Cape: Goukamma Nature Reserve; Dunes near  
                <a title="Search Plazi for locations around (long 22.947443/lat -34.06747)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.947443&amp;materialsCitation.latitude=-34.06747">Rondavell Chalet</a>
                 , 10m 34°04’02,9’’S; 22°56’50,8’’E 24.–26.xi.2013 lg. Ahrens, Eberle, Fabrizi’ (ZFMK)  ,   6 ♂♂ ‘ South Africa Western Cape: Goukamma Nature Reserve; Dunes near  
                <a title="Search Plazi for locations around (long 22.947443/lat -34.06747)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.947443&amp;materialsCitation.latitude=-34.06747">Rondavell Chalet</a>
                 , 10m 34°04’02,9’’S; 22°56’50,8’’E 24.–26.xi.2013 lg. Ahrens, Eberle, Fabrizi’ (ZFMK)  . 
            </p>
            <p>Diagnostic description: Body length: 9.1 mm, elytral length: 6.4 mm, maximum width: 4.6 mm. Labroclypeus wide and short; not incised or narrowed behind labrum. Anterior margin of labroclypeus medially distinctly sinuate. Angle between lateral margin of labroclypeus and ocular canthus distinct. Eyes, ratio diameter/interocular width: 0.42. Antennal club as long as remaining antennomeres combined. Mentum convexly elevated and slightly flattened anteriorly. Basal marginal line of pronotum present. Marginal line of anterior margin of pronotum complete. Hind wings fully developed. Epipleural edge reduced behind anterior half of elytra. Metasternum and metepisernites in length normal. Ratio of length of metepisternum/metacoxa: 1/1.5. Metacoxa glabrous except a few fine lateral setae; its anterior marginal line complete. Protibia, externally tridentate; external margin between apical two teeth deeply sinuated. Metatibia, ratio of width/length: 1/4. Basal group of dorsal spines of metatibia before basal quarter of metatibial length; apical group of external spines about at half of metatibial length. Apex of metatibia interiorly near tarsal articulation concavely sinuate; apical face simple, not elevated or produced. Meso- and metatarsomeres dorsally punctate. Mesotarsomere 1 slightly longer than the subsequent tarsomere. Aedeagus and habitus: Fig. 7A. Female unknown.</p>
            <p>Etymology: The new species is named after its occurrence close to Knysna.</p>
            <p>Variation: Body length: 7.9–9.1 mm, elytral length: 6.1–6.4 mm, maximum width: 4.2–4.6 mm.</p>
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	https://treatment.plazi.org/id/03EA8A534932FFC2AD9DFC38FD08C1FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fabrizi, Silvia;Eberle, Jonas;Ahrens, Dirk	Fabrizi, Silvia, Eberle, Jonas, Ahrens, Dirk (2019): Unexpected diversity of Hyboserica chafers in South African forest remnants: cladistic analysis, new species and the new genus Leoserica (Coleoptera: Scarabaeidae: Melolonthinae). Zoological Journal of the Linnean Society 186: 950-982
