taxonID	type	description	language	source
03EA6A15B10D97573D90FA1F62C54047.taxon	type_taxon	Type Species. Saprositellus denticulatus Balthasar (Designated by Balthasar 1967: 138). 451 Diagnosis. Body elongate oval, convex, feebly shining. Head relatively small, deflexed, weakly convex dorsally; clypeus dilated to cover mouthparts with inflexed triangular process medially; lateral clypeal edge on each side with row of denticles. Eye small, concealed under pronotal margin; mouthparts adapted for soft saprophagy. Prothoracic and clypeal indentations together form cavity to receive fore legs. Pronotum narrow, subquadrate, strongly elevated, sides steep, invisible from above; lateral margin with tooth, posterior angles excavate to basal angulation; surface usually with median longitudinal furrow and small fovea laterally. Scutellum small, parallelsided. Elytra with fine basal bead and with humeral denticles; striae deep, strial punctures coarse, intervals usually narrow. Metathoracic wings functional. Ventral sclerites punctate; posterior prosternal process large, triangular; mesosternum lower than metasternum, widely flattened, punctate; mesocoxae separate; abdominal sternites finely fluted along sutures, punctate, exposed disc of pygidium punctate. Profemur almost circular; meso- and metafemora narrow, posterior edge strongly margined, sinuate; protibia narrow, lateral teeth small; meso- and metatibiae flattened dorsoventrally, apical spurs very fine, located close together below tarsal insertion; tarsi short, segments subquadrate; claws hair-like. Phallobase of male aedeagus without dorsal hump, apical portion of internal sac narrow with minute spicules. Affinity. The closest relatives of Saprositellus known from South America belong to the genus Odontolochus Schmidt (Stebnicka and Howden 1996). Members of Odontolochus differ from those of Saprositellus by having the significantly broader head, the clypeal edge rounded without denticles and the pronotal anterior disc strongly tumid.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10C97543CB9FCB7610E47F9.taxon	description	(Figs. 1, 5)	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10C97543CB9FCB7610E47F9.taxon	description	Description. Length 2.8 – 3.0 mm, greatest width 1.0 mm. Body (Fig. 1) elongate oval, widest at apical third of elytra; color piceus, surface microreticulate, legs dark brown. Clypeus shallowly emarginate, lateral edge with 6 – 7 fine denticles on each side of emargination, surface just above shining and here minutely transversely granulate; remaining surface of head with fine punctures separated by less than one diameter, punctures near eyes forming short longitudinal lines; frontal suture marked by slightly impressed line; gena small, rounded. Pronotum convex, slightly converging posteriorly, side straight, finely margined, lateral tooth small, acutely prominent; base straight, finely crenate without marginal line; pronotal disc with shallow longitudinal furrow and small fovea laterally, surface punctures everywhere close, on disc separated by about one diameter, fine punctures along anterior margin increase in size toward base, those on sides medium-sized, closer. Elytra distinctly arcuate, base as wide as pronotal base, humeral denticles small; striae deep, slightly narrower than intervals with coarse punctures crenating inner margins of intervals, punctures separate each other by about 1 / 3 of their diameter; intervals subcarinate, microreliculate, minute punctures scattered. Ventral sclerites alutaceous; mesosternal surface and lateral area of metasternum distinctly punctate, punctures same size as those on pronotal disc; mestasternal midline shallow, discal punctures finer than those on sides, lateral metasternal triangle marked by very shallow concavity; abdominal sternites finely fluted along sutures, punctate from side to side, punctures separated by about one diameter, finer and closer on penultimate sternite and pygidium. Profemur wide, perimarginal groove deep, surface punctures same size as those of penultimate sternite; meso- and metafemora slender, posterior femoral lines complete, posterior edge slightly lobed medially (Fig. 5); meso- and metatibiae as long as femora, slightly sinuate, minutely and scarcely setigerous; metatibia with fine external spine and very thin apical spurs, tarsus short, segments quadrate; basal tarsomere of metatarsus markedly shorter than upper tibial spur and shorter than following two tarsomeres together. Affinity. The species is most similar to Saprositellus ariquemes n. sp. (see Affinity under that species).	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10C97543CB9FCB7610E47F9.taxon	materials_examined	Type Data. Holotype (sex not determined), labelled ‘‘ Brasilien, Amazonas, ’’ ‘‘ Saprositellus denticulatus m. Dr V. Balthasar, ’’ in BCP. Material Examined. Holotype (seen in 1973) and one specimen, labelled: PERU, Prov. Loreto, 160 km NE Iquitos, Explornapo Camp, 2 km from Rio Napo on Rio Sucusari, 27 – 31. VIII. 1992, black light trap, P. Skelley (FSCA).	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10F97523C5AFB1061454225.taxon	description	(Figs. 2, 6 – 7) Description. Length 2.6 – 2.8 mm, greatest width 1.0 mm. Form (Fig. 2) elongate oval, widest at apical third of elytra; color dark brown, legs reddish brown, surface microreticulate. Clypeus truncate anteriorly, edge on each side with 6 – 7 denticles, surface just above shining and here minutely transversely granulate; remaining surface of head with fine, very close, nearly contiguous punctures; frontal suture marked by slightly thickened line; gena small rounded. Pronotum strongly convex, side slightly arcuate, finely margined, lateral tooth acutely prominent, posterior angle excised, base sinuate, finely crenate with fine marginal line; pronotal disc with shallow longitudinal furrow and small fovea laterally, surface punctures everywhere very close, almost contiguous, rugose. Elytra distinctly arcuate, apex with minute setae visible under high magnification; base as wide as pronotal base, humeral denticle strong, directed upward; striae deep, slightly wider than intervals with deep, coarse punctures crenating inner margins of intervals, punctures separate each other by about 1 / 3 of their diameter; intervals carinate, microreticulate. 6) epipharynx; 7) male genitalia in lateral view; 8) S. peruanus n. sp., right posterior leg. Ventral sclerites alutaceous, surface punctate, punctures gradually decreasing in size from mesosternum to penultimate sternite, generally separated by about one diameter; prosternal process widely triangular, punctate; mesosternum lower than metasternum, mestasternal midline shallow, lateral metasternal triangle marked by very shallow concavity; abdominal sternites finely fluted along sutures, punctate from side to side; disc of pygidium with fine punctures same size as those on penultimate sternite. Profemur wide, perimarginal groove deep, surface punctures same size as those of penultimate sternite; meso- and metafemora narrow, slender, posterior femoral lines strong, complete, posterior edge slightly sinuate; meso- and metatibiae as long as femora, slightly sinuate, minutely and scarcely setigerous; metatibia with fine external spine and seta-like apical spurs, tarsus short, segments subquadrate; basal tarsomere of metatarsus two times shorter than upper tibial spur and equal in length to second tarsomere. Epipharyngeal structures as in Figure 6. In the male, the penultimate abdominal sternite is shorter than in the female, and the disc of pygidium longer; genitalia as in Figure 7. Affinity. The new species is most similar to Saprositellus denticulatus, but differs from that species by having the pronotal punctures denser and pronotal base margined, the elytral humeral denticles significantly larger and the elytral striae wider, with coarser punctures.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10F97523C5AFB1061454225.taxon	materials_examined	Type Material. Holotype male (FSCA), ‘‘ Brazil, Rondonia, 62 km SW Ariquemes nr Fzda Rancho Grande, 8 – 20. XI. 1994, black light, J. Eger & C. O’Brien. Paratype female, same data as holotype’ ’ (FSCA). Paratype male ‘‘ Bolivia, Santa Cruz, Buena Vista 380 m elev., 7.27 S, 63.39 W, 20. II. 1998, light trap, L. Stange’ ’ (ISEA).	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10F97523C5AFB1061454225.taxon	etymology	Etymology. Derived from the name of type locality ‘‘ Ariquemes. ’’	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997523C55FEBE60A94794.taxon	description	(Figs. 4, 8) Description of Male. Length 3.2 mm, greatest width 1.2 mm. Form (Fig. 4) elongate oval, distinctly widened at apical third of elytra; color piceous black, legs dark brown, surface microreticulate. Clypeus very shallowly emarginate anteriorly, edge on each side with 5 – 7 denticles; clypeal surface just above emargination shining and here minutely transversely granulate, punctures above wrinkles up to frontal suture very fine and shallow, slightly elongate, separated by about one diameter; vertex with band of round, close punctures separated by less than one diameter; frontal suture marked by slightly thickened line; gena small rounded. Pronotum strongly convex, diverging posteriorly, side straight toward acutely prominent lateral tooth, posterior angle excised, base margined, sinuate and lobed medially, edge finely crenate; pronotal surface with small fovea laterally, without longitudinal furrow, punctures rather shallow, everywhere close, separated by less than one diameter, fine punctures along anterior margin increase in size toward base, those on sides medium-sized, closer. Elytra distinctly arcuate, apex with minute setae visible under high magnification; base as wide as pronotal base, humeral denticle small, acute; striae moderately deep with moderate punctures crenating inner margins of intervals, punctures separate each other by about ½ diameter; intervals twice as wide as striae, flat on disc, convex laterally and apically, surface microreliculate, minute punctures scattered. Ventral sclerites alutaceous, surface punctate; punctures of mesosternum and metasternum same size as those of pronotal disc, separated by one diameter; disc of metasternum concave in posterior half, midline slightly impressed, lateral metasternal triangle marked by very shallow concavity; abdominal sternites arcuate, very finely fluted along sutures, punctate from side to side, punctures two times smaller than those on mesosternum, disc of pygidium with fine punctures same size as those on penultimate sternite. Profemur wide, perimarginal groove deep, surface shagreened, punctures same size as those of abdominal sternites but denser; meso- and metafemora slender, posterior femoral lines complete, posterior edge lobed at middle (Fig. 8); meso- and metatibiae as long as femora, slightly sinuate, punctate and minutely, scarcely setigerous; metatibia with fine external spine and seta-like apical spurs, tarsus short, segments subquadrate; basal tarsomere of metatarsus only slightly shorter than upper tibial spur and longer than second tarsomere. Female unknown. Affinity. The species is very distinct and may be easily distinguished from all other species in the genus by its size, trapezoid pronotum and wide, flattened elytral intervals.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997523C55FEBE60A94794.taxon	materials_examined	Type Material. Holotype male (CMN): ‘‘ Peru, Madre de Dios, Rio Tambopata Res. 30 mi SW Pto Maldonado, 290 m, 2 – 5. XI. 1979, subtropical moist forest, J. B. Heppner. ’’	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997523C55FEBE60A94794.taxon	etymology	Etymology. Derived from the name of terra typica.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997533C59FAF061254067.taxon	description	(Fig. 3) Description of Male. Length 2.9 mm, greatest width 1.0 mm. Form (Fig. 3) elongate, almost parallel-sided; color dark brown, legs reddish brown, surface microreticulate. Clypeus slightly emarginate anteriorly, edge on each side of emargination with 6 – 7 denticles; surface just above emargination shining and here minutely transversely granulate, punctures above granulate area up to frontal suture very fine and shallow, separated by about one diameter, vertex with band of round, rugose punctures; frontal suture marked by slightly thickened line; gena small rounded. Pronotum strongly convex, side slightly arcuate, finely margined, lateral tooth acutely prominent, posterior angle excised, base straight, finely crenate with distinct marginal line; pronotal disc with shallow longitudinal furrow and small fovea laterally, surface punctures everywhere very close, almost contiguous, rugose. Elytra parallel-sided, apex with minute setae visible under high magnification; base as wide as pronotal base, humeral denticle small, acute; striae deep, slightly wider than intervals with deep coarse punctures crenating inner margins of intervals, punctures separate each other by about 1 / 3 of their diameter; intervals carinate, microreliculate. Ventral sclerites alutaceous, surface punctate; punctures of mesosternum and metasternum equal in size, smaller than those of pronotal disc, generally separated by about one diameter; prosternal process widely triangular, punctate; mestasternal midline shallow, lateral metasternal triangle marked by very shallow concavity; abdominal sternites arcuate, minutely fluted along sutures, punctate from side to side, punctures two times smaller than those on mesosternum, disc of pygidium with fine punctures same size as those on penultimate sternite. Profemur wide, perimarginal groove deep, surface punctures same size as those of penultimate sternite; meso- and metafemora slender, posterior femoral lines strong, complete, posterior edge straight, unlobed; meso- and metatibiae as long as femora, slightly sinuate, minutely and scarcely setigerous; metatibia with fine external spine and seta-like apical spurs, tarsus very short, segments subquadrate; basal tarsomere of metatarsus slightly shorter than upper tibial spur and longer than second tarsomere. Female unknown. Affinity. Saprositellus santaritae is most closely related to S. ariquemes n. sp. sharing with that species a similar sculpture of the pronotum and elytra. It differs from ariquemes by having the less arcuate elytra with distinctly finer humeral denticle.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997533C59FAF061254067.taxon	materials_examined	Type Material. Holotype male (HAHC): ‘‘ Panama, Colón Prov., Santa Rita Ridge, 250 m, 9.23 N, 79.45 W, 10. VI. 1977, H. Stockwell. ’’	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10997533C59FAF061254067.taxon	etymology	Etymology. Derived from the name of type locality ‘‘ Santa Rita. ’’	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
03EA6A15B10B97513C41FD736353422A.taxon	description	Motschulsky (1865) described the genus Dolichodes, diagnosing it from other genera of platynine Carabidae in his key by the more or less depressed body form, rounded pronotal hind angles, and dorsal body surface with opaque, granulate microsculpture. He designated D. geniculatus as the type species, which he described as new in the same work, recording its provenance as Brazil. Dolichodes was listed as a junior synonym of Agonum Bonelli, 1810 in Gemminger and Harold (1868), and has remained a junior generic synonym to the present. Recent taxonomic studies on Agonum showed that many South American species formerly attributed this genus are, in fact, better classified in an endemic South American genus, Incagonum (Liebherr 1994). Removal of species to Incagonum from Agonum leaves five names of South American taxa still combined with Agonum, A. geniculatum (Motschulsky) being one. Comparison of the single known female syntype of Dolichodes geniculatus (Keleinikova 1976) with specimens of Euleptus ooderus Chaudoir, 1850 (J. Schmidt Collection), establishes D. geniculatus as a junior subjective synonym of E. ooderus, thereby making Dolichodes Motschulsky, 1865, a junior generic synonym of Euleptus Klug, 1833. Euleptus ooderus is known from the Himalaya, and the other described species of the genus reside in Africa and Madagascar. Therefore Motschulsky’s type locality of Brazil for D. geniculatus is certainly in error. The type locality is hereby corrected to NEPAL: Karnali Province, Gothichaur Valley, 2,800 m elev., 29812.19 N 82818.59 E, to reflect the locality of compared specimens in the Schmidt collection. Besides the microsculptural and habitus characters alluded to by Motschulsky in his diagnostic generic key, E. ooderus is characterized by the absence of a median mentum tooth (a character shared by all Euleptus spp.), and basally rufotestaceous femora distinctly contrasted with the infuscated femoral apex and tibiae. Taxonomic changes brought about by these findings are summarized as: Euleptus Klug, 1833, Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin, p. 131. Dolichodes Motschulsky, 1865 (‘‘ 1864 ’’), Bulletin de la Société Impériale des Naturalistes de Moscou 37 (4): 371 (NEW SYNONYMY). Euleptus ooderus Chaudoir, 1850, Bulletin de la Société Impériale des Naturalistes de Moscou 23 (2): 365. Dolichodes geniculatus Motschulsky, 1865 (‘‘ 1864 ’’): 321 (NEW SYNONYMY). Corrected type locality: Nepal, Karnali Province, Gothichaur Valley, 2,800 m elev. Dolichodes geuiculatus Motschulsky, 1865 (‘‘ 1864 ’’): 317 (unavailable incorrect original spelling). For D. geniculatus, lectotype hereby designated (in order to fix the identity of this species should additional specimens reflecting a mixed type series be discovered): Female specimen glued on white card, specimen extensively covered with fungal growth, labeled: [green paper isosceles triangle with indecipherable legend] // [green label] Dolichodes geniculatus Brazil Motsch. // [square red label] // [red label] Lectotype Dolichodes geniculatus Motschulsky, det. J. K. Liebherr 2003.	en	Stebnicka, Zdzisława Teresa (2003): The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini). The Coleopterists Bulletin 57 (4): 451-457, DOI: 10.1649/582, URL: http://dx.doi.org/10.1649/582
