taxonID	type	description	language	source
03EB87E51014FFF38E8CF934CCC90C41.taxon	discussion	The following redescription of the type-species, Laphria melanura Wiedemann, 1828, is provided in order to facilitate comparison, and discussion of generic and specific relationships.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51014FFF68E0EF7C9CD3D06A9.taxon	description	Figs 2, 5, 12 - 14	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51014FFF68E0EF7C9CD3D06A9.taxon	description	Redescription: Based on unique holotype specimen. Head: Dark red-brown and yellow areas as indicated. Antenna: Scape and pedicel brown-yellow, ftagellomeres both missing. All setae of scape and pedicel dark red-brown; scape with strong dark red-brown ventral bristle. Face (Fig. 5) width: maximum head width (measured anteriorly) ratio 4,7: 1. Upper five-eighths dark red-brown, lower three-eighths yellow; upper dark area with 2 strong brown bristles centraJly, near antennaI bases; lower paler area with cluster of long yellow bristles and setae, a few of which are slightly dorsoventrally flattened (those occurring laterally) and approaching the condition found in the females of Afrotropical species. Vertex and frons black with few black setae laterally. Ocellar bristles yellow-brown. Occipital area dark red-brown dorsally, yellow-brown ventrally; extensively gold pruinose; major dorsal bristles brown and black; all setae golden yellow. Proboscis yellow but red-brown ventrodistally. Pal pi pale yellow, 2 - segmented, equipped with yellow setae. Thorax: Brown-yellow, uniformly fine gold pruinose. Postpronotal lobes with yellow setae. Mesonotum with tiny black and yellow setae; acrostichal setae undifferentiated; dorsocentrals postsuturally situated only (best developed near hind margin); 3 - 4 notopleurals; 4 supra-alars; 3 postalars (all mesonotal bristles brown-yellow). Scutellum with small black (few yellow) setae on disc, 8 brownyellow marginal bristles. Anatergites (metanotal callosities) gold pruinose, lacking setae. Pleura fine gold pruinose; all setae and bristles brown-yellow. Hind and upper margins of anepisternum with brown-yellow bristles; katatergum with many long brown-yellow bristles. Postmetacoxal area membranous. Wing length 13,5 mm (from humeral crossvein to tip); venation as in Fig. 2; except for small areas of cells c, r l, r 2 + 3, br, bm, cup, a l and discal, wing covered with microtrichia; r 5 open; m 3 closed and stalked; r-m crossvein at about one-third the distance along anterior margin of discal cell; membrane yellow-stained, especially anterobas ally, entire wing tip (from approximately level of r-m crossvein) brown-stained. Legs brownyellow except for tips of hind femora which are dark red-brown and tarsi which are dark brown; setae mostly long yellow but mixed with black on pro- and mesothoracie tarsi and almost dark red-brown on hind tarsi; hind femur with ca 4 strongly developed bristles distally. Abdomen: Terga 1 - 4 brown; T 5 dark red-brown dorsocentrally, brown-yellow laterally; T 6 - 7 dark red-brown. 2 - 4 dis cal bristles on all terga; setae yellow on brown-yellow parts, black on dark red-brown areas. Sl- 5 yellow, S 6 - 8 dark red-brown with longish setae. Genital bulb yellowish, rotated through 90 °. Genitalia as illustrated in Figs 12 - 14; aedeagus with three short terminal filaments (a plesiomorphic condition found in all Laphriinae). Female: Not studied.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51014FFF68E0EF7C9CD3D06A9.taxon	materials_examined	Material examined: BRAZIL: 1 ♂ holotype, ' Brasilien', ' melanura Coli. Winthem', ' Type W. ' (NMW). This inconspicuously labelled specimen has been provided with a modern type label.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51011FFF78E35FA7DC8800DBC.taxon	description	Figs 1, 3, 6, 8 - 9, 15 - 21, 28	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51011FFF78E35FA7DC8800DBC.taxon	description	. Redescription: Based on holotype ♂ (Fig. 1). Head: Dark red-brown but colour largely masked by pruinescence. Antenna (Figs 8 - 9) dark brown: Setae of scape and pedicel yellow and dark red-brown mixed; scape with long yellow ventral bristle; flagellomere uniformly covered with microtrichia except for anterocentral depressed area containing more tightly packed microtrichia (this may be a sense organ); tip with narrow pit containing a single spine. Face width: maximum head width (measured anteriorly) ratio 4,9: 1 (Fig. 6). Face silver pruinose and uniformly covered with silvery glistening dorsoventrally flattened scale-like setae (longer in ventral half); normal setae confined to a row of shiny white setae along lower facial margin and a pair of long shiny white setae at about midway between antennal bases and lower facial margin. Vertex and frons uniform silver pruinose with tiny dark red-brown setae laterally along eye margins. Ocellarium prominent with a pair of long black ocellar bristles. Occiput red-gold pruinose centrally, silver pruinose adjacent to hind margins of eyes; upper occipital bristles black, directed vertically; other occipital setae mostly of moderate length, shiny silver-white. Proboscis and 2 - segmented palpi yellowbrown with fine silver-white setae. Thorax: Brown-yellow, uniformly fine silver pruinose. Postpronotal lobes with gold-yellow setae. Mesonotum with tiny gold-yellow and black setae; acrostichal setae undifferentiated; dorsocentrals weakly defined postsuturally; 1 strongish notopleural bristle accompanied by 2 - 3 longish black setae; 3 gold-yellow supraalars; 2 gold-yellow postalars accompanied by 1 long black seta. Scutellum with small black and gold-yellow setae on disc, 5 gold-yellow marginal bristles accompanied by shorter yellow and black setae. Anatergites with well-developed gold-yellow setae. Pleura fine silver pruinose; all setae and bristles gold-yellow. Hind margin of anepisternum with 3 well-developed gold-yellow bristles; katatergum with many long gold-yellow bristles. Postmetacoxal area membranous. Wing length 7,0 mm (from humeral crossvein to tip); venation as in Fig. 3; except for small narrow areas of cells br, bm, cup and a 1 wing covered with microtrichia; r 1 closed and with long stalk; r 5 open; m 3 closed and stalked; r-m crossvein at about one-third the distance along anterior margin of dis cal cell; membrane transparent. Legs brown-yellow; setae longish gold-yellow; hind femora without obvious distal bristles; pulvilli pale yellow, well developed. Abdomen: First four obvious terga (1 ' 2 - 5) brown with most setae short black, lateral ones gold-yellow; T 6 - 7 yellow, all setae yellow. Gold-yellow dis cal bristles present on all terga; all terga fine gold pruinose. Sterna yellow with gold-yellow setae. Genital bulb yellowish, rotated through 90 °. Genitalia (Nigerian ♂ 0 ' illustrated) as in Figs 15 - 17; aedeagus with three long terminal filaments. Female: Agrees well with ♂ except for following features: Antennae entirely missing. Shiny silver scale-like facial setae limited to 2 small laterally situated groups in lower third of face; most facial setae black, of moderate length; mystacal bristles present on lower facial margin as in ♂ but 4 major central ones broken off leaving sockets. Wing 7,0 mm long; cell r 5 of left wing closed at wing margin. Genitalia as in Figs 18 - 19; cercus with acute apex. Egg (dissected from body cavity) with shape as in Fig. 20; surface with fine pattern of hexagonal reticulation, tiny spine-like structures present at each intersection of reticulation (Fig. 21).	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51011FFF78E35FA7DC8800DBC.taxon	discussion	Variation: The Nigerian ♂ agrees in all important ways with the holotype. Minor differences are as follows: All but one scutellar bristle are black. Wing 5,5 mm long.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51011FFF78E35FA7DC8800DBC.taxon	materials_examined	Material examined: ZAIRE: 1 ♂ holotype, Congo,. Walikale, 1 ° 25 ' S: 28 ° E [1 ° 25 ' S: 28 ° 03 ' E], 7. i. 1915, J. Bequaert (MCZ Type 20137). CENTRAL AFRICAN REPUBLIC: 1 ♀, La Maboke (Rep. Centrafr.) [Maboke-stream 3 ° 54 ' N: 17 ° 53 ' E], 6. vii. 1964, J. Carayon rec. (MNP). NIGERIA: 1 ♂, W. Nigeria, Illaro Forest [lIaro Forest Reserve- 6 ° 47 ' N: 3 ° 04 ' E], 24. iii. 1974, M. A. Comes, 4075 (NMWC). Distribution as in Fig. 28. Other recorded material: Oldroyd (1970: 235) records: 1 ♀, ' W. Kivu, Lubongola, pro Shabunda, 1939, Dr Hautman' housed in KMT. I have not seen this specimen, illustrated (head and tip of tarsus) by Oldroyd, but am confident that it was correctly identified. The locality ' Lubongola' cannot be found in available gazetteers and two places by the name of ' Shabunda' have been found [1 ° 41 ' S: 28 ° 48 ' E & 2 ° 42 ' S: 27 ° 20 ' E]; the first listed has been used to plot this record on Fig. 28.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51013FFFA8E10F815CF0F075A.taxon	description	Figs 4, 7, 10 - 11, 22 - 27, 28	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51013FFFA8E10F815CF0F075A.taxon	etymology	Etymology: Named after Dr Stanley Bromley who was first to record the presence of this genus in Africa.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51013FFFA8E10F815CF0F075A.taxon	description	Description: Based on holotype ♂. Similar to S. pallida except for following details. Head: Antenna (Figs 10 - 11) brown-yellow but distal part of flagellomere darker brown; scape and pedicel with yellow setae only; flagellomere with oval-shaped anterior depressed area. Face fairly narrow, face width: maximum head width ratio 7,2: 1 (Fig. 7). Silvery scale-like mystacal setae narrower than in pallida but otherwise with similar appearance and distribution. Setae and bristles of vertex shiny yellow. Ocellarium with 1 black and 1 yellow bristle. Upper occipital bristles shiny yellow. Proboscis and palpi brown-yellow. Thorax: 1 strong yellow notopleural bristle accompanied by 2 black setae; 2 black supra-alars (1 yellow on left side); 1 - 2 black postalars; ca 8 black scutellar bristles; all scutellar setae black. Wing length 5,7 mm; venation as in ♀; distribution of microtrichia not as extensive as in pallida-cells c, r l, r 2 + 3, r 5 and discal also with small bare areas. Abdomen: All terga similar in colour and with short black setae dorsocentrally. Genitalia as in Figs 22 - 24, quite different from pallida; aedeagus with moderately long terminal filaments. Female: 1 ♀ paratype pinned together with holotype ♂ (captured whilst copulating). Similar to ♂. Antennal ftagellomere somewhat more attenuate. Distribution of scale-like mystacal setae as in ♀ pallida but normal setae all white. Ocellar bristles gold-yellow. All marginal bristles of mesonotum and scutellum yellowbrown except for 2 black ones on scutellar margin. Wing 6,8 mm long, venation as in Fig. 4. Genitalia as in Figs 25 - 26; cerci distally rounded. Egg (Fig. 27) (dissected from body cavity) similar to pallida but tiny spines somewhat smaller.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
03EB87E51013FFFA8E10F815CF0F075A.taxon	materials_examined	Material examined: NIGERIA: 1 ♂ holotype 1 ♀ paratype, Nigeria, Sapoba [6 ° 06 ' N: 5 ° 53 ' E], 17. iv. 1946, M. A. Cornes, Forest, 6622 (NMWC). Distribution as in Fig. 28.	en	Jason G. H. Londt (1989): Afrotropical Asilidae (Diptera) 18. The genus Smeryngolaphria Hermann, 1912 (Laphriinae). Annals of the Natal Museum 30: 147-157, DOI: 10.5281/zenodo.1472869
