taxonID	type	description	language	source
03E887D0FFC4FFBA1593FEEBD1A5FE29.taxon	diagnosis	Genus diagnosis. Moderate size, usually 1 − 3 mm; eyes 8 + 8; four segmented antennae; mucro bidentate and with a basal spine; dentes without spine; scales not narrow and present on body.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC4FFBF1593F99FD09AFE72.taxon	description	Figs 2 − 34, Table 1 Types. Holotype: adult female on slide, China, Gansu Province, Lanzhou City, Yuzhong County, Xinglongshan Mountain, 35 ° 47 ′ 57 ″ N, 104 ° 04 ′ 0 ″ E, altitude 1,900 m, 12, June, 2019, collection number: 1207, by Yitong Ma with entomological aspirator. Paratypes: Eight adult females on slides, same data as holotype. All deposited in NTU. Description. Body length up to 2.73 mm. Ground colour pale yellow or yellow in alcohol. Eye patches dark blue. Antennae with scattered blue pigment. Lateral margin of head with blue pigment and a “ V ” pattern blue stripe present on dorsal head. A pair of longitudinal blue stripes present on lateral margin of body from head to Abd. IV and a third longitudinal blue stripe along midline line from Abd. II to Abd. IV. Coxae, trochanters, femora and ventral side of body with scattered blue pigment (Figs 2 − 5). Scales. Scales spinulate type (Fig. 6), present on Ant. I−II (Fig. 7), head (Fig. 8), terga, legs, ventral tube and ventral side of manubrium. Basal part of ventral side of dens also with scales (Fig. 9). Head. Antennae 0.66 − 0.80 times body length; antennal segments ratio as I: II: III: IV = 1: 1.54 − 1.79: 1.23 − 1.57: 1.87 − 2.57. Distal part of Ant. IV with many sensory chaetae, normal ciliate chaetae and a bilobed apical bulb (Fig. 12). Distal part of Ant. III with 3 − 4 rods (Fig. 13). Distal part of Ant. II with 2 − 3 rods (Fig. 14). Dorsal cephalic chaetotaxy as 8 − 12 antennal (An), four median (M) and nine sutural (S) mac. Ps 2 present. Interocular chaetae as p, s, t mes. Eyes 8 + 8, G and H smaller (Figs 8, 15). Prelabral and labral chaetae as 4 / 5, 5, 4, prelabral chaetae ciliate, other smooth, four labral papillae with one minute denticle each (Fig. 16). Lateral process (l. p.) of labial palp slightly curved, with tip not reaching apex of labial papilla (Fig. 17). Labial triangular chaetae usually as M 1 M 2 REL 1 L 2, all ciliate and R 0.67 − 0.77 length of M 1, rarely R duplicated (Fig. 18). Subapical chaeta of maxillary outer lobe thin, subequal to apical one; three smooth hairs on sublobal plate (Fig. 19). Thorax. Th. II with three (m 1, m 2, m 2 i) mac on T 1 area, seven (a 5, m 4, m 4 i, m 4 p, m 5 and two unknown) mac on T 2 area, 25 − 27 posterior mac, one ms and two se on each side. Th. III with 24 − 27, two se on each side (Fig. 20). Pseudopores on coxa I−III as 2, 3 − 4 (rarely 6), 1, respectively (Figs 21 − 23). Trochanteral organ with 51 − 96 smooth spiny chaetae (Fig. 24). Unguis with four inner teeth, one pair located 0.50 − 0.52 from base of inner edge of unguis, distal unpaired two respectively at 0.75 − 0.76 and 0.88 − 0.90 distance from base. Unguiculus acuminate and outer edge serrate. Tenent hair thick with its tip clavate, longer than inner side of unguis (Figs 10, 25). Abdomen. Abd. IV 4.55 − 6.50 times as long as Abd. III along dorsal midline. Abd. I with seven (m 2, m 2 i, m 3, m 4, a 2, a 3, a 5) mac, one ms and one se, ms not inner to se. Abd. II with two (a 2, a 3) mac on A 1 area, four (m 3, m 3 e, m 3 ea, m 3 ep) mac on A 2 area, one (m 5) lateral mac and two se. Abd. III with two (a 2, a 3) mac on A 4 area, one (m 3) mac on A 5 area, five (am 6, pm 6, p 6, m 7, p 7) lateral mac, one ms and two se (Fig. 26). Abd. IV with 16 − 20 central and 18 − 22 lateral mac (Fig. 27). Abd. V with three se (Figs 11, 28). Ventral tube anteriorly with sacles and 14 − 16 ciliate chaetae on each side, three of them much larger than others (Fig. 29); posteriorly with two smooth chaetae on apical part (Fig. 30); laterally with six smooth and 17 ciliate chaetae (Fig. 31). Manubrial plate dorsally with 11 − 22 ciliate chaetae and 3 − 4 pseudopores (Fig. 32); ventrally with 34 − 43 ciliate chaetae on each side (Fig. 33). Uncrenulated dens 1.07 − 1.45 times as mucro in length. Mucro bidentate, two teeth subequal and tip of basal spine reaching subapical tooth (Fig. 34).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC4FFBF1593F99FD09AFE72.taxon	biology_ecology	Ecology. Found in rotten leaf, mainly composed of Pinus tabuliformis, Quercus liaotungensis and Cotoneaster zabelii.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC4FFBF1593F99FD09AFE72.taxon	etymology	Etymology. Named after its three bands on dorsal body.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC4FFBF1593F99FD09AFE72.taxon	discussion	Remarks. The new species can be distinguished from other known Willowsia species because of its three longitudinal blue stripes on dorsal body. It is similar to W. flava Zhou, Pan & Ma, 2021 (in: Zhou et al. 2021), W. samarkandica Martynova, 1972 and W. variabilis Ma, 2016 in macrochaetotaxy of body. However, there are some differences between them, such as, colour pattern, sculpture and distribution of scales, macrochaetotaxy of head and Th. II−III, apical bulb of Ant. IV and mucro (Table 1).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC3FFBD1593FA2ED67DF9EF.taxon	diagnosis	Genus diagnosis. Moderate size, usually 1 − 2 mm; eyes 8 + 8; four segmented antennae; mucro bidentate and with a basal spine; dentes without spine; chaetae on body long, flattened and leaf-like.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC3FFB21593F975D2D9F8EC.taxon	description	Figs 35 − 59, Table 2 Types. Holotype: adult female on slide, China, Gansu Province, Lanzhou City, Yuzhong County, Xinglongshan Mountain, 35 ° 48 ′ 11 ″ N, 104 ° 03 ′ 56 ″ E, altitude 1,900 m, 12, June, 2019, collection number: 1205, by Yitong Ma with entomological aspirator. Paratypes: Seven adult females on slides, same data as holotype; One adult female on slide, China, Gansu Province, Tianshui City, Qinzhou District, Nanguo Temple, Dengbaoshan Memorial Hall, 34 ° 34 ′ 01 ″ N, 105 ° 44 ′ 36 ″ E, altitude 1,288 m, 13, June, 2019, collection number: 1210, by Yitong Ma with entomological aspirator. All deposited in NTU. Description. Body length up to 1.42 mm. Ground colour pale yellow in ethanol. Distal part of Ant. I−II, Ant. III−IV blackish violet. Eye patches dark blue. Anterior and lateral parts of dorsal head with blue pigment. Blue pigment also present on each tergite and coxa (Figs 35 − 36). Body chaetae long and leaf-like (Fig. 37). Antenna 0.41 − 0.45 times body length; antennal segment ratio as I: II: III: IV = 1: 1.70 − 2.22: 1.60 − 2.00: 2.07 − 2.85. Apical bulb of Ant. IV bilobed (Fig. 41). Distal part of Ant. III with three rods (Fig. 42). Distal part of Ant. II with two rods (Fig. 43). Eyes 8 + 8, G and H smaller than others, interocular chaetae as p, s, t mes (Figs 38, 44). Dorsal cephalic chaetotaxy with four antennal (An), four median (M) and eight sutural (S) mac (Fig. 44). Prelabral and labral chaetae as 4 / 5, 5, 4, prelabral chaetae ciliate, other smooth, four labral papillae with one minute denticle each (Fig. 45). Lateral process (l. p.) of labial papilla E differentiated, thicker than normal chaeta with tip reaching apex of papilla E (Fig. 46). Labial triangular chaetae as MREL 1 L 2, all ciliate and R 0.50 − 0.55 length of M (Fig. 47). Subapical chaeta of maxillary outer lobe thin, subequal to apical one; three smooth hairs on sublobal plate (Fig. 48). Th. II with four (m 1, m 2, m 2 i, m 2 i 2) mac on T 1 area, five (a 5, m 4, m 4 i, m 4 p, m 5) mac on T 2 area, 24 − 30 posterior mac, one ms and two se on each side. Th. III with 32 − 36 mac and two se (Fig. 49). Trochanteral organ with 11 − 13 smooth spiny chaetae (Fig. 50). Unguis with four inner teeth, one pair located 0.57 − 0.59 from base of inner edge of unguis, distal unpaired two respectively at 0.71 − 0.76 and 0.83 − 0.91 distance from base. Unguiculus acuminate and outer edge serrate. Tenent hair thick with its tip clavate, longer than inner side of unguis (Figs 39, 51). Abd. IV 2.50 − 4.28 times Abd. III in dorsal axial length. Abd. I with nine (a 2, a 3, a 5, m 2 − 4, m 2 i, m 4 i, m 4 p), ms antero-external to se. Abd. II with two (a 2, a 3) mac on A 1 arae, four (m 3, m 3 e, m 3 ea, m 3 ep) mac on A 2 area, one (m 5) lateral mac and two se. Abd. III with two (a 2, a 3) mac on A 4 area, one (m 3) mac on A 5 area and three (am 6, pm 6, p 6) lateral mac, one ms and two se (Fig. 52). Abd. IV with seven central mac, 10 lateral mac (Fig. 53). Abd. V with three se (Fig. 54). Anterior face of ventral tube with 11 − 14 ciliate chaetae, line connecting proximal (Pr) and external-distal (Ed) mac oblique to median furrow (Fig. 55); posterior face not clearly seen; lateral flap with 2 − 3 smooth chaetae and 7 − 8 ciliate chaetae (Fig. 56). Manubrial plate dorsally with 6 − 7 ciliate chaetae and three pseudopores (Fig. 57); ventrally with 10 − 11 ciliate chaetae on each side (Fig. 58). Uncrenulated dens 2.43 − 3.21 times as mucro in length. Mucro bidentate, two teeth subequal and tip of basal spine reaching subapical tooth (Figs 40, 59).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC3FFB21593F975D2D9F8EC.taxon	biology_ecology	Ecology. Found in rotten leaf, mainly composed of Pinus tabuliformis, Quercus liaotungensis and Cotoneaster zabelii.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC3FFB21593F975D2D9F8EC.taxon	etymology	Etymology. Named after its locality: Gansu Province.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFC3FFB21593F975D2D9F8EC.taxon	discussion	Remarks. The new species is the second Himalanura species reported from China and can be distinguished from the first Chinese species of the genus, H. shanxiensis Zhou & Ma, 2020 in colour pattern, body macrochaetotaxy and other characters. It is similar to these species in macrochaetotaxy of Abd. III: H. baijali Baquero, Mandal & Jordana, 2014, H. bulunkuli Tshelnokov, 1977, H. gelmani Tshelnokov, 1987, H. himachalensis Baquero & Jordana, 2015 (in: Baquero et al, 2015), H. martynovae Tshelnokov, 1977 and H. tundricola Tshelnokov, 1987, but can be separated from them by apical bulb on antenna, macrochaetotaxy of dorsal head, Th. II−III, Abd I−II and IV (Table 2).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFCFFFB11593F9E4D020F954.taxon	diagnosis	Genus diagnosis. Moderate size, usually 1 − 2 mm; eyes 8 + 8; four segmented antennae; mucro bidentate and with a basal spine, the subapical tooth much larger than the apical one; dentes with spine; scales absent on body; macrochaetae on the anterior part of Abd. IV.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFCFFFAA1593F8CCD327F90E.taxon	description	Figs 60 − 85, Table 3 Types. Holotype: adult female on slide, China, Gansu Province, Tianshui City, Maiji District, Xiaolongshan National Forest Park, Taohuagou Scenic Spot, 34 ° 18 ′ 52 ″ N, 106 ° 34 ′ 17 ″ E, altitude 1,208 m, 17, June, 2019, collection number: 1213, by Yitong MA with entomological aspirator. Paratypes: Three adult females on slides, same data as holotype. All deposited in NTU. Description. Body length up to 2.20 mm. Ground colour pale yellowgreen in ethanol. Eye patches dark blue. Distal part of Ant. III, Ant. IV and tibiotarsi blue pigmented. Coxae and lateral part of Abd. IV with scattered pigment. Posterior part of Abd. IV and V with an irregular transverse stripes, respectively (Figs 60 − 61). Antenna 0.67 − 0.68 times body length; antennal segment ratio as I: II: III: IV = 1: 1.41 − 1.75: 1.18 − 1.83: 1.82 − 2.92. Apical bulb of Ant. IV bilobed (Fig. 62). Distal part of Ant. III with three rods (Fig. 63). Distal part of Ant. II with two rods (Fig. 64). Ant. I with three dorsal, four ventral basal smooth spiny chaetae (Fig. 65). Eyes 8 + 8, G and H smaller than others, interocular chaetae as p, s, t mes. Dorsal cephalic chaetotaxy with four antennal (An), five median (an additional mac between M 2 and M 3) and eight sutural (S) mac (Fig. 66). Prelabral and labral chaetae as 4 / 5, 5, 4, prelabral chaetae ciliate, other smooth, four labral papillae with one minute denticle each (Fig. 67). Lateral process (l. p.) of labial papilla E differentiated, as thick as normal chaeta with tip not reaching apex of papilla E (Fig. 68). Chaetal formula of labial base as MREL 1 L 2, all ciliate, E rarely smooth, R / M as 0.55 − 0.73 (Fig. 69). Subapical chaeta of maxillary outer lobe thin, subequal to apical one; three smooth hairs on sublobal plate (Fig. 70). Th. II with four (m 1, m 2, m 2 i, m 2 i 2) mac on T 1 area, five (a 5, m 4, m 4 i, m 4 p, m 5) mac on T 2 area, 29 − 35 posterior mac, one ms and two se. Th. III with 40 − 43 mac and two se (Fig. 71). Pseudopores on coxa I−III as 1, 2 − 4, 1, respectively; coxal macrochaetal formula as 3 / 4 + 1, 3 / 4 + 2 (Figs 72 − 74). Trochanteral organ with 49 − 66 smooth spiny chaetae (Fig. 75). Tenent hair thick with its tip clavate, almost equal to than inner edge of unguis in length. Unguis with four inner teeth, basal pair located 0.47 − 0.51 distance from base of inner edge of unguis, distal unpaired two respectively at 0.70 − 0.72 and 0.84 − 0.85 distance from base. Unguiculus lanceolate, outer edge slightly serrate (Fig. 76). Abd. IV 5.38 − 6.10 times Abd. III in dorsal axial length. Abd. I with 11 (a 1, a 1 a, a 2, a 3, a 5, m 2 − 4, m 2 i, m 4 i, m 4 p), ms antero-external to se. Abd. II with two (a 2, a 3) mac on A 1 area, four (m 3, m 3 e, m 3 ea, m 3 ep) mac on A 2 area, one (m 5) lateral mac and two se. Abd. III with one (a 2) mac on A 4 area, one (m 3) mac on A 5 area and four (am 6, pm 6, m 7 a, p 6) lateral mac, one ms and two se (Fig. 77). Abd. IV anterior with 6 − 9 mac arranged in irregular transverse row, posterior with 6 − 7 (A 5 − 6, B 5 − 6, Ae 7, Ae 5 pp) central mac, lateral with 19 − 21 mac (Fig. 78). Abd. V with three se (Fig. 79). Anterior face of ventral tube with 35 − 36 ciliate chaetae, line connecting proximal (Pr) and external-distal (Ed) mac oblique to median furrow (Fig. 80); posterior face with four distal smooth and numerous ciliate chaetae (Fig. 81); lateral flap not seen wholly. Manubrial plate dorsally with 9 − 13 ciliate chaetae and three pseudopores (Fig. 82); ventrally with 38 ciliate chaetae on each side (Fig. 83). Dens with 58 − 80 smooth inner spines (Fig. 84). Mucro bidentate with subapical tooth larger than apical one; tip of basal spine slightly exceeding apex of subapical tooth; distal smooth of dens shorter than mucro in length (Fig. 85).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFCFFFAA1593F8CCD327F90E.taxon	biology_ecology	Ecology. Found in rotten leaf, mainly composed of Pinus tabuliformis, Betula Platyphylla and Quercus variabilis.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFCFFFAA1593F8CCD327F90E.taxon	etymology	Etymology. Named after its locality: Maiji District.	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
03E887D0FFCFFFAA1593F8CCD327F90E.taxon	discussion	Remarks. The new species is characterised for its yellowgreen ground colour, labial base chaetae and dental spines. It is similar to H. bilineata Lee & Park, 1984, H. phjongjangica Szeptycki, 1973 and H. rosannae Jordana & Baquero, 2010 in colour pattern, but there are some differences between them, such as, labial base chaetae, macrochaetae on Abd. I and dental spines. It is also similar to H. apigmenta Shi, Pan & Zhang, 2010, H. similis Szeptycki, 1973, H. subcingula Denis, 1948, and H. unichaeta Pan, Shi & Zhang, 2010 in macrochaetotaxy of Abd. II−IV, but can be separated from them by labral papillae, inner teeth on unguis, macrochaetotaxy on Abd. IV and other characters (Table 3).	en	Zhou, Rong, Ma, Yitong (2022): Three new species of Entomobryinae (Collembola, Entomobryidae) from Gansu China. Zootaxa 5155 (3): 371-392, DOI: 10.11646/zootaxa.5155.3.4
