taxonID	type	description	language	source
03E88D171F6CFFD4FDDCFCD22817F814.taxon	type_taxon	type species: Birdantis decens Stål, 1863 by subsequent designation in Distant 1906: 26).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6CFFD4FDDCFCD22817F814.taxon	discussion	Remarks Lallemand (1959) synonymized Myrilla under Birdantis and proposed a key to the species of Birdantis. However, Fennah (1977) considered Myrilla as a subgenus of Birdantis, separating them on characters of the vertex, i. e., anterior margin of vertex straight, without deep transverse sulcus behind it in Myrilla; anterior margin of vertex concave, with a deep transverse sulcus behind it in Birdantis s. str. It is therefore currently formally divided into two subgenera: Birdantis (ten species and one subspecies) and Myrilla (two species) (Fennah 1977; Nagai & Porion 1996). The genus is externally close to the Oriental Polydictya Guérin-Méneville, 1844 and Gebenna Stål, 1863, and the Australian Desudaboides Musgrave, 1927 (Lallemand 1963; Nagai & Porion 1996; Constant 2010, 2011). Nagai & Porion (1996) followed the classification in Fennah (1977), but in my paper on the species of Birdantis from Australia (Constant 2011), I questioned the value of the subgenera as defined by Fennah (1977), because intermediate species exist, and followed the views of Lallemand (1959) but without formally reinstating the synonymy between Birdantis and Myrilla. Accordingly, three species remained without subgeneric affiliation (Bourgoin 2018) within the last formally accepted classification of Fennah (1977). The genus Birdantis is currently placed in the Poiocerina Haupt, 1929 of the Poiocerini Haupt, 1929, in the subfamily Poiocerinae Haupt, 1929 (Lallemand 1963; Constant 2011; Bourgoin 2018). However, Urban & Cryan (2009) have shown, based on molecular data, that Fulgoridae can be separated into two main clades, one grouping all New World taxa with strong support, the other containing all Old World taxa. Hence, as the type genus of the Poiocerinae is the Neotropical genus Poiocera Laporte, 1832, the suprageneric placement of Birdantis needs to be reconsidered. The classifications proposed by Metcalf (1947), Lallemand (1963) and Nagai & Porion (1996) were based on a very small number of characters of the head, especially the presence / absence and shape of the cephalic process. The genus Birdantis is here transferred to the subfamily Aphaeninae, following the conclusions of the DNA study by Urban & Cryan (2009), which place the genus close to Desudaba Walker, 1858. However, the latter study did not contain any species of the genera Polydictya, Gebenna or Desudaboides, which are putatively considered closely related to Birdantis based on morphological characters. Hence, an accurate tribal placement requires a complete study of the suprageneric relations between the genera of Aphaeninae with the inclusion of more Old World taxa in the molecular analysis and the integration of morphological data (Urban & Cryan 2009).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	description	urn: lsid: zoobank. org: act: 3424 BD 5 B- 34 C 2 - 4516 - A 0 CB- 9415 F 485 F 537 Figs 1 – 3	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	diagnosis	Diagnosis This species can be separated from all other species of Birdantis by the following combination of characters: (1) frons entirely yellow-brown without lines or spots (Fig. 1 D) (2) bulge between frons and vertex coloured as vertex (Fig. 1 B) (3) abdomen mostly orange dorsally (Fig. 1 A) and ventrally (Fig. 1 C) (4) tegmina opaque on basal half and infuscate on distal half (Fig. 1 A) (5) hind wings largely smoky, but not hyaline or with a large black area (Fig. 1 A)	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	etymology	Etymology This species is dedicated to Mr Edy Bhaskara (Indonesia) in acknowledgment of his generous contribution to the present work.	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	materials_examined	Material examined Holotype INDONESIA: ♂ (dissected, Figs 1 – 2), Larat Island, 7 ° 09 ′ 17 ″ S, 131 ° 54 ′ 48 ″ E [Coll. I. R. Sc. N. B., Indonesia, Tanimbar islands, Larat Island, xii. 2016, Gift from E. Bhaskara, I. G.: 33.453] (RBINS). Paratypes INDONESIA: 1 ♂, 4 ♀♀, same collection data as for holotype (RBINS). Other material INDONESIA: 2 ♀♀, same collection data as for holotype (EBC).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	description	Description MEASUREMENTS AND RATIOS. LT: ♂ (n = 2): 19.0 mm (18.9 – 19.2); ♀ (n = 6): 21 mm (20.4 – 21.2). LTg / BTg = 3.45; BV / LV = 4.45; LF / BF = 0.83. HEAD (Fig. 1 B, D, F). Pale yellow-brown with small black marking at posterior angles of vertex, a larger black marking behind eyes, median portion of clypeus darker and more reddish, with a yellowish central line, labium pale yellow-brown turning to brown on two apical segments and antennae brown. Vertex with deep transverse groove and all margins carinate, shorter in middle than on sides. Frons finely wrinkled, with two smooth longitudinal carinae slightly diverging towards dorsum and a slight longitudinal groove on each side between carina and lateral margin; broadest near base, above clypeus; convex in dorsal view and with dorsal margin rounded in perpendicular view; bulge between frons and vertex only visible dorsally. Ocelli under eyes. Antennae with scape short and cylindrical, and pedicel inflated and reniform. Clypeus narrower than frons, reaching apex of procoxae. Labium with penultimate segment surpassing hind coxae (Fig. 1 C). THORAX (Fig. 1 B, D, F). Pronotum pale yellow-brown, with median carina and impressed point on each side of latter; small back-brown spot on each side of disc, short black-brown line on median carina on anterior half and brown marking behind eye. Mesonotum dark brown, with median and peridiscal carinae well marked, concolorous; wrinkled in the area limited by peridiscal carinae; mesothoracic sternites pale yellow-brown. Metathoracic sternites dark brown. Tegulae yellow-brown. TEGMINA (Fig. 1 A, C). Elongate, broader at nodal line and acutely rounded apically. Corium and clavus with numerous cross-veins, brown variegated with darker patches and a blackish line along external side of vein Pc + CP; veins and cross-veins yellow. Membrane smoky, roundly extending inside corium medially and with brown spots on distal part and brown markings on nodal line; veins yellow turning to black-brown after nodal line. HIND WINGS (Fig. 1 A, C). Light smoky brown with veins black-brown, darker at basal angle and with anal area with a grey hue and cross-veins whitish; maximal breadth near base; slightly broader than tegmina. LEGS (Fig. 1 A, C). Elongate and slender. Pale yellow-brown with apex of pro- and mesofemora black distally; pro- and mesotarsi black; metatibiae brown apically, with six lateral spines and six apical spines, all spines dark brown apically; metatarsi with first tarsomere yellow-brown, brown apically and with nine apical spines ventrally, second and third tarsomeres dark brown, the second one with eight apical spines ventrally. Metatibiotarsal formula: (6) 6 / 9 / 8. Male genitalia Pygofer higher than long, about 1.4 times as high as maximum length, longer ventrally and abruptly narrowing on dorsal ¼ in lateral view; dorsal margin of pygofer oblique in lateral view (Fig. 2 A). Anal tube elongate, about 2.1 times as long as maximum breadth, curved ventrally near base; lateral margins subparallel, only slightly sinuate and with narrowest portion at basal ¼; apical margin strongly emarginate in dorsal view, acutely rounded in lateral view (Fig. 2 A – B). Gonostyli subtriangular in lateral view, with posterior margin rounded; anterodorsal margin emarginate after lateral process; lateral process laminate, directed laterally and apically pointed, with apical point directed lateroventrally (Fig. 2 A – B). Aedeagus mostly membranous; phallobase with two ventral, elongate processes; aedeagus s. str. strongly reduced but endosoma well developed with six membranous digitiform processes, with posteroventral left process with secondary process directed centrally (Fig. 2 C – F).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F6AFFDFFDD4FEA92CD1FABE.taxon	distribution	Distribution Larat Island (Fig. 3).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F66FFDDFD84FAE12977FCD8.taxon	description	Figs 3 – 4	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F66FFDDFD84FAE12977FCD8.taxon	discussion	Note The specimens examined by Lallemand (1959, 1963) and Fennah (1977), erroneously identified as B. collaris, were actually specimens of B. trilineata (Schmidt, 1926).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F66FFDDFD84FAE12977FCD8.taxon	diagnosis	Diagnosis This species can be separated from all other species of Birdantis by the following combination of characters: (1) frons entirely yellow-brown, without lines or spots (Fig. 4 E) (2) bulge between frons and vertex black (Fig. 4 C) (3) abdomen mostly black dorsally (Fig. 4 A) and brown ventrally (Fig. 4 B) (4) tegmina opaque on basal half and infuscate on distal half (Fig. 4 A) (5) hind wings largely hyaline, with basal angle yellowish (Fig. 4 A)	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F66FFDDFD84FAE12977FCD8.taxon	materials_examined	Material examined Holotype INDONESIA: ♀ (Fig. 4), Morotai Island, 2 ° 20 ′ 52 ″ N, 128 ° 27 ′ 52 ″ E [Morty, Wallace / glued on the reverse: 67 66] [M] [Type] [collaris] [BMNH (E), # 651887] (BMNH).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F66FFDDFD84FAE12977FCD8.taxon	distribution	Distribution Morotai Island (holotype) and Halmahera Island (Breddin 1900) (Fig. 3).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F64FFDDFDD3FC122CC1F86C.taxon	description	Figs 3, 5	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F64FFDDFDD3FC122CC1F86C.taxon	diagnosis	Diagnosis This species can be separated from all other species of Birdantis by the following combination of characters: (1) frons yellowish, with five longitudinal black lines not reaching ventral margin of frons, the three central ones merging together dorsally in a transverse line and central one dilated in middle (Fig. 5 F) (2) bulge between frons and vertex coloured as vertex (Fig. 5 C) (3) abdomen mostly black dorsally (Fig. 5 A) and brown ventrally (Fig. 5 B) (4) tegmina opaque on basal half and hyaline on distal half (Fig. 5 A) (5) hind wings largely hyaline, with basal angle red (Fig. 5 A)	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F64FFDDFDD3FC122CC1F86C.taxon	materials_examined	Material examined Holotype INDONESIA: ♂ (Fig. 5), Aru Island, 6 ° 07 ′ 14 ″ S, 134 ° 30 ′ 00 ″ E [Aru Isl / on the reverse: 58 48] [Birdantis] [Type] [Birdantis decens Stål] [BMNH (E), # 651892] (BMNH).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F64FFDDFDD3FC122CC1F86C.taxon	distribution	Distribution Aru Island (Fig. 3).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFDAFDCBFEA92CABF9F0.taxon	description	Figs 3, 6	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFDAFDCBFEA92CABF9F0.taxon	discussion	Note The specimen mentioned by Kirkaldy (1913) as well as those illustrated by Fennah (1977) and Nagai & Porion (1996), erroneously identified as B. delibuta, were actually specimens of B. trilineata (Schmidt, 1926).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFDAFDCBFEA92CABF9F0.taxon	diagnosis	Diagnosis This species can be separated from all other species of Birdantis by the following combination of characters: (1) frons yellow-brown with three dark brown lines limited dorsally to level of middle of eyes, and with the two lateral ones extending on to clypeus (Fig. 6 B) (2) bulge between frons and vertex dark brown and vertex yellow-brown (Fig. 6 C) (3) tegmina opaque on basal half and infuscate on distal half (Fig. 6 A) (4) hind wings largely smoky, with a basal red marking (Fig. 6 A) (5) anterior margin of frons in dorsal view rounded but not strongly protruding (Fig. 6 C)	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFDAFDCBFEA92CABF9F0.taxon	materials_examined	Material examined Holotype INDONESIA: abdomen lost (Fig. 6), Ternate Island, 0 ° 48 ′ 07 ″ N, 127 ° 20 ′ 41 ″ E [Ternate] [Stevens] [234, 59] [Birdantis delibuta Stål] [Riksmuseum Stockholm] [NHRS-HEMI 000000181] (NHRS).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFDAFDCBFEA92CABF9F0.taxon	distribution	Distribution Ternate Island (Fig. 3).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFC6FDAEF92B2F9DFC11.taxon	description	Figs 3, 7 – 9	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFC6FDAEF92B2F9DFC11.taxon	diagnosis	Diagnosis This species can be separated from all other species of Birdantis by the following combination of characters: (1) frons yellow-brown, with three black-brown lines limited dorsally to level of middle of eyes, and with the two lateral ones extending on clypeus (Figs 7 D, 8 D) (2) bulge between frons and vertex dark brown and vertex yellow-brown (Figs 7 B, 8 B) (3) tegmina opaque on basal half and infuscate on distal half (Figs 7 A, 8 A) (4) hind wings largely smoky, with a basal reddish brown marking (Figs 7 A, 8 A) (5) abdomen mostly dark red dorsally (Figs 7 A, 8 A) and yellow-brown ventrally (Figs 7 C, 8 C) (6) anterior margin of frons in dorsal view rounded and strongly protruding (Figs 7 B, 8 B)	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFC6FDAEF92B2F9DFC11.taxon	materials_examined	Material examined Lectotype INDONESIA: ♀ (designated to improve nomenclatural stability in the group, Fig. 7), Buru Island, 3 ° 23 ′ 55 ″ S, 126 ° 38 ′ 54 ″ E [Buru 1921, Station: 1 leg. L. J. Toxopeus] [E. Schmidt] [Typus] [Myrilla trilineata Schmidt ♀, Edm. Schmidt determ. 1926.] [* trilineata Schmidt 1926] [Mus. Zool. Polonicum Warszawa Typus n. 1211 Myrilla trilineata Schmidt, 1926 Syntypus] [Mus. Zool. Polonicum Warszawa 12 / 45] (ZMPA). Paralectotype INDONESIA: ♀, Buru Island, same collection data as for lectotype (ZMPA). Other material INDONESIA: 1 ♀, Boano Island, 2 ° 59 ′ S, 127 ° 55 ′ E, Aug. 2013 [I. G.: 32.613] (RBINS); 1 ♂, W Seram Island, Waipia, 2 ° 58 ′ 30 ″ S, 129 ° 09 ′ 40 ″ E, 600 – 800 m a. s. l., Jun. 2013 [I. G.: 32.613] (RBINS); 1 spec. (abdomen lost; “ vittiventris ” Walker, in litt.), Ambon Island, 3 ° 38 ′ S, 128 ° 07 ′ E, Wallace leg. (BMNH).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFC6FDAEF92B2F9DFC11.taxon	description	Supplementary description Male genitalia Pygofer higher than long, about 2.0 times as high as maximum length, longer ventrally and abruptly narrowing on dorsal ½ in lateral view; dorsal margin of pygofer horizontal in lateral view (Fig. 9 A). Anal tube elongate, about 1.5 times as long as maximum breadth, nearly straight with ventral margin sligtly curved in lateral view; progressively broadening from base towards ¾ of length in dorsal view, then with lateral margins converging; apical margin strongly, roundly emarginate in dorsal view, acutely rounded in lateral view (Fig. 9 A – B). Gonostyli slightly elongate in lateral view, with ventral margin slightly concave and posterior margin broadly rounded; anterodorsal margin slightly emarginate after lateral process; lateral process laminate, curved laterally and apically pointed, with apical point directed lateroventrally (Fig. 9 A – B). Aedeagus mostly membranous (Fig. 9 C – F).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
03E88D171F63FFC6FDAEF92B2F9DFC11.taxon	distribution	Distribution Buru, Seram, Ambon and Boano Islands (Fig. 3).	en	Constant, Jérôme (2018): Review of the Maluku Islands species of the lanternfly genus Birdantis Stål, 1863, with a new species and identification key (Hemiptera: Fulgoromorpha: Fulgoridae). European Journal of Taxonomy 461: 1-20, DOI: 10.5852/ejt.2018.461
