taxonID	type	description	language	source
03E88D48FF96FFED92F4FEE38BBBFCE9.taxon	materials_examined	Material examined. Host species Rhynchium haemorrhoidale (Vespidae: Eumeninae) for all following reared specimens: April, E 1 U 2, 1, 6; April, E 2 U 3, 1, 8; April, F 4 I 4, 1, 18; April, E 1 U, 1, 19; April, F 1 C 3, 1, 41; April, F 4 I 1, 1, 47; April, F 4 I 4, 1, 97; April, E 1 U 8, 1, 98; April, E 1 U 4, 2, 105; April, E 3 I 2, 1, 137; April, F 4 I 3, 1, 179; April, F 1 U 6, 1, 196; April, F 4 U 2, 1, 222; April, E 1 I 4, 1, 235; April, D 2 U 3, 1, 284; April, F 1 U 7, 1, 349; April, F 4 U 2, 1, 197; Dec, E 2 I 4, 2, 256; Dec, F 2 C 2, 1, 133; Dec, F 4 U 1, 1, 173; Dec, D 3 I 3, 1, 179; Dec, D 4 U 1, 1, 230; Dec, D 2 C 3, 1, 407; Dec, E 2 I 4, 1, 479; Dec, F 4 U 5, 1, 10; Dec, F 4 U 3, 1, 200; Feb, F 1 U 6, 1, 128; Feb, F 1 U 6, 1, 147; Feb, E 1 U 6, 1, 359; Feb, E 3 I 3, 1, 403; Feb, E 4 U 8, 1, 487; Feb, E 3 U 4, 1, 801; Feb, E 1 U 7, 1, 839; Feb, F 4 U 2, 1, 856; Feb, F 2 U 8, 1, 859; Feb, F 1 U 2, 1, 71; Jan, E 1 U 1, 1, 59; Jan, D 5 U 7, 1, 76; Jan, D 4 U 7, 1, 326; Jan, D 4 U 5, 1, 345; Jan, E 3 U 4, 1, 381; Jan, F 4 U 7, 1, 481; Jan, F 4 U 5, 1, 504; Jan, D 4 U 4, 1, 561; Jan, E 1 U 5, 1, 584; Jan, E 2 C 1, 1, 612; Jan, D 4 U 4, 1, 656; Jan, E 1 I 4, 1, 674; Jan, F 4 U 7, 1, 731; July, F 4 I 4, 1, 65; July, F 1 U 8, 1, 71; July, F 1 U 6, 1, 82; July, F 1 C 3, 1, 89; July, E 3 U 6, 1, 119; July, E 3 U 6, 2, 120; July, D 3 C 1, 1, 194; Jun, D 2 U 7, 1, 93; Jun, F 1 U 4, 1, 115; Jun, D 3 U 6, 1, 129; Jun, F 1 U 2, 1, 148; Jun, F 1 U 4, 1, 164; Jun, F 1 C 2, 1, 172; Jun, F 4 C 2, 1, 269; Jun, F 4 C 1, 1, 278; Jun, F 4 U 3, 1, 281; Jun, F 4 I 2, 1, 296; May, F 4 C 1, 2, 4; May, F 4 I 4, 1, 33; May, D 4 U 6, 1, 35; May, E 1 U 1, 3, 59; May, D 2 U 6, 1, 99; May, D 2 U 7, 1, 116; May, E 2 U 5, 1, 133; May, D 2 U 7, 1, 136; March, E 2 I 4, 1, 3; March, E 2 U 3, 1, 64; March, E 2 U 7, 1, 66; March, D 2 U 2, 1, 128; March, E 4 I 1, 1, 217; March, E 4 I 1, 1, 230; March, E 4 U 1, 1, 237; March, E 3 U 4, 1, 245; March, E 3 U 7, 1, 255; March, E 3 U 7, 1, 257; March, E 3 U 8, 1, 258; March, E 3 U 8, 1, 265; March, E 3 U 7, 1, 280; March, E 3 U 7, 1, 281; March, E 1 U 5, 2, 288; March, E 1 U 8, 1, 293; March, E 1 U 8, 1, 294; March, E 1 U 8, 1, 295; March, E 1 U 8, 1, 301; March, E 1 U 8, 1, 302; March, F 1 U 7, 1, 381; March, F 1 U 6, 1, 396; March, F 4 C 3, 1, 420; March, F 4 U 3, 1, 440; March, F 4 U 3, 2, 450; March, D 4 U 7, 1, 527; March, E 3 U 8, 1, 612; March, F 2 U 7, 1, 630; March, E 3 U 8, 1, 656; March, F 4 I 3, 1, 679; March, F 1 U 6, 1, 696; March, F 4 I 3, 2, 708; March, F 3 U 8, 1, 724; March, F 4 U 5, 1, 765; March, F 4 U 5, 1, 769; March, F 3 U 6, 1, 779; March, D 5 U 4, 1, 789; Nov, D 5 C 3, 1, 148; Nov, E 4 U 1, 1, 316; Nov, F 3 U 2, 1, 405; Nov, F 3 U 1, 1, 483; Nov, D 3 C 2, 1, 488; Nov, F 3 U 2, 1, 503; Nov, F 3 U 3, 1, 585; Nov, D 3 C 2, 1, NA; Nov, D 3 I 3, 1, NA; Nov, F 4 U 6, 1, 346; Oct, D 2 C 1, 1, 6; Oct, D 2 C 1, 1, 7; Oct, D 2 C 1, 1, 9; Oct, D 2 C 1, 1, 12; Oct, D 2 C 1, 2, 13; Oct, E 1 U 1, 1, 37; Oct, E 1 U 1, 1, 43; Oct, E 1 I 4, 1, 46; Oct, E 2 U 6, 1, 71; Oct, E 2 U 1, 1, 79; Oct, E 1 I 4, 1, NA; Sept, D 3 U 6, 1, 28; Sept, E 4 U 6, 1, 125; Sept, E 4 U 3, 1, 139; Sept, E 1 I 4, 1, NA (JBCP, PHCG, IPBC; host species PHCG). Parasitism rate was 1.97 % (7,397 individuals (nests) of this eumenid were examined, 146 of which were infested by M. nasuta). Host species Epsilon manifestatum crassipunctatum (Vespidae: Eumeninae): April, D 3 U 3, 1, 136; March, F 3 U 5, 1, 164 (PHCG, IPBC; host species PHCG). Observations. Infested nests of the most abundant host Rhynchium haemorrhoidale had on average 1.93 ± 0.076 breeding cells, 1.07 ± 0.024 of which were parasitized by M. nasuta. Two subspecies of R. haemorrhoidale (R. h. haemorrhoidale and R. h. umeroater Gusenleitner, 1991) were present together in the research area, and it is not possible to identify the host precisely because no adult wasps were present in the infested nests. Additionally, three nests of R. haemorrhoidale were infested each with three individuals of the parasitoid Chrysis (angolensis group) sp., three with Chrysis (smaragdula group) sp. 1 and one with Chrysis (smaragdula group) sp. 2 (Hymenoptera: Chrysididae) and one individual from the family Trigonalidae (Hymenoptera). Macrosiagon nasuta further infested two nests of Epsilon manifestatum crassipunctatum (Gusenleitner, 1991) with one individual each. Macrosiagon nasuta made small holes in the partition walls to get out of the nest, with the dead host larvae remaining in the infested cell. Macrosiagon nasuta usually hatched earlier than the remaining host individuals, perhaps to avoid contact with R. haemorrhoidale in the nest. Nevertheless, we did not observe aggressive behaviour of the host against the parasitoid when both met in the glass tube. In the field we observed M. nasuta many times on the inflorescences of the invasive plant species Hyptis capitata (Jacq.) (Lamiaceae). This native of Central America is common in disturbed areas. Interestingly, H. capitata seems to be highly attractive to R. haemorrhoidale and E. manifestatum crassipunctatum due to the high nectar supply of the plant. Popularity of this plant among eumenid wasps has been observed e. g. by KELLER & ARMBRUSTER (1989) in Panama, Central America. IWATA (1975; fide ITINO 1986, p. 192) assumes that the ‘ ripiphorid beetle, Macrosiagon nasutum [sic!] Thunberg may probably lay many eggs in flowers ... and ... that the hatched beetle larvae are transported into the wasp nest attaching themselves to the host wasp when the wasp visits on flowers’. Thus it is likely that the triungulin larvae of the parasitoid and the host make contact in the inflorescences. However, as the plant is an invasive species, its presence might alter the interaction between the two insects and favour higher parasitism rate of the parasitoid.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF96FFED92F4FEE38BBBFCE9.taxon	discussion	Remarks. Macrosiagon nasuta was first recorded from Sulawesi by a single specimen three years ago (BATELKA 2004). This study provides confirmation of its occurrence on the island. Rhynchium haemorrhoidale had previously been known as a host species of M. nasuta only in Japan (see Conclusions). Epsilon manifestatum crassipunctatum is a new host record for this ripiphorid.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF96FFED92F4FEE38BBBFCE9.taxon	distribution	Distribution. Macrosiagon nasuta is widely distributed in Southeast Asia: China, Indonesia (Borneo, Sumatra, Sulawesi), Japan, Korean Peninsula, Nicobar Islands, Philippines, and Taiwan (BATELKA 2004).	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF91FFEC9212FCCB89EFFDEE.taxon	materials_examined	Material examined. Host species Chalybion sp. (Sphecidae: Sceliphronini): Jun, E 2 U 5, 1, 78 (JBCP; host species PHCG). A total of 827 cells of Chalybion sp. were examined. Host species Pison sp. (Crabronidae: Trypoxylonini): Feb, F 4 U 5, 2, 94 (JBCP, PHCG; host species PHCG). Observations. We found all three individuals of Macrosiagon cf. punctulaticeps (Blackburn, 1899) inside the host’s cocoon. In five breeding cells of one Pison sp. nest, we found two cells occupied by the parasitoid. Chalybion sp. usually has one breeding cell per nest; hence we found one parasitoid in one nest and no surviving host.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF91FFEC9212FCCB89EFFDEE.taxon	discussion	Remarks. The species is recorded from Sulawesi for the first time. No previous host associations for M. punctulaticeps have been made. The species belongs to the Macrosiagon ‘ bifasciata ’ species-group sensu BATELKA (2004). In this group, however, two ‘ subgroups’ can be identified. The first one is characterized by a shortened and robust metatarsomere 2, and includes two species in the Oriental Region, M. bipunctata (Fabricius, 1801) and M. dohertyi Pic, 1949. The former species is widely distributed in the Afrotropical Region; it has been recorded from Yemen and it is known from all over the Indian subcontinent. The latter species, described from Bali, is known only from the type specimen deposited in coll. M. Pic (MNHN). The type was examined to prove its possible conspecificity with the reared specimens; it is a male labeled: ‘ Bali [hw] / Doherty [p] // type [hw] // type [p – red label] // Dohertyi / mihi [hw] ’. However, M. dohertyi is not similar to the reared species. Moreover, no difference between M. bipunctata and M. dohertyi were observed; further research may show that these species are identical. The second ‘ subgroup’ includes all other Palaearctic and Oriental species related to the ‘ bifasciata ’ group, i. e. M. bifasciata (Marseul, 1876), M. medana Pic, 1910, M. medvedevi Iablokoff-Khnzorian, 1973, M. meridionale (Costa, 1859), and M. raffrayi (Fairmaire, 1873). Surprisingly, the second Macrosiagon species reared in this study is identical with a species occurring in Australia and the Solomon Islands. This species differs from other Oriental and Palaearctic members of this group by the strongly acute elytra and poorly visible light markings on a dull black background. We examined the following specimens of this species deposited in BMNH: four females labeled ‘ Australia: N. Q. / Redlynch, 21. x. 1938 / R. G. & J. Ch. Wind. / B. M. 1939 – 572 [p] ’ and one female labeled: ‘ Solomon Is. / Guadalcanal / Gold Ridge / 1 - 2000 ft / P. G. Fenemore / C. I. E. Coll. 16348 [p] / 21. ix. 1958 [hw] // Solomon Is. / Pres. / P. J. M. Greenslade / B. M. 1966 – 477 [p] // Macrosiagon / sp. nr. punctulaticeps / Blackb. [hw] / E. A. J. Duffy det. 1959 [p] ’. For the purpose of this paper we accept Duffy’s determination. However, examination of the type of M. punctulaticeps is needed. In any case, it is the first record of an Australian Macrosiagon species in the Oriental Region.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF90FFEC9203FDCB8E2DFC42.taxon	materials_examined	Material examined. Host species Auplopus sp. (Pompilidae: Auplopodini): July, F 3 U 4, 1, 10; Dec, E 2 U 8, 1, 424 (PHCG, IPBC; host species PHCG). Observations. We found one individual of M. pusilla in each of two nests of Auplopus sp., with on average four cells per nest.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF90FFEC9203FDCB8E2DFC42.taxon	discussion	Remarks. Auplopus sp. is a new host record for this species. The family Pompilidae is recorded as a host group for the genus Macrosiagon for the first time.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
03E88D48FF90FFEC9203FDCB8E2DFC42.taxon	distribution	Distribution. Macrosiagon pusilla is widely distributed in the Palaearctic and Oriental Regions: China (Fujian, Guandong, Hebei, Hunan, Sichuan, Xizang, Yunnan), India (Assam, ‘ Bengale: Barway’, Meghalaya, Uttaranchal), Indonesia (Sumatra), Japan (Hokkaidō, Honshū, Shikoku), Korean Peninsula (including Cheju Do Island), Laos, Malaysia, Nepal, Russia (Far East), Thailand, and Vietnam (BATELKA 2007). This represents the first record from Sulawesi.	en	Batelka, Jan, Hoehn, Patrick (2007): Report on the host associations of the genus Macrosiagon (Coleoptera: Ripiphoridae) in Sulawesi (Indonesia). Acta Entomologica Musei Nationalis Pragae 47: 143-152, DOI: 10.5281/zenodo.5328187
