identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E8277D7E50FF7BFDB5AC26FBBDF9E7.text	03E8277D7E50FF7BFDB5AC26FBBDF9E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra Roth 1995	<div><p>Genus Africalolampra Roth, 1995</p><p>Type species</p><p>Africalolampra ehrmanni Roth, 1995 .</p><p>Diagnosis and history</p><p>The genus Africalolampra was erected to accommodate the new species Africalolampra ehrmanni Roth, 1995 . Later, Anisyutkin (2016) transferred two species of Rhabdoblatta to this genus: A. erubescens and A. punctipennis . According to Roth (1995), the following combination of characters is diagnostic of the genus Africalolampra: weak sexual dimorphism (tegmina and wings completely developed in both sexes – although this is not true for A. punctipennis), metatarsus with 2 rows of spines along lower margin, tarsal claws distinctly serrated, first abdominal tergite of male with medial specialization (this is not true either for A. punctipennis), left stylus absent. This combination of characters is also found in two species from continental Africa belonging to the genus Rhabdoblatta, and we thus propose the following new combinations: Africalolampra camerunensis and Africalolampra stipata . We also reinstate Rhabdoblatta punctipennis because it does not share the apomorphies of Africalolampra . Note that R. lyncea (Gerstaecker, 1883) and R. pallida (Borg, 1902) also have a tergal specialization but on the 5 th and at the junction of the 8–9 th abdominal tergites, respectively. Rhabdoblatta lyncea may also have slightly serrated claws (Kumar 1975) but two styli, whereas R. pallida has a single stylus. Finally, Malagasy species are reported but males of some species have not been observed, so we do not dwell further on their generic affiliation, which would require further examination.</p><p>Below is the updated diagnosis of the genus Africalolampra, slightly modified from Roth (1995) and Anisyutkin (2016):</p><p>Weak sexual dimorphism, with tegmina and wings fully developed in both sexes. Anteroventral margin of front femur type B2; genicular spine absent from fore femur, present on mid and hind femora; spines present on antero- and posteroventral margins of all femora; pulvilli on four proximal tarsomeres; ventral margin of first tarsal segments with two rows of spines; tarsal claws symmetrical and serrated, arolia well-developed. Male with glandular specialization on first abdominal tergite; subgenital plate weakly asymmetrical, with only a single, small, right style (i.e., left style absent).</p></div>	https://treatment.plazi.org/id/03E8277D7E50FF7BFDB5AC26FBBDF9E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E50FF71FE28A843FDC5F9A1.text	03E8277D7E50FF71FE28A843FDC5F9A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra camerunensis (Borg 1902) Mbia & Ngon & Legendre & Menbohan 2024	<div><p>Africalolampra camerunensis (Borg, 1902) comb. nov.</p><p>Figs 3–7; Table 2</p><p>Epilampra camerunensis Borg, 1902: 8 .</p><p>Epilampra camerunensis – Shelford 1909b: 479; 1910: 14. — Anonymous 1910: 579. — Sjöstedt 1933: 7. — Princis 1963a: 136.</p><p>Rhabdoblatta camerunensis – Kumar 1975: 81.</p><p>Material examined</p><p>Lectotype</p><p>CAMEROON • 1 ♀; “Camerun; Sjöstedt; lectotype ”; NHRS, NRM-BLAT 0002100 [pictures only].</p><p>Additional specimens</p><p>CAMEROON – Centre Region • 2 ♀♀, 5 ♂♂, 9 juvs; Mefou watershed, Eloundem; elev. 569– 775 m; Dec. 2019 – Nov. 2020; University of Yaoundé 1 BLA 0001 • 1 ♀; Nkolkoumou; “Adam 350”; MNHNEP7587. – Eastern Region • 1 ♀, 2 ♂♂, 6 juvs; Haut-Nyong watershed, Abong-doum; elev. 670 m; Dec. 2019 – Nov. 2020; University of Yaoundé 1 BLA 0002 • 2 ♂♂, 2 ♀♀ nymphs; same data as for preceding; molecular sample BL946; MNHN-EP7596 to MNHN-EP7599 . – South Region • 1 ♀, 3 ♂♂, 11 juvs; Mvilla watershed, Nonenlam; elev. 579–581 m; Dec. 2019 – Nov. 2020; University of Yaoundé 1 BLA 0003 • 2 ♂♂; same data as for preceding; MNHN-EP7588, MNHN-EP7589 . – Unknown region • 1 ♀; “Camerun; Sjöstedt”; NHRS, NRM-BLAT 0002101 [pictures only] • 1 ♀; “Camerun, Bonge; Sjöstedt”; NHRS, NRM-BLAT 0002102 [pictures only] • 1 ♀; “Deutsch Kamerun, Mundame, Rhode”; NMW 25.718 [pictures only] .</p><p>REPUBLIC OF GUINEA • 1 ♂; “Nimba; 1946; Lamotte”; MNHN-EP7586 • 2 nymphs; “Nimba, brousse secondaire, Br2; 1946; Lamotte”; MNHN • 1 ♂; “Nimba; ??? Septembre, 1969 1493; Lamotte”; LUHM .</p><p>Differential diagnosis</p><p>Contrary to what was written in previous descriptions or keys (e.g., Kumar 1975), A. camerunensis has specialized claws, which contributes to justify its belonging to the genus Africalolampra . This species can be readily differentiated from its congenerics by its pattern of coloration (pronotum, wing, face) and the shape of its tergal gland.</p><p>Redescription</p><p>Male</p><p>Princis (1963a) was the first to describe a male specimen of A. camerunensis . He underlined that it is smaller than the conspecific female and has a hypandrium quite similar to that of A. erubescens, with a single stylus, although its right side is more advanced posteriorly than in A. erubescens . He also described the tergal specialization of the first tergite: triangular and nearly transversely oriented, with tuft of hairs oriented anteriorly on both sides (whereas triangular and vertically oriented in A. erubescens). This description can be complemented as follows:</p><p>HEAD. About as long as wide, dark brown with several black maculae (Fig. 4B). Eyes dark brown, ocellar spots brown with one dark spot below each of them. Labrum and clypeus brown, but lighter than vertex. Scape and pedicel mostly light brown, while other antennal segments dark brown. Distance between eyes similar to distance between antennal sockets.</p><p>PRONOTUM. Light brown with numerous black maculae and spots, and stripes at posterior margin (Fig. 4A). Pronotum roughly triangular but with middle of hind margin distinctly convex.</p><p>TEGMINA AND WINGS. Fully developed, extending beyond end of abdomen (Fig. 4A). Tegmina with rounded apex, light brown with veins darker and numerous dark brown maculae. Sc thickened in ventral view. Hindwings light brown, darker anteriorly than posteriorly, with veins darker (Fig. 4H).</p><p>LEGS. Anteroventral margin of front femur of type B2, with 4–5 strong spines proximally (Fig. 4B). Tibial spines well-developped. Metatarsus of hind leg slightly longer than other segments combined, inner margin with two rows of small spines (Fig. 4F). Arolium and tarsal pulvilli well-developed, the latter bordered by “additional spines” sensu Anisyutkin (2016). Claws symmetrical and finely serrated (Fig. 4G).</p><p>ABDOMEN. Abdominal sternites light brown with several darker maculae (Fig. 4A, C), largest ones near anterolateral corners, and with short stripes along posterior margin (except for last segments). First abdominal tergite specialized: tergal gland with a triangular, longitudinally keeled elevation, with tip directed anteriorly and provided with a posteriorly-oriented tuft of hair; both sides of triangle are also more or less densely covered with hairs (Fig. 4D). Second abdominal tergite with an anterodorsal ridge mesally (Fig. 4D). Supra-anal plate short, transverse and with wrinkles (Fig. 4E); caudal margin concave without medial incision. Subgenital plate asymmetrical, angularly projected on its right side. Cerci multisegmented, pubescent ventrally, with a black tip. Right stylus cylindrical, with sparse setae distally. Left stylus absent (Fig. 4C).</p><p>MALE GENITALIA (Fig. 5). Left phallomere R complex (L1 of MK64 and R70, and R2+R3+N of G96): cleft between R1T and R2 without protuberance; R3 (sensu Anisyutkin 2016) in two parts, very broad; pl.s. sensu Anisyutkin (2016) present. Sclerite L2d shaped as a woodsman’s axe; basal sclerite of L2d slender and rod-shaped (L1 of G96); apical sclerite of L2d (L2vm of MK64, L2v of R70 and L1 of G96) well-developed. Sclerite L3 (R2 of MK64, R70, and L2d of PG96) long, with a hook deeply bent, narrower where it bends, more bulbous at its apex with a distinct “small tooth” sensu Anisyutkin (2016); inner margin of the basal part of L3 crenulated on most of its length (i.e., folded structure f.s. of Anisyutkin 2016). L4U present, trigonal with a very slender vertex (likely homologous to trigonal sclerite of Roth 1995).</p><p>Female The female of A. camerunensis was first described in Borg (1902). It looks like the male, except for the terminalia (Fig. 6C): supra-anal plate extending beyond subgenital plate, with a deep medial incision and with setae on its caudal margin; subgenital plate symmetrical, with several small setae postero-medially and a large black macula medially.</p><p>Measurements of adults</p><p>Male (n = 3): pronotum length ×width: 5.3–6.1 ×7.5–8.7, tegmina length: 21.5–25.4, overall length ×width 23.3–26× 9.1–10.</p><p>Female: pronotum length ×width: 6.8×9.3, tegmina length: 29, overall length ×width: 30 × 11.8.</p><p>Nymphs</p><p>Pictures of nymph are provided in Fig. 7. Spiracular breathing tubes are visible dorsally, posteriorly to the penultimate tergite.</p><p>Molecular data</p><p>A portion of 12S rRNA has been obtained and is available under the GenBank accession number OR589772 (molecular extract BL946).</p><p>Habitat</p><p>Adults and nymphs of this species were found in all the rivers investigated.</p><p>Distribution</p><p>This species is known from Cameroon, Equatorial Guinea and the Republic of Guinea (Fig. 1). Details of localities are provided in Table 2.</p></div>	https://treatment.plazi.org/id/03E8277D7E50FF71FE28A843FDC5F9A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E5AFF6EFE0BA805FDC4FE4C.text	03E8277D7E5AFF6EFE0BA805FDC4FE4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra stipata (Walker 1868) Mbia & Ngon & Legendre & Menbohan 2024	<div><p>Africalolampra stipata (Walker, 1868) comb. nov.</p><p>Figs 8–9; Table 3</p><p>Epilampra stipata Walker, 1868: 208 .</p><p>Epilampra stipata – Kirby 1904: 123. — Shelford 1910: 14. — Princis 1958: 63; 1963a: 138. Rhabdoblatta stipata – Weidner 1969: 101. — Kumar 1975: 80.</p><p>Material examined</p><p>Holotype</p><p>REPUBLIC OF GUINEA • ♀; “ holotype; Epilampra stipata Walker ”; BMNH, # BMNH (E)878073 [pictures only].</p><p>Additional specimens</p><p>REPUBLIC OF GUINEA • 1 ♂; “Nimba (Guinea), 9 Db”; MNHN-EP7593 • 2 ♂♂; “Nimba (Guinée), Yalanzou; II VI 42; M. Lamotte ”; MNHN-EP7594, MNHN-EP7595 • 2 ♀♀; same data as for preceding; MNHN-EP7611, MNHN-EP7612 • 1 ♂, largely damaged; “Nimba (Guinée), Nion; II.VI.42, 1969 1491; M. Lamotte ”; LUHM .</p><p>Differential diagnosis and history</p><p>This species was described initially by Walker (1868), from what has been initially thought to be a male specimen but later identified as a female (Princis 1958). Princis (1963a) complemented this description and described the male for the first time, with a drawing of the male subgenital plate. He notably underlined the strongly specialized tarsal claws and described the tergal gland located on the first tergite, two characters that are, however, found in several species of Africalolampra . This species differs nonetheless from the congeneric species by the very strong intensity of the specialization of the tarsal claws (males and females), the shape of the subgenital plate (asymmetric, concave on the right side but convex on the left side), and the shape of the male tergal gland (i.e., a pit with two transverse fold-like elevations that meet mesally – Fig. 8D). In addition, we here illustrate its male genitalia (Fig. 9). For females, Kumar (1975) underlined that A. stipata can be distinguished from other species by the following combination of characters: wings fully developed, absence of a median brown longitudinal band on the subgenital plate, its size, and the strongly serrated claws.</p><p>Description of male genitalia (Fig. 9)</p><p>Left phallomere R complex (L1 of MK64 and R70, and R2+R3+N of G96): cleft between R1T and R2 with large protuberance at its base. Sclerite L2d with its basal sclerite rod-shaped, broader at its base (L1 of G96); apical sclerite of L2d (L2vm of MK64, L2v of R70 and L1 of G96) in two parts, with a rounded sclerite in the hollow of the crescent-like sclerite (visible in caudal view). Sclerite L3 (R2 of MK64, R70, and L2d of PG96) long, with a slender hook deeply bent and a membranous outgrouth (m.o.); inner margin of the basal part of L3 slightly crenulated along most of its length. L4U large, broadly trapezoidal.</p><p>Measurements of adults</p><p>Male (n = 3): pronotum length ×width: 3.6–4.1 ×4.9–5.3, tegmina length: 15.8–16.5, overall length ×width: 13.1–15.3× 5.4–5.9.</p><p>Female (n = 2): pronotum length ×width: 4.1–4.4 ×5.1–5.3, tegmina length: 18.2–18.9, overall length ×width: 16.3–16.5× 6.4–6.6</p><p>Habitat</p><p>This species has been reported as a water-dwelling cockroach (Weidner 1969).</p><p>Distribution</p><p>This species is known from Sierra Leone, Liberia, Ghana and the Republic of Guinea (Fig. 1). Details of localities are provided in Table 3.</p></div>	https://treatment.plazi.org/id/03E8277D7E5AFF6EFE0BA805FDC4FE4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E45FF6EFDC3AC3EFC71FAE9.text	03E8277D7E45FF6EFDC3AC3EFC71FAE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra ehrmanni Roth 1995	<div><p>Africalolampra ehrmanni Roth, 1995</p><p>Fig. 10</p><p>Africalolampra ehrmanni Roth, 1995: 89 .</p><p>Material examined</p><p>Holotype</p><p>KENYA • ♂; Shima la Tewa, ca 25 km N of Mombasa; 13 Apr.–4 Jul. 1987; R. Ehrmann leg.; MCZENT 00034788 [pictures only].</p><p>Differential diagnosis</p><p>Type species of the genus Africalolampra, it is described with several drawings from five males and one female from the same locality (Roth 1995). Note, howewer, that we were unable to examine the series of specimens and that the MCZ curators reported six male and no female specimens. We here complement the drawings in Roth (1995) with a few pictures obtained from MCZ (Fig. 10). The pronotal and head pattern coloration (Fig. 10A–B), as well as the shape of the tergal specialization, allows one to readily differentiate A. ehrmanni from its conspecifics.</p><p>Habitat</p><p>“The original specimens were collected in dense undergrowth in a damp Galeria woods” (Roth 1995).</p><p>Distribution</p><p>This species is only known from its type locality in Kenya (Fig. 1).</p></div>	https://treatment.plazi.org/id/03E8277D7E45FF6EFDC3AC3EFC71FAE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E45FF6BFE05AB52FC4DFDDF.text	03E8277D7E45FF6BFE05AB52FC4DFDDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra erubescens (Gerstaecker 1883)	<div><p>Africalolampra erubescens (Gerstaecker, 1883)</p><p>Figs 11–13; Table 4</p><p>Epilampra erubescens Gerstaecker, 1883: 54 .</p><p>Epilampra erubescens – Borg 1902: 8. — Shelford 1908: 29; 1909b: 479; 1910: 14. — Rehn 1933: 451. — Morales Agacino 1954: 95. — Princis 1962: 210.</p><p>Heterolampra erubescens – Kirby 1904: 123.</p><p>Rhabdoblatta erubescens – Vanschuytbroeck 1970: 14. — Kumar, 1975: 81. — Grandcolas 1996b: 231.</p><p>Africalolampra erubescens – Anisyutkin 2016: 140.</p><p>Material examined</p><p>Holotype</p><p>CAMEROON • ♂; “Victoria; Buchholz leg.; holotype ”; ZIMG-II-27340a [pictures only].</p><p>Allotype</p><p>CAMEROON • ♀; “Victoria Buchh., Limbar. Buchh.; allotype ”; ZIMG-II-27340b [pictures only]. Note that this female allotype has labels with two different localities (Victoria in Cameroon and Limbareni in Gabon, Fig. 11; see also Rehn (1933) for localities reported by Gerstaecker).</p><p>Paratype</p><p>CAMEROON • 1 ♂; “ paratype Victoria Buchh.”; ZIMG-II-27340c [pictures only] .</p><p>Additional specimens</p><p>CAMEROON • 1 ♂; “Cam. Oberth”; MNHN-EP7613 • 1 ♂; “Camerun; Sjöstedt”; NHRS, NRM-BLAT 0002106 [pictures only] • 1 ♂; “Camerun, Itoki; 31/1 91, 31.I.1891; Sjöstedt”; NHRS, NRM-BLAT 0002107 [pictures only] .</p><p>GABON • 1 ♂; “Gaboun; Coll. Br. v. W.; Standinger; 21.299”; LUHM .</p><p>NIGERIA • 1 ♀; “Biafra, Escalena”; NHRS, NRM-BLAT 0002108 [pictures only] .</p><p>Differential diagnosis and history</p><p>The male and female were described from specimens collected in Victoria, Cameroon, and in Lambarene (= Limbareni), Gabon (Gerstaecker 1883), and largely redescribed from specimens collected in Bitje, Cameroon (Rehn 1933), with drawings of the head and male subgenital plate. Later, Princis (1962) and Anisyutkin (2016) complemented those descriptions, including male and female genitalia as well as the shape of the tergal specialization on the 1 st tergite. Described in the genus Epilampra Burmeister, 1838, it was first transferred to the genus Rhabdoblatta (Princis 1967) and then to the genus Africalolampra (Anisyutkin 2016) . We complement previous descriptions with pictures of the habitus and diagnostic characters of the male – including genitalia – and female (Figs 11–13). Note that in one of the two male specimens dissected, a slightly sclerified sclerite has been observed, ventrally to the apical part of L3; it could be homologous of the ‘trigonal’ sclerite sensu Roth (1995), but it deserves further examination to be ascertained.</p><p>As reported in Kumar’s key (1975), the shape of the tergal specialization (Fig. 11H) is the main diagnostic character for this species, especially by comparison with A. camerunensis . Note that, like A. camerunensis and contrary to Kumar’s key, A. erubescens does have serrated claws (Anisyutkin 2016; Fig. 11I).</p><p>Measurements of adults</p><p>Male (n = 2): pronotum length ×width: 5.1–5.4 ×6.7–7.2, tegmina length: 20.1–21.4, overall length ×width: 21.8–25.6 × 8.6–8.9.</p><p>Habitat</p><p>Unknown, but this species is believed to be relatively frequent in West Africa (Rehn 1933).</p><p>Distribution</p><p>This species is known from Cameroon, Gabon, Togo, Ghana, Nigeria, Central African Republic and Congo Republic (Fig. 1). Details of localities are provided in Table 4.</p></div>	https://treatment.plazi.org/id/03E8277D7E45FF6BFE05AB52FC4DFDDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E40FF6BFF47ACAAFAD6FBC9.text	03E8277D7E40FF6BFF47ACAAFAD6FBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africalolampra Roth 1995	<div><p>Key for males in the genus Africalolampra Roth, 1995</p><p>1. Overall length, from the vertex to the tip of the abdomen, smaller than 20 mm .............................. 2</p><p>– Overall length, from the vertex to the tip of the abdomen, larger than 20 mm ................................ 3</p><p>2. Tegmina and abdominal sternites with several brown maculae, pronotum with uneven dots and maculae ............................................................................ A. camerunensis (Borg, 1902) comb. nov.</p><p>– Tegmina and abdominal sternites mostly unicolored, pronotum with rather uniform pattern of dots ...................................................................................................... A. erubescens (Gerstaecker, 1883)</p><p>3. Face, frons and vertex speckled with dark dots and blotches, tergal specialization fleshy, nonsetose and white ...................................................................................................... A. erhmanni Roth, 1995</p><p>– Face, frons and vertex without conspicuous black dots, tergal specialization as a pit with two transverse fold-like elevations that meet mesally .................... A. stipata (Walker, 1868) comb. nov.</p></div>	https://treatment.plazi.org/id/03E8277D7E40FF6BFF47ACAAFAD6FBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E40FF64FDAFAABDFEC5FE18.text	03E8277D7E40FF64FDAFAABDFEC5FE18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta Kirby 1903	<div><p>Genus Rhabdoblatta Kirby, 1903</p><p>Type species</p><p>Rhabdoblatta praecipua (Walker, 1868) .</p><p>Diagnosis and history</p><p>This genus is composed of more than 150 species distributed in Asia and Africa (Beccaloni 2014), and is known to be polyphyletic (e.g., Legendre et al. 2017; Yang et al. 2019; Wang et al. 2023). Clarifying its delineation is not within the aim of this manuscript. The latest diagnosis of the genus was given in Yang et al. (2019), but that diagnosis might rely mainly on Asian species and may not fit well with some African species (e.g., the widest part of the pronotum is not necessarily in the middle). The following African species share the diagnostic features of Rhabdoblatta but not those of Africalolampra: Rhabdoblata punctulata (Saussure, 1891) has serrated claws but two styli and no tergal gland, R. parva (Chopard, 1952) has two styli, unserrated claws and a tergal specialization on the 7 th tergite, R. lynceae (Gerstaecker, 1883) has slightly serrated claws but two styli and a tergal specialization on the 5 th tergite, R. pluriramosa (Karny, 1915) has two styli, unserrated claws and no tergal gland, and the female has brachypterous wings, R. pallida (Borg, 1902) has one stylus and serrated claws but a tergal specialization at the junction of the 8 th and 9 th tergites, R. punctipennis (Saussure, 1895) has one stylus and slightly serrated claws but no tergal specialization, R. fotoi sp. nov. has serrated claws but two styli and a tergal gland on the 4 th abdominal tergite.</p><p>For the following African species of Rhabdoblatta, their original descriptions did not mention all the diagnostic characters or were based on female specimens only, so that it is premature to envision any taxonomic change or confirmation: Rhabdoblatta cincta (Brunner von Wattenwyl, 1865), R. trilobata (Saussure, 1891), R. usambarensis (Rehn, 1931), R. malagassa (Saussure &amp; Zehntner, 1895) and R. wittei (Jolivet, 1954) . Note that the type specimen of R. usambarensis (Rehn, 1931) is supposed to be deposited in the Berlin Zoological Museum but it was impossible to locate this specimen. As for R. wittei (Jolivet, 1954), the description mentions tegmina that do not extend beyond the 3 rd tergite (sex not specified).</p></div>	https://treatment.plazi.org/id/03E8277D7E40FF64FDAFAABDFEC5FE18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E4FFF60FE79AFE2FD64F956.text	03E8277D7E4FFF60FE79AFE2FD64F956.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta fotoi Nyame Mbia, Legendre & Biram 2024	<div><p>Rhabdoblatta fotoi Nyame Mbia, Legendre &amp; Biram à Ngon sp. nov.</p><p>urn:lsid:zoobank.org:act: 5DDC7759-63FE-4690-B2F2-D77F9BCBE701</p><p>Figs 14–17</p><p>Differential diagnosis</p><p>Males can be readily distinguished from all other congeneric species by the presence and shape of the tergal gland on the 4 th tergite. Males of Rhabdoblatta lyncea also have a tergal gland on the 4 th tergite – although with a different shape – but no bulge on the 5 th one; instead, they show a small, transluscent, posteromedial knob on the 3 rd tergite (compare Fig. 13E with Fig. 30C).</p><p>Etymology</p><p>Named after the Professor Foto Menbohan Samuel, who initiated this work.</p><p>Material examined</p><p>Holotype</p><p>CAMEROON • 1 ♂; Centre Region, Mefou watershed, Eloundem; elev. 569–675 m; Dec. 2019 – Nov. 2020; molecular sample BL943; MNHN-EP7600.</p><p>Allotype</p><p>CAMEROON • ♀; same data as for holotype; MNHN-EP7601.</p><p>Additional specimens</p><p>CAMEROON • 1 ♀, 2 ♂♂; same data as for holotype; MNHN-EP7590 to MNHN-EP7592 .</p><p>Description</p><p>Male HEAD. Triangular, about as long as wide, mostly brown (Fig. 14C). Eyes black, ocellar spots creamish with one dark spot below each of them. Labrum and clypeus brown, lighter than frons and vertex. Scape and pedicel light brown, while other antennal segments darker. Distance between eyes equals distance between antennal sockets.</p><p>PRONOTUM. Light brown, with scattered brown freckles except on its border (Fig. 14D). Pronotum roughly triangular but with middle of hind margin distinctly convex. Laterally deflexed.</p><p>TEGMINA AND WINGS. Fully developed, extending well beyond end of abdomen (Fig. 14A–B). Tegmina with rounded apex, light brown with veins darker, with numerous small dark brown maculae and few other larger. Sc thickened in ventral view. Hindwings light brown, darker anteriorly than posteriorly, with veins darker (Fig. 14B).</p><p>LEGS. Anteroventral margin of front femur of type B2, with 3 strong spines proximally (Fig. 14C). Tibial spines well-developped. Metatarsus of hind leg longer than other segments combined (Fig. 14H), inner margin with two rows of small spines (Fig. 14I). Arolium and tarsal pulvilli well-developed, the latter bordered by “additional spines” sensu Anisyutkin (2016). Claws symmetrical and finely serrated (Fig. 14I).</p><p>ABDOMEN. Coloration of abdominal sternites variable (Fig. 14A, F), dark brown medially but with dark spots of different sizes laterally. Fourth abdominal tergite specialized: tergal gland as in Fig. 14E, with two reddish rod-like processes directed anteriorly and meeting medially in an oval papilla with a deeply emarginated posterior margin. Fifth abdominal tergite also slightly specialized with a little bulge anteromedially (Fig. 14E). Supra-anal plate with its caudal margin with a slight medial incision and with fine, sparse setae (Fig. 14G). Subgenital plate uniformly caramel-colored, symmetrical, convex, with two curled and flattened styli (Fig. 14F). Cerci multisegmented, pubescent ventrally, with a black tip.</p><p>GENITALIA (Fig. 15). Left phallomere R complex (L1 of MK64 and R70, and R2+R3+N of G96); cleft between R1T and R2, very thin, without protuberance; R3 membranous, indistinct (sensu Anisyutkin, 2016), in two parts. Sclerite L2d with its basal sclerite rod-shaped, very tapered, slightly enlarged at its base (L1 of G96); apical sclerite of L2d (L2vm of MK64, L2v of R70 and L1 of G96) crescent-like, its apical vertex more elongated than its basal vertex, and somehow bifid. Sclerite L3 (R2 of MK64, R70, and L2d of PG96) long but sclerotized only at its distal end, with a slender hook extremely bent and elongated, with a subapical incision (groove hge. sensu Klass 1997); inner margin of basal part of L3 not crenulated. No sclerite L4U.</p><p>Remarks</p><p>The male genitalia of Rhabdoblatta fotoi sp. nov. show some similarities with those of the genus Rhabdoblattella known from South-East Asia and India (Anisyutkin 1999; Anisyutkin &amp; Youshkova 2017). However, the new species lacks a median tooth along the hind margin of the subgenital plate, a diagnostic character of the genus Rhabdoblattella . Furthermore, the phylogenetic analysis using the 12S marker does not suggest a close relationship between Rhabdoblattella disparis Wang, Yang &amp; Wang, 2017 and the new species (Fig. 2). We therefore prefer to place the new species in the genus Rhabdoblatta .</p><p>Female</p><p>The female is very similar to the male, although a bit larger and without tergal specialization. Its subgenital plate is symmetrical, semi-circular, without styli but with very sparse setae postero-medially and on its hind margin (Fig. 16).</p><p>Measurements of adults</p><p>Male (n = 1): pronotum length ×width: 4.5× 6.3, tegmina length: 20, overall length × width 15.3 ×7.</p><p>Female (n = 1): pronotum length ×width: 6× 6.8, tegmina length: 24.5, overall length ×width: 28.5 ×11.</p><p>Nymphs</p><p>Pictures of nymph are provided in Fig. 17. In both male and female juveniles, tarsi of all legs with large yellowish bristles on the inner face; supra-anal plate serrated with a deep, rounded, medial incision. Male juveniles have two styli.</p><p>Molecular data</p><p>A portion of 12S rRNA has been obtained and is available under the GenBank accession number OR589771 (molecular extract BL943).</p><p>Habitat</p><p>Adults and nymphs of this species were found in all the rivers investigated.</p><p>Distribution</p><p>This species is only known from Cameroon.</p></div>	https://treatment.plazi.org/id/03E8277D7E4FFF60FE79AFE2FD64F956	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E4BFF62FDEAA911FBE4FBE1.text	03E8277D7E4BFF62FDEAA911FBE4FBE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta punctipennis (Saussure 1895)	<div><p>Rhabdoblatta punctipennis (Saussure, 1895) Fig. 18; Table 5</p><p>Epilampra punctipennis Saussure, 1895: 355 .</p><p>Heterolampra punctipennis – Kirby 1904: 123.</p><p>Epilampra punctipennis – Shelford 1910: 14. — Kevan &amp; Knipper 1961: 409. — Princis 1963b: 200. Rhabdoblatta punctipennis – Princis 1967: 681.</p><p>Africalolampra punctipennis – Anisyutkin 2016: 141.</p><p>Material examined</p><p>TANZANIA • 1 ♂, with cristallized genitalia slide; “ D.O. Afrika (Morogoro); Nachl. Schmitt; 1969; 1492”; LUHM .</p><p>Diagnosis and history</p><p>This species was initially described as Epilampra punctipennis (Saussure 1895) before first being transferred to the genus Rhabdoblatta (Princis 1967) and then to the genus Africalolampra (Anisyutkin 2016) . However, this species does not share the diagnostic characters of Africalolampra as defined by Roth (1995) and adopted here. It notably has a strong sexual dimorphism and no tergal specialization. Anisyutkin (2016) also noted that the structure of the head of R. punctipennis strongly differs from that of the type species, Africalolampra ehrmanni, and of A. erubescens . Therefore, we transfer this species back to the genus Rhabdoblatta .</p><p>Saussure (1895) described and illustrated the habitus of a female specimen, while Princis (1963b) was the first to describe the male of this species. Anisyutkin (2016) complemented this description, including with male genitalia. We complement this description with pictures of the habitus and close-ups on diagnostic characters (Fig. 18), and with a list of published localities for this species (Table 5).</p><p>Measurements of adults</p><p>Male (n = 1): pronotum length ×width: 5.5×7.9, tegmina length: 20.4, overall length ×width: 18.2 ×9.5.</p><p>Habitat</p><p>Reported from understory of coconut plantations (Kevan &amp; Knipper 1961).</p><p>Distribution</p><p>This species is known from Tanzania (including Zanzibar) and Mozambique.</p></div>	https://treatment.plazi.org/id/03E8277D7E4BFF62FDEAA911FBE4FBE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E49FF5DFDB6AA45FE2BFCD0.text	03E8277D7E49FF5DFDB6AA45FE2BFCD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta pallida (Borg 1902)	<div><p>Rhabdoblatta pallida (Borg, 1902)</p><p>Figs 19–21; Table 6</p><p>Epilampra pallida Borg, 1902: 13 .</p><p>Epilampra minuta Borg, 1902: 14 .</p><p>Epilampra borgi Shelford, 1910: 14 (unnecessary replacement name for Epilampra pallida).</p><p>Epilampra minuta – Shelford 1909b: 479; 1910: 14. — Anonymous 1910: 579. — Sjöstedt 1933: 7. Epilampra pallida – Sjöstedt 1933: 7. — Princis 1947: 59; 1962: 210; 1963a: 137.</p><p>Rhabdoblatta pallida – Vanschuytbroeck 1970: 14. — Kumar 1975: 83.</p><p>Material examined</p><p>Holotype</p><p>CAMEROON • ♀; “Camerun; Sjöstedt; Epilampra pallida Borg, det. K. Princis 1957; 123 57; 453 73; Typus ”; NHRS, NRM-BLAT 0002115 [pictures only].</p><p>Additional specimens</p><p>CAMEROON • 1 ♂; “Camerun; 10/2 91, 10.II.1891; Sjöstedt”; NHRS, NRM-BLAT 0002113 [pictures only] • 1 ♂; “Camerun, Itoki; 22/1 91, 22.I.1891; Sjöstedt; typus of minuta Borg; 398 57”; NHRS, NRM-BLAT 0002114 [pictures only] .</p><p>DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♀; “Congo Belge, Kivu Masisi, Le Moult; 1938” LUHM .</p><p>REPUBLIC OF GUINEA • 1 ♀; “ Mt Nimba N.E., 500–700 m; IFAN – 1946; A. VILLIERS”; MNHN-EP7614 • 1 juv.; “Nimba (Guinée), Serengbara; II.VI.42; M. Lamotte; Epilampra pallida Borg juv., det. K. Princis 1960”; MNHN-EP7604 .</p><p>UNKNOWN • 1 ♀; “ Epilampra pallida ♀ Borg, det. K. Princis 1956”; MNHN-EP7615 .</p><p>Diagnosis and history</p><p>This species was described as Epilampra pallida from a female specimen, whose habitus was described and drawn (Borg 1902). Its junior synonum Epilampra minuta was described in the same work, from a male and a female specimen. Note that Borg apparently described the species from two female specimens but he also reported drawing a male specimen, which was later confirmed by Princis (1963a) who illustrated the male subgenital plate and tergal specialization (near the junction of tergites VIII–IX). The shape and localization of this tergal gland is diagnostic (Kumar 1975).</p><p>Measurements of adults</p><p>Female (n = 3): pronotum length ×width: 4.5–5.6 ×5.9–6.9, tegmina length: 19.2–22.2, overall length ×width: 17.2–23.4×7.5–8.7.</p><p>Habitat</p><p>Unknown.</p><p>*nymph</p><p>Distribution</p><p>This species is known from Cameroon, Gabon, Equatorial Guinea and, putatively, the Republic of Guinea (Table 6).</p></div>	https://treatment.plazi.org/id/03E8277D7E49FF5DFDB6AA45FE2BFCD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E75FF59FDDFAF3DFB3BFC10.text	03E8277D7E75FF59FDDFAF3DFB3BFC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta pluriramosa (Karny 1915)	<div><p>Rhabdoblatta pluriramosa (Karny, 1915)</p><p>Figs 22–25; Table 7</p><p>Ischnoptera pluriramosa Karny, 1915: 144 .</p><p>Epilampra pluriramosa – Lamotte &amp; Roy 1961a: 4040; 1961b: 4209; 1962: 26. — Princis 1963a: 138. Rhabdoblatta pluriramosa – Kumar 1975: 81. — Lamotte &amp; Roy 2003: 62.</p><p>Material examined</p><p>REPUBLIC OF GUINEA • 1 ♂; “ Guinée, Nimba, Camp I; 12-13 XII 1951; Lamotte et Roy leg.”; LUHM • 1 ♂; “ Guinée, Nimba; XII 56 –V 57; Lamotte, Amiet, Vanderplaetsen; n°52”; MNHN-EP7616 • 1 ♂; “ Guinée, Nimba, Pr. Camp I; 5 avril 57, XII 56–V 57; Lamotte, Amiet, Vanderplaetsen; R. Roy det. 1961”; MNHN-EP7617 • 1 ♂; “ Nimba (Guinée), Mt To (1600m), Camp 1; II. VI. 42; M. Lamotte ”; MNHN-EP7621 • 3 ♀♀; “ Nimba (Guinée); II. VI. 42; M. Lamotte ”; MNHN-EP7618–20 • 1 juv.; “ Nimba (Guinée), Camp I, ravin; IX 56 – XI 56, 23/10/1956; Lamotte”; MNHN-EP7605 .</p><p>Diagnosis and history</p><p>This species, devoid of tergal specialization, was erroneously described in the genus Ischnoptera from a male specimen. It owes its specific epithet to its hindwing veination (Karny 1915). The type specimen was not found in Vienna (NMW); its location is unkown. Princis (1963a) described male, female and nymph specimens from several localities (Table 7). We complement these descriptions with pictures of male, female and juvenile specimens, as well as male genitalia (Figs 22–25). Females are readily distinguished from all other species except R. wittei because of their short wings. The shape of these wings distinguish the two species (Kumar 1975). Males are distinguished by their absence of a tergal specialization, combined with a pair of styli.</p><p>Description of male genitalia (Fig. 24)</p><p>Genitalia weakly sclerotized overall; left phallomere R complex (L1 of MK64 and R70, and R2+R3+N of G96); cleft between R1T and R2, thin, without protuberance; R3 Y-shaped (sensu Anisyutkin 2016), the base of the Y less sclerotized. Sclerite L2d with its basal sclerite rod-shaped, tapered, thinner and more sclerotized near the apex (L1 of G96); apical sclerite of L2d (L2vm of MK64, L2v of R70 and L1 of G96) quadrangular. Sclerite L3 (R2 of MK64, R70, and L2d of PG96) rather short, with a strong hook; inner margin of the basal part of L3 slightly crenulated. No sclerite ventrally to L3.</p><p>Measurements of adults</p><p>Male(n= 3): pronotum length×width:2.9–3.3 ×4.1–4.7, tegmina length:13.3–12.6, overall length×width: 12.3–12.5 ×4.7–4.8.</p><p>Female (n =3): pronotum length×width: 3.5–3.7×4.7–5.1, tegmina length: 3.5–3.8, overall length×width: 13.4–13.6×6.3–7.</p><p>Habitat</p><p>Orophilous (Lamotte &amp; Roy 1961b, 1962).</p><p>Distribution</p><p>This species is only known from the Republic of Guinea (Table 7). Note that an unidentified specimen at MNHN from the surroundings of Kisangani (Congo) could belong to this species.</p></div>	https://treatment.plazi.org/id/03E8277D7E75FF59FDDFAF3DFB3BFC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E71FF5BFE1BAF3DFC84FB0A.text	03E8277D7E71FF5BFE1BAF3DFC84FB0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta cincta (Brunner von Wattenwyl 1865)	<div><p>Rhabdoblatta cincta (Brunner von Wattenwyl, 1865) Fig. 26; Table 8</p><p>Epilampra cincta Brunner von Wattenwyl, 1865: 172 .</p><p>Epilampra cincta – Walker 1869: 128. — Shelford 1910: 14. — Princis 1965: 122.</p><p>Heterolampra cincta – Kirby 1904: 123.</p><p>Rhabdoblatta cincta – Princis 1967: 681.</p><p>Material examined</p><p>TANZANIA • 1 ♀; “Tanganyika Terr.: Bunduki, Ulguru Mts., 1500 m, gorge Mungula; Forêt transition, dans l’humus; 1/ 6-V-1957; Mission Zoolog. I.R.S.A.C. en Afrique orientale; P. Basilewsky et N. Leleup; coll. mus. congo”; LUHM • 1 ♀; same data as for preceding; RMCA .</p><p>Diagnosis and history</p><p>This rather small species is only known from a few localities within the Ulguru Mountains in Tanzania (Table 8). It was described from a male specimen, without any locality and supposedly deposited at NMW but not found there. This type specimen might be considered lost. Shelford (1910) suspected this species was African, which was later confirmed by Princis (1965) after observation of female specimens matching with the succinct original description. Besides its coloration pattern, this species is readily identifiable from its small size, its laterally deflexed pronotum and relatively short tarsi (Fig. 26).</p><p>Measurements of adults</p><p>Female (n = 2): pronotum length ×width: 4.2–4.5×5.3–5.4, tegmina length: 12.8–13.5, overall length ×width: 15–16.1× 7.5–7.7.</p><p>Habitat</p><p>The labels read: transitional forest, in humus.</p><p>Distribution</p><p>This species is only known from Tanzania (Table 8).</p></div>	https://treatment.plazi.org/id/03E8277D7E71FF5BFE1BAF3DFC84FB0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E70FF54FDDAAAFCFC85FD68.text	03E8277D7E70FF54FDDAAAFCFC85FD68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta usambarensis (Rehn 1931)	<div><p>Rhabdoblatta usambarensis (Rehn, 1931)</p><p>Fig. 27; Table 9</p><p>Epilampra usambarensis Rehn, 1931: 375 .</p><p>Epilampra usambarensis – Princis 1965: 122.</p><p>Rhabdoblatta usambarensis – Princis 1967: 679.</p><p>Material examined</p><p>TANZANIA • 1 ♀; “Tanganyika Terr., Bunduki, Ulguru Mts., moy. Mgeta 1300 m; 30-IV / 11-V-1957; Mission Zoolog. I.R.S.A.C. en Afrique orientale; P. Basilewsky et N. Leleup; Coll. Mus. Congo”; RMCA .</p><p>Diagnosis and history</p><p>Rehn (1931) gave a complete description, apart from the genitalia, of this species from a unique male specimen. Later, the female was found and told to be very similar to the male (Princis 1965). Although the type specimen is mentioned as deposited in the Berlin Zoological Museum, it was not found there and might be lost. We complement Rehn’s description with some pictures of this elongated species, only known from Tanzania (Fig. 26).</p><p>Measurements of adults</p><p>Female (n = 1): pronotum length × width: 4.8×6.3, tegmina length: 21.7, overall length ×width: 20 ×7.9.</p><p>Habitat</p><p>Unknown.</p><p>Distribution</p><p>This species is only known from Tanzania (Table 9).</p></div>	https://treatment.plazi.org/id/03E8277D7E70FF54FDDAAAFCFC85FD68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E7EFF50FDDCAF3DFB7DF933.text	03E8277D7E7EFF50FDDCAF3DFB7DF933.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta lyncea (Gerstaecker 1883)	<div><p>Rhabdoblatta lyncea (Gerstaecker, 1883)</p><p>Figs 28–30; Table 10</p><p>Epilampra lyncea Gerstaecker, 1883: 53 .</p><p>Epilampra electa Borg, 1902: 10 .</p><p>Epilampra infinita Borg, 1902: 11 .</p><p>Epilampra sjostedti Borg, 1902: 12 .</p><p>Heterolampra lyncea – Kirby 1904: 122.</p><p>Epilampra infinita – Shelford 1909b: 479; 1910: 14. — Anonymous 1910: 579. — Sjöstedt 1933: 7.</p><p>Epilampra electa – Shelford 1910: 14. — Sjöstedt 1933: 7.</p><p>Epilampra sjostedti – Shelford 1910: 14. — Sjöstedt 1933: 7. — Chopard 1945: 163. — Princis 1947: 59.</p><p>Epilampra lyncea – Shelford 1910: 14. — Rehn 1933: 408, 451. — Princis 1962: 209.</p><p>Rhabdoblatta lyncea – Vanschuytbroeck 1970: 14. — Kumar 1975: 80.</p><p>Material examined</p><p>Paratype</p><p>CAMEROON • 1 juv.; “Mungo Buchh.; Epilampra lyncea Gerst., juv.; Zool. Mus. Greifswald II 27341”: ZIMG .</p><p>Additional specimens</p><p>CAMEROON • 1 ♂, with genitalia slide; “1969; 1494, 18 512, 237; Nat. Kabinet Stuttgart, Coll. Br. v. W.”; LUHM • 1 ♀; “ Camerun; Sjöstedt; 400 57; type of Epilampra electa Borg; Epilampra lyncea Gerst. ♀, det. K. Princis 1957”; NHRS, NRM-BLAT-0002103 [pictures only] • 1 ♀; “ Camerun; Sjöstedt; 396 57; type of Epilampra infinita Borg; Epilampra lyncea Gerst. ♀, det. K. Princis 1957”; NHRS, NRM-BLAT-0002104 [pictures only] • 1 ♂; “ Camerun, N’dian; 23/5 91, 23.V.1891; Sjöstedt; 397 57; type of Epilampra sjöstedti Borg; Epilampra lyncea Gerst. ♂, det. K. Princis 1957”; NHRS, NRM-BLAT-0002105 [pictures only] .</p><p>Diagnosis and history</p><p>This species was described from a female specimen, with a juvenile as a paratype (Gerstaecker 1883). At ZIMG, only the paratype remains (Fig. 28) and the female holotype is most likely lost, as previously suggested (Rehn 1933). This species has three junior synonyms, described and illustrated by Borg (1902), synonymized by Princis (1962), and whose types are deposited at NHRS (Fig. 29). Kumar (1975) included this species in his key to species of Rhabdoblatta found in West Africa and the Congo Basin; the male specialization on the 4 th adbominal tergite is a diagnostic character (Kumar erroneously mentioned the 5 th tergite in his key; see Fig. 30 and Princis 1962). Genitalia of the male specimen from LUHM were already prepared; they are pictured here (Fig. 30G) for the record although hardly visible.</p><p>Measurements of adults</p><p>Male (n = 1): pronotum length ×width: 4.9 ×6.7, tegmina length: 21.1 [specimen examined, too damaged for other measurements].</p><p>Habitat</p><p>Unknown.</p><p>Distribution</p><p>This species is known from Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Gabon, Nigeria and São Tomé and Príncipe (Fig. 1). Details of localities are provided in Table 10.</p></div>	https://treatment.plazi.org/id/03E8277D7E7EFF50FDDCAF3DFB7DF933	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E7BFF50FDB8A8F6FBD1F84F.text	03E8277D7E7BFF50FDB8A8F6FBD1F84F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta wittei (Jolivet 1954)	<div><p>? Rhabdoblatta wittei (Jolivet, 1954)</p><p>Blatta assiniensis wittei Jolivet, 1954: 16 .</p><p>Rhabdoblatta wittei – Kumar 1975: 82.</p><p>We were unable to observe any specimen from this species, originally described as a subspecies of what is known as Hebardina assiniensis (Bolívar, 1893), a Blattidae, from the Democratic Republic of Congo. Thus, the generic assignation of this species to Rhabdoblatta is doubtful.</p></div>	https://treatment.plazi.org/id/03E8277D7E7BFF50FDB8A8F6FBD1F84F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E7AFF51FE25A874FC48F83D.text	03E8277D7E7AFF51FE25A874FC48F83D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta malagassa (Saussure & Zehntner 1895)	<div><p>Rhabdoblatta malagassa (Saussure &amp; Zehnter, 1895)</p><p>Epilampra malagassa Saussure &amp; Zehntner, 1895: 65 .</p><p>Heterolampra malagassa – Kirby 1904: 123.</p><p>Epilampra malagassa – Shelford 1910: 14.</p><p>Rhabdoblatta malagassa – Princis 1967: 682.</p><p>Remarks</p><p>This species is reported from Madagascar, without further indication.</p></div>	https://treatment.plazi.org/id/03E8277D7E7AFF51FE25A874FC48F83D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E7AFF51FDE7AA96FB67F9B3.text	03E8277D7E7AFF51FDE7AA96FB67F9B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta punctulata (Saussure 1891)	<div><p>Rhabdoblatta punctulata (Saussure, 1891)</p><p>Epilampra punctulata Saussure, 1891: 25 .</p><p>Epilampra punctulata – Saussure 1895: 356; 1899: 578. — Saussure &amp; Zehntner 1895: 63. — Shelford 1910: 14.</p><p>Heterolampra punctulata – Kirby 1904: 123. — Giglio-Tos 1907: 1.</p><p>Rhabdoblatta punctulata – Princis 1967: 682.</p><p>Remarks</p><p>The habitus of male and female specimens were figured in Saussure &amp; Zehntner (1895).</p></div>	https://treatment.plazi.org/id/03E8277D7E7AFF51FDE7AA96FB67F9B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E7AFF51FDD1ACD7FAEAFBD5.text	03E8277D7E7AFF51FDD1ACD7FAEAFBD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta trilobata (Saussure 1891)	<div><p>Rhabdoblatta trilobata (Saussure, 1891)</p><p>Epilampra trilobata Saussure, 1891: 25 .</p><p>Epilampra trilobata – Saussure 1895: 356. — Saussure &amp; Zehntner 1895: 60. — Shelford, 1910: 14. — Chopard 1952: 466.</p><p>Heterolampra trilobata – Kirby 1904: 123.</p><p>Rhabdoblatta trilobata – Princis 1967: 682.</p><p>Remarks</p><p>The habitus and subgenital plate of a female specimen was figured in Saussure &amp; Zehntner (1895). Chopard (1952) reported this species from moss forest (“forêt de mousses”) on Mount Tsaratanana.</p></div>	https://treatment.plazi.org/id/03E8277D7E7AFF51FDD1ACD7FAEAFBD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E79FF52FDC4AA40FEDEFA62.text	03E8277D7E79FF52FDC4AA40FEDEFA62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhabdoblatta parva (Chopard 1952)	<div><p>Rhabdoblatta parva (Chopard, 1952)</p><p>Epilampra parva Chopard, 1952: 466 .</p><p>Rhabdoblatta parva – Princis 1967: 682.</p><p>Remarks</p><p>To our knowledge, this species has not been mentioned in the literature since its description, where the habitus of a male is figured and the species reported from “Mont Tsaratanana, forêt de mousse” (Chopard 1952).</p></div>	https://treatment.plazi.org/id/03E8277D7E79FF52FDC4AA40FEDEFA62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
03E8277D7E79FF4CFF47ABC4FB18FBFE.text	03E8277D7E79FF4CFF47ABC4FB18FBFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blattellidae Karny 1908	<div><p>Unidentified species of the family Blattellidae (Fig. 31)</p><p>Material examined</p><p>CAMEROON • 2 ♀♀; South Region, Mvilla watersheds, exact locality unknown; elev. 570–594 m; Dec. 2019 – Nov. 2020; MNHN, MNHN-EP7602, MNHN-EP7603 .</p><p>Description</p><p>Female</p><p>HEAD. Triangular, longer than wide, mostly light brown, darker near vertex (Fig. 31C). Eyes black, ocellar spots indistinct. Labrum and clypeus light brown. Scape light brown, while other antennal segments darker. Distance between eyes smaller than distance between antennal sockets.</p><p>PRONOTUM. Light brown with darker spots, scattered with sparse setae near its lateral and anterior borders (Fig. 31B). Pronotum roughly trapezoidal but with middle of hind and anterior, although to a lesser extent, margins convex. Laterally deflexed.</p><p>TEGMINA AND WINGS. Fully developed, extending well beyond end of abdomen (Fig. 31A, E). Tegmina with rounded apex, light brown with veins darker. Sc thickened in ventral view. Hindwings light brown, darker anteriorly than posteriorly, with veins darker.</p><p>LEGS. Anteroventral margin of front femur of type A2, with 3–4 stronger spines proximally, distinctly serrated (Fig. 31F). Tibial spines well-developed. Metatarsus of hind leg longer than other segments combined (Fig. 31D), inner margin with two rows of small spines. Arolium and tarsal pulvilli well-developed, the latter bordered by “additional spines” sensu Anisyutkin (2016). Claws symmetrical and very finely crenulated (only visible at higher magnification).</p><p>ABDOMEN. Coloration of abdominal sternites variable, mostly light brown (Fig. 31A, E). Supra-anal plate sub-triangular, its caudal margin with long, fine setae. Subgenital plate light brown, mostly symmetrical, convex (Fig. 31E). Cerci long, multisegmented, pubescent ventrally, with a black tip (Fig. 31E).</p><p>Male</p><p>Although no male was either preserved or pictured, a few of them were observed by one of us (DLNM). Males have two styli and a tergal gland on the 1 st abdominal tergite with a truncated pyramidal shape on its upper part. In genitalia, the apical part of L2d is bifid.</p><p>Measurements of adults</p><p>Female (n = 2): pronotum length ×width: 2.7–2.9 ×3.6–3.8, tegmina length: 13.5–13.7, overall length ×width: 11.1–11.7× 3.9–4.3.</p><p>Habitat</p><p>In all the rivers investigated, adults and nymphs of this species were only found in Sounou and Lo’o rivers.</p><p>Remarks</p><p>This unidentified species was labeled as the morphotype MNCA in Nyame Mbia (2024).</p></div>	https://treatment.plazi.org/id/03E8277D7E79FF4CFF47ABC4FB18FBFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mbia, Donald-L’or Nyame;Ngon, Eric Belmond Biram à;Legendre, Frédéric;Menbohan, Samuel Foto	Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric, Menbohan, Samuel Foto (2024): Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995. European Journal of Taxonomy 946 (1): 1-57, DOI: 10.5852/ejt.2024.946.2615, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981
