taxonID	type	description	language	source
03E8307A6653027B9992FAACC5B9FCF3.taxon	description	(Figs. 1 – 8, 121)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6653027B9992FAACC5B9FCF3.taxon	materials_examined	Type locality. Madagascar, Ambohitantely Special Reserve Type material. Holotype ♂: MADAGASCAR: “ ABT / Nov. 2011 / 11 MADAGASCAR, AMBOHITANTELY Spec. Res.; 1497 m, S 18 ° 10 ’ 52.2 ” E 47 ° 17 ’ 22.5; 18. xi. ” // “ 2011; sifting forest litter, Winkler app. extraction; L. S. Rahanitriniaina lgt. ” // “ HOLOTYPUS Paulianidia densepunctata sp. nov., Janák & Baňař det. 2023 ” (MMBC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6653027B9992FAACC5B9FCF3.taxon	description	Description: Body length 6.4 mm, forebody length 3.6 mm. Dark brown, abdomen dark reddish brown, legs, mouthparts and antennomeres reddish brown (Fig. 1). Head (Fig. 2) moderately convex, a quarter longer than wide (W / L = 1.23). Surface with moderately fine and very dense punctures and with very small interstices. Eyes small, temples about 1.4 times as long as eyes (R = 1.42). Temples markedly roundly narrowed posteriorly. Antennae long and slender, fifth antennomere about 1.3 times as long as wide (L / W = 1.32), tenth antennomere slightly transverse (L / W = 0.89). Pronotum (Fig. 2) egg-shaped, moderately convex, with shallow basal impressions, about 1.1 times as long as wide (L / W = 1.13) and slightly narrower than head (PW / HW = 0.94). Sides slightly roundly widened posteriorly. Surface with moderately fine and very dense punctures and with very narrow, hardly visible interstices, similar as on head, punctation less dense along very narrow, in anterior part of pronotum at most one puncture wide unpunctured midline. Elytra (Fig. 2) elongate with very slightly rounded sides, distinct depression along suture in first quarter behind scutellum, markedly flattened along suture, about 1.1 times longer than wide (R = 1.15). Surface with raised, dense, granular sculpture. Interstices shining, without microsculpture. Abdomen (Fig. 1) slightly widened to segment V, very finely and densely punctured. Interstices on average about 1.5 times larger than diameter of punctures, surface less shining, with distinct microsculpture of transverse fields. Tergite VII with distinct membranous fringe apically. Male. Posterior margin of sternite VII with very short almost invisible middle projection (Fig. 7), sternite VIII with long triangular emargination in posterior third (Fig. 8). Aedeagus (Figs. 4 – 6) moderately long and slender, less than 1 mm long (AL = 0.91 mm). Median lobe in ventral view with wide middle part and moderately wide subrounded apex. Female unknown.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6653027B9992FAACC5B9FCF3.taxon	diagnosis	Differential diagnosis. The new species is arranged in a group of the species with raised, granular punctured elytra together with P. nigra Lecoq, 2008, P. marojejy Lecoq, 2008 and P. densipennis Jarrige, 1978. P. densepunctata Janák & Baňař, sp. nov. differs from P. marojejy by larger eyes and rounded temples, from P. nigra by much densely and coarsely punctate and less anteriorly narrowed pronotum, from P. densipennis (only in female hitherto known) by larger body, larger eyes and head slightly wider than pronotum. The aedeagus of the new species is slender than that of P. marojejy (cf. Fig. 84) and not divided in two projections apically, slightly similar to P. nigra (cf. Figs. 87 – 91), but much more slender and in details different.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6653027B9992FAACC5B9FCF3.taxon	etymology	Etymology. The name refers to the densely punctured elytra.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6653027B9992FAACC5B9FCF3.taxon	distribution	Distribution. Paulianidia densepunctata Janák & Baňař, sp. nov. is currently recorded only from the Ambohitantely Special Reserve in the northern central part of Madagascar (Fig. 121). Bionomics. The type was sifted from forest litter.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6651027D9992FC79C472FCDF.taxon	materials_examined	Type locality. Madagascar, Ranomafana National Park Type material. Holotype ♂: MADAGASCAR: “ RNF / Sept. 2012 / 11 MADAGASCAR, RANOMAFANA N. P.; 8. x. 2012 S 21 ° 15 ’ 54.4 ” E 47 ° 25 ’ 01.8 ”; 1003 m TALATAKELY, sifting litter, Winkler app. extr. L. S. Rahanitriniaina lgt. ” // “ HOLOTYPUS Paulianidia tanala sp. nov., Janák & Baňař det. 2023 ” (MMBC). Paratypes: 5 ♂, 3 ♀: “ RNF / Sept. 2012 / 01 MADAGASCAR, RANOMAFANA N. P.; 27. ix. 2012 S 21 ° 15 ’ 40.4 ” E 47 ° 25 ’ 16.5 ”; 1017 m AMBATOLAHIDIMY; sifting litter, Winkler app. extr.; L. S. Rahanitriniaina lgt. ” (MMBC, JJRC); 3 ♂: “ RNF / Sept. 2012 / 10 MADAGASCAR, RANOMAFANA N. P.; 6. x. 2012 S 21 ° 17 ’ 51.8 ” E 47 ° 26 ’ 35.1 ”; 1008 m VALOHOAKA, sifting litter, Winkler app. extr. L. S. Rahanitriniaina lgt. ” (JJRC); 2 ♂: “ MADAGASCAR — CE, Ranomafana N. P., 16 – 18. xi. 2010; 1079 m // S 21 ° 14 ’ 51 ” E 47 ° 24 ’ 13 ”, sifting; P. Baňař lgt. ” (MMBC); 1 ♂, 1 ♀: “ MADAGASCAR — CE 2010, RANOMAFANA N. P., 16 – 18. xi., S 21 ° 14 ’ 51 ” E 47 ° 24 ’ 13 ”, 1079 m sifting; P. Baňař lgt. ” (MMBC); 1 ♂: “ MADAGASCAR — CE, RANOMAFANA N. P., 17. xi. 2010, ~ 960 m sifting by stream close to main entrance; P. Baňař lgt. ” (MMBC). All paratypes with an additional label “ PARATYPUS Paulianidia tanala sp. nov., Janák & Baňař det. 2023 ”.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6651027D9992FC79C472FCDF.taxon	description	Description: Body length 5.2 – 6.0 mm (M = 5.5 mm, HT = 5.9 mm), forebody length 2.9 – 3.1 mm (M = 3.0 mm, HT = 3.0 mm). Reddish brown to brown, abdomen, mouthparts and antennomeres reddish brown, legs light reddish yellow (Fig. 19). Head (Fig. 20) moderately convex, only slightly longer than wide (L / W = 1.03 – 1.10, M = 1.06, HT = 1.06). Surface between eyes with coarse and dense umbilicate punctures, in centre of disc with punctation markedly sparser and finer and here and there with interstices larger than diameter of punctures. Eyes moderately large, temples on average about 1.5 times as long as eyes (R = 1.46 – 1.68, M = 1.56, HT = 1.46), markedly narrowed posteriorly. Antennae long and slender, fifth antennomere about one and half as long as wide (L / W = 1.34 – 1.65, M = 1.55, HT = 1.55), tenth antennomere about as long as wide (L / W = 0.88 – 1.05, M = 0.96, HT = 1.05). Pronotum (Fig. 20) egg-shaped, moderately convex, with shallow lateral impressions, about 1.1 times as long as wide (L / W = 1.10 – 1.16, M = 1.13, HT = 1.14) and almost as wide as head (PW / HW = 0.94 – 0.99, M = 0.96, HT = 0.97). Sides slightly widened posteriorly. Surface with moderately coarse and sparse punctures and with distinct interstices, punctation coarse and less dense than that on head and less dense along moderately narrow, in anterior part of pronotum more than one puncture wide unpunctured midline. Elytra (Fig. 20) subrectangular, with very slightly rounded sides and distinct depression along suture in first quarter behind scutellum, slightly flattened along suture, about 1.3 times as long as wide (R = 1.29 – 1.42, M = 1.36, HT = 1.33). Surface moderately coarsely and densely punctured. Interstices mostly as large as diameter of punctures, shining, without microscupture. Abdomen (Fig. 19) slightly widened to segment V, very finely and densely punctured. Interstices shining, without microsculpture. Tergite VII with distinct membranous fringe apically. Male. Posterior margin of sternite VII broadly rounded (Fig. 26), sternite VIII with relatively wide triangular emargination in posterior third (Fig. 27). Aedeagus (Figs. 21 – 25) long and slender, with relatively small roof shaped apical structures, about 0.85 mm long (n = 6, AL = 0.84 – 0.90 mm, M = 0.87 mm, HT = 0.88 mm), parameres longer than median lobe. Female. Posterior margin of sternite VIII rounded.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6651027D9992FC79C472FCDF.taxon	diagnosis	Differential diagnosis. The new species is arranged in a small group of species with normally (depressed) punctured elytra, impunctate or sparsely punctate disc of the head and integument of tergites smooth between punctures together with P. hova Lecoq, 2008, P. minima Lecoq, 2008 and a species described below — P. vohimana Janák & Baňař, sp. nov. The aedeagus is similar to that of P. hova, but the apical roof-shaped part of aedeagus is smaller and the species differs also externally by the head markedly narrowed behind eyes (cf. Fig. 29 in P. tanala and Fig. 28 in P. hova) and by different shape of posterior margin of male sternite VII (cf. Fig. 26 in P. tanala and Figs. 16 – 17 in P. hova).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6651027D9992FC79C472FCDF.taxon	etymology	Etymology. This species is named after the Malagasy ethnic group Tanala.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6651027D9992FC79C472FCDF.taxon	distribution	Distribution. Paulianidia tanala Janák & Baňař, sp. nov. is currently recorded only from the Ranomafana National Park in the central eastern part of Madagascar (Fig. 121). Bionomics. The type series was sifted from forest litter.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	description	(Figs. 31 – 37, 121)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	materials_examined	Type locality. Madagascar, Ranomafana, Valohoaka Type material. Holotype ♂: MADAGASCAR: “ RNF / Sept. 2012 / 10 MADAGASCAR, RANOMAFANA N. P.; 6. x. 2012 S 21 ° 17 ’ 51.8 ” E 47 ° 26 ’ 35.1 ”; 1008 m VALOHOAKA, sifting litter, Winkler app. extr. L. S. Rahanitriniaina lgt. ” // “ HOLOTYPUS Paulianidia valohoaka sp. nov., Janák & Baňař det. 2023 ” (MMBC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	description	Description: Body length 5.1 mm, forebody length 3.0 mm. Reddish brown, abdomen, mouthparts and antennomeres reddish, legs yellow (Fig. 31). Head (Fig. 32) moderately convex, slightly longer than wide (L / W = 1.06). Surface with moderately coarse and very dense umbilicate punctures, interstices smaller than diameter of punctures. Eyes small, temples on average about 1.4 times as long as eyes (R = 1.39), slightly roundly narrowed posteriorly. Antennae long and slender, fifth antennomere about 1.4 times as long as wide (L / W = 1.37), tenth antennomere transverse (L / W = 0.73). Pronotum (Fig. 32) egg-shaped, moderately convex, with shallow basal impressions, about 1.2 times as long as wide (L / W = 1.17) and slightly narrower than head (PW / HW = 0.94). Sides slightly roundly widened posteriorly. Surface with moderately coarse and sparse punctures, interstices shining, without microsculpture, about as large or larger than diameter of punctures, punctation more coarse and less dense than that on head and less dense along moderately narrow unpunctured middline; middline enlarged in anterior and basal part of pronotum. Elytra (Fig. 32) subrectangular, with very slightly rounded sides and shallow depression along suture in first quarter behind scutellum, about 1.1 times longer than wide (R = 1.12). Surface moderately coarsely and densely punctured. Interstices mostly as large as diameter of punctures, shining, without microscupture. Abdomen (Fig. 31) slightly widened to segment V, very finely and densely punctured. Interstices shining, without microsculpture. Tergite VII with distinct membranous fringe apically. Female unknown.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	diagnosis	Differential diagnosis. The new species is similar to P. vicina Jarrige, 1978, known only by a female holotype, but differs by the missing microsculpture between punctures on tergites, unpunctured midline on pronotum visible both in anterior and basal parts and shorter and wider elytra (R = 1.12, cf. Fig. 32 in P. valohoaka and R> 1.12, Fig. 115 in P. vicina).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	etymology	Etymology. The name is related to the type locality.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6657027D9992FC95C428F817.taxon	distribution	Distribution. Paulianidia valohoaka Janák & Baňař, sp. nov. is currently recorded only from Valohoaka, a part of the Ranomafana National Park in the southern eastern part of Madagascar (Fig. 121). Bionomics. The type specimen was sifted from forest litter.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	description	(Figs. 38 – 44, 121)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	materials_examined	Type locality. Madagascar, Vohimana Experimental Reserve. Type material. Holotype ♂: MADAGASCAR: “ VOH / Aug. 2012 / 09 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „ Circuit 5, PK 15 “, 22. viii. 2012 ” // “ S 18 ° 55 ’ 43.9 ” E 48 ° 29 ’ 59.8 ”; 891 m, sifting litter; Winkler app. extr.; L. S. Rahanitriniaina & E. M. Rabotoson lgt. ” // “ HOLOTYPUS Paulianidia vohimana sp. nov., Janák & Baňař det. 2023 ” (MMBC). Paratypes: 1 ♀: “ ♀ ” // “ VOH / Aug. 2012 / 01 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „ Circuit 2, borne 18 “, 18. viii. 2012 ” // “ S 18 ° 55 ’ 03.2 ” E 48 ° 29 ’ 37.0 ”; 965 m, sifting litter; Winkler app. extr.; L. S. Rahanitriniaina & E. M. Rabotoson lgt. ” (JJRC); 1 ♀: “ ♀ ” // “ VOH / Aug. 2012 / 08 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „ Circuit 2, PK 12 “, 22. viii. 2012 ” // “ S 18 ° 55 ’ 46.4 ” E 48 ° 30 ’ 02.4 ”; 916 m, sifting litter; Winkler app. extr.; L. S. Rahanitriniaina & E. M. Rabotoson lgt. ” (MMBC). All paratypes with an additional label “ PARATYPUS Paulianidia vohimana sp. nov., Janák & Baňař det. 2023 ”	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	description	Description: Body length 5.7 – 6.0 mm (M = 5.8 mm, HT = 6.0 mm), forebody length 3.0 – 3.1 mm (M = 3.1 mm, HT = 3.0 mm). Dark brown, abdomen reddish brown, mouthparts and antennomeres dark reddish brown, legs reddish yellow (Fig. 38). Head (Fig. 39) moderately convex, about 1.1 times as long as wide (L / W = 1.10 – 1.14, M = 1.12, HT = 1.11). Surface between eyes with coarse and dense umbilicate punctures, centre of disc with punctation markedly sparser and finer and here and there with interstices larger than diameter of punctures. Eyes small, temples on average about 1.7 times as long as eyes (R = 1.48 – 1.84, M = 1.69, HT = 1.48). Temples markedly roundly narrowed posteriorly. Antennae long and slender, fifth antennomere more than one and half as long as wide (L / W = 1.64 – 1.88, M = 1.77, HT = 1.88), tenth antennomere about as long as wide (L / W = 0.93 – 1.07, M = 0.98, HT = 1.07). Pronotum (Fig. 39) egg-shaped, moderately convex, with shallow basal impressions, about 1.1 times as long as wide (L / W = 1.15 – 1.17, M = 1.16, HT = 1.15) and slightly narrower than head (PW / HW = 0.96). Sides slightly roundly widened posteriorly. Surface with moderately coarse and sparse punctures and with distinct interstices between punctures, punctation coarse and less dense than that on head and less dense along moderately narrow unpunctured midline; midline in anterior part of pronotum more than one puncture wide. Elytra (Fig. 39) elongate with rounded sides and distinct depression along suture in first quarter behind scutellum, slightly flattened along suture, about 1.1 times longer than wide (R = 1.13 – 1.14, M = 1.14, HT = 1.13). Surface moderately coarsely and densely punctured. Interstices mostly slightly larger than diameter of punctures, surface shining, without microsculpture. Abdomen (Fig. 38) slightly widened to segment V, finely and sparsely punctured. Interstices shining, without microsculpture. Tergite VII with distinct membranous fringe apically. Male. Posterior margin of sternite VII with short and wide triangular emargination (Fig. 43), sternite VIII with short and moderately wide triangular emargination in posterior tenth (Fig. 44). Aedeagus (Figs. 40 – 42) moderately long and slender, about 0.7 mm long (AL = 0.69 mm). Median lobe in ventral view with moderately wide middle part, narrowed before slightly widened apical part. Female. Sternite VIII largely rounded posteriorly.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	diagnosis	Differential diagnosis. The new species is arranged in a group of species with normally (depressed) punctured elytra, impunctate or sparsely punctate disc of the head and integument of tergites smooth between punctures together with P. hova Lecoq, 2008, P. minima Lecoq, 2008, and P. tanala Janák & Baňař, sp. nov. and differs from all these species by combination of fine and sparse punctation of tergites and head markedly narrowed behind eyes and in males also by very distinct shape of sternites VII and VIII.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	etymology	Etymology. The name is related to the type locality.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A6655027F9992FF64C472F8E3.taxon	distribution	Distribution. Paulianidia vohimana Janák & Baňař, sp. nov. is currently recorded only from the Experimental Reserve Vohimana in the central part of Madagascar (Fig. 121). Bionomics. The type series was sifted from forest litter.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FF2CC32EFD26.taxon	description	(Figs. 51 – 55)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FF2CC32EFD26.taxon	materials_examined	Type locality. Central Madagascar, Eastern Andringitra, Anjavidilava. Material examined (8 specimens). MADAGASCAR, Mt. Ambondrombe, the nearest env. of peak, 1900 – 1936 m, 31. iii. 2001, J. Janák lgt., 1 ♂, 2 ♀ (JJRC); Ranomafana N. P., 16. – 18. xi. 2010, 1079 m, S 21 ° 14 ’ 51 ” E 47 ° 24 ’ 13 ”, sifting, P. Baňař lgt., 1 ♂ (JJRC); Ranomafana N. P., 17. xi. 2010; ~ 960 m; sifting by stream close to main entrance, P. Baňař lgt., 1 ♀ (MMBC); Ranomafana N. P., Vatoharanana, 5. x. 2012, S 21 ° 16 ’ 59.7 ” E 47 ° 25 ’ 30.3 ”, 1126 m, sifting litter, L. S. Rahanitriniaina lgt., 1 ♀ (MMBC); Ambohitantely Spec. Res., 17. xi. 2011, S 18 ° 11 ’ 48.5 ” E 47 ° 17 ’ 09.3 ”, sifting forest litter under Pandanus, very dry material, 1595 m, P. Baňař lgt., 1 ♂, 1 ♀ (MMBC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FF2CC32EFD26.taxon	distribution	Distribution. The species is widely distributed in humid forests from Ambohitantely and region north of lac Alaotra, through environs of Andasibé (Périnet), Ambohimahasoa, Mt. Ambondrombe, Andringitra Mts., Vohitrosa forest near Betroka up to Chaînes Anosyennes in the south-eastern part of Madagascar.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FCCEC38DFA80.taxon	description	(Figs. 58 – 62)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FCCEC38DFA80.taxon	materials_examined	Type locality. Central Madagascar, Ankazobe, Ambohitantely forest. Material examined (34 specimens). MADAGASCAR, Ambohitantely Special Reserve, L. S. Rahanitriniaina lgt.: 1. ix. 2011, S 18 ° 11 ’ 45.1 ” E 47 ° 17 ’ 13.4 ”, 1604 m, sifting forest litter under Pandanus, 1 ♀ (MMBC); 1. ix. 2011, S 18 ° 11 ’ 48.8 ” E 47 ° 17 ’ 10.7 ”, 1621 m, sifting forest litter under palm tree, 3 ♂, 1 ♀ (MMBC); 2. ix. 2011, S 18 ° 10 ’ 52.5 ” E 47 ° 17 ’ 21.2 ”, 1476 m, sifting litter under big rock by stream, 7 ♂, 10 ♀ (MMBC, JJRC); 2. ix. 2011, S 18 ° 10 ’ 52.3 ” E 47 ° 17 ’ 21.3 ”, 1500 m, sifting litter by stream, 4 ♂, 4 ♀ (MMBC, JJRC); 2. ix. 2011, S 18 ° 10 ’ 53.5 ” E 47 ° 17 ’ 22.3 ”, 1490 m, sifting forest litter under Pandanus, close to river, 1 ♀ (MMBC); 2. ix. 2011, S 18 ° 10 ’ 53.9 ” E 47 ° 17 ’ 22.4 ”, 1518 m, sifting forest litter by waterfall, 1 ♂, 2 ♀ (MMBC). Ambohitantely Spec. Res., 18. xi. 2011, sifting forest litter under palm tree, ~ 1600 m, P. Baňař lgt. (MMBC, JJRC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FCCEC38DFA80.taxon	distribution	Distribution. The species is known from Ambohitantely forest in central Madagascar only.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FA6FC34CF91B.taxon	description	(Figs. 9 – 18)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FA6FC34CF91B.taxon	materials_examined	Type locality. Eastern Madagascar, Ranomafana. Material examined (1 specimen). MADAGASCAR, Ranomafana N. P., Valohoaka, 6. x. 2012, S 21 ° 17 ’ 51.8 ” E 47 ° 26 ’ 35.1 ”, 1008 m, sifting litter, L. S. Rahanitriniaina lgt., 1 ♂ (JJRC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02719992FA6FC34CF91B.taxon	distribution	Distribution. The species is distributed in the National Park Ranomafana.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02739992F8D0C474FE8B.taxon	description	(Figs. 73 – 77)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02739992F8D0C474FE8B.taxon	materials_examined	Type locality. Central Madagascar, Ivohibe. Material examined (4 specimens). MADAGASCAR, Mt. Ambondrombe, env. of peak 1579 m, camp 6, 25. iii. – 3. iv. 2001, J. Janák lgt., 1 ♂, 1 ♀ (JJRC); Ranomafana N. P., Talakely, 9. x. 2012, S 21 ° 13 ’ 57.8 ” E 47 ° 23 ’ 48.0 ”, 1141 m, sifting litter, L. S. Rahanitriniaina lgt., 2 ♂ (MMBC, JJRC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665B02739992F8D0C474FE8B.taxon	distribution	Distribution. The species is distributed in forested areas from surroundings of Ambatofitorahana, through Ranomafana, Ranomena and Andringitra Mts. up to Ivohibe.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FF64C69AFE3A.taxon	description	(Figs. 78 – 82)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FF64C69AFE3A.taxon	materials_examined	Type locality. Eastern Madagascar, Anjanaharibe Nord, path Ambodihasina-Ambalarombe. Material examined (1 specimen). MADAGASCAR, Makirovana, - 14.1714, 49.9497, 581 m, 25. xii. 2018, sifting, V. Grebennikov lgt., 1 ♂ (JJRC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FF64C69AFE3A.taxon	distribution	Distribution. The species is distributed in forested areas of Anjanaharibe North and South, from surroundings of Andapa and Makirovana.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FDB2C639FC74.taxon	description	(Figs. 109 – 113)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FDB2C639FC74.taxon	materials_examined	Type locality. Central Madagascar, Ambatofitorahana. Material examined (3 specimens). MADAGASCAR: Madagascar CE, Andasibe N. P., 3. – 6. xi. 2010, ‘ Circuit Indri 2 ’, sifting, P. Baňař & L. S. Rahanitriniaina lgt., 1 ♂, 1 ♀ (MMBC, JJRC); Anjozorobe, - 18.4118, 47.9504, 1424 m, 8. i. 2019, sifting, V. Grebennikov, 1 ♂ (JJRC).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FDB2C639FC74.taxon	distribution	Distribution. The species is hitherto known only from surroundings of Ambatofitorahana, Anjozorobe and Andasibé (Périnet).	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FBE3C27CFA47.taxon	materials_examined	Type locality. Eastern Madagascar, 7 km W Ranomafana, 1100 m a. s. l. Material examined (8 specimens). MADAGASCAR: Madagascar CE, Andasibe N. P., 3. – 6. xi. 2010, ‘ Circuit Indri 2 ’, sifting, P. Baňař & L. S. Rahanitriniaina lgt., 1 ♂, 5 ♀ (MMBC, JJRC); Ranomafana N. P., 16. – 18. xi. 2010, 1079 m, S 21 ° 14 ’ 51 ” E 47 ° 24 ’ 13 ”, sifting P. Baňař lgt., 1 ♂, 2 ♀ (MMBC)	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
03E8307A665F02759992FBE3C27CFA47.taxon	distribution	Distribution. The species is hitherto known from three distant regions — surroundings of Ranomafana in south-eastern, Andasibé (Périnet) in central-eastern and Anjanaharibe Nord in north-eastern part of Madagascar.	en	Janák, Jiří, Baňař, Petr (2024): Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae). Zootaxa 5428 (2): 253-268, DOI: 10.11646/zootaxa.5428.2.5, URL: http://dx.doi.org/10.11646/zootaxa.5428.2.5
