taxonID	type	description	language	source
03E9879FDE5EFFFF70C5FE9EFEA039D6.taxon	description	C o m m e n t: The above specimens represent the first records of this species from Lebanon.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5EFFFF70C5FDB5FD993B99.taxon	description	C o m m e n t: The original description of Stilicus couloni is based on a male holotype and three female paratypes from ̎ Israël: Tel Dan ̎ (DRUGMAND 1989). An earlier attempt of clarifying the status of this doubtful name was unsuccessful because the type material was not found (ASSING 2012 a). Recently, however, ANLAŞ (2017) retrieved this type material on the occasion of a visit to the natural history museum in Brussels. He redescribed the type material, compared it with R. orbiculatus (PAYKULL, 1789), and provided illustrations of external and the male sexual characters. Based on these illustrations, especially those of the aedeagus, S. couloni is neither similar nor closely allied to R. orbiculatus, but undoubtedly conspecific with R. arabs. Remarkably, ANLAŞ (2017) reported specimens of R. arabs from Turkey and Lebanon in the same paper, which raises doubts regarding their identity.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5EFFFF70C5FB3CFD413D1B.taxon	description	C o m m e n t: Rugilus dilutipes is common in Greece.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5EFFFF70C5FAFEFCC13DE2.taxon	description	C o m m e n t: The distribution of R. korbi ranges from East Azerbaijan to North Iran, where it is not uncommon. For a distribution map see ASSING (2011).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5EFFFF70C5FC79FE473C5E.taxon	description	C o m m e n t: The above male represents the first record of this widespread and common species from Pakistan.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5FFFFE70C5FD36FD5E3B18.taxon	description	C o m m e n t: The distribution of this species is confined to East Nepal and West Bengal. For previous records see ASSING (2012 a, 2013).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5FFFFE70C5FCF1FD953C0B.taxon	description	C o m m e n t: Rugilus gracilis is the most widespread Himalayan Rugilus sensu strictu species, its distribution ranging across the South Himalaya from Pakistan to West Bengal (ASSING 2012 a, 2013, 2014).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5FFFFE70C5FB84FB983DDE.taxon	description	C o m m e n t: The above specimens represent the first record since the original description, which is based on type material from two localities near Tashigaon and one near Sheduwa in Khandbari District, East Nepal (ROUGEMONT 1998). Rugilus (Rugilus) prodoni (COIFFAIT, 1982) (Figs 1 - 4) M a t e r i a l e x a m i n e d: Nepal: 1 ♂, Dolakha District, E Ting Sang La, 3100 m, 12 - 13. VI. 2000, leg. Schawaller (SMNS). C o m m e n t: This species had been recorded from only two localities in Bagmati province (ROUGEMONT 1998). The above male was erroneously recorded as R. bagmaticola by ASSING (2012 a). ROUGEMONT (1998) provided rough sketches of the aedeagus. For photos of the male primary and secondary sexual characters see Figs 1 - 4.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5FFFFE70C5FE2AFD383A59.taxon	description	C o m m e n t: Rugilus morvani, like R. smetanai a macropterous species, is endemic to the Dhaulagiri range (ASSING 2012 a, 2013).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5FFFFE70C5FF6BFD18396D.taxon	description	C o m m e n t: The distribution of this macropterous species is confined to the Annapurna and Manaslu ranges (ASSING 2012 a, 2013).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5AFFFA70C5FA03FC01390D.taxon	description	C o m m e n t: The previous records of R. bagmaticola from the Annapurna range (ASSING 2012 a) refer to R. curvatus and R. prodoni (see above). The male primary and secondary sexual characters of R. bagmaticola are illustrated in Figs 5 - 8. The illustration of the aedeagus provided by ASSING (2012 a: figure 36) refers to R. prodoni.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5BFFFA70C5FD80FD2A3B00.taxon	description	C o m m e n t: The distribution of the wing-dimorphic R. confluens includes Gansu and Sichuan, with a female-based record reported also from Northwest Yunnan (ASSING 2012 a, 2013). For a distribution map see ASSING (2013). The above specimens, all of them wingless, were collected with pitfall traps.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5BFFFA70C5FE8BFE2D3A06.taxon	description	C o m m e n t: Rugilus curvatus is endemic to the Annapurna range (ASSING 2013, 2015). The above specimens were erroneously recorded as R. bagmaticola by ASSING (2012 a). The male secondary sexual characters of one of the above specimens are illustrated in Figs 9 - 12.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE5BFFF570C5FC99FC3C3D42.taxon	description	C o m m e n t: The above female was collected together with the material of R. reticulatus recorded by ASSING (2015). In all, 71 specimens of this wing-dimorphic species have been studied so far, 16 (22.5 %) of which are males. Rugilus (Rugilus) reuteri nov. sp. (Figs 13 - 15, 21) T y p e m a t e r i a l: Holotype ♂: ̎ China, N-Sichuan, Huanglong Shan, 2611 m, Sanluogou vill., 32 ° 46 ' 50 '' N, 103 ° 55 ' 25 '' E, 15 - 17. VI. 2018, leg. Reuter / Holotypus ♂ Rugilus reuteri sp. n., det. V. Assing 2018 ̎ (cAss). Paratypes: 1 ♂, 3 ♀♀: same data as holotype (cFel, cAss). E t y m o l o g y: This species is dedicated to Christoph Reuter (Hamburg), who collected the type series. D e s c r i p t i o n: Body length 4.9 - 5.8 mm; length of forebody 2.9 - 3.0 (♂♂) and 2.8 - 3.2 mm (♀♀). Habitus as in Figs 13 - 14. Coloration: body blackish; elytra with weak metallic hue; legs and antennae reddish. Head approximately 1.05 times as broad as long, broadest across eyes; lateral margins behind eyes convering towards posterior constriction in almost straight line in dorsal view, posterior angles obsolete; punctation moderately coarse, very dense, areolate, and partly confluent. Eyes large and bulging, as long as distance from posterior margin of eyes to posterior constriction, or nearly so. Anterior margin of labrum with two teeth on either side of the median incision. Pronotum small in relation to head, approximately 1.25 times as long as broad and 0.65 times as wide as head; midline with more or less distinct glossy longitudinal patch in posterior portion; punctation similar to that of head. Elytra and hind wings sexually dimorphic (Figs 13 - 14); punctures dense, moderately coarse, and of variable size; interstices without microsculpture. Metatarsomere I approximately as long as the combined length of II and III. Abdomen: tergites III-VI with moderately pronounced impressions anteriorly, these impressions with coarse, dense and irregular punctation; punctation of remaining tergal surfaces fine and dense; interstices on tergites III-VI with extremely shallow microsculpture visible only at high magnification (100 x), on tergite VII with slightly more distinct microreticulation; posterior margin of tergite VII with fully developed (♂♂) or rudimentary (♀♀) palisade fringe. ♂: elytra long (Fig. 13), approximately 1.15 times as long as pronotum; hind wings fully developed; sternite VII not distinctly modified; sternite VIII with moderately deep and rather broad posterior excision; aedeagus 0.55 - 0.58 mm long and shaped as in Fig. 15. ♀: elytra (Fig. 14) short, 0.84 - 0.94 times as long as pronotum; hind wings reduced. C o m p a r a t i v e n o t e s: Based on the similar external (including the sexual wing dimorphism) and male sexual characters, R. reuteri is closely allied to R. gansuensis ROUGEMONT, 1998 (Gansu, Shaanxi), R. daxuensis ASSING, 2012 (Sichuan), and R. emeimontis ASSING, 2015 (Sichuan). It is externally identical to R. daxuensis and additionally distinguished from the other two species as follows: from R. gansuensis by smaller body size, smaller and shorter male elytra, paler legs, and a less deep and broader posterior excision of the male sternite VIII; from R. emeimontis by smaller body size, smaller and shorter male elytra, paler legs, a smaller and narrower pronotum with a less pronounced median glossy band on the pronotum, denser and more distinct punctation of the abdomen, and a less deep and broader posterior excision of the male sternite VIII. For illustrations of R. gansuensis, R. daxuensis, and R. emeimontis see ASSING (2012 a, 2015). D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality (Fig. 21) is situated in the Huanglong Shan, North Sichuan, China. The specimens were collected with pitfall traps in a shady moist forest at the foot of a rock wall at an altitude of approximately 2610 m (REUTER pers. comm.). Rugilus (Rugilus) gonggaicus ASSING, 2012 M a t e r i a l e x a m i n e d: China: 1 ♂, 6 ♀♀, Sichuan, Gongga Shan, 29 ° 52 ' N, 102 ° 02 ' E, 3620 m, sifted, 12. VI. 2011, leg. Grebennikov (CAS, cSme, cAss). C o m m e n t: This species is endemic to the Gongga Shan. Only eleven (10.5 %) in a total of 105 specimens studied thus far are males (ASSING 2012 a, 2015).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE54FFF470C5FADFFD4939C4.taxon	description	C o m m e n t: Like most other Chinese representatives of the nominal subgenus, this endemic of the Emei Shan is subject to a sexual wing dimorphism (males macropterous; female brachypterous) and a biased sex ratio. Previously, only two in a total of 45 specimens studied were males (ASSING 2012 a, 2012 b, 2014, 2015). Including the new material, the sex ratio now stands at 0.1 (9 males: 94 females).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE55FFF470C5FE5DFC4F3C5D.taxon	description	C o m m e n t: The known distribution is confined to several localities in the environs of Zhongdian (ASSING 2012 a, 2012 b, 2015). This species, too, is subject to a sexual wing-dimorphism (males macropterous; females brachypterous) and a biased sex ratio. The ratio in the material studied up to today is 0.25 (13 males: 53 females). Rugilus (Rugilus) meilixuensis ASSING, 2012 M a t e r i a l e x a m i n e d: China: Y u n n a n: 5 ♂♂ [1 macropterous], 20 ♀♀, 15 km W Deqin, Mingyong, 28 ° 27 ' N, 98 ° 45 ', 3290 m, sifted, 29. VI. 2012, leg. Grebennikov; 5 ♂♂ [2 macropterous], 17 ♀♀, 15 km W Deqin, Mingyong, 28 ° 27 ' N, 98 ° 45 ', 3260 m, sifted, 29. VI. 2012, leg. Grebennikov (material in CAS, cSme, cAss). C o m m e n t: All known records of R. meilixuensis are from the Meili Xue Shan (ASSING 2012 a, 2012 a, 2013). The above material confirms an earlier hypothesis (ASSING 2012 a) that the males are indeed wing-dimorphic. Thus, this species is subject to a sexual wing-trimorphism (males macropterous or brachypterous with the elytra not dilated posteriorly; females brachypterous with the elytra shorter than in brachypterous males and dilated posteriorly). The sex ratio in the material studied up to today is 0.25 (11 males: 44 females); four males are macropterous, the remainder brachypterous.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE55FFF670C5FB3AFC1B3A3B.taxon	description	Rugilus (Rugilus) gryps nov. sp. (Figs 16 - 20, Map 1) T y p e m a t e r i a l: Holotype ♂: ̎ China: Sichuan Prov., Moxi env., Hailuogou valley, Gonghe vill., 1715 m, 29 ° 37 ' 27 '' N, 102 ° 06 ' 28 '' E, / 17. - 21. VI. 2014, at light, in front of hotel, ruderals and gardens close to margin of mixed forest, J. H ảjek, J. Růžička & M. Tkoč leg. / Holotypus ♂ Rugilus gryps sp. n., det. V. Assing 2015 ̎ (NMP). E t y m o l o g y: The specific epithet (Latin: griffin, also griffon or gryphon, a mythical creature with the head and wings of an eagle) is a noun in apposition. It alludes to the shape of the ventral process of the aedeagus, which somewhat resembles the beak of an eagle in lateral view. D e s c r i p t i o n: Body length 6.2 mm; length of forebody 3.8 mm. Coloration: head, pronotum, and abdomen blackish-brown; elytra dark-brown with slight bronze hue; legs dark-yellowish, with the apices of the meso- and metafemora indistinctly and narrowly darker; antennae pale-reddish, with antennomere I slightly darker red. Head (Fig. 16) weakly transverse, 1.04 times as broad as long, broadest across eyes; margins behind eyes smoothly curving towards posterior constriction in dorsal view, posterior angles obsolete; punctation coarse, largely longitudinally confluent. Eyes large and bulging, approximately 0.7 times as long as distance from posterior margin of eyes to posterior constriction. Anterior margin of labrum with two basally fused teeth on either side of the median incision. Pronotum (Fig. 16) 1.15 times as long as broad and 0.75 times as wide as head; midline without impunctate, glossy band; punctation similar to that of head. Elytra (Fig. 16) 0.97 times as long as pronotum and with pronounced humeral angles; punctation dense, moderately coarse, and defined; interstices glossy. Hind wings present. Metatarsomere I slightly longer than the combined length of II and III. Abdomen approximately 0.9 times as broad as elytra; tergites III-VI with moderately pronounced impressions anteriorly, these impressions with coarse, dense and irregular punctation; punctation of remaining tergal surfaces fine and dense; interstices without distinct microsculpture; posterior margin of tergite VII with palisade fringe. ♂: sternite VII (Fig. 19) strongly transverse, approximately 1.4 times as broad as long, with shallow median impression, posterior margin with deep and broad excision, on either side of this excision acutely produced; sternite VIII (Fig. 20) approximately 1.15 times as broad as long, posterior excision 0.2 times as deep as length of sternite; aedeagus (Figs 17 - 18) small in relation to body size, 0.6 mm long, and of compact shape. C o m p a r a t i v e n o t e s: Based on the similarly derived modifications of the male sternite VII (with large and nearly semi-circular posterior excision), the similarly derived morphology of the aedeagus (small, but robust; ventral process short and forming an angle with the basal portion; internal sac with large black sclerites), as well as the similar external characters, R. gryps belongs to the R. malaisei group, which previously included five species: R. asperrimus ASSING, nov. nom. (Guizhou), R. wuyicus ASSING, 2012 (Jiangxi), R. desectus ASSING, 2012 (Yunnan), R. aquilinus ASSING, 2012 (Sichuan), and R. malaisei (SCHEERPELTZ, 1965) (Burma). It is distinguished from them as follows: from R. asperrimus and R. wuyicus by smaller size, a more slender body, paler antennae, a much less transverse head, much longer elytra, the presence of hind wings, the presence of a palisade fringe at the posterior margin of tergite VII, a deeper posterior excision of the less transverse male sternite VII and more acute projections on either side of this excision, and a slightly smaller aedeagus with a ventral proces of slightly different shape, from R. wuyicus additionally by the more confluent punctation of the head and pronotum; from R. malaisei by smaller size, a more slender head with larger and more bulging eyes, a more slender pronotum, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus; from R. desectus by a slightly smaller body with a less transverse head and a more slender pronotum, a less deep posterior excision of the male sternite VII, and the completely different shape of the ventral process of the aedeagus; from R. aquilinus by somewhat paler legs and antennae, a less transverse head, relatively longer elytra, a less transverse male sternite VII with a deeper posterior excision of different shape, and by the slightly different shape of the ventral process of the aedeagus. For illustrations of the external and male sexual characters of R. asperrimus, R. wuyicus, R. aquilinus, R. desectus, and R. malaisei see ASSING (2012 a, 2012 b, 2015). The distributions of the species of the R. malaisei group are illustrated in Map 1. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated to the southeast of the Gongga Shan in Sichuan, China (Map 1). The holotype was collected with a light trap near the margin of a mixed forest at an altitude of 1715 m.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE57FFF170C5FA3BFD0C3949.taxon	description	C o m m e n t: Rugilus ceylanensis is the most widespread species of the genus, its vast distribution ranging from the Himalaya to the Philippines, Australia, Hawaii, and North America (ASSING 2012 a, 2012 b, 2013, 2014, 2015). The above specimens from Islamabad represent the first records from Pakistan.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE50FFF170C5FB6AFCD33D6D.taxon	description	C o m m e n t: This species had been recorded from Myanmar and China (ASSING 2012 a). The above specimens represent the first records from India.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE50FFF170C5FC59FE0F3CB2.taxon	description	C o m m e n t: Like the three preceding species, R. rufescens is widespread in the East Palaearctic region, its distribution extending from India to Japan and Singapore (ASSING 2012 a, 2013, 2014, 2015).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE50FFF170C5FEC6FC503A35.taxon	description	C o m m e n t: The distribution of R. simlaensis ranges from the Himalaya across China to Laos, Thailand, and Taiwan (ASSING 2012 a, 2012 b, 2013, 2014, 2015).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE50FFF170C5FDD3FC1D3BC5.taxon	description	C o m m e n t: Rugilus velutinus is common and widespread in the southern East Palaearctic and Oriental regions. For a distribution map see (ASSING 2013).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE50FFF070C5FA21FE2B390E.taxon	description	C o m m e n t: Rugilus japonicus is widespread from Sri Lanka across most of the Oriental and southern East Palaearctic regions to South Japan and the Philippines (ASSING 2012 a, 2015).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE51FFF070C5FE86FD0E39C6.taxon	description	C o m m e n t: Rugilus seriatus is widespread, but not common in the Oriental region (Indonesia, Malaysia, Philippines) (ASSING 2012 a).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE51FFF070C5FE5EFDD63A7F.taxon	description	C o m m e n t: This rarely recorded species is currently known only from Nepal, Laos, and Thailand (ASSING 2012 a, 2013, 2014).	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
03E9879FDE51FFF070C5FD18FE1A3B8F.taxon	description	C o m m e n t: This rare minute species had been recorded from North India, Sri Lanka, Thailand, Vietnam, Malaysia, and the Indonesian island Timor (ASSING 2012 a, 2013, 2014, CAMERON 1931). The above specimens represent the first records from Pakistan and Cambodia.	en	Assing, Volker (2019): A revision of Palaearctic and Oriental Rugilus. VI. Two new species Irom China, a new synonymy, and additional records (Coleoptera, Staphylinidae, Paederinae). Linzer biologische Beiträge 51 (1): 5-20, DOI: 10.5281/zenodo.3763624
