identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E987A8FFA5FF9DFCB64BD5F4BA5FC2.text	03E987A8FFA5FF9DFCB64BD5F4BA5FC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epicepon Nierstrasz & Brender	<div><p>Genus Epicepon Nierstrasz &amp; Brender à Brandis, 1931</p><p>Type species: Epicepon japonicum Nierstrasz &amp; Brender à Brandis, 1931 by present designation.</p><p>Diagnosis: Female. Pereomeres without mid-dorsal projections. Pleomeres 1–5 with strongly elongated lateral plates lacking digitation on margins; five pairs of biramous pleopods, endopods much smaller than exopods; uropods well developed, uniramous.</p><p>Male: Pereomeres and pleomeres distinctly segmented. Pleon lacking midventral tubercles, pleopods as low rounded swellings on pleomeres 1–5; pleotelson with large distolateral lobes, lacking true uropods.</p><p>Remarks: Females of species in Epicepon can be distinguished from those of other keponine genera in having oostegite 1 not protruding forward, no mid-dorsal processes on any pereomeres, lacking well-developed digitation on the margins of the lateral plates or pleopods, and small pleopodal endopods and large exopods (Nierstrasz and Brender à Brandis 1931). Nierstrasz and Brender à Brandis (1931) did not designate a type species for Epicepon; we select E. japonicum as the type species herein.</p></div>	https://treatment.plazi.org/id/03E987A8FFA5FF9DFCB64BD5F4BA5FC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barahona-Segovia, Rodrigo M.;Guzmán, Vicente Valdés;Pañinao-Monsálvez, Laura	Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés, Pañinao-Monsálvez, Laura (2021): A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia. Zoological Studies 60 (4): 1-10, DOI: 10.6620/ZS.2021.60-04, URL: http://dx.doi.org/10.5281/zenodo.8055919
03E987A8FFA5FF9FFCA94F15F4645DC2.text	03E987A8FFA5FF9FFCA94F15F4645DC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epicepon japonicum Nierstrasz & Brender	<div><p>Epicepon japonicum Nierstrasz &amp; Brender à Brandis, 1931 (Fig. 1)</p><p>Epicepon japonicum Nierstrasz &amp; Brender à Brandis, 1931: 188–189, figs. 71–73; Dollfus et al. 1933: 76 [list]; Shiino, 1936: 168–170, fig. 3; Danforth, 1963: 8 [list].</p><p>? ”a bopyrid” — Takeda et al. 2005: 108.</p><p>Material examined: Mature female syntype (5.0 mm TL), mature male syntype (2.2 mm TL) parasitizing Tymolus sp. (see Remarks), Misaki, Osaka, Japan, 21.9–146.3 m, coll. T. Mortensen, 10 Jun 1914 (ZMUC CR- 6930). Ovigerous female (7.2 mm TL), mature male (1.8 mm TL), parasitizing right branchial chamber of male Tymolus uncifer (Ortmann, 1892) (11.6 mm CL × 13.4 mm CW), Shizuoka, Suruga Bay, Japan, from aquarium trade, imported into Singapore, obtained by P. Y. C. Ng, 2016 (ZRC 2016.0452).</p><p>Redescription: Female (Fig. 1A–I) length 5.0 mm; maximum width (across pereomere 3) 3.0 mm; head length 0.8 mm; head width 1.4 mm; pleon length 1.8 mm. Body slightly dextral; all segments of body distinct (Fig. 1A).</p><p>Head ovate, wider than long, frontal margin slightly bilobate. Frontal lamina large and extending beyond sides and frontal margin of head (Fig. 1A). Eyes lacking. Antennules and antennae with three and four articles each, respectively, distalmost two articles of antennule and distalmost article of antenna terminally setose (Fig. 1B). Barbula with two slender short, distally rounded lateral projections on each side, outer projections longer and thinner than inner ones, middle region smooth with lateral low lobe near inner projection (Fig. 1G). Maxilliped with prominent anterior segment, palp as low non-articulated projection lacking setae, plectron triangular, short and thin (Fig. 1F).</p><p>Pereon broadest across pereomere 3. Irregularly shaped tergal projections prominent on both sides of pereomeres 1–4 and right pereomere 5, dorsolateral bosses on both sides of pereomeres 1–5 (crescent-shaped on pereomeres 1–3, triangular on pereomeres 4–5), thin coxal plates on both sides of pereomeres 1-3 and right pereomere 4 (Fig. 1A). No mid-dorsal projections on any pereomeres. Oostegites enclosing brood pouch, oostegite 1 (Fig. 1E) with ovate anterior article, longer than posterior article, internal ridge smooth, posterior article rounded distally, triangular proximally. Pereopods subequal in structure, first four shorter than others (Fig. 1A). All pereopods with short, blunt dactyli, cylindrical propodi, carpi and meri fused in posterior pereopods (Fig. 1C, D).</p><p>Pleon with six segments, first five pleomeres each bearing five pairs of biramous pleopods and longer uniramous lateral plates, all margins with slight undulation but not digitate, surfaces smooth (Fig. 1A, H). Stub-like endopods of pleopods no more than 1/4 length of slender, distally tapering exopods (Fig. 1H, I); exopods greater than 1/2 length of lateral plates (Fig. 1H, I). Terminal pleomere ending in uniramous uropods, similar to but slightly longer than fifth pleopodal exopods, surfaces smooth, margins with slight undulation but not digitate (Fig. 1A, H, I).</p><p>Male (Fig. 1J–M) length 2.2 mm; maximum width (across pereomere 3) 0.6 mm; head length 0.25 mm; head width 0.4 mm. All body regions and segments distinct (Fig. 1K).</p><p>Head broadly rounded distally, extending posteriorly to rounded point overlapping approximately 1/3 length of pereomere 1, distinctly separated from first pereomere (Fig. 1K). Thin, slit-like dark eyes near posterolateral margin of head. Antennules and antennae not visible beyond margins of head in dorsal view, of three and four articles, respectively; both bearing setae on all articles (Fig. 1L).</p><p>Pereomeres 3 and 4 subequal in width, other pereomeres only slightly narrower (Fig. 1J, K). All pereomeres tapered laterally, lacking mid-ventral tubercles. First pereopod larger than second, both larger than other five (Fig. 1J); all carpi with setae on distoventral ventral surface. Dactyli of pereopods 1 and 2 elongate, overreaching distal margins of carpi (Fig. 1J, L), dactyli of pereopods 3–7 short, inserted into cup-like indentations at distoventral margin of propodi (Fig. 1J, M).</p><p>Pleon with six pleomeres, rounded laterally, each slightly narrower than preceding one (Fig. 1J, K). Mid-ventral tubercles lacking, pleopods as low ventrolateral swellings on pleomeres 1–5. Pleomere 6 without uropods, posterolaterally extended into two slender rami, each as long as rest of pleotelson, both tipped with short setae; minute anal cone present distomedially (Fig. 1J, K).</p><p>Range: Known from the Pacific coast of Japan, Suruga Bay to Osaka, 21.9–146.3 m depth.</p><p>Remarks: The figure of the female syntype in dorsal view was labeled incorrectly by Nierstrasz &amp; Brender à Brandis (1931: fig. 71): “C II” (Coxalplatten II) is actually the coxal plate on pereomere 1 and “HS III” (Hintere Seitenteile) is actually the tergal projection on pereomere 2. Shiino (1936) reported on a pair of specimens found parasitizing Tymolus japonicus Stimpson, 1858 from Shimoda, Japan; the male and female greatly resemble the syntypes but were likewise incompletely described and figured. The identity of the host of Shiino’s (1936) specimens may be incorrect as at that time T. dromioides (Ortmann, 1892) (see below) was considered a synonym of T. japonicus (Tavares, 1992) .</p><p>The identity of the type host is uncertain as it was not retained with the parasites, but it is likely T. dromioides based on identification of a pair of male specimens collected from the type locality, Misaki, on 30 June 1914 (ZMUC NHMD- 226207). The identity of the bopyrid reported by Takeda et al. (2005) on a female T. uncifer from west of Amami-Oshima Island (28°33.25'N, 126°58.11'E – 28°22.09'N, 126°57.03'E, 334–348 m) is also unknown but is likely E. japonicum based on host choice and the known distribution of the parasite. The type specimens likely came from the right branchial chamber of the host, as the female is slightly dextral (dextral females are found in the right branchial chamber, sinistral ones in the left in all known host species; see Markham 1985).</p></div>	https://treatment.plazi.org/id/03E987A8FFA5FF9FFCA94F15F4645DC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barahona-Segovia, Rodrigo M.;Guzmán, Vicente Valdés;Pañinao-Monsálvez, Laura	Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés, Pañinao-Monsálvez, Laura (2021): A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia. Zoological Studies 60 (4): 1-10, DOI: 10.6620/ZS.2021.60-04, URL: http://dx.doi.org/10.5281/zenodo.8055919
03E987A8FFA7FF94FCB94D14F17C5802.text	03E987A8FFA7FF94FCB94D14F17C5802.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epicepon indicum Nierstrasz & Brender	<div><p>Epicepon indicum Nierstrasz &amp; Brender à Brandis, 1931 (Fig. 2)</p><p>Epicepon indicum Nierstrasz &amp; Brender à Brandis, 1931: 189–191,</p><p>figs. 74–77; Dollfus et al. 1933: 76 [list].</p><p>Material examined: Female syntype (2.8 mm TL), parasitizing Tymolus sp., Kei Islands, Indonesia, St. 56, 05°30'S, 132°51'E, 345 m, coll. T. Mortensen, Danish Expedition to the Kei Islands, 10 May 1922 (ZMUC CR-6788).</p><p>Redescription: Female (Fig. 2A–H) length 2.8 mm; maximum width (across pereomere 3) 1.3 mm; head length 0.5 mm; head width 0.7 mm; pleon length 1.2 mm. Body linear (no asymmetry); all segments of body distinct (Fig. 2A).</p><p>Head ovate, wider than long, frontal margin rounded medially. Frontal lamina relatively thin and extending beyond sides and frontal margin of head (Fig. 2A). Eyes lacking. Antennules and antennae with three and four articles each, respectively, distalmost article of antennule and two distalmost articles of antenna terminally setose (Fig. 2B). Barbula with one slender, distally rounded lateral projection on each side, middle region smooth with low lateral lobe (Fig. 2E, G). Maxilliped with prominent anterior segment, palp as low non-articulated projection with few distal setae, plectron triangular, short and broad (Fig. 2G).</p><p>Pereon broadest across pereomere 3. Irregularly shaped tergal projections prominent on pereomeres 1–5 on right side, 2–4 on left side, ovate dorsolateral bosses on pereomeres 1–5 on right side, 2–3 on left side, thin coxal plates on both sides of pereomeres 1–5 (Fig. 2A). No mid-dorsal projections on any pereomeres. Oostegites enclosing brood pouch, oostegite 1 (Fig. 2F) with ovate anterior article, longer than posterior article, internal ridge smooth, posterior article rounded on lateral margin, extending to triangular point posteriorly. Pereopods subequal in structure, first four shorter than others (Fig. 2A). All pereopods with short, blunt dactyli, cylindrical propodi, carpi and meri fused in posterior pereopods (Fig. 2C, D).</p><p>Pleon with six segments, first five pleomeres each bearing five pairs of biramous pleopods and longer uniramous lateral plates, all margins with slight undulation but not digitate, surfaces smooth (Fig. 2A, H). Short digitiform endopods of pleopods approximately half as long as slender, distally tapering exopods (Fig. 2H). Terminal pleomere ending in uniramous uropods, similar to but slightly longer than fifth pleopodal exopods, surfaces smooth, margins with slight undulation but not digitate (Fig. 2A, H).</p><p>Male (Fig. 2I–J) with all body regions and segments distinct.</p><p>Head broadly rounded distally, extending posteriorly to rounded point overlapping approximately 1/3 length of pereomere 1 (Fig. 2I). Small, ovate dark eyes near posterolateral margins of head. Antennules and antennae not visible beyond margins of head in dorsal view.</p><p>Pereomeres 2 – 4 subequal in width, other pereomeres only slightly narrower (Fig. 2I). All pereomeres tapered laterally, lacking mid-ventral tubercles. All pereopods subequal in structure and size (Fig. 2J).</p><p>Pleon with six pleomeres, rounded laterally, each slightly narrower than preceding one (Fig. 2I). Mid-ventral tubercles lacking, pleopods as ventrolateral swelling on pleomeres 1–5 (Fig. 2J). Pleomere 6 without uropods, posterolaterally extended into two slender rami, each as long as rest of pleotelson; minute anal cone present distomedially (Fig. 2I).</p><p>Range: Known only from the type locality, Kei Islands, Indonesia, 05°30'S, 132°51'E, 345 m depth.</p><p>Remarks: The male syntype was incompletely described and figured by Nierstrasz &amp; Brender à Brandis (1931) but is no longer in the vial with the female syntype and the above redescription of the male is therefore incomplete. The female syntype superficially appears to be immature based on the linear shape of the body but the oostegites are well-developed. The specific identity of the host is unknown as it was not retained with the parasites and the branchial chamber from which the parasites were extracted is not clear as the female is nearly linear.</p><p>Epicepon belema n. sp. (Figs. 3, 4) urn:lsid:zoobank.org:act: 975C18A3-DE0C-4892-AEF0- D47ACEE92297</p><p>Material examined: Female holotype (2.0 mm TL including uropods), male allotype (1.1 mm TL) parasitizing right branchial chamber of Tymolus hirtipes Tan &amp; Huang, 2000, north of Panan, Pott (Phwoc) Island, Belep Islands, New Caledonia, Sta. CP172, 19°01'1.2"S, 163°16'1.2"E, 275–330 m, coll. MUSORSTOM 4, 17 Sept 1985 (ZRC 2016.0424).</p><p>Description: Female (Fig. 3) length 2.0 mm; maximum width (across pereomere 3) 1.4 mm; head length 0.4 mm; head width 0.7 mm; pleon length 0.7 mm. Body slightly dextral; all segments of body distinct (Fig. 3A).</p><p>Head oblong, wider than long, extending into rounded lateral projections, frontal margin entire medially. Frontal lamina thin and extending slightly beyond frontal margin of head (Fig. 3A). Small, irregular black eyes present mediolaterally (Fig. 3A). Antennules and antennae with two and three articles each, respectively, distalmost article of each terminally setose (Fig. 3B). Barbula with hook-shaped rounded lateral projection (Fig. 3G) and one rounded lobe, middle region smooth with low lateral lobe (Fig. 3H). Maxilliped with prominent anterior segment, palp as low non-articulated projection with few distal setae, plectron triangular, short and thin (Fig. 3G).</p><p>Pereon broadest across pereomere 3. Irregularly shaped tergal projections prominent on pereomeres 1–5 on right side and 2–4 on left side, kidney-shaped dorsolateral bosses on both sides of pereomeres 1–4, thin coxal plates on both sides of pereomeres 1–4 (Fig. 3A). No mid-dorsal projections on any pereomeres. Oostegites enclosing brood pouch; oostegite 1 (Fig. 3E, F) with ovate anterior article, subequal in length to posterior article, internal ridge smooth, posterior article rounded distally (smaller posterior lobe is curled and has slight damage in holotype). Pereopods subequal in structure and size (Fig. 3A). All pereopods with short, blunt dactyli, cylindrical propodi, carpi and meri distinct in all pereopods (Fig. 3C, D).</p><p>Pleon with six segments, first five pleomeres each bearing pair of biramous pleopods and longer uniramous lateral plates, all margins with slight undulation but not digitate, surfaces smooth (Fig. 3A, I). Stub-like endopods of pleopods no more than half length of short, distally rounded exopods (Fig. 3I, J); exopods less than ½ length of lateral plates (Fig. 3I, J). Terminal pleomere ending in large uniramous uropods, similar to but longer than fifth pleopodal exopods, surfaces smooth, margins with slight undulation but not digitate (Fig. 3A, I).</p><p>Male (Fig. 4) length 1.1 mm; maximum width (across pereomeres 3 and 4) 0.32 mm; head length 0.2 mm; head width 0.25 mm. All body regions and segments distinct (Fig. 4A).</p><p>Head expanded distally, rounded posteriorly, distinctly separated from first pereomere (Fig. 4A). Large, irregular dark eyes near posterolateral margins of head. Antennules and antennae not visible beyond margins of head in dorsal view, of three articles each; both bearing setae on all articles (Fig. 4C).</p><p>Pereomeres 3 and 4 subequal in width, other pereomeres only slightly narrower except pereomere 7 which is much narrower (Fig. 4A). All pereomeres rounded laterally, lacking mid-ventral tubercles. First pereopod larger than second, both larger than other five (Fig. 4B, C, D); all carpi with setae on distoventral ventral surface. Dactyli of pereopods 1 and 2 elongate, reaching distal margins of carpi (Fig. 4B, C), dactyli of pereopods 3–7 short, inserted into cup-like indentations at distoventral margin of propodi (Fig. 4B, D).</p><p>Pleon with six pleomeres, rounded laterally, each slightly narrower than preceding one (Fig. 4A, B). Mid-ventral tubercles absent, distinct pleopods present laterally on pleomeres 1–5. Pleomere 6 without uropods, posterolaterally extended into two slender rami, each longer than rest of pleotelson, both tipped with short setae; anal cone present distomedially (Fig. 4A, B).</p><p>Range: Known only from the type locality, north of Panan, Pott (Phwoc) Island, Belep Islands, New Caledonia, 275– 330 m.</p><p>Remarks: The female of Epicepon belema n. sp. can be distinguished from those of the other two in the genus in having a total length less than 2 times the maximum width ( E. indicum with total length greater than 2 times the maximum width) and pleopodal endopods greater than 1/4 length of the exopods, exopods less than 1/2 length of lateral plates, and barbula without slender elongated lobes ( E. japonica with pleopodal endopods less than 1/4 length of the exopods, exopods greater than 1/2 length of lateral plates; barbula of E. indicum with one slender elongate lobe, barbula of E. japonica with two slender elongated lobes).</p><p>Etymology: The specific name “ belema ” is both the name inhabitants of the Belep islands call themselves as well as the adjective used for things related to the Belep Islands; the name is used here as a noun in apposition.</p></div>	https://treatment.plazi.org/id/03E987A8FFA7FF94FCB94D14F17C5802	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barahona-Segovia, Rodrigo M.;Guzmán, Vicente Valdés;Pañinao-Monsálvez, Laura	Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés, Pañinao-Monsálvez, Laura (2021): A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia. Zoological Studies 60 (4): 1-10, DOI: 10.6620/ZS.2021.60-04, URL: http://dx.doi.org/10.5281/zenodo.8055919
03E987A8FFACFF94FF444954F3085FF2.text	03E987A8FFACFF94FF444954F3085FF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epicepon	<div><p>Key to species of Epicepon</p><p>Based on females</p><p>1a. Total length&gt; 2x maximum width of pereon ................................ ........................ E. indicum Nierstrasz &amp; Brender à Brandis, 1931</p><p>1b. Total length &lt;2x maximum width of pereon ............................. 2</p><p>2a. Pleopodal endopods &lt;1/4 length of exopods; exopods&gt; 1/2 length of lateral plates; barbula with slender extended lobes ........ ....................... E. japonica Nierstrasz &amp; Brender à Brandis, 1931</p><p>2b. Pleopodal endopods&gt; 1/4 length of exopods; exopods &lt;1/2 length of lateral plates; barbula without slender extended lobes ... .............................................................................. E. belema n. sp.</p></div>	https://treatment.plazi.org/id/03E987A8FFACFF94FF444954F3085FF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barahona-Segovia, Rodrigo M.;Guzmán, Vicente Valdés;Pañinao-Monsálvez, Laura	Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés, Pañinao-Monsálvez, Laura (2021): A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia. Zoological Studies 60 (4): 1-10, DOI: 10.6620/ZS.2021.60-04, URL: http://dx.doi.org/10.5281/zenodo.8055919
