taxonID	type	description	language	source
03E987B6FF880D11FF110B9CFC79842F.taxon	type_taxon	Type species Humerobates rostrolamellatus Grandjean, 1936	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF880D17FF11080CFDF8801B.taxon	description	(Figures 1 – 4) Diagnosis. Body length 357 – 382 μm, width 252 – 273 μm. Rostrum rounded dorsally, with pair of lateral incisions and teeth. Prolamellar ridge absent. All prodorsal setae long, barbed. Bothridial seta composed of short, thin peduncle and large clavate, spiculate head. Lamella narrow, with large lateral tooth-like cusp. Tutorium large, distally not dentate. Notogaster with movable pteromorph, hinge well developed; ten pairs of vestigial notogastral setae; four pairs of porose areas small, circular or subcircular, with indistinct rims. Genital and aggenital setae well developed, six pairs of genital setae; anal and adanal setae represented by their alveoli. Epimeral setae short to medium long, setal formula: 3 – 1 – 3 – 3. Legs heterotridactylous. Measurements. Body length 357 – 382 (371) μm, width 252 – 273 (264) μm, length of notogaster 298 – 319 (311) μm. Integument. Body colour yellowish brown; microtuberculate over whole body and leg segments; excrescences of cerotegument granular; restricted to region between pteromorph, pedotectum I, tutorium and lateral side of body. Prodorsum (Figs. 1 A, C, 2 A, 4 A, D). Rostrum rounded dorsally, but slightly projecting in lateral view; with pair of lateral incisions and teeth; prolamellar ridge absent. Rostral seta (ro) arising slightly mediad of distal margin of tutorium, distinctly barbed, ~ 63 μm long, extending beyond tip of rostrum, curved anteromediad. Lamella narrow, ~ 50 μm long, with distinct cusp like large lateral tooth; translamella absent. Lamellar seta (le) ~ 109 μm long, barbed, inserted on medial end of lamella, extending beyond tip of rostrum. Interlamellar seta (in) ~ 76 μm long, barbed. Distance between bases of setae le and in about 55 and 46 μm, respectively. Exobothridial seta (ex) ~ 29 μm long, barbed. Bothridial seta (bs) ~ 42 μm long, with short peduncle barely extending from bothridium and large, clavate head, with spiculate surface. Bothridium oval, with ventrolateral and ventromedial scales; mostly covered by anterior margin of notogaster. Tutorium large, ~ 109 μm long, narrowed basally, with longitudinal striations, distally with prominent cusp, not dentate. Humerosejugal porose area Aj oval, ~ 21 μm long, hardly visible; porose areas Am, Ah not evident. Genal tooth narrow, pointed distally. Pedotectum I large, with convex dorsal margin, partly covering base of seta ex. Notogaster (Figs. 1 A, 2 A, 4 A, C). Longer than wide, ratio about 1.2: 1.0; anterior margin broadly rounded. Pteromorph large, curved ventrad, with line of desclerotization. Ten pairs of notogastral setae represented by their alveoli. Four pairs of porose areas circular or subcircular, with indistinct rims, subequal in size, 16 – 22 μm in diameter. Opisthonotal gland opening (gla) situated lateral or anterolateral to A 1. Lyrifissures ia, im, ih, ips and ip well developed, in normal positions. Posterior notogastral tectum present.	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF880D17FF11080CFDF8801B.taxon	description	Anogenital region (Figs. 1 B, 4 B). Genital and aggenital setae well developed; six pairs of genital setae, g 1 – g 4 inserted along anterior margin of genital plate, g 5 and g 6 close to posterior margin. Anal and adanal setae represented by their alveoli; ad 1 and ad 2 inserted posterior, ad 3 lateral to anal aperture. Lyrifissure iad located close and parallel to anterolateral margin of anal aperture. Postanal porose area small, oval. Legs (Figs. 2 B, 3). Medial claw thicker than lateral claws, tarsi without pulvillus. Ventral spur on genua II, III and femur II strongly developed. Porose area present proximoventrally on tarsi I – IV; porose areas on femora I – IV and trochanters III and IV large. Lateral antiaxial seta l ” on tibia I and genua I, II thick; seta s of tarsus II similarly barbed to other setae on this segment. Formulas of leg setae: I (1 – 5 – 3 – 4 – 20), II (1 – 5 – 3 – 4 – 15), III (2 – 3 – 1 – 3 – 15), IV (1 – 2 – 2 – 3 – 12); formulas of solenidia: I (1 – 2 – 2), II (1 – 1 – 2), III (1 – 1 – 0), IV (0 – 1 – 0); homology of setae and solenidia indicated in Table 1. Material examined. Holotype (female) and three paratypes (one male and two females): San 171, Taeha-ri, Seo-myeon, Ulleung-gun, North Gyeongsang Province, Korea, 37 ° 29 ’ 35.13 ” N, 130 ° 49 ’ 40.71 ” E, soil and litter of mixed forest composed by Siebold humlock (Tsuga sieboldii Carrière), Japanese white pine (Pinus parviflora Siebold and Zuccarini) and Engler’s beech (Fagus engleriana Seemen ex Diels), 26 May 2022, Coll. T. G. Lee. Etymology. This species is named in memory of the eminent acarologist and soil zoologist, Dr. Jun-ichi Aoki, Professor emeritus of the Yokohama National University, Japan, who has introduced generations of students to the discipline of acarology and soil zoology during his exemplary career as researcher and teacher. His enormous scientific career and influential works have always inspired the generations of soil acarologists, and influenced the creations of many young researchers. Remarks. At a glance of superficial view, both new species proposed here remind us of Chamobates species. The presence of the large lateral tooth distally on lamellae, insertion of the lamellar setae on the surface of prodorsum, absence of the prolamellar ridge, lamellar cusps and translamella all contribute to this appearance. However, both new species have hinged pteromorphs, i. e., with clearly developed line of desclerotization, which is the principal character state that can distinguish the families Chamobatidae and Humerobatidae. It should be noted that most species of Humerobates have a pair of prolamella-like longitudinal ridge on their rostral region (e. g., see Hammer 1967; Pérez-Iñigo & Pérez-Iñigo 1993; Choi 2004; Weigmann 2006), which is absent in both of our new species. Also, most species of Humerobates have short, but distinctly developed lamellar cusps and incomplete transamella, and the lamellar setae are inserted on the lamellar cusps. However, in both of our newly proposed species, lamellar cusps are not developed, but the lamellae have large lateral tooth distally, and the lamellar setae are inserted on the surface of the prodorsum. The present new species clearly differs from all other known species of Humerobates in the relatively small circular or subcircular porose areas with indistinct rims; development of large cusp-like lateral tooth of lamellar cusp; ventrally bent pteromorph with no radiating vein-like ridges, and absence of anal and adanal setae, which represented by their alveoli. Two species previously recorded in Korea, Humerobates flechtmanni Pérez-Iñigo & Pérez-Iñigo, 1993 and Humerobates nudus (Hammer, 1967) are distinguishable from the present new species in the oval and much larger notogastral porose areas, especially Aa (vs. small, round porose areas in the new species); much longer interlamellar setae, which extending beyond rostrum (vs. relatively short setae in not reaching rostrum in the new species); large pteromorphs extended laterally with many radiating vein-like ridges (vs. pteromorphs small, bent down, without vein-like ridges in the new species); lamellar cusps with no lateral tooth (vs. lamellar cusps with large cusp like lateral tooth in the new species); and well-developed anal and adanal setae (anal and adanal setae represented by their alveoli in the new species).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF8E0D1AFF110CB5FEAB80EB.taxon	description	(Figures 5 & 6) Diagnosis. Body length 502 – 567 μm, width 378 – 421. Rostrum bidentate. Prolamellar ridge absent. All prodorsal setae long, distinctly barbed. Bothridial seta with short peduncle and large, clavate head, with spiculate surface. Lamella narrow, with distinct cusp like large lateral tooth; translamella absent. Tutorium large, narrowed basally, with longitudinal striations, distally with prominent cusp, not dentate. Notogastral setae represented by their alveoli; pteromorph large, curved ventrad, with line of desclerotization; porose areas circular or subcircular, with indistinct rims. Anal and adanal setae represented by their alveoli; legs heterotridactylous. Measurements. Body length: 502 – 567 (536) μm, width of notogaster 378 – 421 (396) μm, length of notogaster 454 – 502 (471) μm. Integument. Body colour yellowish brown; microtuberculate over whole body and leg segments; excrescences of cerotegument granular; restricted to region between pteromorph, pedotectum I, tutorium and lateral side of body. Prodorsum (Figs. 5 A, C, 6 A, C). Rostrum bidentate; prolamellar ridge absent. Rostral seta (ro) barbed, ~ 109 μm long. Lamella narrow, with distinct cusp like large lateral tooth; translamella absent. Lamellar seta (le) ~ 147 μm long, barbed. Interlamellar seta (in) ~ 155 μm long, barbed. Exobothridial seta (ex) ~ 38 μm long, barbed. Bothridial seta (bs) ~ 42 μm long, with short peduncle and large, clavate head, with spiculate surface. Bothridium oval, with ventrolateral and ventromedial scales. Tutorium large, ~ 197 μm long, narrowed basally, with longitudinal striations, distally with prominent cusp, not dentate. Humerosejugal porose area AJ present, hardly discernible; porose areas Am, Ah not evident. Genal tooth narrow, pointed distally. Pedotectum I large, with convex dorsal margin, partly covering base of seta ex. Notogaster (Figs. 5 A, 6 A). Longer than wide, ratio about 1.2: 1.0; anterior margin broadly rounded. Pteromorph large, curved ventrad, with line of desclerotization. Ten pairs of notogastral setae represented by their alveoli. Four pairs of porose areas circular or subcircular, with indistinct rims. Opisthonotal gland opening (gla) and lyrifissures ia, im, ih, ips and ip well developed, in normal positions. Posterior notogastral tectum present.	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF8E0D1AFF110CB5FEAB80EB.taxon	description	Anogenital region (Figs. 5 B, 6 B). Genital and aggenital setae well developed; six pairs of genital setae, g 1 – g 4 inserted close to anterior margin of genital plate, g 5 and g 6 close to posterior margin. Anal and adanal setae represented by their alveoli; ad 1 and ad 2 inserted posterior, ad 3 lateral to anal aperture. Lyrifissure iad located close and parallel to anterolateral margin of anal aperture. Postanal porose area small, oval. Legs. Medial claw thicker than lateral claws, tarsi without pulvillus. Structure and setation of legs same as those of previous species. Material examined. Holotype (female) and six paratypes (five females and two males): San 171, Taeha-ri, Seo-myeon, Ulleung-gun, North Gyeongsang Province, Korea, 37 ° 29 ’ 35.13 ” N, 130 ° 49 ’ 40.71 ” E, soil and litter of mixed forest composed by Siebold humlock (Tsuga sieboldii Carrière), Japanese white pine (Pinus parviflora Siebold & Zuccarini) and Engler’s beech (Fagus engleriana Seemen ex Diels), 26 May 2022, Coll. T. G. Lee. Etymology. The specific name “ ulleungdoensis ” refers to the name of the island Ulleungdo, which mainly compose the Ulleung county, located in the east of Korean peninsula, from which we recovered this species. Remarks. The present new species clearly differs from all other known species of Humerobates in the relatively small circular or subcircular porose areas with indistinct rims; development of large cusp-like lateral tooth of lamellar cusp; ventrally bent pteromorph with no radiating vein-like ridges; and absence of anal and adanal setae, which represented by their alveoli. In general appearance, this species is similar to Humerobates aokii sp. nov. However, it differs from the latter species in the distinctly bidentate rostrum; relatively long interlamellar seta, which is reaching tip of rostrum; relatively smaller bothridial seta; proximally conspicuously narrowed and distally not striated tutorium, and much larger body size.	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF830D1AFF110D5DFD6781EE.taxon	type_taxon	Type species Oribata punctum Koch, 1839	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF830D1CFF110DCDFD7E817B.taxon	description	(Figures 7 & 8) Measurements. Body length 352 μm; width 252 μm, length of notogaster 312 μm. Supplementary description. Body colour yellowish brown, body surface smooth. Rostrum rounded. Setae ro 35 µm long, le 28 µm long, both setae smooth, in 72 µm long, finely barbed. Lamella narrow, with fairly broad translamella; lamellar cusp without tooth, pointed distally. Bothridial seta fusiform, its head finely barbed; bothridium mostly covered by anterior edge and tectum of notogaster. Distinct transversal ridge present between paired setae in. Tutorium (tu) broad, blade-like, with rounded cusp, with three teeth on its dorso-distal margin. Pedotectum I broad, with concave dorsal margin. Notogaster with convex anterior tectum partly covering the interlamellar region; pteromorph with line of desclerotization. Ten pairs of fine notogastral setae, of which only p - series well developed, 4 – 6 μm long, other setae represented by their alveoli. Four pairs of porose areas round, their outer margin poorly developed; Aa largest, A 3 smallest, two others subequal in size. Structure of gnathosoma typical for family; formula of palp setation: 0 – 2 – 1 – 3 – 9, palp tarsus with solenidion and three eupathidial setae. Epimeral setae medium long, thin, smooth, setal formula: 3 – 1 – 3 – 4. All ano-genital setae smooth; six pairs of genital and one pair of aggenital setae medium long, two pairs anal and three pairs of adanal setae minute. All legs heterotridactylous, formula of setation: I (1 – 5 – 3 – 4 – 20); II (1 – 5 – 3 – 4 – 15); III (2 – 3 – 1 – 3 – 16); IV (1 – 2 – 2 – 3 – 12); formula of solenidia: I (1 – 2 – 2); II (1 – 1 – 2); III (1 – 1 – 0); IV (0 – 1 – 0). Material examined. One female: Mts. Seoraksan, Sockcho, Gangwon Province, Korea, from a mixed forest, soil and litter under oak trees (Quercus mongolicus), N 38 ° 12 ’ 41 ”, E 128 ° 29 ’ 10 ”, 739 m a. s. l., 21 October 2021, Coll. B. Bayartogtokh; one female: Pyeongchan, Jinbu, Gangweong-do, 920 m a. s. l., 17. IX. 2022, Coll. Y. S. Bae. Distribution. Japan and Korea. Remarks. The characters of the present material mostly correspond well with those of the type material studied by Fujikawa (1982), Nakamura and Nakai (2009). The main characters, such as structure of prodorsum, notogaster and body size are similar. Only a difference we noticed is the smooth lamellar and rostral setae in the Korean specimen, which are finely barbed in the Japanese specimens, and we consider this difference as an intraspecific variation of different populations. So far, three species of this genus have been reported from Korea, P. hexagonus (Berlese, 1908), P. insignis (Berlese, 1910) and P. punctum (C. L. Koch, 1839) (see Choi 1988, 1995; Kim et al. 2013). The first two species are different from P. ezoensis and P. punctum by the anterior notogastral tectum, which has two lateral projections with a concave incision between them forming U-shape; and poorly developed lamella, translamella and lamellar cusp, situated underneath the notogastral tectum. Both P. ezoensis and P. punctum have relatively small, convex anterior notogastral tectum, which has no lateral projections; and well-developed lamella, translamella and lamellar cusp extending well beyond notogastral tectum. These two species are differentiating from each other in the structure of integument, i. e., clearly punctate anterolateral part of notogaster and pteromorphs due to roughened minute granules; structure of translamella, which is relatively wide in P. punctum, while it is narrow in P. ezoensis; and distally pointed tutorium as opposed to the dentate tutorium in P. ezoensis, which has three small teeth (see e. g., Bayartogtokh et al. 2002; Weigmann 2006).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF850D1CFF110DCDFC50867E.taxon	type_taxon	Type species Sphaerozetes (Tectoribates) proximus Berlese, 1910	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF850D1CFF110ABDFE5D8557.taxon	description	(Figures 9 & 10) Measurements. Body length: 300 – 324 μm; width of notogaster: 176 – 200 μm. Supplementary description. Rostral margin with medial notch. Lamellae long and broad, separated; lamellar cusps with lateral and medial dens, subequal in size. Rostral setae long, strongly barbed, curved anteromedially. Lamellar setae strongly barbed, arising between lamellar cusps. Interlamellar setae long, barbed, approximately more than twice as long as lamellar setae, reaching the incision of lamellae. Bothridial seta relatively short (40 – 44 μm), club-shaped. Exobothridial setae long, smooth; porose area Ad not evident. Notogaster densely punctate; octotaxic system expressed as four pairs of small saccules. Ten pairs of notogastral setae thin, smooth, 20 – 24 μm in length. All epimeral setae setiform, smooth; setae 3 b, 4 a, 4 b distinctly longer than other setae; setal formula 3 – 1 – 3 – 3. Six pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae, all setiform, smooth. Discidium and custodium well developed. All legs heterotridactylous, medial claw thicker and larger than lateral claws. Material examined. Two males and two females: SK Forest, Chungju, Chungbuk Province, Korea, from a mixed forest, soil and litter under broad leaved trees, Coll. B. Bayartogtokh, 13 May 2011. Distribution. Palaearctic region (Mediterranean Europe, Iran, Korea). Remarks. The character states of the present material accord well with those of the specimens studied by Bernini (1973). Only the small difference we noticed is the slightly larger inner tooth of lamellar cusp in the Korean specimens, which is contrarily smaller in the European materials, and we consider this as an intraspecific variation of different populations.	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF860D1FFF110EE4FD578367.taxon	type_taxon	Type species: Oribata dentata Berlese, 1883	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF860D01FF110F54FD2E83AF.taxon	description	(Figures 11 & 12) Measurements. Body length 496 – 520 μm, width 304 – 336 μm. Supplementary description. Adult relatively large in size. Rostral and lamellar setae medium long, thin, finely barbed; rostral seta inserted dorsolaterally on prodorsum; lamellar seta inserted on lamellar end. Interlamellar seta long, thin, finely barbed. Bothridial seta long, with unilaterally dilated, apically pointed, ciliate head. Notogastral setae represented by their alveoli. Four pairs of small rounded to oval porose areas. Epimeral and anogenital setae short; adanal setae ad 2, ad 3 and anal setae an 1, an 2 long, all barbed; ad 1 short, thin, smooth; five pairs of genital setae. Legs monodactylous. Material examined. Three females: Mts. Seoraksan, Sockcho, Gangwon Province, Korea, from a mixed forest, soil and litter under oak trees (Quercus mongolicus), Coll. G. Zolzaya and B. Oyunsuren, 14 August 2011; three females: SK Forest, Chungju, Chungbuk Province, Korea, from a mixed forest, soil and litter under broad leaved trees, Coll. B. Bayartogtokh, 13 May 2011. Distribution. Palaearctic region (Portugal, Japan, Korea). Remarks. The characters of the present material correspond well with those of the Asian and European materials studied by Fujikawa (2003) and Weigmann (2012).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF980D03FF110FE9FD2580D7.taxon	description	(Figures 13 & 14) Measurements. Body length 344 – 382 μm, width 191 – 212 μm. Supplementary description. Adult medium in size. Rostral and lamellar setae short, thin, smooth; rostral seta inserted dorsolaterally on prodorsum; lamellar seta inserted on prodorsum, away from distal end of lamella. Interlamellar seta medium long, thin, smooth. Bothridial seta long, with unilaterally dilated, apically pointed, smooth head. Notogastral setae very short, setiform, smooth. Three pairs of small rounded to oval porose areas; porose area A 3 not evident. Epimeral and anogenital setae short to medium long; adanal seta ad 1 and ad 2 long, thick; ad 1 and anal setae short, thin, smooth; five pairs of genital setae. Legs monodactylous. Material examined. Two females: SK Forest, Chungju, Chungbuk Province, Korea, from a mixed forest, soil and litter under broad leaved trees, Coll. B. Bayartogtokh, 13 May 2011; one male: Mts. Seoraksan, Sockcho, Gangwon Province, Korea, from a mixed forest, soil and litter under oak trees (Quercus mongolicus), N 38 º 12 ’ 41 ”, E 128 º 29 ’ 10 ”, elevation 739 m a. s. l., Coll. B. Bayartogtokh, 21 October 2021; two females: Pyeongchan, Jinbu, Gangweong-do, 920 m a. s. l., 17 September 2022, Coll. Y. S. Bae; two males and four females: Pachong, Daegu, Gyeongsangbuk-do, under broad-leaved trees, 15 October 2022, Coll. Y. S. Bae; one female: Wirye park, Seongnamshi, Gyeonggi province, soil and litter under broad-leaved trees, Coll. Y. S. Bae. Distribution. Cosmopolitan, except Antarctic region. Remarks. The character states of our material accord well with those of material studied by Pérez-Iñigo (1974, 1993), Weigmann (2006), and Bayartogtokh (2010).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9A0D03FF110D79FD2B8182.taxon	type_taxon	Type species Oribata peloptoides Berlese, 1888	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9A0D03FF110DE9FD99856B.taxon	description	(Figures 15 & 16) Measurements. Body length: 506 μm, width of notogaster 394 μm, length of notogaster 407 μm. Supplementary description. Body colour reddish brown; body surface foveolate. Rostrum slightly projected in lateral view, but rounded in dorsal view. Rostral seta 71 μm long, barbed. Lamellae relatively wide, but distinctly narrowed distally. Lamellar seta 105 μm long, barbed, inserted at distal end of lamella. Tutorium moderately long, narrow, slightly widening anteriorly, seta ro inserted at its distal end. Interlamellar seta 126 μm long, barbed. Bothridium directed anterolaterad, with small opening; bothridial seta 71 μm long, with minutely barbed clavate head. Notogaster almost circular, anterior margin broadly rounded. Pteromorpha fully hinged, slightly curved downwards; notogastral setae 63 – 84 μm long, all barbed. Sacculi small, round in shape. Lyrifissures ia, im and ih well developed; other lyrifissures and opisthonotal gland opening not evident. Subcapitular mentum wider than long, hypostomal setae a 21 μm long, m 34 μm long, h 42 μm long, all thin, smooth. Chelicera and palp typical for genus; palpal setation: 0 – 2 – 1 – 3 – 9 (+ ω). Epimeral region nearly smooth, apodemes apo. 2, apo. sj and apo. 3 aligned obliquely. Epimeral setae 25 – 29 μm long, thin, smooth, setal formula: 3 – 1 – 3 – 3. Circumpedal carina long, discidium projected laterally. Five pairs of genital setae 21 – 25 μm long, one pair of aggenital setae 12 μm long, two pairs of anal setae 25 – 29 μm long, three pairs of adanal setae 29 – 34 μm long, all setae barbed. Adanal lyrifissure iad situated adjacent and parallel to lateral margin of anal aperture. Legs heterotridactylous, setation of legs typical for genus, formula of leg setation including famulus: I (1 – 5 – 3 – 4 – 21); II (1 – 5 – 3 – 4 – 15); III (1 – 3 – 1 – 3 – 14); IV (1 – 2 – 2 – 2 – 12) and formula of solenidia: I (1 – 2 – 2); II (1 – 1 – 2); III (1 – 1 – 0); IV (0 – 1 – 0). Distribution. Holarctic region. Material examined. One female: Pachong, Daegu, Gyeongsangbuk-do, under broad-leaved trees, 15 October 2022, Coll. Y. S. Bae. Remarks. The character states of the present material accord well with those of the specimens studied by Hammer (1952) and Walter et al. (2014).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9F0D06FF110EE0FD98831F.taxon	type_taxon	Type species Zetes latipes Koch, 1844	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9F0D07FF110F58FB728343.taxon	description	(Figure 17) Measurements. Body length: 416 – 424 μm, width of notogaster 244 – 252 μm, length of notogaster 323 – 332 μm. Supplementary description. Body colour yellowish brown to reddish brown; body surface nearly smooth. Rostrum subtriangular with blunt tip in dorsal view, slightly protruding in lateral view. Lamella located dorsolaterally, slightly longer than half of prodorsum; translamellar line partly developed; sublamella and prolamella well developed. Sublamellar porose area slightly oval. Rostral seta inserted laterally at distal part of prolamella, long, thin, finely barbed. Lamellar seta situated on lamellar end, thin, finely barbed. Interlamellar seta inserted mediad of lamella, finely barbed. Bothridial seta with medium long stalk and finely barbed fusiform head, distally with long tip. Exobothridial seta thin, smooth. Notogaster oval, its anterior margin widely rounded. Ten pairs of notogastral setae; p 1, p 2 and p 3 minute, but clearly developed, other setae represented by their alveoli. Four pairs of sacculi small, oval to rounded. Lyrifissures ia, im, ih, ip, ips and opisthonotal gland opening (gla) clearly developed. Gnathosoma typical for family, subcapitulum longer than wide, subcapitular setae a, m, h setiform, smooth. Palp and chelicera typical for family; palpal setation: 0 – 2 – 1 – 3 – 9 (+ ω). Epimeral setae thin, smooth; epimeral setal formula: 3 – 1 – 3 – 3. Discidium well developed, conspicuously projected laterally; circumpedal carina well developed. All anogenital setae thin, smooth. Four pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae. Adanal lyrifissure iad located in paraanal position, close to anal aperture. Legs heterotridactylous; all leg setae finely barbed; formula of leg setae: I (1 – 5 – 3 – 4 – 20), II (1 – 5 – 3 – 4 – 15), III (2 – 3 – 1 – 3 – 15), IV (1 – 2 – 2 – 3 – 12); formula of solenidia: I (1 – 2 – 2), II (1 – 1 – 2), III (1 – 1 – 0), IV (0 – 1 – 0). Material examined. Two females: Sinhwayeoksa-ro, Andeok-myeon, Seogwipo-si, Jeju-do, Korea, soil under Japanese silverberry (Elaeagnus umbellata), 1 September 2022, Coll. Y. S. Bae. Distribution. Palaearctic, Oriental and Afrotropical regions. Remarks. This is one of the widely distributed species of Scheloribates, which has a Palaearctic and Pantropical geographic range. The character states of the present material accord well with those of the specimens studied by Thor (1930), Mahunka (1987), Bayartogtokh and Akrami (2000), Ivan and Vasiliu (2008).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9E0D07FF110FC5FD588046.taxon	type_taxon	Type species Oribata roubali Berlese, 1910	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF9E0D08FF110CB5FD43801B.taxon	description	(Figures 18 & 20 A, B) Measurements. Body length 539 – 564 μm, width 373 – 390 μm, length of notogaster 456 – 473 µm. Supplementary description. Body colour reddish to deep reddish-brown. Surface of body with relatively thick cerotegument, notogaster and ventral plate microtuberculate. Lamella well developed, slightly arched line between bothridium and insertion of lamellar seta in dorsal view. Sublamellar ridge well developed, short, visible only in lateral view. Rostral seta 78 μm, lamellar seta 102 μm, interlamellar seta 116 μm in length, all these setae finely barbed. Bothridial seta 84 μm long, with thin stalk and slightly expanded fusiform head pointed distally. Notogaster narrowed anteriorly, anterior margin complete; ten pairs of insertion pores of notogastral setae present; four pairs of small sacculi. Only five pairs of epimeral setae developed, other setae not evident. Four pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae thin, smooth. Legs heterotridactylous. Material examined. One male and one female: Mts. Seoraksan, Sockcho, Gangwon Province, Korea, from a mixed forest, soil and litter under oak trees (Quercus mongolicus), N 38 ° 12 ’ 41 ”, E 128 ° 29 ’ 10 ”, 739 m a. s. l., 21 October 2021, Coll. B. Bayartogtokh. Distribution. Holarctic and Oriental regions. Remarks. The characters of the Korean specimens studied here accord well with those of European and Asian materials studied by Aoki (1966), Mahunka (1996), Bayartogtokh and Weigmann (2005), Weigmann (2006). Aoki (1966) mentioned that Japanese specimens have four pairs of genital setae as in the Korean specimens studied here, while Mahunka (1996), Bayartogtokh and Weigmann (2005) observed five pairs of genital setae. Also, Bayartogtokh and Weigmann (2005) noticed nine pairs, while Mahunka (1996) found 10 pairs of epimeral setae. These might be considered as variations in different populations.	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF910D08FF110CEDFD33811E.taxon	type_taxon	Type species Oribatula connexa Berlese, 1904	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF910D08FF110D5DFC81850F.taxon	description	(Figures 19 & 20 C, D) Measurements. Body length 373 – 415 µm, width 249 – 278 µm, length of notogaster 282 – 315 µm. Supplementary description. Body colour yellowish-brown, surface of body and leg segments nearly smooth having small granules on lateral part of prodorsum. Rostrum slightly projected in lateral view, but rounded in dorsal view. Rostral seta 50 µm, lamellar seta 64 µm, interlamellar seta 60 µm long, all thin, conspicuously barbed. Exobothridial seta (ex) 43 µm long, thin, smooth. Lamella wide, conspicuously widened anteriorly; translamella wide, straight to slightly convex centrally. Bothridial seta with densely barbed club-shaped head and thin stalk. Bothridium with large openings, its posterior part concealed under anterior margin of notogaster. Notogaster almost circular in dorsal aspect, dorsosejugal suture slightly arched anteriorly; humeral projection conspicuously developed. Thirteen pairs of short, thin, smooth notogastral setae 16 – 24 µm in length, setae p 1 and p 3 slightly shorter than other notogastral setae. Porose areas small, oval in shape; opisthonotal gland opening (gla) situated posterolateral to lyrifissure im. Subcapitulum wider than long, hypostomal setae short, thin, smooth. Epimeral setae thin, smooth, setal formula: 3 – 1 – 3 – 3. Discidium well developed; circumpedal carina reaching to lateral border of ventral plate. Four pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae short, thin, smooth. Setae ad 3 and adanal lyrifissure iad situated in preanal position. Legs heterotridactylous, formula of setation: I (1 – 5 – 3 – 4 – 21); II (1 – 4 – 2 – 4 – 16); III (2 – 3 – 1 – 3 – 15); IV (1 – 2 – 2 – 3 – 12); formula of solenidia: I (1 – 2 – 2); II (1 – 1 – 2); III (1 – 1 – 0); IV (0 – 1 – 0). Material examined. Four males and four females: Mts. Seoraksan, Sockcho, Gangwon Province, Korea, from a mixed forest, soil and litter under oak trees (Quercus mongolicus), N 38 ° 12 ’ 41 ”, E 128 ° 29 ’ 10 ”, 739 m a. s. l., 21 October 2021, Coll. B. Bayartogtokh. Distribution. Palaearctic region. Remarks. The most important diagnostic character of the species is the shape of the lamellar and translamellar complex with the seta le clearly distant from lateral edge of the very short, but broad cusps. Morphological characters of the present material are similar to those of specimens studied by Mahunka (1994), Weigmann (2006), Bayartogtokh and Smelyansky (2008), and Seniczak et al. (2012).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF920D0BFF110EE0FD53831F.taxon	type_taxon	Type species Oribata nervosa Berlese, 1914	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF920D0DFF110F58FD3E80EB.taxon	description	(Figures 21 & 23 A, B) Measurements. Body length: 714 μm, width of notogaster 556 μm. Supplementary description. Body colour dark brown. Body surface microgranulate. Rostrum rounded; lamellar and sublamellar lines thin, parallel, curving backwards. Rostral (63 μm), lamellar (92 μm) and interlamellar (84 μm) setae setiform, thin, smooth. Bothridial seta (151 μm) setiform, finely barbed in distal half. Dorsosejugal porose area oval; dorsophragma irregular in shape. Dorsosejugal suture absent; ten pairs of notogastral setae represented by their alveoli. Three pairs of elongate oval porose areas (Aa, A 1, A 3) more than twice longer than width, A 2 absent. Median pore absent; opisthonotal gland opening and all lyrifissures distinct. Subcapitulum wider than long; subcapitular setae short, setiform, seta h close to anterior edge of mentum. Chelicera and palp typical for family. Epimeral region smooth; epimeral setal formula: 1 – 0 – 3 – 2, epimeral setae smooth. Pedotectum II subtriangular in ventral aspect. Circumpedal carina not reaching pedotectum I. Genital, anal and adanal setae short, setiform, thin, smooth; g 6 and aggenital setae represented by their alveoli. Adanal lyrifissure located close and parallel to anal plate. Adanal setae ad 1 and ad 2 posteriorly, ad 3 laterally to anal aperture. Postanal porose area absent. Legs heterotridactylous, median claw distinctly thicker than lateral claws; setation of legs typical for family; formulas of leg setation and solenidia: I (1 – 4 – 3 – 4 – 20) II, (1 – 4 – 3 – 4 – 15); III, (1 – 2 – 1 – 3 – 15); IV, (1 – 2 – 2 – 3 – 12), formulas of solenidia I (1 – 2 – 2), II (1 – 1 – 2), III (1 – 1 – 0), IV (0 – 1 – 0). Distribution. Palaearctic and Afrotropical regions. Material examined. One male: Pachong, Daegu, Gyeongsangbuk-do, under broad-leaved trees, 15 October 2022, Coll. Y. S. Bae. Remarks. The characters of the present material correspond well with those of the materials studied by Oudemans (1919), Mahunka (1987), and Pérez-Iñigo (1993).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF940D0DFF110D25FD3C81A6.taxon	type_taxon	Type species Pilogalumna lunai Balogh, 1958	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF940D0DFF110D95FE788557.taxon	description	(Figures 22 A – C & 23 C, D) Measurements. Body length: 280 – 288 μm, width of notogaster 202 – 208 μm. Supplementary description. Body colour deep reddish brown. Body surface microgranulate; pteromorph covered with dense granules; notogaster and ventral plate with transverse granular bands. Rostrum protruding, slightly nasiform, rounded. Tutorial and lamellar ridges (S and L) distinct, curving backwards. Rostral (ro) and lamellar (le) setae setiform, smooth. Interlamellar seta (in) short, thin, smooth. Bothridial seta (bs) having thin stalk and lanceolate head. Dorsosejugal porose area (Ad) rounded, located posterolaterad to seta in. Anterior margin of notogaster not developed. Dorsophragmatic apophysis (D) rounded, medium in size. Notogastral setae, thin, smooth. Four pairs of porose areas, Aa, A 1, A 2 and A 3 nearly circular in shape; A 1 and A 2 closely situated to each other; A 3 situated posterior to seta h 1. Lyrifissures ia, im, ih, ip and opisthonotal gland opening (gla) well developed. Subcapitulum, palp and chelicera typical for genus, subcapitular setae thin, smooth. Only four pairs of short epimeral setae (1 a, 3 a, 3 b, 4 c) observed, other setae not evident. Pedotectum II scale-like, subtriangular in ventral view; circumpedal carina well developed. Genital setae short, thin, smooth; g 1 – g 3 inserted on anterior margin of genital plate, other setae arranged longitudinally. Aggenital, anal and adanal setae represented by their alveoli, ad 1 and ad 2 inserted posterior to anal plates; ad 3 inserted in paraanal position; lyrifissure iad situated anteromediad to ad 3. Postanal porose area (Ap) rounded. Legs heterodactylous, formulas of leg setae: I (1 – 4 – 3 – 4 – 20) II, (1 – 4 – 3 – 4 – 15); III, (1 – 2 – 1 – 3 – 15); IV, (1 – 2 – 2 – 3 – 12), formulas of solenidia I (1 – 2 – 2), II (1 – 2 – 2), III (1 – 1 – 0), IV (0 – 1 – 0). Distribution. Japan and Korea. Material examined. Two females: San 171, Taeha-ri, Seo-myeon, Ulleung-gun, North Gyeongsang Province, Korea, 37 ° 29 ’ 35.13 ” N, 130 ° 49 ’ 40.71 ” E, soil and litter of mixed forest composed by Siebold humlock (Tsuga sieboldii Carrière), Japanese white pine (Pinus parviflora Siebold & Zuccarini) and Engler’s beech (Fagus engleriana Seemen ex Diels), 26 May 2022, Coll. T. G. Lee; one female: Pyeongchan, Jinbu, Gangweong-do, 920 m a. s. l., 17. IX. 2022, Coll. Y. S. Bae, Korea. Remarks. The characters of the present material correspond well with those of the type specimens studied by Hagino et al. (2017).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
03E987B6FF970D0FFF11092EFE788183.taxon	description	(Figure 22 D, E) Measurements. Body length: 316 – 336 μm, width of notogaster 240 – 256 μm. Supplementary description. Body colour brown to dark brown. Body surface faintly microtuberculated; pteromorph striated; notogaster and ventral plate with transverse granular bands. Rostrum protruding, slightly nasiform, rounded. Tutorial and lamellar ridges (S and L) distinct, nearly parallel. Rostral and lamellar setae medium long, smooth. Interlamellar seta very short, thin, smooth. Bothridial seta with long, thin stalk and lanceolate head barbed. Dorsosejugal porose areas rounded, located posterolaterad to seta in. Anterior margin of notogaster not developed; dorsophragmatic apophysis (D) medium in size, rounded. Notogastral setae short to medium long, thin, smooth. Porose areas Aa, A 1, A 2 and A 3 nearly circular; A 1 and A 2 closely situated to each other; A 3 situated posterolateral to seta h 1. Lyrifissures ia, im, ih, ip and opisthonotal gland opening (gla) well developed. Morphology of subcapitulum, palp and chelicera typical for genus. Seven pairs of short, thin epimeral setae (1 a, 1 c, 3 a, 3 c, 4 a, 4 b, 4 c) observed. Pedotectum II scale-like, subtriangular in ventral view. All ano-genital setae short, thin, smooth; g 1, g 2, and g 3 inserted on anterior margin of genital plate, other setae arranged longitudinally; adanal setae ad 1 and ad 2 inserted posterior to anal plates; ad 3 inserted in paraanal position; lyrifissure iad situated lateral to anal plate. Postanal porose area (Ap) oval, transversely oriented. Legs heterotridactylous, lateral claws serrated dorsally, but median claw smooth. Formulas of leg setae: I (1 – 4 – 3 – 4 – 20) II, (1 – 4 – 3 – 4 – 15); III, (1 – 2 – 1 – 3 – 15); IV, (1 – 2 – 2 – 3 – 12), formulas of solenidia I (1 – 2 – 2), II (1 – 2 – 2), III (1 – 1 – 0), IV (0 – 1 – 0). Distribution. Japan and Korea. Material examined. Three males and three females: San 171, Taeha-ri, Seo-myeon, Ulleung-gun, North Gyeongsang Province, Korea, 37 ° 29 ’ 35.13 ” N, 130 ° 49 ’ 40.71 ” E, soil and litter of mixed forest composed by Siebold humlock (Tsuga sieboldii Carrière), Japanese white pine (Pinus parviflora Siebold & Zuccarini) and Engler’s beech (Fagus engleriana Seemen ex Diels), 26 May 2022, Coll. T. G. Lee; 31 females: Sinhwayeoksa-ro, Andeok-myeon, Seogwipo-si, Jeju-do, Korea, soil under Japanese silverberry (Elaeagnus umbellata), 1 September, 2022, Coll. Y. S. Bae. Remarks. The characters of the present material correspond well with those of the type specimens studied by Hagino et al. (2017).	en	Bayartogtokh, Badamdorj, Bae, Yang-Seop (2024): New findings of poronotic oribatid mites (Acari: Oribatida) from Korea. Zootaxa 5405 (2): 151-184, DOI: 10.11646/zootaxa.5405.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5405.2.1
