taxonID	type	description	language	source
03E987D1FFB4FFF7FF178AD23713FA59.taxon	materials_examined	Type material. Holotype: ɗ, China, Sichuan, Xiangcheng Co., 3000 m, 21. VI. 1982, Zhang Xuezhong leg. (IZB). Paratypes: 3 ɗ, 2 Ψ, same data as holotype (IZB, ZISP); 1 ɗ, 2 Ψ, Sichuan, Xiangcheng Co., 2900 m, 18. VI. 1982, Zhang Xuezhong & Wang Shuyong leg. (IZB); 6 ɗ, 3 Ψ, Tibet, Markam, 3250 m, 12. VIII. 1982, Wang Shuyong leg. (IZB).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB4FFF7FF178AD23713FA59.taxon	description	Description. Body length 5.4 – 7.1 mm, width 2.4 – 3.1 mm. Habitus: Fig. 4. Colour: Body dark brown to black, shiny on dorsum; middle portion of mandibles, labrum externally, very narrow margins of pronotum and elytra often reddish brown; femora dark brown, tibiae and tarsi much paler, mid- to light brown, but tibiae usually infuscated apically; palpi and antennae brownish yellow to brown. Head: Medium-sized (WHmax / WPmax = 0.65 – 0.67 in male and 0.64 – 0.68 in female; WHmin / WPmax = 0.52 – 0.56 in male and 0.55 – 0.58 in female), with moderately convex eyes (WHmax / WHmin = 1.17 – 1.25 in male and 1.15 – 1.21 in female), separated from buccal fissure ventrally by distance slightly shorter than width of first antennomere. Tempora rather short, flat or weakly convex. Clypeus with one lateral setigerous pore on each side (one of paratypes with an anomalous additional setigerous pore on left side). Dorsal microsculpture visible on areas around supraorbital pores and behind eyes, usually also on vertex laterally, more distinct in female than in male; meshes fine, more or less isodiametric. Labium (Fig. 1) with prominent mental tooth; submentum with one seta on each side; ligular sclerite narrow, weakly widened apically. Antennae short, not reaching pronotal base; antennomeres 5 – 10 only slightly longer than wide. Pronotum: Moderately wide, 1.44 – 1.51 times as wide as long, widest before middle, with unisetose sides roundly or almost rectilinearly converging basad. Apical margin slightly, arcuately emarginate, bordered only laterally. Apical angles weakly protruding, widely rounded. Basal margin usually widely rounded, sometimes almost straight, bordered throughout, wider than anterior margin; basal edge glabrous. Border along sides narrow throughout. Basal angles obtuse, blunt or rounded. Pronotal disc moderately and rather evenly convex, occasionally slightly flattened at basal angles, impunctate. Median line superficial, anterior transverse depression indistinct. Basal foveae very small and shallow. Microsculpture visible throughout, meshes fine, more or less isodiametric, slightly obliterate on disc medially. Elytra: Moderately convex, 1.34 – 1.44 times as long as wide, 2.35 – 2.55 times as long and 1.15 – 1.23 times as wide as pronotum, widest just behind middle, widely rounded at sides. Base of elytra slightly wider than pronotal base. Humeri angulate, obtuse, each with a small denticle visible dorsally. Preapical sinuation distinct, deeper in female than in male. Sutural angle acute, slightly blunt at apex. Basal border glabrous, weakly sinuate, forming a distinct obtuse angle with lateral margin. Striae impunctate, largely superficial, slightly impressed apically. Scutellar striole long, with a basal pore. Intervals flat, occasionally slightly convex apically. Third interval with a small discal pore in apical third. Fifth and seventh intervals lacking rows of pores before apices. Series umbilicate irregular and not interrupted medially. Microsculpture visible throughout, consisting of fine meshes, isodiametric along margins and weakly transverse on disc. Wings: Fully developed. Ventral surface: Prosternum glabrous medially. Proepisterna impunctate. Metepisterna (Fig. 3) notably longer than wide, narrowed posteriad. Three apical abdominal sternites glabrous, only with obligatory fixed setae. Anal sternite rounded at apex in both sexes, without pronounced sexual dimorphism. Legs: Metacoxa without additional setae. Protibia markedly widened apically, without prominent ventroapical tubercle in male, with one ventroapical spine and with three preapical spines at outer margin isolated from spines on ventral surface. Metafemur ventrally with 4 – 5 setigerous pores along posterior margin and occasionally one pore at anterior margin closer to apex (in addition to an apical pore situated just on anterior margin). Tarsi impunctate and glabrous dorsally. First mesotarsomere in male rather narrow, with two tiny scales apically. Metatarsomeres with sides slightly diverging apicad; first metatarsomere longer than second and shorter than second and third combined together. Female genitalia (Fig. 2): Hemisternite comparatively wide, with 3 – 4 short setae distally. Basal stylomere with two setae distally. Apical stylomere arcuate, relatively wide, with 2 – 5 basal setae on external dorsal margin. Male genitalia: Median lobe (Figs 5 – 6) bent before middle, with almost straight ventral margin in its apical half (lateral aspect). Terminal lamella slightly directed dorsad and weakly bent ventrad just at apex, it moderately long (notably longer than wide), almost parallel-sided, narrowly rounded at apex and with very small dorsoapical marginal flange forming an oblique horseshoe – shaped apical capitulum. Apical orifice shifted to left, extending to middle of median lobe. Internal sac with two groups of small spines: a larger dorsolateral group on left side and a smaller ventrolateral group on right side, without any separate large teeth.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB4FFF7FF178AD23713FA59.taxon	distribution	Distribution. This new species is known only from northwestern Sichuan and eastern Tibet.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB4FFF7FF178AD23713FA59.taxon	etymology	Etymology. The species epithet refers to the shiny dorsum of the new species.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB4FFF7FF178AD23713FA59.taxon	discussion	Remarks. The taxonomic position of this new species is not absolutely clear. In appearance, it is very similar to representatives of the tinctulus group sensu Kataev 1997 (= Zangoharpalus Huang), particularly to some specimens of H. pseudotinctulus Schauberger with impunctate base of pronotum, but differs sharply from them in such important characters as the structure of the aedeagus and the presence of ventral adhesive scales at apex of the fist mesotarsomere of the male. In addition, unlike all known members of the tinctulus group, H. vernicosus sp. n. has the median lobe of aedeagus with horseshoe – shaped apical capitulum and without any individual teeth in the internal sac. Based on total combination of the characters of the male genitalia and external morphology, H. vernicosus sp. n. probably belongs to the tardus group sensu Kataev in Kryzhanovskij et al. 1995, and is most closely related, in our opinion, to H. modestus Dejean, H. bungii Chaudoir, H. chasanensis Lafer, H. tangutorum Kataev, and H. praecurrens Schauberger. The new species shares with these species the following distinctive character states: antennae pale, pronotum with glabrous basal edge, protibia with one ventroapical spine and without prominent ventroapical tubercle in male, mesotibia and anal sternite without pronounced sexual dimorphism, three last abdominal sternites glabrous, elytra with only one discal setigerous pore on each third interval and without rows of pores at apex of fifth and seventh intervals, and internal sac of median lobe with several spiny patches medially and without any large separate teeth. H. vernicosus sp. n. is rather distinct within this group and clearly recognized by its small and relatively narrower body, impunctate base of pronotum, finer elytral striae and specific pattern of spiny patches in the internal sac of the aedeagus. A similar pattern of spiny patches is characteristic also of the species of the vittatus group sensu Kataev 1990 which differ, however, in the dark antennae and clearly prominent ventroapical tuberculum of protibia in male.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB6FFF0FF178DF235E5FEA9.taxon	discussion	Remarks. H. viridanus belongs to the fuscipalpis group which is characterized by the antennae clearly infuscate from first or second antennomere, the pronotum impunctate basally, the abdominal sternites with additional long setae, the last abdominal sternite without distinct sexual dimorphism, the ventroapical tubercle of protibia in male not prominent and the internal sac of the aedeagus with a small median spiny patch and without separate large teeth. The fuscipalpis group is very difficult taxonomically and the number of species to be included has long been a subject of debate among specialists. The last revision of this group was published by Kataev (1989) who recognized within it four species: the Holarctic H. fuscipalpis Sturm, the West Palaearctic H. fuscicornis Ménétriés and H. inexspectatus Kataev, and the Central Asian H. viridanus Motschulsky. H. viridanus is distinguished from the other species by the basal edge of pronotum glabrous, the metafemur with one or several setigerous pores along anterior margin in its apical portion, the protibia usually with two ventroapical spines and the median lobe of the aedeagus more or less straight in dorsal aspect. The colour, body proportions and shape of the terminal lamella of the aedeagus of this species are highly variable both geographically and within populations. This variation was partly described by Mlynář (1974) and Kataev (1989) but no subspecies were recognized. Examination of the additional material from China revealed that at least two forms from the southern boundaries of the species’ geographical range are markedly separated morphologically and geographically and may be treated as subspecies. Further study of variation of H. viridanus is needed.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB0FFF2FF1788823691FAEA.taxon	materials_examined	Type material. Lectotype of H. viridanus (designated by Kataev, 1989: 223): Ψ, labelled " Ooistus viridanus mihi, Transbaic. ", " 5 ", " Transbaic. " (ZMM). Paralectotypes: 1 ɗ, 1 Ψ, from Transbaicalia (ZMM). Lectotype of H. unicolor (designated by Kataev, 1989: 223): ɗ, labelled " Kiachta " (ZMM). Paralectotypes: 1 Ψ (without head and pronotum), labelled " Ooistus unicolor mihi, B. Kiachta ", " Kiachta " (ZMM); 1 Ψ, " Kiachta " (ZMM). Lectotype of H. planatus (designated by Kataev, 1989: 223): Ψ (damaged), labelled " Ooistus planatus mihi, Mongolia ", " Mongol " (ZMM). Holotype of H. cyanellus: ɗ, labelled " Siberia or., G. Zerentui, J. Wagner ", " fuscipalpis St., v. cyaneus m., Typ., Tschitscherin det. " (ZISP). Lectotype of H. Przewalskyi (present designation: see remarks below): ɗ, labelled " Amdo [area between Dulan and Barun, Qinghai, China], 18. IV. – 11. V. 1884, Przevalsky ", " Harpalus Przewalskyi T. Tsch., n. sp., 1894, Typ! ", " v. viridanus Motsch., Tschitscherin det. " (ZISP). Paralectotypes: 1 Ψ, same data (ZISP); also apparently from the type series: 1 ɗ, labelled " Amdo. 1884, Przevalsky / 18. IV – 11. V ", " v. versicolor Reitt., Tschitscherin det. " (ZISP); 1 ɗ, " Amdo. 1884, Przevalsky / 18. IV. – 11. V ", " v. viridanus Motsch., Tschitscherin det. " (ZISP); 1 ɗ, labelled " Amdo. 1884, Przevalsky / 18. IV. – 11. V ", " Fuscipalpis Sturm, Tschitscherin det. " (ZISP); 1 ɗ, labelled " Amdo [“ Dzun-Zasak – Barun-Zasak ” (= Barun env., Qinghai)], 1884, Przevalsky / 2 – 10. V. ", " v. cyaneus m. Tschitscherin det. " (ZISP). Lectotype of H. fuscipalpis var. versicolor (present designation): ɗ, labelled " Quellgebiet des Irkut, Leder ", " v. versicolor m., i. l. (viridanus Motsch.) ", " coll. Reitter ", " Holotypus, 1899, Harpalus fuscipalpis var. versicolor Reitter ", " vidi, 1970, Dr. Z. Mlyná ř det. " (TMB). Holotype of H. csikii: ɗ, " Mongolia, Chentey aimak, Somon Mörön, 1200 m, Exp. Dr. Z. Kaszab, 1965 ", " nr 318, 28. VII. 1965 ", " Holotypus, 1966, Harpalus csikii Jedli ċ ka ", " Holotypus ", " Harpalus csikii sp. n. ", " Harpalus viridanus Mot., Dr. Z. Mlynář det., 1970 " (TMB).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB0FFF2FF1788823691FAEA.taxon	description	Description. Comparatively small: body length 7.6 – 9.8 mm (usually under 9.5 mm). Head relatively small (WHmax / WPmax = 0.61 – 0.69; WHmin / WPmax = 0.53 – 0.58). Pronotum (Fig. 10) with comparatively wide base (WPmax / WPmin = 1.03 – 1.09) and with sides more or less parallel basally, at most only slightly converging posteriad. Pronotal basal angle right-angled or slightly obtuse, usually with somewhat sharp apex. Elytral striae deepened or almost superficial. Elytral intervals more or less convex, occasionally nearly flat. Dorsal microsculpture consisting of more or less isodiametric meshes, either distinct throughout on head, pronotum and elytra or slightly obliterate. Median lobe of aedeagus relatively small (LB / LML = 3.40 – 4.02), its terminal lamella usually rather short, with sides parallel or slightly converging apicad; more rarely sides slightly sinuate (mainly in specimens from China and Kyrgyzstan).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB0FFF2FF1788823691FAEA.taxon	distribution	Distribution. Widely distributed over Mongolia and southern Siberia from Altai Mountains to the Amur River, also in central Yakutia, central Tien Shan (Kyrgyzstan) and northern China from eastern Tien Shan (Xinjiang) to Songhua River (Heilongjiang). In China, the southern boundary runs across northeastern Qinghai (probably to north of Burhan Buda Shan), southern Gansu (to north of Minxian), northern Shanxi (Wutai Shan) and then to Heilongjiang.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB0FFF2FF1788823691FAEA.taxon	discussion	Remarks. Although the nominotypical subspecies is rather variable, it may be recognized by the pronotum with sides more or less parallel and the body comparatively small. According to the original description (Tschitschérine 1894), H. przewalskyi has been described from many specimens collected by N. Przewalsky in China in 1884 – 1885 which were labelled " Amdo, 18. IV. – 11. V. 1884, N. Przewalsky ” (= area between Dulan and Barun, Qinghai) and " Turkestan oriental: Tschertschen [= Qarqan], 11. IV. – 22. V. 1885, N. Przewalsky ” (= Quemo, Xinjiang). Consequently, designation of the male labelled " Amdo [Njam-tzu environs, Dy Qu, source of Yangtze River], 10 – 18. VI. 1884, Przevalsky ” [= Zhi Qu (= Tongtian He), area SE of Qumarleb] and " Harpalus Przewalskyi T. Tsch., n. sp., 1894, Typ! " as the lectotype of H. przewalskyi (Kataev 1989: 223) is invalid since Tschitscherine did not list this specimen among syntypes of H. przewalskyi although labelled it as one of the types of this species (Article 74.2, The International Code of Zoological Nomenclature, Fourth Edition 1999). It should be recorded that this specimen is the single one of all series housed in ZISP and labelled by Tschitschérine as H. przewalskyi, which belongs to the subspecies angustibasis ssp. n. (this specimen is included in the type series of this new subspecies, see below). All other specimens with Tschitschérine’s determinational label " Harpalus Przewalskyi T. Tsch., n. sp., 1894, Typ! ", including all real syntypes, are members of the nominotypical subspecies with the pronotal sides more or less parallel basally and the elytral intervals convex apically. Their body length is 8.2 – 8.5 mm. Here we designate the lectotype of H. przewalskyi from the specimens collected by Przewalsky in 1884 during the period from April 18 till May 11 in the area between Dulan and Barun, Qinghai (see Type material). Interestingly, Tschitschérine (1894) in the original description compared H. przewalskyi only with H. smaragdinus (Duftschmid) but later, according to his additional determination labels on the syntypes, treated his taxon as conspecific with H. fuscipalpis Sturm sensu lato (including H. viridanus as one of its varieties). Harpalus fuscipalpis var. versicolor was described from " Ostsibirien ” and “ Mongolei ". According to the original description (Reitter 1900), this form is characterized by reddish legs combined with black, blue, green or bronze dorsum. Although this taxon was for a long time correctly regarded as a synonym of H. viridanus, a lectotype was not yet designated. Here we designate the male from " Quellgebiet des Irkut, Leder " as the lectotype of Harpalus fuscipalpis var. versicolor (see Type material).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB3FFF3FF178D97369EFD41.taxon	materials_examined	Type material. Holotype: ɗ, China, Garzê Co., 3650 m, 11. VII. 1983, Chen Yuanqing leg. (IZB). Paratypes: China. SICHUAN: 5 ɗ, 2 Ψ, same data as holotype (IZB); 5 ɗ, 2 Ψ, Garzê Co., 3400 m, 29. VI. 1983, Zhang Xuezhong (4 specimens), Wang Shuyong (2 specimens) and Chai Huaicheng (1 specimen) leg. (IZB, ZISP); 1 ɗ, same but 3300 m, 17. VI. 1983, Chai Haicheng leg. (IZB); 1 Ψ, NW Sichuan, Maniganggo western environs, 31 ° 50 ' N 99 ° 07 ' E, cca 4100 m, grassland, 17. VII. 1998, J. Turna leg. (cITO). QING- HAI: 1 ɗ, labelled " Amdo [Njam-tzu environs, Dy Qu, source of Yangtze River], 10 – 18. VI. 1884, Przevalsky " [= Zhi Qu (= Tongtian He), area SE of Qumarleb], " Harpalus Przewalskyi T. Tsch., n. sp., 1894, Typ! ", and " v. viridanus Motsch., Tschitscherin det. " [erroneously designated as lectotype of H. przewalskyi by Kataev in 1989] (ZISP); 1 ɗ, 2 Ψ, Yushu Co., 3750 m, 9. VI. 1964, Wang Shuyong leg. (IZB). TIBET: 3 ɗ, Markam Co., 4030 m, 12. VIII. 1982, Zhang Xuezhong leg. (IZB); 1 ɗ, same but 3700 m, 11. IX. 1981, Wang Shuyong leg. (IZB); 2 ɗ, same but 30. VII. 1984, Jiang Ba leg. (IZB); 1 Ψ, Qamdo Co., 3200 m, 22. VI. 1976, no collector (IZB); 1 ɗ, E Tibet, " Lao Shan " pass, road Markam – Zogang, 10 km W Markam, 4300 m, 29 ° 42 ' N 98 ° 32 ' E, alpine meadow, 27 – 28. VI. 1997, J. Turna leg. (cSCI).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB3FFF3FF178D97369EFD41.taxon	description	Description. Comparatively large or medium-sized (body length 8.8 – 10.0 mm), often with bright green, blue or coppery metallic lustre on dorsum. Head relatively large (WHmax / WPmax = 0.63 – 0.71 in male and 0.65 – 0.67 in female; WHmin / WPmax = 0.56 – 0.62 in male and 0.59 – 0.60 in female). Pronotum (Figs 7 – 8) with comparatively narrow base (WPmax / WPmin = 1.07 – 1.15) and with sides more or less strongly converging posteriad in basal half. Pronotal basal angles obtuse, narrowly rounded at apices. Elytra with superficial striae and flat intervals. Dorsal microsculpture visible throughout on head, pronotum and elytra, consisting of distinct, more or less isodiametric meshes. Median lobe of aedeagus (Figs 11 – 13) relatively large (LB / LML = 3.16 – 3.46); terminal lamella (Fig. 13) comparatively long, with sides more or less notably sinuate before apex.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB3FFF3FF178D97369EFD41.taxon	distribution	Distribution. Occurs in southern Qinghai, eastern Tibet and northwestern Sichuan.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB3FFF3FF178D97369EFD41.taxon	etymology	Etymology. The subspecies epithet refers to the shape of pronotum of this taxon.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB3FFF3FF178D97369EFD41.taxon	discussion	Remarks. Rather distinct form readily recognized by the pronotum clearly narrowed basad and the elytra with superficial striae and absolutely flat intervals even apically.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB2FFFCFF178E863435FCD9.taxon	materials_examined	Type material. Syntypes: 14 ɗ, 17 Ψ labelled " Minchow, China ", " Cotype ", " viridanus Mots., s. staudingerianus Schaub. ", " W. H. Muche, Radeberg, Ankauf ", " Staatl. Museum für Tierkunde, Dresden " (MTD). Additional material examined. China. GANSU: 1 ɗ, 2 Ψ, Minxian Co., 20. VII. 1987, Zhong Tiesen leg. (IZB); 1 ɗ, " Kan – ssu [" Pag. Sche-pu et fl. Po-i-cho, 1937 m ", NE of Nanping near border with Sichuan], 1885, G. Patanin / 13. VII " (ZISP); 1 ɗ, " Kan-ssu [= Jonê, ca 50 km NW of Minxian], 1885, G. Patanin / 10. VI " (ZISP); 1 ɗ, " Kan-ssu, 1885, G. Patain / 18. VI " [= Dangchang, ca 80 km SE of Minxian] (ZISP).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB2FFFCFF178E863435FCD9.taxon	description	Description. Very large: body length 8.7 – 11.0 (usually above 10.0) mm, with or without metallic lustre on dorsum. Head relatively small (WHmax / WPmax = 0.60 – 0.63 in male and 0.60 – 0.65 in female; WHmin / WPmax = 0.53 – 0.55 in male and 0.53 – 0.57 in female). Pronotum (Fig. 9) with comparatively wide base (WPmax / WPmin = 1.03 – 1.08) and with sides more or less parallel basally, sometimes slightly converging posteriad. Pronotal basal angle right-angled or slightly obtuse, usually with somewhat sharp or slightly blunt apex. Elytral striae deepened and intervals convex, particularly markedly near apex. Dorsal microsculpture visible throughout on head, pronotum and elytra, consisting of distinct, more or less isodiametric meshes. Median lobe of aedeagus (Figs 11 – 12) medium – sized (LB / LML = 3.30 – 3.54); terminal lamella (Fig. 14) comparatively long, with sides more or less notably sinuate, sometimes nearly parallel before apex.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB2FFFCFF178E863435FCD9.taxon	distribution	Distribution. This subspecies seems to have a small range in southernmost Gansu.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB2FFFCFF178E863435FCD9.taxon	discussion	Remarks. Described from more than 50 specimens captured in Minxian, Gansu, China. In habitus, it is similar to the nominotypical subspecies but distinguished by larger body. This form seems to be distinct although probably is connected with the nominotypical subspecies by the narrow transition zone to the north of Minxian.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBDFFFFFF178B2A3728FEF9.taxon	materials_examined	Type material. Holotype: ɗ, China, Sichuan, Garzê Co., 3650 m, 11. VII. 1983, Chen Yuanqing leg. (IZB). Paratypes: SICHUAN: 6 ɗ, 6 Ψ, same data as holotype but Chen Yuanqing & Wang Shuyong leg. (IZB, ZISP); 1 ɗ, same locality, 3400 m, 29. VI. 1983, Chen Yuanqing (IZB); 1 Ψ, same but Wang Shuyong leg. (IZB); 1 Ψ, Barkam Co., 2700 m, 7. VIII. 1983, Wang Ruiqi leg. (IZB); 1 ɗ (teneral), Dêgê Co., 3100 – 3250 m, 4. VII. 1983, Zhang Xuezhong leg. (IZB); 2 ɗ, Dêgê Co., 3200 m, 6. VII. 1982, Nin Chunlai leg. (IZB); 1 Ψ, same locality, 3100 – 3300 m, 4. VII. 1983, Nin Chunlai leg. (IZB). QINGHAI: 1 Ψ, Yushu Co., Baitang Township, 4200 – 4500 m, 26. VI. 1964, Wang Shuyong leg. (IZB); 1 ɗ, same locality, 4100 m, 13. VI. 1964, Wang Shuyong leg. (IZB).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBDFFFFFF178B2A3728FEF9.taxon	description	Description. Body length 6.2 – 8.7 mm, width 2.7 – 3.8 mm. Colour: Dark brown to black, weakly shiny on dorsum; tarsi slightly paler; first antennomere, apices of other antennomeres, and apices of palpomeres reddish brown. Pronotum usually not translucent at sides. Head: Medium – sized (WHmax / WPmax = 0.62 – 0.65 in male and 0.62 – 0.67 in female; WHmin / WPmax = 0.53 – 0.55 in male and 0.53 – 0.58 in female), with rather weakly convex eyes (WHmax / WHmin = 1.14 – 1.22), separated from buccal fissure ventrally by distance notably shorter than width of first antennomere. Tempora short and flat. Dorsal microsculpture visible throughout, fine, consisting of isodiametric meshes. Mental tooth distinct, moderately wide, acute at apex. Antennae comparatively short, with apex slightly extending beyond elytral base. Pronotum (Figs 17 – 18): Relatively large, 1.40 – 1.56 times as wide as long, widest before middle (WPmax / WPmin = 1.04 – 1.12), comparatively weakly narrowed basad, with one lateral seta on each side. Apical margin weakly emarginate, bordered only laterally. Apical angles weakly protruding and widely rounded. Basal margin more or less straight, throughout bordered, notably wider than apical margin; basal edge glabrous. Sides more or less clearly sinuate in basal half. Disc moderately convex, slightly obliquely depressed at basal angles. Basal foveae narrow, usually more or less deep and touching base of pronotum, with several coarse punctures inside; often few sparse and coarse punctures present also outside of basal foveae, mainly at basal angles and along sides in basal half; area between basal foveae convex, usually impunctate. Microsculpture throughout distinct, consisting of more or less isodiametric meshes. Elytra: Moderately convex, 1.33 – 1.44 times as long as wide, 2.28 – 2.44 times as long and 1.12 – 1.20 times as wide as pronotum, widest slightly behind middle, widely rounded at sides. Base of elytra slightly wider than pronotal base. Humeri angulate, obtuse, each with a small acute denticle visible dorsally. Preapical sinuation distinct, but not deep, without denticle at its base. Sutural angle acute, rather sharp at apex. Basal edge glabrous, sinuate laterally, forming a distinct, almost right angle with lateral margin. Striae impunctate, throughout superficial or slightly impressed apically. Scutellar striole long, with a basal pore. Intervals weakly convex basally and slightly more strongly convex apically, sometimes intervals almost flat. Third interval with a small discal pore in apical third. Fifth and seventh intervals lacking rows of pores before their apices. Series umbilicate irregular, not interrupted medially. Microsculpture very distinct throughout, isodiametric, nearly granulate. Wings: Fully developed. Ventral surface: Prosternum finely pubescent medially. Proepisterna impunctate. Metepisterna (Fig. 20) longer than wide, notably narrowed posteriad. Three last abdominal sternites glabrous, only with obligatory fixed setae. Anal sternite without pronounced sexual dimorphism, rounded at apex in both sexes. Legs: Metacoxae finely punctate and pubescent medially. Protibia widened apically, with one ventroapical spine, without prominent ventroapical tubercle in male and with three preapical spines at outer margin isolated from spines on ventral surface. Metafemur ventrally with three, sometimes four, setigerous pores along posterior margin and with one to three pores at anterior margin in its apical portion. Tarsi impunctate and glabrous dorsally. Metatarsus longer than WHmin but shorter than WHmax; metatarsomeres with sides diverging apicad; first metatarsomere slightly longer than second, much shorter than second and third combined together. Female genitalia (Fig. 19): Hemisternite comparatively narrow, with two short setae distally. Basal stylomere with two setae distally. Apical stylomere evenly arcuate, relatively narrow, with several basal setae on external dorsal margin. Male genitalia: Median lobe of aedeagus (Figs 21 – 24) comparatively large (LB / LML = 3.45 – 3.70), arcuate, bent between basal bulb and middle of median lobe. Terminal lamella much longer than wide, slightly widened apicad, with oblique, horseshoe – shaped apical capitulum. Apical orifice shifted to left. Internal sac with very narrow, winding spiny patches on left side and with a very small and short separate spine in apical portion of median lobe.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBDFFFFFF178B2A3728FEF9.taxon	distribution	Distribution. North-western Sichuan and southern Qinghai.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBDFFFFFF178B2A3728FEF9.taxon	etymology	Etymology. The species epithet refers to the close relationship of the new species to H. sinuatus.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBDFFFFFF178B2A3728FEF9.taxon	discussion	Remarks. This new species is very similar in habitus to H. sinuatus Tschitschérine, but clearly distinguished from it by the following character states: coloration darker (pronotum usually not or only slightly translucent at margins), body stouter, eyes less convex (in H. sinuatus, WHmax / WHmin = 1.22 – 1.31), elytra, on average, relatively shorter and wider (in H. sinuatus, LE / WE = 1.42 – 1.51, LE / LP = 2.43 – 2.56, and WE / WPmax = 1.15 – 1.20), metepisterna shorter, median lobe of aedeagus relatively larger and less arcuate, bent between basal bulb and middle of median lobe (in H. sinuatus median lobe bent almost in middle and LB / LML = 4.00 – 4.35). It should be noted also that the geographical ranges of both species are widely separated: H. sinuatus is distributed to the north of the geographical range of H. parasinuatus sp. n., over southern Siberia from Altai to Amur Province of Russia, also in central and eastern Mongolia (Selenga, Central, Sukhe – Bator, and East aimaks) and the northeastern part of China (Heilongjiang, Liaoning, Shanxi, Inner Mongolia, and Ningxia provinces). Based on the fact of the geographical vicariance and because the armature of the internal sac is very similar in both species, it is possible that H. sinuatus and H. parasinuatus sp. n. are only subspecies of one species. Now we prefer to treat them as two separate species since the morphological differences between them are distinct and populations with intermediate characters are unknown. Within the genus Harpalus, H. sinuatus and H. parasinuatus sp. n. constitute a separate species group (the sinuatus group) together with H. nigrans Могawitz which, like H. sinuatus, is widely distributed over southern Siberia, Mongolia and northeastern part of China. The members of this group are very similar in the morphological characteristics to the species of the vittatus group (see Kataev 1990), but distinctly differ from them in having the antennae reddish brown, only weakly infuscate on 2 nd – 7 th antennomeres, the protibia of male without prominent ventroapical tubercle, and the aedeagus with different pattern of spiny patches in the internal sac. H. nigrans is easily distinguished from H. parasinuatus sp. n. and H. sinuatus by the densely and coarsely punctate base of pronotum.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBEFFFFFF17899237D8FBCF.taxon	discussion	Remarks. H. kaznakovi was described from the series collected at several localities of northwestern Sichuan (mainly the district of Sertar). This species belongs to the vittatus species group (Kataev 1990) which is characterized by the antennae clearly infuscate from first or second antennomere, the male protibia with prominent ventroapical tubercle bearing one spine at apex, the abdominal sternites usually glabrous, the last abdominal sternite without pronounced sexual dimorphism and the internal sac of the aedeagus with one to three small and narrow median spiny patches and without separate large teeth. H. kaznakovi may easily be distinguished from other related species by the short metepisterna and usually the metallic tinge on dorsum. The study of additional material from Sichuan and Tibet showed that the species is very variable and like many other apterous species consists of several morphologically more or less distinct geographical forms. Although some of these forms at first glance are rather markedly distinguished from each other, they all have very similar male genitalia and most of the distinctive characters appear to demonstrate a clinal variation. On the other hand, the specimens examined from eastern and central Tibet are isolated geographically and differ notably from all other material in shape of metepisterna and some other characters. In our opinion, these specimens represent a taxon which warrants at least a subspecific status. Further investigation of the geographical and intrapopulational variation of H. kaznakovi is much needed.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBEFFF9FF178C6F34BFFC51.taxon	materials_examined	Material examined. In addition to the type series the following specimens were examined: China. SICHUAN: 1 ɗ, same data as holotype: NW Sichuan, road Luhuo – Sertar, pass 35 km NNE Luhuo, alpine region, 3500 – 4000 m, 27. VII. 1994, L. Kalab leg. (cFCCH); 2 ɗ, 2 Ψ, Garzê Co., 3400 m, 29. VI. 1983, Chen Yuanging & Zhang Xuezhong leg. (IZB); 1 ɗ, same but 3300 m, 30. VI. 1983, Chen Yuanging leg. (IZB); 2 ɗ, 2 Ψ, same but 3650 m, 11. VII. 1983, Wang Shuyong (2 specimens) & Chen Yuanqing (2 specimens) leg. (IZB); 2 ɗ, same but 3600 m, 1. VII. 1983, Cheng Yuanqing & Wang Shuyong leg. (IZB); 3 ɗ, 1 Ψ, Wenchuan, 900 – 1000 m, 1. VIII. 1983, Zhang Xuezhong leg. (IZB); 1 ɗ, W Sichuan, 15 km NE Qianning, 30 ° 35 ' N 101 ° 41 ' E, 11. VII. 1998, M. Bocak leg. (cWR); 1 ɗ, 1 Ψ, Kangding, 3700 m, 30. VIII. and 7. IX. 1982, Wang Shuyong leg. (IZB); 4 ɗ, 2 Ψ, Barkam Co., 2500 m, 20. VIII. 1983, Wang Ruqi leg. (IZB); 1 ɗ, Dêgê Co., 3900 m, 6. VII. 1983, Wang Shuyong leg. (IZB); 1 Ψ, Dêgê Co., Maniganggo, 3900 m, 8. VII. 1983, Chen Yuanqing leg. (IZB); 2 Ψ, Dêgê Co., 3800 m, 9. VII. 1983, Wang Shuyong leg. (IZB); 2 ɗ, 2 Ψ, same but 3900 m, 10. VII. 1983, Zhang Xuezhong leg. (IZB); 4 ɗ, 1 Ψ, same but 4200 m, 10. VII. 1983, Niu Chunlai (3 specimens) and Chai Huaicheng (2 specimens) leg. (IZB); 1 Ψ, same but 4000 m, 8. VII. 1983, Wang Shuyong leg. (IZB). TIBET: 1 Ψ, Jomda Co., 3400 m, 27. VII. 1976, Zhang Xuezhong leg. (IZB).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBEFFF9FF178C6F34BFFC51.taxon	description	Description. Habitus: Figs 25 – 26. Metallic tinge on dorsum variable, sometimes recognizable only along margins of pronotum and elytra. Body size very variable: body length 5.9 – 8.4 mm. Head moderate or relatively large (in male WHmax / WPmax = 0.58 – 0.72 and WHmin / WPmax = 0.53 – 0.61), with convex or almost flat eyes (WHmax / WHmin = 1.04 – 1.25). Pronotum variable in proportions and shape, 1.44 – 1.59 as wide as long, with sides rounded or straight, converging posteriad in basal half, sometimes sinuate; basal angles obtuse, somewhat sharp or rounded at apices; basal foveae small, variable in shape, usually elongate and not deep; base of pronotum finely punctate or almost smooth, only with few fine punctures in basal foveae. Elytra relatively short or elongate, 1.26 – 1.42 times as long as wide, 2.14 – 2.43 times as long and 1.09 – 1.21 times as wide as pronotum; parascutellar pore present or absent; each third interval with or without one discal pore. Metepisterna (Fig. 27) wider than long, weakly narrowed posteriad. Metacoxae usually without additional setigerous pores, occasionally with a small such pore medially. Median lobe of aedeagus (Figs 30 – 31, 34 – 37, 39 – 40) more or less strongly bent after basal bulb; terminal lamella (Figs 32 – 33, 38, 41) comparatively long Distribution. Known from north-western Sichuan and eastern Tibet (Jomba County). Variation. The nominotypical subspecies demonstrates geographical variation in several characters, mainly in body size and shape of eyes and pronotum, and apparently represents a complex of several local forms which however seem to be not sharply separated from each other. At least based on the scarce available material delimitation of these forms involves big difficulties. In Sichuan, the specimens from Garzê County, Wenchuang and Kangding are similar to those of the type series; they are, on average, rather large (6.1 – 8.2 mm) and robust but eyes are only scarcely convex, almost flat (Fig. 25; WHmax / WHmin = 1.04 – 1.10). The median lobe of aedeagus of these specimens is usually almost straight from dorsal view or only slightly curved to right (Figs 30 – 31, 34 – 35, 36 – 37); in several specimens from Garzê County the median lobe is narrower and less strongly arcuate (Figs 34 – 35) than that in most other specimens. The pronotum of the specimens from Garzê County, Wenchuang and Kangding is usually with sides rounded or straight, converging posteriad in basal half, sometimes sinuate (more markedly so in the specimens from Wenchuan), and basal angles obtuse and rounded at apices. In one male examined from the environs of Qianning, eyes are scarcely convex but pronotal sides clearly sinuate basally and basal angles nearly right and rather sharp at apices. By contrast, in the specimens from Barkam and Dêgê counties (Sichuan) as well as Jomba County (Tibet), body is, on average, smaller (5.9 – 7.0 mm) and more slender and eyes are more convex (Fig. 26; WHmax / WHmin = 1.11 – 1.25) than those in the most specimens from Garzê, Wenchuang and Kangding counties. The median lobe of aedeagus of these specimens is markedly arcuate and with apical portion more strongly curved to right (Figs 39 – 40). The armature of the internal sac in all forms is very similar.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBEFFF9FF178C6F34BFFC51.taxon	discussion	Remarks. Although the nominotypical subspecies is highly variable, it may be recognized by the short and wide metepisterna, the terminal lamella of the median lobe of aedeagus nearly parallel – sided and the body larger and more robust.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFFBFF178F673486FD69.taxon	discussion	Remarks. This species is a member of the Transpalaearctic anxius species group (Kataev 1989) which is characterized by the pronotum impunctate basally and with basal edge glabrous, the protibia with one ventroapical spine and without prominent ventroapical tubercle in male, the abdominal sternites usually glabrous, the last abdominal sternite without pronounced sexual dimorphism and the internal sac with very characteristic longitudinal spiny patch on left side and usually with two small separate teeth. H. giacomazzoii is the single species of this group distributed in central China and may be distinguished from other related species by the large body, the rounded pronotal basal angles, the short metepisterna and the internal sac of the aedeagus without or with very small separate teeth. This species has been described from the several specimens collected in two localities, both within western Sichuan, China: Batang and 50 km to east of Batang. As noted in the original description (Kataev & Wrase 1996), the specimens from each of these localities differ slightly in body size, shape of pronotum and details of the internal sac of the median lobe of aedeagus. The new material examined from Sichuan and Tibet revealed the marked morphological variability of H. giacomazzoi both within one population and geographically. The species seems to consist of several morphologically recognizable allopatric populations some of which may be treated as subspecies. Based on the available material, now we divide H. giacomazzoi into two subspecies treating the particularly remarkable specimens from Tibet as belonging to a separate subspecies. Further study of the variability of H. giacomazzoi based on more extensive material is necessary.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFF9FF178BCA31CDF8C7.taxon	materials_examined	Type material. Holotype: 1 ɗ, China, Tibet, Zogang Co., 3800 m, 3. VII. 1976, Zhang Xuezhong leg. (IZB). Paratypes: 1 ɗ, same data as holotype, 2. VII. 1976 (IZB); 2 ɗ, Lhünzhub Co., 4200 m, 13. VI. 1960, Wang Chunguang leg. (IZB, ZISP).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFF9FF178BCA31CDF8C7.taxon	description	Description. Size very small: body length 5.2 – 5.8 mm. Habitus: Fig. 28. Head relatively large (in male WHmax / WPmax = 0.69 – 0.71 and WHmin / WPmax = 0.58 – 0.60), with convex eyes (WHmax / WHmin = 1.19 – 1.20). Pronotum relatively narrow, 1.42 – 1.50 as wide as long, with sides converging posteriad in basal half and basal angles rounded at apices; basal foveae small and narrow; base of pronotum usually almost impunctate, at most with few very fine punctures around basal foveae and along lateral furrows basally. Elytra relatively long, 1.32 – 1.42 times as long as wide, 2.40 – 2.62 times as long and 1.17 – 1.34 times as wide as pronotum; humeral denticle small; preapical sinuation moderate, parascutellar pore present; each third interval with one discal pore. Metepisterna (Fig. 29) approximately as long as wide and more strongly than in H. kaznakovi kaznakovi narrowed posteriad. Metacoxae without additional setae. Median lobe of aedeagus (Figs 42 – 43) notably bent just after basal bulb; terminal lamella (Fig. 44) comparatively short, with sides usually slightly converging apicad.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFF9FF178BCA31CDF8C7.taxon	distribution	Distribution. This new subspecies occurs in eastern and central Tibet.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFF9FF178BCA31CDF8C7.taxon	etymology	Etymology. The subspecies epithet refers to the small body of this new taxon.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFB8FFF9FF178BCA31CDF8C7.taxon	discussion	Remarks. The new subspecies distinctly differs from the nominotypical one in the narrower metepisterna, the smaller body and the terminal lamella of aedeagus more clearly converging apicad.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBAFFE4FF178B023696FD41.taxon	materials_examined	Type material. Holotype: ɗ, China, Sichuan, road Litang – Batang, 50 km E Batang, 14 – 16. VII. 1992, E. Giacomazzo leg. (cWR). Paratypes: 1 ɗ, same data as holotype (cSCI); 1 ɗ, 1 Ψ, Batang, 2500 m, 15. VII. 1992, E. Giacomazzo leg. (cSCI). Additional material examined. China. SICHUAN: 5 ɗ, 2 Ψ, Dêgê Co., 3100 – 3300 m, 4. VII. 1983, Niu Chunlai (6 specimens) and Wang Shuyong (1 specimen) leg. (IZB); 1 Ψ, Dêgê Co., 3200 m, 4. VII. 1983, Chen Yuanqing leg. (IZB); 7 Ψ, Batang, 2500 m, 13. VIII. 1982, Wang Shuyong leg. (IZB); 23 ɗ, 6 Ψ, " Yun-nan: Tsekou et Se-Tchouen, Principante de Batang: Yargong, J. A. Soulie, 1907 " (MNHN, ZISP); 5 ɗ, 5 Ψ, Xiangcheng Co., 2900 m, 18. VI. 1982, Wang Shuyong (8 specimens) and Zhang Xuezhong (2 specimens) leg. (IZB). Questionable locality (probably mislabelled and actually Sichuan): 1 ɗ, 1 Ψ, " Peking, Westberge, Exp. Stötzner ", " type " [male] or " cotype " [female], " pekinensis Schaub. " [according to our data, this taxon has never been described by E. Schauberger] (OÖLL).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBAFFE4FF178B023696FD41.taxon	description	Description. Size variable: body length 7.8 – 10.8 mm. Head moderate (WHmax / WPmax = 0.55 – 0.63 and WHmin / WPmax = 0.46 – 0.52). Pronotum variable in form, from nearly oval to more or less trapezium – shaped, but in all cases more strongly narrowed apicad than basad and with base notably wider than apex; basal angles narrowly or widely rounded. Elytra 1.33 – 1.45 times as long as wide, 2.28 – 2.57 times as long and 1.07 – 1.16 times as wide as pronotum; preapical sinuation weak or moderately deep. Wings reduced to scale extending to about 1 / 3 – 3 / 4 of elytra. Three last abdominal sternites glabrous or with scattered additional short setae. Metacoxae with or without additional setae medially. Metafemur with 4 – 10 setigerous pores along posterior margin. Median lobe of aedeagus with terminal lamella notably widened apically (Fig. 47, 50); internal sac with moderately wide longitudinal spiny patch and with or without two extremely small separate teeth (Figs 45 – 46, 48 – 49).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBAFFE4FF178B023696FD41.taxon	distribution	Distribution. The nominotypical subspecies is distributed in western part of Sichuan.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBAFFE4FF178B023696FD41.taxon	description	Variation. In the specimens from Dêgê County, the pronotum is only slightly narrower than elytra (WPmax / WE = 1.07 – 1.08), the metacoxae are with many additional setae medially, the abdominal sternites are covered with scattered additional short setae, the metafemur is with 8 – 10 setigerous pores along posterior margin and the aedeagus has separate teeth in the internal sac larger than in the specimens from other known localities (Figs 45 – 46). In the specimens from Batang County including the type series, the pronotum is variable but usually notably narrower than elytra (WPmax / WE = 1.09 – 1.16), the metacoxae are with or without additional setae medially, the abdominal sternites are usually glabrous, at most with few very short setae on three penultimate sternites, the metafemur possesses 4 – 7 setigerous pores along posterior margin and the separate teeth in the internal sac of the aedeagus are small or absent (Figs 48 – 49). Among the specimens from Batang County, the specimens from “ Yargong ” are characterized by deeper preapical sinuation of elytra. The specimens examined from Xiangcheng County and “ Peking ” are similar to those from Batang County (WPmax / WE = 1.10 – 1.16) but the metacoxae seem to be constantly without additional setae.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFBAFFE4FF178B023696FD41.taxon	discussion	Remarks. The most important distinctive characters of the nominotypical subspecies are the pronotum with comparatively wide base and the terminal lamella of the aedeagus notably widened apically. This taxon is highly variable in many of its morphological characteristics and it is possible that it will be divided into two or several subspecies when more populations will be examined.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE5FF17888237F9FC77.taxon	materials_examined	Type material. Holotype: ɗ, China, Tibet, Markam Co., 3700 m, 11. IX. 1981, Wang Shuyong leg. (IZB). Paratypes: 3 ɗ, 2 Ψ, same data as holotype, Wang Shuyong (4 specimens) and Liao Subai (1 specimen) leg. (IZB, ZISP).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE5FF17888237F9FC77.taxon	description	Description. Medium sized: body length 7.5 – 8.8 mm. Head relatively large (WHmax / WPmax = 0.64 – 0.67 and WHmin / WPmax = 0.52 – 0.56). Pronotum comparatively small, rounded at sides, almost evenly narrowed apicad and basad, with base slightly wider than apex; basal angles widely rounded. Elytra relatively long, 1.36 – 1.40 times as long as wide, 2.45 – 2.52 times as long and 1.11 – 1.20 times as wide as pronotum; preapical sinuation weak. Wings reduced to scale extending to about 1 / 3 of elytra. Three last abdominal sternites glabrous. Metacoxae usually without additional setae, occasionally with one medial seta on right or left metacoxa. Metafemur with 4 – 6 setigerous pores along posterior margin ventrally. Median lobe of aedeagus with terminal lamella slightly widened apically (Fig. 53); internal sac with rather narrow longitudinal spiny patch and two extremely small individual teeth (Figs 51 – 52).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE5FF17888237F9FC77.taxon	distribution	Distribution. This new subspecies is known from Markam County, eastern Tibet, China.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE5FF17888237F9FC77.taxon	etymology	Etymology. The subspecific epithet refers to the slender body of the new subspecies.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE5FF17888237F9FC77.taxon	discussion	Remarks. This new subspecies differs from the nominotypical subspecies in the body more slender, the head relatively larger, the pronotum more strongly narrowed basad, and the terminal lamella of the median lobe of the aedeagus less notably widened apically.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE6FF178C173132FCE1.taxon	materials_examined	Type material. Lectotype (present designation): ɗ, labelled " Syao-Chzhin-kho (= Xiaojin) Valley, 27. VII. 93, Potanin " [in Cyrillic], " Potanini m., typ, Tschtscherin det. ", “ Lectotypus, Harpalus potanini Tschit., Z. Mlynář det. 1976 " (ZISP). Paralectotypes: 3 ɗ, 2 Ψ, same data as lectotype (ZISP); 5 Ψ, " Fu-byan-kho (= Fubian) Valley, 31. VII. – 1. VIII. 93, Potanin " [in Cyrillic] (ZISP). Additional material examined. China. SICHUAN: 1 ɗ, Luding Co., 12. VIII. 1981, Shang Jinwen leg. (IZB); 1 ɗ, Kangding Co., 1800 m, 18. VIII. 1981, Shang Jinwen leg. (IZB); 2 ɗ, 1 Ψ, same data but 1500 m (IZB); 1 ɗ, Kangding Co., 1980 m, 22. VI. 1989, Wang Shuyong leg. (IZB).	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE6FF178C173132FCE1.taxon	distribution	Distribution. This species has originally been described from the series collected in two localities of the Central Sichuan, both to north of Kangding (Xiaojin Valley and Fubian Valley). According to the new examined material, H. potanini seems to be endemic to the central Sichuan.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
03E987D1FFA4FFE6FF178C173132FCE1.taxon	discussion	Remarks. H. potanini is very remarkable species and due to the rather flat body with mat dorsum, widely rounded basal angles of pronotum and rather slender legs slightly similar in appearance to some species of the genus Calathus Bonelli belonging to the fuscipes group. The species was described without comparison with any known species and we did not find additional information about its taxonomy or geographical distribution published since the original description. In our opinion, H. potanini is indeed dissimilar to any known species of the genus and seems to represent a separate species – group within Harpalus s. str. with the following most important distinctive character states: the body length 10.8 – 14.0 mm; the coloration dark, without metallic tinge on dorsum; the head, pronotum, elytra, abdominal sternites and tarsi dorsally glabrous; the pronotal basal margin not ciliate; the elytral basal border glabrous; the elytral preapical sinuation rather deep, with traces of obtuse denticle at its base; the elytral third interval at most with one discal pore; the wings completely reduced; the metepisterna wider than long; the legs comparatively long and slender, with long and narrow, almost parallel – sided metatarsomeres; the protibia with one ventroapical spine and three preapical spines on outer margin isolated from spines on ventral surface; the median lobe of aedeagus with rather long terminal lamella, with oblique horseshoe – shaped apical capitulum and distal orifice shifted to left; the internal sac of aedeagus with two large separate teeth, with a small left spiny patch medially and a larger right spiny patch apically (Figs 54 – 55). Based on the external morphology and especially the male genitalia with characteristic structure of the internal sac, H. potanini probably belongs to the honestus phylogenetic stock (lineage) which includes in addition to the honestus, atratus, aeneipennis, laevipes and major groups distributed in the East Palaearctic (Kataev in Kryzhanovskij et al. 1995: 140) also some species groups endemic to the West Palaearctic. Within this phylogenetic stock, H. potanini seems to be related to the aeneipennis group which includes two apterous species from the West Caucasus: H. aeneipennis (Faldermann) and H. chrysopus Reitter. The species of the aeneipennis group share with H. potanini almost all the distinctive characters of the latter listed above, differing mainly in the smaller and more convex body, the shorter legs and the presence of prominent humeral denticle on elytra. The widely separated distributions of these taxa seem to reflect their relict character. It is known that the species of the aeneipennis group live mainly in deciduous broad – leaved forest that is unusual and probably relict feature for Harpalus most representatives of which occur on open territories. Unfortunately, there is no available information about ecology of H. potanini.	en	Kataev, Boris M., Liang, Hongbin (2007): Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini). Zootaxa 1604: 1-20, DOI: 10.5281/zenodo.178756
