taxonID	type	description	language	source
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	description	Dryini Haliday 1833: 273.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	description	Anteonidae Kieffer 1911 a: 108.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	materials_examined	Type genus. Dryinus Latreille, 1804, by monotypy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	diagnosis	Diagnosis. Hymenoptera Chrysidoidea with antenna composed of 10 antennomeres, inserted on flat or concave surface below middle of head, with torulus a distance less than its width from aboral margin of clypeus; head in lateral view concave or slightly convex anteriorly; ♀ protarsus usually provided with one chela, composed of a laterally protruding fifth tarsomere and one enlarged claw; ♀ protarsus less commonly unmodified (in 94 of 1845 world species).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	biology_ecology	Hosts. Hemiptera Auchenorrhyncha (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013). World species. 1884 species are known, 430 in the Afrotropical region (including four species incertae sedis).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	description	World genera. 50 genera are known, 23 in the Afrotropical region. World subfamilies. 16 subfamilies (12 extant) are known, nine in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFB82A5FFF3ED44EFA9CF9D6.taxon	discussion	Remarks. The following seven subfamilies have not been found in the Afrotropical region: Burmadryininae Olmi, Xu & Guglielmino 2014 (fossil, Burmese amber), Erwiniinae Olmi & Guglielmino 2010 (Neotropical, Olmi & Guglielmino 2010 a), Palaeoanteoninae Olmi & Bechly 2001 (fossil, Baltic amber), Plesiodryininae Olmi 1987 (Nearctic, Olmi 1987 b), Ponomarenkoinae Olmi 2010 (fossil, Baltic and Burmese amber, Olmi 2010 a, Xu et al. 2013), Protodryininae Olmi & Guglielmino 2012 (fossil, Baltic amber), Transdryininae Olmi 1984 (Australian).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8D2A6BFF3ED6D4FA7DFD77.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 15 A, D, 19); head hypognathous; occipital carina complete; ocelli present; frons and partly clypeus without strong median raised carina; clypeus normally large, about as wide as, or smaller or slightly wider than eyes; antenna without ADOs; palpal formula 5 / 2, 5 / 3, 6 / 2 or 6 / 3; forewing usually with costal cell enclosed by pigmented veins (Figs 15 A, D, 18 D), rarely also R cell enclosed by pigmented veins (Figs 18 A, B); forewing with 2 r-rs & Rs vein and pterostigma (Figs 15 A, D, 18 D); course of forewing veins not marked by dark stripes (Figs 15 A, D, 18 D); hind wing (Figs 15 A, D, 19) hyaline, with costal vein, without dark medial longitudinal stripe; protarsus not chelate (Figs 15 A, D, 18 A, B); tibial spurs 1 / 1 / 2. ♂: fully winged (Figs 11 C, 15 B, 19); occipital carina complete; palpal formula 5 / 2, 5 / 3, 6 / 2 or 6 / 3; forewing usually with costal cell enclosed by pigmented veins (Figs 11 C, 15 B, 18 D), rarely also R cell enclosed by pigmented veins (as in Figs 18 A, B; character not recorded in the Afrotropical region); forewing with 2 r-rs & Rs vein and pterostigma (Figs 11 C, 15 B, 18 D); course of forewing veins not marked by dark stripes; hind wing (Figs 11 C, 15 B, 18 D) hyaline, with costal vein, without dark medial longitudinal stripe; basivolsella situated completely below distivolsella apex (Figs 13 A, 20 A); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8D2A6BFF3ED6D4FA7DFD77.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8D2A6BFF3ED6D4FA7DFD77.taxon	biology_ecology	Hosts. Membracidae and Typhlocybinae (Cicadellidae) (Guglielmino & Olmi 1997, 2006; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8D2A6BFF3ED6D4FA7DFD77.taxon	discussion	World species. 95 species are known, nine in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8D2A6BFF3ED6D4FA7DFD77.taxon	discussion	World genera. Two genera are known, both present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	description	(Figs 12 C, D, 16 A – D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	description	Description. ♀ (Figs 16 C, D). Fully winged; body length 1.0 – 1.7 mm. Head black, except mandible testaceous and clypeus occasionally uniformly brown; antenna testaceous-brown, except scape, pedicel and occasionally most part of flagellomere 1 yellow; mesosoma black; metasoma brown; legs yellow, except last tarsomere darkened. Antenna clavate; antennomeres of paratype from Yemen, Al Lahima, in following proportions: 4: 4: 5: 6: 6.5: 5.5: 5: 4.5: 4.5: 8; antennomeres of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, in following proportions: 4: 4: 4: 5: 5.5: 5.5: 5: 5: 4.5: 7. Head dull, granulate; frontal line absent; anterior half of frons with narrow median longitudinal furrow; occipital carina complete; paratype from Yemen, Al Lahima, with POL = 6; OL = 3.5; OOL = 2.5; OPL = 2; TL = 2; specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, with POL = 5; OL = 3; OOL = 2; OPL = 1; TL = 1; greatest breadth of lateral ocelli as long as OOL. Mesoscutum dull, granulate. Notauli short, slightly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels, median area granulate and lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands; in many specimens from Madagascar forewing with one more or less distinct dark spot around 2 r-rs & Rs vein; 2 r-rs & Rs vein regularly curved. Tibial spurs 1 / 1 / 2. ♂ (Figs 16 A, B). Fully winged; body length 1.3 – 1.9 mm. Head black, except mandible testaceous and clyp- eus occasionally uniformly brown; antenna testaceous-brown, except scape, pedicel and occasionally most part of flagellomere 1 yellow; mesosoma black; metasoma brown; legs yellow, except last tarsomere darkened. In two specimens from Madagascar, 23 ° 31.39 ’ S 44 ° 09.16 ’ E, legs yellow, except metacoxa partly darkened. Antenna filiform; antennomeres of holotype in following proportions: 3: 4: 4: 4: 5.5: 6: 6: 5.5: 6: 8; antennomeres of specimen from Cameroon, Messa Bengondo, in following proportions: 4: 4.5: 5: 5: 6.5: 8: 8: 7: 7: 10; antennomeres of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, in following proportions: 3.5: 3.5: 4: 4.5: 5.5: 7: 7: 6.5: 6: 9.5. Head dull, granulate; frontal line absent; occipital carina complete; holotype with POL = 6; OL = 4; OOL = 2.5; OPL = 2; TL = 2; specimen from Cameroon, Messa Bengondo, with POL = 6; OL = 4; OOL = 2; OPL = 2; TL = 2; greatest breadth of lateral ocelli longer than OOL (3: 2). Mesoscutum dull, granulate. Notauli short, slightly visible near anterior margin of mesoscutum. In specimen from Madagascar, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 13 – 16. V. 2001, notauli reaching about 0.3 × length of mesoscutum. In specimens from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, one of two notauli reaching about 0.5 × length of mesoscutum. In specimen from South Africa, 29 ° 56.086 ’ S 29 ° 26.721 ’ E, notauli reaching about 0.6 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels, median area granulate and lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands (in all specimens from Yemen); in many specimens from Madagascar forewing with one more or less distinct dark spot around 2 r-rs & Rs vein; 2 r-rs & Rs vein regularly curved. Basivolsella with 1 – 2 distal bristles situated laterally on area fused with paramere (Figs 12 C, D); distivolsella more or less long (Figs 12 C, D) (figure 1 in Olmi & van Harten 2006 wrong). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	materials_examined	Material examined. Types: ♂ holotype of A. himyarita: YEMEN: Al Lahima, 9. IV – 5. VI. 2001, MT, A. van Harten leg. (MOLC); ♂ holotype of A. incisus: NIGERIA: Abia, Umuahia CRIN (Child Rights Information Network), 9. IV. 1975, J. T. Medler leg. (NHMUK). Paratypes of A. himyarita: same locality label as holotype, 1 ♀, 13 ♂♂ (MOLC). Other material: CAMEROON: East Region, Messa Bengondo, 1 – 15. III. 2003, MT, M. Tussac leg., 3 ♂♂ (2 in HTS, 1 in MOLC). CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 03 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 5 – 6. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 3 – 4. V. 2001, 2 ♂♂ (SAMC); same locality label, 1 – 2. V. 2001, 1 ♂ (SAMC); Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 7 – 8. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 9 ♂♂, 3 ♀♀ (SAMC); Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 22. V. 2001, seep, lowland rainforest, S. van Noort leg., 1 ♀, 2 ♂♂ (SAMC). DEMO- CRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, VII. 2008, MT, T. Chapman leg., 1 ♂ (UCDC); same locality label, 23. IV. 2006, MT along forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (UCDC). IVORY COAST: Vallée du Bandama Region, Bouaké, Mbe. WARDA, MT in rice field, 31. XII. 1991, A. Polaszek leg., 1 ♂ (NHMUK). KENYA: Coast Prov., Taita Hills, Ngangao Forest, 3.36100 ° S 38.34186 ° E, 1848 m, 10 – 24. I. 2012, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 27. VII – 10. VIII. 2011, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Boni Forest area, near Bodhei Village, 1.84752 ° S 40.69307 ° E, 29 m, 6 – 18. VI. 2013, 6 m MT, open canopy forest patch, J. Bukhebi & R. Copeland leg., 3 ♀♀ (NMK); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 3 – 17. V. 2016, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kiboko Sanctuary, 2.20331 ° S 37.71430 ° E, 925 m, 22. IX – 6. X. 2011, MT, edge of indigenous forest, R. Copeland leg., 1 ♂ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 27. V – 3. VI. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Nyanza Prov., Ungoye Field Station, 00 ° 36.91 ’ S 34 ° 05.52 ’ E, 1147 m, 21. IV – 5. V. 2005, MT, R. S. Copeland leg., 1 ♂ (NMK); Rift Valley Prov., Olorgesailie National Monument, 1.57930 ° S 36.44566 ° E, 982 m, 13 – 27. XI. 2011, MT, Acacia-Commiphora savana, R. Copeland leg., 8 ♂♂ (7 in NMK, 1 in MOLC), 1 ♀ (NMK); same locality label, 11 – 25. XII. 2011, 3 ♂♂ (2 in NMK, 1 in MOLC), 1 ♀ (NMK); Rift Valley Prov., Oloitokitok, 2.94456 ° S 37.50714 ° E, 1853 m, 19. VIII – 2. IX. 2011, MT, edge of indigenous forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 11 – 25. XI. 2011, 1 ♀ (NMK); same locality label, 25. XI – 9. XII. 2011, 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest North, Calebs Campground, about 17 km NE Kakamega, 00 ° 22.175 ’ N 34 ° 53.297 ’ E, 5400 feet, XI. 2001, Manfred Kraemer & Co. leg., 1 ♂ (MTC); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, 5400 ft., 2002, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♂ (MTVC). MADAGASCAR: Antananarivo, 3 km 41 ° NE Andranomay, 11.5 km 147 ° SSE Anjozorobe, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 5375, 1 ♂, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 21 – 26. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 26 – 29. I. 2001, MA- 01 - 01 A- 02, 1 ♀ (CAS); same locality label, 19. III – 6. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 09, 2 ♂♂ (CAS); same locality label, 11 – 12. II. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 06, 3 ♂♂ (1 in CAS, 2 in MOLC); same locality label, 5 – 21. IV. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 10, 1 ♂ (CAS); same locality label, 4 – 19. III. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 08, 3 ♂♂ (CAS); An- tsiranana, Montagne d’Ambre National Park, 12 ° 31 S 49 ° 11 ’’ E, 975 m, 19. III – 5. IV. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 07, 11 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 30. V – 6. VI. 2001, MT, MA- 01 - 01 D- 12, R. Harin’Hala leg., 1 ♂ (CAS); same local- ity label, 4 – 19. III. 2001, MA- 01 - 01 D- 05, 2 ♂♂ (CAS); same locality label, 12 – 14. V. 2001, MA- 01 - 01 D- 10, 3 ♂♂ (CAS); same locality label, 21 – 26. IV. 2001, MA- 01 - 01 D- 08, 2 ♂♂ (CAS); same locality label, 6 – 21. IV. 2001, MA- 01 - 01 D- 07, 2 ♂♂ (CAS); same locality label, 29. I – 11. II. 2001, MA- 01 - 01 D- 03, 2 ♂♂ (CAS); same locality label, 11. II – 4. III. 2001, R. Harin’Hala leg., MA- 01 - 01 D- 04, 6 ♂♂ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 27. IV – 13. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 08, M. E. Irwin & R. Harin’Hala leg., 1 ♀, 1 ♂ (CAS); same locality label, 13 – 16. V. 2001, 1 ♂ (MOLC); same locality label, 16 – 31. V. 2001, MA- 01 - 04 B- 10, R. Harin’Hala leg., 1 ♀, 1 ♂ (MOLC); same locality label, 22 – 27. IV. 2001, MA- 01 - 04 B- 07, 1 ♂ (CAS); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 3 ♂♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 14 – 21. II. 2011, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 19, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 4 – 12. II. 2002, MT in high altitude rainforest, R. Harin’Hala leg., 13 specimens (5 ♀♀, 6 ♂♂ in CAS, 1 ♀, 1 ♂ in MOLC); same locality label, 21 – 28. I. 2002, 9 ♀♀ (CAS); same locality label, 22 – 28. XI. 2001, 1 ♂, 1 ♀ (CAS); same locality label, 6 – 15. V. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 15 – 25. VII. 2002, 4 ♀♀ (3 in CAS, 1 in MOLC), 2 ♂♂ (CAS, MOLC); same locality label, 14 – 21. I. 2002, 4 ♂♂ (2 in CAS, 2 in MOLC), 4 ♀♀ (3 in CAS, 1 in MOLC); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 A- 07, 3 ♀♀, 11 ♂♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 5 ♂♂ (4 in CAS, 1 in MOLC), 7 ♀♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 6 ♀♀ (5 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., 3 ♂♂, 17 ♀♀ (CAS); same locality label, 26. II – 10. III. 2003, 2 ♂♂ (CAS, MOLC); same locality label, 19 – 26. II. 2002, 1 ♀ (CAS), 2 ♂♂ (CAS, MOLC); same locality label, 5 – 13. V. 2002, MA- 02 - 09 C- 28, 2 ♀♀ (CAS, MOLC); same locality label, 14 – 22. VII. 2002, MA- 02 - 09 C- 35, 3 ♂♂ (2 in MOLC, 1 in CAS), 1 ♀ (CAS); same locality label, 23. VI – 4. VII. 2002, MA- 02 - 09 C- 33, 2 ♀♀ (CAS); same locality label, 10 – 21. III. 2003, MA- 02 - 09 C- 56, 1 ♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 C- 11, 1 ♀, 2 ♂♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 C- 10, 1 ♂ (CAS); same locality label, 26. II – 4. III. 2002, MA- 02 - 09 C- 18, 1 ♂ (CAS); same lo- cality label, 28. XI – 6. XII. 2001, MA- 02 - 09 C- 05, 2 ♀♀, 7 ♂♂ (CAS); same locality label, 4 – 14. VII. 2002, MA- 02 - 09 C- 34, 2 ♂♂, 3 ♀♀ (CAS); same locality label, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 1 ♂ (CAS); same lo- cality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 3 ♂♂, 6 ♀♀ (CAS); same locality label, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 1 ♂ (CAS); same locality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 1 ♂, 1 ♀ (CAS); same locality label, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♂ (CAS); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 2 ♂♂ (CAS); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 4 ♀♀, 1 ♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 2 ♂♂ (CAS); same locality label, 26 – 31. III. 2002, MA- 02 - 09 C- 22, 1 ♂, 1 ♀ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 2 ♂♂, 4 ♀♀ (CAS); same locality label, 21. III – 12. IV. 2003, MA- 02 - 09 C- 57, 1 ♀, 2 ♂♂ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 5 ♀♀, 12 ♂♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 3 ♂♂, 2 ♀♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, 9 – 20. III. 2003, MT, mixed tropical forest, R. Harin’Hala leg., 7 ♂♂, 3 ♀♀ (CAS); same locality label, 5 – 15. VII. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 9 – 16. IV. 2002, 3 ♀♀ (2 in CAS, 1 in MOLC); same locality label, 4 – 12. III. 2002, 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 B- 15, 2 ♂♂ (CAS); same locality label, 21 – 28. I. 2002, MA- 02 - 09 B- 13, 1 ♀, 1 ♂ (CAS); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 B- 35, 2 ♀♀ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 1 ♂, 2 ♀♀ (CAS); same locality label, 14 – 24. VI. 2002, MA- 02 - 09 B- 32, 1 ♀ (CAS); same locality label, 15 – 27. IV. 2003, MA- 02 - 09 B- 58, 1 ♂, 2 ♀♀ (CAS); same locality label, 12 – 19. III. 2002, MA- 02 - 09 B- 20, 2 ♀♀, 6 ♂♂ (CAS); same locality label, 9 – 20. III. 2003, MA- 02 - 09 B- 55, 4 ♂♂ (CAS); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 1 ♂, 3 ♀♀ (CAS); same locality label, 3 – 15. IV. 2003, MA- 02 - 09 B- 57, 1 ♂ (CAS); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 B- 03, 1 ♀, 5 ♂♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 7 ♀♀, 7 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 6 ♀♀, 8 ♂♂ (CAS); same locality label, 20. III – 3. IV. 2003, MA- 02 - 09 B- 56, 2 ♂♂ (CAS); same locality label, 24. V – 4. VI. 2002, MA- 02 - 09 B- 30, 1 ♀, 2 ♂♂ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 5 ♂♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 4 ♂♂, 6 ♀♀ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 3 ♂♂ (CAS); same locality label, 7 – 17. VII. 2003, MA- 02 - 09 B- 66, 2 ♂♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. XI – 6. XII. 2001, MT near river, R. Harin’Hala leg., MA- 02 - 09 D- 05, 2 ♀♀ (CAS, MOLC), 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 D- 01, 1 ♀ (CAS); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 D- 03, 1 ♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 D- 11, 1 ♀ (CAS); same locality label, 21 – 24. XII. 2001, MA- 02 - 09 D- 08, 3 ♀♀, 4 ♂♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 D- 10, 3 ♀♀ (CAS); Fianarantsoa, Befotaka-Midongy National Park, Papango 27.7 km S Mi- dongy-South Mt. Papango, 23 ° 50.07 ’ S 46 ° 57.49 ’ E, 940 m, 13 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14774, 1 ♀ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 13 – 22. XI. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 68, 1 ♂ (MOLC); Fianarantsoa, Isalo National Park, Sahanafa River, 29.2 km 351 ° N Ranohira, 22 ° 18.48 ’ S 45 ° 17.30 ’ E, 500 m, 10 – 13. II. 2003, MT, gallery forest, BLF 7653, Fisher, Griswold et al. leg., 1 ♂ (CAS); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 2 ♂♂ (CAS); Mahajanga, Ankoririka Reserve, 10.6 km 13 ° NE of Tsaramandroso, 16 ° 16.2 ’ S 46 ° 2.55 ’ E, 210 m, 9 – 14. IV. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3665, 2 ♂♂ (CAS); Mahajanga, Ambovomamy Belembo, 20 km NW Port Berger, 15 ° 27.07 ’ S 47 ° 36.80 ’ E, 33 m, 22 – 27. I. 2007, secondary growth on white sand, MT, R. Harin’Hala, M. Irwin, F. Parker leg., MG- 33 - 04, 1 ♂ (CAS); same locality label, 21 – 22. I. 2007, MG- 33 - 03, 1 ♂ (CAS); Mahajanga, Ankarafantsika National Park, Tsimaloto Forest, 18.3 km 49 ° NE Tsaramandroso, 16 ° 13.41 ’ S 46 ° 08.37 ’ E, 135 M, 2 – 8. IV. 2001, MT in tropical dry forest, Fisher, Griswold et al. leg., BLF 3600, 2 ♂♂ (CAS); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, 43 m, 28. IX – 5. X. 2003, MT in deciduous forest, R. Harin’Hala leg., MA- 25 - 17, 1 ♀ (CAS); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 26. V – 2. VI. 2011, MT, MG- 61 - 15, 5 ♀♀ (3 in CAS, 2 in MOLC), 8 ♂♂ (5 in CAS, 3 in MOLC); same locality label, 13 – 19. XII. 2010, MG- 61 - 01, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 5 – 8. V. 2011, MG- 61 - 11, 3 ♂♂ (MOLC), 1 ♀ (MOLC); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 2 – 8. X. 2001, MT, tropical forest, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 7 – 16. XI. 2001, 1 ♂ (CAS); Toamasina, Anjanaharibe Mt., 19.5 km 27 ° NNE Ambinanitelo, 15 ° 10.42 ’ S 49 ° 38.06 ’ E, 1100 m, MT in montane rainforest, Fisher, Griswold et al. leg., 1 ♂ (CAS); Toamasina, Ankerana, 18 ° 24.37 ’ S 48 ° 49.08 ’ E, 1108 m, 22 – 27. I. 2012, MT, rainforest, B. L. Fisher et al. leg., BLF 27982, 1 ♀ (CAS); Toliara, Sept Lacs, 23 ° 31.39 ’ S 44 ° 09.16 ’ E, 70 m, MT, gallery forest, 9 – 12. III. 2002, California Academy of Sciences, Frontier Project, 2 ♂♂ (CAS); Toliara, Mite Forest, 20.7 km 29 ° WNW Tongobory, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, 75 m, 27. II – 3. III. 2002, MT, in gallery forest, Fisher, Griswold et al. leg., 1 ♂ (CAS); Toliara, Beroboka Forest, 5.9 km 131 ° SE Ankidranoka, 22 ° 13.59 ’ S 43 ° 21.59 ’ E, 80 m, 12 – 16. III. 2002, MT, in tropical dry forest, Fisher, Griswold et al. leg., 2 ♂♂ (CAS); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 1 – 9. VII. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 29, M. Irwin, R. Harin’Hala leg., 1 ♂ (CAS); Toliara, Manombo, 22 ° 48.09 ’ S 43 ° 45.54 ’ E, 227 m, 9 – 14. IV. 2004, MT in dry forest, Frontier Wilderness Project, MGF 102, 6 ♂♂ (4 in CAS, 2 in MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 27. IV – 5. V. 2003, MT in deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 53, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 9 – 23. I. 2003, MT in dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 1 ♂ (CAS). NIGERIA: Oyo, Ibadan, 6. XII. 1962, D. C. Eidt leg., 1 ♂ (CNC); Toliara, Lake Ranobe, 23 ° 02.26 ’ S 43 ° 36.35 ’ E, 30 m, 25 – 30. IV. 2003, MT, spiny forest thicket, MGF 065, Frontier Wilderness Project, 1 ♀ (CAS). SOUTH AFRICA: Eastern Cape, East London, 32 ° 58.9 ’ S 27 ° 53.2 ’ E, 375 ft., 24 – 29. XII. 2004, MT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Coleford Nature Reserve, 16.5 km 216 ° SW Underberg, 29 ° 56.086 ’ S 29 ° 26.721 ’ E, 5 – 8. XII. 2001, MT, moist upland, grassland on sandstone, S. van Noort leg., 3 ♂♂ (2 in SAMC, 1 in MOLC); Western Cape, 45 km SE Sutherland, 32 ° 44.773 ’ S 20 ° 55.624 ’ E, 31. III. 2010, MT, Sweep of green herbaceous verge along dry river course with pools of water, Suculent Karoo, MDK 10 - SUC 1 - S 01, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 30. III – 24. VII. 2010, MT, Renosterveld, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 3 I – 25. IV. 2013, MT, Burnt Mesic Mountain Fynbos, BH 12 - FYN 3 - M 07, S. van Noort leg., 1 ♀ (SAMC). UGANDA: Western Region, Kabarole District, Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.871 ’ S 30 ° 21.355 ’ E, 1495 m, 12 – 26. VIII. 2008, MT, secondary mid-altitude rainforest, S. van Noort leg., 2 ♂♂, 1 ♀ (SAMC); same locality label, 0 ° 33.408 ’ S 30 ° 22.603 ’ E, 1587 m, MT, degraded mid-altitude rainforest, S. van Noort leg., 3 ♀♀, 1 ♂ (SAMC); same locality label, 2 – 12. VIII. 2008, 0 ° 33.784 ’ S 30 ° 22.617 ’ E, 1500 m, MT, primary mid-altitude rainforest, S. van Noort leg., 2 ♀♀, 2 ♂♂ (SAMC); same locality label, 12 – 26. VIII. 2008, 0 ° 33.823 ’’ S 30 ° 21.490 ’ E, 1505 m, MT, primary mid-altitude rainforest, S. van Noort leg., 1 ♂ (SAMC). YEMEN: 12 km NW of Manakhah, 3. VII – 21. VIII. 2001, 15. V – 24. VI. 2003, 4. VIII – 15. IX. 2003, MT, A. van Harten leg., 3 ♀♀, 2 ♂♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	distribution	Distribution. Cameroon, Central African Republic, Democratic Republic of the Congo, Ivory Coast, Kenya, Madagascar, Nigeria, South Africa, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF8B2A60FF3ED4BBFC21FF7C.taxon	discussion	Remarks. Aphelopus incisus Olmi, 1984, is here considered a senior synonym of A. himyarita Olmi & van Harten, 2006, because the length of the notauli is highly variable (previously the two species were separated only by differences in the length of the notauli), and the ♂ genitalia are alike.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF862A64FF3ED0DCFD7FF858.taxon	description	(Figs 12 E, 16 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF862A64FF3ED0DCFD7FF858.taxon	description	883; Olmi & Copeland, 2011: 192; Guglielmino et al. 2013: 79 – 81; Olmi et al. 2015: 337; 2016: 50.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF862A64FF3ED0DCFD7FF858.taxon	materials_examined	Material examined. Types: ♀ holotype of A. mediocarinatus: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Paulis (= Isiro), III. 1947, P. L. G. Benoit leg. (MRAC); ♀ holotype of A. madagascariensis: MADAGAS-CAR: Toliara, III. 1940, A. Seyrig leg. (MNHN). Paratypes of A. madagascariensis: MADAGASCAR: Fianarantsoa, Fianarantsoa, XI. 1936, A. Seyrig leg., 1 ♂ allotype (MRAC). Other material: BOTSWANA: Central District, Serowe, Farmer’s Brigade (USNM); Serowe, III. 1990 (CAS). BURUNDI: Bururi National Forest, 3.93022 ° S 29.61697 ° E, 1955 m, 19. X – 2. XI. 2010, MT, indigenous forest, near stream, R. Copeland leg., 1 ♀ (NMK). CAM- EROON: South Region, near Ebolowa, Nkoemvon, 24. VIII – 7. IX. 1980, cocoa plantation, D. Jackson leg., 1 ♀ (NHMUK); Cameroon (MCZ). CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 03 ° 02 ’ 01 ’’ N 16 ° 24 ’ 57 ’’ E, 510 m, 3 – 4. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♀ (SAMC); Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 22. V. 2001, sweep, lowland rainforest, S. van Noort leg., 1 ♂, 4 ♀♀ (SAMC); Sangha-Mbaéré Pref., Reserve Speciale de Forêt Dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 13. V. 2001, sweep, lowland rainforest, S. van Noort leg., 1 ♀, 4 ♂♂ (SAMC). DEMO- CRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 2. IV. 2006, MT, forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (UCDC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Aff. Butahu, Riv. Katauleko, Kalonge (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo (MRAC); North Kivu, N of Kivu Lake, Rwankwi (MRAC); Haut- Uele (= Haut-Zaire), Garamba National Park (MRAC). ETHIOPIA: Amhara, Zhara Church Forest, 11 ° 47.976 ’ N 37 ° 34.044 ’ E, 1925 m, 6 – 8. I. 2012, forest core, MT, E. McAlister leg., 1 ♀ (NHMUK). GAMBIA: Upper River Divi- sion, Wuli Kantora, Fatoto (KJHC). IVORY COAST: Vallée du Bandama Region, Bouaké, IV. 1980 (CNC). KE- NYA: Coast Prov., Marenje Forest, 4.52814 ° S 39.24028 ° E, 76 m, 25. VII – 8. VIII. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 27 – 30. V. 2013, 6 m MT, in- digenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 27. XII. 2014 – 10. I. 2015, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); same locality label, 5 – 19. XI. 2015, 1 ♂ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 24. VIII – 7. IX. 2011, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♂ (NMK); same locality label, 21. II – 7. III. 2012, 1 ♀ (NMK); same locality label, 27. VII – 10. VIII. 2011, 1 ♀ (NMK); Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 5 – 19. IV. 2012, MT, next to small forest pond, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Matthews Range, 0.97984 ° N 37.34599 ° E, 1459 m, 26. XII. 2015 – 9. I. 2016, MT, riverine forest, near Wamba, R. Copeland leg., 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest Reserve, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, 5400 feet, XI. 2001, Manfred Kraemer & Co. leg., 1 ♂ (MTC). MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW Anivorano Nord, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3011, 2 ♀♀ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 16 – 31. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 10, R. Harin’Hala leg., 1 ♂ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 11. II – 4. III. 2001, MT, MA- 01 - 01 D- 04, R. Harin’Hala leg., 1 ♂ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12.2 km 211 ° SSW of Joffreville, 12 ° 35.47 ’ S 49 ° 9.34 ’ E, 1300 m, 2 – 7. II. 2001, MT, montane rainforest, Fisher, Griswold et al. leg., 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT in high altitude rainforest, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 22 – 28. XI. 2001, 1 ♂ (CAS); same locality label, 19 – 26. II. 2002, 2 ♀♀ (CAS, MOLC); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. I – 4. II. 2002, MT near river, R. Harin’Hala leg., MA- 02 - 09 A- 14, 1 ♀ (CAS); Fi- anarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., 1 ♂ (MOLC); same locality label, 14 – 22. VII. 2002, MA- 02 - 09 C- 35, 1 ♂ (MOLC); same locality label, 10 – 14. I. 2002, MA- 02 - 09 C- 11, 2 ♀♀ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 19, 1 ♂ (MOLC); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 1 ♀ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 1 ♀ (CAS); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, elev 97 m, 27 Jan – 2 Feb 2011, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, M. Irwin, R. Harin’Hala leg, MT, Analagnambe dry degraded forest, MG- 61 - 08, 1 ♀ (CAS). Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., 1 ♂ (CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km 332 ° Toamasina, 17 ° 55.26 ’ S 49 ° 11.59 ’ E, 390 m, 28. XI. 2005, MT, rainforest, Brian L. Fisher et al. leg., BLF 13127, 1 ♂ (CAS); Toliara, 60 km NE of Morondava, Bereboka (MOLC); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 28. III – 8. IV. 2003, MT in transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 20, 2 ♂♂ (CAS, MOLC), 1 ♀ (CAS); same locality label, 30. X – 9. XI. 2003, 2 ♂♂ (CAS, MOLC); same locality label, 26. I – 5. II. 2003, 1 ♂ (MOLC). MOZAMBIQUE: Niassa Prov., Cuamba, Farm of the Catholic University of Mozambique, M. Olmi reared from Molopopterus alfa Dworakowska and Empoascanara ethiopica Dworakowska, parasitized host collected 4. III. 2005, dryinid cocooning 9. III. 2005, dryinid adult emergence 5. IV. 2005, 9 ♀♀, 2 ♂♂ (MOLC), 2 ♀♀ (SAMC). NIGERIA: Abia, Umuahia CRIN (Child Rights Information Network), 9. IV. 1975, J. T. Medler leg., 1 ♂ (NHMUK); Osun, Ile-Ife, X. 1973, J. T. Medler leg., 2 ♂♂ (NHMUK, MOLC); Oyo, Ibadan, I. Amah Pash leg., 2 ♀♀ reared from Empoasca sp. (NHMUK, CNC). SENEGAL: Casamance, 10 km WSW of Velingara (ZIL); Casamance, 20 km N of Kolda (ZIL). SIERRA LEONE: Northern Prov., 5 km W of Kabala, 11 ° 35 ’ W 09 ° 35 ’ N, 28. XI. 1993 (ZIL). SOMALIA: Lower Shebelle, Afgooye (= Afgoi) (AMNH). SOUTH AFRICA: Eastern Cape, Februarie Farm (40.2 km 267 ° W Kirkwood), 33 ° 33.124 ’ S 25 ° 03.043 ’ E, 12 – 14. II. 2001, MT, VB 01 - R 1 N-M 36, Valley bushveld (non-trashed), S. van Noort leg., 2 ♂♂ (SAMC); same locality label, M 22, 10 – 12. II. 2001, 1 ♀ (SAMC); Eastern Cape, Schilpad Laagte Farm (15.4 km 226 ° W Kirkwood), 33 ° 31.653 ’ S 25 ° 22.620 ’ E, 9 – 11. II. 2001, MT, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Marais Hoop Farm (25.6 km 254 ° W Kirkwood), 33 ° 32.635 ’ S 25 ° 13.678 ’ E, 10 – 12. II. 2001, MT, VB 01 - R 4 N-M 26, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♀ (SAMC); Eastern Cape, Somerset East, X. 1930, R. E. Turner leg., 1 ♀ (NHMUK); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 22. II – 9. IV. 2010, MT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♂ (SAMC); same locality label, 22. II – 9. IV. 2010, YPT, 1 ♀ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 23. II – 7. IV. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC); Gauteng, Pretoria, garden, 8 – 11. II. 2005, YPT, V. Kolyada leg., 1 ♂, 2 ♀♀ (NMSA); KwaZulu-Natal, Pietermaritzburg (AEIC); Drakensberg, Van Reenen, 23 – 26. I. 1927, R. E. Turner leg., 1 ♂ (NHMUK); KwaZulu-Natal, Estcourt, Cathedral Peak (UCDC); KwaZulu-Natal, Cathedral Peak Nature Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 3 – 15. XII. 2005, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 23. II – 20. IV. 2007, MT, Kolyada & Mostovski leg., 1 ♀ (NMSA); same locality la- bel, 20. IV – 26. X. 2007, MT, Kolyada & Mostovski leg., 2 ♂♂ (NMSA); KwaZulu-Natal, Royal Natal National Park, Mahai Camp, 28 ° 41.27 ’ S 28 ° 56.86 ’ E, 1450 m, 24. XI – 6. XII. 2005, YPT, M. Mostovski leg., 1 ♂, 1 ♀ (NMSA); same locality label, 25. XI – 12. XII. 2005, MT, 2 ♂♂ (NMSA); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 15 – 30. III. 2006, MT, G. Whiteley leg., 1 ♂ (DJBC); same locality label, 14 – 27. II. 2006, 1 ♂ (DJBC); KwaZulu-Natal, Anyskop Farm, 5.5 km 290 ° W Langebaanweg, 32 ° 57.394 ’ S 18 ° 05.324 ’ E, 2 – 9. X. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Hlogoma, 4.4 km 127 ° SE Underberg, 29 ° 48.308 ’ S 29 ° 32.293 ’ E, 4 – 7. XII. 2001, MT, Moits Upland Grassland on dolerite, livestock grazed, S. van Noort leg., 1 ♂ (SAMC); Limpopo, 30 km E of Nylstroom (PMA); Limpopo, Krüger National Park, Pafuri, Jct. Limpopo River – Levuvhu River (PMA); Limpopo, Trichardtsdal, 15 km W of Malta, 900 m, (PMA); Limpopo, 15 km NW of Klaserie, Guernsey Farm (PMA); Mpumalanga, Krüger National Park, Skukuza (PMA); Mpumalanga, Graskop, 1500 m (PMA); Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.845 ’ S 19 ° 09.443 ’ E, 787 m, 17. III – 21. IV. 2008, MT, Nieuwoudtville Dolerite, Koppie Renosterveld, S. van Noort leg., 1 ♀ (SAMC); same locality label, 22. X – 18. XI. 2008, 1 ♀ (SAMC); Northern Cape, Avontuur Farm, 16 km NW Nieuwoudtville, 1 ° 16.249 ’ S 19 ° 02.900 ’ E, 764 m, 28. IX – 29. XII. 2010, MT, Bokkeveld Sandstone Fynbos, S. van Noort leg., GL 07 - FYN 1 - M 152, SAM-HYM- A 035785, 1 ♀ (SAMC); Western Cape, Mossel Bay, V. 1921, VI. 1921, 1 – 3. VII. 1921, X. 1921, VI – VII. 1930, 28. III. 1939, R. E. Turner leg., 1 ♀, 6 ♂♂ (NHMUK); same locality label, 1 ♂ (AMNH); Western Cape, Ceres, I. 1921, 2 – 21. III. 1921, IV. 1925, R. E. Turner leg., 2 ♀♀, 2 ♂♂ (NHMUK); Western Cape, Cape Town, Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 25. I – 2. II. 1994, MT, Mountain Fynbos on Sandstone, Protea dominated, S. van Noort leg., 1 ♂ (SAMC); same locality label, 12 – 17. IX. 1993, 1 ♂ (SAMC); same locality label, 2 – 9. II. 1994, Mesic Mountain Fynbos, Protea coronata dominated, 1 ♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 14. VI – 12. VII. 1997, 4. X – 1. XI. 1997, 29. XI – 26. XII. 1997, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 1 ♀, 4 ♂♂ (SAMC); same locality label, 11. VIII – 6. IX. 1997, S. van Noort leg., 30 ♂♂, 1 ♀ (SAMC); same locality label, IX – XII. 1997, MT, S. van Noort leg., 6 ♀♀, 10 ♂♂ (SAMC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 6 ♂♂ (SAMC); same locality label, 6. IX – 4. X. 1997, MT, S. van Noort leg., 1 ♀, 2 ♂♂ (CAS); same locality label, 9. VIII – 6. IX. 1997, 1 ♂ (CAS); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 28. XII. 1997 – 21. III. 1998, MT, South Coast Strandveld, 57 m, S. van Noort leg., 2 ♂♂ (SAMC, MOLC); same locality label, 21. III – 18. IV. 1998, 1 ♀ (SAMC); same locality label, 31. V. 1997, sweep, 1 ♀ (MOLC); same locality label, 4. X. 1997, sweep, 1 ♀ (SAMC); same locality label, 21. II – 21. III. 1998, 1 ♂ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37 ’ 622 ’’ S 18 ° 24 ’ 259 ’’ E, 20. III – 17. IV. 1998, MT, S. van Noort leg., 1 ♀ (SAMC): Western Cape, Langberg Farm, 3 km 280 ° W Langebaanweg, 32 ° 58.045 ’ S 18 ° 07.418 ’ E, 9 – 16. X. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 1 ♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 402 ’’ S 18 ° 05 ’ 438 ’’ E, 5.5 km 290 ° W Langebaanweg, 11 – 18. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 817 ’’ S 18 ° 06 ’ 837 ’’ E, 4 km 280 ° E Langebaanweg, 11 – 18. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (MOLC); same local- ity label, 2 – 9. X. 2002, 1 ♀ (SAMC); same locality label, 18 – 25. IX. 2002, 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 10. IX – 4. XI. 2009, MT, Renosterveld, S. van Noort leg., 2 ♀♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, 997 m, 4. X. 2010 – 25. I. 2011, YPT, Rooiberg Sandstone Fynbos, S. van Noort leg., GB 09 - FYN 2 - Y 77, 1 ♂ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.663 ’ S 21 ° 57.600 ’ E, 940 m, 4. X. 2010 – 25. I. 2011, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., GB 09 - FYN 1 - M 52, 1 ♂ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. I – 16. II. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC); same locality label, 16. XI – 16. XII. 1998, 1 ♀ (SAMC); Western Cape, 45 km SE Sutherland, 32 ° 44.773 ’ S 20 ° 55.624 ’ E, 31. III. 2010, MT, Sweep of green herbaceous verge along dry river course with pools of water, Suculent Karoo, MDK 10 - SUC 1 - S 01, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, MT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - M 05, 1 ♂ (SAMC). TANZA- NIA: Arusha, Serengeti National Park Hqts. (USUC); Korogwe, 10 mi. E of Korogwe, 300 m, 11. XI. 1957 (CAS). UGANDA: Wakiso, Entebbe (CNC). YEMEN: Al Kadan, X. 2001, I. 2003, light trap, A. van Harten & T. Abdul-Haq leg., 1 ♀, 1 ♂ (MOLC); Al Lahima, 9. IV – 5. VI. 2001, MT, A. van Harten leg., 1 ♂ (MOLC); Ta’izz, X. 2001, light trap, A. van Harten & A. R. Al Yarimi leg., 1 ♀, 1 ♂ (MOLC). ZIMBABWE: Harare, 23. X. 1970, A. Watsham leg., 1 ♂ (NHMUK); Harare (= Salisbury), Chishawasha, III. 1978, A. Watsham leg., 1 ♂ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF862A64FF3ED0DCFD7FF858.taxon	biology_ecology	Hosts. Typhlocybinae (Cicadellidae) (Guglielmino et al. 2013): in Mozambique: Empoascanara ethiopica Dworakowska and Molopopterus alfa Dworakowska; in Nigeria: Empoasca sp.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF862A64FF3ED0DCFD7FF858.taxon	distribution	Distribution. Botswana, Burundi, Cameroon, Central African Republic, Democratic Republic of the Congo, Ethiopia, Gambia, Ivory Coast, Kenya, Madagascar, Mozambique, Nigeria, Senegal, Sierra Leone, Somalia, South Africa, Tanzania, Uganda, Zimbabwe, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A65FF3ED6D7FE1FFA8D.taxon	description	(Fig. 13 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A65FF3ED6D7FE1FFA8D.taxon	description	Description. ♀. Fully winged; body length 1.6 – 1.9 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; proleg testaceous; mesoleg testaceous, except femur brown; metaleg brown, except coxa and trochanter testaceous. Antenna clavate; antennomeres in following proportions: 5: 4: 4: 4: 4: 4: 4: 4: 4: 6. Head dull, granulate; frontal line incomplete, not present in front of anterior ocellus; occipital carina complete; POL = 5; OL = 3; OOL = 3; OPL = 2; TL = 3; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (3: 2). Occasionally notauli incomplete, reaching about 0.85 – 0.90 × length of mesoscutum. Mesoscutellum and metanotum shiny, slightly granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels and median area less rugose than anterior surface. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curved. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.5 – 1.6 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown. Antenna filiform; antennomeres in following proportions: 5: 4: 4: 4: 4.5: 5: 4.5: 4.5: 4. 5: 7. Head dull, granulate; frontal line incomplete, not present in front of anterior ocellus; occipital carina complete; POL = 5; OL = 3; OOL = 3; OPL = 1.5; TL = 3; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (3: 2). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels and median area shiny, granulate. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curved. Basivolsella (Fig. 13 A) with 2 subdistal bristles and lateral basal process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A65FF3ED6D7FE1FFA8D.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Royal Natal National Park, Gudu Forest, 28 ° 40.90 ’ S 28 ° 55.78 ’ E, 1680 – 1730 m, 18. XI. 2006 – 27. II. 2007, MT, M. Mostovski leg. (NMSA). Paratypes: SOUTH AFRICA: same locality label as holotype, 1 ♀ (NMSA); KwaZulu-Natal, Royal Natal National Park, Mahai Camp, 28 ° 41.27 ’ S 28 ° 56.86 ’ E, 1450 m, 24. XI – 6. XII. 2005, YPT, M. Mostovski leg., 1 ♂ (MOLC); KwaZulu-Natal, Cathedral Peak Nature Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 18. XI. 2006 – 26. II. 2007, MT, M. Mostovski leg., 1 ♂, 2 ♀♀ (NMSA); same locality label, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A65FF3ED6D7FE1FFA8D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A65FF3ED6D7FE1FFA8D.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	description	(Figs 13 B, 14)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	diagnosis	Diagnosis. ♂ of Aphelopus with head mostly black and mesoscutum completely black; notauli reaching approximately 0.6 × length of mesoscutum; basivolsella not fused with paramere, with lateral proximal process; distivolsella with very slender distal part (Fig. 13 B);	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	description	Description. ♂ (Fig. 14). Fully winged; body length 1.9 mm. Head (Fig. 14 C) black, except mandible testa- ceous; clypeus whitish; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs yellow-testaceous, except proximal half of metacoxa brown. Antenna filiform; antennomeres in following proportions: 5: 5: 6: 6: 6: 6: 6: 6: 6: 7. Head dull, granulate; frontal line absent; occipital carina complete; POL = 7; OL = 4; OOL = 3; OPL = 2.5; TL = 4; greatest breadth of lateral ocelli shorter than OL (3: 4); clypeus almost touching antennal toruli. Mesoscutum and mesoscutellum dull, granulate. Notauli incomplete, reaching about 0.6 × length of mesoscutum. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, with transverse posterior keel; propodeal declivity with two complete longitudinal keels. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein curvilinear, shorter than normal 2 r-rs & Rs veins of Aphelopus species; pterostigma brown, as large as in other Aphelopus species. Basivolsella very narrow, with two distal bristles (Fig. 13 B), with lateral proximal process; distivolsella very slender. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	materials_examined	Material examined. Type: ♂ holotype, NMK / INV / T- 637: KENYA: Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 3 – 10. VI. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	etymology	Etymology. This species is named after Mr. Glenn Sequeira, a friend and co-worker of RSC at ICIPE, where he cheerfully and expertly leads the Information Technology unit.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF822A67FF3ED312FA8CFEC4.taxon	discussion	Remarks. This species can be confused with Crovettia species, because the clypeus is almost touching the antennal toruli and the 2 r-rs & Rs vein is shorter than in Aphelopus. However, the 2 r-rs & Rs vein is curvilinear (as in Aphelopus species, though it is shorter than normal) and the pterostigma is less large than in Crovettia species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF802A7AFF3ED3FBFCAFFEC4.taxon	description	(Figs 13 C, 17 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9C2A7CFF3ED6D7FEBEFE0C.taxon	description	(Figs 11 C, 13 D, 15 C, 17 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9C2A7CFF3ED6D7FEBEFE0C.taxon	description	Description. ♀. Fully winged; body length 1.7 – 1.9 mm. Head testaceous (in ♀ from Yemen, Ta’izz, head totally whitish); antenna brown, except scape and pedicel testaceous (in ♀ from Yemen, Ta’izz, antenna totally whit- ish); prothorax, mesoscutum, mesoscutellum, mesopleuron and part of metapleuron testaceous; metanotum brown; metapectal-propodeal disc black; in specimen from Yemen, Ta’izz, mesosoma completely whitish, except brown spot on metapectal-propodeal disc and small brown spot on anterior margin of metanotum; metasoma brown (in specimen from Yemen, Ta’izz, metasoma totally whitish); legs testaceous (in specimen from Yemen, Ta’izz, legs totally whitish). Specimen from Madagascar, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, completely testaceous, except metapectalpropodeal disc darkened, petiole black and metasoma brown. Specimen from Kenya, 4.57776 ° S 39.43825 ° E, with head testaceous, except occiput brown and brown spot between ocelli; antenna brown, except antennomeres 1 – 3 testaceous; mesosoma testaceous, except dorsal region of propleuron, anterior surface of mesoscutum, mesoscutellum, metanotum and metapectal-propodeal disc brown. Antenna clavate; antennomeres in following proportions: 5.5: 5: 6: 6: 6: 5: 4: 4: 4: 6. Head dull, granulate; frontal line absent; occipital carina complete; POL = 6; OL = 4; OOL = 3.5; OPL = 3.5; TL = 3. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum; in ♀ from Yemen, Ta’izz, notauli slightly distinct, apparently reaching approximately 0.8 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curved. Tibial spurs 1 / 1 / 2. ♂ (Figs 17 C, D). Fully winged (Figs 11 C, 15 C); body length 1.4 – 1.8 mm. In holotype head and mesoscutum completely testaceous, antenna testaceous, except antennomeres 5 – 10 darkened, metanotum and metapectal-propodeal disc brown-reddish, metasoma brown, legs testaceous. In ♂ from Tanzania, Amani Malaria Station, head and mesosoma dull and completely testaceous-reddish, except clypeus yellow. In ♂ from Yemen, Ta’izz, colour whit- ish, except mesoscutum centrally and laterally brown, metapectal-propodeal disc laterally testaceous and medially brown, metasoma brown, with anterior third yellow and petiole black. In ♂ from Yemen, Ta’izz, colour whitish, except mesoscutellum centrally darkened, metanotum, metapectal-propodeal disc and metasoma brown. Antenna filiform; antennomeres of holotype in following proportions: 2.5: 4: 4: 5: 4.5: 5: 4.5: 5: 5: 7. Head dull, granulate; head of holotype slightly shiny; frontal line of holotype apparently absent (track visible); in ♂ from Tanzania, Amani Ma- laria Station, frontal line distinctly visible in anterior half of frons (face of this specimen very dull); occipital carina complete; POL = 4; OL = 3; OOL = 3; OPL = 2; TL = 2. Mesoscutum dull, granulate (shiny in holotype). Notauli incomplete, reaching approximately 0.5 × length of mesoscutum in holotype; in ♂ from Tanzania, Amani Malaria Station, notauli reaching approximately 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, unsculptured. Metapectal-propodeal disc and propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curved. Basivolsella with 2 subdistal bristles (Fig. 13 D). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9C2A7CFF3ED6D7FEBEFE0C.taxon	materials_examined	Material examined. Types: ♂ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Équateur Prov., Mbandaka (= Coquilhatville), 1964, A. B. Stam leg. (RMNH). Paratype: SOUTH AFRICA: Gauteng, 25 km W of Pretoria, 29. XI. 1984, H. & A. Howden leg., 1 ♀ (PMA). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 7 – 8. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♂ (SAMC); Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 22. V. 2001, sweep, lowland rainforest, S. van Noort leg., 1 ♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 7 – 14. IV. 2006, MT, field edge, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (UCDC). KENYA: Coast Prov., Funzi Island, 4.57776 ° S 39.43825 ° E, near sea level, 24 – 28. VII. 2012, MT, mixed grass-woodland, R. Copeland leg., 1 ♀ (NMK). MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 25. VII. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 35, 1 ♀ (CAS). TANZANIA: Tanga Region, Usambara Mts., Amani Malaria Station, 3000 ft., 6. I. 1970, M. E. Irwin & E. S. Ross leg., 1 ♂ (CAS); Tanga Region, Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ’ S 38 ° 01.37 ’ E, 16. IV – 2. V. 1996, MT, Commiphora woodland, S. van Noort leg., 1 ♂ (SAMC). YEMEN: Ta’izz, 22 – 24. VIII. 1998, light trap, A. van Harten & A. Awad leg., 1 ♂ (MOLC); Ta’izz, VIII. 1999, light trap, A. van Harten & A. Awad leg., 1 ♀ (MOLC); Ta’izz, V – VI. 2000, light trap, N. 4770, A. van Harten & A. R. Al Yarimi leg., 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9C2A7CFF3ED6D7FEBEFE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9C2A7CFF3ED6D7FEBEFE0C.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Kenya, Madagascar, South Africa, Tanzania, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9B2A7DFF3ED793FD08FCD8.taxon	description	(Figs 13 E, G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9B2A7DFF3ED793FD08FCD8.taxon	description	Description. ♀. Fully winged; body length 1.4 – 1.7 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, occasionally with metacoxa and metafemur darkened. Antenna filiform; antennomeres in following proportions: 5: 4: 4: 4: 4: 4: 4: 4: 4: 6. Head dull, granulate; frontal line absent; occipital carina complete; POL = 6; OL = 4; OOL = 3.5; OPL = 1.5; TL = 3; greatest breadth of posterior ocellus shorter than OL (2: 4). Mesoscutum dull, granulate. Notauli complete and posteriorly separated (minimum distance between notauli about as long as OL) or incomplete and reaching about 0.3 – 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels and median area smooth, not rugose. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein shorter than in other Aphelopus species (as in Fig. 13 G). Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.6 – 2.2 mm (holotype 1.6 mm). Head black (occasionally brown), except man- dible and clypeus testaceous (occasionally mandible, clypeus, malar space and anterior third of frons testaceous; in specimens from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, head black, except mandible testaceous and clypeus brown); antenna brown, except scape testaceous (in ♂ from Madagascar, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, scape and pedicel testa- ceous); mesosoma black, occasionally brown; metasoma brown; legs testaceous, occasionally with stalk of metafemur darkened. Antenna filiform; antennomeres of holotype in following proportions: 3: 3.5: 4: 4: 4: 5: 5: 5: 5: 8 (in large specimens 5: 4.5: 4: 5: 6: 7: 7: 7: 7: 10). Head dull, granulate; in holotype frontal line present only in anterior half of frons (occasionally frontal line complete; in specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, frontal line absent in front of anterior ocellus, present in rest of frons); occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL = 2; greatest breadth of lateral ocelli as long as OPL. Mesoscutum dull, granulate. Notauli incomplete, reaching about 0.3 – 0.5 × length of mesoscutum (in specimen from South Africa, 29 ° 39.23 ’ S 30 ° 27.41 ’ E, notauli reaching about 0.7 × length of mesoscutum; in specimens from South Africa, 28 ° 29.72 ’ S 23 ° 57.85 ’ E, and Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, notauli reaching about 0.8 × length of mesoscutum). Mesoscutellum dull, granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc and propodeal declivity reticulate rugose; occasionally propodeal declivity with two complete longitudinal keels. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein vein shorter than in other Aphelopus species (Fig. 13 G). Basivolsella (Fig. 13 E) with lateral distal process, with 1 – 2 subdistal bristles and distal lateral process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9B2A7DFF3ED793FD08FCD8.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Vernon Crookes Nature Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MT, M. Mostovski leg. (NMSA). Other material: KENYA: Central Prov., Mt. Kenya, Ragati, 6800 ft., 25. VIII. 1983, F. Buna leg., 1 ♂ (AMNH); Coast Prov., Taita Hills, Ngangao Forest, 3.36100 ° S 38.34186 ° E, 1848 m, 22. VIII – 5. IX. 2012, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest Reserve, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, 5400 feet, XI. 2001, Manfred Kraemer & Co., 1 ♂ (MOLC). MADAGASCAR: Antsiranana, Marojejy National Park, Manantenina river, 27.6 km 35 ° NE Andapa, 9.6 km 327 ° NNW Manantenina, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 11 – 14. XII. 2005, MT, rainforest, B. L. Fisher et al. leg., BLF 13383, 1 ♂ (MOLC), 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 21 – 28. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 1 ♂ (MOLC); Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 10 – 21. III. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 56, 1 ♂ (MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 C- 15, 2 ♂♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 C- 10, 1 ♂ (MOLC); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 1 ♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 1 ♂, 1 ♀ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 15 – 22. XI. 2001, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 07, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 1 ♂ (CAS); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 15 – 22. XI. 2001, MT near river, R. Harin’Hala leg., MA- 02 - 09 D- 03, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 25. VII. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 35, 1 ♂ (MOLC); same locality label, 9 – 20. III. 2003, MA- 02 - 09 B- 55, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 3 ♂♂ (CAS), 2 ♀♀ (CAS, MOLC); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 3 ♂♂ (CAS); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 1 ♂ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 2 ♂♂, 1 ♀ (MOLC); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 1 ♂, 1 ♀ (CAS); Toliara, Zombitse Na- tional Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 9 – 19. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 02, 1 ♂ (CAS). SOUTH AFRICA: Free State, Sterkfontein Dam, 28 ° 29.72 ’ S 23 ° 57.85 ’ E, 1780 m, 10. XII. 2005, YPT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Ash- burton, 29 ° 39.23 ’ S 30 ° 27.41 ’ E, 11. IV – 11. V. 2006, MT, G. Whiteley leg., 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9B2A7DFF3ED793FD08FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9B2A7DFF3ED793FD08FCD8.taxon	distribution	Distribution. Kenya, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9A2A73FF3ED5FFFC77FC90.taxon	description	(Figs 13 F, 15 D, E, 17 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9A2A73FF3ED5FFFC77FC90.taxon	description	Description. ♀ (Fig. 17 F). Fully winged (Fig. 15 D); body length 1.6 – 2.6 mm. Head black, except mandible testa- ceous and clypeus whitish; frons black, usually with more or less wide whitish spot; occasionally frons and clypeus completely black (Fig. 15 E); antenna completely testaceous, or brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous; ventral side of body usually black; in specimens with frons partly whitish, ventral side of head and mesosoma completely whitish; occasionally body completely brown or ferruginous. Antenna filiform; antennomeres in following proportions: 5: 5: 6: 8: 8: 7: 5: 5: 5: 7. Head dull, granulate; frontal line incomplete; occipital carina complete; POL = 5.5; OL = 4; OOL = 5; OPL = 4; TL = 5. Mesoscutum, mesoscutellum and metanotum dull, granulate. Notauli incomplete, usually reaching 0.65 – 0.80 × length of mesoscutum, occasionally reaching 0.3 – 0.4 × length of mesoscutum. Mesopleuron and metapleuron smooth, unsculptured or slightly granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity usually with two longitudinal keels, median area usually shiny, smooth, not reticulate rugose, and lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands. Tibial spurs 1 / 1 / 2. ♂ (Fig. 17 E). Fully winged; body length 1.7 – 2.2 mm. Head usually black, except mandible testaceous; clyp- eus black; occasionally frons black, except clypeus whitish; occasionally frons completely whitish, except ocellar region brown, or black, except more or less wide whitish spot; antenna usually brown, except scape and pedicel testaceous; in ♂♂ with head almost completely whitish, antenna completely testaceous; mesosoma and metasoma black (rarely mesosoma brown); legs usually completely testaceous; occasionally stalk of metafemur darkened. Antenna filiform; antennomeres in following proportions: 5.5: 5: 5.5: 6: 7: 7.5: 7: 7: 7: 10. Head dull, granulate; frontal line incomplete; occipital carina complete; POL = 7.5; OL = 4; OOL = 5; OPL = 4; TL = 5. Mesoscutum, mesoscutellum and metanotum dull, granulate; notauli incomplete, usually reaching approximately 0.65 × length of mesoscutum, occasionally reaching 0.4 × length of mesoscutum. Mesopleuron and metapleuron smooth, slightly granulate or unsculptured. Metapectal-propodeal disc reticulate rugose; propodeal declivity usually with two longitudinal keels, median area usually shiny, smooth, unsculptured, and lateral areas reticulate rugose; occasionally propodeal de- clivity without longitudinal keels, completely reticulate rugose. Forewing hyaline, without dark transverse bands. Basivolsella (Fig. 13 F) with 2 subdistal bristles; in specimen from Kenya, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, one of basivolsellae has 2 subdistal bristles and the other one has 1 subdistal bristle. In specimen from South Africa, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, parameres strongly bent and much shorter than penis. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9A2A73FF3ED5FFFC77FC90.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), between Ngesho and Bishakishaki, Kashwa, 2000 m, 7 – 23. I. 1935, Mission G. F. de Witte (MRAC). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Mokoto, Burunga, 2000 m, 17 – 19. III. 1934, Mission G. F. de Witte, ♂ allotype (MRAC); same locality label as allotype, 1 ♂ (NHMUK); North Kivu, Virunga National Park (= Albert National Park), Volc. Mikeno, Rweru, 2400 m, bambous, 26 – 27. VII. 1934, G. F. de Witte leg., 1 ♂ (NHMUK); in MRAC there are further 87 paratypes (not examined) all from North Kivu (see list in Benoit 1951 c). Other material: ANGOLA: Cuanza Sul, 7 mi. W Gabela, 16 – 18. III. 1972, Southern African Expedition, 2 ♂♂, 2 ♀♀ (NHMUK); BOTSWANA: Chobe National Park (USUC). BURUNDI: Muyange, 2100 m, 11. VIII. 1948 (ISNB); Rusizi National Park, 3.34364 ° S 29.27246 ° E, 774 m, 13 – 20. IV. 2010, MT near three small trees, degraded bush / grassland, R. Copeland leg., 1 ♂ (NMK); Bururi National Forest, 3.93022 ° S 29.61697 ° E, 1955 m, 24. VIII – 7. IX. 2010, MT, indigenous for- est, near stream, R. Copeland leg., 1 ♀ (NMK). CAMEROON: East Region, Messa Bengondo, 15 – 30. III. 2003, 13 – 30. IV. 2003, M. Tussac leg., 2 ♂♂ (HTS); Northwest Region, E of Big Babanki, 06 ° 06.698 ’ N 10 ° 15.938 ’ E, 1200 m, 5 – 13. III. 2008, Martin Riha leg., 1 ♀ (OLL); South Region, near Ebolowa, Nkoemvon, XII. 1978 – I. 1979, D. Jack- son leg., 1 ♀ (NHMUK); Southwest Region, Korup, 1981, D. Jackson leg., 1 ♀ (NHMUK). CENTRAL AFRICAN REPUBLIC: Nana-Mambéré Pref., 60 km W Bouar, 05 ° 45 ’ N 15 ° 13 ’ E, 660 m, 23. III. 2010, J. Halada leg., 1 ♂ (OLL); Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 7 – 8. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♀, 4 ♂♂ (SAMC); Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 22. V. 2001, sweep, lowland rainforest, S. van Noort leg., 1 ♀, 12 ♂♂ (SAMC); Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 6. V. 2001, sweep, lowland rainforest, marsh clearing, CAR 01 - S 48, S. van Noort leg., 1 ♂ (SAMC); Sangha-Mbaéré Pref., Reserve Speciale de Forêt Dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 13. V. 2001, sweep, lowland rainforest, S. van Noort leg., 1 ♀, 8 ♂♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Haut- Katanga, Lubumbashi (= Elisabethville) (RMNH, MRAC); Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, VII. 2008, MT, T. Chapman leg., 1 ♂ (UCDC); same locality label, 6 – 17. IV. 2006, MT, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (UCDC); same locality label, 23. IV. 2006, MT along forest edge, 1 ♂ (UCDC); North Kivu, Rut- shuru (MRAC); North Kivu, Munagana (= Mdagana) - Tsheugelero (MRAC); North Kivu, Ngesho (MRAC); North Kivu, Virunga National Park (= Albert National Park), N’Dalage (MRAC); North Kivu, Virunga National Park (= Albert National Park), Magera Lake (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo, Yolohafiri Cave (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo (MRAC, AMNH); North Kivu, Virunga National Park (= Albert National Park), aff. Talia Nord, Secteur Tshiaberimu, Riv. Kalivina (MRAC); North Kivu, Virunga National Park (= Albert National Park), aff. dr. Talia Nord, Secteur Tshiaberimu, Riv. Musabaki (MRAC, AMNH); North Kivu, Virunga National Park (= Albert National Park), Secteur Tshiaberimu, Kirungu (MRAC); North Kivu, Virunga National Park (= Albert National Park), Secteur Tshiaberimu, Mt. Musimba (MRAC); North Kivu, Virunga National Park (= Albert National Park), Secteur Tshiaberimu, Hintumo (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, Aff. Butahu, Riv. Kamahoro (MRAC, AMNH); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, Aff. Butahu, Riv. Kamusonge (MRAC, AMNH); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, Aff. Butahu, Riv. Nyamwamba (MRAC, AMNH); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, Kiondo ya Kwanza (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Aff. Bombi, Riv. Kakalari (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, Kikyo (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Mt. Kyanyamu, Riv. Lusilube (MRAC); North Kivu, Virunga National Park (= Albert National Park), Bitashimva, Mt. Sesero (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kyandolire, Bomboka (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kyandolire, Mt. Ibale (MRAC); further localities listed in Benoit (1951 c); Haut-Uele (= Haut- Zaire) Prov., Garamba National Park (MRAC, AMNH). GABON: Ogoové-Maritime Prov., Doudou Mts. Reserve, 24.3 km 307 ° NW Doussala, 02 ° 13.35 ’ S 10 ° 24.35 ’ E, 370 m, 7. III. 2000, sweep, GA 00 - S 49, Coastal Lowland Rainforest, under-growth, low canopy in forest, S. van Noort leg., 1 ♂ (SAMC); same locality label, 9. III. 2000, 1 ♂ (SAMC). GHANA: Ashanti, Kumasi, Campus of the University of Science and Technology (RNHL). IVORY COAST: Marahoué Region, 8 km NW of Bouaflé, Klébo (DEUW); Marahoué Region, 5 km E of Bouaflé, Congo Aboisso (DEUW); Marahoué Region, Congo Aboisso, 05 ° 40 ’ W 07 ° N, 18. III. 1981 (CNC). KENYA: Coast Prov., Taita Hills, Vuria Forest, 3.41428 ° S 38.29178 ° E, 2162 m, 11 – 25. I. 2012, MT, just inside indigenous forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Taita Hills, Chawia Forest, 3.47901 ° S 38.34134 ° E, 1617 m, 21 – 25. VIII. 2012, MT, next to fallen tree, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Funzi Island, 4.57776 ° S 39.43825 ° E, near sea level, 4 – 10. VII. 2012, MT, near Funzi workshop, ICIPE / NMK Funzi Island Expedition, 2 ♀♀ (NMK); same locality label, 24 – 28. VII. 2012, MT, mixed grass-woodland, R. Copeland leg., 3 ♀♀ (NMK); Coast Prov., Kasigau Mtn., 3.82700 ° S 38.64875 ° E, 1065 m, 5 – 19. X. 2011, MT, indigenous forest, next to campsite in forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Taita Hills, Ngangao Forest, 3.36100 ° S 38.34186 ° E, 1848 m, 18. IX – 2. X. 2011, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Nyambene Hills, Itieni Forest, at bottom, 0.24433 ° N 37.87016 ° E, 2142 m, 7 – 21. VIII. 2011, MT, edge of indigenous forest, near forest station, R. Copeland leg., 1 ♀ (NMK); same locality label, 18. IX – 2. X. 2011, 1 ♀ (NMK); same locality label, 26. VI – 10. VII. 2011, 1 ♂ (NMK); same locality la- bel, 24. VII – 7. VIII. 2011, 1 ♀ (NMK); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi leg., 1 ♂, 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22338 ° S 36.89655 ° E, 1604 m, 24. II – 3. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♂ (NMK); Nyanza Prov., near Victoria Lake, 15 mls. NE of Kisumu, XI. 1979, M. D. Croft leg., 1 ♀ (NHMUK); Nyanza Prov., Ruma National Park, 00 ° 39.121 ’ S 34 ° 19.417 ’ E, 1250 m, 26. VII. 2008, sweep, savanna woodland, S. van Noort leg., 1 ♂ (SAMC); Rift Valley Prov., Molo, Turi, 25. X. 1998, A. Polaszek leg., 1 ♂ (NHMUK); Rift Valley Prov., Saiwa Swamp National Park, near campsite, 1.09417 ° N 35.11833 ° E, 1882 m, 26. II – 12. III. 2006, MT, next to permanent upland swamp, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Mt. Elgon National Park, top of Endebess Bluff, 1.06117 ° N 34.75383 ° E, 2630 m, 16 – 30. I. 2006, MT, forest edge near small stream, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Olorgesailie National Monument, 1.57930 ° S 36.44566 ° E, 982 m, 27. XI – 11. XII. 2011, MT, Acacia-Commiphora savana, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Oloitokitok, 2.94456 ° S 37.50714 ° E, 1853 m, 19. VIII – 2. IX. 2011, MT, edge of indigenous forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 22. VII – 5. VIII. 2011, 3 ♀♀ (NMK); same locality label, 9 – 23. XII. 2011, 1 ♀ (NMK); same local- ity label, 5 – 19. VIII. 2011, 1 ♀ (MOLC); same locality label, 14 – 28. X. 2011, 1 ♀ (NMK); same locality label, 28. X – 11. XI. 2011, 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest Reserve, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, 5400 feet, XI. 2001, Manfred Kraemer & Co., 3 ♂♂, 3 ♀♀ (MTC); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest, Calebs Campground, 00 ° 22.175 ’ N 34 ° 53.297 ’ E, 1580 m, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MTC); Western Prov., Kakamega Forest, 00 ° 14.13 ’ N 34 ° 51.87 ’ E, 4. III. 1999, R. Wharton leg., 1 ♂ (TAMU); Western Prov., Kakamega District, Kakamega Forest (USUC); Western Prov., Kakamega District, Isecheno Nature Reserve, Isecheno, 00 ° 24 ’ N 34 ° 87 ’ E, 1800 m, MT, R. R. Snelling leg., 1 ♂, 2 ♀♀ (UCR, MOLC); Western Prov., 27 mi. NE of Kisumu, Kaimosi Mission, 1650 m (CAS). MADAGAS- CAR: Antananarivo, La Mandraka (MRAC, AMNH); Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 26 – 29. I. 2001, MT, M. E. Irwin, Schlinger E. I. Harin’Hala R. leg. (CAS); same locality label, 19. III – 6. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 09, 2 ♀♀ (CAS); Antsiranana, Marojejy National Park, Manantenina river, 28.0 km 38 ° NE Andapa, 8.2 km 333 ° NNW Manantenina, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, MT, rainforest, B. L. Fisher et al. leg., BLF 8723, 1 ♂, 1 ♀ (CAS), 1 ♀ (MOLC); Fianarantsoa, Rano- mafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., 1 ♀ (CAS); same locality label, 19 – 26. II. 2002, 1 ♀ (CAS); same locality label, 23. VI – 4. VII. 2002, MA- 02 - 09 C- 33, 1 ♂, 1 ♀ (CAS); same locality label, 10 – 21. III. 2003, MA- 02 - 09 C- 56, 1 ♀ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 C- 10, 1 ♀ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 1 ♀ (CAS); same locality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 1 ♂, 1 ♀ (CAS); same locality label, 21. III – 12. IV. 2003, MA- 02 - 09 C- 57, 2 ♀♀ (CAS); same locality la- bel, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 4 – 12. II. 2002, MT in high altitude rainforest, R. Harin’Hala leg., 1 ♀, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 9 – 16. IV. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 24, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 1 ♂, 1 ♀ (CAS); same locality label, 18 – 29. VI. 2003, MA- 02 - 09 B- 64, 1 ♀ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♀ (CAS); same locality label, 20. III – 3. IV. 2003, MA- 02 - 09 B- 56, 1 ♀ (CAS); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 1 ♀ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 1 ♀ (CAS); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 1 ♀ (CAS); Fianarantsoa, Miandritsara Forest, 40 Km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 8 – 17. III. 2007, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 81, 1 ♀ (CAS); same locality label, 16 – 27. VIII. 2005, MA- 29 - 24, 1 ♀ (CAS); Mahajanga, Bemarivo Special Re- serve, 23.8 km 223 ° SW Besalampy, 16 ° 55.30 ’ S 44 ° 22.06 ’ E, 19 – 23. XI. 2002, 30 m, sifted litter (leaf mold, rotten wood), in tropical dry forest, Fisher, Griswold et al. leg., 1 ♂ (CAS); Toamasina, Perinet (AMNH); Toamasina, An- dasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 16 – 24. X. 2001, MT, tropical forest, R. Harin’Hala leg., 1 ♀ (CAS); same locality label, 1 – 7. XI. 2001, 1 ♀ (CAS); Toamasina, Ankerana, 18 ° 24.23 ’ S 48 ° 48.09 ’ E, 1108 m, 19 – 24. I. 2012, MT, montane forest, B. L. Fisher et al. leg., BLF 27860, 1 ♂ (CAS); Toliara, Bekily (MNHN); Toliara, Tsimanampetsotsa Nat. Park, Bemanateza Forest, 20.7 km 81 ° E Efoetse, 23 Km 131 ° SE Beheloka, 23 ° 59.32 ’ S 43 ° 52.50 ’ E, 90 m, 22 – 26. III. 2002, MT, in spiny forest thicket, Fisher, Griswold et al. leg., 1 ♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 30. XI – 11. XII. 2003, MT in transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 47, 1 ♂ (MOLC); Toliara, Manombo, 22 ° 48.25 ’ S 43 ° 45.46 ’ E, 218 m, 23 – 28. IV. 2004, MT in gallery forest, Frontier Wilderness Project, MGF 098, 1 ♀ (CAS). MOZAMBIQUE: Niassa Prov., Cuamba, Farm of the Catholic University of Mozambique, reared from unidentified leafhopper, parasitized host collected 18. II. 2005, dryinid cocooning 20. II. 2005, dryinid adult emergence 8. III. 2005, 1 ♀ (MOLC); same locality label, 17. IV – 2. VI. 2006, MT, M. Olmi leg., 1 ♀ (MOLC). NIGERIA: Osun, Ile-Ife, 2 ♂♂ (NHMUK, AMNH). NORTH SUDAN (= REPUBLIC OF THE SUDAN): Al Jazirah, Wad Medani, 18. XI. 1949, 19. XI. 1949, 22. XI. 1949, G. Morcos leg., reared from Austroasca lybica, 4 ♀ # (BMMH). RWANDA: Kirii Springs, Ruhengeri (MRAC). SENEGAL: Ziguinchor Region, 1 km NE of Djibelor, 7.5 km SW of Ziguinchor (ZIL). SOMALIA: Cotene, 1930, G. Russo leg., reared from Empoasca facialis (Jacobi), 2 ♂♂ (NHMUK). SOUTH AFRICA: Eastern Cape, Nature’s Valley, at Groot River, 33 ° 58 ’ S 23 ° 33 ’ E (MZLU); Eastern Cape, 25 km N Jansenville, 32 ° 49 ’ S 24 ° 44 ’ E, 600 m, 4. I. 2010, J. Halada leg., 1 ♀ (OLL); Eastern Cape, Marais Hoop Farm, 25.6 km 254 ° W Kirkwood, 33 ° 32.635 ’ S 25 ° 13.678 ’ E, 14 – 16. II. 2001, MT, VB 01 - R 4 N- M 52, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♀ (SAMC); Eastern Cape, Blauwe Krans Farm, 12.8 km 216 ° SW Kirkwood, 33 ° 30.747 ’ S 25 ° 24.644 ’ E, 7 – 9. II. 2001, MT, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Mannetjie Farm (31.9 km 262 ° W Kirkwood), 33 ° 32.724 ’ S 25 ° 08.795 ’ E, 12 - 14. II. 2001, MT, VB 01 - R 3 N, M 42, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♂ (SAMC); same locality label, 14 – 16. II. 2001, M 50, 2 ♂♂ (SAMC); same locality label, 8 – 10. II. 2001, M 12, 4 ♂♂, 1 ♀ (SAMC); Eastern Cape, Marais Hoop Farm (25.6 km 254 ° W Kirkwood), 33 ° 32.635 ’ S 25 ° 13.678 ’ E, 14 – 16. II. 2001, MT, VB 01 - R 4 N-M 52, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♀, 5 ♂♂ (SAMC); same locality label, 12 – 14. II. 2001, 1 ♂ (SAMC); same locality label, 8 – 10. II. 2001, M 14, 2 ♂♂ (SAMC); same locality label, 10 – 12. II. 2001, M 26, 5 ♂♂, 1 ♀ (SAMC); Eastern Cape, Schilpad Laagte Farm (15.4 km 226 ° W Kirkwood), 33 ° 31.653 ’ S 25 ° 22.620 ’ E, 7 – 9. II. 2001, MT, Valley Bushveld (non-trashed), S. van Noort leg., 4 ♂♂ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC); Eastern Cape, Februarie Farm (39.9 km 268 ° W Kirkwood), 33 ° 32.813 ’ S 25 ° 03.091 ’ E, 8. II. 2001, sweep, valley bushveld (non-trashed), S. van Noort leg., 1 ♂, 1 ♀ (SAMC); Eastern Cape, Februarie Farm (40.2 km 267 ° W Kirkwood), 33 ° 33.124 ’ S 25 ° 03.043 ’ E, 10 – 17. II. 2001, valley bushveld (non-trashed), H. G. Robertson & R. Tourle leg., 1 ♂ (SAMC); same locality label, Winkler, valley bushveld (goat trashed), 1 ♂ (SAMC); same locality label, 10. II. 2001, sweep, S. van Noort leg., 3 ♂♂ (SAMC); same locality label, 10 – 12. II. 2001, MT, VB 01 - R 1 N-M 22, Val- ley bushveld (non-trashed), S. van Noort leg., 18 m ♂ # (SAMC); same locality label, 14 – 16. II. 2001, M 54, 16 ♂♂ (SAMC); same locality label, 12 – 14. II. 2001, M 36, 14 ♂♂ (SAMC); same locality label, 8 – 10. II. 2001, M 08, 10 ♂♂ (SAMC); Eastern Cape, Blauwe Krans Farm (12.8 km 216 ° SW Kirkwood), 33 ° 30.747 ’ S 25 ° 24.644 ’ E, 7 – 9. II. 2001, MT, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♂ (SAMC); same locality label, 9 – 11. II. 2001, 1 ♂ (SAMC); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 6. X. 2010 – 18. I. 2011, MT, Amathole Mistbelt Grassland, S. van Noort leg., WTB 09 - GRA 1 - M 05, 1 ♀ (SAMC); KwaZulu-Natal, Drakensberg, Cathedral Peak area, Tryme Hill, 1540 m, on Protea caffra, plot No 1, 30. VI. 1989, Pajor István leg., 1 ♂ (SAMC); KwaZulu- Natal, Drakensberg, Cathedral Peak area, Tarn Hill, 1570 m, on Protea caffra, plot No 2, 8. V. 1989, Pajor István leg., 1 ♂ (SAMC); KwaZulu-Natal, Giant’s Castle Game Reserve (FSCA); KwaZulu-Natal, Umlalazi Nature Reserve, 1.5 km E of Mtunzini (PMA); KwaZulu-Natal, 75 km SW of Estcourt, Cathedral Peaks Forest Station (CNC); Kwa- Zulu-Natal, South Drakensberg, Sani Pass, 29 ° 37 ’ S 29 ° 23 ’ E, 2 – 5. III. 2004, MT, Kolyada & Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Pietermaritzburg, Ferncliff Nature Reserve, 29 ° 33.2 ’ S 30 ° 20.6 ’ E, 850 m, 6 – 8. IV. 2005, YPT, V. Kolyada leg., 4 ♀♀ (NMSA); KwaZulu-Natal, Pietermaritzburg, Hilton, 27. IX – 13. X. 2003, 28. X – 12. XI. 2003, 15 – 26. I. 2004, MT, M. Mostovski leg., 1 ♂, 3 ♀♀ (NMSA); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ’ S 30 ° 20.4 ’ E, nr. stream, 9. I – 2. II. 2005, 27. II – 19. III. 2005, 20. III – 25. IV. 2005, M. Mostovski leg., 2 ♀♀, 3 ♂♂ (NMSA); KwaZulu-Natal, Royal Natal National Park, Gudu Forest, 28 ° 40.90 ’ S 28 ° 55.78 ’’ E, 1680 – 1730 m, 18. XI. 2006 – 27. II. 2007, MT, M. Mostovski leg., 1 ♂ (NMSA); same locality label, 22. IX – 17. XI. 2006, 1 ♂ (NMSA); KwaZulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 23. II – 20. IV. 2007, MT, Kolyada & Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Winterskloof, 29 ° 34.56 ’ S 30 ° 17.40 ’ E, 1085 m, 20. IV – 1. IX. 2007. MT, M. Mostovski leg., 1 ♀, 3 ♂♂ (NMSA); KwaZulu-Natal, Vernon Crookes Nat. Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MTr, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Cathedral Peak Nat. Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 18. XI. 2006 – 26. II. 2007, MT, M. Mostovski leg., 1 ♀, 2 ♂♂ (NMSA); Mpumalanga, Graskop, 1500 m (PMA); Western Cape, Matjesfontein, 7 – 13. XI. 1928, R. E. Turner leg., 1 ♂ (NHMUK); same locality label, further specimen (AMNH); Western Cape, De Hoop Nature Reserve, 34 ° 27 ’ S 20 ° 25 ’ E, 0 – 200 m (ZIL); Western Cape, Mossel Bay, V. 1921, VIII. 1921, R. E. Turner leg., 2 ♀♀ (NHMUK); Western Cape, Hermanus, Fernkloof Nature Reserve, Mosselberg, 60 m, 12. III. 1995, MT, Mesic Mountain Fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, 16. VI – 22. VII. 1995, 1 ♀, 1 ♂ (SAMC); Western Cape, Cape Town, Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 27. VIII – 5. IX. 1993, MT, Mountain Fynbos on Sandstone, near stream, bracken present, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 1. VI. 1995, Mesic Mountain Fynbos, near stream, sweep, 1 ♂ (SAMC); same locality label, 10. V – 1. VI. 1995, Mesic Mountain Fynbos, P. coronata dominated, MT, site 2, 1 ♂ (SAMC); Western Cape, Walker Bay Nature Re- serve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 4. X – 1. XI. 1997, YPT, S. van Noort & B. Fisher leg., 1 ♀, 2 ♂♂ (SAMC); same local- ity label, 11. VIII – 6. IX. 1997, S. van Noort leg., 39 ♂♂, 7 ♀♀ (SAMC); same locality label, 9. VIII – 6. IX. 1997, S. van Noort leg., 1 ♀, 7 ♂♂ (SAMC); same locality label, IX – XII. 1997, MT, S. van Noort leg., 9 ♀♀, 17 ♂♂ (SAMC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 26 ♀♀, 73 ♂♂ (SAMC); same locality label, 17. V – 14. VI. 1997, S. van Noort leg., 1 ♀ (CAS); same locality label, 6. IX – 4. X. 1997, MT, S. van Noort leg., 1 ♀, 5 ♂♂ (CAS); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’’ S 19 ° 21.393 ’ E, 28. XII. 1997 – 21. III. 1998, MT, South Coast Strandveld, 57 m, S. van Noort leg., 7 ♂♂, 1 ♀ (SAMC); same locality label, 21. II. 1998, sweep, 1 ♀ (SAMC); same locality label, 29. XI. 1997, sweep, 1 ♀ (SAMC); same locality label, 4. X. 1997, sweep, 1 ♀ (SAMC); same locality label, 16. V. 1998, sweep, 2 ♀♀ (SAMC); same locality label, 14. VI. 1997, sweep, 1 ♂ (SAMC); same locality label, 14. VI. 1997, sweep, 2 ♂♂, 2 ♀♀ (SAMC); same locality label, 18. IV – 16. V. 1998, 6 ♂♂, 2 ♀♀ (SAMC); same locality label, 21. II – 21. III. 1998, 5 ♂♂, 2 ♀♀ (SAMC); Western Cape, Cape Peninsula Nature Reserve, Cape Good Hope, 34 ° 14 ’ S 18 ° 23 ’ E, pitfall trap, 27. II – 21. III. 1999, E. Parker leg., 1 ♂ (SAMC); Western Cape, Cape of Good Hope Nature Reserve, Booi se Skerm, 34 ° 18.27 ’ S 18 ° 27.64 ’ E, 24. II. 1998, sweep, strandveld, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Tsitsikamma Coastal National Park, 34 ° 02 ’ S 23 ° 53 ’ E, XI – XII. 1995, MT, M. Söderlund leg., 2 ♂♂ (MZLU); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 817 ’’ S 18 ° 06 ’ 837 ’’ E (4 km 280 ° E Langebaanweg), 16 – 23. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 3 ♂♂, 1 ♀ (SAMC); same locality label, 18 – 25. IX. 2002, 19 ♂♂, 4 ♀♀ (SAMC); same locality label, 2 – 9. X. 2002, 25 ♂♂ (SAMC); same locality label, 9 – 16. X. 2002, 1 ♀, 30 ♂♂ (SAMC); same locality label, 11 – 18. IX. 2002, 35 ♂♂, 11 ♀♀ (SAMC); same locality label, 23 – 30. X. 2002, 3 ♂♂ (SAMC); same locality label, 4 – 11. IX. 2002, 22 ♂♂, 8 ♀♀ (SAMC); same locality label, 25. IX – 2. X. 2002, 21 ♂♂, 4 ♀♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58 ’ 098 ’’ S 18 ° 07 ’ 244 ’’ E (3 km 280 ° W Langebaanweg), 11 – 18. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58 ’ 020 ’’ S 18 ° 07 ’ 285 ’’ E (3 km 280 ° W Langebaanweg), 23 – 30. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀ (SAMC); same locality label, 16 – 23. X. 2002, 1 ♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 512 ’’ S 18 ° 06 ’ 728 ’’ E (4 km 290 ° E Langebaanweg), 2 – 9. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂, 2 ♀♀ (SAMC); same locality label, 23 – 30. X. 2002, 1 ♂, 1 ♀ (SAMC); same locality label, 9 – 16. X. 2002, 1 ♂ (SAMC); same locality label, 12 – 19. XI. 2003, 1 ♂, 2 ♀♀ (SAMC); same locality label, 6 – 13. XI. 2002, LW 02 - R 7 - M 140, 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 402 ’’ S 18 ° 05 ’ 438 ’’ E (5.5 km 290 ° W Langebaanweg), 11 – 18. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58 ’ 156 ’’ S 18 ° 05 ’ 878 ’’ E (5.5 km 270 ° W Langebaanweg), 16 – 23. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58 ’ 444 ’’ S 18 ° 06 ’ 731 ’’ E (4 km 270 ° W Langebaanweg), 11 – 18. IX. 2002, MT, fynbos dominated by Lebeckia, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58.483 ’ S 18 ° 06.884 ’ E (4 km 270 ° W Langebaanweg), 11 – 18. IX. 2002, MT, fynbos on Lymestone, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Langberg Farm, 32 ° 58 ’ 461 ’’ S 18 ° 07 ’ 344 ’’ E (3 km 270 ° E Langebaanweg), 27. XI – 4. XII. 2002, MT, sand plain fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, 25. IX – 2. X. 2002, 1 ♂ (SAMC); same locality label, 2 – 9. IV. 2003, 1 ♂ (SAMC); same locality label, 11 – 18. IX. 2002, 1 ♂ (SAMC); same locality label, 29. I – 5. II. 2003, 1 ♀ (SAMC); Western Cape, Langberg Farm (3 km 280 ° W Langebaanweg), 32 ° 58.045 ’ S 18 ° 07.418 ’ E, 2 – 9. X. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 9 – 16. X. 2002, 1 ♂ (SAMC); same locality label, 11 – 18. IX. 2002, 1 ♀ (SAMC); Western Cape, Anyskop Farm, 5.5 km 290 ° W Langebaanweg, 32 ° 57.394 ’ S 18 ° 05.324 ’ E, 25. IX – 2. X. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 2. IV – 30. VII. 2010, MT, Boland Granite Fynbos, S. van Noort leg., 1 ♀ (SAMC); same locality label, 28. IX – 28. XII. 2010, DRA 09 - FYN 1 - M 05, 1 ♂ (SAMC); Western Cape, Gamkaberg Nature Re- serve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 21. III – 23. V. 2009, MT, Renosterveld, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 19. IX – 4. XI. 2009, 1 ♂, 1 ♀ (SAMC); same locality label, 19. II – 30. III. 2010, YPT, GB 09 - REN 1 - Y 47, 7 ♂♂, 2 ♀♀ (SAMC); same locality label, 4. XI. 2009 – 19. II. 2010, MT, GB 09 - REN 1 - M 29, 12 ♂♂, 14 ♀♀ (SAMC); same locality label, MT, 30. III – 24. VII. 2010, GB 09 - REN 1 - M 39, 41 ♂♂, 16 ♀♀ (SAMC); same locality label, 23. V – 31. VII. 2009, 2 ♂♂, 3 ♀♀ (SAMC); same locality label, 19. II – 30. III. 2010, GB 09 - REN 1 - M 33, 15 ♂♂, 4 ♀♀ (SAMC); same locality label, 24. VII – 1. X. 2010, YPT, GB 09 - REN 1 - Y 65, 16 ♂♂, 7 ♀♀ (SAMC); same locality label, 24. VII – 4. X. 2010, MT, GB 09 - REN 1 - M 45, 15 ♂♂, 7 ♀♀ (SAMC); same locality label, YPT, 4. X. 2010 – 25. I. 2011, GB 09 - REN 1 - Y 65, 16 ♂♂, 2 ♀♀ (SAMC); same locality label, MT, 4. X. 2010 – 25. I. 2011, GB 09 - REN 1 - M 51, 17 ♂♂, 3 ♀♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, 997 m, 30. III – 24. VII. 2010, YPT, Rooiberg Sandstone Fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, 30. X. 2010 – 25. I. 2011, MT, GB 09 - FYN 2 - M 53, 1 ♂ (MOLC); same locality label, 24. VII – 4. X. 2010, GB 09 - FYN 2 - Y 69, 1 ♂ (SAMC); same locality label, 24. VII – 4. X. 2010, MT, GB 09 - FYN 2 - M 47, 1 ♀ (SAMC); same locality label, 4. X. 2010 – 25. I. 2011, GB 09 - FYN 2 - Y 77, 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 39.504 ’ S 21 ° 53.947 ’ E, 322 m, 30. III – 24. VII. 2010, MT, Gamka Thicket, S. van Noort leg., GB 09 - SUC 4 - M 38, 1 ♂ (SAMC); Western Cape, Kogelberg Nature Researve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. V – 16. VI. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC); same locality label, 16. III – 16. IV. 1999, 1 ♀ (SAMC); same locality label, 16. II – 16. III. 1999, 1 ♀ (SAMC); same locality label, 16. XI – 16. XII. 1998, 1 ♀ (SAMC); same locality label, 16. I – 16. II. 1999, 2 ♀♀, 1 ♂ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37 ’ 622 ’’ S 18 ° 24 ’ 259 ’’ E, 20. III – 17. IV. 1998, MT, West Coast Strandeveld, S. van Noort leg., 1 ♀, 1 ♂ (SAMC); same locality label, 13. VI – 11. VII. 1997, 1 ♂ (SAMC); same locality label, 5. IX – 3. X. 1997, 1 ♂, 2 ♀♀ (SAMC); same locality label, 8. VIII – 5. IX. 1997, 3 ♀♀ (SAMC); same locality label, 3 – 31. X. 1997, 3 ♀♀ (SAMC); same locality label, 28. XI – 27. XII. 1997, 10 ♂♂, 3 ♀♀ (SAMC); same locality label, 23. I – 20. II. 1998, 2 ♀♀ (SAMC); same locality label, 31. X – 28. XI. 1997, 3 ♀♀, 1 ♂ (SAMC); same locality label, 11. VII – 8. VIII. 1997, 2 ♂♂, 1 ♀ (SAMC); same locality label, 27. XII. 1997 – 23. I. 1998, 11 ♂♂, 5 ♀♀ (SAMC); same locality label, 17. IV – 15. V. 1998, 3 ♀♀ (SAMC); same locality label, 20. III – 17. IV. 1998, 3 ♀♀, 2 ♂♂ (SAMC); same locality label, 20. II – 20. III. 1998, 4 ♀♀, 3 ♂♂ (SAMC); same locality label, 16. V – 13. VI. 1997, 3 ♂♂, 4 ♀♀ (SAMC). TANZANIA: Arusha Region, Arusha (HNHM); Tanga Re- gion, West Usambara Mountains, near Lushoto, 1850 m, 04 ° 44.00 ’ S 38 ° 16.06 ’ E, 30. IV. 1996, sweep, mk 41, margins & undergrowth, Wet Montane Forest, S. van Noort leg., 1 ♀ (SAMC); Tanga Region, Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ’ S 38 ° 01.37 ’ E, 2. V. 1996, Sweep, mk 43, Cordia, Solanum shrubs & grasses in Commiphora woodland, S. van Noort leg., 1 ♂ (SAMC); Tanga Region, Mkomazi Game Reserve, Ibaya Camp, NW side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 28. XI. 1995, Sweep, mk 9, Acacia / Commiphora / Combretum bushland, S. van Noort leg., 1 ♂ (SAMC). TOGO: Plateaux Region, Zogbégan, 07 ° 34.50 ’ N 00 ° 40.03 ’ E, 650 m, 20 – 25. IV. 2008, M. von Tschirn- haus leg., 1 ♂ (MTC). UGANDA: Western Region, Kabarole District, Kibale Forest, Kanywara (RNHL); Western Region, Kabarole District, Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 34.405 ’ N 30 ° 21.646 ’ E, 1484 m, 12 – 26. VIII. 2008, MT, primary mid-altitude rainforest, near stream, S. van Noort leg., 1 ♂ (SAMC); same locality label, 00 ° 33.891 ’ N 30 ° 21.468 ’ E, 1506 m, 12 – 26. VIII. 2008, 1 ♀, 5 ♂♂ (SAMC); same locality label, 00 ° 35.442 ’ N 30 ° 21.741 ’ E, 1465 m, 10. VIII. 2008, sweep, primary mid-altitude rainforest, near stream, S. van Noort leg., 7 ♂♂ (SAMC); same locality label, 0 ° 33.871 ’ S 30 ° 21.355 ’ E, 1495 m, 12 – 26. VIII. 2008, MT, secondary mid-altitude rainforest, S. van Noort leg., 2 ♂♂, 1 ♀ (SAMC); same locality label, 0 ° 33.408 ’ S 30 ° 22.603 ’ E, 1587 m, MT, degraded mid-altitude rainforest, S. van Noort leg., 3 ♂♂ (SAMC); same locality label, 2 – 12. VIII. 2008, 0 ° 33.784 ’ S 30 ° 22.617 ’ E, 1500 m, MT, primary mid-altitude rainforest, S. van Noort leg., 1 ♀, 11 ♂♂ (SAMC); same locality label, 12 – 26. VIII. 2008, 0 ° 33.823 ’ S 30 ° 21.490 ’ E, 1505 m, MT, primary mid-altitude rainforest, S. van Noort leg., 2 ♀♀, 2 ♂♂ (SAMC); same locality label, 2 – 12. VIII. 2008, 0 ° 33.836 ’ S 30 ° 21.700 ’ E, 1523 m, MT, primary mid-altitude rainforest, S. van Noort leg., 2 ♀♀, 3 ♂♂ (SAMC); Western Region, Kasese Dis- trict, Ruwenzori Range, Misigo, 8550 ft., 2 – 3. VIII. 1952, D. S. Fletcher leg., 3 ♂♂ (NHMUK, BPBM). ZIMBA- BWE: Harare (= Salisbury), Chishawasha, VI. 1978, A. Watsham leg., 1 ♂ (NHMUK); Harare, Saint Ignatius, V. 1990, MT, A. Watsham leg., 1 ♀ (PMA). YEMEN: Sana’a, I. 1993, MT, A. van Harten leg., 1 ♂ (MOLC); same locality label, 2. III. 1998, MT, 3 ♀♀ (MOLC); same locality label, X. 1998, pitfall-trap, 1 ♀ (MOLC); Ta’izz, X. 1999, light trap, A. van Harten & A. Awad leg., 1 ♀ (MOLC); same locality label, V – VI. 2000, A. van Harten & A. R. Al Yarimi leg., 1 ♀ (MOLC); same locality label, VI. 2002, A. van Harten & A. R. Al Yarimi leg., 1 ♀ (MOLC); Lahj, III. 2001, MT, A. van Harten & A. Sallam leg. (MOLC); Al-Kowd, 1 – 5. IX. 2001, I – III. 2003, light trap, A. van Harten & S. Al Haruri leg., 2 ♂♂ (MOLC); Al Kadan, I. 2003, light trap, A. van Harten & T. Abdul Haq leg., 2 ♂♂ (MOLC); W Ye- men, 20 Km NW Dhawran, 14 ° 40 ’ N 44 ° 13 ’ E, 1800 m, 29. X. 2005, J. Halada leg., 1 ♀ (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9A2A73FF3ED5FFFC77FC90.taxon	biology_ecology	Hosts. Typhlocybinae (Cicadellidae) (Guglielmino et al. 2013): in North Sudan: Jacobiasca lybica (Bergevin & Zanon); in Somalia: Jacobiella facialis (Jacobi).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF9A2A73FF3ED5FFFC77FC90.taxon	distribution	Distribution. Angola, Botswana, Burundi, Cameroon, Central African Republic, Democratic Republic of the Congo, Gabon, Ghana, Ivory Coast, Kenya, Madagascar, Mozambique, Nigeria, North Sudan, Rwanda, Senegal, Somalia, South Africa, Tanzania, Togo, Uganda, Yemen, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF942A73FF3ED527FAB6F8F0.taxon	diagnosis	Diagnosis. ♀: fully winged (Fig. 18 A); epistomal suture touching antennal toruli (as in Fig. 18 C); occipital ca- rina complete; antenna without ADOs; palpal formula 5 / 2, 5 / 3, 6 / 2, or 6 / 3; forewing with only costal cell enclosed by pigmented veins (as in Fig. 18 D) (rarely with C and R cells completely enclosed by pigmented veins: character not present in Afrotropical species (Figs 18 A, B )); forewing with pterostigma present and 2 r-rs & Rs vein straight (as in Fig. 18 D), or angled between proximal and distal parts (character not present in Afrotropical species (Figs 18 A, B )); course of forewing veins not marked by dark stripes; hind wing hyaline, with costal vein, without dark medial longitudinal stripe (Fig. 18 A); protarsus not chelate; tibial spurs 1 / 1 / 2. ♂: fully winged winged (Fig. 19); epistomal suture touching antennal toruli (Fig. 18 C); occipital carina complete; palpal formula 5 / 2, 5 / 3, 6 / 2, or 6 / 3; forewing with only costal cell enclosed by pigmented veins (Figs 18 C) (or with C and R cells completely enclosed by pigmented veins (as in Figs 18 A, B): character not present in Afrotropical species); forewing with pterostigma present and 2 r-rs & Rs vein straight (Fig. 18 D), or angled between proximal and distal parts (character not present in Afrotropical species); course of forewing veins not marked by dark stripes; hind wing (Fig. 19) hyaline, with costal vein, without dark medial longitudinal stripe; basivolsella situated completely below distivolsella apex (Figs 20 A- C); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF942A73FF3ED527FAB6F8F0.taxon	distribution	Distribution. Afrotropical, Australian, Oriental, Nearctic and Neotropical regions. Present at the Oriental bor- der of the Palaearctic region (Nepal).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF942A73FF3ED527FAB6F8F0.taxon	biology_ecology	Hosts. Membracidae (Guglielmino & Olmi 1997).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF942A73FF3ED527FAB6F8F0.taxon	discussion	World species. 16 species are known, one in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF942A73FF3ED527FAB6F8F0.taxon	biology_ecology	Biology. Species of Crovettia are known for their completely endophagous parasitism and poliembriony. Their biology was studied in Nearctic species (Matausch 1909, 1911; Kornhauser 1916, 1917, 1919; Gahan 1918).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	description	(Figs 18 D, E, 19, 20 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	diagnosis	Diagnosis. ♀ and ♂ of Crovettia with head and mesoscutum granulate (Fig. 19); 2 r-rs & Rs vein straight and with apex not broadened (Fig. 18 D); metapectal-propodeal disc and propodeal declivity reticulate rugose.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	description	Description. ♀. Fully winged; body length 1.6 – 1.7 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma black; metasoma brown; legs yellow-testaceous. Antenna clavate; antennomeres in following proportions: 5: 5: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 4: 7. Head dull, granulate; frontal line absent; occipital carina complete; POL = 5; OL = 3; OOL = 3.5; OPL = 3; TL = 4; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum and mesoscutellum dull, granulate. Notauli absent. Metanotum dull, rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity with tracks of two longitudinal keels. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein straight and with apex not broadened (as in Fig. 18 D); pterostigma large and black, occasionally brown. Tibial spurs 1 / 1 / 2. ♂. Fully winged (Fig. 19); body length 1.4 – 1.8 mm (holotype 1.4 mm). Head black, except mandible testaceous; antenna brown or testaceous; mesosoma black; metasoma brown; legs completely yellow-testaceous, or testaceous, except meso- and metafemur, meso- and metatibia and last tarsomere darkened. Antenna filiform; antennomeres in following proportions: 3.5: 3.5: 3: 3: 3.5: 4: 4.5: 5: 5: 7. Head (Fig. 19 C) dull, granulate; frontal line incomplete, present in anterior third of frons; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 1; TL = 3; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum and mesoscutellum dull, granulate. Notauli absent or incomplete (in holotype reaching about 0.5 × length of mesoscutum). Metanotum dull, rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing (Fig. 19 D) hyaline, without dark transverse bands; 2 r-rs & Rs vein straight and with apex not broadened (Fig. 18 D); pterostigma large and black, occasionally brown. Paramere much (Fig. 20 A) or slightly (Figs 20 B, C) longer than penis; basivolsella usually with two distal bristles (Figs 20 A-C), rarely (in specimen from Kenya, 0.62884 ° S 36.42969 ° E) with one distal bristle. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19454): MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ‘ 52 ‘‘ S 49 ° 10 ‘ 53 ‘‘ E, 960 m, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg. (CAS). Paratypes: KENYA: Coast Prov., Arabuko-Sokoke Forest, 3.30958 ° S 39.96538 ° E, 80 m, 15 – 29. XII. 2014, MT in bush outside forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 13 – 27. XII. 2011, MT, below Bura Bluff, riverine forest, R. Copeland leg., MO / 346, 1 ♂ (MOLC); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 1 – 15. XI. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 7 – 21. II. 2015, 1 ♀ (NMK); same locality label, 27. XII. 2015 – 9. I. 2016, 1 ♂ (NMK); same locality label, 5 – 19. XI. 2015, 1 ♀ (NMK); same locality label, 27. IX – 11. X. 2015, 1 ♀ (NMK); Rift Valley Prov., Mlozi, Naivasha, 0.62884 ° S 36.42969 ° E, 1909 m, 5 – 6. VII. 2015, MT, Acacia xanthophloea woodland, R. Copeland leg., MO / 347, 1 ♂ (MOLC). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ‘ S 49 ° 10.45 ‘ E, 1125 m, 6 – 21. IV. 2001, MT, MA- 01 - 01 D- 07, R. Harin’Hala leg., 1 ♂ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 S 49 ° 11 ‘‘ E, 975 m, 4 – 10. III. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 06, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ‘ S 47 ° 22.19 ‘ E, at broken bridge, 1110 m, 15 – 21. XII. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 07, 2 ♂♂ (CAS, MOLC); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ‘ S 47 ° 24.43 ‘ E, 1130 m, 15 – 21. XII. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 07, 1 ♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♂ (CAS), 1 ♀ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 1 ♀ (MOLC); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 8 – 15. XI. 2001, MT, secondary tropical forest, MA- 02 - 09 C- 02, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♀ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	distribution	Distribution. Kenya, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF932A76FF3ED6D7FB4CFB94.taxon	etymology	Etymology. The species is named afra after the collection site in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF912A76FF3ED22BFBDDF95A.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 21 A, B, 30 A, B); occipital carina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); ocelli present; antenna without ADOs; palpal formula 5 / 3 or 6 / 3; pronotal tubercle present; forewing with two cells enclosed by pigmented veins (C and R) (Figs 21 A, B, 30 A, B); forewing with 2 r-rs & Rs vein and pterostigma (Figs 21 A, B, 30 A, B); protarsus chelate (Figs 21 A, B, 30 A, B); chela with rudimentary claw (Figs 26 A, 29 D); tibial spurs 1 / 1 / 2. ♂: fully winged (Figs 21 C, 30 C, D); occipital carina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); palpal formula 5 / 3 or 6 / 3; forewing with two cells enclosed by pigmented veins (C and R) (Figs 21 C, 30 C, D); forewing with 2 r-rs & Rs vein and pterostigma (Figs 21 C, 30 C, D); basivolsella situated completely below distivolsella apex (Figs 27 C, 29 E); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF912A76FF3ED22BFBDDF95A.taxon	distribution	Distribution. Palaearctic, Afrotropical and Oriental regions.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF912A76FF3ED22BFBDDF95A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF912A76FF3ED22BFBDDF95A.taxon	discussion	World species. 19 species are known, 11 in the Afrotropical region. World genera. Two genera are known, both present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF902A08FF3ED395FC20FF7C.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 21 A, B); occipital carina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); antenna without ADOs; palpal formula 5 / 3 or 6 / 3; pronotal tubercle present; forewing with two cells enclosed by pigmented veins (C and R) (Figs 21 A, B); forewing with 2 r-rs & Rs vein and pterostigma (Figs 21 A, B); distal part of 2 r-rs & Rs vein longer than proximal part (Figs 21 A, B); protarsus chelate (Figs 21 A, B); chela with rudimentary claw (Fig. 26 A); tibial spurs 1 / 1 / 2. ♂: fully winged (Fig. 21 C); occipital ca- rina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); palpal formula 5 / 3 or 6 / 3; forewing with two cells enclosed by pigmented veins (C and R) (Fig. 21 C); forewing with 2 r-rs & Rs vein and pterostigma (Fig. 21 C); distal part of 2 r-rs & Rs vein longer than proximal part (Fig. 21 C); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF902A08FF3ED395FC20FF7C.taxon	distribution	Distribution. Afrotropical and Oriental regions. Present at Oriental border of the Palaearctic (Nepal).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF902A08FF3ED395FC20FF7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF902A08FF3ED395FC20FF7C.taxon	discussion	World species. 14 species are known, 10 in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEE2A0AFF3ED3F2FE14FC20.taxon	description	(Figs 24 C, D, 26 A, 27 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEE2A0AFF3ED3F2FE14FC20.taxon	description	Description. ♀ (Fig. 24 C). Fully winged; body length 3.3 mm. Head black, except mandible, clypeus, malar space and anterior third of frons whitish; antenna testaceous, except antennomere 10 slightly darkened; mesosoma black, except pronotal tubercle and part of pronotum sides whitish; metasoma brown; legs completely whitish. Antenna clavate; antennomeres of holotype in following proportions: 12: 6: 15: 14: 11: 9: 8: 6: 5.5: 7. Head dull; frons with anterior half reticulate rugose and posterior half reticulate rugose with numerous longitudinal keels; vertex partly reticulate rugose, with some longitudinal keels behind and on sides of lateral ocelli; temple reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 7; OPL = 8; TL = 9; greatest breadth of lateral ocelli shorter than OL (3: 4). Mesoscutum with anterior third reticulate rugose and remaining surface strongly punctate and unsculptured among punctures. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum punctate, unsculptured among punctures. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hya- line, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 10). Protarsomeres in following proportions: 19: 5: 4: 12: 18. Enlarged claw (Fig. 26 A) without teeth or bristles or setae. Protarsomere 5 (Fig. 26 A) with one row of approximately 49 short and small lamellae; apex with 4 small lamellae, in addition to 3 longer lamellae. Tibial spurs 1 / 1 / 2. ♂ (Fig. 24 D). Fully winged; body length 3.1 – 3.4 mm. Head black, except mandible, clypeus, malar space and anterior third of frons whitish; antenna testaceous, except antennomere 10 slightly darkened; mesosoma black, except pronotal tubercle and part of pronotum sides whitish; metasoma brown; legs completely whitish. Antenna filiform; antennomeres in following proportions: 10: 4.5: 13: 12: 11: 9: 7.5: 6: 5.5: 7. Head dull; frons with anterior half reticulate rugose and posterior half reticulate rugose with more or less numerous longitudinal keels; vertex partly reticulate rugose, with some longitudinal keels behind and on sides of lateral ocelli; temple reticulate rugose; frontal line complete; occipital carina complete; POL = 7; OL = 3; OOL = 8; OPL = 8; TL = 9; greatest breadth of lateral ocelli shorter than POL (3.5: 7). Mesoscutum with anterior third or fifth reticulate rugose and remaining surface strongly punctate and unsculptured among punctures. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum punctate, unsculptured among punctures. Metanotum reticulate rugose. Metapectalpropodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 10). Genitalia in Fig. 27 C. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEE2A0AFF3ED3F2FE14FC20.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Mahajanga, Mahavavy River, 6.2 km 145 ° SE Mitsinjo, 20 m, 1 – 5. XII. 2002, YPT, in gallery forest, Fisher, Griswold et al. leg., BLF 6935 (CAS). Paratypes: same locality label as holotype, 4 ♂♂ (3 in CAS, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEE2A0AFF3ED3F2FE14FC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEE2A0AFF3ED3F2FE14FC20.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	description	(Figs 25, 27 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	diagnosis	Diagnosis. ♂ of Conganteon with head and mesosoma mostly testaceous-whitish (Fig. 25); head and mesoscutum punctate, unsculptured among punctures; paramere (Fig. 27 A) with inner margin of distal part not provided with short sensorial processes.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	description	Description. ♂ (Fig. 25). Fully winged; body length 2.3 – 2.9 mm. Head testaceous-whitish, except brown spots around ocelli; antenna testaceous-whitish; mesosoma testaceous-whitish, except anterior margin of mesoscutellum, most of metanotum (except posterior margin testaceous) and metapectal-propodeal disc brown; metasoma brown; legs completely testaceous-whitish. Antenna filiform; antennomeres in following proportions: 9: 3: 10: 10: 9: 9: 8: 6: 6: 7. Head shiny, strongly punctate, unsculptured among punctures; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 7; OPL = 7; TL = 7; greatest breadth of lateral ocelli shorter than OL (3: 4). Mesoscutum strongly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum punctate, unsculptured among punctures. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 7). Paramere (Fig. 27 A) with inner margin of distal part not provided with short sensorial processes. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	materials_examined	Material examined. Types: ♂ holotype, NMK / INV / T- 639: KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 5 – 19. XI. 2015, MT, just inside isolated woodland patch, R. Copeland leg. (NMK). Paratypes: same locality label as holotype, 4 ♂♂ (3 in NMK, NMK / INV / T- 640, 641, 642; 1 in MOLC, MO / 344).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFED2A0AFF3ED5B7FE89F8F4.taxon	etymology	Etymology. The species is named after William Anderson Hawley, epidemiologist, evolutionary biologist and crackerjack birder.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEC2A0DFF3ED169FE1FFE54.taxon	description	(Figs 21 D, 24 A, 26 B, 27 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEC2A0DFF3ED169FE1FFE54.taxon	description	Description. ♀. Fully winged; body length 2.8 – 3.4 mm (holotype 3.4 mm). Head black, except mandible, clypeus, malar space and anterior third or fourth of frons whitish; antenna testaceous, except antennomeres 7 – 10 slightly darkened; mesosoma of type specimens black, except pronotal tubercle and part of lateral regions of pronotum whitish (in ♀♀ from South Africa, KwaZulu-Natal, mesosoma completely black); metasoma brown; legs completely whitish. Antenna clavate; antennomeres of holotype in following proportions: 10: 5: 14: 11: 9.5: 8: 6.5: 6: 6: 8. Head dull; frons and vertex completely reticulate rugose or sculptured by very large punctures; occasionally frons with more or less large smooth and finely punctured area in front of anterior ocellus; rarely (in ♀ from South Africa, Cathe- dral Peak Nature Reserve) posterior half of frons sculptured by numerous parallel longitudinal and irregular keels; temple reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3.5; OOL = 9; OPL = 7; TL = 9; greatest breadth of lateral ocelli shorter than POL (3: 6). Mesoscutum reticulate rugose, except posterior third or half strongly punctate and unsculptured among punctures. Notauli absent, or shortly present near anterior margin of mesoscutum. In specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, notauli reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing partly hyaline and partly slightly darkened; distal part of 2 r-rs & Rs vein longer than proximal part (14: 9). Protarsomeres of holotype in following proportions: 15: 4: 4: 12: 15. In other ♀♀ from South Africa, KwaZulu-Natal, including some paratypes, protarsomere 1 about twice as long as 4. Enlarged claw (Fig. 26 B) with 1 small subapical tooth, without bristles or setae. Protarsomere 5 (Fig. 26 B) with one row of approximately 31 short and small lamellae; apex with about 5 small lamellae, in addition to 3 longer lamellae. Tibial spurs 1 / 1 / 2. ♂ (Figs 21 D, 22 A). Fully winged; body length 2.7 – 3.3 mm. Head black, except mandible, clypeus, malar space and anterior third of frons whitish; antenna totally testaceous, or with antennomeres 7 – 10 slightly darkened; mesosoma black, except pronotal tubercle and some × part of pronotum sides whitish (in 1 ♂ from South Africa, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, pronotal tubercle black); metasoma brown; legs completely whitish. Antenna filiform; antennomeres in following proportions: 9: 4: 12.5: 10: 10.5: 10.5: 9: 8: 6.5: 9. Head dull; frons and vertex reticulate rugose, or sculptured by very large punctures; occasionally frons with more or less large smooth and punctate area in front of anterior ocellus; temple reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 8.5; OPL = 7; TL = 10; greatest breadth of lateral ocelli shorter than POL (3: 6). Mesoscutum reticulate rugose, except posterior third or half punctate and unsculptured among punctures. Notauli absent, or shortly visible near anterior margin of mesoscutum. In specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing partly hyaline and partly slightly darkened; distal part of 2 r-rs & Rs vein longer than proximal part (13: 10). Genitalia in Fig. 27 B. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEC2A0DFF3ED169FE1FFE54.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, East London, 32 ° 58.9 ’ S 27 ° 53.2 ’ E. 375 ft., 24 – 29. XII. 2004, MT, M. Mostovski leg. (NMSA). Paratypes: SOUTH AFRICA, same locality label as holotype, 1 ♀, 2 ♂♂ (NMSA), 1 ♀, 1 ♂ (MOLC); KwaZulu-Natal, Pietermaritzburg, Karkloof Nature Reserve (East), Unit 4, 22 – 24. IV. 2005, YPT, V. Kolyada leg., 8 ♀♀, 5 ♂♂ (NMSA); KwaZulu-Natal, Pietermaritzburg, Ferncliff Nature Reserve, 29 ° 33.2 ’ S 30 ° 20.6 ’ E, 850 m, 6 – 8. IV. 2005, YPT, V. Kolyada leg., 3 ♀♀ (NMSA). Other material: SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 6. X. 2010 – 18. I. 2011, MT, Camdeboo Escarpment Ticket, S. van Noort leg., PCD 09 - ACA 1 - M 06, 3 ♂♂ (2 in SAMC, 1 in MOLC); Eastern Cape, Huntly Glen Farm, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, 1065 m, 6. X. 2010 – 18. I. 2011, YPT, Great Fish Thicket, HUN 10 - ACA 1 - Y 04, S. van Noort leg., 2 ♀♀, 3 ♂♂ (SAMC), 1 ♂ (MOLC); same locality label, 6. X. 2010 – 18. I. 2011, HUN 10 - ACA 1 - Y 05, 2 ♀♀, 6 ♂♂ (SAMC); Free State, Sterkfontein Dam, 28 ° 29.72 ’ S 23 ° 57.85 ’ E, 1780 m, 10. XII. 2005, YPT, M. Mostovski leg., 5 ♂♂ (NMSA); KwaZulu-Natal, Karkloof, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, 1325 m, 22. I – 6. II. 2007, YPT, Kolyada & Mostovski leg., 2 ♀♀ (NMSA, MOLC); same locality label, 8 – 11. XI. 2007, 1 ♂ (NMSA); same locality label, 28. IX – 3. X. 2005, YPT, M. Mostovski leg., 3 ♀♀, 4 ♂♂ (NMSA), same locality label, 25. VII – 25. IX. 2005, MT, M. Mostovski leg., 2 ♀♀, 1 ♂ (NMSA); KwaZulu-Natal, Louwsberg, Sanyati Farm, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 18 – 20. IV. 2007, YPT, M. Mostovski leg., 1 ♀, 4 ♂♂ (NMSA); same locality la- bel, 1 ♂ (MOLC); same locality label, 30. X – 18. XII. 2005, MT, 9 ♂♂ (NMSA); same locality label, 1 – 24. VI. 2006, MT, 10 ♂♂, 1 ♀ (NMSA); KwaZulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 20. IV – 26. X. 2007, MT, Kolyada & Mostovski leg., 2 ♀♀, 1 ♂ (NMSA); same locality label, 26 – 28. X. 2007, YPT, 10 ♀♀, 18 ♂♂ (NMSA); same locality label, 23 – 24. II. 2007, YPT, 2 ♀♀, 13 ♂♂ (NMSA); KwaZulu-Natal, La Mercy, Site 1, 29 ° 37.41 ’ S 31 ° 06.454 ’ E, 85 m, 6. XII. 2006, YPT, Mostovski leg., 1 ♀, 3 ♂♂ (NMSA); KwaZulu-Natal, Ca- thedral Peak Nature Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 22. IX – 17. XI. 2006, MT, M. Mostovski leg., 4 ♀♀ (NMSA); same locality label, 14. XII. 2005 – 29. I. 2006, MT, 1 ♂ (NMSA); KwaZulu-Natal, Ngoye Forest, Jan. – Apr. 2006, MT, G. Davies leg., 4 ♀♀, 6 ♂♂ (NMSA); KwaZulu-Natal, Pietermaritzburg, Winterskloof, 29 ° 34.56 ’ S 30 ° 17.40 ’ E, 1085 m, 15 – 19. II. 2007, YPT, Kolyada & Mostovski leg., 8 ♂♂, 3 ♀♀ (NMSA); KwaZulu- Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 14 – 27. II. 2006, MT, G. Whiteley leg., 1 ♂ (DJBC); Western Cape Prov., Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 26. X. 2010 – 26. I. 2011, MT, Afromontane Forest, S. van Noort leg., GVB 10 - FOR 1 - M 07, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEC2A0DFF3ED169FE1FFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEC2A0DFF3ED169FE1FFE54.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0DFF3ED46BFE59FACA.taxon	description	(Figs 22 B, C, 24 E, 26 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0DFF3ED46BFE59FACA.taxon	description	Description. ♀ (Figs 22 B, C, 24 E). Fully winged; body length 2.7 mm. Testaceous, except petiole black, one small brown spot on metapectal-propodeal disc near metanotum, brown teeth of mandible and brown borders of ocelli. Antenna clavate; antennomeres in following proportions: 9: 4: 12: 9: 7: 7: 6: 6: 5: 6. Head dull, granulate, sculptured by many hardly visible longitudinal and irregular striae also present behind ocellar triangle; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 6; OPL = 6; TL = 7; greatest breadth of lateral ocelli shorter than POL (3: 6). Pronotum short, transverse, rugose; pronotal tubercle reaching tegula. Mesoscutum with anterior half reticulate rugose and remaining surface unsculptured. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum unsculptured. Metanotum rugose (sculpture hardly visible). Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 7). Protarsomeres in following proportions: 14: 3: 3.5: 7.5: 13. Enlarged claw (Fig. 26 C) with 1 small preapical tooth. Protarsomere 5 (Fig. 26 C) with one row of approximately 37 short and small lamellae; apex with about 5 small lamellae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0DFF3ED46BFE59FACA.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Eastern Prov., Kasaala area, 2.07486 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, 6 m MT, woodland and grass, J. Bukhebi & R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0DFF3ED46BFE59FACA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0DFF3ED46BFE59FACA.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	description	(Figs 27 D, 28, 85 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	diagnosis	Diagnosis. ♂ of Conganteon with head and mesosoma mostly black; frons reticulate rugose, with anterior half whitish; pronotum with pronotal tubercle whitish; inner margin of distal part of paramere with many short sensorial processes (Fig. 27 D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	description	Description. ♂ (Figs 28, 85 A). Fully winged; body length 2.9 mm. Head black, except mandible, clypeus, ma- lar space and anterior half of frons whitish; antenna testaceous, except antennomeres 8 – 10 brown; mesosoma black, except pronotal tubercle whitish; metasoma brown; legs completely whitish. Antenna filiform; antennomeres in following proportions: 10: 4: 12: 13: 11: 10: 7: 6: 6: 7. Head (Fig. 28 C) dull, reticulate rugose; vertex behind ocellar triangle with many longitudinal keels; temple reticulate rugose; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 7; OPL = 8; TL = 10; greatest breadth of lateral ocelli about as long as OL. Mesoscutum (Fig. 28 A) strongly punctate, unsculptured among punctures, except anterior half reticulate rugose. Notauli absent. Mesoscutellum and metanotum punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing (Fig. 28 D) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 12). Paramere (Fig. 27 D) with inner margin of distal part provided with many short sensorial processes. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19453): MADAGASCAR: Mahajanga, Beanka Forest Re- serve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, YPT, tropical dry forest on tsingy, B. L. Fisher et al. leg., BLF 22788 (CAS). Paratype: MADAGASCAR: Mahajanga, Namoroka National Park, 16.40558 S 45.31165 E, 21 – 31. X. 2016, MT 8, C. Villemant leg., 1 ♂ (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFEA2A0EFF3ED3D1FF04FE54.taxon	etymology	Etymology. The species is named sensitivum because its paramere is provided with many short sensorial processes.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE92A0EFF3ED46BFE1FF816.taxon	description	(Figs 26 D, 27 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE92A0EFF3ED46BFE1FF816.taxon	description	Description. ♀. Fully winged; body length 2.8 mm. Head black, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa basally black. Antenna clavate; antennomeres in following proportions: 10: 5: 12: 9: 9: 8: 6: 6: 5: 5. Head dull, completely reticulate rugose; occasionally posterior region of vertex near occipital carina shiny and sculptured by longitudinal keels; frontal line present; occipital carina complete; POL = 7; OL = 4; OOL = 8; OPL = 8; TL = 9. Pronotum setose, very short. Mesoscutum shiny, completely reticulate rugose. Notauli absent. Mesoscutellum shiny, smooth, finely punctate, unsculptured among punctures. Metanotum shiny, strongly punctate, unsculptured among punctures. reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein curvilinear, with distal part longer than proximal part (13: 10). Protarsomeres in following proportions: 15: 3.5: 3: 10: 15; protarsomere 3 produced into hook. Enlarged claw (Fig. 26 D) without teeth, or bristles, or setae, or lamellae. Protarsomere 5 (Fig. 26 D) with one row of approximately 52 small lamellae extending continuously to apex; apex with 3 long lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.7 – 2.9 mm. Head black, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa basally black. In specimen from Madagascar, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, head black, except mandible testaceous; legs testaceous, except coxae and clubs of femora partly brown. Antenna filiform; antennomeres in following proportions: 10: 5: 10: 8: 8: 8: 6: 6: 5: 7. Head dull, completely reticulate rugose; posterior region of vertex near occipital carina shiny and sculptured by longitudinal keels situated among the ocelli; frontal line present; occipital carina complete; POL = 9; OL = 5; OOL = 8; OPL = 9; TL = 10. Pronotum setose, very short. Mesoscutum completely reticulate rugose. Notauli absent (in specimen from Madagascar, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, notauli incomplete, reaching about 0.5 × length of mesoscutum). Mesoscutellum and metanotum strongly punctate, unsculptured among punctures (in specimen from Madagascar, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, mesoscutellum partly rugose). Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 11). Paramere (Fig. 27 E) without proximal membranous process, in lateral view similar to that of Conganteon vulcanicum Benoit. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE92A0EFF3ED46BFE1FF816.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg (AEIC). Para- types: same locality label as holotype, 3 ♂♂ (2 in AEIC, 1 in AMNH). Other material: MADAGASCAR: Toliara, Ifaty, near Hotel Paradisia, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, 9 m, 26. V – 5. VI. 2002, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02 - 16 - 28, 1 ♂ (CAS). SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, Hilton, 24.12.2003 – 14. I. 2004, MT, M. Mostovski leg., 1 ♀ (NMSA); same locality label, 24. XI – 9. XII. 20013, 1 ♀ (SAMC); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 11. V – 24. VII. 2006, MT, G. Whiteley leg., 1 ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE92A0EFF3ED46BFE1FF816.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE92A0EFF3ED46BFE1FF816.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE82A01FF3ED147FE1FFA35.taxon	description	(Fig. 26 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE82A01FF3ED147FE1FFA35.taxon	description	Description. ♀. Fully winged; body length 3.0 mm. Head black, except mandible, clypeus and anterior margin of frons testaceous; antenna testaceous, except antennomeres 7 – 10 brown; mesosoma black; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 13: 8: 7: 6: 6: 6: 5: 8. Head dull, completely reticulate rugose; frontal line complete; frons without lateral keels; occipital carina complete; POL = 8; OL = 4; OOL = 7; OPL = 10; TL = 9. Pronotum very short, reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum with anterior half reticulate rugose and posterior half punctate, unsculptured among punctures. Notauli absent. Me- soscutellum finely punctate, unsculptured among punctures. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 9). Protarsomeres in following proportions: 16: 3: 4: 4: 9. Enlarged claw (Fig. 26 E) without teeth, or bristles, or setae, or lamellae. Pro- tarsomere 5 (Fig. 26 E) with two rows of 13 lamellae; apex with 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE82A01FF3ED147FE1FFA35.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Limpopo, 30 km E Nylstroom, 15. XI – 17. XII. 1984, H. & A. Howden leg. (PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE82A01FF3ED147FE1FFA35.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE82A01FF3ED147FE1FFA35.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE62A02FF3ED04AFE1FFE28.taxon	description	(Figs 22 D – F, 29 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE62A02FF3ED04AFE1FFE28.taxon	description	Description. ♂ (Figs 22 D – F). Fully winged; body length 2.8 mm. Head black, except mandible, clypeus and region between and on sides of antennal toruli testaceous; eye testaceous-whitish; antenna testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 11: 5: 12: 11: 11: 10: 9: 8: 7.5: 10. Antennomere 9 more than 3 × as long as broad (8: 2.4). Head dull, punctate, unsculptured among punctures; punctures very dense on frons; frons with track of complete frontal line; occipital carina complete; POL = 6.5; OL = 4; OOL = 9; OPL = 8; TL = 10. Pronotum very short, transverse, rugose. Mesoscutum and mesoscutellum shiny, smooth, punctate and unsculptured among punctures. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Metanotum dull, reticulate rugose, with smooth and punctate central region. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 11). Genitalia in Fig. 29 A. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE62A02FF3ED04AFE1FFE28.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Cape Town, Tokai State Forest, Con- stantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 27. VIII – 5. IX. 1993, MT, Mesic mountain Fynbos on Sandstone, near stream, bracken present, SAM-HYM-P 024852, S. van Noort leg. (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE62A02FF3ED04AFE1FFE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE62A02FF3ED04AFE1FFE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE42A05FF3ED09CFAACFC20.taxon	description	(Figs 21 A, B, 23 A – F, 26 F, 29 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE42A05FF3ED09CFAACFC20.taxon	description	Description. ♀ (Figs 21 A, B, 23 A – C). Fully winged; body length 2.4 – 3.0 mm. Head black, except mandible testa- ceous (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, head black, except mandible, clypeus and ante- rior third of frons yellowish); antenna brown, except scape and pedicel testaceous (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, antenna totally testaceous); mesosoma and metasoma black; legs testaceous, except coxae and metafemora partly black (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, legs totally yellow). Antenna clavate; antennomeres in following proportions: 9: 5: 10: 7: 7: 6: 6: 6: 5.5: 8. Head shiny, setose, completely reticulate rugose; occipital carina complete; temple distinct, completely reticulate rugose; POL = 2.5; OL = 5; OOL = 8; OPL = 6. Pronotal tubercle reaching tegula. Mesoscutum shiny, setose, completely reticulate rugose (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, mesoscutum reticulate rugose, except posterior third punctate, unsculptured among punctures). Notauli absent. Mesoscutellum shiny, with anterior half unsculptured and posterior half reticulate rugose. In ♀ from Kenya, Kisere Forest, mesoscutellum completely reticulate rugose; in ♀♀ from South Africa, Asante Sana Game Reserve, mesoscutellum unsculptured. Metanotum reticulate rugose. Metapectalpropodeal disc and propodeal declivity completely reticulate rugose, without keels. Forewing hyaline, without dark transverse bands. 2 r-rs & Rs vein angled between proximal and distal parts; distal part slightly longer than proximal part (9: 7); pterostigma brown, short and oval. Protarsomeres of holotype in following proportions: 11: 2: 2: 3: 7; protarsomere 3 produced into hook. In ♀ from Kenya, Kisere Forest, protarsomere 1 twice as long as 4. Enlarged claw (Fig. 26 F) without teeth, or bristles, or setae, or lamellae. Protarsomere 5 (Fig. 26 F) with 1 long row of 27 lamellae extending continuously to apex; apex with 5 lamellae, one of these very long. Tibial spurs 1 / 1 / 2. ♂ (Figs 23 D – F). Fully winged; body length 1.8 – 3.3 mm. Head black; mandible black, except distal half tes- taceous-brown (in ♂♂ from Kenya, 2.07846 ° S 38.22530 ° E and 2.01477 ° S 38.32618 ° E, clypeus, malar space and anterior margin of frons whitish; in ♂♂ from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, clypeus, malar space and anterior surface of frons between antennal toruli testaceous; in ♂♂ from Madagascar, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, clyp- eus totally black); antenna brown (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, antenna testaceous); mesosoma and metasoma black (in ♂♂ from Kenya, 2.07846 ° S 38.22530 ° E and 2.01477 ° S 38.32618 ° E, pronotal tubercle testaceous; in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, lateral regions of pronotum browntestaceous); legs black-brown, except tarsi testaceous (in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, legs testaceous). In ♂♂ from Kenya, 2 ° 27.90 ’ S 37 ° 54.91 ’ E and 2.01477 ° S 38.32618 ° E, clypeus testaceous. An- tenna filiform; antennomeres in following proportions: 6: 3: 8: 7: 7: 7: 6: 5: 6: 7. Head dull, completely reticulate rugose (in ♂♂ from Kenya, 2.07846 ° S 38.22530 ° E and 2.01477 ° S 38.32618 ° E, frons reticulate rugose and with some irregular longitudinal keels); frontal line present; occipital carina complete; POL = 4; OL = 2; OOL = 5; OPL = 4; TL = 7. In specimens from Kenya, 2.05111 ° S 38.32613 ° E, and South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, vertex with longitudinal keels and areolae between lateral ocelli and occipital carina. Mesoscutum, mesoscutellum and metanotum dull, completely reticulate rugose (in ♂♂ from Madagascar, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, posterior half of mesoscu- tum partly granulate and slightly or not rugose). In specimens from Kenya, 2.07846 ° S 38.22530 ° E and 2.01477 ° S 38.32618 ° E, mesoscutum with anterior half reticulate rugose and posterior half strongly punctate. Notauli absent (in specimens from Kenya, 2.07846 ° S 38.22530 ° E and 2.01477 ° S 38.32618 ° E, notauli hardly visible, incomplete, reaching about 0.5 × length of mesoscutum). Occasionally mesoscutellum and metanotum strongly punctate, unsculptured among punctures. In large ♂♂ (3.2 – 3.3 mm) from Madagascar, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, mesoscutum almost smooth near posterior margin, mesoscutellum shiny, finely punctate and unsculptured among punctures, and metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands. 2 r-rs & Rs vein angled between proximal and distal parts; distal part longer than proximal part (6: 5). Paramere without proximal membranous process and without distal inner pointed process (Fig. 29 B). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE42A05FF3ED09CFAACFC20.taxon	materials_examined	Material examined. Type: ♀ holotype: RWANDA: Burambi, Muhavura Volcano, 2325 m, 5. IX. 1934, Mission G. F. de Witte (MRAC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kyandolire, Bomboka (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo, 1280 m, 7 – 15. VII. 1955, P. Vanschuytbroeck leg., 2 ♂♂, 1 ♀ (AMNH); same locality label, many ♂♂, ♀♀ (MRAC). KENYA: Eastern Prov., Kibwezi Forest, 2 ° 27.90 ’ S 37 ° 54.91 ’ E, 13 – 20. XI. 1999, MT, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, 6 m MT, just inside woodland patch, J. Bukhebi & R. Copeland leg., 12 ♂♂, 1 ♀ (NMK), 2 ♂♂ (MOLC); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 5 – 19. XI. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 2 ♂♂ (NMK); same locality label, 13 – 27. XI. 2014, 1 ♂ (NMK); same locality label, 11 – 25. XI. 2016, 2 ♂♂ (NMK); Eastern Prov., Simisi area, 2.01477 ° S 38.32618 ° E, 653 m, 1 – 4. XII. 2013, MT, shru- bland near Kwandula Hill, J. Bukhebi and R. Copeland leg., 2 ♂♂ (NMK, MOLC); Eastern Prov., Simisi area, 2.05111 ° S 38.32613 ° E, 710 m, 28. XI – 1. XII. 2013, MT, near base of Yamalu Hill, J. Bukhebi and R. Copeland leg., 1 ♂ (NMK); Rift Valley Prov., Oloitokitok, 2.94456 ° S 37.50714 ° E, 1853 m, 30. IX – 14. X. 2011, MT, edge of indig- enous forest, R. Copeland leg., 1 ♀ (NMK); Western Prov., Kakamega District, Kisere Forest, 28. VI – 19. VII. 2001, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MOLC). MADAGASCAR: Antananarivo, An- karatra (MNHN); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 S 49 ° 11 ’’ E, 975 m, 4 – 10. III. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 06, 1 ♂ (CAS); Fianarantsoa, Miandritsara Forest, 40 Km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 28. XII. 2006 – 4. I. 2007, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 73, 1 ♂ (CAS); same locality label, 19 – 28. XII. 2006, 1 ♀ (CAS); same locality label, 27. II – 8. III. 2007, MA- 29 - 80, 1 ♀ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic gar- den near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 11, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 1 – 7. XI. 2001, 10 ♂♂ (CAS). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 5 ♂♂ (4 in SAMC, 1 in MOLC); Eastern Cape, Asante Sana Game Reserve, 32 ° 16.762 ’ S 24 ° 57.309 ’ E, 1186 m, 23. II – 7. IV. 2010, MT, Southern Karoo Riviere, S. van Noort leg., 1 ♀, 3 ♂♂ (2 in SAMC, 1 in MOLC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.930 ’ S 24 ° 56.975 ’ E, 1642 m, 23. II – 7. IV. 2010, MT, Southern Karoo Riviere, S. van Noort leg., 6 ♀♀ (5 in SAMC, 1 in MOLC); Eastern Cape, Huntly Glen Farm, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, 1065 m, 6. X. 2010 – 18. I. 2011, YPT, Great Fish Thicket, HUN 10 - ACA 1 - Y 04, S. van Noort leg., 2 ♀♀, 2 ♂♂ (SAMC); same locality label, HUN 10 - ACA 1 - Y 05, 1 ♂ (SAMC). UGANDA: Western Region, Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.891 ’ N 30 ° 21.468 ’ E, 1506 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE42A05FF3ED09CFAACFC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE42A05FF3ED09CFAACFC20.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, Madagascar, Rwanda, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE12A18FF3ED151FD73FA44.taxon	description	(Figs 24 A, B, 27 F, 29 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE12A18FF3ED151FD73FA44.taxon	description	Description. ♀ (Figs 24 A, B). Fully winged; body length 2.3 – 2.6 mm. Head black, except mandible and clypeus testaceous (in ♀ from South Africa, 32 ° 57.732 ’ S 18 ° 05.985 ’ E, clypeus black); antenna of holotype testaceous, except antennomeres 7 – 10 brown; antenna of ♀ paratype brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs of holotype testaceous; legs of ♀ paratype testaceous, except coxae and clubs of meso- and metafemora partly brown. Antenna clavate; antennomeres of holotype in following proportions: 8.5: 5: 8: 6: 5: 4. 5: 4.5: 4.5: 4: 6; antennomeres of ♀ paratype in following proportions: 8: 5: 8.5: 6: 5: 5: 4.5: 4: 4: 5. Head of holotype dull, almost completely sculptured by irregular longitudinal keels, except small punctate area in front of anterior ocellus; head of ♀ paratype shiny, punctate, unsculptured among punctures; sides of frons, temple and vertex behind ocellar triangle slightly or strongly rugose or sculptured by irregular keels; frontal line complete; frons without lateral keels; occipital carina complete. In ♀ from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, head dull, with frons granulate and sculp- tured by many irregular longitudinal keels; vertex and temple granulate and reticulate rugose. Head of holotype with POL = 5; OL = 3; OOL = 7; OPL = 4; TL = 10; greatest breadth of lateral ocelli shorter than OL (2: 3). Head of ♀ paratype with POL = 6; OL = 3; OOL = 7.5; OPL = 5; TL = 9; greatest breadth of lateral ocelli shorter than OL (2.5: 3). Pronotum very short, transverse, reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum punctate, unsculptured among punctures, rugose near anterior margin. In ♀ from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, mesoscutum with anterior half granulate and rugose and posterior half punctate and unsculptured among punctures. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum (0.5 × in ♀ from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E). Mesoscutellum and metanotum finely punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 7 in the holotype; 8.5: 8 in ♀ paratype). Protarsomeres of holotype in following proportions: 11: 2: 3: 4.5: 10.5; protarsomeres of paratype in following proportions: 11: 2.5: 2.5: 5: 11. Enlarged claw (Fig. 27 F) with 1 small subapical tooth, without bristles or setae. Protarsomere 5 (Fig. 27 F) with one row of 26 – 31 lamellae; apex with about 7 – 12 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.1 – 2.8 mm. Head black, except mandible testaceous (in 2 ♂♂ from South Africa, 32 ° 58.611 ’ S 18 ° 06.976 ’ E and 33 ° 37.622 ’ S 18 ° 24.259 ’ E, clypeus testaceous); antenna brown; mesosoma black; metasoma brown; legs brown, occasionally with protibia and protarsus testaceous. Antenna filiform; antennomeres in following proportions: 7: 4: 6.5: 7: 6: 5.5: 5: 5: 4.5: 5.5. Antennomere 9 about twice as long as broad: 4.5: 2.2. Head shiny, punctate, unsculptured among punctures; sides of frons, temple and vertex behind ocellar triangle usually slightly rugose or sculptured by irregular keels (occasionally smooth, not rugose, not sculptured by irregular keels); frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 6; OPL = 5; TL = 8; greatest breadth of lateral ocelli shorter than OL (2.5: 3). In ♂ from South Africa, 32 ° 57.732 ’ S 18 ° 05.985 ’ E, frons punctate and slightly granulate among punctures. Pronotum very short, transverse, rugose. Mesoscutum shiny, punctate, unsculptured among punctures, slightly rugose near anterior margin. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 7). Genitalia in Fig. 29 C. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE12A18FF3ED151FD73FA44.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 11. VIII – 6. IX. 1997, S. van Noort leg., SAM-HYM-P 024855 (SAMC). Paratypes: SOUTH AFRICA: Western Cape, Koeberg Nature Reserve, 33 ° 37.62 ’ S 18 ° 24.26 ’ E, 23. I – 20. II. 1998, YPT, S. van Noort leg., SAM-HYM-P 024853, 1 ♀ paratype (SAMC); same locality label, 20. III – 19. IV. 1998, YPT, S. van Noort leg., SAM- HYM-P 024854, 1 ♂ (SAMC); same locality label, 23. I – 20. II. 1998, YPT, S. van Noort leg., SAM-HYM-P 024853, 1 ♂ (SAMC), 1 ♂ (MOLC). Other material: MADAGASCAR: Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 18 – 28. IX. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 62, 1 ♀ (CAS); same locality label, 27. XII. 2005 – 6. I. 2006, 2 ♀♀ (CAS, MOLC). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 1 ♀ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.930 ’ S 24 ° 56.975 ’ E, 1642 m, 23. II – 7. IV. 2010, MT, Southern Karoo Riviere, S. van Noort leg., 6 ♀♀ (SAMC); Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.182 ’ S 19 ° 08.587 ’ E, 741 m, 27. VIII – 27. X. 2009, MT, Nieuwoudtville Shale, Renosterveld, S. van Noort leg., 1 ♀, 3 ♂♂ (SAMC); same locality label, 31 ° 24.492 ’ S 19 ° 08.978 ’ E, 767 m, 1 ♂ (SAMC); same locality label, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, 749 m, 1 ♀, 2 ♂♂ (SAMC); same locality label, 31 ° 24.293 ’ S 19 ° 09.215 ’ E, 758 m, 1 ♀, 1 ♂ (SAMC); same locality label, 31 ° 24.841 ’ S 19 ° 09.551 ’ E, 797 m, 1 ♀, 1 ♂ (SAMC); same locality label, 31 ° 23.802 ’ S 19 ° 08.799 ’ E, 752 m, 1 ♀, 5 ♂♂ (SAMC); Northern Cape, 6.41 km 295 ° WNW Nieuwoudtville, 31 ° 22.350 ’ S 19 ° 02.752 ’ E, Fynbos 2 Site, 10 – 13. X. 2000, old fynbos on sandstone, MT, S. van Noort leg., 1 ♀ (SAMC); same locality label, 13. X. 2000, sweep, 1 ♀ (SAMC); Northern Cape, 4.66 km 294 ° WNW Nieuwoudtville, 31 ° 22.451 ’ S 19 ° 03.845 ’ E, Fynbos 1 Site, 10. X. 2000, old fynbos on sandstone, sweep, S. van Noort leg., 1 ♀ (SAMC); Northern Cape, Avontuur Farm, 16 km NW Nieuwoudtville, 1 ° 16.249 ’ S 19 ° 02.900 ’ E, 764 m, 27. VIII – 27. X. 2009, YPT, Bokkeveld Sandstone Fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, MT, 1 ♀ (SAMC); Northern Cape, Swaarweerberg, Vredehoek Farm, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, 1613 m, 29. VII – 30. IX. 2010, MT, Succulent Karoo, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 29. VIII – 28. X. 2009, 1 ♀ (SAMC); same locality label, 28. X. 2009 – 9. II. 2010, 1 ♀ (SAMC); Western Cape, Worcester, 17 – 31. VIII. 1928, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, Montagu, 23 – 30. IX. 1924, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, West Coast Fossil Park, 5 km 290 ° E Langebaanweg, 32 ° 57.732 ’ S 18 ° 05.985 ’ E, 16 – 23. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀, 6 ♂♂ (SAMC); same locality label, 9 – 16. X. 2002, 2 ♂♂ (SAMC); same locality label, 11 – 18. IX. 2002, 1 ♀, 7 ♂♂ (SAMC); same locality label, 18 – 25. IX. 2002, 2 ♀♀, 7 ♂♂ (SAMC); same locality label, 4 – 11. IX. 2002, 1 ♀, 3 ♂♂ (SAMC); Western Cape, Langberg Farm, 3 km 280 ° W Langebaanweg, 32 ° 58.045 ’ S 18 ° 07.418 ’ E, 18 – 25. IX. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 8 ♂♂ (SAMC); same local- ity label, 4 – 11. IX. 2002, 2 ♂♂ (SAMC); Western Cape, Anyskop Farm, 4 km 270 ° W Langebaanweg, 32 ° 58.611 ’ S 18 ° 06.976 ’ E, 11 – 18. IX. 2002, MT, Fynbos on Limestone, S. van Noort leg., 7 ♂♂ (SAMC); same locality label, 18 – 25. IX. 2002, 7 ♂♂ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 31. VII – 10. IX. 2009, MT, Renosterveld, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 39.514 ’ S 21 ° 53.947 ’ E, 322 m, 24. VII – 4. X. 2010, MT, Gamka Thicket, GB 09 - SUC 4 - M 44, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Cape Town, Constantiaberg, above Donkerboskloof, 34 ° 2.175 ’ S 18 ° 23.528 ’ E, 460 m, 19. IX – 10. X. 1995, MT, Mesic Fynbos with kloof forest elements, S. van Noort leg., 2 ♀♀ (SAMC, MOLC), 7 ♂♂ (6 in SAMC, 1 in MOLC); Western Cape, Cape Town, Constantiaberg, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.30 ’ E, 460 m, 29. VIII. 1995, sweep, Mesic Mountain Fynbos near stream, Halleria lucida present, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.62 ’ S 18 ° 24.26 ’ E, 5. IX – 3. X. 1997, MT, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 3 – 31. X. 1997, MT, West Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC); same locality label, 27. XII. 1997 – 23. I. 1998, 1 ♀ (SAMC); same local- ity label, 16. V – 13. VI. 1997, 3 ♂♂ (SAMC); same locality label, 11. VII – 8. VIII. 1997, 2 ♂♂ (SAMC); same locality label, 8. VIII – 5. IX. 1997, 9 ♂♂, 3 ♀♀ (SAMC); same locality label, 5. IX – 3. X. 1997, 58 ♂♂, 33 ♀♀ (SAMC); same locality label, 8. VIII – 5. IX. 1997, 43 ♂♂, 30 ♀♀ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 8. VIII – 2. X. 2013, MT, Burnt Mesic Mountain Fynbos, BH 12 - FYN 3 - M 12, S. van Noort leg., 4 ♀♀ (SAMC); same locality label, 23. VIII – 28. IX. 2012, BH 12 - FYN 3 - M 02, 2 ♀♀ (SAMC); same locality label, 2 – 20. X. 2013, BH 12 - FYN 3 - M 13, 2 ♀♀ (SAMC); same locality label, 3 – 25. IV. 2013, BH 12 - FYN 3 - M 07, 2 ♀♀ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.824 ’ S 18 ° 58.123 ’ E, 400 m, 12. IX – 7. X. 2014, MT, Burnt Mesic Mountain Fynbos, BH 12 - FYN 6 - M 21, S. van Noort leg., 1 ♀, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE12A18FF3ED151FD73FA44.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFE12A18FF3ED151FD73FA44.taxon	distribution	Distribution. Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFF2A19FF3ED05CFC02FF7C.taxon	type_taxon	Type species. Fiorianteon junonium Olmi, 1984, by original designation.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFF2A19FF3ED05CFC02FF7C.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 30 A, B); occipital carina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); antenna without ADOs; palpal formula 6 / 3; pronotal tubercle present; forewing with two cells enclosed by pigmented veins (C and R) (Figs 30 A, B); forewing with 2 r-rs & Rs vein and pterostigma present (Figs 30 A, B); distal part of 2 r-rs & Rs vein as long as, or shorter than proximal part (Figs 30 A, B); protarsus chelate (Figs 30 A, B); chela with rudimentary claw (Fig. 29 D); tibial spurs 1 / 1 / 2. ♂: fully winged (Figs 30 C, D); occipital carina complete; mandible quadridentate, including one intermediate rudimentary tooth (Fig. 8 B); palpal formula 6 / 3; forewing with two cells enclosed by pigmented veins (C and R) (Figs 30 C, D); forewing with 2 r-rs & Rs vein and pterostigma present (Figs 30 C, D); distal part of 2 r-rs & Rs vein as long as, or shorter than proximal part (Figs 30 C, D); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFF2A19FF3ED05CFC02FF7C.taxon	distribution	Distribution. Palaearctic, Afrotropical and Oriental regions.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFF2A19FF3ED05CFC02FF7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFF2A19FF3ED05CFC02FF7C.taxon	discussion	World species. Five species are known, one in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFE2A1BFF3ED035FE14FF50.taxon	description	(Figs 29 E, 31 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFE2A1BFF3ED035FE14FF50.taxon	description	Description. ♂ (Figs 31 A – C). Fully winged; body length 2.8 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except most part of coxae black. Antenna filiform; antennomeres in following proportions: 11: 5: 13: 14: 13: 12: 10: 9: 8: 10. Head shiny, completely sculptured by longitudinal subparallel keels on frons, vertex and temple (Fig. 31 C); frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 7; TL = 10; greatest breadth of lateral ocelli about as long as OL. Mesoscutum shiny, with anterior half reticulate rugose; posterior half, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum punctate, unsculptured among punctures. Metanotum dull, rugose. Metapectal-propodeal disc (Fig. 31 B) reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein about as long as proximal part, about as long as antennomere 3. Paramere (Fig. 29 E) with many sensorial processes on inner margin of distal part. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFE2A1BFF3ED035FE14FF50.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Fianarantsoa, Andringitra National Park, Plateau d’Andohariana, 35.9 km 205 ° Ambalavao, 22 ° 09.08 ’ S 46 ° 53.57 ’ E, 2000 m, 15. IV. 2006, MT, B. L. Fisher et al. leg., BLF 13755 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFE2A1BFF3ED035FE14FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFE2A1BFF3ED035FE14FF50.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 31 D, E, 80 A, B, 97 A, B), occasionally brachypterous (Figs 41 A, B) or mi- cropterous (Figs 81 C, D; 84 A); occipital carina complete; mandible quadridentate, with teeth becoming regularly progressively larger from dorsal to ventral tooth (as in Fig. 6 B); antenna without tufts of long setae and without ADOs; ocelli present; palpal formula 6 / 3; pronotal tubercle present; epicnemium exposed; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 31 D, E, 80 A, B, 97 A, B); forewing with 2 r-rs & Rs vein and pterostigma (Figs 31 D, E, 80 A, B, 97 A, B); protarsus chelate (Figs 31 D, E, 80 A, B, 97 A, B); chela without rudimentary claw (Figs 33 C, 83 B, 96 F); protrochanter short and broad; tibial spurs 1 / 1 / 2. ♂: fully winged (Figs 32 A, B, 80 C, D, 97 C, D), rarely brachypterous; occipital carina complete; mandible quadridentate, with teeth becoming regularly progressively larger from dorsal to ventral tooth (Fig. 6 B); palpal formula 6 / 3; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 32 A, B, 80 C, D, 97 C, D); forewing with 2 r-rs & Rs vein and pterostigma (Figs 32 A, B, 80 C, D, 97 C, D); forewing with PostabR 1 shorter than pterostigma (Figs 32 A, B, 80 C, D, 97 C, D); hypopygium not umbrella-shaped (Fig. 8 H); basivolsella completely situated down distivolsella (Figs 35 F, 86 A); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	discussion	World species. 651 species are known, 138 in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	discussion	World genera. Seven genera are known, three in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFFC2A1CFF3ED110FAB6FBA1.taxon	discussion	Remarks. The following four genera have not been recorded from the Afrotropical region: Anteonopsis Olmi, Rasnitsyn & Guglielmino, 2010 (fossil from Russian Far East, Magadan Region), Burmanteon Engel, 2003 (fossil from Burmese amber), Janzeniola Olmi, 2011 (fossil in Baltic amber), Metanteon Olmi, 1984 (Neotropical).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF82A10FF3ED50DFB76FD48.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 31 D, E); rarely brachypterous (Figs 41 A, B); occipital carina complete; palpal formula 6 / 3; antenna without ADOs; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 31 D, E); forewing with 2 r-rs & Rs vein and pterostigma (Figs 31 D, E); distal part of 2 r-rs & Rs vein much shorter than proximal part (Figs 31 D, E), occasionally slightly shorter, as long as, or longer than proximal part; metapectal-propodeal disc usually with transverse posterior keel (Fig. 34 A); protarsus chelate (Figs 31 D, E); inner side of enlarged claw with proximal prominence bearing one long bristle (Fig. 33 C); tibial spurs 1 / 1 / 2. ♂: fully winged (Figs 32 A, B); rarely brachypterous; occipital carina complete; vertex of head usually without two oblique keels connecting lateral ocelli to occipital carina; palpal formula 6 / 3; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 32 A, B); forewing with 2 r-rs & Rs vein and pterostigma (Figs 32 A, B); distal part of 2 r-rs & Rs vein much shorter than proximal part (Figs 32 A, B), occasionally slightly shorter, as long as, or longer than proximal part; pterostigma less than 4 × as long as broad (Figs 32 A, B); metapectal-propodeal disc usually with transverse posterior keel (Fig. 34 C); paramere without large inner branch wrapping penis (Fig. 35 F); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF82A10FF3ED50DFB76FD48.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF82A10FF3ED50DFB76FD48.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013) (Figs 32 C-E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF82A10FF3ED50DFB76FD48.taxon	discussion	World species. 464 species are known, 106 in the Afrotropical region. Remarks. Larvae of Anteon were described by Ponomarenko (1975 b) and more recently by Guglielmino et al. (2017 a). Leafhoppers parasitized by Anteon species usually show dryinid sacs situated in a latero-ventral position between the prothorax and the mesothorax (Fig. 32 C), rarely positioned on the ventral side.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A16FF3ED5EBFE14F956.taxon	description	(Fig. 33 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A16FF3ED5EBFE14F956.taxon	description	Description. ♀. Fully winged; body length 3.1 mm. Head black, except mandible and anterior half of frons tes- taceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 11: 6: 9: 8: 7: 7: 7: 7: 7.5: 10. Head dull, granulate, sculptured by many irregular longitudinal striae; frontal line absent; occipital carina complete; POL = 7; OL = 5; OOL = 5.5; OPL = 6.5; TL = 5. Pronotum shiny, with anterior surface rugose and transversely striate; posterior surface shorter than mesoscutum (8: 17), with anterior half dull and granulate and posterior half shiny and unsculptured; pronotal tubercle reaching tegula. Mesoscutum shiny, unsculptured or very slightly granulate. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 13). Protarsomeres in following proportions: 9: 3: 4: 8: 19. Enlarged claw (Fig. 33 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 33 C) with two rows of approximately 50 lamellae; apex with about 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A16FF3ED5EBFE14F956.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Perinet, 27. IV – 3. V. 1983, J. S. Noyes & M. C. Day leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A16FF3ED5EBFE14F956.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A16FF3ED5EBFE14F956.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	description	(Figs 33 D – F, 34 A – D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	description	Description. ♀ (Figs 34 A, B). Fully winged; body length 1.6 – 2.0 mm. Head black, except mandible testaceous; antenna testaceous (in ♀♀ from Oman, South of Dahariz, antenna darkened); mesosoma black; metasoma brown; legs testaceous; in ♀♀ from Oman, South of Dahariz, legs testaceous, except metacoxa and clubs of femora brown. Antenna clavate; antennomeres in following proportions: 9: 4: 3: 2.5: 2.5: 3: 3: 3: 3: 5. Head dull, completely granulate; in ♀♀ from Dubai, Zabeel, and Afghanistan, near Arghandab River, frons with unsculptured area in front of anterior ocellus; frontal line complete (incomplete in ♀ from Afghanistan, near Arghandab River, and in some ♀♀ from Japan; absent in other ♀♀ from Japan); occipital carina complete; POL = 4; OL = 2.5; OOL = 4; OPL = 3; TL = 5. Pronotum dull, almost completely granulate, only with posterior margin unsculptured; posterior surface shorter than mesoscutum (5: 9); pronotal tubercle reaching tegula. Mesoscutum shiny, slightly granulate; in ♀♀ from Oman, South of Dahariz, mesoscutum dull and strongly granulate. Notauli incomplete, reaching approximately 0.3 – 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticu- late rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 5). Protarsomeres in following proportions: 5: 2: 2.5: 4: 9. Enlarged claw (Figs 33 D, E) with proximal promi- nence bearing one long bristle. Protarsomere 5 (Figs 33 D, E) with two rows of approximately 27 bristles, occasion- ally with two rows of approximately 12 – 13 lamellae in addition to some bristles (occasionally bristles very broad and similar to lamellae); apex with about 4 – 6 lamellae. Tibial spurs 1 / 1 / 2 ♂ (Figs 34 C, D). Fully winged; body length 1.4 – 1.7 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma black (in ♂ from Oman, 23 ° 36 ’ N 58 ° 26 ’ E, mesosoma completely brown-reddish); meta- soma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 6: 4: 4.5: 4.5: 5: 5: 5: 5: 5: 6. Head dull, completely granulate; in ♂ from Oman, 23 ° 36 ’ N 58 ° 26 ’ E, head slightly granulate; frontal line absent (occasionally incomplete); occipital carina complete; POL = 4; OL = 2; OOL = 3.5; OPL = 2; TL = 3. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured. Notauli incomplete, reaching about 0.4 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel (in specimen from Oman, 23 ° 36 ’ N 58 ° 26 ’ E, transverse posterior keel absent); propodeal declivity without longitudinal keels, completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 4). Paramere (Fig. 33 F) without distal inner process, with inner medial process surrounded by membranous band. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	materials_examined	Material examined. Types: ♀ holotype: LEBANON: Fanar, reared from unidentified leafhopper, parasit- ized host collected 23. VII. 1974, dryinid adult emergence 9. VIII. 1974, H. Abdul-Nour leg. (AMNH). Paratypes: LEBANON: same locality label as holotype, parasitized host collected 19. VIII. 1974, 1 ♀ (AMNH); same locality label as holotype, 24. VII. 1975, collected by sweeping, 3 ♀♀ (AMNH); same locality label as holotype, parasit- ized host collected 22. VII. 1974, dryinid adult emergence 6. VIII. 1974, 2 ♀♀ (AMNH); same locality label as holo- type, 20. VIII. 1974, collected by sweeping, 1 ♂ (AMNH); same locality label as holotype, parasitized host collected 18. VII. 1974, dryinid adult emergence 2. VIII. 1974, 1 ♀ (AMNH); same locality label as holotype, 23. VII. 1974, collected by sweeping, 1 ♀ (AMNH); locality label as holotype, 27. VII. 1974, collected by sweeping, 1 ♀ (AMNH); same locality label as holotype, parasitized host collected 23. VII. 1974, dryinid adult emergence 9. VIII. 1974, 1 ♀ (AMNH); locality label as holotype, 19. VIII. 1974, collected by sweeping, 1 ♂ (AMNH); same locality label as holo- type, parasitized host collected 23. VII. 1974, 1 ♀ (AMNH); same locality label as holotype, 1 ♀ (MNHN). TURKEY: Antalya, 9. VII. 1987, M. Olmi leg., 2 ♀♀ (AMNH). Other material: OMAN: Al Khuwayr capital area, 23 ° 36 ’ N 58 ° 26 ’ E (PMA); Salalah, South Dahariz, 22. XI. 1986, J. T. Huber leg., 1 ♀ (AMNH); Al Khaburah, 10. IV. 1985 (ZIL, AMNH); Muscat, 5. IV. 1985 (ZIL). SOUTH AFRICA: Gauteng, Roodeplaat Dam, 21. I. 2006, R. Wharton leg., 1 ♂ (TAMU). UNITED ARAB EMIRATES: Wadi Wurayah farm, 25 ° 23 ’ N 56 ° 19 ’ E, 1. IV – 14. VI. 2009, light trap, 3 ♀♀, 3 ♂♂ (UCDC); Wadi Safad, 25 ° 13 ’ N 56 ° 19 ’ E, 22. IV – 21. VI. 2006, light trap, 1 ♂ (UCDC); same locality label, 4. V – 15. VII. 2006, 2 ♂♂ (UCDC); Dubai, Zabeel, light trap, 27 – 29. I. 1985, 1 ♂ (UCDC). YEMEN: Lahj, IX. 1998, MT, No 3304, A. van Harten & A. Sallam leg., 2 ♂♂, 9 ♀♀ (AMNH); same locality label, VI – VII. 1998, 1 ♀ (AMNH); same locality label, X. 1998, IV. 2000, 15 – 30. VI. 2000, VIII. 2000, XI. 2000, I. 2001, III – V. 2002, MT, A. van Harten & A. Sallam leg., 5 ♀♀, 8 ♂♂ (MOLC); Al Lahima, 16. X – 31. XII. 2000, MT, A. van Harten & A. M. Hager leg., 2 ♀♀, 3 ♂♂ (MOLC); Al-Kadan, IV. 2002, light trap, A. van Harten & T. Abdul-Haq leg., 1 ♂ (MOLC). Palaearctic. Known from Afghanistan, China, Hungary, Italy, Japan, Lebanon, Turkey (Mita et al. 2009; Olmi 1999, 2014; Olmi & van Harten 2000, 2006).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Japan: Psammotettix alienus (Dahlbom) (= Psammotettix striatus (L. )); in Lebanon: Aconurella prolixa (Lethierry); in Turkey: Exitianus capicola (Stål).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	distribution	Distribution. Oman, South Africa, United Arab Emirates, Yemen, in addition to many Palaearctic countries (from Italy to Japan).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFF12A29FF3ED16DFC50F9D4.taxon	discussion	Remarks. The ♂ from South Africa (Roodeplaat Dam) is similar to the ♂♂ known from the Arabian Peninsula, but its antenna is less slender: in ♂♂ from the Arabian Peninsula, antennomere 8 is ~ 3 × as long as broad (5: 1.5); in the ♂ from South Africa, antennomere 8 is ~ 2 × as long as broad (4: 2).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCE2A2AFF3ED0EBFE1FFC20.taxon	description	(Fig. 35 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCE2A2AFF3ED0EBFE1FFC20.taxon	description	Description. ♂. Fully winged; body length 2.4 – 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa partly black, stalk of metafemur brown, distal half of metatibia brown. Antenna filiform; antennomeres in following proportions: 9: 5: 6: 6: 6: 7: 6: 6: 6: 10. Head dull, granulate and reticulate rugose (in specimen from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, area in front of anterior oclelus punctate, unsculptured among punctures, not rugose); frontal line incomplete, shortly present in front of anterior ocellus (in specimen from South Africa, Butterfly Sanctuary, frontal line complete); occipital carina complete; POL = 6; OL = 4; OOL = 7; OPL = 4; TL = 3. Mesoscutum shiny, granulate, with posterior region punctate and unsculptured among punctures (in specimne from South Africa, Pietermaritzburg, Hilton, anterior third of mesoscutum slightly rugose and lateral regions of propodeal declivity slightly granulate; in specimen from South Africa, Butterfly Sanctuary, mesoscutum punctate, unsculptured among punctures, except narrow rugose stripe near anterior margin). Notauli incomplete, reaching about 0.3 – 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, with two longitudinal keels and median area sculptured by areolae less wide than those of the lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part. Paramere (Fig. 35 E) without distal inner process and without proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCE2A2AFF3ED0EBFE1FFC20.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg (AEIC). Paratypes: SOUTH AFRICA: KwaZulu-Natal, Hluhluwe Game Reserve, 16. XI. 1970, H. Townes leg., 2 ♂♂ (AEIC, AMNH). Other material: SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 6 – 14. III. 2005, YPT, V. Kolyada leg., 1 ♂ (NMSA); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ‘ S 30 ° 20.4 ‘ E, MT near stream, 1. XI – 2. XII. 2004, M. Mostovski leg., 1 ♂ (NMSA); same locality label, 27. IV – 27. VII. 2005, 1 ♂ (NMSA); KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ‘ S 32 ° 44.792 ‘ E, 50 m, 18 – 20. X. 2010, MT, Maputaland Coastal Belt, SA 10 - MAB-M 01, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCE2A2AFF3ED0EBFE1FFC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCE2A2AFF3ED0EBFE1FFC20.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCD2A2DFF3ED5B7FD73F83C.taxon	description	(Figs 35 A, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCD2A2DFF3ED5B7FD73F83C.taxon	description	Description. ♀. Fully winged; body length 1.9 – 3.7 mm. Testaceous, except petiole black and antennomeres 4 – 10 or 8 – 10 brown; occasionally pronotum with two lateral brown spots; occasionally metapectal-propodeal disc brown, or black, or darkened. Occasionally head black, except anterior region of frons, clypeus and mandible testaceous; mesosoma and metasoma totally black. Specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, testaceous-brown, except metapectal-propodeal disc black. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 63, head black, except mandible, clypeus and anterior region of frons testaceous; antenna yellow, except antennomeres 9 – 10 brown; mesosoma black, except lateral regions partly ferruginous; metasoma brown; legs yellow. In very large specimen (3.7 mm) from Madagascar, 21 ° 17.24 ’ S 47 ° 26.00 ’ E, BLF 8397, head black, except mandible, clypeus, malar space, area between antennal toruli and part of ventral side yellow-testaceous; antenna testaceous, except antennomeres 5 – 10 darkened; mesosoma black; forewing completely darkened; metasoma brown; legs yellowtestaceous. Occasionally head and mesosoma reddish, except metapectal-propodeal disc black. Antenna clavate; antennomeres in following proportions: 8: 4.5: 5: 5: 5: 5: 5: 5: 5.5: 9. Head shiny, more or less granulate, sculptured by numerous irregular slight longitudinal striae; frontal line absent; occipital carina complete; holotype with POL = 5; OL = 3; OOL = 4; OPL = 2.5; TL = 3. In ♀ from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, POL = 4; OL = 2; OOL = 3; OPL = 2.5; TL = 2.5; greatest breadth of lateral ocelli about as long as OOL. In ♀ from Madagascar, 18 ° 16 ’ 00 ’’ S 45 ° 24 ’ 24 ’’ E, head reticulate rugose, with many slight longitudinal striae in front of anterior ocellus. In ♀ from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, head rugose among ocelli and sculptured by irregular striae on sides of ocelli and on anterior half of frons, unsculptured in front of anterior ocellus. In specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, head completely unsculptured. Pronotum shiny, transversely striate; posterior surface flat, shorter than mesoscutum (6: 13 or 8: 10); pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 63, mesoscutum shiny, slightly granulate. In specimen from South Africa, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, head completely granulate, not rugose. Notauli incomplete, reaching approximately 0.3 – 0.5 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands (large black specimens from Madagascar, 21 ° 17.24 ’ S 47 ° 26.00 ’ E, with forewing completely darkened); distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 10). Protarsomeres in following proportions: 5: 2: 3: 6: 14. In specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, protarsomeres in following proportions: 5: 2: 3: 4.5: 14. Enlarged claw (Fig. 35 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 35 A) with two rows of approximately 50 lamellae; apex with about 3 – 4 lamellae; usually apex short. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.9 – 2.7 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except club of metafemur brown. Antenna filiform; antennomeres in following proportions: 7: 5: 6: 6.5: 6: 6.5: 6.5: 6: 5.5: 9. Head dull, granulate and sculptured by numerous irregular striae (occasionally head granulate and reticulate rugose; rarely (in specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 36, head almost completely unsculptured )); frontal line complete; frons without lateral keels along orbits; occipital carina complete; POL = 4; OL = 3; OOL = 4; OPL = 2; TL = 2.5. Mesoscutum, mesoscutellum and metanotum shiny, smooth, finely punctate, unsculptured or slightly granulate among punctures. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels, completely reticulate rugose. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, propodeal declivity with one complete longitudinal keel. Forewing hyaline, without dark transverse bands (occasionally forewing darkened, mainly in large specimens); distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Paramere (Fig. 35 F) without distal inner process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCD2A2DFF3ED5B7FD73F83C.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toamasina, Perinet, 27. IV – 3. V. 1983, B. M. 1983 – 201, J. S. Noyes & M. C. Day leg. (NHMUK). Paratype: same locality label as holotype, 1 ♀ (AMNH). Other material: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT in montane rainforest, Fisher, Griswold et al. leg., BLF 2375, 4 ♂♂ (3 in CAS, 1 in MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 01, 3 m # m # (CAS); same locality label, 4 – 10. III. 2001, MA- 01 - 01 B- 06, 2 ♀♀, 8 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 29. I. 2001, netted by hand, M. E. Irwin leg., MA- 01 - 01 A- 05, 5 ♂♂ (CAS); same locality label, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg., 2 ♀♀, 2 ♂♂ (CAS); same locality label, 11 – 12. II. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 06, 1 ♀ (CAS); same locality label, 12. II – 4. III. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 07, 1 ♂ (CAS); same lo- cality label, 4 – 19. III. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 08, 1 ♂ (CAS); same locality label, 26 – 29. I. 2001, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 02, 1 ♀, 2 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 21 – 26. IV. 2001, MT, MA- 01 - 01 D- 08, R. Harin’Hala leg., 1 ♀, 1 ♂ (CAS); same locality label, 4 – 19. III. 2001, MA- 01 - 01 D- 05, 1 ♀ (CAS); same locality label, 6 – 21. IV. 2001, MA- 01 - 01 D- 07, 4 ♀♀ (CAS); same locality label, 29. I – 11. II. 2001, R. Harin’Hala leg., MA- 01 - 01 D- 03, 1 ♂ (CAS); same locality label, 11. II – 4. III. 2001, R. Harin’Hala leg., MA- 01 - 01 D- 04, 3 ♂♂ (CAS); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 11 – 18. II. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 11, 1 ♂ (CAS); same locality label, 18 – 25. III. 2005, MA- 31 - 16, 1 ♂ (CAS); same locality label, MA- 31 - 22, 1 ♂ (CAS); Antsiranana, Marojejy National Park, Manantenina river, 27.6 km 35 ° NE Andapa, 9.6 km 327 ° NNW Manantenina, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 11 – 14. XII. 2005, MT, rainforest, B. L. Fisher et al. leg., BLF 13383, 3 ♂♂ (CAS); Antsiranana, Marojejy National Park, Manantenina river, 28.0 km 38 ° NE Andapa, 8.2 km 333 ° NNW Manantenina, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, MT, rainforest, B. L. Fisher et al. leg., BLF 8723, 1 ♂ (CAS); Antsiranana, Binara Forest, 7.5 km 230 ° SW Daraina, 13 ° 15.18 ’ S 49 ° 37.00 ’ E, 375 m, 1. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9557, 1 ♀ (CAS); same locality label, YPT, BLF 9559, 1 ♀ (CAS); Antsiranana, Ambondrobe, 41.1 km 175 ° Vohemar, 13 ° 42.55 ’ S 50 ° 06.06 ’ E, 29. XI. 2004, 10 m, YPT, littoral rainforest, B. L. Fisher leg., BLF 11098, 2 ♂♂ (CAS); Antsiranana, Montagne des Français, 12 ° 18.8 ’ S 49 ° 38.51 ’ E, 150 m, along forested limestone ridge, 16. II – 8. III. 2001, MT, R. Harin’Hala leg., MA- 01 - 06 - 06, 1 ♀, 2 ♂♂ (CAS); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 16 – 31. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 10, R. Harin’Hala leg., 1 ♂, 1 ♀ (CAS); same locality label, 15. II – 6. III. 2001, MA- 01 - 04 B- 10, 6 ♂♂, 1 ♀ (CAS); Antsiranana, SAVA Region, Vohemar Dis- trict, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 19 – 26. XII. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 12, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, in dry wash east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 7 – 22. IX. 2002, MT, in open area, MA- 02 - 11 B- 39, R. Harin’Hala leg., 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Talatakely, 21 ° 15 ’ S 47 ° 25 ’ E, 14. IV. 1998, 2 ♂♂ (CAS); Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 5 ♂♂ (CAS); same locality label, 26. II – 10. III. 2003, 1 ♀ (CAS), 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 19 – 26. II. 2002, MA- 02 - 09 C- 17, 1 ♀ (CAS), 6 ♂♂ (4 in CAS, 2 in MOLC); same locality label, 5 – 13. V. 2002, MA- 02 - 09 C- 28, 1 ♀ (CAS); same locality label, 14 – 24. VII. 2002, 1 ♂ (CAS); same locality label, 6 – 18. VI. 2003, MA- 02 - 09 C- 63, 2 ♀♀ (CAS, MOLC), 2 ♂♂ (CAS, MOLC); same locality label, 10 – 21. III. 2003, MA- 02 - 09 C- 56, 1 ♀ (MOLC); same locality label, 26. II – 4. III. 2002, MA- 02 - 09 C- 18, 2 ♂♂ (CAS); same locality label, 23 – 28. IV. 2002, MA- 02 - 09 C- 26, 5 ♂♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 C- 06, 2 ♂♂ (CAS); same locality label, 4 – 14. VII. 2002, MA- 02 - 09 C- 34, 3 ♂♂ (CAS); same locality label, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 1 ♂ (MOLC); same locality label, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 1 ♀ (CAS); same locality label, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 6 ♂♂ (CAS); same lo- cality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 3 ♂♂ (CAS); same locality label, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♂ (CAS); same locality label, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 2 ♀♀ (CAS, MOLC), 2 ♂♂ (CAS); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 1 ♂ (CAS); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 1 ♀ (MOLC); same locality label, 26 – 31. III. 2002, MA- 02 - 09 C- 22, 3 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 2 ♂♂ (CAS); same locality label, 21. III – 12. IV. 2003, MA- 02 - 09 C- 57, 1 ♀ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♂ (CAS); same lo- cality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 5 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 1 ♂ (CAS); same locality label, 14 – 21. I. 2002, 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 26 – 31. III. 2002, MA- 02 - 09 A- 22, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 4 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 A- 16, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 12 – 19. III. 2002, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 20, 1 ♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 A- 04, 4 ♂♂ (CAS); same locality label, 19 – 26. II. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 1 ♀ (MOLC), 3 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 2 ♂♂ (CAS, MOLC); same local- ity label, 7 – 18. VI. 2003, 1 ♀ (CAS), 2 ♂♂ (CAS, MOLC); same locality label, 9 – 16. IV. 2002, 1 ♂ (CAS); same local- ity label, 15 – 25. VII. 2002, MA- 02 - 09 B- 35, 1 ♀ (CAS); same locality label, 14 – 24. VI. 2002, MA- 02 - 09 B- 32, 1 ♂ (CAS); same locality label, 15 – 27. IV. 2003, MA- 02 - 09 B- 58, 1 ♀ (CAS); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 3 ♂♂ (CAS); same locality label, 18 – 29. VI. 2003, MA- 02 - 09 B- 64, 1 ♂ (CAS); same locality label, 20. III – 3. IV. 2003, MA- 02 - 09 B- 56, 1 ♂ (CAS); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 1 ♂ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 5 ♂♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 1 ♂ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 4 ♀♀ (2 in CAS, 2 in MOLC); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 1 ♂ (CAS); same locality label, 7 – 17. VII. 2003, MA- 02 - 09 B- 66, 5 ♂♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 2 ♀♀ (CAS), 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ranomafana National Park, Vatoha- ranana River, 4.1 Km 231 ° SW Ranomafana, 21 ° 17.24 ’ S 47 ° 26.00 ’ E, 1100 m, YPT, montane rainforest, 27 – 31. III. 2003, Fisher, Griswold et al. leg., BLF 8399, 3 ♀♀ (2 in CAS, 1 in MOLC), 1 ♂ (CAS); same locality la- bel, 27 – 31. III. 2003, MT, BLF 8397, 2 ♀♀ (MOLC); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. I – 4. II. 2002, MT near river, R. Harin’Hala leg., MA- 02 - 09 A- 14, MA- 02 - 09 D- 14, 2 ♂♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 D- 11, 2 ♂♂ (CAS); same locality label, 21 – 24. XII. 2001, MA- 02 - 09 D- 08, 1 ♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 D- 10, 1 ♂ (CAS); Fianarantsoa, Befotaka-Mi- dongy National Park, Papango 27.7 km S Midongy-South Mt. Papango, 23 ° 50.07 ’ S 46 ° 57.49 ’ E, 940 m, 13 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14774, 2 ♂♂ (CAS, MOLC); same locality label, 13 – 15. XI. 2006, BLF 14775, 1 ♀ (CAS); Fianarantsoa, Befotaka-Midongy National Park, Papango 28.5 km S Midongy-South Mt. Papango, 23 ° 50.27 ’ S 46 ° 57.27 ’ E, 1250 m, 17 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14944, 1 ♀ (CAS); Fi- anarantsoa, Atsirakamblaty Forest, 7.6 km 285 ° WNW Itremo, 20 ° 35.36 ’ S 46 ° 33.48 ’ E, 1550 m, 22 – 26. I. 2003, YPT, montane rainforest, Fisher, Griswold et al. leg., BLF 7154, 1 ♀ (CAS); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, YPT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7385, 4 ♀♀ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT in tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 2 ♂♂ (CAS); Mahajanga, Namoroka National Park, 17.8 km 329 ° WNW Vilanandro, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, 100 m, 8 – 12. XI. 2002, YPT, in tropical dry forest, Fisher, Griswold et al. leg., BLF 6511, 1 ♀ (CAS); Mahajanga, Namoroka National Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, 100 m, 12 – 16. XI. 2002, MT in tropical dry forest, Fisher, Griswold et al. leg., BLF 6581, 3 ♀♀, 3 ♂♂ (CAS); same locality label, YPT in tropical dry forest, BLF 6536, 64 ♀♀ (54 in CAS, 10 in MOLC), 16 ♂♂ (14 in CAS, 2 in MOLC); Mahajanga, Namoroka National Park, 16.40818 S 45.31437 E, 25 – 27. X. 2016, YPT, C. Villemant leg., 1 ♀ (MNHN); same locality label, 23 – 25. X. 2016, YPT, 1 ♀ (MNHN); same locality label, 27 – 30. X. 2016, YPT, 1 ♀ (MNHN); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 27 – 30. X. 2016, YPT, C. Villemant leg., 1 ♀ (MNHN); same locality label, 23 – 25. X. 2016, 2 ♀♀ (MNHN); Mahajanga, Namoroka National Park, 16.40558 S 45.31165 E, 25 – 27. X. 2016, YPT, C. Villemant leg., 3 ♀♀ (MNHN); same locality label, 23 – 25. X. 2016, YPT, 7 ♀♀ (MNHN); same locality label, 27 – 30. X. 2016, YPT, 2 ♀♀ (MNHN); Mahajanga, Namoroka National Park, 23 – 25. X. 2016, YPT, C. Villemant leg., 11 ♀, 1 ♂ (MNHN); same locality label, 25 – 27. X. 2016, YPT, 2 ♀♀ (MNHN); same locality label, 21 – 22. X. 2016, YPT, 1 ♀ (MNHN); same locality label, 27 – 30. X. 2016, YPT, 6 ♀♀ (MNHN); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, YPT, tropical dry forest on tsingy, B. L. Fisher et al. leg., BLF 22788, 2 ♀♀ (CAS); Toamasina, 9.2 km WSW Befingotra, Anjanaharibe-sud Reserve, 14 ° 45 ’ S 49 ° 28 ’ E, 1200 m, 9. XI. 1994, montane rainforest, sifted litter (leaf mold, rotten wood), B. L. Fisher leg., 1 ♀ (UCDC); Toamasina, Manakambahiny-E, 17 ° 45 ’’ S 48 ° 43 ’ E, 18 – 25. I. 1991 (FSAE); Toamasina, Parcelle E 3 Tampolo, 17 ° 17.00 ’ S 49 ° 26.00 ’ E, 10 m, 14. IV. 2004, YPT, littoral forest, Malagasy ant team leg., BLF 10730, 1 ♀ (CAS); Toamasina, Parcelle E 3 Tampolo, 17 ° 17.00 ’ S 49 ° 26.00 ’ E, 10 m, 18 – 21. IV. 2004, YPT, littoral forest, Malagasy ant team leg., BLF 108340, 1 ♀ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 8 – 16. X. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 15, 2 ♀♀ (CAS); same locality label, 16 – 23. XI. 2001, MA- 01 - 08 B- 20, 1 ♂ (CAS); same locality label, 1 – 5. IX. 2001, MA- 01 - 08 B- 11, 2 ♀♀, 8 ♂♂ (CAS); same locality label, 1 – 7. XI. 2001, 5 ♂♂ (4 in CAS, 1 in MOLC); same locality label, 7 – 16. XI. 2001, 1 ♀ (MOLC); Toamasina, Mananara-Nord National Park, 7.1 km 261 ° Antanambe, 16 ° 27.18 ’ S 49 ° 47.15 ’ E, 225 m, 14. XI. 2005, MT, rain forest, Brian L. Fisher et al. leg., BLF 12557, 2 ♂♂ (CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km 332 ° Toamasina, 17 ° 55.26 ’ S 49 ° 11.59 ’ E, 390 m, 28. XI. 2005, YPT, rainforest, Brian L. Fisher et al. leg., BLF 13124, 1 ♂ (CAS), 2 ♀♀ (1 in CAS, 1 in MOLC); Toamasina, Zahamena National Park, Onibe River, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, 780 m, 21 – 23. II. 2009, rainforest, YPT, BLF 22170, B. L. Fisher et al. leg., 1 ♀ (CAS); Toamasina, Zahamena National Park, Tetezambatana Forest, near junction of Nosivola and Manakam- bahiny Rivers, 17 ° 44 ’ 35 ’’ S 48 ° 43 ’ 46 ’’ E, 860 m, 18 – 19. II. 2009, rainforest, MT, BLF 21975, B. L. Fisher et al. leg., 1 ♀ (CAS); Toamasina, Ankerana, 18 ° 24.23 ’ S 48 ° 48.09 ’ E, 1108 m, 21 – 26. I. 2012, MT, rainforest, B. L. Fisher et al. leg., BLF 27929, 1 ♂ (CAS); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Marofandilla, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, 100 m, 28. XI – 3. XII. 2001, MT in tropical dry forest, Fisher, Griswold et al. leg., BLF 4601, 4 ♂♂ (CAS); Toliara, Am- bohijanahary Special Reserve, Ankazotslhitafototra Forest, 35.2 km 312 ° NW Ambaravaranata, 18 ° 16 ’ 00 ’’ S 45 ° 24 ’ 24 ’’ E, 1050 m, Montane rainforest, MT, Fisher, Griswold et al. leg., BLF 7023, 1 ♀, 3 ♂♂ (CAS); Toliara, Andohahela National Park, Col de Tanatana, 33.3 km NW Tolagnaro, 24 ° 45.31 ’ S 46 ° 51.13 ’ E, 275 m, 22 – 24. XI. 2006, MT, B. L. Fisher et al. leg., BLF 15102, 2 ♂♂ (CAS); Toliara, Kalambatritra Special Reserve, Ampanihy, 23 ° 27 ’ 49 ’’ S 46 ° 27 ’ 47 ’’ E, 1270 m, 9 – 10. II. 2009, YPT, montane rainforest, B. L. Fisher et al. leg., BLF 21567, 1 ♀ (CAS); Toliara, Ambohijanahary Special Reserve, Ankazotsyhitafototra Forest, 35.2 km 312 ° NW Ambaravaranala, 18 ° 16.00 ’ S 45 ° 24.24 ’ E, 1050 m, 13 – 17. I. 2003, YPT, montane rainforest, BLF 7019, Fisher, Griswold et al. leg., 1 ♂, 6 ♀♀ (CAS); Toliara, Analavelona Classified Forest, 29.2 km 343 ° NNW Mahaboboka, 22 ° 40.30 ’ S 44 ° 11.24 ’ E, 1100 m, 18 – 22. II. 2003, YPT, montane rainforest, BLF 7822, Fisher, Griswold et al. leg., 2 ♀♀ (CAS). SOUTH AFRICA: Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - Y 10, 1 ♀ (SAMC); same locality label, DRA 09 - FYN 1 - Y 09, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCD2A2DFF3ED5B7FD73F83C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFCD2A2DFF3ED5B7FD73F83C.taxon	distribution	Distribution. Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC92A2FFF3ED0B6FA6EFA44.taxon	description	(Figs 35 B, G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC92A2FFF3ED0B6FA6EFA44.taxon	description	Description. ♀. Fully winged; body length 2.0 – 2.7 mm. Head black, except mandible yellow-testaceous; antenna yellow-testaceous; mesosoma and metasoma black; tegula and legs yellow-testaceous. Antenna clavate; antenno- meres in following proportions: 8: 4: 2: 2: 2.5: 2: 2: 3: 2.5: 4.5. Head dull, smooth, completely granulate; frontal line absent; frons without lateral keels; occipital carina complete; POL = 4; OL = 3; OOL = 4; OPL = 2.5; TL = 4. Pronotum dull, granulate; posterior surface longer than anterior surface and shorter than mesoscutum (5: 7); pronotal tubercle reaching tegula. Mesoscutum granulate. Notauli incomplete, reaching about 0.5 × length of mesoscutum (in specimen from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, right notaulus reaching about 0.5 × length of mesoscutum, left notaulus absent). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity granulate, without longitudinal keels, not reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 4). Protarsomeres in following proportions: 4: 1.5: 2: 3: 9. Enlarged claw (Fig. 35 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 35 B) with two rows of 13 – 16 lamellae; apex with 3 – 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.4 – 1.9 mm. Head black, except mandible testaceous; antenna testaceous; me- sosoma and metasoma black; tegula and legs testaceous. Antenna filiform; antennomeres in following proportions: 7: 5: 5: 5: 6: 5: 5: 5: 5: 7. Head dull, granulate; frontal line incomplete and slight; frons without lateral keels; occipital carina complete; POL = 5; OL = 3; OOL = 4.5; OPL = 3; TL = 4. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures, with some very slightly granulate areas near margins. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels, completely finely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 5). Paramere (Fig. 35 G) without distal inner pointed process and without proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC92A2FFF3ED0B6FA6EFA44.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: Okahandja, 19 – 29. XII. 1927, R. E. Turner leg. (NHMUK). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: Orientale Prov. (= Haut-Zaire), Garamba National Park, 1 ♀ (AMNH); same locality label, 1 ♂ (MRAC). Other material: KENYA: Nyanza Prov., Ungoye, ICIPE Field Station, 0.61517 ° S 34.09200 ° E, 1147 m, 12 – 26. II. 2005, MT, inside seasonally swampy forest, R. Copeland leg., 1 ♀ (NMK). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 6 – 21. IV. 2001, MT, MA- 01 - 01 D- 07, R. Harin’Hala leg., 1 ♀ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 7 – 17. III. 2003, MT, gallery forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 22 - 18, 2 ♂♂ (CAS). MOZAMBIQUE: Maputo Prov., Maputo (AMNH) Maputo Prov. 3 Km N Um- beluzi (AMNH); Maputo Prov. 5 km E of Manhiça (AMNH); Maputo Prov., Manhiça District, Manhiça, Vasco’s Farm, 9. IX – 24. X. 1995, MT, M. Olmi leg., 1 ♀ (MOLC); same locality label, 1 – 8. IX. 1996, 1 ♀ (MOLC). NAMIB- IA: Okahandja, 19 – 29. XII. 1927, R. E. Turner leg., 1 ♂ (NHMUK). SOUTH AFRICA: KwaZulu-Natal, Eshowe, 1 – 3. V. 1926, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, West Coast Fossil Park, 5 km 290 ° E Langebaanweg, 32 ° 57.732 ’ S 18 ° 05.985 ’ E, 16 – 23. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC92A2FFF3ED0B6FA6EFA44.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Nephotettix afer Ghauri; Nephotettix modulatus Melichar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC92A2FFF3ED0B6FA6EFA44.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, Madagascar, Mozambique, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC82A20FF3ED05BFC0BFAFC.taxon	description	(Figs 35 C, 38 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC82A20FF3ED05BFC0BFAFC.taxon	description	Description. ♀. Fully winged; body length 2.6 – 3.2 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black, except pronotum reddish or brown; metasoma black; legs testaceous, except metacoxa occasionally partly brown. Antenna filiform; antennomeres in following proportions: 10: 6: 9: 10: 8: 8: 8: 8: 8: 11. Head shiny, smooth, slightly punctate, unsculptured among punctures, more strongly punctate behind ocelli; frontal line absent; frons with no lateral keels; occipital carina complete; POL = 5; OL = 5; OOL = 6; OPL = 5; TL = 7. Pronotum shiny, smooth, almost flat, finely punctate, unsculptured among punctures, with pos- terior surface longer than mesoscutum (14: 11); pronotal tubercle reaching tegula. Mesoscutum shiny, smooth, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Protarsomeres in following proportions: 9: 3: 3.5: 12: 21; protarsomere 4 much longer than basal part of 5 (12: 8). Enlarged claw (Fig. 35 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 35 C) with basal part shorter than distal part (8: 13), with two rows of 28 lamellae; apex with 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.7 – 2.7 mm. Head black, except mandible testaceous; antenna brown; occasion- ally antennomere 1 testaceous; mesosoma and metasoma black; tegula testaceous; legs testaceous, except metacoxa proximally darkened. Antenna filiform; antennomeres in following proportions: 12: 6: 10: 10: 10: 10: 8: 9: 10: 10. Head shiny, strongly punctate, unsculptured among punctures; frontal line absent; frons without lateral keels along orbits directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 3; TL = 5; ocellar triangle with slight keels joining ocelli. Mesoscutum, mesoscutellum and metanotum shiny, smooth, strongly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two longitudinal keels and median area approximately as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 12). Paramere (Fig. 38 A) with distal inner pointed process and proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC82A20FF3ED05BFC0BFAFC.taxon	materials_examined	Material examined. Types: ♀ holotype: CAMEROON: Mt. Cameroon, Mann’s Quelle, 7400 ft., 1. II. 1932, M. Steele leg. (NHMUK). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Magera Lake, 2000 m, 6. III. 1934, Mission G. F. de Witte, ♂ allotype of Chelogynus kivuanus (MRAC); North Kivu, Virunga National Park (= Albert National Park), Mt. Ruwenzori, Kalonge, 1 ♀ (MRAC), 1 ♂ (AMNH); North Kivu, Virunga National Park (= Albert National Park), Mikeno Volcano, Rweru, 2400 m, bambous, 26 – 27. VII. 1934, Mission G. F. de Witte, 2 ♂♂ (MRAC); North Kivu, Virunga National Park (= Albert National Park), Tshiaberimu, Talia N River, Mbulikerere River, 1 ♀ (AMNH). SOUTH AFRICA: Eastern Cape, Katberg, 15 – 30. I. 1933, R. E. Turner leg., 1 ♂ (NHMUK). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park, Between Ngesho and Bishakishaki, Kashwa, 2000 m, 7 – 23. I. 1935, Mission G. F. de Witte, 1 ♂ (Benoit 1951 c: misid. Chelogynus kivuanus Benoit) (MRAC); North Kivu, Virunga National Park, Rutshuru, 1285 m, 13. VII. 1935, Mission G. F. de Witte, 1 ♂ (Benoit 1951 c: misid. Chelogynus kivuanus Benoit) (MRAC); Orientale Prov. (= Haut-Zaire), Garamba National Park (MRAC). KENYA: Western Prov., 27 mi. NE Kisumu, Kaimosi Mission, 1650 m, 29. XI. 1957, E. S. Ross & R. E. Leech leg., 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC82A20FF3ED05BFC0BFAFC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC82A20FF3ED05BFC0BFAFC.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC72A22FF3ED383FE59FF50.taxon	description	(Figs 32 F, 35 D, 36 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC72A22FF3ED383FE59FF50.taxon	description	Description. ♀ (Figs 32 F, 36 A). Fully winged; body length 3.5 mm. Head black, except mandible testaceous; antenna brown, except antennomeres 1 – 3 testaceous; mesosoma black; metasoma brown; tegula testaceous; legs testaceous, except basal region of metacoxa brown. Antennomeres in following proportions: 14: 7: 8: 6: 7: 7: 8: 7: 7: 9. Head dull, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 8 OL = 5; OOL = 6; OPL = 8; TL = 5; greatest breadth of lateral ocelli shorter than TL (3: 5). Pronotum shiny, reticulate rugose, except posterior margin smooth; posterior surface much shorter than mesoscutum (5: 20); pronotal tubercle reaching tegula. Mesoscutum shiny, rugose and irregularly striate, except median region smooth, punctate, unsculptured among punctures, with anterior third reticulate rugose. Notauli incomplete, hardly visible, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectalpropodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 12). Protarsomeres in fol- lowing proportions: 12: 2: 3: 3: 12; protarsomeres 3 and 4 produced into hooks; protarsomere 4 less than 0.5 × as long as basal part of 5 (3: 9). Enlarged claw (Fig. 35 D) slightly longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 35 D) with basal part much longer than apical part (9: 3), with some proximal and medial bristles in addition to one lamella; apex with 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC72A22FF3ED383FE59FF50.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Nyanza Prov., Ruma National Park, near Kamato Gate, 0.64725 ° S 34.33595 ° E, 1264 m, 18. XII. 2005 – 1. I. 2006, MT, in open grass-woodland, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC72A22FF3ED383FE59FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC72A22FF3ED383FE59FF50.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC52A23FF3ED767FA88FDE4.taxon	description	(Figs 37, 38 B, 39 A) Diagnosis. ♂ of Anteon with head granulate and reticulate rugose; propodeal declivity with two longitudinal keels and median area as rugose as lateral areas (Fig. 39 A); paramere with rounded distal inner process provided with small papillae on outer margin (Fig. 38 B). Description. ♂ (Figs 37, 39 A). Fully winged; body length 2.2 – 2.5 mm. Head black, except mandible testa- ceous; antenna brown-testaceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa and stalk of metafemur partly brown. Antenna filiform; antennomeres in following proportions: 10: 5: 5: 5: 5: 5: 5: 5: 5 (last antennomere missing in holotype). Head (Fig. 37 C) dull, granulate and reticulate rugose; frons without lateral keels near orbits directed towards antennal toruli; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 4; OPL = 3; TL = 2; greatest breadth of lateral ocelli shorter than OL (2: 4). Mesoscutum (Figs 37 A, 39 A) granulate, dull. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, with strong transverse posterior keel; propodeal declivity with two longitudinal keels and median area as rugose as lateral areas. Forewing (Fig. 37 D) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Paramere (Fig. 38 B) with distal inner rounded process provided with small papillae on outer margin; proximal membranous process with distal extremity setose. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC52A23FF3ED767FA88FDE4.taxon	materials_examined	Material examined. Type: ♂ holotype (CASTYPE 19439): MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 6 – 21. IV. 2001, MT, MA- 01 - 01 D- 07, R. Harin’Hala leg. (CAS). Paratypes: MADAGASCAR: Antsiranana, same locality label as holotype, 4 – 19. III. 2001, MA- 01 - 01 D- 05, 1 ♂ (MOLC); same locality label, 11. II – 4. III. 2001, MA- 01 - 01 D- 04, 1 ♂ (MOLC); Fianarantsoa, Ranomafana Na- tional Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 12 – 19. II. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 16, 2 ♂♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 1 ♂ (CAS); Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 8 – 15. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 02, 1 ♂ (MOLC); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC52A23FF3ED767FA88FDE4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC52A23FF3ED767FA88FDE4.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC52A23FF3ED767FA88FDE4.taxon	etymology	Etymology. The species is named ambrense after the collection site in Montagne d’Ambre National Park.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC32A25FF3ED3B9FE1FFE28.taxon	description	(Fig. 38 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC32A25FF3ED3B9FE1FFE28.taxon	description	Description. ♀. Fully winged; body length 2.7 mm. Completely testaceous, except petiole black. Antenna clavate; antennomeres in following proportions: 10: 5: 5: 5: 5: 5: 5: 5: 5: 7. Head shiny, setose, completely slightly reticulate rugose; frontal line present, slight; frons without lateral keels along orbits directed towards antennal toruli; occipital carina complete; POL = 5; OL = 4; OOL = 4; OPL = 4; TL = 2. Pronotum shiny, slightly rugose; posterior surface much shorter than mesoscutum (5: 15); pronotal tubercle reaching tegula. Mesoscutum shiny, slightly punctate. Notauli slightly visible, incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron and metapleuron dull, granulate and slightly reticulate rugose. Metapectal-propodeal disc dull, strongly reticulate rugose, with transverse posterior keel; propodeal declivity completely granulate, without longitudinal keels. Forewing with one faint preapical dark transverse band; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 10). Protarsomere 1 much longer than 4 (9: 2.5); protarsomere 4 produced into hook, about 0.5 × as long as basal part of 5. Enlarged claw (Fig. 38 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 38 D) with inner side almost rectilinear and with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC32A25FF3ED3B9FE1FFE28.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 1 – 14. V. 1923, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC32A25FF3ED3B9FE1FFE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC32A25FF3ED3B9FE1FFE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A25FF3ED44FFE5DFAA9.taxon	description	(Figs 38 E, 39 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A25FF3ED44FFE5DFAA9.taxon	description	Description. ♀ (Fig. 39 B). Fully winged; body length 2.4 mm. Head black, except mandible testaceous; antenna black, except scape and pedicel testaceous; mesosoma and metasoma black; legs black, except tarsi brown. Antenna clavate; antennomeres in following proportions: 10: 4: 3.5: 4: 3: 4: 4: 4: 4: 5. Head dull, granulate, with slight short irregular keels; frontal line present; frons with two lateral incomplete keels along orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 4; OOL = 4; OPL = 3; TL = 3.5. Pronotum dull, granulate, with numerous transverse keels; posterior surface very short, shorter than anterior surface and shorter than mesoscutum (5: 14); pronotal tubercle reaching tegula. Mesoscutum shiny, very slightly alutaceous, with numerous longitudinal slight keels or furrows. Notauli absent. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Protarsomeres in following proportions: 6: 1.5: 1.5: 1.5: 7. Enlarged claw (Fig. 38 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 38 E) with basal part longer than apical part (5: 2), with four lamellae and some bristles. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A25FF3ED44FFE5DFAA9.taxon	materials_examined	Material examined. Type: ♀ holotype: YEMEN: Aden, Jebel Jihaf, about 7000 ft., 7. X. 1937, at edge of culti- vation, H. Scott & E. B. Britton leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A25FF3ED44FFE5DFAA9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A25FF3ED44FFE5DFAA9.taxon	distribution	Distribution. Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	description	(Figs 38 F, 39 C, 40)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with head reticulate rugose (Fig. 39 C); clypeus and frons black; posterior surface of pronotum longer than mesoscutum, almost completely granulate; forewing with one dark spot around 2 r-rs & Rs vein (Fig. 39 C); propodeal declivity with two complete longitudinal keels; protarsomere 4 longer than basal part of 5.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	description	Description. ♀ (Figs 39 C, 40). Fully winged; body length 3.1 – 4.6 mm (3.1 mm in holotype). Head black, except mandible testaceous; antenna testaceous, except antennomeres 5 – 10 brown (in holotype) (in paratype from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, antenna testaceous except antennomere 10 brown; in paratype from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, antenna testaceous, except antennomeres 7 – 10 brown); mesosoma black; metasoma brown; legs of holotype and paratype from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, brown, except tarsi testaceous; legs of paratype from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, yellow-testaceous, except basal third of metacoxa and stalk of metafemur brown. Antenna clavate; antennomeres in following proportions: 11: 5: 6: 4: 5: 5: 5: 4.5: 5: 7.5. Head (Fig. 40 C) dull, reticulate rugose; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 5; TL = 4; greatest breadth of lateral ocelli about as long as OL. Pronotum dull, granulate, except posterior margin smooth, unsculptured; posterior sur- face longer than mesoscutum (16: 13); pronotal tubercle reaching tegula. Mesoscutum (Fig. 40 A) shiny, finely punc- tate, very slightly granulate among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesos- cutum in holotype and in paratype from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E (0.3 × in paratype from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two longitudinal keels and median area as ru- gose as lateral areas. Forewing (Fig. 40 D) with one slightly dark spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 8: 2.5: 3: 6.5: 15; protarsomere 4 longer than basal part of 5 (6.5: 5). Enlarged claw (Fig. 38 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 38 F) with basal part shorter than distal part (5: 10), with one row of 18 – 24 lamellae; apex with about 11 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19440): MADAGASCAR: Mahajanga, Beanka Forest Re- serve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, YPT, tropical dry forest on tsingy, B. L. Fisher et al. leg., BLF 22788 (CAS). Paratypes: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 21 – 26. IV. 2001, MT, MA- 01 - 01 D- 08, R. Harin’Hala leg., 1 ♀ (MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 27. IV. 2003, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 58, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC22A26FF3ED3CEFAE0F82B.taxon	etymology	Etymology. The species is named beankanum after the collection site in Beanka Forest Reserve.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED6D7FE24FC6D.taxon	description	(Fig. 38 G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED6D7FE24FC6D.taxon	description	Description. ♀. Fully winged; body length 3.9 mm. Head black, except mandible testaceous; antenna brown, except antennomeres 1 – 3 testaceous; mesosoma black, except pronotal tubercle and lateral regions of pronotum testaceous; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 21: 7: 11: 8: 8: 9: 9: 9: 9: 11. Head dull, reticulate rugose, with two keels near orbits; frontal line complete; occipital carina complete; vertex with two strong complete oblique keels connecting lateral ocelli to occipital carina (as in many Deinodryinus); POL = 8; OL = 6; OOL = 9; OPL = 8; TL = 10. Pronotum shiny, crossed by strong transverse impression; posterior surface shorter than mesoscutum (13: 25), with anterior declivity strongly sculptured by transverse keels and posterior region flat, partly granulate and partly sculptured by transverse keels; pronotal tubercle reaching tegula. Mesoscutum dull, granulate, with anterior fourth shiny, unsculptured. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4.5: 9.5). Protarsomeres in following proportions: 7: 3: 5: 11: 28. Enlarged claw (Fig. 38 G) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 38 G) with basal part shorter than apical part (10: 18), with two rows of approximately 18 lamellae; apex with about 17 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED6D7FE24FC6D.taxon	materials_examined	Material examined. Type: ♀ holotype: ZAMBIA: Lusaka, 13 – 18. II. 1980, MT, R. A. Beaver leg. (LACM).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED6D7FE24FC6D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED6D7FE24FC6D.taxon	distribution	Distribution. Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED272FE59F8A6.taxon	description	(Figs 36 B, C, 38 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED272FE59F8A6.taxon	description	Description. ♂ (Figs 36 B, C). Fully winged; body length 2.4 mm. Head black, except distal half of mandible tes- taceous; antenna brown; mesosoma black; metasoma brown-black; legs brown. Antenna filiform; antennomeres in following proportions: 12: 7: 6: 7: 7: 7: 7: 7: 7: 10. Head dull, reticulate rugose and granulate; frontal line complete (irregular in front of anterior ocellus); occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 2.5; TL = 4; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum shiny, irregularly rugose. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum rugose. Metapectalpropodeal disc reticulate rugose, with transverse posterior keel; propodeal declivity completely reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 8). Paramere (Fig. 38 C) with one large distal inner process showing many transverse folds. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED272FE59F8A6.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Eastern Prov., Nyambene Hills, Itieni Forest, at bottom, 0.24433 ° N 37.87016 ° E, 2142 m, 27. XI – 11. XII. 2011, MT, edge of indigenous forest, near forest station, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED272FE59F8A6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A27FF3ED272FE59F8A6.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A38FF3ED13DFE32FD00.taxon	description	(Fig. 42 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A38FF3ED13DFE32FD00.taxon	description	Description. ♂. Fully winged; body length 1.7 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown. Antenna filiform; antennomeres in following proportions: 5: 4: 7: 7: 6: 7: 7: 6: 6: 7. Head shiny, smooth, swollen, punctate, unsculptured among punctures; frontal line absent; frons with median longitudinal furrow; occipital carina complete; POL = 4; OL = 2; OOL = 6; OPL = 4; TL = 6. Mesoscutum, mesoscutellum and metanotum shiny, smooth, finely punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Metapectal-propodeal disc dull, completely reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Paramere (Fig. 42 A) much shorter than penis. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A38FF3ED13DFE32FD00.taxon	materials_examined	Material examined. Type: ♂ holotype: TANZANIA: East Entrance of Serengeti National Park, 24. II. 1976, G. E. Bohart leg. (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A38FF3ED13DFE32FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFC02A38FF3ED13DFE32FD00.taxon	distribution	Distribution. Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A38FF3ED480FDBCF90C.taxon	description	(Figs 41 A, B, 42 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A38FF3ED480FDBCF90C.taxon	description	Description. ♀ (Figs 41 A, B). Brachypterous, with forewing very reduced, slightly longer than mesoscutum + me- soscutellum + metanotum. Length 2.5 – 3.4 mm. Head black, except mandible testaceous-brown; antenna brown, except antennomeres 1 – 3 testaceous; mesosoma and metasoma black; tegula testaceous; legs of holotype testaceous, except coxae and clubs of femora partly brown; legs of specimen from Mozambique, Cuamba, yellow, except part of meso- and metacoxa and stalks of meso- and metafemur brown. Antenna clavate; antennomeres in following proportions: 10: 6: 7: 7: 7: 6: 6: 6: 6: 8. Head dull, granulate; frontal line present; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 4.5; TL = 5. Pronotum dull, setose, granulate, approximately twice as long as mesoscutum (13: 5); pronotal tubercle reaching tegula. Mesoscutum dull, very reduced, granulate and reticulate rugose. Notauli absent. Mesoscutellum shiny, small, unsculptured, slightly shorter than mesoscutum (4: 5). Metanotum short, approximately 0.5 × as long as mesoscutellum (2: 4). Mesopleuron and metapleuron transversely striate. Metapectal-propodeal disc dull, reticulate rugose and granulate, with strong transverse posterior keel; propodeal declivity granulate, with few irregular keels, not reticulate rugose. Protarsomeres in following proportions: 7: 2: 2: 4: 10; no protarsomeres produced into hook. Chela without rudimentary claw. Enlarged claw (Fig. 42 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 42 B) with one row of 14 – 20 lamellae; apex with 4 – 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A38FF3ED480FDBCF90C.taxon	materials_examined	Material examined. Type: ♀ holotype: ERITREA: Ghinda (ZMUH). Other material: MOZAMBIQUE: Ni- assa, Cuamba, Catholic University of Mozambique, Campus of Agriculture Faculty, M. Olmi reared from a nymph of an unknown species of Cicadellidae showing two thylacia, parasitized host collected 21. III. 2008, dryinid cocooning 22. III. 2008, dryinid adult emergence in 2009, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A38FF3ED480FDBCF90C.taxon	biology_ecology	Hosts. In Mozambique: nymphs of unidentified species of Cicadellidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A38FF3ED480FDBCF90C.taxon	distribution	Distribution. Eritrea, Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A3AFF3ED093FE59F895.taxon	description	(Figs 36 D, E, 42 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A3AFF3ED093FE59F895.taxon	description	Description. ♀ (Figs 36 D, E). Fully winged; body length 3.1 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma black; metasoma brown; tegula testaceous; legs testaceous. Antenna clavate; antennomeres in following proportions: 13: 8: 6: 5: 4.5: 6: 6: 6: 6: 8. Head dull, strongly reticulate rugose; frontal line present; frons with- out lateral keels; occipital carina complete; POL = 7.5; OL = 4; OOL = 6; OPL = 6; TL = 5; greatest breadth of lateral ocelli shorter than OL (3: 4). Pronotum rugose, with posterior surface shiny and almost completely smooth; posterior surface shorter than mesoscutum (5: 18). Mesoscutum shiny, very weakly granulate. Notauli very short, reaching about 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron proximally reticulate rugose, distally unsculptured. Metapleuron rugose, sculptured by many transverse keels. Metapectal-pro- podeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, with two longitudinal keels and median area sculptured by areolae less wide than those of lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 8: 2: 3: 4: 15. Protarsomeres 2 and 3 produced into hook. Protarsomere 4 much shorter than basal part of 5 (5: 11). Enlarged claw (Fig. 42 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 42 D) with two rows of 8 + 9 proximal and medial lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A3AFF3ED093FE59F895.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Taita Hills, Mbololo Forest, 03 ° 20.00 ’ S 38 ° 26.85 ’ E, 1550 m, MT, 5 – 13. IV. 1999, Taita Biodiversity project (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A3AFF3ED093FE59F895.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDF2A3AFF3ED093FE59F895.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	description	(Figs 42 C, 43 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	synonymic_list	Hirtanteon canabensis Benoit 1951 b: 164.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	description	Anteon canabense (Benoit): Olmi 1984: 370; Olmi & van Harten 2006: 318; Olmi et al. 2015: 342. Description. ♀. Fully winged; body length 2.3 – 3.5 mm. Completely testaceous-reddish, except petiole black; in specimen from Sierra Leone, 08 ° 31 ’ N 13 ° 17 ’ W, metapectal-propodeal disc darkened. Antenna clavate; antennomeres in following proportions: 12: 4: 7: 6: 5: 6: 5: 6: 5: 8. Head shiny, setose, granulate and reticulate rugose; frontal line present; frons with two lateral keels near orbits; occipital carina complete; POL = 7; OL = 5; OOL = 5 OPL = 4; TL = 4. Pronotum dull, granulate, with numerous transverse keels; posterior surface longer than OPL (7: 4) and shorter than mesoscutum (7: 14); pronotal tubercle reaching tegula. Mesoscutum dull, granulate, reticulate rugose near lateral and posterior margins; occasionally mesoscutum granulate and sculptured by numerous irregular keels. Notauli incomplete, reaching about 0.5 × length of mesoscutum; occasionally notauli hardly visible among areolae and keels. Mesoscutellum shiny, unsculptured. Metanotum dull, not smooth, sculptured by numerous longitudinal keels. Mesopleuron and metapleuron granulate and sculptured by short transverse keels. Metapectal-propodeal disc reticulate rugose, dull, with transverse posterior keel; propodeal declivity completely granulate, without longitudinal keels, with few irregular keels near margins. Forewing hyaline, without dark transverse bands (in specimens from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, and Yemen, Ta’izz, forewing with small dark spot around 2 r-rs & Rs vein); distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7) (distal part longer than proximal part, according to Benoit 1954). Protarsomere 1 slightly longer than 4 (7: 6); protarsomere 3 produced into hook. Protarsomere 4 approximately as long as basal part of 5. Protarsomere 5 (Figs 42 C, 43 A) with basal part shorter than distal part (7: 8), with inner side curvilinear and with apical region distinct. Enlarged claw (Figs 42 C, 43 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 42 C) with one row of 23 lamellae (specimen from Yemen, Ta’izz, with two rows of 11 + 17 lamellae (Fig. 43 A); specimen from South Africa, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, with two rows of lamellae); apex with 4 – 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Ubangi, Likimi, Mundjungani, 4. X. 1927, A. Collart leg. (MRAC). Other material: BOTSWANA: Serowe, II. 1990, 1 ♀ (CAS). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 6 – 14. IV. 2006, MT, forest, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD). KENYA: Rift Valley Prov., road to Nguruman Town, 1.83216 ° S 36.08156 ° E, 676 m, 27 – 29. III. 2013, 6 m MT, edge of indigenous forest, R. Copeland leg., 1 ♀ (NMK). NIGERIA: Edo State, Udo Forest Reserve, 2 ♀♀ (AEIC, AMNH). SENEGAL: Niokolo Koba National Park, 13 ° 01.1 ’ N 13 ° 18.5 ’ W, 15. VII. 2004, Marek Halada leg., 2 ♀♀ (OLL). SIERRA LEONE: Freetown, Cape Sierra Hotel area, 08 ° 31 ’ N 13 ° 17 ’ W, 19. XI. 1993, 1 ♀ (ZIL). SOUTH AFRICA: KwaZ ulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 6. I – 1. II. 2006, MT, G. Whiteley leg., 1 ♀ (DJBC); Limpopo, 15 km E of Klaserie, Guern- sey Farm, 2 ♀♀ (AMNH, CNC). YEMEN: Ta’izz, V – VI. 2000, Light trap, N. 4770, A. van Harten & A. R. Al Yarimi leg., 1 ♀ (MOLC); same locality label, VII. 2002, N. 7316, 1 ♀ (MOLC); 20 km S Ta’izz, 13 ° 30 ’ N 43 ° 57 ’ E, 1200 m, 24. X. 2005, J. Halada leg., 1 ♀ (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDD2A3CFF3ED117FEA3FC04.taxon	distribution	Distribution. Botswana, Democratic Republic of the Congo, Kenya, Nigeria, Senegal, Sierra Leone, South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDB2A3CFF3ED59BFCE1F8CE.taxon	description	(Figs 43 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDB2A3CFF3ED59BFCE1F8CE.taxon	description	Description. ♂. Fully winged; body length 1.9 – 3.1 mm. Black, except mandible testaceous, antenna and legs brown. Antenna filiform; antennomeres in following proportions: 8: 5: 10: 11.5: 11.5: 11.5: 10: 10: 9: 9. Head shiny, smooth, slightly punctate, unsculptured among punctures (in specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, head unsculptured except vertex and temple granulate); frontal line absent; frons without lateral keels, with slight median longitudinal furrow; occipital carina complete; POL = 7; OL = 4; OOL = 6; OPL = 3; TL = 5. Mesoscutum shiny, slightly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc completely reticulate rugose, without transverse posterior keel. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 9). Paramere (Fig. 43 B) without distal inner process and proximal membranous process. In specimen from South Africa, Kogelberg Forest Reserve, paramere with small basal area sculptured with mosaic pattern (Fig. 43 C). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDB2A3CFF3ED59BFCE1F8CE.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Matjesfontein, 6 – 15. X. 1928, R. E. Turner leg. (NHMUK). Other material: MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 17 – 24. III. 2003, MT, spiny forest, MA- 02 - 22 - 19, M. Irwin, F. Parker & R. Harin’Hala leg., 1 ♂ (CAS). NAMIBIA: Namib / Naukluft Park, near Gobabeb, Kuiseb River, 1 ♂ (SANC). SOUTH AFRICA: Western Cape, De Hoop Nature Reserve, 34 ° 27 ’ S 20 ° 25 ’ E, 0 – 200 m, 2 ♂♂ (1 in ZIL, in AMNH); Kogelberg Forest Reserve, 4 – 22. III. 1983, D. M. Kroon leg., 1 ♂ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDB2A3CFF3ED59BFCE1F8CE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDB2A3CFF3ED59BFCE1F8CE.taxon	distribution	Distribution. Madagascar, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDA2A3FFF3ED6D7FF17F964.taxon	description	(Figs 43 D – F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDA2A3FFF3ED6D7FF17F964.taxon	description	Description. ♂. Fully winged; body length 1.4 – 2.9 mm. Head black, except mandible testaceous; antenna tes- taceous, occasionally brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa, club of metafemur and metatibia partly brown; in two specimens from Madagascar, 17 ° 17.00 ’ S 49 ° 26.00 ’ E, and three specimens from Kenya, Vuria Forest, legs completely yellow-testaceous. Antenna filiform; antennomeres in following proportions: 7: 4: 4: 4: 4: 4.5: 4: 4: 4: 6. Head dull or shiny, reticulate rugose and granulate (occasionally area in front of anterior ocellus punctate and unsculptured among punctures; in few small specimens from Kenya, 2.07846 ° S 38.22530 ° E, head almost completely unsculptured and shiny, except some lateral and anterior areas granulate); frontal line complete (absent in three specimens from Kenya, Vuria Forest); occipital carina complete; POL = 6; OL = 3; OOL = 3.5; OPL = 3; TL = 3. In specimen from Kenya, Vuria Forest, POL = 6; OL = 3; OOL = 5; OPL = 2; TL = 3; greatest breadth of lateral ocelli about as long as OL. Mesoscutum shiny, weakly or strongly granulate. In two specimens from Madagascar, 18 ° 55.58 ’ S 48 ° 24.47 ’ E and Ranomafana, mesoscutum finely punctate and unsculptured among punctures (as in Anteon emeritum Olmi). In specimens from Kenya, 2.07846 ° S 38.22530 ° E, mesoscutum almost completely unsculptured and shiny, except small lateral areas granulate. In specimen from Uganda, 0 ° 33.859 ’ S 30 ° 21.630 ’ E, mesoscutum slightly granulate, except small surface near anterior margin rugose. Notauli incomplete, reaching about 0.3 – 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, unsculptured or finely punctate. Metapectal-propodeal disc reticulate rugose, with transverse posterior keel; propodeal declivity completely reticulate rugose, with tracks of two longitudinal keels; in specimens from Madagascar (21 ° 13.57 ’ S 47 ° 22.19 ’ E, 17 ° 17.00 ’ S 49 ° 26.00 ’ E and 18 ° 55.58 ’ S 48 ° 24.47 ’ E), propodeal declivity with two distinct longitudinal keels and median area as rugose as lateral areas (in specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, median area partly shiny and unsculptured). Usually largest specimens with distinct longitudinal keels on propodeal declivity, smallest specimens with tracks of longitudinal keels. Occasionally propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 7). Paramere (Figs 43 D – F) with distal inner rounded (in small specimens) or pointed (in large specimens) process; dorsal membranous process with distal area bare (Fig. 43 D). In specimens from Madagascar, inner side of paramere with membranous band transversely striate (Fig. 43 F). In specimen from Madagascar, 17 ° 41 ’ 36 ’’ S 49 ° 27 ’ 37 ’’ E, MG- 37 B- 12, distal region of paramere inner process only visible as darker region. In some specimens from Madagascar, distal inner process faint (then genitalia very similar to those of A. zairense, but with dorsal membranous process short (long in A. zairense (Figs 79 F, G )). In specimen from Democratic Republic of the Congo, 01 ° 56 ’ 29.82 ’’ N 30 ° 02 ’ 19.17 ’’ E, distal inner process of paramere very large. Tibial spurs 1 / 1 / 2. ♀. Unknown (see remarks under Anteon zairense Benoit).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDA2A3FFF3ED6D7FF17F964.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Toliara, 12 km NW Amboasary, Berenty, 5 – 15. V. 1983, J. S. Noyes & M. C. Day leg. (NHMUK). Paratypes: MADAGASCAR: same locality label as holotype, 4 ♂♂ (1 in NHMUK, 3 in AMNH). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Reserve speciale de Forêt dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 17. V. 2001, sweep, lowland rainforest, S. van Noort leg., 4 ♂♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Ituri, Djugu Territory, Mongbwalu mining headquarters, behind Titanic building, 01 ° 56 ’ 29.82 ’’ N 30 ° 02 ’ 19.17 ’’ E, 1224 m, Yellow Pan Traps, agroforestry, 2 – 9. III. 2015, A. Gumovsky leg., 1 ♂ (SIZK); Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, VII. 2008, MT, T. Chapman leg., 1 ♂ (DEUCD); same local- ity label, 13. IV. 2006, forest pantrap, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (DEUCD); same locality label, 6 – 14. IV. 2006, forest MT, 1 ♂ (DEUCD); same locality label, 04 ° 16 ’ S 17 ° 11 ’ E, 3. IV. 2006, sweep in forest along river, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (DEUCD). KENYA: Coast Prov., Funzi Island, 4.57776 ° S 39.44127 ° E, near sea level, 24 – 28. VII. 2012, MT, mixed grass- woodland, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 24. VII – 7. VIII. 2011, MT, next to small forest pond, R. Copeland leg., 1 ♂ (NMK); same locality label, 28. XI – 12. XII. 2011, 1 ♂ (NMK); same locality label, 4 – 18. IX. 2011, 1 ♂ (NMK); Coast Prov., Taita Hills, Vuria Forest, 3.41428 ° S 38.29178 ° E, 2162 m, MT just inside indigenous forest, 5 – 19. IX. 2012, R. Copeland leg .. 1 ♂ (NMK); same locality label, 21. VIII – 4. IX. 2011, R. Copeland leg .. 2 ♂♂ (NMK, MOLC); same locality label, 7 – 21. VIII. 2011, 1 ♂ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 26. VII – 9. VIII. 2015, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg., 9 ♂♂ (8 in NMK, 1 in MOLC); Western Prov., Kakamega District, Kakamega Forest, 19. V – 4. VI. 2000, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 3 ♂♂ (1 in MOLC, 2 in MTC). MADAGASCAR: Antsira- nana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 16 – 31. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 10, R. Harin’Hala leg., 1 ♂ (CAS); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 3 ♂♂ (CAS); Antsiranana, Marojejy National Park, Manantenina river, 28.0 km 38 ° NE Andapa, 8.2 km 333 ° NNW Manantenina, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, MT, rainforest, B. L. Fisher et al. leg., BLF 8723, 4 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 27. I – 11. II. 2007, MT, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 01 - 01 A- 17, 1 ♂ (CAS); same locality label, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg., 6 ♂♂ (CAS); same locality label, 4 – 19. III. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 08, 3 ♂♂ (CAS); same locality label, 21 – 28. I. 2001, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 2 ♂♂ (CAS); Antsiranana, Mon- tagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 900 m, 29. I. 2001, netted by hand, M. E. Irwin leg., MA- 01 - 01 A- 05, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 6 – 21. IV. 2001, MT, MA- 01 - 01 D- 07, R. Harin’Hala leg., 1 ♂ (MOLC); same locality label, 29. I – 11. II. 2001, R. Harin’Hala leg., MA- 01 - 01 D- 03, 2 ♂♂ (CAS); same locality label, 11. II – 4. III. 2001, MA- 01 - 01 D- 04, 3 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, 11. II – 4. III. 2001, MT, MA- 01 - 01 B- 05, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 4 – 10. III. 2001, MA- 01 - 01 B- 06, 2 ♂♂ (CAS); An- tsiranana, Manongarivo Special Reserve, 14.5 km 220 ° SW Antanambao, 13 ° 59.9 ’ S 48 ° 25.7 ’ E, 1175 m, 20. X. 1998, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher leg., 1938, 1 ♂ (CAS); Antsiranana, SAVA Region, Vo- hemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 4 – 10. X. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 01, 1 ♂ (CAS); Fianarantsoa, 45 km S Ambalavao, 22 ° 13 ’ S 47 ° 01 ’ E, 785 m, 5. IX – 2. X. 1993, B. L. Fisher leg., 1 ♂ (DEUCD); Fianarantsoa, Ranomafana National Park, Vo- hiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 4 – 12. II. 2002, 1110 m, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 2 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 A- 04, 3 ♂♂ (CAS); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 6 ♂♂ (4 in CAS, 2 in MOLC); same locality label, 14 – 21. I. 2002, 2 ♂♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 A- 07, 2 ♂♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 5 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 4 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 2 ♂♂ (CAS); same lo- cality label, 26. II – 10. III. 2003, 2 ♂♂ (CAS, MOLC); same locality label, 19 – 26. II. 2002, 4 ♂♂ (2 in MOLC, 2 in CAS); same locality label, 5 – 13. V. 2002, MA- 02 - 09 C- 28, 2 ♂♂ (CAS, MOLC); same locality label, 14 – 24. VII. 2002, 2 ♂♂ (CAS); same locality label, 6 – 18. VI. 2003, MA- 02 - 09 C- 63, 3 ♂♂ (1 in MOLC, 2 in CAS); same locality label, 23. VI – 4. VII. 2002, MA- 02 - 09 C- 33, 1 ♂ (CAS); same locality label, 10 – 21. III. 2003, MA- 02 - 09 C- 56, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 10 – 14. I. 2002, MA- 02 - 09 C- 11, 4 ♂♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 C- 10, 2 ♂♂ (CAS); same locality label, 26. II – 4. III. 2002, MA- 02 - 09 C- 18, 2 ♂♂ (CAS); same locality la- bel, 28. XI – 6. XII. 2001, MA- 02 - 09 C- 05, 1 ♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 C- 06, 1 ♂ (CAS); same locality label, 4 – 14. VII. 2002, MA- 02 - 09 C- 34, 3 ♂♂ (CAS); same locality label, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 1 ♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 1 ♂ (CAS); same locality label, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 4 ♂♂ (CAS); same locality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 10 ♂♂ (CAS); same locality label, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 5 ♂♂ (CAS); same locality label, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 3 ♂♂ (CAS); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 2 ♂♂ (MOLC); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 3 ♂♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 3 ♂♂ (CAS); same locality label, 26 – 31. III. 2002, MA- 02 - 09 C- 22, 3 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 8 ♂♂ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 8 ♂♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 10 ♂♂ (9 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 3 ♂♂ (1 in CAS, 2 in MOLC); same locality label, 7 – 18. VI. 2003, 4 ♂♂ (3 in CAS, 1 in MOLC); same local- ity label, 9 – 16. IV. 2002, 1 ♂ (CAS); same locality label, 4 – 12. III. 2002, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 B- 15, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 1 ♂ (CAS); same locality label, 14 – 24. VI. 2002, MA- 02 - 09 B- 32, 4 ♂♂ (CAS); same locality label, 15 – 27. IV. 2003, MA- 02 - 09 B- 58, 2 ♂♂ (CAS); same locality label, 12 – 19. III. 2002, MA- 02 - 09 B- 20, 6 ♂♂ (CAS); same locality label, 9 – 20. III. 2003, MA- 02 - 09 B- 55, 4 ♂♂ (CAS); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 1 ♂ (CAS); same locality label, 3 – 15. IV. 2003, MA- 02 - 09 B- 57, 2 ♂♂ (CAS); same locality label, 18 – 29. VI. 2003, MA- 02 - 09 B- 64, 4 ♂♂ (CAS); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 B- 03, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 4 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 5 ♂♂ (CAS); same locality label, 20. III – 3. IV. 2003, MA- 02 - 09 B- 56, 4 ♂♂ (CAS); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 3 ♂♂ (CAS); same locality label, 24. V – 4. VI. 2002, MA- 02 - 09 B- 30, 6 ♂♂ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 2 ♂♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 13 ♂♂ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 3 ♂♂ (CAS); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 7 ♂♂ (CAS); same locality label, 7 – 17. VII. 2003, MA- 02 - 09 B- 66, 4 ♂♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 3 ♂♂ (CAS); Ranomafana, 27. I. 2004, Brantlová & Kubik leg., 1 ♂ (OLL); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. XI – 6. XII. 2001, MT near river, R. Harin’Hala leg., MA- 02 - 09 D- 05, 1 ♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 A- 11, 2 ♂♂ (CAS); same locality label, 21 – 24. XII. 2001, MA- 02 - 09 D- 08, 1 ♂ (CAS); Fianarantsoa, Manombo Special Reserve, 24.5 km 228 ° SW Farafangana, 23 ° 00.57 ’ S 47 ° 43.08 ’ E, 30 m, 20. IV. 2006, MT, rainforest, B. L. Fisher et al. leg., BLF 13960, 3 ♂♂ (2 in CAS, 1 in MOLC); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 25. X – 4. XI. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 75, 1 ♂ (CAS); same locality label, 28. XII. 2006 – 4. I. 2007, MA- 29 - 73, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 750 m, 15. X – 9. XI. 2001, MT, MA- 02 - 11 A- 01, M. E. Irwin, F. D. Parker, R. Harin’Hala leg., 1 ♂ (CAS); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♂ (MNHN); Toamasina, Parcelle E 3 Tampolo, 17 ° 17.00 ’ S 49 ° 26.00 ’ E, 10 m, 14. IV. 2004, MT, littoral forest, Malagasy ant team leg., BLF 10729, CASLOT 021823, 3 ♂♂ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 8 – 16. X. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 15, 5 ♂♂ (CAS); same locality label, 16 – 23. XI. 2001, MA- 01 - 08 B- 20, 7 ♂♂ (CAS); same locality label, 1 – 7. XI. 2001, 8 ♂♂ (7 in CAS, 1 in MOLC); Toamasina, Mobot site, Analalava, 7 km SW of Foulpointe, 17 ° 41 ’ 36 ’’ S 49 ° 27 ’ 37 ’’ E, 60 ft., 7 – 14. XII. 2007, MT on sand in low altitude dense humid forest, MG- 37 B- 13, 1 ♂ (CAS); same locality label, 1 – 8. II. 2008, MG- 37 B- 21, 1 ♂ (CAS); same locality label, 9 – 16. XI. 2007, MG- 37 B- 09, 1 ♂ (MOLC); same locality label, 30. XI – 7. XII. 2007, MG- 37 B- 12, 1 ♂ (MOLC); same locality label, 18 – 25. I. 2008, MG- 37 B- 19, 1 ♂ (CAS); Toamasina, Ambatovaky Special Reserve, Sandrangato River, 16 ° 49.03 ’ S 49 ° 17.42 ’ E, 360 m, 25 - 27. II. 2010, MT, rainforest, B. L. Fisher et al. leg., BLF 24749, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 9 – 20. IX. 2002, MT in dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 1 ♂ (MOLC); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.09 ’ S 044 ° 39.38 ’ E, 10 – 17. XII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 58, 2 ♂♂ (CAS); same locality label, 5 – 12. XII. 2007, BLF 18285 _ 05, 1 ♂ (CAS). SOUTH AFRICA: KwaZulu-Natal, Karkloof, 29 ° 19.1 ‘ S 30 ° 15.5 ‘ E, 1325 m, 28. IX – 3. X. 2005, YPT, M. Mostovski leg., 3 ♂♂ (2 in DJBC, 1 in MOLC); KwaZulu-Natal, Ashburton, 29 ° 39 ‘ 23 ‘‘ S 30 ° 27 ‘ 41 ‘‘ E, 8. XII. 2005 – 6. I. 2006, MT, G. Whiteley leg., 1 ♂ (DJBC); Western Cape, Cape of Good Hope Nature Reserve, Booi se Skerm, 34 ° 18.27 ‘ S 18 ° 27.64 ‘ E, 24. II. 1998, sweep, strandveld, S. van Noort leg., 2 ♂♂ (SAMC). UGANDA: Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 33.859 ’ S 30 ° 21.630 ’ E, 1505 m, 5 – 12. VIII. 2005, MT, primary mid-altitude rainforest, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDA2A3FFF3ED6D7FF17F964.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFDA2A3FFF3ED6D7FF17F964.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Kenya, Madagascar, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD82A30FF3ED17BFE14FD74.taxon	description	(Fig. 44 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD82A30FF3ED17BFE14FD74.taxon	description	Description. ♀. Fully winged; body length 2.7 mm. Head black, except mandible testaceous with teeth brown; antenna brown, except antennomeres 1 – 4 testaceous; mesosoma and metasoma black; tegula testaceous; legs testaceous, except metacoxa and club of metafemur partly brown. Antenna clavate; antennomeres in following proportions: 11: 4: 5: 4: 4: 4: 4: 4: 4: 7. Head dull, granulate, with numerous irregular transverse keels; occipital carina complete; POL = 4; OL = 5; OOL = 4; OPL = 5; TL = 2. Pronotum (damaged in holotype) sculptured by transverse keels, with smooth area near posterior margin. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Protarsomere 1 longer than 4 (9: 2); protarsomere 4 produced into hook, approximately 0.5 × as long as basal part of 5. Enlarged claw (Fig. 44 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 A) with basal part much longer than apical part, with some proximal and medial bristles; apex with six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD82A30FF3ED17BFE14FD74.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, I. 1940, A. Seyrig leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD82A30FF3ED17BFE14FD74.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD82A30FF3ED17BFE14FD74.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED6D7FE59FBD0.taxon	description	(Figs 36 F, 39 D, 46 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED6D7FE59FBD0.taxon	description	Description. ♀ (Figs 36 F, 39 D). Fully winged; body length 2.5 mm. Head black, except mandible testaceous; an- tenna testaceous, except antennomeres 7 – 10 or 6 – 10 brown; mesosoma black; metasoma brown; legs testaceous, except stalk of metafemur darkened. Antennomeres in following proportions: 13: 6: 7: 4: 5: 6: 6: 6: 6: 7. Head shiny, reticulate rugose; frontal line complete; frons with two lateral longitudinal keels near orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 6; OOL = 4.5; OPL = 7; TL = 3; greatest breadth of lateral ocelli about as long as TL. Pronotum shiny, rugose; posterior surface very short, shorter than mesoscutum (6: 17); pronotal tubercle reaching tegula. Mesoscutum shiny, reticulate rugose. Notauli absent. Mesoscutellum partly smooth, punctate, unsculptured among punctures, with some areolae. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels, with areolae about as large as those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 11). Protarsomeres in following proportions: 10: 2: 2: 3: 12. Basal part of protarsomere 5 much longer than distal part (10: 2). Arolium very large, slightly shorter than basal part of protarsomere 5 (8: 10). Enlarged claw (Fig. 46 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 46 A) with five proximal and medial bristles; apex with about five lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED6D7FE59FBD0.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Arabuko-Sokoke Forest, 03 ° 25.21 ’ S 39 ° 53.81 ’ E, 29. X – 5. XI. 1999, MT, R. Copeland leg. (NMK). Other material. KENYA: Coast Prov., Marenje Forest, 4.52814 ° S 39.24028 ° E, 76 m, 3 – 17. X. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (MOLC); Coast Prov., Buda Forest, 4.46109 ° S 39.40446 ° E, 98 m, 25. III – 8. IV. 2016, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED6D7FE59FBD0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED6D7FE59FBD0.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED2E6FE59F87E.taxon	description	(Figs 41 C, D, 44 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED2E6FE59F87E.taxon	description	Description. ♀ (Figs 41 C, D). Fully winged; body length 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown. Antenna clavate; antennomeres in following proportions: 14: 6: 4.5: 3.5: 4: 4.5: 4.5: 4: 5: 7. Head convex, dull, strongly reticulate rugose; frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 5; OOL = 6; OPL = 4; TL = 3; greatest breadth of posterior ocellus about as long as TL. Pronotum anteriorly crossed by slight transverse impression, dull, rugose; posterior surface rugose, much shorter than mesoscutum (3: 15); pronotal tubercle eaching tegula. Mesoscutum, mesoscutellum and metanotum dull, completely strongly reticulate rugose. Notauli absent. Metapectal-propodeal disc dull, reticulate rugose, with strong transverse posterior keel; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 8). Protarsomeres in following proportions: 8: 2: 2: 3: 10. Enlarged claw (Fig. 44 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 B) with basal region much longer than distal region (8: 2), with many proximal and medial bristles (some bristles are thicker than others); apex with one lamella. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED2E6FE59F87E.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Western Prov., Kakamega Forest, near Rondo Guesthouse, 0.22767 ° N 34.88533 ° E, 1630 m, 17 – 31. XII. 2006, MT, set across small permament stream, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED2E6FE59F87E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD62A31FF3ED2E6FE59F87E.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	description	(Fig. 46 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	description	Description. ♂. Fully winged; body length 1.4 mm. Head black, except mandible testaceous; antenna testaceousbrown; mesosoma black; metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 7: 4.5: 5: 5: 5: 5.5: 6: 5.5: 5: 7. Head dull, granulate; frontal line absent; occipital carina complete; POL = 4; OL = 2; OOL = 3.5; OPL = 2.5; TL = 4. Mesoscutum granulate, with stripe near posterior margin shiny and unsculptured. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels, completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 4). Paramere (Fig. 46 B) with distal inner rounded process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, 17 km NE of Empangeni, Nseleni River, 28 ° 42 ’ S 32 ° 01 ’ E, 24. X. 1994, loc. 32, R. Danielsson leg. (ZIL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A32FF3ED6D7FE5DFC25.taxon	discussion	Remarks. Anteon danielssoni may be confused with A. inane Olmi, 2004. Some specimens of A. inane in fact have a granulate head and slightly evanescent striae. However, in A inane the frontal line is complete, whereas in A. danielssoni it is absent.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A33FF3ED5BAFC9DFE55.taxon	description	(Figs 44 C, 46 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A33FF3ED5BAFC9DFE55.taxon	description	Description. ♀. Fully winged; body length 2.4 – 3.0 mm. Head black, except mandible testaceous; antenna brown, except antennomeres 1 – 5 testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa brownish; occasionally also meso- and metacoxa, femora and tibiae brown. Antenna clavate; antennomeres in following proportions: 6: 5: 5: 4: 4: 4: 5: 5: 5: 7. Head dull, granulate, with slight areolae near clypeus; frontal line present; frons without lateral keels; occipital carina complete; POL = 7; OL = 5; OOL = 5 OPL = 5; TL = 4. Pronotum flat, dull, granulate; anterior surface shorter than posterior surface (5: 10); posterior surface shorter than mesoscutum (10: 14); pronotal tubercle reaching tegula. Mesoscutum shiny, unsculptured, with lateral regions granulate. Notauli absent, or slightly visible and incomplete, reaching almost 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity slightly reticulate rugose, without longitudinal keels. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Protarsomeres in following proportions: 6: 2: 3: 8: 16. Enlarged claw (Fig. 44 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 C) with basal part shorter than apical part (6: 10), with two rows of 26 lamellae; apex with 7 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.1 – 1.2 mm. Head black, except mandible testaceous; antenna brown, except antennomeres 1 or 1 – 2 testaceous; mesosoma and metasoma black; legs brown, except tarsi and protibia testaceous. Antenna filiform, almost moniliform; antennomeres in following proportions: 6: 3: 3: 3: 3: 3: 3: 3: 3: 4. Head dull, completely granulate; frontal line complete; occipital carina complete; POL = 5.5; OL = 3; OOL = 3; OPL = 1; TL = 2. Mesoscutum dull, granulate. Notauli incomplete, shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, strongly reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 rrs & Rs vein much shorter than proximal part (1: 5). Paramere (Fig. 46 C) without distal inner pointed process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A33FF3ED5BAFC9DFE55.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: 15 mi. SSW Windhoek, Regenstein, 9. IV. 1972, Southern African Expedition (NHMUK). Paratype: NAMIBIA: Okahandja, 2 – 18. III. 1928, R. E. Turner leg., 1 ♀ (NHMUK). Other material: BOTSWANA: Serowe, Farmer’s Brigade (USNM). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ‘ 52 ‘‘ S 49 ° 10 ‘ 53 ‘‘ E, 960 m, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg., 1 ♂ (MOLC); Toamasina, Andasibe National Park, 18 ° 55.58 ‘ S 48 ° 24.47 ‘ E, botanic garden near entrance to park, 1025 m, 8 – 16. X. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 15, 1 ♂ (CAS). SOUTH AFRICA: Limpopo, 15 km NW Klaserie, Guernsey Farm (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A33FF3ED5BAFC9DFE55.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD52A33FF3ED5BAFC9DFE55.taxon	distribution	Distribution. Botswana, Madagascar, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A33FF3ED46AFD68FA66.taxon	description	(Figs 44 D, 47 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A33FF3ED46AFD68FA66.taxon	description	Description. ♀ (Figs 47 A, B). Fully winged; body length 2.2 mm. Head black, except mandible testaceous; an- tenna testaceous, except dorsal side of antennomeres 6 – 10 brownish; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 10: 5: 6: 5: 4: 5: 6: 5.5: 5.5: 7. Head dull, granulate and reticulate rugose (Fig. 47 C); frontal line complete; frons without two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 3: OPL = 4; TL = 4; greatest breadth of lateral ocelli about as long as OL. Pronotum dull, reticulate rugose; posterior surface longer than OPL (7: 4) and shorter than mesoscutum (7: 14); pronotal tubercle reaching tegula. Mesoscutum dull, granulate and reticulate rugose, except short unsculptured area near anterior margin. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Mesopleuron and metapleuron dull, reticulate rugose. Metapectal-propodeal disc dull, with transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 9). Protarsomeres in following proportions: 5: 2: 3: 4: 13; protarsomere 2 produced into hook. Protarsomere 4 slightly shorter than basal part of 5 (4: 6). Protarsomere 5 (Fig. 44 D) with basal part slightly shorter than distal part (6: 7), with inner side curvilinear and with apical region distinct. Enlarged claw (Fig. 44 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 D) with one row of 16 lamellae; apex with 6 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A33FF3ED46AFD68FA66.taxon	materials_examined	Material examined. Type: ♀ holotype: CENTRAL AFRICAN REPUBLIC: angha-Mbaéré Prefecture, Réserve Spéciale de Forêt Dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 03 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 11 – 17. V. 2001, YPT, lowland rainforest, CAR 01 - Y 26, S. van Noort leg. (SAMC)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A33FF3ED46AFD68FA66.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A33FF3ED46AFD68FA66.taxon	distribution	Distribution. Central African Republic.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	description	(Figs 44 E, 45, 47 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon (Fig. 47 D) with head completely reticulate rugose; forewing hyaline, without dark transverse bands, with distal part of 2 r-rs & Rs vein less than 0.5 × as long as proximal part; propodeal declivity reticulate rugose, without longitudinal keels; protarsomere 4 approximately as long as basal part of 5; protarsomere 5 (Fig. 44 E) with inner side curvilinear, with distinct apical region, with basal part shorter than distal part, with less than 20 proximal and medial lamellae, with distal region very long and slender, with distal group of lamellae located far from medial lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	description	Description. ♀ (Figs 45, 47 D). Fully winged; body length 2.9 mm. Head black, except mandible, clypeus and transverse stripe between antennal toruli testaceous; antenna brown, except proximal half of scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 12: 5: 9: 7: 6: 7: 7: 7: 7: 9. Head (Fig. 45 C) dull, completely reticulate rugose; frontal line complete; frons with two lateral keels along orbits; occipital carina complete; POL = 5; OL = 3; OOL = 3; OPL = 3; TL = 4; greatest breadth of lateral ocelli as long as OPL. Pronotum dull, reticulate rugose; posterior surface shorter than mesoscutum (8: 16); pronotal tubercle reaching tegula. Mesoscutum (Fig. 45 A) shiny, punctate and with many irregular folds and striae. Notauli incomplete, reaching about 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculp- tured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticu- late rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 10). Protarsomeres in following proportions: 5: 2: 3: 11: 18. Enlarged claw (Fig. 44 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 E) with one row of 13 lamel- lae and distal region very long and slender; apex with three lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19441): MADAGASCAR: Mahajanga, Marotandrano Spe- cial Reserve, Marotandrano, 48.3 km S Mandritsara, 16 ° 17 ’ 00 ’’ S 8 ° 48 ’ 52 ’’ E, 865 m, 7 – 8. XII. 2007, YPT, transition humid forest, B. L. Fisher et al. leg., BLF 19159 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD42A34FF3ED07DFB0BF896.taxon	etymology	Etymology. The species is named elongatum because its body is particularly long and slender.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD32A36FF3ED12DFCB6FD00.taxon	description	(Fig. 46 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD32A36FF3ED12DFCB6FD00.taxon	materials_examined	Material examined. Type: ♂ holotype: NAMIBIA: Okahandja, Camping Place, 4. II. 1972, MT, riverside vegetation, Southern Africa Expedition (NHMUK). Other material: KENYA: Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 17 – 31. X. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (RSC); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 30. V – 19. VI. 2013, six-meter MT, indigenous forest, R. Copeland leg., 6 ♂♂ (2 in NMK, 3 in RSC, 1 in MOLC). MADAGASCAR: Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14 ° 26.29 ’ S 49 ° 46.44 ’’ E, 490 m, 28. VIII – 11. IX. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 29, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 2 ♂♂ (CAS). SOUTH AFRICA: Limpopo, Gravelotte, 20 km NW Beacon Ranch (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD32A36FF3ED12DFCB6FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD32A36FF3ED12DFCB6FD00.taxon	distribution	Distribution. Kenya, Madagascar, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD12A37FF3ED497FDF9FCBC.taxon	description	(Figs 44 F, 46 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD12A37FF3ED497FDF9FCBC.taxon	description	Description. ♀. Fully winged; body length 2.0 – 2.4 mm. Head black, except mandible testaceous; occasionally head brown-reddish, except dark ocellar spot; antenna completely testaceous, or brown, except antennomeres 1 – 3 testaceous; mesosoma black, except prothorax yellow or orange and some reddish spots on pleuron; metasoma brown or testaceous; legs testaceous, except stalk of metafemur darkened. In specimen from Kenya, Kakamega Forest, head and prothorax completely testaceous, except malar space brown; mesosoma black, except anterior half of mesopleuron, mesoscutum and mesoscutellum brown; metasoma testaceous, partly darkened dorsally; legs testaceous, except stalk of metafemur darkened. In specimen from Democratic Republic of the Congo, 01 ° 56 ’ 29.82 ’’ N 30 ° 02 ’ 19.17 ’’ E, head and mesosoma completely black; antenna brown, except antennomeres 1 – 2 testaceous; metasoma brown,. Antenna clavate; antennomeres in following proportions: 5: 3: 5: 4: 4: 4: 4: 4: 4: 6. Head dull, completely granulate; frontal line complete; occipital carina complete; POL = 4; OL = 3; OOL = 5; OPL = 4; TL = 4. Pronotum dull, completely granulate; posterior surface flat, slightly shorter than mesoscutum (8: 9). Mesoscutum dull, completely granulate, with some evanescent areolae near lateral margins. Notauli absent. Mesoscutellum dull, strongly or slightly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Protarsomeres in following proportions: 5: 1.5: 2: 6: 12. Enlarged claw (Fig. 44 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 44 F) with two rows of 22 – 23 lamellae; apex with five-six lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.4 – 1.7 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs completely testaceous; occasionally meso- and metatibia and clubs of femora brown. Antenna filiform; antennomeres in following proportions: 5: 3: 4: 5: 4: 4.5: 4.5: 4.5: 4: 6. Head dull, granulate; frontal line complete; occipital carina complete; POL = 4; OL = 3; OOL = 3; OPL = 2; TL = 3. In specimen from Madagascar, 22 ° 50.36 ’ S 44 ° 42.36 ’ E, OPL slightly longer than TL. In specimen from Kenya, 2.32031 ° N 37.98595 ° E, OPL as long as TL. Mesoscutum dull, granulate. Notauli very short, shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 5). Paramere (Fig. 46 D) without distal inner pointed process, usually without papillae on inner side (in specimen from Kenya, 2.32031 ° N 37.98595 ° E, paramere with few papillae on inner side). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD12A37FF3ED497FDF9FCBC.taxon	materials_examined	Material examined. Types: ♀ holotype: IVORY COAST: 3 km S Katiola, 26. XII. 1980, gallery forest, MT, J. W. Everts leg. (DEUW). Paratypes: IVORY COAST: Klébo, 8 km NW Bouaflé, 31. XII. 1980, MT, J. W. Everts leg., 1 ♂ [DEUW); same locality label, 7. I. 1981, 1 ♂ [DEUW]; same locality label, 18. II. 1981, 1 ♀ (AMNH); same locality label, 11. III. 1981, 1 ♂ (AMNH). Other material: CAMEROON: Messa Bengondo, 15 – 30. III. 2003, M. Tussac leg., 1 ♂ (HTS). CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 23. V. 2001, sweep, lowland rainforest, S. van Noort leg., 4 ♂♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Ituri, Djugu Territory, Mongbwalu mining headquarters, behind Titanic building, 01 ° 56 ’ 29.82 ’’ N 30 ° 02 ’ 19.17 ’’ E, 1224 m, Yellow Pan Traps, agroforestry, 2 – 9. III. 2015, A. Gumovsky leg., 1 ♀ (SIZK). GAMBIA: Bakau, Cape St. Mary, 5. XI. 1977 (ZIL). KENYA: Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 24. I – 7. II. 2012, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Marsabit Forest, 2.32031 ° N 37.98595 ° E, 1380 m, 16 – 30. XI. 2015, MT, near campsite, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Western Prov., Kakamega District, Kakamega Forest, 19. V – 4. VI. 2000, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MOLC). MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 28. III – 8. IV. 2003, MT in transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 20, 2 ♂♂ (CAS, MOLC); Toliara, Zom- bitse National Park, 19.8 km 84 ° E Sakaraha, 22 ° 50.36 ’ S 44 ° 42.36 ’ E, 770 m, 5 – 9. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7506, 1 ♂ (CAS). SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 6. X. 2010 – 18. I. 2011, MT, Amathole Mistbelt Grassland, S. van Noort leg., WTB 09 - GRA 1 - M 05, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD12A37FF3ED497FDF9FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD12A37FF3ED497FDF9FCBC.taxon	distribution	Distribution. Cameroon, Central African Republic, Democratic Republic of the Congo, Gambia, Ivory Coast, Kenya, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02A37FF3ED5C3FE1FF9AE.taxon	description	(Fig. 49 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02A37FF3ED5C3FE1FF9AE.taxon	description	Description. ♂. Fully winged; body length 2.2 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa basally black and femora partly black. Antenna filiform; antennomeres in following proportions: 9: 5: 5: 5.5: 5: 5: 5: 5: 5: 8. Head dull, granulate and punctate; frontal line present; occipital carina complete; POL = 7; OL = 5; OOL = 5; OPL = 4; TL = 4. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures, granulate near anterior margin. Notauli incomplete, reaching about 0.65 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels, with lateral areas reticulate rugose and median area shiny, smooth and sculptured by some keels near margins. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6.5). Paramere (Fig. 49 A) with distal inner process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02A37FF3ED5C3FE1FF9AE.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Stellenbosch, Jonkershoek (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02A37FF3ED5C3FE1FF9AE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02A37FF3ED5C3FE1FF9AE.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	description	(Figs 49 B, 50 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	description	Description. ♀. Fully winged; body length 1.9 – 2.4 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown. Antenna clavate; antennomeres in following proportions: 10: 5: 4: 3.5: 4: 4: 4: 4: 4: 6. Head convex, dull, granulate and slightly and irregularly rugose; frontal line complete; frons without lateral keels; occipital carina complete; POL = 7; OL = 4; OOL = 4.5; OPL = 3; TL = 3. Pronotum anteriorly crossed by slight transverse impression, dull, rugose; posterior surface rugose, much shorter than mesoscutum (4: 15); pronotal tubercle reaching tegula. Mesoscutum dull, completely granulate, irregularly rugose on sides and near anterior margin. In specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, mesoscutum rugose and with many irregular longitudinal keels. Notauli shortly visible near anterior margin of mesoscutum. Mesoscutellum shiny, slightly granulate. In specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, mesoscutellum unsculptured. Metanotum shiny, rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing with one or two dark transverse bands (two in holotype); distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Protarsomeres in following proportions: 7: 1.5: 1.5: 1.5: 7. Enlarged claw (Fig. 50 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 50 A) with one row of eight bristles (three bristles thicker than others); apex with one lamella. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.8 mm. Head black, except mandible testaceous; antenna brown, except proxi- mal third of scape and pedicel testaceous; mesosoma and metasoma black; legs brown, except protarsus, protibia and stalk of profemur testaceous. Antenna filiform, covered of setae longer than breadth of antennomeres; antennomeres in following proportions: 8: 5.5: 4: 5: 5: 4: 5: 5: 4.5: 7. Head shiny, slightly granulate and slightly rugose; vertex without two oblique keels from lateral ocelli to occipital carina; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL = 2; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum shiny, mostly punctate and unsculptured among punctures, with slight irregular striae on lateral regions and near anterior margin. Notauli thin, incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose; propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Paramere (Fig. 49 B) without distal inner process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	materials_examined	Material examined. Types: ♀ holotype: YEMEN: Ar Rujum, 9. IV – 5. VI. 2001, MT, A. van Harten leg. (MOLC). Paratype: same locality label as holotype, 1 ♂ (MOLC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 6 – 14. IV. 2006, MT, forest, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD). SOUTH AFRICA: KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 11. V – 24. VII. 2006, MT, G. Whiteley leg., one ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FFD02AC8FF3ED032FD89FA88.taxon	discussion	Remarks. The ♂ described above was tentatively attributed to A. felice, because it was not reared. This attribution was based mainly on the similarity of the mesoscutal sculpture in both sexes and on the same collection locality. Anteon occultum Olmi & van Harten, 2006 was also collected in the same locality, however, the ♀ of this species has a very different mesoscutal sculpture.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	description	(Fig. 50 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	description	Description. ♀. Fully winged; body length 3.0 – 3.6 mm. In typical specimens, head testaceous-reddish (occasion- ally head with small black spots around ocelli; occasionally clypeus and anterior margin of frons brown) antenna testaceous-reddish, except antennomeres 6 – 10 or 7 – 10 brown; mesosoma testaceous-reddish (occasionally ventral surface of mesosoma totally or partially black; metapectal-propodeal disc black or with brown spots; propodeal declivity almost entirely black; metanotum and part of metapectal-propodeal disc black); petiole black; metasoma testaceous-reddish (occasionally distal half of metasoma brown); legs testaceous-reddish. In A. proteicolum, head black, except mandible testaceous; head with few testaceous reddish spots situated on clypeus, around antennal toruli, on vertex behind ocellar triangle and on temple; antenna testaceous, except antennomeres 6 – 10 brown; propleuron black; pronotum black, except lateral margins and posterior tubercle testaceous-reddish; rest of mesosoma black, except meso-metapleural suture testaceous-reddish; metasoma brown; legs brown, except coxae, trochanters, articulations and part of tarsi testaceous. In specimen from Kenya, 1.09417 ° N 35.11833 ° E, head ferruginous, with frons and vertex partly darkened; mesosoma black, except part of pleuron, pronotum, mesoscutum and mesoscutellum ferruginous-dark. Specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, with head and meso- soma totally black, antenna brown, except antennomeres 1 – 4 testaceous, gaster brown, legs mostly brown. Antenna clavate. Antennomeres of holotype of A. fiorii in following proportions: 13: 6: 6.5: 5: 5: 5: 4: 5: 5: 7; antennomeres of holotype of A. copelandi in following proportions: 17: 5: 8: 7: 7: 7: 6.5: 6.5: 6.5: 9. Head dull, reticulate rugose; frontal line complete; frons with two lateral keels near orbits directed towards antennal toruli; occipital carina complete; vertex without two oblique keels connectin lateral ocelli to occipital carina (occasionally oblique keels present and incomplete). Holotype of A. fiorii with POL = 8; OL = 5; OOL = 7; OPL = 7; TL = 7. Holotype of A. proteicolum with POL = 8; OL = 4; OOL = 6; OPL = 7; TL = 8. Pronotum humped, shiny, slightly granulate, with numerous transverse keels, with anterior surface longer than posterior surface (14: 7 in holotype of A. fiorii; 11: 7 in holotype of A. proteicolum); posterior surface shorter than mesoscutum (3: 12 in holotype of A. fiorii; 7: 17 in holotype of A. proteicolum); pronotal tubercle reaching tegula. Mesoscutum shiny, usually strongly granulate, occasionally very slightly granulate and almost or completely unsculptured. Notauli incomplete, reaching approximately 0.2 – 0.3 × length of mesoscutum. Mesoscutellum very humped, shiny, unsculptured. Metanotum very short, dull, smooth, or rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity smooth, without areolae, without longitudinal keels; occasionally propodeal declivity partly slightly sculptured by transverse curvilinear striae, or slightly granulate, or slightly rugose; in specimens with metapectal-propodeal disc black, propodeal declivity strongly reticulate rugose. Forewing with two dark transverse bands; distal part of 2 rrs & Rs vein shorter than proximal part (6.5: 9 in the holotype of A. fiorii; 4: 8 in other specimens; 4.5: 9 in holotype of A. proteicolum; 7: 9 in paratype of A. proteicolum). Protarsomeres of holotype of A. fiorii in following proportions: 6: 3: 5: 9: 25. Protarsomere 2 produced into hook. Protarsomeres of holotype of A. proteicolum in following proportions: 4.5: 3.5: 4: 8: 24. Enlarged claw (Fig. 50 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 50 B) with inner side curvilinear, with distal region distinct from median and proximal regions, with basal part shorter than apical part, with two rows of 28 – 31 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	materials_examined	Material examined. Types: ♀ holotype of A. fiorii: SOUTH AFRICA: Eastern Cape, Port St. Johns, 5 – 30. IV. 1923, R. E. Turner leg. (NHMUK); ♀ holotype of A. proteicolum: SOUTH AFRICA: Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ N 18 ° 23.5 ’ E, 460 m, 14 – 23. III. 1994, Mesic Mountain Fynbos on Sandstone, Protea dominated, MT, SAM-HYM-PO 03805, S. van Noort leg. (SAMC) Paratypes of A. fiorii: SOUTH AFRICA: same locality label as holotype, 1 ♀ (NHMUK); same locality label as holotype, XII. 1923, 1 ♀ (NHMUK); same locality label as holotype, 2 ♀♀ (AMNH); KwaZulu-Natal, Durban, 1 ♀ (AMNH); Limpopo: Tzaneen, Magoeba’s Kloof, 1 ♀ (AEIC). Paratype of A. proteicolum: SOUTH AFRICA: same locality label as holotype, 10 – 17. III. 1995, Protea coronata dominated, 1 ♀ (SAMC). Other material: BOTSWA- NA: Serowe, Farmer’s Brigade, 1 ♀ (USNM); Serowe, IV. 1990, IX. 1989, 2 ♀♀ (CAS). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 2 – 5. IV. 2006, pantrap along forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD); same locality label, 10. IV. 2006, PT, 2 ♀♀ (DEUCD, MOLC); Ituri Prov., Mongbwalu AGK camp, 01 ° 56 ‘ 32.81 ‘‘ N 30 ° 02 ‘ 25.62 ‘‘ E, 21 - 28. II. 2015, 75 YPT, behind the former President Mobutu’s Doctor’s house, Alex Gumovsky leg., 1 ♀ (SIZK). ETHIOPIA: Abijata Shala Lake Na- tional Park, 1 – 10. X. 2012, YPT, Alain Pauly leg., 1 ♀ (FSAE). KENYA: Eastern Prov., Tsavo East National Park, near Athi River, 02 ° 38.51 ’ S 38 ° 21.98 ’ E, 28. XII. 1998 – 1. I. 1999, MT, R. Copeland leg., 1 ♀ (NMK); Nyanza Prov., Gwasi Hill, Ungoye side, 0.6167 ° S 34.1017 ° E, 1500 m, 26. VIII – 9. IX. 2005, MT, near hilltop next to indigenous forest, R. Copeland leg., 2 ♀♀ (NMK); same locality label, 3 – 17. XI. 2005, 1 ♀ (NMK); Rift Valley Prov., Saiwa Swamp National Park, near campsite, 1.09417 ° N 35.11833 ° E, 1882 m, 26. II – 12. III. 2006, MT, next to permanent upland swamp, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Nguruman, 25 km W of Magadi, 01 ° 50.248 ’ S 36 ° 05.215 ’ E, 1670 m, 23. VII. 2008, sweep, savanna woodland, S. van Noort leg., 1 ♀ (SAMC); Rift Valley Prov., near Lake Naivasha, 0.62884 ° S 36.42969 ° E, 1909 m, 19. VII – 2. VIII. 2015, MT, in understory of yellow Acacia patch, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: Gauteng, Pretoria, garden, 8 – 11. II. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Hluhluwe Groundcamp, 1 ♀ (PMA); KwaZulu-Natal, Cedara (SANC); KwaZu- lu-Natal, Pietermaritzburg, Ferncliff Nature Reserve, 29 ° 33.2 ’ S 30 ° 20.6 ’ E, 850 m, 6 – 8. IV. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Pietermaritzburg, Cumberland Nature Reserve, 29 ° 30.8 ’ S 30 ° 30.3 ’ E, 640 m, 21 – 22. II. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 24. XII. 2003 – 14. I. 2004, 19 – 21. IV. 2005, YPT, V. Kolyada leg., 3 ♀♀ (NMSA); KwaZulu-Natal, Louwsberg, Sanyati Farm, 1090 m, 27 ° 34 ’ S 31 ° 17.9 ’ E, 30. X – 18. XII. 2005, MT, M. Mostovski leg., 2 ♀♀ (1 in NMSA, 1 in MOLC); same locality label, 1 – 24. VI. 2006, 1 ♀ (NMSA); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 8. XII. 2005 – 6. I. 2006, MT, G. Whiteley leg., 1 ♀ (DJBC); same locality label, 29. X – 9. XI. 2005, 1 ♀ (MOLC); Limpopo, 15 km NW Klaserie, Guernsey Farm, 1 ♀ (PMA); Limpopo, Klaserie, 1 ♀ (PMA); Limpopo, Rustenburg Nature Reserve, 1 ♀ (SANC); Mpumalanga, Pilgrims Rest, 1400 m, 1 ♀ (PMA); Mpumalanga, Graskop, 1500 m, 1 ♀ (AMNH, PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	distribution	Distribution. Botswana, Democratic Republic of the Congo, Ethiopia, Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2F2ACAFF3ED32FFEDEFD48.taxon	discussion	Remarks. Anteon fiorii was separated from A. proteicolum based only on the different colour of the ♀ (the ♂ is unknown): mostly testaceous-reddish in A. fiorii; mostly black in A. proteicolum (Olmi 2006). However, subsequently a study of many specimens from South Africa showed that the colour is very variable, suggesting that keeping the two species separate is unsustainable. For this reason, Anteon fiorii is here considered the senior synonym of A. proteicolum. Anteon fiorii is also very similar to A. kivuanum (Benoit). The only difference between A. fiorii and A. kivuanum is also in respect to the colour of the ♀: mostly black in kivuanum, mostly testaceous-reddish in fiorii. The only known ♂ is that of A. kivuanum. Both species were collected in South Africa (Ashburton) during an temporally extensive Malaise trap survey. The chela of the two species is similar and hence the probability that the two species are conspecific is very high. However, in this case the ♂ of only one of the two species is known, and hence we prefer to wait for the discovery of the ♂ of A. fiorii before reaching a decision on species delimitation of these two taxa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2D2ACBFF3ED56FFD16FA88.taxon	description	(Fig. 49 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2D2ACBFF3ED56FFD16FA88.taxon	description	Description. ♂. Fully winged; body length 1.4 – 1.8 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; pro- and mesoleg testaceous; metaleg brown in holotype, testaceous in two further specimens from Madagascar, 19 ° 08.31 ’ S 44 ° 49.41 ’ E. Antenna filiform; antennomeres in following proportions: 8.5: 5: 5: 5: 5: 5.5: 5: 5.5: 5: 7. Head shiny, punctate, unsculptured among punctures; vertex of head (ocellar triangle, temple and region behind lateral ocelli) dull and rugose; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 1.5; TL = 3. Mesoscutum, mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.25 – 0.50 × length of mesoscutum (0.25 × in the holotype; 0.5 × in two further specimens from Madagascar, 19 ° 08.31 ’ S 44 ° 49.41 ’ E). In specimen from South Africa, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, notauli very short, hardly visible near anterior margin of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. In specimen from South Africa, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, propodeal declivity with left longitudinal keel; right longitudinal keel absent. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6.5). Paramere (Fig. 49 C) without distal inner pointed process; tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2D2ACBFF3ED56FFD16FA88.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape Prov., Table Mnt., Orange Kloof Nat. Reserve, 34 ° 00 ’ S 18 ° 23.7 ’ E, 125 m, 18. XII. 1997, B. L. Fisher leg., 1546, sifted litter (leaf mold, rotted wood), montane rainforest (originally in CAS, now in SAMC). Other material: MADAGASCAR: Antsiranana, Ambato Forest, 26.6 km 33 ° NE Ambanja, 13 ° 27.52 ’ S 48 ° 33.06 ’ E, 150 m, 8. XII. 2004, MT, rainforest, B. L. Fisher leg., BLF 11518, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 1 ♂ (CAS); Fian- arantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 17, 1 ♂ (CAS); same locality label, 28. XI – 6. XII. 2001, MA- 02 - 09 C- 05, 2 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 19, 2 ♂♂ (1 in CAS, 1 in MOLC); Mahajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 2 ♂♂ (CAS, MOLC); Mahajanga, Ambatofolaka, Namoroka, 53 km from Soalala, 3 km N Vilanan- dro village, 16 ° 28 ’ 04 ’’ S 45 ° 23 ’ 48 ’’ E, 400 ft., 9 – 18. XII. 2007, dense dry forest, MT, M. Irwin, R. Harin’Hala leg., MG- 40 A- 11, 1 ♂ (CAS); Mahajanga, Namoroka village, Befatika Andranovory, 16 ° 28 ’ 04 ’’ S 45 ° 23 ’ 48 ’’ E, 400 ft., 30. XI – 9. XII. 2007, dense dry forest, MT, M. Irwin, R. Harin’Hala leg., MG- 40 B- 11, 1 ♂ (CAS); same locality label, 9 – 16. XI. 2007, MG- 40 B- 08, 1 ♂ (CAS); Toamasina, Zahamena National Park, Onibe River, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, 780 m, 21 – 23. II. 2009, rainforest, MT, BLF 22168, B. L. Fisher et al. leg., 1 ♂ (CAS); Toliara, Tsimanampetsotsa Nat. Park, Bemanateza Forest, 20.7 km 81 ° E Efoetse, 23 km 131 ° SE Beheloka, 23 ° 59.32 ’ S 43 ° 52.50 ’ E, 90 m, 22 – 26. III. 2002, MT, in spiny forest thicket, Fisher, Griswold et al. leg., BLF 6257, 1 ♂ (CAS); Toliara, Zombitse National Park, 19.8 km 84 ° E Sakaraha, 22 ° 50.36 ’ S 44 ° 42.36 ’ E, 770 m, 5 – 9. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7506, 2 ♂♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 16 – 27. XII. 2002, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 10, 1 ♂ (CAS). SOUTH AFRICA: KwaZulu-Natal, Karkloof, 1325 m, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, 25. VII – 25. IX. 2005, MT, M. Mostovski leg., 1 ♂ (NMSA); Royal Natal National Reserve, Mahai Camp, 28 ° 41.27 ’ S 28 ° 56.86 ’ E, 1450 m, 25. XI – 12. XII. 2005, MT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Cathedral Peak Natural Reserve, Rain- bow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 14. XII. 2005 – 29. I. 2006, MT, M. Mostovski leg., 3 ♂♂ (NMSA); same locality label, 18. XI. 2006 – 26. II. 2007, 5 ♂♂ (NMSA); Western Cape Prov., Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 11. VIII – 6. IX. 1997, S. van Noort leg., 1 ♂ (SAMC); same locality label, IX – XII. 1997, MT, S. van Noort leg., 10 ♂♂ (SAMC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 11 ♂♂ (SAMC); Western Cape Prov., Cape Town, Constantiaberg, Tokai Forest, pine plant, X – XI. 1995, MT, S. van Noort leg., 1 ♂ (SAMC); Western Cape Prov., Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 5. XI. 2009 – 27. II. 2010, MT, Afromontane Forest, S. van Noort leg., 1 ♂ (SAMC); same locality label, 26. X. 2010 – 26. I. 2011, GVB 10 - FOR 1 - M 07, 5 ♂♂ (SAMC); same locality label, 11. XII. 2012 – 26. I. 2013, GVB 10 - FOR 1 - M 14, 1 ♂ (SAMC); Western Cape Prov., Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. X – 16. XI. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 6 ♂♂ (SAMC); same locality label, 16. XI – 16. XII. 1999, 2 ♂♂ (SAMC); same locality label, 16. II – 16. III. 1999, 1 ♂ (SAMC); same locality label, 16. XI – 16. XII. 1998, 2 ♂♂ (SAMC); Western Cape Prov., Koe- berg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 20. II – 20. III. 1998, MT, West Coast Strandveld, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, MT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - M 05, 1 ♂ (SAMC). UGANDA: Western Region, Kabarole District, Kibale Forest, Kanywara (RNHL); Western Region, Kabarole District, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 34.390 ’ S 30 ° 21.658 ’ E, 1587 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, near stream, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2D2ACBFF3ED56FFD16FA88.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2D2ACBFF3ED56FFD16FA88.taxon	distribution	Distribution. Madagascar, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2C2ACCFF3ED32FFE5BFE28.taxon	description	(Fig. 50 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2C2ACCFF3ED32FFE5BFE28.taxon	description	Description. ♀. Fully winged; body length 2.6 mm. Head black, except mandible testaceous; clypeus almost to- tally testaceous; antenna yellow; mesosoma black; metasoma brown; legs testaceous, except metafemur slightly darkened. Antenna clavate; antennomeres in following proportions: 11: 5: 7: 5.5: 5: 5: 5: 5: 5: 6.5. Head flat, setose, dull, granulate; frons almost completely slightly reticulate rugose; frontal line incomplete, not present in anterior third of frons; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3.5; OOL = 3.5; OPL = 3.5; TL = 5. Pronotum anteriorly crossed by slight transverse impression, shiny, unsculptured, except few slight irregular keels; posterior surface slightly shorter than mesoscutum (13: 16); pronotal tubercle reaching tegula. Mesoscutum shiny, slightly granulate, with many irregular striae and keels. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without complete longitudinal keels, with areolae approximately as large as those of metapectalpropodeal disc. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 8). Protarsomeres in following proportions: 7: 2: 3: 12: 19. Enlarged claw (Fig. 50 C) with proximal promi- nence bearing one long bristle. Protarsomere 5 (Fig. 50 C) with basal part much shorter than apical part, with two rows of 34 lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2C2ACCFF3ED32FFE5BFE28.taxon	materials_examined	Material examined. Type: ♀ holotype: GABON: Ogoové-Maritime Prov., Moukalaba-Dougoua Reserve, 12.2 km 305 ° NW Doussala, 02 ° 17.00 ’ S 10 ° 29.83 ’ E, 110 m, 2. III. 2000, Sweep, GA 00 - S 41, coastal lowland rainforest, elephant path, S. van Noort leg., SAM-HYM-P 023732 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2C2ACCFF3ED32FFE5BFE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2C2ACCFF3ED32FFE5BFE28.taxon	distribution	Distribution. Gabon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED44FFC6AFA8E.taxon	description	(Figs 49 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED44FFC6AFA8E.taxon	description	Description. ♂. Fully winged; body length 1.6 – 1.7 mm. Black, except mandible testaceous; antenna brown, ex- cept scape and pedicel testaceous; legs testaceous, except clubs of femora, meso- and metatibia, metacoxa partly brown. Antenna filiform; antennomeres in following proportions: 7: 4: 3: 3.5: 3: 3.5: 3.5: 3.5: 3.5: 6. Head dull, completely granulate; frontal line present (no frontal line in specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E); frons without lateral keels; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 1.5; TL = 3. Mesoscutum shiny, unsculptured. Notauli incomplete, reaching about 0.3 × length of mesoscutum (0.5 × in specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc completely reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (1: 4). Paramere (Figs 49 D, E) with distal inner process, without proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED44FFC6AFA8E.taxon	materials_examined	Material examined. Type: ♂ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Garamba Nation- al Park (MRAC). Other material: MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 24. III – 4. IV. 2004, M. Irwin, F. Parker & R. Harin‘Hala, MT, spiny forest, MA- 02 - 22 - 01, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED44FFC6AFA8E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED44FFC6AFA8E.taxon	distribution	Distribution. Democratic Republic of the Congo, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED312FE1FF87F.taxon	description	(Fig. 49 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED312FE1FF87F.taxon	description	Description. ♂. Fully winged; body length 2.2 mm. Head black, except mandible testaceous; antenna brown; me- sosoma and metasoma black; legs brown, except tarsi testaceous. Antenna filiform; antennomeres in following proportions: 7: 4: 5: 5: 5: 5: 5: 5: 5: 6. Head dull, granulate; frontal line absent; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 3; TL = 2. Mesoscutum shiny, unsculptured. Notauli incomplete, reaching about 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels, lateral areas reticulate rugose and median area centrally smooth and laterally rugose. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Paramere (Fig. 49 F) without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED312FE1FF87F.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape Prov., George, 15 – 17. XI. 1921, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED312FE1FF87F.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF2B2ACCFF3ED312FE1FF87F.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED6D7FE1FFC25.taxon	description	(Fig. 50 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED6D7FE1FFC25.taxon	description	Description. ♀. Fully winged; body length 3.1 mm. Head black, except mandible testaceous; antenna black, except antennomeres 1 – 4 testaceous; mesosoma and metasoma black; legs testaceous, except coxae and clubs of femora darkened. Antenna clavate; antennomeres in following proportions: 12: 6: 6: 4.5: 5: 5: 5: 5: 5: 7. Head completely reticulate rugose; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 4. Pronotum rugose, with posterior surface less than 0.5 × as long as mesoscutum (7: 15). Mesoscutum shiny, smooth, punctate, unsculptured among punctures. Notauli absent. Mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Mesopleuron and metapleuron reticulate rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, less than 0.5 × as long as mesoscutellum + metanotum (6: 15); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with one dark transverse band; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 11: 2: 2.5: 3: 9; protarsomere 4 less than 0.5 × as long as basal part of 5. Enlarged claw (Fig. 50 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 50 D) with some bristles and two distal lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED6D7FE1FFC25.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape Prov., Port St. Johns, IX. 1923, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED6D7FE1FFC25.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED6D7FE1FFC25.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED5BAFE27F97A.taxon	description	(Figs 48 A – D, 51 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED5BAFE27F97A.taxon	description	Description. ♂ (Figs 48 A – D). Fully winged; body length 1.7 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 8: 5: 7: 7: 7: 7: 7: 7: 7: 8. Head granulate; frontal line complete; frons without lateral keels along orbits directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 3; TL = 4; greatest breadth of posterior ocellus slightly shorter than OPL (2: 3). Mesoscutum dull, granulate and slightly reticulate rugose. Notauli very short, hardly visible near anterior margin of mesoscutum. Mesoscutellum shiny, slightly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc completely reticulate rugose, with strong; propodeal declivity without longitudinal keels, with areolae about as large as those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 6). Paramere (Fig. 51 A) with small distal inner rounded process; proximal membranous process without mosaic sculpture and sensorial setae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED5BAFE27F97A.taxon	materials_examined	Material examined. Type: ♂ holotype: UGANDA: Western Region, Kabarole District, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 33.996 ’ N 30 ° 21.262 ’ E, 1495 m, 12 – 18. VIII. 2005, MT, secondary mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 023258 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED5BAFE27F97A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292ACEFF3ED5BAFE27F97A.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292AC1FF3ED101FE14FE0C.taxon	description	(Figs 47 E, 51 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292AC1FF3ED101FE14FE0C.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, BLF 2375, Fisher, Griswold et al. leg. (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 14 – 30. V. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 11, 1 ♂ (CAS); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 28. VIII – 11. IX. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 29, 1 ♂ (MOLC); Antsiranana, Marojejy National Park, Manantenina river, 28.0 km 38 ° NE Andapa, 8.2 km 333 ° NNW Manantenina, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, MT, rainforest, B. L. Fisher et al. leg., BLF 8723, 3 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 17, 1 ♂ (MOLC); same locality la- bel, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (MOLC); Toamasina, Tamatave, 5.3 km SSE Ambanizana, Andranobe Field Station, 15 ° 40 ’ S 49 ° 58 ’ E, 425 m, 20 – 27. XI. 1993, MT, B. Fisher leg., 1 ♂ (DEUCD); Toliara, Berenty Spe- cial Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 24. III – 4. IV. 2004, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 - 01, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292AC1FF3ED101FE14FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF292AC1FF3ED101FE14FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF262AC4FF3ED13EFDE5F98C.taxon	description	(Figs 50 E, 51 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF262AC4FF3ED13EFDE5F98C.taxon	materials_examined	Material examined. Types: ♀ holotype of X. gutturnium: SOUTH AFRICA: Western Cape Prov., Mossel Bay, I. 1922, R. E. Turner leg. (NHMUK); ♀ holotype of X. gutturnium rutshuricum: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Rutshuru, 1285 m, VII. 1935, Mission G. F. de Witte (MRAC). Paratypes of X. gutturnium: SOUTH AFRICA: same locality label as holotype, XII. 1921, 1 ♀ (MRAC); Eastern Cape Prov., Port St. Johns, 15 – 31. V. 1923, R. E. Turner leg., 1 ♀ (NHMUK); Eastern Cape Prov., Katberg, 4000 ft., XII. 1932, R. E. Turner leg., ♂ allotype (NHMUK). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Reserve Special de Forêt dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 17. V. 2001, sweep, lowland rainforest, S. van Noort leg., 3 ♂♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo, Ikonongi Cave, 2 ♀♀ (MRAC, MOLC). KENYA: Coast Prov., Diani Beach, VIII. 1951, N. L. H. Krauss leg., 1 ♀ (NHMUK); Coast Prov., Kaya Ki- nondo, 4.39382 ° S 39.54567 ° E, 10 m, 27. XI – 11. XII. 2011, MT, coral rag canopy forest, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Ngangao Forest, 3.36100 ° S 38.34186 ° E, 1848 m, 10 – 24. I. 2012, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 21. II – 7. III. 2012, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mu- haka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 27. XII. 2015 – 9. I. 2016, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK). MADAGASCAR: Antsiranana, Marojejy National Park, Manantenina river, 28.0 km 38 ° NE Andapa, 8.2 km 333 ° NNW Manantenina, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, MT, rainforest, B. L. Fisher et al. leg., BLF 8723, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 5 – 21. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 10, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 14 – 21. I. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 12, 1 ♂ (CAS); same locality label, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 1 ♂ (CAS); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 1 ♂ (MOLC); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 1 ♂ (CAS); Fianarantsoa, Rano- mafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 6 ♂♂ (5 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 6 – 15. XII. 2001, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 06, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 25. VII. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 35, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 4 ♂♂ (2 in CAS, 2 in MOLC); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 21 – 24. XII. 2001, MT near river, R. Harin’Hala leg., MA- 02 - 09 D- 08, 1 ♂ (CAS); Toliara, Fiherenana, 23 ° 14.07 ’ S 43 ° 52.15 ’ E, 50 m, 1 – 4. XII. 2002, MT. degraded gallery forest, MGF 049, Frontier Wilderness Project, 1 ♂ (CAS); Toliara, Zombitse National Park, 19.8 km 84 ° E Sakaraha, 22 ° 50.36 ’ S 44 ° 42.36 ’ E, 770 m, 5 – 9. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7506, 1 ♂ (CAS). SENEGAL: Casamance, Fafakourou, 4. XI. 1992 (ZIL). SOUTH AFRICA: Eastern Cape, Katberg, 4000 ft., 15 – 30. I. 1933, R. E. Turner leg., 1 ♂ (NHMUK); Eastern Cape, Port Elizabeth, Settlers Park, 1 ♂ (PMA); Eastern Cape, Grahamstown, 1 ♀ (AEIC); Eastern Cape, Kenton-on-Sea, 2 ♂♂ (AEIC, AMNH); Eastern Cape, 8 km N of Fort Beaufort, Blinkwater, 32 ° 43 ’ S 26 ° 35 ’ E, 750 m, 1 ♀ (ZIL); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 22. II – 9. IV. 2010, MT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♂ (SAMC); Free State, Harrismith, SW foot of Platberg, 28 ° 15 ’ S 29 ° 09 ’ E, 2000 m, 1 ♀ (NHMUK); Kwa- Zulu-Natal, Hluhluwe Game Reserve, 1 ♂ (AEIC, MOLC); KwaZulu-Natal Royal Natal Nat. Reserve, Gudu Forest, 28 ° 40.90 ’ S 28 ° 55.78 ’ E, 1680 – 1730 m, 18. XI. 2006 – 27. II. 2007, MT, M. Mostovski leg., 1 ♂ (NMSA); Mpuma- langa, Pilgrims Rest, 1400 m, 1 ♂ (PMA); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 29. XI – 26. XII. 1997, YPT, S. van Noort & B. Fisher leg., 1 ♂ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. X – 16. XI. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 26. XI – 10. XII. 1995, MT, Acacia / Commiphora / Combretum bushland, SAM-HYM-PO 14827, S. van Noort leg., 1 ♂ (SAMC). UGANDA: Western Region, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 33.836 ’ N 30 ° 21.700 ’ E, 1523 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., 3 ♂♂ (SAMC); same locality label, 0 ° 33.891 ’ N 30 ° 21.468 ’ E, 1506 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., 2 ♂♂ (SAMC); Western Region, Bushenyi District, Bushenyi, III. 1939, H. C. Taylor leg., 1 ♀ (NHMUK); Central Region, Masaka District, Kawanda, 2 ♀♀ (NHMUK, AMNH); Central Region, Kampala District, Kampala, V. 1938, H. C. Taylor leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF262AC4FF3ED13EFDE5F98C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF262AC4FF3ED13EFDE5F98C.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Kenya, Madagascar, Senegal, South Africa, Tanzania, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF232AC5FF3ED013FE24FC90.taxon	description	(Fig. 50 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF232AC5FF3ED013FE24FC90.taxon	description	Description. ♀. Fully winged; body length 4.1 mm. Head black, except mandible, clypeus and anterior region of frons near clypeus and along inner margin of eyes testaceous; antenna testaceous; propleuron brown, except posterior half and part of lateral regions testaceous; pronotum brown, except anterior margin, lateral regions and posterior tubercle testaceous; rest of mesosoma black, except posterior third of mesoscutum, part of lateral regions of mesoscutum, mesoscutellum and part of metanotum ferruginous; metasoma brown-testaceous; legs testaceous. Antenna clavate; antennomeres in following proportions: 20: 7: 10: 9: 7: 7: 8: 8: 7: 10.5. Head dull, reticulate rugose; frontal line absent; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 6; TL = 5; greatest breadth of lateral ocelli about as long as OL. Pronotum shiny, strongly crossed by transverse keels, with posterior third smooth, finely punctate, unsculptured among punctures, with posterior surface shorter than mesoscutum (14: 18); pronotal tubercle reaching tegula. Mesoscutum shiny, smooth, finely punctate, unsculptured among punctures, with many large punctures situated near anterior margin. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4.5: 19). Protarsomeres in following proportions: 10: 3.5: 6: 17: 30; protarsomere 4 much longer than basal part of 5 (17: 8). Enlarged claw (Fig. 50 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 50 F) with basal part shorter than apical part (8: 22), with two rows of 22 (short) + 12 (long) lamellae; apex with five lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF232AC5FF3ED013FE24FC90.taxon	materials_examined	Material examined. Type: ♀ holotype: ZAMBIA: Central Prov., 15 km S Kapiri Mposhi, 16 – 17. I. 2003, J. Halada leg. (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF232AC5FF3ED013FE24FC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF232AC5FF3ED013FE24FC90.taxon	distribution	Distribution. Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC5FF3ED527FE14F912.taxon	description	(Figs 47 F, 51 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC5FF3ED527FE14F912.taxon	description	Description. ♂ (Fig. 47 F). Fully winged; body length 2.1 – 2.2 mm. Head black, except mandible testaceous; an- tenna testaceous-brown; mesosoma black; metasoma brown; legs testaceous, except club of metafemur darkened. Antenna filiform; antennomeres in following proportions: 9: 4.5: 5: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 8. Head dull, completely granulate and reticulate rugose; frontal line complete; frons without two lateral keels directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 2.5; TL = 2; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, completely strongly granulate and reticulate rugose. Notauli absent. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 9). Paramere (Fig. 51 D) without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC5FF3ED527FE14F912.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohipa- rara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04 (CAS). Paratypes: MADAGASCAR: Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 11, 3 ♂♂ (2 in CAS, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC5FF3ED527FE14F912.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC5FF3ED527FE14F912.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC6FF3ED0A6FDA6FD2C.taxon	description	(Fig. 51 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC6FF3ED0A6FDA6FD2C.taxon	description	Description. ♂. Fully winged; body length 2.5 – 2.8 mm. Head black, except mandible testaceous (with teeth brown) and clypeus partly brown (occasionally totally black); antenna of holotype brown, except antennomeres 1 – 3 testaceous; mesosoma and metasoma black; legs of holotype testaceous, except coxae black and clubs of femora partly brown. Antenna filiform; antennomeres in following proportions: 6.5: 4.5: 9: 8: 8: 8: 7.5: 7.5: 7: 9. Head shiny, strongly punctate, unsculptured among punctures; frontal line absent; frons with short and narrow median furrow in front of anterior ocellus; occipital carina complete; POL = 5; OL = 2.5; OOL = 7; OPL = 5; TL = 8. Mesoscutum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc without transverse posterior keel, reticulate rugose; propodeal declivity partly granulate and reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Paramere (Fig. 51 E) approximately as long as penis, without distal inner process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC6FF3ED0A6FDA6FD2C.taxon	materials_examined	Material examined. Types: ♂ holotype: CAPE VERDE ISLANDS: Fogo Island, São Jorge dos Orgãos (AMNH). Paratypes: same locality label as holotype, 12 – 13. IX. 1987, A. van Harten leg., 2 ♂♂ (AMNH); same locality label as holotype, IX. 1988, suction trap, 1 ♂ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC6FF3ED0A6FDA6FD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF222AC6FF3ED0A6FDA6FD2C.taxon	distribution	Distribution. Cape Verde Islands.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	description	(Figs 51 F, 52)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	description	Diagnosis. ♂ of Anteon with head and mesoscutum granulate; propodeal declivity with two longitudinal keels, lateral areas reticulate rugose and median area shiny and granulate.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	description	Description. ♂ (Fig. 52). Fully winged; body length 1.6 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma and legs brown. Antenna filiform; antennomeres in following proportions: 4: 3.5: 4: 5: 6: 7: 6: 6: 6: 8. Head dull, granulate; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 3; OPL = 2; TL = 3; greatest breadth of posterior ocellus about as long as OPL. Mesoscutum, mesoscutellum and metanotum dull, granulate. Notauli absent. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels, lateral areas reticulate rugose and median area shiny and granulate. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Paramere (Fig. 51 F) much shorter than penis, without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 22. II – 9. IV. 2010, MT, WTB 09 - GRA 1 - M 02, Amathole Mistbelt Grassland, S. van Noort leg., SAM-HYM-P 038921 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF212AC7FF3ED4B3FB82FD48.taxon	etymology	Etymology. The species is named hoekense after the collection site in Hoek Farm.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF202AD8FF3ED56FFE14FF34.taxon	description	(Fig. 53 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF202AD8FF3ED56FFE14FF34.taxon	description	Description. ♀. Fully winged; body length 2.2 – 3.0 mm. Head black, except mandible testaceous; antenna tes- taceous; mesosoma black, except tegula testaceous; metasoma brown; legs yellow, occasionally with metacoxa basally brown. Antenna clavate; antennomeres in following proportions: 9: 4: 3: 3: 3: 3: 3: 3: 3.5: 6.5. Head of holotype missing; head of specimen from Madagascar, 21 ° 25.041 ’ S 47 ° 41.945 ’ E, convex, shiny, punctate and sculptured by numerous irregular striae; head of specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, strongly rugose; frontal line complete; occipital carina complete; POL = 5; OL = 4; OOL = 4; OPL = 3; TL = 3. Pronotum of holotype shiny, unsculptured, except transverse keels on anterior surface; pronotum of specimen from Madagascar, 21 ° 25.041 ’ S 47 ° 41.945 ’ E, dull, rugose, except posterior half of posterior surface unsculptured; pronotal tubercle reaching tegula; posterior surface of pronotum much shorter than mesoscutum (4: 12). Mesoscutum, mesoscutellum and metanotum shiny, unsculptured in holotype, finely punctate and unsculptured among punctures in specimen from Madagascar, 21 ° 25.041 ’ S 47 ° 41.945 ’ E. Notauli incomplete, reaching approximately 0.4 – 0.5 × length of mesoscutum. Mesopleuron with anterior surface rugose and posterior surface smooth and unsculptured. Metapleuron rugose. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity with two complete longitudinal keels; median area mostly smooth, with some keels around margins; lateral areas rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7 in holotype; 2: 9 in specimen from Madagascar, 21 ° 25.041 ’ S 47 ° 41.945 ’ E). Protarsomeres in following proportions: 5: 1.5: 1.5: 2: 7.5; protarsomere 4 produced into hook. Enlarged claw (Fig. 53 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 A) with two rows of five bristles, in addition to five very narrow lamellae; apex with 4 – 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF202AD8FF3ED56FFE14FF34.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, X. 1936, A. Seyrig leg. (MNHN). Paratype: same locality label as holotype, 1 ♀ (MRAC). Other material: MADAGASCAR: Fianarantsoa, Ranomafana Nat. Park, Talatakely area, 21 ° 25.041 ’ S 47 ° 41.945 ’ E, 900 m, flight trap, mixed tropical forest, 9 – 19. I. 2001, D. H. & K. M. Kavanaugh, R. L. Brett, E. Elsom & F. Vargas leg., 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 15 – 21. XII. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 07, 1 ♀ (MOLC); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatake- ly, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 28. IV – 5. V. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 27, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF202AD8FF3ED56FFE14FF34.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF202AD8FF3ED56FFE14FF34.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3F2AD9FF3ED046FCBDFD2C.taxon	description	(Fig. 53 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3F2AD9FF3ED046FCBDFD2C.taxon	description	Description. ♀. Fully winged; body length 2.5 – 3.5 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel or also flagellomere 1 testaceous; mesosoma and metasoma black; legs completely testaceous (occasionally metacoxa and metafemur partly brown; in specimen from Uganda, 00 ° 33.836 ’ N 30 ° 21.700 ’ E, meso- and metatibia and meso- and metafemur partly brown). Antenna clavate; antennomeres in following proportions: 12: 5: 7: 5: 5: 6: 6: 6: 6: 8. Head shiny, reticulate rugose; frontal line present; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 4; TL = 2. Pronotum dull, reticulate rugose; posterior surface approximately as long as anterior surface, shorter than mesoscutum (7: 15); pronotal tubercle reaching tegula. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures (in specimen from Uganda, 00 ° 33.836 ’ N 30 ° 21.700 ’ E, mesoscutum shiny, sculptured by many deep punctures, similar to areolae, unsculptured among punctures). Notauli very short, faintly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 9). Protarsomeres in following proportions: 10: 1.5: 2: 3: 8. Enlarged claw (Fig. 53 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 B) with basal part much longer than apical part (7: 1), with some bristles and four distal lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3F2AD9FF3ED046FCBDFD2C.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Mt. Hoyo, Kabambi Cave (MRAC). Paratype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Rutshuru, 1285 m, VII. 1935, G. F. de Witte leg., 1 ♀ (AMNH )). Other material: UGANDA: Western Region, Kabarole District, Kibale National Park, Kanyawara, Makerere University Biological Field Station, 00 ° 33.836 ’ N 30 ° 21.700 ’ E, 1523 m, 6. VIII. 2008, sweep, primary mid-altitude rainforest, S. Van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3F2AD9FF3ED046FCBDFD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3F2AD9FF3ED046FCBDFD2C.taxon	distribution	Distribution. Democratic Republic of the Congo, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	description	(Figs. 54 A, 55 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	description	Description. ♂ (Fig. 55 A). Fully winged; body length 1.4 – 2.2 mm. Head black, except mandible testaceous; an- tenna testaceous-brown; mesosoma black; metasoma brown; legs brown or testaceous, except coxae and clubs of femora partly brown. Antenna filiform; antennomeres in following proportions: 11: 4.5: 5: 5: 5: 5: 5.5: 6: 6: 8. Head shiny, granulate, with vertex rugose and anterior half of frons sculptured by many transverse striae and keels (in specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, head granulate, with slight evanescent striae); frontal line complete; occipital carina complete; POL = 7; OL = 3.5; OOL = 4.5; OPL = 3; TL = 3. Mesoscutum dull, usually strongly granulate, occasionally slightly granulate. Notauli incomplete, reaching approximately 0.25 – 0.40 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, finely punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels. In specimens from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 36 and MA- 02 - 09 C- 62, propodeal declivity with two longitudinal keels and median area rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 6.5). Pterostigma partly unpigmented. Paramere (Fig. 54 A) with large distal inner rounded process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, BLF 2375, Fisher, Griswold et al. leg. (CAS). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 1 ♂ (CAS); same locality label, 21 – 28. I. 2002, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 C- 15, 1 ♂ (CAS); same locality label, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 1 ♂ (MOLC); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 2 ♂♂ (CAS); same locality la- bel 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 10 ♂♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 2 ♂♂ (CAS); Fianarantsoa, Rano- mafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04, 2 ♂♂ (CAS); same locality label, 21 – 28. I. 2002, 1 ♂ (CAS); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 7 – 18. VI. 2003, 1 ♂ (CAS); same locality label, 4 – 12. III. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 B- 15, 1 ♂ (CAS); same locality label, 12 – 19. III. 2002, MA- 02 - 09 B- 20, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 6 ♂♂ (4 in CAS, 2 in MOLC); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 2 ♂♂ (CAS); same locality label, 24. V – 4. VI. 2002, MA- 02 - 09 B- 30, 2 ♂♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 1 ♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ranomafana, JIRAMA wa- ter works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. XI – 6. XII. 2001, MT, near river, R. Harin’Hala leg., MA- 02 - 09 D- 05, 1 ♂ (MOLC); same locality label, 21 – 24. XII. 2001, MA- 02 - 09 D- 08, 2 ♂♂ (CAS); Fianarantsoa, Manombo Special Reserve camp site, 32 km SSE of Farafangana, 23 ° 01.31 ’ S 47 ° 43.20 ’ E, 36 m, 13 – 26. III. 2005, MT, lowland rainfor- est, M. Irwin, R. Harin’Hala leg., MA- 28 - 15, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3E2ADAFF3ED4B3FC06F825.taxon	discussion	Remarks. Anteon inane may possibly be the male of A. mingoae Olmi.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3C2ADDFF3ED162FC4AFCD8.taxon	description	(Figs 53 C, 54 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3C2ADDFF3ED162FC4AFCD8.taxon	description	Description. ♀. Fully winged; body length 3.0 – 3.7 mm. Head black, except mandible yellow (with teeth brown); antenna completely testaceous; occasionally antennomeres 3 – 10, or 4 – 10, 6 – 10, or 8 – 10 completely brown or with dorsal surface brown; mesosoma of holotype and other specimens black, except prothorax ferruginous; in other specimens (including one paratype in NHMUK) mesosoma completely black, or with anterior margin of pronotum reddish dark; metasoma black; tegula testaceous; legs completely testaceous; occasionally clubs of mesofemur partly brown, or metacoxa, club of metafemur and metatibia brown. Antenna clavate; antennomeres in following proportions: 9: 5: 5: 4: 4: 5: 5: 5: 5: 6. Head dull, granulate and slightly or strongly reticulate rugose (in specimen from South Africa, Butterfly Ridge, frons unsculptured among areolae and keels; in specimen from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, head almost completely smooth, with very slight tracks of areolae); frontal line complete; frons without lateral keels (in specimen from South Africa, Butterfly Ridge, frons with two lateral keels near orbits directed towards antennal toruli); occipital carina complete; POL = 7; OL = 4; OOL = 5; OPL = 5; TL = 4; in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, POL = 7; OL = 5; OOL = 5; OPL = 6; TL = 3; greatest breadth of lateral ocelli shorter than OL (4: 5). Pronotum rugose, with posterior surface shiny and smooth; posterior surface shorter than mesoscutum (5: 12), approximately as long as OPL. Mesoscutum shiny, smooth, finely punctate, unsculptured among punctures (in specimen from Zimbabwe, Bvumba, mesoscutum punctate and slightly granulate among punctures; in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, mesoscutum strongly granulate). Notauli absent; occasionally incomplete and reaching about 0.25 – 0.30 × length of mesoscutum (in specimen from South Africa, Butterfly Ridge, notauli shortly and slightly visible near anterior margin of mesoscutum). Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron and metapleuron rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. Forewing with one more or less dark transverse band around 2 r-rs & Rs vein (in specimens from South Africa, Butterfly Ridge and Mahai Camp, forewing completely hyaline, without dark transverse band); distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Protarsomere 1 longer than 4 (7: 2.5); protarsomere 4 produced into hook, approximately 0.5 × as long as basal part of 5. Enlarged claw (Fig. 53 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 C) with proximal and medial bristles (in two specimens from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E and 32 ° 57.394 ’ S 18 ° 05.324 ’ E, with one slender lamella and three bristles); apex with 3 – 9 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.1 – 2.4 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown, except junctions and tarsi testaceous. Antennomeres in following proportions: 10: 6: 7: 6: 6: 6: 6: 6: 6: 7. Head shiny, reticulate rugose (in specimens from South Africa, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, head almost completely granulate, except lateral regions of frons slightly rugose); frontal line present; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 4; OOL = 6; OPL = 5; TL = 3. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two complete longitudinal keels and median area as rugose as lateral areas; occasionally longitudinal keels incomplete. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 10). Paramere (Fig. 54 B) much shorter than penis, without distal inner pointed process and without proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3C2ADDFF3ED162FC4AFCD8.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape Prov., Port St. Johns, XII. 1923, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, X. 1923, ♂ allotype (NHMUK); same locality label as holotype, IX. 1923, X. 1923, 1 – 11. VI. 1923, 15 – 31. VIII. 1923, R. E. Turner leg., 2 ♀♀, 3 ♂♂ (NHMUK); same locality label as holotype, 2 ♀♀, 2 ♂♂ (MRAC); KwaZulu-Natal, Eshowe, 6 – 31. V. 1926, R. E. Turner leg., 1 ♀ (NHMUK). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 4. V. 2001, sweep, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♀ (SAMC). SOUTH AFRICA: Eastern Cape, Asante Sana Game Re- serve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Natal Drakensberg, Royal Natal National Park, Butterfly Ridge, 400 m, on Protea nubigena, 30. III. 1991, Pajor István leg., 1 ♀ (NMSA); KwaZulu-Natal, Royal Natal Nat. Reserve, Mahai Camp, 28 ° 41.27 ’ S 28 ° 56.86 ’ E, 1450 m, 25. XI – 12. XII. 2005, MT, M. Mostovski leg., 4 ♀♀ (NMSA); KwaZulu- Natal, Eshowe, VII. 1926, R. E. Turner leg., 1 ♀ (AMNH); KwaZulu-Natal, Eshowe, garden, 21 – 23. III. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Eshowe, near Ntumeni N. Res., 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg., 2 ♀♀ (NMSA); same locality label, 23 – 24. II. 2007, YPT, 1 ♀ (NMSA); KwaZulu-Natal, Louwsberg, Sanyati Farm, 1090 m, 27 ° 34 ‘ S 31 ° 17.9 ‘ E, 1 – 24. VI. 2006, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Cathedral Peak Natural Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 14. XII. 2005 – 29. I. 2006, MT, M. Mostovski leg., 1 ♀ (NMSA); same locality label, 18. XI. 2006 – 26. II. 2007, 10 ♀♀ (NMSA); Western Cape, near Cape Town, Milnerton, I. 1926, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Stellenbosch, Jonkershoek Nature Reserve, 2 ♀♀ (AEIC, PMA); Western Cape, Koomplanskloof, 10 km S of Cit- rusdal, 32 ° 40 ’ S 19 ° 01 ’ E, 200 – 270 m, 1 ♀ (ZIL); Western Cape, Koeberg Nature Reserve, 33 ° 37.62 ’ S 18 ° 24.26 ’ E, 23. I – 20. II. 1998, YPT, S. van Noort leg., 2 ♀♀ (SAMC); Western Cape, Anyskop Farm, 5.5 km 290 ° W Langebaan- weg, 32 ° 57.394 ’ S 18 ° 05.324 ’ E, 18 – 25. IX. 2002, MT, sand plain fynbos, S. van Noort leg., 1 ♀ (SAMC); West- ern Cape, Langberg Farm, 3 km 270 ° W Langebaanweg, 32 ° 58.461 ’ S 18 ° 07.344 ’ E, 13 – 20. XI. 2002, MT, sand plain fynbos, S. van Noort leg., 1 ♀, 2 ♂♂ (SAMC); Western Cape, Langberg Farm, 3 km 280 ° W Langebaanweg, 32 ° 58.045 ’ S 18 ° 07.418 ’ E, 11 – 18. IX. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. XII. 1999 – 16. I. 2000, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC); same locality label, 16. XI – 16. XII. 1999, 2 ♂♂ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 8. VIII – 2. X. 2013, MT, Burnt Mesic Mountain Fynbos, BH 12 - FYN 3 - M 12, S. van Noort leg., 1 ♀ (SAMC); same locality label, 22. X – 27. XI. 2013, BH 12 - FYN 3 - M 14, 1 ♀ (MOLC); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 26. X. 2010 – 26. I. 2011, GVB 10 - FYN 1 - M 07, MT, South Langeberg Sandstone Fynbos, S. van Noort leg., 2 ♀♀ (SAMC, MOLC). ZIMBABWE: Mutare, Bvumba, 21 – 24. XII. 1998, M. Snizek leg., 1 ♀ (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3C2ADDFF3ED162FC4AFCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3C2ADDFF3ED162FC4AFCD8.taxon	distribution	Distribution. Central African Republic, South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADDFF3ED5FFFC30F815.taxon	description	(Fig. 54 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADDFF3ED5FFFC30F815.taxon	description	Description. ♂. Fully winged; body length 2.1 – 2.5 mm. Head black, except mandible testaceous; antenna testa- ceous; mesosoma black; metasoma and legs testaceous. Antenna clavate, very setose, with setae longer than breadth of antennomeres; antennomeres in following proportions: 10: 5: 5: 5: 5: 5: 5: 5: 5: 8. Head convex, dull, finely setose, completely reticulate rugose; frontal line complete; frons with two lateral longitudinal keels near orbits and directed towards antennal toruli; vertex with two strong oblique keels connecting lateral ocelli to occipital carina (as in many Deinodryinus); occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 3; TL = 3; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum, mesoscutellum and metanotum shiny, smooth, finely setose, finely punctate, unsculptured among punctures (in specimen from Kenya, 4.57749 ° S 39.43825 ° E, mesoscutum weakly granulate). Notauli incomplete, reaching about 0.4 × length of mesoscutum (in specimen from Kenya, 4.57749 ° S 39.43825 ° E, notauli reaching about 0.25 × length of mesoscutum). Metapectal-propodeal disc with (erroneously “ without ” in original description) strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels, with areolae about as large as those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 7.5). Paramere (Fig. 54 C) without inner process, with numerous papillae on inner side. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADDFF3ED5FFFC30F815.taxon	materials_examined	Material examined. Type: ♂ holotype: CAMEROON: South, near Ebolowa, Nkoemvon, 16 – 30. III. 1980, MT, unshaded cocoa plantation, D. Jackson leg., NHMUK (E) 2007 - 19 (NHMUK). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 30. III – 5. IV. 2006, MT, S. L. Heydon & S. E. Stevenson leg., 2 ♂♂ (DEUCD, MOLC). KENYA: Coast Prov., Funzi Island, 4.57749 ° S 39.43825 ° E, near sea level, 4 – 10. VII. 2012, MT, near Funzi workshop, ICIPE / NMK Funzi Island Expedition leg., 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADDFF3ED5FFFC30F815.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADDFF3ED5FFFC30F815.taxon	distribution	Distribution. Cameroon, Democratic Republic of the Congo, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADEFF3ED18EFC9BFB8F.taxon	description	(Fig. 53 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADEFF3ED18EFC9BFB8F.taxon	description	Description. ♀. Fully winged; body length 3.7 – 5.6 mm. Head black, except mandible testaceous-brown; antenna brown, except antennomeres 1 – 4 or 1 – 6 testaceous; mesosoma black, except pronotal tubercle and lateral margins of pronotum reddish; tegula testaceous; metasoma brown-testaceous; legs testaceous, except femora and metatibia partly or completely brown, with junctions, trochanters and tarsi testaceous. Antennomeres in following proportions: 24: 8: 12: 8: 7: 7: 8: 7: 7: 10. Head dull, granulate, sculptured by numerous irregular keels; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 12; OPL = 8; TL = 11. Pronotum dull, granulate, with numerous transverse keels on anterior surface; posterior surface shorter than mesoscutum (13: 18). Mesoscutum shiny, unsculptured or slightly alutaceous. Notauli reduced, reaching approximately 0.2 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesoscutellum convex. Metapectal-propodeal disc dull, with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (8: 13). Protarsomeres in following proportions: 7: 5: 6: 14: 39; protarsomere 2 produced into prominent hook. Enlarged claw (Fig. 53 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 D) with two rows of about 34 lamellae extending continuously to apex; apex slightly prominent. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADEFF3ED18EFC9BFB8F.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Kasai, 1921, J. Ghes- quière leg. (MRAC). Other material: CAMEROON: Victoria, 16. I. 1959, at light, 1 ♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADEFF3ED18EFC9BFB8F.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF3A2ADEFF3ED18EFC9BFB8F.taxon	distribution	Distribution. Cameroon, Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	description	(Figs 53 E, 54 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	description	Description. ♀. Fully winged; body length 1.5 – 2.0 mm. Head black, except mandible testaceous; antenna brown, some × completely testaceous; mesosoma and metasoma black; legs brown, some × completely testaceous. Antenna clavate; antennomeres in following proportions: 6: 3.5: 2: 2: 2: 2: 2: 2: 2.5: 5. Head dull, completely granulate; frontal line present or absent; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 4; OL = 3; OOL = 5; OPL = 3; TL = 5. Pronotum shiny, with anterior surface granulate and posterior surface unsculptured; posterior surface shorter than mesoscutum (3: 9); pronotal tubercle reaching tegula. Mesoscutum shiny, smooth, finely punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.3 – 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 4.5). Protarsomeres in following proportions: 4.5: 1: 1: 3: 7. Enlarged claw (Fig. 53 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 E) with one row of about 10 lamellae; apex with about 7 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.1 – 1.4 mm. Head black, except mandible testaceous; antenna brown or yellow; mesosoma and metasoma black; legs brown or yellow. Antenna filiform; antennomeres in following proportions: 6: 4: 4: 4: 3: 4: 4: 4: 5: 6. Head shiny, granulate, with more or less wide unsculptured frontal region; frontal line absent; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 4; OL = 2; OOL = 3; OPL = 3; TL = 4. Mesoscutum, mesoscutellum and metanotum shiny, smooth, slightly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.3 – 0.6 × length of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 4). Paramere (Fig. 54 D) without distal inner pointed process and proximal membranous process. In specimen from Malawi, Kasungu National Park, paramere with inner membranous band. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Central Region, Masaka District, Kawanda, V. 1943, T. H. C. Taylor leg. (NHMUK). Paratypes: same locality label as holotype, 13. VII. 1943, VII. 1943, 15. VII. 1943, 19. VII. 1943, 5 ♂♂, 1 ♀ (NHMUK); same locality label as holotype, V. 1943, T. H. C. Taylor leg., 1 ♂ (AMNH); same locality label as holotype, 4. VI. 1943, T. H. C. Taylor leg., 1 ♀ (AMNH); Eastern Region, Serere District, Serere, VII. 1943, T. H. C. Taylor leg., 10 ♀♀ (NHMUK); same locality label, VII. 1943, T. H. C. Taylor leg., 1 ♀ (AMNH). SOMALIA: Lower Shebelle, Afgooye (= Afgoi), 28. IV – 7. V. 1980, MT, M. Olmi leg., 1 ♀ (AMNH). Other material: BENIN: Gomé, 12. XI. 1993, 1 ♀ (CIRAD). BOTSWANA: Kweneng District, Kolobeng River near Kumark- wane, 25 km W of Gaborone, 1 ♀ (ZIL); Gaborone, 3. II. 1980, 1 ♂ (CNC). BURKINA FASO: Bobo-Dioulasso, 27. IX. 1979, A. Pauly leg., 2 ♂♂ (FSAE, AMNH). CAMEROON: Maroua, Djarengol, 1 ♂ (CIRAD). DEMOCRAT- IC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 6. IV. 2006, sweeping on airstrip, S. L. Heydon & S. E. Stevenson leg., 15 ♀♀ (14 in DEUCD, 1 in MOLC). GAMBIA: Bakau, 1 ♀ (KJHC); Bakau, at Tropic Bungalow, 4. XI. 1977, 1 ♀ (ZIL); Lower River Division, Upper Baddibu, Farafenni, 1 ♂ (KJHC); Outside Abuko Nature Reserve, 1 ♂ (ZIL); S-side of Maccarthy Island, 1 ♀ (ZIL). MADAGASCAR: Antsiranana, Montagne des Français, 12 ° 18.8 ’ S 49 ° 38.51 ’ E, 150 m, along forested limestone ridge, 23 – 27. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 06 - 01, 2 ♀♀ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 20 – 28. VII. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 15, M. E. Irwin & R. Harin’Hala leg., 1 ♀ (CAS). Antsiranana, Orangea, 3 km E of Ramena near fort, 12 ° 14.49 ’ S 49 ° 22.17 ’ E, 65 m, 23 – 27. I. 2001, MT, littoral forest on sand, MA- 01 - 05 - 02, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 12 – 19. III. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 20, 1 ♂ (CAS); Fianarantsoa, Rano- mafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04, 3 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’’ E, 1130 m, 27. IV – 7. V. 2003, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 59, 1 ♀ (MOLC); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 11, 4 ♂♂ (CAS); Toliara, 12 km NW of Amboasary, Berenty, 5 – 15. V. 1983, J. S. Noyes & M. C. Day leg., 2 ♂♂, 1 ♀ (AMNH); Toliara, Berenty Special Reserve, 8 km NW Amboasary, elev 35 m, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 24. III – 4. IV. 2004, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 A- 15, 3 ♀♀ (2 in CAS, 1 in MOLC); same locality label, 22 – 30. XI. 2003, 4 ♀♀ (2 in CAS, 2 in MOLC), 2 ♂♂ (CAS, MOLC); same locality label, 7 – 17. X. 2004, 4 ♀♀ (3 in CAS, 1 in MOLC), 1 ♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 1 ♀ (CAS), 7 ♂♂ (4 in CAS, 3 in MOLC); same locality label, 16 – 27. XII. 2002, 3 ♀♀ (2 in CAS, 1 in MOLC), 2 ♂♂ (CAS, MOLC); same locality label, 17 – 26. I. 2003, 1 ♂ (CAS); same locality label, 27. XII. 2002 – 7. I. 2003, 4 ♀♀ (2 in CAS, 2 in MOLC), 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 25. II – 7. III. 2003, MA- 02 - 22 - 17, 3 ♀♀ (2 in CAS, 1 in MOLC), 6 ♂♂ (3 in CAS, 3 in MOLC); same locality label, 17 – 24. III. 2003, MA- 02 - 22 - 19, 1 ♀ (CAS); same local- ity label, 5 – 15. II. 2003, MA- 02 - 22 - 15, 4 ♂♂ (CAS); same locality label, 15 – 25. II. 2003, MA- 02 - 22 - 16, 2 ♀♀, 6 ♂♂ (CAS); same locality label, 30. XI – 7. XII. 2002, MA- 02 - 22 - 07, 1 ♀ (CAS); same locality label, 30. XI – 7. XII. 2002, MA- 02 - 22 - 07, 1 ♀, 3 ♂♂ (CAS); same locality label, 17 – 26. I. 2003, MA- 02 - 22 - 13, 7 ♀♀, 4 ♂♂ (CAS); same lo- cality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 1 ♀, 1 ♂ (CAS); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 3 ♀♀, 3 ♂♂ (CAS). MALAWI: Kasungu National Park, Lifupa Lodge, 23. IX. 1993, M. von Tschirnhaus leg., 1 ♀, 1 ♂ (MTC). MOZAMBIQUE: Maputo Prov., Maputo, 1 – 18. III. 1985, MT, M. Olmi leg., 2 ♀♀ (AMNH, CNC), 1 ♂ (AMNH); Nampula Prov., Nampula, 28. V – 11. VI. 1986, MT, M. Olmi leg., 1 ♂ (AMNH); Niassa Prov., Cuamba, Catholic University Campus, C. 23. III. 2003, B. 24. III. 2003, Sf. ♀ 9. IV. 2003, Sf. ♂ 11. IV. 2003, M. Olmi leg., reared from a nymph of an unidentified species of Cicadellidae showing two thylacia, 1 ♀, 1 ♂ (MOLC); same locality label, C. 17. III. 2012, B. 18. III. 2012, Sf. ♀ 7. IV. 2012, Sf. ♀ 8. IV. 2012, M. Olmi leg., reared from a nymph of an unidentified species of Cicadellidae showing two thylacia, 2 ♀♀ (MOLC); Niassa Prov., Cuamba, Mozambique Catholic University Farm, 17. IV – 2. VI. 2006, MT, M. Olmi leg., 1 ♂ (MOLC); Niassa Prov., Cuamba, Camboja, 28. III – 4. IV. 2009, MT, 1 ♀ (MOLC). SIERRA LEONE: S. L. Peninsula, Mac Donald, 08 ° 16 ’ N 13 ° 04 ’ W, 25. XI. 1993, 1 ♀ (ZIL); Freetown, Cape Sierra Hotel area, 08 ° 31 ’ N 13 ° 17 ’ W, 1 ♂ (ZIL); S of Freetown, Crossing to Guma Dam, 08 ° 22 ’ N 13 ° 13 ’ W, 20. XI. 1993, swept along roadsides, L. Cederholm, R. Danielsson & R. Hall leg., 1 ♀ (ZIL). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 8. X. 2010 – 18. I. 2011, MT, Camdeboo Escarpment Thick-et, S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Marais Hoop Farm (25.6 km 254 ° W Kirkwood), 33 ° 32.635 ’ S 25 ° 13.678 ’ E, 14 – 16. II. 2001, MT, VB 01 - R 4 N-M 52, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 30. X. 2009 – 22. II. 2010, YPT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♂ (SAMC); KwaZulu-Natal, Otobotini, 1 ♀ (SANC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, MT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - M 05, 1 ♂ (SAMC). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, NW side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, sweep, mk 34, 24. IV. 1996, Acacia / Commiphora / Combretum bushland, SAM-HYM-PO 18169, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Mzukune River in Mbono Valley, 03 ° 58.57 ’ S 37 ° 49.34 ’ E, 27. IV. 1996, sweep, mk 37, grasses & shrubs, riverine bushland along dry river course, SAM-HYM-PO 20415, SAM-HYM-PO 20416, S. van Noort leg., 1 ♀, 1 ♂ (SAMC). TOGO: Plateaux Region, N of Kpédomé along national road No. 1 at river Yoto, 06 ° 58.58 ’ N 01 ° 10.14 ’ E, 650 m, 29. IV. 2008, M. von Tschirnhaus leg., 7 ♀♀ (MTC). YEMEN: Al Lahima, 1. I – 9. IV. 2001, MT, A. van Harten & A. M. Hager leg., 1 ♂ (MOLC); same locality label, 16. X – 31. XII. 2000, 1 ♂, 2 ♀♀ (MOLC); Lahj, 17. XII. 2001 – 31. I. 2002, MT, A. van Harten & A. Sallam leg., 2 ♀♀ (MOLC); same locality label, 1. X – 17. XII. 2001, 1 ♀ (MOLC); same locality label, IX. 2001, 3 ♀♀ (MOLC); same locality label, VII. 2001, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	biology_ecology	Hosts. In Mozambique: unidentified Cicadellidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	distribution	Distribution. Benin, Botswana, Burkina Faso, Cameroon, Democratic Republic of the Congo, Gambia, Madagascar, Malawi, Mozambique, Sierra Leone, Somalia, South Africa, Tanzania, Togo, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF392AD0FF3ED214FEFAFC2F.taxon	discussion	Remarks. The above description of the ♂ is based on specimens reared in Mozambique (Cuamba) and Uganda (Kawanda).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD0FF3ED5B7FDC4F959.taxon	description	(Fig. 54 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD0FF3ED5B7FDC4F959.taxon	description	Description. ♂. Fully winged; body length 2.0 – 2.2 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tibiae and tarsi testaceous; occasionally metatarsus brown. Antenna filiform; antennomeres in following proportions: 12: 6: 5: 5: 5.5: 6: 6: 6: 6: 9. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 6; OPL = 2.5; TL = 3. Mesoscutum shiny, with surface near anterior margin rugose; rest of surface sculptured by slight irregular striae and unsculptured among striae. Notauli reduced, slightly and shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Paramere (Fig. 54 E) much shorter than penis, without distal inner process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD0FF3ED5B7FDC4F959.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Western Prov., 27 mi. NE Kisumu, Kaimosi Mission, 1650 m, 29. XI. 1957, E. S. Ross & R. E. Leech leg. (CAS). Other material: ETHIOPIA: Shewa (= Shoa) Prov., about 130 km S Addis Abeba, Marakò, 1 ♂ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD0FF3ED5B7FDC4F959.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD0FF3ED5B7FDC4F959.taxon	distribution	Distribution. Ethiopia, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD1FF3ED17EFE27FC04.taxon	description	(Figs 48 E, F, 53 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD1FF3ED17EFE27FC04.taxon	description	Description. ♀. Fully winged (Figs 48 E, F); body length 2.1 – 3.3 mm. Head black, except mandible brown; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations, trochanters, procoxa and mesocoxa testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 7: 6: 5: 6: 6: 6: 6: 8. Head slightly convex, dull, completely reticulate rugose; frontal line complete; frons without two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 8; OL = 4; OOL = 6; OPL = 5; TL = 5; greatest breadth of lateral ocelli shorter than OPL (3: 5). Pronotum anteriorly crossed by slight transverse impression, dull, with anterior surface transversely striate; posterior surface with anterior half transversely striate and posterior half slightly granulate, not striate; posterior surface shorter than mesoscutum (7: 14); pronotal tubercle reaching tegula. Mesoscutum granulate, rugose and irregularly striate. Notauli absent. Mesoscutellum shiny, unsculptured. Metanotum rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels, with areolae about as large as those of metapectal-propodeal disc. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 8: 3: 4: 6: 16. Enlarged claw (Fig. 53 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 53 F) with basal part slightly shorter than distal part (7: 9), with one row of 16 lamellae; apex bent and with 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD1FF3ED17EFE27FC04.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Western Region, Kibale National Park, Kanyawara Mak- erere University Biological Field Station, 0 ° 33.996 ’ S 30 ° 21.262 ’ E, 1495 m, 29 – 31. VII. 2005, YPT, secondary midaltitude rainforest, S. van Noort leg., SAM-HYM-A 023416 (SAMC). Paratypes: same locality label as holotype, SAM-HYM-A 023278, 1 ♀ (SAMC); same locality label as holotype, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD1FF3ED17EFE27FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF372AD1FF3ED17EFE27FC04.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	description	(Figs 54 F, 56 A, 57 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	description	Description. ♀. Fully winged; body length 2.7 – 3.9 mm. Head black, except mandible testaceous (in specimen from South Africa, 1.5 km E Mtunzini, anterior margin of clypeus testaceous); antenna black, except antennomeres 1 – 5 brown or testaceous (in specimen from South Africa, 1.5 km E Mtunzini, antenna testaceous, except antennomeres 7 – 10 darkened); mesosoma and metasoma black (in specimen from South Africa, 1.5 km E Mtunzini, metasoma testaceous); proleg brown or testaceous; meso- and metaleg brown-black, except tarsi testaceous and coxae partly testaceous; in specimen from South Africa, 1.5 km E Mtunzini, legs completely testaceous. Antenna clavate; antennomeres in following proportions: 12: 5: 6: 5: 5: 5: 5: 5: 5.5: 7. Head granulate and reticulate rugose; frontal line present; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 5; OOL = 8; OPL = 7; TL = 7. In some specimens (from Kenya, South Africa and Uganda) vertex behind lateral ocelli with two oblique keels connecting lateral ocelli to occipital carina. Pronotum dull, granulate and punctate, with strong transverse keels; posterior surface shorter than mesoscutum (8: 20). Mesoscutum shiny, very weakly granulate (strongly granulate in specimen from South Africa, 1.5 km E Mtunzini). Notauli incomplete, very reduced, shortly present near anterior margin of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum shiny, sculptured by longitudinal keels. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (4: 6). Protarsomeres in following proportions: 6: 3: 4: 8: 25. Enlarged claw (Fig. 57 A) very long, longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 A) curvilinear, with basal part shorter than apical part, with two rows of about 10 – 12 lamellae; apex of protarsomere 5 with about 10 – 17 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.2 – 3.1 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous, except part of metacoxa and part of club of metafemur brown. Immature specimens completely brown. Antennomeres in following proportions: 10: 4: 4: 4: 4: 4: 4: 4: 4: 5. Head shiny, granulate, with slight areolae; frontal line complete; in specimens from South Africa, 29 ° 34.56 ’ S 30 ° 17.40 ’ E, and Kenya, 3.30958 ° S 39.96538 ° E, head strongly granulate and reticulate rugose; frons with or without two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 7; OL = 4; OOL = 5; OPL = 2.5; TL = 3. In large specimens, vertex with two oblique keels connecting lateral ocelli to occipital carina; small specimens without above oblique keels. Mesoscutum shiny, unsculptured or slightly granulate or punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.25 – 0.40 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose (occasionally with left longitudinal keel). In two specimens from Madagascar, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, propodeal declivity with two incomplete longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Paramere (Fig. 54 F) (figure 222 in Olmi 1984 erroneous) without distal inner pointed process and with small and broad proximal membranous process (Fig. 54 F); in two specimens from South Africa, Graskop and 29 ° 34.56 ’ S 30 ° 17.40 ’ E, paramere with some transverse folds on their inner sides (Fig. 56 A); in specimens from South Africa, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, inner side of paramere with transverse folds or mosaic pattern. In specimens from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’’ E, and Kenya, 3.30958 ° S 39.96538 ° E, paramere much shorter than penis. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Rutshuru, 1285 m, G. F. De Witte Mission, VII. 1935 (MRAC). Paratype: same locality label as holotype, 13. VII. 1935, 1 ♀ (MRAC). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 4. V. 2001, sweep, lowland rainforest, marsh clearing, S. van Noort leg., 3 ♂♂ (SAMC); Sangha- Mbaéré Prefecture, Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 23. V. 2001, sweep, lowland rainforest, S. van Noort leg., 5 ♂♂ (SAMC); Sangha-Mbaéré Prefecture, Reserve Spe- ciale de Forêt dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 17. V. 2001, sweep, lowland rainforest, S. van Noort leg., 4 ♂♂ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Haut- Lomami, Upemba National Park, Kiamokoto-Kiwakishi, 2 ♀♀ (MRAC, AMNH); Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 30. III – 5. IV. 2006, MT, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (DEUCD; Ituri Prov., Mongbwalu AGK camp, 01 ° 56 ‘ 32.81 ‘‘ N 30 ° 02 ‘ 25.62 ‘‘ E, 21 - 28. II. 2015, 75 YPT, behind the former President Mobutu’s Doctor’s house, A. Gumovsky leg., 1 ♀ (SIZK); North Kivu, Virunga National Park (= Albert Na- tional Park), Rutshuru, 1285 m, G. F. De Witte Mission, 2 ♀♀ (ISNB, AMNH); North Kivu, Virunga National Park (= Albert Nat. Park), Mt. Ruwenzori, Bombi River, Kakalari River, 1 ♀ (MRAC); North Kivu, Virunga National Park (= Albert Nat. Park), Tshiaberimu, Talia N River, Mbulikerere River, 1 ♀ (MRAC); North Kivu, Virunga Na- tional Park (= Albert Nat. Park), Mt. Hoyo, 2 ♀♀ (MRAC, AMNH); North Kivu, Virunga National Park (= Albert Nat. Park), Kyandolire, Bomboka, 1 ♀ (MRAC); South Kivu, Lwiro River, 47 Km N of Bukavu, 1650 m, 1 ♀ (CAS). KENYA: Coast Prov., Arabuko-Sokoke Forest, 3.30958 ° S 39.96538 ° E, 80 m, 22. IX – 6. X. 2014, MT in bush outside forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Kaya Kinondo, 4.39382 ° S 39.54567 ° E, 10 m, 22. VIII – 5. IX. 2011, indigenous forest, MT, coral rag canopy forest, R. Copeland leg., 1 ♂ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest, Calebs Campground, 00 ° 22.175 ’ N 34 ° 53.297 ’ E, 1580 m, 2002, KE 1633, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MOLC). MADAGASCAR: Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 15. II – 6. III. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 03, M. E. Irwin & R. Harin’Hala leg., 3 ♂♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 21 – 26. IV. 2001, MT, MA- 01 - 01 D- 08, R. Harin’Hala leg., 1 ♂ (CAS); Antsiranana, Diana Re- gion, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 1 – 8. VI. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 25, 1 ♂ (CAS); Fianarantsoa, Ranomafana Na- tional Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 4 – 12. II. 2002, 1110 m, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 3 ♂♂ (CAS); same locality label, 21 – 28. I. 2002, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 14 – 22. VII. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 35, 1 ♂ (MOLC); same locality label, 2 – 10. I. 2002, MA- 02 - 09 C- 10, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana Na- tional Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 19, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 21 – 24. XII. 2001, MT, near river, R. Harin’Hala leg., MA- 02 - 09 D- 08, 1 ♂ (MOLC); Fianarantsoa, Ambalagoavy Nord Forest, Ikongo, Ambatombe, 21 ° 49.39 ’ S 47 ° 20.20 ’ E, 625 m, XI. 2000, MT, Harin’Hala R. & M. E. Irwin leg., MA- 01 - 12 - 01, 1 ♂ (CAS); Fianarantsoa, Vevembe Forest, 66.6 km 293 ° WNW Farafangana, 22 ° 47.28 ’ S 47 ° 10.55 ’ E, 600 m, 23. IV. 2006, MT, rainforest transitioning to montane forest, B. L. Fisher et al. leg., BLF 14118, 1 ♂ (CAS); Toamasina, Ankerana, 18 ° 24.37 ’ S 48 ° 49.08 ’ E, 1108 m, 22 – 27. I. 2012, MT, rainforest, B. L. Fisher et al. leg., BLF 27982, 1 ♂ (CAS); Toamasina, Zahamena National Park, Tetezambatana Forest, mear junction of Nosivola and Manakambahiny Rivers, 17 ° 44 ‘ 35 ‘‘ S 48 ° 43 ‘ 46 ‘‘ E, 860 m, 18 – 19. II. 2009, rainforest, MT, BLF 21972, B. L. Fisher et al. leg., 2 ♂♂ (CAS); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Ma- rofandilia, 20 ° 04.09 ‘ S 044 ° 39.38 ‘ E, 10 – 17. XII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 58, 1 ♂ (CAS); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.07 ’ S 044 ° 39.34 ’ E, 26. XI – 37. XII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18284 _ 56, 1 ♂ (MOLC); Toliara, Tsifota, 20 km N Manombo, 1 km E of Tsifota village, 22 ° 49.08 ’ S 43 ° 22.36 ’ E, 15 m, 29. XII. 2008 – 8. I. 2009, MT, spiny forest, M. Itwin, R. Harin’Hala leg., MG- 48 - 09, 1 ♂ (CAS). SOUTH AFRICA: Eastern Cape, Somerset East, 2 ♀♀ (NHMUK, AMNH); Port St. Johns, XI. 1923, R. E. Turner leg., 1 ♂ (NHMUK); Eastern Cape, Pearston, Plains of Camdeboo Game Re- serve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 2 ♂♂ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 9. IV – 26. VII. 2010, YPT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♂ (SAMC); same locality label, 3. IX – 30. X. 2009, 1 ♂ (SAMC); Kwazulu-Natal, Pietermaritzburg, 2 ♂♂ (AEIC, AMNH); Kwazulu-Natal, 1.5 Km E Mtunzini, Umlalazi Nature Reserve, 1 ♂ (PMA); Kwazulu-Natal, 1.5 km E Mtunzini, Umlalazi Nature Reserve, 2831 Dd, II. 1979, Indigenous forest, MT, R. M. Miller leg., 1 ♀ (PMA); Kwa- zulu-Natal, Cedara, 1 ♀ (SANC); Kwazulu-Natal, Pietermaritzburg, Hilton, garden, 6 – 14. III. 2005, YPT, V. Kolyada leg., 1 ♂ (NMSA); Kwazulu-Natal, Winterskloof, 29 ° 34.56 ‘ S 30 ° 17.40 ‘ E, 1085 m, 20. IV – 1. IX. 2007, MT, M. Mo- stovski leg., 1 ♂ (NMSA); Kwazulu-Natal, Royal Natal National Reserve, Gudu Forest, 28 ° 40.90 ‘ S 28 ° 55.78 ‘ E, 1680 – 1730 m, 13. XII. 2005 – 28. I. 2006, MT, M. Mostovski leg., 2 ♂♂ (NMSA); Kwazulu-Natal, Ashburton, 29 ° 39 ‘ 23 ‘‘ S 30 ° 27 ‘ 41 ‘‘ E, 24. XI – 8. XII. 2005, MT, G. Whiteley leg., 1 ♀ (DJBC); same locality label, 29. X – 9. XI. 2005, 1 ♂ (DJBC); same locality label, 15 – 30. III. 2006, 1 ♂ (MOLC); same locality label, 24. XI – 8. XII. 2005, 8 ♂♂ (7 in DJBC, 1 in MOLC); same locality label, 6. I – 1. II. 2006, 1 ♂ (DJBC); same locality label, 8. XII. 2005 – 6. I. 2006, 4 ♂♂ (DJBC); same locality label, 9 – 24. XI. 2005, 11 ♂♂ (DJBC); Limpopo, 30 km E Nylstroom, 1 ♀ (PMA); Mpuma- langa, E Transvaal, 11 km SE Pilgrims Rest, 1400 m, 1 ♂ (PMA; Mpumalanga, Graskop, 1500 m, 1 ♂ (PMA); Mpumalanga, Krüger National Park, Skukuza Malt., 1 ♀ (PMA); Western Cape, Constantiaberge, 34 ° 02.5 ‘ S 18 ° 23.5 ‘ E, 640 m, above road to mast overlooking Hout Bay, 14 – 23. III. 1994, Mesic Mountain Fynbos, MT, SAM- HYM-PO 03801, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ‘ N 18 ° 23.5 ‘ E, 460 m, 17 – 27. III. 1995, Site 2, Mesic Mountain Fynbos, Protea coronata dominated, MT, SAM-HYM-PO 13731, S. van Noort leg., 1 ♂ (SAMC); same locality label, 10 – 17. III. 1995, SAM-HYM-PO 13722, 1 ♂ (SAMC); same locality label, 6 – 13. XII. 1994, SAM-HYM-PO 13730, 1 ♂ (SAMC); same locality label, Protea dominated, 9 – 15. II. 1994, SAM-HYM-PO 05920, SAM-HYM-PO 03791, SAM-HYM- PO 05921, 5 ♂♂ (SAMC); same locality label, Protea dominated, 25. I – 2. II. 1994, 1 ♂ (SAMC); Western Cape, Ce- res, 2 – 21. III. 1921, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, Kirstenbosch, 1 ♂ (AEIC); Western Cape, Stel- lenbosch, Jonkershoek Reserve, 1 ♂ (PMA); Western Cape, Cape of Good Hope Nat. Reserve, Da Gama Peak, 34 ° 21 ’ S 18 ° 29 ’ E, 28 – 31. III. 1995, MT, Old Mesic Mountain Fynbos, SAM-HYM-PO 13726, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Hermanus, Fernkloof Nat. Reserve, Mosselberg, 60 m, 12. III. 1995, Mesic Mountain Fynbos, MT, SAM-HYM-PO 13726, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 11. VIII – 6. IX. 1997, S. van Noort leg., 22 ♂♂ (SAMC); same locality label, IX – XII. 1997, MT, S. van Noort leg., 7 ♂♂ (SAMC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 43 ♂♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 21. II – 21. III. 1998, MT, South Coast Strandveld, 57 m, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 18. IV – 16. V. 1998, 3 ♂♂ (SAMC); Western Cape, Die Dam, C. 23. II. 2006, B. 24. II. 2006, M. Olmi reared from a nymph of Iassomorphus drakensteini (Naudé), 1 ♀ (MOLC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 8. II – 2. IV. 2010, MT, Boland Granite Fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, 28. IX – 28. XII. 2010, GRA 09 - FYN 1 - M 05, 3 ♂♂ (SAM); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 58.888 ’ S 20 ° 48.885 ’ E, 454 m, 29. III – 23. VII. 2010, MT, South Langeberg Sandstone Fynbos, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 26. X. 2010 – 25. I. 2011, GVB 10 - FYN 1 - M 08, 2 ♂♂ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 20. III – 17. IV. 1998, MT, West Coast Strandveld, S. van Noort leg., 1 ♂ (SAMC); same locality label, 20. II – 20. III. 1998, 5 ♂♂ (SAMC); same locality label, 17. IV – 15. V. 1998, 1 ♂ (SAMC); same locality label, 3 – 31. X. 1997, 5 ♂♂ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 25. IV – 16. V. 2013, MT, Mesic Mountain Fynbos, BH 12 - FYN 3 - M 08, S. van Noort leg., 2 ♂♂ (SAMC). UGANDA: Western Region, Kibale Forest, Kanyawara, 1 ♂ (RNHL); Western Region, Kibale National Park, Kan- yawara Makerere University Biological Field Station, 0 ° 34.390 ’ S 30 ° 21.658 ’ E, 1587 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, near stream, S. van Noort leg., 1 ♀ (SAMC); same locality label, 0 ° 33.836 ’ N 30 ° 21.700 ’ E, 1523 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., 2 ♀♀, 2 ♂♂ (SAMC); same locality label, 0 ° 33.891 ’ N 30 ° 21.468 ’ E, 1506 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., 1 ♀, 1 ♂ (SAMC); same locality label, 0 ° 33.996 ’ S 30 ° 21.262 ’ E, 1495 m, 29 – 31. VII. 2005, YPT, second- ary mid-altitude rainforest, S. van Noort leg., 1 ♀, 1 ♂ (MOLC); same locality label, 0 ° 33.784 ’ S 30 ° 21.617 ’ E, 1500 m, 12 – 26. VIII. 2008, MT, primary mid-altitude rainforest, S. van Noort leg., 1 ♂ (SAMC). YEMEN: Ar Rujum, 16. X. 2000 – 15. I. 2001, 15. I – 9. IV. 2001, MT, A. van Harten, 5 ♂♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in South Africa: Iassomorphus drakensteini (Naudé).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Kenya, Madagascar, South Africa, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF362AD5FF3ED59BFB17F8E5.taxon	discussion	Remarks. Anteon fiorii Olmi is very similar to A. kivuanum (Benoit). The only difference is in respect of the colour: mostly black in kivuanum, mostly testaceous-reddish in fiorii. The only known male is that of A. kivuanum. Both species were collected in South Africa, Ashburton, by a temporally extensive survey using Malaise traps. The chelae of the two species are alike. The probability that the two species are conspecific is very high.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	description	(Figs 56 B – D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	description	Description. ♂. Fully winged; body length 1.4 – 1.7 mm. Head black (occasionally brown), except mandible testa- ceous; antenna testaceous; occasionally testaceous-darkened, except scape and pedicel testaceous; mesosoma black (occasionally brown); metasoma brown; legs testaceous; occasionally stalk of metafemur darkened. Antenna filiform; antennomeres in following proportions: 6: 3.5: 4.5: 4: 4: 4.5: 4.5: 4: 4.5: 6. Head dull, completely strongly granulate, or with frons granulate and very slightly rugose; frontal line absent; frons without lateral keels; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 2; TL = 3; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum dull, completely strongly granulate, with region strictly close to anterior margin slightly rugose. Notauli shortly present near anterior margin of mesoscutum; in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, notauli reaching about 0.25 × length of mesoscutum. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, notauli reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels; areolae of propodeal declivity less wide than those of dorsal region. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 62, propodeal declivity with two complete longitudinal keels. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, propodeal declivity with two complete longitudinal keels and median region showing central unsculptured area. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 6.5). Paramere (Figs 56 B-D) without distal inner pointed process, with more or less large proximal or subdistal inner lobe having mosaic pattern and not covered with papillae (extension of lobe variable). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Queen Elizabeth Park Reserve, 833 m, nr. stream, 1. XII. 2003, J. Londt & M. Mostovski leg. (NMSA). Paratypes: SOUTH AFRICA: KwaZulu- Natal, Royal Natal Nature Reserve, 28 ° 41.4 ’ S 28 ° 56.3 ’ E, 1420 m, 10 – 13. XII. 2004, MT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ‘ S 30 ° 20.4 ‘ E, MT, near stream, 1. XI – 2. XII. 2004, M. Mostovski leg., 1 ♂ (NMSA); same locality label, 3 – 26. II. 2005, M. Mostovski leg., 2 ♂♂ (NMSA); same local- ity label, 20. III – 25. IV. 2005, 1 ♂ (MOLC); KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 6 – 14. III. 2005, YPT, V. Kolyada leg., 1 ♂ (NMSA). Other material: KENYA: Coast Prov., Kasigau Mtn., 3.82700 ° S 38.64875 ° E, 1065 m, 19. X – 2. XI. 2011, MT, indigenous forest, next to campsite in forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 3 – 17. X. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (MOLC); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 24. X – 7. XI. 2015, MT, indigenous forest, R. Copeland leg., 3 ♂♂ (2 in NMK, 1 in MOLC); same locality label, 6 – 20. IX. 2014, 2 ♂♂ (NMK); same locality label, 16 – 26. VII. 2014, 1 ♂ (NMK); same locality label, 26. VII – 9. VIII. 2015, 1 ♂ (NMK); same locality label, 1 – 15. XI. 2014, 3 ♂♂ (NMK); same locality label, 11 – 25. X. 2015, 3 ♂♂ (NMK); same locality label, 23. VIII – 6. IX. 2015, 2 ♂♂ (NMK); same locality label, 27. XII. 2015 – 9. I. 2016, 1 ♂ (NMK); same locality label, 5 – 19. XI. 2015, 1 ♂ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 8 – 22. I. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 3 ♂♂ (2 in NMK, 1 in MOLC); Eastern Prov., Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bot- tom of forest, 24. XII. 2011 – 7. I. 2012, MT, inside indigenous forest, R. Copeland leg., 1 ♂ (NMK). MADAGASCAR: Fianarantsoa, 38 km S Ambalavao, Andringitra Reserve, 22 ° 12 ’ S 46 ° 58 ’ E, 1680 m, 22 – 29. X. 1993, B. L. Fisher leg., 1 ♂ (DEUCD); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 8 – 15. XI. 2001, MT, secondary tropical forest, MA- 02 - 09 C- 02, R. Harin’Hala leg., 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 1 ♂ (CAS); same locality label, 28. V – 6. VI. 2003, MA- 02 - 09 C- 62, 2 ♂♂ (MOLC); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 2 ♂♂ (CAS); same local- ity label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 4 ♂♂ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 28 ♂♂ (26 in CAS, 2 in MOLC); same locality label, 21. III – 12. IV. 2003, MA- 02 - 09 C- 57, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 10 ♂♂ (8 in CAS, 2 in MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 24. V – 4. VI. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 30, 3 ♂♂ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 1 ♂ (CAS); same lo- cality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 1 ♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 1 ♂ (MOLC). SOUTH AFRICA: KwaZulu-Natal, Eshowe, near Ntumeni N. Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 23. II – 20. IV. 2007, MT, Kolyada & Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, La Mercy, Site 1, 29 ° 37.41 ‘ S 31 ° 06.454 ‘ E, 85 m, 6. XII. 2006, YPT, Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Vernon Crookes Nat. Reserve, 250 m, 30 ° 17.4 ‘ S 30 ° 36.9 ‘ E, 24. I – 10. V. 2006, MT, M. Mostovski leg., 4 ♂♂ (3 in NMSA, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	distribution	Distribution. Kenya, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF312AD7FF3ED6F3FBF7FE54.taxon	discussion	Remarks. Deinodryinus sabaeus Olmi & van Harten 2006, is similar to Anteon kwazuluense, but the inner lobe of the paramere does not have a mosaic pattern (Fig. 88 E) as in A. kwazuluense.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	description	(Fig. 57 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	description	Description. ♀. Fully winged; body length 3.7 mm. Head black, except mandible testaceous (with teeth brown); an- tenna testaceous, except antennomeres 9 – 10 brown; mesosoma and metasoma black; tegula testaceous; legs brown, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 17: 6: 7: 5: 5: 6: 6: 6: 6: 11. Head shiny, granulate and reticulate rugose; frontal line present; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 5; OOL = 4; OPL = 6; TL = 4. Pronotum shiny, slightly granulate, unsculptured near the posterior margin; posterior surface very long, more than 0.5 × as long as mesoscutum (9: 16); pronotal tubercle reaching tegula. Mesoscutum shiny, slightly granulate. Notauli absent. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron shiny, unsculptured. Metapleuron with numerous irregular keels. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity granulate, with numerous slight transverse keels, with slight tracks of two longitudinal keels. Forewing with distal wide dark transverse band; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 10). Protarsomere 1 longer than 4 (7: 4); protarsomere 4 approximately 0.5 × as long as basal part of 5. Enlarged claw (Fig. 57 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 B) with basal part much longer than apical part, with 22 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Fanovana, VII. 1933, Seyrig leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AD7FF3ED46BFC6FFA5A.taxon	discussion	Remarks. According to Ceballos (1936), the type locality is Fanovari. However, this locality apparently does not exist (see Viette 1991). The correct name is probably Fanovana.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	description	(Figs 55 F, 57 C, 64)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	diagnosis	Diagnosis. ♀ of Anteon macropterous, with head and mesosoma completely testaceous (Fig. 55 F); pronotum with posterior surface longer than anterior surface, more than 0.5 × as long as mesoscutum; posterior surface of pronotum with raised carina on both anterior and lateral margins; forewing with distal part of 2 r-rs & Rs vein more than 0.5 × as long as proximal part; propodeal declivity without longitudinal keels; protarsomere 4 much longer than basal part of 5; protarsomere 5 (Fig. 57 C) with inner side curvilinear, with distinct apical region, with basal part much shorter than distal part.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	description	Description. ♀ (Figs 55 F, 64). Fully winged; body length 1.8 – 2.6 mm. Completely testaceous, except petiole black. Antenna clavate; antennomeres in following proportions: 10: 5: 5: 5: 3: 4: 4: 4: 4: 6. Head dull, granulate; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 5; OOL = 4; OPL = 4; TL = 5; greatest breadth of lateral ocelli shorter than OL (2: 5). Pronotum dull, with posterior surface granulate, surrounded by raised carina on both anterior and lateral margins (Fig. 55 F); posterior surface slightly shorter than mesoscutum (12: 13) and much longer than anterior surface (12: 4); pronotal tubercle reaching tegula. Mesoscutum dull, weakly granulate. Notauli incomplete, hardly visible, reaching approximately 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with areolae very small (almost granulate), without longitudinal keels. Forewing (Fig. 64 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (4: 5). Protarsomeres in following proportions: 5: 2: 3: 9: 18; protarsomere 4 much longer than basal part of 5 (9: 4). Enlarged claw (Fig. 57 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 C) with basal part much shorter than apical part (4: 14), with two rows of 17 + 21 lamellae; apex with four lamellae, one of which very long. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, YPT, Maputaland Coastal Belt, SA 10 - MAB-Y 01, S. van Noort leg., SAM- HYM-A 035626 (SAMC). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 14. IV. 2006, PT along forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD). SOUTH AFRICA: same locality label as holotype, 1 ♀ (MOLC); KwaZulu-Natal, Ithala Game Reserve, Thalu Research Camp, 27 ° 36.520 ’ S 31 ° 12.263 ’ E, 610 m, 10 – 14. X. 2010, YPT, Maputaland Coastal Belt, S. van Noort leg., SA 10 - ITH-Y 02, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF302AE8FF3ED060FB07FC04.taxon	etymology	Etymology. The species is named after Mabibi, the locality where the holotype was collected.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	description	(Figs 56 E, F, 57 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	description	Description. ♀. Fully winged; body length 2.5 – 3.0 mm. Head black, except mandible testaceous (with teeth brown); antenna brown, except antennomeres 1 – 6 or 1 – 4 testaceous (occasionally antenna completely brown or testaceous, except antennomere 10 darkened); mesosoma and metasoma black; tegula testaceous; legs brown, except tarsi testaceous, or testaceous, except metacoxa brown. Antenna clavate; antennomeres in following proportions: 13: 5: 4: 4: 4: 4: 5: 5: 5: 7. Head shiny, granulate and reticulate rugose; frontal line present; frons with or without lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 7; OL = 4; OOL = 5; OPL = 6; TL = 4. Pronotum shiny, granulate; posterior surface very long, more than 0.5 × as long as mesoscutum (10: 13); pronotal tubercle reaching tegula. Mesoscutum shiny, granulate. Notauli absent; occasionally notauli present and incomplete, reaching about 0.2 – 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, slightly granulate. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity granulate and with numerous irregular keels or sculptured by small areolae, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 9). Protarsomeres in following proportions: 7: 2: 3: 3: 12. Protarsomere 4 produced into hook, about 0.5 × as long as basal part of 5. Enlarged claw (Fig. 57 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 D) with two rows of 10 – 14 lamellae, among which one very long distal lamella. Tibial spurs 1 / 1 / 2. ♂ (not reared; tentatively associated to the above ♀). Fully winged; body length 1.0 – 1.9 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 3 – 10 darkened (in holotype of A. brooksi, antenna totally testaceous; in other specimens, antenna brown, except scape and pedicel testaceous); mesosoma and metasoma black; occasionally metasoma brown; legs testaceous, except coxae, clubs of femora and tibiae partly brown (in holotype of A. brooksi, pro- and mesoleg completely testaceous). Antenna filiform; antennomeres in following proportions: 6: 3.5: 3: 3: 3: 3: 3: 3: 3: 4.5. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL = 3. Mesoscutum shiny, slightly or strongly granulate. Notauli incomplete, reaching about 0.3 – 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose (in specimen from Madagascar, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, propodeal declivity with tracks of two complete longitudinal keels). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 5.5). Paramere (Figs 56 E, F) candelabrum shaped, with distal part very slender, with an inner lobe large or small, covered or not with papillae. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	materials_examined	Material examined. Types: ♀ holotype of A. madagascolum: MADAGASCAR: Toliara, Bekily, XI. 1938, A. Seyrig leg. (MNHN); ♂ holotype of A. brooksi: MADAGASCAR: Antananarivo, Antananarivo, 25 – 27. X. 1984, YPT, Robert W. Brooks leg. (NAMUK). Paratype of A. madagascolum: MADAGASCAR: same locality label as holotype, IX. 1938, A. Seyrig leg., 1 ♀ (MRAC); Antsiranana, Antsiranana, 1893, Ch. Alluaud leg., 1 ♀ (not exam- ined) (Benoit 1954); Fianarantsoa, Ranomafana, X. 1938, A. Seyrig leg., 1 ♀ (not examined) (Benoit 1954). Other material: COMOROS ISLANDS: Mohéli Island, Ouallah, 12 ° 18 ’ 13 ’’ S 43 ° 40 ’ 06 ’’ E, 500 m, 18 – 22. I. 2009, MT, rainforest, B. L. Fisher et al. leg., BLF 20683, 1 ♂ (CAS). MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., code BLF 2375, 2 ♂♂ (CAS); Antananarivo, 46 km NE of Ankazobe, Ambohitan- tely, 18 ° 11.88 ’ S 47 ° 16.89 ’ E, 700 m, 17 – 24. X. 2003, MT, sclerophyl forest, M. Irwin & R. Harin’Hala leg., MA- 27 - 02, 1 ♂ (CAS); Antsiranana, Ankarana, IX. 1986, MT, Jane Wilson leg., 1 ♀ (NHMUK); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 23. II – 2. III. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 11, 1 ♂ (CAS); same locality label, 15 – 22. VI. 2011, MG- 62 - 27, 1 ♂ (CAS); same locality label, 18 – 26. V. 2011, MG- 62 - 23, 1 ♂ (CAS); Antsiranana, Orangea, 3 km E of Ramena near fort, 12 ° 14.49 ’ S 49 ° 22.17 ’ E, 65 m, 23 – 27. I. 2001, MT, littoral forest on sand, MA- 01 - 05 - 02, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 1 ♂ (CAS); Antsiranana, Montagne d’Ambre Nat. Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, Antsiranana, 960 m, 26 – 29. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 02, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 7 – 27. I. 2007, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 01 - 01 A- 16, 1 ♂ (CAS); same locality label, 7 – 27. I. 2007, MA- 01 - 01 A- 16, 1 ♂ (CAS); same locality label, 21 – 28. I. 2001, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 1 ♂ (CAS); same locality label, 27. I – 11. II. 2007, MA- 01 - 01 A- 17, 1 ♀ (MOLC); same locality label, 21 – 26. I. 2001, MA- 01 - 01 A- 01, 4 ♂♂ (3 in CAS, 1 in MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, 19. III – 5. IV. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 07, 1 ♂ (CAS); Antsirinana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 29. I – 11. II. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 03, 2 ♂♂ (CAS); same locality label, 14 – 30. V. 2001, MA- 01 - 01 D- 11, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 01, 1 ♂ (CAS); Antsiranana, Marojejy National Park, Manantenina river, 27.6 km 35 ° NE Andapa, 9.6 km 327 ° NNW Manantenina, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 11 – 14. XII. 2005, MT, rainforest, B. L. Fisher et al. leg., BLF 13383, 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 19 – 26. XII. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 12, 1 ♂ (CAS); same locality label, 4 – 10. X. 2010, MG- 57 - 01, 1 ♂ (CAS); Fianarantsoa, 7 km W of Ranomafana, 900 m, 1 – 7. III. 1990, W. E. Steiner leg., 1 ♀, 1 ♂ (USNM); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 10 – 14. I. 2002, secondary tropical forest, MT, R. Harin’Hala leg., 1 ♀ (MOLC); same locality label, 4 – 12. II. 2002, MA- 02 - 09 C- 15, 2 ♂♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 C- 11, 2 ♂♂ (CAS); same locality label, 26. II – 4. III. 2002, MA- 02 - 09 C- 18, 1 ♂ (CAS); same locality label, 28. XI – 6. XII. 2001, MA- 02 - 09 C- 05, 3 ♂♂ (CAS); same locality label, 4 – 14. VII. 2002, MA- 02 - 09 C- 34, 1 ♂ (CAS); same locality label, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 5 ♂♂ (CAS); same locality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 1 ♂ (MOLC); same locality label, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 2 ♂♂ (CAS); same locality label, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 1 ♂ (CAS); same locality label, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 1 ♂ (CAS); same locality label, 19 – 26. II. 2002, MA- 02 - 09 C- 17, 13 ♂♂ (8 in CAS, 5 in MOLC); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 C- 03, 1 ♂ (CAS); same locality label, 23. VI – 4. VII. 2002, MA- 02 - 09 C- 33, 2 ♂♂ (CAS); same locality label, 10 – 21. III. 2003, MA- 02 - 09 C- 56, 3 ♂♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 1 ♂ (CAS); same locality label, 8 – 15. XI. 2001, MA- 02 - 09 C- 02, 2 ♂♂ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 3 ♂♂ (MOLC); same locality label, 26 – 31. III. 2002, MA- 02 - 09 C- 22, 3 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 4 ♂♂ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 9 ♂♂ (7 in CAS, 2 in MOLC); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 6 ♂♂ (CAS); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 28. XI – 6. XII. 2001, MT, near river, R. Harin’Hala leg., MA- 02 - 09 D- 05, 4 ♂♂ (1 in CAS, 3 in MOLC); same locality label, 10 – 14. I. 2002, MA- 02 - 09 D- 11, 1 ♂ (CAS); same locality label, 21 – 24. XII. 2001, MA- 02 - 09 D- 08, 1 ♂ (CAS); same locality label, 2 – 10. I. 2002, MA- 02 - 09 D- 10, 3 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 2 ♂♂ (CAS); same locality label, 19 – 26. II. 2002, MA- 02 - 09 A- 17, 2 ♂♂ (CAS, MOLC); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 1 ♂ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 A- 12, 7 ♂♂ (5 in CAS, 2 in MOLC); same lo- cality label, 21 – 28. I. 2002, MA- 02 - 09 A- 13, 2 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 A- 04, 3 ♂♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 A- 07, 2 ♂♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 5 ♂♂ (3 in CAS, 2 in MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 19, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 B- 01, 9 ♂♂ (8 in CAS, 1 in MOLC); same locality label, 7 – 18. VI. 2003, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 3 ♂♂ (CAS); same locality label, 14 – 24. VI. 2002, MA- 02 - 09 B- 32, 3 ♂♂ (CAS); same locality label, 15 – 27. IV. 2003, MA- 02 - 09 B- 58, 1 ♂ (CAS); same locality label, 12 – 19. III. 2002, MA- 02 - 09 B- 20, 4 ♂♂ (CAS); same locality label, 9 – 20. III. 2003, MA- 02 - 09 B- 55, 1 ♀ (MOLC), 2 ♂♂ (CAS); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 5 ♂♂ (CAS); same locality label, 18 – 29. VI. 2003, MA- 02 - 09 B- 64, 3 ♂♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 2 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 2 ♂♂ (CAS); same locality label, 20. III – 3. IV. 2003, MA- 02 - 09 B- 56, 1 ♂ (CAS); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 2 ♂♂ (CAS); same locality label, 24. V – 4. VI. 2002, MA- 02 - 09 B- 30, 1 ♂ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 2 ♂♂ (CAS); same local- ity label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 10 ♂♂ (8 in CAS, 2 in MOLC); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 2 ♂♂ (CAS); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 2 ♂♂ (CAS, MOLC); same local- ity label, 7 – 17. VII. 2003, MA- 02 - 09 B- 66, 1 ♂ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 3 ♂♂ (CAS); Fianarantsoa, Manombo Special Reserve camp site, 32 km SSE of Farafangana, 23 ° 01.31 ’ S 47 ° 43.20 ’ E, 36 m, 22. V – 6. VI. 2005, MT, lowland rainforest, MA- 28 - 20, M. Irwin, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 30. X – 12. XI. 2004, MA- 28 - 03, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, in dry wash east of Interpre- tive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 12 – 22. VI. 2002, MT, in open area, MA- 02 - 11 B- 32, R. Harin’Hala leg., 1 ♂ (MOLC); same locality label, 26. I – 3. II. 2002, MA- 02 - 11 B- 13, 1 ♂ (CAS); same locality label, 7 – 15. IV. 2002, MA- 02 - 11 B- 23, 1 ♀ (CAS); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 1 ♂ (CAS); Mahajanga, Marotan- drano Special Reserve, Marotandrano, 48.3 km S Mandritsara, 16 ° 17 ’ 00 ’’ S 8 ° 48 ’ 52 ’’ E, 865 m, 7 – 8. XII. 2007, transition humid forest, MT, B. L. Fisher et al. leg., BLF 19157, 1 ♂ (MOLC); Mahajanga, Sofia District, 45 km S Antso- hihy, 5 km W of Anjiamangirana, elev. 97 m, 27 I – 2. II. 2011, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, M. Irwin, R. Harin’Hala, MT, Analagnambe dry degraded forest, MG- 61 - 08, 1 ♂ (MOLC); same locality label, 2 – 6. I. 2011, MG- 61 - 04, 1 ♂ (CAS); Mahajanga, Besalampy District, Analangidro dry forest, 7 km NE Besalampy, 16 ° 41.49 ’ S 44 ° 31.41 ’ E, 200 ft., 9 – 16. X. 2007, MT, dry forest on sand, M. Irwin & R. Harin’Hala leg., MG- 41 - 03, 2 ♂♂ (CAS); same locality label, 28. X – 4. XI. 2007, MG- 41 - 06, 4 ♂♂ (CAS); same locality label, 4 – 11. XI. 2007, 1 ♂ (CAS); same locality label, 11 – 18. XI. 2007, 1 ♂ (CAS); same locality label, 18 – 25. XI. 2007, 4 ♂♂ (CAS); same locality label, 24. IX – 2. X. 2007, 5 ♂♂ (CAS); Mahajanga, Maintirano District, Asondrodava dry forest, 15 km N of Maintirano, 17 ° 57.92 ’ S 44 ° 2.13 ’ E, 200 ft., 21 – 28. I. 2008, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 43 A- 16, 1 ♂ (CAS); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 1 ♂ (CAS); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 15 – 22. I. 2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 14, 1 ♂ (CAS); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19 ’ 16 ’’ S 46 ° 48 ’ 80 ’’ E, 43 m, 24 – 31. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 12, 1 ♂ (CAS); same locality label, 9 – 20. XI. 2003, MA- 25 - 23, 1 ♂ (CAS); same locality label, 14 – 22. IX. 2003, MA- 25 - 15, 1 ♂ (CAS); same locality label, 7 – 14. IX. 2003, MA- 25 - 14, 2 ♂♂ (CAS); same locality label, 31. VIII – 7. IX. 2003, MA- 25 - 13, 1 ♂ (CAS); Mahajanga, Namoroka National Park, 9.8 km 300 ° WNW Vilanandro, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, 140 m, 4 – 8. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445, 1 ♂ (CAS); Toamasina, Ivondro, 1 ♀ (AMNH); Mahajanga, Sofia Dis- trict, 45 km S Antsohihy, 5 km W Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 26. V – 2. VI. 2011, MT, MG- 61 - 15, 2 ♂♂ (CAS); Toliara, 12 km NW of Amboasary, Berenty, 5 – 15. V. 1983, J. S. Noyes & M. C. Day leg., 1 ♂ (NHMUK); same locality label, 18 – 23. V. 1983, 1 ♂ (AMNH); Toamasina, Mobot site, Analalava, 7 km SW of Foulpointe, 17 ° 41 ’ 36 ’’ S 49 ° 27 ’ 37 ’’ E, 60 ft., 16 – 23. XI. 2007, MT, sand in low altitude dense humid forest, MG- 37 B- 10, 1 ♂ (MOLC); Toliara, 60 km NE of Morondava, Bereboka, 18 – 23. V. 1983, J. S. Noyes & M. C. Day leg., 4 ♂♂ (AMNH, NHMUK); Toliara, Tsimanampetsotsa Nat. Park, Bemanateza Forest, 20.7 km 81 ° E Efoetse, 23 km 131 ° SE Beheloka, 23 ° 59.32 ’ S 43 ° 52.50 ’ E, 90 m, 22 – 26. III. 2002, beating low vegetation, in spiny forest thicket, Fisher, Griswold et al. leg., BLF 6256, 1 ♀ (CAS); same locality label, 22 – 26. III. 2002, BLF 6257, 1 ♂ (CAS); same locality label, 22 – 26. III. 2002, BLF 6257, 1 ♂ (CAS); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 50.09 ’ S 46 ° 29.21 ’ E, 80 m, 26. XII. 2002 – 5. I. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 05, 1 ♀ (CAS); same locality label, 7 – 17. XII. 2003, MA- 02 - 21 - 41, 1 ♂ (CAS); same locality label, 28. IV – 6. V. 2003, MA- 02 - 21 - 17, 1 ♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 16 – 17. XII. 2002, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 09, 2 ♂♂ (CAS, MOLC); same locality label, 28. III – 8. IV. 2003, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 10 – 20. VII. 2003, 1 ♂ (CAS); same locality label, 17 – 24. VIII. 2003, MA- 02 - 20 - 36, 1 ♂ (CAS); same locality label, 22 – 29. VI. 2003, MA- 02 - 20 - 30, 3 ♂♂ (CAS); same locality label, 30. XI – 11. XII. 2003, MA- 02 - 20 - 47, 2 ♂♂ (CAS); Toliara, Fiherenana, 23 ° 14.07 ’ S 43 ° 52.15 ’ E, 50 m, 16. X – 16. XI. 2002, MT. degraded gallery forest, MGF 047, Frontier Wilderness Project, 1 ♀ (CAS); Toliara, Fiherenana, 23 ° 10.37 ’ S 43 ° 57.39 ’ E, 100 m, 22 – 28. X. 2002, MT. gallery forest, MGF 041, Frontier Wilderness Project, 1 ♂ (MOLC); Toliara, Ranobe, 23 ° 02.22 ’ S 43 ° 36.377 ’ E, 30 m, 5 – 9. II. 2003, MT, spiny forest thicket, MGF 058, Frontier Wilderness Project, 1 ♂ (CAS); Toli- ara, Ranobe, 23 ° 02.367 ’ S 43 ° 36.617 ’ E, 20 m, 8 – 15. XI. 2003, MT, gallery forest, MGF 084, Frontier Wilderness Project, 1 ♀ (MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 6 – 16. I. 2002, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 09, 1 ♀ (CAS); same locality label, 6 – 16. I. 2002, MA- 02 - 18 A- 09, 1 ♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 18 – 29. IV. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 18 B- 20, 1 ♂ (CAS); same locality label, 11 – 24. II. 2003, MA- 02 - 18 A- 46, 1 ♂ (CAS); Toliara, Zombitse National Park, near national road, 22 ° 50.43 ’ S 44 ° 43.87 ’ E, 825 m, 15. X – 9. XI. 2001, MT, deciduous spiny forest, R. Harin’Hala leg., MA- 02 - 13 B- 01, 1 ♂ (CAS); same locality label, 7 – 13. VII. 2002, 1 ♂ (CAS); same locality label, 13 – 20. III. 2002, 1 ♂ (MOLC); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 15. X – 9. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 01, 2 ♂♂ (CAS, MOLC); Toliara, Beza Mahafaly Reserve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 20 – 24. XII. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 45, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue, 23 ° 41.39 ’ S 44 ° 34.53 ’ E, 180 m, 15. X – 10. XI. 2001, MT, spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 14 B- 01, 1 ♂ (CAS); Toliara, Antafoky, 23 ° 28.43 ’ S 44 ° 03.51 ’ E, 55 m, 25 – 28. I. 2002, MT, gallery forest, Frontier Project leg., MGF 003, 1 ♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 1 ♂ (CAS); same locality label, 16 – 27. XII. 2002, 2 ♂♂ (CAS); same locality label, 25. II – 7. III. 2003, MA- 02 - 22 - 17, 2 ♂♂ (CAS, MOLC); same locality label, 17 – 24. III. 2003, MA- 02 - 22 - 19, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 5 – 15. II. 2003, MA- 02 - 22 - 15, 4 ♂♂ (CAS); same locality label, 7 – 17. III. 2003, MA- 02 - 22 - 18, 5 ♂♂ (CAS); same locality label, 15 – 25. II. 2003, MA- 02 - 22 - 16, 3 ♂♂ (CAS); same locality label, 17 – 26. I. 2003, MA- 02 - 22 - 13, 4 ♂♂ (CAS); same locality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 2 ♂♂ (CAS); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 3 ♂ # ♂ (CAS); To- liara, Berenty Special Reserve, 8 km NW Amboasary, elev 35 m, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 22 – 30. XI. 2003, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 A- 04, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 7 – 17. X. 2004, 1 ♂ (CAS); Toliara, Berenty Reserve, 80 km W Fort Dauphin, 25 ° 00 ’ S 46 ° 17 ’ E, 7. IV. 1994, MT, M. Wasbauer leg., 1 ♂ (DEUCD); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 8 – 16. VI. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 26, M. Irwin, R. Harin’Hala leg., 3 ♂♂ (CAS); same lo- cality label, 430 ft., 22 – 30. V. 2008, MG- 45 A- 24, 1 ♂ (MOLC); same locality label, 1 – 9. VII. 2008, MG- 45 A- 29, 2 ♂♂ (CAS); Toliara, Mite Forest, 20.7 km 29 ° WNW Tongobory, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, 75 m, 27. II – 3. III. 2002, MT, in gallery forest, Fisher, Griswold et al. leg., BLF 5849, 1 ♂ (CAS); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.09 ’ S 044 ° 39.38 ’ E, 6 – 13. VIII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 40, 1 ♂ (MOLC); same locality label, 15 – 22. X. 2008, BLF 18285 _ 50, 1 ♂ (MOLC); Toliara, Androimpano Forest, 5 km E of Itampolo, 24 ° 39.02 ’ S 43 ° 57.79 ’ E, 130 m, 3 – 16. VII. 2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54 C- 33, 1 ♂ (CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 9 – 15. I. 2009, MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG- 53 C- 12, 1 ♂ (CAS); same locality label, 24 – 29. X. 2008, MG- 53 C- 01, 1 ♂ (CAS); same locality label, 4 – 12. XII. 2008, MG- 53 C- 07, 1 ♂ (CAS); same locality label, 24 – 29. X. 2008, across trail at base of escarpment, dense dry forest, MG- 53 B- 01, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	distribution	Distribution. Comoros Islands, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0F2AECFF3ED59BFDAFFE70.taxon	discussion	Remarks. The record from Diego Suarez Island reported by Olmi (1984) is erroneous, because it was based on a misinterpretation of a paper of Benoit (1954), and instead refers to the town of Diégo-Suarez situated in Madagascar. The new synonymy of Anteon brooksi is justified because the small difference in the ♂ genitalia of A. madagascolum (the only difference between these two species) falls within the range of variability of the species, as shown by the study of hundreds of specimens.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	description	(Figs 58 A, 59, 60 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	diagnosis	Diagnosis. ♂ of Anteon with head granulate; OPL slightly shorter than TL; antenna almost moniliform (Fig. 58 A); antennomere 4 about as long as broad; mesoscutum dull, granulate; notauli very short, reaching about 0.3 × length of mesoscutum; metapectal-propodeal disc with strong transverse posterior keel; propodeal declivity without longitudinal keels; paramere with inner side provided with numerous papillae (Fig. 60 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	description	Description. ♂ (Figs 58 A, 59). Fully winged; body length 1.6 mm. Head black, except mandible testaceous; antenna brown-testaceous, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs brown, except tarsi and part of coxae, tibiae and femora testaceous. Antenna almost moniliform; antennomeres in following proportions: 6: 4: 2: 2: 3: 3: 3: 3.5: 3: 4; antennomere 4 about as long as broad. Head (Fig. 59 D) dull, granulate; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL = 2.5; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum (Fig. 59 B) dull, granulate. Notauli very short, reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels; areolae of propodeal declivity less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (1: 4). Paramere (Fig. 60 A) with inner side provided with numerous papillae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	materials_examined	Material examined. Type: ♂ holotype (CASTYPE 19442): MADAGASCAR: Mahajanga, Ampijoroa Nation- al Park, 160 km N of Maevatanana on RN 04, 16 ° 19 ’ 16 ’’ S 46 ° 48 ’ 80 ’’ E, 43 m, 17 – 24. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 11 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AECFF3ED407FA6AFA44.taxon	etymology	Etymology. The species is named majunganum after the collection site in Majunga (= Mahajanga) Province.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	description	(Figs 55 B – E, 60 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	description	Description. ♂ (Figs 55 B – E). Fully winged; body length 1.7 mm. Head black, except mandible testaceous; an- tenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous-dark. Antenna filiform; antennomeres in following proportions: 9: 4: 4: 4: 4: 4: 4: 4: 4.5: 7. Head completely reticulate rugose and granulate; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 4; OPL = 1.5; TL = 2; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum dull, with anterior third reticulate rugose; remaining surface sculptured by many longitudinal subparallel irregular keels. Notauli very short, hardly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, sculptured by areolae less wide than those of metapectal-propodeal disc, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Paramere (Fig. 60 B) without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	materials_examined	Material examined. Type: ♂ holotype: UGANDA: Western Region, Kibale National Park, Kanyawara, Mak- erere University Biological Field Station, 0 ° 33.408 ’ S 30 ° 22.603 ’ E, 1587 m, 30. VII – 5. VIII. 2005, MT, degraded mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 023413 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF0B2AEEFF3ED05BFB95FA45.taxon	discussion	Remarks. The species is named after Makerere University Biological Field Station.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	description	(Figs 58 B, 60 C – E, 62)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	diagnosis	Diagnosis. ♂ of Anteon with head completely strongly reticulate rugose (Fig. 58 B); propodeal declivity reticulate rugose, with one or two longitudinal keels, with median area rugose; paramere with proximal inner lobe provided or not with mosaic sculpture (Figs 60 C, D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	description	Description. ♂ (Figs 58 B, 62). Fully winged; body length 2.1 – 2.2 mm. Head black, except mandible testaceous; antenna testaceous (in paratype brown); mesosoma black; metasoma brown; legs testaceous (in paratype metacoxa and club of metafemur partly brown). Antenna filiform; antennomeres in following proportions: 9: 5: 5: 5: 5: 5: 5: 5: 5: 7. Head (Fig. 62 D) dull, completely strongly granulate and reticulate rugose; frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 3; OOL = 5; OPL = 2; TL = 3; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum shiny, completely slightly granulate. In specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, MA- 02 - 09 A- 06, mesoscutum punctate, unsculptured among punctures. Notauli (Fig. 62 B) incomplete, reaching about 0.3 – 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc (Fig. 62 B) with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, with one or two longitudinal keels; areolae of propodeal declivity about as large as those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Paramere (Figs 60 C-E) without distal in- ner pointed process, with inner more or less large proximal lobe not covered with papillae and having (Fig. 60 C) or not having (Figs 60 D, E) a mosaic pattern. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	materials_examined	Material examined. Type: ♂ holotype (CASTYPE 19443): MADAGASCAR: Fianarantsoa, Ranomafana Na- tional Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 14 – 24. VII. 2002, MT, secondary tropical for- est, R. Harin’Hala leg., MA- 02 - 09 C- 35 (CAS). Paratypes: MADAGASCAR: same locality label as holotype, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 1 ♂ (CAS); same locality label as holotype, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♂ (CAS); same locality label as holotype, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (CAS); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 2 – 10. I. 2002, MT, near river, R. Harin’Hala leg., MA- 02 - 09 D- 10, 1 ♂ (MOLC); Fianarantsoa, Ranomafana Na- tional Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 22. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 03, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 1 ♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 1 ♂ (CAS); same locality la- bel, 7 – 17. VII. 2003, MA- 02 - 09 B- 66, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 15 – 21. XII. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 07, 2 ♂♂ (CAS, MOLC); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	etymology	Etymology. The species is named malagasy after the collection site in Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF092AE0FF3ED05AFCE7FCBC.taxon	discussion	Remarks. Anteon malagasy is similar to A. kwazuluense Olmi, 2007, but the propodeal declivity has areolae as large as those of the metapectal-propodeal disc (the areolae are less wide than those of the metapectal-propodeal disc in A. kwazuluense). Anteon malagasy is also similar to A. madagascolum (Benoit, 1954), but the shape of the inner process of the paramere is different (Figs 56 E, F).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF072AE1FF3ED5C3FCC0FAFC.taxon	description	(Figs 57 F, 60 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF072AE1FF3ED5C3FCC0FAFC.taxon	description	Description. ♀. Fully winged; body length 2.5 – 3.2 mm. Head black, except mandible testaceous (with teeth brown); antenna testaceous, except antennomeres 7 – 10, or 8 – 10, or 9 – 10 darkened; propleuron reddish or brown, except dorsal surface testaceous; pronotum completely reddish; occasionally pronotum totally black, or brown, except anterior and lateral margins testaceous; pronotal tubercle testaceous; tegula testaceous; rest of mesosoma and metasoma black; legs brown, except tarsi, procoxa and protibia testaceous. Antenna clavate; antennomeres in following proportions: 15: 5: 6: 5: 5: 5: 5: 5: 4: 5. Head shiny; frons sculptured by numerous transverse keels, with frontal line complete and two lateral keels near orbits; vertex with two oblique keels connecting lateral ocelli to occipital carina, unsculptured between and on sides of above keels; occipital carina complete; POL = 7; OL = 4; OOL = 6; OPL = 5; TL = 5. In specimen from Kenya, 2.99615 ° S 38.45988 ° E, head with frons completely reticulate rugose, not sculptured by transverse keels. Pronotum shiny, setose, unsculptured, with raised transverse carina between anterior and posterior surface; anterior surface approximately as long as OPL (6: 5); posterior surface shorter than mesoscutum (5: 15). Mesoscutum shiny, setose, unsculptured. Notauli incomplete, reaching approximately 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part. Protarsomere 1 approximately as long as 4; protarsomere 4 slightly shorter than basal part of 5; protarsomere 3 produced into hook. Enlarged claw (Fig. 57 F) very long, longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 F) with basal part longer than apical part, with about 23 lamellae, with inner side almost straight, without separate group of distal lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.0 – 2.9 mm. Head black, except mandible testaceous (with teeth brown); antenna brown, except scape and pedicel yellow (in five ♂♂ from South Africa, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, antenna totally testaceous); mesosoma and metasoma black; tegula testaceous; legs brown, except tarsi and protibia testaceous. Head shiny, slightly granulate, usually with numerous irregular transverse keels; in some male specimens, head completely granulate and reticulate rugose; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 2; TL = 2. In specimen from Kenya, 2.05111 ° S 38.32613 ° E, frons mostly punctate, unsculptured among punctures; vertex granulate and partly rugose. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured. Occasionally (in ♂ from South Africa, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, mesoscutum weakly granulate). Notauli incomplete, reaching approximately 0.25 – 0.30 × length of mesoscutum. Mesopleuron with posterior surface partly smooth and shiny; anterior surface dull, with numerous irregular keels. Metapleuron dull, with numerous irregular keels. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 5). Paramere (Fig. 60 F) with distal inner pointed process (margin of process with short setae) and without proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF072AE1FF3ED5C3FCC0FAFC.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Milnerton, II. 1926, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, 2 ♀♀, 2 ♂♂ (NHMUK). Other material: BURUNDI: Rusizi National Park, 3.34364 ° S 29.27246 ° E, 774 m, 13 – 20. IV. 2010, MT, near three small trees, degraded bush / grassland, R. Copeland leg., 1 ♂ (NMK). DEMOCRATIC REPUBLIC OF THE CONGO: Orientale Prov. (= Haut-Zaire), Ga- ramba National Park, 1 ♀ (AMNH). GABON: Ntoum, 1 ♀ (FSAE). KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 25. XII. 2014 – 8. I. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♂ (NMK); same locality label, 13 – 27. III. 2015, 1 ♂ (NMK); same locality label, 8 – 22. I. 2015, 2 ♂♂ (NMK); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg., 2 ♂♂ (NMK); Eastern Prov., Simisi area, 2.05111 ° S 38.32613 ° E, 710 m, 28. XI – 1. XII. 2013, MT, near base of Yamalu Hill, Acacia shrubland, J. Bukhebi and R. Copeland leg., 1 ♂ (NMK); Rift Valley Prov., Tsavo West National Park, Riverine woodland, 2.99615 ° S 38.45988 ° E, 464 m, 9 – 23. IX. 2008, MT, bank of Tsavo River, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Nguruman, near Sampu River, 1.90117 ° S 36.05040 ° E, 723 m, 5 – 19. XI. 2011, MT, near base of Nguruman Escarpment, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 15 – 30. III. 2006, MT, G. Whiteley leg., 5 ♂♂ (4 in DJBC, 1 in MOLC); same locality label, 11. IV – 11. V. 2006, 1 ♂ (DJBC); same locality label, 24. XI – 8. XII. 2005, 10 ♂♂ (9 in DJBC, 1 in MOLC); same locality label, 6. I – 1. II. 2006, 6 ♂♂ (DJBC); same locality label, 14 – 27. II. 2006, 5 ♂♂ (DJBC); same locality label, 29. X – 9. XI. 2005, 9 ♂♂ (DJBC); same locality label, 9 – 24. XI. 2005, 19 ♂♂ (DJBC); Western Cape, Milnerton, II. 1926, many specimens (MRAC); same locality label, 1 ♀ (AMNH); same locality label, I. 1926, 6 ♀♀, 7 ♂♂ (NHMUK); Western Cape, Ceres, II. 1921, 2 – 21. III. 1921, R. E. Turner leg., 2 ♀♀, 1 ♂ (NHMUK); Western Cape, Mossel Bay, XII. 1921, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 23. IX – 12. X. 2006, 2 ♂♂ (DJBC); same locality label, 15. XI. 2006 – 5. I. 2007, 1 ♂ (DJBC); same locality label, 27. II – 15. III. 2006, 2 ♂♂ (DJBC); same locality label, 8. XII. 2005 – 6. I. 2006, 11 ♂♂ (DJBC). ZIMBABWE: Harare, Chishawasha, X. 1982, A. Watsham leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF072AE1FF3ED5C3FCC0FAFC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF072AE1FF3ED5C3FCC0FAFC.taxon	distribution	Distribution. Burundi, Democratic Republic of the Congo, Gabon, Kenya, South Africa, Zimbabwe. Not present in Madagascar (record of Azevedo et al. 2010 wrong).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF062AE2FF3ED383FCAFFDE4.taxon	description	(Figs 57 E, 58 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF062AE2FF3ED383FCAFFDE4.taxon	description	Description. ♀ (Figs 58 C, D). Fully winged; body length 2.3 – 2.4 mm. Head brown-ferruginous, except mandible testaceous; antenna testaceous-brown, except scape and pedicel testaceous; propleuron and pronotum yellow-testaceous; rest of mesosoma black; metasoma testaceous-brown; legs testaceous, except stalk of metafemur brown. Antenna clavate; antennomeres in following proportions: 8: 4.5: 6: 5: 5: 5: 4.5: 4.5: 4.5: 5.5. Head convex, dull, almost completely bare, strongly granulate; frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 5.5; OL = 3; OOL = 4; OPL = 4.5; TL = 5; greatest breadth of posterior ocellus shorter than Ol (3: 2). Pronotum anteriorly crossed by weak transverse impression, dull, granulate; posterior surface granulate, dull, slightly shorter than mesoscutum (10: 11); posterior surface of pronotum with lateral sharp carinas separating disc from lateral regions; pronotal tubercle reaching tegula. Mesoscutum dull, completely granulate, with some rugosities near posterior margin and on lateral regions. Notauli very shortly visible near anterior margin of mesoscutum. Mesoscutellum dull, granulate. Metanotum shiny, rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Protarsomeres in following proportions: 6: 2: 3: 7: 15. Enlarged claw (Fig. 57 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 57 E) with basal part much shorter than distal part, with two rows of 10 + 13 lamellae; apex with four-six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF062AE2FF3ED383FCAFFDE4.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Nyanza Prov., Ruma National Park, near Kamato Gate, 0.64725 ° S 34.33595 ° E, 1264 m, 10 – 18. XII. 2005, MT, open grass- woodland, R. Copeland leg. (NMK). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 23. IV. 2006, MT along forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF062AE2FF3ED383FCAFFDE4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF062AE2FF3ED383FCAFFDE4.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF052AE3FF3ED4FBFCB5FE0C.taxon	description	(Figs 63 A, 65 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF052AE3FF3ED4FBFCB5FE0C.taxon	description	Description. ♀. Fully winged; body length 2.2 – 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous (in two specimens from Tanzania, Mkomazi Game Reserve, and South Africa, Ukulinga, antenna testaceous-brown, except scape, pedicel and flagellomeres 1 – 2 testaceous); mesosoma and metasoma black; tegula testaceous; legs brown, except tibiae and tarsi testaceous (in two specimens from Tanzania, Mkomazi Game Reserve, and South Africa, Ukulinga, legs testaceous, except metacoxa slightly darkened). Antenna clavate; antennomeres in following proportions: 9: 5: 6: 5: 5: 5: 5: 5: 5: 7. Head dull, granulate, with few slight areolae near clypeus (in two specimens from Tanzania, Mkomazi Game Reserve, and South Africa, Ukulinga, head granulate, with slightly distinct areolae present on entire surface); frontal line present; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 5; OL = 4; OOL = 5; OPL = 6; TL = 6. Pronotum dull, granulate, with smooth and unsculptured stripe at posterior margin; anterior half with numerous transverse keels; posterior surface very long, more than 0.5 × as long as mesoscutum; pronotal tubercle reaching tegula. Mesoscutum dull, granulate (in specimen from South Africa, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, mesoscutum granulate and with some slight areolae and irregular keels on lateral regions). Notauli absent. In specimen from Cameroon, Nkoemvon, notauli shortly present near anterior margin. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 5). Protarsomeres in following proportions: 6: 2: 2: 2.5: 8. Enlarged claw (Fig. 63 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 A) with basal part longer than apical part (5: 3), with one row of 7 – 8 lamellae; apex with 5 – 6 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.4 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa, metafemur and distal region of metatibia brownish. Antenna filiform; antennomeres in following proportions: 6: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 6. Head dull, granulate; frontal line absent; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 4; OL = 2.5; OOL = 4; OPL = 2; TL = 4. Mesoscutum shiny, smooth, slightly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 4). Paramere (Fig. 65 A) with distal inner rounded process, without proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF052AE3FF3ED4FBFCB5FE0C.taxon	materials_examined	Material examined. Types: ♀ holotype: NIGERIA: W State, Ile-Ife, X. 1973, J. T. Medler leg. (NHMUK). Paratype: same locality label as holotype, V. 1973, 1 ♂ (NHMUK). Other material: CAMEROON: South Region, Nkoemvon, 1 ♀ (CIRAD); same locality label, 2. XI – 13. XII. 1980, first forest path, D. Jackson leg., 1 ♀ (NHMUK). SOUTH AFRICA: KwaZulu-Natal, 10 km SE Pietermaritzburg, Ukulinga, 6 – 12. II. 1986, grassland, MT, R. M. Miller leg., 1 ♀ (DJBC); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 15 – 30. III. 2006, MT, G. Whiteley leg., 1 ♀ (MOLC). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, north west side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 18 – 22. IV. 1996, Acacia / Commiphora / Combretum bushland, YPT, SAM-HYM-PO 19460, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF052AE3FF3ED4FBFCB5FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF052AE3FF3ED4FBFCB5FE0C.taxon	distribution	Distribution. Cameroon, Nigeria, South Africa, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE3FF3ED793FE14F9CA.taxon	description	(Figs 58 E, 63 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE3FF3ED793FE14F9CA.taxon	description	Description. ♀ (Fig. 58 E). Fully winged; body length 1.6 – 2.2 mm. Head black, except mandible testaceous; clyp- eus of holotype testaceous-brown; clypeus of specimen from Madagascar, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, black; antennomeres 1 – 7 of holotype yellow (antennomeres 8 – 10 missing in holotype); antenna of specimen from Madagascar, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, yellow, except antennomere 10 darkened; mesosoma black; metasoma brown; legs yellow. Antenna clavate; antennomeres in following proportions: 9: 4.5: 5: 3.5: 4: 4: 4 (antennomeres 8 – 10 missing in unique known specimen). Head convex, dull, granulate and with many irregular striae; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 4.5; OPL = 3; TL = 3.5. Pronotum anteriorly crossed by slight transverse impression, dull, rugose; posterior surface rugose, much shorter than mesoscutum (5: 11), with posterior margin shiny and smooth; pronotal tubercle reaching tegula. Mesoscutum shiny, slightly granulate or with some irregular striae. Notauli incomplete, reaching approximately 0.5 – 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without complete longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Protarsomeres in following proportions: 6: 2: 2: 4: 10. Enlarged claw (Fig. 63 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 B) with inner side straight, with basal part slightly longer than apical part (5.5: 5), with distal extremity of extended part divided into two lobes (incision between them receiving distal extremity of enlarged claw), one of them bearing one very large lamella; second lobe with one row of lamellae continuing in the extended part of protarsomere 5 (reaching a total of 10 – 17 lamellae). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE3FF3ED793FE14F9CA.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375 (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 4 – 19. III. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 08, 1 ♀ (CAS); Antsira- nana, Marojejy National Park, Manantenina River, 27.6 km 327 ° NNW Manantenina, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 15 – 18. XI. 2003, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher et al. leg., BLF 8872, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE3FF3ED793FE14F9CA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE3FF3ED793FE14F9CA.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE4FF3ED0EEFE14F880.taxon	description	(Fig. 63 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE4FF3ED0EEFE14F880.taxon	description	Description. ♀. Fully winged; body length 1.9 – 3.4 mm. Head black, except mandible testaceous; antenna testa- ceous, except antennomeres 6 – 10 or 7 – 10 brown; mesosoma and metasoma black; legs testaceous, except metacoxa partly brown; occasionally legs testaceous, except stalk of metafemur and all arolia brown. In specimen from Madagascar, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, legs and antenna testaceous (except antennomere 10 slightly darkened). Antenna clavate; antennomeres in following proportions: 10: 4: 3: 3: 3: 4: 4: 3.5: 4: 5. Head shiny, reticulate rugose; frontal line present; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 4; OL = 3; OOL = 6; OPL = 3; TL = 3. Pronotum rugose; posterior surface almost completely smooth and unsculptured in holotype, slightly granulate in specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, rugose in specimen from Madagascar, 20 ° 10 ’ 40 ’’ S 47 ° 05 ’ 18 ’’ E; posterior surface shorter than mesoscutum (3: 12 in holotype; 3: 9 in specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E; 3: 14 in specimen from Madagascar, 20 ° 10 ’ 40 ’’ S 47 ° 05 ’ 18 ’’ E); pronotal tubercle reaching tegula. Mesoscutum shiny, slightly (in holotype) or strongly (in specimens from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E and 18 ° 55.58 ’ S 48 ° 24.47 ’ E) granulate. In specimens from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, MA- 02 - 09 A- 35, and 20 ° 10 ’ 40 ’’ S 47 ° 05 ’ 18 ’’ E, mesoscutum granulate and with very slight irregular longitudinal folds. In specimen from Madagascar, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, mesoscutum alutaceous, with slight irregular folds, not granulate. Notauli absent. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (1: 7). Protarsomeres in following proportions: 6: 1.5: 1.5: 1.5: 6. Enlarged claw (Fig. 63 C) longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 C) with basal part much longer than apical part (4.5: 1.5), with 6 – 17 lamellae (6 in the holotype; 17 in specimen from Madagascar, 21 ° 13.57 ’ S 47 ° 22.19 ’ E) and some bristles (in specimen from Madagascar, 20 ° 10 ’ 40 ’’ S 47 ° 05 ’ 18 ’’’ E, lamellae present on apex; proximal and medial region with bristles, some of which very broad and similar to lamellae). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE4FF3ED0EEFE14F880.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, Ankaratra (MNMS). Other material: MADAGASCAR: Antananarivo, Ankokoy Forest, 3 km E of Ibity, 20 ° 04.05 ’ S 46 ° 59.97 ’ E, 1700 m, 14 – 24. XI. 2008, MT, in Uapaca forest, M. Irwin, R. Harin’Hala leg., MG- 56 - 02, 1 ♀ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 16 – 31. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 10, R. Harin’Hala leg., 1 ♀ (MOLC); same locality label, 15. II – 6. III. 2001, MA- 01 - 04 B- 10, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg., 1 ♀ (CAS); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantel- la, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 18 – 25. III. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 16, 1 ♀ (MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 1 ♀ (CAS); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 1 ♀ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 A- 12, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 1 – 12. VII. 2006, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 154, 1 ♀ (MOLC); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 1 ♀ (CAS); same locality label, 15 – 22. XI. 2001, MA- 02 - 09 B- 03, 1 ♀ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 1 ♀ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♀ (MOLC); same locality label, 18 – 27. II. 2003, MA- 02 - 09 B- 53, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 4 – 16. V. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 60, 1 ♀ (CAS); same locality label, 13 – 23. VI. 2002, MA- 02 - 09 C- 32, 1 ♀ (MOLC); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♀ (CAS); Fianarantsoa, Italaviana, 35 km SSE of Antsir- abe, 20 ° 10 ’ 40 ’’ S 47 ° 05 ’ 18 ’’ E, 1360 m, 30. III – 9. IV. 2003, MT, in Uapacca forest, M. Irwin, R. Harin’Hala leg., MA- 24 - 12, 1 ♀ (CAS); same locality label, 5 – 16. X. 2003, MA- 24 - 31, 1 ♀ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 8 – 16. X. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 15, 1 ♀ (CAS); same locality label, 24. X – 1. XI. 2001, MA- 01 - 08 B- 17, 1 ♀ (CAS); same locality label, 1 – 7. XI. 2001, 1 ♀ (MOLC); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.07 ’ S 44 ° 39.34 ’ E, 24 – 31. XII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18284 _ 60, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE4FF3ED0EEFE14F880.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF042AE4FF3ED0EEFE14F880.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF032AE5FF3ED117FD85FD00.taxon	description	(Fig. 65 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF032AE5FF3ED117FD85FD00.taxon	description	Description. ♂. Fully winged; body length 2.2 – 3.4 mm. Black, except mandible and tegula testaceous. Antenna filiform; antennomeres in following proportions: 8: 5: 10: 9.9: 9: 9: 9: 9: 9. Head shiny, strongly punctate, unsculptured among punctures; occipital carina complete; POL = 8; OL = 4; OOL = 7; OPL = 3; TL = 5. Mesoscutum shiny, strongly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc completely reticulate rugose, without transverse posterior keel. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 9). Paramere (Fig. 65 B) without distal inner process and proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF032AE5FF3ED117FD85FD00.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, X. 1933, R. E. Turner leg. (NHMUK). Paratypes: SOUTH AFRICA: Western Cape, near Stellenbosch, Jonkershoek Reserve, 14 ♂♂ (10 in AEIC, 4 in AMNH). Other material: SENEGAL: near Kaolack, Diedieng, 1 ♂ (FSAE).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF032AE5FF3ED117FD85FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF032AE5FF3ED117FD85FD00.taxon	distribution	Distribution. Senegal, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE5FF3ED497FCBDF886.taxon	description	(Figs 63 D, 68 A – D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE5FF3ED497FCBDF886.taxon	description	Description. ♀ (Figs 68 A – D). Fully winged; body length 2.6 mm. Head black, except mandible and part of anterior half of clypeus testaceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 8: 6: 7: 6: 5: 5: 6: 6: 6: 8. Head dull, reticulate rugose, except vertex behind lateral ocelli and temple granulate; frontal line complete, partly continuing also behind the anterior ocellus; frons with two lateral keels along orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 4; OOL = 4 OPL = 4; TL = 5; greatest breadth of lateral ocelli shorter than OPL (3: 4). Pronotum with slight transverse anterior impression; anterior surface short, transversely striate, hidden behind head; posterior surface slightly shorter than mesoscutum (11: 14), shiny, granulate, except few irregular keels near anterior margin; posterior surface with raised carina on anterior and lateral margins; pronotal tubercle reaching tegula. Mesoscutum shiny, slightly granulate, with some irregular keels on lateral surfaces. Notauli absent. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with with areolae about as large as those of metapectal-propodeal disc, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 6: 2: 3: 10: 18; protarsomere 2 produced into hook; protarsomere 4 much longer than basal part of 5 (10: 5). Enlarged claw (Fig. 63 D) with a proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 D) with basal part much shorter than distal part (5: 13), with two rows of 8 + 12 lamellae; apex with 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE5FF3ED497FCBDF886.taxon	materials_examined	Material examined. Type: ♀ holotype: UGANDA: Western Region, Kibale National Park, Kanyawara Mak- erere University Biological Field Station, 0 ° 35.405 ’ S 30 ° 21.646 ’ E, 1484 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, near stream, S. van Noort leg., SAM-HYM-A 023421 (SAMC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Ituri Prov., Djugu Territory, confluence of Ituri and Kelebo rivers, Uphill “ Camp Kelebo ”, 30. III – 6. IV. 2015, primary / secondary forest, 75 YPTs, A. Gumovsky leg., 1 ♀ (SIZK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE5FF3ED497FCBDF886.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE5FF3ED497FCBDF886.taxon	distribution	Distribution. Democratic Republic of the Congo, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	description	(Figs 58 F, 61, 63 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with head granulate (Fig. 58 F); propodeal declivity with two complete longitudinal keels; protarsomere 4 much longer than basal part of 5.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	description	Description. ♀ (Figs 58 F, 61). Fully winged; body length 2.2 mm. Head reddish-brown-, except mandible partly testaceous; antenna testaceous; mesosoma reddish-brown, except dorsal region of metapectal-propodeal disc and propodeal declivity black; metasoma and legs testaceous. Antenna clavate; antennomeres in following proportions: 9: 5: 5: 5: 3.5: 3.5: 3.5: 4: 4: 6. Head (Fig. 61 C) dull, granulate; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 4; OL = 4; OOL = 4; OPL = 4; TL = 5; greatest breadth of lateral ocelli shorter than OL (2: 4). Pronotum dull, strongly crossed by transverse keels, with posterior surface weakly granulate, shorter than mesoscutum (10: 14) and much longer than anterior surface (10: 4); pronotal tubercle reaching tegula. Mesoscutum dull, weakly granulate. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two complete longitudinal keels, lateral areas reticulate rugose and median area rugose and with areolae less wide than those of lateral areas. Forewing (Fig. 61 D) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Protarsomeres in following proportions: 6: 2: 3: 10: 18; protarsomere 4 much longer than basal part of 5 (10: 5). Enlarged claw (Fig. 63 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 E) with basal part much shorter than apical part (5: 13), with two rows of 5 + 17 lamellae; apex with three lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	materials_examined	Material examined. Type: ♀ holotype, NMK / INV / T- 636: KENYA: Coast Prov., Marenje Forest, 4.52814 ° S 39.24028 ° E, 76 m, 18. IX – 2. X. 2014, MT, indigenous forest, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF022AE6FF3ED107FD45FC4C.taxon	etymology	Etymology. The species is named in honour of a longtime friend of one of the authors (RSC), Jeffrey L. Musman, legal expert and exceptional 3 rd baseman.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF002AF8FF3ED35DFD5CFF50.taxon	description	(Fig. 63 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF002AF8FF3ED35DFD5CFF50.taxon	description	Description. ♀. Fully winged; body length 2.0 – 2.2 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; tegula testaceous; mesosoma and metasoma black; proleg testaceous; meso- and metaleg brown. Antennomeres in following proportions: 11: 4: 4: 3: 3.5: 3.5: 3.5: 3.5: 3.5: 4.5. Head dull, granulate; frontal line complete; frons with two lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 8; OL = 4; OOL = 5; OPL = 3; TL = 2. Pronotum granulate; posterior surface much shorter than mesoscutum (2: 15); pronotal tubercle reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, very short, faintly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity granulate, slightly rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Protarsomeres in following proportions: 6.5: 1.5: 1: 1.5: 5.5. Enlarged claw (Fig. 63 F) very long, longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 63 F) with four lamellae and some bristles. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF002AF8FF3ED35DFD5CFF50.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: Aus, 8 – 30. XI. 1929, R. E. Turner leg. (NHMUK). Para- types: NAMIBIA: same locality label as holotype, 1 ♀ (AMNH). SOUTH AFRICA: Free State, Glen, X. 1958, D. P. Annecke leg., 1 ♀ (NHMUK). Other material: BENIN: Gomé, 12. XI. 1993, 1 ♀ (CIRAD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF002AF8FF3ED35DFD5CFF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF002AF8FF3ED35DFD5CFF50.taxon	distribution	Distribution. Benin, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1F2AF9FF3ED383FB91FD2C.taxon	description	(Fig. 67 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1F2AF9FF3ED383FB91FD2C.taxon	description	Description. ♀. Fully winged; body length 2.6 – 2.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous. In specimen from Zimbabwe, Harare, antenna brown, except antennomeres 1 – 3 testaceous; proleg testaceous, except coxa partly brown; meso- and metaleg brown, except tarsi testaceous. Antennomeres in following proportions: 13: 4.5: 4: 4: 4: 5: 5: 5: 5: 7. Head shiny, reticulate rugose; occipital carina complete; POL = 8; OL = 5; OOL = 5; OPL = 4; TL = 3. Pronotum shiny, rugose; posterior surface very short, shorter than mesoscutum (5: 16); pronotal tubercle reaching tegula. Mesoscutum shiny, reticulate rugose, with or without numerous strong longitudinal keels; in specimen from Zimbabwe, Harare, mesoscutum granulate, with slight areolae and irregular and longitudinal keels. Notauli absent. In specimen from Zimbabwe, Harare, notauli slightly visible, incomplete and reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with areolae less wide than those of the metapectal-propodeal disc. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 9). Protarsomeres in following proportions: 7: 1.5: 2: 2.5: 8. Enlarged claw (Fig. 67 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 A) with 5 – 8 very slender lamellae and some bristles. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1F2AF9FF3ED383FB91FD2C.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Drakensberg, Van Reenen, 1 – 22. I. 1927 (NHMUK). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: Orientale Prov. (= Haut-Zaire), Garamba National Park, 2 ♀♀ (MRAC, AMNH). Other material: KENYA: Nairobi Prov., ICIPE Campus, Kasara- ni, 1.22317 ° S 36.89653 ° E, 1600 m, 8 – 15. IV. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Co- peland leg., 1 ♀ (NMK). SOUTH AFRICA: Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC). ZIMBABWE: Harare (= Salisbury), X. 1981, MT, A. Watsham leg., 1 ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1F2AF9FF3ED383FB91FD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1F2AF9FF3ED383FB91FD2C.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1E2AFAFF3ED16FFC9DFC4C.taxon	description	(Fig. 65 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1E2AFAFF3ED16FFC9DFC4C.taxon	description	Description. ♂. Fully winged; body length 1.3 – 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa, metafemur and metatibia partly brown. In specimen from Kenya, 01 ° 06.117 ’ N 34 ° 75.383 ’ E, metaleg testaceous, except coxa brown. Antenna filiform; antennomeres in following proportions: 9: 4: 3.5: 4: 4: 4: 4: 4: 4: 6. Head dull, completely strongly granulate, and slightly rugose (in specimen from Central African Republic, 03 ° 02 ’ 01 ’’ N 16 ° 24 ’ 57 ’’ E, head weakly granulate); frontal line complete; frons with two lateral keels along orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 4; OPL = 2; TL = 4; greatest breadth of lateral ocelli about as long as OL. Mesoscutum, mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Notauli short, reaching about 0.25 × length of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels; central region of propodeal declivity with areolae less wide than those of dorsal region. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Paramere (Fig. 65 C) without distal inner pointed process, with inner proximal lobe with mosaic pattern and not covered with papillae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1E2AFAFF3ED16FFC9DFC4C.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Ngoye Forest, I – IV. 2006, MT, G. Davies leg. (NMSA). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga- Ndoki National Park, Mabéa Bai, 21.4 Km 53 ° NE Bayanga, 03 ° 02 ’ 01 ’’ N 16 ° 24 ’ 57 ’’ E, 510 m, 4 – 5. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♂ (SAMC). KENYA: Rift Valley Prov., Mount Elgon National Park, Top of Endebess Bluff, 01 ° 06.117 ’ N 34 ° 75.383 ’ E, 2630 m, MT, forest edge near small stream, 16 – 30. I. 2006, R. Copeland leg., 1 ♂ (NMK). SOUTH AFRICA: Eastern Cape, Huntly Glen Farm, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, 1065 m, 6. X. 2010 – 18. I. 2011, YPT, Great Fish Thicket, HUN 10 - ACA 1 - Y 05, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1E2AFAFF3ED16FFC9DFC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1E2AFAFF3ED16FFC9DFC4C.taxon	distribution	Distribution. Central African Republic, Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1D2AFAFF3ED253FE14F816.taxon	description	(Figs 65 D, 68 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1D2AFAFF3ED253FE14F816.taxon	description	Description. ♂ (Fig. 68 E). Fully winged; body length 2.2 – 2.9 mm. Head black, except mandible testaceous; anten- na brown; mesosoma black; metasoma brown; legs testaceous, except metacoxa, stalk of metafemur and distal half of metatibia darkened. Antenna filiform; antennomeres in following proportions: 9: 5.5: 6.5: 6.5: 6: 7: 6: 6.5: 6: 10. Head dull, rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 4; TL = 3. Mesoscutum, mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.3 × length of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity with two longitudinal keels, median area shiny, smooth, not rugose, and lateral areas slightly rugose. Forewing darkened; distal part of 2 r-rs & Rs vein much shorter than proximal part (5: 11). Paramere (Fig. 65 D) with distal inner rounded process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1D2AFAFF3ED253FE14F816.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 11.5 km 147 ° SSE of Andranomahy, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375 (CAS). Paratypes: same locality label as holotype, 8 ♂♂ (6 in CAS, 2 in MOLC). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 22 – 28. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 04, 1 ♂ (CAS); Toamasina, 6.9 km NE Ambanizana, Ambohitsitondroina, 15 ° 34 ’ S 50 ° 00 ’ E, 750 m, 2 – 9. XII. 1993, MT, B. Fisher leg., 2 ♂♂ (DEUCD); Toliara, Ivohibe Forest, 55 km N Tolagnaro, 24 ° 34.08 ’ S 47 ° 12.14 ’ E, 200 m, 2 – 4. XII. 2006, MT, B. L. Fisher et al. leg., BLF 15448, 2 ♂♂ (CAS, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1D2AFAFF3ED253FE14F816.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1D2AFAFF3ED253FE14F816.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	description	(Fig. 67 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with colour mostly black; head granulate, with slight irregular striae and areolae; pronotum with posterior surface much shorter than mesoscutum; mesoscutum granulate; forewing hyaline, without dark transverse bands; propodeal declivity rugose, without longitudinal keels; protarsomere 4 at most 0.5 × as long as basal part of 5; protarsomere 5 with basal part much longer than distal part (Fig. 67 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	description	Description. ♀. Fully winged; body length 2.1 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs brown-testaceous. Antenna clavate; antennomeres in following proportions: 10: 4: 4: 4: 4: 5: 4: 4: 5: 6. Head dull, granulate, with slight irregular striae and areolae; frontal line absent; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 7; OL = 3; OOL = 4; OPL = 5; TL = 4; greatest breadth of posterior ocellus shorter than OPL (1: 5). Pronotum dull, transversely striate, except posterior surface almost completely unsculptured; posterior surface very short, shorter than anterior surface and shorter than mesoscutum (3: 12); pronotal tubercle reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Protarsomeres in following proportions: 9: 1: 2: 2: 6. Enlarged claw (Fig. 67 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 B) with basal part much longer than apical part (5: 1), with two medial bristles and one distal lamella. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 22. II – 9. IV. 2010, MT, Amathole Mistbelt Grassland, S. van Noort leg., SAM- HYM-A 035775 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1C2AFCFF3ED02DFBA3FD48.taxon	etymology	Etymology. The species is named nigropictum, because mostly black coloured.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	description	(Figs 66, 67 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with colour almost completely black; head granulate and reticulate rugose; mesoscutum granulate, except unsculptured surface near anterior margin; forewing with two dark transverse bands; pronotum with posterior surface much shorter than mesoscutum; propodeal declivity rugose, without longitudinal keels; protarsomere 4 about 0.5 × as long as basal part of 5.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	description	Description. ♀. Fully winged (Fig. 66); body length 2.2 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 6 – 10 darkened; mesosoma black; metasoma brown; legs testaceous, except meso- and metacoxa, clubs of meso- and metafemur, meso- and metatibia brownish. Antenna clavate; antennomeres in following proportions: 12: 4: 5: 4: 4: 5: 5: 5: 5: 7. Head slightly convex, dull, granulate and reticulate rugose; frontal line complete; frons with two lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 5; OPL = 5; TL = 5; greatest breadth of lateral ocelli shorter than OPL (3: 5). Pronotum anteriorly crossed by slight transverse impression, dull, granulate, with anterior surface rugose and posterior surface sculptured by longitudinal keels; posterior surface much shorter than mesoscutum (5: 16), slightly shorter than anterior surface (5: 7); border between anterior and posterior surface marked by transverse carina; pronotal tubercle reaching tegula. Mesoscutum dull, completely granulate, except anterior surface near pronotum shiny and unsculptured. Notauli very shortly visible near anterior margin of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum dull, sculptured by many longitudinal keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 8). Protarsomeres in following proportions: 5: 2: 2: 4: 16. Enlarged claw (Fig. 67 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 C) with basal part about as long as distal part, twice as long as protarsomere 4 (8: 4), with one row of 7 lamellae; apex with about 7 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	materials_examined	Material examined. Type: ♀ holotype (no registration number): GUINEA: Nzérékoré Region, Mount Nimba, Gouan River, 7 ° 42 ’ N 8 ° 23 ’ W, 514 m, 1 – 15. XII. 1990, FIT, rainforest, L. LeBlanc leg. (PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	distribution	Distribution. Guinea.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1B2AFCFF3ED56FFB6DF8C8.taxon	etymology	Etymology. The species is named nimbense after the collection site on Mount Nimba.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1A2AFDFF3ED340FE2AF874.taxon	description	(Figs 65 E, 68 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1A2AFDFF3ED340FE2AF874.taxon	description	Description. ♂ (Fig. 68 F). Fully winged; body length 2.5 mm. Head black, except mandible testaceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous, except proximal extremity of metacoxa darkened. Antenna filiform; antennomeres in following proportions: 11: 7: 5: 5: 5: 6: 6: 7: 7: 10. Head dull, completely granulate and reticulate rugose; frontal line complete; frons with two lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 9; OL = 4; OOL = 5; OPL = 5; TL = 5; greatest breadth of posterior ocellus shorter than OPL (3: 5). Mesoscutum dull, completely strongly granulate and reticulate rugose, with tracks of notauli reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels, with areolae much less wide than those of metapectalpropodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Paramere (Fig. 65 E) without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1A2AFDFF3ED340FE2AF874.taxon	materials_examined	Material examined. Type: ♂ holotype: BURUNDI: Rusizi National Park, 3.34364 ° S 29.27246 ° E, 774 m, 13 – 20. IV. 2010, MT, near three small trees, degraded bush / grassland, R. Copeland leg. (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1A2AFDFF3ED340FE2AF874.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF1A2AFDFF3ED340FE2AF874.taxon	distribution	Distribution. Burundi.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF192AFFFF3ED3A7FE5DFE54.taxon	description	(Fig. 67 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF192AFFFF3ED3A7FE5DFE54.taxon	description	Description. ♀. Fully winged; body length 1.8 – 2.1 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 9: 4.5: 3.5: 3: 3: 4: 4: 4: 4: 4.5. Head convex, dull, almost completely bare, granulate and with very slight tracks of irregular striae; frontal line absent; no lateral keels on frons; occipital carina complete; POL = 5; OL = 3.5; OOL = 4; OPL = 3; TL = 2. Pronotum anteriorly crossed by slight transverse impression, dull, granulate, with few long setae on posterior surface; posterior surface rugose, much shorter than mesoscutum (2: 11); pronotal tubercle reaching tegula. Mesoscutum of holotype dull, completely granulate, with few long setae; mesoscutum of paratype shiny, unsculptured, except for very slight irregular striae near anterior margin and on lateral regions. Notauli very shortly visible near anterior margin of mesoscutum. Mesoscutellum shiny, slightly granulate. Metanotum shiny, rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 5). Protarsomeres in following proportions: 6: 1.5: 1.5: 1.5: 5. Enlarged claw (Fig. 67 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 D) with one row of four bristles; apex with one lamella. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF192AFFFF3ED3A7FE5DFE54.taxon	materials_examined	Material examined. Types: ♀ holotype: YEMEN: Ar Rujum, 15. I – 9. IV. 2001, MT, A. van Harten leg. (MOLC). Paratype: YEMEN: Al-Kowd, IV – V. 2001, light trap, A. van Harten & S. Al Haruri leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF192AFFFF3ED3A7FE5DFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF192AFFFF3ED3A7FE5DFE54.taxon	distribution	Distribution. Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF172AF1FF3ED078FDB8FE54.taxon	description	(Fig. 65 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF172AF1FF3ED078FDB8FE54.taxon	description	Description. ♂. Fully winged; body length 1.6 – 2.1 mm. Head black, except mandible testaceous; antenna testa- ceous, except antennomeres 5 – 10 darkened; mesosoma black; metasoma brown; legs testaceous, except coxae and clubs of meso- and metafemur partly brown. Antenna filiform; antennomeres in following proportions: 10: 5: 6: 5: 5: 6: 5.5: 5.5: 5.5: 8. Head dull, punctate, unsculptured among punctures; temple and vertex behind ocelli very slightly rugose or with large punctures; vertex with two short oblique keels connecting lateral ocelli to occipital carina; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 4.5; OPL = 2.5; TL = 3. Mesoscutum, mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, re-ticulate rugose, dull; propodeal declivity dull, with two longitudinal keels and with median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 8). Paramere (Fig. 65 F) without distal inner process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF172AF1FF3ED078FDB8FE54.taxon	materials_examined	Material examined. Type: ♂ holotype: YEMEN: 12 km NW of Manakhah, 3. VII – 21. VIII. 2001, MT, A. van Harten leg. (MOLC). Other material: SOUTH AFRICA: Western Cape, Grootvadersbosch Nature Reserve, 33 ° 58.888 ’ S 20 ° 48.885 ’ E, 454 m, 29. III – 23. VII. 2010, YPT, Afromontane Forest, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF172AF1FF3ED078FDB8FE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF172AF1FF3ED078FDB8FE54.taxon	distribution	Distribution. South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF162AF2FF3ED17DFE06FD74.taxon	description	(Figs 69 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF162AF2FF3ED17DFE06FD74.taxon	description	Description. ♂. Fully winged; body length 2.0 – 2.2 mm. Head black, except mandible testaceous (with teeth brown); antenna testaceous; mesosoma and metasoma black; tegula and legs testaceous. Antenna filiform; antennomeres in following proportions: 20: 11: 13: 13: 11: 11: 11: 12: 12: 19. Head shiny, strongly punctate, unsculptured among punctures; frontal line present; frons with two lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 2; TL = 2. Mesoscutum, mesoscutellum and metanotum shiny, smooth, very slightly punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.4 – 0.5 × length of mesoscutum. Mesopleuron and metapleuron rugose, with posterior surface unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels, median area shiny, smooth and sculptured by some keels near margins, lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Paramere (Figs 69 A, B) without distal inner process, with large dorsal proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF162AF2FF3ED17DFE06FD74.taxon	materials_examined	Material examined. Type: ♂ holotype: SEYCHELLES: Mahé Island, top of Mount Sebert, about 2000 ft., 16. I. 1909 (NHMUK). Other material: SEYCHELLES: Mahé Island, Mont Copolia, 04 ° 39.04 ’ S 55 ° 27.30 ’ E, 520 m, 8 – 11. II. 2010, MT in forest, BLF 24027, B. L. Fisher et al. leg., 5 ♂♂ (3 in CAS, 2 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF162AF2FF3ED17DFE06FD74.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF162AF2FF3ED17DFE06FD74.taxon	distribution	Distribution. Seychelles.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	description	(Figs 67 E, 71, 72 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with forewing with one dark spot around 2 r-rs & Rs vein (Fig. 58 A); propodeal declivity with two strong longitudinal keels and median area mostly smooth, with keels only around margins; protarsomere 4 at most 0.5 × as long as basal part of 5.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	description	Description. ♀ (Figs 71, 72 A). Fully winged; body length 2.7 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 9 – 10 brown; mesosoma black, except tegula testaceous; metasoma brown; legs brown. Antenna clavate; antennomeres in following proportions: 13: 5: 5: 5: 4: 5: 4: 5: 5: 7.5. Head (Fig. 71 C) convex, shiny, punctate and sculptured by numerous irregular striae; frons with two lateral longitudinal keels near orbits and directed towards antennal toruli; vertex with two longitudinal keels connecting lateral ocelli to occipital carina; frontal line complete; occipital carina complete; POL = 4; OL = 4; OOL = 3; OPL = 3; TL = 3; greatest breadth of lateral ocelli shorter than OPL (2: 3). Pronotum shiny, unsculptured, except transverse keels on anterior surface; pronotal tubercle reaching tegula; posterior surface of pronotum much shorter than mesoscutum (5: 12). Mesoscutum, mesoscutellum and metanotum (Fig. 71 A) shiny, unsculptured. Notauli incomplete, reaching approximately 0.3 × length of mesoscutum. Mesopleuron shiny, unsculptured. Metapleuron sculptured by slight transverse keels. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity with two complete longitudinal keels, median area shiny, unsculptured and lateral areas shiny and very slightly rugose. Forewing (Fig. 71 D) hyaline, with one dark spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 8). Protarsomeres in following proportions: 7: 2: 2: 2: 10; protarsomere 3 produced into hook. Enlarged claw (Fig. 67 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 E) with basal part much longer than distal part (7: 3), with two rows of six bristles, in addition to setae; apex with 12 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19444): MADAGASCAR: Fianarantsoa, Ranomafana Na- tional Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 16 – 26. II. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 54, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF152AF2FF3ED50BFB15F8C8.taxon	etymology	Etymology. The species is named pseudohova, because of its similarity to Anteon hova Benoit.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	description	(Fig. 69 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	description	Description. ♂. Fully winged; body length 2.0 mm. Head black, except mandible testaceous (with teeth brown); antenna brown; mesosoma and metasoma black; legs testaceous, except metacoxa brown. Antenna filiform; antennomeres in following proportions: 11: 5: 5: 5.5: 6: 7: 7: 6: 6: 8. Head completely reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 6; OPL = 2; TL = 2. Mesoscutum dull, with anterior half reticulate rugose and posterior half unsculptured or slightly granulate. Notauli incomplete, reaching about 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc completely reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 8). Paramere (Fig. 69 C) without distal inner pointed process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	materials_examined	Material examined. Type: ♂ holotype: LA RÉUNION: Mare Longue (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	distribution	Distribution. La Réunion.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF142AF3FF3ED35FFBE6F855.taxon	discussion	Remarks. The record from Mare hongue (Olmi 1987 a) is erroneous, because of a misinterpretation of the handwritten locality label of the holotype; the correct type locality is Mare Longue.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF132AF5FF3ED115FE1FFC4C.taxon	description	(Fig. 67 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF132AF5FF3ED115FE1FFC4C.taxon	description	Description. ♀. Fully winged; body length 3.6 – 4.4 mm. Head reddish, except wide brown-black spot on vertex; oc- casionally head black, except clypeus reddish; mandible testaceous; antenna testaceous; mesosoma of holotype reddish, except brown nuances on mesoscutum, mesoscutellum and metanotum and black metapectal-propodeal disc; occasionally mesoscutum, mesoscutellum, metanotum, mesopleuron and metapleuron completely black; metasoma brown-testaceous; tegula testaceous; legs testaceous. Antennomeres in following proportions: 16: 7: 8: 6: 6: 7: 8: 7: 7: 10. Head dull, completely reticulate rugose; frontal line present; occipital carina complete; POL = 10; OL = 7; OOL = 6; OPL = 8; TL = 3. Pronotum setose, shiny, reticulate rugose; posterior surface shorter than mesoscutum (8: 20); pronotal tubercle reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli absent. Mesoscutellum dull or shiny, slightly or strongly granulate, with some punctures. Metanotum reticulate rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (5: 11). Protarsomeres in following proportions: 12: 2.5: 3: 4: 14; protarsomere 4 produced into hook, approximately 0.3 as long as basal part of 5. Enlarged claw (Fig. 67 F) very long, longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 67 F) with basal part much longer than apical part, with some bristles and 5 – 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF132AF5FF3ED115FE1FFC4C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 10 – 31. VII. 1923, R. E. Turner leg. (NHMUK). Other material: SOUTH AFRICA: Eastern Cape, same locality label as holotype, 1 ♀ (AMNH); Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23 ’ E, 460 m, 25 – 31. VIII. 1994, Mesic Mountain Fynbos, Protea coronata dominated, MT, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF132AF5FF3ED115FE1FFC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF132AF5FF3ED115FE1FFC4C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	description	(Figs 70 A, 72 B, 73)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	diagnosis	Diagnosis. Macropterous ♀ of Anteon with propodeal declivity with two incomplete longitudinal keels, reaching about 0.5 × length of propodeal declivity; protarsomere 4 longer than basal part of 5; protarsomere 5 (Fig. 70 A) with inner side curvilinear, with distal region distinct from median and proximal regions, with basal part much shorter than apical part.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	description	Description. ♀ (Figs 72 B, 73). Fully winged; body length 2.3 mm. Head ferruginous-dark; antenna testaceous; mesosoma ferruginous-dark, except metapectal-propodeal disc black; petiole black; metasoma brown-testaceous; legs testaceous. Antenna clavate; antennomeres in following proportions: 11: 6: 8: 5: 4: 5: 5: 5: 5: 7. Head (Fig. 73 C) dull, sculptured by many slightly irregular keels; frontal line complete; frons with two slight lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 4; OL = 4; OOL = 3; OPL = 4; TL = 6; greatest breadth of lateral ocelli shorter than OPL (2.5: 4). Pronotum flat, shiny, sculptured by many very slight transverse striae, with anterior surface very short, shorter than posterior surface (2: 11); posterior surface shorter than mesoscutum (11: 14), with anterior and lateral margins sharp; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured (Fig. 73 A). Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two incomplete longitudinal keels reaching about 0.5 × length of propodeal declivity. Forewing (Fig. 73 D) with one small dark spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein shorter than proximal part (4.5: 9). Protarsomeres in following proportions: 6: 2: 4: 9: 18. Protarsomere 2 produced into hook. Protarsomere 4 longer than basal part of 5. Enlarged claw (Fig. 70 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 70 A) with inner side curvilinear, with distal region distinct from median and proximal regions, with basal part much shorter than apical part, with one row of 33 lamellae; apex with about 9 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19445): MADAGASCAR: Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 15. II – 6. III. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 03, M. E. Irwin & R. Harin’Hala leg., 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF122AF6FF3ED253FB56FE98.taxon	etymology	Etymology. The species is named sakalavense after the collection site in Sakalava Beach.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF112AF7FF3ED3DEFDB4FB94.taxon	description	(Fig. 69 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF112AF7FF3ED3DEFDB4FB94.taxon	description	Description. ♂. Fully winged; body length 2.2 – 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous (antenna totally testaceous in specimen from Kenya, Funzi Island); mesosoma and metasoma black; legs testaceous, except metacoxa partly black or darkened and club of metafemur brown or darkened. Antenna filiform; antennomeres in following proportions: 11: 6: 7: 6: 5.5: 6: 6: 6: 6: 8. Head dull, granulate and strongly reticulate rugose; frontal line complete; occipital carina complete; POL = 8; OL = 4; OOL = 5; OPL = 4; TL = 3; greatest breadth of lateral ocelli about as long as TL. Mesoscutum dull, granulate and irregularly rugose. Notauli hardly visible, absent or incomplete and reaching about 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels and median area sculptured by areolae less wide than those of the lateral areas (except large areolae along margins); in specimen from Kenya, Funzi Island, median area almost completely sculptured by large areolae. In specimen from Kenya, 4.32664 ° S 39.52462 ° E, propodeal declivity with two incomplete longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 9). Paramere (Fig. 69 D) without distal inner process and with proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF112AF7FF3ED3DEFDB4FB94.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Louwsberg, Sanyati Farm, 1090 m, 27 ° 34 ‘ S 31 ° 17.9 ‘ E, 1 – 24. VI. 2006, MT, M. Mostovski leg. (NMSA). Other material: KENYA: Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 17 – 31. V. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (NMK); same locality label, 27. VI – 11. VII. 2011, 1 ♂ (NMK); same locality label, 3 – 17. X. 2011, 1 ♂ (MOLC); Coast Prov., Taita Hills, Ngangao Forest, 3.36100 ° S 38.34186 ° E, 1848 m, 24. VII – 7. VIII. 2001, MT, indigenous forest, R. Copeland leg., 2 ♂♂ (NMK); Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 24. VII – 7. VIII. 2011, MT, next to small forest pond, R. Copeland leg., 1 ♂ (NMK); same locality label, 16 – 30. X. 2011, 1 ♂ (NMK); Coast Prov., Funzi Island, 4.57749 ° S 39.43825 ° E, near sea level, 4 – 10. VII. 2012, MT, near Funzi workshop, ICIPE / NMK Funzi Island Expedition leg., 4 ♂♂ (NMK); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 30. V – 19. VI. 2013, 6 m MT, indigenous forest, R. Copeland leg., 8 ♂♂ (7 in NMK, 1 in MOLC); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 9 – 23. VIII. 2015, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); same locality label 5 – 19. XI. 2015, 1 ♂ (NMK); Coast Prov., Kasigau Mtn., 3.82700 ° S 38.64875 ° E, 1065 m, 19. X – 2. XI. 2011, MT, indigenous forest, next to campsite in forest, R. Copeland leg., 2 ♂♂ (NMK); Coast Prov., Kasigau Mtn., 3.82080 ° S 38.64178 ° E, 737 m, 7 – 21. IX. 2011, MT, woodland with grass, R. Copeland leg., 2 ♂♂ (NMK); Coast Prov., Kasigau Mtn., 3.82700 ° S 38.64875 ° E, 1065 m, 21. IX – 5. X. 2011, MT, indigenous forest, next to campsite in forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 13 – 27. XII. 2011, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kiri- miri Forest, near top of hill, 0.42563 ° S 37.54460 ° E, 1664 m, 12 – 26. V. 2011, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, 3 – 17. IX. 2011, MT, inside indigenous forest, R. Copeland leg., 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF112AF7FF3ED3DEFDB4FB94.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF112AF7FF3ED3DEFDB4FB94.taxon	distribution	Distribution. Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	description	(Figs 69 E, 72 C – F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	description	Description. ♂ (Figs 72 C – F). Fully winged; body length 1.2 – 2.3 mm. Head black, except mandible testaceous; an- tenna testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 8: 4: 4: 4: 4: 5: 5: 5: 5: 7. Head granulate and reticulate rugose (rugosity hardly visible); frontal line complete; frons with two hardly visible lateral keels along orbits directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 3.5; OPL = 2.5; TL = 2.5; greatest breadth of posterior ocellus slightly shorter than OPL (2: 2.5). Mesoscutum shiny, alutaceous, with lateral regions and occasionally median region granulate; area near anterior margin rugose. In specimen from Kenya, 2.07846 ° S 38.22530 ° E, mesoscutum punctate, unsculptured among punctures, rugose near anterior margin. Notauli incomplete, reaching about 0.4 × length of mesoscutum (in specimen from Uganda, 0 ° 33.871 ’ S 30 ° 21.355 ’ E, notauli invisible among areolae located near anterior margin of mesoscutum; in specimen from Kenya, 2.07846 ° S 38.22530 ° E, notauli incomplete, reaching about 0.25 × length of mesoscutum). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc completely reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands or spots; distal part of 2 rrs & Rs vein much shorter than proximal part (2: 6). Paramere (Fig. 69 E) with distal inner pointed process; proximal membranous process long, with mosaic sculpture and many sensorial setae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 30. V – 19. VI. 2013, 6 m MT, indigenous forest, R. Copeland leg. (NMK). Paratypes: KENYA: same locality label as holotype, 5 ♂♂ (NMK); Coast Prov., Kasigau Mtn., 3.82700 ° S 38.64875 ° E, 1065 m, 19. X – 2. XI. 2011, MT, indigenous forest, next to campsite in forest, R. Copeland leg., 1 ♂ (MOLC); Coast Prov., Mrima Hill Forest 4.48576 ° S 39.25845 ° E, 212 m, 17 – 31. X. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Gede Forest, 3.30946 ° S 40.01941 ° E, 19 m, 27. XI – 11. XII. 2011, indigenous forest, MT, secondary forest, R. Copeland leg., 3 ♂♂ (2 in NMK, 1 in MOLC). UGANDA: Western Region, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 33.871 ’ S 30 ° 21.355 ’ E, 1495 m, 12 – 26. VIII. 2008, MT, secondary mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 024095, SAM-HYM-A 024103, 2 ♂♂ (SAMC). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 23. V. 2001, sweep, lowland rainforest, S. van Noort leg., 2 ♂♂ (SAMC). KENYA: Coast Prov., Gede Forest, 3.30946 ° S 40.01941 ° E, 19 m, 22. VIII – 5. IX. 2011, indigenous forest, MT, secondary forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 27. XI – 11. XII. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 20. IX – 4. X. 2014, MT, indigenous forest, R. Copeland leg., 2 ♂♂ (NMK); same local- ity label, 27. XII. 2014 – 10. I. 2015, 1 ♂ (NMK); same locality label, 13 – 27. XII. 2014, 1 ♂ (NMK); same locality label, 1 – 15. XI. 2014, 1 ♂ (NMK); same locality label, 11 – 25. X. 2015, 2 ♂♂ (NMK); same locality label, 15 – 29. XI. 2014, 3 ♂♂ (2 in NMK, 1 in MOLC); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg., 1 ♂ (NMK). SOUTH AFRICA: Gauteng, [no locality], 20 – 27. I. 2006, MT, R. Wharton leg., 1 ♂ (TAMU); KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, MT, Maputaland Coastal Belt, SA 10 - MAB-M 01, 4 ♂♂ (3 in SAMC, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	distribution	Distribution. Central African Republic, Kenya, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF102A88FF3ED22BFC3CFC4C.taxon	discussion	Remarks. There is only a single morphological difference between A. semajanna and A. whartoni Olmi 2011: in A. semajanna, the propodeal declivity does not have longitudinal keels; A. whartoni has two longitudinal keels or only one keel on the left. It is possible that the two species are conspecific.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6F2A89FF3ED253FDB4FEC4.taxon	description	(Figs 70 B, 74 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6F2A89FF3ED253FDB4FEC4.taxon	description	Description. ♀ (Figs 74 A, B). Fully winged; body length 2.9 – 5.0 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 5 – 10 or 6 – 10 brown; mesosoma black; metasoma brown; tegula testaceous; legs testaceous or testaceous-ferruginous. Antennomeres in following proportions: 15: 6: 8: 5.5: 6: 6: 6: 6: 6: 7. Head dull, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 9; OL = 6; OOL = 5; OPL = 8; TL = 3; greatest breadth of lateral ocelli longer than TL (4: 3). Pronotum shiny, reticulate rugose, except posterior margin smooth; posterior surface shorter than mesoscutum (9: 22); pronotal tubercle reaching tegula. Mesoscutum dull, granulate and reticulate rugose (occasionally reticulate rugose and not granulate). Notauli absent in holotype; in large specimen (5.0 mm) from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, notauli reaching about 0.5 × length of mesoscutum). Mesoscutellum shiny, sculptured by large punctures. Metanotum sculptured by large punctures, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels and median area as rugose as lateral areas. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 11). Protarsomeres in following proportions: 12: 2.5: 3: 5: 13; protarsomere 4 produced into hook, approximately 0.5 × as long as basal part of 5 (5: 11). Enlarged claw (Fig. 70 B) very long, slightly longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 70 B) with basal part much longer than apical part (11: 2), with some bristles and 5 – 6 distal lamellae. Arolium very large, about as large as protarsomere 5. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6F2A89FF3ED253FDB4FEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Shimba Hills National Park, near artificial pond, 04 ° 22.752 ’ S 39 ° 43.197 ’ E, 335 m, MT, mixed grassland-shrubland, 27. XII. 2005 – 10. I. 2006, R. Copeland leg. (NMK). Other material: KENYA: Coast Prov., Kasigau Mtn., 3.82080 ° S 38.64178 ° E, 737 m, 7 – 21. IX. 2011, MT, woodland with grass, R. Copeland leg., 1 ♀ (MOLC); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 10 – 24. I. 2012, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, YPT, Maputaland Coastal Belt, SA 10 - MAB-Y 01, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6F2A89FF3ED253FDB4FEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6F2A89FF3ED253FDB4FEC4.taxon	distribution	Distribution. Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8AFF3ED6D7FE1FF9C9.taxon	description	(Fig. 70 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8AFF3ED6D7FE1FF9C9.taxon	description	Description. ♀. Fully winged; body length 2.1 – 2.8 mm. Head black, except mandible testaceous; antenna tes- taceous, except antennomeres 7 – 10 brownish; mesosoma and metasoma black; legs testaceous, except clubs of femora, metacoxa, meso- and metatibia darkened. Antenna clavate; antennomeres in following proportions: 10: 5: 6: 4.5: 4.5: 5: 5: 5.5: 5: 7. Head completely granulate and reticulate rugose (in specimen from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, frons granulate and very slightly rugose); frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 9; OL = 6; OOL = 5; OPL = 5.5; TL = 4; greatest breadth of lateral ocelli shorter than OL (2: 6). Pronotum with anterior surface rugose; posterior surface less than 0.5 × as long as mesoscutum (6: 15), rugose, with posterior half unsculptured. Mesoscutum shiny, smooth, punctate, unsculptured among punctures. In specimen from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, mesoscutum very slightly granulate. Notauli reaching approximately 0.3 × length of mesoscutum; in paratype and specimen from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, notauli almost completely absent; in specimens from South Africa, 32 ° 57.512 ’ S 18 ° 06.728 ’ E, notauli reaching about 0.2 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron with anterior half reticulate rugose and posterior half smooth, punctate, unsculptured among punctures; metapleuron completely reticulate rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, more than 0.5 × as long as mesoscutellum + metanotum (9: 14); propodeal declivity reticulate rugose, without longitudinal keels (occasionally with two narrow longitudinal keels). Forewing with dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 8). Protarsomeres in following proportions: 10: 2: 2: 2.5: 8.5; protarsomere 4 less than 0.5 × as long as basal part of 5. Enlarged claw (Fig. 70 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 70 C) with some medial and proximal bristles and five distal lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8AFF3ED6D7FE1FF9C9.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, IX – XII. 1997, MT, S. van Noort leg., SAM-HYM-A 024103 (SAMC). Paratypes: same locality label as holotype, SAM-HYM-A 024103, 1 ♀ (SAMC), 1 ♀ (MOLC). Other material: SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 3. IX – 30. X. 2009, YPT, WTB 09 - GRA 1 - Y 01, Amathole Mistbelt Grassland, S. van Noort leg., 3 ♀♀ (2 in SAMC, 1 in MOLC); Western Cape, West Coast Fossil Park, 4 km 290 ° E Langebaanweg, 32 ° 57.512 ’ S 18 ° 06.728 ’ E, 20 – 27. XI. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 5. XI. 2009 – 27. II. 2010, MT, Afromontane Forest, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. X – 16. XI. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC); same locality label, 16. XII. 1999 – 16. I. 2000, 1 ♀ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 31. X – 28. XI. 1997, MT, West Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 20. XII. 2012 – 24. I. 2013, MT, Mesic Mountain Fynbos, BH 12 - FYN 3 - M 05, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8AFF3ED6D7FE1FF9C9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8AFF3ED6D7FE1FF9C9.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8BFF3ED0EEFD82FDB8.taxon	description	(Fig. 69 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8BFF3ED0EEFD82FDB8.taxon	description	Description. ♂. Fully winged; body length 2.9 – 3.4 mm. Head black, except mandible testaceous; antenna tes- taceous-brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa partly darkened. Antenna filiform; antennomeres in following proportions: 18: 7: 7: 7: 8: 8: 8: 9: 8.5: 11.5. Head dull, granulate and reticulate rugose; frontal line complete; occipital carina complete; POL = 10; OL = 3; OOL = 5; OPL = 4; TL = 5. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.25 – 0.40 × length of mesoscutum (0.25 × in holotype). Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum rugose. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity without longitudinal keels, reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 rrs & Rs vein much shorter than proximal part (5: 12). Paramere (Fig. 69 F) with distal inner rounded process; in holotype, inner side of paramere with proximal mosaic sculpture; in specimen from South Africa, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, mosaic sculpture of paramere absent. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8BFF3ED0EEFD82FDB8.taxon	materials_examined	Material examined. Type: ♂ holotype: UGANDA: Central Region, Mubende District, Mulange, XI. 1922, R. Dummer leg., SAM-HYM-PO 03772 (SAMC). Other material: SOUTH AFRICA: KwaZulu-Natal, Karkloof, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, 1325 m, 25. VII – 25. IX. 2005, MT, M. Mostovski leg., 1 ♂ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8BFF3ED0EEFD82FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6D2A8BFF3ED0EEFD82FDB8.taxon	distribution	Distribution. South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8BFF3ED4DFFE14F9CA.taxon	description	(Figs 74 C, 75 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8BFF3ED4DFFE14F9CA.taxon	description	Description. ♂ (Fig. 74 C). Fully winged; body length 1.6 – 1.9 mm. Head black, except mandible testaceous; an- tenna brown, except scape and pedicel slightly testaceous; mesosoma black; metasoma brown; legs of holotype testaceous, except metafemur and metatibia darkened; legs of paratype brown, except tarsi testaceous (with tarsomere 5 brown). Antenna filiform, with setae partly longer than breadth of antennomeres; antennomeres in following proportions: 8.5: 4: 4: 4: 4: 4.5: 4.5: 4.5: 4.5: 7.5. Head dull, granulate and reticulate rugose; frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 6; OL = 3; OOL = 4; OPL = 2; TL = 2; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum dull, granulate, with anterior region near pronotum rugose. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity without longitudinal keels, completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Paramere (Fig. 75 A) with distal inner rounded lobe not covered with papillae, with few papillae situated on inner margin (Fig. 75 B). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8BFF3ED4DFFE14F9CA.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohipa- rara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13 (CAS). Paratype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 1 ♂ (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 26. IV – 12. IV. 2001, MT, MA- 01 - 01 D- 09, R. Harin’Hala leg., 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8BFF3ED4DFFE14F9CA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8BFF3ED4DFFE14F9CA.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8DFF3ED0EEFD73FCD8.taxon	description	(Figs 75 C – E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8DFF3ED0EEFD73FCD8.taxon	description	Description. ♂. Fully winged; body length 1.6 – 2.4 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations and protibia testaceous. In specimen from Madagascar, 12 ° 20 ’ S 49 ° 15 ’ E, antenna testaceous-brown; legs yellow-testaceous, except basal extremity of metacoxa brown. In specimen from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, legs testaceous, except part of metacoxa and stalk of metafemur black. Antenna filiform; antennomeres in following proportions: 6.5: 2.5: 4: 4.5: 4.5: 5: 4.5: 4: 4.5: 5.5. Head dull, punctate, unsculptured among punctures (occasionally vertex slightly rugose behind and on sides of lateral ocelli); frons with track of complete frontal line, without lateral keels near orbits directed towards antennal toruli; occipital carina complete; holotype with POL = 6; OL = 3; OOL = 3.5; OPL = 3; TL = 3; in specimen from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, POL = 4; OL = 2; OOL = 6; OPL = 3; TL = 3; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum shiny, smooth, finely punctate, unsculptured among punctures. Notauli incom- plete, reaching approximately 0.4 – 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity with two irregular longitudinal keels, with median area almost completely shiny, smooth, not rugose and lateral areas rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Paramere (Fig. 75 C) without distal inner process, with slender dorsal membranous process. Occasionally paramere with small rounded or slightly pointed distal inner process (Figs 75 D, E). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8DFF3ED0EEFD73FCD8.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ N 18 ° 23.5 ’ E, 460 m, 15 – 23. II. 1994, Mesic Mountain Fynbos on Sandstone, Protea dominated, MT, SAM-HYM-PO 03788, S. van Noort leg. (SAMC). Paratype: SOUTH AFRICA: Western Cape, Hermanus, Fernkloof Nature Reserve, Die Mond se Kop, above Voëlklip, 60 m, 19 – 20. II. 1994, Mesic Mountain Fynbos, MT, S. van Noort leg., SAM-HYM-P 003787, 1 ♂ (SAMC). Other material: MADAGAS- CAR: Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 4 ♂♂ (3 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 14 – 21. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 12, 1 ♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 A- 07, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. II. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 15, 1 ♂ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 25. X – 4. XI. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 66, 1 ♂ (MOLC). SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 1 ♂ (SAMC); Western Cape Prov., Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 26. X. 2010 – 26. I. 2011, MT, Afromontane Forest, S. van Noort leg., GVB 10 - FOR 1 - M 07, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8DFF3ED0EEFD73FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6C2A8DFF3ED0EEFD73FCD8.taxon	distribution	Distribution. Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6A2A8DFF3ED5FFFD86F85E.taxon	description	(Figs 75 F, 77 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6A2A8DFF3ED5FFFD86F85E.taxon	description	Description. ♂. Fully winged; body length 2.5 – 2.9 mm. Head black, except mandible testaceous; antenna of ho- lotype brown; antenna of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, completely testaceous-darkened; mesosoma black; metasoma brown; pro- and mesolegs testaceous; metalegs of holotype testaceous, except stalks of femora, tibiae and tarsi darkened; metalegs of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, completely testaceous. Antenna filiform; antennomeres in following proportions: 13: 7: 12: 11: 11: 11: 10: 10: 10: 13. Head dull, reticulate rugose, with anterior half or third of frons sculptured by numerous longitudinal and about parallel keels; clypeus sculptured by numerous longitudinal and parallel keels; in specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, clypeus completely smooth and without parallel keels; frontal line complete; occipital carina complete; POL = 7; OL = 4; OOL = 7; OPL = 5; TL = 4. Mesoscutum of holotype shiny, smooth, slightly granulate and punctate, with some little distinct irregular keels; mesoscutum of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, punctate and unsculptured among punctures; mesoscutum of specimen from Madagascar, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, strongly granulate. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 12). Paramere with (Fig. 75 F) or without (Fig. 77 A) small distal inner rounded process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6A2A8DFF3ED5FFFD86F85E.taxon	materials_examined	Material examined. Type: ♂ holotype: TANZANIA: Tanga District, W Usambara Mnts., Mazumbai, 04 ° 49 ’ S 38 ° 30 ’ E, 1400 – 1600 m, 11 – 19. XI. 1995, C. E. Griswold, N. Scharff & D. Ubick leg., 1546, MT, in forest (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 21 – 26. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 2 ♂♂ (CAS, MOLC); Fian- arantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 4 – 12. II. 2002, 1110 m, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 4 ♂♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6A2A8DFF3ED5FFFD86F85E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF6A2A8DFF3ED5FFFD86F85E.taxon	distribution	Distribution. Madagascar, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8EFF3ED6AFFDFCFBF8.taxon	description	(Fig. 70 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8EFF3ED6AFFDFCFBF8.taxon	description	Description. ♀. Fully winged; body length 2.7 – 3.0 mm. Head black, except mandible testaceous; antenna tes- taceous, except antennomeres 7 – 10 brown; mesosoma and metasoma black; tegula testaceous; legs testaceous, except metacoxa, metafemur and metatibia partly brown. Antenna clavate; antennomeres in following proportions: 15: 6: 6: 6: 6: 6: 5: 5: 5: 8. Head dull, granulate and reticulate rugose; frontal line complete; frons with two lateral keels near orbits directed towards antennal toruli; vertex with two oblique keels connecting inner side of lateral ocelli to occipital carina; occipital carina complete; POL = 7; OL = 4; OOL = 7; OPL = 5; TL = 4. Pronotum humped, dull, granulate, with numerous transverse keels; posterior surface shorter than anterior surface (7: 11), about 0.5 × as long as mesoscutum (8: 16); pronotal tubercle reaching tegula. Mesoscutum shiny, smooth, granulate. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum sculptured by some longitudinal keels. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, sculptured by strong transverse keels, not reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 8). Protarsomeres in following proportions: 5: 3: 2.5: 6: 20. Enlarged claw (Fig. 70 D) very long, longer than protarsomere 5, with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 70 D) with basal part shorter than apical part (8: 12), with one row of 9 – 11 lamellae; apex with 8 – 9 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8EFF3ED6AFFDFCFBF8.taxon	materials_examined	Material examined. Type: ♀ holotype: UGANDA: Central Region, Masaka District, Kawanda, XI. 1942, T. H. C. Taylor leg. (NHMUK). Other material. KENYA: Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 24. II – 10. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8EFF3ED6AFFDFCFBF8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8EFF3ED6AFFDFCFBF8.taxon	distribution	Distribution. Kenya, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8FFF3ED29FFD17FD9C.taxon	description	(Fig. 70 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8FFF3ED29FFD17FD9C.taxon	description	Description. ♀. Fully winged; body length 2.6 – 4.0 mm. Head black, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous; mesosoma black, except lateral margins of pronotum testaceous; metasoma brown; legs testaceous. Specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, darker, with head totally black, except mandible brown; antenna brown; mesosoma black; metsoma brown; legs brown. Antenna clavate; antennomeres in following proportions: 11: 5.5: 6: 4.5: 5: 5: 5: 4: 5: 7. Head dull, granulate; frontal line complete, partly continuing behind anterior ocellus (frontal line incomplete, only present in posterior half of frons, in specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E); frons with two lateral keels along orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 5; OOL = 5 OPL = 5; TL = 5; greatest breadth of lateral ocelli shorter than OL (2: 5). Pronotum smooth, with slight transverse anterior impression; anterior surface shiny, punctate, unsculptured among punctures; posterior surface approximately as long as mesoscutum (12.5: 12), dull, granulate, except few transverse keels situated at anterior margin (keels forming raised transverse carina); pronotal tubercle reaching tegula. Anterior surface of pronotum shorter than posterior surface (7: 12.5). Mesoscutum shiny, granulate, unsculptured medially near anterior margin. Notauli incomplete, reaching about 0.3 × length of mesoscutum (0.2 × in specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity with small areolae, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 6). Protarsomeres in following proportions: 7: 2.5: 3.5: 12.5: 21; protarsomere 2 produced into hook. Enlarged claw (Fig. 70 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 70 E) with two rows of 25 – 35 lamellae; apex very slender, with group of 3 – 5 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8FFF3ED29FFD17FD9C.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Tete Prov., 20 km N Tete, 16 ° 02 ’ S 33 ° 35 ’ E, 260 m, 1. XII. 2005, J. Halada leg. (OLL). Other material: KENYA: Eastern Prov., Kasaala area, Mulu Musingila’s Farm, 13 – 27. XII. 2016, MT, near gully, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 6. X. 2010 – 18. I. 2011, YPT, Camdeboo Escarpment Ticket, S. van Noort leg., PCD 09 - ACA 1 - Y 06, 1 ♀ (SAMC); same locality label, MT, PCD 09 - ACA 1 - M 06, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8FFF3ED29FFD17FD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF692A8FFF3ED29FFD17FD9C.taxon	distribution	Distribution. Kenya, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF682A80FF3ED423FB9FFC20.taxon	description	(Fig. 78 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF682A80FF3ED423FB9FFC20.taxon	description	Description. ♀. Fully winged; body length 3.0 – 3.6 mm. Completely testaceous-reddish or testaceous-yellow, except petiole black; occasionally head and thorax brown-reddish or brown; occasionally metapectal-propodeal disc partly or completely black or brown. In specimen from South Africa, 34 ° 02 ’ S 18 ° 23.5 ’ E, head, mesosoma and metasoma brown-reddish dark; antenna and legs brown. Antennomeres in following proportions: 10: 6: 8: 7: 6: 6: 6: 6: 6: 7. Head shiny, granulate and slightly reticulate rugose (in ♀♀ from South Africa, 34 ° 27.41 ’ S 19 ° 21.39 ’ E and 28 ° 57.60 ’ S 29 ° 13.61 ’ E, head completely granulate and not rugose); frontal line incomplete; frons with two incomplete lateral keels directed towards antennal toruli; occipital carina complete; POL = 5.5; OL = 5; OOL = 6.5 OPL = 6; TL = 6 (occasionally POL = 6; OL = 4; OOL = 4; OPL = 5; TL = 5). Pronotum dull, granulate, with slight transverse anterior impression; posterior surface approximately as long as mesoscutum; pronotal tubercle reaching tegula. Mesoscutum shiny, unsculptured, with slight tracks of irregular keels on sides; in specimens from South Africa, 34 ° 02 ’ N 18 ° 23.5 ’ E, 34 ° 27.41 ’ S 19 ° 21.39 ’ E and 28 ° 57.60 ’ S 29 ° 13.61 ’ E, mesoscutum slightly or strongly granulate. Notauli incomplete, faintly and shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity with small areolae and with or without irregular transverse keels, without longitudinal keels; in specimen from South Africa, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, propodeal declivity with transverse keels situated only near apex. In specimen from Uganda, 0 ° 35.442 ’ S 30 ° 21.741 ’ E, propodeal declivity with two irregular and incomplete longitudinal keels, median area granulate and lateral areas reticulate rugose. In specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, propodeal declivity almost completely unsculptured, shiny. Forewing with two dark transverse bands; distal part of 2 rrs & Rs vein much shorter than proximal part (4: 9). Protarsomeres in following proportions: 8: 2: 3.5: 10: 20. Enlarged claw (Fig. 78 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 A) with two rows of 29 – 40 lamellae; apex with 3 – 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF682A80FF3ED423FB9FFC20.taxon	materials_examined	Material examined. Types: ♀ holotype of A. townesi: SOUTH AFRICA: KwaZulu-Natal, Royal Natal National Park (AEIC); ♀ holotype of A. decellei: DEMOCRATIC REPUBLIC OF THE CONGO: Orientale Prov. (= Haut-Zaire), Garamba National Park (MRAC). Paratype of A. townesi: SOUTH AFRICA: Eastern Cape, Kentonon-Sea, 1 ♀ (AMNH). Other material: KENYA: Eastern Prov., base of Endau Mtn., 1.30026 ° S 38.52805 ° E, 531 m, 28. XI – 12. XII. 2016, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 3 – 17. XII. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 2 ♀♀ (NMK, MOLC). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 2 ♀♀ (SAMC); Eastern Cape, Winter- berg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 22. II – 9. IV. 2010, MT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♀ (SAMC); Gauteng, Onderstepoort, 1986, schlupfalle, I. Pajor leg., 1 ♀ (ZSM); KwaZulu-Natal, Louwsberg, Sanyati Farm, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 18 – 20. IV. 2007, YPT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, La Mercy, Site 1, 29 ° 37.41 ’ S 31 ° 06.454 ’ E, 85 m, 6. XII. 2006, YPT, Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Cathedral Peak Natural Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 18. XI. 2006 – 26. II. 2007, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Itala Game Reserve, Ngubhu House, 27 ° 32.99 ’ S 31 ° 13.34 ’ E, 720 m, 30. XI – 7. XII. 1999, YPT, closed Euclea / Acacia Natal Lowveld Bushveld, S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, YPT, Maputa- land Coastal Belt, SA 10 - MAB-Y 01, 1 ♀ (SAMC); Western Cape, Stellenbosch, Jonkershoek Reserve, 1 ♀ (PMA); Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 7 – 21. IV. 1995, Site 2, Mesic Mountain Fynbos, Protea coronata dominated, MT, SAM-HYM-PO 20893, S. van Noort leg., 1 ♀ (SAMC); same locality label, 6. XII. 1994, sweep, 1 ♀ (SAMC); Western Cape, Constantiaberg, Donkerboskloof, 34 ° 02.28 ’ S 18 ° 23.75 ’ E, 370 m, 10. III. 1995, sweep, Kloof forest, S. van Noort leg., 2 ♀♀ (SAMC, MOLC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 1 ♀ (SAMC); Western Cape, Cape of Good Hope Nature Reserve, Booi se Skerm, 34 ° 18.27 ’ S 18 ° 27.64 ’ E, 24. II. 1998, sweep, strandveld, S. van Noort leg., 2 ♀♀ (SAMC, MOLC); Western Cape, Hermanus, Fernkloof Nature Reserve, Mosselberg, south slope, 100 m, 4 – 5. II. 1995, MT, Mesic Mountain Fynbos, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Table Mountain National Park, Tokai S, Prinskasteel, Site 29, 34 ° 03.561 ’ S 18 ° 24.054 ’ E, 19. I. 2009, Pitfall Trap 8, Afrotemperate Forest, Fynbos, C. Uys leg., 1 ♀ (SAMC); Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - Y 09, 1 ♀ (SAMC). UGANDA: Western Region, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 35.442 ’ S 30 ° 21.741 ’ E, 1465 m, 10. VIII. 2008, sweep, primary mid-altitude rainforest, near stream, S. van Noort leg., 1 ♀ (SAMC). ZIMBABWE: Harare, 1 ♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF682A80FF3ED423FB9FFC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF682A80FF3ED423FB9FFC20.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF672A80FF3ED5B7FE1FF8EA.taxon	description	(Fig. 78 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF672A80FF3ED5B7FE1FF8EA.taxon	description	Description. ♀. Fully winged; body length 2.9 mm. Head testaceous-reddish; antenna testaceous, except anten- nomeres 7 – 10 darkened; mesosoma testaceous-reddish; petiole and metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 6.5: 4.5: 5: 6: 5: 5: 5: 7.5. Head shiny, sparsely setose, completely strongly reticulate rugose; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 4; OOL = 3; OPL = 5; TL = 2.5; greatest breadth of lateral ocelli about as long as OOL. Pronotum shiny, reticulate rugose, except posterior margin smooth; posterior surface much shorter than mesoscutum (5: 15); pronotal tubercle reaching tegula. Mesoscutum dull, granulate. Notauli slightly visible, incomplete, reaching about 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron and metapleuron dull, reticulate rugose. Metapectal-propodeal disc dull, with a transverse posterior keel, strongly reticulate rugose; propodeal declivity completely strongly reticulate rugose, with two tracks of complete longitudinal keels. Forewing with very dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much shorter than proximal part (4.5: 10). Protarsomeres in following proportions: 11: 2: 2: 3: 10. Protarsomere 4 produced into hook, less than 0.5 × as long as basal part of 5 (3: 8). Enlarged claw (Fig. 78 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 B) with inner side almost rectilinear and four lamellae, in addition to few bristles. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF672A80FF3ED5B7FE1FF8EA.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 12 – 30. VI. 1923, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF672A80FF3ED5B7FE1FF8EA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF672A80FF3ED5B7FE1FF8EA.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A81FF3ED6D7FCA2FBB5.taxon	description	(Fig. 78 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A81FF3ED6D7FCA2FBB5.taxon	description	Description. ♀. Fully winged; body length 1.6 – 2.6 mm. Completely testaceous, except petiole black. Antenna cla- vate; antennomeres in following proportions: 7: 4: 5: 3.5: 3.5: 3.5: 4: 4: 4: 6. Head shiny, smooth, finely setose, punctate, unsculptured among punctures; frontal line complete; occipital carina complete; POL = 3; OL = 4; OOL = 4; OPL = 3.5; TL = 6. Pronotum shiny, smooth, finely setose, punctate, unsculptured among punctures; posterior surface approximately as long as mesoscutum (9: 9); pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, smooth, finely setose, punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.7 – 0.8 × length of mesoscutum. Metapectal-propodeal disc shiny, reticulate rugose; propodeal declivity with two complete longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Protarsomeres in following proportions: 4.5: 1.5: 2: 6: 11.5. Enlarged claw (Fig. 78 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 C) with two rows of 2 – 3 + 13 – 17 lamellae (3 + 13 lamellae in holotype); apex with small group of four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A81FF3ED6D7FCA2FBB5.taxon	materials_examined	Material examined. Types: ♀ holotype: BURKINA FASO: Centre-Ouest Region, Boulkiemdé Prov., Kou- dougou Dept., Saria, S. Traoré leg. (MNHN). Paratypes: same locality label as holotype, 6 ♀♀ (AMNH). Other material: MOZAMBIQUE: Maputo Prov., 5 km E Manhiça, 1 ♀ (AMNH). SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 27. I – 16. II. 2004, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Umtam- vuna Nature Reserve, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A81FF3ED6D7FCA2FBB5.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Burkina Faso: Cicadulina mbila (Naudé); Cicadulina similis China.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A81FF3ED6D7FCA2FBB5.taxon	distribution	Distribution. Burkina Faso, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	description	(Figs 74 D, 76, 77 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	diagnosis	Diagnosis. ♂ of Anteon with head granulate (Fig. 74 D); OPL as long as TL; antenna filiform, with antennomere 4 more than twice as long as broad; mesoscutum granulate; notauli very short, hardly visible near anterior margin of mesoscutum; metapectal-propodeal disc with strong transverse posterior keel; propodeal declivity without longitudinal keels; paramere without distal inner pointed or rounded process, with inner side provided with numerous papillae (Fig. 77 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	description	Description. ♂ (Figs 74 D, 76). Fully winged; body length 1.5 – 1.6 mm. Head black, except mandible tes- taceous; antenna brown-testaceous; mesosoma black; metasoma brown; legs testaceous, except part of coxae, club of metafemur and distal half of metatibia brown. Antenna filiform; antennomeres in following proportions: 8: 4: 5: 5: 5: 5: 5: 5: 5: 7; antennomere 4 more than twice as long as broad (5: 2). Head (Fig. 76 C) dull, granulate; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 3; TL = 3; greatest breadth of lateral ocelli shorter than OPL (2: 3). Mesoscutum (Fig. 76 A) dull, granulate. Notauli very short, hardly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels; areolae of propodeal declivity less wide than those of metapectal-propodeal disc. Forewing (Fig. 76 D) hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 5). Paramere (Fig. 77 B) with inner side almost completely provided with numerous papillae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19446): MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 26. I – 5. II. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 14 (CAS). Paratype: KENYA: Nairobi Prov., ICIPE Campus, Kasarani, 1.22338 ° S 36.89655 ° E, 1604 m, 24. II – 3. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	distribution	Distribution. Kenya, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF662A82FF3ED237FF2CFE98.taxon	etymology	Etymology. The species is named tulearense after the collection site of the holotype in Tulear (= Toliara) Prov- ince.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	description	(Figs 74 E, 78 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	description	Description. ♀ (Fig. 74 E). Fully winged; body length 2.4 – 3.4 mm. Completely testaceous-reddish, except petiole black, metanotum and metapectal-propodeal disc partly black, first segment of metasoma partly black (in two ♀♀ from South Africa, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, metasoma completely testaceous). In specimen from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, BH 12 - FYN 3 - M 05, head brown, except some testaceous spots on vertex; antenna testaceous, except antennomeres 1 – 3 brown; mesosoma and metasoma testaceous, except many brown spots; legs testaceous, except coxae, femora and tibiae partly brown. Antennomeres in following proportions: 10: 5: 6: 5: 5: 5: 5: 5: 4.5: 6. Head dull, granulate; frontal line present; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 4; OL = 5; OOL = 5 OPL = 5; TL = 5. Pronotum shiny, setose, slightly granulate; posterior surface much longer than anterior surface and slightly shorter than mesoscutum (10: 11); pronotal tubercle reaching tegula. Mesoscutum shiny, granulate. Notauli incomplete, faintly and shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, with transverse posterior keel, reticulate rugose; propodeal declivity completely granulate, without longitudinal keels, not reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2.5: 7). Protarsomeres in following proportions: 6: 2: 3: 7: 16. Enlarged claw (Fig. 78 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 D) with two rows of 23 – 30 lamellae (30 in holotype); apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, XII. 1921, R. E. Turner leg. (NHMUK). Other material: SOUTH AFRICA: Eastern Cape, Februarie Farm (39.9 km 268 ° W Kirkwood), 33 ° 32.813 ’ S 25 ° 03.091 ’ E, 10 – 12. II. 2001, MT, VB 01 - R 2 N-M 24, Valley bushveld (non-trashed), S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 24. XI – 9. XII. 2003, MT, M. Mostovski leg., 1 ♀ (NMSA); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 2 ♀♀ (SAMC); same locality label, 18. IV – 16. V. 1998, 1 ♀ (SAMC); same locality label, IX – XII. 1997, MT, S. van Noort leg., 6 ♀♀ (5 in SAMC, 1 in MOLC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 5 ♀♀ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 28. XII. 1997 – 21. III. 1998, MT, South Coast Strandveld, 57 m, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 15 ’ S 19 ° 05 ’ E, 16. IV. 1999, pitfall trap, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. I – 16. II. 2000, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.62 ’ S 18 ° 24.26 ’ E, 23. I – 20. II. 1998, YPT, S. van Noort leg., 1 ♀ (SAMC); Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 3 – 31. X. 1997, MT, West Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 58.888 ’ S 20 ° 48.885 ’ E, 454 m, 29. III – 23. VII. 2010, MT, South Langeberg Sandstone Fynbos, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ‘ S 18 ° 23 ‘ E, 460 m, 10 – 17. III. 1995, Mesic Mountain Fynbos, Protea coronata dominated, MT, S. van Noort leg., 2 ♀♀ (SAMC); Western Cape, Goukamma Nature Reserve, Eastern section, near Buffelsbaai, 34 ° 4.566 ’ S 22 ° 57.670 ’ E, 30 m, 29. XII. 1994 – 15. I. 1995, Dune Fynbos, MT, GOU 94 - M 01, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 20. X – 27. XI. 2013, MT, Mesic Mountain Fynbos, BH 12 - FYN 3 - M 14, S. van Noort leg., 1 ♀ (MOLC); same locality label, 10. XII. 2012 – 24. I. 2013, BH 12 - FYN 3 - M 05, 1 ♀ (SAMC); same locality label, 3 – 25. IV. 2013, BH 12 - FYN 3 - M 07, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF652A84FF3ED3C4FB45FC04.taxon	discussion	Remarks. Anteon turneri is very similar to A. townesi Olmi 1984; the unique difference regards the forewing colouration (A. townesi has two dark transverse bands; whereas A. turneri has completely hyaline wings). There is a possibility that these species are conspecific. They were both collected in the same locality (Walker Bay Nature Reserve). The discovery of the ♂♂ of both species should elucidate the species delimitation.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF632A87FF3ED59BFCDEFE70.taxon	description	(Figs 77 C, 78 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF632A87FF3ED59BFCDEFE70.taxon	description	Description. ♀. Fully winged; body length 1.9 – 2.5 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous (in specimens from Uganda, Mali and Zimbabwe), or completely yellow (in specimens from South Africa and Democratic Republic of the Congo); mesosoma and metasoma black; legs brown, except tarsi testaceous (in specimens from Uganda), or more or less yellow (in specimens from Mali, Democratic Republic of the Congo and South Africa). Occasionally (in specimens from South Africa, 29 ° 37.41 ’ S 31 ° 06.454 ’ E) antenna brown, except antennomeres 1 – 4 or 1 – 5 testaceous. In specimens from Mali, Katibougou, antenna brown, except scape and pedicel testaceous. Antenna clavate; antennomeres in following proportions: 6: 3.5: 3.5: 3: 2.5: 3: 3.5: 3.5: 3.5: 4.5. Head dull, granulate, with frons completely or partly granulate and reticulate rugose (occasionally very weakly rugose); frontal line present; frons without lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 3; TL = 4. Pronotum dull, granulate, with few transverse keels; posterior surface shorter than mesoscutum (5: 9); pronotal tubercle reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, reaching about 0.25 × length of mesoscutum; in series of specimens from Tanzania, Mkomazi Game Reserve, few specimens with notauli reaching about 0.25 × length of mesoscutum and other specimens with shorter notauli, slightly visible near anterior margin of mesoscutum; two specimens from Mali, Katibougou, and one specimen from Uganda, Kawanda, with notauli very shortly present near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands, occasionally with very slight dark spot situated across 2 R 1 cell; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Protarsomeres in following proportions: 5: 2: 2: 3: 9. Enlarged claw (Fig. 78 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 E) with one row of 10 – 15 lamellae; apex with 4 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.7 – 1.9 mm. Head black, except mandible testaceous; antenna completely tes- taceous; occasionally antennomeres 3 – 10 slightly darkened; mesosoma and metasoma black; legs completely testaceous; occasionally coxae, femora and tibiae brown. In specimen from South Africa, Ashburton, 9 – 24. XI. 2005, head brown, except irregular testaceous spots on frons, vertex and occiput. Antenna filiform; antennomeres in following proportions: 9: 5: 5: 6: 5: 6: 6: 5: 5: 8. Head dull, completely granulate; frontal line complete; frons without lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 4; OOL = 4; OPL = 2.5; TL = 2.5; in some specimens from Congo, POL = 5; OL = 4; OOL = 5; OPL = 3; TL = 5. Mesoscutum dull, granulate, with some slight areolae near anterior margin. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with or without strong transverse posterior keel, reticulate rugose; propodeal declivity without longitudinal keels, completely reticulate rugose, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Paramere (Fig. 77 C) without distal inner pointed process and proximal membranous process; in specimen from South Africa, Guernsey Farm, inner side of paramere with some transverse folds. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF632A87FF3ED59BFCDEFE70.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Central Region, Masaka District, Kawanda, 16. VII. 1943, T. H. C. Taylor leg. (NHMUK). Paratypes: DEMOCRATIC REPUBLIC OF THE CONGO: DEMOCRATIC RE- PUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert Nat. Park), Mt. Hoyo, 1 ♂ (MRAC); North Kivu, Virunga National Park (= Albert Nat. Park), Mt. Ruwenzori, Kakalari River, affl. Bombi, 1725 m, P. Vanschuytbroeck & H. Synave leg., 2 ♂♂ (AMNH, MRAC); North Kivu, Virunga National Park (= Albert Nat. Park), Mt. Ruwenzori, Mid Lume, Kiribata, 1760 m, P. Vanschuytbroeck & J. Kekenbosch leg., 2 ♂♂ (AMNH, MRAC); North Kivu, Virunga National Park (= Albert Nat. Park), May ya Moto River, affl. Talya, 1180 m, 6. V. 1957, P. Vanschuytbroeck leg., 2 ♂♂ (AMNH, MRAC); Orientale Prov., Haut-Uele, Isiro (= Paulis), 1 ♂ (MRAC); Ori- entale Prov. (= Haut-Zaire), Garamba Nat. Park, 26. VI. 1952, H. De Saeger leg., 1 ♀ (AMNH); same locality label, 4. IV. 1951, 2 ♀♀ (AMNH, MRAC); same locality label, 20. IX. 1951, 1 ♂ (AMNH). NAMIBIA: 19 mi. NW Karibib, Ameib Farm, 31. I – 2. II. 1972, Southern African Expedition, 1 ♂ (NHMUK). UGANDA: same locality label as holo- type, 15. VII. 1943, 1 ♀ (NHMUK); same locality label as holotype, VIII. 1943, 1 ♀ (NHMUK); same locality label as holotype, X. 1942, 1 ♀ (AMNH); same locality label as holotype, 10. VIII. 1943, 1 ♀ (AMNH); same locality label as holotype, IV. 1943, 1 ♀ (AMNH); Eastern Region, Serere District, Serere, VII. 1943, T. H. C. Taylor leg., 6 ♀♀ (NHMUK). Other material: BOTSWANA: Serowe, Farmer’s Brigade, 1 ♀ (USNM). CAMEROON: SW Cameroon, Buea, 24. XII. 1981, S. Compton leg., 1 ♀ (NHMUK). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Yelingi Grassland, 5 km E Kikongo Mission, 04 ° 19 ’ S 17 ° 08 ’ E, 8. IV. 2006, sweeping grass, S. L. Heydon & S. E. Stevenson leg., 1 ♂ (DEUCD). GUINEA BISSAU: Cacheu Region, 2 km E of Bigene, 1. XI. 1992, 1 ♀ (ZIL); Gabú Region, 7 km NW of Gabú, 3. XI. 1992, 1 ♀ (ZIL). KENYA: Coast Prov., Muhaka Forest, 4.32490 ° S 39.52349 ° E, 44 m, 28 – 30. V. 2013, YPT, next to indigenous forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 27 – 30. V. 2013, 6 m MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 16 – 26. VII. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 26. VII – 9. VIII. 2015, 1 ♀ (NMK); same locality label, 1 – 15. XI. 2014, 2 ♀♀ (NMK); same locality label, 11 – 25. X. 2015, 3 ♀♀ (NMK); same locality label, 15 – 29. XI. 2014, 2 ♀♀ (NMK); same locality label, 23. VIII – 6. IX. 2015, 4 ♀♀ (3 in NMK, 1 in MOLC); same locality label, 27. XII. 2015 – 9. I. 2016, 1 ♀ (NMK); Coast Prov., Kaya Kinondo, 4.39382 ° S 39.54567 ° E, 10 m, 8 – 22. VIII. 2011, MT, coral rag canopy for- est, R. Copeland leg., 1 ♀ (NMK); same locality label, 22. VIII – 5. IX. 2011, 1 ♀ (NMK); Coast Prov., Taita Hills, Mwatate area, 3.48444 ° S 38.33251 ° E, 1011 m, 7 – 21. II. 2012, MT, below Bura Bluff, riverine forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 21. II – 7. III. 2012, 1 ♀ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 8 – 22. I. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♀ (MOLC); Eastern Prov., Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, 15 – 29. X. 2011, MT, inside indigenous forest, R. Copeland leg., 1 ♂ (NMK); KENYA: Eastern Prov., Kasaala area, Mulu Musingila’s Farm, 13 – 27. XII. 2016, MT, near gully, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Marsabit Forest, 2.32167 ° N 37.98409 ° E, 1418 m, 1 – 15. VI. 2016, MT, near Ahmed Gate, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest, Calebs Campground, 00 ° 22.175 ‘ N 34 ° 53.297 ‘ ’ E, 1580 m, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MTC). MALI: Koulikoro Region, Katibougou, VIII. 2008, MT, D. Sommaggio leg., 2 ♀♀ (MOLC). NAMIBIA: Aus, I. 1930, R. E. Turner leg., 1 ♂ (NHMUK); Otjiwaronge, 1 ♂ (SANC). SENEGAL: Casamance, 20 km N of Kolda, 5. XI. 1992, 1 ♀ (ZIL). SOUTH AFRICA: Gauteng, Donkerhoek, 1 ♂ (SANC); Gauteng, Pretoria, garden, 8 – 11. II. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu- Natal, Drakensberg, Van Reenen, 1 – 22. I. 1927, R. E. Turner leg., 1 ♂ (NHMUK); KwaZulu-Natal, Riveryiew, 1 ♂ (SANC); KwaZulu-Natal, Pietermaritzburg, Ferncliff Nature Reserve, 29 ° 33.2 ’ S 30 ° 20.6 ’ E, 850 m, 6 – 8. IV. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Pietermaritzburg, Cumberland Nature Reserve, 29 ° 30.8 ’ S 30 ° 30.3 ’ E, 640 m, 21 – 22. II. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, Royal Natal Nature Re- serve, 28 ° 41.4 ’ S 28 ° 56.3 ’ E, 1420 m, 10 – 13. XII. 2004, MT, M. Mostovski leg., 3 ♀♀ (NMSA); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ’ S 30 ° 20.4 ’ E, MT, near stream, 26 – 28. III. 2004, M. Mostovski leg., 1 ♀ (NMSA); same locality label, 3. XII. 2004 – 8. I. 2005, 1 ♀ (NMSA); same locality label, 3 – 26. II. 2005, 1 ♀ (NMSA); same locality label, 27. II – 19. III. 2005, 1 ♀ (NMSA); KwaZulu-Natal, Umtamvuna Nature Reserve, 1 ♀ (SAMC); KwaZulu-Natal, Eshowe, near Ntumeni N. Res., 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg., 2 ♀♀ (NMSA); KwaZulu-Natal, La Mercy, Site 1, 29 ° 37.41 ’ S 31 ° 06.454 ’ E, 85 m, 6. XII. 2006, YPT, Mostovski leg., 9 ♀♀ (8 in NMSA, 1 in MOLC); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 24. XI – 8. XII. 2005, MT, G. Whiteley leg., 1 ♀ (DJBC); same locality label, 9 – 24. XI. 2005, 5 ♂♂ (3 in DJBC, 2 in MOLC); KwaZulu-Natal, Coleford Nature Reserve, 18.8 km 207 ° SW Underberg, 29 ° 57.677 ’ S 29 ° 28.123 ’ E, 7. XII. 2001, sweep, moist upland, grassland on dolerite, S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, 16.5 km 216 ° SW Underberg, 29 ° 56.086 ’ S 29 ° 26.721 ’ E, grassland on sandstone, 1 ♀ (SAMC); Limpopo, 15 km E Klaserie, Guernsey Farm, 19 – 31. XII. 1985, black MT, M. Sanborne leg., 1 ♀ (PMA); Limpopo, Klaserie, 24 – 31. XII. 1985, W. R. Mason leg., 1 ♂ (PMA). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, north west side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 26 – 30. XI. 1995, Acacia / Commiphora / Combretum bushland, YPT, SAM-HYM-PO 19694, S. van Noort leg., 6 ♀♀ (SAMC); same locality label, 24. IV. 1996, sweep, mk 34, SAM-HYM-PO 18158, 1 ♀ (SAMC); same local- ity label, 26. XI. 1995, YPT, open Combretum bushland, SAM-HYM-PO 19588, S. van Noort leg., 2 ♀♀ (SAMC); same locality label, 30. XI – 3. XII. 1995, SAM-HYM-PO 19353, 1 ♀ (SAMC); Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ‘ S 37 ° 48 ‘ E, 9 – 10. XII. 1995, Acacia bushveld, YPT, SAM-HYM-PO 14830, SAM-HYM-PO 14831, S. van Noort leg., 3 ♀♀ (SAMC); Mkomazi Game Reserve, Kisima Plot, 04 ° 06.06 ‘ S 38 ° 05.58 ‘ E, 25. XI – 8. XII. 1995, MT, Acacia / Commiphora bushland, SAM-HYM-PO 16496, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, near Viteweni Ridge, 03 ° 55.09 ‘ S 37 ° 49.63 ‘ E, 22 – 23. IV. 1996, YPT, open Combretum / Dichrostachys bushland, burnt Nov. 1995, SAM-HYM-PO 19219, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Dindera Dam, 03 ° 56 ‘ S 37 ° 48 ‘ E, 22 – 23. IV. 1996, YPT, open bushland on dam wall, SAM-HYM-PO 15580, S. van Noort leg., 2 ♀♀ (SAMC). UGANDA: Busoga Kingdom, Bussu, 1909, E. Bayon leg., 1 ♀ (MCSNG). ZIMBABWE: Harare (= Salis- bury), 24. II. 1974, A. Watsham leg., 2 ♀♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF632A87FF3ED59BFCDEFE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF632A87FF3ED59BFCDEFE70.taxon	distribution	Distribution. Botswana, Cameroon, Democratic Republic of the Congo, Guinea Bissau, Kenya, Mali, Namibia, Senegal, South Africa, Tanzania, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A87FF3ED407FE1FFAAA.taxon	description	(Fig. 77 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A87FF3ED407FE1FFAAA.taxon	description	Description. ♂. Fully winged; body length 1.4 mm. Head brown, except mandible testaceous; antenna testa- ceous; mesosoma and metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 7: 4: 4: 4: 4: 3.5: 4: 4: 4.5: 6. Head dull, completely strongly granulate; frontal line absent; frons without lateral keels near orbits directed towards antennal toruli; occipital carina complete; POL = 4; OL = 2; OOL = 4; OPL = 3; TL = 3.5; greatest breadth of lateral ocelli shorter than OPL (1.5: 3). Mesoscutum shiny, punctate, unsculptured among punctures, with lateral regions weakly granulate. Notauli incomplete, reaching about 0.65 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part. Paramere (Fig. 77 D) without distal inner pointed process, with inner proximal lobe with mosaic pattern and not covered with papillae. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A87FF3ED407FE1FFAAA.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Eastern Cape, Umtata, 18. II – 18. III. 1923, R. E. Turner leg., Brit. Mus. 1923 – 189 (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A87FF3ED407FE1FFAAA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A87FF3ED407FE1FFAAA.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A98FF3ED3CEFC16FCBC.taxon	description	(Fig. 78 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A98FF3ED3CEFC16FCBC.taxon	description	Description. ♀. Fully winged; body length 2.0 – 4.3 mm. Head black, except mandible testaceous; antenna of holo- type testaceous, except antennomeres 6 – 10 brown; antenna of paratype brown, except scape and pedicel testaceous; mesosoma black; in specimen from South Africa, Pietermaritzburg, Hilton, pronotum and mesoscutum black, with reddish-dark nuances; metasoma brown; holotype with proleg testaceous, mesoleg testaceous (except coxae, femora, tibiae and tarsomere 5 darkened), metaleg brown (except trochanters and tarsomeres 1 – 4 testaceous); paratype with legs brown, except tarsi, articulations, protibia and stalks of femora testaceous. Antenna clavate; antennomeres in following proportions: 10: 5: 6: 5: 3.5: 5: 5: 6: 5: 6.5. Head dull, granulate, with numerous slight and irregular areolae; frontal line complete; POL = 8; OL = 5; OOL = 4; OPL = 6; TL = 6; occipital carina complete. Pronotum shiny, smooth, with anterior surface transversely striate; posterior surface slightly granulate, very long, shorter than mesoscutum (11: 18 in holotype; 14: 18 in large specimen (4.3 mm) from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E), slightly broader than long (17: 11); pronotal tubercle reaching tegula. Mesoscutum slightly granulate. Notauli incomplete, shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity without longitudinal keels, dull, reticulate rugose, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7); in specimen from South Africa, Pietermaritzburg, Hilton, forewing with two slightly infuscate transverse bands and distal part of 2 rrs & Rs vein more than 0.5 × as long as proximal part (5: 3). In specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, forewing with two dark transverse bands. Protarsomeres in following proportions: 6: 2: 3: 9: 18. Enlarged claw (Fig. 78 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 78 F) with basal part much shorter than distal part (6: 13), with two rows of about 46 lamellae; apex with approximately 16 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A98FF3ED3CEFC16FCBC.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Gauteng, Pretoria, 21. XI. 1994, A. J. Urban leg. (SANC). Paratype: ZAMBIA: 45 km SE Kitwe, 12 – 15. I. 2003, J. Halada leg., 1 ♀ (OLL). Other material: DEM- OCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 24. VI. 2006, sweeping, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (MOLC); same locality label, 24. IV. 2006, PT at forest edge, 1 ♀ (DEUCD). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 6. X. 2010 – 18. I. 2011, YPT, Camdeboo Escarpment Ticket, S. van Noort leg., PCD 09 - ACA 1 - Y 06, 1 ♀ (SAMC); KwaZulu-Natal, Pietermaritzburg, Hilton, garden, 27. I – 16. II. 2004, MT, M. Mostovski leg., 1 ♀ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A98FF3ED3CEFC16FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF602A98FF3ED3CEFC16FCBC.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa, Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7F2A98FF3ED5C3FE1FF8EA.taxon	description	(Fig. 79 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7F2A98FF3ED5C3FE1FF8EA.taxon	description	Description. ♀. Fully winged; body length 2.4 mm. Head black, except mandible testaceous; occiput black, except testaceous spot behind ocellar triangle; antenna brown, except tarsomeres 7 – 10 testaceous; propleuron black; pronotum black, except lateral regions, posterior tubercle, posterior margin and median spot on anterior collar testaceous; rest of mesosoma black; metasoma brown; proleg brown, except tarsus testaceous; meso- and metaleg brown, except tibia and tarsus testaceous. Antenna clavate; antennomeres in following proportions: 8: 5: 5.5: 4.5: 4: 4: 4: 4.5: 4: 6. Head flat, dull, bare, completely granulate; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5; OL = 4; OOL = 4.5; OPL = 5; TL = 5. Pronotum anteriorly crossed by slight transverse impression, dull, granulate, almost bare, with posterior margin unsculptured; posterior surface approximately as long as mesoscutum (10: 11), with lateral borders rounded; pronotal tubercle reaching tegula. Mesoscutum dull, completely granulate. Notauli incomplete, slightly visible, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum shiny, unsculptured, partly rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 7). Protarsomeres in following proportions: 6: 2: 3: 6: 15. Enlarged claw (Fig. 79 A) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 79 A) with two rows of 9 + 13 lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7F2A98FF3ED5C3FE1FF8EA.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Constantiaberge, 34 ° 02.5 ’ S 18 ° 23.5 ’ E, 640 m, above road to mast overlooking Hout Bay, 15 – 23. II. 1994, Mesic Mountain Fynbos, MT, SAM- HYM-PO 03801, S. van Noort leg. (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7F2A98FF3ED5C3FE1FF8EA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7F2A98FF3ED5C3FE1FF8EA.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7E2A9AFF3ED17AFD01FBF8.taxon	description	(Fig. 77 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7E2A9AFF3ED17AFD01FBF8.taxon	description	Description. ♂. Fully winged; body length 1.9 – 2.2 mm. Head black, except mandible testaceous; antenna testa- ceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa and club of metafemur partly darkened. In specimen from Kenya, 00 ° 20.56 ’ N 34 ° 52.12 ’ E, legs totally testaceous. Antenna filiform, with setae longer than breadth of antennomeres; antennomeres in following proportions: 10: 7: 7.5: 7: 7: 7: 7: 7: 6.5: 10. Head dull, granulate and reticulate rugose; frontal line absent; occipital carina complete; POL = 6; OL = 3; OOL = 4; OPL = 2.5; TL = 3; greatest breadth of lateral ocelli about as long as OL. Mesoscutum shiny, finely punctate, unsculptured among punctures, with surface near anterior margin weakly (in holotype) or strongly (in specimen from Kenya, 00 ° 20.56 ’ N 34 ° 52.12 ’ E) rugose; occasionally mesoscutum partly granulate. Notauli incomplete, reaching about 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with transverse posterior keel, reticulate rugose; propodeal declivity completely reticulate rugose, usually with left longitudinal keel hardly visible and right longitudinal keel absent, rarely with two complete longitudinal keels; usually longitudinal keels of propodeal declivity irregular. Forewing hyaline, without dark transverse bands or spots; distal part of 2 rrs & Rs vein much shorter than proximal part (3: 10). Paramere (Fig. 77 E) with distal inner rounded process; dorsal membranous process with distal area setose (Fig. 77 E). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7E2A9AFF3ED17AFD01FBF8.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Western Prov., Kakamega District, Kakamega Forest, 00 ° 14.13 ’ N 34 ° 51.87 ’ E, 4. III. 1999, R. Wharton leg. (TAMU (it will be deposited in NMK )). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Reserve Speciale de Forêt dense de Dzanga- Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 17. V. 2001, sweep, lowland rainforest, S. van Noort leg., 6 ♂♂ (5 in SAMC, 1 in MOLC); Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 4. V. 2001, sweep, lowland rainforest, marsh clearing, S. van Noort leg., 2 ♂♂ (SAMC). KENYA: Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 27. V – 10. VI. 2012, MT, next to small forest pond, R. Copeland leg., 1 ♂ (NMK); same locality label, 15 – 29. V. 2011, 1 ♂ (NMK); same locality label, 10 – 24. VII. 2011, 1 ♂ (NMK); same locality label, 3.47901 ° S 38.34134 ° E, 1617 m, 21 – 25. VIII. 2012, 6 m MT, next to fallen tree, R. Copeland leg., 1 ♂ (NMK); Western Prov., Kakamega District, Kakamega Forest, 00 ° 20.56 ’ N 34 ° 52.12 ’ E, 22. V – 3. VI. 2000, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7E2A9AFF3ED17AFD01FBF8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7E2A9AFF3ED17AFD01FBF8.taxon	distribution	Distribution. Central African Republic, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7D2A9BFF3ED29FFC57FAB4.taxon	description	(Figs 77 F, 79 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7D2A9BFF3ED29FFC57FAB4.taxon	description	Description. ♀. Fully winged; body length 1.9 – 2.2 mm. Head black, except mandible testaceous; antenna tes- taceous, except distal extremity of antennomere 10 darkened; mesosoma black; metasoma brown; legs yellow. Antenna clavate; antennomeres in following proportions: 9: 4: 5: 4: 4: 5: 5: 5: 5: 6.5. Head convex, dull, covered with short thin setae, granulate, with many very evanescent tracks of rugosities; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 4; TL = 5. Pronotum anteriorly crossed by slight transverse impression, dull, granulate, covered with long thin setae; pronotal tubercle reaching tegula; posterior surface shorter than mesoscutum (6: 12). Mesoscutum dull, as sculptured as head. Notauli incomplete, very shortly visible near anterior margin of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc without strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 6). Protarsomeres in following proportions: 50: 16: 20: 20: 68. Enlarged claw (Fig. 79 B) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 79 B) with two rows of 1 + 7 or 2 + 5 lamellae; apex with four lamellae. tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.5 – 1.9 mm. Head black, except mandible testaceous; antenna of holotype brown, except scape and pedicel yellow; antenna of paratype from Yemen, Lahj, completely brown-testaceous; mesosoma black; metasoma brown; legs testaceous, except metacoxa, club of metafemur and metatibia partly darkened. Antenna filiform; antennomeres in following proportions: 6: 4: 4.5: 4.5: 4.5: 5: 4.5: 5: 5: 7. Head dull, granulate and with tracks of rugosities; frontal line complete; occipital carina complete; POL = 4.5; OL = 3; OOL = 4; OPL = 2; TL = 3. Mesoscutum of holotype dull, slightly granulate, with granulation more evident on anterior half and with posterior half punctate and very slightly granulate; mesoscutum of paratype from Yemen, Lahj, completely and strongly granulate; in two specimens from South Africa, Pretoria, mesoscutum almost completely unsculptured and partly very slightly granulate. Notauli of holotype incomplete, reaching approximately 0.5 × length of mesoscutum; in paratype from Yemen, Lahj, notauli faintly visible. Mesoscutellum and metanotum shiny, smooth, punctate, unsculptured among punctures. Metapectal-propodeal disc without transverse posterior keel; in specimen from Tanzania, Mkomazi Game Reserve, metapectal-propodeal disc with irregular transverse keel separating metapectal-propodeal disc and propodeal declivity; metapectal-propodeal disc completely reticulate rugose, dull; propodeal declivity without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (2: 4). Paramere (Fig. 77 F) without inner process, with numerous papillae on inner side. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7D2A9BFF3ED29FFC57FAB4.taxon	materials_examined	Material examined. Types: ♂ holotype: YEMEN: Ta’izz, 13 ° 35 ’ N 44 ° 02 ’ E, V. 2000, N. 4631, light trap, A. van Harten & A. R. Al Yarimi leg. (MOLC). Paratypes: YEMEN: Lahj, 13 ° 04 ’ N 44 ° 53 ’ E, V – VI. 2000, MT, A. van Harten & A. Sallam leg., 1 ♂ (MOLC); same locality label, XI. 2000, 1 ♂, 1 ♀ (MOLC); same locality label, IX. 2000, 1 ♀ (MOLC); same locality label, IX. 2001, 2 ♀♀ (MOLC); same locality label, 1. X – 17. XII. 2001, 1 ♀ (MOLC); same locality label, III – V. 2002, 2 ♀♀, 1 ♂ (MOLC). SOUTH AFRICA: Limpopo, Klaserie, MT, 18 – 24. XII. 1985, W. R. Mason leg., 1 ♂ (PMA); Mpumalanga, Krüger National Park, Lower Sabie, 250 m, 14. XII. 1985, W. Mason leg., 1 ♂ (PMA). Other material: KENYA: Eastern Prov., base of Ukasi Hill, 0.82103 ° S 38.54443 ° E, 613 m, 21. XI – 5. XII. 2011, MT, Acacia-Commiphora savanna, R. Copeland leg., 1 ♂ (NMK). MADAGASCAR: Fianara- ntsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04, 1 ♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 26. I – 5. II. 2003, MT in transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 14, 1 ♂ (MOLC). SOUTH AFRICA: Eastern Cape, Ndawana, 21.5 km 208 ° SW Underberg, 29 ° 59.102 ’ S 29 ° 27.756 ’ E, 29. XI. 2001, sweep, moist upland grassland on dolerite, livestock grazed, S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Ndawana, 20.8 km 208 ° SW Underberg, 29 ° 58.759 ’ S 29 ° 27.812 ’ E, 29. XI. 2001, sweep, moist upland grassland on sandstone, livestock grazed, S. van Noort leg., 1 ♂ (MOLC); Gauteng, Donkerhoek, 1 ♂ (SANC); Gauteng, Pretoria, garden, 8 – 11. II. 2005, YPT, V. Kolyada leg., 2 ♂♂ (NMSA); KwaZulu- Natal, Umtamvuna Nature Reserve, 1 ♂ (SAMC). TANZANIA: Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ’ S 38 ° 01.37 ’ E, 2. V. 1996, sweep, mk 43, Cordia, Solanum shrubs & grasses in Commiphora woodland, SAM-HYM- PO 18082, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7D2A9BFF3ED29FFC57FAB4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7D2A9BFF3ED29FFC57FAB4.taxon	distribution	Distribution. Kenya, Madagascar, South Africa, Tanzania, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	description	(Figs 74 F, 79 C, F, G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	description	Description. ♀ (tentatively associated with the ♂ below; see remarks). Fully winged; body length 2.6 – 2.7 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 6 – 10 brown (occasionally antennomere 5 partly brown); mesosoma and metasoma black; legs testaceous, except metacoxa partly brown. Antenna clavate; antennomeres in following proportions: 9: 5: 6: 5: 5: 5: 5: 5: 5: 8.5. Head convex, dull, granulate and reticulate rugose (in specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, head slightly rugose, not granulate, with a large unsculptured area in front of anterior ocellus); frons with two lateral longitudinal keels near orbits and directed towards antennal toruli; frontal line complete; POL = 6; OL = 4; OOL = 4; OPL = 3; TL = 3; greatest breadth of lateral ocelli shorter than OL (3: 4); occipital carina complete. Occasionally POL about as long as OL. Pronotum dull, granulate and rugose, with posterior margin shiny, unsculptured; posterior surface shorter than mesoscutum (5: 14); pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two complete longitudinal keels, with median area dull, as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4.5: 9). Protarsomeres in following proportions: 9: 2: 1.5: 2.5: 9.5. Enlarged claw (Fig. 79 C) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 79 C) with about 8 – 9 bristles, without proximal and medial lamellae; apex with 7 – 9 lamellae. Tibial spurs 1 / 1 / 2. ♂ (Fig. 74 F). Fully winged; body length 1.7 – 3.4 mm. Head black; mandible yellow, with teeth brown; antenna completely brown, or totally testaceous, or brown, except antennomeres 1 – 3 testaceous; mesosoma black; metasoma black; legs completely brown, or testaceous, except metacoxa, club of metafemur and metatibia brown. Antenna filiform; antennomeres of specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, in following proportions: 10: 6: 7: 6: 6: 6: 6: 6.5: 6.5: 9. Antennomeres of A. terminale holotype in following proportions: 8.5: 4: 5: 5: 5: 5: 5: 5: 4.5: 8. Head dull or shiny, reticulate rugose and granulate; clypeus completely smooth, without parallel keels; frontal line complete; frons without lateral keels directed towards antennal toruli (in specimen from Gabon, Mts. Doudou Reserve, frons with two tracks of lateral keels along orbits and directed towards antennal toruli; in specimen from Guinea, 7 ° 42 ’ N 8 ° 23 ’ W, frons with two strong lateral longitudinal keels along orbits and directed towards antennal toruli, and vertex with two oblique keels from lateral ocelli to occipital carina; in specimen from South Africa, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, frons unsculptured and with very slightly rugose, without lateral keels directed towards antennal toruli); occipital carina complete; POL = 8; OL = 4; OOL = 6; OPL = 4; TL = 4. In holotype of A. terminale, POL = 4.5; OL = 3; OOL = 4.5; OPL = 2; TL = 2.5. Rarely head almost completely punctate, unsculptured among punctures (as in Anteon fisheri Olmi, 2003). Mesoscutum shiny, punctate and unsculptured among punctures, strongly rugose near anterior margin or in anterior third of mesoscutum; in specimen from Gabon, Doudou Mts. Reserve, mesoscutum completely punctate, unsculptured among punctures; in specimen from South Africa, 29 ° 39 ‘ 23 ‘‘ S 30 ° 27 ‘ 41 ‘‘ E, mesoscutum weakly granulate, except anterior region rugose and central area unsculptured. Notauli incomplete, reaching approximately 0.25 – 0.50 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose, dull; propodeal declivity dull, with two longitudinal keels and with median area as rugose as lateral areas; occasionally median area rugose and with numerous longitudinal keels; occasionally (specimens previously attributed to A. terminale) median area partly shiny, unsculptured. In specimen from Zimbabwe, Chishawasha, propodeal declivity with two hardly visible longitudinal keels; in specimens from South Africa longitudinal keels absent. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (5: 10; 2.5: 7 in A. terminale holotype). Paramere without distal inner pointed or rounded process, with long and slender dorsal proximal membranous process (Figs 79 F, G (Fig. 255 in Olmi 1984 erroneous )). In specimen from South Africa, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, inner side of paramere with some papillae. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	materials_examined	Material examined. Types: ♂ holotype of A. zairense: DEMOCRATIC REPUBLIC OF THE CONGO: Basoko, V. 1949, P. L. G. Benoit leg. (MRAC); ♂ holotype of A. terminale: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13 (CAS). Paratypes of A. terminale: MADAGASCAR: same locality label as holotype, 12 – 19. III. 2002, MA- 02 - 09 A- 20, 1 ♂ (CAS). SOUTH AFRICA: KwaZulu-Natal, Rams- gate, Butterfly Sanctuary, 30 ° 53.3 ‘ S 30 ° 20.4 ‘ E, MT near stream, 9. VIII – 2. IX. 2004, M. Mostovski leg., 1 ♂ (NMSA); same locality label, 3. IX – 2. X. 2004, 1 ♂ (NMSA); same locality label, 20. III – 25. IV. 2005, 1 ♂ (NMSA). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, 38.6 km 173 ° S Lidjombo, 2 ° 21.60 ’ N 16 ° 03.20 ’ E, 350 m, 23. V. 2001, sweep, lowland rainforest, S. van Noort leg., 6 ♂♂ (SAMC). GABON: Ogoové-Maritime Prov., Mts. Doudou Reserve, 25.2 km 304 ° NW Doussala, 02 ° 13.63 ’ S 10 ° 23.67 ’ E, 660 m, 15. III. 2000, Sweep, GA 00 - S 98, Coastal Lowland Rainforest, undergrowth, low canopy in forest, SAM-HYM- PO 023741, S. van Noort leg. 1 ♂ (SAMC). GUINEA: Nzérékoré Region, Mount Nimba, Gouan River, 7 ° 42 ’ N 8 ° 23 ’ W, 514 m, 15 – 22. I. 1991, FIT, rainforest, L. LeBlanc leg., 1 ♂ (PMA). KENYA: Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 11 – 25. X. 2015, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Vuria Forest, 3.41428 ° S 38.29178 ° E, 2162 m, MT just inside indigenous forest, 21. VIII – 4. IX. 2011, R. Copeland leg., 1 ♂ (RSC); Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 24. VII – 7. VIII. 2011, MT, next to small forest pond, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Taita Hills, Ngangao Forest, 3.36930 ° S 38.34495 ° E, 1736 m, 26. VI – 10. VII. 2011, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); same locality label, 3.36100 ° S 38.34186 ° E, 1848 m, 13 – 27. V. 2012, 2 ♂♂ (NMK); Coast Prov., Taita Hills, Ngangao For- est, 3.36100 ° S 38.34186 ° E, 1848 m, 10. IX – 2. X. 2011, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 17 – 31. X. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Diani Beach area, 4.27559 ° S 39.59337 ° E, 10 m, 2 – 16. XII. 2014, MT, shrubland off Diani Beach Rd., R. Copeland leg., 1 ♂ (NMK); Coast Prov., Gede Forest, 3.30946 ° S 40.01941 ° E, 19 m, 22. VIII – 5. IX. 2011, indigenous forest, MT, secondary forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Marenje Forest, 4.52814 ° S 39.24028 ° E, 76 m, 22. VIII – 5. IX. 2014, MT, indigenous forest, R. Copeland leg., 1 ♂ (MOLC); Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, 26. X – 10. XII. 2011, MT, inside indigenous forest, R. Copeland leg., 1 ♂ (NMK); Nairobi Prov., ICIPE Cam- pus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 29. IV – V. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♂ (NMK); same locality label, 3 – 10. VI. 2014, 1 ♂ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22338 ° S 36.89655 ° E, 1604 m, 24. II – 3. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♂ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere Forest Reserve, 00 ° 23.73 ’ N 34 ° 53.165 ’ E, 5400 feet, XI. 2001, Manfred Kraemer & Co. Leg., 1 ♂ (MTC). MADAGASCAR: An- tsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 29. I. 2001, netted by hand, M. E. Irwin leg., MA- 01 - 01 A- 05, 1 ♂ (CAS); same locality label, 19. III – 6. IV. 2001, MT, MA- 01 - 01 A- 09, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 26 – 29. I. 2001, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 02, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 6 – 21. IV. 2001, MT, MA- 01 - 01 D- 07, R. Harin’Hala leg., 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry for- est, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 14 – 21. II. 2011, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 19, 2 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 4 – 12. II. 2002, 1110 m, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 3 ♂♂ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 A- 04, 12 ♂♂ (CAS); same locality label, 6 – 15. V. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 A- 12, 2 ♂♂ (CAS); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 1 ♂ (CAS); same locality label, 21 – 28. I. 2002, MA- 02 - 09 A- 13 (CAS); Fianarantsoa, Ranomafana Na- tional Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 3 ♂♂, 1 ♀ (CAS); same locality label, 5 – 13. V. 2002, MA- 02 - 09 C- 28, 1 ♀ (MOLC); same locality label, 14 – 22. VII. 2002, MA- 02 - 09 C- 35, 1 ♀ (MOLC), 1 ♂ (CAS); same locality label, 23. VI – 4. VII. 2002, MA- 02 - 09 C- 33, 1 ♂ (CAS); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 C- 16, 8 ♂♂ (6 in CAS, 2 in MOLC); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 7 ♂♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 3 ♂♂ (2 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT in high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13, 1 ♀ (CAS); same locality label, 14 – 21. I. 2002, 6 ♂♂ (2 in CAS, 4 in MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 1 ♀ (MOLC), 5 ♂♂ (4 in CAS, 1 in MOLC); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 B- 35, 2 ♂♂ (CAS); same locality label, 12 – 19. II. 2002, MA- 02 - 09 B- 16, 2 ♂♂ (CAS); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 1 ♂ (MOLC); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♀ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 2 ♀♀ (CAS, MOLC), 2 ♂♂ (CAS); same locality label, 4 – 12. III. 2002, MA- 02 - 09 B- 19, 1 ♂ (MOLC); same locality label, 21 – 28. I. 2002, MA- 02 - 09 B- 13, 1 ♂ (MOLC); same locality label, 27. II – 9. III. 2003, MA- 02 - 09 B- 54, 1 ♂ (CAS); same locality label, 18 – 29. VI. 2003, MA- 02 - 09 B- 64, 1 ♂ (MOLC); Fianarantsoa, Rano- mafana, JIRAMA water works, 21 ° 14.91 ‘ S 47 ° 27.13 ‘ E, 690 m, 21 – 24. XII. 2001, MT, near river, R. Harin’Hala leg., MA- 02 - 09 D- 08, 1 ♂ (CAS); same locality label, 28. I – 4. II. 2002, MA- 02 - 09 A- 14, 2 ♂♂ (CAS, MOLC); same local- ity label, 10 – 14. I. 2002, MA- 02 - 09 A- 11, 1 ♂ (CAS); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W Anjia- mangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 26. V – 2. VI. 2011, MT, MG- 61 - 15, 3 ♂♂ (2 in CAS, 1 in MOLC); Toamasina, Ankerana, 18 ° 24.23 ’ S 48 ° 48.09 ’ E, 1108 m, 16 – 21. I. 2012, MT, rainfor- est, B. L. Fisher et al. leg., BLF 27665, 2 ♂♂ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 2 – 8. X. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 14, 1 ♂ (CAS); same locality label, 1 – 7. XI. 2001, MA- 01 - 08 B- 18, 1 ♂ (CAS); same locality label, 1 – 5. IX. 2001, MA- 01 - 08 B- 11, 10 ♂♂ (CAS); same locality label, 7 – 16. XI. 2001, 1 ♂ (CAS). SOUTH AFRICA: Eastern Cape, Port St. Johns, I. 1924, R. E. Turner leg., 1 ♂ (NHMUK); same locality label, XII. 1923, 1 ♂ (NHMUK); Eastern Cape, Gra- hamstown, 1 ♂ (AEIC); Eastern Cape, Gonubie, 32 ° 56 ’ S 28 ° 02 ’ E, 3 – 12. I. 1995, MT, Coastal Dune Forest near River Mouth, SAM-HYM-P 013717, S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♂ (SAMC); Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 9. IV – 26. VII. 2010, YPT, Amathole Mistbelt Grassland, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 3. IX – 30. X. 2009, WTB 09 - GRA 1 - Y 01, 13 ♂♂ (SAMC); same locality label, 6. X. 2010 – 18. I. 2011, MT, WTB 09 - GRA 1 - M 05, 1 ♂ (MOLC); KwaZulu-Natal, Ubombo, Lake Sibaya, 1 ♂ (PMA); KwaZulu-Natal, 75 km WSW Estcourt, Cathedral Peak Forest Station, 1 ♂ (CNC); KwaZulu-Natal, Cathedral Peak Natural Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 14. XII. 2005 – 29. I. 2006, MT, M. Mostovski leg., 3 ♂♂ (NMSA); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ‘ S 30 ° 20.4 ‘ E, MT near stream, 6. VI – 9. VII. 2004, 9. VII – 9. VIII. 2004, 9. VIII – 2. IX. 2004, 3. IX – 2. X. 2004, 3 – 30. X. 2004, 1. XI – 2. XII. 2004, 9. I – 2. II. 2005, 27. II – 19. III. 2005, 20. III – 25. IV. 2005, 27. IV – 27. VII. 2005, 18. XI. 2006 – 26. II. 2007, M. Mostovski leg., 37 ♂♂ (NMSA); same locality label, 27. IV – 27. VII. 2005, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Karkloof, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, 1325 m, 28. IX – 3. X. 2005, YPT, M. Mostovski leg., 1 ♂ (NMSA); same locality label, 25. VII – 25. IX. 2005, MT, M. Mostovski leg., 8 ♂♂ (NMSA); KwaZulu-Natal, Vernon Crookes Nat. Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Royal Natal National Reserve, Gudu Forest, 28 ° 40.90 ’ S 28 ° 55.78 ’ E, 1680 – 1730 m, 13. XII. 2005 – 28. I. 2006, MT, M. Mostovski leg., 1 ♂ (NMSA); KwaZulu-Natal, Ashburton, 29 ° 39 ’ 23 ’’ S 30 ° 27 ’ 41 ’’ E, 29. X – 9. XI. 2005, MT, G. Whiteley leg., 1 ♂ (DJBC); Mpumalanga, Krüger National Park, Skukuza Malt., 1 ♂ (PMA); Western Cape, Stellenbosch, Jonkershoek, 3 ♂♂ (PMA, AEIC, AMNH); Western Cape, Ceres, IV. 1925, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, Cape Peninsula, Hout Bay, Skoorsteenkop, 1 ♂ (ZIL); Western Cape, Cape Town, above Tokai Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ N 18 ° 23.5 ’ E, 460 m, 10 – 17. III. 1995, Mesic Mountain Fynbos, Protea coronata dominated, MT, SAM-HYM-PO 13732, S. van Noort leg., 1 ♂ (SAMC); same locality label, 12 – 17. IX. 1993, Mesic Mountain Fynbos on Sandstone, SAM-HYM-PO 03786, 1 ♂ (SAMC); Western Cape, Constantiaberg, above Donkerboskloof, 34 ° 2.175 ‘ N 18 ° 23.528 ‘ E, 460 m, 5 – 13. X. 1993, MT, Mesic Mountain Fynbos with kloof forest elements, S. van Noort leg., 1 ♂ (SAMC); same locality label, 19. IX – 10. X. 1995, 1 ♂ (SAMC); Western Cape, Constantiaberge, 34 ° 02.5 ‘ S 18 ° 23.5 ‘ E, 640 m, above road to mast overlooking Hout Bay, 25. I – 2. II. 1994, MT, mesic mountain fynbos, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Wolfgat Nature Reserve, 34 ° 04 ‘ S 18 ° 39 ‘ E, 5. IX. 1995, sweep, S Coast Strandveld, SAM-HYM-PO 11635, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ‘ S 19 ° 21.39 ‘ E, 11. VIII – 6. IX. 1997, S. van Noort leg., 1 ♂ (SAMC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ‘ S 19 ° 21.393 ‘ E, 21. II – 21. III. 1998, MT, South Coast Strandveld, 57 m, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Anyskop Farm, 5.5 km 290 ° W Langebaanweg, 32 ° 57.394 ‘ S 18 ° 05.324 ‘ E, 11 – 18. IX. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 18 – 25. IX. 2002, 1 ♂ (SAMC); Western Cape, Langberg Farm, 3 km 270 ° W Langebaanweg, 32 ° 58.461 ‘ S 18 ° 07.344 ‘ E, 6 – 13. XI. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 3 ♂♂ (SAMC); Western Cape, Langberg Farm, 3 km 280 ° W Langebaan- weg, 32 ° 58.045 ‘ S 18 ° 07.418 ‘ E, 18 – 25. IX. 2002, MT, Sand Plain Fynbos, S. van Noort leg., 4 ♂♂ (SAMC); same locality label, 4 – 11. IX. 2002, 4 ♂♂ (SAMC); same locality label, 11 – 18. IX. 2002, 20 ♂♂ (SAMC); Western Cape, West Coast Fossil Park, 3 km 280 ° W Langebaanweg, 32 ° 58.020 ‘ S 18 ° 07.285 ‘ E, 18 – 25. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 3 km 270 ° W Langebaanweg, 32 ° 58.445 ‘ S 18 ° 07.236 ‘ E, 11 – 18. IX. 2002, MT, sand plain fynbos, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.663 ‘ S 21 ° 57.600 ‘ E, 940 m, 23. V – 31. VII. 2009, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ‘ S 19 ° 01.033 ‘ E, 16. III – 16. IV. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 3 ♂♂ (SAMC); same locality label, 16. X – 16. XI. 1999, 6 ♂♂ (SAMC); same locality label, 16. XII. 1999 – 16. I. 2000, 2 ♂♂ (SAMC); same locality label, 16. XI – 16. XII. 1999, 2 ♂♂ (SAMC); same locality label, 16. I – 16. II. 2000, 4 ♂♂ (SAMC); same locality label, 16. II – 16. III. 1999, 9 ♂♂, 2 ♀♀ (SAMC); same locality label, 16. XII. 1998 – 16. I. 1999, 3 ♂♂ (SAMC); same locality label, 16. XI – 16. XII. 1998, 3 ♂♂, 1 ♀ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ‘ S 18 ° 24.259 ‘ E, 31. X – 28. XI. 1997, MT, West Coast Strandveld, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Fynbos Estate, Dragon- ridge, 33 ° 34.092 ‘ S 18 ° 47.680 ‘ E, 635 m, 28. IX – 28. XII. 2010, MT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - M 05, 1 ♂ (SAMC); Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ‘ S 18 ° 57.711 ‘ E, 410 m, 20. X – 27. XI. 2013, MT, Mesic Mountain Fynbos, BH 12 - FYN 3 - M 14, S. van Noort leg., 1 ♂ (MOLC). UGANDA: Western Region, Kibale National Park, Kanyawara Makerere University Biological Field Station, 0 ° 33.784 ’ S 30 ° 21.617 ’ E, 1500 m, 2 – 12. VIII. 2008, MT, primary mid-altitude rainforest, S. van Noort leg., 1 ♂ (SAMC). ZIMBABWE: Harare, Chishawasha, III. 1978, A. Watsham leg., 1 ♂ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Gabon, Guinea, Kenya, Madagascar, South Africa, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF7C2A9FFF3ED3CBFB0FFD9C.taxon	discussion	Remarks. The above description of the ♀ is based on a few specimens collected in Madagascar (21 ° 15.99 ’ S 47 ° 25.21 ’ E, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 21 ° 15.05 ’ S 47 ° 24.43 ’ E), Kenya (4.48576 ° S 39.25845 ° E) and South Africa (34 ° 16.481 ’ S 19 ° 01.033 ’ E); the specimens were not reared and were tentatively assigned to Anteon zairense Benoit (based on ♂ description). Within the same locality in Madagascar, Anteon cautum Olmi (only known from ♂♂) was also collected, consequently it is possible that the above ♀♀ represent the opposite sex of A. cautum. Anteon terminale is here synonymized with A. zairense, based on the fact that within the same localities in Madagascar, mixed populations comprising ♂♂ with either a rugose or unsculptured median area of the metapectal-propodeal disc, and either a rugose or unsculptured propodeal declivity, were collected. As the unique difference between the two species is the sculpture of the metapectal-propodeal disc median area, they were synonymized.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A9FFF3ED423FE24F9F6.taxon	description	(Fig. 79 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A9FFF3ED423FE24F9F6.taxon	description	Description. ♀. Fully winged; body length 3.6 mm. Black; mandible testaceous; clypeus black, except ferruginous spot; antenna testaceous, except antennomeres 5 – 10 darkened; tegula testaceous; metasoma brown; legs yellow. Antenna clavate; antennomeres in following proportions: 13: 8: 15: 12: 11: 10: 10.5: 10: 10: 14. Head slightly convex, shiny, punctate, unsculptured among punctures, with anterior two thirds of frons dull and reticulate rugose; vertex with many short longitudinal keels situated around occipital carina; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 5.5; OL = 4.5; OOL = 5; OPL = 5; TL = 4.5; greatest breadth of lateral ocelli shorter than OL (3.5: 4.5). Pronotum with anterior surface dull and strongly transversely striate; posterior surface shiny, finely punctate, unsculptured among punctures; pronotal tubercle reaching tegula; posterior surface of pronotum much shorter than mesoscutum (10: 21). Mesoscutum, mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, very long, almost reaching posterior margin of mesoscutum. Mesopleuron with anterior surface rugose and propodeal declivity punctate, unsculptured among punctures. Metapleuron strongly transversely striate. Metapectal-propodeal disc with a transverse posterior keel, reticulate rugose; propodeal declivity with two complete longitudinal keels; median area mostly smooth, with some keels around margins; lateral areas rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (7: 13.5). Protarsomeres in following proportions: 12: 4: 4: 7: 25; protarsomere 3 produced into a hook. Enlarged claw (Fig. 79 D) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 79 D) with two rows of 36 (larger) + 14 (slender) lamellae; apex with about 9 lamellae. Basal part of protarsomere 5 longer than distal part (16: 9) and than protarsomere 4 (16: 7). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A9FFF3ED423FE24F9F6.taxon	materials_examined	Material examined. Type: ♀ holotype: ZAMBIA: Central Prov., 15 km S Kapiri Mposhi, 16 – 17. I. 2003, J. Halada leg. (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A9FFF3ED423FE24F9F6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A9FFF3ED423FE24F9F6.taxon	distribution	Distribution. Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A90FF3ED08AFE06FD74.taxon	description	(Fig. 79 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A90FF3ED08AFE06FD74.taxon	description	Description. ♀. Fully winged; body length 3.7 mm. Head black, except mandible brown; antenna brown; propleuron brown; pronotum brown, except posterior tubercle and anterior margin brown-testaceous; rest of mesosoma brown, except axilla brown-testaceous; metasoma brown; legs brown, except articulations, distal extremities of procoxa and mesocoxa testaceous. Antenna clavate; antennomeres in following proportions: 13: 7: 11: 8: 6: 7: 6.5: 6: 6: 9. Head flat, dull, with thin setae, completely reticulate rugose and granulate; frontal line complete; frons with two lateral keels near orbits and directed towards antennal toruli; occipital carina complete; POL = 6; OL = 4; OOL = 5; OPL = 5; TL = 4. Pronotum anteriorly crossed by slight transverse impression, dull, with thin setae, reticulate rugose, with posterior margin unsculptured; posterior surface approximately as long as mesoscutum (14: 15); pronotal tubercle reaching tegula. Mesoscutum shiny, punctate, unsculptured among punctures, laterally slightly irregularly striate. Notauli incomplete, hardly visible, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum shiny, partly unsculptured, partly rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels, with areolae less wide than those of metapectal-propodeal disc. Forewing with two slight dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (4: 12). Protarsomeres in following proportions: 8: 3: 4: 9: 21. Enlarged claw (Fig. 79 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 79 E) with two rows of 17 + 18 lamellae; apex very bent and with six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A90FF3ED08AFE06FD74.taxon	materials_examined	Material examined. Type: ♀ holotype: ZIMBABWE: Eastern Zimbabwe, Mutare, Bvumba, 21 – 24. XII. 1998, M. Snizek leg. (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A90FF3ED08AFE06FD74.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF782A90FF3ED08AFE06FD74.taxon	distribution	Distribution. Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF772A93FF3ED50BFA7DFF6D.taxon	diagnosis	Diagnosis. ♀: macropterous (Figs 80 A, B), or micropterous (Fig. 84 A); palpal formula 6 / 3; occipital carina complete; vertex of head frequently with two strong oblique keels connecting lateral ocelli to occipital carina; pronotum with distinct anterior collar and posterior disc (Figs 80 A, B); in macropterous ♀♀, forewing usually with distal part of 2 r-rs & Rs vein longer than proximal part (Figs 80 A, B), less frequently as long as, or shorter than proximal part; enlarged claw without inner proximal prominence bearing bristles (Fig. 83 A), with one – two bristles or peg-like setae located further distally than proximal prominence (Fig. 83 A); tibial spurs 1 / 1 / 2. ♂ (Figs 80 C, D): always macropterous (even with micropterous ♀) (Figs 80 C, D); palpal formula 6 / 3; vertex of head frequently with two strong oblique keels connecting lateral ocelli to occipital carina; antennal setae usually much longer than breadth of antennomeres (Fig. 80 C), less frequently shorter; forewing usually with distal part of 2 r-rs & Rs vein longer than proximal part (Figs 80 C, D), less frequently as long as, or shorter than proximal part; forewing usually with pterostigma than 4 × as long as broad or more; paramere without dorsal process, usually with one more or less large inner branch wrapping penis (Fig. 88 E), less frequently with one reduced inner branch (Figs 86 C, D); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF772A93FF3ED50BFA7DFF6D.taxon	distribution	Distribution. Worlwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF772A93FF3ED50BFA7DFF6D.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF772A93FF3ED50BFA7DFF6D.taxon	discussion	World species. 165 species are known, 28 in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF772A93FF3ED50BFA7DFF6D.taxon	discussion	Remarks. Fossil species of Deinodryinus were described by Olmi et al. (2010) and Guglielmino & Olmi (2011). Larvae of Deinodryinus were described by Guglielmino et al. (2017 a). Leafhoppers parasitized by Deinodryinus species usually show dryinid sacs dorsally situated between the head and the prothorax (Fig. 81 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	description	(Figs 81 B, 82, 83 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	diagnosis	Diagnosis. Macropterous ♀ of Deinodryinus with head strongly punctate, unsculptured among punctures; frons without frontal line and lateral keels joining lateral ocelli to antennal toruli; mesosoma without strong constriction between prothorax and mesothorax (Fig. 81 B); posterior surface of pronotum with sharp lateral margins; protarsomere 1 much shorter than 4; distal region of protarsomere 5 very slender (Fig. 83 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	description	Description. ♀ (Figs 81 B, 82). Fully winged; body length 3.4 mm. Head ferruginous; antenna testaceous, except antennomere 10 brown; mesosoma ferruginous, except lateral regions of pronotum brown, metanotum and metapectal-propodeal disc black; petiole black; metasoma testaceous, except anerior half of first sclerite brown; legs testaceous, except brown spot on outer side of metacoxa. Antenna clavate; antennomeres in following proportions: 9: 5: 16: 10: 7: 7: 6: 6: 6: 8. Head (Fig. 82 C) slightly swollen, shiny, strongly punctate, unsculptured among punctures; frontal line absent; frons without lateral keels; vertex behind ocellar triangle without two oblique keels connecting lateral ocelli to occipital carina; occipital carina complete; POL = 5; OL = 4; OOL = 8; OPL = 7; TL = 6; greatest breadth of lateral ocelli shorter than OL (3: 4). Pronotum anteriorly crossed by strong transverse impression, shiny, smooth, punctate, unsculptured among punctures; posterior surface of pronotum slightly broader than long (21: 16), slightly shorter than mesoscutum (16: 18), with lateral margins sharp; pronotal tubercle reaching tegula. Mesoscutum and mesoscutellum shiny, punctate, unsculptured among punctures. Notauli (Fig. 82 A) incomplete, reaching approximately 0.7 × length of mesoscutum. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, with- out transverse posterior keel; propodeal declivity reticulate rugose, except posterior third sculptured by transverse keels. Forewing (Fig. 82 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (15: 8). Protarsomeres in following proportions: 10: 3: 6: 18: 28. Enlarged claw (Fig. 83 A) with one lamella situated further distally than proximal prominence. Protarsomere 5 (Fig. 83 A) with two rows of 26 (short) + 8 (long) lamel- lae; apex very slender, with 3 lamellae, one of which very long. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19455): MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 900 m, 29. I. 2001, netted by hand, M. E. Irwin leg., MA- 01 - 01 A- 05 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF742A94FF3ED330FA8EFDB8.taxon	etymology	Etymology. The species is named ambrensis after the collection site in Montagne d’Ambre National Park.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF732A95FF3ED0E5FE14FD48.taxon	description	(Fig. 83 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF732A95FF3ED0E5FE14FD48.taxon	description	Description. ♀. Fully winged; body length 6.6 mm. Head black, except mandible, clypeus and anterior region of frons (mainly along orbits) testaceous; antenna testaceous, except antennomeres 9 – 10 brown; mesosoma black, except margins of pronotum and apex of metapectal-propodeal disc testaceous; tegula testaceous; petiole black; metasoma black, except segment 1 mostly testaceous; legs testaceous-reddish, except distal region of clubs of pro- and metafemur brown. Antenna filiform; antennomeres in following proportions: 18: 7: 30: 19: 16: 14: 13: 12: 12: 17. Head shiny, punctate, unsculptured among punctures; frons anteriorly strongly punctate, without lateral keels; frontal line incomplete; occipital carina complete; POL = 6; OL = 5; OOL = 12; OPL = 13; TL = 9. Pronotum shiny, finely punctate, with transverse keels, crossed by transverse impression separating anterior collar from posterior disc; lateral margin of posterior surface rounded; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel. Forewing hyaline, with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (18: 13). Protarso- meres in following proportions: 15: 5: 10: 32: 48. Enlarged claw (Fig. 83 B) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 83 B) with two rows of approximately 90 lamellae (all about same length); apex with approximately eight lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF732A95FF3ED0E5FE14FD48.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Périnet, 19. XII. 1955, in forest, E. McC. Callan leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF732A95FF3ED0E5FE14FD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF732A95FF3ED0E5FE14FD48.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712A96FF3ED6D7FE1FFA1D.taxon	description	(Fig. 83 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712A96FF3ED6D7FE1FFA1D.taxon	description	Description. ♀. Micropterous; forewing very reduced, reaching transverse furrow behind metanotum, slightly lon- ger than mesoscutellum (6: 5). Body length 3.2 – 4.2 mm. Holotype testaceous, except antennomeres 8 – 10, ocellar area, prosternum, lateral regions of pronotum, posterior half of metasoma darkened; mesoscutum, metanotum, transverse furrow behind metanotum, posterior half of metapectal-propodeal disc and petiole black; mesopleuron and metapleuron partly darkened; legs testaceous, except meso- and metacoxa and club of metafemur darkened. In two paratypes from South Africa, Kogelberg Nature Reserve, colour broadly darkened or black, as follows: head testaceous, with ocellar region darkened; mesosoma black, except propleuron, disc of pronotum, posterior collar of pronotum, mesoscutellum, part of mesopleuron testaceous; metasoma partly testaceous and partly darkened or brown. Antenna clavate; antennomeres of holotype in following proportions: 7: 5: 15: 11: 10: 8: 7.5: 7: 6: 9.5; antennomeres of paratype in following proportions: 7: 3: 12.5: 9: 7: 7: 6: 6: 5: 6.5. Head shiny, slightly granulate; occipital carina complete; frontal line absent; frons with slender longitudinal furrow from anterior ocellus to clypeus; anterior region of frons with tuft of long setae. Head of holotype with POL = 2; OL = 3; OOL = 8.5; OPL = 5; TL = 10; greatest breadth of lateral ocelli shorter than OL (2: 3). Head of paratype with POL = 2; OL = 2; OOL = 7; OPL = 4.5; TL = 9; greatest breadth of lateral ocelli shorter than OL (1.5: 2). Pronotum shiny, unsculptured, not crossed by transverse impressions, with short anterior collar; pronotal tubercle reaching tegula. Mesoscutum very reduced, rugose; notauli complete, posteriorly joint. Mesoscutellum flat, shiny, unsculptured. Metanotum very short, shiny, unsculptured. Metapectal-propodeal disc dull, with strong anterior transverse and rugose furrow situated behind metanotum; metapectal-propodeal disc smooth, granulate; propodeal declivity reticulate rugose. Mesopleuron and metapleuron granulate, dull, with few transverse keels. Meso-metapleural suture distinct and complete. Mesopleuron with tuft of long setae situated on sides of transverse furrow behind metanotum. Protarsomeres of holotype in following proportions: 10: 2: 5: 13: 22. Protarsomeres of paratype in following proportions: 9: 2: 4: 10: 17. Enlarged claw (Fig. 83 C) with two peg-like lamellae situated further distally than proximal prominence. Protarsomere 5 (Fig. 83 C) with two rows of at least 32 – 40 lamellae; apex with four lamellae, among which one very long. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712A96FF3ED6D7FE1FFA1D.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 29. XI – 26. XII. 1997, YPT, S. van Noort & B. Fisher leg., SAM-HYM-P 024848 (SAMC). Paratypes: SOUTH AFRICA: Western Cape, Kogelberg Nature Reserve, 34 ° 15 ’ S 19 ° 05 ’ E, 16. XII. 1999 – 16. I. 2000, pitfall trap, S. van Noort leg., SAM-HYM-P 024849, 1 ♀ (SAMC); same locality label, 16. IV. 1999, S. van Noort leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712A96FF3ED6D7FE1FFA1D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712A96FF3ED6D7FE1FFA1D.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712AA9FF3ED3A2FE1FFE98.taxon	description	(Figs 81 C, D, 83 D, 84 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712AA9FF3ED3A2FE1FFE98.taxon	description	Description. ♀. Micropterous (Figs 81 C, D, 84 A); forewing very reduced, approximately as long as mesoscutel- lum, reaching transverse furrow behind mesoscutellum (Fig. 84 A). Body length 3.2 – 4.0 mm. Head black; mandible testaceous with teeth brown; malar space, clypeus and frontal longitudinal median area yellowish; occasionally only anterior margin of frons near clypeus yellowish (in specimen from South Africa, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, head brown, except mandibles, area between toruli, stripes along orbits and frontal longitudinal median area yellowish); antenna brown, with scape and pedicel partly testaceous (in specimen from South Africa, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, antenna brown, except scape, pedicel and flagellomeres 1 – 2 testaceous); mesosoma black, except posterior collar and lateral margins of pronotum testaceous (in specimen from South Africa, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, pronotum, mesoscutum and mesoscutellum brown, except posterior collar of pronotum brown-yellowish); tegula testaceous; metasoma brown-reddish; legs brown, except trochanters, articulations and tarsi testaceous (in specimen from South Africa, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, legs testaceous, except coxae and clubs of femora partly brown). Antenna clavate; antennomeres in following proportions: 8: 5: 13: 9: 8: 7: 5: 6: 5: 9. Head shiny, slightly punctate, unsculptured among punctures; occipital carina complete; usually with long longitudinal median furrow from anterior ocellus to clypeus (occasionally furrow reduced to small impression); POL = 3.5; OOL = 9; OPL = 5; TL = 8. Pronotum shiny, with lateral region slightly punctate; disc of pronotum very long and humped; posterior transversal impression separating pronotum disc from posterior collar narrower than disc; posterior margin of collar almost straight, reaching tegula, without posterior lobe directed towards tegula. Mesoscutum excavated, rugose, very reduced, approximately as long as posterior collar of pronotum. Mesoscutellum shiny, humped, unsculptured, approximately as long and as broad as mesoscutum. Metanotum not visible. Metapectal-propodeal disc humped, separated from mesoscutellum by deep transverse furrow (impression sculptured by transverse keels); metapectal-propodeal disc shiny, unsculptured; propodeal declivity dull, reticulate rugose. Meso-metapleural suture distinct, strong, complete. Mesopleuron and metapleuron dull, granulate, with slight transverse keels. Protarsomeres in following proportions: 9: 2: 5: 12: 21; protarsomere 2 produced into small hook. Enlarged claw (Fig. 83 D) with two bristles (one bristle in specimen from South Africa, 31 ° 20.602 ’ S 19 ° 11.695 ’ E) situated further distally than proximal prominence. Protarsomere 5 (Fig. 83 D) with one row of about 33 lamellae; apex with approximately three lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712AA9FF3ED3A2FE1FFE98.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, near Cape Town, Milnerton, II. 1926, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, 6 ♀♀ (NHMUK); same locality label as ho- lotype, I. 1926, 1 ♀ (NHMUK); same locality label as holotype, 4 ♀♀ (AMNH). Other material: SOUTH AFRICA: Eastern Cape, N. of Matatiele, 20 mi. from Qacha’s Nek, 1 ♀ (ZIL); Northern Cape, Hantam National Botanical Garden, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, 749 m, 28. IX – 29. XII. 2010, YPT, Hantam Succulent Karoo, S. van Noort leg., GL 07 - SUC 1 - Y 147, SAM-HYM-A 035618 (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712AA9FF3ED3A2FE1FFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF712AA9FF3ED3A2FE1FFE98.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4E2AABFF3ED138FB6CFBF8.taxon	description	(Figs 83 E, 86 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4E2AABFF3ED138FB6CFBF8.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: Western Cape, Koomplanskloof, 10 km S Citrus- dal, 32 ° 40 ’ S 19 ° 01 ’ E, 200 - 270 m, MT, 4 – 8. X. 1994, loc. 6, R. Danielsson leg. (ZIL). Paratype: SOUTH AFRICA: Western Cape, Cape Peninsula, Hout Bay, Skoorsteenkop, XII. 1950, n. 78, Swedish South Africa Expedition, 1950 – 1951, Brinck-Rudebeck, Insect trap, 1 ♂ (AMNH). Other material: SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 6. X. 2010 – 18. I. 2011, PCD 09 - ACA 1 - M 06, 1 ♂ (SAMC); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 23. II – 7. IV. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♂ (SAMC); same locality label, 29. X. 2009 – 23. II. 2010, 1 ♂ (SAMC); KwaZulu-Natal, Hlogoma, 4.92 km 127 ° SE Underberg, 29 ° 48.40 ’ S 29 ° 32.60 ’ E, 4. XII. 2001, sweep, S. van Noort leg., 1 ♂ (SAMC); Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.492 ’ S 19 ° 08.978 ’ E, 767 m, 27. X. 2009 – 9. II. 2010, MT, Nieuwoudtville Shale, Renosterveld, S. van Noort leg., 1 ♂ (SAMC); same locality label, 18. XI – 25. XII. 2008, 3 ♂♂ (SAMC); same locality label, 31 ° 24.182 ’ S 19 ° 08.587 ’ E, 741 m, 10. XII. 2007 – 13. I. 2008, 1 ♂ (SAMC); same locality label, 22. X – 18. XI. 2008, 7 ♂♂ (SAMC); same local- ity label, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, 749 m, 27. VIII – 27. X. 2009, 2 ♂♂ (SAMC); Northern Cape, Hantam National Botanical Garden, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, 749 m, 28. IX – 29. XII. 2010, MT, Hantam Succulent Karoo, S. van Noort leg., GL 07 - SUC 1 - M 153, SAM-HYM-A 035762, 4 ♂♂ (SAMC); Northern Cape, Swaarweerberg, Vredehoek Farm, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, 1613 m, 28. X. 2009 – 9. II. 2010, MT, Succulent Karoo, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 30. IX – 30. XII. 2010, SWA 09 - SUC 1 - M 06, 9 ♂♂ (SAMC); same locality label, YPT, 30. IX – 30. XII. 2010, SWA 09 - SUC 1 - Y 06, 1 ♂ (SAMC); Northern Cape, Avontuur Farm, 16 km NW Nieuwoudt- ville, 31 ° 16.249 ’ S 19 ° 02.900 ’ E, 764 m, 27. X. 2009 – 9. II. 2010, MT, Bokkeveld Sandstone Fynbos, S. van Noort leg., 7 ♂♂ (SAMC); same locality label, 28. IX – 29. XII. 2010, GL 07 - FYN 1 - M 152, 56 ♂♂ (SAMC); Northern Cape, West Driefontein farm, Succulent Karoo Site 1, 13.1 km 106 ° ESE Nieuwoudtville, 31 ° 22.178 ’ S 19 ° 15.064 ’ E, 13 – 16. X. 2000, MT, Succulent Karoo on shales, S. van Noort leg., 1 ♂ (SAMC); Northern Cape, Gamkaberg Na- ture Reserve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 4. IX. 2009 – 19. II. 2010, MT, Renosterveld, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Kogelberg Nature Reserve, C. 17. I. 2003, B. 30. I. 2003, Sf. 16. II. 2003, reared from an adult of Colistra parvulus (Linnavuori) feeding on Berzelia lanuginosa (Bruniaceae), M. Olmi reared (MOLC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 4. X – 1. XI. 1997, 29. XI – 26. XII. 1997, 8. IV – 16. V. 1998, YPT, S. van Noort & B. Fisher leg., 5 ♂♂ (SAMC); same locality label, 11. VIII – 6. IX. 1997, S. van Noort leg., 21 ♂♂ (20 in SAMC, 1 in MOLC); same locality label, 17 – 31. V. 1997, MT, S. van Noort leg., 2 ♂♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 26. XII. 1997, sweep, South Coast Strandveld, 57 m, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 732 ’’ S 18 ° 05 ’ 985 ’’ E, 5 km 290 ° E Langebaanweg, 16 – 23. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); same locality label, 18 – 25. IX. 2002, 1 ♂ (SAMC); same locality label, 9 – 16. X. 2002, 1 ♀ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 817 ’’ S 18 ° 06 ’ 837 ’’ E, 4 km 290 ° E Langebaanweg, 23 – 30. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 58 ’ 483 ’’ S 18 ° 06 ’ 884 ’’ E, 4 km 270 ° W Langebaanweg, 16 – 23. X. 2002, MT, Fynbos on limestone, S. van Noort leg., 4 ♂♂ (SAMC); same locality label, 9 – 16. X. 2002, 1 ♂ (SAMC); Western Cape, West Coast Fossil Park, 32 ° 57 ’ 732 ’’ S 18 ° 05 ’ 985 ’’ E, 5 km 290 ° E Langebaanweg, 11 – 18. IX. 2002, MT, rehabilitated mine dump, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Anyskop Farm, 4 km 270 ° W Langebaanweg, 32 ° 58.611 ’ S 18 ° 06.976 ’ E, 9 – 16. X. 2002, MT, Fynbos on Limestone, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Kamferskraal Farm, Site K 4, 32 ° 16.92 ’ S 23 ° 01.13 ’ E, 23. IV. 2001, MT, Nama Karoo on dolerite soils, S. van Noort & H. G. Robertson leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4E2AABFF3ED138FB6CFBF8.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in South Africa: Colistra parvulus (Linnavuori). Biology. The only parasitized host collected in Kogelberg Nature Reserve (Western Cape Province, South Africa) showed a dryinid thylacium in the dorsal side between head and pronotum. This dorsal position is usual in Deinodryinus species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4E2AABFF3ED138FB6CFBF8.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4E2AABFF3ED138FB6CFBF8.taxon	discussion	Remarks. The above described ♀ specimen was not reared. It was associated tentatively with ♂♂ collected together in South Africa, 32 ° 57 ’ 732 ’’ S 18 ° 05 ’ 985 ’’ E. Only a single ♀ specimen is known.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	description	(Figs 83 F, 84 B, C, 85 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	diagnosis	Diagnosis. Micropterous ♀ of Deinodryinus with mesoscutellum humped (Fig. 84 C); propodeal declivity granulate, not rugose, not transversely striate (Fig. 84 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	description	Description. ♀ (Figs 84 B, C, 85 A). Micropterous; forewing very reduced, slightly longer than mesoscutellum (5: 4), reaching transverse furrow behind mesoscutellum. Body length 2.8 mm. Head testaceous, except large brown spot in front of anterior ocellus and vertex and temple testaceous-darkened; mandible and clypeus testaceous; antenna with antennomeres 1 – 4 testaceous (rest of antenna missing); protorax brown, except disc and posterior surface of pronotum testaceous; mesoscutum brown; mesoscutellum testaceous; metanotum, metapectal-propodeal disc and propodeal declivity black, except mesopleuron brown; tegula testaceous; metasoma brown; legs testaceous, except metacoxa brown and clubs of femora slightly darkened. Antenna broken; antennomeres 1 – 4 in following proportions: 7: 3: 13: 10. Head shiny, granulate; frontal line absent; occipital carina complete; vertex flat; POL = 2; OL = 2; OOL = 8; OPL = 4; TL = 7; greatest breadth of posterior ocellus slightly shorter than POL (1.5: 2). Pronotum shiny, unsculptured, except lateral regions granulate; anterior collar of pronotum rugose; pronotum with anterior transverse furrow and humped and long disc; disc separated from posterior collar of pronotum by strong transverse impression; posterior collar narrower than disc; posterior margin of posterior collar curved, laterally reaching tegula. Mesoscutum concave, rugose, very reduced, approximately as long as posterior collar of pronotum. Mesoscutellum shiny, humped, unsculptured, slightly shorter and narrower than mesoscutum. Metanotum very short, reduced to transverse strip behind mesoscutellum. Metapectal-propodeal disc humped, separated from metanotum by transverse deep furrow sculptured by transverse keels; metapectal-propodeal disc shiny, unsculptured; disc and propodeal declivity dull, granulate. Meso-metapleural suture distinct, strong, complete. Mesopleuron and meta- pleuron granulate. Protarsomeres in following proportions: 8: 1.5: 5: 10: 16; no protarsomeres produced into hooks. Enlarged claw (Fig. 83 F) with two peg-like setae situated further distally than proximal prominence. Protarsomere 5 (Fig. 83 F) with two rows of about 28 lamellae; apex with approximately five lamellae (one of which very long). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. XII. 1999 – 16. I. 2000, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., SAM- HYM-P 036375 (SAMC). Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4C2AACFF3ED29FFBC5F8F5.taxon	etymology	Etymology. The species is named granulatus, referring to its head sculpture.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4A2AAEFF3ED0FAFE21FCBC.taxon	description	(Figs 91 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4A2AAEFF3ED0FAFE21FCBC.taxon	description	Description. ♀. Fully winged; body length 3.4 mm. Head black, with lateral margins, anterior surface of frons, part of vertex, temple, mandible and anterior half of clypeus brown-reddish; antenna testaceous, except antennomeres 5 – 7 darkened and 8 – 10 brown; mesosoma brown-reddish, with mesopleuron, metapleuron, metanotum and metapectalpropodeal disc black; metasoma brown; legs brown-reddish. Antenna clavate; antennomeres in following propor- tions: 12: 5: 8: 6.5: 3.5: 5.5: 6: 6: 5.5: 9. Head shiny, with frons sculptured by three median longitudinal keels (complete frontal line + two keels from lateral ocelli to antennal toruli; surface among these three keels and on sides sculptured by strong transverse subparallel keels); vertex and temple punctate, unsculptured among punctures; occipital carina complete; area behind lateral ocelli sculptured by two oblique keels connecting lateral ocelli to occipital carina; POL = 6; OL = 4; OOL = 8; OPL = 6.5; TL = 8. Pronotum shiny, setose, punctate, unsculptured among punctures, crossed by two strong transverse impressions (Figs 91 A, B); disc ovoidal, flat and surrounded by sharp margin (Figs 91 A, B). Mesosoma with strong constriction between prothorax and mesothorax (Fig. 91 A). Pronotal tubercle reaching tegula. Mesoscutum and mesoscutellum shiny, setose, finely punctate, unsculptured among punctures. Notauli absent. Metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with median area unsculptured, except few irregular keels near anterior margin. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Protarsomeres in following proportions: 6: 2: 6: 11: 24; Protarsomere 2 produced into hook. Enlarged claw (Fig. 91 C) with one long bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 91 C) with two rows of 3 + 28 lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4A2AAEFF3ED0FAFE21FCBC.taxon	materials_examined	Material examined. Type: ♀ holotype: GUINEA: Mt. Nimba, Gouan River, 07 ° 42 ’ N 08 ° 23 ’ W, 514 m, 7 – 15. I. 1991, FIT rainforest, L. Leblanc leg. (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4A2AAEFF3ED0FAFE21FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF4A2AAEFF3ED0FAFE21FCBC.taxon	distribution	Distribution. Guinea.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAEFF3ED5CCFE14F970.taxon	description	(Figs 85 B, 86 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAEFF3ED5CCFE14F970.taxon	description	Description. ♂ (Fig. 85 B). Fully winged; body length 2.4 – 2.7 mm. Head black, except mandible testaceous; an- tenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa partly brown. Antenna filiform, with setae about as long as, or shorter than breadth of antennomeres; antennomeres in following proportions: 9: 4: 8: 6: 5.5: 5.5: 5.5: 5.5: 5.5: 8. Head shiny, strongly punctate, unsculptured among punctures; frontal line absent; occipital carina complete; POL = 7; OL = 4; OOL = 4.5; OPL = 2.5; TL = 4; greatest breadth of the lateral ocelli shorter than OL (3: 4). Mesoscutum shiny, as sculptured as head. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculpured among punctures. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (12: 8). Paramere (Fig. 86 B) with two large and long apical inner branches (one of these branches with few papillae situated mainly on inner margin; second branch with mosaic pattern; volsellae located between two above branches). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAEFF3ED5CCFE14F970.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohipa- rara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04 (CAS). Other material: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 16. X – 8. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 01, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAEFF3ED5CCFE14F970.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAEFF3ED5CCFE14F970.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAFFF3ED170FE14FB24.taxon	description	(Figs 91 D – F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAFFF3ED170FE14FB24.taxon	description	Description. ♀. Fully winged; body length 5.0 – 5.6 mm. Testaceous-reddish, except ocellar triangle black, peti- ole black, metasoma partly brown and antennomeres 8 – 10 darkened. In specimen from Madagascar, 24 ° 45.31 ’ S 46 ° 51.13 ’ E, head and mesosoma ferruginous, except ocellar triangle darkened. Antenna clavate; antennomeres in following proportions: 18: 5: 18: 10: 8: 8: 9: 9: 9: 11. Head shiny, strongly reticulate rugose (in specimen from Madagascar, 24 ° 45.31 ’ S 46 ° 51.13 ’ E, vertex behind ocellar triangle and temple strongly punctate, unsculptured among punctures); frontal line present; ocellar triangle convex, with keels among ocelli; occipital carina complete; POL = 5; OL = 3; OOL = 10; OPL = 10; TL = 8. Pronotum (Figs 91 D, E) shiny, strongly punctate, with transverse keels, crossed by transverse impression separating anterior collar from posterior disc; lateral margin of posterior surface of pronotum sharp; pronotal tubercle reaching the tegula. Mesoscutum shiny, strongly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum shiny, slightly punctate, unsculptured among punctures. Metanotum shiny, unsculptured, with anterior margin very convex and raised into carina. Mesopleuron and metapleuron rugose, with slight keels; occasionally mesopleuron and metapleuron shiny, punctate, unsculptured among punctures; occasionally anterior half of mesopleuron very setose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels, reticulate rugose, except central smooth area. Forewing entirely darkened, or with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 7). Protarsomeres in following proportions: 10: 4: 6: 11: 26. Protarsomere 2 produced into hook. Enlarged claw (Fig. 91 F) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 91 F) with two-three rows of approximately 27 lamellae (all about same length) (in specimen from Madagascar, 24 ° 45.31 ’ S 46 ° 51.13 ’ E, three rows of about 48 lamellae, many lamellae longer than others); apex with approximately 6 – 11 lamellae, among which one much longer than others. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAFFF3ED170FE14FB24.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toamasina, Ivondro, V. 1940, A. Seyrig leg. (MNHN). Paratype: MADAGASCAR: Antananarivo, La Mandraka, IV. 1944, A. Seyrig leg., 1 ♀ (MRAC). Other material: MADAGASCAR: Toamasina, Périnet, 1 ♀ (AMNH); Toamasina, Botanic garden near entrance to An- dasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, 1025 m, 1 – 7. XI. 2001, MT, tropical forest, R. Harin’Hala leg., 2 ♀♀ (MOLC); Toliara, Andohahela National Park, Col de Tanatana, 33.3 km NW Tolagnaro, 24 ° 45.31 ’ S 46 ° 51.13 ’ E, 275 m, 22 – 24. XI. 2006, YPT, B. L. Fisher et al. leg., BLF 15099, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAFFF3ED170FE14FB24.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF492AAFFF3ED170FE14FB24.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF482AA0FF3ED2BBFE14FD9C.taxon	description	(Figs 86 C – E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF482AA0FF3ED2BBFE14FD9C.taxon	description	Description. ♂. Fully winged; body length 1.9 – 3.1 mm. Head black; mandible testaceous; antenna brown; meso- soma black; metasoma brown; legs testaceous, except metacoxa partly brown, club of metafemur and metatibia partly darkened. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 6.5: 4: 5: 5: 4.5: 4.5: 4: 4: 4: 7. Head shiny, very strongly punctate, unsculptured among punctures (punctures mostly strong and close on frons, similar to areolae); frontal line absent; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 3; TL = 3. Mesoscutum shiny, punctate, unsculptured among punctures (punctures very strong and close near anterior margin of mesoscutum). Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal or transverse keels. In specimen from Madagascar, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, propodeal declivity with two tracks of longitudinal keels (area betweense two tracks rugose). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (8: 7). Paramere (Figs 86 C-E) with distal region broadened and with small inner distal branch not wrapping penis. Tibial spurs 1 / 1 / 2. ♀. Uknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF482AA0FF3ED2BBFE14FD9C.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Toliara, Bereboka, 60 km NE Morondava, Bere- boka, 18 – 23. V. 1983, J. S. Noyes & M. C. Day leg., B. M. 1983 – 201 (NHMUK). Other material: MADAGASCAR: Antsiranana, Bekaraoka Forest, 6.8 km 60 ° ENE Daraina, 13 ° 00.00 ’ S 49 ° 42.36 ’ E, 150 m, 7. XII. 2003, tropical dry forest, general collecting night spiders, B. L. Fisher leg., BLF 9876, 4 ♂♂ (3 in CAS, 1 in MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 29. I – 11. II. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 03, 1 ♂ (MOLC); Mahajanga, Tsingy de Bemaraha Nat. Park, 10.6 km ESE 123 ° Antsalova, 19 ° 42.34 ’ S 44 ° 43.5 ’ E, 150 m, 16 – 20. XI. 2001, MT, tropical dry forest on Tsingy, Fisher, Griswold et al. leg., BLF 4462, 4 ♂♂ (3 in CAS, 1 in MOLC); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Marofandilla, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, 100 m, 28. XI – 3. XII. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4601, 1 ♂ (CAS); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 28. IV – 6. V. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 21, M. Irwin, R. Harin’Hala leg., 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF482AA0FF3ED2BBFE14FD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF482AA0FF3ED2BBFE14FD9C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF472AA1FF3ED423FE14FF50.taxon	description	(Fig. 91 G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF472AA1FF3ED423FE14FF50.taxon	description	Description. ♀. Fully winged; body length 2.7 – 5.0 mm (usually 4.1 – 5.0 mm). Testaceous; antenna testaceous, except antennomeres 3 – 10, or 5 – 10, or 6 – 10 brown; petiole black; metasoma totally testaceous, or partly brown. Small specimen (2.7 mm) from Madagascar, Ankarana, totally testaceous, except petiole black, metanotum brown and last antennomere darkened. In specimen from Madagascar, Périnet, antenna testaceous, except antennomeres 8 – 10 brown, frons and profemur brown-ferruginous, petiole black, stalks of meso- and metafemur brown. Antenna clavate; antennomeres in following proportions: 12: 7: 25: 10: 8: 8: 8: 8: 8: 11. Head shiny, with frons strongly punctate (almost reticulate rugose); vertex and temple strongly punctate, not reticulate rugose; frontal line complete or incomplete (then present only on anterior half of frons); occipital carina complete; ocellar triangle convex; POL = 5; OL = 4; OOL = 13; OPL = 12; TL = 8. Pronotum shiny, strongly punctate, unsculptured among punctures, crossed by transverse impression separating anterior collar from posterior disc; lateral margin of posterior surface of pronotum rounded; pronotal tubercle reaching tegula. Mesoscutum shiny, strongly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.4 – 0.5 × length of mesoscutum (in specimen from Madagascar, Périnet, notauli reaching about 0.8 × length of mesoscutum). Mesoscutellum and metanotum shiny, unsculptured. Anterior margin of metanotum very convex and raised into carina. Mesopleuron strongly punctate, with tuft of setae on anterior third. Metapleuron strongly punctate, not reticulate rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels, reticulate rugose, except central smooth area; in specimen from Madagascar, 24 ° 39.13 ’ S 43 ° 59.48 ’ E, propodeal declivity with few transverse keels near posterior extremity; in small specimen (2.7 mm) from Madagascar, Ankarana, propodeal declivity completely reticulate rugose. Forewing hyaline, with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (12: 9 in specimen from Madagascar, 24 ° 39.13 ’ S 43 ° 59.48 ’ E). Protarsomeres in following proportions: 11: 4: 7: 16: 30; no protarsomeres produced into hooks. Enlarged claw (Fig. 91 G) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 91 G) with two-three rows of lamellae (about 30 – 40 small and 7 – 9 longer); apex with approximately 9 – 15 lamellae, among which one much longer than others and pointed. Tibial spurs 1 / 1 / 2. ♂. Uknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF472AA1FF3ED423FE14FF50.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, Antananarivo (= Tananarive), Ch. Lamberton leg. (MRAC). Other material: MADAGASCAR: Antsiranana, Ankarana, IX. 1986, MT, Jane Wilson leg., 1 ♀ (NHMUK); Mahajanga, Tsimembo Forest, 8.7 km 336 ° NNW of Soatana, 19 ° 01.17 ’ S 44 ° 26.26 ’ E, 20 m, MT, tropical dry forest, 21 – 25. XI. 2001, Fisher, Griswold et al. leg., BLF 4507, 1 ♀ (MOLC); Toamasina, Périnet, 1 ♀ (MNHN); Toliara, Mahafaly Plateau, 6.2 km 74 ° ENE of Itampolo, 24 ° 39.13 ’ S 43 ° 59.48 ’ E, 80 m, MT, spiny forest thicket, 21 – 25. II. 2002, Fisher, Griswold et al. leg., BLF 5759, 1 ♀ (MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 23. VII – 6. VIII. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 18 B- 29, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF472AA1FF3ED423FE14FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF472AA1FF3ED423FE14FF50.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF462AA3FF3ED0F0FE14FC68.taxon	description	(Figs 86 F, 91 H)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF462AA3FF3ED0F0FE14FC68.taxon	description	Description. ♀. Fully winged; body length 3.0 – 5.0 mm. Reddish-ferruginous or reddish-darkened; mandible testa- ceous, except teeth brown; antenna testaceous, except antennomeres 7 – 10 brown; metapectal-propodeal disc black; metasoma brown. In specimen from Madagascar, 23 ° 25.82 ’ S 43 ° 50.05 ’ E, antenna totally brown. Antenna clavate; antennomeres in following proportions: 11: 5: 11: 6: 6: 6: 6: 6: 5: 7. Head shiny, strongly punctate, especially in anterior region, unsculptured among punctures; occipital carina complete; POL = 7; OL = 5; OOL = 9; OPL = 7; TL = 7. Pronotum shiny, strongly punctate, unsculptured among punctures, crossed by transverse impression separating anterior collar from posterior disc; lateral margins of posterior surface of pronotum rounded; pronotal tubercle reaching tegula. Mesoscutum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures; anterior margin of metanotum shiny, unsculptured, with anterior margin very convex and raised into carina. Mesopleuron shiny, strongly punctate, with tuft of setae on anterior third. Metapleuron partly unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity without longitudinal keels, reticulate rugose, except central smooth area. In specimen from Madagascar, 23 ° 25.82 ’ S 43 ° 50.05 ’ E, propodeal declivity without central smooth area. Forewing hyaline, with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 8). Protarsomeres in following proportions: 8: 3: 4: 8: 19; no protarsomeres produced into hooks. Enlarged claw (Fig. 91 H) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 91 H) with two rows of lamellae (about eight longer and 21 shorter); apex with approximately seven lamellae, among which one much longer than others; apex forming obtuse angle with rest of protarsomere 5. Tibial spurs 1 / 1 / 2. ♂ (supposed, because not reared; referred to D. madecassus by Benoit (1954 )): fully winged; body length 2.4 – 3.4 mm. Head black; mandible testaceous, except teeth brown; antenna brown, except scape and pedicel or only scape testaceous; mesosoma black; tegula testaceous; metasoma brown; legs brown, except tarsi and meso- and metatibia testaceous; occasionally legs totally brown. Antenna filiform, with setae at most as long as breadth of antennomeres; antennomeres in following proportions: 8: 4: 7: 5: 5: 5: 5: 5: 5: 8. Head shiny, strongly punctate (finely punctate in specimen from Madagascar, 13 ° 42.55 ’ S 50 ° 06.06 ’ E), unsculptured among punctures; frontal line incomplete, not present in front of anterior ocellus; occipital carina complete; POL = 8; OL = 4; OOL = 6; OPL = 5; TL = 4. Mesoscutum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 – 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity completely reticulate rugose, or with more or less large smooth area situated near distal extremity or in centre. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (10: 9). Paramere (Fig. 86 F) with large apical hatchet blade shaped branch wrapping penis, without dorsal proximal membranous process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF462AA3FF3ED0F0FE14FC68.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, IV. 1938, A. Seyrig leg. Paratypes: same locality label as holotype, VI. 1936, 1 ♀ (MNHN); same locality label as holotype, X. 1936, 1 ♀ (MNHN); same locality label as holotype, X. 1938, 1 ♀ (MRAC); same locality label as holotype, X. 1940, 1 ♀ (MRAC); same locality label as holotype, IX. 1938, ♂ allotype (MNHN). Other material: MADAGASCAR: Antsiranana, Ambondrobe, 41.1 km 175 ° Vohemar, 13 ° 42.55 ’ S 50 ° 06.06 ’ E, 10 m, 29. XI. 2004, littoral rainforest, MT, BLF 11099, B. L. Fisher leg., 1 ♂ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 21 – 28. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 1 ♂ (CAS); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 1 – 8. VI. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 25, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 2 ♂♂ (CAS); same locality la- bel, 24. VII – 4. VIII. 2002, MA- 02 - 09 C- 36, 1 ♂ (CAS); same locality label, 21. III – 12. IV. 2003, MA- 02 - 09 C- 57, 3 ♂♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 4 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 22 – 28. XI. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 04, 1 ♂ (CAS); same locality label, 15 – 25. VII. 2002, MA- 02 - 09 A- 35, 1 ♂ (MOLC); same locality label, 14 – 21. I. 2002, 2 ♂♂ (1 in CAS, 1 in MOLC); same locality label, 6 – 15. XII. 2001, MA- 02 - 09 A- 06, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 19, 1 ♂ (CAS); same locality label, 4 – 12. II. 2002, MA- 02 - 09 B- 15, 2 ♂♂ (CAS); same locality label, 14 – 24. VI. 2002, MA- 02 - 09 B- 32, 1 ♂ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 3 ♂♂ (CAS); Fianarantsoa, Befotaka-Midongy National Park, Pa- pango 27.7 km S Midongy-South Mt. Papango, 23 ° 50.07 ’ S 46 ° 57.49 ’ E, 940 m, 13 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14774, 2 ♂♂ (1 in CAS, 1 in MOLC); Fianarantsoa, Ranomafana, JIRAMA water works, 21 ° 14.91 ’ S 47 ° 27.13 ’ E, 690 m, 10 – 14. I. 2002, MT near river, R. Harin’Hala leg., MA- 02 - 09 A- 11, 1 ♂ (CAS); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, in dry wash east of In- terpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 7 – 15. IV. 2002, MT, in open area, MA- 02 - 11 B- 23, R. Harin’Hala leg., 1 ♂ (CAS); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 1 ♂ (CAS); Mahajanga, Sofia Dis- ctrict, 45 km S Antsohihy, 5 km W of Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 13 – 19. XII. 2010, MT, M. Irwin, R. Harin’Hala leg., MG- 61 - 01, 1 ♂ (CAS); Toliara, Andohahela Nat. Park, 1.7 km 61 ° ENE of Tsimelahy, 36.1 km 308 ° NW of Tolagnaro, 24 ° 55.48 ’ S 46 ° 38.44 ’ E, 300 m, 16 – 20. I. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4917, 1 ♂ (CAS); Toliara, Zombitse National Park, near national road, 22 ° 50.43 ’ S 44 ° 43.87 ’ E, 825 m, 13 – 20. III. 2002, MT, deciduous spiny forest, R. Harin’Hala leg., MA- 02 - 13 B- 20, 1 ♂ (MOLC); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 9 – 19. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 02, 1 ♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 6 – 16. I. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 18 B- 09, 1 ♂ (MOLC); Toliara, Ankazomena, 8 km N of Ambohimahavelona village, 23 ° 25.82 ’ S 43 ° 50.05 ’ E, 122 m, 21 – 31. X. 2008, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 51 - 01, 1 ♀ (CAS); Toliara, Fiherenana, 23 ° 10.37 ’ S 43 ° 57.39 ’ E, 100 m, 22 – 31. X. 2002, MT, gallery forest, MGF 042, Frontier Wilderness Project, 2 ♂♂ (CAS); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 49.85 ’ S 46 ° 32.17 ’ E, 80 m, 13 – 23. IX. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 32, 1 ♀ (MOLC); Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 26. II – 2. III. 2009, MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG- 53 C- 19, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF462AA3FF3ED0F0FE14FC68.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF462AA3FF3ED0F0FE14FC68.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA3FF3ED20FFE2AF959.taxon	description	(Fig. 87 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA3FF3ED20FFE2AF959.taxon	description	Description. ♂. Fully winged; body length 2.4 mm. Head black; mandible testaceous; antenna brown; mesosoma and metasoma black; legs testaceous, except metacoxa and metafemur partly brown. Antenna filiform, with setae at most as long as breadth of antennomeres; antennomeres in following proportions: 9: 5: 6: 6: 6: 6: 5.5: 5: 6: 8. Head dull, granulate and reticulate rugose; frontal line present; occipital carina complete; POL = 7; OL = 3; OOL = 5; OPL = 2; TL = 3. Mesoscutum shiny, setose, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels and median and lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3.5: 8.5). Paramere (Fig. 87 A) with large apical branch wrapping penis, without dorsal proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA3FF3ED20FFE2AF959.taxon	materials_examined	Material examined. Type: ♂ holotype: RWANDA: Kisenyi (= Gisenyi), Kayove (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA3FF3ED20FFE2AF959.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA3FF3ED20FFE2AF959.taxon	distribution	Distribution. Rwanda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA4FF3ED17EFE59FD2C.taxon	description	(Figs 85 C, D, 87 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA4FF3ED17EFE59FD2C.taxon	description	Description. ♂ (Figs 85 C, D). Fully winged; body length 2.1 mm. Head black; mandible testaceous; clypeus yellow- whitish; antenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa brown. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 7: 4: 5.5: 6: 6: 6: 5: 6: 5: 7. Head shiny, punctate, unsculptured among punctures; frontal line absent; frons with longitudinal median furrow; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 4; TL = 3; greatest breadth of posterior ocellus about as long as OL. Mesoscutum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc and propodeal declivity completely reticulate rugose, without longitudinal or transverse keels; propodeal declivity partly sculptured by areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (5: 7.5). Paramere (Fig. 87 B) with large inner distal branch wrapping penis. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA4FF3ED17EFE59FD2C.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Eastern Prov., Kasaala area, 2.07846 ° S 38.22530 ° E, 740 m, 28. XI – 4. XII. 2013, MT, just inside isolated woodland patch, J. Bukhebi & R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA4FF3ED17EFE59FD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF442AA4FF3ED17EFE59FD2C.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA4FF3ED4B3FE14F8A2.taxon	description	(Figs 85 E, 87 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA4FF3ED4B3FE14F8A2.taxon	description	Description. ♂ (Fig. 85 E). Fully winged; body length 2.0 – 2.6 mm. Head black, except mandible testaceous; an- tenna testaceous; mesosoma black; metasoma brown; legs brown, except articulations, tarsi, tibiae and trochanters testaceous. Antenna filiform; antennomeres in following proportions: 9: 4: 8: 6: 5.5: 6: 5: 5: 5: 8; antennomere 6 about twice as long as broad (6: 3); antennomere 8 less than 3 × as long as broad (5: 2.5). Head convex, dull, reticulate rugose; frontal line absent; occipital carina complete; POL = 7; OL = 4; OOL = 5; OPL = 7.5; TL = 4; greatest breadth of lateral ocelli about as long as OL. Mesoscutum shiny, strongly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum shiny, unsculptured. Mesopleuron shiny, punctate, unsculptured among punctures. Metapleuron shiny, unsculptured. Metapectal-propodeal disc without strong transverse posterior keel (however, under good light a hardly visible transverse keel is visible); metapectal-propodeal disc very short, dull, reticulate rugose; propodeal declivity dull, with two complete irregular longitudinal keels, with median and lateral areas completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (12: 13); 2 R 1 cell open. Paramere much shorter than penis in holotype, slightly shorter in paratype, with two inner branches, one of them long and narrow, the second one shorter (second shorter branch hardly visible in holotype) (Figs 87 C, D; fig. 8 in Olmi 2010 b wrong). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA4FF3ED4B3FE14F8A2.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Mahajanga, Namoroka National Park, 9.8 km 300 ° WNW Vilanandro, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, 140 m, 4 – 8. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445 (CAS). Paratype: same locality label as holotype, 1 ♂ (CAS). Other material: MADAGASCAR: Antsiranana, Ampondrabe Forest, 26.3 km 10 ° NNE Daraina, 12 ° 58 ’ 12 ’’ S 49 ° 42 ’ 00 ’’ E, 175 m, 10. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9976, 1 ♂ (CAS); Mahajanga, Mahavavy River, 6.2 km 145 ° SE Mitsin- jo, 16 ° 03 ’ 06 ’’ S 45 ° 54 ’ 30 ’’ E, 20 m, 1 – 5. XII. 2002, MT, gallery forest, Fisher, Griswold et al. leg., BLF 6930, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA4FF3ED4B3FE14F8A2.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA4FF3ED4B3FE14F8A2.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	description	(Figs 85 F, 90 A – E, 92 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	diagnosis	Diagnosis. Micropterous ♀ of Deinodryinus with head mostly black (Fig. 85 F 90 A – D); mesosoma black; pronotum granulate, except median region of posterior collar unsculptured; mesoscutellum humped (Fig. 90 B); metapectal- propodeal disc granulate; propodeal declivity granulate and reticulate rugose. Description. ♀ (Figs 85 F, 90 A – E). Micropterous; forewing very reduced, reaching transverse furrow behind metanotum, slightly longer than mesoscutellum (8.5: 8). Body length 4.4 – 5.0 mm. Head black, except mandible, clypeus and anterior region of frons between antennal toruli testaceous; antenna brown; mesosoma and petiole black; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 9: 5: 15: 12: 10: 8: 7: 7: 7: 9. Head shiny, punctate, very slightly granulate among punctures; temple and vertex behind ocellar triangle granulate; occipital carina complete; frontal line absent; frons with slender longitudinal furrow from anterior ocellus to clypeus; POL = 2; OL = 2; OOL = 11; OPL = 7; TL = 8; greatest breadth of lateral ocelli about as long as POL. Pronotum dull, granulate, except median region of posterior collar unsculptured; pronotum not crossed by transverse impressions, with short anterior collar; pronotal tubercle reaching tegula. Mesoscutum very reduced, rugose. Notauli absent. Mesoscutellum slightly humped, shiny, unsculptured. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, granulate, with strong anterior transverse and rugose furrow situated behind metanotum; propodeal declivity granulate and reticulate rugose. Mesopleuron and metapleuron granulate and rugose, dull. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 12: 3: 6: 13: 26. Enlarged claw (Fig. 92 A) with two peg-like lamellae situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 A) with two rows of about 52 lamellae; apex with four lamellae, among which one very long. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Blauwe Krans Farm, 12.8 km 216 ° SW Kirkwood, 33 ° 30.747 ’ S 25 ° 24.644 ’ E, 9 – 16. II. 2001, YPT, Valley Bushveld (non-trashed), S. van Noort leg., SAM-HYM-A 024057 (SAMC). Paratype: SOUTH AFRICA: Northern Cape, Swaarweerberg, Vredehoek Farm, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, 1613 m, 30. IX – 30. XII. 2010, YPT, Roggeveld Shale Renosterveld, SWA 09 - SUC 1 - Y 06, S. van Noort leg., 1 ♀ (MOLC); Western Cape, Gamkaberg Nature Reserve, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, 997 m, 4. X. 2010 – 25. I. 2011, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., GB 09 - FYN 2 - M 53, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF432AA6FF3ED136FBD9FC04.taxon	etymology	Etymology. The species is named nigropictus, because mostly black coloured.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF412AA7FF3ED584FE14F847.taxon	description	(Figs 90 F, 92 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF412AA7FF3ED584FE14F847.taxon	description	Description. ♀ (Fig. 90 F). Fully winged; body length 3.75 – 6.50 mm. Completely testaceous-reddish, except petiole black, antennomeres 5 – 10 or 7 – 10 brown and small black spots on ocellar triangle. Specimen from Madagascar, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, with head brown, except anterior third of frons, clypeus and mandible testaceous; antenna testaceous-brown, except antennomeres 7 – 10 brown; mesosoma brown, except part of lateral regions of prothorax and mesopleuron ferruginous, metanotum, metapectal-propodeal disc and propodeal declivity black; metasoma and legs brown. Very large specimen (6.5 mm) from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, testaceous-reddish, except head completely black, antennomeres 7 – 10 brown, club of profemur brown, stalk of metafemur black. In specimen from Madagascar, 22 ° 49.08 ’ S 43 ° 30.13 ’ E, head and mesosoma reddish-brown, except metapectal-propodeal disc and propodeal declivity black. Antenna clavate; antennomeres in following proportions: 9: 6: 15: 7: 7: 7: 7: 7: 7: 8. Head flat, shiny, strongly punctate, unsculptured among punctures; anterior third of frons very strongly punctate, almost reticulate rugose; frontal line incomplete, only present in anterior half of frons; frons without lateral keels near orbits directed towards antennal toruli; vertex behind ocellar triangle without two oblique keels connecting lateral ocelli to occipital carina; occipital carina complete; holotype with POL = 4; OL = 2; OOL = 10; OPL = 10; TL = 7.5. In specimen from Madagascar, 22 ° 49.08 ’ S 43 ° 30.13 ’ E, POL = 5; OPL = 6. Pronotum anteriorly crossed by strong transverse impression, shiny, finely punctate, unsculptured among punctures; posterior surface approximately as long as broad, shorter than mesoscutum (10: 23), with lateral margins sharp; pronotal tubercle reaching tegula Mesoscutum and mesoscutellum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.60 – 0.65 × length of mesoscutum. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity almost completely dull and reticulate rugose, except very small central unsculptured area. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (16: 11). Protarsomeres of holotype in following proportions: 10: 3: 6: 11: 24. Protarsomeres of specimen from Madagascar, 13 ° 27.52 ’ S 48 ° 33.06 ’ E, in following proportions: 12: 4: 6: 12: 28. In specimen from Madagascar, 22 ° 49.08 ’ S 43 ° 30.13 ’ E, protarsomere 1 longer than 4 (12: 8). Enlarged claw (Fig. 92 B) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 B) with three rows of approximately 44 lamellae (many lamellae much longer than others); apex with approximately 15 lamellae, among which one much longer than others. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF412AA7FF3ED584FE14F847.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antsiranana, Orangea Forest, 3.6 km 128 ° SE of Ra- mena, 12 ° 15.32 ’ S 49 ° 22.29 ’ E, 90 m, 22 – 28. II. 2001, beating low vegetation, in littoral rainforest, Fisher, Griswold et al. leg., BLF 3202 (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 4 – 19. III. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 08, 1 ♀ (CAS); Antsiranana, Ambato Forest, 26.6 km 33 ° NE Ambanja, 13 ° 27.52 ’ S 48 ° 33.06 ’ E, 150 m, 8. XII. 2004, MT, rainforest, B. L. Fisher leg., BLF 11518, 1 ♀ (CAS); Antsiranana, Ampasindava, Ambilanivy Forest, 3.9 km 181 ° S Ambaliha, 13 ° 47.55 ’ S 48 ° 9.42 ’ E, 800 m, 4 – 9. III. 2001, MT, rainforest, Fisher, Griswold et al. leg., BLF 3251, 1 ♀ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 26. XII. 2004 – 5. I. 2005, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 05, 1 ♀ (MOLC); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 1 – 12. VII. 2006, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 154, 1 ♀ (MOLC); Fianarantsoa, near Isalo National Park, in dry wash east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 7 – 22. IX. 2002, MT, in open area, MA- 02 - 11 B- 39, R. Harin’Hala leg., 1 ♀ (MOLC); Toliara, 5 km SE Manombo, Andranovato, 22 ° 49.08 ’ S 43 ° 30.13 ’ E, 18 m, 20 – 30. X. 2008, MT, in Euphorbia forest, M. Irwin, R. Harin’Hala leg., MG- 49 - 01, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF412AA7FF3ED584FE14F847.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF412AA7FF3ED584FE14F847.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	description	(Figs 81 A, 87 E, 92 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	description	Description. ♀. Micropterous; forewing very reduced, reaching transverse furrow behind mesoscutellum. Body length 2.5 – 2.9 mm. Completely yellow-testaceous, except antennomeres 6 – 10 or 7 – 10, petiole and distal half of metasoma brown-dark. Antenna clavate; antennomeres in following proportions: 7: 4: 18: 8: 8: 8: 7: 7: 6: 8. Head shiny, punctate, unsculptured among punctures; occipital carina complete; frontal line absent; POL = 3; OL = 2; OOL = 8; OPL = 5; TL = 9. Pronotum shiny, finely punctate, unsculptured among punctures, not crossed by transverse impressions, with short anterior collar; pronotal tubercle reaching tegula. Mesoscutum very reduced, unsculptured. Notauli present, posteriorly joint. Mesoscutellum flat, unsculptured. Metanotum absent. Metapectal-propodeal disc and propodeal declivity dull, reticulate rugose. Enlarged claw (Fig. 92 C) with one peg-like lamella situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 C) with two rows of about 18 – 28 lamellae; apex with 3 – 5 lamellae, among which one very long. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.9 – 2.2 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tarsi and protibia testaceous. Rarely body brown-reddish. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 6: 4: 7: 7: 7: 7: 6: 6: 6: 7. Head shiny, punctate, unsculptured among punctures (occasionally frons slightly granulate); frontal line absent; frons with longitudinal median furrow; occipital carina complete; head of holotype with POL = 4; OL = 2.5; OOL = 6; OPL = 5; TL = 5.5; in specimen from Senegal, 15 km S Guignol, POL = OPL; in specimen from Namibia, 23 ° 34 ’ S 15 ° 03 ’ E, POL = 5; OL = 2.5; OOL = 5. Mesoscutum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.3 – 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc and propodeal declivity completely reticulate rugose, without longitudinal or transverse keels; propodeal declivity sculptured by areolae less wide than those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3: 7). Parameres (Fig. 87 E) with inner proximal branch wrapping penis. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	materials_examined	Material examined. Types: ♀ holotype of P. paulyi: SENEGAL: Tattaguine (FSAE); ♂ holotype of D. nigerensis: NIGER: Maradi, Tarna goulbin, 2 – 6. IX. 1985, G. J. Steck leg. (USNM). Paratype of P. paulyi: CAPE VERDE ISLANDS: Saõ Tiago Island, Moia-Moia, 1 ♀ (AMNH). Paratypes of D. nigerensis: NIGER: same locality label as holotype, 10 – 13. IX. 1985, 2 ♂♂ (AMNH, TAMU). Other material: KENYA: Eastern Prov., base of Ukasi Hill, 0.82103 ° S 38.54443 ° E, 613 m, 21. XI – 5. XII. 2011, MT, Acacia - Commiphora savana, R. Copeland leg., 2 ♂♂ (NMK); same locality label, 7 – 21. XI. 2011, 3 ♂♂ (NMK); Rift Valley Prov., Olorgesailie National Monu- ment, 1.57930 ° S 36.44566 ° E, 982 m, 27. XI – 11. XII. 2011, MT, Acacia - Commiphora savana, R. Copeland leg., 1 ♂ (NMK); same locality label, 11 – 25. XII. 2011, 4 ♂♂ (NMK); Rift Valley Prov., Olorgesailie National Monument, 1.57962 ° S 36.44730 ° E, 979 m, 7 – 21. V. 2011, MT, near Ol Keju Nyiro River, R. Copeland leg., 1 ♂ (MOLC); Rift Valley Prov., 6 km S Kapedo, 750 m, 2 – 3. VI. 1980, B. Lamoral leg., 7 ♂♂ (PMA). NAMIBIA: 40 km W Keetman- shoop, along Rd B 4, C. 10. III. 2006, B. 16. III. 2006, Sf. 7. IV. 2006, reared from adult of Exitianus nanus (Distant), M. Olmi leg., 1 ♀ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 14. III. 2006, Sf. 7. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♀ (SMWN); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 10. III. 2006, Sf. 3. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 11. III. 2006, Sf. 4. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 12. III. 2006, Sf. 5. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (SAMC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 18. III. 2006, Sf. 6. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 18. III. 2006, Sf. 7. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (SAMC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 18. III. 2006, Sf. 7. IV. 2006, reared from adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♂ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 13. III. 2006, Sf. 5. IV. 2006, reared from adult of Exitianus okahandia Ross, M. Olmi leg., 1 ♂ (MOLC); Namib / Naukluft Park, Kuiseb River near Gobabeb, 23 ° 34 ’ S 15 ° 03 ’ E, 18. II – 20. III. 1983, 1 ♂ (SANC); Namib-Naukluft Park, Namib Des- ert Research Station, Kuiseb River, 23 ° 33.45 ’ S 15 ° 02.38 ’ E, 420 m, 17 – 24. II. 1998, MT, riparian vegetation, Kapofi & Irwin leg., 1 ♂ (TAMU); Windhoek, Hoffnung Farm, 7. II. 1972, lark side vegetation, Southern African Expedi- tion, B. M. 1972 – 1, 1 ♂ (NHMUK). NIGER: Maradi, Tarna goulbin, 5. VIII. 1985, G. J. Steck leg. (TAMU). SEN- EGAL: 15 km S of Guignol, 6. X. 1978, 1 ♂ (USNM). SOUTH AFRICA: Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. XII. 1998 – 16. I. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 1 ♂ (SAMC); same locality label, 16. XI – 16. XII. 1998, 1 ♂ (SAMC); same locality label, 16. III – 16. IV. 1999, 1 ♂ (SAMC); same locality label, 16. X – 16. XI. 1999, 5 ♂♂ (SAMC); same locality label, 16. XII. 1999 – 16. I. 2000, 6 ♂♂ (SAMC); same locality label, 16. XI – 16. XII. 1999, 2 ♂♂ (SAMC); same locality label, 16. I – 16. II. 2000, 12 ♂♂ (SAMC); same locality label, 16. II – 16. III. 1999, 21 ♂♂ (SAMC); same locality label, 16. XII. 1998 – 16. I. 1999, 9 ♂♂ (SAMC); same locality label, 16. XI – 16. XII. 1998, 16 ♂♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Namibia: Aconurella compta (Naudé); Exitianus nanus (Distant) and E. okahandia Ross.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	distribution	Distribution. Cape Verde Islands, Kenya, Namibia, Niger, Senegal, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5F2ABAFF3ED042FBD5FE98.taxon	discussion	Remarks. The leafhopppers captured in Namibia and parasitized by P. paulyi possessed a single large black thylacium situated dorsally between the head and the pronotum (Fig. 81 A). Based on the presence of a proximal inner branch wrapping the aedeagus, the ♂ of this species is very similar to Deinodryinus tussaci Olmi from Morocco. However, the distal part of 2 r-rs & Rs vein is shorter in D. tussaci than in D. paulyi.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	description	(Figs 92 D, 93, 94 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	diagnosis	Diagnosis. Macropterous ♀ of Deinodryinus with head strongly punctate, unsculptured among punctures (Fig. 94 A); frontal line absent; frons without lateral keels joining lateral ocelli to antennal toruli; posterior surface of pro- notum with sharp lateral margins; mesosoma without strong constriction between prothorax and mesothorax (Fig. 94 A); protarsomere 1 slightly shorter than 4; distal region of protarsomere 5 robust (Fig. 92 D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	description	Description. ♀ (Figs 93, 94 A). Fully winged; body length 3.4 – 3.5 mm. Head brown, except mandible, clypeus and anterior half of frons ferruginous; antenna testaceous, except antennomeres 7 – 10 brown; mesosoma brownblack, except lateral and posterior margins of pronotum and pronotal tubercle ferruginous; posterior half of median region of mesoscutum yellow, with margins ferruginous; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 7: 4: 12: 5: 4: 4: 5: 5: 5: 6.5. Head (Fig. 93 C) slightly swollen, shiny, strongly punctate, unsculptured among punctures; frontal line absent; frons without lateral keels near orbits directed towards antennal toruli; vertex behind ocellar triangle without two oblique keels connecting lateral ocelli to occipital carina; occipital carina complete; POL = 4; OL = 2; OOL = 6.5; OPL = 5; TL = 7; greatest breadth of lateral ocelli longer than OL (3: 2). Pronotum anteriorly crossed by strong transverse impression, shiny, finely punctate, unsculptured among punctures; posterior surface of pronotum approximately as long as broad, as long as mesoscutum, with lateral margins sharp; pronotal tubercle reaching tegula. Mesoscutum and mesoscutellum shiny, punctate, unsculptured among punctures (Fig. 93 A). Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, except posterior half sculptured by transverse keels. Forewing (Fig. 93 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 7). Protarsomeres in following proportions: 7: 2: 5: 10: 21. Enlarged claw (Fig. 92 D) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 D) with three rows of approximately 50 lamellae (many lamellae longer than others); apex with at least 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	materials_examined	Material examined. Types: ♀ holotype (CASTYPE 19456): MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 28. III – 8. IV. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 20 (CAS). Paratypes: same locality label as holotype, 1 ♀ (MOLC); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 30. XI – 7. XII. 2002, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 07, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5D2ABBFF3ED73FFF11FCBB.taxon	etymology	Etymology. The species is named piceus, because of the dark colour (piceus is a Latin adjective meaning dark brown).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5C2ABBFF3ED5C3FE1FF80B.taxon	description	(Figs 92 E, 94 B – E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5C2ABBFF3ED5C3FE1FF80B.taxon	description	Description. ♀ (Figs 94 B – E). Micropterous; forewing very reduced, approximately as long as mesoscutellum, reaching transverse furrow behind mesoscutellum. Hind wing absent. Body length 4.0 – 4.4 mm. Head black, except mandible, clypeus and anterior margin of frons testaceous; antenna black, except scape, pedicel and part of flagellomere 1 testaceous; mesosoma black, except posterior collar of pronotum testaceous; metasoma black; legs brown, except tarsi and meso- and metatibia testaceous. Antenna clavate; antennomeres in following proportions: 8: 5: 17: 11: 11: 9: 8: 7: 7: 9. Head dull, flat, strongly punctate, unsculptured among punctures, or very slightly granulate; frontal line absent; frons without median furrow; occipital carina complete; POL = 5; OL = 3; OOL = 10; OPL = 5; TL = 8. Pronotum shiny, very slightly granulate, with very humped and long anterior disc and posterior transverse impression; posterior collar short; posterior margin of posterior collar almost straight, reaching tegula, without posterior lobe directed towards tegula. Mesoscutum excavated, rugose, very reduced, longer than posterior collar of pronotum. Mesoscutellum shiny, humped, unsculptured, approximately as long and as broad as mesoscutum. Metanotum not visible. Metapectal-propodeal disc humped, separated from mesoscutellum by deep transverse furrow (impression sculptured by transverse keels). Metapectal-propodeal disc and propodeal declivity reticulate rugose. Meso-metapleural suture distinct, strong, complete. Mesopleuron and metapleuron reticulate rugose. Protarsomeres in following proportions: 11: 1.5: 8: 13: 25; protarsomere 2 produced into small hook. Enlarged claw (Fig. 92 E) with two bristles situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 E) with one row of about 44 lamellae; apex with approximately three lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5C2ABBFF3ED5C3FE1FF80B.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Alexandria, 33 ° 39 ’ S 26 ° 25 ’ E, G. L. Prinsloo leg. (SANC). Other material: SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 30. X. 2009 – 22. II. 2010, YPT, Amathole Mistbelt Grassland, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5C2ABBFF3ED5C3FE1FF80B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5C2ABBFF3ED5C3FE1FF80B.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	description	(Figs 87 F, 89 F, 92 F, 94 F, 95 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	description	Description. ♀ (Fig. 94 F). Fully winged; body length 4.1 – 5.1 mm. Testaceous-reddish, except head brown-reddish, median region of metasoma brown and petiole black (in specimen from Madagascar, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, head ferruginous, except ocellar triangle black, and metasoma totally ferruginous); antenna testaceous-reddish, except antennomere 10 brown (in specimen from Madagascar, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, antenna testaceous, except antennomeres 9 – 10 brown). Specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, testaceous-reddish, except head black and posterior half of metasoma darkened. In specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, head black, except part of mandible, part of malar space and occiput ferruginous. Antenna clavate; antennomeres in following proportions: 13: 7: 17: 9: 7: 8: 8: 8: 7.5: 11. Head convex, shiny, with frons completely reticulate rugose and vertex and temple strongly punctate, unsculptured among punctures; frontal line complete; frons without lateral keels near orbits directed towards antennal toruli; vertex behind ocellar triangle with tracks of two incomplete oblique keels connecting lateral ocelli to occipital carina; occipital carina complete; POL = 4; OL = 3; OOL = 7; OPL = 9; TL = 5; greatest breadth of lateral ocelli longer than OL (4.5: 3). Pronotum anteriorly crossed by strong transverse impression, shiny, finely punctate, unsculptured among punctures; posterior surface broader than long, shorter than mesoscutum (11: 24), with lateral margins sharp; pronotal tubercle reaching tegula. Mesoscutum and mesoscutellum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.9 × length of mesoscutum (0.8 × in specimen from Madagascar, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E). Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity almost completely dull and reticulate rugose, except very small central smooth area punctate and unsculptured among punctures. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (20: 14). Protarsomeres in following proportions: 11: 3.5: 6: 9: 22. Enlarged claw (Fig. 92 F) with one bristle situated further distally than proximal prominence. Protarsomere 5 (Fig. 92 F) with three rows of approximately 31 lamellae (many lamellae much longer than others); apex with approximately eight lamellae, among which one much longer than others; tibial spurs 1 / 1 / 2. ♂ (Fig. 95 A). Fully winged; body length 2.1 – 2.4. Head black, except mandible testaceous; antenna browntestaceous; mesosoma and metasoma black; legs testaceous, except metacoxa partly brown. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 6: 3.5: 6.5: 5: 5: 5: 5: 5: 4: 8. Head shiny, finely punctate, unsculptured among punctures; vertex without two oblique keels from lateral ocelli to occipital carina; frontal line absent; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 2; TL = 3; greatest breadth of lateral ocelli slightly shorter than OL (2.5: 3). Mesoscutum shiny, as sculptured as head. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculpured among punctures. Metapectal-propodeal disc dull, reticulate rugose, with transverse posterior keel (keel dorsally extending to anterior margin of metapectal-propodeal disc; metapectal-propodeal disc very reduced (Fig. 89 F )); propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (9: 8). Paramere (Fig. 87 F) with one very large distal inner branch (margins of branch showing many small papillae). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohipa- rara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 21 – 28. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 13 (CAS). Paratype: same locality label as holotype, 1 ♂ (CAS). Other material: MADA- GASCAR: Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 21. XII. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 07, 1 ♀ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♀ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 B- 12, 2 ♂♂ (CAS); Fi- anarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 4 – 14. VII. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 34, 1 ♂ (MOLC); same locality label, 16 – 26. II. 2003, MA- 02 - 09 C- 54, 1 ♀ (CAS); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality la- bel 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (CAS); Toamasina, Zahamena National Park, Onibe River, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, 780 m, 21 – 23. II. 2009, YPT, rainforest, BLF 22170, B. L. Fisher et al. leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF5B2ABDFF3ED128FAF0F8F4.taxon	discussion	Remarks. There is a possibility that D. pulcher is junior synonym of D. insulanus (Benoit, 1954).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF592ABFFF3ED30AFE1FFE70.taxon	description	(Fig. 96 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF592ABFFF3ED30AFE1FFE70.taxon	description	Description. ♀. Micropterous; forewing very reduced, approximately as long as mesoscutellum, not reaching trans- verse furrow behind mesoscutellum. Body length 4.7 mm. Head black; mandible testaceous with teeth brown; clypeus testaceous; antenna brown, except scape, pedicel and proximal half of flagellomere 1 testaceous; mesosoma black, except posterior collar and anterior margin of pronotum testaceous; tegula testaceous; metasoma black; legs brown, except distal part of coxae, trochanters, stalks of femora, tibiae and tarsi testaceous. Antenna clavate; antennomeres in following proportions: 8: 5: 17: 12: 10: 10: 8: 7: 7: 9. Head shiny, strongly punctate, unsculptured among punctures; occipital carina complete; vertex flat; POL = 4; OL = 3; OOL = 12; OPL = 6; TL = 11. Pronotum shiny, slightly punctate, unsculptured among punctures, with lateral regions granulate; pronotum with very humped and long anterior disc; posterior transversal impression separating pronotum disc from posterior collar narrower than disc; posterior margin of collar almost straight, reaching tegula, without posterior lobe directed towards tegula. Mesoscutum excavated, rugose, very reduced, approximately as long as posterior collar of pronotum. Mesoscutellum shiny, humped, unsculptured, approximately as long and as broad as mesoscutum. Metanotum not visible. Metapectal-propodeal disc humped, slightly granulate, separated from mesoscutellum by deep transverse furrow (impression sculptured by transverse keels); propodeal declivity strongly transversely striate. Meso-metapleural suture distinct, strong, complete. Mesopleuron and metapleuron strongly transversely striate. Protarsomeres in following propor- tions: 11: 3: 7: 12: 24; protarsomere 2 produced into small hook. Enlarged claw (Fig. 96 A) with two bristles situated further distally than proximal prominence. Protarsomere 5 (Fig. 96 A) with one row of about 43 lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF592ABFFF3ED30AFE1FFE70.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Drakensberg, Van Reenen, XI. 1926, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF592ABFFF3ED30AFE1FFE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF592ABFFF3ED30AFE1FFE70.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF582AB2FF3ED121FC98F8AC.taxon	description	(Figs 88 A – D, 95 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF582AB2FF3ED121FC98F8AC.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Mahajanga, Ankoririka Reserve, 10.6 km 13 ° NE of Tsaramandroso, 16 ° 16.2 ’ S 46 ° 2.55 ’ E, 210 m, 9 – 14. IV. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3665 (CAS). Paratypes: MADAGASCAR: Antananarivo, 11.5 km 47 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375, 6 ♂♂ (4 in CAS, 2 in MOLC). Other material: MADAGASCAR: Antananarivo, 46 km NE of Ankazobe, Ambohitantely, 18 ° 11.88 ’ S 47 ° 16.89 ’ E, 700 m, 15. X – 1. XI. 2004, MT, sclerophyl forst, M. Irwin & R. Harin’Hala leg., MA- 27 - 19, 1 ♂ (MOLC); Antsiranana, 3 km W of Sakalava Beach, white dunes, 12 ° 17.17 ’ S 49 ° 22.00 ’ E, 40 m, 27 – 30. I. 2001, MT, MA- 01 - 03 - 03, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 21 – 23. I. 2001, MA- 01 - 03 - 01, 3 ♂♂ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 27. IV – 13. V. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 08, M. E. Irwin & R. Harin’Hala leg., 1 ♂ (CAS); Antsiranana, Binara Forest, 7.5 km 230 ° SW Daraina, 13 ° 15.18 ’ S 49 ° 37.00 ’ E, 375 m, 1. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9557, 7 ♂♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Antsahabelela rain forest, 9 km SW Daraina, Binara, 13 ° 15.03 ’ S 49 ° 37.00 ’ E, 182 m, 17 – 24. XI. 2010, MT, in humid forest, M. Irwin, R. Harin’Hala leg., MG- 58 - 07, 1 ♂ (CAS); same locality label, 8 – 15. XII. 2010, MG- 58 - 10, 1 ♂ (CAS); same locality label, 11 – 19. I. 2011, MG- 58 - 15, 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 14 – 21. XI. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 07, 1 ♂ (CAS); same locality label, 19 – 26. XII. 2010, MG- 57 - 12, 2 ♂♂ (CAS, MOLC); same locality label, 14 – 21. II. 2011, MG- 57 - 19, 1 ♂ (CAS); same locality label, 11 - 18. IV. 2011, MG- 57 - 27, 3 ♂♂ (2 in CAS, 1 in MOLC); Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 1 – 8. VI. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 25, 1 ♂ (CAS); same locality label, 12 – 19. I. 2011, MG- 62 - 05, 1 ♂ (CAS); Antsiranana, Ambato Forest, 26.6 km 33 ° NE Ambanja, 13 ° 27.52 ’ S 48 ° 33.06 ’ E, 150 m, 8. XII. 2004, MT, rainforest, B. L. Fisher leg., BLF 11518, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 12 – 14. V. 2001, MT, MA- 01 - 01 D- 10, R. Harin’Hala leg., 3 ♂♂ (CAS); same locality label, 14 – 30. V. 2001, R. Harin’Hala leg., MA- 01 - 01 D- 11, 1 ♂ (CAS); Antsiranana, Orangea, 3 km E of Ramena near fort, 12 ° 14.49 ’ S 49 ° 22.17 ’ E, 65 m, 23 – 27. I. 2001, MT, littoral forest on sand, MA- 01 - 05 - 02, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 4 – 12. III. 2002, 1 ♂ (CAS); same locality label, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 15 – 25. VII. 2002, MA- 02 - 09 B- 35, 1 ♂ (CAS); same locality label, 24. V – 4. VI. 2002, MA- 02 - 09 B- 30, 5 ♂♂ (CAS); same locality label, 7 – 17. V. 2003, MA- 02 - 09 B- 60, 1 ♂ (CAS); same locality label, 27. VI – 7. VII. 2003, MA- 02 - 09 B- 65, 1 ♂ (CAS); Fianarantsoa, Rano- mafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 17, 3 ♂♂ (CAS); same locality label, 15 – 28. V. 2003, MA- 02 - 09 C- 61, 1 ♂ (MOLC); same locality label, 28. IV – 5. V. 2002, MA- 02 - 09 C- 27, 1 ♂ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 2 ♂♂ (CAS, MOLC); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (CAS); Fianarant- soa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 15 – 25. VII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 35, 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 14 – 21. I. 2002, 2 ♂♂ (CAS, MOLC); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 A- 07, 1 ♂ (CAS), 1 gynander specimen (MOLC); same locality label, 26 – 31. III. 2002, MA- 02 - 09 A- 22, 3 ♂♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 25. VII – 3. VIII. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 36, 1 ♂ (CAS); Fianarantsoa, Befotaka-Midongy National Park, Papango 27.7 km S Midongy-South Mt. Papango, 23 ° 50.07 ’ S 46 ° 57.49 ’ E, 940 m, 13 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14774, 1 ♂ (CAS); Mahajanga, Namoroka National Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, 100 m, 12 – 16. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6581, 6 ♂♂ (CAS); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♂ (MNHN); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, elev 97 m, 5 – 8 May 2011, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, M. Irwin, R. Harin’Hala, MT, Analagnambe dry degraded forest, MG- 61 - 11, 1 ♂ (CAS); same locality label, 2 – 6. I. 2011, MG- 61 - 04, 1 ♂ (CAS); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 2 ♂♂ (CAS); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 23 – 30. X. 2007, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 06, 1 ♂ (CAS); same locality label, 12 – 19. II. 2008, MG- 38 - 18, 1 ♂ (CAS); Mahajanga, Bemarivo Special Reserve, 23.8 km 223 ° SW Besalampy, 16 ° 55.30 ’ S 44 ° 22.06 ’ E, 19 – 23. XI. 2002, 30 m, sifted litter (leaf mold, rotten wood), in tropical dry forest, Fisher, Griswold et al. leg., BLF 6693, 4 ♂♂ (CAS); Mahajanga, Ankarafantsika National Park, Tsimaloto Forest, 18.3 km 49 ° NE Tsaramandroso, 16 ° 13 ’ 41 ’’ S 46 ° 08 ’ 37 ’’ E, 135 M, 2 – 8. IV. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3600, 5 ♂♂ (4 in CAS, 1 in MOLC); Mahajanga, Besalampy District, Marofototra palm forest, 17 km W of Besalampy, 16 ° 43.30 ’ S 44 ° 25.42 ’ E, 35 ft., 24. IX – 1. X. 2007, MT, palm trees on sand, M. Irwin, R. Harin’Hala leg., MG- 42 B- 01, 1 ♂ (CAS); same locality label, Marofototra dry forest, 170 ft., 19 – 26. XI. 2007, dry wash in forest, MG- 42 A- 09, 1 ♂ (CAS); same locality label, Marofototra dry forest, 170 ft., 1 – 8. X. 2007, dry wash in forest, MG- 42 A- 02, 1 ♂ (MOLC); Mahajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 1 ♂ (CAS); Toama- sina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 16 – 23. XI. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 20, 1 ♂ (CAS); Toliara, Mahafaly Plateau, 6.2 km 74 ° ENE of Itampolo, 24 ° 39.13 ’ S 43 ° 59.48 ’ E, 80 m, MT, spiny forest thicket, 21 – 25. II. 2002, Fisher, Griswold et al. leg., BLF 5759, 2 ♂♂ (CAS, MOLC); Toliara, Andohahela Nat. Park, Manantalinjo Forest, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, 24 ° 49.1 ’ S 46 ° 36.36 ’ E, 150 m, 12 – 16. I. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 4811, 15 ♂♂ (13 in CAS, 2 in MOLC); Toliara, Beza Mahafaly Reserve, parcelle I near research sta- tion, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 1 – 8. II. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 2 ♂♂ (CAS, MOLC); same locality label, 25. XII. 2001 – 2. I. 2002, 1 ♂ (CAS); same locality label, 9 – 20. IX. 2002, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 23. III – 2. IV. 2003, 1 ♂ (CAS); same locality label, 29. III – 10. IV. 2002, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue, 23 ° 41.39 ’ S 44 ° 34.53 ’ E, 180 m, 15. X – 10. XI. 2001, MT, spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 14 B- 01, 3 ♂♂ (1 in CAS, 2 in MOLC); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 16 – 17. XII. 2002, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 09, 1 ♂ (CAS); same locality label, 10 – 20. VII. 2003, 2 ♂♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 20 – 27. XI. 2001, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 03, 1 ♂ (CAS); same locality label, 23. IX – 3. X. 2002, MA- 02 - 18 A- 34, 2 ♂♂ (MOLC); same locality label, 19. II – 1. III. 2002, 2 ♂♂ (CAS, MOLC); same local- ity label, 23. VII – 6. VIII. 2002, 1 ♂ (CAS); same locality label, 6 – 16. I. 2002, MA- 02 - 18 A- 09, 1 ♂ (CAS); same locality label, 17 – 28. I. 2002, MA- 02 - 18 A- 11, 3 ♂♂ (2 in CAS, 1 in MOLC); Toliara, Mikea Forest, NW of Manom- bo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 23. VII – 6. VIII. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 18 B- 29, 1 ♂ (CAS); same locality label, 8 – 18. III. 2002, MA- 02 - 18 B- 16, 1 ♂ (CAS); Toliara, Andohahela National Park, Ihazo- fotsy Parcelle III, 24 ° 49.85 ’ S 46 ° 32.17 ’ E, 80 m, 20. V – 3. VI. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 20, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 13 – 24. II. 2003, MA- 02 - 21 - 10, 1 ♂ (CAS); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 50.09 ’ S 46 ° 29.21 ’ E, 80 m, 7 – 17. XII. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 41, 1 ♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 2 ♂♂ (CAS); same locality label, 27. XII. 2002 – 7. I. 2003, 1 ♂ (CAS); same locality label, 25. II – 7. III. 2003, 2 ♂♂ (CAS); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 1 ♂ (CAS); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 9 – 19. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 02, 1 ♂ (CAS); Toliara, Ivohibe Forest, 55 km N Tolagnaro, 24 ° 34.08 ’ S 47 ° 12.14 ’ E, 200 m, 2 – 4. XII. 2006, MT, B. L. Fisher et al. leg., BLF 15448, 4 ♂♂ (3 in CAS, 1 in MOLC); Toliara, Vohidava Forest, 88.9 km N Amboasary, 24 ° 14.26 ’ S 46 ° 17.16 ’ E, 500 m, 6 – 8. XII. 2006, B. L. Fisher et al. leg., BLF 15694, 1 ♂ (MOLC); Toliara, Androimpano Forest, 3 km E of Itampolo, 24 ° 39.47 ’ S 43 ° 57.37 ’ E, 45 m, 25. XII. 2008 – 1. I. 2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54 B- 09, 2 ♂♂ (CAS); same locality label, 6 – 13. XI. 2008, MG- 54 B- 03, 8 ♂♂ (6 in CAS, 2 in MOLC); Toliara, Androimpano Forest, 5 km E of Itampolo, 24 ° 39.02 ’ S 43 ° 57.79 ’ E, 130 m, 14 – 21. V. 2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54 C- 27, 7 ♂♂ (CAS); Toliara, Ankazomena, 8 km N of Ambohimahavelona village, 23 ° 25.82 ’ S 43 ° 50.05 ’ E, 122 m, 21 – 31. X. 2008, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 51 - 01, 2 ♂♂ (CAS); Toliara, Ranobe, 23 ° 02.783 ’ S 43 ° 36.617 ’ E, 20 m, 27. X – 3. XI. 2003, MT, spiny forest thicket, MGF 082, Frontier Wilderness Project, 3 ♂♂ (CAS); Toliara, Ifaty, near Hotel Paradisia, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, 9 m, 26. V – 5. VI. 2002, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02 - 16 - 28, 1 ♂ (CAS); same locality label, 13. X- 12. XI. 2001, MA- 02 - 16 - 01, 1 ♂ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI – 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 1 ♂ (CAS); Toliara, Tsifota, 20 km N Manombo, 1 km E of Tsifota village, 22 ° 49.08 ’ S 43 ° 22.36 ’ E, 15 m, 30. X – 9. XI. 2008, MT, spiny forest, M. Itwin, R. Harin’Hala leg., MG- 48 - 03, 2 ♂♂ (CAS); Toliara, 5 km SE Manombo, Andranovato, 22 ° 49.08 ’ S 43 ° 30.13 ’ E, 18 m, 20 – 30. X. 2008, MT, in Euphorbia forest, M. Irwin, R. Harin’Hala leg., MG- 49 - 01, 3 ♂♂ (CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 24 – 29. X. 2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 53 A- 01, 6 ♂♂ (CAS); same locality label, 4 – 12. XII. 2008, transitional forest, MG- 53 C- 07, 2 ♂♂ (CAS); same local- ity label, 24 – 29. X. 2008, across trail at base of escarpment, dense dry forest, MG- 53 B- 01, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF582AB2FF3ED121FC98F8AC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF582AB2FF3ED121FC98F8AC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF582AB2FF3ED121FC98F8AC.taxon	discussion	Remarks. A gynander specimen has been collected in the following locality: MADAGASCAR: Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 15 – 21. XII. 2001, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 07, one gynander specimen (MOLC). Description: general aspect of ♂; fully winged; body length 3.9 mm; head black, except mandible testaceous, clypeus testaceous, exept posterior margin black, one testaceous spot on malar space; antenna testaceous-ferruginous; mesosoma black; metasoma brown; legs testaceous-ferruginous. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 9: 6: 8: 11: 10: 10: 8: 8: 7: 11. Head dull, strongly rugose and sculptured by many irregular longitudinal keels; vertex without two oblique keels from lateral ocelli to the occipital carina; frontal line absent; occipital carina complete; POL = 11; OL = 5; OOL = 9; OPL = 8.5; TL = 6; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, as sculptured as head. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum; notauli hardly visible among longitudinal keels. Mesoscutellum shiny, partly rugose and partly sculptured by deep punctures, similar to areolae. Metanotum rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (30: 13). ♂ genitalia absent. Ovipositor present. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	description	(Figs 88 E, 96 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	description	Description. ♂. Fully winged; body length 1.1 – 2.4 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown, except tarsi, protibia and stalk of profemur testaceous. Antenna filiform, with setae longer than breadth of antennomeres; antennomeres in following proportions: 7: 4.5: 4.5: 4.5: 4.5: 4.5: 4: 4.5: 4.5: 6.5. Head shiny, slightly granulate and rugose or with irregular striae (in many specimens strongly granulate and rugose); vertex without two oblique keels from lateral ocelli to occipital carina; frontal line complete; occipital carina complete; POL = 7; OL = 3.5; OOL = 5; OPL = 2; TL = 2. Mesoscutum shiny, partly very slightly granulate and partly punctate and unsculptured among punctures (rarely mesoscutum completely unsculptured). Notauli thin, incomplete, reaching approximately 0.25 – 0.30 × length of mesoscutum (0.4 × in specimen from Kenya, 0.22121 ° N 37.11998 ° E). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc (Fig. 96 B) dull, reticulate rugose, with transverse posterior keel; propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (1.5: 7). Paramere (Fig. 88 E) with one very large and long inner branch not showing mosaic pattern. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	materials_examined	Material examined. Types: ♂ holotype: YEMEN: Ar Rujum, 15 ° 26 ’ N 43 ° 40 ’ E, 9. IV – 5. VI. 2001, MT, A. van Harten leg. (MOLC). Paratypes: YEMEN: same locality label as holotype, 13 ♂♂ (MOLC). Other material: KE- NYA: Rift Valley Prov., Lolldaiga Ranch, 0.22121 ° N 37.11998 ° E, 2061 m, 10 – 14. XII. 2013, MT, in meadow near small dam, R. Copeland & J. Muriuki leg., 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	distribution	Distribution. Kenya, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF552AB3FF3ED133FC44FC68.taxon	discussion	Remarks: Deinodryinus sabaeus is similar to Anteon madagascolum (Benoit 1954), but with the inner branch of the paramere much longer and more transverse (Fig. 88 E). The record from Madagascar reported by Azevedo et al. (2010) is incorrect: it should be referred to Anteon madagascolum.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF542AB4FF3ED20FFE14FE28.taxon	description	(Figs 88 F, 95 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF542AB4FF3ED20FFE14FE28.taxon	description	Description. ♂ (Fig. 95 C). Fully winged; body length 1.6 – 1.9 mm. Head black, except mandible testaceous; an- tenna testaceous-brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; pro- and mesoleg yellow; metaleg testaceous-brown, except coxa brown. Antenna filiform, with setae about as long as breadth of antennomeres; antennomeres in following proportions: 5: 3.5: 6: 5: 5: 4: 5: 5: 4.5: 7. Head shiny, very strongly punctate, unsculptured among punctures; areolae of frons and vertex very deep (head almost rugose); frontal line absent; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 2.5; TL = 2; greatest breadth of posterior ocellus as long as OPL. Mesoscutum and mesoscutellum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal or transverse keels, with usually median area partly smooth, shiny and unsculptured (occasionally median area dull, completely rugose). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 7). Paramere (Fig. 88 F) with two large and long apical inner branches (one of these branches showing few papillae situated mainly on inner margin; second branch showing mosaic sculpture; volsellae situated between two above branches; inner apical branch of paramere with apex very large). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF542AB4FF3ED20FFE14FE28.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, elev 97 m, 5 – 8. V. 2011, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, M. Irwin, R. Harin’Hala leg., MT, Analagnambe dry degraded forest, MG- 61 - 11 (CAS). Paratypes: 2 ♂♂ (CAS, MOLC). Other material: MADA- GASCAR: Antsiranana, Ankarana Special Reserve, 22.9 km 224 ° SW Anivorano Nord, 12 ° 54.32 ’ S 49 ° 6.35 ’ E, 80 m, 10 – 18. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 2856, 1 ♂ (CAS); Mahajanga, Sofia Dis- ctrict, 45 km S Antsohihy, 5 km W of Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 13 – 19. XII. 2010, MT, M. Irwin, R. Harin’Hala leg., MG- 61 - 01, 2 ♂♂ (CAS, MOLC); Toliara, Manombo, 22 ° 48.44 ’ S 43 ° 44.22 ’ E, 165 m, 20 – 25. V. 2004, MT, gallery forest, Frontier Wilderness Project, MGF 104, 1 ♂ (CAS); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 8 – 16. VI. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 26, M. Irwin, R. Harin’Hala leg., 2 ♂♂ (CAS, MOLC); same locality label, 1 – 9. VII. 2008, MG- 45 A- 29, 1 ♂ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI – 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF542AB4FF3ED20FFE14FE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF542AB4FF3ED20FFE14FE28.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	description	(Figs 89 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	description	Description. ♂. Fully winged; body length 2.9 – 3.2 mm. Head black, except mandible testaceous; antenna testa- ceous; mesosoma black; metasoma brown; legs testaceous, except pro- and metacoxa partly black; occasionally legs testaceous, except metacoxa partly black. Antenna filiform, with setae at most as long as breadth of antennomeres; antennomeres in following proportions: 11: 6: 11.5: 8: 8: 7: 7: 7.5: 7: 12. Head shiny, strongly punctate (punctures large and deep, similar to areolae; some regions of head seem reticulate rugose; in specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, frons completely reticulate rugose); frontal line complete; occipital carina complete; POL = 8.5; OL = 4; OOL = 7; OPL = 6; TL = 3. Mesoscutum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.7 – 0.8 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Mesopleuron punctate, unsculptured among punctures. Metapleuron shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without strong transverse posterior keel; propodeal declivity without longitudinal keels, with central shiny unsculptured area. Forewing with dark distal transverse band; distal part of 2 r-rs & Rs vein longer than proximal part (17: 13). Paramere (Figs 89 A-C) with one large apical band wrapping penis. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Fianarantsoa, 7 km W Ranomafana, 900 m, 1 – 7. III. 1990, MT, small clearing, montane rain forest, W. E. Steiner leg. (USNM). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 960 m, 21 – 26. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 1 ♂ (CAS); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 28. IV – 7. V. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 20, 1 ♂ (MOLC); same locality label, 28. VIII – 11. IX. 2005, MA- 31 - 29, 1 ♂ (CAS); same locality label, 18 – 25. III. 2005, MA- 31 - 16, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 1 ♂ (CAS); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 2 ♂♂ (CAS, MOLC); Fian- arantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 14 – 21. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 12, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 12 – 19. II. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 16, 1 ♂, 1 gynander (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 11, 2 ♂♂ (CAS); Toamasina, Sainte Marie Island, Ambohidena Forest, 22.8 km 44 ° NW Ambodifotatra, 16 ° 49.28 ’ S 49 ° 57.51 ’ E, 20 m, 21 – 24. XI. 2005, MT, littoral rainforest, Brian L. Fisher et al. leg., BLF 12842, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF532AB5FF3ED44FFBB3FDB8.taxon	discussion	Remarks. One gynander specimen of D. steineri is known from Madagascar, Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E (CAS). Description: general aspect of a ♂, without chelae. Fully winged; body length 3.7 mm. Head black, except mandible testaceous (with brown teeth) and anterior half of clyp- eus testaceous; antenna testaceous, except antennomeres 8 – 10 brown; mesosoma black; metasoma brown-black; legs testaceous-ferruginous. Antenna filiform, setose; antennomeres in following proportions: 10: 5: 6: 11: 10: 10: 9: 8: 8: 11. Head dull, reticulate rugose and with many irregular longitudinal keels; frontal line absent; occipital carina complete; POL = 10; OL = 5; OOL = 7; OPL = 8; TL = 7; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, reticulate rugose and with many irregular longitudinal keels. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing completely slightly darkened. Distal part of 2 r-rs & Rs vein much longer than proximal part (29: 11). Specimen with ovipositor, without ♂ genitalia. This specimen is attributed tentatively to D. steineri, because collected together with one normal ♂ specimen in the same locality. The reticulate rugose sculpture of the mesoscutum is not present in any species collected in the same locality (D. harinhalai, madecassus, pulcher, rusticus, steineri, suavis). The above sculpture is possibly present in the still unknown ♀ of D. steineri.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB5FF3ED4DFFE14F886.taxon	description	(Figs 89 D, 95 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB5FF3ED4DFFE14F886.taxon	description	Description. ♂ (Fig. 95 D). Fully winged; body length 2.7 – 2.9 mm. Head black, except mandible testaceous; an- tenna testaceous; mesosoma and metasoma black; legs testaceous, except metacoxa partly brown (occasionally all coxae partly brown; occasionally club of metafemur and metatibia brown). Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 9: 6: 8: 6.5: 6.5: 7: 7: 7.5: 7: 11. Head shiny, finely punctate, unsculptured among punctures; vertex without two oblique keels connecting lateral ocelli to occipital carina; frontal line very thin, almost complete, missing in front of anterior ocellus; occipital carina complete; POL = 7.5; OL = 3; OOL = 6; OPL = 4; TL = 2. Mesoscutum shiny, as sculptured as head. Notauli incomplete, reaching approximately 0.5 – 0.6 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculpured among punctures. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity almost completely reticulate rugose, with small central smooth and shiny area, without longitudinal or transverse keels. Forewing hyaline, usually with slight small dark spot just beneath 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein slightly longer than proximal part (13: 12). Paramere (Fig. 89 D) with one very large apical inner branch. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB5FF3ED4DFFE14F886.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375 (CAS). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 4 – 12. II. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 1 ♂ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 27. XII. 2005 – 5. I. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 39, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 14 – 22. VII. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 35, 1 ♂ (CAS); same locality label, 10 – 14. I. 2002, MA- 02 - 09 C- 11, 1 ♂ (CAS); Toliara, Ifaty, near Hotel Paradisia, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, 9 m, 26. V – 5. VI. 2002, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02 - 16 - 28, 3 ♂♂ (2 in CAS, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB5FF3ED4DFFE14F886.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB5FF3ED4DFFE14F886.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB6FF3ED11AFE1FFC90.taxon	description	(Fig. 89 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB6FF3ED11AFE1FFC90.taxon	description	Description. ♂. Fully winged; body length 1.6 – 1.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous. Antenna filiform, with setae at most as long as breadth of antennomeres; antennomeres in following proportions: 7.5: 5: 6: 6: 6: 6: 5.5: 5.5: 5.5: 7.5. Head shiny, punctate, unsculptured among punctures; frontal line absent; occipital carina complete; POL = 5; OL = 3; OOL = 4; OPL = 2.5; TL = 4; greatest breadth of lateral ocelli shorter than OL (2: 3). Mesoscutum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc with strong transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels and with median and lateral areas reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much shorter than proximal part (3.5: 8). Paramere (Fig. 89 E) with one large apical branch wrapping penis and one large dorsal proximal membranous process. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB6FF3ED11AFE1FFC90.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Umtamvuna Nature Reserve, 31 ° 03.509 ’ S 30 ° 10.484 ’ E, 160 m, 11 – 12. xi. 2000, MT, KW 00 - M 08, Coastal forest / Pondoland Coastal Plateau, sour Grassland margin, S. van Noort, SAM-HYM-P 024850 (SAMC). Paratype: same locality label as holotype, SAM- HYM-P 024850, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB6FF3ED11AFE1FFC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF522AB6FF3ED11AFE1FFC90.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	type_taxon	Type species. Lonchodryinus tricolor Kieffer 1905, by monotypy (Kieffer 1905).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	diagnosis	Diagnosis. ♀ (Figs 97 A, B): macropterous (Figs 97 A, B), or rarely brachypterous (only known in the Palaearctic re- gion); palpal formula 6 / 3; occipital carina complete; antenna without ADOs; pronotum with distinct anterior collar and posterior disc; metapectal-propodeal disc usually without transverse posterior keel (Fig. 97 A); in macropterous ♀♀, forewing with distal part of 2 r-rs & Rs vein as long as, or longer than proximal part (Figs 97 A, B); rarely distal part of 2 r-rs & Rs vein slightly shorter than proximal part; enlarged claw with inner proximal prominence bearing one long bristle (Fig. 96 F); tibial spurs 1 / 1 / 2. ♂ (Figs 97 C, D): always macropterous (even if ♀ brachypterous) (Figs 97 C, D); palpal formula 6 / 3; occipital carina complete; metapectal-propodeal disc usually without a transverse posterior keel (Fig. 97 D); forewing with distal part of 2 r-rs & Rs vein as long as, or longer than proximal part (Figs 97 C, D); rarely distal part of 2 r-rs & Rs vein slightly shorter than proximal part; forewing with pterostigma less than 4 × as long as broad (Figs 97 C, D); paramere without dorsal process, usually without inner branch wrapping penis (Fig. 96 C); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	distribution	Distribution. Worlwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	discussion	World species. 18 species are known, four in the Afrotropical region. Remarks. Larvae of Lonchodryinus were described by Guglielmino et al. (2017 a).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF512AB6FF3ED527FB88F89D.taxon	discussion	Remarks. Larvae of Lonchodryinus were described by Guglielmino et al. (2017 a).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	description	(Figs 96 C, D, 98 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	description	Description. ♂ (Fig. 98 A). Fully winged; body length 2.4 – 2.6 mm. Head black, except mandible testaceous; anten- na testaceous; mesosoma black; metasoma brown; legs of holotype testaceous, except metacoxa almost completely brown; legs of specimen from Madagascar, 23 ° 41.39 ’ S 44 ° 34.53 ’ E, brownish; in specimens from Madagascar, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, legs completely yellow, except metacoxa basally black. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 8: 4: 8: 6: 5: 5: 5: 5: 5: 8. Head dull, with vertex strongly punctate, unsculptured among punctures (vertex completely reticulate rugose in specimen from Madagascar, 23 ° 41.39 ’ S 44 ° 34.53 ’ E); in holotype vertex punctures very strong (head almost rugose); frons completely reticulate rugose; vertex without two oblique keels from lateral ocelli to the occipital carina; clypeus sculpured by many longitudinal keels, with anterior margin rounded, not bidentate; mandible quadridentate, with four teeth becoming regularly progressively larger from dorsal to ventral tooth; frontal line incomplete, missing in anterior half of frons; occipital carina complete; POL = 6; OL = 4; OOL = 4.5; OPL = 5; TL = 3. Mesoscutum shiny, very strongly punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity almost completely reticulate rugose, with central smooth and shiny area, without longitudinal or transversal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (10: 11). Paramere with (Fig. 96 C) or without (Fig. 96 D; in specimens from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E and 12 ° 19.22 ’ S 49 ° 20.17 ’ E) distal inner process and with many papillae on inner side. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW Anivorano Nord, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3011 (CAS). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 30. V – 6. VI. 2001, MT, MA- 01 - 01 D- 12, R. Harin’Hala leg., 1 ♂ (MOLC); Antsiranana, Montagne des Français, 7.2 km 142 ° SE Antsiranana, 12 ° 19.22 ’ S 49 ° 20.17 ’ E, 180 m, 22 – 28. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3130, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue, 23 ° 41.39 ’ S 44 ° 34.53 ’ E, 180 m, 15. X – 10. XI. 2001, MT, spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 14 B- 01, 1 ♂ (MOLC); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 19 – 26. XII. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 12, 1 ♂ (MOLC); same locality label, 14 – 21. II. 2011, MG- 57 - 19, 4 ♂♂ (2 in CAS, 2 in MOLC); same locality label, 11 - 18. IV. 2011, MG- 57 - 27, 2 ♂♂ (CAS, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FF502B48FF3ED1C5FB8DF87B.taxon	discussion	Remarks. It is possible that L. hovanus (for which only the ♂ is known) is the opposite sex of L. seyrigi (Benoit, 1954) or L. pauliani (Benoit, 1954). Only ♀ specimens are known of these two species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	description	(Figs 96 E, 98 B, 99)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	diagnosis	Diagnosis. ♀ of Lonchodryinus with head and prothorax mostly black (Fig. 98 B); forewing with two dark transverse bands; protarsomere 5 with three rows of lamellae, one of which composed of lamellae about as long as breadth of protarsomere 5 (Fig. 96 E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	description	Description. ♀ (Figs 98 B, 99). Fully winged; body length 4.6 mm. Head black, except mandible and clyp- eus ferruginous; antenna testaceous, except antennomeres 8 – 10 brown; prothorax black, except lateral margins of pronotum and pronotal tubercle ferruginous; rest of mesosoma ferruginous-reddish, except proximal extremity of mesopleuron black; tegula testaceous; metasoma and legs ferruginous-reddish. Antenna clavate; antennomeres in following proportions: 14: 8: 21: 12: 9: 8: 8: 9: 9: 11. Head (Fig. 99 C) shiny, strongly punctate, unsculptured among punctures; frontal line complete; frons without keels along orbits directed towards antennal toruli; POL = 5; OL = 3; OOL = 11; OPL = 11; TL = 7; greatest breadth of lateral ocelli shorter than OPL (5: 11); occipital carina complete. Pronotum shiny, with few punctures, unsculptured among punctures, with strong anterior transverse furrow; pronotal tubercle reaching tegula. Posterior surface of pronotum shorter than mesoscutum (15: 27). Mesoscutum (Fig. 99 A) shiny, with few punctures, unsculptured among punctures. Notauli incomplete, reaching approximately 0.8 × length of mesoscutum. Mesoscutellum and metanotum swollen, shiny, unsculptured. Mesopleuron shiny, unsculptured, except anterior black area and posterior third rugose. Metapleuron shiny, unsculptured. Metapectal-propodeal disc without transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with central area unsculptured and shiny. Forewing (Fig. 99 D) hyaline, with two dark transverse bands; distal part of 2 r-rs & Rs vein about as long as proximal part; 2 R 1 cell open. Protarsomeres in following proportions: 13: 4: 6: 15: 31; protarsomere 3 produced into hook; protarsomere 2 not produced into hook. Enlarged claw (Fig. 96 E) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 96 E) with three rows of approximately 60 lamellae (one row is composed of 11 lamellae about as long as breadth of protarsomere 5); apex with approximately 12 lamellae (among which one very long lamella). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19468): MADAGASCAR: Fianarantsoa, Ranomafana Na- tional Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 26. II – 10. III. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 55 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAE2B4AFF3ED6D7FED6FD9C.taxon	etymology	Etymology. The Latin adjective madagascolus means “ inhabiting Madagascar ”, country where the new species was collected.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4AFF3ED423FE14F9CF.taxon	description	(Fig. 96 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4AFF3ED423FE14F9CF.taxon	description	Description. ♀. Fully winged; body length 5.5 mm. Head yellow-testaceous; antenna testaceous, except anten- nomeres 8 – 10 brown; mesosoma reddish-dark, except metapleuron, metapectal-propodeal disc and propodeal declivity black; mesopleuron black, with reddish spot on anterior half; tegula testaceous; metasoma black; proleg completely testaceous-reddish; meso- and metaleg black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 14: 7: 27: 12: 8: 8: 8: 8: 8: 11. Head shiny, strongly punctate mainly on frons), unsculptured among punctures; frontal line complete; frons without keels near orbits directed towards antennal toruli; POL = 5; OL = 3; OOL = 14; OPL = 13; TL = 11; occipital carina complete. Pronotum shiny, finely punctate, unsculptured among punctures, with strong anterior transverse impression; pronotal tubercle reaching tegula. Mesoscutum shiny, finely punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.75 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Mesopleuron reticulate rugose. Metapleuron shiny, unsculptured. Metapectal-propodeal disc without transverse posterior keel, reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels, with central unsculptured area. Forewing hyaline, with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part, almost reaching margin of wing; pterostigma more than 4 × as long as broad (19: 4). Protarsomeres in following proportions: 13: 4: 7: 16: 29; protarsomere 3 produced into hook; protarsomere 2 not produced into hook. Enlarged claw (Fig. 96 F) with proximal prominence bearing one long bristle. Protarsomere 5 (Fig. 96 F) with three rows of approximately 45 lamellae (some of these very long); apex with approximately seven lamellae (among which one very long lamella). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4AFF3ED423FE14F9CF.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antsiranana, Mt. Tsaratanana, 1500 m, forêt de mousses, X. 1949, R. Paulian leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4AFF3ED423FE14F9CF.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4AFF3ED423FE14F9CF.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4BFF3ED0D4FE14FE28.taxon	description	Description. ♀. Fully winged; body length 3.0 mm. Holotype (only known specimen) very damaged, without head, antennae and prolegs. According to the original description, head black, except palpi, mandible and clypeus yellow; antenna yellow, except antennomeres 7 – 10 black. Mesosoma black, except margins of pronotum reddish; metasoma black. Prolegs missing in holotype. According to the original description, proleg yellow. Meso- and metaleg completely testaceous. Pronotum shiny, very strongly punctate; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Metapectal-propodeal disc dull, completely reticulate rugose, without keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 7). Chela missing in holotype. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4BFF3ED0D4FE14FE28.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, III. 1938, A. Seyrig leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4BFF3ED0D4FE14FE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAD2B4BFF3ED0D4FE14FE28.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	type_taxon	Type genus. Bocchus Ashmead 1893, designated by Olmi (1984).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	diagnosis	Diagnosis. ♀ (Figs 100 A, B): fully winged (Figs 100 A, B); occasionally brachypterous or micropterous or apterous; occipital carina complete; ocelli present; mandible with one, two, three or four teeth (Figs 8 A, B, C, D); quadridentate mandible usually with three large teeth and one rudimentary tooth between two posterior teeth (Fig. 8 B); rarely quadridentate mandible with teeth becoming regularly progressively larger from dorsal to ventral tooth (as in Fig. 6 B); palpal formula 6 / 3; antenna without tufts of long setae and without ADOs; pronotal tubercle present; epicnemium not visible, because lateral regions of prothorax continuous with mesopleura (as in Fig. 7 C); forewing of fully winged specimens with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 100 A, B); forewing with 2 r-rs & Rs vein (Figs 100 A, B); rarely 2 r-rs & Rs vein absent; pterostigma of fully winged specimens present (Fig. 100 B), often very reduced (Fig. 100 A); protarsus chelate (Figs 100 A, B); chela with rudimentary claw (Fig. 101 A); protrochanter short and slightly longer than broad; tibial spurs 1 / 1 / 1, or 1 / 1 / 2. ♂ (Figs 100 C, D): fully winged (Figs 100 C, D), rarely micropterous (Fig. 107 E) or apterous (Mystrophorus apterus Ponomarenko, not present in the Afrotropical region); occipital carina complete; mandible with one, two, three or four teeth (Figs 8 A, B, C, D); quadridentate mandible usually with three large teeth and one rudimentary tooth between two posterior teeth (Fig. 8 B); palpal formula 6 / 3; epicnemium invisible, because lateral regions of prothorax continuous with mesopleura (Fig. 7 C); in fully winged specimens forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 100 C, D); in fully winged specimens forewing with 2 r-rs & Rs vein (Figs 100 C, D); pterostigma present (Figs 100 C, D); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	biology_ecology	Hosts. Caliscelidae, Tropiduchidae, Cicadellidae (except Idiocerinae, Macropsinae and Typhlocybinae) (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	distribution	World species. 112 species are known, 33 in the Afrotropical region. World genera. Three genera are known, one in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEAC2B4BFF3ED44FFD25F918.taxon	discussion	Remarks. The following genera have not been recorded from the Afrotropical region: Mystrophorus Förster 1856 (Western Palaearctic) and Mirodryinus Ponomarenko 1972 (Western Palaearctic). Larvae of Mystrophorus were described by Guglielmino & Bückle (2010).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA92B41FF3ED0B9FEB0FE98.taxon	description	(Figs 101 A, 102 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA92B41FF3ED0B9FEB0FE98.taxon	description	Description. ♀. Fully winged; body length 3.1 – 4.1 mm. Head completely reddish or black, except mandible and occasionally also clypeus testaceous; occasionally head black, except region between ocelli and occipital carina reddish; antenna completely testaceous, or brown, except scape and pedicel or also flagellomere 1 testaceous; mesosoma completely reddish, occasionally with metapectal-propodeal disc and propodeal declivity completely or partly black; metasoma black; legs testaceous. Head dull, setose, completely reticulate rugose; occasionally head granulate on sides of frontal line; occipital carina complete; ocellar triangle equilateral. Pronotum setose. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli complete, posteriorly separated; in paratype with head reddish (in MOLC) notauli almost invisible. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels. Forewing with two dark transverse bands; pterostigma short and narrow. Protarsomeres 1 – 4 in following proportions: 13: 2: 2: 8. Enlarged claw (Fig. 101 A) with one row of five-six teeth and one bristle. Protarsomere 5 (Fig. 101 A) with one preapical lamella and two bristles. Tibial spurs 1 / 1 / 1. ♂. Fully winged; body length 2.0 – 3.0 mm (2.0 – 2.6 mm in African specimens; 2.9 – 3.0 mm in specimens from Ye- men). Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 13: 7: 8: 8: 8: 9: 8: 8: 8: 10.5. Head swollen, dull, granulate and reticulate rugose; frontal line complete; occipital carina complete. Head of specimen from Uganda, Kichwamba, with POL = 6; OL = 3; OOL = 8; OPL = 5; TL = 2.5; head of specimen from Yemen, 12 km NW Manakhah, with POL = 6; OL = 3.5; OOL = 7.5; OPL = 8; TL = 7; head of specimen from Madagascar, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, with POL = 3; OL = 3; OOL = 7; OPL = 7; TL = 4. Mesoscutum, mesoscutellum and metanotum dull, completely granulate and reticulate rugose; occasionally mesoscutellum partly granulate. Notauli complete and posteriorly separated; in some specimens from Madagascar mesoscutum with slight hardly visible tracks of notauli. In specimens from Uganda, Kichwamba, minimum distance between notauli longer than greatest breadth of lateral ocelli (8: 3); in specimens from Yemen, 12 km NW Manakhah, minimum distance between notauli as long as temple: 7: 7. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with two complete longitudinal keels (occasionally hardly visible); in specimens from Uganda, Kichwamba, and Tanzania, Kikolo Plot, propodeal declivity with median and lateral areas completely reticulate rugose; in specimens from Yemen, 12 km NW Manakhah, and from Tanzania, Mkomazi Game Reserve, propodeal declivity with median area partly shiny and unsculptured. In specimen from Kenya, 2.07836 ° S 38.22517 ° E, propodeal declivity with two incomplete longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (10: 12 in specimens from Uganda, Kichwamba; 7: 12 in specimens from Yemen, 12 km NW Manakhah); 2 R 1 cell open. In specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, apex of basivolsella broadened and with subdistal inner pointed apophysis (Fig. 102 A); in specimen from Uganda, Kichwamba, apex of basivolsella not broadened (Fig. 102 B). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA92B41FF3ED0B9FEB0FE98.taxon	materials_examined	Material examined. Types: ♀ holotype: SOMALIA: Lower Shebelle, Afgooye (= Afgoi), 1 – 15. II. 1977, MT, N. Bin leg. (AMNH). Paratypes: GHANA: Kumasi, Kwadaso, 28. III. 1977, CIBC, Scheibelreiter leg., 1 ♀ (with black head) (AMNH). NIGERIA: Oyo State, Ibadan, 1 ♀ (CNC); Ogun State, Ikenne, IX. 1974, J. T. Medler leg., 1 ♀ (NHMUK). SOMALIA: same locality label as holotype, 14 – 18. IV. 1977, N. Bin leg., 1 ♀ (AMNH); same locality label as holotype, 30. V – 7. VI. 1978, G. Fiori leg., 1 ♀ (AMNH). Other material: BOTSWANA: Smiti Grassland, 19 – 26. IX. 1975, A. R. Smith leg., 1 ♂ (NHMUK). KENYA: Coast Prov., Boni Forest area, near Bodhei Village, 1.85602 ° S 40.69880 ° E, 31 m, 6 – 18. VI. 2013, 2 m Mt, mixed grass- and woodland, J. Bukhebi & R. Copeland leg., 1 ♂ (MOLC); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 8 – 22. I. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 2 ♂♂ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 8 – 15. IV. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♂ (NMK); Rift Valley Prov., Lake Nakuru National Park, 0.47203 ° S 36.06388 ° E, 1908 m, 6 – 20. I. 2006, MT, mixed Olea, Cussonia, Vepris, R. Copeland leg., 1 ♂ (NMK); Rift Valley Prov., 15 mi. E Talec River, near Mara Game Ref., 11. VIII. 1984, D. E. Russell leg., 1 ♂ (DEUCD). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 14 – 30. V. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 11, 1 ♂ (MOLC); Antsiranana, Diana Re- gion, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 26. I – 2. II. 2007, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 07, 2 ♂♂ (CAS, MOLC); same locality label, 1 – 8. VI. 2011, MG- 62 - 25, 2 ♂♂ (MOLC); Antsiranana, SAVA Region, Vohemar District, Antsahabelela rain for- est, 9 km SW Daraina, Binara, 13 ° 15.03 ’ S 49 ° 37.00 ’ E, 182 m, 17 – 24. XI. 2010, MT, in humid forest, M. Irwin, R. Harin’Hala leg., MG- 58 - 07, 4 ♂♂ (2 in CAS, 2 in MOLC); Antsiranana, SAVA Region, Vohemar District, An- dranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 14 – 21. XI. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 07, 1 ♂ (CAS); same locality label, 19 – 26. XII. 2010, MG- 57 - 12, 2 ♂♂ (CAS, MOLC); same locality label, 14 – 21. II. 2011, MG- 57 - 19, 1 ♂ (CAS); same locality label, 11 - 18. IV. 2011, MG- 57 - 27, 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 19 – 26. XII. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 12, 1 ♂ (CAS); same locality label, 4 – 10. X. 2010, MG- 57 - 01, 2 ♂♂ (CAS); same locality label, 11 - 18. IV. 2011, MG- 57 - 27, 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 1 ♂ (CAS); Fianarantsoa, Isalo National Park, Sahanafa River, 29.2 km 351 ° N Ranohira, 22 ° 18.48 ’ S 45 ° 17.30 ’ E, 500 m, 10 – 13. II. 2003, MT, gallery forest, BLF 7653, Fisher, Griswold et al. leg., 1 ♂ (CAS); Mahajanga, Bemarivo Special Reserve, 23.8 km 223 ° SW Besalampy, 16 ° 55.30 ’ S 44 ° 22.06 ’ E, 19 – 23. XI. 2002, 30 m, sifted litter (leaf mold, rotten wood), tropical dry forest, Fisher, Griswold et al. leg., BLF 6693, 3 ♂♂ (2 in CAS, 1 in MOLC); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, 43 m, 17 – 24. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 11, 2 ♂♂ (CAS); Toliara, Andohahela National Park, Manantalinjo Forest, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, 24 ° 49.1 ’ S 46 ° 36.36 ’ E, 150 m, 12 – 16. I. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 4811, 1 ♂ (CAS); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Marofandilla, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, 100 m, 28. XI – 3. XII. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4601, 1 ♂ (CAS); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.09 ’ S 044 ° 39.38 ’ E, 14 – 21. XI. 2007, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 02, 1 ♂ (MOLC); Toliara, Kirindy Mite National Park, 16.3 km 127 ° SE Belo sur Mer, 20 ° 47.43 ’ S 44 ° 08.49 ’ E, 80 m, 6 – 10. XII. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4730, 4 ♂♂ (CAS); Toliara, Antafoky, 23 ° 28.43 ’ S 44 ° 03.51 ’ E, 55 m, 25 – 28. I. 2002, MT, gallery forest, Frontier Project leg., MGF 003, 2 ♂♂ (CAS); Toliara, Berenty Private Reserve, Malaza Forest, Man- draré River, 8.6 km 314 ° NW Amboasary, 25 ° 00.28 ’ S 46 ° 18.22 ’ E, 40 m, 6. II. 2002, sifted litter (leaf mold, rotten wood), gallery forest, Fisher, Griswold et al. leg., BLF 5426, 1 ♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 27. XII. 2002 – 7. I. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 11, 13 ♂♂ (8 in CAS, 5 in MOLC); same locality label, 25. II – 7. III. 2003, MA- 02 - 22 - 17, 2 ♂♂ (CAS, MOLC); same locality label, 5 – 15. II. 2003, MA- 02 - 22 - 15, 9 ♂♂ (8 in CAS, 1 in MOLC); same locality label, 7 – 17. III. 2003, MA- 02 - 22 - 18, 7 ♂♂ (CAS); same locality label, 15 – 25. II. 2003, MA- 02 - 22 - 16, 7 ♂♂ (CAS); same locality label, 30. XI – 7. XII. 2002, MA- 02 - 22 - 07, 1 ♂ (CAS); same locality label, 16 – 23. XI. 2002, MA- 02 - 22 - 05, 3 ♂♂ (CAS); same locality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 15 ♂♂ (11 in CAS, 4 in MOLC); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 30 ♂♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 35 m, 7 – 17. X. 2004, MT, spiny forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 A- 28, 2 ♂♂ (CAS, MOLC); Toliara, Ambohimahavelona village, 33 km NE of Tulear, 23 ° 26.45 ’ S 43 ° 53.98 ’ E, 46 m, 10 – 21. VIII- 2009, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 50 B- 40, 1 ♂ (MOLC). NAMIBIA: Namib / Naukluft Park, near Gobabeb, Kuiseb River, 1 ♀ (SANC); Toliara, Beroboka village, 45 km NE Moron- dava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 430 ft., 22 – 30. V. 2008, MT, Antsarongaza gallery forest, MG- 45 A- 24, M. Irwin, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 22 – 30. XII. 2007, MG- 45 A- 14, 1 ♂ (CAS); Toliara, Tsimanam- petsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 24 – 29. X. 2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 53 A- 01, 1 ♂ (CAS). NIGERIA: East Central State, Udo FR, 11. IV. 1973, J. T. Medler leg., 1 ♂ (NHMUK). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 6. X. 2010 – 18. I. 2011, MT, Camdeboo Escarpment Ticket, S. van Noort leg., PCD 09 - ACA 1 - M 06, 13 ♂♂ (11 in SAMC, 2 in MOLC); Gauteng, Pretoria District, Soutpan, 1 ♀ (SANC); Limpopo, 15 km E Kla- serie, Guernsey Farm, 19 – 31. XII. 1985, 1 ♀ (CNC). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ‘ S 37 ° 48 ‘ E, 25. XII. 1995 – 29. I. 1996, MT, Acacia / Commiphora / Combretum bushland, SAM-HYM-PO 15909, S. van Noort leg., 1 ♂ (SAMC); same locality label, 11. III – 5. IV. 1996, SAM-HYM-PO 17644, 1 ♂ (SAMC); same locality label, 16. V. 1996, H. G. Robertson leg. from light, SAM-HYM-PO 20422, 1 ♂ (SAMC); Mkomazi Game Reserve, Ibaya Camp, SSE side, 03 ° 57.91 ‘ S 37 ° 48.09 ‘ E, 26. XI. 1995, Sweep mk 5, mixed grasses & shrubs in open Combretum bushland, SAM-HYM-PO 16701, S. van Noort leg., 1 ♂ (SAMC); Mkomazi Game Reserve, Ibaya Camp, NW side, 03 ° 57.91 ‘ S 37 ° 48.09 ‘ E, 28. XI. 1995, Sweep mk 9, Acacia / Commiphora / Combretum bushland, SAM- HYM-PO 16839, S. van Noort leg., 1 ♂ (SAMC); Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ‘ S 38 ° 01.37 ‘ E, 16. IV – 2. V. 1996, MT, Commiphora woodland, SAM-HYM-PO 16286, S. van Noort leg., 2 ♂♂ (SAMC); same lo- cality label, 2. V. 1996, Sweep, mk 43, Cordia, Solanum shrubs & grasses in Commiphora woodland, SAM-HYM- PO 18080, S. van Noort leg., 1 ♂ (SAMC). UGANDA: Ankole, Kichwamba, 23 – 29. IV. 1968; 1 ♂ (AMNH), 1 ♀ (♀ with black head) (USNM). YEMEN: 12 km NW of Manakhah, 15 ° 05 ’ N 43 ° 42 ’ E, 3. VII – 21. VIII. 2001, MT, A. van Harten leg., 2 ♂♂ (MOLC); same locality label, 15. V – 24. VI. 2003, N. 7770, 3 ♂♂ (MOLC); same locality label, 24. VI – 4. VIII. 2003, N. 7915, 2 ♂♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA92B41FF3ED0B9FEB0FE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA92B41FF3ED0B9FEB0FE98.taxon	distribution	Distribution. Botswana, Ghana, Kenya, Namibia, Nigeria, Madagascar, Somalia, South Africa, Tanzania, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA62B42FF3ED03CFB1AF87C.taxon	description	(Figs 101 B, 102 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA62B42FF3ED03CFB1AF87C.taxon	description	Description. ♀. Fully winged; body length 4.0 – 4.4 mm. Head and mesosoma testaceous-reddish; antenna testa- ceous, except antennomeres 6 – 10 darkened; petiole testaceous; metasoma black; legs testaceous, except metafemur and metatibia dark. Antenna clavate; antennomeres in following proportions: 16: 7: 11: 8: 6: 6: 6: 5: 5: 8. Head dull, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 5; OL = 4; OOL = 10; OPL = 10; TL = 10. Pronotum setose, crossed by strong transverse impression, dull, sculptured by short longitudinal keels; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. No- tauli incomplete, shortly visible near anterior margin of mesoscutum. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two longitudinal keels, median area rugose and lateral areas reticulate rugose. Forewing hyaline, with one slight dark transverse band; distal part of 2 r-rs & Rs vein approximately as long as proximal part (11: 12). Protarsomeres in following proportions: 19: 3: 4: 12: 17. Enlarged claw (Fig. 101 B) with one row of eight teeth and one bristle. Protarsomere 5 (Fig. 101 B) without preapical lamella, with one row of eight bristles. Tibial spurs 1 / 1 / 1. ♂. Fully winged; body length 2.0 – 2.7 mm. Head black, except mandible testaceous; antenna testaceous-dark; mesosoma and metasoma black; legs black, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 16: 5.5: 6: 7: 6: 6: 6: 5.5: 5: 8. Head convex, dull, reticulate rugose, except unsculptured or granulate areas around toruli; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 3.5; TL = 5.5. In specimen from Kenya, 0.82103 ° S 38.54443 ° E, POL = 6; OL = 3. Mesoscutum, mesoscutellum and metanotum dull, completely reticulate rugose. Notauli incomplete, reaching approximately 0.2 – 0.4 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two tracks of longitudinal keels and median area shiny, less rugose than lateral areas. In two specimens from South Africa, 27 ° 34 ’ S 31 ° 17.9 ’ E and 30 ° 17.4 ’ S 30 ° 36.9 ’ E, propodeal declivity with two distinct longitudinal keels and dull and reticulate rugose median area. In specimen from Madagascar, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, propodeal declivity with two distinct longitudinal keels and median area almost completely shiny and unsculptured. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter (7: 9 in holotype) or slightly longer than proximal part (12: 11 in specimen from Madagascar, 18 ° 28.24 ’ S 47 ° 57.36 ’ E). Genitalia in Fig. 102 C. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA62B42FF3ED03CFB1AF87C.taxon	materials_examined	Material examined. Types: ♂ holotype: BOTSWANA: Serowe, Farmer’s Brigade, IX. 1987, MT, Per Forch- hammer leg. (USNM). Paratypes: same locality label as holotype, XII. 1987, 1 ♂ (AMNH); same locality label as holotype, Serowe, IX. 1984, MT, Per Forchhammer leg., 1 ♀ (NHMUK). Other material: BOTSWANA: Serowe, Farmer’s Brigade, XII. 1987, MT, Per Forchhammer leg., 1 ♂ (USNM). DEMOCRATIC REPUBLIC OF THE CON- GO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 30. III – 5. IV. 2006, MT, S. L. Heydon & S. E. Stevenson leg., 2 ♂♂ (DEUCD, MOLC); same locality label, 04 ° 15 ’ S 17 ° 10 ’ E, 24. IV. 2006, MT just inside forest, 1 ♂ (DEUCD); same locality label, 04 ° 15 ’ S 17 ° 10 ’ E, 20. IV. 2006, MT forest, 2 ♂♂ (DEUCD, MOLC). KENYA: Coast Prov., Buda Forest, 4.46109 ° S 39.40446 ° E, 98 m, 8 – 22. IV. 2016, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 27. XI – 11. XII. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 8 – 22. I. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♂ (NMK); same locality label, 22. X – 5. XI. 2015, 2 ♂♂ (NMK, MOLC); Eastern Province, Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, 24. XII. 2011 – 7. I. 2012, MT, inside indigenous forest, R. Copeland leg., 1 ♂ (NMK); Eastern Prov., Simisi area, 2.05111 ° S 38.32613 ° E, 710 m, 28. XI – 1. XII. 2013, MT, near base of Yamalu Hill, Acacia shrubland, J. Bukhebi and R. Copeland leg., 8 ♂♂ (7 in NMK, 1 in MOLC); Eastern Prov., base of Ukasi Hill, 0.82103 ° S 38.54443 ° E, 613 m, 5 – 10. XII. 2011, MT, Acacia / Commiphora savanna, R. Copeland leg., 2 ♂♂ (NMK); same locality label, 10 – 17. XII. 2011, 1 ♂ (NMK). MADA- GASCAR: Antananarivo, 11.5 km 17 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 5 – 13. XII. 2000, 1300 m, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375, 1 ♂ (CAS); Fianarant- soa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 13 – 22. XI. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 68, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 750 m, 17 – 24. III. 2002, MT, in open area, MA- 02 - 11 A- 20, R. Harin’Hala leg., 1 ♂ (CAS); Mahajanga, Tsimembo Forest, 8.7 km 336 ° NNW of Soatana, 19 ° 01.17 ’ S 44 ° 26.26 ’ E, 20 m, MT, tropical dry forest, 21 – 25. XI. 2001, Fisher, Griswold et al. leg., BLF 4507, 1 ♂ (MOLC); Toamasina, Mobot site, Analalava, 7 km SW of Foulpointe, 17 ° 42.32 ’ S 49 ° 27.29 ’ E, 80 ft., 12 – 19. X. 2007, MT, low altitude dense humid forest, MG- 37 A- 05, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelled I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 1 – 8. II. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 2 ♂♂ (CAS, MOLC); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI- 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 2 ♂♂ (CAS, MOLC). SOUTH AFRICA: KwaZulu-Natal, Umlalazi Nature Reserve, 1.5 km E of Mtunzini (PMA); KwaZulu-Natal, Kosi Bay, 26 ° 58 ’ S 32 ° 48 ’ E (SANC); KwaZulu-Natal, Louwsberg, Gwala Gwala, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 2 – 3. VI. 2005, YPT, M. Mostovski leg., 1 ♂ (NMSA); Kwa- Zulu-Natal, Vernon Crookes Nat. Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MT, M. Mostovski leg., 1 ♂ (NMSA); Limpopo, Ellisras District, D’Nyala Nature Reserve, 23 ° 45 ’ S 27 ° 49 ’ E (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA62B42FF3ED03CFB1AF87C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA62B42FF3ED03CFB1AF87C.taxon	distribution	Distribution. Botswana, Democratic Republic of the Congo, Kenya, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA42B44FF3ED02AFDADF98C.taxon	description	(Fig. 101 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA42B44FF3ED02AFDADF98C.taxon	description	Description. ♀. Fully winged; body length 4.1 – 4.2 mm. Head and mesosoma completely testaceous-reddish; an- tenna of holotype testaceous, except antennomeres 9 – 10 brown; antenna of specimen from Madagascar, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, testaceous, except antennomeres 5 – 10 brown; antenna of two ♀♀ from Madagascar, 16 ° 00.36 ’ S 45 ° 15.54 ’ E, completely testaceous; metasoma brown-black; legs testaceous, except stalk of metafemur with inner side brown; legs of two ♀♀ from Madagascar, 16 ° 00.36 ’ S 45 ° 15.54 ’ E, totally testaceous. In specimen from Kenya, 0.32442 ° N 38.05038 ° E, frons and mesoscutum partly darkened. In specimen from Kenya, 2.07836 ° S 38.22517 ° E, head testaceous-reddish, except ocellar triangle brown; antenna testaceous, except antennomeres 4 – 10 brown. Antenna distally thickened, short, usually with antennomeres 5 – 9 approximately as long as broad, occasionally (in specimen from Kenya, 2.07836 ° S 38.22517 ° E) with antennomeres 5 – 9 slightly longer than broad; antennomeres in following proportions: 18: 8: 12: 8: 8: 7: 6: 6: 6: 10. Head convex, dull, covered with short setae, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 4; OL = 3.5; OOL = 12; OPL = 12; TL = 11. Pronotum crossed by strong transverse impression, dull, reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli much longer than greatest breadth of lateral ocelli (10: 4). In specimen from Kenya, 0.32442 ° N 38.05038 ° E, notauli not visible among areolae of mesoscutum. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and slightly rugose); propodeal declivity dull, with two complete longitudinal keels and median and lateral areas sculptured by numerous transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein approximately as long as proximal part (11: 11); 2 R 1 cell open. Petiole long, much shorter than half of metasoma. Protarsomeres in following proportions: 65: 12: 10: 44: 60. Enlarged claw (Fig. 101 C) with one row of 5 – 11 teeth and one bristle. Protarsomere 5 (Fig. 101 C) with one preapical lamella, inner band and three-eight bristles (in specimen from Kenya, 2.07836 ° S 38.22517 ° E, two bristles). Tibial spurs 1 / 1 / 1 (for mistake 1 / 1 / 2 in original description). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA42B44FF3ED02AFDADF98C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Beza Mahafaly Reserve, 23 ° 42 ’ S 41 ° 42 ’ E, 21. XI. 1984, MT, Robert W. Brooks leg. (NAMUK). Other material: KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 24. IV – 10. V. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 4 ♀♀ (3 in NMK, 1 in MOLC); same locality label, 10 – 24. IV. 2015, 1 ♀ (NMK); same locality label, 3 – 17. V. 2016, 2 ♀♀ (NMK, MOLC); Eastern Prov., Kasaala area, Mulu Musingila’s Farm, 13 – 27. XII. 2016, MT, near gully, R. Copeland leg., 1 ♀ (NMK); Eastern Province, Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, 23. VII – 6. VIII. 2011, MT, inside indigenous forest, R. Copeland leg., 1 ♀ (NMK); same locality label, 3 – 17. IX. 2011, 1 ♀ (NMK); same locality label, 8 – 22. VI. 2011, 1 ♀ (MOLC); same locality label, 9 – 23. VII. 2011, 1 ♀ (NMK); Rift Valley Prov., Matthews Range, 1.00661 ° N 37.38631 ° E, 1006 m, 26. XII. 2015 – 9. I. 2016, MT, Sarara campsite, shrubland, R. Copeland leg., 1 ♀ (NMK). MADAGASCAR: Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, 15 – 22. VI. 2011, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 27, 1 ♀ (CAS); Fianarantsoa, Italaviana, 35 km SSE of Antsirabe, 20 ° 10.40 ’ S 47 ° 05.16 ’ E, 1360 m, 6 – 20. V. 2004, MT in Uapacca Forest, M. Irwin & R. Harin’Hala leg., MA- 24 - 47, 1 ♀ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 1 ♀ (CAS); Mahajanga, Baie de Baly National Park, 12.4 km 337 ° NNW Soalala, 16 ° 00.36 ’ S 45 ° 15.54 ’ E, 10 m, 26 – 30. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6811, 2 ♀♀ (MOLC); Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 26. II – 2. III. 2009, MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG- 53 C- 19, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA42B44FF3ED02AFDADF98C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA42B44FF3ED02AFDADF98C.taxon	distribution	Distribution. Kenya, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA32B45FF3ED013FD63FCD8.taxon	description	(Fig. 102 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA32B45FF3ED013FD63FCD8.taxon	description	Description. ♂. Fully winged; body length 1.9 – 2.4 mm. Head black, except mandible testaceous; antenna testa- ceous-brown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 10: 5: 7: 6: 6: 6: 6: 6: 5.5: 8. Head convex, dull, covered with short setae, completely reticulate rugose and granulate; frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 5; TL = 5. Mesoscutum dull, completely granulate and sculptured by numerous irregular keels. Notauli not visible among numerous keels; in few paratypes mesoscutum with slight tracks of notauli. Me- soscutellum shiny, reticulate rugose, with anterior half unsculptured (in specimen from Namibia, 21 ° 55 ’ S 18 ° 11 ’ E, mesoscutellum almost completely smooth, unsculptured). Metanotum dull, reticulate rugose (in specimen from Namibia, 21 ° 55 ’ S 18 ° 11 ’ E, metanotum shiny, unsculptured). Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with or without two complete longitudinal keels (in holotype propodeal declivity with two incomplete and short longitudinal keels); propodeal declivity with median and lateral areas completely reticulate rugose (in holotype median area shiny, punctate and unsculptured among punctures). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (6: 9); 2 R 1 cell open. Genitalia in Fig. 102 D. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA32B45FF3ED013FD63FCD8.taxon	materials_examined	Material examined. Types: ♂ holotype: YEMEN: 12 km NW of Manakhah, 15 ° 05 ’ N 43 ° 42 ’ E, 3. VII – 21. VIII. 2001, MT, A. van Harten leg. (MOLC). Paratypes: same locality label as holotype, 8 ♂♂ (MOLC); same locality label as holotype, 6. VII – 21. VIII. 2002, A. van Harten leg., 3 ♂♂ (MOLC). Other material: KENYA: Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 27 – 30. V. 2013, 6 m MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK). NAMIBIA: Gobabis, Farm Vendetta 202, Swart-Nossob River, 21 ° 55 ’ S 18 ° 11 ’ E, 1620 m, 26 – 28. VI. 1996, MT, dry vegetated wash, M. E. Irwin, E. I. Schlinger & E. Marias leg., 1 ♂ (DEUCD). YEMEN: 12 km NW of Manakhah, 15 ° 05 ’ N 43 ° 42 ’ E, 15. V – 24. VI. 2003, N. 7770, 1 ♂ (MOLC); same locality label, 24. VI – 4. VIII. 2003, N. 7915, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA32B45FF3ED013FD63FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA32B45FF3ED013FD63FCD8.taxon	distribution	Distribution. Kenya, Namibia, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	description	(Figs 102 E, 103, 104 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	diagnosis	Diagnosis. ♂ of Bocchus with OPL much longer than POL; antennomere 8 less than 3 × as long as broad (Fig. 104 A); mesoscutum granulate, slightly rugose near anterior margin; Notauli incomplete, reaching approximately 0.6 × length of mesoscutum (with obsolete tracks of notauli in posterior third of mesoscutum); propodeal declivity reticulate rugose, without longitudinal keels.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	description	Description. ♂ (Figs 103, 104 A). Fully winged; body length 2.4 mm. Head black, except mandible brown; antenna brown; mesosoma black; metasoma brown; legs brown, except tarsi brown-testaceous. Antenna filiform; antennomeres in following proportions: 9: 6: 7: 6: 6: 6: 5: 5: 5: 8; antennomere 6 twice as long as broad; antennomere 8 less than 3 × as long as broad (5: 3). Head convex, dull, granulate and rugose; frontal line complete; occipital carina complete; POL = 3; OL = 2.5; OOL = 6; OPL = 5; TL = 4; greatest breadth of lateral ocelli shorter than OPL (2: 5). Mesoscutum dull, granulate, slightly rugose near anterior margin. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum (with obsolete tracks of notauli in posterior third of mesoscutum). Mesoscutellum dull, granulate. Metanotum granulate and rugose. Mesopleuron dull, granulate. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, without longitudinal keels. Forewing (Fig. 103 D) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (10: 14); 2 R 1 cell open. Genitalia in Fig. 102 E. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	materials_examined	Material examined. Type: ♂ holotype (CASTYPE 19448): MADAGASCAR: Fianarantsoa, Miandritsara For- est, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 8 – 17. III. 2007, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 81 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA22B45FF3ED5FFFB62F8C8.taxon	etymology	Etymology. The new species is named forestalis, because it was collected in a forest.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA12B58FF3ED35BFE1FFE28.taxon	description	(Fig. 102 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA12B58FF3ED35BFE1FFE28.taxon	description	Description. ♂. Fully winged; body length 3.1 – 3.4 mm. Head black, except mandible testaceous; mesosoma black; metasoma brown; antenna brown; legs brown, exceept articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 15: 8: 10: 10: 10: 10: 9.5: 10: 10: 13; antennomere 8 at least 3 × as long as broad (10: 3 in the holotype; 9: 3 in other specimen). Head convex, dull, covered with short setae, completely reticulate rugose and granulate; occasionally head granulate and with numerous slight irregular keels; frontal line complete (reduced to irregular track in front of anterior ocellus); occipital carina complete; POL = 7; OL = 4; OOL = 10; OPL = 2.5; TL = 5. In series of specimens from South Africa, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, POL slightly shorter than OOL (7: 8). Mesoscutum dull, completely granulate and sculptured by numerous irregular keels; occasionally mesoscutum granulate and without irregular keels; in some specimens from South Africa, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, mesoscutum granulate, strongly rugose near posterior margin. Notauli incomplete, reaching approximately 0.4 – 0.6 × length of mesoscutum (0.4 × in holotype). In some specimens from South Africa, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, notauli almost complete (distal part hardly visible among areolae present near posterior margin of mesoscutum). Mesoscutellum and metanotum dull, granulate, with few short irregular keels. Mesopleuron dull, granulate and partly reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, reticulate rugose, with tracks of two complete longitudinal keels (tracks visible only under special conditions of light) and median area dull and as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (14: 13); 2 R 1 cell open. Genitalia in Fig. 102 F. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA12B58FF3ED35BFE1FFE28.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Cape Town, Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 6 – 13. XII. 1994, MT, Mesic mountain Fynbos, Protea coronata dominated, SAM-HYM-PO 13715, S. van Noort leg. (SAMC). Other material: SOUTH AFRICA: Western Cape, Cape Town, Constantiaberg, Tokai Forest, pine plant, X – XI. 1995, MT, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 11. VIII – 6. IX. 1997, S. van Noort leg., 2 ♂♂ (SAMC, MOLC); Western Cape, Mossel Bay, 18 – 30. XI. 1921, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. XI – 16. XII. 1998, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 8 ♂♂ (7 in SAMC, 1 in MOLC); same locality label, 16. XI – 16. XII. 1999, 4 ♂♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA12B58FF3ED35BFE1FFE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEA12B58FF3ED35BFE1FFE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	description	(Figs 104 B – D, 108 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	diagnosis	Diagnosis. ♂ of Bocchus with mesoscutum completely granulate (Fig. 104 D); notauli complete and posteriorly separated; mesoscutellum finely punctate, unsculptured among punctures; distal part of 2 r-rs & Rs vein about as long as proximal part; propodeal declivity without longitudinal keels.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	description	Description. ♂ (Figs 104 B – D). Fully winged; body length 2.4 – 2.6 mm. Head black, except mandible testa- ceous; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomere 6 more than twice as long as broad (8: 2); antennomeres in following proportions: 11: 6: 8: 8: 7: 8: 8: 8: 7: 11. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 5; OL = 3.5; OOL = 6; OPL = 3; TL = 5; greatest breadth of posterior ocellus about as long as OPL. Mesoscutum dull, completely granulate. Notauli complete and posteriorly separated; minimum distance between notauli longer than greatest breadth of posterior ocellus (8: 3). Notauli of paratype hardly visible. Mesoscutellum shiny, finely punctate, unsculptured among punctures. Metanotum unsculptured, shiny. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal and transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein about as long as proximal part. Petiole short. Genitalia in Fig. 108 A. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.845 ’ S 19 ° 09.443 ’ E, 787 m, 27. X. 2009 – 9. II. 2010, MT, Nieuwoudtville Dolerite, Koppie Renosterveld, S. van Noort leg., SAM-HYM-A 022248 (SAMC). Paratype: SOUTH AFRICA: Western Cape, West Coast Fossil Park, 32 ° 58.020 ’ S 18 ° 07.285 ’ E, 3 km 280 ° E Langebaanweg, 23 – 30. X. 2002, MT, rehabilitated mine dump, S. van Noort leg., SAM-HYM-A 023974, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B58FF3ED44FFAE3FAFB.taxon	etymology	Etymology. The specific name granulatus refers to the sculpture of the mesoscutum (granulate).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	description	(Figs 104 E, 105, 108 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	diagnosis	Diagnosis. ♂ of Bocchus with antennomere 6 less than twice as long as broad; mesoscutum reticulate rugose (Fig. 104 E); notauli incomplete, reaching approximately 0.6 × length of mesoscutum; propodeal declivity with median area as reticulate rugose as lateral areas.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	description	Description. ♂ (Figs 104 E, 105). Fully winged; body length 2.6 mm. Head black, except mandible testaceous; antenna brown-testaceous; mesosoma black; metasoma brown-black; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 11: 5: 7: 7: 6: 6: 6: 5: 5: 8; antennomere 6 less than twice as long as broad (6: 4). Head (Fig. 1059 D) convex, dull, covered with short setae, completely strongly reticulate rugose; frontal line complete, weakly convex between antennal toruli; occipital carina complete; POL = 5; OL = 2; OOL = 7.5; OPL = 5; TL = 5; greatest breadth of lateral ocelli longer than OL (3: 2). Mesoscutum (Fig. 1059 C) dull, completely strongly reticulate rugose. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum dull, reticulate rugose. Metanotum rugose. Mesopleuron dull, strongly reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with two complete longitudinal keels, median area as rugose as lateral areas. Forewing (Fig. 105 E) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (8: 9); 2 R 1 cell open. Apex of basivolsella slender and with subdistal inner pointed apophysis (Fig. 108 B). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	materials_examined	Material examined. Type: ♂ holotype (CASTYPE 19449): MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 10 – 20. VII. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 32 (CAS). Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBF2B5AFF3ED383FB63FF50.taxon	etymology	Etymology. The new species is named after one of the collectors, Mr. R. Harin’Hala.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBD2B5BFF3ED752FE14FF50.taxon	description	(Figs 104 F, 108 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBD2B5BFF3ED752FE14FF50.taxon	description	Description. ♂ (Fig. 104 F). Fully winged; body length 1.6 – 1.7 mm. Head black, except mandible testaceous and clypeus brown; mesosoma black; metasoma brown; antenna testaceous-brown; legs testaceous, except club of metafemur and metatibia slightly darkened; in ♂ specimens from Madagascar, 18 ° 28.24 ’ S 47 ° 57.36 ’ E and 23 ° 02.667 ’ S 43 ° 36.917 ’ E, legs testaceous, except part of metacoxa brown; in three specimens from Madagascar, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, legs testaceous, except metacoxa, metafemur and metatibia brown. Antenna filiform; antennomeres in following proportions: 8: 4: 3.5: 3.5: 4: 4: 4: 4: 4: 5.5. Head convex, dull, covered with short setae, completely granulate; frontal line complete; occipital carina complete; POL = 5.5; OL = 3; OOL = 3.5; OPL = 2; TL = 3. Mesoscutum dull, completely granulate. Notauli complete, posteriorly separated; minimum distance between notauli much longer than greatest breadth of lateral ocelli (7: 2). Mesoscutellum shiny, finely punctate, unsculptured among punctures; in specimen from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, mesoscutellum slightly granulate. Metanotum shiny, unsculptured. Mesopleuron dull, granulate. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with two complete longitudinal keels and median and lateral areas completely reticulate rugose (in three specimens from Madagascar, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, median area partly smooth). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (4: 8); 2 R 1 cell open; veins surrounding R an 1 Cu cells diaphane (apparently only costal cell enclosed by pigmented veins). Volsella slightly shorter than paramere (Fig. 108 C). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBD2B5BFF3ED752FE14FF50.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW Anivorano Nord, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3011 (CAS). Paratypes: same locality label as holotype, 2 ♂♂ (CAS, MOLC). Other material: MADAGASCAR: Antananarivo, 11.5 km 147 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 15 – 22. XI. 2001, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 03, 1 ♂ (CAS); same locality label, 4 – 14. VII. 2002, MA- 02 - 09 C- 34, 1 ♂ (CAS); same locality label, 26 – 31. III. 2002, MA- 02 - 09 C- 22, 1 ♂ (MOLC); same locality label, 16. X – 8. XI. 2001, MA- 02 - 09 C- 01, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 1 ♂ (CAS); Fianarantsoa, Radio tower 22 km SW of Hakaka, near Fianarantsoa / Tulear border, 22 ° 46.75 ’ S 45 ° 01.50 ’ E, 1100 m, 16 – 27. II. 2002, MT, Uapacca forest, M. Irwin, R. Harin’Hala leg., MA- 02 - 12 - 09, 1 ♂ (MOLC); Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 2 ♂♂ (CAS); Mahajanga, Ambovomamy Belembo, 20 km NW Port Berger, 15 ° 27.07 ’ S 47 ° 36.80 ’ E, 33 m, 19 – 25. IV. 2007, secondary growth on white sand, MT, R. Harin’Hala, M. Irwin, F. Parker leg., MG- 33 - 18, 1 ♂ (CAS); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 12 – 19. II. 2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 18, 1 ♂ (CAS); Mahajan- ga, Sofia Disctrict, 45 km S Antsohihy, 5 km W of Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 13 – 19. XII. 2010, MT, M. Irwin, R. Harin’Hala leg., MG- 61 - 01, 2 ♂♂ (CAS, MOLC). Toli- ara, Mite Forest, 20.7 km 29 ° WNW Tongobory, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, 75 m, 27. II – 3. III. 2002, MT, gallery forest, Fisher, Griswold et al. leg., BLF 5849, 1 ♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 30. X – 9. XI. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 44, 1 ♂ (MOLC); same locality label, 26. I – 5. II. 2003, 1 ♂ (MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 8 – 18. III. 2002, MT, spiny forest, M. Irwin & R. Harin’Hala leg., MA- 02 - 18 B- 16, 1 ♂ (MOLC); Toliara, Ranobe, 23 ° 02.667 ’ S 43 ° 36.917 ’ E, 20 m, 26 – 29. I. 2004, MT, spiny forest thicket, MGF 093, Frontier Wilderness Project, 3 ♂♂ (2 in CAS, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBD2B5BFF3ED752FE14FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBD2B5BFF3ED752FE14FF50.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	description	(Figs 101 D; 106 A, 108 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	description	Description. ♀ (Fig. 106 A). Fully winged; body length 2.7 – 3.8 mm. Yellow-whitish, except ocelli slightly dark- ened. Occasionally mesothoracic axilla and sutures behind mesoscutum, mesoscutellum and metanotum darkened. In specimen from the United Arab Emirates, 25 ° 24 ’ N 56 ° 17 ’ E, mesosoma ferruginous and metasoma brown; in specimen from Kenya, 0.82103 ° S 38.54443 ° E, mesosoma ferruginous and metasoma partly brown and partly ferruginous. Antenna clavate, short, with antennomeres 5 – 9 longer than broad (antennomere 5: 6: 3; antennomere 9: 4.5: 3.5); antennomeres in following proportions: 11: 6: 9: 7: 6: 5.5: 5: 5: 4.5: 7. Head dull, completely granulate and reticulate rugose; frontal line complete; occipital carina complete; POL = 4; OL = 2; OOL = 7; OPL = 7; TL = 7. Pronotum crossed by strong transverse impression, reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum dull, completely granulate and reticulate rugose. Notauli complete, posteriorly separated. Occasionally notauli hardly visible. In specimen from the United Arab Emirates, 25 ° 24 ’ N 56 ° 17 ’ E, mesoscutum strongly reticulate rugose, notauli almost completely invisible. In specimen from Kenya, 0.82103 ° S 38.54443 ° E, mesoscutum strongly reticulate rugose, notauli completely invisible Mesoscutellum slightly granulate. In specimen from the United Arab Emirates, 25 ° 24 ’ N 56 ° 17 ’ E, mesoscutellum with anterior half unsculptured and posterior half reticulate rugose. In specimen from Kenya, 0.82103 ° S 38.54443 ° E, mesoscutellum granulate. Metanotum short and rugose. Metapectalpropodeal disc reticulate rugose; propodeal declivity dull and rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (11: 13). Petiole very long, much shorter than metasoma (6: 27). Protarsomeres in following proportions: 15: 3: 3.5: 14: 17. Enlarged claw (Fig. 101 D) with one row of four-nine teeth (nine in holotype) and one bristle. Protarsomere 5 (Fig. 101 D) with one preapical lamella, two bristles and an inner band. Tibial spurs 1 / 1 / 1. ♂. Fully winged; body length 1.5 – 2.2 mm. Head black, except mandible testaceous; mesosoma black; meta- soma brown; antenna testaceous; legs testaceous, except coxae and clubs of femora partly darkened. Antennae filiform; antennomeres in following proportions: 8: 5: 5.5: 5: 5: 5: 4.5: 4.5: 4: 7; antennomere 8 less than 3 × as long as broad (4.5: 2). Head convex, dull, covered with short setae, granulate and reticulate rugose; frontal line complete; occipital carina complete; POL = 5; OL = 2; OOL = 4.5; OPL = 3; TL = 2; greatest breadth of lateral ocelli about as long as OPL. Mesoscutum dull, completely granulate and reticulate rugose. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Mesopleuron dull, granulate. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with two complete longitudinal keels, lateral areas completely reticulate rugose and median area partly shiny and unsculptured. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curved with no distinct separation between distal and proximal part; 2 R 1 cell open. Apex of basivolsella without subdistal inner pointed apophysis (Fig. 108 D). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	materials_examined	Material examined. Type: ♀ holotype: OMAN: 4 km S of Dhagmar, 23 ° 09 ’ N 58 ° 59 ’ E, 80 m, 14. XI. 1991, dry stony plateau, 8422, M. D. Gallagher leg. (CNC). Other material: KENYA: Eastern Province, base of Ukasi Hill, 0.82103 ° S 38.54443 ° E, 613 m, 21. XI – 5. XII. 2011, MT, Acacia / Commiphora savanna, R. Copeland leg., 1 ♀ (NMK). OMAN: Al-Uyaina – Samail, 30. IV. 2006, ex Ommatissus lybicus De Bergevin on date palm, K. Al- Miqbali leg., 1 ♀, 5 ♂♂ (NHMUK); SE of Súr, 16. III. 2015, M. Snizek leg., 2 ♀♀ (OLL). UNITED ARAB EMIR- ATES: Fujairah, 25 ° 08 ’ N 56 ° 21 ’ E, 13 – 19. IV. 2005, LT, 1316, A. van Harten leg., 2 ♀♀ (MOLC); Wadi Maidaq, 25 ° 18 ’ N 56 ° 07 ’ E, 22 – 24. XII. 2005, yellow and white water-traps, A. van Harten leg., 1 ♀ (MOLC); same locality label, 27. IV – 4. V. 2006, LT, A. van Harten leg., 4 ♀♀ (MOLC); same locality label, 2 – 16. II. 2006, LT, A. van Harten leg., 1 ♀ (MOLC); Wadi Safad, 25 ° 13 ’ N 56 ° 19 ’ E, 26 – 29. XII. 2005, yellow and white water-traps, A. van Harten leg., 1 ♀ (MOLC); same locality label, 17 – 24. VI. 2006, LT, A. van Harten leg., 1 ♀ (MOLC); same locality label, 14 – 21. V. 2006, LT, A. van Harten leg., 3 ♀♀ (MOLC); same locality label, 25. III – 1. IV. 2006, LT, 11626, A. van Harten leg., 2 ♀♀ (MOLC); Hatta, 24 ° 49 ’ N 56 ° 07 ’ E, 8 – 26. IV. 2006, LT, A. van Harten leg., 1 ♀ (MOLC); same locality label, 19 – 28. III. 2006, LT, A. van Harten leg., 1 ♀ (MOLC); Sharhah x Khor Kalba, 24 ° 59 ’ N 56 ° 09 ’ E, 16 – 31. I. 2006, LT, A. van Harten leg., 2 ♀♀ (MOLC); same locality label, 7 – 22. III. 2006, LT, A. van Harten leg., 9 ♀♀ (MOLC); same locality label, 24 – 30. V. 2006, LT, A. van Harten leg., 5 ♀♀ (MOLC); Wadi Wurayah, 25 ° 24 ’ N 56 ° 17 ’ E, 26. XI. 2006, in water, 5695, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	biology_ecology	Hosts. Tropiduchidae (Guglielmino et al. 2013): in Oman: Ommatissus lybicus De Bergevin.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	distribution	Distribution. Kenya, Oman, United Arab Emirates.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBC2B5CFF3ED767FE40F8C4.taxon	discussion	Remarks. ♀♀ of B. hyalinus show two varieties based on colour and sculpture as follows: variety A (very light and slightly sculptured): as represented by the holotype, in a specimen from Oman, Al-Uyaina – Samail, and in almost all known specimens from the UAE (except for a specimen from 25 ° 24 ’ N 56 ° 17 ’ E); variety B (dark and strongly sculptured): known in a specimen from the UAE, 25 ° 24 ’ N 56 ° 17 ’ E, and in a specimen from Kenya, 0.82103 ° S 38.54443 ° E.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBA2B5FFF3ED14CFE32FF50.taxon	description	(Fig. 108 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBA2B5FFF3ED14CFE32FF50.taxon	description	Description. ♂. Fully winged; body length 1.7 mm. Head black, except mandible testaceous; antenna testaceousbrown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 10: 4: 4.5: 4.5: 4.5: 4.5: 4.5: 4.5: 5: 5; antennomere 8 less than 3 × as long as broad (4.5: 2). Head convex, dull, covered with short setae, granulate and sculptured by slight short irregular keels and areolae; frontal line complete; occipital carina complete; POL = 3; OL = 2; OOL = 6; OPL = 2; TL = 3.5. Mesoscutum dull, completely granulate and sculptured by slight irregular keels and areolae. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, slightly rugose. Meso- pleuron dull, granulate. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, without longitudinal keels (under special conditions of light two tracks of complete longitudinal keels visible) and median and lateral areas completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (7: 8); 2 R 1 cell open. Genitalia in Fig. 108 E. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBA2B5FFF3ED14CFE32FF50.taxon	materials_examined	Material examined. Type: ♂ holotype: TANZANIA: Mkomazi Game Reserve, Kisima Plot, 04 ° 06.6 ’ S 38 ° 05.58 ’ E, 25. XI – 8. XII. 1995, MT, Acacia / Commiphora bushland, SAM-HYM-PO 16495, S. van Noort leg. (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBA2B5FFF3ED14CFE32FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEBA2B5FFF3ED14CFE32FF50.taxon	distribution	Distribution. Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B5FFF3ED767FE14FB1A.taxon	description	(Fig. 108 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B5FFF3ED767FE14FB1A.taxon	description	Description. ♂. Fully winged; body length 2.1 – 2.9 mm. Head black, except mandible testaceous; antenna browntestaceous, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs brown-testaceous. Antenna filiform; antennomeres in following proportions: 9: 6: 7: 6: 5.5: 5: 5: 4.5: 4.5: 7. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 6; OPL = 6; TL = 4; greatest breadth of posterior ocellus as long as POL. Mesoscutum, mesoscutellum and metanotum reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (7: 3). Metapectalpropodeal disc reticulate rugose; propodeal declivity with two complete longitudinal keels, lateral areas rugose and median area almost completely shiny and unsculptured or with small central unsculptured area (less frequently median area completely rugose, without unsculptured area). Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (10: 9); 2 r-rs & Rs vein not S-shaped. Petiole short. Apex of paramere pointed (Fig. 108 F); apex of basivolsella broadened (Fig. 108 F). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B5FFF3ED767FE14FB1A.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W of Anjiamangirana, elev 97 m, 27. I – 2. II. 2011, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, M. Irwin, R. Harin’Hala, MT, Analagnambe dry degraded forest, MG- 61 - 08 (CAS). Paratypes: same locality label as holotype, 3 ♂♂ (2 in CAS, 1 in MOLC). Other material: MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 5 ♂♂ (2 in MOLC, 3 in CAS); same locality label, 16 – 27. XII. 2002, 12 ♂♂ (7 in CAS, 5 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B5FFF3ED767FE14FB1A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B5FFF3ED767FE14FB1A.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B50FF3ED2BEFE59FF50.taxon	description	(Figs 106 B, C, 109 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B50FF3ED2BEFE59FF50.taxon	description	Description. ♂ (Figs 106 B, C). Fully winged; body length 2.5 mm. Head black, except mandible testaceous; an- tenna brown, except scape, pedicel and ventral side of flagellomeres 1 – 3 testaceous; mesosoma amd metasoma black; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 11: 7: 9: 9: 8: 8: 8: 8: 7: 9; antennomere 6 more than twice as long as broad (8: 3.5); antennomere 8 less than 3 × as long as broad (8: 3). Head convex, dull, covered with short setae, completely strongly reticulate rugose; frontal line complete, slightly swollen between antennal toruli; occipital carina complete; POL = 7; OL = 3; OOL = 9; OPL = 4; TL = 6; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, completely strongly reticulate rugose. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum dull, with anterior half granulate and posterior half rugose. Metanotum rugose. Mesopleuron dull, strongly reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity dull, with two complete longitudinal keels, median area crossed by numerous transverse keels and lateral areas rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (8: 15); 2 R 1 cell open. Apex of basivolsella without subdistal inner pointed apophysis (Fig. 109 A). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B50FF3ED2BEFE59FF50.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Coast Prov., Kasigau Mtn., 3.82080 ° S 38.64178 ° E, 737 m, bottom of forest, woodland with grass, MT, 7 – 21. IX. 2011, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B50FF3ED2BEFE59FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB82B50FF3ED2BEFE59FF50.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB72B51FF3ED0E2FE27FDE4.taxon	description	(Figs 106 D, 107 A – C, 109 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB72B51FF3ED0E2FE27FDE4.taxon	description	Description. ♂ (Figs 106 D, 107 A – C). Fully winged; body length 2.9 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel ferruginous; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 14: 6: 8: 8: 7: 7: 7: 7: 7: 10; antennomere 6 less than twice as long as broad (7: 5). Head convex, dull, covered with short setae, completely reticulate rugose; clypeus reticulate rugose; frontal line complete, with pointed protrusion between antennal toruli; occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 7; TL = 7; greatest breadth of lateral ocelli shorter than POL (4: 5). Mesoscutum, mesoscutellum and metanotum dull, completely reticulate rugose. Notauli absent. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectalpropodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity with two complete longitudinal keels, median area crossed by many transverse keels and lateral areas completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein about as long as proximal part. Genitalia in Fig. 109 B. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB72B51FF3ED0E2FE27FDE4.taxon	materials_examined	Material examined. Type: ♂ holotype: UGANDA: Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.836 ’ N 30 ° 21.700 ’ E, 1523 m, 12 – 26. VIII. 2008, MT, primary mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 024122 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB72B51FF3ED0E2FE27FDE4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB72B51FF3ED0E2FE27FDE4.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B51FF3ED4FBFE31F8A2.taxon	description	(Fig. 101 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B51FF3ED4FBFE31F8A2.taxon	description	Description. ♀. Fully winged; body length 3.7 – 4.4 mm. Head ferruginous, except brown spot on ocellar region; antenna testaceous; pronotum ferruginous; mesoscutum black, except regions along notauli and posterior third of median region ferruginous; mesoscutellum black, with ferruginous nuances; mesopleuron and metapleural-suture ferruginous; rest of mesosoma black; petiole testaceous; metasoma brown, except posterior third testaceous; legs testaceous. Antenna clavate, short, with antennomeres 5 – 9 longer than broad; antennomeres in following proportions: 16: 8: 12: 7.5: 6.5: 6: 5.5: 5: 5: 8. Head convex, dull, covered with short setae, completely reticulate rugose and granulate; frontal line complete; occipital carina complete; POL = 6; OL = 2; OOL = 8; OPL = 9; TL = 9. Pronotum crossed by strong transverse impression, dull, setose, with sides sculptured by many longitudinal keels, with anterior collar granulate and with many short longitudinal keels on transverse furrow; pronotal tubercle reaching tegula. Mesoscutum completely granulate and reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (10: 4). Mesoscutellum partly granulate and partly sculptured by numerous longitudinal keels. Metanotum reticulate rugose. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and rugose); propodeal declivity dull, with two complete longitudinal keels and median and lateral areas reticulate rugose. Forewing with one dark spot situated around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (15: 17); 2 R 1 cell open. Petiole long, much shorter than metasoma (14: 72). Protarsomeres in following proportions: 20: 3: 4: 16: 17. Enlarged claw (Fig. 101 E) with one row of seven teeth and one bristle. Protarsomere 5 (Fig. 101 E) with one preapical lamella, two bristles and an inner band. Tibial spurs 1 / 1 / 1 (for mistake 1 / 1 / 2 in original description). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B51FF3ED4FBFE31F8A2.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: Brandberg, Hungarob River, 21 ° 13.5 ’ S 14 ° 31.1 ’ E, 1200 m, 3. XI. 1998, LT, A. H. Kirk-Spriggs leg. (SMWN). Paratypes: NAMIBIA: Brandberg, Inside Hungorob ravine, 21 ° 11.30 ’ S 14 ° 31.40 ’ E, 1200 m, 25. X. 1998, at light 1900 – 0700, R. Butlin & J. Altringham leg., 2 ♀♀ (MOLC, SMWN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B51FF3ED4FBFE31F8A2.taxon	biology_ecology	Hosts. Unknown	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B51FF3ED4FBFE31F8A2.taxon	distribution	Distribution. Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B52FF3ED136FE1FFC4D.taxon	description	(Figs 82 F, 86 D – F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B52FF3ED136FE1FFC4D.taxon	description	Description. ♀ (Figs 107 D – F). Micropterous; forewing about twice as long as tegula (13: 6). Body length 3.1 – 3.5 mm. Head black, except mandible testaceous and clypeus ferruginous; antenna testaceous, except antennomeres 5 – 10 brown; mesosoma and metasoma black; legs testaceous, except coxae and clubs of femora darkened. Antenna clavate; antennomeres in following proportions: 14: 7: 9: 7: 7: 6: 6: 5.5: 6: 8.5. Head (Figs 107 D, F) dull, setose, granulate and with many irregular longitudinal keels; frontal line complete; occipital carina complete; dorsal side of head without two lateral keels around orbits; POL = 5; OL = 3.5; OOL = 10; OPL = 8.5; TL = 10; greatest breadth of lateral ocelli shorter than OL (2: 3.5). Pronotum setose, shiny, crossed by transverse impression between anterior and posterior collar; anterior collar partly slightly granulate and partly unsculptured; posterior collar reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum (Fig. 107 D) setose, dull, reticulate rugose and sculptured by strong irregular longitudinal keels. Notauli absent. Mesoscutellum dull, reticulate rugose. Metanotum very short, reduced. Metapectal-propodeal disc and propodeal declivity setose, completely reticulate rugose; propodeal declivity without longitudinal keels. Mesopleuron and metapleuron dull, setose, reticulate rugose and sculptured by transverse keels. Protarsomeres in following proportions: 16: 3: 3.5: 12: 17. Enlarged claw (Fig. 101 F) with one row of six teeth and one long bristle. Protarsomere 5 (Fig. 101 F) with one preapical lamella, an inner band and one row of three bristles. Tibial spurs 1 / 1 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B52FF3ED136FE1FFC4D.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Kogelberg Nature Reserve, 34 ° 15 ’ S 19 ° 05 ’ E, 16. XII. 1999 – 16. I. 2000, pitfall trap, S. van Noort leg., SAM-HYM-P 024851 (SAMC). Paratype: same locality label as holotype, SAM-HYM-P 024851, 1 ♀ (SAMC). Other material: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 57 m, 26. XII. 1997, Sweep, WB 97 - S 40, South Coast Strandveld, S. van Noort leg., SAM-HYM-A 022123, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B52FF3ED136FE1FFC4D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB62B52FF3ED136FE1FFC4D.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B52FF3ED252FE3FF8A1.taxon	description	(Fig. 110 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B52FF3ED252FE3FF8A1.taxon	description	Description. ♀. Fully winged; body length 2.7 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 6 – 10 brown; mesosoma black; legs brown, except tarsi testaceous. Antenna clavate, except antennomeres 5 – 9 approximately as long as broad; antennomeres in following proportions: 10: 6: 5: 4: 4: 4.5: 4: 4: 3.5: 6. Head dull, granulate; frons sculptured by slight areolae; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 4; OPL = 7; TL = 7. Mesoscutum dull, granulate, with slight areolae near anterior margin. Notauli very hardly visible, apparently reaching posterior margin of mesoscutum. Mesoscutellum slightly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, with areolae very broad; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with one dark transverse band situated on distal half; distal part of 2 r-rs & Rs vein shorter than proximal part (4: 12). Petiole very long, approximately 0.25 as long as metasoma (13: 45). Protarsomeres in following proportions: 12: 2: 4: 10: 12. Enlarged claw (Fig. 110 A) with one preapical tooth and one bristle; Protarsomere 5 (Fig. 110 A) with one preapical lamella and two bristles. Tibial spurs 1 / 1 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B52FF3ED252FE3FF8A1.taxon	materials_examined	Material examined. Type: ♀ holotype: BOTSWANA: Serowe, Farmer’s Brigade, III. 1986, MT, Per Forch- hammer leg. (USNM).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B52FF3ED252FE3FF8A1.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B52FF3ED252FE3FF8A1.taxon	distribution	Distribution. Botswana.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B53FF3ED139FDADFC20.taxon	description	(Fig. 109 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B53FF3ED139FDADFC20.taxon	description	Description. ♂. Fully winged; body length 2.4 – 2.9 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomere 6 more than twice as long as broad (7: 3); antennomeres in following proportions: 10: 6: 7: 7: 7: 7: 6.5: 6: 5.5: 8. Head dull, reticulate rugose; frontal line complete; occipital carina complete; holotype with POL = 6; OL = 3; OOL = 7; OPL = 5; TL = 5; in specimen from Kenya, 0.47203 ° S 36.06388 ° E, POL = 6; OOL = 9. Mesoscutum dull, with notauli complete and posteriorly separated; area between notauli reticulate rugose; lateral areas of mesoscutum granulate. Minimum distance between notauli longer than greatest breadth of lateral ocelli (6: 2.5). In specimen from Madagascar, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, mesoscutum completely granulate and reticulate rugose. Mesoscutellum dull, granulate. Metanotum rugose. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal and transverse keels. In specimen from Kenya, 0.47203 ° S 36.06388 ° E, propodeal declivity with median region shiny, granulate, not rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (7: 11). Petiole short. Genitalia in Fig. 109 C. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B53FF3ED139FDADFC20.taxon	materials_examined	Material examined. Type: ♂ holotype: MADAGASCAR: Toliara, 60 km NE of Morondava, Bereboka, 18 – 23. V. 1983, B. M. 1983 – 201, J. S. Noyes & M. C. Day leg. (NHMUK). Other material: KENYA: Rift Valley Prov., Lake Nakuru National Park, 0.47203 ° S 36.06388 ° E, 1908 m, 6 – 20. I. 2006, MT, mixed Olea, Cussonia, Vepris, R. Copeland leg., 1 ♂ (NMK). MADAGASCAR: Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 9 – 15. I. 2009, MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG- 53 C- 12, 1 ♂ (CAS); same locality label, 4 – 12. XII. 2008, MG- 53 C- 07, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B53FF3ED139FDADFC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB52B53FF3ED139FDADFC20.taxon	distribution	Distribution. Kenya, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB42B53FF3ED5B7FE3FF831.taxon	description	(Figs 110 B, 112 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB42B53FF3ED5B7FE3FF831.taxon	description	Description. ♀ (Fig. 112 A). Fully winged; body length 3.1 – 4.1 mm. Head and mesosoma completely testaceousreddish; metasoma brown-black; legs testaceous; antenna testaceous-ferruginous. Antenna distally thickened, short, with antennomeres 5 – 9 longer than broad; antennomeres in following proportions: 20: 8: 13: 8: 7: 7: 6.5: 6.5: 6.5: 9. Head convex, dull, covered with short setae, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 6.5; OL = 3; OOL = 12; OPL = 10; TL = 9. Pronotum crossed by strong transverse impression, dull, setose, mainly sculptured by short longitudinal keels, except transverse striae on anterior collar; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, setose, reticulate rugose. Notauli absent. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and slightly rugose); propodeal declivity dull, with two complete longitudinal keels and median and lateral areas reticulate rugose. Forewing of holotype and specimen from Zimbabwe, Naletale ruins environs, with one dark spot situated around 2 r-rs & Rs vein; forewing of specimen in BM (from Botswana, Serowe) hyaline, without dark spots. Distal part of 2 r-rs & Rs vein slightly shorter than proximal part (13: 14); 2 R 1 cell open. Petiole long, much shorter than metasoma (12: 70). Protarsomeres in following proportions: 20: 3: 3: 15: 20. Enlarged claw (Fig. 110 B) with one row of seven-nine teeth and one bristle. Protarsomere 5 (Fig. 110 B) with one preapical lamella, sixeight bristles and an inner band. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB42B53FF3ED5B7FE3FF831.taxon	materials_examined	Material examined. Type: ♀ holotype: BOTSWANA: Central District, Serowe, IV. 1990, MT, P. Forchhammer leg. (CAS). Other material: BOTSWANA: same locality label as holotype, IX. 1985, 1 ♀ (NHMUK). ZIMBA- BWE: Bulawayo, Shangani, Naletale ruins environs, 10. XI. 2014, Snízek leg., 1 ♀ (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB42B53FF3ED5B7FE3FF831.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB42B53FF3ED5B7FE3FF831.taxon	distribution	Distribution. Botswana.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB32B55FF3ED0D4FD01FD48.taxon	description	(Fig. 109 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB32B55FF3ED0D4FD01FD48.taxon	description	Description. ♂. Fully winged; body length 1.9 – 3.6 mm. Head black, except mandible testaceous; antenna testa- ceous-brown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 12: 5.5: 7: 7: 7: 7: 6: 6: 5.5: 8; antennomere 8 less than twice as long as broad (6: 3.5); antennomere 6 less than twice as long as broad (7: 4). In specimen from Republic of the Congo, Lésio-Louna Reserve, antennomere 6 less than twice as long as broad (8: 5). Head convex, dull, covered with short setae, completely reticulate rugose; clypeus reticulate rugose; frontal line complete; occipital carina complete; POL = 7; OL = 3; OOL = 8; OPL = 5; TL = 5. In specimen from Republic of the Congo, Lésio-Louna Reserve, POL = 8; OL = 3. Mesoscutum, mesoscutellum and metanotum dull, completely reticulate rugose. Notauli absent. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectalpropodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity with two complete irregular longitudinal keels, median area almost completely shiny, unsculptured, rugose only near margins, and lateral areas completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (10: 11); 2 R 1 cell open. Genitalia in Fig. 109 D. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB32B55FF3ED0D4FD01FD48.taxon	materials_examined	Material examined. Types: ♂ holotype: TANZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 5 – 14. IV. 1996, MT, Acacia / Commiphora / Combretum bushland, SAM-HYM-PO 17700, S. van Noort leg. (SAMC). Paratype: same locality label, 11. III – 5. IV. 1996, SAM-HYM-PO 17643, 1 ♂ (SAMC). Other material: REPUBLIC OF THE CONGO: Pool Department, Abio, Lésio-Louna Reserve, 03 ° 06.020 ’ S 15 ° 31.440 ’ E, 330 m, MT, 21 – 28. X. 2008, Sharkey & Braet leg., 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB32B55FF3ED0D4FD01FD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB32B55FF3ED0D4FD01FD48.taxon	distribution	Distribution. Republic of the Congo, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB22B55FF3ED56FFE14F816.taxon	description	(Figs 110 C, 112 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB22B55FF3ED56FFE14F816.taxon	description	Description. ♀ (Fig. 112 B). Fully winged; body length 4.1 mm. Head black, except mandible and posterior half of clypeus testaceous; antenna brown, except antennomeres 6 – 10 black; mesosoma black, except lateral margins of pronotum testaceous; petiole yellow; metasoma brown, except apex testaceous; legs brown, except tarsi, trochanters and protibia testaceous. Antenna clavate, short, with antennomeres 5 – 9 approximately as long as broad; antennomeres in following proportions: 15: 9: 14: 8: 7: 6: 6: 6: 6: 7. Head convex, dull, covered with short setae, completely reticulate rugose (sculpture stronger on vertex than on frons); frontal line complete; occipital carina complete; POL = 2; OL = 2; OOL = 13; OPL = 13; TL = 16. Pronotum crossed by strong transverse impression, dull, setose, reticulate rugose; pronotal tubercle reaching tegula. Mesoscutum shiny, with anterior third slightly reticulate rugose and remaining surface partly granulate and partly punctate and unsculptured among punctures. Notauli complete, posteriorly separated; minimum distance between notauli much longer than greatest breadth of lateral ocelli (19: 5). Mesoscutellum very swollen, shiny, sculptured by few punctures, unsculptured among punctures. Metanotum shiny, with three very large areolae. Mesopleuron dull, setose, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with many very large areolae (inner surface of areolae shiny and slightly rugose); propodeal declivity dull, with two complete longitudinal keels and median area laterally and anteriorly reticulate rugose (rest of surface shiny, almost completely punctate and unsculptured among punctures); lateral areas of propodeal declivity reticulate rugose. Forewing with one large dark transverse band around 2 r-rs & Rs vein and one small brown spot on basal cells; distal part of 2 r-rs & Rs vein approximately as long as proximal part (20: 20); 2 R 1 cell open. Petiole long, much shorter than metasoma (7: 29). Protarsomeres in following proportions: 23: 3: 4: 20: 23. Enlarged claw (Fig. 110 C) with one row of 13 teeth and one bristle. Protarsomere 5 (Fig. 110 C) with one preapical lamella, two bristles and a medial prominence. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB22B55FF3ED56FFE14F816.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, 11.5 km 17 ° SSE of Anjozorobe, 3 km 41 ° NE of Andranomay, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 5 – 13. XII. 2000, 1300 m, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2375 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB22B55FF3ED56FFE14F816.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB22B55FF3ED56FFE14F816.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	description	(Figs 110 D, 111, 112 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	diagnosis	Diagnosis. Macropterous ♀ of Bocchus with head mostly yellow-testaceous and completely granulate (Fig. 112 C); OPL about twice as long as OOL; mesoscutum granulate; forewing hyaline, without dark transverse bands or spots; enlarged claw with long row of teeth (Fig. 110 D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	description	Description. ♀ (Figs 111, 112 C). Fully winged; body length 2.3 mm. Head yellow-testaceous, except black margins of ocelli; antenna whitish; mesosoma black; petiole yellow; metasoma brown; legs yellow-testaceous. Antenna clavate, short, with antennomeres 5 – 9 longer than broad; antennomeres in following proportions: 9: 4: 6: 4: 4: 4: 4: 3: 3: 5. Head (Fig. 111 C) convex, dull, completely granulate, not rugose; frontal line complete; occipital carina complete; POL = 3; OL = 4; OOL = 3; OPL = 6; TL = 6; greatest breadth of lateral ocelli shorter than OL (2: 4). Pronotum crossed by strong transverse impression, dull, granulate; pronotal tubercle reaching tegula. Mesoscutum (Fig. 111 A) dull, completely granulate. Notauli complete, posteriorly separated; minimum distance between notauli much longer than breadth of lateral ocelli (7: 2). Mesoscutellum slightly swollen, shiny, slightly granulate. Metanotum shiny, unsculptured. Mesopleuron dull, granulate. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae slightly rugose); propodeal declivity dull, with two complete longitudinal keels, median area transversely striate and lateral areas reticulate rugose. Forewing (Fig. 111 D) hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein shorter than proximal part (5: 8); 2 R 1 cell open. Petiole long, much shorter than metasoma (13: 41). Protarsomeres in following proportions: 10: 3: 3: 8: 11. Enlarged claw (Fig. 110 D) with one row of 5 teeth and 1 bristle. Protarsomere 5 (Fig. 110 D) with one preapical lamella, 3 bristles and medial prominence. Tibial spurs 1, 1, 2 (one of the two spurs of the hind tibiae is shorter). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19450): MADAGASCAR: Antsiranana, Orangea, 3 km E of Ramena near fort, 12 ° 14.49 ’ S 49 ° 22.17 ’ E, 65 m, 23 – 27. I. 2001, MT, littoral forest on sand, MA- 01 - 05 - 02, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg. (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB12B57FF3ED6D7FCADFE37.taxon	etymology	Etymology. The new species is named nigroflavus, a Latin adjective meaning partly black and partly yellow- testaceous (head mostly yellow-testaceous; mesosoma black).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB02B68FF3ED09FFE14FB94.taxon	description	(Figs 109 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB02B68FF3ED09FFE14FB94.taxon	description	Description. ♂. Fully winged; body length 2.1 – 2.9 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown, except tarsi and protibia testaceous. Antenna filiform, with antennomere 6 less than twice as long as broad (5: 3.5); antennomeres in follow- ing proportions: 9: 5: 7: 6: 5: 5: 4.5: 4: 4: 6.5. Head dull, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 3; OL = 2.5; OOL = 6; OPL = 6; TL = 4. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (6: 2.5). In specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, notauli hardly visible. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels (occasionally one or two tracks of longitudinal keels visible). In specimen from Madagascar, 21 ° 13.22 ’ S, 45 ° 19.296 ’ E, propodeal declivity with two tracks of longitudinal keels and median area shiny, unsculptured. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein as long as proximal part (10: 10). Genitalia in Fig. 109 E. In specimens from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E and 22 ° 48.09 ’ S 43 ° 45.54 ’ E, apex of basivolsella with inner pointed apophysis (Fig. 109 F). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB02B68FF3ED09FFE14FB94.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Toliara, 12 km NW of Amboasary, Berenty, 5 – 15. V. 1983, B. M. 1983 – 201, J. S. Noyes & M. C. Day leg. (NHMUK). Paratypes: same locality label as holotype, 13 ♂♂ (5 in NHMUK, 8 in AMNH). Other material: MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 1 ♂ (MOLC); same locality label, 25. II – 7. III. 2003, MA- 02 - 22 - 17, 8 ♂♂ (5 in CAS, 3 in MOLC); same locality label, 30. XI – 7. XII. 2002, MA- 02 - 22 - 07, 1 ♂ (CAS); same locality label, 17 – 26. I. 2003, MA- 02 - 22 - 13, 4 ♂♂ (CAS); Toliara, Manombo, 22 ° 48.09 ’ S 43 ° 45.54 ’ E, 227 m, 9 – 14. IV. 2004, MT, dry forest, Frontier Wilderness Project, MGF 102, 3 ♂♂ (2 in CAS, 1 in MOLC); Toliara, Fiherenana, 23 ° 14.07 ’ S 43 ° 52.15 ’ E, 50 m, 16. X – 16. XI. 2002, MT, degraded gallery forest, MGF 047, Frontier Wilderness Project, 1 ♂ (CAS); Toliara, Fiherenana, 23 ° 10.37 ’ S 43 ° 57.39 ’ E, 100 m, 31. X – 6. XI. 2002, MT, gallery forest, MGF 045, Frontier Wilderness Project, 1 ♂ (CAS); Toliara, Makay Mts., 21 ° 13.22 ’ S, 45 ° 19.296 ’ E, 490 m, 24. XI- 1. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25846, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB02B68FF3ED09FFE14FB94.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEB02B68FF3ED09FFE14FB94.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	description	(Figs 112 D, 113, 114 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	diagnosis	Diagnosis. ♂ of Bocchus with head and mesoscutum reticulate rugose (Fig. 112 D); notauli complete; propodeal declivity with two complete longitudinal keels; genitalia with volsella slightly shorter than paramere; apex of basivolsella with one subdistal inner pointed apophysis (Fig. 114 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	description	Description. ♂ (Figs 112 D, 113). Fully winged; body length 2.2 – 3.4 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, partly testaceous. Antenna filiform; antennomeres in following proportions: 11: 7: 9: 8: 7: 7: 6: 6: 5: 7. Head (Fig. 113 C) dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 4; OL = 2; OOL = 9; OPL = 8; TL = 5; greatest breadth of posterior ocellus longer than OL (3.5: 2). Mesoscutum (Fig. 113 A), mesoscutellum and metanotum reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (9: 3.5). In specimens from Madagascar, 16 ° 13.41 ’ S 46 ° 08.37 ’ E and 16 ° 19.16 ’ S 46 ° 48.80 ’ E, notauli hardly visible. Metapectal-propodeal disc reticulate rugose; propodeal declivity with two complete longitudinal keels, median area as rugose as lateral areas. Forewing (Fig. 113 D) hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (12: 11); 2 r-rs & Rs vein not S-shaped. Petiole short. Apex of paramere rounded (Fig. 114 A); apex of basivolsella broadened and with subdistal inner pointed apophysis (Fig. 114 A). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19461): MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, elev 35 m, 24. III – 4. IV. 2004, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 A- 15 (CAS). Paratypes: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 22.9 km 224 ° SW Anivorano Nord, 12 ° 54.32 ’ S 49 ° 6.35 ’ E, 80 m, 10 – 18. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 2856, 3 ♂♂ (2 in CAS, 1 in MOLC); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Mahajanga, Ankarafantsika National Park, Tsimaloto Forest, 18.3 km 49 ° NE Tsaramandroso, 16 ° 13.41 ’ S 46 ° 08.37 ’ E, 135 m, 2 – 8. IV. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3600, 1 ♂ (CAS; Mahajanga, Sofia District, 45 km S Antso- hihy, 5 km W Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 5 – 8. V. 2011, MT, MG- 61 - 11, 1 ♂ (MOLC); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, 43 m, 28. IX – 5. X. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 17, 1 ♂ (CAS); Toliara, same locality label as holotype, 2 ♂♂ (CAS, MOLC); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 24. III – 4. IV. 2004, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 - 01, 2 ♂♂ (CAS, MOLC); same locality label, 17 – 24. III. 2003, MA- 02 - 22 - 19, 12 ♂♂ (7 in CAS, 5 in MOLC); Toliara, Andro- impano Forest, 3 km E of Itampolo, 24 ° 39.47 ’ S 43 ° 57.37 ’ E, 45 m, 25. XII. 2008 – 1. I. 2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54 B- 09, 2 ♂♂ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI- 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 1 ♂ (MOLC). Other material: MADAGASCAR: Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 23 – 30. III. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 15, 3 ♂♂ (2 in CAS, 1 in MOLC); Antsiranana, SAVA Region, Vohemar District, Antsahabelela rain forest, 9 km SW Daraina, Binara, 13 ° 15.03 ’ S 49 ° 37.00 ’ E, 182 m, 8 – 15. XII. 2010, MT, in humid forest, M. Irwin, R. Harin’Hala leg., MG- 58 - 10, 1 ♂ (CAS); same locality label, 25. V – 1. VI. 2011, MG- 58 - 34, 1 ♂ (MOLC); same locality label, 16 - 23. II. 2011, MG- 58 - 20, 1 ♂ (CAS); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W Anjiamangirana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 5 – 8. V. 2011, MT, MG- 61 - 11, 9 ♂♂ (4 in CAS, 5 in MOLC); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 17 – 26. I. 2003, M. Irwin, F. Parker & R. Harin’Hala, MT, gallery forest, MA- 02 - 22 - 13, 6 ♂♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8F2B6AFF3ED22BFBC5F9D4.taxon	etymology	Etymology. The new species is named after one the collectors, Dr. F. Parker.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8D2B6BFF3ED0EBFE21FC90.taxon	description	(Fig. 110 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8D2B6BFF3ED0EBFE21FC90.taxon	description	Description. ♀. Fully winged; body length 4.8 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 6 – 10 brown; mesosoma black, except part of lateral regions of pronotum testaceous; petiole and metasoma black; legs testaceous, except meso- and metacoxa, clubs of meso- and metafemur and meso- and metatibia brown. Antenna clavate, short, with antennomeres 5 – 9 approximately as long as broad (6: 5); antenno- meres in following proportions: 21: 8: 14: 9: 6: 6: 6: 6: 6: 9. Head convex, dull, covered with short setae, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 13; OPL = 11; TL = 11; greatest breadth of lateral ocelli shorter than OL (3: 4). Pronotum crossed by strong transverse furrow, dull, with dorsal region slightly rugose and lateral regions crossed by strong transverse keels; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Areolae of metanotum broader than those of mesoscutum and mesoscutellum. Notauli absent. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and punctate); propodeal declivity shiny, with two complete longitudinal keels and median and lateral areas sculptured by numerous transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (17: 15); 2 R 1 cell open. Petiole long, much shorter than metasoma (13: 67). Protarsomeres in following proportions: 22: 3.5: 4: 15: 20. Enlarged claw (Fig. 110 E) with one row of eight teeth and one bristle. Protarsomere 5 (Fig. 110 E) with one preapical lamella, two bristles, a medial prominence and one long proximal lamella. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8D2B6BFF3ED0EBFE21FC90.taxon	materials_examined	Material examined. Type: ♀ holotype: GUINEA: 10 km W Mamou, 10 ° 22.18 ’ N 12 ° 07.75 ’ W, 5. VII. 2004, Marek Halada leg. (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8D2B6BFF3ED0EBFE21FC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8D2B6BFF3ED0EBFE21FC90.taxon	distribution	Distribution. Guinea.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6BFF3ED527FE5DF959.taxon	description	(Fig. 110 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6BFF3ED527FE5DF959.taxon	description	Description. ♀. Fully winged; body length 2.9 mm. Completely testaceous; antenna testaceous, with antennomeres 7 – 10 brown. Antenna distally thickened, short, with antennomeres 5 – 9 longer than broad; antennomeres in following proportions: 12: 6: 10: 7: 7: 7: 7: 7: 6.5: 9. Head convex, dull, covered with short setae, granulate, sculptured by irregular keels and rugose; frontal line complete; occipital carina complete; POL = 4; OL = 2.5; OOL = 10; OPL = 8; TL = 9. Pronotum crossed by strong transverse impression, shiny, setose, partly smooth and partly slightly rugose; pronotal tubercle reaching tegula. Mesoscutum setose, granulate, sculptured by numerous irregular longitudinal keels and slightly rugose. Notauli absent. Mesoscutellum and metanotum rugose. Mesopleuron and metapleuron dull, strongly reticulate rugose. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and slightly rugose); propodeal declivity dull, with two complete longitudinal keels, median area slightly rugose and lateral areas strongly reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (7: 9); 2 R 1 cell open. Petiole long, much shorter than metasoma (5: 48). Protarsomeres in following proportions: 19: 3: 3: 11: 17.5. Enlarged claw (Fig. 110 F) of right chela with one row of nine teeth and one bristle; enlarged claw of left chela with one row of six teeth and one bristle. Protarsomere 5 (Fig. 110 F) with apex very large, without preapical lamella, with two bristles and an inner band. Tibial spurs 1 / 1 / 1 (for mistake 1 / 1 / 2 in original description). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6BFF3ED527FE5DF959.taxon	materials_examined	Material examined. Type: ♀ holotype: YEMEN: Al Lahima, 9. IV – 5. VI. 2001, MT, A. van Harten leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6BFF3ED527FE5DF959.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6BFF3ED527FE5DF959.taxon	distribution	Distribution. Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6CFF3ED17EFE1FFD00.taxon	description	(Fig. 115 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6CFF3ED17EFE1FFD00.taxon	description	Description. ♀. Micropterous; forewing about as long as tegula. Body length 3.3 mm. Completely reddish-dark. Antenna clavate; antennomeres in following proportions: 15: 6: 9: 6: 6: 6: 5: 5: 5: 8. Head shiny, granulate and with slight areolae; frontal line complete; occipital carina complete; dorsal side of head with two lateral keels around orbits; POL = 5; OL = 3; OOL = 12; OPL = 8; TL = 10. Pronotum crossed by transverse impression between anterior and posterior collar, shiny, setose, slightly granulate; pronotal tubercle reaching tegula. Mesoscutum dull, granulate, with slight areolae and sculptured by strong longitudinal keels. Notauli absent. Mesoscutellum dull, sculptured by longitudinal keels. Metanotum very short, reduced. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, not sculptured by longitudinal keels. Protarsomeres in following proportions: 15: 3: 3: 11: 14. Enlarged claw (Fig. 115 A) with one row of nine teeth and one bristle. Protarsomere 5 (Fig. 115 A) with one preapical lamella and one row of eight bristles. Tibial spurs 1 / 1 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6CFF3ED17EFE1FFD00.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Cape Town, Milnerton, I. 1926, R. E. Turner leg. (NHMUK). Paratype: SOUTH AFRICA: Western Cape, Cape Town, 1 ♀ (USNM).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6CFF3ED17EFE1FFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8C2B6CFF3ED17EFE1FFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	description	(Figs 114 B, 116, 117 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	diagnosis	Diagnosis. ♂ of Bocchus with mesoscutum reticulate rugose (Fig. 117 A); notauli incomplete, reaching approxi- mately 0.5 – 0.8 × length of mesoscutum; propodeal declivity with median area at least partly shiny, unsculptured; apex of basivolsella with one subdistal inner pointed apophysis (Fig. 114 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	description	Description. ♂ (Figs 116, 117 A). Fully winged; body length 2.2 – 2.9 mm. Head black, except mandible testaceous; antenna ferruginous; mesosoma black; metasoma brown; legs brown, except procoxa and tarsi testaceous. Antenna filiform; antennomeres in following proportions: 11: 6: 7: 7: 7: 7: 7: 6: 6: 8; antennomere 8 less than 3 × as long as broad (6: 3). Head (Fig. 116 C) convex, dull, covered with short setae, granulate and reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 3; OOL = 8; OPL = 6; TL = 3; greatest breadth of lateral ocelli longer than OL (4: 3). Mesoscutum (Fig. 116 A), mesoscutellum and metanotum dull, completely strongly reticulate rugose. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum (in specimens from Madagascar, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, notauli reaching about 0.8 × length of mesoscutum). Mesopleuron dull, strongly reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity with two complete longitudinal keels, lateral areas completely reticulate rugose and median area partly shiny and unsculptured. In paratype from Madagascar, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, median area of propodeal declivity almost completely reticulate rugose, with unsculptured central area very small. Forewing (Fig. 116 D) hyaline, without dark transverse bands; 2 r-rs & Rs vein with distinct separation between distal and proximal part; distal part of 2 r-rs & Rs vein slightly shorter than proximal part (10: 11). Apex of basivolsella with one subdistal inner pointed apophysis (Fig. 114 B). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19452): MADAGASCAR: Toliara, Beza Mahafaly Re- serve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 9 – 20. IX. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 35 (CAS). Paratypes: MADAGASCAR: Fianarantsoa, near Isalo National Park, in dry wash east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 9 – 17. XI. 2001, MT, in open area, MA- 02 - 11 B- 02, R. Harin’Hala leg., 1 ♂ (MOLC); Mahajanga, Besalampy District, Analangidro dry forest, 7 km NE Besalampy, 16 ° 41.49 ’ S 44 ° 31.41 ’ E, 200 ft., 9 – 16. X. 2007, MT, dry forest on sand, M. Irwin & R. Harin’Hala leg., MG- 41 - 03, 2 ♂♂ (CAS); same locality label, 24. IX – 2. X. 2007, MG- 41 - 01, 2 ♂♂ (CAS); Mahajanga, Ambovomamy Belembo, 20 km NW Port Berger, 15 ° 27.07 ’ S 47 ° 36.80 ’ E, 33 m, 26. III – 1. IV. 2007, secondary growth on white sand, MT, R. Harin’Hala, M. Irwin, F. Parker leg., MG- 33 - 14, 2 ♂♂ (CAS, MOLC); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 11 – 18. XII. 2007, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 11, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 9 – 20. IX. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 35, 1 ♂ (CAS); same locality label, 29. III – 10. IV. 2002, MA- 02 - 14 A- 22, 1 ♂ (MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 11 – 24. II. 2003, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 46, 1 ♂ (CAS); same locality label, 27. VII – 3. VIII. 2003, MA- 02 - 18 A- 63, 1 ♂ (CAS); same locality label, 6 – 16. I. 2002, MA- 02 - 18 A- 09, 2 ♂♂ (CAS). Other material: MADAGASCAR: Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 23. II – 2. III. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 11, 3 ♂♂ (MOLC); same locality label, 15 – 22. VI. 2011, MG- 62 - 27, 1 ♂ (MOLC); same locality label, 18 – 26. V. 2011, MG- 62 - 23, 5 ♂♂ (3 in CAS, 2 in MOLC); same locality label, 12 – 19. I. 2011, MG- 62 - 05, 1 ♂ (CAS); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♂ (MNHN); Mahajanga, Namoroka National Park, 16.40682 S 45.31377 E, 21 – 31. X. 2016, MT, C. Villemant leg., 2 ♂♂ (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	etymology	Etymology. The Latin adjective ruvidus (= rugose) refers to the rugose sculpture of head, mesoscutum, mesoscutellum and metanotum.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE8B2B6EFF3ED117FB5AFBDC.taxon	discussion	Remarks. In the paratype from Madagascar, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, the median area of the propodeal declivity is almost completely reticulate rugose, with a very small unsculptured central area, with the result that it may be confused with Bocchus johanssoni Olmi, Copeland & Guglielmino, that has the median area completely reticulate rugose. However, in B. johanssoni the antenna is more slender than in B. ruvidus (antennomere 8 much more than twice as long as broad (8: 3) in B. johanssoni; about twice as long as broad (6: 3) in B. ruvidus).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE892B6FFF3ED2E3FE14F9F0.taxon	description	(Figs 114 C, 115 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE892B6FFF3ED2E3FE14F9F0.taxon	description	Description. ♀. Fully winged; body length 4.6 – 5.5 mm. Head reddish, except brown ocellar spot; antenna brown, except scape and pedicel or also flagellomeres 1 – 2 reddish; mesosoma reddish; metasoma black-brown or browntestaceous; legs reddish. In specimen from Madagascar, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, colour completely testaceousreddish, except antenna brown, ocellar triangle black and proximal half of metasoma black. In specimen from Madagascar, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, metasoma brown, except distal third and ventral side of segment 1 testaceous. In specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, legs ferruginous, except one black spot on metacoxa and metatibia and club of metafemur slightly darkened; metasoma with proximal half brown and distal half testaceous. Antenna short, clavate, with antennomeres 5 – 9 approximately as long as broad; antennomeres in following proportions: 18: 7: 15: 7: 5.5: 5: 6: 6: 5: 7. Head dull, completely reticulate rugose, setose; frontal line complete; occipital carina complete; POL = 2; OL = 2; OOL = 15; OPL = 19; TL = 14; greatest breadth of lateral ocelli longer than OL (4: 2). Mesoscutum, mesoscutellum and metanotum dull, setose, completely reticulate rugose. Notauli absent. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Petiole long. Forewing with two dark transverse bands; pterostigma short and narrow. Protarsomeres 1 – 4 in following proportions: 12: 2: 3: 12. Enlarged claw (Fig. 115 B) with one row of 11 – 13 teeth and one bristle. Protarsomere 5 (Fig. 115 B) with one preapical lamella and two bristles. Tibial spurs 1 / 1 / 1. ♂. Fully winged; body length 2.4 – 2.6 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel partly testaceous; mesosoma black; metasoma brown; legs brown, except tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 11: 6: 7: 7: 6.5: 6: 5: 5: 5: 8; antennomere 6 less than twice as long as broad (6: 3.5); antennomere 8 less than twice as long as broad (5: 3). Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 8; OPL = 7; TL = 4; greatest breadth of lateral ocelli about as long as OL. Mesoscutum, mesoscutellum and metanotum reticulate rugose. Notauli absent. In specimen from Madagascar, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, right notaulus slightly distinct and complete; in specimens from Madagascar, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, mesoscutum with tracks of notauli hardly visible under special conditions of light. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two irregular and incomplete longitudinal keels, median area partly shiny and punctate and lateral areas reticulate rugose. In specimen from Madagascar, 20 ° 04.07 ’ S 44 ° 39.34 ’ E, propodeal declivity with median area completely rugose, without smooth area. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 10). Petiole short. Genitalia in Fig. 114 C. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE892B6FFF3ED2E3FE14F9F0.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Behara, XI. 1940, A. Seyrig leg. (MNHN). Paratype: MADAGASCAR: Toliara, Bekily, VII. 1937, A. Seyrig leg., 1 ♀ (MRAC). Other material: MADAGAS- CAR: Fianarantsoa, Ananalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7383, 1 ♂ (CAS); Fianarantsoa, near Isalo National Park, in dry wash east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 7 – 22. IX. 2002, MT, in open area, MA- 02 - 11 B- 39, R. Harin’Hala leg., 1 ♀ (CAS); Mahajanga, Namoroka National Park, 9.8 km 300 ° WNW Vilanandro, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, 140 m, 4 – 8. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445, 1 ♀, 2 ♂♂ (CAS); Mahajanga, Namoroka National Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, 100 m, 12 – 16. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6581, 1 ♂ (CAS); Mahajanga, Ambato- folaka, Namoroka, 53 km from Soalala, 3 km N Vilanandro village, 16 ° 28.04 ’ S 45 ° 23.48 ’ E, 400 ft., 2 – 9. XI. 2007, dense dry forest, MT, M. Irwin, R. Harin’Hala leg., MG- 40 A- 06, 1 ♂ (CAS); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 3 ♂♂ (2 in CAS, 1 in MOLC); Mahajanga, Ambovomamy Belembo, 20 km NW Port Berger, 15 ° 27.07 ’ S 47 ° 36.80 ’ E, 33 m, 26. III – 1. IV. 2007, secondary growth on white sand, MT, R. Harin’Hala, M. Irwin, F. Parker leg., MG- 33 - 14, 2 ♂♂ (CAS); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, 43 m, 17 – 24. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 11, 1 ♂ (CAS); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.09 ’ S 044 ° 39.38 ’ E, 10 – 17. IX. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 45, 1 ♀, 1 ♂ (CAS); same locality label, 14 – 21. XI. 2007, BLF 18285 _ 02, 1 ♀ (MOLC); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.07 ’ S 44 ° 39.34 ’ E, 17 – 24. IX. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18285 _ 45, 1 ♂ (MOLC); same locality label, 26. XI – 3. XII. 2008; BLF 18284 _ 56, 1 ♂ (MOLC); same locality label, 5 – 12. XII. 2007, BLF 18284 _ 05, 1 ♂ (CAS); Toliara, Berenty Spe- cial Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 24. III – 3. IV. 2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 20, 1 ♀ (MOLC); same locality label, 27. XII. 2002 – 7. I. 2003, MA- 02 - 22 - 11, 1 ♀ (MOLC); same locality label, 7 – 17. III. 2003, MAQ- 02 - 22 - 18, 1 ♀ (CAS); same locality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 1 ♀ (MOLC); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 1 ♀ (MOLC); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 20. I – 13. II. 2003, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 48, 1 ♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 17 – 27. VII. 2003, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 62, 1 ♂ (CAS); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 430 ft., 30. X – 6. XI. 2007, Antsarongaza dry forest, MT, MG- 45 A- 07, M. Irwin, R. Harin’Hala leg., 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE892B6FFF3ED2E3FE14F9F0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE892B6FFF3ED2E3FE14F9F0.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE882B60FF3ED087FE1FFD00.taxon	description	(Fig. 114 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE882B60FF3ED087FE1FFD00.taxon	description	Description. ♂. Fully winged; body length 3.6 mm. Head black, except mandible testaceous; antenna brown; me- sosoma black; metasoma brown; legs brown. Antenna filiform, setose; antennomeres in following proportions: 16: 7: 10: 11: 9: 10: 9: 10: 8: 10; antennomere 6 more than 3 × as long as broad (10: 3). Head dull, reticulate rugose and granulate; frontal line complete; frons without two lateral keels around orbits; occipital carina complete; POL = 8; OL = 4.5; OOL = 8 OPL = 5; TL = 7; greatest breadth of lateral ocelli shorter than OL (2: 4.5). Mesoscutum dull, granulate and sculptured by many irregular longitudinal keels. Notauli complete, posteriorly separated; minimum distance between notauli much longer than greatest breadth of lateral ocelli (10: 2). Mesoscutellum dull, granulate and sculptured by some irregular longitudinal keels. Metanotum shiny, weakly granulate, slightly rugose near anterior margin. Metapectal-propodeal disc dull, with transverse posterior keel, reticulate rugose; propodeal declivity with two longitudinal keels, lateral areas reticulate rugose and median area partly rugose and partly (posterior half) shiny, unsculptured. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein about as long as proximal part. Genitalia in Fig. 114 D. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE882B60FF3ED087FE1FFD00.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Milnerton, I. 1926, R. E. Turner leg., Brit. Mus. 1926 – 71 (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE882B60FF3ED087FE1FFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE882B60FF3ED087FE1FFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	description	(Figs 114 E, 115 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	description	Description. ♀. Fully winged; body length 3.6 – 4.1 mm. Head and mesosoma of holotype completely testaceousreddish, except ocellar triangle brown and metapectal-propodeal disc partly brown (in three paratypes head black, except clypeus, mandible and posterior margin of vertex ferruginous; in specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, head totally ferruginous-testaceous); mesosoma ferruginous, except mesosternum totally or partly black and part of metapectal-propodeal disc and propodeal declivity black. Petiole testaceous. Metasoma brownblack. Legs testaceous. Antenna of holotype testaceous-darkened, except scape and pedicel testaceous. Antenna of three paratypes brown, except scape and pedicel testaceous. In specimen from Kenya, 00 ° 64.725 ’ S 34 ° 33.595 ’ E, head black, except clypeus and mandible testaceous; antennae brown, except scape and pedicel testaceous; mesosoma black, except prothorax ferruginous; petiole testaceous; metasoma brown-testaceous; legs testaceous. In specimen from Kenya, 1.90103 ° S 36 ° 04804 ° E, head completely black, except mandible testaceous; mesosoma ferruginous, except small black spot on central region of metapectal-propodeal disc. Antenna clavate, short, with antennomeres 5 – 9 longer than broad (length / breadth ratio of 9 th antennomere: 8: 3.5); antennomeres in following proportions: 13: 7: 16: 11: 10: 10: 9.5: 8: 8: 12. Head convex, dull, covered with short setae, reticulate rugose, except two large granulate areas of frons on sides of frontal line; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 14; OPL = 9; TL = 13. In specimen from Kenya, 1.90103 ° S 36 ° 04804 ° E, head completely reticulate rugose, without granulate areas on sides of frontal line. Pronotum crossed by strong transverse impression, dull, setose, with anterior collar unsculptured and lateral regions and posterior collar reticulate rugose or sculptured by irregular keels; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, setose, reticulate rugose (in specimen from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, median region of mesoscutum granulate and reticulate rugose (rugosity hardly visible )). Notauli complete, posteriorly separated; minimum distance between notauli longer than POL (9: 3). Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, rugose, with few very large areolae (inner surface of areolae shiny and slightly rugose); propodeal declivity dull, with two complete longitudinal keels and median and lateral areas reticulate rugose. Forewing with two dark transverse bands (occasionally proximal band reduced to small and hardly visible spot); distal part of 2 r-rs & Rs vein shorter than proximal part (9: 12); 2 R 1 cell open. Petiole long, much shorter than metasoma (14: 54). Protarsomeres in following proportions: 23: 3: 4: 14: 20. Enlarged claw (Fig. 115 C) of holotype with one row of eight teeth (in addition to pointed apex) and one broad lamella-like bristle. Enlarged claw of other specimens with one row of five-seven teeth. Protarsomere 5 (Fig. 115 C) with one preapical lamella, two bristles and an inner band. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 1.9 – 2.5 mm. Head black, except mandible testaceous; antenna brown, or browntestaceous, except scape and pedicel partly testaceous; mesosoma and metasoma black; legs brown, except tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 9: 5: 7: 7: 7: 7: 6: 6: 5.5: 8. Head shiny, reticulate rugose; frontal line complete; occipital carina complete; head of holotype with POL = 5; OL = 3; OOL = 7; OPL = 5; TL = 6; head of specimen from Botswana, Farmer’s Brigade, with POL = 5; OL = 3.5; OOL = 5; OPL = 5; TL = 6. Mesoscutum, mesoscutellum and metanotum reticulate rugose (in some specimens from South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, mesoscutellum with anterior half granulate and posterior half reticulate rugose). Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (5: 3 in holotype; 6: 3 in specimen from Botswana, Farmer’s Brigade). Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (7: 11). Petiole short. Genitalia (Fig. 114 E) with apex of paramere flat and apex of basivolsella slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	materials_examined	Material examined. Types: ♂ holotype of B. watshami: NAMIBIA: Gobabis, XI. 1973, A. Watsham leg. (NHMUK); ♀ holotype of B. simoni: TANZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 5 – 25. XII. 1995 – 29. I. 1996, MT, Acacia / Commiphora / Combretum bushland, SAM-HYM-PO 15907, S. van Noort leg. (SAMC). Paratypes of B. simoni: TANZANIA: same locality label as holotype, 11. III – 5. IV. 1996, SAM-HYM- PO 17643, 1 ♀ (SAMC); Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ’ S 38 ° 01.37 ’ E, 16. IV – 2. V. 1996, MT, Commiphora bushland, SAM-HYM-PO 16285, S. van Noort leg., 2 ♀♀ (SAMC, MOLC); Mkomazi Game Reserve, Kikolo Plot, 04 ° 06.72 ’ S 38 ° 01.37 ’ E, 25. XI – 8. XII. 1995, MT, Commiphora bushland, SAM-HYM-PO 17870, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Kisima Plot, 04 ° 06.06 ’ S 38 ° 05.58 ’ E, 25. XI – 8. XII. 1995, MT, Acacia / Commiphora bushland, SAM-HYM-PO 16498, S. van Noort leg., 1 ♀ (SAMC). Other material: BOTSWA- NA: Serowe, Farmer’s Brigade, XII. 1987, MT, Per Forchhammer leg., 1 ♂ (AMNH). GUINEA-BISSAU: Cacheu Region, 2 km E of Bigene, 1. XI. 1992, M. Söderlund leg., 2 ♂♂ (AMNH, ZIL). KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 3 – 17. VII. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♀ (NMK); same locality label, 10 – 24. IV. 2015, 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 8 – 15. IV. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22338 ° S 36.89655 ° E, 1604 m, 24. II – 3. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Nyanza Prov., Ruma National Park, near Kamato Gate, 00 ° 64.725 ’ S 34 ° 33.595 ’ E, 1264 m, MT, open grassland-woodland, 4 – 18. III. 2006, R. Copeland leg., 1 ♀ (NMK); Nyanza Prov., Gembe Hills, 00 ° 29.36 ’ S 34 ° 14.60 ’ E, 1362 m, 12 – 26. II. 2005, MT, R. Copeland leg., 1 ♀ (MOLC); Rift Valley Prov., Nguruman, near Sampu River, 1.90103 ° S 36 ° 04804 ° E, 753 m, MT, base of Ngu- ruman Escarpment, 8 – 22. XII. 2007, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Olorgesailie National Monu- ment, 1.57962 ° S 36.44730 ° E, 979 m, 4 – 18. VI. 2011, MT, near Ol Keju Nyiro River, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Matthews Range, 0.97984 ° N 37.34599 ° E, 1459 m, 16 – 26. XII. 2015, MT, riverine forest, near Wamba, R. Copeland leg., 1 ♂ (NMK). MOZAMBIQUE: Niassa, Cuamba, Mozambique Catholic University Farm, 17. IV – 2. VI. 2006, MT, M. Olmi leg., 1 ♂ (MOLC). SOUTH AFRICA: Eastern Cape, Pearston, Plains of Camdeboo Game Reserve, 32 ° 32.033 ’ S 25 ° 14.267 ’ E, 969 m, 30. X. 2009 – 22. II. 2010, MT, Camdeboo Escarpment Ticket, S. van Noort leg., 1 ♀ (SAMC), 21 ♂♂ (20 in SAMC, 1 in MOLC); same locality label, 6. X. 2010 – 18. I. 2011, PCD 09 - ACA 1 - M 06, 1 ♀ (SAMC); Limpopo, Ellisras District, D’Nyala Nature Reserve, 23 ° 45 ’ S 27 ° 49 ’ E, 8 – 12. XII. 1989, 1 ♂ (SANC); Mpumalanga, Krüger National Park, Skukuza, 12 – 15. XII. 1985, thornscrub forest, S. & I. Peck leg., 1 ♂ (PMA); same locality label, 11. I. 1985, M. Olmi leg., 1 ♂ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	distribution	Distribution. Botswana, Guinea-Bissau, Kenya, Mozambique, Namibia, South Africa, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE872B61FF3ED497FC15F964.taxon	discussion	Remarks. Bocchus simoni Olmi (described on the basis of a ♀) is here tentatively synonymized with B. watshami (described on the basis of a ♂), because both sexes were collected in the same locality (South Africa, 32 ° 32.033 ’ S 25 ° 14.267 ’ E). Their head, mesoscutum, mesoscutellum and metanotum have the same sculpture, so that it is likely that they are the opposite sexes of B. watshami, though they were not reared.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE862B62FF3ED17BFD08FA60.taxon	description	(Fig. 114 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE862B62FF3ED17BFD08FA60.taxon	description	Description. ♂. Fully winged; body length 1.6 – 3.2 mm. Head black, except mandible testaceous; mesosoma black; metasoma brown; antenna testaceous-brown; legs brown, except articulations, tarsi and protibia testaceous. Antenna filiform; antennomeres in following proportions: 13: 7: 9: 8: 8: 8: 8: 8: 7.5: 9.5; antennomere 6 more than twice as long as broad (8.5: 3.5). In specimens from Kenya, 0.32442 ° N 38.05038 ° E, antennomere 6 twice as long as broad. Head convex, dull, covered with short setae, completely reticulate rugose; clypeus reticulate rugose; frontal line complete or incomplete (then only present in anterior third or half of frons near clypeus); occipital carina complete; POL = 6; OL = 4; OOL = 9; OPL = 4; TL = 6. In specimen from Kenya, 0.32442 ° N 38.05038 ° E, POL = 4; OOL = 9. Mesoscutum, mesoscutellum and metanotum dull, completely reticulate rugose. Notauli absent. In small specimen (1.56 mm) from Madagascar, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, mesoscutellum granulate. Mesopleuron dull, reticulate rugose. Metapleuron dull, sculptured by numerous strong transverse keels. Metapectal-propodeal disc with strong transverse posterior keel, dull, reticulate rugose; propodeal declivity with two complete longitudinal keels, median area almost completely shiny and unsculptured, rugose only near margins, and lateral areas completely reticulate rugose. In specimens from Kenya, 0.32442 ° N 38.05038 ° E and 4.48576 ° S 39.25845 ° E, propodeal declivity with median area dull, completely reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (13: 10); 2 R 1 cell open. Genitalia in Fig. 114 F. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE862B62FF3ED17BFD08FA60.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Ramsgate, butterfly sanctuary, 30 ° 53.3 ’ S 30 ° 20.4 ’ E, 26 – 28. III. 2004, YPT, M. Mostovski leg. (NMSA). Paratypes: same locality label as holotype, 6. VI – 9. VII. 2004, 2 ♂♂ (NMSA, MOLC); same locality label as holotype, 9. VII – 9. VIII. 2004, 1 ♂ (NMSA); same locality label as holotype, 9. VIII – 2. IX. 2004, 1 ♂ (NMSA); same locality label as holotype, 3. IX – 2. X. 2004, 2 ♂♂ (NMSA). Other material: KENYA: Coast Prov., Buda Forest, 4.46109 ° S 39.40446 ° E, 98 m, 15 – 29. I. 2016, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); same locality label, 11 – 25. III. 2016, 1 ♂ (NMK); same locality label 8 – 22. IV. 2016, 1 ♂ (NMK); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 30. V – 19. VI. 2013, 6 m MT, indigenous forest, R. Copeland leg., 2 ♂♂ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 6 – 20. IX. 2014, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 19. IX – 3. X. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Ka- sigau Mtn., 3.82700 ° S 38.05038 ° E, 1065 m, 5 – 19. X. 2011, MT, inside indigenous forest, R. Copeland leg., 7 ♂♂ (NMK); Eastern Province, Ngaia Forest, 0.32442 ° N 38.05038 ° E, 1057 m, bottom of forest, MT, inside indigenous forest, 22. VI – 6. VII. 2011, R. Copeland leg., 2 ♂♂ (NMK); same locality label, 11 – 25. V. 2011, 3 ♂♂ (NMK); same locality label, 12 – 26. XI. 2011, 2 ♂♂ (NMK); same locality label, 8 – 22. VI. 2011, 15 ♂♂ (14 in NMK, 1 in MOLC); same locality label, 24. XII. 2011 – 7. I. 2012, 11 ♂♂ (5 in NMK, 2 in MOLC); same locality label, 13 – 27. IV. 2011, 4 ♂♂ (NMK); same locality label, 7 – 21. I. 2012, 2 ♂♂ (NMK); same locality label, 10 – 24. XII. 2011, 1 ♂ (NMK); same locality label, 9 – 23. VII. 2011, 1 ♂ (NMK). MADAGASCAR: Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 1 ♂ (CAS). SOUTH AFRICA: KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, MT, Maputaland Coastal Belt, SA 10 - MAB-M 01, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE862B62FF3ED17BFD08FA60.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE862B62FF3ED17BFD08FA60.taxon	distribution	Distribution. Kenya, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE852B63FF3ED077FD62FE54.taxon	description	(Fig. 115 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE852B63FF3ED077FD62FE54.taxon	description	Description. ♂. Fully winged; body length 2.2 – 2.6 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except tarsi testaceous. Antenna filiform; antennomeres in following proportions: 10: 5: 5.5: 6: 6: 6: 5.5: 6.5: 5.5: 8. Head dull, completely reticulate rugose and granulate; frontal line complete; occipital carina complete; head of holotype with POL = 5; OL = 2.5; OOL = 6; OPL = 3; TL = 5. Mesoscutum completely granulate and with numerous irregular keels. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (4: 2). Mesoscutellum shiny, with anterior third unsculptured and posterior surface granulate and with irregular keels. Metanotum shiny, slightly granulate. Metapectal-propodeal disc reticulate rugose, with strong transverse posterior keel; propodeal declivity with two longitudinal keels and median area as rugose as lateral areas. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (7: 10). Genitalia in Fig. 115 D. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE852B63FF3ED077FD62FE54.taxon	materials_examined	Material examined. Type: ♂ holotype: ZIMBABWE: Gokwe, Sengwa, I. 1983, B. Bell leg. (PMA). Other material: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 57 m, 4. X – 1. XI. 1997, MT, South Coast Strandveld, S. van Noort leg., 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE852B63FF3ED077FD62FE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE852B63FF3ED077FD62FE54.taxon	distribution	Distribution. South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	type_taxon	Type genus. Dryinus Latreille, 1804 (type genus of the family Dryinidae).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	diagnosis	Diagnosis. ♀ (Figs 117 B, C, 154 C, D): fully winged (Figs 117 B, C); ocelli present; occipital carina complete, or incomplete, or absent; palpal formula usually 6 / 3; occasionally palpal formula different (4 / 2, or 5 / 3 in Gonadryinus Olmi; 3 / 2, or 4 / 2, or 5 / 3 in Pseudodryinus Olmi); mandible usually with four teeth becoming regularly progressively larger from dorsal to ventral tooth (as in Fig. 6 B); occasionally (in Pseudodryinus Olmi) mandible with four irregular teeth; antenna without tufts of long setae (Figs 117 B, C); pronotal tubercle present (Fig. 117 C); epicnemium visible, because lateral regions of prothorax not continuous with mesopleura (as in Fig. 7 D); forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 117 B, C); forewing with 2 r-rs & Rs vein and pterostigma (Figs 117 B, C); protarsus chelate (Figs 117 B, C); chela with rudimentary claw (Fig. 126 F); protrochanter long and slender (Fig. 118 C); usually tibial spurs 1 / 1 / 2; rarely (in some species of Dryinus Latreille) 1 / 1 / 1. ♂ (Figs 117 D, 118 A): fully winged (Figs 117 D, 118 A); palpal formula 6 / 3; occipital carina complete or incomplete; mandible usually with one – three teeth (three teeth becoming regularly progressively larger from dorsal to ventral tooth (Fig. 8 A )); occasionally (in Pseudodryinus Olmi) mandible with four irregular teeth; epicnemium visible, because lateral regions of prothorax not continuous with mesopleura (Fig. 7 D); mesosternum fused with mesopleura and not distinct; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 117 D, 118 A); forewing with 2 r-rs & Rs vein and pterostigma (Figs 117 D, 118 A); forewing with PostabR 1 as long as, or longer than pterostigma (Fig. 117 D); hypopygium umbrella-shaped (as in Fig. 8 G); dorsal process of paramere absent (Fig. 119 D); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	biology_ecology	Hosts. Acanaloniidae, Cixiidae, Dictyopharidae, Flatidae, Fulgoridae, Issidae, Lophopidae, Ricaniidae, Tropiduchidae (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	distribution	World species. 344 species are known, 62 in the Afrotropical region. World genera. Nine genera are known, three in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE842B63FF3ED46BFBC2F853.taxon	discussion	Remarks. Haliday is considered to be the author of Dryininae based on the principle of coordination (Art. 36 ICZN; International Commission on Zoological Nomenclature 1999). The author of the family is also author of the nominate subfamily whose name is derived from the family name. This holds even though Kieffer was the first author to introduce the subfamily division. The following five genera have not been recorded from the Afrotropical region: Cretodryinus Ponomarenko 1975 (fossil in amber from Russia, Taimyr Peninsula), Harpactosphecion Haupt 1944 (fossil in amber from the Dominican Republic and Baltic area), Hybristodryinus Engel 2005 (fossil in Myanmar amber), Megadryinus Richards 1953 (Neotropical), Palaeodryinus Olmi & Bechly 2001 (fossil in Baltic amber).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	synonymic_list	Dryinus Latreille 1804: 176; 1805: 228; Haliday 1833: 273; Kieffer in Kieffer & Marshall 1905: 69, partim; 1906: 497, partim; Richards 1935: 144; 1937: 118; 1939: 229; Olmi 1984: 728; 1991: 259; 1993 b: 61; 1994: 63; 1995 g: 5; 1999: 180; He & Xu 2002: 249; Virla & Olmi 2008: 370; Moya-Raygoza & Olmi 2010: 95; Lelej 2012: 397; Xu et al. 2013: 254; Olmi & Virla 2014: 214; Olmi & Xu 2015: 129; Olmi et al. 2016: 73. Campylonyx Westwood 1835: 52 (synonymized by Olmi 1984); type species: Campylonyx ampuliciformis Westwood 1835, by monotypy; Kieffer in Kieffer & Marshall 1905: 84; Kieffer 1914 b: 43. Chelothelius Reinhard 1863: 409 (synonymized by Olmi 1993 b); type species: Chelothelius gryps Reinhard 1863, by monotypy; Kieffer in Kieffer & Marshall 1905: 125; Kieffer 1914 b: 56; Berland 1928: 143; Olmi 1984: 604; 1993 b: 61; 1993 c: 47. Paradryinus R. Perkins 1905: 53 (synonymized by Olmi 1984); type species: Paradryinus koebelei R. Perkins, 1905, by original designation. Chlorodryinus R. Perkins 1905: 57 (synonymized by Olmi 1984); type species: Chlorodryinus pallidus R. Perkins 1905, by original designation. Plastodryinus Kieffer in Kieffer & Marshall 1906: 496 (synonymized by Móczár 1965); type species: Plastodryinus szepligetii (Kieffer in Kieffer & Marshall 1906), by original designation. Mesodryinus Kieffer in Kieffer & Marshall 1906: 497 (synonymized by Olmi 1993 b); type species: Mesodryinus niger (Kieffer 1904), by original designation; Kieffer 1914 b: 29; Muesebeck & Walkley 1951: 1034; Krombein 1979: 1243; Olmi 1984: 1000; 1993 b: 61; 1993 c: 47. Hesperodryinus R. Perkins 1907: 40 (synonymized by Olmi 1984); type species: Hesperodryinus arizonicus R. Perkins 1907, by original designation; Krombein 1979: 1242. Perodryinus R. Perkins 1907: 43 (synonymized by Olmi 1993 b); type species: Perodryinus amoenus R. Perkins 1907, by monotypy; Kieffer 1914 b: 48; Olmi 1984: 1033; 1993 b: 61. Lestodryinus Kieffer 1911 a: 108 (new name for Dryinus Latreille, 1804; synonymized by Richards 1937); type species: Lestodryinus collaris (Linnaeus 1767), by automatic designation (see Dryinus Latreille 1804); Kieffer 1913 a: 189; 1914 a: 90; 1914 b: 20. Tridryinus Kieffer 1913 c: 325 (synonymized by Olmi 1993 b); type species: Tridryinus striaticeps (Kieffer 1909), by original designation; Olmi 1984: 932; 1993 b: 61. Bocchoides Benoit 1954: 424 (synonymized by Olmi 1993 b); type species: Bocchoides bekilyensis Benoit 1954, by monotypy; Olmi 1984: 601; 1993 b: 61; 1993 c: 47. Richardsidryinus Móczár 1965: 376 (synonymized by Olmi 1991); type species: Richardsidryinus corsicus (Marshall 1874), by original designation; Olmi 1984: 907; 1991: 259. Avodryinus Ponomarenko 1981: 143 (synonymized by Olmi 1984, as synonym of Richardsidryinus Móczár 1965); type species: Avodryinus canadensis Ponomarenko 1981, by monotypy and original designation. Alphadryinus Olmi 1984: 973 (synonymized by Olmi 1993 b); type species: Alphadryinus asiaticus Olmi 1984, by original designation; Olmi 1990 a: 137; 1993 b: 61; 1993 c: 47.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	diagnosis	Diagnosis. ♀ (Figs 117 B, C); fully winged (Figs 117 B, C); occipital carina complete, or incomplete, or absent; palpal formula 6 / 3; mandible with 1 – 4 teeth; antenna usually with ADOs, rarely without; antenna without tufts of long setae (Figs 117 B, C); antennomere 3 usually less than 5 × as long as 2, occasionally more than 5 × (in this case, notauli often complete, then mesoscutum completely sculptured by numerous and parallel longitudinal keels); pronotal tubercle reaching or not reaching tegula; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 117 B, C); chela with rudimentary claw (Fig. 126 F); protarsomere 5 less than twice as broad as enlarged claw (Fig. 119 E); enlarged claw as long as, or shorter than protibia; tibial spurs 1 / 1 / 2, rarely 1 / 1 / 1. ♂ (Figs 117 D, 118 A): fully winged (Figs 117 D, 118 A); occipital carina complete or incomplete; mandible with 1 – 3 teeth; palpal formula 6 / 3; epicnemium visible, because lateral regions of prothorax not continuous with mesopleura (Fig. 7 D); mesosternum fused with mesopleura and not distinct; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 117 D, 118 A); paramere without dorsal process (Fig. 119 D); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	biology_ecology	Hosts. Acanaloniidae, Cixiidae, Dictyopharidae, Flatidae, Fulgoridae, Issidae, Lophopidae, Ricaniidae, Tropiduchidae (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	distribution	World species. 295 species are known, 59 in the Afrotropical region, including two species incertae sedis.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE822B66FF3ED010FE00F90D.taxon	discussion	Remarks. Larvae of Dryinus were described by Abril-Ramírez (1992) and Guglielmino et al. (2015). Fossil species of Dryinus were described by Ponomarenko (1981), Olmi (1995 e), Olmi et al. (2011), Olmi & Guglielmino (2011), Guglielmino et al. (2018) and Perkovsky & Olmi (2018). Dryinus ♀♀ were divided for convenience by Olmi (1993 b) into four groups on the basis of the characters detailed in the following key:	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9E2B7BFF3ED15BFBE8FD2C.taxon	description	(Figs 118 B, C, 119 B – E, 120 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9E2B7BFF3ED15BFBE8FD2C.taxon	description	Description. ♀ (Figs 118 B, C). Fully winged; body length 5.0 – 6.4 mm. Head black, except mandible, clypeus and anterior region of frons (more along orbits) testaceous; antenna brown, except scape and pedicel yellow; mesosoma black, except lateral regions of pronotum and posterior collar of pronotum reddish; metasoma brown; legs brown, except trocanters and part of procoxa yellow. In specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, head brown, except mandible, clypeus and anterior region of frons testaceous-brown; antenna testaceous; mesosoma brown, except lateral regions of pronotum and epicnemium brown-testaceous; metasoma brown; legs brown-testaceous. Head (Fig. 119 B) dull, granulate; occipital carina complete; lateral ocelli situated in front of virtual straight line joining posterior edges of eyes. Pronotum (Fig. 119 C) dull, granulate, with numerous longitudinal keels on lateral regions and on posterior transverse furrow; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow (in specimen from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, anterior transverse impression deeper than in type specimens); posterior collar long, about as long as distance from tegula to pronotal tubercle; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with numerous short longitudinal keels on posterior third. Notauli hardly visible, reduced to tracks, incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate and reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two brown transverse bands; distal part of 2 r-rs & Rs vein rectilinear, much longer than proximal part (18: 8). Protarsomere 1 about as long as 4. Enlarged claw (Fig. 119 E) with one large subdistal tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 119 E) with two rows of about 38 lamellae; apex with approximately 13 lamellae. Tibial spurs 1 / 1 / 2. ♂ (Figs 120 A, B). Fully winged; body length 2.7 – 3.1 mm. Head black, except mandible yellow; antenna yel-low-brown; mesosoma black; metasoma brown; legs brown, except tarsi and pro- and mesotibia testaceous. Head dull, completely reticulate rugose; frons flat in front of anterior ocellus and humped between antennal toruli; oc- cipital carina incomplete; POL = 6; OL = 3; OOL = 4; OPL = 1; temple absent. Mesoscutum dull, reticulate rugose, setose. Notauli hardly visible, incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate and punctate. Metapectal-propodeal disc and propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part; 2 R 1 cell closed. Genitalia in Fig. 119 D. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9E2B7BFF3ED15BFBE8FD2C.taxon	materials_examined	Material examined. Types: ♀ holotype: SÃO TOMÉ AND PRINCIPE: São Tomé Island, Canavial, Derron leg. (AMNH). Paratypes: same locality label as holotype, 3 ♂♂ (AMNH). Other material: DEMOCRATIC RE- PUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 2. IV. 2006, MT, forest edge, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9E2B7BFF3ED15BFBE8FD2C.taxon	biology_ecology	Hosts. Ricaniidae (Guglielmino et al. 2013): in São Tomé Island: Ricania insularis Lallemand. Natural enemies. In São Tomé Island this species may be parasitized by Helegonatopus saotomensis Prinsloo 1979 (Hymenoptera, Encyrtidae) (Prinsloo 1979).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9E2B7BFF3ED15BFBE8FD2C.taxon	distribution	Distribution. Democratic Republic of the Congo, São Tomé and Principe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7CFF3ED6D7FC9BFAAE.taxon	description	(Figs 121 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7CFF3ED6D7FC9BFAAE.taxon	description	Description. ♀. Fully winged; body length 4.5 – 5.0 mm. Head ferruginous, except one elongate brown spot on ocellar region and partly on frons (in specimen from Congo, 01 ° 56 ’ 49.49 ’’ N 30 ° 02 ’ 19.58 ’’ E, elongate spot on ocellar triangle and along frontal line); ventral side of head ferruginous, with brown spot (in specimen from Congo, 01 ° 56 ’ 49.49 ’’ N 30 ° 02 ’ 19.58 ’’ E, ventral side completely ferruginous); antenna testaceous; propleuron black, with dorsal reddish spot; pronotum ferruginous; rest of mesosoma black; metasoma brown; legs testaceous, except meso- and metacoxa partly brown. Head (Fig. 121 A) excavated, dull, granulate, with temple very prominent and posterior margin of vertex excavated; frontal line complete; occipital carina complete; TL about 3 × as long as POL; OL slightly longer than POL; lateral ocelli touching occipital carina in holotype, almost touching in specimen from Congo, 01 ° 56 ’ 49.49 ’’ N 30 ° 02 ’ 19.58 ’’ E). Pronotum dull, granulate; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slight; posterior collar distinct; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with some areolae near posterior margin. Notauli incomplete, approximately reaching 0.75 × length of mesoscutum (in specimen from Congo, 01 ° 56 ’ 49.49 ’’ N 30 ° 02 ’ 19.58 ’’ E, notauli almost reaching posterior margin of mesoscutum). Mesoscutellum dull, granulate. Metanotum rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three brown transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (17: 4). Protarsomeres in following proportions: 20: 4: 6: 19: 30. Protarsomere 3 produced into hook. Enlarged claw (Fig. 121 B) with one large subdistal tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 121 B) with two rows of 24 – 30 lamellae; apex with approximately 10 – 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7CFF3ED6D7FC9BFAAE.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Rutsh- uru-Kilinga, 1. IV. 1936, L. Lippens leg. (MRAC). Paratype: CAMEROON: Littoral Region, Moungo Department, N’Kongsamba, pont du N’Kam, 1939, P. Lepesme, R. Paulian & A. Villiers leg., 1 ♀ (MNHN). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, VII. 2008, MT, T. Chapman leg., 1 ♀ (DEUCD); Ituri, Djugu Territory, Mongbwalu mining camp, Mr. Beguin’s house, 01 ° 56 ’ 49.49 ’’ N 30 ° 02 ’ 19.58 ’’ E, 75 YPT traps, 1 – 8. III. 2015, A. Gumovsky & C. Dhendro leg., 1 ♀ (SIZK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7CFF3ED6D7FC9BFAAE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7CFF3ED6D7FC9BFAAE.taxon	distribution	Distribution. Cameroon, Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7DFF3ED335FBFFFDE4.taxon	description	(Figs 120 C, D, 121 C, D, 123 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7DFF3ED335FBFFFDE4.taxon	description	Description. ♀ (Figs 120 C, D, 123 A – C). Fully winged; body length 4.8 – 7.5 mm. Head black, except mandible, clypeus and anterior region of frons (more along orbits) yellow or testaceous (clypeus with posterior half brown in specimen from Central African Republic, 3 º 02.01 ’ N 16 º 24.57 ’ E); antenna testaceous, except first two antennomeres yellow; propleuron black; pronotum black, except lateral margins and posterior collar reddish or testaceous; rest of mesosoma black (with brown nuances in specimen from Central African Republic, 3 º 02.01 ’ N 16 º 24.57 ’ E); metasoma yellow – brown; legs yellow-testaceous. Head (Figs 121 C, 123 B) shiny, unsculptured, except many slight striae on frons and vertex; occipital carina complete; lateral ocelli situated very close to occipital carina; POL = OL; OPL shorter than OL. Pronotum (Fig. 123 A) shiny, with many striae around disc; pronotum bare, except posterior collar very setose; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar hardly visible; posterior collar short, about 0.5 × as long as distance from tegula to pronotal tubercle (4: 9); pronotal tubercle not reaching tegula. Mesoscutum (Fig. 123 A) dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing (Fig. 123 C) with two dark transverse bands; distal part of 2 r-rs & Rs vein more than 3 × as long as proximal part (17: 5). Protarsomere 1 about as long as 4 (17: 16); protarsomere 3 produced into hook. Enlarged claw (Fig. 121 D) with one large subdistal tooth and one row of 11 lamellae (lamellae situated near subdistal tooth very long and similar to peg – like setae). Protarsomere 5 (Fig. 121 D) with two rows of about 30 lamellae; apex with approximately 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7DFF3ED335FBFFFDE4.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Congo Central Prov. (= Bas Congo Prov.), Mbanza-Ngungu (= Thysville), 30. XI. 1952, P. Basilewsky leg. (MRAC). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Parc National de Dzanga-Ndoki, Mabéa Bai, 21.4 km 53 º NE Bayanga, 3 º 02.01 ’ N 16 º 24.57 ’ E, 510 m, 7. V. 2001, Sweep, CAR 01 - S 99, Lowland Rainforest, marsh clearing, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7DFF3ED335FBFFFDE4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9B2B7DFF3ED335FBFFFDE4.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	description	(Figs 121 E, 122, 123 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head testaceous-ferruginous, except ocellar triangle brown (Fig. 123 D); mesosoma totally testaceous-ferruginous; head (dorsally viewed) with lateral ocelli located in front of imaginary straight line joining posterior edges of eyes; notauli almost complete, posteriorly separated; protarsomere 1 less than twice as long as 4; enlarged claw (Fig. 121 E) slightly shorter than protarsomere 5 and with one big subapical tooth.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	description	Description. ♀ (Figs 122, 123 D). Fully winged; body length 6.6 mm. Head testaceous-ferruginous, except ocellar triangle brown; antenna testaceous, except antennomere 6, distal half of antennomere 5 and proximal half of antennomere 7 black; mesosoma testaceous-ferruginous; petiole black; metasoma testaceous ferruginous; legs testaceous. Antenna clavate; antennomeres in following proportions: 13: 7: 56: 32: 25: 16: 11: 11: 10: 12. ADOs present on antennomeres 5 – 10. Head (Fig. 122 C) dull, granulate and slightly irregularly striate; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 12; OPL = 2; TL = 2; greatest breadth of lateral ocelli longer than OPL (3.5: 2). Head (dorsally viewed) with lateral ocelli located in front of imaginary straight line joining posterior edges of eyes. Pronotum shiny, unsculptured, crossed by slight anterior transverse impression and strong posterior transverse furrow (Fig. 122 B); anterior collar slightly visible; posterior collar long, with two long lateral longitudinal keels (reaching sides of disc) and few short medial longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum (Fig. 122 A) shiny, punctate and very slightly irregularly striate, with surface close to posterior margin reticulate rugose. Notauli almost complete, hardly visible among areolae located near posterior margin of mesoscutum. Notauli posteriorly separated. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, about twice as long as propodeal declivity; propodeal declivity reticulate rugose, without longitudinal keels. Forewing (Fig. 122 D) with two brown transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (27: 13). Protarsomeres in following proportions: 32: 4: 9: 26: 41. Protarsomere 3 produced into hook. Enlarged claw (Fig. 121 E) with one large subdistal tooth and one row of 16 lamellae. Protarsomere 5 (Fig. 121 E) with two rows of 51 lamellae; apex with approximately 24 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19457): MADAGASCAR: Fianarantsoa, Ranomafana Na- tional Park, Vohiparara, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 14 – 21. I. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 12 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE9A2B7FFF3ED126FA96FF7C.taxon	etymology	Etymology. The species is named bellicosus (Latin adjective; = bellicose) because of its aggressive aspect.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE982B71FF3ED33CFC3DFEC4.taxon	description	(Figs 125 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE982B71FF3ED33CFC3DFEC4.taxon	description	Description. ♀. Fully winged; body length 6.0 – 8.4 mm. Head black, except mandible and clypeus testaceous; occasionally clypeus black; antenna totally testaceous, or black, except antennomeres 8 – 10 testaceous; propleuron black; pronotum totally black or brown – reddish, occasionally black, except posterior collar brownish; rest of mesosoma black; metasoma brown; legs brown-reddish, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 13: 8: 52: 29: 21: 14: 11: 10: 9: 11. Head (Fig. 125 A) dull, granulate; occipital carina complete; lateral ocelli not touching occipital carina; POL = 2.5; OL = 2.5; OOL = 12; OPL = 1.5; temple absent. Pronotum (Fig. 125 B) dull, granulate, with many longitudinal keels on lateral regions; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slight; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, completely sculptured by many longitudinal keels. Notauli almost complete, almost reaching posterior margin of mesoscutum. Mesoscutellum and metanotum dull, with many longitudinal keels. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein curvilinear, much longer than proximal part (24: 10). Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 125 C) with one large subdistal tooth and one row of 13 lamellae. Protarsomere 5 (Fig. 125 C) with two rows of about 37 lamellae; apex with approximately 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE982B71FF3ED33CFC3DFEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Ituri Prov., Moto, L. Burgeon leg. (MRAC). Other material: CAMEROON: South Region, Ebolowa, Nkoemvon, MT, 1978, D. Jackson leg., 1 ♀ (NHMUK); same locality label, IX. 1979, 1 ♀ (NHMUK); same locality label, X. 1980, 1 ♀ (NHMUK); same locality label, 1 ♀ (AMNH). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Lomami, Upemba Na- tional Park, Kankunda, 1 ♀ (MRAC). NIGERIA: Osun State, Ife (= Ilé – Ifè), 1 ♀ (AIEC); Oyo State, Ibadan, Cocoa Research Institute, 3. VII. 1963, on cocoa, 2 ♀♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE982B71FF3ED33CFC3DFEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE982B71FF3ED33CFC3DFEC4.taxon	distribution	Distribution. Cameroon, Democratic Republic of the Congo, Nigeria.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	description	(Figs 124 A, B, 125 D-G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	description	Richardsidryinus cariniceps (Cameron): Olmi 1984: 918. Description. ♀ (Figs 124 A, B). Fully winged; body length 3.4 – 7.5 mm. Head ferruginous-testaceous; occasionally ventral side of head and central region of clypeus darkened; antenna brown, except antennomeres 9 – 10 yellow, distal part of antennomere 4 and proximal part of antennomere 5 whitish; propleuron and pronotum ferruginous-testaceous or brown-reddish; rest of mesosoma ferruginous, or brown-reddish, or completely black; occasionally mesosoma ferruginous, with metanotum and metapectal-propodeal disc and propodeal declivity black; petiole black; metasoma brown-reddish; legs brown-reddish. Head flat, granulate and slightly rugose or with irregular longitudinal keels; posterior margin of vertex convex; occipital carina complete, hardly visible on sides of occiput; OPL slightly shorter than OL; POL = OL. Pronotum (Fig. 125 D) dull, granulate, with slight keels around disc; pronotum crossed by strong posterior transverse furrow (anterior transverse impression absent); posterior collar long, longer than length of ocellar region; pronotal tubercle not reaching tegula. Mesoscutum dull, reticulate rugose, or sculptured by numerous irregular and longitudinal keels. Notauli slightly visible, incomplete, reaching approximately 0.5 – 0.7 × length of mesoscutum. Mesoscutellum dull, reticulate rugose. Metanotum dull, punctate, or rugose. Metapectal-propodeal disc (Fig. 125 E) reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein curvilinear, much longer than proximal part. Protarsomere 1 longer than 4 (15: 12); protarsomere 3 produced into hook. Enlarged claw (Fig. 125 F) with one large subdistal tooth and one row of 9 – 11 lamellae. Protarsomere 5 (Fig. 125 F) with two rows of about 17 – 37 lamellae; apex with at least 16 lamellae. Tibial spurs 1 / 1 / 1. ♂ (see remarks). Fully winged; body length 3.6 – 4.2 mm. Head black, except mandible, clypeus, malar space and palpi testaceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous, except meso- and metacoxa partly brown. Antenna filiform; antennomeres in following proportions: 8: 7: 21: 14: 15.5: 15: 13: 12: 10: 12. Head dull, convex, completely reticulate rugose; frontal line complete; occipital carina incomplete, absent in temple; POL = 9; OL = 3.5; OOL = 4.5; OPL = 2; TL = 0; greatest breadth of lateral ocelli slightly longer than OL (4: 3.5). Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, with strong irregular transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (11: 10); 2 R 1 cell open. Paramere with large dorsal process (Fig. 125 G). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns (NHMUK). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele (= Haut-Zaire), Garamba National Park, 2 ♀♀ (MRAC); same locality label, 1 ♀ (AMNH). GABON: La Makaude, 10. III. 1999, on canopée, 1 ♀ (MOLC). IVORY COAST: Lagunes District, Dabou, 25. VII. 1963, 1 ♀ (MNHN). KENYA: Coast Prov., Funzi Island, 4.53577 S 39.46042 ’ E, near Mliani, near sea level, by sweeping, 6. VII. 2012, ICIPE / NMK, Funzi Island Expedition leg., 1 ♀ (ICIPE). SOUTH AFRICA: Eastern Cape, same locality label as holotype, 15 – 31. V. 1923, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 3 ♀♀ (MRAC). YEMEN: 60 km E Aden, Al-Kowd, 13 ° 05 ’ N 45 ° 22 ’ E, 17 – 21. VII. 2001, LT, A. van Harten & S. Al Haruri leg., 1 ♂ (MOLC); same locality label, IV. 1993, MT, A. van Harten leg., 1 ♀ (MOLC); 12 km NW of Manakhah, 3. VII – 21. VIII. 2001, MT, A. van Harten leg., 4 ♂♂ (MOLC); same locality label, 15. V – 24. VI. 2003, N. 7770, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	distribution	Distribution. Democratic Republic of the Congo, Gabon, Ivory Coast, Kenya, South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE962B72FF3ED7DBFE70FEE1.taxon	discussion	Remarks. The ♂ above described was attributed doubtfully to Dryinus cariniceps Cameron, because a ♀ and a ♂ specimen of this species were both collected from the same locality (Yemen, Al-Kowd). However, in a further locality of Yemen (12 km NW of Manakhah) two ♀ specimens of Dryinus yemenensis Olmi & van Harten were collected by a Malaise trap together with 4 ♂ specimens similar to those collected in Al-Kowd. There is the pos- sibility that these ♂♂ are the opposite sex of Dryinus yemenensis. Only rearings of paired sexes, or DNA analysis will resolve this issue.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE952B73FF3ED3ADFE14FC04.taxon	description	(Fig. 126 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE952B73FF3ED3ADFE14FC04.taxon	description	Description. ♀. Fully winged; body length 6.0 – 6.2 mm. Holotype with head reddish, except brown spot on frons; antenna testaceous; propleuron black; mesosoma brown-reddish, except metapectal-propodeal disc and propodeal declivity black; metasoma brown-reddish; legs ferruginous. In specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, head brown, except mandible, clypeus and anterior region of frons ferruginous; antenna testaceous; propleuron, pronotum and mesoscutum brown, with ferruginous nuances; rest of mesosoma black; metasoma brown; legs brown, except tarsi, articulations and part of procoxa testaceous. Antenna clavate; antennomeres in following proportions: 10: 5: 48: 27: 22: 15: 12: 10: 8: 10. Head shiny, slightly punctate (in specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, punctate and slightly granulate), with many irregular longitudinal keels on frons; frontal line complete; occipital carina complete; lateral ocelli not touching occipital carina; POL = 1; OL = 1; OOL = 13; OPL = 1; TL = 6; greatest breadth of lateral ocelli longer than OPL (3: 1). Pronotum shiny, unsculptured, except lateral regions sculptured by numerous longitudinal keels; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slight; posterior collar long; disc humped; pronotal tubercle not reaching tegula. Mesoscutum shiny, punctate, unsculptured among punctures in holotype, slightly granulate among punctures in specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E. Notauli incomplete, approximately reaching 0.65 – 0.70 × length of mesoscutum. Mesoscutellum shiny, punctate, with numerous curvilinear longitudinal keels. Metanotum punctate and rugose. Metapectal-propodeal disc reticulate rugose, about twice as long as propodeal declivity; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein about twice as long as proximal part. Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 126 A) with one large subdistal tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 126 A) with two rows of about 42 lamellae; apex with approximately 12 – 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown. Benoit (1954) described a ♂ allotype; however it was not obtained by rearing (Benoit 1954), so that it is not sure that it is the opposite sex of the described ♀ (Olmi 1984).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE952B73FF3ED3ADFE14FC04.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Behara, II. 1938, A. Seyrig leg. (MNHN). Paratypes: MADAGASCAR: Toamasina, Ivondro, III. 1940, A. Seyrig leg., 1 ♀ (MRAC); samel locality label, II. 1940, 1 ♀ (MNHN); Toliara, same locality label as holotype, III. 1937, II. 1938, III. 1939, 8 ♀♀ (5 in MNHN, 3 in MRAC); Toliara, Bekily, IV. 1937, A. Seyrig leg., 1 ♂ allotype (MNHN); same locality label, IV. 1937, 1 ♀ (MNHN); same locality label, X. 1936, IV. 1942, 2 ♀♀ (MRAC). Other material: MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 16 – 17. XII. 2002, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA – 02 – 20 – 09, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE952B73FF3ED3ADFE14FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE952B73FF3ED3ADFE14FC04.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE932B75FF3ED001FE59FCBC.taxon	description	(Figs 123 E, F, 126 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE932B75FF3ED001FE59FCBC.taxon	description	Description. ♀ (Figs 123 E, F). Fully winged; body length 3.9 – 5.2 mm. Head black, except mandible, clypeus and part of malar space brown or testaceous; occasionally frons with testaceous stripe along orbits; antenna brown, except antennomeres 7 – 10 testaceous-whitish; mesosoma black; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 11: 5: 25: 9: 6: 6: 6: 6: 5: 9. Head dull, granulate and reticulate rugose (with areolae very small, so that superficially the frons seems granulate), occasionally with some irregular keels on frons; occipital carina complete; occiput concave; temple prominent; frontal line complete; POL = 6; OL = 4; OOL = 8; OPL = 2; TL = 1; greatest breadth of lateral ocelli longer than OPL. Occasionally POL twice as long as OL. Pronotum dull, crossed by two weak transverse impressions; anterior collar granulate; lateral regions granulate and partly reticulate rugose; disc granulate; posterior collar very short. Mesoscutum dull, granulate and reticulate rugose. Notauli complete, posteriorly separated (occasionally notauli hardly visible among roughness near posterior margin of mesoscutum); minimum distance between notauli longer than antennomere 2 (7: 5). Mesoscutellum dull, granulate. Metanotum shiny, granulate. Metapectal-propodeal disc dull, reticulate rugose, slightly longer than propodeal declivity (22: 20), without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (17: 11). Protarsomeres in following proportions: 18: 3: 5: 15: 24. Enlarged claw (Fig. 126 B) with one small subdistal tooth and one row of five peg-like setae. Protarsomere 5 (Fig. 126 B) with one row of 13 – 17 lamellae; apex with about 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE932B75FF3ED001FE59FCBC.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Arabuko-Sokoke Forest, 03 ° 25.21 ’ S 39 ° 53.81 ’ E, 25 – 29. VI. 1999, MT, R. Copeland leg. (NMK). Other material: KENYA: Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 12 – 26. VII. 2014, MT, indigenous forest, R. Copeland leg., 2 ♀♀ (NMK, MOLC); same locality label, 27. XII. 2015 – 9. I. 2016, 1 ♀ (NMK); Coast Prov., Buda Forest, 4.46109 ° S 39.40446 ° E, 98 m, 12 – 26. II. 2016, MT, indigenous forest, R. Copeland leg., 1 ♀ (MOLC); same locality label, 20. V – 3. VI. 2016, 1 ♀ (NMK); same lo- cality label, 26. II – 11. III. 2016, 1 ♀ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE932B75FF3ED001FE59FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE932B75FF3ED001FE59FCBC.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B75FF3ED5CCFE1FF89C.taxon	description	(Fig. 126 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B75FF3ED5CCFE1FF89C.taxon	description	Description. ♀. Fully winged; body length 4.1 mm. Head black, except mandible testaceous; clypeus black, except lateral regions testaceous; antenna brown-testaceous, except antennomere 10 testaceous; mesosoma and metasoma black; legs brown, except coxae and clubs of femora black. Antenna clavate; antennomeres in following proportions: 13: 6: 26: 12: 9: 8: 6.5: 6: 5.5: 8; ADOs on antennomeres 5 – 10. Head dull, granulate, with few longitudinal and irregular keels on frons; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 9; OPL = 1; TL = 3; greatest breadth of lateral ocelli longer than OPL (2: 1); lateral ocelli situated on virtual straight line joining posterior edges of eyes; posterior margin of vertex excavated. Pronotum shiny, slightly granulate, with numerous slight striae on lateral regions; pronotum crossed by strong posterior transverse furrow and slight anterior transverse impression; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum dull, strongly reticulate rugose, except anterior third of median region shiny, sculptured by few slight irregular striae. Notauli incomplete, reaching about 0.6 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose, about as long as propodeal declivity, without longitudinal or transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 6). Protarsomeres in following proportions: 17: 3: 8: 18: 28. Protarsomere 3 produced into hook. Enlarged claw (Fig. 126 C) with one large subapical tooth and one row of 10 lamellae. Protarsomere 5 (Fig. 126 C) with two rows of 9 + 17 lamellae; apex with about 18 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B75FF3ED5CCFE1FF89C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Ngoye Forest, I – IV. 2006, MT, G. Davies leg. (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B75FF3ED5CCFE1FF89C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B75FF3ED5CCFE1FF89C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B76FF3ED12CFE14FCD8.taxon	description	(Fig. 126 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B76FF3ED12CFE14FCD8.taxon	description	Description. ♀. Fully winged; body length 8.0 mm. Reddish, except metasoma and petiole black; antenna brown, except scape, pedicel and flagellomeres 5 – 8 testaceous; legs reddish, except black spots on coxae, femora and tibiae. Antenna not distally thickened; antennomeres in following proportions: 11: 6: 52: 29: 24: 17: 13: 11: 9: 11. Head flat, shiny, granulate and with numerous irregular keels on frons and on vertex around ocellar triangle; occipital carina complete; frontal line complete; POL = 2; OL = 2; OOL = 12; OPL = 1.5; TL = 7. Pronotum shiny, crossed by slight anterior transverse impression and strong posterior transverse furrow; disc humped and unsculptured; lateral regions of pronotum granulate and sculptured by irregular striae; posterior collar granulate; pronotal tubercle not reaching tegula. Mesoscutum shiny, punctate, unsculptured among punctures, with numerous short keels near posterior margin. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Mesoscutellum shiny, sculptured by irregular longitudinal keels. Metanotum shiny, rugose, with posterior half smooth. Metapectal-propodeal disc dull, reticulate rugose, more than twice as long as propodeal declivity (42: 16); propodeal declivity reticulate rugose, with two short longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (20: 7). Protarsomeres in following proportions: 31: 5: 9: 26: 40. Enlarged claw (Fig. 126 D) with one large subdistal tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 126 D) with two rows of about 32 lamellae; apex with approximately 14 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B76FF3ED12CFE14FCD8.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Beroboka, 60 km NE Morondava, 18 – 23. V. 1983, J. S. Noyes & M. C. Day leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B76FF3ED12CFE14FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE922B76FF3ED12CFE14FCD8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE912B77FF3ED5FFFD5BFE54.taxon	description	(Figs 125 H, 126 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE912B77FF3ED5FFFD5BFE54.taxon	description	Description. ♀. Fully winged; body length 6.0 – 6.6 mm. Yellow – testaceous, except petiole black and metasoma brown – reddish. In specimen from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, distal extremity of metasoma black. Occasionally ocellar triangle brown and metasoma totally yellow-testaceous. Antenna clavate; antennomeres in following proportions: 6: 3: 28: 17: 12: 8: 6: 5: 5: 5. Head (Fig. 125 H) flat, dull, granulate; frontal line complete; occipital carina complete; OPL shorter than POL; OL = POL; temple short. Pronotum dull, granulate, with numerous keels on lateral regions; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum shiny, granulate, occasionally with few irregular keels (in specimen from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, mesoscutum slightly rugose). Notauli incomplete, approximately reaching 0.5 – 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, granulate. Metapectal-propodeal disc slightly reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. In specimen from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, mesoscutellum rugose. Forewing with two brown transverse bands (in specimen from South Africa, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, three brown transverse bands); distal part of 2 r-rs & Rs vein much longer than proximal part. Protarsomere 1 shorter than 4 (18: 22); protarsomeres 2 and 3 produced into hook. Enlarged claw (Fig. 126 E) with one large subdistal tooth and one row of 12 – 15 lamellae. Protarsomere 5 (Fig. 126 E) with two rows of 34 – 37 lamellae; apex with approximately 19 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.4 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous, except coxae, clubs of femora, mid and hind tibiae brown. Antenna filiform; antennomeres in following proportions: 5: 3: 11: 7: 6: 6: 6: 6: 5: 7. Head dull, granulate, with slight tracks of areolae; frontal line absent; frons with median longitudinal hump from anterior ocellus to antennal toruli; occipital carina incomplete; POL = 5.6; OL = 2; OOL = 4; OPL = 1; TL = 0. Mesoscutum, mesoscutellum and metanotum dull, granulate, with slight tracks of areolae. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part; 2 R 1 cell closed. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE912B77FF3ED5FFFD5BFE54.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 6 – 25. II. 1924, R. E. Turner leg. (NHMUK). Paratype: same locality label as holotype, XII. 1923, 1 ♀ (NHMUK). Other material: KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 24. IV – 10. V. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: KwaZulu-Natal, La Mercy, Site 1, 29 ° 37.41 ’ S 31 ° 06.454 ’ E, 85 m, 6. XII. 2006, YPT, Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, YPT, Maputaland Coastal Belt, SA 10 - MAB-Y 01, 1 ♀ (SAMC). UGANDA: Kampala (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE912B77FF3ED5FFFD5BFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE912B77FF3ED5FFFD5BFE54.taxon	distribution	Distribution. Kenya, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B77FF3ED46BFE59F981.taxon	description	(Figs 124 C, 126 F, 129 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B77FF3ED46BFE59F981.taxon	description	Description. ♀ (Figs 124 C, 129 A). Fully winged; body length 5.9 mm. Head black, except mandible testaceous, clypeus and malar space ferruginous-dark and testaceous strie along orbits; antenna brown, except antennomeres 7 – 10 and distal region of 6 whitish; mesosoma black; metasoma brown; legs brown, except tarsi testaceous-dark. Antenna clavate; antennomeres in following proportions: 12: 6: 29: 10: 9: 8: 6: 7: 7: 10. ADOs present in antennomeres 5 – 10. Head convex, dull, granulate, with some hardly visible irregular keels on frons; frontal line absent; occipital carina incomplete, only present behind and on sides of lateral ocelli, laterally not reaching eyes; lateral ocelli situated in front of virtual straight line joining posterior edges of eyes; posterior margin of vertex weakly excavated; POL = 5; OL = 4; OOL = 8; OPL = 3; temple absent; greatest breadth of posterior ocellus about as long as OPL. Pronotum crossed by strong anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar very short; pronotum dull, granulate and sculptured by many longitudinal keels and striae; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose, with areolae very small, larger near posterior margin. Notauli apparently complete and posteriorly separated, hardly visible among areolae situated near posterior margin of mesoscutum; minimum distance between notauli longer than POL (8: 5). Mesoscutellum and metanotum granulate. Metapectal-propodeal disc dull, reticulate rugose, slightly longer than propodeal declivity (24: 22), without transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (20: 11) Protarsomeres in following proportions: 21: 3: 6: 18: 26. Enlarged claw (Fig. 126 F) with one large subdistal tooth and one row of 5 very slender lamellae. Protarsomere 5 (Fig. 126 F) with one row of 21 lamellae; apex with about 11 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B77FF3ED46BFE59F981.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Kaya Kinondo, 4.39382 ° S 39. 54567 ° E, 10 m, 25. XII. 2011 – 8. I. 2012, indigenous forest, MT, coral rag, canopy forest, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B77FF3ED46BFE59F981.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B77FF3ED46BFE59F981.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	description	(Figs 127, 128 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head mostly black; frons sculptured by many irregular longitudinal striae; lateral ocelli not touching occipital carina; POL approximately twice as long as OL; temple absent; head (dorsally viewed) with lateral ocelli located in front of imaginary straight line joining posterior edges of eyes; mesosoma black; mesoscutum shiny, with lateral areas sculptured by many subparallel longitudinal keels, and median area medially punctate, unsculptured among punctures; forewing with distal part of 2 r-rs & Rs vein less than 3 × as long as proximal part; metapectal-propodeal disc short, about as long as propodeal declivity; protarsomere 1 less than twice as long as 4; enlarged claw slightly shorter than protarsomere 5, with one big subapical tooth big (Fig. 128 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	description	Description. ♀ (Figs 127 A, B). Fully winged; body length 6.0 mm. Head black, except mandible and ante- rior margin of clypeus testaceous-yellow; antenna brown, except scape, pedicel and flagellomeres 4 – 8 testaceouswhitish; mesosoma black; metasoma brown; proleg brown, except part of coxa and trochanter testaceous-whitish; meso- and metaleg brown, except distal extremity of coxae, trochanters, proximal extremity of tibiae, tarsomere 2 and distal half of tarsomere 1 testaceous-whitish. Antenna clavate; antennomeres in following proportions: 10: 5: 3 5: 20: 14: 10: 8: 8: 8: 12. ADOs present in antennomeres 5 – 10. Head (Fig. 127 C) weakly excavated, shiny, with frons sculptured by many irregular longitudinal striae, unsculptured among striae; occipital carina incomplete, only present behind ocellar triangle and on sides of lateral ocelli, laterally not reaching eyes, reaching about 0.7 × length of OOL; occiput little concave; temple absent; frontal line complete; POL = 4; OL = 2; OOL = 7; OPL = 1; greatest breadth of lateral ocelli longer than OPL (4: 1). Pronotum shiny, crossed by two weak transverse impressions; anterior collar and disc punctate, unsculptured among punctures; lateral regions sculptured by many longitudinal striae, unsculptured among striae; posterior collar very short; pronotal tubercle not reaching tegula. Mesoscutum (Fig. 127 A) shiny, with lateral areas sculptured by many subparallel longitudinal keels; median area medially punctate, unsculptured among punctures, laterally sculptured by longitudinal keels. Notauli incomplete, reaching about 0.8 × length of mesoscutum, hardly visible near posterior margin of mesoscutum. Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum shiny, with anterior half rugose and posterior half unsculptured. Metapectal-propodeal disc dull, reticulate rugose, about as long as propodeal declivity (Fig. 127 B), without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing (Fig. 127 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (20: 12). Hind wing with distal third darkened. Protarsomeres in following proportions: 22: 4: 7: 17: 28. Enlarged claw (Fig. 128 A) with one large subdistal tooth and one row of nine lamellae. Protarsomere 5 (Fig. 128 A) with two rows of 19 lamellae; apex with about 12 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	materials_examined	Material examined. Type: ♀ holotype, DAF 2200: IVORY COAST: Taï National Park, UTM 29 N 0682666 0.646862, III. 2014, MT, Erena Dupont leg. (DAFS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	distribution	Distribution. Ivory Coast.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE902B09FF3ED016FC0CF8A6.taxon	etymology	Etymology. The species is named after the collector, Ms Erena Dupont.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEE2B0BFF3ED13CFCECFB94.taxon	description	(Figs 128 B – G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEE2B0BFF3ED13CFCECFB94.taxon	description	Description. ♀. Fully winged; body length 3.4 – 6.0 mm. Head black, except mandible and clypeus yellow (in small specimen (3.4 mm) from Kenya, 3.82080 ° S 38.64178 ° E, head black, except mandible, clypeus and malar space testaceous); antenna testaceous, except antennomeres 7 – 9 or 3 – 7 slightly darkened (in specimen from Uganda, 0 ° 33.823 ’ N 30 ° 21.490 ’ E, antenna brown, except scape, pedicel and flagellomere 8 testaceous); propleuron black; pronotum black, with anterior margin, lateral regions and posterior collar reddish; mesosoma and petiole black; metasoma brown; legs testaceous-reddish, some × with coxae darkened. Antenna clavate; antennomeres in following proportions: 10: 5: 15: 8: 6: 5: 4.5: 4: 4: 6. Head shiny, granulate, occasionally with some longitudinal keels on frons (in specimen from Kenya, 1.22328 ° S 36.89674 ° E, frons granulated and partly rugose); occipital carina complete; POL = 3.5; OL = 3; OOL = 7; OPL = 2; TL = 7. In specimens from South Africa, 33 ° 16.84 ’ S 24 ° 22.78 ’ E, and Kenya, 0.97984 ° N 37.34599 ° E, lateral ocelli very close to occipital carina (almost touching it). Pronotum (Fig. 128 B) shiny, granulate, with longitudinal keels on lateral regions and on posterior transverse impression (in small specimens, above keels usually hardy visible or absent; in specimens from Namibia, 21 ° 09.00 ’ S 14 ° 34.51 ’ E, pronotum shiny and unsculptured, except longitudinal striae on lateral regions); pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow. Mesoscutum dull, as sculptured as head, with posterior half reticulate rugose (occasionally in small specimens, mesoscutum completely rugose; in small specimen (3.4 mm) from Kenya, 3.82080 ° S 38.64178 ° E, mesoscutum granulate, except some areolae near posterior margin). Notauli incomplete, approximately reaching 0.5 – 0.7 × length of mesoscutum. In small specimen (3.4 mm) from Kenya, 3.82080 ° S 38.64178 ° E, notauli reaching about 0.9 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum rugose. Metapectal-propodeal disc (Fig. 128 G) reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three distinct brown transverse bands (rarely transverse bands hardly visible); distal part of 2 rrs & Rs vein usually less than 3 × as long as proximal part (rarely distal part 3 × as long as proximal part). Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. In small specimen (3.4 mm) from Kenya, 3.82080 ° S 38.64178 ° E, protarsomere 1 about twice as long as 4 (14: 7.5). Enlarged claw with one small (Fig. 128 C) or large (Fig. 128 D) subdistal tooth and one row of 6 – 11 lamellae. Protarsomere 5 (Figs 128 C, D) with two rows of 19 – 24 lamellae; apex with approximately 16 lamellae. Tibial spurs 1 / 1 / 1 (in holotype), or 1 / 1 / 2. ♂. Fully winged; body length 2.2 – 3.5 mm. Head black, except mandible yellow; antenna brown-yellow; meso- soma black; metasoma brown or testaceous; legs testaceous, except metacoxa partly brown. Antenna filiform; antennomeres in following proportions: 5: 3: 11: 7: 6: 6: 6: 6: 5: 7. Head dull, granulate; occipital carina incomplete; temple absent; lateral ocelli almost touching occipital carina; POL = 7; OL = 2.5; OOL = 4; OPL = 1; TL = 3; greatest breadth of lateral ocelli longer than OPL (3: 1). Mesoscutum dull, granulate, with slight keels near posterior margin. Notauli incomplete, reaching approximately 0.65 – 0.75 × length of mesoscutum. Mesoscutellum and metanotum granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (12: 9). Genitalia in Fig. 128 E. Hypopygium in Fig. 128 F. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEE2B0BFF3ED13CFCECFB94.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, XI. 1922, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, X. 1921, 1 ♀ (NHMUK); same locality label as holotype, 18 – 30. XI. 1921, 1 ♀ (NHMUK); same locality label as holotype, VI – VII. 1930, 1 ♂ allotype (not valid because des- ignated by Benoit (1951 b) after the original description) (NHMUK). Other material: ANGOLA: Namibe Province, 10 mi. NE Namibe (= Moçamedes), 27 – 29. II. 1972, R. Giraul leg., 1 ♀ (NHMUK). DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Mt. Hoyo, 1 ♂ (MRAC). ER- ITREA: Southern Red Sea Region, Asseb (= Assab), 1 ♀ (HNHM). KENYA: Coast Prov., Kasigau Mtn., 3.82080 ° S 38.64178 ° E, 737 m, bottom of forest, MT, woodland with grass, 28. XII. 2011 – 11. I. 2012, R. Copeland leg., 1 ♀ (RSC); Nairobi Prov., ICIPE Campus, Kasarani, 1.22328 ° S 36.89674 ° E, 1613 m, 17 – 24. II. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Marich Pass field station, 1.53633 ° N 35.45800 ° E, 917 m, 20. III – 4. IV. 2005, MT, low canopy riverine forest, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Nguruman, near Sampu River, 1.90117 ° S 36.05040 ° E, 723 m, 27. VIII – 10. IX. 2011, MT, near base of Nguruman Escarpment, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Matthews Range, 0.97984 ° N 37.34599 ° E, 1459 m, 19. IV – 3. V. 2016, MT, riverine forest, near Wamba, R. Copeland leg., 1 ♀ (NMK). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 21 – 28. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA – 01 – 01 – 01, 1 ♀ (CAS); Mahajanga, Maintirano District, Asondrodava dry forest, 15 km N of Maintirano, 17 ° 57.92 ’ S 44 ° 2.13 ’ E, 200 ft., 10 – 17. XII. 2007, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG – 43 A – 101, 1 ♀ (CAS); same locality label, 14 – 21. I. 2008, MG – 43 A – 15, 1 ♀ (MOLC); Toliara, Andohahela Na- tional Park, Ihazofotsy Parcelle III, 24 ° 49.85 ’ S 46 ° 32.17 ’ E, 80 m, 20. V – 3. VI. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA – 02 – 21 – 20, 1 ♀ (CAS); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 50.09 ’ S 46 ° 29.21 ’ E, 80 m, 21 – 29. VI. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA – 02 – 21 – 23, 1 ♀ (CAS). NAMIBIA: Brandberg, Königstein, 21 ° 09.00 ’ S 14 ° 34.51 ’ E, 2470 m, 11 – 14. IV. 1999, NA 99 – M 08, MT, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg., 1 ♀ (SMWN); Brandberg, Wasserfallfläche, 21 ° 10.77 ’ S 14 ° 32.87 ’ E, 1960 m, 7 – 10. IV. 1999, NA 99 – M 06, MT, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg., 1 ♀ (SMWN). SOMALIA: Benadir, Afgoi, M. Olmi leg, 2 ♀♀ (AMNH). SOUTH AFRICA: Eastern Cape, 6 km N Steytlerville, 33 ° 16.84 ’ S 24 ° 22.78 ’ E, 500 m, 16 – 23. XI. 1999, M. E. Irwin et al. leg., SA – 05, 1 ♀ (CNC); KwaZulu-Natal: Mt. Edgecombe, 27. I. 1970, 5. III. 1970, A. Carnegie leg., 4 ♀♀, 4 ♂♂ (NHMUK); same locality label, many specimens of both sexes (SASAES, 9 in AMNH); KwaZulu-Natal, Pongola, 1 ♀, 1 ♂ (SASAES); Limpopo, Ellisras Distric, Mogol Nature Reserve, 1 ♀ (SANC). SWAZILAND: Big Bend, 1 ♀ (AMNH); Swaziland, 13. IX. 1965, reared from Numicia viridis Muir, C. I. E. 4382, no collector (NHMUK). TAN- ZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 29. I – 11. III. 1996, Acacia / Commiphora / Combretum bushland, MT, SAM – HYM – PO 15605, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 25. XII. 1995 – 29. I. 1996, Acacia / Commiphora / Combretum bushland, MT, SAM – HYM – PO 15906, S. van Noort leg., 2 ♀♀ (SAMC). UGANDA: Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.823 ’ N 30 ° 21.490 ’ E, 1505 m, 4 – 26. VIII. 2008, YPT, primary mid – altitude rainforest, S. van Noort leg., 1 ♀ (SAMC). ZIMBABWE: Harare (= Salisbury), Chishawasha, VIII. 1979, A. Watsham leg., 1 ♀ (NHMUK); Harare (= Salisbury), IV. 1975, 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEE2B0BFF3ED13CFCECFB94.taxon	biology_ecology	Hosts. Tropiduchidae (Guglielmino et al. 2013): in South Africa and Swaziland: Numicia viridis Muir.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEE2B0BFF3ED13CFCECFB94.taxon	distribution	Distribution. Angola, Democratic Republic of the Congo, Eritrea, Kenya, Madagascar, Namibia, Somalia, South Africa, Swaziland, Tanzania, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEC2B0BFF3ED22BFE13F8E9.taxon	description	(Fig. 132 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEC2B0BFF3ED22BFE13F8E9.taxon	description	Description. ♀. Fully winged; body length 4.4 mm. Testaceous – reddish, except mesothorax, metathorax, metapec- tal-propodeal disc and propodeal declivity black; metasoma brown. Antenna clavate; antennomeres in following proportions: 5: 5: 30: 12: 11: 7: 7: 6: 5: 7. Head flat, dull, granulate; occipital carina complete; frontal line complete; POL = 1.5; OL = 2; OOL = 10; OPL = 1; TL = 6. Pronotum dull, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum dull, completely granulate. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel, much longer than propodeal declivity (20: 15); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 7). Protarsomeres in following proportions: 17: 3: 5: 15: 22. Enlarged claw (Fig. 132 A) with one large subdistal tooth and one row of 11 lamellae. Protarsomere 5 (Fig. 132 A) with two rows of about 20 lamellae; apex with approximately 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEC2B0BFF3ED22BFE13F8E9.taxon	materials_examined	Material examined. Type: ♀ holotype: IVORY COAST: Vallée du Bandama, Katiola Dept., 20 km E Katiola, Logbonou, 21. XII. 1980, in savanna, MT, J. W. Everts leg. (DEUW).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEC2B0BFF3ED22BFE13F8E9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEC2B0BFF3ED22BFE13F8E9.taxon	distribution	Distribution. Ivory Coast.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEB2B0DFF3ED148FE14FC20.taxon	description	(Figs 129 B, 132 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEB2B0DFF3ED148FE14FC20.taxon	description	Description. ♀ (Fig. 129 B). Fully winged; body length 6.7 mm. Head black, except mandible testaceous; clypeus black, with lateral and anterior margins testaceous; antenna brown, except antennomeres 9 – 10 whitish; mesosoma black, with lateral margins of pronotum testaceous; metasoma black; legs brown, except inner side of protibia and femora testaceous-darkened and with protrochanter and procoxa partly testaceous and whitish. Antenna clavate; antennomeres in following proportions: 13: 7: 39: 17: 15: 13: 12: 10: 9: 12.5; ADOs present on antennomeres 5 – 10. Head flat, dull, granulate; anterior third of frons slightly rugose; ocellar triangle slightly rugose among ocelli; frontal line complete; occipital carina complete; POL = 3.5; OL = 3.5; OOL = 10; OPL = 2; TL = 2.5. Pronotum shiny, smooth, unsculptured, humped, crossed by weak anterior transverse impression and strong posterior transverse furrow; anterior collar long; posterior collar very short; posterior transverse furrow of pronotum crossed by short longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with areas near posterior margin reticulate rugose, lateral regions more strongly granulate and median region slightly granulate. Notauli almost complete and approximately reaching posterior margin of mesoscutum. Mesoscutellum shiny, slightly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel, slightly longer than propodeal declivity (13: 10); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two very broad dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (23: 11). Protarsomeres in following proportions: 26: 4: 10: 25: 39. Enlarged claw (Fig. 132 B) with one large subdistal tooth and one row of 17 lamellae. Protarsomere 5 (Fig. 132 B) with two rows of approximately 42 lamellae; apex with at least 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEB2B0DFF3ED148FE14FC20.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, 3 km 41 ° NE Andranomay, 11.5 km 147 ° SSE Anjozorobe, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, beating / sweeping, montane rainforest, Fisher, Griswold et al. leg., BLF 2544 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEB2B0DFF3ED148FE14FC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEB2B0DFF3ED148FE14FC20.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEA2B0EFF3ED02CFC63FC20.taxon	description	(Fig. 132 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEA2B0EFF3ED02CFC63FC20.taxon	description	Description. ♀. Fully winged; body length 4.2 – 6.0 mm. Head black, except mandible and clypeus testaceous; oc- casionally also malar space testaceous; antenna testaceous, except antennomeres 5 – 9 or 6 – 9 darkened; propleuron black; pronotum totally black, except posterior collar and lateral regions testaceous; rest of mesosoma black; metasoma partly black and partly testaceous; legs testaceous, occasionally except meso- and metatibia darkened and procoxa partly black. Antenna clavate; antennomeres of holotype in following proportions: 13: 6: 29: 13: 11: 10: 7: 6: 6: 8; in specimen from Botswana, Farmer’s Brigade, antennomeres in following proportions: 10: 4.5: 19: 8.5: 7: 6: 5.5: 5: 4.5: 7. Head dull, granulate and with some irregular keels on frons; occasionally head completely reticulate rugose; occipital carina complete; occiput concave; temple prominent; frontal line complete; holotype with POL = 6; OL = 4; OOL = 11; OPL = 1; TL = 6; in specimen from Botswana, Farmer’s Brigade, POL = 4.5; OL = 2.5; OOL = 7; OPL = 0.5; TL = 3; in specimen from Cameroon, Messabengondo, POL = 4; OL = 2. Pronotum shiny, crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar and lateral regions of pronotum unsculptured or with longitudinal striae; disc slightly or strongly granulate, sculptured by some striae; posterior collar very short. Mesoscutum dull, reticulate rugose, anteriorly smooth. Notauli incomplete, reaching approximately 0.5 – 0.6 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum rugose. Metapectal-propodeal disc dull, reticulate rugose, about as long as propodeal declivity; propodeal declivity reticulate rugose. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (12.6: 6). Protarsomeres of holotype in following proportions: 20: 5: 9: 24: 37; Protarsomeres of specimens from Botswana, Farmer’s Brigade, in following proportions: 14: 3: 6: 12: 21. Enlarged claw (Fig. 132 C) with one large subdistal tooth and one row of 7 – 15 lamellae. Protarsomere 5 (Fig. 132 C) with two rows of about 24 – 28 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 1 / 2 (in holotype), or 1 / 1 / 1 (in other specimens). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEA2B0EFF3ED02CFC63FC20.taxon	materials_examined	Material examined. Type: ♀ holotype: ZIMBABWE: Harare (MCZ). Other material: BOTSWANA: Serowe, Farmer’s Brigade, XI. 1992, MT, P. Forchhammer leg., 1 ♀ (CNC); same locality label, IV. 1992, 1 ♀ (AMNH). CAMEROON: Messabengondo, 1 – 15. III. 2003, M. Tussac leg., 1 ♀ (HTS). SOUTH AFRICA: Western Cape, Walk- er Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 57 m, 26. XII. 1997, sweep, South Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEA2B0EFF3ED02CFC63FC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEEA2B0EFF3ED02CFC63FC20.taxon	distribution	Distribution. Botswana, Cameroon, South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE92B0FFF3ED5B7FDADFB24.taxon	description	(Figs 132 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE92B0FFF3ED5B7FDADFB24.taxon	description	Description. ♀. Fully winged; body length 5.0 – 5.3 mm. Head black, except mandible and clypeus testaceous; an- tenna totally testaceous in holotype, or with antennomeres 5 – 10 brown in other specimens; mesosoma black, except lateral margins and anterior collar of pronotum reddish; metasoma black; legs testaceous, except coxae and clubs of femora brown; occasionally also tibiae and meso- and metatarsomere 5 brown. Antenna clavate; antennomeres in following proportions: 9: 5: 24: 10: 9: 7: 6: 6: 5: 7.5. Head dull, granulate, with numerous short transverse keels on anterior half or third of frons (occasionally frons granulate and with irregular keels or areolae); frontal line complete; occipital carina complete; lateral ocelli situated on virtual straight line joining posterior edges of eyes; lateral ocelli touching occipital carina; POL = 3.5; OL = 3; OOL = 9; OPL = 0; TL = 2; in holotype POL = OL. Pronotum dull, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slight; posterior collar very short; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with some strong punctures. Notauli incomplete, reaching 0.3 – 0.7 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum rugose or unsculptured. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with 2 – 3 dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (15: 8); 2 R 1 cell open. Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. Enlarged claw (Figs 132 D, E) with one small subdistal tooth and one row of 6 – 8 lamellae. Protarsomere 5 (Figs 132 D, E) with two rows of approximately 13 – 23 lamellae; apex with at least 12 – 27 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE92B0FFF3ED5B7FDADFB24.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, III. 1933, A. Seyrig leg. (MNHN). Other material: KENYA: Eastern Prov., Tsavo East National Park, Nutter’s Farm, near Athi River, 02 ° 38.51 ’ S 38 ° 21.98 ’ E, 1 – 4. II. 1999, MT, R. Copeland leg., 1 ♀ (NMK). MADAGASCAR: Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 20 – 28. VII. 2001, dwarf littoral forest, MT, across sandy trail, MA – 01 – 04 B – 15, M. E. Irwin & R. Harin’Hala leg., 1 ♀ (MOLC); Antsiranana, Ambato Forest, 26.6 km 33 ° NE Ambanja, 13 ° 27.52 ’ S 48 ° 33.06 ’ E, 150 m, 8. XII. 2004, MT, rainforest, B. L. Fisher leg., BLF 11518, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 21 – 26. IV. 2001, MT, MA – 01 – 01 D – 08, R. Harin’Hala leg., 1 ♀ (CAS); Antsiranana, Montagne des Français, 7.2 km 142 ° SE Antsiranana, 12 ° 19.22 ’ S 49 ° 20.17 ’ E, 180 m, 22 – 28. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3130, 1 ♀ (CAS); Antsiranana, SAVA Region, Vohemar District, 43 km E of Daraina, Analabe Sahaka littoral forest, 13 ° 04.76 ’ S 49 ° 54.14 ’ E, 182 m, 14 – 21. XII. 2010, MT, wooded grassland / bushland, MG- 59 - 11, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA – 02 – 09 C – 17, 1 ♀ (CAS); same locality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 1 ♀ (MOLC); Ma- hajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 2 ♀♀ (CAS, MOLC); Ma- hajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19 ’ 16 ’’ S 46 ° 48 ’ 80 ’’ E, 43 m, 14 – 22. IX. 2003, MT, deciduous forest, R. Harin’Hala leg., MA – 25 – 15, 2 ♀♀ (CAS, MOLC); same locality label, 28. IX – 5. X. 2003, MA- 25 - 17, 3 ♀♀ (CAS); same locality label, 7 – 14. IX. 2003, MA- 25 - 14, 1 ♀ (CAS); same locality label, 22 – 28. IX. 2003, MA- 25 - 16, 3 ♀♀ (CAS); Mahajanga, Namoroka National Park, 9.8 km 300 ° WNW Vilanan- dro, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, 140 m, 4 – 8. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445, 1 ♀ (CAS); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♀ (MNHN); same locality label, YPT, 1 ♀ (MNHN); Mahajanga, Ambovomamy Belambo, 20 km NW of Port Berger, 15 ° 27 ’ 07 ’’ S 47 ° 36 ’ 80 ’’ E, 33 m, 1 – 28. X. 2007, MT, secondary growth on white sand, R. Harin’Hala, M. Irwin, F. Parker leg., MG- 33 - 35, 1 ♀ (CAS); Mahajanga, Maintirano District, 50 km E of Maintirano, 18 ° 00.24 ’ S 44 ° 27.12 ’ E, 900 ft., 25. X – 1. XI. 2007, MT, dry forest on sand, MG- 44 - 05, 1 ♀ (MOLC); Mahajanga, Besalampy District, Ma- rofototra dry forest, 17 km W of Besalampy, 16 ° 43.30 ’ S 44 ° 25.42 ’ E, 170 ft., 1 – 8. X. 2007, MT, dry wash in forest, M. Irwin, R. Harin’Hala leg., MG- 42 A- 02, 1 ♀ (MOLC); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 6 – 16. I. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 18 B- 09, 1 ♀ (CAS); Toliara, Cap S. te Marie Special Reserve, 74 km S of Tsihombe, 25 ° 35 ’ 26 ’’ S 45 ° 09 ’ 78 ’’ E, 37 m, 13 – 20. VII. 2003, MT, spiny bush, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 23 - 35, 1 ♀ (CAS); Toliara, Ankazomena, 8 km N of Ambohimahave- lona village, 23 ° 25.82 ’ S 43 ° 50.05 ’ E, 122 m, 21 – 31. X. 2008, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 51 - 01, 1 ♀ (CAS); Toliara, Ifaty, near Hotel Paradisia, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, 9 m, 13. X – 12. XI. 2001, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02 - 16 - 01, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE92B0FFF3ED5B7FDADFB24.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE92B0FFF3ED5B7FDADFB24.taxon	distribution	Distribution. Kenya, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE82B00FF3ED2A4FE14FE70.taxon	description	(Figs 129 C, 132 F, 133 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE82B00FF3ED2A4FE14FE70.taxon	description	Description. ♀ (Fig. 129 C). Fully winged; body length 9.8 mm. Head ferruginous, except vertex and almost entire frons brown; antenna with antennomeres 1 – 4 ferruginous (except distal extremity of antennomere 4 brown), antennomeres 5 – 7 brown (except distal half of antennomere 7 whitish) and 8 – 10 whitish; mesosoma ferruginous, except metanotum, metapectal-propodeal disc, propodeal disc, axillae, epicnemial sulcus, pleural sulcus and meso-metapleural suture black; petiole black; metasoma brown-ferruginous; legs ferruginous. Antenna clavate; antennomeres in following proportions: 18: 9: 67: 44: 35: 25: 19: 17: 14: 17; ADOs present in antennomeres 5 – 10. Head (Fig. 132 F) dull, granulate; occipital carina incomplete, present only behind and on sides of lateral ocelli, laterally not reaching eyes, not present on temple and on latero-ventral side of head; frontal line incomplete, not present near clypeus; clypeus bidentate; lateral ocelli not touching occipital carina; posterior margin of vertex straight (Fig. 132 F); temple absent; POL = 5; OL = 4; OOL = 12; OPL = 3; greatest breadth of lateral ocelli longer than OPL (6: 3). Pronotum (Fig. 133 A) shiny, unsculptured, except some longitudinal keels on lateral regions; pronotum crossed by two deep transverse furrows; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum shiny, very weakly granulate (sculpture hardly visible). Notauli incomplete, reaching about 0.7 × length of mesoscutum. Mesoscutellum shiny, very weakly granulate (sculpture hardly visible). Metanotum shiny, unsculptured, with dense tuft of setae. Metapectal-propodeal disc reticulate rugose, much longer than propodeal declivity (55: 30); propodeal declivity reticulate rugose, without longitudinal keels; median area of propodeal declivity dull, less rugose than rest of metapectal-propodeal disc + propodeal declivity. Mesopleuron and metapleuron shiny, very weakly granulate. Forewing with two large dark transverse bands; distal part of 2 r-rs & Rs vein rectilinear, much longer than proximal part (35: 19). Protarsomeres in following proportions: 31: 4: 13: 40: 65. Protarsomeres 2 and 3 produced into hook. Enlarged claw (Fig. 133 B) with one large subapical tooth and one row of 15 lamellae. Protarsomere 5 (Fig. 133 B) with two rows of about 20 + 21 lamellae; apex very long, with approximately 50 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE82B00FF3ED2A4FE14FE70.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antsiranana, Sakalava Beach, 12 ° 15 ’ 46 ’’ S 49 ° 23 ’ 51 ’’ E, dwarf littoral forest, 10 m, 15. II – 6. III. 2001, MT, across sandy trail, M. Irwin & R. Harin’Hala leg., MA- 01 - 04 B- 03 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE82B00FF3ED2A4FE14FE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE82B00FF3ED2A4FE14FE70.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE72B01FF3ED024FE27FC90.taxon	description	(Figs 129 D – F, 130 A, 133 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE72B01FF3ED024FE27FC90.taxon	description	Description. ♀ (Figs 129 D – F, 130 A). Fully winged; body length 7.3 mm. Head black, except mandible and clypeus testaceous; antenna brown, except scape, pedicel and flagellomere 8 testaceous, distal extremity of antennomere 4 and proximal third of 5 whitish; propleuron brown-black; pronotum black, except posterior collar ferruginous; rest of mesosoma black; metasoma brown-testaceous; legs testaceous. Antenna clavate; antennomeres in following proportions: 12: 6: 56: 33: 21: 14: 10: 7: 8: 10; ADOs present in antennomeres 6 – 10. Head (Figs 129 F, 130 A) dull, granulate and reticulate rugose, without longitudinal keels; occipital carina complete; lateral ocelli almost touching occipital carina, situated just behind virtual straight line joining posterior edges of eyes; POL = 2; OL = 1.5; OOL = 12; OPL = 0.3; TL = 6; greatest breadth of lateral ocelli longer than POL (3: 2). Pronotum (Figs 129 E, F) dull, granulate, with numerous keels around disc, on lateral regions and anterior collar; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar little distinct; posterior collar long; pronotal tu- bercle not reaching tegula. Mesoscutum (Fig. 129 F) granulate and reticulate rugose, not sculptured by longitudinal keels. Notauli incomplete, reaching about 0.8 × length of mesoscutum. Mesoscutellum granulate, weakly rugose. Metanotum rugose. Metapectal-propodeal disc (Fig. 129 F) reticulate rugose, much longer than propodeal declivity; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (22: 8). Protarsomeres in following proportions: 27: 5: 9: 26: 41. Protarsomere 3 produced into hook. Enlarged claw (Fig. 133 C) with one large subdistal tooth and one row of 13 lamellae. Protarsomere 5 (Fig. 133 C) with two rows of about 48 lamellae; apex with at least 40 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE72B01FF3ED024FE27FC90.taxon	materials_examined	Material examined. Type: ♀ holotype: UGANDA: Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.836 ’ N 30 ° 21.700 ’ E, 1523 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 023431 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE72B01FF3ED024FE27FC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE72B01FF3ED024FE27FC90.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE52B03FF3ED34EFE06FC87.taxon	description	(Figs 133 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE52B03FF3ED34EFE06FC87.taxon	description	Description. ♀. Fully winged; body length 4.7 mm. Testaceous, except petiole black and metasoma darkened. An- tenna clavate; antennomeres in following proportions: 21: 13: 71: 28: 24: 20: 18: 15: 14: 20. Head shiny, slightly punctate, setose; frontal line very thin, complete; occipital carina complete, almost invisible on lateral regions of occiput; POL = 3; OL = 3; OOL = 24; OPL = 3; TL distinct. Pronotum shiny, unsculptured, crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slightly visible; posterior collar present; pronotal tubercle not reaching tegula. Mesoscutum shiny, slightly punctate. Notauli incomplete, approximately reaching 0.65 × length of mesoscutum. Mesoscutellum and metanotum shiny, slightly punctate. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two brown transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part. Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 133 D) with one large subdistal tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 133 D) with two rows of 30 lamellae; apex with approximately 15 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.1 – 3.4 mm (holotype 3.4 mm). Head black, except mandible yellow (in speci- men from Mahé Island, 04 ° 39.04 ’ S 55 ° 27.30 ’ E, clypeus brown); antenna yellow-brown; mesosoma black; meta- soma brown; legs yellow, except metacoxa partly black. Antenna filiform, very setose; antennomeres in following proportions: 14: 13: 38: 26: 29: 28: 26: 23: 20: 28. Head dull, granulate; temple absent; frontal line absent; occipital carina incomplete, laterally reaching posterior margin of eyes; holotype with POL = 13; OL = 8; OOL = 5; OPL = 3; greatest breadth of lateral ocelli longer than OL (11: 8); small specimen (2.1 mm) from Mahé Island, 04 ° 39.04 ’ S 55 ° 27.30 ’ E, with POL = 4; OL = 2; OOL = 2.5; OPL = 0.5; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, granulate, with slight areolae on lateral regions. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum shiny, slightly granulate. Metanotum dull, rugose. Metapectal-propodeal disc and propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 rrs & Rs vein much longer than proximal part. Paramere (Fig. 133 E) with distal half sculptured by many transverse striae. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE52B03FF3ED34EFE06FC87.taxon	materials_examined	Material examined. Types: ♀ holotype of M. mahensis: SEYCHELLES: Mahé Island, above Port Glaud, 500 – 1000 ft., 5. XI. 1908, Percy Sladen Trust Expedition (NHMUK); ♂ holotype of L. saxetanus: SEYCHELLES: Mahé Island, top of Mount Sebert, about 2000 ft., 16. I. 1909, Percy Sladen Trust Expedition (NHMUK). Other material: SEYCHELLES: Mahé Island, Mont Copolia, 04 ° 39.04 ’ S 55 ° 27.30 ’ E, 520 m, 8 – 11. II. 2010, MT in forest, BLF 24027, B. L. Fisher et al. leg., 1 ♂ (CAS); Silhouette Island, forest above Mare aux Cochons, IX. 1908, Percy Sladen Trust Expedition, 1 ♂ (Kieffer 1912).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE52B03FF3ED34EFE06FC87.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE52B03FF3ED34EFE06FC87.taxon	distribution	Distribution. Seychelles.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B03FF3ED51EFC74F92E.taxon	description	(Figs 133 F, 135 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B03FF3ED51EFC74F92E.taxon	description	Description. ♀. Fully winged; body length 8.1 – 10.0 mm. Head black, except mandible, clypeus, malar space and anterior region of frons (more along orbits) yellow; occasionally head with some small reddish spots along orbits; propleuron black; pronotum black, except lateral margins and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs brown-black, except tarsi testaceous. Head dull, slightly reticulate rugose; occipital carina complete; lateral ocelli not touching occipital carina; OL shorter than POL; OL = OPL; TL slightly shorter than POL (8: 9). Pronotum (( Fig. 133 F) dull, granulate, with many striae around disc; pronotum crossed by slight anterior transverse impression and very strong posterior transverse furrow; anterior collar almost absent; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum very setose, dull, granulate, with some slight irregular keels. Notauli incomplete, approximately reaching 0.65 × length of mesoscutum. Mesoscutellum as sculptured as mesoscutum. Metanotum rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two short longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein about twice as long as proximal part. Protarsomere 1 about as long as 4. Protarsomere 3 produced into hook. Enlarged claw (( Fig. 135 A) with one large subdistal tooth and one row of 19 lamellae. Protarsomere 5 (Fig. 135 A) with two rows of about 38 lamellae; apex with approximately 14 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B03FF3ED51EFC74F92E.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Paulis (= Isiro), VII. 1947, P. L. G. Benoit leg. (MRAC). Other material: SIERRA LEONE: Freetown, 1 ♀ (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B03FF3ED51EFC74F92E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B03FF3ED51EFC74F92E.taxon	distribution	Distribution. Democratic Republic of the Congo, Sierra Leone.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	description	(Figs 135 B, 136)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head mostly black; head completely and strongly reticulate rugose, with region of vertex along occipital carina sculptured by numerous short longitudinal keels; mesosoma black, except prothorax red (Fig. 136); protarsomere 1 almost 3 × as long as 4; enlarged claw (Fig. 135 B) approximately as long as protarsomere 5, with one row of peg-like setae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	description	Description. ♀ (Figs 136). Fully winged; body length 5.5 mm. Head black, except mandible, clypeus, malar space and narrow stripe between antennal toruli and between eyes and antennal toruli testaceous-reddish; antenna testaceous, except antennomere 3 slightly darkened; mesosoma black, except prothorax red; metasoma testaceousreddish, except distal third black; proleg testaceous; meso- and metaleg testaceous, except femur, tibia and part of coxa brown. Antenna clavate; antennomeres in following proportions: 15: 6: 27: 14: 12: 10: 10: 8: 8: 10; antennomere 3 about ten × as long as broad (27: 2.5). ADOs present on antennomeres 5 – 10. Head (Fig. 136 C) dull, slightly convex, granulate and completely reticulate rugose (with region of vertex along occipital carina sculptured by numerous short longitudinal keels); frontal line complete; occipital carina complete; POL = 5; OL = 3; OOL = 10; OPL = 5; TL = 7; greatest breadth of lateral ocelli shorter than POL (4: 5). Pronotum shiny, punctate and slightly rugose, crossed by transverse furrow; disc slightly humped; posterior collar very short; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli incomplete, reaching about 0.7 × length of mesoscutum. Mesoscutellum strongly punctate, unsculptured among punctures. Metanotum granulate and rugose. Metapectal-propodeal disc reticulate rugose, approximately as long as propodeal declivity; propodeal declivity reticulate rugose, with two short incomplete longitudinal keels. Forewing (Fig. 136 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (19: 12). Protarsomeres in following proportions: 31: 3: 6: 11: 19. Enlarged claw (Fig. 135 B) with one large subdistal tooth and one row of 6 peg-like setae. Protarsomere 5 (Fig. 135 B) with one row of 15 lamellae; apex with about 6 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	materials_examined	Material examined. Type: ♀ holotype, NMK / INV / T- 643: KENYA: Nairobi Prov., ICIPE Campus, Kasara- ni, 1.22281 ° S 36.89714 ° E, 1600 m, 17 – 25. I. 2017, MT, near stream, shaded area under trees, R. Copeland leg. (NMK);	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE42B04FF3ED0B6FD4FFBB0.taxon	etymology	Etymology. This species is named in honour of Scott Miller, formerly Programme Leader, Biodiversity and Conservation, and Principal Scientist at the International Centre of Insect Physiology and Ecology in Nairobi, on whose campus this striking wasp was collected.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	description	(Figs 134, 135 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head mostly black, granulate, except some irregular longitudinal keels along frontal line; mesosoma black; pronotum weakly excavated, not humped (Fig. 134 B); forewing with distal half darkened; propodeal declivity with two complete longitudinal keels; protarsomere 1 about twice as long as 4; enlarged claw (Fig. 135 C) slightly shorter than protarsomere 5, with big subapical tooth.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	description	Description. ♀ (Fig. 134). Fully winged; body length 4.7 mm. Head black, except mandible testaceous and anterior half of clypeus yellow; antenna testaceous, except antennomeres 7 – 10 brown; mesosoma black; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 13: 6: 22: 12: 10: 8: 8: 7: 6: 7. Head (Fig. 134 C) dull, granulate and with few longitudinal keels along frontal line; frontal line complete; occipital carina complete; POL = 6; OL = 4; OOL = 7; OPL = 1.5; TL = 3; greatest breadth of lateral ocelli about as long as TL. Pronotum dull, granulate, weakly excavated, with posterior collar hardly visible. Mesoscutum and mesoscutellum dull, granulate (Fig. 134 A). Notauli incomplete, approximately reaching 0.5 × length of mesoscutum. Metanotum unsculptured. Metapectal-propodeal disc reticulate rugose, slightly shorter than propodeal declivity (18: 25); propodeal declivity reticulate rugose, with two complete longitudinal keels and median area sculptured by transverse keels, except small central unsculptured area. Forewing (Fig. 134 D) with distal half darkened; distal part of 2 r-rs & Rs vein about twice as long as proximal part (14: 7). Protarsomeres in following proportions: 21: 2.5: 4: 10: 17. Enlarged claw (Fig. 135 C) with one large subapical tooth and one row of five bristles. Protarsomere 5 (Fig. 135 C) with two rows of 10 lamellae; apex with approximately 17 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19458): MADAGASCAR: Toamasina, Mobot site, Anala- lava, 7 km SW of Foulpointe, 17 ° 42 ’ 32 ’’ S 49 ° 27 ’ 29 ’’ E, 80 ft., 12 – 19. X. 2007, MT, low altitude dense humid forest, MG- 37 A- 05 (CAS.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE32B05FF3ED2C7FF2BFE98.taxon	etymology	Etymology. The species is named mobotensis after the collection locality in Mobot site (Toamasina, Madagas- car).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE22B06FF3ED3DCFE14FD2C.taxon	description	(Figs 135 D, E, 137 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE22B06FF3ED3DCFE14FD2C.taxon	description	Description. ♀. Fully winged; body length 3.4 – 5.0 mm. Head black, except mandible and partly clypeus yellow; propleuron black; pronotum black, except lateral margins and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs completely testaceous, or with coxae and clubs of femora brown. Antenna clavate; antennomeres of specimen from Aldabra, Takamaka Grove, in following proportions: 19: 12: 55: 26: 23: 15: 13: 12: 11: 16. Head (Fig. 135 D) dull, flat, granulate; occipital carina complete; lateral ocelli not touching occipital carina; OPL shorter than OL; OL = POL; TL shorter than POL. Pronotum shiny, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar slight; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, approximately reaching 0.75 × length of mesoscutum. Mesoscutellum as sculptured as mesoscutum. Metanotum rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (11: 5). Protarsomere 1 longer than 4 (12: 9); protarsomere 3 produced into hook. Enlarged claw (Fig. 135 E) with one large subdistal tooth and one row of 10 lamellae. Protarso- mere 5 (Fig. 135 E) with two rows of about 20 lamellae; apex with approximately 11 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.0 – 2.2 mm. Head black, except mandible testaceous; antenna black; mesosoma and metasoma black; legs brown, except tarsi testaceous. Antenna filiform; antennomeres in following proportions: 6: 5: 9: 7: 7: 7: 6: 5: 5: 7. Head dull, granulate and partly rugose; frontal line complete; occipital carina incomplete, laterally not reaching eyes and temple; POL = 5; OL = 2; OOL = 3.5; OPL = 0.5. Mesoscutum, mesoscutellum and metanotum granulate, slightly rugose. Notauli incomplete, reaching approximately 0.70 – 0.75 × length of mesoscutum. Metapectal-propodeal disc and propodeal declivity dull, reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 7); 2 R 1 cell open. Genitalia in Fig. 137 A. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE22B06FF3ED3DCFE14FD2C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily (MRAC). Other material: SEY- CHELLES: Aldabra Archipelago, Michel Island, 1 ♀ (NHMUK); Aldabra Archipelago, South Island, Takama- ka Grove, 1 – 17. II. 1968, B. Cogan & A. Hutson leg., 1 ♀, 1 ♂ (NHMUK); South Island, Dune Jean – Louis, 1 ♀ (NHMUK). MADAGASCAR: Antsiranana, Nosy Bé Island, 1 ♀ (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE22B06FF3ED3DCFE14FD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE22B06FF3ED3DCFE14FD2C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	description	(Figs 137 D, 138)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head mostly black; frons granulate and rugose; lateral ocelli not touching occipital carina; POL approximately twice as long as OL; temple absent; head (dorsally viewed) with lateral ocelli located in front of imaginary straight line joining posterior edges of eyes; mesosoma black; mesoscutum shiny, with lateral areas sculptured by many subparallel longitudinal keels, and median area punctate, unsculptured among punctures; forewing with distal part of 2 r-rs & Rs vein less than 3 × as long as proximal part; metapectal-propodeal disc short, about as long as propodeal declivity; protarsomere 1 slightly shorter than 4; enlarged claw slightly shorter than protarsomere 5, with one big subapical tooth big (Fig. 137 D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	description	Description. ♀ (Fig. 138). Fully winged; body length 6.3 mm. Head black, except mandible and anterior margin of clypeus testaceous-yellow; antenna brown, except antennomeres 1 and 6 – 10 testaceous-whitish; mesosoma black; metasoma brown; legs brown, except mesotarsomere 1 of meso- and metalegs testaceous-whitish. Antenna clavate; antennomeres in following proportions: 14: 7: 51: 32: 24: 17: 13: 11: 9: 12. ADOs present in antennomeres 5 – 10. Head weakly excavated, dull, granulate and rugose (Fig. 138 C); occipital carina incomplete, only present behind ocellar triangle and on sides of lateral ocelli, laterally not reaching eyes, reaching about 0.5 length of OOL; occiput slightly concave; temple absent; frontal line complete; POL = 4; OL = 2; OOL = 10; OPL = 1; greatest breadth of lateral ocelli longer than OPL (4: 1). Pronotum shiny, crossed by two weak transverse impressions; anterior collar sculptured by many longitudinal striae; lateral regions granulate and sculptured by many longitudinal striae; disc sculptured by many longitudinal striae; posterior collar very short; pronotal tubercle not reaching tegula. Mesoscutum shiny, with lateral areas sculptured by many subparallel longitudinal keels; median area punctate, unsculptured among punctures. Notauli incomplete, reaching about 0.8 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum shiny, with anterior half rugose and posterior half unsculptured. Metapectal-propodeal disc dull, with about as long as propodeal declivity, without transverse posterior keel, sculptured by many subparallel longitudinal keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (21: 18). Hind wing with distal third darkened. Protarsomeres in following proportions: 27: 5: 11: 29: 46. Enlarged claw (Fig. 137 D) with one large subdistal tooth and one row of 19 lamellae. Protarsomere 5 (Fig. 137 D) with two rows of 22 lamellae; apex with at least 25 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	materials_examined	Material examined. Type: ♀ holotype, DAF 2201: IVORY COAST: Taï National Park, UTM 29 N 0682666 0.646862, IV. 2014, MT, Erena Dupont leg. (DAFS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	distribution	Distribution. Ivory Coast.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEE12B18FF3ED090FBE6FEC4.taxon	etymology	Etymology. The species is named nigrithorax, because its thorax is black.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFF2B1AFF3ED065FBF0FB94.taxon	description	(Figs 137 B, C, E, F, 139 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFF2B1AFF3ED065FBF0FB94.taxon	description	Description. ♀. Fully winged; body length 2.2 – 5.3 mm. Head black, except clypeus and mandible testaceous; an- tenna totally testaceous, or brown, except flagellomere 8 testaceous, or brown, except scape, pedicel (occasionally also flagellomere 1) and flagellomeres 6 – 8 testaceous; propleuron black; pronotum black, except lateral margins and posterior collar testaceous or reddish; rest of mesosoma black; metasoma brown or black; legs testaceous, except coxae and clubs of femora brown; occasionally legs totally testaceous; rarely legs brown, except protrochanter, profemur and protibia testaceous-darkened. Antenna clavate; antennomeres of specimen from Nigeria, Ife, in following proportions: 23: 12: 72: 27: 18: 17: 15: 15: 12: 15. Head (Fig. 137 B) dull, granulate, with few longitudinal and irregular keels on frons and vertex; occasionally head completely granulate, without keels on frons and vertex (except for frontal line); frontal line complete; occipital carina complete; OPL = OL or shorter than OL; POL = OL or longer than OL; TL longer or shorter than POL; lateral ocelli situated on virtual straight line joining the posterior edges of eyes. Pronotum (Fig. 137 C) shiny, granulate, with numerous slight striae around disc; pronotum crossed by strong posterior transverse furrow and slight anterior transverse impression; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with short irregular keels on posterior half (often entire posterior half or fourth of mesoscutum reticulate rugose). Notauli incomplete, hardly distinct in posterior half of mesoscutum, reaching at least 0.65 – 0.75 × length of mesoscutum; occasionally notauli almost reaching posterior margin of mesoscutum. Mesoscutellum dull, granulate. Metanotum reticulate rugose, occasionally granulate. Metapectalpropodeal disc and propodeal declivity reticulate rugose, without longitudinal or transverse keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 9 in holotype of L. orophilus; 7: 3 in holotype of D. incertus). Protarsomere 1 longer than 4 (21: 15 in holotype of L. orophilus; 9: 7 in holotype of D. incertus); protarsomere 3 produced into hook. Enlarged claw (Figs 137 E, F) with one large subapical tooth and one row of 7 – 11 lamellae (occasionally in addition to one bristle). Protarsomere 5 (Figs 137 E, F) with two rows of 15 – 31 lamellae; apex with 10 – 18 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.8 – 3.6 mm. Head black, except mandible yellow; antenna yellow-brown; me- sosoma black; metasoma brown; legs yellow, except coxae partly brown. Antenna filiform; antennomeres of specimen from Nigeria, Ife, in following proportions: 13: 9: 25: 16: 18: 18: 15: 13: 13: 17. Head dull, granulate and slightly reticulate rugose; occipital carina incomplete; OL shorter than OOL; OPL shorter than POL; OL shorter than POL. Mesoscutum dull, as sculptured as head. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum and metanotum as sculptured as mesoscutum. Metapectal-propodeal disc and propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part; 2 R 1 cell open. Genitalia in Figs 139 A, B. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFF2B1AFF3ED065FBF0FB94.taxon	materials_examined	Material examined. Types: ♀ holotype of L. orophilus: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), Rutshuru, 7. IV. 1937, Miss. Prophylactique (MRAC); ♀ holotype of D. incertus: SOUTH AFRICA: Eastern Cape, Grahamstown, VIII. 1958, suction trap, D. Annecke leg. (NHMUK). Paratype of L. orophilus: same locality label as holotype, ♂ allotype of L. orophilus (not valid, because designated after the original description). Other material: ANGOLA: Moxico Prov., Luena (= Vila Luso), 1 ♀ (CAS). CAMEROON: South-West Region, Limbe (= Victoria), 1 ♀ (CAS); Messabengondo, 1 – 15. III. 2003, M. Tussac leg., 1 ♀ (HTS). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Katanga Prov., Shaba, Mweru Lake, Pweto, 1918, P. Gérard leg., 1 ♀ (MRAC) (Benoit 1951 c); Haut-Lomami, Upemba National Park, Lusinga (MRAC); Haut-Uele (= Haut-Zaire), Garamba National Park (MRAC); North Kivu, Virunga National Park (= Albert National Park), Kalinga- Bitshumbi (MRAC). GABON: Estuaire Prov., Ntoum, 1 ♀ (FSAE); Nyanga Prov., Tchibanga (FSAE); Ogooué-Maritime Prov., Moukalaba-Dougoua Reserve, 7 km NW Doussala, 02 ° 19.84 ’ S 10 ° 32.65 ’ E, 110 m, 21. III. 2000, sweep, GA 00 - S 151, Savanna, grasses & Nauclea latifolia, SAM-HYM-PO 023755, S. van Noort leg., 1 ♀ (SAMC); Woleu-Ntem Prov., Assok-Ngoum, 01 ° 42 ’ N 11 ° 39 ’ E, 1 ♀ (FSAE). GHANA: Ashanti Region, Kumasi Metropolitan, Kwadaso, 2 ♀♀ (NHMUK, AMNH). KENYA: Coast Prov., Shimba Hills National Park, 04 ° 23.456 ’ S 39 ° 41.687 ’ E, 389 m, MT, just inside Longomwagandi Forest, 24. IX – 8. X. 2008, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Umani Springs camp, near Chyulu Hills, 02 ° 27.97 ’ S 37 ° 54.80 ’ E, 21 – 25. XII. 1998, MT, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Njuki – ini Forest, near Forest Station, 00 ° 51.660 ’ S 37 ° 41.843 ’ E, 1455 m, 4. IV – 2. V. 2007, MT, just inside indigenous forest, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Samburu Na- tional Reserve, near Ewaso Ng’iro River, 00 ° 56.797 ’ N 37 ° 53.563 ’ E, 874 m, 12 – 26. VI. 2006, MT, riverine forest, next to headquarters, R. Copeland leg., 2 ♀♀ (NMK); Nyanza Prov., Ungoye, ICIPE Field Station, 00 ° 61.517 ’ S 34 ° 09.200 ’ E, 1147 m, 12 – 26. II. 2005, MT, inside seasonally swampy forest, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Masai Mara National Reserve, 01 ° 54.638 ’ S 35 ° 30.672 ’ E, 1756 m, MT, Riverine forest, near KWS Research Station, 25. VII – 8. VIII. 2008, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Marich Pass field station, 01 ° 53.633 ’ N 35 ° 45.800 ’ E, MT, low canopy riverine forest, 18. IV – 2. V. 2005, R. Copeland leg., 1 ♀ (NMK). MA- LAWI: Central Region, Kasungu District, Mtunthama, VIII. 1982, J. Feehan leg., 2 ♀♀ (DJBC); same locality label, VII. 1982, 1 ♀ (CNC). MOZAMBIQUE: Maputo Prov., Maputo, Costa do Sol, reared from Dictyopharidae, 1 ♀ (AMNH); Maputo Prov., 9 km N of Ponta de Ouro, reared from Paroxychara cowla Medler, 1 ♀ (AMNH); Gaza Prov., Praia do Bilene Airport, reared from Paroxychara cowla Medler, 1 ♀ (AMNH); Niassa Prov., Cuamba, Farm of the Catholic University of Mozambique, C. 4. III. 2005, B. 9. III. 2005, Sf. 26. III. 2005, M. Olmi reared from a nymph of Elasmoscelis cimicoides Spinola, 1 ♀ (MOLC); Niassa Prov., Cuamba, Mituque, 6 – 22. V. 2008, MT, M. Olmi leg., 1 ♀ (MOLC). NIGERIA: Kogi State, Lokoja, Kabba, 1 ♀ (CAS); Ogun State, Ikenne, 1 ♀ (NHMUK); Ondo State, Akure, 2 ♀♀ (AMNH, NHMUK); Osun State, Ife (= Ilé – Ifè), VII. 1973, X. 1973, IX. 1974, 9. II. 1975, J. T. Medler leg., 2 ♀♀, 2 ♂♂ (NHMUK); Oyo State, Ibadan, 1 ♀ (CNC). OMAN: Dhofar, Ain Razat, 17 ° 07.75 ’ N 54 ° 14.24 ’ E, 110 m, 6. X. 2001, dishes, F. Strumia leg., 1 ♀ (MOLC). SOUTH AFRICA: Western Cape, Cape Town, Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 12 – 17. IX. 1993, MT, Mesic mountain Fynbos on Sandstone, Protea dominated, SAM-HYM-PO 03806, S. van Noort leg., 1 ♀ (SAMC); same locality label, 2 – 9. II. 1994, 1 ♀ (SAMC). UNITED ARAB EMIRATES: Sharjah Desert Park, 25 ° 17 ’ N 55 ° 42 ’ E, 22. III – 5. IV. 2009, LT, 1 ♀ (MOLC); Wadi Shawkah, 25 ° 06 ’ N 56 ° 01 ’ E, 5. V – 24. VI. 2007, WT, 2 ♀♀ (MOLC). YE- MEN: Al Lahima, 16. X – 31. XII. 2000, MT, A. van Harten & A. M. Hager leg., 1 ♀ (MOLC); same locality label, 9. IV – 5. VI. 2001, 1 ♀ (MOLC). ZIMBABWE: near Harare (= Salisbury), Chishawasha, X – XI. 1978, A. Watsham leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFF2B1AFF3ED065FBF0FB94.taxon	biology_ecology	Hosts. Dictyopharidae, Flatidae and Lophopidae (Guglielmino et al. 2013): in Mozambique: Elasmoscelis cimicoides Spinola (Lophopidae); Paroxychara cowla Medler (Flatidae); unidentified species of Dictyopharidae. Natural enemies. In Mozambique: Cheiloneurus caesar Guerrieri 2006 (Hymenoptera, Encyrtidae) (Guerrieri 2006). The reared specimens have the following locality labels: Mozambique, Maputo Prov., Maputo, municipal plant nursery, parasitized planthopper collected on 26. XI. 1993; cocoon spun on 28. XI. 1993; encyrtids emergence on 13. XII. 1993: 3 ♂♂, 18 ♀♀. A further unidentified species of Cheiloneurus was reared in Mozambique, Niassa Prov., Cuamba, Campus of the Faculty of Agriculture, from a nymph of Dictyopharidae parasitized by a Dryinus. The host is probably Dryinus orophilus: C. 24. IV. 2012; B. 28. IV. 2012; Encyrtids emergence 1. VI. 2012: 5 ♀♀, 14 ♂♂.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFF2B1AFF3ED065FBF0FB94.taxon	distribution	Distribution. Angola, Cameroon, Democratic Republic of the Congo, Gabon, Ghana, Kenya, Malawi, Mozambique, Nigeria, Oman, South Africa, United Arab Emirates, Yemen, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFD2B1BFF3ED22BFE5BFE98.taxon	description	(Fig. 139 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFD2B1BFF3ED22BFE5BFE98.taxon	description	Description. ♀. Fully winged; body length 6.2 mm. Head black, except mandible and part of clypeus testaceous; antenna brown, except scape, pedicel and flagellomeres 5 – 8 testaceous; mesosoma black; metasoma brown; legs black, except part of procoxa, part of protrochanter and part of meso- and metatarsus testaceous. Antenna clavate; antennomeres in following proportions: 12: 7: 50: 28: 19: 13: 12: 11: 9.5: 13. Head excavated, dull, granulate and sculptured by small areolae; frontal line complete; occipital carina complete; lateral ocelli situated in front of virtual straight line joining posterior edges of eyes; POL = 4; OL = 3; OOL = 10; OPL = 2; TL = 3; temple slightly visible. Pronotum crossed by strong anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar very short; pronotum sculptured by numerous longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum shiny, with lateral regions sculptured by numerous parallel and longitudinal keels; median region partly smooth and partly sculptured by irregular keels (in specimen from Tanzania, Mkomazi Game Reserve, median region completely sculptured by numerous longitudinal keels). Notauli apparently complete and posteriorly separated; near posterior margin of mesoscutum, notauli hardly visible among longitudinal keels. Mesoscutellum and metanotum shiny, punctate, unsculpured among punctures. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel, slightly longer than propodeal declivity (30: 25); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (19: 14) Protarsomeres in following proportions: 28: 4: 10: 23: 36. Enlarged claw (Fig. 139 C) with one large subdistal tooth and one row of 11 lamellae. Protarsomere 5 (Fig. 139 C) with two rows of 13 + 15 lamellae; apex with approximately 22 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFD2B1BFF3ED22BFE5BFE98.taxon	materials_examined	Material examined. Type: ♀ holotype: GABON: Estuaire Prov., Kango, 17. XI. 1985, A. Pauly leg. (FSAE). Other material: TANZANIA: Mkomazi Game Reserve, Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 26. XI – 10. XII. 1995, Aca- cia / Commiphora / Combretum bushland, MT, SAM-HYM-PO 14829, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFD2B1BFF3ED22BFE5BFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFD2B1BFF3ED22BFE5BFE98.taxon	distribution	Distribution. Gabon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFC2B1CFF3ED3C5FE14FE54.taxon	description	(Figs 130 B, 139 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFC2B1CFF3ED3C5FE14FE54.taxon	description	Description. ♀ (Fig. 130 B). Fully winged; body length 3.9 mm. Head black, except mandible, clypeus and frontal region between antennal toruli testaceous; antenna testaceous; mesosoma black, except lateral and posterior margins of pronotum testaceous; metasoma black; legs testaceous, except coxae, clubs of femora and protibia darkened. Antenna clavate; antennomeres in following proportions: 7: 5.5: 25: 12: 9: 7: 5.5: 6: 5: 7.5; ADOs present on antennomeres 5 – 10. Head flat, dull, granulate; frontal line complete; occipital carina complete; POL = 2; OL = 2; OOL = 7; OPL = 0.5; TL = 3. Pronotum shiny, smooth, humped, crossed by weak anterior transverse impression and strong posterior transverse furrow; anterior collar long; posterior collar very short; anterior collar of pronotum unsculptured; disc of pronotum slightly granulate; lateral regions of pronotum with few longitudinal striae; posterior transverse furrow of pronotum crossed by short longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, approximately reaching 0.65 × length of mesoscutum. Mesoscutellum shiny, slightly granulate. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel, approximately as long as propodeal declivity (20: 20); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 7). Protarsomeres in following proportions: 19: 2.5: 6.5: 12: 19. Enlarged claw (Fig. 139 D) with one large subdistal tooth and one row of eight lamellae. Protarsomere 5 (Fig. 139 D) with two rows of 6 + 15 lamellae (shortest row composed of very long lamellae); apex with at least 17 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFC2B1CFF3ED3C5FE14FE54.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW Anivorano Nord, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, MT, tropical dry forest, Fisher, Gris- wold et al. leg., BLF 3011 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFC2B1CFF3ED3C5FE14FE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFC2B1CFF3ED3C5FE14FE54.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B1DFF3ED6D7FE1FFA8D.taxon	description	(Figs 139 E, F, 140 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B1DFF3ED6D7FE1FFA8D.taxon	description	Description. ♀. Fully winged; body length 6.2 mm. Head testaceous-reddish, with brown spot on frons; antenna brown, except scape and pedicel reddish and flagellomere 8 whitish; prothorax, mesoscutum and mesoscutellum reddish-dark; rest of mesosoma black; metasoma brown, except segment 1 partly reddish; legs testaceous, except chela and metatarsomere 2 and partly 3 whitish. Antenna clavate; antennomeres in following proportions: 9: 7: 49: 29: 21: 14: 10: 9: 7: 8. Head dull, flat, granulate; frontal line present; occipital carina complete; POL = 4.5; OL = 3.5; OOL = 11; OPL = 2; TL = 7; lateral ocelli situated behind virtual straight line joining posterior edges of eyes; posterior margin of vertex convex. Pronotum dull, setose, granulate, crossed by strong transverse impression; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, granulate, with numerous slight irregular keels. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Metapectal-propodeal disc dull, reticulate rugose, with two median longitudinal keels; propodeal declivity reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (15: 7). Protarsomeres in following proportions: 20: 3.5: 8: 17: 28. Enlarged claw (Fig. 139 E) with one subdistal tooth and one row of 10 lamellae. Protarsomere 5 (Fig. 139 E) with two rows of about 45 lamellae extending continuously to apex. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.4 – 2.6 mm. Head black, except mandible testaceous; antenna black, except scape and pedicel testaceous; mesosoma and metasoma black; legs brown, except articulations and tarsi testaceous. Antenna filiform; antennomeres in following proportions: 4: 4: 10: 8: 8: 8: 8: 6: 6: 8. Head dull, granulate, with some slight areolae on lateral regions; frontal line incomplete; occipital carina incomplete; POL = 6; OL = 3; OOL = 4; OPL = 1; TL = 3. Mesoscutum and mesoscutellum dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Metanotum dull, granulate, with numerous areolae near anterior margin. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 7). Genitalia in Fig. 139 F. Hypopygium in Fig. 140 A. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B1DFF3ED6D7FE1FFA8D.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Gauteng, Pretoria (AEIC). Paratypes: same local- ity label as holotype, 2 ♂♂ (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B1DFF3ED6D7FE1FFA8D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B1DFF3ED6D7FE1FFA8D.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B10FF3ED312FB5FF8F4.taxon	description	(Figs 140 B – D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B10FF3ED312FB5FF8F4.taxon	description	Description. ♀. Fully winged; body length 4.0 – 5.9 mm. Head black, except mandible and clypeus yellow; rarely clypeus black or brown; occasionally anterior region of frons testaceous; occasionally malar space with testaceous spot (in specimen from Botswana, Serowe); antenna testaceous; occasionally antennomeres 7 – 8, or 3 – 10, or 3 – 8 brown; propleuron black; pronotum black, with lateral regions and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs testaceous, except coxae and clubs of femora partly brown. Antenna clavate; antennomeres in following proportions: 10: 5: 21: 10: 9: 8: 6: 6: 4.5: 8. Head (Fig. 140 B) shiny or dull, completely or partly granulate, occasionally partly unsculptured; occasionally head partly granulate and partly sculptured by a few irregular keels; frontal line complete; occipital carina complete; lateral ocelli usually touching occipital carina, occasionally very close, but not touching occipital carina; OL = POL. In specimen from Madagascar, 17 ° 45 ’ S 47 ° 59 ’ E, POL = 4; OL = 2. Pronotum dull, partly or completely granulate, with numerous slight striae around disc; occasionally pronotum shiny, almost completely unsculptured; posterior collar of pronotum short, about 0.5 × as long as distance from tegula to pronotal tubercle; pronotal tubercle not reaching tegula. Mesoscutum dull, slightly or strongly granulate, with short keels or areolae near posterior margin; occasionally median region of mesoscutum shiny, unsculptured (in specimen from Cameroon, Nkoemvon, mesoscutum completely granulate, with no areolae or keels; in specimen from South Africa posterior half of mesoscutum completely reticulate rugose; in specimen from Botswana, Serowe, mesoscutum almost completely reticulate rugose; in specimen from Cameroon, Messabengondo, mesoscutum completely granulate and rugose). Notauli incomplete, approximately reaching 0.65 – 0.90 × length of mesoscutum (rarely 0.5 × length of mesoscutum; rarely notauli complete and posteriorly separated). Mesoscutellum dull, granulate, or shiny and unsculptured. Metanotum dull, rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with three brown transverse bands (two proximal bands almost joint, occasionally completely joint); distal part of 2 r-rs & Rs vein about twice as long as proximal part (18: 8). Hind wing hyaline, without dark spots; occasionally (in very large specimens) with one dark spot. Protarsomere 1 approximately as long as 4; in specimens from Madagascar, 19 ° 07.56 ’ S 44 ° 48.53 ’ E and 18 ° 01.35 ’ S 44 ° 03.02 ’ E, protarsomere 1 longer than 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 140 D) with one big subdistal tooth and one row of 9 – 13 lamellae (in specimen from Madagascar, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, subapical tooth distinctly divided into two teeth (Fig. 140 C), so that apparently this specimen has to be included in group 3 of Dryinus). Protarsomere 5 (Fig. 140 D) with two rows of 16 – 39 lamellae; apex with 11 – 16 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.6 – 2.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae, club of metafemur and metatibia brown. Antenna filiform; antennomeres in following proportions: 8: 7: 17: 12: 13: 13: 11: 10: 10: 11. Head dull, granulate; frons flat in front of anterior ocellus and humped between antennal toruli; frontal line incomplete; occipital carina incomplete; POL = 9; OL = 3; OOL = 3; lateral ocelli touching occipital carina; temple absent. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum almost completely dull, granulate, with narrow posterior region smooth and shiny. Metanotum with anterior half rugose and posterior half shiny and smooth. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (14: 13); 2 R 1 cell open. Paramere without dorsal process. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B10FF3ED312FB5FF8F4.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Rogez (= Andekaleka), I. 1931, A. Sey- rig leg. (MNMS). Other material: BOTSWANA: Central District, Serowe, XI. 1985, MT, P. Forchhammer leg., 1 ♀ (NHMUK). CAMEROON: South Region, near Ebolowa, Nkoemvon, 30. XI. 1979 – 5. I. 1980, MT in forest, D. Jackson leg., NHMUK (E) 2007 – 19, 1 ♀ (NHMUK); same locality label, 23. IX – 25. X. 1980, 1 ♀ (NHMUK); Messaben- gondo, 1 – 15. III. 2003, M. Tussac leg., 1 ♀ (HTS). ETHIOPIA: Shoa Prov., 95 km W Addis Ababa, Ambo, near Plant Protection Research Institute, 2400 – 2500 m, 19. VI. 1990, A. Emel’yanov leg., 1 ♀ (ZISP). MADAGASCAR: Anta- nanarivo, La Mandraka, 1 ♀ (MRAC); same locality label, II. 1953, Pierre Andria Robinson leg., 1 ♀ (MNHN); An- tananarivo, Analamanga Region, Ambatolaona, VI. 1957, Pierre Andria Robinson leg., 1 ♀ (MNHN); Antananarivo, Andebadeba, 4 km from Arivanimamo, 19 ° 00 ’ 29 ’’ S 47 ° 08 ’ 55 ’’ E, 1600 m, 7 – 14. X. 2007, Uapaca forest, MT, M. Irwin, R. Harin’Hala leg., MA- 46 - 02, 1 ♀ (CAS); Antananarivo, Ankokoy Forest, 3 km E of Ibity, 20 ° 04.05 ’ S 46 ° 59.97 ’ E, 1700 m, 12 – 22. XII. 2008, MT, in Uapaca forest, M. Irwin, R. Harin’Hala leg., MG- 56 - 05, 1 ♀ (CAS); Antsiranana, Nosy Bé Island, 1 ♀ (HNHM); Antsiranana, Manongativo Reserve, 19.8 km 229 ° SW Antanambao, 13 ° 58 ’ S 48 ° 26 ’ E, 400 m, 8 – 13. XI. 1998, B. L. Fisher leg., 1 ♀ (DEUCD); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 28. IV – 7. V. 2005, MT, low altitude rainforest, M. Irwin & R. Harin’Hala leg., MA- 31 - 20, 1 ♀ (CAS); Antsiranana, Marojejy National Park, Manantenina River, 27.6 km 35 ° NE Andapa, 9.6 km NNW Manantenina village, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 15 – 18. XI. 2003, YPT, rainforest, BLF 8873, B. L. Fisher et al. leg., 1 ♀ (CAS); Antsiranana, Marojejy National Park, Manantenina River, 28.0 km 38 ° NE Andapa, 8.2 km NNW Manantenina village, 14 ° 26.12 ’ S 49 ° 46.30 ’ E, 450 m, 12 – 25. XI. 2003, YPT, rainforest, BLF 8725, B. L. Fisher et al. leg., 2 ♀♀ (CAS, MOLC); Antsiranana, 7 km N of Joffreville, 12 ° 27 ’ S 49 ° 14 ’ E, 426 m, dryforest, MT, M. Irwin, R. Harin’Hala leg., MA- 01 - 07 - 16, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 19. III – 6. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 09, 1 ♀ (MOLC); same locality label, 4 – 19. III. 2001, MA- 01 - 01 A- 08, 1 ♀ (CAS); same locality label, 21 – 28. I. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 A- 01, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 14 – 30. V. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 11, 1 ♀ (CAS); Fianarantsoa, Haute-Matsiatra Region, Andringitra-Ambalavao, Cirque Boby, 2500 m, 13. I. 1958, R. Paulian leg., 1 ♀ (MNHN); Fianarantsoa, Ranomafana National Park, Talatakely, 21 ° 15 ’ S 47 ° 25 ’ E, 850 m, 14. IV. 1998, 1 ♀ (CAS); Fianarantsoa, Ranomafana, 15. III. 1994, 1 ♀ (ISNB); 7 km W of Ranomafana, 900 m, 2 ♀♀ (FSAE, USNM); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 4 – 12. II. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 15, 1 ♀ (CAS); Fianarantsoa, Alakamisy, 21. XI. 2006, in forest, C. Canepari leg., 1 ♀ (MCSNG); Fianarantsoa, Ranomafana National Park, Belle Vue at Ta- latakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 26. II – 10. III. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 55, 1 ♀ (MOLC); same locality label, 14 – 21. I. 2002, MA- 02 - 09 C- 12, 1 ♀ (CAS); same locality label 22 – 28. XI. 2001, MA- 02 - 09 C- 04, 1 ♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 12 – 19. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 20, 1 ♀ (MOLC); same locality label, 15 – 21. XII. 2001, MA- 02 - 09 B- 07, 1 ♀ (CAS); same locality label, 22 – 28. XI. 2001, MA- 02 - 09 B- 04, 1 ♀ (MOLC); same locality label, 27. IV – 7. V. 2003, MA- 02 - 09 B- 59, 1 ♀ (CAS); Fianarantsoa, Be- fotaka – Midongy National Park, Papango, 27.7 km S Midongy-South Mt. Papango, 23 ° 50.07 ’ S 46 ° 57.49 ’ E, 940 m, 13 – 19. XI. 2006, MT, B. L. Fisher et al. leg., BLF 14774, 1 ♀ (CAS); Fianarantsoa, Agnalazaha Forest Reserve, Ma- habo, 42.9 km 215 ° SW of Farafangana, 23 ° 11.38 ’ S 47 ° 43.23 ’ E, 20 m, 18. IV. 2006, MT, littoral rainforest, B. L. Fisher et al. leg., BLF 13822, 1 ♀ (CAS); Fianarantsoa, 50 km S of Farafangana, Mahabo Mananivo, Ampitavanan- ima Forest, 23 ° 07 ’ 79 ’’ S 47 ° 43 ’ 02 ’’ E, 34 m, 26. I – 4. II. 2007, MT, in low altitude littoral forest, M. Irwin, F. Parker, R. Harin’Hala leg., MG- 35 - 03, 1 ♀ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 25. X – 4. XI. 2006, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 56, 1 ♀ (CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 750 m, 10 – 17. III. 2002, MT, in open area, MA- 02 - 11 A- 19, R. Harin’Hala leg., 1 ♀ (CAS); same locality label, 15 – 22. IV. 2002, MA- 02 - 11 A- 24, 1 ♀ (CAS); same locality label, 30. XI – 7. XII. 2001, MA- 02 - 11 A- 05, 1 ♀ (CAS); same locality label, 19 – 26. I. 2002, MA- 02 - 11 A- 12, 1 ♀ (CAS); Fianarantsoa, near Isalo National Park, dry wash east of Interpretive Center, 22 ° 37.60 ’ 45 ° 21.49 ’ E, 885 m, 9 – 23. VIII. 2002, MT, in open area, MA- 02 - 11 B- 37, R. Harin’Hala leg., 1 ♀ (CAS); same locality label, 7 – 22. IX. 2002, MA- 02 - 11 B- 39, 2 ♀♀ (CAS); same locality label, 3 – 17. X. 2002, MA- 02 - 11 B- 41, 1 ♀ (CAS); Fianarantsoa, Analalava Forest, 29.6 km 280 ° W Ranohira, 22 ° 35.30 ’ S 45 ° 07.42 ’ E, 700 m, 1 – 5. II. 2003, tropical dry forest, beating low vegetation, Fisher, Griswold et al. leg., BLF 7384, 1 ♀ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 2.5 km 62 ° ENE Bekopaka, Ankidrodroa River, 19 ° 07.56 ’ S 44 ° 48.53 ’ E, 100 m, 11 – 15. XI. 2001, sifted litter, tropical dry forest on Tsingy, Fisher, Griswold et al. leg., BLF 4340, 1 ♀ (CAS); Maha- janga, Bale de Baly National Park, 12.4 km 337 ° NNW Soalala, 16 ° 00.36 ’ S 45 ° 15.54 ’ E, 10 m, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6811, 2 ♀♀ (MOLC); Mahajanga, Beanka Forest Reserve, 50.2 km E Main- tirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, YPT, tropical dry forest on tsingy, B. L. Fisher et al. leg., BLF 22788, 3 ♀♀ (2 in CAS, 1 in MOLC); Mahajanga, Maintirano District, Asondrodava dry forest, 15 km N of Maintirano, 17 ° 57.92 ’ S 44 ° 2.13 ’ E, 200 ft., 7 – 14. XII. 2007, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 41 - 11, 1 ♀ (CAS); same locality label, 3 – 10. XII. 2007, MG- 43 A- 09, 1 ♀ (CAS); same locality label, 8 – 15. X. 2007, 1 ♀ (CAS); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 23 – 30. X. 2007, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 06, 1 ♀ (CAS); same locality label, 20. XI – 4. XII. 2007, MG- 38 - 09, 2 ♀♀ (CAS); Mahajanga, Ambatofolaka, Namoroka, 53 km from Soalala, 3 km N Vilanandro village, 16 ° 28 ’ 04 ’’ S 45 ° 23 ’ 48 ’’ E, 400 ft., 21 – 28. IX. 2007, dense dry forest, MT, M. Irwin, R. Harin’Hala leg., MG- 40 A- 01, 1 ♀ (CAS); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19 ’ 16 ’’ S 46 ° 48 ’ 80 ’’ E, 43 m, 17 – 24. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 11, 1 ♀ (CAS); same locality label, 12 – 19. X. 2003, MA- 25 - 19, 1 ♀ (CAS); same locality label, 28. IX – 5. X. 2003, MA- 25 - 17, 2 ♀♀ (CAS); Mahajanga, Namoroka Na- tional Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, 100 m, 12 – 16. XI. 2002, YPT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6536, 2 ♀♀ (CAS, MOLC); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♀ (MNHN); Mahajanga, Namoroka National Park, 23 – 25. X. 2016, YPT, C. Villemant leg., 1 ♀ (MNHN); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W Anjiaman- girana, Analagnambe dry degraded forest, 15 ° 09.42 ’ S 47 ° 47.05 ’ E, 97 m, 2 – 6. I. 2011, MT, MG- 61 - 04, 1 ♀ (MOLC); Toamasina, Manakambahiny-E, 17 ° 45 ’ S 48 ° 43 ’ E, 1 ♀ (FSAE); Toamasina, 25 km W of Morarano- Chrome, 17 ° 45 ’ S 47 ° 59 ’ E, 1 ♀ (FSAE); Toamasina, Morarano-Chrome, II. 1992, 1 ♀ (ISNB); Toamasina, Foulpointe, 1 ♀ (FSAE); Toamasina, Ivondro, 1 ♀ (MNHN); Toamasina, Rogez (= Andekaleka), VI. 1957, Pierre Andria Robinson leg., 1 ♀ (MNHN); Toamasina, Akirindro Mt., 7.6 km 341 ° NNW Ambinanitelo, 15 ° 17.18 ’ S 49 ° 32.54 ’ E. 600 m, 17 – 21. III. 2003, YPT, rainforest, Fisher, Griswold et al. leg., BLF 8255, 2 ♀♀ (CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km 332 ° Toamasina, 17 ° 55.26 ’ S 49 ° 11.59 ’ E, 390 m, 28. XI. 2005, YPT, rainforest, Brian L. Fisher et al. leg., BLF 13124, 1 ♀ (CAS); Toamasina, Zahamena National Park, Onibe River, 17 ° 45 ’ 33 ’’ S 48 ° 51 ’ 17 ’’ E, 780 m, 21 – 23. II. 2009, rainforest, YPT, BLF 22170, B. L. Fisher et al. leg., 1 ♀ (CAS); Toamasina, 9.2 km WSW Befingotra, Anjanaharibe-sud Reserve, 14 ° 45 ’ S 49 ° 28 ’ E, 1280 m, 13 – 21. XI. 1994, B. L. Fisher leg., 1 ♀ (DEUCD); Toliara, Bekily, 2 ♀♀ (MNHN, MRAC); Toliara, Beroboka, 1 ♀ (NHMUK); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 3 – 11. VIII. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 33, M. Irwin, R. Harin’Hala leg., 1 ♀ (MOLC); Toliara, Belalanda, 1 ♀ (FSAE); Toliara, Mahafaly Plateau, 6.2 km 74 ° ENE of Itampolo, 24 ° 39.13 ’ S 43 ° 59.48 ’ E, 80 m, MT, spiny forest thicket, 21 – 25. II. 2002, Fisher, Griswold et al. leg., BLF 5759, 1 ♀ (MOLC); Toliara, Androimpano Forest, 5 km E of Itampolo, 24 ° 39.02 ’ S 43 ° 57.79 ’ E, 130 m, 3 – 16. VII. 2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54 C- 33, 1 ♀ (CAS); Toliara, Behara, 1 ♀ (MRAC); Toliara, Fort-Dauphin, 1 ♀ (MRAC); Toliara, Cap S. te Marie Special Reserve, 74 km S of Tsihombe, 25 ° 35 ’ 26 ’’ S 45 ° 09 ’ 78 ’’ E, 37 m, 18 – 26. V. 2003, MT, spiny bush, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 23 - 27, 1 ♀ (CAS); same locality label, 25. II – 7. III. 2003, 3 ♀♀ (2 in CAS, 1 in MOLC); Toliara, Forêt de Kirindy, 15.5 km 64 ° ENE Marofandilia, 10 – 17 September 2008, 20 ° 04 ’ 09 ’’ S 044 ° 39 ’ 38 ’’ E, B. L. Fisher, MT, tropical dry forest, 30 m, BLF 18285 _ 45, 2 ♀♀ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 23. IX – 3. X. 2002, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 34, 1 ♀ (CAS); same locality label, 6 – 17. VII. 2003, MA- 02 - 18 A- 61, 1 ♀ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 30. X – 9. XI. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 44, 1 ♀ (CAS); same locality label, 5 – 15. I. 2004, MA- 02 - 20 - 51, 1 ♀ (CAS); Toliara, Beza Mahafaly Reserve, par- celle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 23. III – 2. IV. 2003, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 54, 1 ♀ (MOLC); same locality label, 1 – 8. II. 2002, MA- 02 - 14 A- 14, 1 ♀ (CAS); same locality label, 8 - 15. II. 2002, MA- 02 - 14 A- 15, 1 ♀ (CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Cave, 6.4 km 77 ° ENE Efoetse, 17.4 km 170 ° S Beheloka, 24 ° 02.50 ’ S 43 ° 45.11 ’ E, 40 m, 18 – 22. III. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 6159, 1 ♀ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 17 – 24. III. 2003, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 - 19, 1 ♀ (CAS); same locality label, 5 – 15. II. 2003, MA- 02 - 22 - 15, 1 ♀ (CAS); same locality label, 15 – 25. II. 2003, MA- 02 - 22 - 16, 1 ♀ (CAS); same locality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 1 ♀ (CAS); Toliara, Berenty, 1 ♀ (NHMUK); Toliara, Berenty, 12 km NW Amboasary, 5 – 15. V. 1983, J. S. Noyes & M. C. Day leg., 2 ♀♀ (NHMUK); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 9 – 19. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 02, 1 ♀ (CAS); Toliara, Kalambatritra Special Reserve, Ampanihy, 23 ° 27 ’ 49 ’’ S 46 ° 27 ’ 47 ’’ E, 1270 m, 9 – 10. II. 2009, YPT, montane rainforest, B. L. Fisher et al. leg., BLF 21567, 1 ♀ (CAS); Toliara, Ranobe, 23 ° 02.783 ’ S 43 ° 36.617 ’ E, 20 m, 27. X – 2. XI. 2003, MT, spiny forest thicket, MGF 081, Frontier Wilderness Project, 1 ♀ (CAS); Toliara, Ranobe, 23 ° 02.367 ’ S 43 ° 36.617 ’ E, 20 m, 8 – 15. XI. 2003, MT, gallery forest, MGF 084, Frontier Wilderness Project, 1 ♀ (CAS); Toliara, Fiherenana, 23 ° 13.351 ’ S 43 ° 52.853 ’ E, 65 m, 5 – 10. VIII. 2003, MT, degraded riparian forest close to water, MGF 075, Frontier Wilderness Project, 1 ♀ (CAS); Toliara, Fiherenana, 23 ° 10.37 ’ S 43 ° 57.39 ’ E, 100 m, 31. X – 6. XI. 2002, MT, gallery forest, MGF 045, Frontier Wilderness Project, 4 ♀♀ (CAS); same locality label, 22 – 31. X. 2002, MGF 042, 2 ♀♀ (CAS, MOLC); Toliara, Manderano, 23 ° 31.39 ’ S 44 ° 5.18 ’ E, 70 m, 15 – 29. V. 2002, MT, edge of marsh, 5 m from road, gallery forest, MGF 032, Frontier Wilderness Project, 2 ♀♀ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI- 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 1 ♀ (CAS); Toliara, Vohi- basia National Park, 44 km NE of Sakaraha, Ambakaka gallery / palm forest, 22 ° 40 ’ S 44 ° 50 ’ E, 600 m, 26. XII. 2010 - 3. I. 2011, M. Irwin, R. Harin’Hala leg., MG- 60 - 12, 1 ♀ (CAS). SOUTH AFRICA: Limpopo, 13 km W of Beitbridge, Altyddroog Farm, 22 ° 11 ’ S 29 ° 53 ’ E, 8. III. 1990, C. D. Eardley leg. (SANC); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.663 ‘ S 21 ° 57.600 ‘ E, 940 m, 10. IX – 4. XI. 2009, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., 1 ♀ (SAMC). UGANDA: Uganda, 1 ♀ (OLL); Kibale National Park, Kanyawara, Makerere Uni- versity Biological Field Station, 0 ° 33.836 ‘ N 30 ° 21.700 ‘ E, 1523 m, 4 – 26. VIII. 2008, YPT, primary mid – altitude rainforest, near stream, S. van Noort leg., 2 ♀♀ (SAMC). YEMEN: 12 km NW Manakhah, 21. VIII – 28. X. 2002, MT, N. 7143, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B10FF3ED312FB5FF8F4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEFA2B10FF3ED312FB5FF8F4.taxon	distribution	Distribution. Botswana, Cameroon, Ethiopia, Madagascar, South Africa, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	description	(Figs 140 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	description	Description. ♀. Fully winged; body length 6.7 – 9.4 mm. Head reddish or reddish-dark, except mandible and clypeus testaceous; antenna black, except scape, pedicel and flagellomeres 6 – 8 testaceous; prothorax reddish or brown-reddish; rest of mesosoma black; metasoma brown; legs brown-black, except tarsi testaceous. In Madagascar there are two different forms: first form almost totally testaceous-reddish, except petiole black, metasoma brown and occasionally mesothoracic axilla blackish; second form almost totally black, except mandible, clypeus, malar space, anterior region of frons near clypeus, lateral regions of pronotum and posterior collar of pronotum testaceous; both forms with antenna testaceous, except antennomere 3 darkened. Antenna clavate; antennomeres of specimen from Zimbabwe, Chishawasha, in following proportions: 8: 6: 36: 20: 17: 11: 9: 9: 7: 7. Head dull, granulate and slightly reticulate rugose; frontal line complete; occipital carina complete; lateral ocelli not touching occipital carina, situated on virtual straight line joining posterior edges of eyes; POL = OL = OPL; TL much longer than OPL. Pronotum (Fig. 140 E) dull, granulate, with many keels around disc; pronotum crossed by strong anterior transverse impression and very strong posterior transverse furrow; anterior collar distinct; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, granulate, usually with numerous slight longitudinal keels (in specimen from Madagascar, 15 ° 55 ’ S 45 ° 57 ’ E, MG- 38 - 20, mesoscutum very slightly granulate and without longitudinal keels; in specimen from Madagascar, 16 ° 08 ’ S 45 ° 42 ’ E, mesoscutum completely unsculptured). Notauli incomplete, approximately reaching 0.65 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, about 3 × as long as propodeal declivity; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein about twice as long as proximal part. Protarsomere 1 about as long as 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 140 F) with one large subdistal tooth and one row of 18 lamellae. Protarsomere 5 (Fig. 140 F) with two rows of about 34 lamellae; apex with about 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Low Congo, Banana, 6. VIII. 1920, H. Schouteden leg. (MRAC). Other material: MADAGASCAR: Antananarivo, Tsimbazaza Park, 1 – 6. XI. 1984, YPT, Robert W. Brooks leg. (NAMUK); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 12 – 19. II. 2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 18, 1 ♀ (CAS); same locality label, 16 ° 08 ’ S 45 ° 42 ’ E, 26. II – 4. III. 2008, MG- 38 - 20, 1 ♀ (MOLC); Toamasina, near Vavatène, Manakam- bahiny, in forest, II. 1995, 1 ♀ (FSAE); Toamasina, 25 km W of Morarano-Chrome, in forest, 10 – 20. IV. 1992, 1 ♀ (FSAE); Toamasina, Alaotra Lake, 3. XII. 1990, 1 ♀ (AMNH); Toamasina, Garden of Alaotra Station, I. 1991, 1 ♀ (FSAE). SOUTH AFRICA: Gauteng, Pretoria, Faerie Glen, 25 ° 46 ’ S 28 ° 17 ’ E, 21. II. 1995, 1 ♀ (CAS). ZIMBABWE: Harare, Chishawasha, V. 1979, A. Watsham leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	distribution	Distribution. Democratic Republic of the Congo, Madagascar, South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF62B12FF3ED03EFC61F9F0.taxon	discussion	Remarks. The ♀ population from Madagascar is apparently characterized by a colour different from that of other African populations. The specimens from continental Africa, have the body partly testaceous-reddish (head and prothorax) and partly black, whereas the Madagascar population has the body totally testaceous-reddish or totally black. Only one totally black specimen is known (from Manakambahiny), whereas five testaceous-reddish specimens are known from Tsimbazaza Park, Alaotra Lake, 25 km W of Morarano-Chrome and Alaotra Station, and from 15 ° 55 ’ S 45 ° 57 ’ E. The small number of known specimens means that we need to postpone assessment of this population. These specimens from Madagascar have a morphology similar to that of the continental population; the only differences are in the colour. Possibly they are a different species. It is prudent to wait for further specimens, before reaching a decision on the status of this population. The testaceous-reddish specimens from Madagascar are 8.1 mm long, whereas the black specimen is 6.7 mm long. The main characters may be summarized as follows: antennomeres in following proportions: 12: 6: 53: 30: 23: 16: 12: 10.5: 8.5: 10; POL = 2.5; OL = 2; OOL = 12; OPL = 2; TL = 6; distal part of 2 r-rs & Rs vein longer than proximal part (19: 10); Protarsomeres in following proportions: 31: 4: 9: 27: 41; enlarged claw with one large subapical tooth and one row of 15 lamellae; protarsomere 5 with two rows of approximately 41 lamellae; apex with at least 20 lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF52B13FF3ED087FE14FD48.taxon	description	(Figs 140 G, 141 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF52B13FF3ED087FE14FD48.taxon	description	Description. ♀. Fully winged; body length 3.7 mm. Head black, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous, except antennomeres 6 – 9 brown; propleuron black; pronotum black, except lateral regions and posterior collar reddish; rest of mesosoma black; metasoma missing in unique known specimen; legs testaceous, except coxae and clubs of femora brown. Antenna clavate; antennomeres in following proportions: 10: 5: 21: 10: 9: 8: 6: 6: 4.5: 8. Head shiny, granulate; occipital carina complete; OL = POL; temple prominent. Prono- tum (Fig. 140 G) shiny, granulate, crossed by strong posterior transverse impression and slight anterior transverse impression; disc humped; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with short keels near posterior margin. Notauli incomplete, approximately reaching 0.5 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum short, with numerous longitudinal keels. Metapectal-propodeal disc re- ticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with three brown trans- verse bands; distal part of 2 r-rs & Rs vein much longer than proximal part. Protarsomere 1 approximately twice as long as 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 141 A) with one small subdistal tooth and one row of three lamellae. Protarsomere 5 (Fig. 141 A) much longer than enlarged claw, with inner margin straight, without distal prominence, with two rows of 22 lamellae extending continuously to apex. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF52B13FF3ED087FE14FD48.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Fianarantsoa, Ranomafana, X. 1938, A. Seyrig leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF52B13FF3ED087FE14FD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF52B13FF3ED087FE14FD48.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF32B14FF3ED6D7FC4CF85D.taxon	description	(Figs 131 A, B, 141 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF32B14FF3ED6D7FC4CF85D.taxon	description	Description. ♀ (Figs 131 A, B). Fully winged; body length 6.2 – 7.2 mm. Head black, except mandible testaceous and clypeus brown with margins testaceous (in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E, clypeus and malar space totally testaceous, frons black, except two short testaceous stripes along orbits); antenna brown, except scape and pedicel partly testaceous and flagellomeres 6 – 8 whitish (in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E, antenna brown, except ventral side of scape and flagellomeres 4 – 8 whitish); mesosoma black; metasoma brown; legs black, except distal extremity of coxae, proximal half of trochanters, proximal extremity of meso- and metatibia, metatarsomere 2 and part of 1 whitish; chelae, meso- and metarsomere 2 and part of 1 testaceous. Antenna clavate; antennomeres in following proportions: 15: 7: 47: 29: 22: 16: 13: 12: 9: 12. ADOs present in antennomeres 6 – 10. Head flat, dull, granulate and sculptured by small areolae and irregular keels; frontal line complete; occipital carina incomplete, only present behind and on sides of lateral ocelli, laterally not reaching eyes; lateral ocelli situated in front of virtual straight line joining posterior edges of eyes; posterior margin of vertex weakly excavated; POL = 3.5; OL = 4; OOL = 10; OPL = 1; temple absent; greatest breadth of lateral ocelli longer than OPL (3.5: 1). in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E, head weakly excavated. In specimen from Zambia, Chati, POL = 4, OL = 4. Pronotum crossed by strong anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar very short; pronotum granulate and sculptured by numerous longitudinal keels and striae (pronotum punctate, unsculptured among punctures in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E); pronotal tubercle not reaching tegula. Mesoscutum of holotype shiny, completely sculptured by numerous irregular, parallel and longitudinal keels. In specimen from Zambia, Chati, median region of mesoscutum reticulate rugose. Notauli apparently complete and posteriorly separated, hardly visible among longitudinal keels. Mesoscutellum and metanotum shiny, smooth, punctate, without sculpure among punctures (granulate in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E). Metapectalpropodeal disc dull, without transverse keels, reticulate rugose and sculptured by few longitudinal keels (completely reticulate rugose in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E), slightly longer than propodeal declivity (15: 10); propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (22: 14). Protarsomeres in following proportions: 28: 5: 8: 23: 37. Enlarged claw (Fig. 141 B) with one large subdistal tooth and one row of 13 – 15 lamellae. Protarsomere 5 (Fig. 141 B) with two rows of 18 – 33 lamellae (6 + 12 in holotype, 8 + 10 in specimen from Zambia, Chati, 33 in specimen from Central African Republic, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E); apex with at least 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF32B14FF3ED6D7FC4CF85D.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Shimba Hills National Park, near artificial pond, 04 ° 22.752 ’ S 39 ° 43.197 ’ E, 335 m, MT, mixed grassland- shrubland, 6 – 20. VI. 2006, R. Copeland leg. (NMK). Other material: BURUNDI: Ruvubu National Park, 2.98144 ° S 30.45531 ° E, 1382 m, 20. I – 4. II. 2010, MT, near Ruvubu River, R. Copeland leg., 1 ♀ (NMK). CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Sangha Dense Forest Special Reserve, 12.7 km 326 ° NW Bayanga, 03 ° 00 ’ 27 ’’ N 16 ° 11 ’ 55 ’’ E, 420 m, 14 – 15. V. 2001, MT, lowland rainforest, S. van Noort leg., 1 ♀ (SAMC). KENYA: Coast Prov., Mrima Hill Forest, 4.48576 ° S 39.25845 ° E, 212 m, 13 – 27. VI. 2011, MT, indigenous forest edge, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Gede Forest, 3.30946 ° S 40.01941 ° E, 19 m, 31. X – 13. XI. 2011, MT, secondary forest, 1 ♀ (NMK); Coast Prov., Kaya Kinondo, 4.39382 ° S 39. 54567 ° E, 10 m, 17 – 31. V. 2011, indigenous forest, MT, coral rag, canopy forest, R. Copeland leg., 1 ♀ (NMK); Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 30. V – 19. VI. 2013, 6 m MT, indigenous forest, R. Copeland leg., 3 ♀♀ (2 in NMK, 1 in MOLC); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 12 – 26. VII. 2014, MT, indigenous forest, R. Copeland leg., 3 ♀♀ (2 in NMK, 1 in MOLC); same locality label, 5 – 19. XI. 2015, 1 ♀ (NMK); Coast Prov., Marenje Forest, 4.52814 ° S 39.24028 ° E, 76 m, 5 – 19. IX. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK). ZAMBIA: Copperbelt Prov., Kitwe, Chati, 6. III. 1980, K. Loyttyniemi leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF32B14FF3ED6D7FC4CF85D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF32B14FF3ED6D7FC4CF85D.taxon	distribution	Distribution. Burundi, Central African Republic, Kenya, Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B15FF3ED6D7FE1EFBB4.taxon	description	(Fig. 141 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B15FF3ED6D7FE1EFBB4.taxon	description	Description. ♀. Fully winged; body length 8.7 mm. Head black, except mandible, clypeus, occiput and anterior region of frons reddish; antenna black, except scape and pedicel reddish, junction between flagellomeres 1 – 2 testaceous, distal extremity and proximal half of flagellomere 2 yellow, flagellomere 8 yellow; prothorax brown-reddish; mesoscutum black, except lateral regions brown-reddish; mesosternum reddish-dark; rest of mesosoma black; metasoma black, except some reddish regions; legs black, except junctions and tarsi reddish. Antenna clavate; antennomeres in following proportions: 10: 7: 58: 33: 24: 14: 11: 8: 8: 10. Head dull, granulate, with tracks of irregular keels on frons; frontal line present; occipital carina complete; POL = 5; OL = 3.5; OOL = 11; OPL = 2; TL = 7; lateral ocelli situated behind virtual straight line joining posterior edges of eyes; posterior margin of vertex convex. Pronotum dull, granulate, with numerous longitudinal striae around disc, crossed by strong posterior transverse impression and strong anterior transverse impression; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate, with numerous slight longitudinal keels. Notauli incomplete, slightly visible, almost reaching posterior margin of mesoscutum. Mesoscutellum dull, granulate and with numerous longitudinal keels. Metanotum dull, reticulate rugose. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, with two incomplete longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part. Protarsomeres in following proportions: 28: 5: 10: 23: 38. Enlarged claw (Fig. 141 C) with one small subdistal tooth and one row of 20 lamellae. Protarsomere 5 (Fig. 141 C) with two rows of 49 lamellae; apex with 14 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B15FF3ED6D7FE1EFBB4.taxon	materials_examined	Material examined. Type: ♀ holotype: SIERRA LEONE: Freetown (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B15FF3ED6D7FE1EFBB4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B15FF3ED6D7FE1EFBB4.taxon	distribution	Distribution. Sierra Leone.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B16FF3ED2CAFE81FB40.taxon	description	(Figs 131 C, D, 141 D, 143 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B16FF3ED2CAFE81FB40.taxon	description	Description. ♀ (Figs 131 C, D). Fully winged; body length 5.6 – 8.4 mm. Usually completely testaceous or ferrugi- nous, except ocellar triangle and petiole black (occasionally ocellar region testaceous or ferruginous, mesothoracic axilla and anterior margin of metapectal-propodeal disc black); antenna completely testaceous, or with antennomeres 4 – 5 partly whitish and 6 – 8 brown; occasionally antenna brown, except scape, pedicel, flagellomeres 8, part of 7, distal third of 2 and proximal half of 3 whitish. Antenna clavate; antennomeres in following proportions: 10: 7: 57: 36: 24: 14: 11: 8: 7: 10. Head (Fig. 143 A) flat, dull, granulate, slightly rugose or sculptured by many longitudinal striae; occipital carina complete, but slightly visible on lateral regions of occiput; POL = 3; OL = 2; OOL = 13; OPL = 2; TL = 7; lateral ocelli situated behind virtual straight line joining posterior edges of eyes; posterior margin of vertex convex. Pronotum dull, granulate and with numerous longitudinal striae on lateral regions; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar very long, much longer than pronotal tubercle; pronotal tubercle reaching tegula. Mesoscutum dull, granulate, with lateral regions sculptured by numerous longitudinal striae; occasionally longitudinal striae present also on median region of mesoscutum. Notauli complete, occasionally slightly visible near mesoscutellum. In specimen from Madagascar, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, notauli reaching about 0.7 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate, occasionally rugose and striate. Metapectal-propodeal disc and propodeal declivity shiny, completely reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part. Protarsomeres in following proportions: 27: 3: 8: 22: 33. Enlarged claw (Fig. 141 D) with one large subdistal tooth and one row of 12 lamellae. Protarsomere 5 (Fig. 141 D) with two rows of approximately 33 lamellae; apex with approximately 23 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B16FF3ED2CAFE81FB40.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Mombasa (USNM). Other material: GHANA: Bia National Park, Research Camp, 23 – 26. VI. 2005, fogging, 1 ♀ (MOLC). GUINEA-BISSAU: Cacheu Region, 2 km E of Bigene, 1. XI. 1992, 1 ♀ (ZIL). MADAGASCAR: Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 1 – 8. VI. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, 152 m, MG- 62 - 25, 1 ♀ (CAS). MOZAMBIQUE: Maputo Prov., Inhaca Island, 15. XI – 3. XII. 1995, Mal- aise trap, M. Olmi leg., 1 ♀ (MOLC); Nampula Prov., Nampula, Neighbourhood of Nampula Catholic Seminary, C. 21. XI. 1999, B. 26. XI. 1999, Sf. 22. XII. 1999, M. Olmi reared from Paranotus rufilineus (Walker), 1 ♀ (MOLC) (Olmi 2004 a). NAMIBIA: Etoshe National Park, Vredekoppies Perdekamp, 19 ° 20 ’ S 16 ° 54 ’ E, 15. I. 1987, J. Irish & E. Marais leg., 1 ♀ (SANC). NIGERIA: Gashaka-Gumti National Park, 25 km SW Serti, 07 ° 20 ’ N 11 ° 13 ’ E, 400 m, 2011, J. Halada leg., 1 ♀ (OLL). SOUTH AFRICA: Gauteng, Pretoria, garden, 8 – 11. II. 2005, YPT, V. Kolyada leg., 1 ♀ (NMSA); KwaZulu-Natal, St. Lucia Estuary, 2 ♀♀ (misid. Dryinus deceptor (Turner) partim: Olmi 1984: 762) (AEIC, AMNH); KwaZulu-Natal, 10 km SE Pietermaritzburg, Ukulinga, 20 – 26. XI. 1985, grassland, MT, R. M. Miller leg., 1 ♀ (DJBC); KwaZulu-Natal, Ramsgate, Butterfly Sanctuary, 30 ° 53.3 ’ S 30 ° 20.4 ’ E, 26 – 28. III. 2004, YPT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Mabibi Campsite, 27 ° 19.818 ’ S 32 ° 44.792 ’ E, 50 m, 18 – 20. X. 2010, YPT, Maputaland Coastal Belt, SA 10 - MAB-Y 02, 1 ♀ (SAMC); Limpopo, Ellisras District, Mogol Na- ture Reserve, 1 ♀ (SANC); Mpumalanga, Krüger National Park, Satara, 1 ♀ (CNC). TANZANIA: Mkomazi Game Reserve, Ibaya Hill, 03 ° 58.70 ’ S 37 ° 47.19 ’ E, 27. XI. 1995, sweep, mk 7, mixed grasses & shrubs in open Combretum bushland, SAM-HYM-PO 16932, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, valley NW of Ibaya Camp, 03 ° 57 ’ S 37 ° 48 ’ E, 20. IV. 1996, sweep, mk 29, grasses & shrubs in riverine bushland in rocky gorge, SAM-HYM-PO 17241, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Ibaya Camp, north west side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 24 – 27. IV. 1996, Acacia / Commiphora / Combretum bushland, YPT, SAM-HYM-PO 19153, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, between Zange gate & Kisiwani, 04 ° 05.99 ’ S 37 ° 54.00 ’ E, 25. XI. 1995, sweep, shrubs & bushes of forest margin along road, SAM-HYM-PO 15584, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, between Ngurungu Dam & Plot, 04 ° 00.73 ’ S 37 ° 54.13 ’ E, 6. XII. 1995, Acacia / Commiphora / Combretum bushland, Sweep, grasses, shrubs, SAM-HYM-PO 14833, S. van Noort leg., 1 ♀ (SAMC); Morogoro, Schmitt leg., 1 ♀ (NHMW). ZIMBABWE: Midlands Province, Gokwe, Sengwa, 1 ♀ (PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B16FF3ED2CAFE81FB40.taxon	biology_ecology	Hosts. Flatidae (Guglielmino et al. 2013): in Mozambique: Paranotus rufilineus (Walker).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF22B16FF3ED2CAFE81FB40.taxon	distribution	Distribution. Ghana, Guinea-Bissau, Kenya, Madagascar, Mozambique, Namibia, Nigeria, South Africa, Tanzania, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF12B17FF3ED357FF17FC04.taxon	description	(Fig. 141 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF12B17FF3ED357FF17FC04.taxon	description	Description. ♀. Fully winged; body length 6.0 – 7.5 mm. Head black, except mandible, clypeus and anterior region of frons (more along orbits) yellow (in specimen from Kenya, 1.00661 ° N 37.38631 ° E, clypeus brown-black); antenna testaceous, except antennomeres 3 and 6 – 8 and dorsal side of scape and pedicel brown (in specimen from Mozambique, 8 km N of Maúa, antenna black, except articulation between flagellomeres 1 and 2, flagellomere 8, distal half of flagellomere 2 and proximal half of flagellomere 3 whitish; in specimen from Uganda, Kampala, antenna brown, except articulation between flagellomeres 1 and 2, distal third of flagellomere 2, flagellomeres 5 and 8 whitish); propleuron black; pronotum black, except margins slightly reddish; rest of mesosoma black; metasoma brown; legs brown-black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 6: 3.5: 32: 19: 12: 6: 6: 4.5: 4: 6. Head dull, granulate and with numerous slight longitudinal keels on frons and vertex; occipital carina complete; lateral ocelli not touching occipital carina, situated behind virtual straight line joining posterior edges of eyes; OPL shorter than OL; POL = OL; TL more than twice as long as POL. In specimen from Kenya, 00 ° 04.364 ’ N 36 ° 97.554 ’ E, POL = 4; OL = 3; OOL = 14; OPL = 1; TL = 9; greatest breadth of lateral ocelli shorter than POL (2: 4). In specimen from Central African Republic, 04 ° 03 ’ N 17 ° 02 ’ E, POL = 4, TL = 7. Pronotum dull, granulate, with numerous keels around disc; pronotum crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar absent; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum and mesoscutellum dull, granulate, sculptured by numerous slight longitudinal keels. Notauli incomplete, almost reaching posterior margin of the mesoscutum. Metanotum rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (10: 4). Protarsomere 1 slightly longer than 4 (15: 13); protarsomere 3 produced into hook. Enlarged claw (Fig. 141 F) with one large subdistal tooth and one row of 9 lamellae (in specimen from Kenya, 00 ° 04.364 ’ N 36 ° 97.554 ’ E, 8 lamellae + 1 seta). Protarsomere 5 (Fig. 141 F) with two rows of about 39 lamellae; apex with at least 60 lamellae. Tibial spurs 1 / 1 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF12B17FF3ED357FF17FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Kivu, Nialopul, 21. III. 1929, A. Collart leg. (MRAC). Other material: BURUNDI: Bururi, 1 ♀ (CAS). CENTRAL AFRICAN RE- PUBLIC: 150 km NWW Mbaiki, 04 ° 03 ’ N 17 ° 02 ’ E, 620 m, 14. VI. 2009, J. Halada leg., 1 ♀ (OLL). KENYA: Rift Valley Prov., Ol Pejeta Conservancy, 00 ° 04.364 ’ N 36 ° 97.554 ’ E, 1825 m, MT, near Pelican Hse, next to Euclea stand, 13 – 27. I. 2006, R. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Matthews Range, 1.00661 ° N 37.38631 ° E, 1006 m, 9 – 23. I. 2016, MT, Sarara campsite, shrubland, R. Copeland leg., 1 ♀ (NMK). MOZAMBIQUE: Niassa Prov., approximately 8 km N of Maúa, Coração Sagrado de Jesús de Maúa Mission, reared from and adult of Elasmoscelis cimicoides Spinola, C. 11. XI. 1999, B. 12. XI. 1999, Sf. 1. XII. 1999, M. Olmi reared, 1 ♀ (MOLC) (Olmi 2004 a). UGANDA: Kampala, IV. 1936, T. H. C. Taylor leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF12B17FF3ED357FF17FC04.taxon	biology_ecology	Hosts. Lophopidae (Guglielmino et al. 2013): in Mozambique: Elasmoscelis cimicoides Spinola.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF12B17FF3ED357FF17FC04.taxon	distribution	Distribution. Burundi, Central African Republic, Democratic Republic of the Congo, Kenya, Mozambique, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	description	(Figs 141 E, 142)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head and mesosoma totally testaceous, except ocellar area black (Fig. 142); head granulate, with lateral ocelli located in front of imaginary straight line joining posterior edges of eyes; notauli incomplete, reaching about 0.65 × length of mesoscutum; mesoscutellum granulate; metapectal-propodeal disc about twice as long as propodeal declivity; protarsomere 1 slightly shorter than 4; enlarged claw (Fig. 141 E) with one large subdistal tooth and one row of 15 lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	description	Description. ♀ (Fig. 142). Fully winged; body length 5.9 – 6.2 mm. Completely testaceous, except ocellar area, petiole and distal extremity of metasoma black. Antenna clavate; antennomeres in following proportions: 12: 7: 43: 24: 18: 12: 9: 7: 6: 9. Head (Fig. 142 E) flat, dull, granulate; frontal line complete; occipital carina complete; lateral ocelli situated behind virtual straight line joining posterior edges of eyes; POL = 3; OL = 2.5; OOL = 12; OPL = 1; TL = 4. Pronotum dull, granulate, with many longitudinal keels on lateral regions; pronotum crossed by strong anterior transverse impression and very strong posterior transverse furrow; anterior collar distinct; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum granulate and reticulate rugose. Notauli incomplete, approximately reaching 0.65 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum dull, granulate and rugose. Metapectal-propodeal disc reticulate rugose, about twice as long as propodeal declivity (30: 17); propodeal declivity reticulate rugose, without longitudinal keels. Forewing (Fig. 142 F) with two dark transverse bands; distal part of 2 r-rs & Rs vein more than twice as long as proximal part (23: 9). Protarsomeres in following proportions: 21: 4: 10: 22: 38. Protarsomere 3 produced into hook. Enlarged claw (Fig. 141 E) with one large subdistal tooth and one row of 15 lamellae. Protarsomere 5 (Fig. 141 E) with two rows of about 36 lamellae; apex with about 30 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	materials_examined	Material examined. Type: ♀ holotype, NMK / INV / T- 644: KENYA: Rift Valley Prov., Matthews Range, 1.00661 ° N 37.38631 ° E, 1006 m, 9 – 23. I. 2016, MT, Sarara campsite, shrubland, R. Copeland leg. (NMK). Paratype: same locality label as holotype, 1 ♀ (MOLC, MO / 345).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEF02B17FF3ED584FA6BF87F.taxon	etymology	Etymology. The new species is named in honour of Rick and Tina Whittle, both for their friendship with one of the authors (RSC) during their Kenya days and in recognition of their passion for exploration and natural history.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	description	(Fig. 143 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	description	Description. ♀. Fully winged; body length 3.6 – 4.6 mm. Head black, except mandible, clypeus, anterior region of frons between antennal toruli and malar spacee testaceous-reddish; ventral side of head brown, except anterior half testaceous-reddish; antenna testaceous, except antennomeres 5 – 9 darkened and 10 whitish; prothorax testaceous- reddish; mesothorax, metathorax, metapectal-propodeal disc, propodeal declivity and petiole black; metasoma and legs testaceous-reddish, except stalks of meso- and metafemur and distal extremities of tibiae darkened. Specimen from Oman, 17 ° 01.02 ’ N 54 ° 09.32 ’ E, (length 3.7 mm) with slightly different colour (head black, except mandible, clypeus and part of malar space testaceous; antenna testaceous, except antennomeres 4 – 9 darkened; pronotum brown, except margins and posterior collar testaceous; legs testaceous, except meso- and metatibia and stalks of meso- and metafemur darkened). Antenna clavate; antennomeres in following proportions: 12: 5: 21: 10: 8: 7: 6: 6: 4: 8. Head flat, dull, covered with many short setae, granulate, with frons sculptured by few irregular and longitudinal striae; frontal line complete; occipital carina complete; POL = 3; OL = 3; OOL = 7; OPL = 1; TL = 4. Pronotum shiny, unsculptured, humped, crossed by slight anterior transverse impression and strong posterior transverse furrow; anterior collar long; posterior collar very short; pronotal tubercle not reaching tegula. Mesoscutum dull, covered with short setae (so dense that sculpture hardly visible); mesoscutum granulate, with lateral regions and posterior surface reticulate rugose. Notauli almost complete and approximately reaching posterior margin of mesoscutum. Mesoscutellum and metanotum dull, granulate, covered with short setae. Metapectal-propodeal disc reticulate rugose, approximately as long as propodeal declivity; propodeal declivity with anterior half reticulate rugose and posterior half sculptured by transverse keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (14: 6). Protarsomeres in following proportions: 16: 3: 6: 11: 18. Enlarged claw (Fig. 143 B) with one small subdistal tooth and one row of 4 – 5 very narrow lamellae similar to bristles. Protarsomere 5 (Fig. 143 B) with three rows of 15 lamellae (one row composed of very long lamellae); apex with at least 18 – 25 lamellae. In specimen from Oman, 17 ° 01.02 ’ N 54 ° 09.32 ’ E, enlarged claw with one small subdistal tooth and seven very narrow lamellae similar to bristles + 1 seta; protarsomere 5 with two rows of 8 (slightly longer) + 10 lamellae and apex with at least 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown (see remarks).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	materials_examined	Material examined. Types: ♀ holotype: YEMEN: Al Lahima, 16. X – 31. XII. 2000, MT, N. 5322, A. van Harten & A. M. Hager leg. (MOLC). Paratypes: YEMEN: 12 km NW of Manakhah, 3. VII – 21. VIII. 2001, MT, N. 5966, A. van Harten leg., 2 ♀♀ (MOLC); Al-Kowd, 1 – 5. IX. 2001, LT, A. van Harten & S. Al Haruri leg., 1 ♀ (MOLC). Other material: YEMEN: 12 km NW of Manakhah, MT, 24. VI – 4. VIII. 2003, N. 7915, A. van Harten leg., 1 ♀ (MOLC); 12 km NW of Manakhah, MT, 15. IX – 22. X. 2003, N. 8167, A. van Harten leg., 3 ♀♀ (MOLC). Palaearctic. Known from Oman (new record): Dhofar, Salalah East, Dahariz, 17 ° 01.02 ’ N 54 ° 09.32 ’ E, 23. IX. 2001, dishes-RP, F. Strumia leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	distribution	Distribution. Yemen, in addition to the Palaearctic Oman.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECF2B2AFF3ED0A2FB53FDB8.taxon	discussion	Remarks. The ♂ is unknown. However, it is possible that the ♂ attributed provisionally to Dryinus cariniceps Cameron is actually a ♂ of D. yemenensis. See remarks under the treatment for D. cariniceps.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECD2B2AFF3ED28EFE1FF86B.taxon	description	(Fig. 143 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECD2B2AFF3ED28EFE1FF86B.taxon	description	Description. ♀. Fully winged; body length 4.1 mm. Head testaceous-reddish, except anterior third of frons and clypeus testaceous; antenna testaceous-reddish; prothorax testaceous-reddish; rest of mesosoma black; metasoma and legs testaceous-reddish. Antenna clavate; antennomeres in following proportions: 6: 6: 30: 14: 12: 9: 8: 7: 6: 8. Head dull, excavated, granulate, with frons sculptured by numerous longitudinal striae; frontal line complete; occipital carina complete; occiput excavated; POL = 2.5; OL = 2.5; OOL = 11; OPL = 1; TL = 5; greatest breadth of lateral ocelli longer than OPL (2: 1). Pronotum dull, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum, mesoscutellum and metanotum setose, dull, granulate and reticulate rugose. Notauli absent. Metapectal-propodeal disc dull, reticulate rugose, without transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, with two distinct dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (16: 5). Protarsomeres in following proportions: 18: 2: 6: 14: 23. Enlarged claw (Fig. 143 C) with one large subdistal tooth and one row of 13 lamellae. Protarsomere 5 (Fig. 143 C) with two rows of approximately 28 lamellae; apex with about 18 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECD2B2AFF3ED28EFE1FF86B.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, 37 km NW Willowmore, Grootrivi- erberg Range, 33 ° 11.46 ’ S 24 ° 09.51 ’ E, 700 m, 19 – 24. XI. 1999, MT, M. E. Irwin et al. leg. (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECD2B2AFF3ED28EFE1FF86B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECD2B2AFF3ED28EFE1FF86B.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2BFF3ED6D7FE1FFC46.taxon	description	(Fig. 143 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2BFF3ED6D7FE1FFC46.taxon	description	Description. ♀. Fully winged; body length 4.4 mm. Head and antenna testaceous reddish; prothorax testaceous-red- dish; rest of mesosoma black; metasoma and legs testaceous-reddish. Antenna clavate; antennomeres in following proportions: 7: 7: 32: 14: 13: 10: 8: 7: 6: 8. Head dull, excavated, completely granulate; frontal line complete; occipital carina complete; occiput excavated; POL = 2; OL = 3; OOL = 11; OPL = 1; TL = 6. Pronotum dull, granulate, with slight striae on lateral regions, crossed by slight anterior transverse impression and strong posterior transverse furrow; disc humped; posterior collar long; pronotal tubercle not reaching tegula. Mesoscutum setose, dull, granulate and slightly rugose. Notauli absent. Mesoscutellum dull, granulate. Metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing almost completely hyaline, only with two slight dark spots; distal part of 2 r-rs & Rs vein longer than proximal part (14: 6). Protarsomeres in following proportions: 20: 3: 7: 15: 25. Enlarged claw (Fig. 143 D) with one large subdistal tooth and one row of 10 lamellae. Protarsomere 5 (Fig. 143 D) with two rows of approximately 25 lamellae; apex with about 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2BFF3ED6D7FE1FFC46.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Limpopo, 30 km E of Nylstroom, 15. XI – 17. XII. 1984, H. and A. Howden leg. (PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2BFF3ED6D7FE1FFC46.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2BFF3ED6D7FE1FFC46.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2DFF3ED25DFD6BFD2C.taxon	description	(Figs 143 E, F, 144 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2DFF3ED25DFD6BFD2C.taxon	description	Description. ♀. Fully winged; body length 3.1 – 6.6 mm. Testaceous, except petiole black, ocellar triangle brown or black and axillae brown or black; occasionally tegula black. Antenna clavate; antennomeres in following proportions: 9: 6: 29: 16: 14: 11: 9: 8: 6.5: 9 (holotype); 9: 4: 16: 7: 7: 6: 5: 5: 4: 6 (in small specimen (4.0 mm) from Namibia, 21 ° 10.77 ’ S 14 ° 32.87 ’ E); ADOs present on antennomeres 5 – 10. Head flat, shiny, reticulate rugose or granulate and with slight tracks of irregular striae; vertex around the occipital carina sculptured by short and slight longitudinal striae; frontal line complete; occipital carina complete; holotype with POL = 3.5; OL = 2.5; OOL = 11; OPL = 2.5; TL = 5; in small specimen (4.0 mm) from Namibia, 21 ° 10.77 ’ S 14 ° 32.87 ’ E, POL = 2.5; OL = 1.5; OOL = 7; OPL = 2; TL = 3; in specimen from Uganda, Kampala, POL = 5; OL = 9. Pronotum (Fig. 143 E) shiny, smooth, slightly humped, crossed by slight anterior and posterior transverse impressions; anterior collar long; posterior collar very short; anterior collar and lateral regions of pronotum with few slight longitudinal striae, unsculptured among striae; disc of pronotum unsculptured; posterior transverse impression of pronotum crossed by short longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum dull, slightly granulate, completely reticulate rugose or with tracks of very slight areolae on lateral regions. Notauli usually absent; occasionally notauli hardly visible, incomplete, approximately reaching 0.65 × length of mesoscutum (notauli hardly visible in large specimens with mesoscutum strongly reticulate rugose; notauli distinct in small specimens with mesoscutum slightly rugose). Mesoscutellum dull, reticulate rugose or slightly granulate. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, without transverse keels, slightly shorter than propodeal declivity (13: 17); propodeal declivity reticulate rugose, with two incomplete longitudinal keels. Forewing hyaline, completely yellowish, usually without dark bands or spots; occasionally forewing with two small dark transverse bands or with one small dark spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein slightly longer than proximal part (10: 7 in holotype; 8: 7 in small specimen (4.0 mm) from Namibia, 21 ° 10.77 ’ S 14 ° 32.87 ’ E). Protarsomeres of holotype in following proportions: 36: 4: 8: 16: 26; 20: 3: 6: 10: 18 in small specimen (4.0 mm) from Namibia, 21 ° 10.77 ’ S 14 ° 32.87 ’ E. Enlarged claw (Fig. 143 F) with one large subdistal tooth and one row of 6 – 8 lamellae. Protarsomere 5 (Fig. 143 F) with 1 – 2 rows of 8 – 13 lamellae; apex with at least 10 – 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 3.0 – 5.2 mm. Head black, except mandible, malar space and anterior part of frons testaceous; antenna testaceous, except antennomeres 3 – 5 darkened; mesosoma black; metasoma black; legs testaceous. Antenna filiform; antennomeres in following proportions: 9: 6: 20: 13: 15: 15: 13: 11: 10: 11. Head dull, reticulate rugose; frontal line complete; occipital carina complete; POL = 9; OL = 4.5; OOL = 6; OPL = 2; TL = 2; temple very short. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose, with areolae less large than ocelli. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose and propodeal declivity, without keels. Forewing hyaline, without dark transverse bands; distal part of 2 rrs & Rs vein longer than proximal part (12: 10). Genital armature (Fig. 144 A) with paramere approximately as long as penis. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2DFF3ED25DFD6BFD2C.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 12 – 30. VI. 1923, R. E. Turner leg. (NHMUK). Paratype: same locality label as holotype, 1 ♀ (NHMUK). Other material: BOTSWA- NA: Central District, Serowe, VIII. 1989, MT, P. Forchhammer leg., 1 ♀ (CAS); same locality label, XI. 1985, 1 ♀ (NHMUK); Central District, Serowe, Farmer’s Brigade, 2 ♀♀ (CNC, USNM); Central District, 25 mi. W Gweta, 20 ° 17 ’ S 24 ° 54 ’ E, 21 – 22. IV. 1972, 1 ♀ (NHMUK); Ghanzi District, Kalahari, Kuke Pan, 2 ♀♀ (AMNH, NHMUK). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Lomami, Upemba Nat. Park, Zaziba, 1 ♀ (MRAC); Orien- tale Prov. (= Haut-Zaire), Garamba National Park, 1 ♀ (MRAC). ETHIOPIA: Amhara, Gelawdios Church Forest, 11 ° 38 ’ N 37 ° 48 ’ E, 2549 m, 9. I. 2012, flight intercept, C. Ozanne leg., 1 ♀ (NHMUK). KENYA: Central Prov., Nai- robi, Karen, 2 ♀♀ (AEIC, AMNH); Eastern Prov., At Athi River, 02 ° 38.51 ‘ S 38 ° 21.98 ‘ E, 19 – 24. VII. 1999, MT, R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Samburu National Reserve, near Ewaso Ng’iro River, 00 ° 56.797 ’ N 37 ° 53.563 ’ E, 874 m, 10 – 24. VII. 2006, MT, riverine forest, next to headquarters, R. Copeland leg., 3 ♀♀ (NMK); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 27. VIII – 10. IX. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♀ (NMK); same locality label, 22. III – 5. IV. 2016, 1 ♀ (NMK); Eastern Prov., Kibwezi Forest, 2.41687 ° S 37.95388 ° E, 945 m, 9 – 23. III. 2016, MT, indigenous forest, R. Copeland leg., 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22328 ° S 36.89674 ° E, 1613 m, 17 – 24. II. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.22317 ° S 36.89653 ° E, 1600 m, 24. II – 10. III. 2014, MT, near stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (NMK); Nyanza Prov., Ungoye Field Station, 0 ° 36 ’ 91 ’’ S 34 ° 05 ’ 52 ’’ E, 1147 m, 27. II – 6. III. 2005, MT, R. S. Copeland leg., 1 ♀ (NMK); same locality label, 6 – 13. III. 2005, 1 ♀ (NMK); Rift Valley Prov., Marich Pass, 1 ° 32 ’ 18 ’’ N 35 ° 27 ’ 48 ’’ E, 917 m, 23. I – 6. II. 2005, MT, R. S. Copeland leg., 1 ♀ (NMK); Rift Valley Prov., Nguruman, near Sampu River, 1.90117 ° S 36.05040 ° E, 723 m, 27. VIII – 10. IX. 2011, MT, near base of Nguruman Escarpment, R. Copeland leg., 1 ♀ (NMK); same locality label, 13 – 27. VIII. 2011, 1 ♀ (NMK); Western Prov., Kakamega District, Kakamega Forest, Kisere For- est, Calebs Campground, 00 ° 22.175 ’ N 34 ° 53.297 ’ E, 1580 m, Manfred Kraemer leg., ex Michael von Tschirnhaus collection, 1 ♀ (MTC). NAMIBIA: Aus, I. 1930, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 1 ♀ (AMNH); Gobabeb, Kuiseb River Bed, 1 ♀ (ZMK); Brandberg, Wasserfallfläche, 21 ° 10.77 ’ S 14 ° 32.87 ’ E, 1960 m, 12. IV. 1999, NA 99 - S 04, sweep grass, shrubs trees, Bushy Karoo-Namib shrubland, S. van Noort leg., 1 ♀ (SMWN); Brandberg, Wasserfallfläche, 21 ° 10.77 ’ S 14 ° 32.87 ’ E, 1960 m, 7 – 10. IV. 1999, NA 99 - M 05, MT, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg., 1 ♀ (SMWN); Brandberg, Hungorob Valley, 21 ° 11.40 ’ S 14 ° 31.69 ’ E, 1180 m, 5 – 16. IV. 1999, NA 99 - M 09, MT, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg., 1 ♀ (MOLC); Brandberg, Messum Valley, 21 ° 13.29 ’ S 14 ° 30.98 ’ E, 700 m, 5 – 17. IV. 1999, NA 99 - M 12, MT, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg., 1 ♀ (SMWN). NIGERIA: Lagos, Ikoyi Park, 9 – 19. I. 1973, J. C. Deeming leg., 1 ♀ (NHMUK). SOMALIA: Benadir Prov., Afgoi, 3 ♀♀ (AMNH). SOUTH AFRICA: Eastern Cape, Port St. Johns, R. E. Turner leg., 1 – 14. V. 1923, 15 – 31. V. 1923, 1 – 11. VI. 1923, 12 – 30. VI. 1923, 1 – 9. VII. 1923, 10 – 31. VII. 1923, 7 – 13. VIII. 1923, 13 ♀♀ (NHMUK); Eastern Cape, Addo Valley, Sundays River Valley, 4. III. 1953, C. N. Smithers leg., 6 ♀♀ (NHMUK); Eastern Cape, Februarie Farm (40.2 km 267 ° W Kirkwood), 33 ° 33.124 ’ S 25 ° 03.043 ’ E, 8 – 10. II. 2001, MT, VB 01 - R 1 N-M 08, Valley bushveld (non-trashed), S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Ithala Game Reserve, Craig Adam Dam, 27 ° 28 ’ S 31 ° 25 ’ E, I – II. 1991, MT, R. M. Miller leg., 1 ♀ (DJBC); KwaZulu-Natal, Umtentweni, VII. 1954, A. L. Capener leg., 1 ♀ (NHMUK); KwaZulu-Natal, Louwsberg, Sanyati Farm, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 11 – 13. VII. 2006, YPT, M. Mostovski leg., 1 ♀ (NMSA); Limpopo, 15 km E Klaserie, Guernsey Farm, 1 ♀ (CNC); Limpopo, Krüger National Park, Levuvhu-Limpopo River, 1 ♀ (AEIC); Limpopo, Krüger National Park, Pafuri, 22 ° 26 ’ S 31 ° 12 ’ E, 20 – 27. X. 1984, 12 – 24. III. 1986, L. E. O. Braack leg., 2 ♀♀ (SANC); Limpopo, Krüger National Park, Pafuri, 22 ° 26 ’ S 31 ° 12 ’ E, 264 m, 20 – 24. I. 1985, G. L. Prinsloo leg., 2 ♀♀ (SANC); Western Cape, Gamkaberg Nature Reserve, 33 ° 39.570 ’ S 21 ° 53.957 ’ E, 328 m, 4. XI. 2009 – 19. II. 2010, MT, Gamka Thicket, S. van Noort leg., 1 ♀ (SAMC); same locality label, 31. VII – 10. IX. 2009, 2 ♀♀ (SAMC). TANZA- NIA: Mkomazi Game Reserve, Ibaya Camp, north west side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 27 – 28. IV. 1996, Acacia / Commiphora / Combretum bushland, YPT, SAM-HYM-PO 19510, S. van Noort leg., 1 ♀ (SAMC). UGANDA: Kampala, 1 ♀ (AEIC). ZIMBABWE: Harare, 1 ♀ (NHMUK); same locality label, IV. 1982, A. Watsham leg., 1 ♀ (DJBC); same locality label, X. 1981, A. Watsham leg., 1 ♀ (DJBC); near Harare, VIII. 1982, YPT, A. Watsham leg., 1 ♀ (NHMUK); Harare, Chishawasha, XI. 1980, A. Watsham leg., 1 ♀ (NHMUK); same locality label, XII. 1980, 1 ♀ (NHMUK); same locality label, X. 1982, 1 ♀, 1 ♂ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2DFF3ED25DFD6BFD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FECC2B2DFF3ED25DFD6BFD2C.taxon	distribution	Distribution. Botswana, Democratic Republic of the Congo, Ethiopia, Kenya, Namibia, Nigeria, Somalia, South Africa, Tanzania, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	description	(Figs 144 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	description	Description. ♀. Fully winged; body length 4.6 – 5.1 mm. Head black, except mandible, clypeus and some × malar space and narrow stripe between antennal toruli and between eyes and antennal toruli testaceous; antenna totally testaceous, or brown, except scape and pedicel testaceous, or testaceous, except flagellomeres 1 – 2 slightly darkened; mesosoma black (occasionally pronotum with lateral margins, posterior collar and pronotal tubercle testaceous); metasoma brown or brown – testaceous; legs testaceous, except coxae, clubs of femora and protrochanter partly brown (occasionally legs brown, except coxae, trochanters, clubs of femora and tibiae partly testaceous; last tarsomere of meso- and metalegs brown); in specimen from South Africa, 33 ° 30.747 ’ S 25 ° 24.644 ’ E, legs testaceous, except metacoxa brown. Antenna clavate; antennomeres of holotype of D. ugandanus in following proportions: 13: 6: 28: 14: 10: 9: 6: 6: 6: 8; antennomeres of holotype of D. constantiae in following proportions: 13: 5: 32: 21: 16: 12: 9: 8: 6: 9. ADOs present on antennomeres 5 – 10. Head dull, slightly convex or flat, granulate and completely reticulate rugose (with region of vertex along occipital carina sculptured by numerous short longitudinal keels); frontal line complete; occipital carina complete; holotype of D. ugandanus with POL = 4; OL = 2; OOL = 9; OPL = 3; TL = 4; holotype of D. constantiae with POL = 3.5; OL = 3; OOL = 9; OPL = 3; TL = 6. Pronotum dull, granulate (occasionally partly unsculptured or punctate), with slight irregular striae, crossed by transverse furrow; disc slightly humped; posterior collar very short or absent; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose. In holotype of D. constantiae, mesoscutum with median region almost completely granulate, not reticulate rugose. Notauli absent in holotype of D. ugandanus (in holotype of D. constantiae, notauli incomplete, reaching about 0.5 × length of mesoscutum; in specimen from South Africa, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, notauli almost reaching posterior margin of mesoscutum). Mesoscutellum granulate and reticulate rugose (in holotype of D. constantiae, mesoscutellum granulate, not rugose). Metanotum granulate and rugose. Metapectal-propodeal disc reticulate rugose, approximately as long as propodeal declivity; propodeal declivity reticulate rugose, with two short incomplete longitudinal keels. Forewing with 1 – 2 transverse bands very slightly darkened (in specimen from South Africa, 33 ° 30.747 ’ S 25 ° 24.644 ’ E, forewing hyaline, without dark transverse bands or spots; in specimen from South Africa, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, forewing hyaline, with very small dark spot around 2 r-rs & Rs vein); distal part of 2 r-rs & Rs vein longer than proximal part (15: 9 in holotype of D. ugandanus; 13: 9 in holotype of D. constantiae). Protarsomeres of holotype of D. ugandanus in following proportions: 26: 4: 7: 11: 20. Protarsomeres of holotype of D. constantiae in following proportions: 33: 3: 6: 12: 20. Enlarged claw (Figs 144 B, C) with one large subdistal tooth and one row of 5 – 8 lamellae. Protarsomere 5 (Figs 144 B, C) with one row of 8 – 13 lamellae; apex with about 9 – 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	materials_examined	Material examined. Types: ♀ holotype of T. ugandanus: UGANDA: Kampala (AEIC); ♀ holotype of D. constantiae: SOUTH AFRICA: Western Cape, Cape Town, above Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’’ E, 460 m, 2 – 9. II. 1994, MT, Mesic mountain Fynbos on Sandstone, Protea dominated, SAM-HYM-PO 03806, S. van Noort leg. (SAMC). Paratypes of T. ugandanus: SIERRA LEONE: Freetown, 1 ♀ (AEIC); SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, 1 ♀ (AMNH). Other material: SOUTH AFRICA: Eastern Cape, Blauwe Krans Farm, 12.8 km 216 ° SW Kirkwood, 33 ° 30.747 ’ S 25 ° 24.644 ’ E, 9 – 16. II. 2001, YPT, Valley Bushveld (non-trashed), S. van Noort leg., 1 ♀ (SAMC); Northern Cape, Swaarweerberg, Vredehoek Farm, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, 1613 m, 30. IX – 30. XII. 2010, MT, Succulent Karoo, S. van Noort leg., SWA 09 - SUC 1 - M 06, 1 ♀ (MOLC); Western Cape, Cape Good Hope Nature Reserve, 7 – 10. III. 1968, 1 ♀ (SAMC); Western Cape, Cape Town, above Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 17 – 27. III. 1995, MT, Mesic mountain Fynbos Protea coronata dominated, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	distribution	Distribution. Sierra Leone, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC92B2FFF3ED6D7FECAFF50.taxon	discussion	Remarks. Dryinus ugandanus is here considered as a senior synonym of D. constantiae, because a study of ♀♀ attributed to D. constantiae showed a variability concerning the notauli. Some specimens have the notauli almost or totally invisible, as in D. ugandanus, so that they can be identified as D. ugandanus and attributed to group 2. For this reason, D. ugandanus can be attributed to both groups, 1 and 2. The conspecificity of these two species is highly likely.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC82B20FF3ED0C3FE14FCBC.taxon	description	(Fig. 144 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC82B20FF3ED0C3FE14FCBC.taxon	description	Description. ♀. Fully winged; body length 5.0 – 5.2 mm. Head black, except mandible and clypeus testaceous; antenna testaceous; propleuron black; pronotum black, except margins and posterior collar reddish; rest of mesosoma black; metasoma brown; legs testaceous, except coxae and clubs of femora brown; occasionally tibiae brown. Antenna clavate; antennomeres in following proportions: 9: 4: 25: 13: 9: 5.5: 5: 5: 4.5: 6. Head dull, granulate; occipital carina complete; lateral ocelli touching occipital carina; holotype with POL = OL; in specimen from Madagascar, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, POL = 4; OL = 3; OOL = 7; OPL = 0; TL = 3; greatest breadth of lateral ocelli about as long as OL. Pronotum dull, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum shiny, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum and metanotum dull, granulate. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein about twice as long as proximal part; in specimen from Madagascar, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, 2 r-rs & Rs vein with distal part less than twice as long as proximal part (9: 6). Holotype with protarsomere 1 longer than 4 (17: 10); protarsomere 3 produced into hook. In specimen from Madagascar, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, Protarsomeres in following proportions: 18: 3: 6: 10: 18. Enlarged claw (Fig. 144 E) with two small subdistal teeth and one row of five lamellae. Protarsomere 5 (Fig. 144 E) with two rows of approximately 15 – 17 lamellae; apex with approximately 9 – 19 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC82B20FF3ED0C3FE14FCBC.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, X. 1938, A. Seyrig leg. (MNHN). Paratype: same locality label as holotype, IV. 1937, 1 ♀ (MRAC). Other material: MADAGASCAR: Mahajanga, Namoroka National Park, 17.8 km 329 ° WNW Vilanandro, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, 100 m, 8 – 12. XI. 2002, YPT, in tropical dry forest, Fisher, Griswold et al. leg., BLF 6511, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC82B20FF3ED0C3FE14FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC82B20FF3ED0C3FE14FCBC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC72B21FF3ED5C3FC9CFEE0.taxon	description	(Figs 144 D, 145 A, B, 147 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC72B21FF3ED5C3FC9CFEE0.taxon	description	Description. ♀ (Figs 145 A, B). Fully winged; body length 5.3 – 6.9 mm. Head black, except mandible and clypeus ferruginous; propleuron black; pronotum black, except margins and posterior collar reddish; rest of mesosoma black; metasoma black-brown, some × with apex reddish; legs testaceous, except coxae and clubs of femora black or brown. Antenna clavate; antennomeres in following proportions: 16: 6: 27: 19: 14: 11: 9: 9: 8: 10. Head shiny, completely reticulate rugose; occipital carina complete; OL shorter than POL; OPL shorter than OL. In specimen from Kenya, 00 ° 61.517 ’ S 34 ° 09.200 ’’ E, POL = 6; OL = 5; OOL = 11; OPL = 2; temple absent; greatest breadth of lateral ocelli longer than OPL (3: 2). Pronotum dull, granulate, not humped, crossed by slight anterior and posterior transverse impressions; posterior collar short, slightly visible; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli absent. Mesoscutum with two longitudinal keels on place of notauli and reaching approximately 0.5 × length of mesoscutum. Mesoscutellum dull, granulate and partly reticulate rugose. Metanotum completely reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two long longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (20: 13). In holotype, protarsomere 1 about as long as 4; protarsomere 3 produced into hook. In specimen from Kenya, 00 ° 61.517 ’ S 34 ° 09.200 ’ E, Protarsomeres in following proportions: 16: 4: 8: 16: 31. Protrochanter about twice as long as broad. Enlarged claw (Fig. 144 D) without subdistal teeth and with three distal lamellae. Protarsomere 5 (Fig. 144 D) with two rows of approximately 42 lamellae extending continuously to apex (one row is composed of very long lamellae). Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 4.4 mm. Head black, except mandible and clypeus testaceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae and clubs of femora black. Antenna filiform; antennomeres in following proportions: 9: 6: 20: 13: 13: 14: 12: 11: 9: 12. Head shiny, reticulate rugose; occipital carina incomplete; lateral ocelli not touching occipital carina; OL shorter than POL; OPL shorter than OL; temple very short. Mesoscutum shiny, reticulate rugose. Notauli absent. Mesoscutellum dull, granulate, with some areolae. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (17: 9). Genitalia in Fig. 147 A. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC72B21FF3ED5C3FC9CFEE0.taxon	materials_examined	Material examined. Types: ♀ holotype: SIERRA LEONE: Njala, VII. 1928, E. Hargreaves leg. (NHMUK). Paratypes: CENTRAL AFRICAN REPUBLIC: Boukoko, 1 ♀, 1 ♂ (MNHN). SIERRA LEONE: same locality label as holotype, 11. VII. 1925, on cotton leaf, 1 ♀ (NHMUK); Freetown, 1 ♀ (AEIC). Other material: KENYA: Nyanza Prov., Ungoye, ICIPE Field Station, 00 ° 61.517 ’ S 34 ° 09.200 ’ E, 1147 m, MT, inside seasonally swampy forest, 25. V – 8. VI. 2006, R. Copeland leg., 1 ♀ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC72B21FF3ED5C3FC9CFEE0.taxon	biology_ecology	Hosts. In Central African Republic: unidentified Dictyopharidae (Olmi 1984).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC72B21FF3ED5C3FC9CFEE0.taxon	distribution	Distribution. Central African Republic, Kenya, Sierra Leone.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC62B22FF3ED0FDFE14FCD8.taxon	description	(Figs 145 C, D, 146 A, B, 147 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC62B22FF3ED0FDFE14FCD8.taxon	description	Description. ♀ (Figs 145 C, D). Fully winged; body length 6.2 – 6.5 mm. Head black, except mandible and clyp- eus testaceous; antenna testaceous, except antennomeres 8 – 10 darkened; mesosoma black; metasoma black; legs brown-testaceous, except coxae and clubs of femora dark. Antenna clavate; antennomeres in following proportions: 14: 7: 33: 14: 12: 10: 9: 9: 8: 10. Head dull, setose, rugose and sculptured by short irregular keels; occipital carina complete, laterally reaching posterior margin of eyes; temple absent. Pronotum shiny, unsculptured, setose. Mesoscutum shiny, unsculptured. Notauli absent. Mesoscutellum unsculpured. Metanotum smooth, with some areolae near anterior margin. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two long longitudinal keels and median area rugose. Forewing hyaline, without dark transverse bands; pterostigma about 5 × as long as broad (14: 2.5). Protarsomere 1 shorter than 4 (11: 14). Protrochanter about twice as long as broad (8: 4). Enlarged claw (Fig. 147 B) without subdistal teeth and lamellae, with many bristles. Protarsomere 5 (Fig. 147 B) with 2 – 3 rows of approximately 63 lamellae extending continuously to apex. Tibial spurs 1 / 1 / 2. ♂ (Figs 146 A, B): fully winged; body length 3.1 mm. Head black, except mandible yellow; antenna with antennomeres 1 – 6 yellow (antennomeres 7 – 10 missing in unique known ♂); mesosoma black; metasoma brownblack; legs yellow. Antennomeres 1 – 6 in following proportions: 5: 4: 14: 10: 10: 9. Head reticulate rugose; mandible tridentate, with one rudimentary intermediate tooth (Fig. 8 D); occipital carina complete, reaching posterior margins of eyes; temple absent. Mesoscutum reticulate rugose. Notauli very slightly visible only on anterior half of mesoscutum. Mesoscutellum granulate. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; pterostigma more than 3 × as long as broad (25: 7). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC62B22FF3ED0FDFE14FCD8.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, III. 1951, A. Seyrig leg. (MNHN). Paratypes: same locality label as holotype, X. 1936, 1 ♀ (MRAC), 1 ♂ allotype (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC62B22FF3ED0FDFE14FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC62B22FF3ED0FDFE14FCD8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC42B24FF3ED331FDD4FDB8.taxon	description	(Fig. 147 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC42B24FF3ED331FDD4FDB8.taxon	description	Description. ♀. Fully winged; body length 5.1 – 5.3 mm. Head black, except mandible, clypeus and some small spots on malar space testaceous; antenna ferruginous, except antennomeres 6 – 9 darkened; propleuron black; pronotum black, except margins and posterior collar reddish; rest of mesosoma black; metasoma black; legs ferruginous, except coxae partly black and clubs of femora partly darkened. In specimen from Kenya, 04 ° 23.783 ’ S 39 ° 39.567 ’ E, malar space completely black, antenna brown, except scape and pedicel partly testaceous and flagellomeres 7 – 8 totally testaceous, legs brown, except tarsi partly testaceous. Antenna clavate; antennomeres in following proportions: 14: 6: 28: 14: 11: 9: 7: 6: 5: 9. Head flat, dull, granulate, with irregular keels on frons and vertex; frontal line complete; occipital carina complete; lateral ocelli touching occipital carina; POL = 5; OL = 3; OOL = 9; OPL = 0; TL = 4. In specimen from Kenya, 04 ° 23.783 ’ S 39 ° 39.567 ’ E, lateral ocelli very close to occipital carina, not touching it. Pronotum shiny, alutaceous, crossed by slight anterior transverse impression and strong posterior transverse furrow; posterior collar short; disc humped; pronotal tubercle not reaching tegula. In specimen from Kenya, 04 ° 23.783 ’ S 39 ° 39.567 ’ E, pronotum dull, granulate and sculptured by longitudinal striae around and on disc. Mesoscutum dull, reticulate rugose. Notauli incomplete, reaching approximately 0.6 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum dull, rugose. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal and transverse keels. Forewing with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (15: 9). Protarsomeres in following proportions: 20: 5: 9: 20: 33. Protarsomere 3 produced into hook. Enlarged claw (Fig. 147 C) with two small subdistal teeth and one row of 13 – 18 lamellae. Protarsomere 5 (Fig. 147 C) with two rows of approximately 23 – 28 lamellae; apex with approximately 30 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC42B24FF3ED331FDD4FDB8.taxon	materials_examined	Material examined. Type: ♀ holotype: BOTSWANA: Serowe, Farmer’s Brigade, IX. 1988, MT, Per Forchham- mer leg. (USNM). Other material: KENYA: Coast Prov., Shimba Hills National Park, 04 ° 23.783 ’ S 39 ° 39.567 ’ E, 436 m, MT, just inside Makadara Forest, 22. X – 5. XI. 2008, R. Copeland leg., 1 ♀ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC42B24FF3ED331FDD4FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC42B24FF3ED331FDD4FDB8.taxon	distribution	Distribution. Botswana, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	description	(Figs 147 D, 148)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	diagnosis	Diagnosis. ♀ of Dryinus with head granulate, not sculptured by irregular keels; notauli absent; forewing with two dark transverse bands; enlarged claw (Fig. 147 D) spatulate, with one long row of 13 teeth extending from proximal region to apex and one row of 11 lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	description	Description. ♀ (Fig. 148). Fully winged; body length 3.4 mm. Head black, except mandible testaceous and clypeus brown; antenna brown, except scape and pedicel testaceous; mesosoma black, except lateral margins of pronotum testaceous; metasoma brown; legs testaceous, except metatibia and inner side of metafemur club darkened. Antenna clavate; antennomeres in following proportions: 11: 5: 22: 12: 10: 8: 8: 6: 5: 8. Head weakly excavated, dull, granulate; occiput slightly granulate; frontal line complete; occipital carina incomplete, only present behind and on sides of lateral ocelli; temple absent; lateral ocelli touching occipital carina; POL = 5; OL = 3; OOL = 8; greatest breadth of posterior ocellus shorter than OL (2: 3). Pronotum dull, granulate, crossed by deep anterior transverse furrow and strong posterior transverse furrow; posterior collar short; disc humped; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli absent. Mesoscutum with two longitudinal keels similar to notauli. Mesoscutellum dull, granulate and slightly rugose. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 6). Hind wing hyaline. Protarsomeres in following proportions: 18: 3: 7: 13: 22. Enlarged claw (Figs 147 D, 148 D) spatulate, with one long row of 13 teeth extending from proximal region to apex and one row of 11 lamellae. Protarsomere 5 (Figs 147 D, 148 D) with three rows of 5 (very long) + 27 lamellae; apex with group of about 13 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Gamkaberg Nature Reserve, 33 ° 39.941 ’ S 21 ° 53.505 ’ E, 315 m, 8. II – 21. III. 2009, MT, Gamka Thicket, S. van Noort leg., SAM-HYM-A 023302 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC32B25FF3ED4DFFE9EFC4C.taxon	etymology	Etymology. The species is named dentatiforceps (= pince with teeth) because of the long row of teeth present in the enlarged claw.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC22B25FF3ED253FE1FF80B.taxon	description	(Fig. 149 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC22B25FF3ED253FE1FF80B.taxon	description	Description. ♀. Fully winged; body length 3.2 mm. Head black, except mandible testaceous; antenna with scape and pedicel testaceous and flagellomere 1 black (other flagellomeres missing in unique known specimen); mesosoma black; metasoma missing in unique known specimen; legs brown, except tarsus and protibia testaceous. First three antennomeres in following proportions: 11: 5: 23 (other antennomeres missing in unique known specimen). Head flat, dull, granulate and reticulate rugose; frontal line complete; occipital carina incomplete, laterally not reaching eyes; temple absent; lateral ocelli touching occipital carina; POL = 6; OL = 4; OOL = 9. Pronotum dull, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; posterior collar absent; disc humped; pronotal tubercle almost reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli absent. Mesoscutellum dull, granulate. Metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose; propodeal declivity with central area sculptured by irregular and longitudinal keels (area surrounded by distinct keel). Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 6). Protarsomeres in following proportions: 13: 3: 6: 9: 20. Enlarged claw (Fig. 149 A) without subdistal teeth, with one row of eight bristles. Protarsomere 5 (Fig. 149 A) with two rows of approximately 18 lamellae; apex with approximately 12 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC22B25FF3ED253FE1FF80B.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, 24 km NE of Montagu, Ouberg Pass, 33 ° 48 ’ S 20 ° 12 ’ E (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC22B25FF3ED253FE1FF80B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC22B25FF3ED253FE1FF80B.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC12B27FF3ED032FE1FFC04.taxon	description	(Figs 146 C, D, 149 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC12B27FF3ED032FE1FFC04.taxon	description	Description. ♀ (Figs 146 C, D). Fully winged; body length 5.3 mm. Head testaceous-reddish; antenna testaceous, except antennomeres 8 – 10 brown (antennomeres 6 – 7 missing in holotype); mesosoma testaceous-reddish, except fore axilla black and small median black spot near anterior margin of mesoscutum; metasoma testaceous-reddish; legs testaceous. Antenna clavate; antennomeres in following proportions: 17: 8: 26: 15: 13:?:?: 8.5: 7: 9 (antennomeres 6 – 7 missing in holotype); ADOs present in antennomeres 8 – 10 (antennomeres 6 – 7 missing in holotype). Head slightly convex (Fig. 146 C), dull, setose, granulate and reticulate rugose; frontal line incomplete, not visible in an- terior third of frons; occipital carina incomplete, not visible on temple, laterally not reaching eyes; occipital carina not visible on sides of occiput; POL = 7; OL = 3.5; OOL = 10; OPL = 2; TL = 1. Propleuron almost completely hidden under pronotum; pronotum dull, setose, granulate, slightly humped, crossed by anterior slight transverse impression and posterior deep furrow; posterior collar of pronotum very short, almost absent; pronotal tubercle not reaching tegula. Mesoscutum dull, setose, completely granulate, with lateral regions sculptured by many transverse keels. Notauli almost absent (two very short tracks of notauli shortly visible near anterior margin of mesoscutum). Mesoscutellum dull, granulate. Metanotum dull, reticulate rugose. Metapectal-propodeal disc completely reticulate rugose, shorter than propodeal declivity (20: 25); propodeal declivity with two longitudinal keels, with median area dull, reticulate rugose (with areolae less strong than those of lateral regions). Mesopleuron dull, reticulate rugose. Metapleuron dull, granulate and transversely striate. Forewing with one small dark spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein slightly longer than proximal part (13: 11). Protarsomeres in following proportions: 14: 4: 7: 13: 26. Enlarged claw (Fig. 149 B) without subdistal teeth, with one row of seven bristles in addition to two distal lamellae. Protarsomere 5 (Fig. 149 B) with two rows of approximately 30 lamellae present continuously to apex (shortest row composed of three very long lamellae, longest row composed of short lamellae forming three rows at apex. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC12B27FF3ED032FE1FFC04.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Limpopo, Guernsey (central farm) near Klaserie, 24 ° 28.8 ’ S 31 ° 05.4 ’ E, 550 m, 18. I. 1997, parasitoid of Homoptera on Acacia nilotica, Grobbelaar / Millar / Stals leg. (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC12B27FF3ED032FE1FFC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC12B27FF3ED032FE1FFC04.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC02B38FF3ED59BFD18FEC4.taxon	description	(Figs 149 C – E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC02B38FF3ED59BFD18FEC4.taxon	description	Description. ♀. Fully winged; body length 3.7 – 4.4 mm. Head black, with mandible testaceous; occasionally clyp- eus partly testaceous; antenna brown, except antennomeres 1, 2 and part of 3 testaceous; mesosoma black, except lateral margins of pronotum testaceous; metasoma black or brown, except distal extremity testaceous; legs testaceous, except clubs of femora darkened, meso- and metacoxa partly black and last tarsomere of meso- and metalegs brown. Antenna clavate; antennomeres in following proportions: 8: 5: 25: 13: 10: 9: 7: 7: 5: 7. Head flat, shiny, granulate and completely sculptured by irregular keels; occiput granulate; frontal line complete; occipital carina incomplete, only present behind and on sides of lateral ocelli; temple absent; lateral ocelli touching occipital carina; POL = 4; OL = 3; OOL = 7. Pronotum shiny, granulate, crossed by slight anterior transverse impression and strong posterior transverse furrow; posterior collar short; disc humped; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate and reticulate rugose. Notauli absent. Mesoscutum with two longitudinal keels similar to notauli. Mesoscutellum dull, granulate. Metanotum reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with two dark transverse bands (in specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, forewing with one dark transverse band around 2 r-rs & Rs vein); distal part of 2 r-rs & Rs vein longer than proximal part (14: 7 in holotype; 8: 7 in specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E). Hind wing hyaline. Protarsomeres in following proportions: 15: 3: 5: 13: 21. Enlarged claw (Fig. 149 C) spatulate, with seven very small teeth around apex (in specimen from Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, enlarged claw with 11 medial and distal teeth (Fig. 149 D), in addition to one row of 6 – 7 proximal and medial lamellae; in specimens from Tanzania, 03 ° 58.57 ’ S 37 ° 49.34 ’ E, and Madagascar, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, enlarged claw with two distal lamellae (Fig. 149 E )). Protarsomere 5 (Figs 149 C-E) with two rows of approximately 2 (very long) + 18 – 22 lamellae extending continuously to apex. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC02B38FF3ED59BFD18FEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: IVORY COAST: Vallée du Bandama, Katiola Dept., 20 km E Katiola, Logbonou, MT, savanna, 8. V. 1981, J. W. Everts leg. (DEUW). Other material: MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 28. III – 8. IV. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 20, 1 ♀ (CAS). TANZANIA: Mkomazi Game Reserve, Mzukune River in Mbono Valley, 03 ° 58.57 ’ S 37 ° 49.34 ’ E, 27. IV. 1996, sweep, mk 37, grasses & shrubs, riverine bushland along dry river corse, SAM-HYM-PO 20414, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC02B38FF3ED59BFD18FEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEC02B38FF3ED59BFD18FEC4.taxon	distribution	Distribution. Ivory Coast, Madagascar, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B38FF3ED7DBFE14FB62.taxon	description	(Fig. 149 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B38FF3ED7DBFE14FB62.taxon	description	Description. ♀. Fully winged; body length 4.2 mm. Head black, except mandible, part of clypeus and part of malar space testaceous; antenna testaceous, except antennomeres 6 – 10 darkened; propleuron black; pronotum black, except lateral regions and pronotal tubercle testaceous; rest of mesosoma black; metasoma brown; legs brown, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 25: 13: 12: 8: 7: 6: 5: 8; ADOs in antennomeres 5 – 10. Head dull, smooth, granulate; frontal line complete; occipital carina incomplete, laterally reaching eyes; temple absent; POL = 5; OL = 3; OOL = 6; OPL = 1. Pronotum dull, granulate, crossed by slight anterior transverse impression; disc slightly humped; pronotal tubercle not reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Mesoscutellum dull, granulate. Metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose; propodeal declivity with two complete longitudinal keels, with median area granulate and lateral areas rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 8). Protarsomeres in following proportions: 17: 3: 7: 17: 30. Protarsomere 3 produced into hook. Enlarged claw (Fig. 149 F) without subdistal teeth, with one row of 6 bristles, with apex rounded and provided with two lamellae and few bristles. Protarsomere 5 (Fig. 149 F) with two rows of 8 (longer) + 15 (shorter) lamellae; apex with approximately 16 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B38FF3ED7DBFE14FB62.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, 12 km NW of Amboasary, Berenty, 5 – 15. V. 1983, J. S. Noyes & M. C. Day leg., B. M. 1983 – 201 (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B38FF3ED7DBFE14FB62.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B38FF3ED7DBFE14FB62.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B39FF3ED376FE14FEC4.taxon	description	(Fig. 150 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B39FF3ED376FE14FEC4.taxon	description	Description. ♀. Fully winged; body length 10.6 mm. Head and prothorax testaceous-reddish; antenna testaceous, except antennomeres 7 – 10 and distal part of antennomere 6 darkened; rest of mesosoma black; metasoma black; legs testaceous. Antenna clavate; antennomeres in following proportions: 18: 7: 54: 25: 20: 16: 12: 11: 9: 11. Head flat, dull, reticulate rugose; frontal line complete; occipital carina incomplete, laterally reaching eyes; occipital carina not visible on sides of occiput, present on ventral side of head; POL = 8; OL = 3; OOL = 10; OPL = 2; temple absent. Pronotum shiny, slightly granulate, with disc very humped, crossed by anterior transverse impression; anterior collar with anterior margin emarginated; posterior collar absent; pronotal tubercle not reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli absent. Mesoscutum with two longitudinal keels, reaching approximately 0.5 × length of mesoscutum and similar to notauli. Metapectal-propodeal disc dull, reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing yellow, with one black transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein longer than proximal part (23: 16). Hind wing hyaline, with distal region darkened. Protarsal in following proportions: 24: 4: 13: 28: 50. Enlarged claw (Fig. 150 A) spatulate, without teeth and lamellae. Protarsomere 5 (Fig. 150 A) with some rows of approximately 72 lamellae; apex with approximately 18 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B39FF3ED376FE14FEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Berenty, 12 km NW Amboasari, 5 – 15. V. 1983, J. S. Noyes leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B39FF3ED376FE14FEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDF2B39FF3ED376FE14FEC4.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	description	(Figs 150 B, 151)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	diagnosis	Diagnosis. ♀ of Dryinus with mesoscutum granulate, not reticulate rugose; notauli incomplete, reaching approximately 0.7 × length of mesoscutum; enlarged claw with apex rounded (Fig. 150 B), without subdistal teeth, with one row of lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	description	Description. ♀ (Fig. 151). Fully winged; body length 4.2 – 4.3 mm. Head black, except mandible and clypeus ferruginous (in paratype, clypeus brown, except lateral regions ferruginous); antenna testaceous; mesosoma black, except posterior collar and lateral and anterior margins of pronotum testaceous; metasoma brown; legs testaceous- darkened. Antenna clavate; antennomeres in following proportions: 9: 6: 24: 13: 10: 8: 6: 5: 5: 8; ADOs on antennomeres 6 – 10. Head (Fig. 151 A, C) dull, granulate; frontal line complete; occipital carina complete; temple distinct; lateral ocelli touching occipital carina; POL = 3; OL = 2; OOL = 8; TL = 3; greatest breadth of lateral ocelli about as long as OL. Pronotum granulate, except lateral regions partly unsculptured, crossed by slight anterior transverse impression; disc slightly humped; posterior collar short; pronotal tubercle not reaching tegula. Mesoscutum and mesoscutellum dull, granulate (Fig. 151 A). Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose, about as long as propodeal declivity; propodeal declivity reticulate rugose, without longitudinal keels. Forewing (Fig. 151 D) with three dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 6). Protarsomeres in following proportions: 16: 3: 6: 18: 30. Protarsomere 3 produced into hook. Enlarged claw (Fig. 150 B) without subdistal teeth, with one row of 10 lamellae, with apex rounded and not provided with lamellae. Protarsomere 5 (Fig. 150 B) with two rows of 8 (longer) + 16 (shorter) lamellae (5 (longer) + 14 (shorter) in paratype); apex with approximately 25 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	materials_examined	Material examined. Types: ♀ holotype (CASTYPE 19459): MADAGASCAR: Toliara, Fiherenana, 23 ° 14.07 ’ S 43 ° 52.15 ’ E, 50 m, 1 – 4. XII. 2002, MT, degraded gallery forest, GF 051, Frontier Wilderness Project (CAS). Paratype: MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 22 – 29. VI. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 30, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDE2B3AFF3ED0F9FED8F877.taxon	etymology	Etymology. The species is named teres (latin adjective meaning “ rounded ”) because of the rounded apex of the enlarged claw.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDC2B3CFF3ED6D4FF2BFE0C.taxon	description	(Figs 150 C, 152)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDC2B3CFF3ED6D4FF2BFE0C.taxon	diagnosis	Diagnosis. ♀ of Dryinus with OL more than 4 × as long as POL; notauli absent; enlarged claw (Fig. 150 C) without subdistal teeth, with one row of seven medial lamellae, in addition to four distal lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDC2B3CFF3ED6D4FF2BFE0C.taxon	description	Description. ♀ (Fig. 152). Fully winged; body length 3.2 mm. Head brown, except mandible, clypeus and frons testaceous; antennomeres 1 – 9 testaceous (10 th antennomere missing); mesosoma black; metasoma brown; legs brown, except articulations, part of tibiae, protarsus, meso- and metatarsomeres 1 – 4 testaceous. Antenna clavate; antennomeres 1 – 9 in following proportions: 7: 4: 21: 9: 8: 6: 6: 5.5: 4.5 (antennomere 10 missing). Head (Fig. 152 A, C) shiny, very excavated, with frons slightly granulate and vertex sculptured by very slight longitudinal striae; occipital carina incomplete, only present behind and shortly on sides of lateral ocelli; lateral ocelli touching occipital carina; POL = 1; OL = 5; OOL = 8; temple absent; greatest breadth of lateral ocelli longer than POL (2: 1). Pronotum dull, granulate, slightly humped, crossed by slight anterior and posterior transverse impressions (Fig. 152 B); posterior collar very short, hardly visible; pronotal tubercle not reaching tegula. Mesoscutum (Fig. 152 A) dull, granulate and slightly reticulate rugose. Notauli absent. Mesoscutellum dull, granulate. Metanotum reticulate rugose. Metapec- tal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two incomplete longitudinal keels. Forewing (Fig. 152 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 7). Protarsomeres in following proportions: 10: 2: 5: 11: 20. Enlarged claw (Fig. 150 C) without subdistal teeth, with one row of seven medial lamellae, in addition to four distal lamellae. Protarsomere 5 (Fig. 150 C) with one row of seven lamellae; apex with about 18 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDC2B3CFF3ED6D4FF2BFE0C.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19460): MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 15 – 28. I. 2004, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 52 (CAS). Hosts. Unknown. Distribution. Madagascar. Etymology. The species is named tulearensis after the collection site in Tulear (= Toliara) Province (Madagascar).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3CFF3ED793FE1FF93E.taxon	description	(Fig. 150 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3CFF3ED793FE1FF93E.taxon	description	Description. ♀. Fully winged; body length 3.5 – 4.1 mm. Head black, except clypeus and mandible testaceous; ventral side of head black; occiput black; antenna brown, except scape and pedicel testaceous; propleuron black; mesosoma black, except lateral regions of pronotum slightly testaceous; metasoma brown, except posterior third testaceous-brown; prolegs testaceous, except most part of coxae brown and clubs of femora darkened; mesolegs testaceous, except coxae partly black and tibiae and clubs of femora darkened; metalegs brown, except articulations, tarsi, part of trochanters and distal extremity of coxae testaceous. Antenna clavate; antennomeres in following proportions: 8: 6: 19: 10: 10: 7: 6.5: 6: 6: 8.5. Head flat, dull, setose, granulate and sculptured by numerous longitudinal and irregular keels; frontal line complete; occipital carina incomplete, present on sides of occiput, laterally not reaching eyes, absent on temple; occiput dull, granulate; POL = 5; OL = 3; OOL = 8; OPL = 0.5; TL = 1. Propleuron almost completely hidden under pronotum; pronotum dull, setose, granulate, slightly humped, crossed by anterior slight transverse furrow and posterior deep furrow; posterior collar of pronotum very short, almost absent, sculptured by few short longitudinal keels; pronotal tubercle not reaching tegula. Mesoscutum dull, setose, completely granulate and strongly or slightly reticulate rugose. Notauli absent. Mesoscutum with two incomplete longitudinal keels reaching almost posterior margin of mesoscutum and very similar to notauli. Mesoscutellum dull, granulate. Metanotum dull, granulate and reticulate rugose. Metapectal-propodeal disc completely reticulate rugose, shorter than propodeal declivity (16: 20); propodeal declivity reticulate rugose, with two longitudinal keels and median area dull, rugose. Mesopleuron dull, reticulate rugose. Metapleuron dull, granulate and transversely striate. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much longer than proximal part (11: 6). Protarsomeres in following proportions: 13: 2.5: 4.5: 9: 15. Enlarged claw (Fig. 150 D) without subdistal teeth, with two distal bristles, in addition to two proximal long and slender lamellae (most proximal lamella very long). Protarsomere 5 (Fig. 150 D) with three rows of approximately 31 lamellae extending continuously to apex (one lamella very long). Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3CFF3ED793FE1FF93E.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Cape Town, Tokai State Forest, Constantiaberge, above Donkerboskloof, 34 ° 02 ’ S 18 ° 23.5 ’ E, 460 m, 12 – 17. IX. 1993, MT, Mesic mountain Fynbos on Sandstone, SAM-HYM-PO 03804, S. van Noort leg. (SAMC). Other material: SOUTH AFRICA: Western Cape, Kogelberg Nature Researve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. XI – 16. XII. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 2 ♀♀ (SAMC, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3CFF3ED793FE1FF93E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3CFF3ED793FE1FF93E.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3DFF3ED142FDE2FA60.taxon	description	(Figs 150 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3DFF3ED142FDE2FA60.taxon	description	Description. ♀. Fully winged; body length 6.9 mm. Head black, except clypeus and mandible testaceous; ventral side of head black; occiput black; antenna testaceous, except antennomeres 7 – 10 brown; propleuron black; meso- soma black, except sides and posterior collar of pronotum slightly testaceous; metasoma black, except posterior half testaceous-reddish; proleg testaceous, with most part of coxa, part of trochanter and proximal extremity of femur black; meso- and metalegs black, except articulations and tarsi (except arolium) testaceous. Antenna clavate; antennomeres in following proportions: 14: 7: 23: 17: 14: 11: 9: 8: 7: 9; ADOs present in antennomeres 7 – 10. Head convex, dull, setose, granulate and reticulate rugose; frontal line incomplete, absent in anterior third of frons; occipital carina incomplete, shortly present on sides of lateral ocelli, laterally not reaching eyes, absent on temple; POL = 5; OL = 4; OOL = 9; OPL = 1; TL = 2; greatest breadth of lateral ocelli about as long as OL. Propleuron almost completely hidden under pronotum. Pronotum dull, setose, granulate, slightly humped, crossed by anterior slight transverse furrow and posterior deep furrow; posterior collar of pronotum very short, almost absent, slightly rugose; pronotal tubercle not reaching tegula. Mesoscutum dull, setose, completely granulate and reticulate rugose. Notauli absent. Mesoscutum with two incomplete longitudinal keels reaching 0.5 × length of mesoscutum and very similar to notauli. Mesoscutellum dull, reticulate rugose, except small median granulate area. Metanotum dull, reticulate rugose. Metapectal-propodeal disc completely reticulate rugose, shorter than propodeal declivity (16: 25); propodeal declivity reticulate rugose, with two longitudinal keels and median area dull, rugose. Mesopleuron dull, reticulate rugose. Metapleuron dull, granulate and transversely striate. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein much longer than proximal part (18: 9). Protarsomeres in following proportions: 14: 3: 8: 14: 25. Enlarged claw (Fig. 150 E) without subdistal teeth, with one row of five bristles in addition to two distal lamellae. Protarsomere 5 (Fig. 150 E) with two rows of 12 + 4 lamellae (shortest row composed of very long lamellae; longest row composed of very short lamellae); apex with at least 19 lamellae, among which one very long lamella. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 4.5 mm. Head black, except mandible and clypeus testaceous; antenna testaceous; mesosoma black; metasoma black, except posterior half testaceous-reddish; proleg testaceous, except coxa mostly black, trochanter partly brown and brown spot on club of femur; meso- and metalegs with coxae mostly black, trochanters partly black and partly whitish, femora black, tibiae testaceous-darkened, tarsi testaceous, except arolia brown, articulations testaceous. Antenna filiform; antennomeres in following proportions: 8: 6: 18: 12: 11: 12: 10: 9: 9: 10.5. Head dull, convex, completely reticulate rugose; frontal line complete; occipital carina complete; POL = 8; OL = 4; OOL = 6; OPL = 3; TL = 3; greatest breadth of lateral ocelli slightly longer than OL (5: 4). Mesoscutum, mesoscutellum and metanotum dull, reticulate rugose. Notauli slightly visible, incomplete, reaching approximately 0.5 × length of mesoscutum. Metapectal-propodeal disc reticulate rugose, without transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (17: 10); 2 R 1 cell open. Paramere with dorsal side showing mosaic sculpture (Fig. 150 F). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3DFF3ED142FDE2FA60.taxon	materials_examined	Material examined. Types: ♀ holotype: MOZAMBIQUE: Zambezia Prov., Gurué, 580 m, ex Philotheria talassio Fennah, C. 20. XI. 2002, B. 26. XI. 2002, Sf. 16. XII. 2002, M. Olmi reared (MOLC). Paratype: same locality label as holotype, C. 19. XI. 2002, B. 24. XI. 2002, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3DFF3ED142FDE2FA60.taxon	biology_ecology	Hosts. Dictyopharidae (Guglielmino et al. 2013): in Mozambique: adults of Philotheria talassio Fennah (feeding on Triumfetta sp. (Malvaceae) growing in tea fields).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDB2B3DFF3ED142FDE2FA60.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDA2B3EFF3ED17BFF2BFE70.taxon	description	Description. ♀. Fully winged; body length 7.0 mm (according to the original description). The holotype is a speci- men in very bad conditions: the mesosoma is completely missing. According to the original description, colour black, except mouthparts, antennomere 1 and legs brown (except meso- and metacoxa black). ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDA2B3EFF3ED17BFF2BFE70.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Périnet, III. 1935, east coast forest, Ol- soufieff leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDA2B3EFF3ED17BFF2BFE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDA2B3EFF3ED17BFF2BFE70.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FEDA2B3EFF3ED17BFF2BFE70.taxon	discussion	Remarks. The original description is not sufficient to permit identification of this species, or to include it in the key to the ♀♀ of the Afrotropical region. It is inevitable that it has to be included among the incertae sedis species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED407FE70FAF2.taxon	description	Description. ♂. Fully winged; body length 4.2 mm. Head black, except mandible testaceous; antenna with scape and pedicel testaceous, flagellomeres 1 – 4 darkened, 5 – 8 missing in holotype; mesosoma black; metasoma brown; legs yellow. Antenna filiform; antennomeres 1 – 6 in following proportions: 11: 8: 26: 15: 16: 15 (other antennomeres missing in holotype). Head dull, granulate and reticulate rugose, with vertex sculptured by irregular keels; frontal line absent; occipital carina complete, laterally reaching eyes; temple absent; POL = 5; OL = 2; OOL = 3; OPL = 1. Mesoscutum, mesoscutellum and metanotum dull, granulate and reticulate rugose. Notauli incomplete, reaching approximately 0.5 × length o mesoscutum. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (25: 16). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED407FE70FAF2.taxon	materials_examined	Material examined. Type: ♂ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Sankuru, Komi, I. 1930, J. Ghesquière leg. (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED407FE70FAF2.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED407FE70FAF2.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED407FE70FAF2.taxon	discussion	Remarks. The complete occipital carina and the characters of the forewing place this species within Dryinus. For this reason, a new combination for Lestodryinus luweli is presented above (Lestodryinus and Dryinus are synonyms). In the Afrotropical region, the genus Dryinus is divided into three groups based on ♀♀. The ♀ of this species is unknown, so that it is impossible to place it in a group. Out of necessity it has to be included among the incertae sedis species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED389FC03F837.taxon	diagnosis	Diagnosis. ♀ (Fig. 153): macropterous (Fig. 153); forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Fig. 153); protarsus chelate (Fig. 153); chela with rudimentary claw (Fig. 155 A); protarsomere 5 less than twice as broad as enlarged claw (Fig. 155 A); enlarged claw much longer than protibia, without subapical teeth (Fig. 155 A); antenna without tufts of long setae on antennomeres 5 – 10 (Fig. 153); palpal formula 4 / 2, 5 / 3; mandible with four teeth becoming regularly progressively larger from dorsal to ventral tooth (Fig. 6 B); tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED389FC03F837.taxon	distribution	Distribution. Afrotropical, Neotropical.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED389FC03F837.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED92B3EFF3ED389FC03F837.taxon	discussion	World species. Two species are known, one in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED82B31FF3ED28AFE14FAAD.taxon	description	(Figs 154 A, B, 155 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED82B31FF3ED28AFE14FAAD.taxon	description	Description. ♀ (Figs 154 A, B). Fully winged; body length 3.5 – 4.4 mm. Head testaceous, except large brown trans- verse band on ocellar region, temple and posterior half of frons; antenna of holotype with first two antennomeres testaceous, other antennomeres missing; antenna of paratype with scape and pedicel testaceous, flagellomeres 1 – 6 brown (except ADOs whitish), flagellomeres 7 – 8 whitish; propleuron brown, except ventral side testaceous; pronotum brown, with testaceous spots on posterior collar and pronotal tubercle; mesoscutum brown, except lateral regions partly testaceous; rest of mesosoma black, except mesopleuron partly testaceous; petiole brown; metasoma brown, except ventral side testaceous; legs yellow-testaceous, except club of profemur darkened. In paratype from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 60, mesosoma brown-black, except testaceous spots on propleuron, posterior surface of pronotum, lateral regions of mesoscutum, mesopleuron; metasoma brown; legs totally yellow-testaceous. Antenna of holotype with only the first two antennomeres, other antennomeres missing; antennomeres 1 - 2 in following proportions: 9: 6. Antenna of paratype from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 60, very slender, filiform; antennomeres in following proportions: 9: 7: 45: 35: 20: 14: 9: 6: 6: 10. Head slightly swollen, shiny, unsculptured; frontal line of holotype complete (frontal line of paratype incomplete, present only in posterior half of frons); occipital carina complete; holotype with POL = 4; OL = 2; OOL = 6; TL = 5; greatest breadth of lateral ocelli about as long as POL; posterior ocelli touching occipital carina; paratype from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 60, with POL = 4; OL = 2; OOL = 6; OPL = 0.5; TL = 5; greatest breadth of lateral ocelli longer than OL (4: 2). Palpal formula 5 / 3. Pronotum shiny, punctate, unsculptured among punctures, crossed by two transverse impressions, anterior one very deep and posterior one weak; anterior collar very long; disc very short and humped; posterior collar short; pronotal tubercle reaching tegula. Mesoscutum (Fig. 154 B) shiny, with lateral regions unsculptured and median region finely punctate, unsculptured among punctures. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of lateral ocelli (2: 4). Mesoscutellum and metanotum almost flat, shiny, unsculptured. Metapectal-propodeal disc dull, with granulate, except two median subparallel keels connecting anterior margin with transverse keel separating metapectal-propodeal disc and propodeal declivity; propodeal declivity with two median longitudinal keels, median area shiny and slightly granulate (in paratype from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 60, with some distal transverse keels), lateral areas with two large shiny areolae. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (31: 11); 2 R 1 cell open (almost closed). Protarsomeres of holotype in following proportions: 15: 4: 6: 29: 38. Protarsomeres of paratype in following proportions: 16: 3: 7: 30: 38. Protarsomere 3 produced into hook. Proleg of holotype with the following proportions: 41 (coxae): 41 (trochanters): 45 (femora): 43 (tibiae): 31 (enlarged claw). Enlarged claw (Figs 155 A, B) without teeth, with one row of 9 – 11 lamellae + 1 – 2 distal lamellae. Holotype with protarsomere 5 (Fig. 155 A) provided with one row of seven lamellae (with two rows of 2 (proximal) + 7 lamellae in paratype from Madagascar, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, MA- 02 - 09 C- 60; with two rows of 1 (proximal) + 6 lamellae in paratype from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ‘ E) (Fig. 155 B); apex with group of about 8 – 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED82B31FF3ED28AFE14FAAD.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 26. II – 10. III. 2003, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 55 (CAS). Paratypes: same locality label as holotype, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♀ (MOLC); Fi- anarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ‘ E, 1130 m, 18 – 27. II. 2003, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 53, 1 ♀ (CAS); same locality label, 17 – 30. V. 2003, MA- 02 - 09 B- 61, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED82B31FF3ED28AFE14FAAD.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED82B31FF3ED28AFE14FAAD.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED62B32FF3ED332FD2AFDD0.taxon	diagnosis	Diagnosis. ♀ (Figs 154 C, D): fully winged (Figs 154 C, D); palpal formula 3 / 2, 4 / 2, 4 / 3, 5 / 2, or 5 / 3 (for 6 / 3, see remarks about Pseudodryinus townesi); antenna with many long scattered setae on antennomeres 6 – 10 (setae not forming tufts) (Fig. 154 C); forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 154 C, D); protarsus chelate (Figs 154 C, D); chela with rudimentary claw (Fig. 155 C); protarsomere 5 less than twice as broad as enlarged claw (Fig. 155 C); enlarged claw with one subapical tooth (Fig. 155 C )); tibial spurs 1 / 1 / 2. ♂ (Figs 156 A, B): fully winged (Figs 156 A, B); palpal formula 6 / 3; occipital carina complete; mandible with four irregular teeth; notauli joint or separated at posterior margin of mesoscutum; if separated, minimum distance between notauli much shorter than greatest breadth of lateral ocelli; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 156 A, B); forewing with PostabR 1 as long as, or longer than pterostigma (Fig. 156 B); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED62B32FF3ED332FD2AFDD0.taxon	distribution	Distribution. Palaearctic, Afrotropical, Oriental and Australian regions.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED62B32FF3ED332FD2AFDD0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED62B32FF3ED332FD2AFDD0.taxon	discussion	World species. Ten species are known, two in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED62B32FF3ED332FD2AFDD0.taxon	discussion	Remarks. Two fossil species of Pseudodryinus are known: Pseudodryinus parisiensis Peinado et al. 2006 (amber from Le Quesnoy deposit (Oise, France )); P. burmensis Olmi, Xu & Guglielmino 2014 (amber from Myanmar).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B32FF3ED4E6FE5BF9C7.taxon	description	(Fig. 155 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B32FF3ED4E6FE5BF9C7.taxon	description	Description. ♀. Fully winged; body length 5.6 mm. Head testaceous, except transverse black band on vertex; antenna brown, except scape, pedicel and flagellomeres 5 – 8 testaceous; mesosoma black, except propleuron and lateral regions of pronotum partly testaceous; metasoma brown; legs completely testaceous. Antenna very slender, filiform; antennomeres in following proportions: 11: 6: 70: 59: 32: 15: 11: 9: 7: 13. Head shiny, alutaceous; frontal line incomplete, present only in front of anterior ocellus; frons with median longitudinal furrow; anterior half of frons strongly setose; occipital carina incomplete, not present on temple; POL = 4; OL = 2; OOL = 9; OPL = 1; TL = 6. Palpal formula 3 / 2. Pronotum shiny, setose, unsculptured, crossed by strong transverse impression; disc humped; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of lateral ocelli (3.5: 5). Metapectal-propodeal disc with strong transverse posterior keel, granulate, with numerous irregular keels and areolae and with two longitudinal and parallel keels situated in median region; propodeal declivity reticulate rugose, with two complete longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (40: 17); 2 R 1 cell almost closed. Protarsomere 3 produced into hook. Protarsomeres in following proportions: 23: 5: 10: 47: 60. Enlarged claw (Fig. 155 C) with one large subdistal tooth and one row of 13 lamellae and six long distal bristles. Protarsomere 5 (Fig. 155 C) with two rows of 2 (proximal) + 12 lamellae; apex with approximately 20 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B32FF3ED4E6FE5BF9C7.taxon	materials_examined	Material examined. Type: ♀ holotype: GABON: Woleu N’Tem District, Foulebeng, 19. III. 1987, forest, near a river, A. Pauly leg. (FSAE).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B32FF3ED4E6FE5BF9C7.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B32FF3ED4E6FE5BF9C7.taxon	distribution	Distribution. Gabon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	description	(Figs 154 C, D, 155 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	description	Description. ♀ (Figs 154 C, D). Fully winged; body length 3.4 – 5.6 mm. Completely testaceous, except petiole black; antenna brown, except scape and pedicel testaceous. In specimen from Democratic Republic of the Congo, 04 ° 16 ’ S 17 ° 11 ’ E, head brown, except mandible, clypeus, anterior half of frons, ventral side and part of occiput testaceous; antenna testaceous-brown, except scape and pedicel testaceous; mesosoma testaceous, except scutum and metanotum brown; metapectal-propodeal disc and propodeal declivity black; metanotum testaceous, except petiole and part of first tergite brown; legs testaceous. Antenna very slender (Figs 154 C, D), clavate; holotype with antennomeres 1 – 4 in following proportions: 7: 5: 50: 40 (antennomeres 5 – 10 missing in holotype). Head flat, shiny, slightly alutaceous; occiput strongly excavated; frontal line absent; frons without lateral keels, with short median furrow in front of anterior ocellus; occipital carina complete; POL = 6; OL = 3.5; OOL = 8; OPL = 1.5; TL = 7. Palpal formula of type material 4 / 2; in specimen from Central African Republic, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by transverse impression; pronotal tubercle reaching tegula. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured. Notauli complete, posteriorly separated, very converging. Metapectal-propodeal disc dull, granulate, with some areolae or tracks of areolae, with transverse posterior keel; propodeal declivity with two longitudinal keels and median area dull, granulate. In specimen from Central African Republic, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, metapectal-propodeal disc sculptured by large areolae, not granulate, with two median parallel irregular longitudinal keels; propodeal declivity with two longitudinal keels and large areolae similar to those of metapectal-propodeal disc. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part; 2 R 1 cell closed. Hind wings hyaline, not darkened. Protrochanter about as long as profemur (44: 46), procoxa (44: 46) and protibia (44: 42). Protarsomeres in following proportions: 16: 4: 8: 31: 41. Enlarged claw (Fig. 155 D) with one large subdistal tooth and one row of six lamellae. Protarsomere 5 (Fig. 155 D) with one row of nine lamellae; apex with 10 lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown (see remarks).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Kampala (AEIC). Paratypes: SOUTH AFRICA: Lim- popo, Tzaneen, 1 ♀ (AEIC). UGANDA: same locality label as holotype, 1 ♀ (AMNH). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 6. V. 2001, sweep, lowland rainforest, marsh clearing, CAR 01 - S 48, S. van Noort leg., 1 ♀ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 18. IV. 2006, seeping in riparian forest, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED52B33FF3ED0DCFB6FFAD0.taxon	discussion	Remarks. The ♂ tentatively attributed to Ps. townesi by Olmi (1984) cannot belong to teh genus Pseudodryinus, because the minimum distance between the notauli is longer than the greatest breadth of the lateral ocelli. For this reason it is now attributed to Thaumatodryinus Perkins (Th. sokokensis Olmi 2007). In the specimen from Central African Republic, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, both chelae are missing, so that it is not possible to check if it is a new species. The sculpture of the metapectal-propodeal disc is very different from that of the type material. In addition, the palpal formula 6 / 3 is not known in Pseudodryinus, being typical of Harpactosphecion. It is likely that the two genera are synonyms and Harpactosphecion gains priority. It is necessary to wait for additional specimens with complete chelae before reaching an informed decision about this potential synonymy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	diagnosis	Diagnosis. ♀ (Figs 156 C, D): fully winged (Figs 156 C, D); forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 156 C, D); usually in forewing also 2 Cu cell completely enclosed by pigmented veins; protarsus chelate (Figs 156 C, D); chela with a rudimentary claw (Fig. 158 A); antenna with tufts of long setae (“ tiloïdes ” of Benoit 1954) on antennomeres 5 – 10 (antennomere 10 with two tufts) (Figs 156 C, D, 157 C); pronotal tubercle reaching tegula; mandible with four teeth becoming regularly progressively larger from dorsal to ventral tooth (Fig. 6 B); occipital carina complete; antenna without ADOs; palpal formula 6 / 3; tibial spurs 1 / 1 / 2. ♂ (Figs 157 A, B): fully winged; forewing with three cells enclosed by pigmented veins (C, R and 1 Cu) (Figs 157 A, B); forewing with PostabR 1 as long as, or longer than pterostigma (Figs 157 A, B); occipital carina complete; paramere without inner branch wrapping penis (Figs 159 A, C); mandible usually with four teeth becoming regularly progressively larger from dorsal to ventral tooth (Fig. 6 B); occasionally mandible with four teeth of different length; never quadridentate mandibles including one intermediate rudimentary tooth; notauli always separated at posterior margin of mesoscutum; minimum distance between notauli as long as, or longer than greatest breadth of lateral ocelli; pal- pal formula 6 / 3; hypopygium not umbrella-shaped (as in Fig. 8 H); tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	distribution	Distribution. Worldwide, except Palaearctic region. Brues (1923, 1933) described three fossil species of Thau- matodryinus from Baltic amber; they are now attributed to Harpactosphecion Haupt.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	biology_ecology	Hosts. Flatidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	discussion	World species. 33 species are known, ten in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	discussion	World genera. One genus is known.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED42B34FF3ED3E7FDD3FDB8.taxon	discussion	Remarks. After the original description by Olmi (1984), the subfamily Thaumatodryininae was considered a junior synonym of Dryininae Haliday 1833 (Xu et al. 2013; Olmi & Virla 2014). Based on molecular analysis, Tribull (2015) resurrected Thaumatodryininae to establish the monophyly of Dryininae and retain Gonatopodinae as a separate subfamily. The defined morphological synapomorphy for Thaumatodryininae is the presence of tufts of long setae on antennomeres 5 – 10.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	type_taxon	Type species. Thaumatodryinus koebelei R. Perkins 1905, by original designation.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	diagnosis	Diagnosis. ♀ (Figs 156 C, D) and ♂ (Figs 157 A, B): see above diagnosis of the subfamily Thaumatodryininae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	distribution	Distribution. Worldwide, except Palaearctic region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	biology_ecology	Hosts. Flatidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	discussion	World species. 33 species are known, ten in the Afrotropical region. Remarks. One fossil species (Thaumatodryinus miocenicus Olmi 1995) was described in amber from the Dominican Republic. Brues (1923, 1933) described three fossil species of Thaumatodryinus from Baltic amber; they are now attributed to Harpactosphecion Haupt.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED22B35FF3ED056FD48F86B.taxon	discussion	Remarks. One fossil species (Thaumatodryinus miocenicus Olmi 1995) was described in amber from the Dominican Republic. Brues (1923, 1933) described three fossil species of Thaumatodryinus from Baltic amber; they are now attributed to Harpactosphecion Haupt.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED12B36FF3ED21BFE1FF8BD.taxon	description	(Fig. 158 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED12B36FF3ED21BFE1FF8BD.taxon	description	Description. ♀. Fully winged; body length 3.9 mm. Ferruginous, except petiole black; mesoscutellum brown; mesopleuron, metapleuron and metapectal-propodeal disc darkened. Antenna clavate; antennomeres in following proportions: 16: 9: 28: 34: 42: 38: 30: 13: 9: 10. Head dull, convex, granulate; frontal line complete, reduced to track; occipital carina complete; POL = 3; OL = 2; OOL = 11; OPL = 2; TL = 3. Pronotum setose, crossed by strong transverse impression, granulate; posterior tubercle of pronotum reaching tegula. Mesoscutum, mesoscutellum and metanotum dull, granulate. Notauli incomplete, reaching approximately 0.4 × length of mesoscutum. Metapectalpropodeal disc and propodeal declivity reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (35: 13); 2 R 1 cell open. Protarsomeres in following proportions: 27: 4.5: 7: 24: 33. Enlarged claw (Fig. 158 A) with two subapical teeth and one row of 35 lamellae. Protarsomere 5 (Fig. 158 A) with two rows of approximately 43 lamellae; apex with approximately six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED12B36FF3ED21BFE1FF8BD.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu – Natal, Umlalazi Nature Reserve, 1.5 km E of Mtunzini, X. 1978, indigenous forest, R. M. Miller leg. (PMA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED12B36FF3ED21BFE1FF8BD.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED12B36FF3ED21BFE1FF8BD.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED02BC9FF3ED376FD85FDB8.taxon	description	(Figs 158 B, 159 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED02BC9FF3ED376FD85FDB8.taxon	description	Description. ♀. Fully winged; body length 4.6 – 5.6 mm. Testaceous, except petiole black. Specimen from Madagas- car, Ankarana, quite different from type material, because of following characters: head mostly testaceous, except brown area around occipital hole and two brown stripes in front of antennal toruli; antenna testaceous; mesosoma mostly testaceous, except isolated brown stripes on propleuron, along posterior margin of pronotum, on lateral regions of pronotum, along anterior margin of mesoscutum, suture between mesoscutum and mesoscutellum, isolated stripes on mesopleuron; metanotum totally brown; metapectal-propodeal disc and propdeal declivity partly testaceous and partly brown; metasoma testaceous, except some brown spots; legs testaceous, except metatibia and club of profemur partly brown. Antenna clavate; antennomeres in following proportions: 17: 7: 34: 38: 39: 15: 10: 19: 11.5: 10. Head shiny, completely granulate and reticulate rugose; frontal line absent; occipital carina complete; ocellar region swollen; POL = 7; OL = 3; OOL = 11; OPL = 3; TL = 4. Pronotum shiny, crossed by transverse impression, granulate; sides of pronotum sculptured by numerous longitudinal striae; posterior tubercle of pronotum reaching tegula. Mesoscutum shiny, granulate and slightly rugose; posterior third of mesoscutum sculptured by slight longitudinal keels. Notauli complete, posteriorly separated; minimum distance between notauli approximately twice as long as greatest breadth of lateral ocelli (10: 5). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with one dark transverse band situated around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein longer than proximal part (34: 23). Hind wing hyaline, without dark transverse bands. Protarsomeres in following propor- tions: 29: 5: 8: 30: 45. Enlarged claw (Fig. 158 B) with two subapical teeth and one row of 35 lamellae. Protarsomere 5 (Fig. 158 B) with two rows of about 57 lamellae; apex with approximately eight lamellae. Tibial spurs 1 / 1 / 2. ♂. Fully winged; body length 2.7 mm. Totally testaceous. Antenna filiform; antennomeres in following propor- tions: 6: 5: 10: 15.5: 14: 13.5: 11: 10: 9: 9. Head shiny, convex, with many slight irregular striae, unsculptured among striae; frontal line absent; occipital carina complete; POL = 8; OL = 2; OOL = 6; OPL = 2; TL = 4; greatest diameter of lateral ocelli longer than OL (4: 2). Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of the ocelli (6.5: 4). Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc reticulate rugose, without strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (23: 13); 2 R 1 cell open. Inner side of paramere with characteristic mosaic sculpture (Fig. 159 A). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED02BC9FF3ED376FD85FDB8.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: Brandberg, Wasserfallfläche, 1960 m, 21 ° 10.77 ’ S 14 ° 32.87 ’ E, 7. IV. 1999, NA 99 – L 03, UV light trap, Bushy Karoo-Namib shrubland, S. van Noort & S. G. Compton leg. (SMWN). Paratypes: same locality label as holotype, 3 ♀♀, 1 ♂ (SMWN), 1 ♀ (MOLC). Other material: MAD- AGASCAR: Antsiranana, Sakalava Beach, 12 ° 15 ’ 46 ’’ S 49 ° 23 ’ 51 ’’ E, dwarf littoral forest, 10 m, 22 – 27. IV. 2001, MT, across sandy trail, M. Irwin & R. Harin’Hala leg., MA- 01 - 04 B- 07, 1 ♀ (CAS); Antsiranana, Ankarana, IX. 1986, MT, Jane Wilson leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED02BC9FF3ED376FD85FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FED02BC9FF3ED376FD85FDB8.taxon	distribution	Distribution. Madagascar, Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	description	(Fig. 158 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	description	Description. ♀. Fully winged; body length 4.5 – 4.8 mm. Head yellow-testaceous, except occiput partly black, ocel- lar region black and frons with four black spots (two in front and two behind antennal toruli); antenna testaceous, except antennomeres 1 and 4 – 10 dark; propleuron yellow; pronotum yellow-testaceous, except lateral regions partly brown; mesoscutum yellow-testaceous, except anterior margin black; mesoscutellum yellow-testaceous; furrow between mesoscutellum and mesoscutum brown; metanotum black; regions on sides of mesoscutellum and metanotum black; metapectal-propodeal disc black-brown, except anterior third reddish; petiole black; metasoma brownyellow; legs testaceous, except clubs of pro- and metafemora brown. In specimen from Madagascar, 13 ° 10.00 ’ S 49 ° 42.36 ’ E, head testaceous, except two small brown spots behind antennal toruli; antenna testaceous, except antennomere 2 yellow; mesosoma testaceous, except propleuron partly brown, two brown longitudinal stripes in lateral regions of pronotum, two small brown spots on mesopleuron near tegula, metanotum mostly brown, central brown longitudinal stripe on propodeal declivity; petiole brown; metasoma yellow, except part of segment 1 brown and brown transverse stripe on every segment from segment 2 to distal extremity; legs yellow, except brown spots on clubs of pro- and metafemur. Antenna clavate; antennomeres in following proportions: 15: 9: 28: 33: 41: 40: 28: 13: 9: 11. Head shiny, granulate; occipital carina incomplete, only present behind ocelli and on sides of vertex; POL = 3.5; OL = 1; OOL = 8; OPL = 2; greatest breadth of lateral ocelli longer than OPL (3: 2). In specimen from Madagascar, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, OPL as long as OL. Pronotum shiny, setose, crossed by transverse impression, granulate; posterior tubercle of pronotum reaching tegula. Mesoscutum shiny, setose, granulate, sculptured by slight longitudinal striae. Notauli almost complete, posteriorly separated, slightly longer than POL (4: 3.5). Mesoscutellum shiny, unsculptured. Metanotum rugose. Metapectal-propodeal disc reticulate rugose, with anterior third almost smooth (rugose in specimen from Madagascar, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E). Forewing hyaline, without dark transverse bands in specimen from Madagascar, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, forewing with small dark spot around 2 r-rs & Rs vein); distal part of 2 r-rs & Rs vein much longer than proximal part (36: 14); 2 R 1 cell open; pterostigma brown-light. Protarsomere 1 much longer than 4 (26: 19). Protarsomere 3 produced into hook. Enlarged claw (Fig. 158 C) with two subapical teeth and one row of 36 lamellae. Protarsomere 5 (Fig. 158 C) with two rows of approximately 63 lamellae extending continuously to apex. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, XI. 1936, A. Seyrig leg. (MNHN). Paratypes: same locality label as holotype, IX. 1936, 1 ♀ (MNHN); same locality label as holotype, X. 1936, 1 ♀ (MNHN); same locality label, I. 1940, 2 ♀♀ (MRAC). Other material: MADAGASCAR: Antsiranana, Bekaraoka Forest, 6.8 km 60 ° ENE Daraina, 13 ° 10.00 ’ S 49 ° 42.36 ’ E, 150 m, 7. XII. 2003, YPT, tropical dry forest, B. L. Fisher leg., BLF 9873, 1 ♀ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 12. II – 4. III. 2001, MT, R. Harin’Hala leg., MA- 1 - 01 ° - 07, 1 ♀ (CAS); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 1 ♀ (CAS); Mahajanga, Namoroka National Park, 16.40682 S 45.31377 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♀ (MNHN); Toliara, Fiherenana, 23 ° 13.351 ’ S 43 ° 52.853 ’ E, 65 m, 5 – 10. VIII. 2003, MT, degraded riparian forest close to water, MGF 076, Frontier Wilderness Project, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2E2BCAFF3ED4DFFC41FE53.taxon	discussion	Remarks. One ♀ of this species is also known in copal from Madagascar. It is deposited in MOLC. It is pos- sible that Thaumatodryinus merinus is the opposite sex of T. dentatus Benoit (because one ♀ and six ♂♂ have been collected in the same site in Mahajanga, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, and 1 ♀ and 7 ♂♂ have been collected in the same site in Antsiranana, Bekaraoka Forest, 13 ° 10.00 ’ S 49 ° 42.36 ’ E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCAFF3ED46BFE14FA8E.taxon	description	(Fig. 158 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCAFF3ED46BFE14FA8E.taxon	description	Description. ♀. Fully winged; body length 4.7 mm. Testaceous-reddish, except petiole and mesoscutellum black. An- tenna of holotype broken, with only antennomeres 1 – 5; antennomeres 1 – 5 in following proportions: 39: 15: 73: 92: 83. Head shiny, slightly reticulate rugose; occipital carina complete; POL = 14; OL = 6; OOL = 28; OPL = 7; TL = 10. Pronotum shiny, setose, crossed by transverse impression, slightly granulate, with slight transverse striae; posterior tubercle of pronotum reaching tegula. Mesoscutum shiny, completely sculptured by numerous longitudinal keels. Notauli complete, posteriorly separated. Mesoscutellum and metanotum punctate. Metapectal-propodeal disc and propodeal declivity reticulate rugose. Forewing with one dark transverse band; distal part of 2 r-rs & Rs vein much longer than proximal part; 2 R 1 cell closed; pterostigma brown-light. Hind wing with distal brown transverse band. Protarsomere 1 approximately as long as 4. Protarsomere 3 produced into hook. Enlarged claw (Fig. 158 D) with two subapical teeth and one row of 32 lamellae. Protarsomere 5 (Fig. 158 D) with two rows of approximately 43 lamellae; apex with approximately six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCAFF3ED46BFE14FA8E.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Saint-Augustin, 29. III. 1968, K. M. G. & P. D. leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCAFF3ED46BFE14FA8E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCAFF3ED46BFE14FA8E.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	description	(Figs 157 D, 159 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	description	Description. ♂ (Fig. 157 D). Fully winged; body length 2.2 – 3.0 mm. Head testaceous, except ocellar triangle black or brown; antenna testaceous-darkened, except antennomere 1 testaceous-light; mesosoma testaceous, except brown spot on anterior surface of mesoscutum; mesoscutellum and metanotum black or brown; large brown spot on metapectal-propodeal disc and propodeal declivity; anterior surface of mesopleuron and meso-metapleuron suture brown; metasoma brown; legs testaceous. Some specimens from Madagascar, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, are more darkened, with part of occiput and entire mesopleuron and metapleuron brown. Occasionally mesosoma testaceous, except mesoscutum, mesoscutellum, part of mesopleuron and median region of propodeal declivity brown. In specimen from Madagascar, 16 ° 41.49 ’ S 44 ° 31.41 ’ E, head black, except mandible, clypeus, part of malar space and stripe between antennal toruli ferruginous-dark; antenna brown, except scape and pedicel testaceous; prothorax ferruginous-dark; mesoscutum totally black; rest of mesosoma black; metasoma brown; legs testaceous. In specimen from Madagascar, 23 ° 13.351 ’ S 43 ° 52.853 ’ E, head black, except mandible, clypeus, malar space and stripe along orbits testaceous, area between antennal toruli ferruginous-dark; antenna brown, except antennomere 1 testaceous; mesosoma black, except prothorax (dorsally brown), posterior third of mesoscutum and basal third of metapleuron testaceous; metasoma brown; legs testaceous. In specimen from Madagascar, 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, head and mesosoma completely testaceous-reddish, except ocellar triangle and axilla brown. Antenna filiform; antennomeres in following proportions: 7: 5: 9: 7: 11: 12: 11: 10: 8: 11. Head dull, convex, granulate and slightly rugose; frontal line absent; occipital carina complete; POL = 5.5; OL = 2; OOL = 6; OPL = 1; TL = 4; greatest breadth of lateral ocelli longer than OL (4: 2). In dark specimens head with some irregular keels on frons. Mesoscutum dull, as sculpured as head (in dark specimens mesoscutum with some irregular keels). Notauli complete, posteriorly separated; minimum distance between notauli as long as greatest breadth of lateral ocelli (4: 4), shorter than breadth of each lateral region of mesoscutum measured at posterior margin of mesoscutum (Fig. 159 B); in two specimens from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, minimum distance between notauli longer than greatest breadth of lateral ocelli (4: 3). Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without strong transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (19: 14); 2 R 1 cell open. Inner side of paramere without mosaic sculpture (Fig. 159 C); tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW Anivorano Nord, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., code BLF 3011, (CAS). Paratypes: same locality label as holotype, 9 ♂♂ (CAS), 2 ♂♂ (MOLC). Other material: MADAGASCAR: Antsiranana, Diana Region, Orangea dry forest, Ramena, Baie de dune, 900 m E of Camp Minier, 15 – 22. VI. 2011, 12 ° 13.97 ’ S 49 ° 21.99 ’ E, M. Irwin, R. Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62 - 27, 1 ♂ (CAS); same locality label 23 – 31. III. 2011, 1 ♂ (MOLC); Antsiranana, Orangea, 3 km E of Ra- mena near fort, 12 ° 14.49 ’ S 49 ° 22.17 ’ E, 65 m, 23 – 27. I. 2001, MT, littoral forest on sand, MA- 01 - 05 - 02, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., 4 ♂♂ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 20 – 28. VII. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 15, M. E. Irwin & R. Harin’Hala leg., 2 ♂♂ (CAS); Antsiranana, 1 km W Sakalava Beach, 12 ° 15.59 ’ S 49 ° 23.42 ’ E, brushy area, 30 m, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 04 A- 01, 2 ♂♂ (CAS); Antsiranana, Ampondrabe Forest, 26.3 km 10 ° NNE Daraina, 12 ° 58 ’ 12 ’’ S 49 ° 42 ’ 00 ’’ E, 175 m, 10. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9976, 1 ♂ (CAS); Antsiranana, Bekaraoka Forest, 6.8 km 60 ° ENE Daraina, 13 ° 10.00 ’ S 49 ° 42.36 ’ E, 150 m, 7. XII. 2003, tropical dry forest, general collecting night spiders, B. L. Fisher leg., BLF 9876, 7 ♂♂ (6 in CAS, 1 in MOLC); An- tsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 19. III – 6. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 09, 1 ♂ (CAS); same locality label, 11 – 12. II. 2001, R. Harin’Hala leg., MA- 01 - 01 A- 06, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 26. IV – 12. IV. 2001, MT, MA- 01 - 01 D- 09, R. Harin’Hala leg., 1 ♂ (CAS); same locality label, 14 – 30. V. 2001, MA- 01 - 01 D- 11, 1 ♂ (CAS); Antsira- nana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, 19. III – 5. IV. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 07, 1 ♂ (CAS); same locality label, 4 – 10. III. 2001, MA- 1 - 01 B- 06, 1 ♂ (CAS); Antsiranana, Ambre Special Reserve, 3.5 km 235 ° SW Sakaramy, 325 m, 12 ° 28.8 ’ S 49 ° 14.32 ’ E, 26 – 31. I. 2001, MT, tropical dry forest, BLF 2661, Fisher, Griswold et al. leg., 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Antsahabelela rain forest, 9 km SW Daraina, Binara, 13 ° 15.03 ’ S 49 ° 37.00 ’ E, 182 m, 25. V – 1. VI. 2011, MT, in humid forest, M. Irwin, R. Harin’Hala leg., MG- 58 - 34, 2 ♂♂ (CAS, MOLC); same locality label, 16 – 23. II. 2011, MG- 58 - 20, 1 ♂ (CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimaty dry forest, 9 km NE Daraina, 13 ° 10.17 ’ S 49 ° 42.04 ’ E, 90 m, 19 – 26. XII. 2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG- 57 - 12, 2 ♂♂ (CAS); same locality label, 14 – 21. II. 2011, MG- 57 - 19, 2 ♂♂ (CAS). Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 16. X – 8. XI. 2001, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 01, 3 ♂♂ (CAS); Fianarantsoa, near Isalo National Park, in dry wash east of Interpre- tive Center, 22 ° 37.60 ’ S 45 ° 21.49 ’ E, 885 m, 6 – 14. XII. 2002, MT, in open area, MA- 02 - 11 B- 49, R. Harin’Hala leg., 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 26 – 31. III. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 22, 1 ♂ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 2.5 km 62 ° ENE Bekopaka, Ankidrodroa River, 19 ° 07.56 ’ S 44 ° 48.53 ’ E, 100 m, 11 – 15. XI. 2001, MT, tropical dry forest on Tsingy, Fisher, Griswold et al. leg., BLF 4345, 2 ♂♂ (CAS); Mahajanga, Namoroka National Park, 9.8 km 300 ° WNW Vilanandro, 16 ° 28.00 ’ S 45 ° 21.00 ’ E, 140 m, 4 – 8. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6445, 9 ♂♂ (3 in CAS, 6 in MOLC); Mahajanga, Namoroka Na- tional Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.24 ’ S 45 ° 18.36 ’ E, 100 m, 12 – 16. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6581, 7 ♂♂ (CAS); Mahajanga, Namoroka National Park, 17.8 km 329 ° WNW Vilanandro, 16 ° 22.36 ’ S 45 ° 19.36 ’ E, 100 m, 8 – 12. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6508, 1 ♂ (CAS); Mahajanga, Namoroka National Park, 16.40818 S 45.31437 E, 21 – 31. X. 2016, MT, C. Ville- mant leg., 1 ♂ (MNHN); Mahajanga, Namoroka National Park, 16.40494 S 45.31294 E, 21 – 31. X. 2016, MT, C. Vil- lemant leg., 1 ♂ (MNHN); Mahajanga, Namoroka National Park, 16.40682 S 45.31377 E, 21 – 31. X. 2016, MT, C. Villemant leg., 2 ♂♂ (MNHN); Mahajanga, Namoroka National Park, 21 – 31. X. 2016, MT, C. Villemant leg., 2 ♂♂ (MNHN); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 6 ♂♂ (5 in CAS, 1 in MOLC); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15 ° 55 ’ S 45 ° 57 ’ E, 65 ft., 23 – 30. X. 2007, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 38 - 06, 1 ♂ (CAS); same locality label, 15 – 22. I. 2008, MG- 38 - 14, 1 ♂ (CAS); same locality label, 2 – 9. X. 2007, MG- 38 - 03, 1 ♂ (CAS); same locality label, 20. XI – 4. XII. 2007, MG- 38 - 09, 3 ♂♂ (CAS); same locality label, 22 – 29. I. 2008, MG- 38 - 15, 3 ♂♂ (CAS); same locality label, 26. II – 4. III. 2008, MG- 38 - 20, 1 ♂ (CAS); Mahajanga, Ambatofolaka, Namoroka, 53 km from Soalala, 3 km N Vilanandro village, 16 ° 28 ’ 04 ’’ S 45 ° 23 ’’ 48 ’’ E, 400 ft., 9 – 18. XII. 2007, dense dry forest, MT, M. Irwin, R. Harin’Hala leg., MG- 40 A- 11, 1 ♂ (CAS); Mahajanga, Paro, Ankarafantsika National Park, Ampijoroa Forest Station, 40 km 306 ° NW Andranotasica, 16 ° 10 ’ 15 ’’ S 46 ° 49 ’ 38 ’’ E, 130 m, 20. III – 1. IV. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 3520, 2 ♂♂ (CAS); Mahajanga, Bsalampy District, Analangidro dry forest, 7 km NE Besalampy, 16 ° 41.49 ’ S 44 ° 31.41 ’ E, 200 ft., 4 – 11. XI. 2007, MT, dry forest on sand, M. Irwin & R. Harin’Hala leg., MG- 41 - 07, 1 ♂ (MOLC); Toliara, Andohahela National Park, Manantalinjo Forest, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, 24 ° 49.1 ’ S 46 ° 36.36 ’ E, 150 m, 12 – 16. I. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 4811, 2 ♂♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelled II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 16 – 17. XII. 2002, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 09, 8 ♂♂ (6 in CAS, 2 in MOLC); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Marofandilia, 20 ° 02.42 ’ S 44 ° 39.44 ’ E, 100 m, 28. XI – 3. XII. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4601, 2 ♂♂ (CAS); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Ma- rofandilia, 20 ° 04.09 ’ S 44 ° 39.38 ’ E, 30 m, 11 – 18. III. 2009, MT, tropical dry forest, B. L. Fisher leg., BLF 18285 _ 71, 1 ♂ (CAS); same locality label, 9 – 16. I. 2008, BLF 18285 _ 10, 1 ♂ (MOLC); same locality label, 27. II – 5. III. 2008, BLF 18285 _ 17, 1 ♂ (CAS); Toliara, Kirindy Forest, 15.5 km 64 ° ENE Marofandilia, 20 ° 04.07 ’ S 44 ° 39.34 ’ E, 24 – 31. XII. 2008, B. L. Fisher, MT, tropical dry forest, el 30 m, BLF 18284 _ 60, 1 ♂ (CAS); same locality label, 25. III – 1. IV. 2009, BLF 18284 _ 73, 1 ♂ (CAS); same locality label, 14 – 21. I. 2009, BLF 18284 _ 63, 1 ♂ (CAS); Toliara, Antaf- oky, 23 ° 28.43 ’ S 44 ° 03.51 ’ E, 55 m, 25 – 28. I. 2002, MT, Gallery forest, Frontier Project leg., MGF 003, 1 ♂ (CAS); Toliara, Antafoky, 23 ° 29.19 ’ S 44 ° 04.44 ’ E, 60 m, 8 – 10. II. 2002, MT, Gallery forest, Frontier Project leg., MGF 006, 1 ♂ (CAS); Toliara, Antafoky, 23 ° 28.44 ’ S 44 ° 03.56 ’ E, 60 m, 25 – 28. I. 2002, MT, Gallery forest, Frontier Project leg., MGF 002, 1 ♂ (CAS); Toliara, Zombitse National Park, near national road, 22 ° 50.43 ’ S 44 ° 43.87 ’ E, 825 m, 5 – 12. I. 2002, MT, deciduous spiny forest, R. Harin’Hala leg., 2 ♂♂ (CAS, MOLC); Toliara, Zombitse National Park, near ANGAP office, 22 ° 53.19 ’ S 44 ° 41.53 ’ E, 840 m, 15. X – 9. XI. 2001, MT, deciduous spiny forest, M. E. Irwin, F. D. Parker & R. Harin’Hala leg., MA- 02 - 13 A- 01, 1 ♂ (CAS); same locality label, 9 – 19. XI. 2001, 4 ♂♂ (3 in CAS, 1 in MOLC); Toliara, Zombitse National Park, near national road, 22 ° 50.43 ’ S 44 ° 43.87 ’ E, 825 m, 15. X – 9. XI. 2001, MT, deciduous spiny forest, R. Harin’Hala leg., MA- 02 - 13 B- 01, 2 ♂♂ (CAS, MOLC); Toliara, Zombitse National Park, 19.8 km 84 ° E Sakaraha, 22 ° 50.36 ’ S 44 ° 42.36 ’ E, 770 m, 5 – 9. II. 2003, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 7506, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 1 – 8. II. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 20 – 24. XII. 2002, 1 ♂ (CAS); same locality label, 25. XII. 2001 – 2. I. 2002, 1 ♂ (CAS); same locality label, 29. III – 10. IV. 2002, 1 ♂ (CAS); same locality label, 8 – 15. II. 2002, MA- 02 - 14 A- 15, 1 ♂ (CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue, 23 ° 41.39 ’ S 44 ° 34.53 ’ E, 180 m, 1 – 8. II. 2002, MT, spiny forest, R. Harin’Hala leg., MA- 02 - 14 B- 14, 1 ♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.22 ’ S 43 ° 28.53 ’ E, 30 m, 20 – 27. XI. 2001, MT, deciduous dry forest, R. Harin’Hala leg., MA- 02 - 18 A- 03, 1 ♂ (CAS); same locality label, 23. IX – 3. X. 2002, MA- 02 - 18 A- 34, 4 ♂♂ (3 in CAS, 1 in MOLC); same locality label, 11 – 24. II. 2003, 17 ♂♂ (7 in CAS, 10 in MOLC); same locality label, 19. II – 1. III. 2002, 1 ♂ (CAS); same locality label, 27. IV – 5. V. 2003, MA- 02 - 18 A- 53, 3 ♂♂ (2 in CAS, 1 in MOLC); same locality label, 6 – 16. I. 2002, MA- 02 - 18 A- 09, 1 ♂ (CAS); Toliara, Mikea Forest, NW of Manombo, 22 ° 54.80 ’ S 43 ° 28.93 ’ E, 37 m, 8 – 18. III. 2002, MT, spiny forest, M. Irwin & R. Harin’Hala leg., MA- 02 - 18 B- 16, 1 ♂ (CAS); Manombo, 22 ° 48.25 ’ S 43 ° 45.50 ’ E, 222 m, 23 – 28. IV. 2004, MT, gallery forest, Frontier Wilderness Project, MGF 099, 1 ♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, 85 m, 16 – 27. XII. 2002, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 - 0, 1 ♂ (CAS); same locality label, 7 – 17. III. 2003, MA- 02 - 22 - 18, 2 ♂♂ (CAS); same locality label, 15 – 25. II. 2003, MA- 02 - 22 - 16, 1 ♂ (CAS); same locality label, 17 – 26. I. 2003, MA- 02 - 22 - 13, 2 ♂♂ (CAS); same lo- cality label, 16 – 27. XII. 2002, MA- 02 - 22 - 10, 1 ♂ (CAS); same locality label, 26. I – 5. II. 2003, MA- 02 - 22 - 14, 2 ♂♂ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 35 m, 7 – 17. X. 2004, MT, spiny forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 A- 28, 1 ♂ (CAS); Toliara, Cap Ste Marie Special Reserve, 74 km S Tsihombe, 25 ° 35.26 ’ S 45 ° 09.78 ’ E, 37 m, 25. II – 7. III. 2003, MT, spiny bush, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 23 - 18, 3 ♂♂ (2 in CAS, 1 in MOLC); Toliara, Ranobe, 23 ° 02.783 ’ S 43 ° 36.617 ’ E, 20 m, 27. X – 2. XI. 2003, MT, spiny forest thicket, MGF 081, Frontier Wilderness Project, 2 ♂ m ♀ (CAS); Toliara, Fiher- enana, 23 ° 13.351 ’ S 43 ° 52.853 ’ E, 65 m, 5 – 10. VIII. 2003, MT, degraded riparian forest close to water, MGF 076, Frontier Wilderness Project, 1 ♂ (MOLC); Toliara, Fiherenana, 23 ° 14.07 ’ S 43 ° 52.15 ’ E, 50 m, 1 – 4. XII. 2002, MT, degraded gallery forest, MGF 051, Frontier Wilderness Project, 1 ♂ (CAS); Toliara, Ifaty, near Hotel Paradisia, 23 ° 10.78 ’ S 43 ° 37.01 ’ E, 9 m, 26. V – 5. VI. 2002, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02 - 16 - 28, 9 ♂♂ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI – 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 2 ♂♂ (CAS, MOLC); Toliara, Makay Mts., 21 ° 20.32 ’ S, 45 ° 10.59 ’ E, 410 m, 27 – 30. XI. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25354, 3 ♂♂ (CAS); Toliara, Makay Mts., 21 ° 18.36 ’ S, 45 ° 07.46 ’ E, 590 m, 3. XII. 2010, MT, dry forest, sandy soil, B. L. Fisher et al. leg., BLF 25550, 2 ♂♂ (CAS); Toliara, Tsifota, 20 km N Manombo, 1 km E of Tsifota village, 22 ° 49.08 ’ S 43 ° 22.36 ’ E, 15 m, 30. X – 9. XI. 2008, MT, spiny forest, M. Irwin, R. Harin’Hala leg., MG- 48 - 03, 1 ♂ (CAS); same locality label, 29. XII. 2008 – 8. I. 2009, MG- 48 - 09, 1 ♂ (CAS); same locality label, 14 – 24. II. 2009, MG- 48 - 14, 2 ♂♂ (CAS); Toliara, Beroboka village, 45 km NE Morondava, 19 ° 58.65 ’ S 44 ° 39.92 ’ E, 420 ft., 3 – 11. VIII. 2008, MT, Antsarongaza gallery forest, MG- 45 B- 33, M. Irwin, R. Harin’Hala leg., 2 ♂♂ (CAS, MOLC); same locality label, 430 ft., 20 – 28. IV. 2008, MG- 45 A- 20, 1 ♂ (MOLC); same locality label, 430 ft., 22 – 30. V. 2008, MG- 45 A- 24, 2 ♂♂ (CAS); same locality label, 420 ft., 28. IV – 6. V. 2008, MG- 45 B- 21, 1 ♂ (CAS); Toliara, Ambohimahavelona village, 33 km NE of Tulear, 23 ° 26.45 ’ S 43 ° 53.98 ’ E, 46 m, 20 – 27. II. 2009, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG- 50 B- 22, 1 ♂ (CAS); Tsimanampetsotsa National Park, Mitoho Forest, 24 ° 02.91 ’ S 43 ° 45.14 ’ E, 120 m, 9 – 15. I. 2009, MT, transitional forest, M. Irwin, R. Harin’Hala leg., MG- 53 C- 12, 2 ♂♂ (CAS). MAYOTTE: Mount Combani, 12 ° 48 ’ 23 ’’ S 45 ° 09 ’ 11 ’’ E, 370 m, 25. XI – 4. XII. 2007, rainforest, MT, B. L. Fisher et al. leg., BLF 18635, 1 ♂ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	distribution	Distribution. Madagascar, Mayotte.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2D2BCDFF3ED312FB6FFB6B.taxon	discussion	Remarks. It is possible that Thaumatodryinus merinus is the opposite sex of T. dentatus Benoit (because one ♀ and six ♂♂ have been collected in the same site in Mahajanga, 18 ° 01 ’ 35 ’’ S 44 ° 03 ’ 02 ’’ E, and 1 ♀ and 7 ♂♂ have been collected in the same site in Antsiranana, Bekaraoka Forest, 13 ° 10.00 ’ S 49 ° 42.36 ’ E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2A2BCDFF3ED37CFE14F874.taxon	description	(Fig. 161 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2A2BCDFF3ED37CFE14F874.taxon	description	Description. ♀. Fully winged; body length 3.5 mm. Testaceous-reddish, except ocellar region slightly brownish and petiole black; antenna testaceous, except antennomeres 4 – 10 darkened. Antenna clavate; antennomeres in following proportions: 7: 5: 14: 15: 22: 23: 20: 10: 10: 8. Head shiny, slightly punctate; occipital carina incomplete, only present behind lateral ocelli and on sides of vertex; OPL as long as OL. Pronotum shiny, setose, crossed by transverse impression, slightly punctate; posterior tubercle of pronotum reaching tegula. Mesoscutum shiny, setose, punctate. Notauli almost complete, posteriorly separated. Mesoscutellum and metanotum shiny, slightly punctate. Metapectal-propodeal disc reticulate rugose, with anterior third almost smooth. Propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part; 2 R 1 cell closed; pterostigma brown-light. Protarsomere 1 approximately as long as 4. Protarsomere 3 produced into hook. Enlarged claw (Fig. 161 A) with two subapical teeth and one row of 21 lamellae. Protarsomere 5 (Fig. 161 A) with two rows of approximately 38 lamellae; apex with four lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2A2BCDFF3ED37CFE14F874.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, XI. 1936, A. Seyrig leg. (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2A2BCDFF3ED37CFE14F874.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE2A2BCDFF3ED37CFE14F874.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCEFF3ED6D4FE14FB89.taxon	description	(Fig. 161 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCEFF3ED6D4FE14FB89.taxon	description	Description. ♀. Fully winged; body length 6.2 – 7.0 mm. Head black, except malar space, clypeus and mandible yellow; occasionally occiput partly brown-testaceous; antenna testaceous; pronotum and propleuron yellow; rest of mesosoma black, except propodeal spiracles brown; petiole black; metasoma reddish; legs testaceous, except club of profemur black and clubs of meso- and metafemora brown. Antenna clavate; antennomeres in following proportions: 20: 7: 40: 45: 44: 45: 35: 25: 19: 15. Head dull, reticulate rugose; occipital carina complete; OPL longer than OL. Pronotum shiny, setose, crossed by transverse impression, slightly punctate, with slight transverse striae; posterior tubercle of pronotum reaching tegula. Mesoscutum dull, completely reticulate rugose. Notauli complete, posteriorly separated. Mesoscutellum reticulate rugose. Metanotum shiny, with few anterior furrows. Metapectalpropodeal disc reticulate rugose. Propodeal declivity reticulate rugose. Forewing with one dark transverse band situated around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein longer than proximal part (34: 23). Hind wing hyaline, with one brown transverse band; distal part of 2 r-rs & Rs vein much longer than proximal part; 2 R 1 cell closed; pterostigma brown-light. Hind wing with distal brown transverse band. Protarsomere 1 approximately as long as 4. Protarsomere 3 produced into hook. Enlarged claw (Fig. 161 B) with two subapical teeth and one row of 32 lamellae. Protarsomere 5 (Fig. 161 B) with two rows of approximately 59 lamellae; apex with approximately five lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCEFF3ED6D4FE14FB89.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toamasina, Route Lakato, Ankasoka (MNHN). Paratype: same locality label as holotype, 1 ♀ (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCEFF3ED6D4FE14FB89.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCEFF3ED6D4FE14FB89.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCFFF3ED22EFCAFFE28.taxon	description	(Figs 159 D, 160 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCFFF3ED22EFCAFFE28.taxon	description	Description. ♂ (Figs 160 A, B). Fully winged; body length 2.1 – 3.5 mm. Head brown, except mandible, clypeus and anterior half of frons testaceous; antenna testaceous-dark; mesosoma brown, except pronotum and small area behind hind tegula testaceous (in paratype, small area behind fore tegula and hind tegula testaceous); metasoma brown; legs testaceous. In large specimens (more than 3.0 mm) from Kenya, 4.32664 ° S 39.52462 ° E, head testaceous-ferruginous, except transverse brown band on vertex and ocellar region; antenna testaceous-ferruginous; mesosoma testaceous-ferruginous, except posterior surface of median region of mesoscutum brown, mesoscutellum and metanotum black, mesoaxilla black (in specimen from Democratic Republic of the Congo, 04 ° 16 ’ S 17 ° 11 ’ E, mesosoma totally brown); metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 6: 6: 7: 9: 11: 11: 10: 9: 8: 10. Head shiny, convex, punctate and granulate; frontal line absent; occipital carina complete; POL = 5; OL = 2; OOL = 4.5; OPL = 0.5; TL = 2; greatest breadth of lateral ocelli as long as TL. Mesoscutum dull, granulate. Notauli incomplete, reaching about 0.7 – 0.8 × length of mesoscutum. Mesoscutellum granulate and punctate. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without strong transverse posterior keel; propodeal declivity reticulate rugose, with tracks of two median longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (18: 13). Inner side of paramere without mosaic sculpture (Fig. 159 D). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCFFF3ED22EFCAFFE28.taxon	materials_examined	Material examined. Types: ♂ holotype: KENYA: Coast Prov., Gede Forest, 3.30946 ° S 40.01941 ° E, 19 m, 22. VIII – 5. IX. 2011, MT, secondary Forest, R. Copeland leg. (NMK). Paratype: KENYA: Coast Prov., Muhaka Forest, 4.32530 ° S 39.52345 ° E, 52 m, 27 – 30. V. 2013, 6 m MT, indigenous forest, R. Copeland leg., 1 ♂ (MOLC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 30. III – 5. IV. 2006, MT, S. L. Heydon & S. E. Stevenson leg., 2 ♂♂ (DEUCD, MOLC). KENYA: Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 9 – 23. VIII. 2014, MT, indigenous forest, R. Copeland leg., 1 ♂ (NMK); same locality label, 11 – 25. X. 2015, 2 ♂♂ (NMK, MOLC); same locality label, 20. IX – 4. X. 2014, 1 ♂ (NMK); same locality label, 4 – 18. X. 2014, 1 ♂ (NMK); same locality lable, 19. VI – 3. VII. 2015, 1 ♂ (MOLC); same locality label, 15 – 29. XI. 2014, 1 ♂ (NMK); same locality label, 6 – 20. IX. 2014, 1 ♂ (NMK); same locality label, 1 – 15. XI. 2014, 1 ♂ (NMK); same locality label, 23. VIII – 6. IX. 2015, 1 ♂ (NMK); same locality label, 5 – 19. XI. 2015, 2 ♂♂ (NMK); same locality label, 9 – 23. VIII. 2015, 1 ♂ (NMK); same locality label, 27. XII. 2014 – 10. I. 2015, 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCFFF3ED22EFCAFFE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE292BCFFF3ED22EFCAFFE28.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	description	(Figs 159 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	description	Description. ♂. Fully winged; body length 1.6 – 3.0 mm. Head black or brown, except mandible testaceous; antenna testaceous, except antennomeres 3 – 10 darkened; prothorax testaceous; mesoscutum black, except notauli and medi- an region between notauli testaceous; rest of mesosoma black; metasoma brown; legs testaceous. In specimens from Kenya, 3.47908 ° S 38.34162 ° E, mesosoma black, except pronotum brown. Occasionally mesosoma brown. Antenna filiform; antennomeres in following proportions: 6: 6: 9: 12: 11: 12: 11: 10: 8: 10. Head dull, granulate and reticulate rugose; frontal line absent; occipital carina complete; POL = 6; OL = 2; OOL = 6; OPL = 1; TL = 4. Mesoscutum dull, granulate and slightly reticulate rugose. Notauli complete, posteriorly separated; minimum distance between notauli as long as POL, longer than greatest breadth of lateral ocelli (5: 4), longer than breadth of each lateral region of mesoscutum measured at posterior margin of mesoscutum (Fig. 159 F). Mesoscutellum and metanotum shiny, unsculptured (occasionally punctate, unsculptured among punctures). Metapectal-propodeal disc dull, reticulate rugose, without transverse keels; propodeal declivity reticulate rugose without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part. Inner side of paramere (Fig. 159 E) without mosaic pattern. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	materials_examined	Material examined. Type: ♂ holotype: KENYA: Coast Prov., Sokoke Forest, 8. V. 1976, Ian Bampton leg. (AEIC). Other material: KENYA: Coast Prov., Taita Hills, Chawia Forest, 3.47908 ° S 38.34162 ° E, 1614 m, 24. VII – 7. VIII. 2011, MT, next to small forest pond, R. Copeland leg., 1 ♂ (NMK); same locality label, 7 – 21. VIII. 2011, 1 ♂ (NMK); same locality label, 27. V – 10. VI. 2012, 1 ♂ (MOLC); Coast Prov., Taita Hills, Vuria Forest, 3.41428 ° S 38.29178 ° E, 2162 m, 11 – 25. I. 2012, MT, just inside indigenous forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Kaya Kinondo, 4.39382 ° S 39.54567 ° E, 10 m, 22. VIII – 5. IX. 2011, MT, coral rag canopy forest, R. Copeland leg., 1 ♂ (NMK); Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 12 – 26. VII. 2014, MT, indigenous forest, R. Copeland leg., 2 ♂♂ (NMK, MOLC); same locality label, 23. VIII – 8. IX. 2015, 1 ♂ (NMK); same locality label, 18. V – 1. VI. 2016, 1 ♂ (NMK); same locality label, 6 – 20. IX. 2015, 1 ♂ (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE282BC0FF3ED079FCF8FC20.taxon	discussion	Remarks. The ♂ tentatively attributed to Pseudodryinus townesi by Olmi (1984) cannot be a Pseudodryinus, because the minimum distance between notauli is longer than the greatest breadth of the lateral ocelli. For this reason, it is now associated with Thaumatodryinus sokokensis. However, it may be the opposite sex of one of the other African species known only on the basis of ♀ specimens.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE272BC0FF3ED5A0FE59F874.taxon	description	(Figs 160 C, D, 161 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE272BC0FF3ED5A0FE59F874.taxon	description	Description. ♀ (Figs 160 C, D). Fully winged; body length 3.4 – 3.7 mm. Head black, except mandible testaceous and clypeus partly testaceous and brown; antenna testaceous; mesosoma black, except disc and lateral regions of pronotum, including pronotal tubercle, testaceous; petiole black; mesosoma brown; legs testaceous-whitish, except clubs of femora brown. Antenna clavate; antennomeres in following proportions: 14: 7: 26: 29: 35: 34: 25: 14: 11: 13. Head dull, granulate and reticulate rugose; occipital carina incomplete, only present behind ocellar triangle and on sides of lateral ocelli, laterally not reaching eyes; POL = 8; OL = 3; OOL = 12; OPL = 2; temple absent; greatest breadth of lateral ocelli about as long as OL. In specimen from Kenya, 4.32664 ° S 39.52462 ° E, POL = 6; OOL = 10. Pronotum dull, crossed by one transverse furrow, granulate and transversely striate; posterior tubercle reaching tegula. Mesoscutum dull, completely granulate and slightly reticulate rugose (in specimen from Kenya, 4.32664 ° S 39.52462 ° E, mesoscutum strongly reticulate rugose and granulate). Notauli complete, posteriorly separated; minimum distance between notauli slightly longer than POL (9: 8). Mesoscutellum shiny, slightly reticulate rugose. Metanotum shiny, unsculptured. Metapectal-propodeal disc reticulate rugose. Propodeal declivity reticulate rugose. Forewing with one dark transverse band around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein longer than proximal part (32: 15). Hind wing hyaline, without brown transverse bands Protarsomeres in following proportions: 23: 3: 7: 24: 37. Protarsomere 3 produced into hook. Enlarged claw (Fig. 161 C) with two subapical teeth and one row of 27 lamellae. Protarsomere 5 (Fig. 161 C) with two rows of approximately 38 lamellae; apex with approximately six lamellae. Tibial spurs 1 / 1 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE272BC0FF3ED5A0FE59F874.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Coast Prov., Taita Hills, Vuria Forest, 3.41428 ° S 38.29178 ° E, 2162 m, 28. XII. 2011 – 11. I. 2012, MT, just inside indigenous forest, R. Copeland leg. (NMK). Other material. KENYA: Coast Prov., Muhaka Forest, 4.32664 ° S 39.52462 ° E, 41 m, 18. X – 1. XI. 2014, MT, indigenous forest, R. Copeland leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE272BC0FF3ED5A0FE59F874.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE272BC0FF3ED5A0FE59F874.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE202BC7FF3ED6D7FC3CFDB2.taxon	type_taxon	Type species. Adryinus cerrutii (Benoit 1951), by original designation.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE202BC7FF3ED6D7FC3CFDB2.taxon	diagnosis	Diagnosis. ♀: Fully winged (Figs 162 A, B); palpal formula 6 / 3; antenna with ADOs; pronotum crossed by strong transverse furrow (Figs 162 B, 164 C); notauli absent; enlarged claw with one subapical tooth and lamellae, or peg – like setae, or bristles (Figs 163 A, B); tibial spurs 1 / 0 / 1. ♂: Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE202BC7FF3ED6D7FC3CFDB2.taxon	distribution	Distribution. Afrotropical, Oriental, Nearctic, Neotropical.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE202BC7FF3ED6D7FC3CFDB2.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE202BC7FF3ED6D7FC3CFDB2.taxon	discussion	World species. 12 species are known, eight in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	description	(Fig. 163 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	description	Description. ♀. Fully winged; body length 5.0 – 5.6 mm. Head black, except mandible reddish; antenna testaceous, except antennomeres 7 – 10 black; occasionally antenna totally black or brown; mesosoma black; occasionally lateral regions and posterior margin of pronotum reddish; metasoma brown or brown-reddish; legs black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 40: 14: 58: 41: 28: 20: 18: 14: 13: 19. Head shiny, excavated, granulate, with many parallel keels on frons, vertex and occiput; occipital carina absent; POL = 14; OL = 6; OOL = 30. Palpal formula 6 / 3. Pronotum crossed by transverse impression, dull, granulate and sculptured by some striae on collar and around disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, granulate and with longitudinal keels. In specimen from Uganda, 01 ° 45 ’ N 31 ° 35 ’ E, mesoscutum with many longitudinal keels crossed by many transverse keels (so mesoscutum apparently reticulate rugose). Notauli absent. In specimen from Uganda, 01 ° 45 ’ N 31 ° 35 ’ E, mesoscutum with two complete longitudinal keels similar to notauli. Mesoscutellum granulate and with many longitudinal keels; in specimen from Uganda, 01 ° 45 ’ N 31 ° 35 ’ E, mesoscutum completely reticulate rugose. Metanotum very reduced and short. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately 1.4 – 1.5 as long as proximal part. Protrochanter approximately 4 × as long as broad. Protarsomere 1 longer than 4 (24: 14). Enlarged claw (Fig. 163 A) with one subapical tooth and 11 peg-like setae. Protarsomere 5 (Fig. 163 A) with two rows of 16 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown (see remarks).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Loashi Valley, 01 ° 22 ’ S 28 ° 47 ’ E, VIII. 1937, J. Ghesquière leg. (MRAC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Lomami, Upemba National Park, Kaziba, 2 ♀♀ (MRAC, AMNH). UGANDA: Ruzumbusa, V. 1939, T. H. C. Taylor leg., 1 ♀ (NHMUK); Namutamba, 2 ♀♀ (AMNH, NHMUK); Masindi District, Budongo Forest near Sonso, 01 ° 45 ’ N 31 ° 35 ’ E, VI. 1995, Th. Wagner leg., 1 ♀ (CNC). ZIMBABWE: Harare (= Salisbury), Chishawasha, II. 1979, A. Watsham leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	distribution	Distribution. Democratic Republic of the Congo, Uganda, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3F2BD8FF3ED529FE51F870.taxon	discussion	Remarks. The ♂ allotype of A. bellicosus, described by Benoit (1951 c), belongs to a different genus, because the palpal formula is 4 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3E2BDAFF3ED32BFE1FFD00.taxon	description	(Fig. 163 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3E2BDAFF3ED32BFE1FFD00.taxon	description	Description. ♀. Fully winged; body length 6.2 – 6.6 mm. Head black, except mandible (except brown teeth) and clypeus testaceous (except brown median stripe); antenna of holotype black, except scape, pedicel and flagellomeres 4 – 8 testaceous; antenna of paratype brown, except scape, pedicel and flagellomeres 3 – 8 testaceous; mesosoma black; metasoma brown; proleg brown, except tarsus, articulations and part of coxa testaceous; meso- and metalegs brown, except tarsi and distal extremities of coxae and trochanters testaceous. Antenna clavate; antennomeres in following proportions: 13: 8: 38: 25: 21: 17: 13: 11: 9: 11; antennal ADOs present in antennomeres 5 – 10. Head dull, weakly excavated, granulate and reticulate rugose; frontal line complete; occipital carina incomplete, only present behind and shortly on sides of lateral ocelli (on sides reaching about 0.6 × length of OOL); POL = 5; OL = 5; OOL = 17; OPL = 2 (but branch of occipital carina curving and reaching inner side of lateral ocelli); temple distinct. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, dull, setose, granulate and reticulate rugose. with many longitudinal keels on sides; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum and mesoscutellum dull, granulate and reticulate rugose. Notauli hardly visible among areolae of mesoscutum, apparently complete and posteriorly very close, but separated. Metanotum dull, short, transverse, rugose. Metapectal-propodeal disc dull, reticulate rugose, without strong transverse posterior keel; Propodeal declivity with two long, complete and subparallel longitudinal keels (keels begining at posterior apex and reaching anterior margin of metapectal-propodeal disc); median and lateral areas of propodeal declivity reticulate rugose; median area composed of areolae less wide than those of lateral areas. Mesopleuron and metapleuron dull, granulate and reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately as long as proximal part (10: 10). Hind wing hyaline. Protarsomeres in following proportions: 28: 5: 11: 37: 55. Enlarged claw (Fig. 163 B) with one strong subapical tooth and one row of 14 lamellae. Protarsomere 5 (Fig. 163 B) with two rows of 19 + 23 lamellae; apex with at least 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3E2BDAFF3ED32BFE1FFD00.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: KwaZulu – Natal, near Hluhluwe, Zulu Nyala Pri- vate Game Reserve, 18. XI. 1985, Eragrostis patch, Acacia veld (DJBC). Paratype: SOUTH AFRICA: KwaZulu – Natal, near Hluhluwe, Umziki Pan Reserve, 16 m, XII. 1990, P. E. Reavell leg., 1 ♀ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3E2BDAFF3ED32BFE1FFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3E2BDAFF3ED32BFE1FFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDAFF3ED497FDDFF987.taxon	description	(Figs 162 A, B, 163 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDAFF3ED497FDDFF987.taxon	description	Description. ♀ (Figs 162 A, B). Fully winged; body length 6.0 mm. Head black, except mandible partly reddish; antenna black, except ventral side of scape yellow; mesosoma and metasoma black; legs black, except tarsi testaceous. Head dull, excavated, setose, granulate, with temple prominent, with many parallel keels on frons, vertex and occiput; frontal line incomplete, absent near clypeus; occipital carina absent; POL = 17; OL = 9; OOL = 37. Palpal formula 6 / 3. Pronotum (Fig. 162 B) crossed by transverse impression, granulate and sculptured by many keels around disc and collar; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, completely reticulate rugose. Notauli absent. Mesoscutellum completely reticulate rugose, unsculptured among areolae. Metanotum very reduced and short. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Mesopleuron very prominent. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein slightly longer than proximal part (10: 9). Protrochanter approximately 4 × as long as broad. Protarsomere 1 longer than 4 (30: 15). Enlarged claw (Fig. 163 C) with one subapical tooth and three bristles. Protarsomere 5 (Fig. 163 C) with two rows of 13 lamellae; apex with 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDAFF3ED497FDDFF987.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Central Prov., Nairobi, III. 1945, S. Patrizi leg. (MRAC). Other material: ZIMBABWE: Victoria Falls, 25. IX, J. O. leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDAFF3ED497FDDFF987.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDAFF3ED497FDDFF987.taxon	distribution	Distribution. Kenya, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDBFF3ED01CFE1FFD00.taxon	description	(Figs 162 C, 163 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDBFF3ED01CFE1FFD00.taxon	description	Description. ♀ (Fig. 162 C). Fully winged; body length 4.0 mm. Head black, except mandible testaceous; antenna testaceous, except antennomeres 6 – 10 brown; mesosoma black, except sides of pronotum testaceous; metasoma brown, except distal extremity testaceous; legs brown, except articulations and tarsi testaceous. Antenna clavate; antennomeres in following proportions: 12: 5: 19: 12: 9: 7: 5.5: 5: 4.5: 7; antennal ADOs present on antennomeres 6 – 10. Head dull, weakly excavated, granulate and with many longitudinal and parallel keels on frons, vertex and occiput; frontal line complete; occipital carina absent; POL = 5; OL = 3; OOL = 11; temple distinct, short. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, dull, granulate and sculptured by many longitudinal and paral- lel striae on anterior collar, lateral regions and disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, granulate and sculptured by many parallel and longitudinal keels. Notauli absent. Mesoscutellum dull, granulate, completely reticulate rugose. Metanotum dull, short, transverse, rugose. Metapectal-propodeal disc dull, without transverse posterior keel, sculptured by many parallel and longitudinal keels (with many transverse keels among longitudinal keels); propodeal declivity reticulate rugose, without longitudinal keels. Mesopleuron and metapleuron dull and reticulate rugose. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (9: 6). Hind wing hyaline. Protarsomeres in following proportions: 19: 2: 6.5: 10: 17. Enlarged claw (Fig. 163 D) without subapical tooth and with one row of three bristles. Protarsomere 5 (Fig. 163 D) with two rows of 4 – 6 lamellae; apex with at least 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDBFF3ED01CFE1FFD00.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Gauteng, Roodeplaat Horticultural Research Sta- tion, 25 ° 37 ’ S 28 ° 20 ’ E, 2. IX. 1986, J. Donaldson leg. (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDBFF3ED01CFE1FFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3D2BDBFF3ED01CFE1FFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDBFF3ED497FE1FF9AE.taxon	description	(Fig. 163 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDBFF3ED497FE1FF9AE.taxon	description	Description. ♀. Fully winged; body length 4.7 mm. Head black, except mandible and clypeus ferruginous-reddish; antenna testaceous, except antennomeres 7 – 10 brown; mesosoma black, except lateral regions, anterior and posterior margin of pronotum, one dorsal stripe of propleuron reddish; metasoma ferruginous, except a few brown areas; legs ferruginous-testaceous. Antenna clavate; antennomeres in following proportions: 15: 6: 22: 15: 11: 8: 6: 5: 5: 7. Head dull, excavated, granulate and reticulate rugose, without many parallel keels on frons and vertex; occiput granulate and sculptured by many longitudinal keels; frontal line complete; occipital carina absent. POL = 6; OL = 2.5; OOL = 12. Palpal formula 6 / 3. Pronotum crossed by transverse impression, dull, granulate and sculptured by many longitudinal striae on collar and around the disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, granulate and with many longitudinal keels. Notauli absent. Mesoscutellum granulate and reticulate rugose. Metanotum very reduced and short, reticulate rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two short longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately 1.6 as long as proximal part (8: 5). Protarsomeres in following proportions: 22: 3: 7: 12: 22. Enlarged claw (Fig. 163 E) with one subapical tooth and four bristles. Protarsomere 5 (Fig. 163 E) with two rows of 7 + 6 lamellae; apex with at least 30 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDBFF3ED497FE1FF9AE.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Royal Natal Nature Reserve, 28 ° 41.4 ’ S 28 ° 56.3 ’ E, 1420 m, 10 – 13. XII. 2004, MT, M. Mostovski leg. (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDBFF3ED497FE1FF9AE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDBFF3ED497FE1FF9AE.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDCFF3ED035FE22FDB8.taxon	description	(Fig. 164 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDCFF3ED035FE22FDB8.taxon	description	Description. ♀. Fully winged; body length 4.1 mm. Head black, except mandible reddish; antenna testaceous; me- sosoma black, except lateral regions of pronotum reddish; metasoma brown-black; legs brown-black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 24: 17: 62: 36: 28: 23: 19: 15: 14: 19. Head dull, weakly excavated, granulate and completely reticulate rugose; occiput granulate, without areolae or keels; occipital carina incomplete, reaching approximately 0.5 × length of OOL; POL = 7; OL = 6; OOL = 25. Palpal formula 6 / 3. Pronotum crossed by transverse impression, dull, granulate and sculptured by some striae on collar around disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum completely reticulate rugose. Notauli absent. Mesoscutellum completely reticulate rugose. Metanotum very reduced and short. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately 1.66 as long as proximal part. Protrochanter approximately 4 × as long as broad. Protarsomere 1 longer than 4 (22: 14). Enlarged claw (Fig. 164 A) with one sub- apical tooth and 12 peg-like setae. Protarsomere 5 (Fig. 164 A) with two rows of nine lamellae; apex with at least six lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDCFF3ED035FE22FDB8.taxon	materials_examined	Material examined. Type: ♀ holotype: NIGERIA: Niger State, near Mokwa, Zugurma, 26. XII. 1971, J. G. Deemingkurmi leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDCFF3ED035FE22FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3C2BDCFF3ED035FE22FDB8.taxon	distribution	Distribution. Nigeria.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3B2BDDFF3ED026FDB9FD9C.taxon	description	(Fig. 164 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3B2BDDFF3ED026FDB9FD9C.taxon	description	Description. ♀. Fully winged; body length 5.5 – 6.0 mm. Head black, except mandible distally testaceous; antenna black; mesosoma and metasoma black; legs black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 22: 7: 29: 17: 13: 10: 7: 5: 5: 8. Head shiny, excavated, granulate, with many areolae and longitudinal keels; occiput sculptured by longitudinal keels; frontal line present; occipital carina absent; POL = 7; OL = 3; OOL = 16; temple very prominent. Palpal formula 6 / 3. Pronotum crossed by transverse impression, dull, granulate and sculptured by irregular keels; anterior margin of collar strongly excavated; disc humped, but not pointed; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, with many longitudinal and transverse keels. Notauli absent. Mesoscutellum reticulate rugose. Metanotum very short, reticulate rugose. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal and transverse keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately as long as proximal part. Protarsomeres in following proportions: 29: 4: 8: 13: 26. Enlarged claw (Fig. 164 B) with one subapical tooth and five bristles. Protarsomere 5 (Fig. 164 B) with two rows of 32 lamellae extending continuously to apex. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3B2BDDFF3ED026FDB9FD9C.taxon	materials_examined	Material examined. Type: ♀ holotype: UGANDA: Kampala (AEIC). Other material: IVORY COAST: Bouaké Department, Bouaké, 11. X. 1980, Pitfall trap, 1 ♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3B2BDDFF3ED026FDB9FD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3B2BDDFF3ED026FDB9FD9C.taxon	distribution	Distribution. Ivory Coast, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDDFF3ED423FE1FFA7C.taxon	description	(Figs 164 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDDFF3ED423FE1FFA7C.taxon	description	Adryinus rugidorsalis (Turner): Olmi 1984: 1135; 2006: 44. Description. ♀. Fully winged; body length 4.7 mm. Head black, except clypeus and mandible reddish; antenna testaceous-reddish; mesosoma black, except lateral regions and posterior margin of pronotum reddish; metasoma testaceous, except tergite 1 partly black; legs testaceous-reddish. Antenna clavate; antennomeres in following proportions: 35: 13: 51: 33: 24: 15: 13: 12: 11: 17. Head shiny, weakly excavated, reticulate rugose on frons and vertex; occiput sculptured by many keels; occipital carina absent. POL = 11; OL = 6; OOL = 29. Palpal formula 6 / 3. Pronotum (Fig. 164 C) crossed by transverse impression, dull, granulate and sculptured by some striae on collar and around the disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, granulate and reticulate rugose. Notauli absent. Mesoscutellum granulate and reticulate rugose. Metanotum very reduced and short. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein approximately as long as proximal part. Protrochanter approximately 4 × as long as broad. Protarsomere 1 longer than 4 (21: 8). Enlarged claw (Fig. 164 D) with one subapical tooth and five bristles. Protarsomere 5 (Fig. 164 D) with two rows of 14 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDDFF3ED423FE1FFA7C.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, XI. 1923, R. E. Turner leg. (NHMUK). Paratype: same locality label as holotype, XII. 1923, 1 ♀ (NHMUK). Other material: same locality label as holotype, 7 – 13. VIII. 1923, 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDDFF3ED423FE1FFA7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDDFF3ED423FE1FFA7C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDEFF3ED003FC2EFEC4.taxon	type_taxon	Type species. Echthrodelphax fairchildii R. Perkins 1903, by monotypy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDEFF3ED003FC2EFEC4.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 162 D, 165 A, B); rarely brachypterous (only in one specimen of E. rufus Olmi 1984 from China, Guangdong, Yunjishan Nature Reserve); palpal formula 3 / 2, 4 / 2, 5 / 2, 5 / 3, or 6 / 3; antenna with ADOs; pronotum crossed by strong transverse furrow (Fig. 162 D); notauli complete (Figs 165 A, B); enlarged claw with one subapical tooth and one row of lamellae (Figs 166 A, D); protarsomere 5 with less than 15 medial lamellae (Figs 166 A, D); tibial spurs 1 / 0 / 1. ♂: fully winged (Figs 165 C, D); head with distinct complete occipital carina; oc- ciput concave; temple present; palpal formula 3 / 2, 4 / 2, 5 / 2, 5 / 3, or 6 / 3; tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDEFF3ED003FC2EFEC4.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDEFF3ED003FC2EFEC4.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino & Olmi 1997, 2006; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE3A2BDEFF3ED003FC2EFEC4.taxon	discussion	World species. 15 species are known, two in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	description	(Figs 165 B, 166 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	description	Description. ♀ (Fig. 165 B). Fully winged; body length 2.2 – 2.3 mm. Head testaceous, except brown transverse band on vertex; antenna testaceous; propleuron black, occasionally testaceous; pronotum totally black or with collar black and disc testaceous or reddish (occasionally pronotum completely testaceous); rest of mesosoma black; metasoma black; legs testaceous. Antenna clavate. Head shiny, unsculptured, except occiput and anterior region of frons granulate; occipital carina absent; ocellar triangle strongly isosceles, with median keel; OL much longer than POL. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, with disc granulate and collar unsculptured; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, unsculptured. Notauli complete, joint at posterior margin of mesoscutum. Mesoscutellum shiny, unsculptured. Metanotum very reduced and short. Metapectal-propodeal disc reticulate rugose, with transverse keels; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands. In specimen from Kenya, Kasigau Mtn., forewing with small dark spot around 2 r-rs & Rs vein. Protrochanter approximately 4 × as long as broad. Enlarged claw (Fig. 166 A) with one large subapical tooth and one row of four lamellae. Protarsomere 5 (Fig. 166 A) with one row of 11 lamellae; apex with approximately 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.3 – 1.4 mm. Head brown-black, except mandible and clypeus testaceous; antenna brown; mesosoma black; metasoma brown-testaceous; legs testaceous. Antenna filiform; antennomeres in following proportions: 4: 3.5: 7: 7: 7: 7: 7: 7: 6.5: 8. Head flat, shiny, finely punctate, unsculptured among punctures; frontal line absent; occipital carina very thin and incomplete, not present behind ocellar triangle; temple very prominent; POL = 3; OL = 1; OOL = 2.5; TL = 2.5; lateral ocelli touching occipital carina. Mesoscutum shiny, finely punctate, unsculptured among punctures. Notauli complete, posteriorly joint or separated. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, completely reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (11: 4). Paramere (Fig. 166 B) with dorsal process very long and sculptured with a mosaic pattern (figure 9 in Olmi (1989 b) was erroneously associated with E. migratorius, whereas it is actually related to E. tauricus Ponomarenko). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, IV. 1942, A. Seyrig leg. (MRAC). Other material: BOTSWANA: Serowe, Farmer’s Brigade, 1 ♀ (USNM). BURKINA FASO: Kou River Sources, 1 ♀ (FSAE). CAMEROON: South, near Ebolowa, Nkoemvon, 21. I – 6. II. 1980, cocoa plantation, D. Jackson leg., 1 ♀ (NHMUK); Southwest, Mt. Cameroun, Buea, 1200 m, 1 ♀ (ISNB). CAPE VERDE ISLANDS: Santiago Island, São Jorge dos Orgãos, A. van Harten leg., 2 ♀♀ (AMNH). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele (= Haut-Zaire), Garamba National Park, 2 ♀♀ (MRAC, AMNH); North Kivu, Edward Lake, 1 ♀ (MRAC). IVORY COAST: 8 km NW of Bouaflé, Klébo, 1 ♀ (DEUW); 10 km SE of Bouaflé, Koudougou, 1 ♀ (DEUW); 20 km E Katiola, Logbonou, 1 ♀ (DEUW). KENYA: Coast Prov., Kasigau Mtn., 3.82667 ° S 38.64982 ° E, 1117 m, indigenous forest, MT, next to spring in forest, 30. VI – 13. VII. 2011, R. Copeland leg., 1 ♀ (RSC); Coast Prov., Boni Forest area, near Bodhei Village, 1.85551 ° S 40.69889 ° E, 30 m, 6 – 18. VI. 2013, 2 m MT, mixed grass- and woodland, J. Bukhebi & R. Copeland leg., 1 ♀ (NMK); Eastern Prov., Ngaia Forest, bottom of forest, 0.32442 ° N 38.05038 ° E, 1057 m, 24. XII. 2011 – 7. I. 2012, MT, inside indigenous forest, R. Copeland leg., 2 ♀♀ (NMK). MADAGASCAR: Anta- nanarivo, Tsimbazaza Park, 21 – 25. X. 1984, MT, Robert W. Brooks leg., 1 ♀ (NAMUK); Antsiranana, Montagne d’Ambre National Park, 12 ° 30 ’ 52 ’’ S 49 ° 10 ’ 53 ’’ E, 960 m, 19. III – 6. IV. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 A- 09, 1 ♀ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 14 – 30. V. 2001, R. Harin’Hala leg., MT, MA 01 - 01 D- 11, 1 ♀ (MOLC); Antsiranana, Montagne des Français, 12 ° 18.8 ’ S 49 ° 38.51 ’ E, 150 m, 15. II – 8. III. 2001, MT, along forested limestone ridge, R. Harin’Hala leg., MA- 01 - 06 - 06, 1 ♀ (MOLC); Antsiranana, 7 km N of Joffreville, 12 ° 20 ’ S 49 ° 15 ’ E, 300 m, 7 – 27. IV. 2001, dry forest, MT, R. Harin’Hala leg., MA- 01 - 07 - 10, 1 ♂ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 3.4 km 93 ° E Bekopaka, Tombeau Vazimba, 19 ° 08.31 ’ S 44 ° 49.41 ’ E, 50 m, 6 – 10. XI. 2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233, 2 ♂♂ (CAS); Toliara, Bekily, IV. 1942, A. Seyrig leg., 1 ♀ (MNHN); Toliara, 60 km NE of Morondava, Bere- boka, 1 ♀ (AMNH); Beza Mahafaly Reserve, parcelled I near research station, 23 ° 41.19 ’ S 44 ° 35.46 ’ E, 165 m, 25. XII. 2001 – 2. I. 2002, MT, dry deciduous forest, R. Harin’Hala leg., MA- 02 - 14 A- 14, 1 ♀ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, elev 35 m, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 22 – 30. XI. 2003, M. Irwin, F. Parker & R. Harin’Hala, MT, spiny forest, MA- 02 - 22 A- 04, 1 ♀ (CAS); Toliara, Berenty Reserve, 80 km W Fort Dauphin, 25 ° 00 ’ S 46 ° 17 ’ E, 7. IV. 1994, MT, M. Wasbauer leg., 1 ♀ (DEUCD); Toliara, Manderano, 23 ° 31.39 ’ S 44 ° 5.18 ’ E, 70 m, 23. VI – 18. VII. 2002, MT, edge of marsh, 5 m from road, gallery forest, MGF 035, Frontier Wilderness Project, 1 ♀ (CAS); same locality label, 23. VI – 27. VII. 2002, MT, edge of marsh, 5 m from road, gallery forest, MGF 036, Frontier Wilderness Project, 2 ♀♀ (CAS). MALI: Sikasso, 1 ♀ (MRAC). MOZAMBIQUE: Gaza Prov., between Macia and Palmeira, 2 km E of Incomati River Bridge along Road EN 1, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 3 km N of Umbelúzi along Maputo-Boane Road, M. Olmi leg., 2 ♀♀ (AMNH); Maputo Prov., Manhiça District, Manhiça, Vasco’s Farm, 1 – 8. IX. 1996, MT, M. Olmi leg., 2 ♀♀ (MOLC); same locality label, 9. IX – 24. X. 1995, 2 ♀♀ (MOLC); Niassa Prov., Cuamba, Catholic University of Mozambique Campus, C. 7. IV. 2003, B. 9. IV. 2003, Sf. 24. IV. 2003, M. Olmi reared ex nymph of an unknown species of Delphacidae, 1 ♂ (MOLC); Niassa Prov., Cuamba, Catholic University of Mozambique Farm, 17. IV – 2. VI. 2006, MT, M. Olmi leg., 1 ♀ (MOLC). NIGER: Maradi, Tarna goul- bin, 5 – 10. X. 1985, MT, G. J. Steck leg., 1 ♂ (TAMU). OMAN: Muscat, 5. IV. 1985, 1 ♀ (ZIL). SENEGAL: 14 km S of Ross Bethio, 7. X. 1978, 1 ♀ (USNM). SIERRA LEONE: 6 km SW of Freetown, 08 ° 26 ’ N 13 ° 16 ’ W, 26. XI. 1993, 1 ♀ (ZIL). SOMALIA: Benadir Prov., Afgoi, M. Olmi leg., 3 ♀♀ (AMNH). SOUTH AFRICA: Eastern Cape, 8 km N of Fort Beaufort, Blinkwater, 32 ° 43 ’ S 26 ° 35 ’ E, 1 ♀ (ZIL); Eastern Cape, Port St. Johns, 5 – 30. IV. 1923, R. E. Turner leg., 1 ♂ (NHMUK); same locality label, 1 – 5. IV. 1923, 25 – 31. III. 1923, 2 ♀♀ (NHMUK); same locality label, 1 ♀ (DASB); Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ‘ S 24 ° 55.962 ‘ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC); KwaZulu-Natal, Pietermaritzburg, Hilton, gar- den, 19 – 24. IV. 2005, YPT, V. Kolyada leg., 1 ♂ (NMSA); KwaZulu-Natal, Pietermaritzburg, Fernkliff, 22. II. 2004, sweeping, Kolyada & Mostovski leg., 1 ♀ (SAMC); KwaZulu-Natal, Louwsberg, Sanyati Farm, 1090 m, 27 ° 34 ’ S 31 ° 17.9 ’ E, 30. X – 18. XII. 2005, MT, M. Mostovski leg., 1 ♂ (NMSA); Limpopo, Klaserie, 1 ♀ (PMA); Limpopo, 15 km NW of Klaserie, Guernsey Farm, 1 ♀ (PMA); North West, Derdepoort, III. 1958, D. P. Annecke leg., 1 ♀ (NHMUK); Western Cape, Salmanslaagte, 32 ° 03.228 ‘ S 19 ° 06.258 ‘ E, 459 m, 26. II. 2009, sweep, Cederbeg Sandstone Fynbos, S. van Noort leg., 1 ♂ (SAMC); same locality label, 26 – 28. II. 2009, YPT, 2 ♂♂, 1 ♀ (SAMC). TANZANIA: Arusha, 1 ♀ (HNHM). UGANDA: Kawanda, X. 1942, T. H. C. Taylor leg., 1 ♂ (NHMUK). UNITED ARAB EMIR- ATES: Abu Dhabi, Houbara Protected Area, 1. XI – 10. XII. 2015, MT, N. 635, A. Saji & A. van Harten leg., 1 ♂ (MOLC). YEMEN: Qa Al Boun, IV. 1993, M. Knapp leg., 1 ♀ (AMNH); ZIMBABWE: Harare, 1 ♀ (CNC); Harare, Chishawasha, V. 1978, A. Watsham leg., 1 ♀ (NHMUK). Palaearctic. Known from EGYPT (Hendawy 2001): Kafr El-Sheikh Governorate, near Kafr El-Sheikh, 11. VIII. 1999, in rice fields, A. S. Hendawi leg., many ♀♀ and ♂♂ (MOLC); Gharbia Governorate, near Tanta, A. Hendawy leg., 1 ♀ (AHC); Damietta Governorate, Shirbin, S of El- Serwy, XI. 2000, P. Skidmore leg. (JTBC) (Burn 2011); Sharkia Governorate, Kafr Sakr, June 2006, in rice field, A. S. Hendawy leg. (AHC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	biology_ecology	Hosts. Delphacidae Delphacinae (Guglielmino et al. 2013): in Egypt: nymphs of Sogatella furcifera (Horváth) and S. vibix (Haupt); in Mozambique: Sogatella petax Fennah; in South Africa: Megamelus scutellaris Berg (new record: reared from Megamelus scutellaris colonies of Neotropical origin (imported from Argentina via USA) that are being mass reared at Rhodes University in Grahamstown for biocontrol against water hyacinth, Eichhornia crassipes Mart. (Solms) (Pontederiaceae )).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	distribution	Distribution. Botswana, Burkina Faso, Cameroon, Cape Verde Islands, Democratic Republic of the Congo, Ivory Coast, Kenya, Madagascar, Mali, Mozambique, Niger, Oman, Senegal, Sierra Leone, Somalia, South Africa, Tanzania, Uganda, United Arab Emirates, Yemen, Zimbabwe, in addition to Egypt (Palaearctic).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE392BD0FF3ED5BBFB6CFC9F.taxon	discussion	Remarks. Echthrodelphax migratorius was recorded by mistake from many further localities in Yemen (Olmi & Van Harten 2000, 2006). These records (all ♂♂) are actually of E. tauricus (see below). The unique locality in Yemen, where E. migratorius was definitely collected (one ♀), is Qa Al Boun (see above).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE372BD4FF3ED527FD0EFE0C.taxon	description	(Figs 162 D, 165 A, C, D, 166 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE372BD4FF3ED527FD0EFE0C.taxon	description	Description. ♀ (Figs 162 D, 165 A). Fully winged; body length 1.9 – 3.1 mm. Head testaceous, occasionally except brown spot on vertex; antenna testaceous; propleuron, pronotum and mesoscutum testaceous, occasionally brownreddish; rest of mesosoma black, except distal extremity of propodeal declivity testaceous; occasionally sides of mesoscutellum and metanotum brown; occasionally metapectal-propodeal disc with brown spot; in specimen from South Africa, 25 ° 48 ’ S 27 ° 34 ’ E, mesosoma testaceous-reddish, except axillae, metapectal-propodeal disc and propodeal declivity brown; in two specimens from South Africa, 34 ° 15 ’ S 18 ° 21.5 ’ E, mesoscutum testaceous, except posterior half black; petiole black; metasoma yellow, except tergite 1 partly black and with some brown transverse bands; legs testaceous. Specimen from Mozambique, Mituque (12. IV – 6. V. 2008), completely testaceous, except petiole black and axillae darkened. Antenna clavate. Head shiny, unsculptured; occasionally occiput and part of vertex granulate; occipital carina absent; ocellar triangle strongly isosceles, with median keel; frontal line present; POL = 1; OL = 4; OOL = 7. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, with disc dull and granulate; collar and lateral regions of pronotum smooth, shiny and unsculptured; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, smooth, unsculptured or slightly punctate. Notauli complete, joint at posterior margin of mesoscutum. Occasionally notauli incomplete, reaching about 0.7 × length of mesoscutum. In testaceous specimen from Mozambique, Mituque (12. IV – 6. V. 2008), notauli absent. Mesoscutellum shiny, unsculptured or slightly puntate. Metanotum very reduced and short. Metapectal-propodeal disc and propodeal declivity reticulate rugose, without longitudinal keels. Forewing with 1 – 2 more or less large dark transverse bands; occasionally dark transverse bands little distinct and hardly visible. Protrochanter approximately 4 × as long as broad. Enlarged claw (Fig. 166 D) with one large subapical tooth and one row of 4 – 5 lamellae. Protarsomere 5 (Fig. 166 D) with one row of 7 – 11 lamellae; apex with 6 – 11 lamellae. Tibial spurs 1 / 0 / 1. Description. ♂ (Figs 165 C, D). Fully winged; body length 1.2 – 1.9 mm. Head black, except mandible testa- ceous; clypeus black, occasionally testaceous; antenna brown, occasionally except scape and pedicel testaceous; mesosoma black; metasoma brown or brown-reddish; legs testaceous. Antenna filiform, with antennomere 3 four or more than 4 × as long as broad; antennomeres in following proportions: 4: 5: 8: 8: 8: 7.5: 7.5: 7.5: 7: 10. Head shiny, unsculptured or granulate; frontal line absent; occipital carina very thin and complete; temple very prominent; POL = 4.5; OL = 2; OOL = 4; OPL = 1; TL = 5.5. Palpal formula 4 / 2 or 5 / 2. Mesoscutum shiny, slightly granulate. Notauli complete, posteriorly joint or separated. Specimen from Yemen (Lahj, III. 2001, N. 5578) with incomplete notauli reaching approximately 0.7 × length of mesoscutum (erroneously tentatively attributed to E. migratorius by Olmi 1989 b, Olmi & van Harten 2000, 2006). Mesoscutellum and metanotum shiny, unsculptured. Metapectalpropodeal disc dull, completely reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 6). Paramere (Fig. 166 C) with dorsal process short and lobe-shaped, without mosaic pattern. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE372BD4FF3ED527FD0EFE0C.taxon	materials_examined	Material examined. Types: ♀ holotype of E. tauricus: UKRAINE: Crimea, Sevastopol, 20 – 23. IX. 1968, N. G. Ponomarenko leg. (ZMM); ♀ holotype of P. hortusensis: FRANCE: Hérault, Hortus, H. Abdul-Nour leg. (MNHN); ♀ holotype of E. afer: ZIMBABWE: Harare (= Salisbury), A. Watsham leg. (NHMUK). Paratypes of P. hortusensis: same locality label as holotype, 1 ♀ (MNHN); Héralt, Montarnaud, H. Abdul-Nour leg., 1 ♀ (MNHN); paratypes of E. afer: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele (= Haut-Zaire), Garamba National Park, 3 ♀♀ (1 in AMNH, 2 in MRAC); SOMALIA: Benadir, Afgoi, M. Olmi leg., 2 ♀♀ (AMNH); SOUTH AFRICA: East- ern Cape, Katberg, 4000 ft., XII. 1932, R. E. Turner leg., 2 ♀♀ (AMNH, NHMUK); Eastern Cape, Port St. Johns, 1 – 11. VI. 1923, R. E. Turner leg., 1 ♀ (NHMUK). Other material: BOTSWANA: Serowe, Farmer’s Brigade, 1 ♀ (USNM); Kgatleng District, Odi, 17 km NE of Gaborone, 1 ♀ (ZIL). CAMEROON: South, near Ebolowa, Nko- emvon, 21. I – 6. II. 1980, cocoa plantation, D. Jackson leg., 1 ♀ (NHMUK). CAPE VERDE ISLANDS: Santiago Is- land, São Jorge dos Orgãos, A. Van Harten leg., 1 ♀ (AMNH). CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 7. V. 2001, sweep, lowland rainforest, marsh clearing, S. van Noort leg., 19 ♂♂ (18 in SAMC, 1 in MOLC). IVORY COAST: Bouaké, 1 ♀ (CNC); 3 km S of Katiola, 1 ♀ (DEUW). KENYA: Coast Prov., Kasigau Mtn., 3.82667 ° S 38.64982 ° E, 1117 m, 16 – 30. VI. 2011, indigenous forest, MT, next to spring in forest, R. Copeland leg., 1 ♀ (NMK) (new for Kenya). MADAGASCAR: Toliara, Mite Forest, 20.7 km 29 ° WNW Tongobory, 23 ° 31.27 ’ S 44 ° 07.17 ’ E, 75 m, 27. II – 3. III. 2002, MT, in gallery forest, Fisher, Griswold et al. leg., BLF 5849, 1 ♂ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 26. I – 5. II. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 14, 1 ♀ (CAS); Toliara, Lake Ranobe, 23 ° 02.22 ’ S 43 ° 36.37 ’ E, 30 m, 25 – 28. IV. 2003, MT, spiny forest thicket, MGF 063, Frontier Wilderness Project, 1 ♀, 1 ♂ (CAS); same locality label, 25 – 29. I. 2003, MGF 053, 2 ♂♂ (CAS); Toliara, Lake Ranobe, 23 ° 02.26 ’ S 43 ° 36.35 ’ E, 30 m, 25 – 30. IV. 2003, MT, spiny forest thicket, MGF 065, Frontier Wilderness Project, 1 ♂ (CAS). MOZAMBIQUE: Maputo Prov., Ma- puto, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 26 km S of Maputo along Maputo-Namaacha Road, Umbelúzi, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 3 km N of Umbelúzi along Maputo-Namaacha Road, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., Ponta de Ouro, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 6 km N of Palmeira along Maputo-Xai Xai road, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., Manhiça District, Manhiça, Vasco’s Farm, 1 – 8. IX. 1996, MT, M. Olmi leg., 1 ♀ (MOLC); same locality label, 24. X – 6. XII. 1996, 2 ♀♀ (MOLC); same local- ity label, 9. IX – 24. X. 1995, 4 ♀♀ (MOLC); same locality label, 1 – 8. IX. 1996, 1 ♂ (MOLC); Niassa Prov., Cuamba, Mituque, 22. VII – 5. VIII. 2008, MT, M. Olmi leg., 1 ♀ (MOLC); same locality label, 12. IV – 6. V. 2008, 1 ♀ (MOLC); same locality label, 12. IV – 6. V. 2008, MT, M. Olmi leg., 2 ♂♂ (MOLC). NIGERIA: Oyo State, Ibadan, IITA, B. R. Critchley leg., 1 ♀ (NHMUK). OMAN: Salalah, South Dahariz, 1 ♀ (PMA); 24 km E of Salalah, 23. II. 1986, 1 ♀ (CNC). SOUTH AFRICA: Eastern Cape, Katberg, 4000 ft., XII. 1932, R. E. Turner leg., 1 ♀ (AMNH); Eastern Cape, 8 km N of Fort Beaufort, Blinkwater, 32 ° 43 ’ S 26 ° 35 ’ E, 1 ♀ (ZIL); Eastern Cape, Somerset East, IX. 1930, R. E. Turner leg., 1 ♂ (NHMUK); Eastern Cape, Port St. Johns, 25 - 31. III. 1923, 5 – 30. IV. 1923, 1 – 14. V. 1923, 1 – 11. VI. 1923, R. E. Turner leg., 8 ♂♂ (NHMUK); KwaZulu-Natal: Louwsberg, Sanyati Farm, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 1 – 24. III. 2006, MT, M. Mostovski leg., 1 ♀ (NMSA); Mpumalanga, Krüger National Park, Lower Sabie, 14. XII. 1985, 1 ♀ (CNC); Northern Cape, Sterboom Farm, 32 ° 36.794 ’ S 20 ° 44.660 ’ E, 1599 m, 9. II – 31. III. 2010, MT, Succulent Karoo, S. van Noort leg., 1 ♀ (SAMC); North West, Brits district, Magaliesberg, Castle Gorge, 25 ° 48 ’ S 27 ° 34 ’ E, 9. XI. 2003, sweep river course grass & forbs, regrowth after fire, M. Stiller leg., 1 ♀ (SANC); Western Cape, Cape Peninsula National Park, 34 ° 15 ’ S 18 ° 21.5 ’ E, 100 m, 14 – 31. XII. 1999, fynbos, FIT, S. & J. Peck leg., 99 – 141, 3 ♀♀ (CNC); Western Cape, Ceres, III. 1925, R. E. Turner leg., 1 ♂ (NHMUK); Western Cape, West Coast Fossil Park, 4 km 270 ° W Langebaanweg, 32 ° 58.444 ’ S 18 ° 06.731 ’ E, 9 – 16. X. 2002, MT, Sand plain fynbos dominated by Lebeckia, S. van Noort leg., 1 ♂ (SAMC). UNITED ARAB EMIRATES: Dubai, Nakhalai, 1 ♀ (CDAE); Wadi Maidaq, 25 ° 18 ’ N 56 ° 07 ’ E, 29. III – 10. IV. 2006, yellow and white water-traps, A. van Harten leg., 1 ♀ (MOLC); al-Ajban, 24 ° 36 ’ N 55 ° 01 ’ E, 2 – 9. IV. 2006, MT, A. van Harten leg., 2 ♀♀ (MOLC); same locality label, 26. II – 2. IV. 2006, MT, A. van Harten leg., 6 ♀♀ (MOLC); same locality label, 2 – 9. IV. 2006, MT, A. van Harten leg., 3 ♀♀ (MOLC); same locality label, 9. IV – 2. V. 2006, MT, A. van Harten leg., 1 ♂ (MOLC). YEMEN: Wadi Bana, 13 ° 21 ’ N 45 ° 18 ’ E, 100 m, 20. III. 1993, A. van Harten leg., 1 ♀ (MOLC); Al-Kadan, 15 ° 16 ’ N 43 ° 12 ’ E, 200 m, 3. XII. 1997 – 17. II. 1998, MT, A. van Harten & H. M. Naser leg., 1 ♀ (MOLC); Near Sana’a, 15 ° 21 ’ N 44 ° 13 ’ E, 2300 m, 9. IV. 1998, A. van Harten leg., 1 ♀ (MOLC); Lahj, 13 ° 04 ’ N 44 ° 53 ’ E, 150 m, X. 1998, IV. 2000, V – VI. 2000, I. 2001, III. 2001, 1. X – 7. XII. 2001, 17. XII. 2001 – 31. I. 2002, III – V. 2002, MT, A. van Harten & A. Sallam leg., 6 ♀♀, 3 ♂♂ (MOLC); Al-Kowd, 13 ° 05 ’ N 45 ° 22 ’ E, 20 m, 01 – 15. I. 1993, A. van Harten leg., 1 ♀ (MOLC); Al Lahima, 15 ° 24 ’ N 43 ° 32 ’ E, 1200 m, 16. X – 31. XII. 2000, A. van Harten & A. M. Hager leg., 1 ♀ (MOLC); Ja’ar, 13 ° 13 ’ N 45 ° 18 ’ E, 50 m, 15. I. 2001, A. van Harten leg., 1 ♀ (MOLC). ZAMBIA: Lusaka, 1 ♀ (DEUCD). ZIMBABWE: Harare (= Salisbury), A. Watsham leg., 2 ♀♀ (NHMUK); Harare, Chishawasha, IX. 1979, A. Watsham leg., 1 ♀ (NHMUK). Palaearctic. Known from many Palaearctic countries (Algeria, Azerbaijan, Egypt, France, Germany, Hungary, Israel, Italy, Lebanon, Morocco, Spain, Sweden, Turkey, Ukraine).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE372BD4FF3ED527FD0EFE0C.taxon	biology_ecology	Hosts. Delphacidae Delphacinae (Guglielmino et al. 2013): in the Palaearctic region: in France: Ribautodelphax imitans (Ribaut); in Germany: Ribautodelphax albostriata (Fieber); in Italy: Falcotoya minuscula (Horváth), Ribautodelphax imitans (Ribaut), Laodelphax striatella (Fallén), Megadelphax sordidula (Stål). In the Afrotropical region: in the Cape Verde Islands: Peregrinus maidis (Ashmead); in Mozambique: Nycheuma endymion (Fennah), Metadelphax propinqua (Fieber). Natural enemies. Hymenoptera Encyrtidae: in Mozambique: Cheiloneurus gonatopodis R. Perkins. This Encyrtid is one of the most common natural enemies of Dryinids in Mozambique (see also Gonatopus amoenus Olmi, Gonatopus mossambicus Olmi and Gonatopus incognitus Olmi). The reared specimens of C. gonatopodis have the following locality labels: Mozambique, Maputo Prov., 6 km N Palmeira, parasitized planthopper col- lected on 3. IX. 1995; cocoon spun on 7. IX. 1995; encyrtids emergence on 25. IX. 1995, 3 ♂♂, 1 ♀ (MOLC); Mozam- bique, Maputo Prov., 3 km N Umbelúzi, parasitized planthopper collected on 11. XI. 1992; encyrtids emergence on 26. XI. 1992, 1 ♂, 2 ♀♀ (MOLC); Mozambique, Maputo Prov., 3 km N Umbelúzi, parasitized planthopper collected on 11. XI. 1992; encyrtids emergence on 26. XI. 1992, 1 ♂, 2 ♀♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE372BD4FF3ED527FD0EFE0C.taxon	distribution	Distribution. Botswana, Cameroon, Cape Verde Islands, Central African Republic, Democratic Republic of the Congo, Ivory Coast, Kenya, Madagascar, Mozambique, Nigeria, Oman, Somalia, South Africa, Yemen, Zambia, Zimbabwe, in addition to many Palaearctic countries.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE332BD5FF3ED793FE55F96C.taxon	diagnosis	Diagnosis. ♀: apterous (Fig. 167 A) or fully winged (Fig. 247 A); palpal formula 3 / 2, 4 / 2, 4 / 3, 5 / 2, 5 / 3, or 6 / 3; antenna with (Fig. 6 A) or without ADOs; pronotum crossed (Fig. 171 E) or not (Fig. 198 F) by transverse furrow; enlarged claw with apex pointed and with one large (Fig. 168 F) or small (Fig. 198 G) subapical tooth (occasionally subapical tooth absent, then enlarged claw with distal group of lamellae); in fully winged forms, protarsomere 5 with more than 20 lamellae (Fig. 244 E); tibial spurs 1 / 0 / 1. ♂: fully winged (Figs 167 B, C); occipital carina absent (Fig. 6 C) or incomplete (in this last case, only present behind and shortly on sides of lateral ocelli); occiput concave (Fig. 6 C); temple present; palpal formula 3 / 2, 4 / 2, 4 / 3, 5 / 2, 5 / 3, or 6 / 3; tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE332BD5FF3ED793FE55F96C.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE332BD5FF3ED793FE55F96C.taxon	biology_ecology	Hosts. Acanaloniidae, Caliscelidae, Cicadellidae (except Typhlocybinae, Idiocerinae, Macropsinae), Delphacidae, Flatidae, Issidae, Lophopidae, Tropiduchidae (Guglielmino & Olmi 1997, 2006, 2007; Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE332BD5FF3ED793FE55F96C.taxon	discussion	World species. 442 species are known, 133 in the Afrotropical region, including two species incertae sedis.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE332BD5FF3ED793FE55F96C.taxon	discussion	Remarks. Larvae of Gonatopus were described by Ponomarenko (1975 b), Carcupino et al. (1996, 1998), Guglielmino & Virla (1998), Virla & Mangione (2000), Guglielmino (2002), Mangione & Virla (2004) and Guglielmino et al. (2006, 2007, 2008).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0F2BEBFF3ED6D7FA72FC68.taxon	description	(Figs 167 A – C, 168 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0F2BEBFF3ED6D7FA72FC68.taxon	description	Description. ♀ (Fig. 167 A). Apterous; body length 2.0 – 4.0 mm. Head black or brown, except mandible, clypeus and anterior region of frons yellow or testaceous; antenna black or brown, except antennomeres 1 – 3 yellow or testaceous; occasionally antenna testaceous-brown; mesosoma, petiole and metasoma usually black, except posterior apex of propodeal declivity testaceous; occasionally mesosoma and metasoma brown-testaceous or testaceous-reddish, except petiole black; occasionally mesosoma brown, except mesoscutum yellow; legs yellow or testaceous, except coxae and clubs of femora almost completely black, or darkened, or brown-reddish. ♀♀ with testaceousreddish mesosoma observed only in the USA (Texas: Lost Maples State Park; Inks Lake State Park). Antenna clavate; antennomeres of holotype of P. nearcticus in following proportions: 4: 4: 9: 5.5: 5: 5: 4: 4: 4: 7; antennomeres of holotype of A. sinensis in following proportions: 5: 3: 7: 4: 4: 4: 4: 4: 3.5: 5. Head flat or slightly convex, dull, slightly or strongly granulate; frontal line complete; occipital carina absent; temple distinct; POL = 3; OL = 2; OOL = 5. Palpal formula 5 / 2, or 4 / 2, or 3 / 2, or 2 / 2. Pronotum crossed by strong transverse impression, shiny, slightly or strongly granulate. Mesoscutum dull, granulate. Metanotum granulate and transversely striate, about coplanar to mesoscutellum. Metapectal-propodeal disc dull, granulate; propodeal declivity usually transversely striate, occasionally almost completely granulate and not transversely striate. Meso-metapleural suture obsolete. Mesopleuron and metapleuron granulate, with or without transverse striae. Protarsomere 1 about twice as long as 4 (14: 7; or 15: 7; or 12: 7). Protarsomeres of holotype of A. sinensis in following proportions: 15: 2: 3: 7: 13. Enlarged claw (Fig. 168 A) with one large subapical tooth and one row of 3 – 6 peg-like bristles. Protarsomere 5 (Fig. 168 A) with two rows of 6 – 22 lamellae; apex with approximately 6 – 9 lamellae. Tibial spurs 1 / 0 / 1. ♂ (Figs 167 B, C). Fully winged; body length 1.6 – 2.8 mm. Head black or brown, except mandible testaceous or yellow; antenna brown or black; mesosoma and metasoma black or brown; legs banded, brown or black, except trochanters, stalks of femora, extremities of tibiae and tarsi yellow. Antenna setose, filiform; antennomeres of ♂ paratype of A. bouceki in following proportions: 5: 4: 8: 7: 7: 7: 6: 6: 6: 8; antennomere 3 four or more than 4 × as long as broad. Antennomeres of specimen from P. R. China, Ruyang, in following proportions: 4: 4: 7: 7: 6.5: 6.5: 5.5: 5: 5: 7. Head shiny, alutaceous, slightly punctate; vertex laterally without two shiny and smooth areas situated between lateral ocelli and eyes; frontal line incomplete, only present between antennal toruli; occipital carina absent; occiput concave; temple distinct; POL = 6; OL = 4; OOL = 2 (or POL = 6; OL = 2; OOL = 2; or POL = 7; OL = 3; OOL = 1.5); greatest breadth of lateral ocelli about twice as long as OOL. Palpal formula 3 / 2, or 4 / 2. Mesoscutum dull, setose, slightly granulate. Notauli incomplete, reaching approximately 0.4 – 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc and propodeal declivity completely shiny, unsculptured; metapectal-propodeal disc with or without median longitudinal furrow. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part much longer than proximal part. Dorsal process of paramere (Fig. 168 B) more or less long and slender, with distal third parallel to penis. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0F2BEBFF3ED6D7FA72FC68.taxon	materials_examined	Material examined. Types: ♀ holotype of P. nearcticus: USA: Iowa, Woodbury Co., Sioux City, IX. 1919, reared from a jassid, C. N. Ainslie leg. (USNM); ♀ holotype of E. americanus: USA: Iowa, Story Co., Ames, 25. VII. 1919, E. D. Ball leg. (USNM); ♀ holotype of P. ugandanus: UGANDA: Bululo, 1909, E. Bayon leg. (MCSNG); ♀ holotype of A. bouceki: SPAIN: Malaga, Nerja, 3. VII. 1974, Z. Boucek leg. (NHMUK); ♀ holotype of A. sinensis: P. R. CHI-NA: Macau (BPBM). Paratypes of E. americanus: USA: same locality label as holotype, 4 ♀♀ (USNM). Paratypes of A. bouceki: SPAIN: Malaga, Nerja, 23. VI. 1973, Z. Boucek leg., 13 ♀♀ (9 in NHMUK, 4 in DISAFA); Granada, La Herradura, 24. VI. 1973, Z. Boucek leg., 1 ♀, 1 ♂ (NHMUK); Granada, Calahonda, 22. VI. 1973, Z. Boucek leg., 2 ♀♀ (NHMUK); Castellón, Benicasim, 13 – 15. VI. 1973, Z. Boucek leg., 1 ♀ (NHMUK); same locality label, 22 – 24. VI. 1974, 1 ♂ (NHMUK); Granada, Calahonda, 22. VI. 1973, Z. Boucek leg., 2 ♀♀ (NHMUK); Barcelona, Calella de la Costa, VI. 1974, Z. Boucek leg., 1 ♀ (NHMUK); Paratypes of A. sinensis: CHINA: Macau, 3 ♀♀ (1 in BPBM, 2 in AMNH); Chin San, 1 ♀ (BPBM). VIET NAM: Lam Dong Prov., Di Linh, Plateau of Djiring, 1 ♀ (AMNH). Other material: BENIN: Gomé, 12. XI. 1993, 1 ♀ (CIRAD). BOTSWANA: SE of Gaborone District, at roadsides in town, 9. I. 1982, 1 ♀ (ZIL). ETHIOPIA: Kefa Province, Jima, 1 ♀ (NHMUK). KENYA: Central Prov., Nairobi, 1 ♀ (MRSNT); Central Prov., Olgasalic, 1 ♀ (MRAC); Central Prov., Mount Kenya Safari Club, 1 ♀ (USUC); Rift Valley Prov., Subukia, 00 ° 0.429 ’ N 36 ° 14.523 ’ E, 24. III – 11. IV. 2010, Dupopnt & Braet leg., 1 ♀ (FSAE). MADA- GASCAR: Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20 ° 47.56 ’ S 47 ° 10.54 ’ E, 825 m, 11 – 20. II. 2007, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 29 - 78, 1 ♀ (CAS). MOZAMBIQUE: Nampula Prov., Nampula, approximately 500 m from Nampula Catholic Seminary, 24. XI. 1999, M. Olmi leg., 1 ♀ (MOLC); Ni- assa, Cuamba, Catholic University of Mozambique, Faculty of Agriculture Campus, 26. V. 2007, M. Olmi leg., 2 ♀♀ (MOLC). NAMIBIA: 134 km E Swakopmund, along Rd. B 2, 4. III. 2006, M. Olmi reared ex Paradorydium spatulatum (Naudé), 2 ♀♀, 1 ♂ (MOLC). SOUTH AFRICA: Eastern Cape, Port St Johns, 1 – 14. V. 1923, 15 – 3. V. 1923, 16 – 28. IV. 1924, R. E. Turner leg., 9 ♀♀ (7 in NHMUK, 2 in AMNH); Free State, Harrismith, II. 1927, R. E. Turner leg., 1 ♀ (NHMUK); Free State, Sasolburg, Wonderwater Colliery, 26 ° 46.11 ’ S 27 ° 46.59 ’ E, 1450 m, 12. V. 1999, D- Vac, IS- 08 b, M. Stiller leg., 1 ♀ (SANC); Gauteng, Tierpoort, 1 ♀ (NCIP); Gauteng, Pretoria Dist., Soutpan, 25 ° 24 ’ S 28 ° 06 ’ E, 1 ♀ (SANC); Gauteng, Pretoria, EKH, V. 1946, A. R. I. Pretoria, SAM-HYM-PO 21006, 1 ♀ (SAMC); Kwa- Zulu-Natal, Richards Bay, 28 ° 46 ’ S 32 ° 04 ’ E, 1 ♀ (ZIL); KwaZulu-Natal, Gingindhlovu, 9. VI. 1926, R. E. Turner leg., 1 ♀ (NHMUK, AMNH); KwaZulu-Natal, Empangeni, 26. IV. 1926, R. E. Turner leg., 1 ♀ (NHMUK); KwaZulu-Natal, Kloof, VIII. 1926, IX. 1926, R. E. Turner leg., 10 ♀♀ (NHMUK); same locality label, 1 ♀ (DASB); KwaZulu-Natal, Drakensberg, Van Reenen, 23 – 26. I. 1927, R. E. Turner leg., 2 ♀♀ (AMNH, NHMUK); KwaZulu-Natal, Champagne Castle, Hotel vicinity, 29 ° 04 ’ S 29 ° 25 ’ E, 16 – 20. X. 2000, swept off grass (Oplismenus sp., Poaceae) and herb. shrubs under tree canopy, M. Stiller leg., 2 ♀♀ (SANC); KwaZulu-Natal, Northern Drakensberg, 4 km S of Cavern Berg Re- sort, along the road from R 74 to Cavern Berg Resort, 9. IV. 2007, M. Olmi leg., 2 ♀♀ (MOLC, SAMC); same locality label, C. 9. IV. 2007, B. 10. IV. 2007, M. Olmi reared from Balclutha frontalis (Ferrari), 1 ♂ (MOLC); KwaZulu-Natal, Winterton, along Rd 600, 5 km S of junction with Rd to Loskop, C. 10. IV. 2007, B. 12. IV. 2007, M. Olmi reared from Balclutha frontalis (Ferrari), 1 ♂ (MOLC); KwaZulu-Natal, Winterton, along Rd 600, grass in front of the gate of Cathrin Farm, 10. IV. 2007, M. Olmi leg., 2 ♀♀ (MOLC); same locality label, C. 10. IV. 2007, B. 11. IV. 2007, M. Olmi reared from Balclutha frontalis (Ferrari), 1 ♀ (MOLC); same locality label, C. 10. IV. 2007, B. 12. IV. 2007, M. Olmi reared from Balclutha frontalis (Ferrari), 1 ♀ (SAMC), 1 ♂ (MOLC); KwaZulu-Natal, Drakensberg, Cathedral Peak, inside the park, grass near the gate of Cathedral Peak Hotel, C. 11. IV. 2007, B. 12. IV. 2004, M. Olmi reared from Balclutha rosea (Scott), 1 ♂ (MOLC); KwaZulu-Natal, Drakensberg, Cathedral Peak, inside the park, grass near the gate of Cathedral Peak Hotel, C. 11. IV. 2007, B. 12. IV. 2007, M. Olmi reared from Maiestas angustisecta (Linnavuori), 1 ♂ (MOLC); Western Cape, Koornplanskloof, 10 km S of Citrusdal, 32 ° 40 ’ S 19 ° 01 ’ E, 200 – 270 m, 1 ♀ (ZIL). SWAZILAND: 2 km N of Loyengo, 26 ° 33 ’ S 31 ° 11 ’ E, 1 ♀ (ZIL). UGANDA: Kawanda, V. 1943, VI. 1943, T. H. C. Taylor leg., 3 ♀♀, 3 ♂♂ (NHMUK); same locality label, 1 ♀, 1 ♂ (AMNH); Serere, VII. 1943, T. H. C. Taylor leg., 1 ♀ (NHMUK). ZIMBABWE: Harare (= Salisbury), A. Watsham leg., 2 ♀♀, 1 ♂ (NHMUK); Harare, Chisha- washa, V. 1980, A. Watsham leg., 1 ♀ (NHMUK). Palaearctic: G. nearcticus is known from the following countries: Bulgaria (new record: Oblast Shumen, Bajkovo, 16. VIII. 2016, A. Guglielmino and C. Bückle leg., 1 ♀ (MOLC )), Croatia, Cyprus, France, Greece, Italy, Montenegro, Portugal, Spain (including Canary Islands) and Turkey (Olmi 1999). Oriental region: G. nearcticus is known from the following countries: India, P. R. China, Singapore, Thailand, Viet Nam (Xu et al. 2013). Nearctic region: G. nearcticus is known from the USA (Olmi 1984, 1999).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0F2BEBFF3ED6D7FA72FC68.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in the Palaearctic region: in France: Balclutha frontalis (Ferrari); in Italy (Pantelleria Island): Balclutha frontalis (Ferrari). In the Afrotropical region: in Namibia: Paradorydium spatulatum (Naudé); in South Africa: Balclutha frontalis (Ferrari) (= Balclutha rosea (Scott )), Maiestas angustisecta (Linnavuori). In the Nearctic region: in the USA: Balclutha neglecta (DeLong & Davidson); Balclutha impicta (Van Duzee).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0F2BEBFF3ED6D7FA72FC68.taxon	distribution	Distribution. Benin, Botswana, Ethiopia, Kenya, Madagascar, Mozambique, Namibia, South Africa, Swaziland, Uganda, Zimbabwe, in addition to many countries of the Oriental, Western Palaearctic and Nearctic regions.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0C2BECFF3ED2BBFE4BFC6E.taxon	discussion	Remarks. The ♀ mesosoma of Gonatopus festivus Olmi can vary in colour, from totally testaceous to partly black or brown. If the colour is partly black or brown, ♀♀ of G. festivus can be confused with G. harteni Olmi. In this case, ♀♀ of these two species are indistinguishable. However, the two species are distinct, because their ♂♂ show strong differences.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0B2BEDFF3ED275FCA8FAFC.taxon	description	(Figs 168 C, D, 171 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0B2BEDFF3ED275FCA8FAFC.taxon	description	Description. ♀. Apterous; body length 1.9 – 3.5 mm. Head black or brown, except clypeus, mandible and occasion- ally anterior region of frons yellow; antenna brown, except scape yellow and flagellomere 8 whitish; mesosoma black or brown-reddish, with posterior extremity of propodeal declivity testaceous; occasionally posterior half of mesoscutum yellow; petiole black; metasoma black or brown-black; legs yellow, except coxae, clubs of femora and tibiae brownish. Head shiny, excavated (Fig. 168 C), granulate; POL = 3; OL = 10; OOL = 20. Palpal formula 4 / 2. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum shiny, slightly granulate, without lateral pointed apophyses. Metanotum weakly excavated behind mesoscutellum, transversely striate, with sides protruding; lateral protrusions of metanotum pointed. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate; top of metapectal-propodeal disc with track of median longitudinal furrow. Meso-metapleural suture distinct and complete. Mesopleuron and metapleuron transversely striate. Protarsomeres in following proportions: 13: 2: 4: 13: 20. Enlarged claw (Fig. 168 D) with one large subapical tooth and one row of 6 – 11 lamellae. Protarsomere 5 (Fig. 168 D) with two rows of 10 – 21 lamellae (occasionally shortest row only situated in proximal half of protarsomere 5); apex with 8 – 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.4 – 1.7 mm. Head black, except mandible and part of clypeus testaceous; an- tenna brown; mesosoma and metasoma black; legs brown, except trochanters, articulations, part of profemora, part of meso- and metatarsus testaceous. Antenna setose, filiform; antennomeres in following proportions: 4: 4: 5: 4.5: 4.5: 5: 4.5: 4.5: 4: 6; antennomere 3 less than 3 × as long as broad: 5: 2. Head shiny, granulate; frontal line absent; occipital carina absent; POL = 5; OL = 2; OOL = 3; vertex with two oval smooth and shiny areas situated on sides of lateral ocelli and occupying all areas between lateral ocelli and eyes (oval areas anteriorly surrounded by high carina). Palpal formula 4 / 2. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.6 – 0.7 × length of mesoscutum. Mesoscutellum dull or shiny, more or less strongly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc shiny, unsculptured, with strong longitudinal median furrow; propodeal declivity shiny, with irregular keels or striae, unsculptured among striae and keels. Forewing hyaline, without dark transverse bands, with only costal cell surrounded by well pigmented veins (other basal cells surrounded by obsolete veins); 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part much longer than proximal part (approximately 14: 4). Dorsal process of paramere (Fig. 171 A) very long and slender, with apex very pointed. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0B2BEDFF3ED275FCA8FAFC.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, XII. 1934, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, 15 – 28. III. 1922, 1 ♀ (NHMUK); same locality label as holotype, 1 – 13. III. 1922, 1 ♀ (NHMUK); same locality label as holotype, 30. XII. 1938, 1 ♀ (NHMUK); same local- ity label as holotype, 30. XI. 1938, 1 ♀ (NHMUK); same locality label as holotype, 1. I. 1939, 1 ♀ (NHMUK); same locality label as holotype, 2 ♀♀ (AMNH). Other material: BENIN: Dassa, 12. XI. 1993, 1 ♀ (CIRAD). GABON: Woleu-Mntem Prov., 31.3 km ESE Minvoul, 02 ° 04.8 ’ N 12 ° 24.4 ’ E, 600 m, 7. II. 1998, 1648, sifted litter (leaf mold, rotted wood), rainforest, Brian L. Fisher leg., 1 ♀ (CAS). MOZAMBIQUE: Niassa, Cuamba, Camboja, reared from Caliscelis sp., C. 15. XI. 1999; B. 18. XI. 1999; Sf. 12. XII. 1999, M. Olmi reared, 1 ♀, 2 ♂♂ (MOLC); Niassa, Cuamba, Catholic University Campus, C. 17. III. 2012, B. 19. III. 2012, Sf. 1. IV. 2012, M. Olmi leg., 1 ♂ (MOLC); same locality label, C. 17. III. 2012, B. 20. III. 2012, Sf. 9. IV. 2012, M. Olmi leg., 1 ♂ (MOLC). SOUTH AFRICA: KwaZulu-Natal, 40 km S of Harrismith, Van Reenen, C. 26. I. 1997, M. Olmi leg., 1 ♀ (AMNH); KwaZulu-Natal, Port Edward, along road from R 61 to Umtamvuna Nature Reserve, C. 13. IV. 2007, B. 15. IV. 2007, M. Olmi reared from unidentified adult of Delphacid, 2 ♀♀, 2 ♂♂ (MOLC); same locality label, C. 13. IV. 2007, B. 16. IV. 2007, 3 ♀♀ (MOLC); same locality label, C. 13. IV. 2007, B. 17. IV. 2007, 1 ♀ (MOLC); same locality label, C. 14. IV. 2007, 1 ♀ (MOLC). Lim- popo, 15 km NW of Klaserie, Guernsey Farm, 1 ♀ (PMA); Western Cape, near Citrusdal, The Baths, reared from Toya tuberculosa (Distant), C. 22. I. 2003, B. 24. I. 2003, Sf. 10. II. 2003, M. Olmi reared, 1 ♀, 1 ♂ (MOLC); same local- ity label, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0B2BEDFF3ED275FCA8FAFC.taxon	biology_ecology	Hosts. Caliscelidae and Delphacidae (Guglielmino et al. 2013): in Mozambique: Caliscelidae: Caliscelis sp.; in South Africa: Delphacidae: Matutinus vitticollis (Stål); Sogatella kolophon (Kirkaldy); Toya ceresensis (Muir) and T. tuberculosa (Distant); Biology. In 1999 in Cuamba (Mozambique) this species was reared from an unknown species of Caliscelis (Caliscelidae), a parasitized specimen with five larval thylacia. Only four larvae were able to spin their cocoons and only three adults emerged, one ♀ and two ♂♂. In 2003 at The Baths (South Africa) this species was reared from Toya tuberculosa (Distant) (Delphacidae). The parasitized specimen was a ♀ with two thylacia, from which two larvae emerged; the two larvae pupated and from the cocoons one ♀ and one ♂ emerged. These are the only two cases of rearing of this species before 2007. This species is probably specialized in superparasitism, because in 2007 in Port Edward, South Africa, many delphacid adults with two thylacia were reared and each time one ♀ and one ♂ emerged from the same host; ♀ and ♂ specimens were very small. In one case in South Africa, Port Edward, a single ♂ was reared from a nymph of an unidentified delphacid having only one thylacium.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0B2BEDFF3ED275FCA8FAFC.taxon	distribution	Distribution. Benin, Gabon, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0A2BEFFF3ED383FC4BFDB9.taxon	description	(Figs 168 E, 171 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0A2BEFFF3ED383FC4BFDB9.taxon	description	Description. ♀. Apterous; body length 2.4 – 4.0 mm. Head brown, except mandible, clypeus and anterior region of frons testaceous; antenna brown, except scape yellow; propleuron brown; pronotum testaceous-reddish or brown; mesoscutum yellow, except anterior margin darkened; rest of mesosoma testaceous-reddish or brown; metasoma brown; legs testaceous, except tibiae, clubs of femora and occasionally coxae more or less darkened. Antenna clavate; antennomeres in following proportions: 6: 5: 13: 7: 6: 6: 5.5: 5: 4.5: 7. Head excavated, shiny, smooth, alutaceous or unsculptured; frontal line complete; occipital carina absent; temple distinct; POL = 2.5; OL = 2.5; OOL = 7. Palpal formula 4 / 2. Pronotum shiny, unsculptured or slightly granulate, crossed by strong transverse impression; disc of pronotum without posterior contraction near posterior margin. Mesoscutum shiny or dull, unsculptured or granulate, without lateral pointed apophyses. Metanotum flat, granulate (in large specimens) or unsculptured (in small specimens), about coplanar to mesoscutellum; sides of metanotum rounded. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity granulate and transversely striate; anterior surface unsculptured; top with track of median longitudinal furrow. Mesopleuron and metapleuron dull, granulate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 15: 3: 5: 12: 19. Enlarged claw (Fig. 168 E) with one large subapical tooth and one row of four lamellae. Protarsomere 5 (Fig. 168 E) with two rows of approximately 20 lamellae; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.0 – 2.4 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs yellow, except meso- and metatibia and clubs of femora slightly darkened. Antenna filiform; antennomeres in following proportions: 4: 4: 8: 6: 6: 6: 5.5: 6: 5: 7; antennomere 3 more than 4 × as long as broad: 8: 1.5. Head shiny, unsculptured or very slightly granulate; frontal line absent; occipital carina absent; POL = 6; OL = 2.5; OOL = 2.5; temple distinct; vertex with track of two oval areas situated between lateral ocelli and eyes (areas anteriorly not surrounded by high carina). Palpal formula 3 / 2 or 4 / 2. Mesoscutum dull, alutaceous; notauli complete, usually posteriorly joint or situated very close; occasionally (in two specimens from South Africa, Cape Recife Natural Reserve, and Mozambique, Cuamba, Campus of UCM) notauli posteriorly distinctly separated. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull and completely reticulate rugose; in specimen from Mozambique, 6 km N of Palmeira, metapectal-propodeal disc with two smooth areas on lateral regions. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part (14: 6). Dorsal process of paramere (Figs 171 B, C) long and with broad and more or less rounded distal region. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0A2BEFFF3ED383FC4BFDB9.taxon	materials_examined	Material examined. Types: ♀ holotype: MOZAMBIQUE: Maputo Prov., 26 km S of Maputo, Umbelúzi, in the farm of the Instituto de Investigaciones Agronomicas (INIA), 20. X. 1992, M. Olmi reared (AMNH). Paratypes: BURKINA FASO: N Bobo, Mare-aux-Hippopotames, 3. X. 1979, A. Pauly leg., 1 ♀ (AMNH). MOZAMBIQUE: same locality label as holotype, X. 1992, Xi. 1992, 12 ♀♀, 13 ♂♂ (AMNH); Maputo Prov., 3 km N of Umbelúzi (23 km S of Maputo), in the fields of a private farm, C. 11. XI. 1992, Sf. 26. XI. 1992, M. Olmi reared, 1 ♂ (AMNH); same locality label, 28. XI. 1992, 1 ♂ (AMNH); Maputo Prov., Maputo, in the fields of the Municipal Nursery in Costa do Sol, 25. IX. 1992, M. Olmi reared, 1 ♀ (AMNH); same locality label, C. 31. X. 1992, Sf. 15. XI. 1992, 1 ♂ (AMNH). Other material: GAMBIA: Upper River, Basse (= Basse Santa Su), 1 ♀ (KJHC). MOZAMBIQUE: Maputo Prov., 2 km S of Umbelúzi, in SEMOC farm, C. 4. XI. 1995, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 6 km N of Palmeira along Road EN 1, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 5 km E Manhiça, C. 25. IX. 1996, M. Olmi leg., 1 ♀ (AMNH); Gaza Prov., Between Macia and Palmeira, 2 km E of River Incomati Bridge along Road EN 1, M. Olmi leg., 1 ♀ (AMNH); Niassa Prov., Cuamba, Mozambique Catholic University Campus, C. 4. IV. 003, B. 7. IV. 2003, Sf. 24. IV. 2003, M. Olmi reared, ex adult ♀ of an unknown species of Delphacidae, 1 ♂ (MOLC); same locality label, C. 7. IV. 003, B. 11. IV. 2003, Sf. 25. IV. 2003, 1 ♂ (MOLC); same locality label, C. 7. IV. 003, B. 10. IV. 2003, Sf. 25. IV. 2003, 1 ♂ (MOLC); same locality label, C. 6. III. 2005, B. 8. III. 2005, Sf. 22. III. 2005, 1 ♂ (MOLC); same locality label, C. 17. III. 2012, B. 19. III. 2012, Sf. 1. IV. 2012, M. Olmi leg., 2 ♀♀ (MOLC); same locality label, C. 17. III. 2012, B. 19. III. 2012, Sf. 2. IV. 2012, M. Olmi leg., 2 ♀♀, 2 ♂♂ (MOLC); same locality label, C. 17. III. 2012, B. 18. III. 2012, Sf. 3. IV. 2012, M. Olmi leg., 1 ♀ (MOLC); Niassa Prov., Cuamba, Farm of the Catholic University of Mozambique, C. 18. II. 2005, B. 20. II. 2005, Sf. 8. III. 2005, M. Olmi reared from an adult specimen of Meenoplidae, M. Olmi reared, 1 ♂ (MOLC). SOUTH AFRICA: Eastern Cape, near Port Elizabeth, Cape Recife Nature Reserve, collected and reared in front of the gate of the reserve, C. 30. I. 1997, M. Olmi leg., 1 ♀ (AMNH); Eastern Cape, 16 km E of Port Elizabeth, S. George’s Strand, C. 31. I. 1997, M. Olmi reared, 1 ♀ (AMNH); Eastern Cape, Grahamstown, XII. 1956, C. B. Coytrell leg., 1 ♀ identified in the past as Pseudogonatopus ruens Olmi (Olmi 1984) (NHMUK); KwaZulu-Natal, Drakensberg, Cathedral Peak, inside the park, grass near the gate of Cathedral Peak Hotel, C. 11. IV. 2007, B. 13. IV. 2004, M. Olmi reared, 1 ♀ (MOLC); Western Cape, about 18 km S of Citrusdal, The Baths, 22. I. 2003, sweeping, M. Olmi leg., 1 ♀ (MOLC); same locality label, reared by M. Olmi from adults of Metadelphax propinqua (Fieber) and Toya tuberculosa (Distant) (Delphacidae) as follows: C. 22. I. 2003, B. 24. I. 2003, Sf. 8.2.2003, 1 ♂ (MOLC); C. 22. I. 2003, B. 24. I. 2003, Sf. 7.2.2003, 1 ♀ (MOLC); C. 22. I. 2003, B. 23. I. 2003, Sf. 7.2.2003, 1 ♀ (MOLC); C. 22. I. 2003, B. 24. I. 2003, Sf. 7.2.2003, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0A2BEFFF3ED383FC4BFDB9.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013) and Meenoplidae: Delphacidae: in Mozambique: Metadelphax propinqua (Fieber) (= Toya propinqua (Fieber )), Sogatella kolophon (Kirkaldy) and S. petax Fennah,; in South Africa: Matutinus vitticollis (Stål); Metadelphax propinqua (Fieber) (= Toya propinqua (Fieber )); Toya ceresensis (Muir) and T. tuberculosa (Distant). In Mozambique: unidentified species of Meenoplidae. Natural enemies. Hymenoptera Encyrtidae: in Mozambique: Cheiloneurus gonatopodis R. Perkins and Cheiloneurus exitiosus (Perkins). Ch. gonatopodis is one of the most common natural enemies of Dryinids in Mozambique (see also Echthrodelphax tauricus Ponomarenko, Gonatopus mossambicus Olmi and Gonatopus incognitus Olmi). The reared specimens have the following locality labels: E 3: Mozambique, Maputo Prov., Umbelúzi, para- sitized planthopper collected on 20. X. 1992, M. Olmi reared, 2 ♂♂, 3 ♀♀ of Cheiloneurus gonatopodis Perkins. 28: Mozambique, Maputo Prov., 5 km E Manhiça, parasitized planthopper collected on 25. IX. 1996; cocoon spun on 26. IX. 1996; encyrtids emergence on 13. X. 1996; M. Olmi reared, 1 ♂, 2 ♀♀ of Cheiloneurus exitiosus (Perkins). 30: Mozambique, Maputo Prov., 5 km E Manhiça, parasitized planthopper collected on 25. IX. 1996; cocoon spun on 26. IX. 1996; encyrtids emergence on 14. X. 1996, M. Olmi reared, 2 ♂♂, 2 ♀♀ of Cheiloneurus gonatopodis Perkins. 56: Mozambique, Maputo Prov., 6 km E Manhiça, parasitized planthopper collected on 25. IX. 1996; cocoon spun on 25. IX. 1996; encyrtids emergence on 11. XI. 1996, M. Olmi reared, 1 ♂, 3 ♀♀ of Cheiloneurus gonatopodis Perkins.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE0A2BEFFF3ED383FC4BFDB9.taxon	distribution	Distribution. Burkina Faso, Gambia, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BEFFF3ED4DEFE05F982.taxon	description	(Figs 168 F, 171 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BEFFF3ED4DEFE05F982.taxon	description	Description. ♀. Apterous; body length 3.6 mm. Head brown, except malar space, clypeus and mandible testaceous brown; antenna brown, except scape testaceous; mesosoma brown, except mesoscutum and metanotum testaceousbrown; metasoma brown; legs brown. Antenna clavate, with ADOs in antennomeres 6 – 10; antennomeres in following proportions: 10: 5: 14: 6: 5: 5: 5: 5: 4.5: 9. Head very excavated (Fig. 171 D), dull, granulate; occiput very excavated; frontal line complete; occipital carina absent; POL = 0.5; OL = 1; OOL = 8; temple absent. Palpal formula 4 / 2. Pronotum crossed by strong and broad transverse impression, with anterior collar and sides shiny, unsculptured; disc of pronotum sculptured by numerous longitudinal striae. Mesoscutum shiny, smooth, granulate, without lateral points. Mesoscutellum small and unsculptured, slightly sloping anteriorly. Metanotum weakly excavated behind mesoscutellum, transversely striate, with lateral sides protruding; lateral protrusions of metanotum rounded. Metapectalpropodeal disc dull, with anterior surface and top sculptured by numerous longitudinal striae; propodeal declivity transversely striate; top of metapectal-propodeal disc with strong median longitudinal furrow. Meso-metapleural suture distinct and complete. Mesopleuron and metapleuron shiny, transversely striate. Protarsomeres in following proportions: 14: 2.5: 5: 13: 23. Enlarged claw (Fig. 168 F) with one large subapical tooth and one row of nine lamellae + one bristle. Protarsomere 5 (Fig. 168 F) with two rows of 20 lamellae; apex with approximately 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BEFFF3ED4DEFE05F982.taxon	materials_examined	Material examined. Type: ♀ holotype: CAMEROON: South Province, Campo Ma’an National Park, 43.3 km 108 ° ESE Campo, 290 m, 7. IV. 2000, sifted litter in rainforest, B. L. Fisher leg., BLF 2298 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BEFFF3ED4DEFE05F982.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BEFFF3ED4DEFE05F982.taxon	distribution	Distribution. Cameroon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BE0FF3ED019FE14FE0C.taxon	description	(Fig. 168 G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BE0FF3ED019FE14FE0C.taxon	description	Description. ♀. Apterous; body length 2.7 mm. Head yellow-testaceous; antenna yellow-testaceous, except an- tennomeres 3 – 10 slightly brownish; mesosoma yellow-testaceous; petiole black; metasoma partly brownish; legs yellow-testaceous. Head excavated. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression. Mesometapleural suture obsolete. Propodeal declivity transversely striate. Protarsomere 1 longer than 4 (13: 10). Enlarged claw (Fig. 168 G) with one large subapical tooth and one row of 4 – 5 lamellae. Protarsomere 5 (Fig. 168 G) with two rows of 19 – 30 lamellae extending continuously to apex. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BE0FF3ED019FE14FE0C.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Tolagnaro (= Fort-Dauphin), XII. 1942, A. Seyrig leg. (MNHN). Paratype: same locality label as holotype, 1 ♀ (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BE0FF3ED019FE14FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE082BE0FF3ED019FE14FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	description	(Figs 169 C, 170, 171 E, 172 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 3 / 2; pronotum crossed by strong transverse furrow; metanotum with lateral pointed protrusions (Fig. 169 C); meso-metapleural suture obsolete.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	description	Description. ♀ (Fig. 169 C, 170). Apterous; body length 4.4 mm. Completely testaceous-ferruginous, except antennomeres 3 – 10 brown, posterior margin of mesoscutellum and petiole black. Antenna clavate; antennomeres in following proportions: 10: 6: 16: 9: 8: 7: 6: 6: 5: 10. ADOs present in antennomeres 6 – 10. Head (Fig. 170 D) shiny, unsculptured; frontal line complete; occipital carina very short, only present behind ocellar triangle; POL = 1; OL = 2; OOL = 8. Palpal formula 3 / 2. Pronotum shiny, bare, unsculptured, crossed by strong transverse furrow (Fig. 170 C). Mesoscutum shiny, sculptured by few longitudinal keels, without lateral pointed apophyses. Mesoscutellum shiny, unsculptured. Metanotum flat, sculptured by transverse keels, about coplanar to mesoscutellum (Fig. 171 E), with lateral pointed protrusions (Figs 169 C, 170 B). Metapectal-propodeal disc (Fig. 169 C) shiny, unsculptured, with track of median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron strongly transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 16: 3: 6: 15: 26. Enlarged claw (Fig. 172 A) with one large subapical tooth and one row of seven lamellae + one bristle. Protarsomere 5 (Fig. 172 A) with two rows of 16 lamellae; apex with about 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19462): UNION OF THE COMOROS: Nzwani (= Anjo- uan) Island, Mount Ntringui, 12.19641 ° S 44.41791 ° E, 550 m, 29. I. 2009, rainforest, B. L. Fisher et al., BLF 21326 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	distribution	Distribution. Union of the Comoros.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE072BE1FF3ED3D6FB03FA5A.taxon	etymology	Etymology. The species is named comorensis after the collection site in the Comoros Islands.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	description	(Figs 171 F, 172 B, 173 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	description	Description. ♀. Apterous; body length 2.4 – 3.3 mm. Head brown, except mandible, clypeus and anterior region of frons testaceous; antenna brown, except scape testaceous; mesosoma reddish-darkened; in specimens from South Africa, Port Edward, mesosoma more or less broadly darkened or black (in some cases almost completely black, except mesoscutum partly yellow); metasoma brown; legs brown-reddish, except stalks of meso- and metafemur darkened and distal extremities of meso- and metatibia brown. Antenna clavate; antennomeres in following proportions: 8: 5.5: 12: 7.5: 6: 6: 5.5: 5.5: 5.5: 8. Head excavated, dull, granulate; frontal line incomplete; occipital carina absent; POL = 1; OL = 2; OOL = 7.5. Palpal formula 4 / 2. Pronotum shiny, unsculptured, crossed by strong transverse impression; disc of pronotum with posterior contraction near posterior margin (Fig. 171 F). Mesoscutum dull, sculptured by numerous longitudinal keels, without lateral pointed apophyses. Metanotum flat, sculptured by transverse striae; sides of metanotum rounded (Fig. 171 F). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 13: 3: 4.5: 13: 19. Enlarged claw (Fig. 172 B) with one large subapical tooth and one row of seven lamellae. Protarsomere 5 (Fig. 172 B) with two rows of approximately 24 lamellae; apex with approximately seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.3 – 2.5 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs yellow, except metacoxa darkened. Antenna filiform; antennomeres in following proportions: 6: 5: 9: 9: 9: 9: 8.5: 8.5: 8: 11; antennomere 3 more than 4.5 × as long as broad (9: 1.7). Head shiny, convex, slightly granulate; frontal line absent; occipital carina incomplete, only present behind lateral ocelli; temple distinct; POL = 6; OL = 2.5; OOL = 3.5. Palpal formula 4 / 2. Mesoscutum shiny, very slightly granulate. Notauli complete, posteriorly joint. Mesoscutellum and metanotum shiny, very slightly granulate. Metapectal-propodeal disc reticulate rugose, with median longitudinal furrow; propodeal declivity reticulate rugose. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; distal part of 2 r-rs & Rs vein longer than proximal part (18: 6). Dorsal process of paramere (Fig. 173 A) short and transverse, with pointed apex. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Maputo Prov., Maputo, Costa do Sol, Municipal Agricultural Nursery, reared from Nisia nervosa (Mots.), C. 11. X. 1992, Sf. 29. X. 1992, M. Olmi reared (AMNH). Paratypes: same locality label as holotype, C. 11. X. 1992, Sf. 30. X. 1992, 1 ♀ (AMNH); same locality label as ho- lotype, C. 11. X. 1992, Sf. 5. XI. 1992, 1 ♀ (AMNH); same locality label as holotype, C. 11. X. 1992, Sf. 30. X. 1992, 1 ♂ (AMNH); same locality label as holotype, C. 6. X. 1992, Sf. 25. X. 1992, 1 ♂ (AMNH). Other material: SOUTH AFRICA: KwaZulu-Natal, Port Edward, along road from R 61 to Umtamvuna Nature Reserve, C. 13. IV. 2007, B. 14. IV. 2007; C. 13. IV. 2007, B. 15. IV. 2007; C. 13. IV. 2007, B. 17. IV. 2007; M. Olmi reared from adults of Sogatella kolophon (Kirkaldy), 4 ♀♀, 3 ♂♂ (3 ♀♀, 2 ♂♂ in MOLC, 1 ♀, 1 ♂ in SAM) (Olmi 2009 b).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	biology_ecology	Hosts. Meenoplidae and Delphacidae (Guglielmino et al. 2013): in Mozambique: Nisia nervosa (Motschulsky) (Meenoplidae); in South Africa: Sogatella kolophon (Kirkaldy) (Delphacidae).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	distribution	Distribution. Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE062BE3FF3ED07EFE4BFE70.taxon	discussion	Remarks. The ♀ mesosoma of Gonatopus festivus Olmi can vary in colour, from totally testaceous to partly black or brown. If the colour is partly black or brown, ♀♀ of G. festivus can be confused with G. harteni Olmi. In this case, ♀♀ of these two species are indistinguishable. However, the two species are distinct, because their ♂♂ show strong differences.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE042BE4FF3ED0FBFE5BFD2C.taxon	description	(Figs 172 C, 174 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE042BE4FF3ED0FBFE5BFD2C.taxon	description	Description. ♀. Apterous; body length 3.3 mm. Testaceous, except petiole black; antenna brown, except antenno- meres 1 – 3 testaceous and 9 – 10 whitish. Antenna clavate, with ADOs in antennomeres 6 – 10; antennomeres in following proportions: 10: 5: 16: 9: 8.5: 7.5: 7: 6.5: 6: 11. Head very excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 0.5; OL = 1; OOL = 7; temple distinct. Palpal formula 4 / 2. Pronotum crossed by strong and broad transverse impression, unsculptured, shiny. Mesoscutum very slender, shiny, unsculptured, without lateral pointed apophyses (Fig. 174 A). Mesoscutellum small and unsculptured. Metanotum excavated behind mesoscutellum (Fig. 174 B), transversely striate, with sides protruding; lateral protrusions of metanotum rounded (Fig. 174 A). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity with transverse striae; top of metapectal-propodeal disc with track of median longitudinal furrow. Meso-metapleural suture distinct and complete. Mesopleuron shiny, unsculptured. Metapleuron shiny, almost completely unsculptured, partly transversely striate. Protarsomeres in following proportions: 13: 3: 4.5: 12: 23. Enlarged claw (Fig. 172 C) with one large subapical tooth and one row of nine lamellae + one bristle. Protarsomere 5 (Fig. 172 C) with two rows of 16 lamellae; apex with approximately 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE042BE4FF3ED0FBFE5BFD2C.taxon	materials_examined	Material examined. Type: ♀ holotype: GABON: Estuaire Prov., 21.331 km NNW of Libreville, 00 ° 34.6 ’ N 09 ° 20.1 ’ E, 10 m, 24. II. 1998, 1742, Brian L. Fisher leg., sifted litter (leaf mold, rotten wood), littoral rainforest (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE042BE4FF3ED0FBFE5BFD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE042BE4FF3ED0FBFE5BFD2C.taxon	distribution	Distribution. Gabon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	description	(Fig. 172 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	description	Description. ♀. Apterous; body length 3.1 mm. Completely testaceous, except petiole black. Antenna clavate; an- tennomeres in following proportions: 6: 5: 14: 7: 6: 6: 6: 5: 4.5: 7. Head excavated, shiny, very slightly granulate; frontal line complete; occipital carina absent; POL = 1.5; OL = 2; OOL = 7. Palpal formula 4 / 2. Pronotum shiny, granulate, crossed by strong transverse impression. Mesoscutum dull, granulate. Mesoscutellum shiny, unsculptured. Metanotum flat, unsculptured, about coplanar to mesoscutellum; sides of metanotum rounded. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 15: 3: 5: 13: 20. Enlarged claw (Fig. 172 D) with one large subapical tooth and one row of five lamellae. Protarsomere 5 (Fig. 172 D) with two rows of 17 lamellae; apex with eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: GHANA: Greater Accra Region, Dawhenya, 3. VII. 1972, on rice, Sche- belreiter leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	distribution	Distribution. Ghana.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE022BE6FF3ED3B0FBC6FE0C.taxon	discussion	Remarks. Gonatopus ghanensis was described under the genus Pseudogonatopus, that is now a junior synonym of Gonatopus. This is the reason for the new combination presented above.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	description	(Figs 172 E, F, 173 B, 174 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	description	Description. ♀. Apterous; body length 2.0 – 3.2 mm. Head black, except mandible, anterior region of frons and clypeus testaceous; antenna brown, except scape and part of flagellomere 1 testaceous. Mesosoma black, except pronotum brown, distal extremity of propodeal declivity and most part of mesoscutum testaceous (anterior margin of mesoscutum darkened); petiole black; metasoma black; legs testaceous, except coxae and clubs of femora partly black. Antenna clavate; antennomeres in following proportions: 6: 4: 10: 7: 5: 5: 5: 5: 4.5: 6.5. Head shiny, unsculptured, except occiput granulate; frontal line complete; occipital carina absent; POL = 1.5; OL = 2; OOL = 7. Palpal formula 4 / 2. Pronotum shiny, bare, crossed by strong transverse impression, with collar unsculptured and disc slightly granulate. Mesoscutum granulate, without lateral pointed apophyses. Mesoscutellum unsculptured. Metanotum flat, granulate, about coplanar to mesoscutellum; sides of metanotum rounded. Metapectal-propodeal disc shiny, slightly granulate, with track of median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 13: 3: 4: 12: 18. Enlarged claw (Figs 172 E, F) with one large subapical tooth and one row of five lamellae. Protarsomere 5 (Figs 172 E, F) with two rows of 17 – 18 lamellae; apex with 5 – 6 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 – 1.7 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tarsomeres 1 – 4 whitish. Antenna filiform; antennomeres in following proportions: 2: 3: 4: 4: 4: 3.5: 3.5: 3.5: 3.5: 4.5; antennomere 3 less than 3 × as long as broad (4: 2). Head dull and granulate; clypeus very prominent, shallowly emarginated, sticking out as bill (Fig. 174 C); frontal line absent; occipital carina absent; POL = 5; OL = 3; OOL = 4; vertex with two oval shiny areas situated between lateral ocelli and eyes (areas not anteriorly surrounded by high carina). Palpal formula 4 / 2. Mesoscutum shiny, granulate. Notauli incomplete, reaching approximately 0.75 × length of mesoscutum. Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc shiny, unsculptured, with median longitudinal furrow; propodeal declivity unsculptured. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 6). Dorsal process of paramere (Fig. 173 B) long and with broad apex. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	materials_examined	Material examined. Types: ♀ holotype: CAPE VERDE ISLANDS: Santiago Island, São Jorge dos Orgãos, 19. XI. 1985, on lucerne, A. van Harten leg. (AMNH). Paratypes: same locality label as holotype, 2 ♂♂ (AMNH). Other material: GAMBIA: Kombo North District, Abuko Nature Reserve, 10. XI. 1992, 1 ♀ (ZIL). SOUTH AF- RICA: KwaZulu-Natal, Nhlazatshe, NNW Denny Dalton, roadside S of hill, 28 ° 11 ’ S 31 ° 12 ’ E, 1000 m, 1. XII. 1999, ex emergence box with stems of Mikania capensis, DEL 12.03, O. C. Neser, J. Balciunas & Grobbelaar leg., 1 ♀ (SANC); Western Cape, near George, N 12 - Road to Montagu Pass junction, C. 25. I. 2003, B. 27. I. 2003, Sf. 19. II. 2003; C. 25. I. 2003, B. 28. I. 2003, Sf. 19. II. 2003; M. Olmi reared from Syndelphax capellana (Jacobi), 2 ♀♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in South Africa: Syndelphax capellana (Jacobi).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	distribution	Distribution. Cape Verde Islands, Gambia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE012BE6FF3ED793FE4BF837.taxon	discussion	Remarks. The ♀ mesosoma of Gonatopus festivus Olmi can vary in colour, from totally testaceous to partly black or brown. If the colour is partly black or brown, ♀♀ of G. festivus can be confused with G. harteni Olmi. In this case, ♀♀ of these two species are indistinguishable. However, the two species are distinct, because their ♂♂ show strong differences.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BE7FF3ED6D7FE05FC6D.taxon	description	(Figs 174 D-F, 175 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BE7FF3ED6D7FE05FC6D.taxon	description	Description. ♀. Apterous; body length 3.6 mm. Head brown-testaceous, except anterior half of frons, malar space and clypeus testaceous; antenna brown, except antennomeres 1 and 9 testaceous and 10 whitish; mesosoma browntestaceous, except mesoscutum and posterior extremity of propodeal declivity testaceous; petiole black; metasoma brown; legs brown-testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 15: 7: 6: 5: 5: 5.5: 5.5: 10. Head very excavated (Fig. 174 D), almost bare, shiny, finely punctate, unsculptured among punctures; frontal line complete; occipital carina almost completely absent (only short track behind ocellar triangle); POL = 1; OL = 1.5; OOL = 7. Palpal formula 4 / 2. Pronotum bare, crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, unsculptured, laterally without pointed apophyses (Fig. 174 E). Mesoscutellum shiny, smooth, flat. Metanotum short, transversely striate, about coplanar to mesoscutellum (Fig. 174 F); sides of metanotum protruding, with pointed protrusions (Fig. 174 E). Metapectal-propodeal disc shiny, almost bare, with top and anterior surface unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 2.5: 4: 14: 24. Enlarged claw (Fig. 175 E) with one large subapical tooth and one row of eight lamellae + one bristle. Protarsomere 5 (Fig. 175 E) with two rows of 5 + 9 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BE7FF3ED6D7FE05FC6D.taxon	materials_examined	Material examined. Type: ♀ holotype: CAMEROON: South, near Ebolowa, Nkoemvon, 30. XI. 1979 – 5. I. 1980, MT, in forest, D. Jackson leg., NHMUK (E) 2007 – 19 (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BE7FF3ED6D7FE05FC6D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BE7FF3ED6D7FE05FC6D.taxon	distribution	Distribution. Cameroon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BF8FF3ED272FDE2FE70.taxon	description	(Figs 173 C, 175 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BF8FF3ED272FDE2FE70.taxon	description	Description. ♀. Apterous; body length 2.3 mm. Head brown-black, except mandible, anterior margin of frons, malar space, clypeus and short median longitudinal stripe near clypeus testaceous; antenna brown, except scape testaceous; mesosoma black, except apex of propodeal declivity testaceous; petiole black; metasoma brown; legs brown, except part of coxae, trochanters, articulations and tarsi testaceous. Antenna clavate; antennomeres in following proportions: 5: 4.5: 12: 7: 5.5: 5: 5: 5: 5: 7. Head excavated, shiny, slightly granulate; frontal line incomplete, not present in anterior third; occipital carina absent; POL = 1.5; OL = 1.5; OOL = 6.5. Palpal formula 4 / 2. Pronotum almost bare, crossed by strong transverse impression, with anterior collar shiny, unsculptured and with disc and sides slightly granulate. Mesoscutum dull, granulate, laterally without lateral pointed apophyses. Mesoscutellum shiny, smooth, flat. Metanotum short, dull, granulate and rugose, about coplanar to mesoscutellum; sides of metanotum rounded, not laterally protruding. Metapectal-propodeal disc shiny, almost bare, with anterior surface sculptured by numerous slight longitudinal striae; top of metapectal-propodeal disc with track of median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 12: 2: 3.5: 10: 16. Enlarged claw (Fig. 175 F) with one large subapical tooth and one row of five lamellae. Protarsomere 5 (Fig. 175 F) with two rows of 15 lamellae; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.7 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tarsi and protibia testaceous. Antenna setose, filiform; antennomeres in following proportions: 5: 3: 6: 5: 5: 5: 5: 5: 5: 7; antennomere 3 about 3 × as long as broad: 6: 2. Head setose, dull and granulate; frontal line absent; occipital carina absent; POL = 5; OL = 2; OOL = 3; vertex with track of two oval areas situated between lateral ocelli and eyes (areas not anteriorly surrounded by high carina). Palpal formula 4 / 2. Mesoscutum dull and strongly granulate. Notauli incomplete, reaching approximately 0.7 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc dull and completely reticulate rugose; propodeal declivity reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part approximately as long as proximal part. Dorsal process of paramere (Fig. 173 C) long and with broad distal region; distal margin of dorsal process slightly serrate. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BF8FF3ED272FDE2FE70.taxon	materials_examined	Material examined. Types: ♀ holotype: MOZAMBIQUE: Niassa, Cuamba, Campus of the Catholic Univer- sity of Mozambique, reared from parasitized unidentified Delphacid, C. 23. III. 2003, B. 25. III. 2003, Sf. 14. IV. 2003, M. Olmi reared (MOLC). Paratype: same locality label as holotype, C. 21. III. 2003, B. 22. III. 2003, Sf. 11. IV. 2003, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BF8FF3ED272FDE2FE70.taxon	biology_ecology	Hosts. In Mozambique: unidentified species of Delphacidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE002BF8FF3ED272FDE2FE70.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1F2BF9FF3ED3B1FE1FFDB8.taxon	description	(Fig. 175 G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1F2BF9FF3ED3B1FE1FFDB8.taxon	description	Description. ♀. Apterous; body length 3.4 mm. Head brown, except mandible, anterior margin of frons, malar space, clypeus and short median longitudinal stripe near clypeus testaceous; antenna brown, except scape testaceous and flagellomere 10 whitish; prothorax testaceous-brown; mesoscutum testaceous, except anterior third dark; rest of mesosoma brown-testaceous and partly brown; metasoma brown; legs brown, except part of coxae, trochanters, articulations and tarsi testaceous ot testaceous-brown. Antenna clavate; antennomeres in following proportions: 9: 5: 10: 5: 4: 4: 4: 4: 4: 9. Head excavated, dull, granulate; frontal line complete, present in ocellar triangle (ocellar triangle divided into two halfs by median longitudinal keel); occipital carina incomplete, shortly present behind lateral ocelli; POL = 1; OL = 3; OOL = 6.5. Palpal formula 4 / 2. Pronotum bare, crossed by strong transverse impression, shiny, unsculptured. Mesoscutum sculptured by many longitudinal striae, laterally without pointed apophyses. Mesoscutellum shiny, smooth, flat. Metanotum short, dull, sculptured by many transverse striae, little excavated behind mesoscutellum; sides of metanotum rounded, not laterally protruding. Metapectal-propodeal disc dull, with top rugose and anterior surface sculptured by numerous irregular striae; anterior surface of metapectal-propodeal disc with track of median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 13: 3: 5: 14: 22. Enlarged claw (Fig. 175 G) with one large subapical tooth and one row of six lamellae. Protarsomere 5 (Fig. 175 G) with two rows of 12 + 8 lamellae; apex with approximately 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1F2BF9FF3ED3B1FE1FFDB8.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Eshowe, near Ntumeni N. Res., 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg. (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1F2BF9FF3ED3B1FE1FFDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1F2BF9FF3ED3B1FE1FFDB8.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BF9FF3ED4DFFE14FA5B.taxon	description	(Figs 174 G, H)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BF9FF3ED4DFFE14FA5B.taxon	description	Description. ♀. Apterous; body length 3.5 mm. Head and antenna missing in the only known specimen (head black and antenna yellow, except antennomere 10 white, according to the original description); pronotum yellow, except collar black; mesoscutum yellow, except anterior surface black; mesoscutellum yellow; rest of mesosoma black, except distal extremity of propodeal declivity testaceous; petiole black; metasoma yellow-reddish; legs yellow (prolegs missing in the only known specimen). Antenna distally thickened (according to fig. 6 in Benoit 1963). Palpal formula 4 / 2 (according to figs 4 - 5 in Benoit 1963, and the original description). Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum and mesoscutellum shiny, unsculptured. Mesoscutum without lateral pointed apophyses (Fig. 174 G). Metanotum transversely striate, with sides rounded (Fig. 174 G), about coplanar to mesoscutellum (Fig. 174 H). Meso-metapleural suture obsolete. Metapectal-propodeal disc shiny, smooth, bare, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Prolegs missing in the only known specimen, so that the chela is unknown. Chela not described in the original paper of Benoit. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BF9FF3ED4DFFE14FA5B.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Brickaville, reared from a parasit- ized macropterous adult of Perkinsella saccharicida Kirkaldy, B. 24. IX. 1960, Sf. 17. X. 1960, M. Sigwalt reared (MHNG).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BF9FF3ED4DFFE14FA5B.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in Madagascar: Perkinsiella saccharicida Kirkaldy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BF9FF3ED4DFFE14FA5B.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BFBFF3ED060FD72FF50.taxon	description	(Fig. 176 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BFBFF3ED060FD72FF50.taxon	description	Description. ♀. Apterous; body length 3.8 – 5.4 mm. Head black, except mandible and clypeus testaceous or yellow; antenna black or brown-black, except scape testaceous and flagellomere 8 whitish; mesosoma and metasoma black; legs brown-black. Antenna distally thickened; antennomeres in following proportions: 9: 7: 23: 14: 14: 10: 11: 10: 10: 14. Body covered with long sparse setae. Head excavated, shiny; in holotype, frons almost completely unsculptured, except few short irregular keels on sides of frontal line; frons of holotype with two parallel complete very close frontal lines; in two specimens from Sierra Leone, 8 ° 53 ’ N 12 ° 3 ’ W, frons partly rugose on sides of frontal line (less rugose in the smallest specimen, more rugose in the largest specimen (5.4 mm )); frontal line complete; occipital carina absent; temple distinct; POL = 1; OL = 2; OOL = 9. Palpal formula 4 / 2. Pronotum shiny, crossed by strong transverse impression; in holotype and the smallest specimen from Sierra Leone, 8 ° 53 ’ N 12 ° 3 ’ W, pronotum completely unsculptured; in the largest specimen from Sierra Leone, 8 ° 53 ’ N 12 ° 3 ’ W, disc rugose. Mesoscutum rugose, with numerous longitudinal keels, without lateral pointed apophyses. Mesoscutellum small, shiny, smooth. Metanotum dull, transversely striate, with sides rounded. Meso-metapleural suture almost completely obsolete (only few parts visible). Metapectal-propodeal disc with track of median longitudinal furrow; top and anterior surface of metapectal-propodeal disc shiny, unsculptured (partly rugose in the smallest specimen from Sierra Leone, 8 ° 53 ’ N 12 ° 3 ’ W; completely and strongly rugose in the largest specimen from Sierra Leone, 8 ° 53 ’ N 12 ° 3 ’ W); propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Protarsomeres in following proportions: 16: 3: 6: 16.5: 26. Enlarged claw (Fig. 176 A) with one large subapical tooth and one row of 12 – 14 lamellae. Protarsomere 5 (Fig. 176 A) with two rows of 22 – 33 lamellae; apex with approximately 12 – 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BFBFF3ED060FD72FF50.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Isoanala, III. 1938, A Seyrig leg. (MNHN). Other material: SIERRA LEONE: Makeni, 08 ° 53 ’ N 12 ° 03 ’ W, 27. XI. 1993, LT, L. Cederholm, R. Danielsson, R. Hall leg., 2 ♀♀ (ZIL, AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BFBFF3ED060FD72FF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1E2BFBFF3ED060FD72FF50.taxon	distribution	Distribution. Madagascar, Sierra Leone.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1C2BFCFF3ED767FBDAFE0C.taxon	description	(Figs 176 B, 177 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1C2BFCFF3ED767FBDAFE0C.taxon	description	Description. ♀ (Figs 177 A, B). Apterous; body length 3.0 – 3.6 mm. Completely yellow-reddish, except petiole black. Head excavated, shiny, slightly granulate, with frons unsculptured. Palpal formula 4 / 2. Pronotum shiny, granulate, crossed by strong transverse impression. Mesoscutum and mesoscutellum granulate. Mesoscutum without lateral pointed apophyses (Fig. 177 A). Metanotum with sides rounded (Fig. 177 A), about coplanar to mesoscutellum (Fig. 177 B). Metapectal-propodeal disc granulate, with track of median longitudinal furrow; propodeal declivity transversely striate. Meso-metapleural suture obsolete. Mesopleuron and metapleuron not transversely striate; occasionally striate. In specimen from Niger, Tarna goulbin, body almost completely unsculptured, except transverse striae on propodeal declivity and pleuron. Protarsomeres in following proportions: 13: 2: 4: 11: 18. Enlarged claw (Fig. 176 B) with one large subapical tooth and one row of 4 – 5 lamellae. Protarsomere 5 (Fig. 176 B) with two rows of 20 – 37 lamellae extending continuously to apex. Tibial spurs 1 / 0 / 1.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1C2BFCFF3ED767FBDAFE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Maniema, Kalombo, 23. VI. 1947, M. Poll leg., by sweeping on grass on banks of Lualaba River near Kisale Lake (MRAC). Other material: NIGER: Maradi, Tarna goulbin, 1 ♀ (TAMU). SOUTH AFRICA: Eastern Cape, Somerset East, 1 ♀ (AMNH); Western Cape, Mossel Bay, 30. XII. 1938, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, II. 1922, 1 ♀ (NHMUK); Western Cape, Ceres, IV. 1925, R. E. Turner leg., 1 ♀ (NHMUK). YEMEN: Sana’a, 9. V. 1991, in rad- ish field, A. van Harten leg., 2 ♀♀ (MOLC); Wadi Dhar, 10. VII. 1991, ex cocoon on Sorghum, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1C2BFCFF3ED767FBDAFE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1C2BFCFF3ED767FBDAFE0C.taxon	distribution	Distribution. Democratic Republic of the Congo, Niger, South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	description	(Figs 173 D, 174 I, M, 176 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	description	Description. ♀. Apterous; body length 2.8 – 3.1 mm. Head reddish, except ocellar region brownish and anterior region of frons yellow-reddish; antenna brown, except antennomeres 1 – 3 yellow; mesosoma reddish; petiole black; metasoma black, except tergite 1 reddish; legs yellow-reddish, except clubs of femora brown. Head excavated, shiny, smooth, granulate; area in front of anterior ocellus unsculptured; occiput granulate. Palpal formula 2 / 2, or 3 / 2, or 4 / 2. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum dull, with some longitudinal striae, without lateral pointed apophyses (Fig. 174 I). Metanotum flat, transversely striate, as long as mesoscutum, with sides rounded (Fig. 174 I). Metapectal-propodeal disc dull, without track of median longitudinal furrow; propodeal declivity transversely striate. Meso-metapleural suture obsolete. Mesopleuron and metapleuron transversely striate. Protarsomeres in following proportions: 9: 2: 3: 6: 10. Enlarged claw (Fig. 176 C) with one large subapical tooth and one row of five lamellae. Protarsomere 5 (Fig. 176 C) with two rows of 15 – 17 lamellae; apex with 5 – 8 lamellae. Tibial spurs 1 / 0 / 1. ♂ (Fig. 174 M): fully winged; body length 2.5 mm. Head black, except mandible yellow; antenna brown; me- sosoma black; metasoma brown; legs yellow. Antennomeres in following proportions: 5.5: 5.5: 12: 11: 10: 10: 10: 10: 9: 13; antennomere 3 more than 4 × as long as broad (4.8). Head dull, setose, granulate; frontal line absent between antennal toruli; occipital carina absent; POL = 8; OL = 4; OOL = 3; greatest breadth of lateral ocelli about as long as OOL. Palpal formula 2 / 2. Mesoscutum setose, granulate. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of lateral ocelli (2: 3). Mesoscutellum and metanotum setose, granulate. Metapectal-propodeal disc dull, with longitudinal median furrow; propodeal declivity reticulate rugose; top of metapectal-propodeal disc with short irregular keels. Forewing hyaline, without dark transverse bands. Dorsal process of paramere (Fig. 173 D) long and broad, distally sharp. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: Swakopmund, 26 – 30. I. 1972, Southern African Expedi- tion, B. M. 1972 (NHMUK). Paratypes: same locality label as holotype, 7 ♀♀, 1 ♂ (NHMUK), 2 ♀♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	distribution	Distribution. Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFCFF3ED79CFF2BF91E.taxon	discussion	Remarks. The record from South Africa (Grahamstown) by Olmi (1984) is an error; it is in fact a specimen of Gonatopus amoenus Olmi (see account of this species above). The ♀ from Grahamstown has a track of a median longitudinal furrow on the top of metapectal-propodeal disc, which correctly assigns this specimen to the latter species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	description	(Figs 173 E, 174 J – L, 176 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	description	Description. ♀. Apterous; body length 2.5 – 2.8 mm. Head yellow-testaceous; antenna brown, except scape and flag- ellomere 8 white or yellow; occasionally flagellomere 8 brown; mesosoma yellow-testaceous; petiole black; metasoma brownish or yellow-testaceous; legs yellow-testaceous. Head (Fig. 174 L) excavated. Palpal formula 3 / 2 or 4 / 2. Pronotum crossed by strong transverse impression. Mesoscutum without lateral pointed apophyses (Fig. 174 J). Metanotum transversely striate, about coplanar to mesoscutellum (Fig. 174 K), with sides protruding (Fig. 174 J); in holotype lateral protrusions of metanotum rounded (Fig. 174 J); in specimen from South Africa, 26 ° 45.58 ’ S 27 ° 48.22 ’ E, lateral protrusions of metanotum pointed. Metapectal-propodeal disc shiny, unsculptured. Mesopleuron, metapleuron and propodeal declivity transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 slightly longer than 4 (12: 11). Enlarged claw (Fig. 176 D) with one large subapical tooth and one row of 8 – 11 lamellae. Protarsomere 5 (Fig. 176 D) with two rows of 16 – 19 lamellae; apex with 9 – 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.1 mm. Head brown; antenna testaceous; mesosoma and metasoma brown; legs testaceous and partly darkened. Antenna filiform; antennomeres in following proportions: 3.5: 3: 4: 3: 3: 3.5: 3.5: 3: 4: 6; third antennomere 4 × as long as broad (4: 1). Head shiny, granulate, with two smooth areas on sides of lateral ocelli; frontal line absent; occipital carina absent; POL = 5; OL = 2; OOL = 3. Palpal formula 4 / 2. Mesoscutum shiny, granulate. Notauli complete, posteriorly joint. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc shiny, dorsally unsculptured, laterally rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (10: 5). Dorsal process of paramere (Fig. 173 E) long and slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	materials_examined	Material examined. Types: ♀ holotype of G. similis: SOUTH AFRICA: Eastern Cape, Algoa Bay, 25. V. 1898, Brauns leg. (MCPM); ♀ holotype of P. mauritianus: MAURITIUS: Palmyre, IV. 1955, ex nymphs of Perkinsiella saccharicida Kirkaldy, J. R. Williams reared (NHMUK). Paratypes of P. mauritianus: same locality label as holotype, 2 ♀♀ (NHMUK). Other material: GAMBIA: Lower River Division, Upper Baddibu, Farafenni, 1 ♀ (KJHC). KENYA: Rift Valley Prov., Nguruman, Oloibortoto River irrigation scheme, 01 ° 48 ’ S 36 ° 04 ’ E, 12 – 16. VI. 1999, MT, R. Copeland leg., 1 ♀ (NMK). LA REUNION: La Reunion, 1 ♀ (AMNH). LESOTHO: Mamathes, 1 ♀ (AMNH). MOZAMBIQUE: Gaza Prov., 2 km E of Incomati River bridge, along road EN 1 between Macia and Palmeira, 1 ♀ (AMNH); Maputo Prov., 6 km N of Palmeira, along road EN 1 between Maputo and Macia, 1 ♀ (AMNH). SOUTH AFRICA: Eastern Cape, Port St. Johns, 1 – 17. III. 1924, R. E. Turner leg., 1 ♀ (NHMUK); Eastern Cape, about 16 km E Port Elizabeth, S. George’s Strand, 1 ♀ (AMNH); Free State, Sasolburg, Wonderwater Colliery, 26 ° 45.58 ’ S 27 ° 48.22 ’ E, 1450 m, 23. XI. 1999, D-Vac, IS 09 b, seeded rehabilitation, east burm, slope, M. Stiller leg., 1 ♀ (SANC); KwaZulu-Natal, Port Edward, along road from R 61 to Umtamvuna Nature Reserve, C. 13. IV. 2007, B. 15. IV. 2007, M. Olmi reared from adults of Sogatella kolophon (Kirkaldy), 1 ♀ (MOLC); same locality label, C. 13. IV. 2007, B. 17. IV. 2007, 1 ♀ (MOLC); Limpopo: 15 km E Klaserie, Guernsey Farm, 1 ♀ (CNC); Western Cape, Mossel Bay, 10 – 12.1938, R. E. Turner leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in La Reunion: Peregrinus maidis Ashmead; in Mauritius: Perkinsiella saccharicida Kirkaldy and Numata muiri (Kirkaldy) (= Dicranotropis muiri Kirkaldy); in Mozambique: unidentified Delphacidae; in South Africa: Sogatella kolophon (Kirkaldy). Biology. The biology of this species was partly studied in Mauritius by Williams (1956). The pupal stage in Mauritius lasts 18 – 24 days in March-April. Natural enemies. In Mauritius this species may be parasitized by Cheiloneurus gonatopodis R. Perkins 1906 (Encyrtidae) (Williams 1956, 1958; Prinsloo 1983; Ganeshan & Williams 2000; Guerrieri & Viggiani 2005).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	distribution	Distribution. Gambia, Kenya, La Reunion, Lesotho, Mauritius, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1B2BFDFF3ED0AFFEF3F964.taxon	discussion	Remarks. Gonatopus similis Brues and Pseudogonatopoides mauritianus Williams were considered synonyms by Olmi (1984) after comparison of ♀♀. However, only the study of ♂ specimens from Mauritius can confirm this synonymy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1A2BFEFF3ED17AFE14FD49.taxon	description	(Figs 175 A – C, 176 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1A2BFEFF3ED17AFE14FD49.taxon	description	Description. ♀. Apterous; body length 3.0 – 3.4 mm. Completely yellow-testaceous, except petiole black. Body shiny, bare, unsculptured. Head (Fig. 175 A) excavated. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression. Mesoscutum without lateral pointed apophyses (Fig. 175 B). Metanotum about coplanar to mesoscutellum (Fig. 175 C), with sides protruding (Fig. 175 B); lateral protrusions of metanotum rounded (Fig. 175 B). Metapectal-propodeal disc with median longitudinal furrow; propodeal declivity transversely striate. Meso-metapleural suture distinct and complete. Mesopleuron and metapleuron transversely striate. Protarsomere 1 slightly shorter than 4 (13: 14). Enlarged claw (Fig. 176 E) with one large subapical tooth and one row of seven lamellae. Protarsomere 5 (Fig. 176 E) with two rows of approximately 20 lamellae; apex with approximately 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1A2BFEFF3ED17AFE14FD49.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toamasina, Brickaville, reared from Perkinsella saccharicida Kirkaldy, B. 3. VI. 1961, Sf. 3. VII. 1961, M. Sigwalt reared (MHNG). Paratypes: same locality label as holotype, Sf. III. 1961, 1 ♀ (MHNG); same locality label as holotype, Sf. 23. V. 1961, 1 ♀ (MHNG); same locality label as holotype, 3. VI. 1961, 1 ♀ (MRAC); same locality label as holotype, Sf. 31. VII. 1961, 1 ♀ (MRAC) (accord- ing to the original description the type series is composed of 6 ♀♀).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1A2BFEFF3ED17AFE14FD49.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in Madagascar: Perkinsiella saccharicida Kirkaldy.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE1A2BFEFF3ED17AFE14FD49.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE192BFFFF3ED56EFE1FFE28.taxon	description	(Figs 173 F, 175 D, 176 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE192BFFFF3ED56EFE1FFE28.taxon	description	Description. ♀. Apterous; body length 2.0 mm. Head brown-reddish, except anterior half of frons, malar space and clypeus testaceous; antenna brown, except scape testaceous and flagellomere 8 light; mesosoma brown-reddish, except mesoscutum almost completely yellow; petiole black; metasoma brown; legs testaceous. Antenna clavate; antennomeres in following proportions: 7: 4.5: 7: 4: 4: 3.5: 3.5: 3.5: 3.5: 6. Head excavated (Fig. 175 D), shiny, slightly granulate; frontal line complete; occipital carina almost completely absent (short track present behind ocellar triangle); POL = 1; OL = 2.5; OOL = 6. Palpal formula 3 / 2. Pronotum almost bare, crossed by strong transverse impression, with anterior collar shiny, unsculptured and with disc slightly granulate. Mesoscutum shiny, unsculptured, laterally without pointed apophyses. Mesoscutellum shiny, smooth, slightly sloping anteriorly. Metanotum short, granulate and about coplanar to mesoscutellum; sides of metanotum protruding, with pointed protrusions. Metapectal-propodeal disc shiny, setose, with top and anterior surface slightly granulate and punctate; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 9: 2: 3.5: 9: 16. Enlarged claw (Fig. 176 F) with one large subapical tooth and one row of eight lamellae. Protarsomere 5 (Fig. 176 F) with two rows of 7 + 8 lamellae; apex of protarsomere 5 with 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 – 2.3 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except articulations testaceous. Antenna setose, filiform; antennomeres in following proportions: 3: 3: 4.5: 4: 4: 4: 4: 4: 4: 5.5; antennomere 3 more than 3 × as long as broad: 5: 1. Head dull and granulate in large ♂ paratype (2.00 mm length); in smaller paratypes head shiny and almost completely unsculptured; frontal line absent; occipital carina absent; POL = 4; OL = 2; OOL = 2; vertex with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas not surrounded anteriorly by high carina). Palpal formula 4 / 2. Mesoscutum dull and strongly granulate in large paratype (length 2.0 mm); in smaller paratypes mesoscutum shiny and punctate or very slightly granulate. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of lateral ocelli (1: 1.5). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull and completely reticulate rugose in large paratype (length 2.0 mm); in smaller paratypes metapectal-propodeal disc shiny, partly slightly reticulate rugose; propodeal declivity without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; in the forewing only costal cell surrounded by well pigmented veins, other basal cells surrounded by less pigmented veins; 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part. Dorsal process of paramere (Fig. 173 F) short and rising from medial region of paramere, with apex rounded. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE192BFFFF3ED56EFE1FFE28.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Buffalo Bay, Goukamma Nature Reserve, C. 26. I. 2003, B. 29. I. 2003, Sf. 16. II. 2003, M. Olmi reared from Syndelphax capellanus (Jacobi), SAM- HYM-P 024846 (SAMC). Paratypes: same locality label as holotype, C. 26. I. 2003, B. 29. I. 2003, Sf. 19. II. 2003, 2 ♂♂ (MOLC); same locality label as holotype, C. 26. I. 2003, B. 30. I. 2003, Sf. 23. II. 2003, 1 ♂ (SAMC); Western Cape, near George, N 12 - Road to Montagu Pass junction, C. 25. I. 2003, B. 29. I. 2003, Sf. 19. II. 2003, M. Olmi reared from Syndelphax capellanus (Jacobi), 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE192BFFFF3ED56EFE1FFE28.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in South Africa: Syndelphax capellana (Jacobi); Syndelphax sp.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE192BFFFF3ED56EFE1FFE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE182BF2FF3ED106FC0BFE54.taxon	description	(Figs 177 C, 178 A, 179 A, 180 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE182BF2FF3ED106FC0BFE54.taxon	description	♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE182BF2FF3ED106FC0BFE54.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Lomami, Up- emba National Park, Lusinga, right tributary of Kenia (right tributary of Lufwa), 1585 m, 19. XII. 1947, G. F. de Witte Mission (MRAC). Other material: KENYA: Coast Prov., Shimba Hills National Park, 04 ° 23.456 ’ S 39 ° 41.687 ’ E, 389 m, MT, just inside Longomwagandi Forest, 19. XI – 3. XII. 2008, R. Copeland leg., 1 ♀ (NMK); Nairobi Prov., ICIPE Campus, Kasarani, 1.2233 ° S 36.8967 ° E, 1600 m, 25. II – 6. III. 2014, MT, along stream, meadow in degraded shrub- / grassland, R. Copeland leg., 1 ♀ (MOLC). SOUTH AFRICA: Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - Y 10, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE182BF2FF3ED106FC0BFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE182BF2FF3ED106FC0BFE54.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF2FF3ED454FE1FFAE2.taxon	description	(Fig. 179 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF2FF3ED454FE1FFAE2.taxon	description	Description. ♀. Apterous; body length 3.6 mm. Head reddish-dark, occasionally brown-black; antenna reddishdark, except antennomere 10 whitish; prothorax and mesoscutum testaceous, or brown-reddish; rest of mesosoma black, except distal extremity of propodeal declivity reddish; petiole black; metasoma reddish-dark; legs testaceous. Antenna clavate; antennomeres of holotype in following proportions: 21: 14: 35: 22: 17: 13: 11: 12: 11: 18. Head excavated, dull, granulate; frontal line complete; occipital carina incomplete, only very shortly visible on sides of lateral ocelli; POL = 3; OL = 6; OOL = 25. Palpal formula 5 / 3. Pronotum shiny, smooth, with some longitudinal keels, crossed by strong transverse impression. Mesoscutum with two pointed lateral apophyses, situated on sides of mesoscutellum. Metanotum short, excavated behind mesoscutellum, with sides rounded. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (15: 17). Enlarged claw (Fig. 179 B) with one large subdistal tooth and one row of 6 – 8 lamellae. Protarsomere 5 (Fig. 179 B) with two rows of 12 – 30 lamellae; apex with 10 – 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF2FF3ED454FE1FFAE2.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Matjesfontein, 1 – 6. XI- 1928, R. E. Turner leg. (NHMUK). Other material: SOUTH AFRICA: Western Cape, near Cape Town, Milnerton, I. 1926, R. E. Turner leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF2FF3ED454FE1FFAE2.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF2FF3ED454FE1FFAE2.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF3FF3ED3E3FBE7FD48.taxon	description	(Figs 179 C, 180 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF3FF3ED3E3FBE7FD48.taxon	description	Description. ♀. Apterous; body length 3.3 – 3.7 mm. Completely testaceous-reddish, except petiole black. Antenna clavate; antennomeres of holotype in following proportions: 19: 11: 34: 21: 22: 14: 13: 12: 12: 19. Head excavated, dull, granulate; frontal line almost complete, absent only near clypeus; occipital carina absent; POL = 5; OL = 6; OOL = 20. Palpal formula 5 / 3. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum dull, granulate, without lateral pointed apophyses (Fig. 180 B). Metanotum transversely striate, excavated behind mesoscutellum, with sides protruding; lateral protrusions of metanotum slightly pointed (Fig. 180 B). Metapectal-propodeal disc dull, granulate, sculptured by many slight longitudinal striae; propodeal declivity, mesopleuron and metapleuron transversely striate. Occasionally propodeal declivity mostly granulate and rugose, with distal third transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 slightly shorter than 4 (13: 16). Enlarged claw (Fig. 179 C) with one large subdistal tooth and one row of 6 – 12 lamellae. Protarsomere 5 (Fig. 179 C) with two rows of 13 – 23 lamellae; apex with 12 – 18 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF3FF3ED3E3FBE7FD48.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Ceres, III. 1925, R. E. Turner leg. (NHMUK). Paratype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 15 – 31. V. 1923, R. E. Turner leg., 1 ♀ (NHMUK). Other material: MOZAMBIQUE: Nampula Prov., Nampula, near Nampula Catholic Seminary, sweep- ing, 21. XI. 1999, M. Olmi leg., 1 ♀ (MOLC); Niassa, Cuamba, Camboja, near University Dormitory, reared from a ♀ adult of Caliscelis sp., C. 15. XI. 1999, B. 19. XI. 1999, Sf. 18. XII. 1999, M. Olmi reared, 1 ♀ (MOLC); same locality label, reared from an adult of Eumecurus hottentottus (Stål), C. 15. XI. 1999, B. 17. XI. 1999, Sf. 18. XII. 1999, M. Olmi reared, 1 ♀ (MOLC). NAMIBIA: Namib / Naukluft Park, Kuiseb River near Gobabeb, 23 ° 34 ’ S 15 ° 03 ’ E, 18. II – 20. III. 1983 (AMNH). SOUTH AFRICA: Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 26. X. 2009 – 8. II. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., 4 ♀♀ (3 in SAMC, 1 in MOLC); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 3 – 31. X. 1997, MT, West Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC). TANZANIA: Chambezi, VII. 1987, on coconut, A. M. Varela leg., 1 ♀ (NHMUK). ZIMBABWE: Sawmills, 9. VII. 1923, Rhodesia Museum, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF3FF3ED3E3FBE7FD48.taxon	biology_ecology	Hosts. Caliscelidae and Cixiidae (Guglielmino et al. 2013): Caliscelidae: in Mozambique: Caliscelis sp.; Cixiidae: in Mozambique: Eumecurus hottentottus (Stål).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE152BF3FF3ED3E3FBE7FD48.taxon	distribution	Distribution. Mozambique, Namibia, South Africa, Tanzania, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE142BF3FF3ED56FFE1FF87F.taxon	description	(Figs 179 D, 180 C, D, I)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE142BF3FF3ED56FFE1FF87F.taxon	description	Description. ♀ (Fig. 180 I). Apterous; body length 3.1 – 4.0 mm. Completely testaceous-reddish, or reddish-dark, or brown-reddish, except petiole black; occasionally metasoma brownish. Antenna clavate; antennomeres of holotype in following proportions: 20: 14: 36: 22: 17: 15: 14: 13: 12: 19. Head (Fig. 180 C) flat, dull, setose, granulate; frontal line incomplete, absent in anterior half of frons; occipital carina incomplete, only very shortly present on sides of lateral ocelli; POL = 4; OL = 6; OOL = 24. Palpal formula 5 / 3. Pronotum dull, setose, granulate, with numerous slight longitudinal keels, crossed by strong transverse impression. Mesoscutum dull, granulate and with some longitudinal keels, without lateral pointed apophyses. Metanotum dull, transversely striate, weakly excavated behind mesoscutellum (Fig. 180 D), with sides protruding; lateral protrusions of metanotum more or less pointed (Fig. 180 I). Metapectal-propodeal disc dull, granulate, sculptured by many longitudinal keels; propodeal declivity, mesopleuron and metapleuron not or slightly transversely striate. In paratype from South Africa, Katberg, anterior surface of metapectal-propodeal disc rugose and without longitudinal keels. In specimen from South Africa, Umtamvuna Nature Reserve, propodeal declivity completely strongly transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 slightly shorter than 4 (14: 15). Enlarged claw (Fig. 179 D) with apex broad, one large subdistal tooth and one row of 5 – 9 lamellae. Protarsomere 5 (Fig. 179 D) with two rows of 12 – 17 lamellae (shortest row extending or not beyond 0.5 length of protarsomere); apex with approximately 10 – 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE142BF3FF3ED56FFE1FF87F.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, 18 – 30. XI- 1921, R. E. Turner leg. (NHMUK). Paratypes: SOUTH AFRICA: Eastern Cape, Katberg, 4000 ft., XII. 1932, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 1 ♀ (AMNH); KwaZulu-Natal, Drakensberg, Van Reenen, 23 – 26. I. 1927, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, same locality label as holotype, 1 – 2. I. 1922, 1 ♀ (NHMUK). Other material: SOUTH AFRICA: KwaZulu-Natal, Cathedral Peak area, 1 ♀ (AEIC); KwaZulu-Natal, Umtamvuna Na- ture Reserve, 1 ♀ (SAMC); KwaZulu-Natal, Louwsberg, Gwala Gwala, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 2 – 3. VI. 2005, YPT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Louwsberg, Sanyati Farm, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 18 – 20. IV. 2007, YPT, M. Mostovski leg., 2 ♀♀ (NMSA, MOLC); KwaZulu-Natal, Vernon Crookes Nature Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Royal Natal National Park, Mahai Camp, 28 ° 41.27 ’ S 28 ° 56.86 ’ E, 1450 m, 25. XI – 12. XII. 2005, MT, M. Mostovski leg., 1 ♀ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE142BF3FF3ED56FFE1FF87F.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE142BF3FF3ED56FFE1FF87F.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE132BF5FF3ED6D7FA77FA1E.taxon	description	(Figs 180 E, F, 181, 182 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE132BF5FF3ED6D7FA77FA1E.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 5 / 3; mesosoma black; pronotum crossed by strong trans- verse furrow; mesoscutum with two strong lateral pointed apophyses; meso-metapleural suture distinct and com- plete; metanotum without lateral protrusions; metapectal-propodeal disc fusiform, without strong median longitudi- nal furrow; enlarged claw (Fig. 182 A) with one large subdistal tooth, one row of lamellae, apex tapering.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE132BF5FF3ED6D7FA77FA1E.taxon	description	Description. ♀ (Fig. 181). Apterous; body length 3.7 mm. Head black, except mandible, lateral surfaces of clypeus and part of malar space (short stripe along orbits) testaceous; antenna brown, except antennomeres 7 – 10 testaceous; mesosoma black; metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 10: 6: 15: 10: 8: 7: 5: 5: 5: 8. Head excavated, shiny, unsculptured, partly slightly granulate; frontal line complete; oc- cipital carina absent; POL = 1; OL = 2; OOL = 9. Palpal formula 5 / 3. Pronotum shiny, unsculptured, crossed by strong transverse furrow. Mesoscutum with two lateral pointed apophyses, situated on sides of anterior margin of mesoscutellum (Fig. 180 E). Mesoscutellum sloping anteriorly. Metanotum short, excavated behind mesoscutellum (Fig. 180 F), with sides rounded (Fig. 180 E). Metapectal-propodeal disc fusiform, shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete, step shaped. Protarsomeres in following proportions: 15: 3: 5: 15: 26. Enlarged claw (Fig. 182 A) with one large subdistal tooth and one row of seven lamellae and one bristle. Protarsomere 5 (Fig. 182 A) with two rows of 9 + 13 lamellae; apex with about 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE132BF5FF3ED6D7FA77FA1E.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.182 ’ S 19 ° 08.587 ’ E, 741 m, 27. VIII – 27. X. 2009, MT, Nieuwoudtville Shale Renosterveld, S. van Noort leg., SAM-HYM-A 022635, 1 ♀ (SAMC). Hosts. Unknown. Distribution. South Africa. Etymology. The species is named hantamensis after the collection site in Hantam National Botanical Garden.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE122BF6FF3ED3A5FD81FD74.taxon	description	(Figs 180 G, 182 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE122BF6FF3ED3A5FD81FD74.taxon	description	Description. ♀. Apterous; body length 4.2 – 6.0 mm. Head brown, except mandible, clypeus and most part of frons testaceous (occasionally anterior half of frons testaceous); antenna brown, except antennomeres 9 – 10 yellow-whitish (in specimen from Mayotte, antenna brown, except scape and flagellomere 8 testaceous); mesosoma black, except lateral margins of pronotum brown-testaceous (in specimen from Mayotte, mesosoma totally black); metasoma brown; legs brown, except tarsi testaceous (in specimen from Mayotte, legs brown, except articulations and part of trochanters whitish, part of meso- and metafemur whitish). Antenna clavate; antennomeres of holotype in following proportions: 13: 7: 32: 16: 13: 10: 10: 9: 8: 14. Head excavated, bare, shiny, finely punctate, unsculptured among punctures; frontal line complete; occipital carina absent; temple distinct; POL = 2.5; OL = 2.5; OOL = 12. Palpal formula 5 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, sculptured by many longitudinal keels, with two lateral pointed apophyses situated on sides of margin between mesoscutum and mesoscutellum (Fig. 180 G). Mesoscutellum sloping anteriorly, shiny, unsculptured. Metanotum sloping anteriorly, shiny, unsculptured, weakly excavated behind mesoscutellum, with two lateral pointed apophyses (Fig. 180 G); lateral regions of metanotum transversely striate. Metapectal-propodeal disc shiny, strongly transversely striate, except anterior surface unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron strongly transversely striate. Meso-metapleural suture obsolete, slightly visible only in posterior half. Protarsomeres in following proportions: 20: 4: 8: 20: 33. Protarsomere 3 produced into hook. Enlarged claw (Fig. 182 B) with one large subdistal tooth and one row of seven lamellae. Protarsomere 5 (Fig. 182 B) with two rows of 4 + 15 - 17 lamellae; apex with at least 16 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE122BF6FF3ED3A5FD81FD74.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Foulpointe (= Mahavelona), XI. 1995, forêt / sable, A. Pauly leg. (FSAE). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vatoharanana River, 4.1 km 231 ° SW Ranomafana, 21 ° 17.24 ’ S 47 ° 26.00 ’ E, 1100 m, YPT, montane rainforest, 27 – 31. III. 2003, Fisher, Griswold et al. leg., BLF 8399, 1 ♀ (MOLC); Toamasina, Périnet, 27. IV – 3. V. 1983, J. S. Noyes & M. C. Day leg., 1 ♀ (NHMUK). MAYOTTE: Dapani, 12 ° 57 ’ 46 ’’ S 45 ° 09 ’ 01 ’’ E, 135 m, 2 – 4. XII. 2007, sifted litter, rainforest, B. L. Fisher et al. leg., BLF 19060, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE122BF6FF3ED3A5FD81FD74.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE122BF6FF3ED3A5FD81FD74.taxon	distribution	Distribution. Madagascar. Mayotte.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE112BF7FF3ED574FE14FDB8.taxon	description	(Figs 178 B, C, 182 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE112BF7FF3ED574FE14FDB8.taxon	description	Description. ♀ (Figs 178 B, C). Apterous; body length 3.4 – 3.5 mm. Testaceous, except petiole black; antenna of specimen from Madagascar, 15 ° 11.18 ’ S 49 ° 36.54 ’ E, brown, except scape testaceous and flagellomeres 6 – 8 whitish; antenna of holotype testaceous, with last two antennomeres whitish; antenna of specimen from Madagascar, 18 ° 51 ’ 29 ’’ S 48 ° 17 ’ 06 ’’ E, brown, except antennomere 10 whitish. Antenna clavate; ADOs in antennomeres 6 – 10; antennomeres of specimen from Madagascar, 15 ° 11.18 ’ S 49 ° 36.54 ’ E, in following proportions: 10: 5: 15: 7: 6: 5.5: 5.5: 5.5: 5: 10; antennomeres of holotype in following proportions: 10: 4: 12: 5.5: 5: 5: 5: 5: 5: 9. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; temple distinct; head of specimen from Madagascar, 15 ° 11.18 ’ S 49 ° 36.54 ’ E, with POL = 0.5; OL = 1.5; OOL = 7; head of holotype with POL = 1; OL = 2; OOL = 6. Palpal formula 5 / 3 or 6 / 3 (5 / 3 in holotype; 6 / 3 in specimen from Madagascar, 7 km W Ranomafana). Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, unsculptured, with two large lateral pointed apophyses (Fig. 178 B). Mesoscutum very slender, about twice as long as broad (9: 4.5: length calculated from anterior margin to imaginary line joining two lateral pointed apophyses of mesoscutum). Mesoscutellum shiny, smooth, flat. Metanotum about coplanar to mesoscutellum (Fig. 178 C), flat, almost completely unsculptured, with or without few short keels. Metanotum with two large lateral pointed protrusions (Fig. 178 B). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. In specimens from Madagascar, 16 ° 46.22 ’ S 49 ° 15.56 ’ E, metapectal-propodeal disc slightly transversely striate. Mesopleuron and metapleuron shiny, unsculptured. Mesometapleural suture distinct and complete. In specimen from Madagascar, 15 ° 11.18 ’ S 49 ° 36.54 ’ E, Protarsomeres in following proportions: 13: 2.5: 5: 15: 25; Protarsomeres of holotype in following proportions: 10: 2: 4: 12: 20. Enlarged claw (Fig. 182 C) with one large subdistal tooth and one row of 4 – 6 lamellae + 1 bristle. Protarsomere 5 (Fig. 182 C) with two rows of 11 – 15 lamellae; apex with approximately 10 – 11 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE112BF7FF3ED574FE14FDB8.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohipa- rara area, 21 ° 24.032 ’ S 47 ° 39.399 ’ E, 1150 m, Pitfall trap, mixed tropical forest, 8 – 19. I. 2001, D. H. & K. M. Kavanaugh, R. L. Brett, E. Elsom & F. Vargas leg. (CAS). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vatoharanana River, 4.1 km 231 ° SW Ranomafana, 21 ° 17.24 ’ S 47 ° 26.00 ’ E, 1100 m, sifted litter, montane rainforest, 27 – 31. III. 2003, Fisher, Griswold et al. leg., BLF 8400, 3 ♀♀ (2 in CAS, 1 in MOLC); Fianrant- soa, 7 km W Ranomafana, 900 m, 20 – 31. I. 1990, flight intercept-yellow pan trap in Malaise trap in small clearing montante rainforest, W. E. Steiner leg., 1 ♀ (USNM); Fianarantsoa, 38 km S Ambalavao, 22 ° 12 ’ S 46 ° 58 ’ E, 1680 m, 23. X. 1993, B. L. Fisher leg., 2 ♀♀ (DEUCD, MOLC); Toamasina, Montagne d’Anianaharibe, 18.0 km 21 ° NNE Ambinanitelo, 15 ° 11.18 ’ S 49 ° 36.54 ’ E, 470 m, 8 – 12. III. 2003, sifted litter (leaf mold, rotten wood) in rainforest, Fisher, Griswold et al. leg., BLF 8002, 1 ♀ (CAS); Toamasina, 11 km WSW Befingotra, Anjanaharibe-sud Reserve, 14 ° 45 ’ S 49 ° 27 ’ E, 1565 m, 16. XI. 1994, montane rainforest, sifted litter (leaf mold, rotten wood), B. L. Fisher leg., 2 ♀♀ (DEUCD); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18 ° 51 ’ 29 ’’ S 48 ° 17 ’ 06 ’’ E, 1040 m, 5 – 8. III. 2007, pitfall trap, B. L. Fisher et al. leg., BLF 16919, 1 ♀ (CAS); Toamasina, Ambatovaky Special Reserve, Sandrangato River, 16 ° 46.22 ’ S 49 ° 15.56 ’ E, 450 m, 20 – 22. II. 2010, sifted litter, rainforest, B. L. Fisher et al. leg., 1 ♀ (MOLC); Toliara, 9.2 km N Tolanaro, Ilapiry Mt., 24 ° 51.27 ’ S 47 ° 00.38 ’ E, 400 m, 31. I. 1995, K. C. Emberton et al. leg., rainforest, sifted litter (leaf mold, rotten wood), 1 ♀ (DEUCD); Toliara, 13 km W Enakara, Andohahela Reserve, 24 ° 33 ’ S 46 ° 48 ’ E, 1250 m, 30. XI. 1992, montane rainforest, sifted litter (leaf mold, rotten wood), B. L. Fisher leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE112BF7FF3ED574FE14FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE112BF7FF3ED574FE14FDB8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102BF7FF3ED4DFFE3FFA39.taxon	description	(Figs 180 H, 182 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102BF7FF3ED4DFFE3FFA39.taxon	description	Description. ♀. Apterous; body length 3.9 mm. Completely ferruginous, except metasoma partly brown. Antenna clavate; antennomeres in following proportions: 10: 6: 19: 10: 9: 7: 6: 5: 5: 8. Head excavated, dull, granulate; frontal line absent; frons with short narrow longitudinal furrow situated in front of anterior ocellus and directed towards antennal toruli, reaching approximately mid length of frons; occipital carina absent; temple distinct; occiput dull, granulate; POL = 1.5; OL = 3; OOL = 11. Palpal formula 5 / 3. Pronotum shiny, sculptured by numerous longitudinal striae, crossed by strong transverse impression. Mesoscutum shiny, sculptured by many longitudinal striae, without lateral pointed apophyses. Mesoscutellum sloping anteriorly. Metanotum sculptured by short longitudinal keels, deeply excavated behind mesoscutellum (Fig. 180 H); sides of metanotum protruding; lateral protrusions of metanotum rounded. Metapectal-propodeal disc dull, sculptured by many longitudinal striae; top of metapectal-propodeal disc with strong median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 17: 4: 6: 16: 28. Enlarged claw (Fig. 182 D) with one large subdistal tooth and one row of seven lamellae and one bristle. Protarsomere 5 (Fig. 182 D) with two rows of approximately 13 lamellae; apex with approximately 18 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102BF7FF3ED4DFFE3FFA39.taxon	materials_examined	Material examined. Type: ♀ holotype: BOTSWANA: Central District, 30 km W of Palapye, 10. IV. 1997, by sweeping, M. Olmi leg. (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102BF7FF3ED4DFFE3FFA39.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102BF7FF3ED4DFFE3FFA39.taxon	distribution	Distribution. Botswana.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102B88FF3ED05EFE14FB87.taxon	description	(Figs 182 E, 186 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102B88FF3ED05EFE14FB87.taxon	description	Description. ♀ (Fig. 186 A). Apterous; body length 2.7 – 3.2 mm. Colour of holotype: head brown, except mandible, clypeus and anterior half of frons testaceous; antenna brown, except antennomere 1 testaceous (antennomeres 9 – 10 missing in holotype); mesosoma brown-black; metasoma brown, except posterior third whitish; legs brown, except part of coxae, part of trochanters, part of meso- and metafemur testaceous. Colour of paratypes from Madagascar, 12 ° 51.49 ’ S 49 ° 13.33 ’ E and 12 ° 54.32 ’ S 49 ° 6.35 ’ E: totally testaceous-reddish, except petiole black and antennomeres 3 – 8 darkened; antennomeres 9 – 10 yellow-whitish. Antenna clavate; antennomeres of holotype in following proportions: 9: 5: 13: 6.5: 5: 5: 5: 5 (antennomeres 9 and 10 missing in holotype); antennomeres of paratype from Madagascar, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, in following proportions: 9: 6: 14: 7: 6: 6: 5: 5: 5: 9. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; temple distinct; ocellar triangle of holotype with POL = 1.5; OL = 2; OOL = 8; ocellar triangle of paratype from Madagascar, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, with POL = 1; OL = 1; OOL = 7. Palpal formula 5 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, unsculptured, with two large lateral pointed apophyses (Fig. 186 A). Mesoscutum as long as broad (length calculated from anterior margin to imaginary line joining two lateral pointed apophyses of mesoscutum). Mesoscutellum shiny, smooth, flat. Metanotum about coplanar to mesoscutellum, flat, partly smooth and partly rugose; metanotum with two large lateral rounded protrusions (Fig. 186 A). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron shiny, unsculptured. Meso-metapleural suture distinct and complete. Protarsomeres of holotype in following proportions: 12.5: 2.5: 5: 14: 24; Protarsomeres of paratype from Madagascar, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, in following proportions: 12: 2.5: 6: 14: 23. Enlarged claw (Fig. 182 E) with one large subdistal tooth and one row of 5 – 6 lamellae and one bristle. Protarsomere 5 (Fig. 182 E) with two rows of 12 – 14 lamellae; apex with approximately 10 lamellae. In paratype from Madagascar, 12 ° 54.32 ’ S 49 ° 6.35 ’ E, protarsomere 5 with two rows of nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102B88FF3ED05EFE14FB87.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toamasina, F. C. Sandranantitra, 18 ° 02.9 ’ S 49 ° 05.5 ’ E, 450 m, sifted litter (leaf mold, rotten wood), rainforest, 18 – 21. I. 1999, H. J. Ratsirarson leg. (CAS). Paratypes: MADAGASCAR: Antsiranana, Ankarana Special Reserve, 13.6 km 192 ° SSW of Anivorano North, 12 ° 51.49 ’ S 49 ° 13.33 ’ E, 210 m, 16 – 20. II. 2001, sifted litter tropical dry forest, Fisher, Griswold et al. leg., BLF 3012, 1 ♀ (MOLC); Antsiranana, Ankarana Special Reserve, 22.9 km 224 ° SW of Anivorano North, 12 ° 54.32 ’ S 49 ° 6.35 ’ E, 80 m, 10 – 16. II. 2001, sifted litter in tropical dry forest, Fisher, Griswold et al. leg., BLF 2858, 1 ♀ (CAS); Antsiranana, Ambre Special Reserve, 3.5 km 235 ° SW of Sakaramy, 12 ° 28.8 ’ S 49 ° 14.32 ’ E, 325 m, 26 – 31. I. 2001, sifted litter in tropical dry forest, Fisher, Griswold et al. leg., code BLF 2654, 1 ♀ (CAS); Toamasina, 5.3 km SSE Ambanizana, Andranobe Field Station, 15 ° 40 ’ S 49 ° 58 ’ E, 425 m, 21. XI. 1993, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102B88FF3ED05EFE14FB87.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE102B88FF3ED05EFE14FB87.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	description	(Figs 182 F, 183 A, B, G, 184)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 5 / 3; pronotum crossed by strong transverse furrow; mesoscutum without lateral pointed apophyses (Fig. 183 A); metanotum laterally rounded (Fig. 183 A); meso-metapleural suture distinct and complete; metapectal-propodeal disc without strong median longitudinal furrow; enlarged claw (Figs 182 F, 183 G) with apex tapering, one big subdistal tooth located very far from most distal lamella and one row of lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	description	Description. ♀ (Fig. 184). Apterous; body length 2.7 – 3.0 mm (holotype 2.7 mm). Head brown, except man- dible, clypeus, malar space and anterior region of frons testaceous (in paratypes from South Africa, 33 ° 43.745 ’ S 21 ° 56.972 ’ E and 33 ° 39.941 ’ S 21 ° 53.505 ’ E, head black, except mandible, part of clypeus and two short frontal stripes along orbits testaceous-whitish); antenna brown, except ventral side of scape and flagellomeres 6 – 8 whitish (in paratype from South Africa, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, antenna brown, except ventral side of scape and distal half of flagellomere 8 whitish; in paratype from South Africa, 33 ° 39.941 ’ S 21 ° 53.505 ’ E, antenna brown, except ventral side of scape and flagellomeres 7 – 8 whitish )); mesosoma and metasoma brown (( in paratype from South Africa, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, mesosoma black, except lateral regions and posterior margin of pronotum and great part of mesoscutum testaceous); legs brown. Antenna clavate; antennomeres in following proportions: 9: 4: 9: 4: 4: 4: 4: 4: 4: 7. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 7; greatest breadth of lateral ocelli longer than POL (2: 1). Palpal formula 5 / 3. Pronotum shiny, crossed by strong transverse impression, unsculptured. Mesoscutum dull, unsculptured, laterally without pointed apophyses (Fig. 183 A). Metanotum narrow, excavated behind mesoscutellum (Fig. 183 B), with sides protruding; lateral protrusions of metanotum rounded (Fig. 183 A). In paratypes metanotum little protruding, so that metapectalpropodeal disc almost fusiform. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity and metapleuron transversely striate. Mesopleuron unsculptured. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 9: 2: 4: 11: 19. Enlarged claw (Figs 182 F, 183 G) with one large subdistal tooth and one row of 6 – 13 lamellae (six in holotype). Protarsomere 5 (Figs 182 F, 183 G) with two rows of 15 – 21 lamellae (15 in holotype); apex very large (Figs 182 F, 183 G) and provided with at least 50 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Schilpad Laagte Farm, 14.7 km 229 ° SW Kirkwood, 33 ° 31.104 ’ S 25 ° 22.353 ’ E, 9 – 16. II. 2001, H. G. Robertson & R. Tourle leg., Winkler, VB 01 - A 2 TN-W 04, Valley Bushveld (non-trashed), SAM-HYM-A 022727 (SAMC). Paratypes: SOUTH AFRICA: Western Cape, Gamkaberg Nature Reserve, 33 ° 43.745 ’ S 21 ° 56.972 ’ E, 1000 m, 19. II – 30. III. 2010, MT, Renosterveld, S. van Noort leg., 1 ♀ (MOLC); Western Cape, Gamkaberg Nature Reserve, 33 ° 39.941 ’ S 21 ° 53.505 ’ E, 315 m, 23. XII. 2008 – 16. I. 2009, MT, Gamka Thicket, S. van Noort leg., SAM-HYM-A 023311, 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 43.663 ’ S 21 ° 57.600 ’ E, 940 m, 31. VII – 10. IX. 2009, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., SAM-HYM-P 039007, 1 ♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, 997 m, 4. X. 2010 – 25. I. 2011, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., GB 09 - FYN 2 - M 53, SAM-HYM-A 026532, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6F2B8AFF3ED20BFAACFD00.taxon	etymology	Etymology. The species is named after one of the collectors of the holotype, Dr. Hamish G. Robertson.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8AFF3ED497FE1FFA5B.taxon	description	(Fig. 183 H)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8AFF3ED497FE1FFA5B.taxon	description	Description. ♀. Apterous; body length 3.4 mm. Completely brown-reddish, except petiole dark and antenna and tarsi testaceous. Antenna clavate; antennomeres in following proportions: 20: 12: 24: 15: 14: 13: 12: 12: 12: 17. Head excavated, shiny, bare, unsculptured; frontal line complete; occipital carina incomplete, only shortly visible on sides of lateral ocelli; occiput setose; POL = 3; OL = 5; OOL = 18. Palpal formula 5 / 3. Pronotum shiny, bare, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, with some longitudinal keels, with two pointed lateral apophyses situated on sides of mesoscutellum. Metanotum smooth, not transversely striate, excavated behind mesoscutellum, with sides rounded, not protruding. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsal segmens in following proportions: 14: 2: 4: 11: 17. Enlarged claw (Fig. 183 H) with one large subdistal tooth and one row of five lamellae. Protarsomere 5 (Fig. 183 H) with two rows of approximately 13 lamellae; apex with approximately 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8AFF3ED497FE1FFA5B.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Milnerton, I. 1926, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8AFF3ED497FE1FFA5B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8AFF3ED497FE1FFA5B.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8BFF3ED060FE14FD00.taxon	description	(Figs 186 B, C, 187 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8BFF3ED060FE14FD00.taxon	description	Description. ♀ (Figs 186 B, C). Apterous; body length 4.9 mm. Head brown, except mandible, clypeus and anterior region of frons (with stripes along orbits) testaceous; antenna brown, except ventral side of scape, flagellomere 8, ventral side of flagellomeres 4 – 9 testaceous-whitish; mesosoma black, except lateral margins of pronotum testaceous; metasoma brown, with testaceous spots; legs brown, except meso- and metatrochanter, part of coxae, part of protrochanter, part of meso- and metafemur testaceous-whitish; tarsi testaceous. Antenna clavate; ADOs present in antennomeres 6 – 10; antennomeres in following proportions: 12: 6: 19: 8: 7: 6: 6: 5: 5: 10. Head excavated, dull, with occiput unsculptured and frons sculptured by many longitudinal irregular striae; vertex with many transverse irregular striae situated on sides of ocelli; frontal line complete; occipital carina absent; temple distinct; POL = 1.5; OL = 2; OOL = 10. Palpal formula 5 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, sculptured by many longitudinal striae, with two large lateral pointed apophyses (Fig. 186 B). Mesoscutum as long as broad (length calculated from anterior margin to imaginary line joining two lateral pointed apophyses of mesoscutum). Mesoscutellum shiny, smooth, flat. Metanotum about coplanar to mesoscutellum (Fig. 186 C), flat, unsculptured, with two large lateral rounded protrusions (Fig. 186 B). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron shiny, unsculptured. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 15: 3.5: 5: 21: 34. Enlarged claw (Fig. 187 A) with one large subdistal tooth and one row of six lamellae; Protarsomere 5 (Fig. 187 A) with two rows of 4 + 16 lamellae; apex with at least 40 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8BFF3ED060FE14FD00.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Analamay, 18 ° 48.22 ’ S 48 ° 20.13 ’ E, 1068 m, 21. III. 2004, sifted litter (leaf mold, rotten wood), montane rainforest, Malagasy ant tem, BLF 10502 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8BFF3ED060FE14FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6D2B8BFF3ED060FE14FD00.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6C2B8DFF3ED497FF35FC04.taxon	diagnosis	(Figs 183 C, D, 185, 187 B) Diagnosis. Apterous ♀ of Gonatopus with palpal formula 5 / 3; pronotum crossed by strong transverse furrow; me- soscutum without two lateral pointed apophyses; meso-metapleural suture distinct and complete; metapectal-propo- deal disc without strong median longitudinal furrow; enlarged claw (Fig. 187 B with apex tapering, one big subdistal tooth and one row of lamellae (subdistal tooth located very close to more distal lamella). Description. ♀ (Fig. 185). Apterous; body length 2.8 mm. Head brown, except mandible and clypeus testa- ceous; antenna brown; mesosoma, metasoma and legs brown. Antenna clavate; antennomeres in following proportions: 7: 5: 15: 10: 8: 6: 6: 5: 5: 7. Head excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 2; OL = 2; OOL = 10; greatest breadth of lateral ocelli about as long as OL. Palpal formula 5 / 3. Pronotum shiny, crossed by strong transverse impression, unsculptured, except many longitudinal striae on disc and lateral regions. Mesoscutum dull, sculptured by many longitudinal striae, laterally without two pointed apophyses (Fig. 183 C). Metanotum rugose, deeply excavated behind mesoscutellum (Fig. 183 D), with sides protruding; lateral protrusions slightly pointed (Fig. 183 C). Metapectal-propodeal disc dull, sculptured by many slight longitudinal striae; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 3: 6: 14: 23. Enlarged claw (Fig. 187 B with one big subdistal tooth and one row of seven lamellae (subdistal tooth located very close to more distal lamella). Protarsomere 5 (Fig. 187 B) with two rows of 2 + 6 lamellae; apex with at least 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6C2B8DFF3ED497FF35FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 27. X. 2009 – 9. II. 2010, MT, Hantam Succulent Karoo, S. van Noort leg., SAM-HYM-A 035765 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6C2B8DFF3ED497FF35FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6C2B8DFF3ED497FF35FC04.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6C2B8DFF3ED497FF35FC04.taxon	etymology	Etymology. The species is named after Prof. Clarke Scholtz (University of Pretoria), beetle specialist, on whose farm, Kelkiewyn, one of the authors (SVN) collected the specimen. He kindly allowed SVN to run traps on the farm.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6A2B8DFF3ED59BFDB4F85E.taxon	description	(Figs 183 E, F, 187 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6A2B8DFF3ED59BFDB4F85E.taxon	description	Description. ♀. Apterous; body length 3.0 mm. Head black, except mandible, malar space and anterior stripes on frons along orbits and near clypeus testaceous; antenna brown, except scape and flagellomeres 6 – 8 testaceous; mesosoma black or brown-black, except pronotum brown, posterior half of mesoscutellum occasionally yellow, metanotum brown, apex of propodeal declivity yellow; metasoma black; proleg testaceous, except part of coxa, club of femur and tibia brown; meso- - and metaleg testaceous, except part of coxae and clubs of femora brown. Antenna clavate; antennomeres in following proportions: 7: 5: 15: 7: 7: 5: 6: 4: 4.5: 8. Head excavated, dull, slightly or strongly granulate; frontal line incomplete, absent near clypeus; occipital carina absent; POL = 1; OL = 2; OOL = 8. Palpal formula 5 / 3. Pronotum shiny, crossed by strong transverse impression, unsculptured, except some longitudinal striae on disc. In specimen from South Africa, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, pronotum dull, sculptured by many longitudinal striae. Mesoscutum dull, sculptured by many longitudinal striae, laterally without strong pointed apophyses, with three small lateral prominences corresponding to transverse carinae present on lateral regions of mesoscutum (Fig. 183 E). Metanotum rugose, excavated behind mesoscutellum (Fig. 183 F), with sides protruding; lateral protrusions of metanotum pointed (Fig. 183 E). Metapectal-propodeal disc dull, sculptured by many slight longitudinal striae; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 15: 2.5: 6: 15: 23. Enlarged claw (Fig. 187 C) with one large subdistal tooth and one row of 6 – 10 lamellae. Protarsomere 5 (Fig. 187 C) with two rows of 11 – 19 lamellae; apex with about 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6A2B8DFF3ED59BFDB4F85E.taxon	materials_examined	Material examined. Type: ♀ holotype: GHANA: Eastern Region, NW Juapong, 9 km E Akosombo, 06 ° 16 ’ N 00 ° 07 ’ E, 31. VIII. 2012, M. & J. Schwarz leg. (OLL). Other material: SOUTH AFRICA: Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 28. IX – 28. XII. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., DRA 09 - FYN 1 - Y 10, 2 ♀♀ (SAMC, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6A2B8DFF3ED59BFDB4F85E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE6A2B8DFF3ED59BFDB4F85E.taxon	distribution	Distribution. Ghana, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	description	(Figs 186 D, 188, 189 A, 191 A; 192 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; pronotum crossed by strong transverse furrow; metanotum about coplanar to mesoscutellum (Fig. 192 B); metapectal-propodeal disc brown-black, with disc slightly humped (Fig. 189 A); enlarged claw with apex very broad (Fig. 191 A) and one big subdistal tooth; protarsomere 5 with one short row of lamellae (Fig. 191 A).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	description	Description. ♀ (Figs 186 D, 188, 189 A). Apterous; body length 3.7 mm. Head brown-black, except mandible testaceous and one ferruginous longitudinal stripe between antennal toruli; antenna brown (antennomere 10 missing); mesosoma brown-black, except mesoscutum testaceous; metasoma brown; legs brown, except tarsi, part of procoxa and protrochanter testaceous. Antenna clavate; antennomeres in following proportions: 11: 6: 25: 16: 13: 12: 9: 8: 7 (antennomere 10 missing in holotype). Head (Fig. 188 D) excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 1; OL = 2.5; OOL = 9; greatest breadth of lateral ocelli longer than POL (2: 1). Palpal for-mula 6 / 3. Pronotum shiny, finely punctate, unsculptured among punctures, crossed by strong transverse furrow. Mesoscutum shiny, unsculptured, with two strong lateral pointed apophyses (Figs 188 B, 192 A). Mesoscutellum shiny, flat. Metanotum about coplanar to mesoscutellum (Figs 188 C, 192 B). Metapectal-propodeal disc swollen, shiny, finely punctate, unsculptured among punctures; propodeal declivity transversely striate. Mesopleuron shiny, unsculptured. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 19: 3: 7: 20: 34. Enlarged claw (Fig. 191 A) with apex very broad, one big subdistal tooth and one row of eight lamellae + two bristles. Protarsomere 5 (Fig. 191 A) with one row of 10 lamellae situated in distal half of protarsomere 5; apex with at least 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19451): MADAGASCAR: Antsiranana, Marojejy National Park, Antranohofa, 26.6 km 31 ° NNE Andapa, 10.7 km 318 ° NW Manantenina, 14 ° 26.36 ’ S 49 ° 44.36 ’ E, 1325 m, 18. XI. 2003, on low vegetation, montane rainforest, B. L. Fisher leg. (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE692B8FFF3ED056FA71FD2C.taxon	etymology	Etymology. The species is named bellicosus (Latin adjective; = aggressive), because of its aggressive aspect.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE682B80FF3ED15CFE1FFC4C.taxon	description	(Figs 191 B, 192 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE682B80FF3ED15CFE1FFC4C.taxon	description	Description. ♀. Apterous; body length 3.0 – 3.4 mm. Head black, except mandible testaceous; clypeus testaceous, except darkened central area; antenna brown, except ventral side of scape whitish (in specimen from South Africa, Kogelberg Nature Reserve, antenna brown, except ventral side of scape and distal half of flagellomere 8 whitish); propleuron black; pronotum black, except lateral regions and posterior margin of disc testaceous-reddish; mesoscutum testaceous-reddish, except anterior third brown; rest of mesosoma black, except posterior extremity of propodeal declivity testaceous-reddish; metasoma black, except apex brown; legs brown. Antenna clavate; antennomeres in following proportions: 8: 5: 11: 5: 5: 5: 4: 5: 4.5: 7. Head excavated, shiny, granulate; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 8. Palpal formula 6 / 3. Pronotum finely punctate, unsculptured among punctures or very slightly granulate, crossed by strong transverse impression. Mesoscutum without lateral pointed apophyses (Fig. 192 C). Mesoscutellum strongly sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 192 D); sides of metanotum rounded. Metapectal-propodeal disc shiny and unsculptured; metapleuron and propodeal declivity transversely striate; mesopleuron granulate and not transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 10: 3: 6: 14: 25. Enlarged claw (Fig. 191 B) with one large subdistal tooth and one row of 7 – 8 lamellae in addition to one bristle. Protarsomere 5 (Fig. 191 B) with two rows of 14 – 29 lamellae; apex very broad and with at least 25 – 50 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE682B80FF3ED15CFE1FFC4C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, De Hoop Nature Reserve, 34 ° 27 ’ S 20 ° 25 ’ E, loc. 12, 10 – 13. X. 1994, R. Danielsson leg. (ZIL). Other material: SOUTH AFRICA: Northern Cape, Avontuur Farm, 16 km NW Nieuwoudtville, 31 ° 16.249 ’ S 19 ° 02.900 ’ E, 764 m, 27. VIII – 27. X. 2009, YPT, Bokkeveld Sandstone Fynbos, S. van Noort leg., 1 ♀ (SAMC); Western Cape, Kogelberg Nature Reserve, C. 17. I. 2003, B. 29. I. 2003, M. Olmi leg., reared from a nymph of an unidentified brachypterous species of Tropiduchidae feeding on Berzelia lanuginosa (Bruniaceae) in fynbos, M. Olmi reared, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE682B80FF3ED15CFE1FFC4C.taxon	biology_ecology	Hosts. In South Africa: unidentified species of Tropiduchidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE682B80FF3ED15CFE1FFC4C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE672B80FF3ED253FD91F832.taxon	description	(Figs 191 C, 192 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE672B80FF3ED253FD91F832.taxon	description	Description. ♀. Apterous; body length 2.6 – 4.1 mm. Completely yellow-testaceous, except petiole black. Antenna clavate; antennomeres in following proportions: 7: 5: 12: 7: 5: 5: 4.5: 4.5: 4.5: 6.5. Head excavated, shiny, with frons almost completely unsculptured (granulate only near clypeus) and occiput granulate; frontal line complete; occipital carina absent; temple distinct; POL = 1; OL = 1; OOL = 7; greatest breadth of lateral ocelli about as long as OL. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, unsculptured, without lateral pointed apophyses. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum deeply excavated behind mesoscutellum (Fig. 192 E), flat, strongly transversely striate. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Metapectal-propodeal disc laterally rounded. Mesopleuron shiny, very slightly granulate. Metapleuron dull, very slightly transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 12: 2: 4.5: 11: 18. Enlarged claw (Fig. 191 C) with one large subdistal tooth and one row of 6 – 8 lamellae + one subdistal bristle. Protarsomere 5 of holotype (Fig. 191 C) with two rows of 2 + 8 lamellae; apex with one lamella. Protarsomere 5 of specimen from Mozambique, Cuamba, with two rows of 3 + 12 lamellae; apex with at least 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE672B80FF3ED253FD91F832.taxon	materials_examined	Material examined. Type: ♀ holotype: ETHIOPIA: Shewa (= Shoa) Province, 95 km W Addis Abeba, Ambò (= Hagere Hiywot), near Plant Protection Research Institute, 2400 – 2500 m, 31. V. 1990, A. Emelyanov leg. (ZISP). Other material: MOZAMBIQUE: Niassa, Cuamba, Farm of the Catholic University of Mozambique, C. 11. III. 2005, B. 12.3.2005, Sf. 2. IV. 2005, M. Olmi reared from a nymph of an unidentified species of Dictyopharidae, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE672B80FF3ED253FD91F832.taxon	biology_ecology	Hosts. In Mozambique: unidentified species of Dictyopharidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE672B80FF3ED253FD91F832.taxon	distribution	Distribution. Ethiopia, Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE662B82FF3ED17AFDE2FD00.taxon	description	(Figs 191 E, 192 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE662B82FF3ED17AFDE2FD00.taxon	description	Description. ♀. Apterous; body length 4.2 mm. Completely ferruginous, except petiole black and distal half of metasoma darkened. Antenna clavate; antennomeres in following proportions: 11: 6: 17: 11: 8: 7: 6: 5: 5: 8. Head (Fig. 192 F) flat, dull, granulate; frontal line complete; occipital carina incomplete, only present behind and shortly on sides of lateral ocelli; temple distinct; POL = 2; OL = 2; OOL = 11. Palpal formula 6 / 3. Pronotum shiny, sculptured by many longitudinal striae, crossed by strong transverse impression. Metanotum excavated behind mesoscutellum, with sides protruding; lateral protrusions of metanotum pointed. Metapectal-propodeal disc sculptured by irregular longitudinal striae; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 17: 3.5: 5.5: 16: 28. Enlarged claw (Fig. 191 E) with one large subdistal tooth and one row of seven lamellae. Protarsomere 5 (Fig. 191 E) with two rows of 4 + 12 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE662B82FF3ED17AFDE2FD00.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Gaza Prov., Praia do Bilene, Airport, 16. IX. 1995, sweeping on grass, M. Olmi leg. (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE662B82FF3ED17AFDE2FD00.taxon	biology_ecology	Hosts. Flatidae (Guglielmino et al. 2013): in Mozambique: Paroxychara cowla Medler.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE662B82FF3ED17AFDE2FD00.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B82FF3ED497FE32F9CA.taxon	description	(Fig. 191 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B82FF3ED497FE32F9CA.taxon	description	Description. ♀. Apterous; body length 2.9 – 3.4 mm. Holotype completely testaceous, except petiole black. Paratype testaceous, except petiole black and part of mesocoxa brown. Antenna clavate; antennomeres in following proportions: 9: 5.5: 15: 9: 8: 7: 5: 5: 4.5: 7. Head excavated, shiny, with frons almost completely unsculptured (granulate only near clypeus) and occiput granulate; frontal line complete; occipital carina absent; temple distinct; POL = 1; OL = 2; OOL = 8. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression, except for many longitudinal striae on disc and on part of lateral region. Mesoscutum shiny, granulate, without lateral pointed apophyses. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum weakly excavated behind mesoscutellum, flat, strongly irregularly striate. Metapectal-propodeal disc shiny, strongly sculptured by many longitudinal striae; propodeal declivity transversely striate; Metapectal-propodeal disc laterally protruding and forming a slightly pointed protrusion. Mesopleuron and metapleuron dull, transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 3.5: 5: 16: 26. Enlarged claw (Fig. 191 D) with one large subdistal tooth and one row of 8 – 9 lamellae + one bristle. Protarsomere 5 (Fig. 191 D) with two rows of 16 – 18 lamellae; apex with approximately 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B82FF3ED497FE32F9CA.taxon	materials_examined	Material examined. Types: ♀ holotype: TANZANIA: Mkomazi Game Reserve, Ibaya Camp, NW side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 24. IV. 1996, Sweep, mk 34, Acacia / Commiphora / Combretum bushland, SAM-HYM- PO 18157, S. van Noort leg. (SAMC). Paratype: Mkomazi Game Reserve, valley NW of Ibaya Camp, 03 ° 58 ’ S 37 ° 48 ’ E, 30. XI. 1995, riverine bushland, sweep, SAM-HYM-PO 14832, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B82FF3ED497FE32F9CA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B82FF3ED497FE32F9CA.taxon	distribution	Distribution. Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	description	(Figs 190, 192 G, H, 194 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; head excavated; pronotum crossed by strong transverse furrow; metapectal-propodeal disc and propodeal declivity totally ferruginous; metapectal-propodeal disc sculptured by irregular longitudinal keels; protarsomere 1 longer than 4; enlarged claw (Fig. 194 A) with one large subdistal tooth and one row of lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	description	Description. ♀ (Fig. 190). Apterous; body length 3.0 mm. Ferruginous, except petiole black and metasoma brown. Antenna clavate; antennomeres in following proportions: 11: 7: 14: 9: 7: 5: 5: 4: 4: 7. Head excavated, shiny, granulate, except two large unsculptured areas on sides of frontal line; frontal line complete; occipital carina absent; temple distinct; POL = 1; OL = 3; OOL = 9; greatest breadth of lateral ocelli shorter than OL (2: 3). Palpal formula 6 / 3. Pronotum shiny, with many slight longitudinal striae on disc and lateral regions; rest of surface unsculptured; pronotum crossed by strong transverse furrow. Mesoscutum shiny, sculptured with some longitudinal striae and with two strong, pointed apophyses laterally (Fig. 192 G). Mesoscutellum shiny, smooth, sloping strongly anteriorly. Metanotum deeply excavated behind mesoscutellum (Fig. 192 H), flat, strongly rugose, with sides protruding (lat- eral protrusions slightly pointed (Fig. 192 G )). Metapectal-propodeal disc shiny, strongly sculptured by numerous irregular longitudinal keels and partly rugose; propodeal declivity transversely striate. Mesopleuron partly granulate and partly transversely striate. Metapleuron dull, transversely striate. Meso-metapleural suture distinct and com- plete, step shaped. Protarsomeres in following proportions: 17: 3: 5: 14: 22. Enlarged claw (Fig. 194 A) with one large subdistal tooth and one row of five lamellae. Protarsomere 5 (Fig. 194 A) with two rows of 14 lamellae; apex with at least 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 27. X. 2009 – 9. II. 2010, MT, Hantam Succulent Karoo, S. van Noort leg., SAM-HYM-A 022287 (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE652B83FF3ED0EEFB84FCBC.taxon	etymology	Etymology. The species is named karooensis after the collection site in the Karoo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE622B86FF3ED6D7FAA6FA20.taxon	description	(Figs 189 B, C, 193, 194 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE622B86FF3ED6D7FAA6FA20.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; pronotum crossed by strong transverse furrow; meta- notum about coplanar to mesoscutellum (Fig. 189 C); metapectal-propodeal disc black, very humped (Fig. 189 C); enlarged claw (Fig. 194 B) with apex slender, one big subdistal tooth and one row of lamellae; protarsomere 5 with long rows of lamellae (Fig. 194 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE622B86FF3ED6D7FAA6FA20.taxon	description	Description. ♀ (Figs 189 B, C, 193). Apterous; body length 4.3 mm. Head brown-black, except mandible, clypeus, anterior half of frons and ventral side whitish; antenna testaceous-brown, except scape yellow; mesosoma black, except dorsal stripe on propleuron testaceous; metasoma brown; legs brown, except part of coxae, trochanters and part of femora whitish, tarsi testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 19: 9: 7: 5: 6: 5.5: 5.5: 9. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1.5; OOL = 10; greatest breadth of lateral ocelli longer than POL (2: 1). Palpal formula 6 / 3. Pronotum shiny, unsculp- tured, crossed by strong transverse furrow. Mesoscutum shiny, unsculptured, with two strong pointed apophyses laterally (Figs 189 B, 193 B). Mesoscutellum shiny, flat. Metanotum about coplanar to mesoscutellum (Fig. 189 C). Metapectal-propodeal disc very swollen (Fig. 189 C), shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron shiny, unsculptured. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 16: 3: 6: 22: 33. Enlarged claw (Fig. 194 B) with apex slender, one big subdis- tal tooth and one row of six lamellae + one bristle. Protarsomere 5 (Fig. 194 B) with two rows of 18 lamellae; apex with at least 30 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE622B86FF3ED6D7FAA6FA20.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19465): MADAGASCAR: Antsiranana, Marojejy National Park, Manantenina River, 27.6 km 327 ° NNW Manantenina, 14 ° 26.06 ’ S 49 ° 45.36 ’ E, 775 m, 15 – 18. XI. 2003, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher et al. leg., BLF 8872 (CAS). Hosts. Unknown. Distribution. Madagascar. Etymology. The species is named marojejyanus after the collection site in the Marojejy National Park.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE612B87FF3ED3B7FD73FCD8.taxon	description	(Figs 192 I, 194 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE612B87FF3ED3B7FD73FCD8.taxon	description	Description. ♀. Apterous; body length 2.9 – 3.9 mm. Head black, except mandible testaceous; antenna brown, except ventral side of scape testaceous; mesosoma and metasoma black; legs brown, except distal extremities of coxae, trochanters, stalks of femora, proximal half of tibiae and tarsi testaceous. In specimen from South Africa, Kruispad Farm, head black, except mandible and part of clypeus testaceous; antenna brown, except flagellomere 8 and ventral side of scape whitish; mesosoma black, except lateral regions of pronotum, posterior margin of disc of pronotum, part of mesoscutum and posterior extremity of propodeal declivity testaceous; metasoma brown; legs brown, except articulations testaceous. Antenna clavate; antennomeres in following proportions: 7: 6: 12: 7: 6.5: 6.5: 5.5: 5: 5.5: 8. Head excavated, shiny, with occiput granulate and frons unsculptured, except narrow and slightly rugose area near clypeus; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 9. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, sculptured by numerous longitudinal striae, with two strong lateral pointed apophyses (Fig. 192 I). Mesoscutellum shiny, sloping anteriorly. Metanotum deeply excavated behind mesoscutellum. In specimen from South Africa, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, metanotum weakly excavated behind mesoscutellum. Metapectal-propodeal disc swollen, shiny, unsculptured; metapleuron and propodeal declivity transversely striate; mesopleuron partly transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 13: 2.5: 4.5: 13.5: 22. Enlarged claw (Fig. 194 C) with one large subdistal tooth and one row of 8 – 9 lamellae. Protarsomere 5 (Fig. 194 C) with two rows of 16 – 24 lamellae; apex with 10 – 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE612B87FF3ED3B7FD73FCD8.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Free State, 9 km S of Harrismith, 300 m after Junc- tion R 74 – 712, along R 74 to Sterkfontein, 25. I. 1997, sweeping on grass, M. Olmi leg. (AMNH). Other material: MADAGASCAR: Mahajanga, Tsimembo Forest, 8.7 km 336 ° NNW Soatana, 19 ° 01.17 ’ S 44 ° 26.26 ’ E, 20 m, 21 – 25. XI. 2001, beating low vegetation, tropical dry forest, Fisher, Griswold et al. leg., BLF 4510, 1 ♀ (CAS). SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 6. X. 2010 – 18. I. 2011, YPT, Amathole Mistbelt Grassland, S. van Noort leg., MTB 09 - GRA 1 - Y 06, 1 ♀ (MOLC); Gauteng, near Roode- poort, Discovery, about 26 ° 09.5 ’ S 27 ° 53.5 ’ E, 28. IX. 1961, H. N. Empey leg., 1 ♀ (DJBC); Western Cape, 10 km SE of Velddrif, Kruispad Farm, 32 ° 50 ’ S 18 ° 15 ’ E, 10. XI. 1992, M. Picker leg., swept from Salicornia, SAM-HYM- PO 03778, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE612B87FF3ED3B7FD73FCD8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE612B87FF3ED3B7FD73FCD8.taxon	distribution	Distribution. Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B87FF3ED5FFFE1FF97B.taxon	description	(Figs 194 D, 195 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B87FF3ED5FFFE1FF97B.taxon	description	Description. ♀. Apterous; body length 4.8 mm. Head reddish-dark; antenna testaceous, except antennomere 10 whitish; pronotum, mesoscutum and mesoscutellum reddish-dark; metanotum reddish; rest of mesosoma blackbrown, except distal extremity of propodeal declivity reddish; petiole black; metasoma testaceous; legs testaceous. Antenna clavate; antennomeres in following proportions: 22: 15: 47: 32: 25: 21: 16: 15: 15: 22. Head excavated, dull, granulate; frontal line complete; occipital carina incomplete, shortly present on sides of lateral ocelli; POL = 9; OL = 8; OOL = 27. Palpal formula 6 / 3. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum shiny, with some longitudinal keels, with two lateral pointed apophyses situated on sides of mesoscutellum. Metanotum transversely striate, excavated behind mesoscutellum (Fig. 195 A). Metapectal-propodeal disc dull, granulate, sculptured by some slight longitudinal keels; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (9: 13). Enlarged claw (Fig. 194 D) strongly curved, with one large subdistal tooth, one row of four lamellae and one proximal tooth. Protarsomere 5 (Fig. 194 D) with two rows of approximately 32 lamellae; apex with approximately 40 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B87FF3ED5FFFE1FF97B.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, 28. XI. 1938, R. E. Turn- er leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B87FF3ED5FFFE1FF97B.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B87FF3ED5FFFE1FF97B.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B98FF3ED100FE1FFC90.taxon	description	(Figs 195 B, C, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B98FF3ED100FE1FFC90.taxon	description	Description. ♀. Apterous; body length 3.7 – 4.8 mm. Completely ferruginous, except petiole black. Antenna cla- vate; antennomeres in following proportions: 24: 13: 40: 24: 20: 17: 15: 12: 12: 16. Head (Fig. 195 B) excavated, dull, granulate; frontal line complete; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 4; OL = 5; OOL = 21. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, with collar shiny and slightly granulate and disc dull and strongly granulate. Mesoscutum dull, granulate, without lateral pointed apophyses. Metanotum dull, transversely striate, weakly excavated behind mesoscutellum (Fig. 195 C), laterally slightly protruding (protrusions slightly pointed). Metapectal-propodeal disc dull, granulate, very sloping anteriorly (Fig. 195 C); propodeal declivity transversely striate. Mesopleuron and metapleuron not transversely striate. Meso- metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (14: 16). Enlarged claw (Fig. 195 E) with one large subdistal tooth and one row of eight lamellae. Protarsomere 5 (Fig. 195 E) with two rows of approximately 22 – 24 lamellae; apex with approximately 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B98FF3ED100FE1FFC90.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 29. I – 5. II. 1924, R. E. Turner leg. (NHMUK). Paratype: same locality label as holotype, XII. 1923, 1 ♀ (NHMUK). Other mate- rial: SOUTH AFRICA: KwaZulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg., 1 ♀ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B98FF3ED100FE1FFC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE602B98FF3ED100FE1FFC90.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7F2B99FF3ED11BFE1FFC20.taxon	description	(Figs 189 D – F, 195 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7F2B99FF3ED11BFE1FFC20.taxon	description	Description. ♀ (Figs 189 D-F). Apterous; body length 3.6 – 3.8 mm. Head brown-ferruginous, except anterior region of frons and mandible testaceous; clypeus yellow, except brown central spot; antenna brown, except ventral side of scape whitish and flagellomere 8 testaceous-brown; mesosoma brown-ferruginous, except posterior margin of pronotum, posterior half of mesoscutum and apex of propodeal declivity testaceous; metasoma brown; legs brown, except junctions, meso- and metatarsus and chelae testaceous. Antenna clavate; antennomeres in following propor- tions: 8: 6: 16: 10: 8: 7: 6: 6: 6: 9. Head excavated, dull, granulate; frontal line complete; occipital carina absent; temple distinct; POL = 2; OL = 3; OOL = 10. Palpal formula 6 / 3. Pronotum dull, granulate, crossed by strong transverse impression, with many longitudinal striae on sides and anterior collar. Mesoscutum dull, granulate, with many longitudinal striae, without two lateral pointed apophyses. Mesoscutellum shiny, smooth, sloping anteriorly. Meta- notum excavated behind mesoscutellum, flat, strongly irregularly striate. Metapectal-propodeal disc dull, strongly sculptured by many longitudinal striae; propodeal declivity transversely striate; Metapectal-propodeal disc laterally protruding and forming slightly pointed protrusion. Mesopleuron and metapleuron dull, transversely striate. Meso- metapleural suture distinct and complete. Protarsomeres in following proportions: 17: 3.5: 7: 19: 30. Enlarged claw (Fig. 195 F) with one large subdistal tooth and one row of 13 – 14 lamellae + one bristle. Protarsomere 5 (Fig. 195 F) with two rows of 27 – 29 lamellae; apex with approximately 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7F2B99FF3ED11BFE1FFC20.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, WSW Grahamstown, Faraway, 33 ° 19.52 ’ S 26 ° 28.53 ’ E, 8. III. 1979, C. F. Jacot-Guillarmod leg. (AMG). Other material: SOUTH AFRICA: Kwa- Zulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg., 1 ♀ (NMSA).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7F2B99FF3ED11BFE1FFC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7F2B99FF3ED11BFE1FFC20.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9AFF3ED6D7FE32FC41.taxon	description	(Figs 195 D, G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9AFF3ED6D7FE32FC41.taxon	description	Description. ♀. Apterous; body length 3.6 mm. Completely ferruginous-dark, except petiole black. Antenna cla- vate; antennomeres in following proportions: 10: 6.5: 19: 12: 11: 8: 7: 6: 5.5: 9.5. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; temple distinct; POL = 1.5; OL = 1; OOL = 9. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, unsculptured, with few longitudinal keels on sides, without lateral pointed apophyses. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum weakly excavated behind mesoscutellum (Fig. 195 D), flat, with few transverse striae. Metapectal-propodeal disc shiny, unsculptured, sloping anteriorly (Fig. 195 D); propodeal declivity transversely striate; metapectal-propodeal disc protruding laterally and forming a slightly pointed protrusion. Mesopleuron and metapleuron shiny, almost completely unsculptured, partly transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 16: 3: 6: 19: 29. Enlarged claw (Fig. 195 G) with one large subdistal tooth and one row of seven lamellae + one bristle. Protarsomere 5 (Fig. 195 G) with two rows of 2 (proximal) + 14 lamellae; apex with approximately 19 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9AFF3ED6D7FE32FC41.taxon	materials_examined	Material examined. Type: ♀ holotype: TANZANIA: South Pare Mountains, peak above Kisiwani, 04 ° 08 ’ S 37 ° 53 ’ E, 29. XI. 1995, winkler bag leaf litter extraction, montane forest, SAM-HYM-PO 15170, H. G. Robertson leg. (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9AFF3ED6D7FE32FC41.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9AFF3ED6D7FE32FC41.taxon	distribution	Distribution. Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	description	(Figs 197, 198 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 4 / 2; head and mesosoma completely yellow-testaceous; pronotum not crossed by transverse furrow; protarsomere 1 slightly longer than 4; enlarged claw (Fig. 198 B) with one large subdistal tooth and one row of lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	description	Description. ♀ (Fig. 197). Apterous; body length 2.7 mm. Completely yellow-testaceous, except petiole black. Head (Fig. 197 C) excavated, shiny, unsculptured; frontal line incomplete, present in anterior half of frons; occipital carina absent; POL = 1; OL = 2; OOL = 7. Palpal formula 4 / 2. Pronotum shiny, unsculptured, not crossed by transverse impression. Mesoscutum shiny, unsculptured, without lateral pointed apophyses. Metanotum shiny, unsculptured, flat, with sides rounded (Fig. 197 A), about coplanar to mesoscutellum (Figs 197 B, 198 A). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity with posterior half transversely striate. Mesopleuron and metapleuron shiny, unsculptured. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 14: 2: 3: 11: 16. Enlarged claw (Figs 197 D, 198 B) with one large subdistal tooth and one row of five lamellae. Protarsomere 5 (Figs 197 D, 198 B) with two rows of 15 lamellae; apex with about eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19463): MADAGASCAR: Fianarantsoa, 20 km SSW Am- bositra, Ankazomivady, 20 ° 46.6 ’ S 47 ° 09.9 ’ E, 1700 m, 7. I. 1998, B. L. Fisher # 1590, disturbed montane rainforest edge, sifted litter (leaf mold, rotten wood) (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7D2B9BFF3ED39FFB57FE0C.taxon	etymology	Etymology. The species is named flavotestaceus, because of its colour yellow-testaceous.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7C2B9CFF3ED04FFE1FFE0C.taxon	description	(Fig. 198 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7C2B9CFF3ED04FFE1FFE0C.taxon	description	Description. ♀. Apterous; body length 3.4 mm. Head brown, except mandible, clypeus and anterior region of frons reddish; antenna testaceous; pronotum reddish-dark; rest of mesosoma testaceous-reddish; petiole black; metasoma testaceous-reddish; legs yellow. Head excavated, shiny, unsculptured or slightly granulate; frontal line complete; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 3; OL = 7; OOL = 16. Palpal formula 3 / 2. Pronotum shiny, granulate, not crossed by transverse impression or very slightly impressed. Mesoscutum dull, with some longitudinal keels, without lateral pointed apophyses. Metanotum transversely striate, with sides rounded, about coplanar to mesoscutellum. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 approximately twice as long as 4. Enlarged claw (Fig. 198 C) with one large subdistal tooth and one row of six lamellae. Protarso- mere 5 (Fig. 198 C) with two rows of 11 lamellae; apex with eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7C2B9CFF3ED04FFE1FFE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Katberg, 4000 ft., 1 – 15. I. 1933, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7C2B9CFF3ED04FFE1FFE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7C2B9CFF3ED04FFE1FFE0C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7A2B9EFF3ED3ACFF20FCBC.taxon	description	(Figs 198 D – G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7A2B9EFF3ED3ACFF20FCBC.taxon	description	Description. ♀. Apterous; body length 2.4 – 2.6 mm. Head brown-black, except mandible, clypeus and anterior re- gion of frons yellow; antenna brown, except scape yellow; pronotum brown-black, partly reddish; rest of mesosoma black; metasoma brown-black or black; legs yellow or yellow-brown, except clubs of femora, coxae and tibiae brown. Head (Fig. 198 D) excavated, shiny, granulate. Palpal formula 4 / 2. Pronotum (Fig. 198 F) shiny, granulate, not crossed by transverse impression or very slightly impressed. Mesoscutum granulate, with some longitudinal keels, without lateral pointes apophyses (Fig. 198 E). Metanotum not striate. Metapectal-propodeal disc dull, granulate, with track of median longitudinal furrrow; propodeal declivity, meso- and metapleuron not transversely striate. Meso-metapleural suture obsolete or slightly distinct. Protarsomeres in following proportions: 9: 2: 3: 9: 16. Enlarged claw (Fig. 198 G) with one small subapical tooth and a 4 – 6 peg-like setae. Protarsomere 5 (Fig. 198 G) with one row of 17 – 20 lamellae; apex with at least nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7A2B9EFF3ED3ACFF20FCBC.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele (= Haut- Zaire), Garamba National Park (MRAC). Paratype: same locality label as holotype, 1 ♀ (MRAC). Other material: ETHIOPIA: Shoa Province, 95 km W Addis Ababa, Ambo, near Plant Protection Research Institute, 2400 – 2500 m, 4. V. 1990, A. Emel’yanov leg., 1 ♀ (ZISP); same locality label, 25. V. 1990, 2 ♀♀ (ZISP); same locality label, 4. VI. 1990, 1 ♀ (MOLC). MADAGASCAR: Toamasina, Rogez, 1 ♀ (MRAC). MOZAMBIQUE: Nampula Prov., Nampula, ex Exitianus frontalis (Distant), C. 22. XI. 1999, B. 25. XI. 1999, Sf. 23. XII. 1999, M. Olmi reared, 1 ♀ (MOLC). SOUTH AFRICA: Free State, Royal Natal National Park, 11 km E of Park Gate along Road from R 74 to Park, grass along the road, C. 9. IV. 2007, B. 11. IV. 2007, M. Olmi reared, 1 ♀ (MOLC); Western Cape, Cederberg Mts., Algeria, 6. II. 2006, M. Olmi reared ex Cicadulina theroni Van Rensburg, 2 ♀♀ (MOLC, SAMC). TANZANIA: Mkomazi Game Reserve, Kisima Plot, 04 ° 06.06 ’ S 38 ° 05.58 ’ E, 25. XI – 8. XII. 1995, MT, Acacia / Commiphora bushland, SAM-HYM-PO 16497, S. van Noort leg., 1 ♀ (SAMC); Mkomazi Game Reserve, Kavateta Dam, 03 ° 54.90 ’ S 38 ° 02.55 ’ E, 16. IV. 1996 sweep, mk 22, grasses and shrubs in Acacia / Commiphora bushland, SAM-HYM-PO 17442, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7A2B9EFF3ED3ACFF20FCBC.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Exitianus frontalis (Distant); in South Africa: Cicadulina theroni Van Rensburg; Exitianus taeniaticeps (Kirschbaum); Maiestas angustisecta (Linnavuori).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE7A2B9EFF3ED3ACFF20FCBC.taxon	distribution	Distribution. Democratic Republic of the Congo, Ethiopia, Madagascar, Mozambique, South Africa, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE792B9FFF3ED5C3FDBDFB40.taxon	description	(Figs 200 A, 201 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE792B9FFF3ED5C3FDBDFB40.taxon	description	Description. ♀. Apterous; body length 1.7 – 2.9 mm. Completely testaceous, except petiole black, posterior third of metasoma black and antennomeres 3 – 10 brown or black. Head rarely darkened. Antenna clavate; antennomeres in following proportions: 7: 4.5: 6: 4.5: 4.5: 4.5: 4.5: 4: 4: 6. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; temple distinct; POL = 1; OL = 1; OOL = 6. Palpal formula 3 / 2. Pronotum shiny, unsculptured, not crossed by transverse impression or very slightly impressed. Metanotum very sloping anteriorly, about coplanar to mesoscutellum. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Metapectal-propodeal disc usually unsculptured, occasionally granulate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 10: 2: 4: 9: 15. Enlarged claw (Fig. 201 A) with one small subapical tooth and a two peg-like setae + one bristle. Protarsomere 5 (Fig. 201 A) with one row of 1 (proximal) + 12 lamellae; apex with seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.7 – 2.0 mm. Head black, except mandible testaceous; antenna and mesosoma black; metasoma brown; legs brown, except tarsi and stalks of femora testaceous. Antenna filiform, with antennomere 3 less than 3 × as long as broad (5: 2); antennomeres in following proportions: 4: 3.5: 5: 5: 5: 5: 5: 5: 4.5: 7. Head dull, granulate; frontal line absent; occipital carina absent; POL = 6; OL = 3; OOL = 4; vertex with smooth ovoidal shiny area situated between eyes and lateral ocelli (area surrounded anteriorly by strong prominent process). Palpal formula 3 / 2. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.3 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc dull, completely granulate. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (9: 6). Dorsal process of paramere (Fig. 200 A) much shorter than paramere. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE792B9FFF3ED5C3FDBDFB40.taxon	materials_examined	Material examined. Types: ♀ holotype: MOZAMBIQUE: Maputo Prov., along Maputo-Boane road, 3 km N of Umbelúzi, C. 29. XI. 1993, B. 1. XII. 1993, Sf. 18. XII. 1993, M. Olmi reared from Cicadellidae (AMNH). Para- types: MOZAMBIQUE: same locality label as holotype, 29 ♀♀, 8 ♂♂ (AMNH) and 1 ♀, 1 ♂ (USNM), reared as follows: C. 29. XI. 1993, B. 30. XI. 1993, Sf. 17. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 1. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 3. XII. 1993, Sf. 19. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 20. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 29. XI. 1993, Sf. 20. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 30. XI. 1993, Sf. 17. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 20. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 1. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 20. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 1. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 29. XI. 1993, B. 2. XII. 1993, Sf. 18. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 23. XI. 1993, Sf. 10. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 7. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 7. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♀; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♀; C. 19. X. 1993, B. 23. X. 1993, Sf. 13. XI. 1993, 1 ♀; C. 19. X. 1993, B. 20. X. 1993, Sf. 11. XI. 1993, 1 ♀; C. 19. X. 1993, B. 22. X. 1993, Sf. 11. XI. 1993, 1 ♀; C. 19. X. 1993, B. 20. X. 1993, Sf. 11. XI. 1993, 1 ♀; C. 29. XI. 1993, B. 1. XII. 1993, Sf. 18. XII. 1993, 1 ♂; C. 29. XI. 1993, B. 30. XI. 1993, Sf. 16. XII. 1993, 1 ♂; C. 29. XI. 1993, B. 1. XII. 1993, Sf. 17. XII. 1993, 1 ♂; C. 19. X. 1993, B. 22. X. 1993, Sf. 12. XI. 1993, 1 ♂; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 5. XII. 1993, 1 ♂; C. 15. XI. 1993, B. 16. XI. 1993, Sf. 5. XII. 1993, 1 ♂; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♂; C. 15. XI. 1993, B. 18. XI. 1993, Sf. 6. XII. 1993, 1 ♂; C. 15. XI. 1993, B. 17. XI. 1993, Sf. 5. XII. 1993, 1 ♂; Maputo Prov., near Magude, Chobela, C. 29. X. 1993, B. 2. XI. 1993, Sf. 24. XI. 1993, 1 ♀; C. 29. X. 1993, B. 2. XI. 1993, Sf. 24. XI. 1993, 1 ♀; C. 29. X. 1993, B. 1. XI. 1993, Sf. 22. XI. 1993, 1 ♀; C. 29. X. 1993, B. 31. X. 1993, Sf. 22. XI. 1993, 1 ♀; C. 29. X. 1993, B. 4. XI. 1993, Sf. 26. XI. 1993, 1 ♀. Other material: GABON: Prefecture Ogoové-Maritime, Reserve de la Moukalaba-Dougoua, 7 km NW Doussala, 2 ° 19.84 ’ S 10 ° 32.65 ’ E, 110 m, 21. III. 2000, sweep, savanna, grasses & Nauclea latifolia, S. van Noort leg., 1 ♀ (SAMC). MOZAMBIQUE: Maputo Prov., Maputo, Maputo Race-course, M. Olmi leg., 2 ♀♀ (AMNH); Maputo Prov., 6 km E of Manhiça, C. 25. IX. 1996, M. Olmi leg., 3 ♀♀ (AMNH); Niassa, Cuamba, reared from Doratulina instabilis (Ribaut), M. Olmi reared, C. 15. XI. 1999, B. 18. XI. 1999, Sf. 5. XII. 1999, 1 ♀ (MOLC); same locality label, reared from Stirellus flavovirescens (Stål), C. 4. IV. 2003, B. 8. IV. 2003, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE792B9FFF3ED5C3FDBDFB40.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Anaconura instabilis (Ribaut) (= Doratulina instabilis (Ribaut )); Stirellus flavovirescens (Stål). Natural enemies. Hymenoptera Encyrtidae: in Mozambique: Cheiloneurus gonatopodis R. Perkins. This Encyrtid is one of the most common natural enemies of Dryinids in Mozambique (see also Echthrodelphax tauricus Ponomarenko, Gonatopus amoenus Olmi and Gonatopus incognitus Olmi). The reared specimens have the following locality labels: 10: Mozambique, Maputo Prov., 5 km E Manhiça, parasitized leafhopper collected on 17. IX. 1996; cocoon spun on 19. IX. 1996; encyrtids emergence on 7. X. 1996; 1 ♂, 3 ♀♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE792B9FFF3ED5C3FDBDFB40.taxon	distribution	Distribution. Gabon, Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE782B90FF3ED357FC72FEE0.taxon	description	(Fig. 201 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE782B90FF3ED357FC72FEE0.taxon	description	Description. ♀. Apterous; body length 2.1 – 2.2 mm. Head brown-black, except mandible, clypeus and region of frons along orbits yellow; antenna brown, except scape yellow; mesosoma and metasoma black; legs testaceous, except clubs of femora and meso- and metacoxa brown; occasionally extremities of meso- and metatibia brown. In specimen from Democratic Republic of the Congo, 04 ° 19 ’ S 17 ° 08 ’ E, head totally testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous. Head excavated, shiny, bare, unsculptured; frontal line complete; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 2.5; OL = 2.5; OOL = 14. Palpal formula 4 / 2. Pronotum shiny, setose, unsculptured, not crossed by transverse impression or very slightly impressed. Mesoscutum shiny, with some longitudinal keels. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, meso- and metapleuron slightly transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 slightly longer than 4 (8: 7). Enlarged claw (Fig. 201 B) with one small subapical tooth and 4 – 5 lamellae or peg-like setae. Protarsomere 5 (Fig. 201 B) with one row of 18 – 21 lamellae; apex with 5 – 11 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE782B90FF3ED357FC72FEE0.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Drakensberg, Van Reenen, 1 – 22. I. 1927 (NHMUK). Paratype: same locality label as holotype, XI. 1926, 1 ♀ (NHMUK). Other material: DEMO- CRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Yelingi Grassland, 5 km E Kikongo Mission, 04 ° 19 ’ S 17 ° 08 ’ E, 8. IV. 2006, sweeping grass, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD). SOUTH AF- RICA: Free State, Sasolburg, Roadside verges, Wonderfontein Road, 26 ° 47.20 ’ S 27 ° 47.33 ’ E, 1450 m, 23. XI. 1999, D-Vac, IS 22, seeded mainly with Eragrostis tef., M. Stiller leg., 1 ♀ (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE782B90FF3ED357FC72FEE0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE782B90FF3ED357FC72FEE0.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE772B92FF3ED036FC71FEC4.taxon	description	(Figs 200 B, C, 201 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE772B92FF3ED036FC71FEC4.taxon	description	Description. ♀. Apterous; body length 2.0 – 2.2 mm. Completely yellow-testaceous, except petiole black. Head al- most flat, shiny, unsculptured or slightly granulate; frontal line incomplete, absent near clypeus; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 2; OL = 3; OOL = 11. Palpal formula 3 / 2. Pronotum shiny, granulate, not crossed by transverse impression or very slightly impressed. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, meso- and metapleuron transversely striate. In specimen from Mozambique, Cu- amba, metapectal-propodeal disc dull and slightly granulate. Meso-metapleural suture obsolete or slightly visible on posterior half; in specimens from Cameroon, Djarengol, and Namibia, 38 km W Keetmanshoop, meso-metapleural suture distinct and complete. Protarsomere 1 as long as 4 (in holotype, or slightly shorter (7: 8, in specimens from Cameroon, Djarengol). Enlarged claw (Fig. 201 C) with one small subapical tooth and few bristles or four peg-like setae. Protarsomere 5 (Fig. 201 C) with inner proximal serrate region and one row of 16 lamellae or two rows of 1 (proximal) + 12 – 14 lamellae (extending beyond 0.5 × length of protarsomere); apex with approximately 9 – 11 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.1 – 1.6 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tarsi, protibia and articulations testaceous. Antenna filiform, with antennomere 3 less than 3 × as long as broad (3.5: 1.7); antennomeres in following proportions: 2: 3: 3.5: 3.5: 3: 3: 3: 3: 3: 4.5. Head dull, granulate; frontal line absent; occipital carina absent; POL = 4; OL = 2; OOL = 3; vertex with smooth ovoidal shiny area situated between eyes and lateral ocelli (area surrounded anteriorly by strong prominent process (Fig. 200 C )). Temple without sharp carina. Palpal formula 3 / 2. Mesoscutum, mesoscutellum and metanotum dull, granulate. Notauli incomplete, reaching approximately 0.3 – 0.6 × length of mesoscutum. Metapectal-propodeal disc shiny, not reticulate rugose, with numerous irregular striae, unsculptured among striae. Forewing hyaline, without dark transverse bands. Dorsal process of paramere (Fig. 200 B) long and with distal region broadened. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE772B92FF3ED036FC71FEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: NAMIBIA: Okahandja, 27. I – 2. II. 1928, R. E. Turner leg. (NHMUK). Other material: CAMEROUN: Maroua, Djarengol, 1 ♀ (CIRAD), 2 ♀♀, 1 ♂ (AMNH). MOZAMBIQUE: Maputo Prov., 27 km S of Maputo, Umbelúzi, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 22 km N of Maputo, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 6 km N of Palmeira, M. Olmi leg., 1 ♀ (AMNH); Nampula Prov., Nampula, near Nampula Catholic Seminary, 21. XI. 1999, sweeping, M. Olmi leg., 1 ♀ (MOLC); same locality label, reared from a nymph of unidentified Cicadellidae, C. 21. XI. 1999, B. 24. XI. 99, Sf. 16. XII. 99, M. Olmi reared, 1 ♂ (MOLC); Ni- assa, 8 km N of Maúa, Catholic Mission of “ Coração Sagrado de Jesus de Maúa ”, ex nymphs and adults of Exitianus capicola (Stål), M. Olmi reared (MOLC) (reared specimens as follows: C. 11. XI. 1999, B. 13. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 13. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 12. XI. 99, Sf. 2. XII. 99, 1 ♀; C. 11. XI. 1999, B. 14. XI. 99, Sf. 4. XII. 99, 1 ♀; C. 11. XI. 1999, B. 14. XI. 99, Sf. 4. XII. 99, 1 ♀; C. 11. XI. 1999, B. 14. XI. 99, Sf. 4. XII. 99, 1 ♀; C. 11. XI. 1999, B. 13. XI. 99, Sf. 1. XII. 99, 1 ♀; C. 11. XI. 1999, B. 19. XI. 99, Sf. 3. XII. 99, 1 ♂; C. 11. XI. 1999, B. 12. XI. 99, Sf. 1. XII. 99, 1 ♂; C. 11. XI. 1999, B. 12. XI. 99, Sf. 2. XII. 99, 1 ♂; C. 11. XI. 1999, B. 13. XI. 99, Sf. 2. XII. 99, 1 ♂; C. 11. XI. 1999, B. 13. XI. 99, Sf. 1. XII. 99, 1 ♂; C. 11. XI. 1999, B. 12. XI. 99, Sf. 2. XII. 99, 1 ♂; C. 11. XI. 1999, B. 13. XI. 99, Sf. 1. XII. 99, 1 ♂); Niassa, Cuamba, Mozambique Catho- lic University Campus, 26. V. 2007, sweeping, M. Olmi leg., 1 ♀ (MOLC); same locality label, C. 29. V. 2007, B. 31. V. 2007, Sf. 22. VI. 2007, M. Olmi reared from an adult of Cicadellidae, 1 ♀ (SAMC); same locality label, C. 29. V. 2007, B. 1. VI. 2007, Sf. 1. VII. 2007, M. Olmi reared from a nymph of Cicadellidae, 1 ♀ (MOLC); same local- ity label, sweeping, 11. IV. 2003, 6 ♀♀ (MOLC); same locality label, reared from nymphs of unidentified Cicadel- lidae, C. 7. IV. 2003, B. 10. IV. 2003, Sf. 26. IV. 2003, 2 ♀♀ (MOLC); same locality label, reared from a ♀ adult of an Aconurella species (Cicadellidae), C. 5. IV. 2003, B. 8. IV. 2003, Sf. 24. IV. 2003, 1 ♀ (MOLC); Niassa, near Cuamba, pastures near a bridge at the beginning of Maúa Road, C. 8. XI. 1999, B. 9. XI. 99, Sf. 1. XII. 99, reared from a nymph of unidentified Cicadellid, 1 ♀ (MOLC). NAMIBIA: 38 km W Keetmanshoop, along Rd B 4, 10. III. 2006, sweeping, M. Olmi leg., 3 ♀♀ (MOLC); same locality label, C. 10. III. 2006, B. 13. III. 2006, Sf. 3. IV. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 26 C, 1 ♂ (MOLC); same locality label, C. 10. III. 2006, B. 14. III. 2006, Sf. 5. IV. 2006, M. Olmi leg., 26 D, 1 ♀ (MOLC); same locality label, C. 10. III. 2006, B. 13. III. 2006, Sf. 5. IV. 2006, M. Olmi leg., 26 E, 1 ♀ (MOLC); same locality label, C. 10. III. 2006, B. 11. III. 2006, Sf. 5. IV. 2006, M. Olmi leg., 26 A, 1 ♀ (MOLC); 89 Km S Keetmanshoop, along Rd B 1, C. 1. III. 2006, B. 4. III. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 18 M, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 2. III. 2006, Sf. 27. III. 2006, M. Olmi leg., 18 B, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 3. III. 2006, Sf. 27. III. 2006, M. Olmi leg., 18 D, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 3. III. 2006, Sf. 22. III. 2006, M. Olmi leg., 18 F, 1 ♀ (SAMC); same locality label, C. 1. III. 2006, B. 4. III. 2006, Sf. 27. III. 2006, M. Olmi leg., 18 H, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 2. III. 2006, Sf. 22. III. 2006, M. Olmi leg., 18 E, 1 ♀ (MOLC); 40 Km W Keet- manshoop, along Rd B 4, C. 10. III. 2006, B. 14. III. 2006, Sf. 5. IV. 2006, reared from an adult of Aconurella compta (Naudé), M. Olmi leg., 27 C, 1 ♀ (MOLC). SOUTH AFRICA: Eastern Cape, 16 km E of Port Elizabeth, S. George’s Strand, M. Olmi leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE772B92FF3ED036FC71FEC4.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Aconurella shaba Ghauri; Exitianus capicola (Stål); in Namibia: Aconurella compta (Naudé).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE772B92FF3ED036FC71FEC4.taxon	distribution	Distribution. Cameroon, Mozambique, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE752B93FF3ED3BEFD7EFCBC.taxon	description	(Figs 200 D, 201 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE752B93FF3ED3BEFD7EFCBC.taxon	description	Description. ♀. Apterous; body length 2.2 – 3.6 mm. Completely reddish, except petiole black and metasoma brown- ish. Head excavated, shiny, granulate; frontal line complete; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 5; OL = 5; OOL = 22. Palpal formula 4 / 2. Pronotum dull, unsculptured or slightly alutaceous, not crossed by transverse impression or very slightly impressed. Mesoscutum without lateral pointed apophyses. Metanotum with sides rounded. Metapectal-propodeal disc dull, granulate; propodeal declivity transversely striate. Mesopleuron and metapleuron not transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 approximately as long as 4. Enlarged claw (Fig. 201 D) with one small subapical tooth and 4 – 5 peg-like setae. Protarsomere 5 (Fig. 201 D) with inner proximal serrate region and one row of approximately 16 lamellae; apex with approximately 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.5 – 2.1 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except tarsi and articulations testaceous. Antenna filiform, with antennomere 3 less than 3 × as long as broad (4: 1.5); antennomeres in following proportions: 3.5: 3.5: 4: 3.5: 3.5: 3.5: 3.5: 3.5: 3: 5. Head swollen, shiny, granulate; frontal line absent; occipital carina absent; POL = 6; OL = 3; OOL = 2; temple distinct; vertex with smooth ovoidal shiny area situated between eyes and lateral ocelli (area surrounded anteriorly by strong prominent process). Palpal formula 4 / 2. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc and propodeal declivity shiny, unsculptured, slightly granulate near metanotum. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; distal part of 2 r-rs & Rs vein longer than proximal part (9: 6). Dorsal process of paramere (Fig. 200 D) short, transverse and with apex rounded. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE752B93FF3ED3BEFD7EFCBC.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Mtunzini District, Lr. Umhlatuzi R., 6. VI. 1926, R. E. Turner leg. (NHMUK). Other material: MOZAMBIQUE: Maputo Prov., along Macia-Praia do Bilene road, 23.6 km S of Macia, M. Olmi reared, C. 4. X. 1996, B. 6. X. 1996, Sf. 3. XI. 1996, 1 ♂ (AMNH); same locality label, C. 4. X. 1996, B. 7. X. 1996, Sf. 3. XI. 1996, 1 ♂ (AMNH); same locality label, C. 4. X. 1996, B. 6. X. 1996, Sf. 4. XI. 1996, 1 ♂ (AMNH); same locality label, C. 4. X. 1996, B. 5. X. 1996, Sf. 4. XI. 1996, 1 ♂ (AMNH); same lo- cality label, C. 4. X. 1996, B. 8. X. 1996, Sf. 5. XI. 1996, 1 ♂ (AMNH); same locality label, C. 4. X. 1996, B. 8. X. 1996, Sf. 4. XI. 1996, 1 ♀ (AMNH); same locality label, sweeping, 4. X. 1996, 5 ♀♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE752B93FF3ED3BEFD7EFCBC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE752B93FF3ED3BEFD7EFCBC.taxon	distribution	Distribution. Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE742B94FF3ED5C3FBE9FE28.taxon	description	(Figs 200 E, 201 E, F, 202 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE742B94FF3ED5C3FBE9FE28.taxon	description	Description. ♀. Apterous; body length 2.3 – 2.9 mm. Head yellow-reddish, occasionally with vertex darkened; an- tenna yellow-reddish, occasionally with antennomeres 4 – 10 darkened; pronotum black, except lateral regions reddish (in specimen from Gambia, Perai Memadi, prothorax completely testaceous); rest of mesosoma black or brown (in specimen from Gambia, Perai Memadi, mesoscutum and mesoscutellum testaceous, metapectal-propodeal disc brown, except anterior surface and metanotum testaceous); metasoma brown; legs yellow, occasionally except clubs of femora and meso- and metacoxa brown. Head (Fig. 202 A) excavated and shiny, with occiput granulate. Palpal formula 4 / 2. Pronotum shiny, granulate, not crossed by transverse impression or very slightly impressed (Fig. 202 C). Mesoscutum without lateral pointed apophyses (Fig. 202 B). Mesoscutum, mesoscutellum and metapectal-propodeal disc dull, granulate; propodeal declivity transversely striate. Mesopleuron and metapleuron not transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 9: 2: 3: 10: 15. Enlarged claw (Figs 201 E, F) with one small subapical tooth and few peg-like setae or bristles. Protarsomere 5 (Figs 201 E, F) with inner proximal serrate region and one row of approximately 26 lamellae, or 2 rows of 2 (proximal) + 22 lamellae; apex with 6 – 8 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 – 1.9 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except articulations, proximal extremities of tibiae and tarsi whitish. Antenna filiform, with antennomere 3 less than 3 × as long as broad (4.5: 2); antennomeres in following proportions: 4: 4: 4.5: 4.5: 4.5: 4.5: 4: 4: 4: 5.5. Head dull, granulate; frontal line absent; occipital carina absent; POL = 7; OL = 3; OOL = 2.5; vertex with smooth ovoidal shiny area situated between eyes and lateral ocelli (area surrounded anteriorly by strong prominent process). Palpal formula 4 / 2. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum. Mesoscutellum and metanotum shiny, setose, unsculptured. Metapectal-propodeal disc shiny, unsculptured, with anterior half granulate, median longitudinal furrow and two short longitudinal keels on sides of furrow; propodeal declivity unsculptured. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; distal part of 2 r-rs & Rs vein longer than proximal part. Dorsal process of paramere (Fig. 200 E) short and with apex rounded. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE742B94FF3ED5C3FBE9FE28.taxon	materials_examined	Material examined. Type: ♀ holotype: ZIMBABWE: Eastern Mashonaland, Harare (= Salisbury), G. A. K. Marshall leg. (MCPM). Other material: BOTSWANA: Gaborone, 3. II. 1980, 1 ♀ (CNC). GAMBIA: Wuli District, Perai Memadi, 1 ♀ (ZIL). MOZAMBIQUE: Maputo Prov., Maputo Bay, Inhaca Island, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 22 km N of Maputo, M. Olmi leg., 1 ♀ (AMNH); Maputo Prov., 6 km N of Palmeira along the road Maputo-Xai Xai, C. 24. VIII. 1996, M. Olmi leg., 1 ♀ (AMNH); Niassa, Cuamba, Campus of the Catholic University of Mozambique, 10. IV. 2002, M. Olmi leg., 1 ♀ (MOLC); Niassa, Cuamba, Farm of the Catholic University of Mo- zambique, 18. II. 2005, M. Olmi leg., 1 ♀ (MOLC). SOUTH AFRICA: Gauteng, Pretoria District, Soutpan, 25 ° 24 ’ S 28 ° 06 ’ E, 21. III. 1990, N. C. Pienaar leg., 2 ♀♀ (SANC); KwaZulu-Natal, Umkomaas Game Ranch, 1 ♀ (AEIC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE742B94FF3ED5C3FBE9FE28.taxon	biology_ecology	Hosts. In Mozambique, unidentified Cicadellidae (Olmi 1994 a).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE742B94FF3ED5C3FBE9FE28.taxon	distribution	Distribution. Botswana, Gambia, Mozambique, South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE732B95FF3ED3C4FCD2FBF8.taxon	description	(Figs 200 F, 202 D – F, 203 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE732B95FF3ED3C4FCD2FBF8.taxon	description	Description. ♀. Apterous; body length 1.7 – 2.4 mm. Head brown-reddish, except mandible, clypeus and anterior region of frons yellow; antenna yellow-brown; pronotum brown-reddish; mesoscutum yellow; rest of mesosoma brown-reddish or black, except distal extremity of propodeal declivity yellow; petiole black; metasoma brown-reddish; legs yellow, except club of profemur brownish. In specimen from Madagascar, Morondava, head and mesosoma completely testaceous-reddish. In two small specimens (1.7 mm) from South Africa, 17 th km of R 315 from Malmesbury to Darling, body totally yellow-testaceous, except petiole black. Head (Fig. 202 D) excavated, shiny, unsculptured; frontal line complete; occipital carina incomplete, only shortly present on sides of lateral ocelli; POL = 3; OL = 5; OOL = 15. Palpal formula 3 / 2. Pronotum shiny, unsculptured, not crossed by transverse impression or very slightly impressed (Fig. 202 F). Mesoscutum without lateral pointed apophyses (Fig. 202 E). Metanotum sloping anteriorly, about coplanar to mesoscutellum, with sides rounded. Metapectal-propodeal disc shiny, unsculptured, with or without track of median longitudinal furrow; propodeal declivity, meso- and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 8: 2: 3: 7: 13. Enlarged claw (Fig. 203 D) with one small subapical tooth and 4 – 6 peg-like setae. Protarsomere 5 (Fig. 203 D) with inner proximal serrate region and one row of 12 – 15 lamellae; apex with 6 – 8 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 mm. Head, antenna and mesosoma black; metasoma and legs brown. An- tenna filiform, with antennomere 3 less than 3 × as long as broad (4: 2); antennomeres in following proportions: 3.5: 3: 4: 4: 4: 4: 4: 4: 3.5: 5. Head swollen, shiny, granulate; frontal line absent; occipital carina absent; POL = 6; OL = 3; OOL = 3; temple distinct; vertex with smooth ovoidal shiny area situated between eyes and lateral ocelli (area surrounded anteriorly by strong prominent process). Palpal formula 3 / 2. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc and propodeal declivity dull, reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (8: 4). Dorsal process of paramere (Fig. 200 F) short and slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE732B95FF3ED3C4FCD2FBF8.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape Prov., Mossel Bay, 30. XII. 1938, R. E. Turner leg. (NHMUK). Paratypes: same locality label as holotype, 6. XII. 1938, 17. XII. 1938, 20. XII. 1938, 30. XII. 1938, 24. I. 1939, 3. II. 1939, 1. II. 1939, 6. II. 1939, 37 ♀♀ (NHMUK); same locality label, 7 ♀♀ (AMNH). Other material: MADAGASCAR: Toliara, Morondava, 2 ♀♀ (AMNH). MOZAMBIQUE: Niassa, Cuamba, Mozambique Catholic University Campus, reared from a nymph of an unidentified species of Cicadellidae, C. 30. III. 2003, B. 1. IV. 2003, Sf. 20. IV. 2003, M. Olmi leg., 1 ♀ (MOLC). SOUTH AFRICA: Eastern Cape, near Port Elizabeth, Cape Recife Nature Reserve, reared from an unidentified species of Cicadellidae, 1 ♀ (AMNH); Free State, Wonderwater Coal Strip Mine, near Sasolburg, 26 ° 46.11 ’ S 27 ° 47.00 ’ E, 1450 m, 17. XII. 1998, Dvac grass in seeded rehabilitation, West burm, M. Stiller leg., 1 ♀ (SANC); Free State, Sasolburg, Wonderwater Colliery, 1450 m, 23. XI. 1999, D-Vac, near IS 08, 08 a, 20, mainly on Cynodon dactylon, regrowth after fire, west burm, M. Stiller leg., 1 ♀ (SANC); Western Cape, 17 th km of R 315 from Malmesbury to Darling, C. 4. II. 2006, B. 7. II. 2006, reared from Exitianus taeniaticeps (Kirschbaum), 3 A, M. Olmi reared, 2 ♀♀ (SAMC); Western Cape, Cederberg Mts., Algeria, 6. II. 2006, M. Olmi reared from Aconurella aethiopica (Cogan), 4 ♀♀, 1 ♂ (MOLC), 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE732B95FF3ED3C4FCD2FBF8.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: nymphs of unidentified species of Cicadellidae; in South Africa: Aconurella aethiopica (Cogan); Exitianus taeniaticeps (Kirschbaum).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE732B95FF3ED3C4FCD2FBF8.taxon	distribution	Distribution. Madagascar, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	description	(Figs 202 G – I, 203 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	description	Description. ♀. Apterous; body length 3.0 mm. Head brown, except clypeus, malar space, mandible and part of oc- ciput testaceous; propleuron brown; pronotum brown, except part of disc and sides reddish-testaceous; mesoscutum, mesoscutellum, mesopleuron and metapectal-propodeal disc brown, except some reddish-testaceous nuances; metasoma brown; legs brown, except tarsi, trochanters and part of protibia testaceous. Antenna clavate; antennomeres in following proportions: 8: 5: 16: 6: 6: 6: 5: 5.5: 5: 8. Head flat (Fig. 202 G), dull, granulate, covered with thin short setae; frontal line complete; occipital carina almost completely absent (short track present behind ocellar triangle); POL = 1; OL = 2; OOL = 6; temple very long (Fig. 202 H). Palpal formula 4 / 2. Pronotum almost bare, crossed by track of transverse impression (Fig. 202 I), with anterior collar shiny, unsculptured; disc dull and slightly alutaceous. Mesoscutum shiny, with numerous longitudinal striae, without lateral pointed apophyses. Mesoscutellum shiny, smooth, flat. Metanotum short, transversely striate and about coplanar to mesoscutellum (Fig. 202 I). Metapectal-propodeal disc shiny, covered with thin short setae, with top and anterior surface unsculptured; propodeal declivity, meso- and metapleuron transversely striate; top of metapectal-propodeal disc with short track of longitudinal furrow. Mesometapleural suture obsolete. Protarsomeres in following proportions: 12: 2: 3: 15: 21. Enlarged claw (Fig. 203 F) with one small subapical tooth and one row of seven peg-like setae + one bristle. Protarsomere 5 (Fig. 203 F) with short serrated inner proximal portion and one row of 25 lamellae; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: YEMEN: Ja’ar, 13 ° 13 ’ N 45 ° 18 ’ E, 50 m, 15. I. 2001, A. van Harten leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	distribution	Distribution. Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE722B96FF3ED29FFAF9FE0C.taxon	discussion	Remarks. Because of the presence of the track of the transverse furrow on the pronotum, G. yemenensis can also be included in group 7, where it is similar to Gonatopus mosseli Olmi 1984. However, the subapical tooth of the enlarged claw of G. yemenensis is farther from the apex (Fig. 203 F) than in G. mosseli (Fig. 233 E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE712B97FF3ED384FD16FE98.taxon	description	(Figs 203 A – C, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE712B97FF3ED384FD16FE98.taxon	description	Description. ♀. Apterous; body length 2.2 mm. Head brown, except mandible, clypeus and anterior region of frons yellow; antenna brown, except scape and part of flagellomere 1 yellow; mesosoma and metasoma black; legs yellow, except coxae and clubs of femora brownish. Head (Fig. 203 A) excavated, shiny, granulate. Palpal formula 4 / 2. Pronotum (Figs 203 B, C) granulate, not crossed by transverse impression or very slightly impressed. Mesoscutum granulate, with some keels, without lateral pointed apophyses (Fig. 203 C). Mesoscutellum granulate. Metapectalpropodeal disc granulate, very sloping anteriorly (Fig. 203 C); propodeal declivity, meso- and metapleuron not transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 7: 2: 3.5: 8: 14. Enlarged claw (Fig. 203 E) with one small subapical tooth and six peg-like setae. Protarsomere 5 (Fig. 203 E) with one row of approximately 19 lamellae; apex with at least nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE712B97FF3ED384FD16FE98.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Uele (= Haut- Zaire), Garamba National Park (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE712B97FF3ED384FD16FE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE712B97FF3ED384FD16FE98.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4E2BAAFF3ED0B9FE1FFBB0.taxon	description	(Fig. 204 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4E2BAAFF3ED0B9FE1FFBB0.taxon	description	Description. ♀. Apterous; body length 2.2 – 3.6 mm. Holotype reddish-dark, except head and metasoma blackish. In specimen from South Africa, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, head brown, except mandible, clypeus and short fron- tal stripe along orbits yellow-testaceous; mesosoma brown-reddish. In specimen from South Africa, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, head brown, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous, except last two antennomeres brown; mesosoma brown, except most part of mesoscutum testaceous; metasoma browntestaceous; legs mostly brown. Antenna clavate; antennomeres in following proportions: 10: 5: 12: 8: 6: 6: 5: 5: 5: 7. Head excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 1.5; OL = 3; OOL = 9. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, granulate. Mesoscutum with two lateral pointed apophyses. Metanotum rugose, excavated behind mesoscutellum. Metapectal-propodeal disc fusiform, reticulate rugose (in specimens from South Africa, 34 ° 27.414 ’ S 19 ° 21.393 ’ E and 32 ° 21.260 ’ S 26 ° 23.001 ’ E, slightly granulate); propodeal declivity and metapleuron transversely striate. Mesopleuron granulate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 4: 7: 18: 29. Enlarged claw (Fig. 204 A) with one small subapical tooth and 3 – 6 proximal setae. Protarsomere 5 (Fig. 204 A) with two rows of 3 (proximal) + 17 – 21 lamellae; apex with 12 – 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4E2BAAFF3ED0B9FE1FFBB0.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, near Cape Town, Milnerton, I. 1926, R. E. Turner leg. (NHMUK). Paratype: same locality label as holotype, 1 ♀ (NHMUK). Other material: SOUTH AFRICA: Eastern Cape, Winterberg, The Hoek Farm, 32 ° 21.260 ’ S 26 ° 23.001 ’ E, 1879 m, 6. X. 2010 – 18. I. 2011, MT, Amathole Mistbelt Grassland, S. van Noort leg., WTB 09 - GRA 1 - M 05, 1 ♀ (SAMC); Western Cape, Uitsig Farm near Clanwilliam, 31 ° 58 ’ S 19 ° 07 ’ E, 380 m, 28. XI. 1996, collected by D-Vac from 4 – yr old Aspalathus linearis plantation, Neser, Oberprieler & Stiller leg., 1 ♀ (SANC); Walker Bay Nature Reserve, 34 ° 27.414 ’ S 19 ° 21.393 ’ E, 57 m, 29. XI. 1997, sweep, South Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC); Gamkaberg Nature Reserve, 33 ° 39.941 ’ S 21 ° 53.505 ’ E, 315 m, 21. III – 5. V. 2009, MT, Gamka Thicket, S. van Noort leg., 1 ♀ (SAMC); Koeberg Nature Reserve, 33 ° 37.52 ’ S 18 ° 25.33 ’ E, 8. VIII. 1997, sweep 1, Metalasia muricata and Restio sp., Sand Plain Fynbos, S. van Noort leg., 1 ♀ (SAMC); Koeberg Nature Reserve, 33 ° 37.517 ’ S 18 ° 25.325 ’ E, 31. X. 1997, sweep, Sand Plain Fynbos, S. van Noort leg., 4 ♀♀ (3 in SAMC, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4E2BAAFF3ED0B9FE1FFBB0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4E2BAAFF3ED0B9FE1FFBB0.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4D2BAAFF3ED2C7FE2FF816.taxon	description	(Figs 204 B, 207 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4D2BAAFF3ED2C7FE2FF816.taxon	description	Description. ♀. Apterous; body length 4.8 mm. Head black, except mandible yellow; antenna brown, except an- tennomere 1 yellow; mesosoma black, except two reddish spots on lateral regions of pronotum collar; metasoma black; legs testaceous-reddish, except coxae and clubs of femora brownish. Head excavated, shiny, unsculptured, except occiput and anterior region of frons granulate; POL = 7; OL = 7; OOL = 25. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, setose, unsculptured, except lateral regions granulate. Mesoscutum shiny, with some longitudinal keels without lateral pointed apophyses (Fig. 207 A). Metanotum excavated behind mesoscutellum. Metapectal-propodeal disc shiny, unsculptured, with median longitudinal furrow, sculptured by longitudinal keels; propodeal declivity strongly transversely striate. Mesopleuron not transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 20: 4: 6: 26: 38. Enlarged claw (Fig. 204 B) with one small subapical tooth and three peg-like setae. Protarsomere 5 (Fig. 204 B) with inner side proximally not serrate and two rows of 3 + 27 lamellae; apex with at least 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4D2BAAFF3ED2C7FE2FF816.taxon	materials_examined	Material examined. Type: ♀ holotype: ETHIOPIA: Shewa (= Shoa) Prov., Fiche, 1430 m, V. 1947, K. M. Guichard leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4D2BAAFF3ED2C7FE2FF816.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4D2BAAFF3ED2C7FE2FF816.taxon	distribution	Distribution. Ethiopia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4C2BACFF3ED3E8FE14FE0C.taxon	description	(Fig. 204 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4C2BACFF3ED3E8FE14FE0C.taxon	description	Description. ♀. Apterous; body length 3.2 mm. Head black, except mandible and clypeus brown; antenna black, except antennomeres 5 – 6 brown; mesosoma and metasoma black; legs black, except tarsi brown. Antennomeres in following proportions: 8: 4: 11: 5: 5: 5: 4: 4: 4: 6. Head flat, dull, granulate; temple prominent. Palpal formula 6 / 3 (palps actually missing in holotype; according to Ceballos (1936), maxillary palps 6 - segmented; according to Benoit (1954), labial palps 3 - segmented). Pronotum crossed by strong transverse impression, shiny, unsculptured, with some slight longitudinal striae on disc. Mesoscutum shiny, with some slight longitudinal striae, with two lateral pointed apophyses. Metanotum very reduced, smooth, about coplanar to mesoscutellum. Metapectal-propodeal disc dull, with many longitudinal striae; propodeal declivity strongly transversely striate near petiole. Mesopleuron and metapleuron slightly transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 10: 2: 5: 13: 21. Enlarged claw (Fig. 204 C) with one small subapical tooth and five peg-like setae. Protarsomere 5 (Fig. 204 C) with inner side proximally not serrate and two rows of 2 (proximal) + 22 lamellae; apex with approximately six lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4C2BACFF3ED3E8FE14FE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, Ankaratra, III. 1932, A. Seyrig leg. (MNMS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4C2BACFF3ED3E8FE14FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4C2BACFF3ED3E8FE14FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BACFF3ED793FE28FB1A.taxon	description	(Fig. 204 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BACFF3ED793FE28FB1A.taxon	description	Description. ♀. Apterous; body length 2.8 mm. Head brown-black, except mandible, clypeus and anterior region of frons (more along orbits) yellow; antenna brown, except scape and part of flagellomere 1 yellow; mesosoma brown-reddish; petiole black; metasoma brown-black; legs testaceous, except club of profemur dark. Head excavated, shiny, unsculptured; POL = 2; OL = 7; OOL = 15. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum dull. Metanotum transversely striate, excavated behind mesoscutellum. Metapectal-propodeal disc dull, with median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (10: 12). Enlarged claw (Fig. 204 D) with one small subapical tooth, series of small teeth and eight peg-like setae. Protarsomere 5 (Fig. 204 D) with inner side proximally not serrate and two rows of 4 + 13 lamellae; apex with four lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BACFF3ED793FE28FB1A.taxon	materials_examined	Material examined. Type: ♀ holotype: LESOTHO: Mamathes, 21. III. 1954, C. Jacot Guillarmod leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BACFF3ED793FE28FB1A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BACFF3ED793FE28FB1A.taxon	distribution	Distribution. Lesotho.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BADFF3ED2BEFB71FF7C.taxon	description	(Fig. 204 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BADFF3ED2BEFB71FF7C.taxon	description	Description. ♀. Apterous; body length 4.3 – 5.0 mm. Head black, except mandible, clypeus, malar space and a nar- row stripe along orbits yellow; antenna brown, except scape yellow; mesosoma and metasoma black; legs yellow, except coxae partly black and clubs of femora partly brown. Head excavated, shiny, granulate, with slight or strong longitudinal keels on frons. Palpal formula 6 / 3. Pronotum shiny, smooth, granulate, crossed by strong transverse impression; in some specimens disc of pronotum unsculptured. Mesoscutum granulate, with some longitudinal keels, with two lateral pointed apophyses. Metapectal-propodeal disc slightly granulate, sculptured by many transverse or longitudinal striae, with top smooth; metanotum, propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 shorter than 4 (15: 19). Enlarged claw (Fig. 204 E) with one small subapical tooth and one row of 5 – 8 lamellae. Protarsomere 5 (Fig. 204 E) with inner side proximally serrate, with two rows of at least 17 – 22 lamellae extending beyond 0.5 × length of protarsomere; apex with at least 13 – 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BADFF3ED2BEFB71FF7C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, IV. 1933, A. Seyrig leg. (MNHN). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Katanga, Mitwaba, 1 ♀ (CAS). LESOTHO: Mamathes, 2. XI. 1952, C. Jacot Guillarmod leg., 1 ♀ (NHMUK). MADAGASCAR: Toliara, Bekily, IV. 1933, A. Seyrig leg., 1 ♀ (MRAC). SOUTH AFRICA: Western Cape, Cape Town, Milnerton, I. 1926, II. 1926, R. E. Turner leg., 5 ♀♀ (4 in NHMUK, 1 in AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BADFF3ED2BEFB71FF7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4B2BADFF3ED2BEFB71FF7C.taxon	distribution	Distribution. Democratic Republic of the Congo, Lesotho, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4A2BAEFF3ED703FB25FD00.taxon	description	(Figs 204 F, 205, 207 B, C) Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; temple without sharp carina; mesosoma mostly black; pronotum crossed by transverse furrow; mesoscutum less than twice as long as broad, without lateral pointed prominences; meso-metapleural suture distinct and complete; metanotum deeply excavated behind mesoscutellum; mesopleuron without lateral pointed prominence; metapectal-propodeal disc with slight track of median longitudi- nal furrow; mesopleuron, metapleuron and propodeal declivity granulate and strongly transversely striate; protarso- mere 1 slightly shorter than 4; enlarged claw (Fig. 204 F) with small subapical tooth; protarsomere 5 without inner proximal serrate region, with one row of 12 lamellae and apex unusually long. Description. ♀ (Fig. 205). Apterous; body length 3.1 mm. Head black, except mandible brown-testaceous; antenna brown; mesosoma black, except lateral margins brown; metasoma brown; legs brown-testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 17: 11: 9: 8: 7: 6: 6: 8. Head weakly excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 2; OL = 1.5; OOL = 9; greatest breadth of posterior ocellus about as long as POL. Palpal formula 6 / 3. Pronotum shiny, crossed by transverse furrow, with disc smooth, unsculptured, and anterior collar and lateral regions granulate. Mesoscutum less than twice as long as broad, granulate, without lateral pointed apophyses (Fig. 207 B). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 207 C). Metapectal-propodeal disc (Fig. 207 B) fusiform, dull, with anterior surface and top granulate; propodeal declivity granulate and strongly transversely striate; disc with track of median longitudinal furrow. Mesopleuron and metapleuron dull, granulate and transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 15: 3: 6: 16: 26. Enlarged claw (Fig. 204 F) with one small subapical tooth and two bristles. Protarsomere 5 (Fig. 204 F) without inner proximal serrate region, with one row of 12 lamellae; apex unusually long (Fig. 204 F), with about 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4A2BAEFF3ED703FB25FD00.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Northern Cape, Avontuur Farm, 16 km NW Nieu- woudtville, 31 ° 16.249 ’ S 19 ° 02.900 ’ E, 764 m, 9. II – 1. IV. 2010, YPT, Bokkeveld Sandstone Fynbos, S. van Noort leg., SAM-HYM-P 039012 (SAMC). Paratypes: same locality label as holotype, SAM-HYM-P 039012, 1 ♀ (SAMC); same locality label as holotype, 27. X. 2009 – 9. II. 2010, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4A2BAEFF3ED703FB25FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4A2BAEFF3ED703FB25FD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE4A2BAEFF3ED703FB25FD00.taxon	etymology	Etymology. The species is named avontuurensis after the collection site in Avontuur Farm.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BAEFF3ED497FE59F9AD.taxon	description	(Figs 206 A, 207 D, E, 217 E, F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BAEFF3ED497FE59F9AD.taxon	description	Description. ♀ (Figs 217 E, F). Apterous; body length 4.6 mm. Head black, except mandible, clypeus, part of malar space and anterior margin of frons testaceous; antenna brown, except scape testaceous; mesosoma and metasoma black; legs brown, except trochanters, protibia and part of tarsi testaceous-reddish. Antenna clavate; antennomeres in following proportions: 10: 6: 22: 12: 10: 10: 9: 8: 7: 12. Head excavated, dull, granulate, with frons and vertex rugose; occiput dull, granulate and sculptured by many irregular longitudinal striae; frontal line complete; occipital carina absent; POL = 2; OL = 3; OOL = 13. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum about twice as long as broad, dull, rugose, with two lateral pointed apophyses situated at border of mesoscutellum (Fig. 207 D). Mesoscutellum shiny, unsculptured, sloping anteriorly. Metanotum rugose, about coplanar to mesoscutellum (Fig. 207 E). Metapectal-propodeal disc shiny, with track of median longitudinal furrow, partly smooth and partly rugose; propodeal declivity strongly transversely striate. Mesopleuron and metapleuron rugose. Meso-metapleural suture thin, distinct and complete. Protarsomeres in following proportions: 18: 4: 6: 20: 34. Enlarged claw (Fig. 206 A) with one small subapical tooth and one row of eight lamellae in addition to one bristle. Protarsomere 5 (Fig. 206 A) with two rows of approximately 22 lamellae; apex with at least 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BAEFF3ED497FE59F9AD.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Nyanza Prov., Ungoye Field Station, 00 ° 36.91 ’ S 34 ° 05.52 ’ E, 1147 m, 15 – 29. I. 2005, MT, R. S. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BAEFF3ED497FE59F9AD.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BAEFF3ED497FE59F9AD.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BA0FF3ED032FC6BFC90.taxon	description	(Figs 206 B, 219 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BA0FF3ED032FC6BFC90.taxon	description	Description. ♀. Apterous; body length 2.9 – 3.0 mm. Head black, except clypeus and mandible yellow; antenna brown, except scape and pedicel or only scape yellow; mesosoma and metasoma black; legs yellow, except coxae and clubs of femora brown-black. Head excavated, shiny, bare, unsculptured, except frons and occiput granulate. Palpal formula 3 / 2 or 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, with some longitudinal keels, without lateral pointed apophyses. Metanotum transversely striate. Metapectalpropodeal disc shiny, unsculptured; propodeal declivity transversely striate. Meso- and metapleuron transversely striate. Meso-metapleural suture obsolete. In two specimens from Mozambique, Cuamba, metapectal-propodeal disc with deep median longitudinal furrow (furrow not present in other specimens reared in same locality). Enlarged claw (Fig. 206 B) with small subapical tooth and 5 – 9 peg-like setae. Protarsomere 5 (Fig. 206 B) with inner side proximally serrate and one row of 15 – 24 lamellae; apex with 8 – 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations and tarsomeres 1 – 4 testaceous. Antenna setose, filiform; antennomeres of ♂ from Mozambique, Cuamba, in following proportions: 4: 3: 4.5: 4: 3.5: 4: 3.5: 4: 3: 6; antennomere 3 less than 3 × as long as broad: 4.5: 2 (in ♂ specimen from Mozambique, Cuamba). Head dull, granulate; fron- tal line absent; occipital carina absent; in ♂ specimen from Mozambique, Cuamba, POL = 5; OL = 3; OOL = 4; greatest breadth of lateral ocelli shorter than OL (2: 3). Vertex of head with two oval, dull, granulate areas situated between lateral ocelli and eyes (areas surrounded anteriorly by very high carina; posteriorly not surrounded by carina). Temple distinct, granulate, not crossed by carina. Palpal formula 4 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli slightly shorter than greatest breadth of lateral ocelli (1.5: 2). Mesoscutellum and metanotum shiny, slightly punctate, unsculptured among punctures. Metapectalpropodeal disc dull, slightly reticulate rugose, without median longitudinal furrow; areolae of metapectal-propodeal disc small; propodeal declivity without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands, with only costal cell surrounded by strongly pigmented veins (other basal cells surrounded by less pigmented veins); 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part. Dorsal process of paramere (Fig. 219 A) short and slender, with apex pointed. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BA0FF3ED032FC6BFC90.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, III. 1943, A. Seyrig leg. (MNHN). Paratypes: same locality label as holotype, VII. 1937, A. Seyrig leg., 11 ♀♀ (7 in MNHN, 4 in MRAC). Other material: KENYA: Central Prov., Mt. Kenya, 1 ♀ (DEUW). MADAGASCAR: Toamasina, Tamatave, 1 ♀ (FSAE); Toamasina, Rogez, 2 ♀♀ (MRAC, AMNH). Toliara, Ifaty, 23 ° 09 ’ S 43 ° 37 ’ E, 17. IX. 1993, under seaweed drift on sand beach, W. E. Steiner & R. Andriamasimanana leg., 1 ♀ (USNM). MOZAMBIQUE: Niassa, Cuamba, Mo- zambique Catholic University Campus, reared from adults and nymphs of Exitianus capicola (Stål) and Exitianus taeniaticeps (Kirschbaum), M. Olmi leg., C. 19. IV. 2002, B. 21. IV. 2002, Sf. 12. V. 2002, 1 ♀ (MOLC); same locality label, C. 19. IV. 2002, B. 21. IV. 2002, Sf. 12. V. 2002, 1 ♀ (MOLC); same locality label, C. 22. III. 2003, B. 23. III. 2003, Sf. 14. IV. 2003, 1 ♀ (MOLC); same locality label, C. 23. III. 2003, B. 27. III. 2003, Sf. 18. IV. 2003, 1 ♀ (MOLC); same locality label, C. 29. V. 2007, B. 30. V. 2007, Sf. 30. VI. 2007, 1 ♀ (MOLC); same locality label, C. 29. V. 2007, B. 31. V. 2007, Sf. 1. VII. 2007, 1 ♀ (SAMC); same locality label, C. 19. IV. 2007, B. 20. IV. 2007, Sf. 14. V. 2007, 1 ♀ (SAMC); same locality label, C. 3. II. 2011, B. 4. II. 2011, Sf. 23. II. 2011, 1 ♀ (MOLC); same locality label, C. 6. II. 2011, B. 8. II. 2011, Sf. 27. II. 2011, 1 ♀ (MOLC); same locality label, C. 3. II. 2011, B. 4. II. 2011, Sf. 23. II. 2011, 1 ♀ (MOLC); same locality label, C. 6. II. 2011, B. 7. II. 2011, Sf. 1. III. 2011, 1 ♂ (MOLC); same locality label, C. 6. II. 2011, B. 10. II. 2011, Sf. 1. III. 2011, 1 ♀ (MOLC); same locality label, C. 1. IV. 2009, B. 7. IV. 2009, 1 ♀ (MOLC). SOUTH AFRICA: Gauteng, near Pretoria, Discovery, 28. IX. 1961, Empey leg., 1 ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BA0FF3ED032FC6BFC90.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Exitianus capicola (Stål) and Exitianus taeniaticeps (Kirschbaum). Biology. Biological cycle studied by Olmi in Mozambique (Niassa Prov., Cuamba): egg 27. V. 2007, first instar larva 31. V. 2007, second instar larva 1. VI. 2007, third instar larva 3. VI. 2007, fourth instar larva 5. VI. 2007, fifth instar larva 6. VI. 2007, cocoon from 7. VI. 2007, ♀ adult emerged 30. VI. 2007 (specimens in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE492BA0FF3ED032FC6BFC90.taxon	distribution	Distribution. Kenya, Madagascar, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE472BA1FF3ED08FFE14FC04.taxon	description	(Figs 206 C, 207 F – H)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE472BA1FF3ED08FFE14FC04.taxon	description	Description. ♀. Apterous; body length 2.6 – 3.7 mm. Yellow-reddish or ferruginous or yellow-testaceous, except petiole black and metasoma brown (in holotype) or testaceous; antenna testaceous, except antennomeres 6 – 10 or 9 – 10 or only 10 darkened (6 – 10 darkened in holotype). Antenna clavate; antennomeres in following proportions: 9: 6: 20: 11: 9: 8.5: 7: 6: 5.5: 8. Head excavated, shiny, unsculptured, or with tracks of slight longitudinal striae on frons (in holotype); frontal line incomplete and visible only in posterior half of frons; occipital carina almost completely absent (shortly present behind ocellar triangle); occiput shiny, unsculptured; POL = 2; OL = 3; OOL = 8. Palpal formula 6 / 3. Pronotum, crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, unsculptured, except for some slight longitudinal striae, laterally with two pointed apophyses (Fig. 207 F). Mesoscutellum shiny, smooth, flat. Length of mesoscutum + mesoscutellum twice as long as breadth of mesoscutum (16: 8). Metanotum smooth, with sides protruding (protrusions rounded) (Fig. 207 F), not (Fig. 207 G) or weakly excavated behind mesoscutellum (Fig. 207 H). Metapectal-propodeal disc shiny, unsculptured. Metapleuron transversely striate. Propodeal declivity with posterior half transversely striate. Mesopleuron smooth, shiny, transversely not striate. Meso-metapleural suture distinct and complete, occasionally (in testaceous specimens) hardly visible. Protarsomeres in following proportions: 12: 2: 4: 16: 23. Enlarged claw (Fig. 206 C) with one small subapical tooth and one row of 6 – 7 peg-like setae or eight small lamellae, in addition to one bristle. Protarsomere 5 (Fig. 206 C) with two rows of 3 – 4 (proximal) + 23 – 33 lamellae; apex with approximately 11 – 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE472BA1FF3ED08FFE14FC04.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Ivondro (MRAC). Other material: MADAGASCAR: Antsiranana, Manongarivo Special Reserve, 17.3 km 218 ° SW Antanambao, 14 ° 01.3 ’ S 48 ° 25.1 ’ E, 1580 m, montane rainforest, beating low vegetation, 27. X. 1998, Brian L. Fisher leg., 3 ♀♀ (2 in CAS, 1 in MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, 19. III – 5. IV. 2001, MT, MA- 01 - 01 B- 07, M. E. Irwin, Schlinger E. I. leg., 1 ♀ (MOLC). No locality, maybe from Desjardins’ collection, 2 ♀♀ (MNHN);	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE472BA1FF3ED08FFE14FC04.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE472BA1FF3ED08FFE14FC04.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE462BA1FF3ED59BFCEAF8EA.taxon	description	(Figs 206 D, 207 I, 208 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE462BA1FF3ED59BFCEAF8EA.taxon	description	Description. ♀. Apterous; body length 2.9 – 3.1 mm. Head brown, except mandible, clypeus, malar space, anterior half of frons and ventral side partly or totally testaceous; antenna brown-testaceous, except antennomeres 1 – 3 testaceous; mesosoma black, except lateral regions of pronotum testaceous; metasoma brown; legs brown-yellow. Head (Fig. 207 I) excavated, dull, granulate; frontal line incomplete, only present in posterior half of frons; POL = 1; OL = 2; OOL = 8; temple distinct; occipital carina incomplete, shortly present on sides of lateral ocelli. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, dull, granulate. Mesoscutum dull, granulate, less than twice as long as broad (11: 6) with two lateral pointed apophyses (Fig. 208 A). Mesoscutellum sloping anteriorly, shiny, unsculptured. Metanotum dull, flat, transversely striate, excavated behind mesoscutellum (Fig. 208 B), with sides protruding; lateral protrusions slightly pointed (Fig. 208 A). Metapectal-propodeal disc dull, granulate; propodeal declivity granulate, with posterior half transversely striate. Mesopleuron granulate and partly transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 13: 2: 4: 18: 25. Enlarged claw (Fig. 206 D) with one small subapical tooth and three small lamellae or peg-like bristles + one bristle. Protarsomere 5 (Fig. 206 D) with inner side proximally serrate and two rows of 3 (proximal) + 20 – 21 lamellae; apex with about eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE462BA1FF3ED59BFCEAF8EA.taxon	materials_examined	Material examined. Type: ♀ holotype: CAMEROON: South, near Ebolowa, Nkoemvon, 25. IX – 19. XI. 1979, MT, in forest, D. Jackson leg., NHMUK (E) 2007 – 19 (NHMUK). Other material: CENTRAL AFRICAN REPUB- LIC: Sangha-Mbaéré Prefecture, Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 03 ° 02 ’ 01 ’’ N 16 ° 24 ’ 57 ’’ E, 510 m, 6 – 7. V. 2001, MT, lowland rainforest, marsh clearing, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE462BA1FF3ED59BFCEAF8EA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE462BA1FF3ED59BFCEAF8EA.taxon	distribution	Distribution. Cameroon, Central African Republic.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE442BA3FF3ED6D7FCE7FC6D.taxon	description	(Figs 210 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE442BA3FF3ED6D7FCE7FC6D.taxon	description	Description. ♀. Apterous; body length 5.1 – 5.6 mm. Head brown; antenna reddish; mesosoma and metasoma black; legs reddish, except coxae black and clubs of femora brown. Head excavated, shiny, granulate, bare; POL = 5; OL = 10; OOL = 25. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, granulate, bare. Mesoscutum bare, with two lateral pointed apophyses. Metanotum transversely striate. Metapectal-propodeal disc granulate, with or without longitudinal striae; propodeal declivity strongly transversely striate. Mesopleuron and metapleuron trans- versely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 17: 4: 7: 30: 42. Enlarged claw (Figs 210 A, B) with one small subapical tooth and eight small lamellae in addition to one bristle. Protarsomere 5 (Fig. 210 A) with inner side proximally serrate and two rows of 1 (proximal) + 23 – 25 lamellae; apex with at least 6 – 10 lamellae. In specimen from Mozambique, Praia do Bilene, protarsomere 5 with two rows o 5 (proximal) + 13 lamellae (Fig. 210 B). Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE442BA3FF3ED6D7FCE7FC6D.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Willowmore (MCPM). Other mate- rial: GAMBIA: Bakau, 1 ♀ (AMNH). MOZAMBIQUE: Gaza Prov., Praia do Bilene airport, reared from Hecalus dubius Melichar, M. Olmi reared, 1 ♀ (AMNH). SOUTH AFRICA: Eastern Cape, Willowmore, 13 km from Ful- lerton, 3. X. 1971, C. Jacot – Guillarmod leg., 1 ♀ (AMG); Eastern Cape, Grahamstown, Strowan, 4. XII. 1966, A. Ja- cot – Guillarmod leg., 1 ♀ (AMG); Western Cape, Worcester, I. 1929, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Little Karoo, 38 mi E of Ceres, 17 – 25. XI. 1924, R. E. Turner leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE442BA3FF3ED6D7FCE7FC6D.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Hecalus dubius Melichar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE442BA3FF3ED6D7FCE7FC6D.taxon	distribution	Distribution. Gambia, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA4FF3ED6D7FDE2F959.taxon	description	(Figs 210 C, 219 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA4FF3ED6D7FDE2F959.taxon	description	Description. ♀. Apterous; body length 3.1 – 3.9 mm. Head black, except mandible, clypeus, anterior region of frons (more along orbits) and part of ventral side testaceous; antenna brown, except antennomere 1 testaceous; prothorax black, except posterior margin of pronotum testaceous; mesoscutum testaceous-dark, except anterior and posterior surface black; rest of mesosoma black; metasoma brown; proleg brown, except tarsus testaceous; meso- and metaleg brown, except tarsomeres 1 – 4 testaceous. Antenna clavate; antennomeres in following proportions: 8: 6: 16: 15: 14: 12: 10: 9: 7: 8. Head excavated, dull, granulate; frontal line incomplete, absent on anterior third of frons; occipital carina absent; temple prominent, granulate and crossed by longitudinal keel; POL = 1; OL = 1; OOL = 7; greatest breadth of lateral ocelli about as long as POL. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, granulate. Mesoscutum very long, about 4 × as long as broad, dull, granulate. Mesoscutellum hardly visible, dull, granulate. Stalk between pronotum and metanotum composed of mesoscutum + mesoscutellum, more than 4 × as long as broad (19: 4). Metanotum very long, granulate, not transversely striate, about coplanar to mesoscutellum. Metapectal-propodeal disc dull, completely granulate; propodeal declivity granulate, not transversely striate. Mesopleuron and metapleuron granulate, not transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 13: 2.5: 4: 14: 21. Enlarged claw (Fig. 210 C) with one small subapical tooth and one row of five peg-like setae + one bristle. Protarsomere 5 (Fig. 210 C) with one row of 10 – 14 lamellae; apex with about 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.9 mm. Head black, except mandible testaceous; antenna brown; mesosoma, metasoma and legs black. Antenna setose, filiform; antennomeres in following proportions: 4: 4: 5: 5: 5: 5: 5: 4: 4: 7; antennomere 3 less than 3 × as long as broad: 5: 2. Head dull, setose, granulate; frontal line complete; occipital carina absent; POL = 4; OL = 2; OOL = 4; greatest breadth of lateral ocelli about as long as OL. Vertex of head with two oval smooth, granulate areas situated between lateral ocelli and eyes (areas surrounded anteriorly by very high carina and posteriorly by lower carina not continuing on temple and behind ocellar triangle). Temple distinct, granulate, not crossed by carina. Palpal formula 4 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between the notauli about 0.5 × as long as greatest breadth of lateral ocelli. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc dull, slightly reticulate rugose, with areolae small, with median longitudinal furrow; propodeal declivity without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands, with only costal cell surrounded by strongly pigmented veins (other basal cells surrounded by less pigmented veins); 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part. Dorsal process of paramere absent; paramere with inner membranous band along entire length (Fig. 219 B). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA4FF3ED6D7FDE2F959.taxon	materials_examined	Material examined. Types: ♀ holotype: MOZAMBIQUE: Niassa, Mecanhelas District, Entre-Lagos, bank of Lake Chirua, C. 6. III. 2011, B. 9. III. 2011, Sf. 22. IV. 2011, M. Olmi reared from a nymph of Nicolaus attenuatus Still- er (MOLC). Paratypes: same locality label as holotype, C. 13. III. 2011, B. 15. III. 2011, Sf. 12. V. 2011, 1 ♀ (MOLC); same locality label as holotype, C. 13. III. 2011, B. 14. III. 2011, Sf. 14. V. 2011, 1 ♀ (MOLC); same locality label as holotype, C. 13. III. 2011, B. 14. III. 2011, Sf. 14. V. 2011, 1 ♂ (MOLC); same locality label as holotype, C. 13. III. 2011, B. 15. III. 2011, Sf. 15. V. 2011, M. Olmi reared from an adult of Nicolaus attenuatus Stiller, 1 ♀ (MOLC); same lo- cality label as holotype, C. 13. III. 2011, B. 16. III. 2011, Sf. V. 2011, 1 ♀ (MOLC); same locality label as holotype, C. 13. III. 2011, B. 16. III. 2011, dead in cocoon, reared from a ♀ adult of Nicolaus attenuatus Stiller, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA4FF3ED6D7FDE2F959.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Nicolaus attenuatus Stiller.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA4FF3ED6D7FDE2F959.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA6FF3ED17EFEE5FE70.taxon	description	(Figs 208 C, 210 D, 219 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA6FF3ED17EFEE5FE70.taxon	description	Description. ♀. Apterous; body length 2.9 – 3.1 mm. Head completely testaceous; occasionally head brown, except occiput, clypeus, mandible and anterior region of frons yellow; antenna yellow, except distal extremity slightly brownish; pronotum yellow or brown; mesoscutum yellow, except anterior margin black or brown; mesoscutellum, metapectal-propodeal disc and propodeal declivity yellow or reddish; in some specimens metapectal-propodeal disc yellow or reddish, with black or brown nuances on metanotum, mesoscutellum and metapleuron; mesopleuron yellow or reddish, occasionally with brown or black nuances; petiole black; metasoma brown-reddish or yellow; legs yellow; in some specimens club of profemur brownish. Head (Fig. 208 C) excavated, shiny, unsculptured; POL = 3; OL = 2.5; OOL = 12. Palpal formula 4 / 2 or 5 / 2. Pronotum crossed by strong transverse impression, shiny, bare, unsculptured. Mesoscutum dull, granulate, without lateral pointed apophyses. Metanotum slightly rugose. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Enlarged claw (Fig. 210 D) with one small subapical tooth and 3 – 5 peg-like setae or bristles. Protarsomere 5 (Fig. 210 D) with inner side proximally serrate (occasionally not serrate) and one row of 18 – 26 lamellae; apex with approximately 9 – 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.1 – 1.4 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown. Antenna filiform; antennomeres in following proportions: 3: 4: 6: 5: 5: 5: 5: 5: 5: 7; third antennomere 3 × or more than 3 × as long as broad (6: 2; 6: 1.5). Head shiny, swollen, granulate and rugose; frontal line complete; occipital carina absent; POL = 5; OL = 2; OOL = 2.5; vertex with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas anteriorly surrounded by more or less high carina). Palpal formula 5 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between the lateral ocelli as long as greatest breadth of lateral ocelli (2: 2). Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc dull, reticulate rugose, with areolae very small. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curvilinear, with distal part longer than proximal part (15: 5). Dorsal process of paramere (Fig. 219 C) slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA6FF3ED17EFEE5FE70.taxon	materials_examined	Material examined. Types: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Mt. Ruwenzori, Kiribata (MRAC). Paratypes: SOUTH AFRICA: Eastern Cape, Somerset East, 1 – 26. I. 1937, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Mossel Bay, 3. II. 1939, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 30. XII. 1938, 1 ♀ (NHMUK); same locality label, 2 ♀♀ (AMNH). TANZANIA: Zanzibar Island, Zanzibar, 16. III. 1955, L. F. Brown leg., 1 ♀ (NHMUK). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: Ituri Prov., Mongbwalu AGK camp, 01 ° 56 ‘ 32.81 ‘‘ N 30 ° 02 ‘ 25.62 ‘‘ E, 21 - 28. II. 2015, 75 YPT, behind the former President Mobutu’s Doctor’s house, A. Gumovsky leg., 1 ♀ (SIZK). ETHIOPIA: Kefa Province, Jima, 1 ♀ (DEUW). KENYA: West- ern Prov., Kakamega District, Kakamega Forest, 00 ° 22.12 ‘ N 34 ° 51.48 ‘ E, 1628 m, 5. V. 2006, Manfred Kraemer & Co., 1 ♀ (MTC). MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12.2 km 211 ° SSW of Joffre- ville, 12 ° 35.47 ‘ S 49 ° 9.34 ‘ E, 1300 m, 2 – 7. II. 2001, MT, montane rainforest, Fisher, Griswold et al. leg., BLF 2853, 1 ♀ (CAS). MOZAMBIQUE: Maputo Prov., Maputo, 3 ♀♀ (AMNH); Maputo Prov., 3 km N of Umbeluzi, 2 ♀♀ (AMNH); Maputo Prov., 6 km N of Palmeira, C. 30. XI. 1995, B. 5. XII. 1995, Sf. 21. XII. 1995, F. Finocchi reared from Cicadellid, 1 ♀ (AMNH); same locality label, C. 11. IX. 1996, B. 14. IX. 1996, SF. 10. XI. 1996, M. Olmi reared from Cicadellid, 1 ♀ (AMNH); Maputo Prov., along Palmeira-Macia road, 2 km E of Incomati River bridge, EN N. 1, 2 ♀♀ (AMNH); Nampula Prov., Nampula, M. Olmi reared from Cicadellid, C. 21. XI. 1999, B. 24. XI. 1999, Sf. 16. XII. 1999, 1 ♀ (MOLC); same locality label, C. 21. XI. 1999, B. 23. XI. 1999, Sf. 10. XII. 1999, 1 ♂ (MOLC); same locality label, C. 21. XI. 1999, B. 21. XI. 1999, Sf. 24. XII. 1999, 1 ♂ (MOLC); Zambézia, Gurué, C. 19. II. 2011, B. 21. II. 2011, Sf. 10. III. 2011, M. Olmi reared from Balclutha saltuella (Kirschbaum), 1 ♀ (MOLC). SOUTH AFRICA: Eastern Cape, 16 km E of Port Elizabeth, S. George’s Strand, 1 ♀ (AMNH); Free State, Royal Natal National Park, 11 km E of Park Gate along Road from R 74 to Park, grass along the road, C. 9. IV. 2007, B. 10. IV. 2007, M. Olmi reared, 1 ♀ (MOLC); same locality label, C. 9. IV. 2007, B. 13. IV. 2007, Sf. 2009, 1 ♀ (MOLC); KwaZulu-Natal, Ver- non Crookes Nature Reserve, 30 ° 17.4 ’ 30 ° 36.9 ’ E, 250 m, 18. II – 18. III. 2005, MT, M. Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Northern Drakensberg, The Ledges Guest Farm, along the road from R 74 to Cavern Berg Resort, grass in front of the farm, C. 9. IV. 2007, B. 12. IV. 2007, M. Olmi reared, 1 ♀ (MOLC); Western Cape, Buffalo Bay, Goukamma Nature Reserve, C. 26. I. 2003, B. 28. I. 2003, Sf. 19. II. 2003, M. Olmi reared from nymphs of Bonaspeia eriocephala (Cogan), 1 ♀ (MOLC); Western Cape, near Lottering, Road 103 - N 2 E Junction, C. 28. I. 2003, B. 30. I. 2003, Sf. 19. II. 2003. M. Olmi reared from adults of Platentomus sobrinus (Stål), 2 ♀♀ (MOLC, SAMC); West- ern Cape, near George, N 12 - Road to Montagu Pass junction, C. 25. I. 2003, B. 29. I. 2003, Sf. 19. II. 2003, M. Olmi reared from Recilia lobata (Linnavuori), 1 ♀ (MOLC); Western Cape, De Hoop Nature Reserve, near main office, 16. II. 2006, M. Olmi reared ex Recilia cotula (Cogan), 1 ♀, 2 ♂♂ (MOLC), 1 ♀, 1 ♂ (SAMC); Western Cape, West Coast National Park, Abrahamskraal Water Hole, M. Olmi reared from nymphs of unidentified Cicadellidae, C. 4. IV. 2007, B. 6. IV. 2007, 1 ♀ (MOLC); same locality label, C. 4. IV. 2007, B. 7. IV. 2007, 1 ♂ (MOLC); same locality label, C. 4. IV. 2007, B. 8. IV. 2007, 1 ♀ (MOLC); same locality label, C. 4. IV. 2007, B. 10. IV. 2007, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA6FF3ED17EFEE5FE70.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Balclutha saltuella (Kirschbaum); in South Africa: Bonaspeia eriocephala (Cogan); Deltocephalus cotulus (Cogan) (= Recilia cotula (Cogan )); Exitianus taeniaticeps (Kirschbaum); Maiestas angustisecta (Linnavuori); Maiestas lobata (Linnavuori) (= Recilia lobata (Linnavuori )); Platentomus sobrinus (Stål).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE432BA6FF3ED17EFEE5FE70.taxon	distribution	Distribution. Democratic Republic of the Congo, Ethiopia, Kenya, Madagascar, Mozambique, South Africa, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE412BA7FF3ED407FD8DFF7C.taxon	description	(Figs 208 D-G, 211 A, 219 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE412BA7FF3ED407FD8DFF7C.taxon	description	Description. ♀. Apterous; body length 2.3 mm. Completely yellow-testaceous, except petiole black and eye grey. Antenna clavate; antennomeres in following proportions: 7: 4: 6: 6: 5: 5: 4: 3.5: 3.5: 6. Head (Fig. 208 D) excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 5. Palpal formula 4 / 2. Pronotum shiny, unsculptured, crossed by transverse furrow. Mesoscutum less than twice as long as broad, without lateral pointed apophyses (Fig. 208 E). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 208 F). Metapectal-propodeal disc shiny, unsculptured, without track of median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron shiny, unsculptured. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 7: 1.5: 2.5: 8: 14. Enlarged claw (Fig. 211 A) with one small subapical tooth, one peg-like seta and one bristle. Protarsomere 5 (Fig. 211 A) with inner short proximal serrate region and one row of 14 lamellae; apex unusually very long (Fig. 211 A), with four lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.7 – 2.0 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown – reddish; legs brown-testaceous. Antenna setose, filiform; antennomeres of specimen from South Africa, Uitkyk Pass, in following proportions: 5: 4.5: 5.5: 5.5: 4.5: 5: 4.5: 5: 4.5: 6.5; antennomere 3 less than 3 × as long as broad: 5.5: 2 (in ♂ specimen from South Africa, Uitkyk Pass). Head shiny, setose, strongly punctate, unsculptured among punctures; frontal line of holotype incomplete, short, visible only in front of anterior ocellus; frontal line of ♂ specimen from South Africa, Uitkyk Pass, absent; occipital carina absent; in ♂ specimen from South Africa, Uitkyk Pass, POL = 4; OL = 1.3; OOL = 4.5; greatest breadth of lateral ocelli lobger than OL (2: 1.3). Vertex of head with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas surrounded anteriorly by very high carina (Fig. 208 G) and posteriorly by lower carina continuing on temple and behind ocellar triangle (Fig. 208 G )). Temple distinct, crossed by sharp carina parallel to posterior margin of eyes. Palpal formula 4 / 2. Mesoscutum shiny, strongly punctate. Notauli complete, posteriorly separated; minimum distance between notauli as long as (in holotype) or shorter (1.3: 2; in specimen from South Africa, Uitkyk Pass) than greatest breadth of lateral ocelli. Mesoscutellum and metanotum shiny, slightly punctate, unsculptured among punctures. Metapectal-propodeal disc dull, slightly reticulate rugose, with median longitudinal furrow; areolae of metapectal-propodeal disc small; propodeal declivity without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; with only costal cell surrounded by strongly pigmented veins; other basal cells surrounded by less pigmented veins); 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part (approximately 11: 4). Dorsal process of paramere (Fig. 219 D) short and slender, with apex pointed. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE412BA7FF3ED407FD8DFF7C.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: Western Cape, Swellendam, 17. XII. 1931 – 18. I. 1932 (NHMUK). Other material: NAMIBIA: 89 km S Keetmanshoop, along Rd B 1, C. 1. III. 2006, B. 2. III. 2006, Sf. 30. III. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 18 C, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 5. III. 2006, Sf. 2. IV. 2006, 18 S, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 7. III. 2006, Sf. 6. IV. 2006, 18 Z, 1 ♀ (MOLC). SOUTH AFRICA: Western Cape, Cederberg Mts., Uitkyk Pass, 7. II. 2006, M. Olmi leg., 1 ♀ (MOLC); same locality label, C. 7. II. 2006, B. 9. II. 2006, 6 P, M. Olmi reared from Elginus saltus (Naudé), 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE412BA7FF3ED407FD8DFF7C.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Namibia: Aconurella compta (Naudé); in South Africa: Elgi- nus saltus (Naudé).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE412BA7FF3ED407FD8DFF7C.taxon	distribution	Distribution. Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE402BB8FF3ED291FD7EFE98.taxon	description	(Figs 208 H, 211 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE402BB8FF3ED291FD7EFE98.taxon	description	Description. ♀. Apterous; body length 5.9 – 6.9 mm. Head black, except clypeus, mandible and narrow stripe along orbits and on malar space testaceous; antenna brown, except scape and pedicel testaceous; mesosoma and metasoma black; legs black, except tarsi testaceous. Antenna clavate; antennomeres in following proportions: 10: 7: 25: 14: 12: 1 2: 11: 10: 10: 12. Head very slightly excavated, almost flat, dull, with short sparse setae, granulate and slightly rugose; temple distinct; frontal line complete; occipital carina absent; POL = 3; OL = 2; OOL = 10. Palpal formula 6 / 3. Pronotum dull, crossed by strong transverse impression, strongly granulate, with long sparse setae. Mesoscutum dull, rugose, granulate and sculptured by strong longitudinal keels, without lateral pointed apophyses, with long sparse setae. Mesoscutellum dull, granulate, sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 208 H), flat, granulate and reticulate rugose. Metapectal-propodeal disc dull, with long sparse setae, granulate; propodeal declivity granulate and with slight tracks of few transverse striae (striae more visible on sides of propodeal declivity). Meso- and metapleuron dull, granulate and rugose. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 18: 4: 6: 15: 25. Enlarged claw (Fig. 211 B) without subdistal tooth, with one row of seven bristles, with proximal half very broad. Protarsomere 5 (Fig. 211 B) with three rows of 24 lamellae; apex with at least 40 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE402BB8FF3ED291FD7EFE98.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Niassa, Cuamba, Mozambique Catholic University Campus, reared from an adult of Hecalus virescens (Distant), C. 26. IV. 2002, B. 30. IV. 2002, Sf. 15. V. 2002, M. Olmi leg. (MOLC). Other material: SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, Chase Valley, 29 ° 34.28 ’ S 30 ° 21.59 ’ E, 24. V. 1987, D. J. Brothers leg., 1 ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE402BB8FF3ED291FD7EFE98.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Hecalus virescens (Distant).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE402BB8FF3ED291FD7EFE98.taxon	distribution	Distribution. Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB8FF3ED73FFE53FB32.taxon	description	(Figs 208 I, 209 A, 211 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB8FF3ED73FFE53FB32.taxon	description	Description. ♀. Apterous; body length 3.2 – 3.3 mm. Head and mesosoma brown-reddish; antenna brown, except scape and pedicel testaceous; metasoma black; legs brown, except tarsi light; antenna clavate; antennomeres in following proportions: 9: 6: 12: 9: 7: 6.5: 6: 5: 5: 7. Head excavated, shiny, unsculptured, except occiput, temple and lower frons granulate; frontal line complete; occipital carina absent; POL = 1; OL = 3; OOL = 7. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured, except few striae in transversal furrow. Mesoscutum very slender, shiny, approximately twice as long as broad (10: 5), with few longitudinal striae, without lateral pointed apophyses (Fig. 208 I). Mesoscutellum flat, shiny, unsculptured. Meso-metapleural suture distinct and complete. Metanotum flat, about coplanar to mesoscutellum (Fig. 209 A). Metapectal-propodeal disc with strong median longitudinal furrow, with slight irregular striae; mesopleuron, metapleuron, propodeal declivity and lateral regions of metapectal-propodeal disc strongly transversely striate; top of metapectal-propodeal disc partly transversely striate. Protarsomeres in following proportions: 14: 3: 4: 13: 20. Enlarged claw (Fig. 211 C) with one small subapical tooth and one row of one bristle + six peg-like bristles. Protarsomere 5 (Fig. 211 C) with one row of 20 lamellae; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB8FF3ED73FFE53FB32.taxon	materials_examined	Material examined. Type: ♀ holotype: BENIN: 5 km W Ouidah, Asobonou, 14. XI. 1993, G. Delvare leg. (MNHN). Paratype: same locality label as holotype, 1 ♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB8FF3ED73FFE53FB32.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB8FF3ED73FFE53FB32.taxon	distribution	Distribution. Benin.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB9FF3ED346FF2EFEC4.taxon	description	(Figs 209 B, 211 D, 212 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB9FF3ED346FF2EFEC4.taxon	description	Description. ♀. Apterous; body length 3.0 – 3.5 mm. Yellow-reddish, except petiole black. Antenna clavate; anten- nomeres in following proportions: 8: 6: 18: 13: 10: 9.5: 8: 8: 8: 8. Head slightly excavated, shiny, slightly granulate; frontal line complete; occipital carina absent; POL = 2; OL = 3; OOL = 10. Palpal formula 4 / 2 or 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, slightly rugose, without lateral pointed apophyses. Metanotum shiny, not transversely striate, about coplanar to mesoscutellum (Fig. 209 B). Metapectalpropodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 15: 3: 4: 18: 28. Enlarged claw (Figs 211 D, 212 A) with one small subapical tooth and 5 – 7 bristles or peg-like setae. Protarsomere 5 (Fig. 211 D) with one row of 21 – 23 lamellae (in specimen from the United Arab Emirates, 25 ° 23 ’ N 56 ° 10 ’ E, one row of 10 – 11 lamellae (Fig. 212 A )); apex of protarsomere 5 with 9 – 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB9FF3ED346FF2EFEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: South Kivu, Mt. Shamulamba (AMNH). Other material: GAMBIA: Lower River Division, Upper Baddibu, Farafenni, 1 ♀ (KJHC). MOZAMBIQUE: Maputo Prov., 6 km E of Manhiça, 1 ♀ (AMNH). TANZANIA: Mkomazi Game Reserve, Ibaya Camp, north west side, 03 ° 57.91 ’ S 37 ° 48.09 ’ E, 30. IV – 2. V. 1996, YPT, Acacia / Commiphora / Combretum bushland, SAM – HYM – PO 19289, S. van Noort leg., 1 ♀ (SAMC). UNITED ARAB EMIRATES: near Tayyibah, 25 ° 23 ’ N 56 ° 10 ’ E, 10. III. 2005, hand collecting, 927, A. van Harten leg., 1 ♀ (MOLC). YEMEN: Madinat – ash – Shirq, 28. II. 2001, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB9FF3ED346FF2EFEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5F2BB9FF3ED346FF2EFEC4.taxon	distribution	Distribution. Democratic Republic of the Congo, Gambia, Mozambique, Tanzania, United Arab Emirates, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5E2BBAFF3ED037FD7EF9D4.taxon	description	(Figs 209 C, 212 C, 219 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5E2BBAFF3ED037FD7EF9D4.taxon	description	Description. ♀. Apterous; body length 3.6 – 4.8 mm. Completely ferruginous, except petiole black and metasoma darkened. Antenna clavate; antennomeres in following proportions: 7: 4: 17: 11: 8.5: 8: 7: 7: 6.5: 8.5. Head excavated, shiny, strongly granulate on occiput, slightly or strongly granulate on frons; temple distinct; frontal line complete; occipital carina incomplete, only present behind lateral ocelli; POL = 1; OL = 1.5; OOL = 9. Palpal formula 5 / 2. Pronotum dull, crossed by strong transverse impression, granulate. Mesoscutum dull, granulate, without lateral pointed apophyses. Metanotum about coplanar to mesoscutellum, with sides rounded, without lateral pointed pro- trusions (Fig. 209 C), occasionally transversely striate. Metapectal-propodeal disc granulate; propodeal declivity transversely striate; Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 17: 3.5: 5: 14: 22. Enlarged claw (Fig. 212 C) without subdistal tooth, with one row of 4 – 5 peg-like setae. Protarsomere 5 (Fig. 212 C) with two rows of 3 – 5 + 15 – 20 lamellae (shortest row extended beyond 0.5 × length of protarsomere); apex with at least 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.2 – 2.5 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs brown, except articulations, tarsi and part of tibiae whitish. Antenna filiform; antennomeres in following proportions: 4: 4: 7.5: 6: 6: 6.5: 6: 6: 6: 8; antennomere 3 more than 3 × as long as broad (7.5: 2). Head shiny, convex, granulate and rugose; frontal line complete; occipital carina absent; POL = 7; OL = 3; OOL = 3; vertex with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas not surrounded anteriorly by high carina). Palpal formula 4 / 2 or 5 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of lateral ocelli (3.5: 2). Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc shiny, mostly unsculptured, partly rugose. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; 2 r-rs & Rs vein regularly curvilinear, with distal part longer than proximal part (13: 9). Dorsal process of paramere (Fig. 219 F) very broad. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5E2BBAFF3ED037FD7EF9D4.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Maputo Prov., 6 km N of Palmeira along Maputo- Xai Xai road, C. 5. IX. 1995, B. 1. XII. 1995, M. Olmi reared from adult of Cicadellidae (AMNH). Paratypes: MO- ZAMBIQUE: same locality label as holotype, C. 24. VIII. 1996, B. 4. IX. 1996, Sf. 29. IX. 1996, F. Finocchi reared, 1 ♀ (AMNH); same locality label as holotype, C. 30. XI. 1995, B. 3. XII. 1995, 1 ♀ (AMNH); same locality label as holotype, C. 30. XI. 1995, B. 2. XII. 1995, Sf. 23. XII. 1995, F. Finocchi reared, 1 ♂ (AMNH); same locality label as holotype, C. 30. XI. 1995, B. 2. XII. 1995, Sf. 21. XII. 1995, F. Finocchi leg., 1 ♂ (AMNH); same locality label as ho- lotype, C. 24. VIII. 1996, B. 27. VIII. 1996, Sf. 24. IX. 1996, F. Finocchi reared, 1 ♂ (AMNH); same locality label as holotype, C. 24. VIII. 1996, B. 25. VIII. 1996, Sf. 29. IX. 1996, F. Finocchi reared, 1 ♂ (AMNH); Maputo prov., along Bairro dos Piscadores (Maputo) - Marracuene road, S. Maria, M. Olmi reared from Doratulina instabilis (Ribaut, 1948), 5. IX. 1996, B. 6. IX. 1996, Sf. 9. X. 1996, 1 ♀ (AMNH); same locality label, C. 5. IX. 1996, B. 10. IX. 1996, Sf. 14. X. 1996, 2 ♀♀ (AMNH); same locality label, C. 6. IX. 1996, B. 12. IX. 1996, Sf. 14. X. 1996, 1 ♀ (AMNH); same locality label, C. 6. IX. 1996, B. 7. IX. 1996, Sf. 10. X. 1996, 1 ♂ (AMNH); same locality label, C. 5. IX. 1996, B. 8. IX. 1996, Sf. 10. X. 1996, 1 ♂ (AMNH); same locality label, C. 5. IX. 1996, B. 9. IX. 1996, Sf. 10. X. 1996, 1 ♂ (AMNH); same locality label, C. 6. IX. 1996, B. 11. IX. 1996, Sf. 13. X. 1996, 1 ♂ (AMNH); same locality label, C. 5. IX. 1996, B. 11. IX. 1996, Sf. 13. X. 1996, 1 ♂ (AMNH); same locality label, C. 6. IX. 1996, B. 10. IX. 1996, Sf. 14. X. 1996, 1 ♂ (AMNH); same locality label, C. 5. IX. 1996, B. 12. IX. 1996, Sf. 14. X. 1996, 1 ♂ (AMNH); same locality label, C. 6. IX. 1996, B. 12. IX. 1996, Sf. 14. X. 1996, 1 ♂ (AMNH); same locality label, C. 5. IX. 1996, B. 10. IX. 1996, Sf. 15. X. 1996, 1 ♂ (AMNH); same locality label, C. 6. IX. 1996, B. 12. IX. 1996, Sf. 15. X. 1996, 1 ♂ (AMNH). Other material: MOZAMBIQUE: Nampula Prov., Nampula, about 500 m from the Bishop Seminary, C. 24. XI. 1999, B. 25. XI. 1999, Sf. 21. XII. 1999, M. Olmi reared from unidentified cicadellid, 1 ♀ (MOLC). SOUTH AFRICA: Limpopo, Thabazimbi and vicinity, 24 ° 41 ’ S 27 ° 21 ’ E, 2. IV. 2000, swept off Cenchrus ciliaris (Poaceae), M. Stiller leg., 1 ♀ (SANC); Mpumalanga, Krüger National Park, Satara, 16. XII. 1985, sweep, M. Sanborne leg., 1 ♀ (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5E2BBAFF3ED037FD7EF9D4.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Anaconura instabilis (Ribaut) (= Doratulina instabilis (Ribaut )).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5E2BBAFF3ED037FD7EF9D4.taxon	distribution	Distribution. Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5D2BBBFF3ED0EBFE31FBDC.taxon	description	(Figs 209 D, E, 212 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5D2BBBFF3ED0EBFE31FBDC.taxon	description	Description. ♀. Apterous; body length 3.2 – 3.3 mm. Head black, except mandible, clypeus, anterior margin of frons and part of malar space testaceous; antenna black, except scape and pedicel testaceous; mesosoma black, except lateral margins of pronotum brown-testaceous and mesoscutellum almost completely yellow; occasionally mesosoma completely black; metasoma brown; legs brown, except tibiae, tarsi and trochanters testaceous-brown. Head excavated, shiny, bare, unsculptured; frontal line complete; occipital carina absent; temple prominent, unsculptured, crossed by sharp carina (Fig. 209 D). Antenna clavate; antennomeres in following proportions: 8: 6.5: 10.5: 9: 8: 7: 6.5: 6.5: 6: 7. Antennomere 3 as broad as 2. POL = 1; OL = 2; OOL = 7; greatest breadth of lateral ocelli about as long as OL. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum very long, about twice as long as broad (11: 5.5), shiny, with few short longitudinal striae, without lateral pointed apophyses (Fig. 209 E). Mesoscutellum flat, shiny, smooth, circular. Stalk between pronotum and metanotum (composed of mesoscutum + mesoscutellum) about 3 × as long as broad. Metanotum very long, transversely striate, about coplanar to mesoscutellum. Metapectal-propodeal disc shiny, unsculptured, with track of median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron shiny, with posterior half smooth and unsculptured and anterior half transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 15: 2.5: 4: 13: 21. Enlarged claw (Fig. 212 D) with one small subapical tooth, four peg-like setae and one bristle. Protarsomere 5 (Fig. 212 D) with one row of 19 lamellae; apex with about 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5D2BBBFF3ED0EBFE31FBDC.taxon	materials_examined	Material examined. Types: ♀ holotype: NAMIBIA: 51 km S Grünau, along Rd B 1, C. 1. III. 2006, B. 3. III. 2006, Sf. 1. IV. 2006, reared from a ♀ adult of Aconurella compta (Naudé), M. Olmi leg., 16 A (MOLC). Paratypes: 89 km S Keetmanshoop, along Rd B 1, C. 1. III. 2006, B. 4. III. 2006, Sf. 1. IV. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 18 A, 1 ♀ (AMNH); same locality label, C. 1. III. 2006, B. 3. III. 2006, Sf. 31. III. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 18 P, 1 ♀ (MOLC); same locality label, C. 1. III. 2006, B. 3. III. 2006, Sf. 3. IV. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 18 N, SAM-HYM-P 025117, 1 ♀ (SAMC); 42 km N Kalkrand, along Rd. B 1, 9. III. 2006, M. Olmi leg., 25 A, 1 ♀ (MOLC); 38 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 11. III. 2006, Sf. 11. IV. 2006, reared from an adult of Aconurella compta (Naudé), M. Olmi leg., 26 F, 1 ♀ (MOLC); 38 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 18. III. 2006, Sf. 11. IV. 2006, reared from a nymph of Aconurella compta (Naudé), M. Olmi leg., 26 H, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5D2BBBFF3ED0EBFE31FBDC.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Namibia: Aconurella compta (Naudé).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5D2BBBFF3ED0EBFE31FBDC.taxon	distribution	Distribution. Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5C2BBBFF3ED2EEFE14F87E.taxon	description	(Figs 212 E, 217 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5C2BBBFF3ED2EEFE14F87E.taxon	description	Description. ♀. Apterous; body length 2.7 mm. Head brown, except mandible, clypeus and anterior part of frons (with stripe along orbits) testaceous; antenna brown, except scape, pedicel and flagellomeres 7 – 8 testaceous; mesosoma brown, except lateral regions of pronotum testaceous, part of mesoscutum yellow and mesoscutellum, metapectal-propodeal disc and propodeal declivity brown-black; metasoma brown; legs brown, except trochanters testaceous. Antenna clavate; antennomeres in following proportions: 7: 3.5: 8: 5: 4.5: 4: 3: 3: 3: 5. Head excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 1.5; OL = 1.5; OOL = 6. Palpal formula 5 / 3. Pronotum crossed by strong transverse impression, completely shiny, unsculptured. Mesoscutum shiny, unsculptured, with two small pointed apophyses laterally (Fig. 217 C). Mesoscutellum shiny, smooth, slightly sloping anteriorly. Metanotum short, punctate, unsculptured among punctures, weakly excavated behind mesoscutellum (Fig. 217 D). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron shiny, unsculptured. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 10: 2: 3: 9: 15. Enlarged claw (Fig. 212 E) with one small subapical tooth and two proximal bristles. Protarsomere 5 (Fig. 212 E) with two rows of 3 + 11 lamellae situated in distal half; apex with approximately 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5C2BBBFF3ED2EEFE14F87E.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Beroboka Forest, 5.9 km 131 ° SE Ankidra- noka, 22 ° 13.59 ’ S 43 ° 21.59 ’ E, 80 m, 12 – 16. III. 2002, pitfall trap, in tropical dry forest, Fisher, Griswold et al. leg., BLF 6068, (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5C2BBBFF3ED2EEFE14F87E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5C2BBBFF3ED2EEFE14F87E.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBCFF3ED6D7FC72FBB5.taxon	description	(Fig. 212 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBCFF3ED6D7FC72FBB5.taxon	description	Description. ♀. Apterous; body length 2.1 – 2.4 mm. Head testaceous, except vertex darkened; antenna testaceous; mesosoma testaceous; petiole black; metasoma testaceous, except posterior region black; legs testaceous. In specimen from Democratic Republic of the Congo, 04 ° 16 ’ S 17 ° 11 ’ E, head completely testaceous; antenna and mesosoma testaceous; metasoma brown, except first tergite testaceous-whitish; legs testaceous, except club of profemur and protibia partly brown-testaceous. Antenna clavate; antennomeres in following proportions: 6: 4: 4.5: 3.5: 3: 3: 3: 3.5: 3.5: 5.5. Head flat, shiny, unsculptured; temple very prominent; frontal line absent; occipital carina incomplete, only present behind posterior ocell; POL = 1.5; OL = 1; OOL = 6. Palpal formula 4 / 2. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum dull, granulate, without lateral pointed apophyses. Metanotum sloping anteriorly, shiny, unsculptured, without lateral pointed protrusions. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron partly transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 9: 2: 3.5: 7: 13. Enlarged claw (Fig. 212 F) of holotype with one small subapical tooth and 2 – 5 short bristles. In specimen from Democratic Republic of the Congo, 04 ° 16 ’ S 17 ° 11 ’ E, enlarged claw without subapical tooth. Protarsomere 5 (Fig. 212 F) with one row of bristles and 1 – 2 subdistal lamellae; apex with approximately 4 – 5 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBCFF3ED6D7FC72FBB5.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Gauteng, Pretoria Dist., Soutpan, 25 ° 24 ’ S 28 ° 06 ’ E, N. C. Grobbelaar leg. (SANC). Paratype: same locality label as holotype, 1 ♀ (AMNH). Other material: DEM- OCRATIC REPUBLIC OF THE CONGO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 16 ’ S 17 ° 11 ’ E, 6. IV. 2006, seeping on airstrip, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBCFF3ED6D7FC72FBB5.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBCFF3ED6D7FC72FBB5.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBDFF3ED2CAFCE5FA88.taxon	description	(Figs 209 F, G, 215 A, 219 E, 220 A, 224 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBDFF3ED2CAFCE5FA88.taxon	description	Description. ♀ (Fig. 224 A). Apterous; body length 4.5 – 6.2 mm. Head black, except mandible, clypeus and ante- rior part of frons yellow; antenna black, except scape and pedicel yellow; mesosoma black; metasoma black, with brown nuances; legs yellow, except brown spots on coxae, protrochanter, clubs of femora and outer side of protibia. Antennomeres in following proportions: 12: 12: 33: 27: 21: 19: 15: 15: 14: 18. Head (Fig. 209 F) excavated; frons with many longitudinal keels; POL = 3; OL = 4; OOL = 15. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, smooth and setose. Mesoscutum alutaceous, with two lateral pointed apophyses laterally. Metanotum transversely striate, about coplanar to mesoscutellum (Fig. 209 G). Metapectal-propodeal disc shiny, smooth; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete or partly visible. Enlarged claw (Fig. 215 A) with one small subapical tooth and seven peg-like setae. Protarsomere 5 (Fig. 215 A) with inner side proximally serrate and two rows of 1 – 2 (proximal) + 23 – 26 lamellae; apex with approximately 16 – 18 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.4 – 2.8 mm. Head black, except mandible testaceous; antenna brown; mesosoma black; metasoma brown; legs testaceous, except femora and meso- and metatibia partly brown. Antenna filiform; antennomeres in following proportions: 4.5: 5.5: 8: 7: 6.5: 6: 6: 5.5: 6: 10; antennomere 3 less than 3 × as long as broad (8: 3). Head dull, swollen, completely reticulate rugose; frontal line absent; occipital carina absent; POL = 7; OL = 3; OOL = 4; vertex with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas not surrounded anteriorly by high carina). Palpal formula 6 / 3. Mesoscutum dull, granulate. Notauli complete, posteriorly sepa- rated; minimum distance between notauli almost as long as greatest breadth of lateral ocelli (3: 2.8). Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Metapectal-propodeal disc dull, reticulate rugose. Forewing hyaline, without dark transverse bands; 2 r-rs & Rs vein regularly curvilinear, with distal part longer than proximal part (16: 9). Dorsal process of paramere (Figs 219 E, 220 A) long and slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBDFF3ED2CAFCE5FA88.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Busoga Kingdom, Bussu, 1910, E. Bayon leg. (MCSNG). Paratype: same locality label as holotype, 1 ♀ (MRAC) (collecting data has been wrongly reported as “ Bussu- 1909 ” in the original description of Benoit (1951 a) (see Penati & Olmi 2011 )). Other material: MOZAMBIQUE: Gaza Prov., along Palmeira-Macia Road, 2 Km E of Incomati River Bridge, M. Olmi reared from Cicadellid, C. 11. X. 1996, B. 15. X. 1996, Sf. 14. XI. 1996, 1 ♀ (AMNH); same locality label, C. 11. X. 1996, B. 15. X. 1996, Sf. 18. XI. 1996, 1 ♀ (AMNH); same locality label, C. 11. X. 1996, B. 16. X. 1996, Sf. 18. XI. 1996, 1 ♂ (AMNH); same locality label, C. 11. X. 1996, B. 15. X. 1996, Sf. 13. XI. 1996, 1 ♂ (AMNH); same locality label, C. 11. X. 1996, B. 16. X. 1996, Sf. 18. XI. 1996, 1 ♂ (AMNH); Maputo Prov., Catembe, C. 26. XI. 1999, B. 27. XI. 1999, Sf. 7. I. 2000, M. Olmi reared from Cicadellid, 1 ♂ (AMNH); Nampula Prov., Nampula, approximately 500 m from Nampula Catholic Seminary, C. 24. XI. 1999, B. 28. XI. 1999, Sf. 1. I. 2000, reared from nymphs and adults of Exitianus zuluensis Ross, M. Olmi reared, 1 ♀ (MOLC); Niassa, Cuamba, campus of the Faculty of Agriculture of Catholic University of Mozambique, C. 29. V. 2007, B. 5. VI. 2007, Sf. 7. VII. 2007, M. Olmi reared from an adult of Exitianus, 1 ♀ (MOLC). SOUTH AFRICA: Free State, near Bethulie, Tussen Die Riviere Reserve, 30 ° 30 ’ S 26 ° 12 ’ E, 30. III – 3. IV. 1987, 1 ♀ (SANC); Free State, Odendaalsrus caravan park, 27 ° 52 ’ S 26 ° 41 ’ E, 28. I. 1979, B. Lamoral leg., 1 ♀ (DJBC); KwaZulu-Natal, Manguzi Forest edge, 5. I. 1988, day flight, P. Reavell leg., 1 ♀ (NMSA); KwaZulu-Natal, Northern Drakensberg, The Ledges Guest Farm, along the road from R 74 to Cavern Berg Resort, grass in front of the farm, C. 9. IV. 2007, B. 12. IV. 2007, M. Olmi reared from Cicadellid, 1 ♀ (MOLC); KwaZulu-Natal, Winterton, along Rd 600, grass in front of the gate of Cathrin Farm, 10. IV. 2007, M. Olmi reared, 1 ♀ (SAMC); Limpopo, 30 km W Hoedspruit, 27. XI. 1978, D. Brothers & C. Jacot-Guillarmod leg., 1 ♀ (DJBC); Limpopo, Ellisras District, D’Nyala Nature Reserve, 23 ° 45 ’ S 27 ° 49 ’ E, 8 – 12. XII. 1989, 1 ♀ (MOLC); Western Cape, near Citrusdal, The Baths, M. Olmi reared from adult of Exitianus taeniaticeps (Kirschbaum), C. 22. I. 2003, B. 25. I. 2003, Sf. 18. II. 2003, 1 ♀ (MOLC); same locality label, C. 22. I. 2003, B. 26. I. 2003, Sf. 19. II. 2003, 1 ♀ (MOLC); same locality label, C. 22. I. 2003, B. 3. II. 2003, Sf. 25. II. 2003, 1 ♀ (MOLC); Western Cape, Cederberg Mts., Algeria, 6. II. 2006, M. Olmi reared ex Exitianus taeniaticeps (Kirschbaum); 15.5 km N Viliersdorp, along Rd. 43, 13. II. 2006, M. Olmi reared ex Exitianus taeniaticeps (Kirschbaum), 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBDFF3ED2CAFCE5FA88.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Exitianus zuluensis Ross; in South Africa: Exitianus natalensis Ross; Exitianus taeniaticeps (Kirschbaum).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5B2BBDFF3ED2CAFCE5FA88.taxon	distribution	Distribution. Mozambique, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5A2BBFFF3ED32FFEA4FE0C.taxon	description	(Figs 213, 214)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5A2BBFFF3ED32FFEA4FE0C.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; temple without sharp carina; pronotum crossed by strong transverse impression (Fig. 213 B); mesoscutum less than twice as long as broad, with two lateral pointed apophyses (Fig. 213 C); meso-metapleural suture distinct and complete; mesopleuron with lateral pointed apophysis (Fig. 213 C); metapectal-propodeal disc without strong median furrow; protarsomere 1 shorter than 4; enlarged claw with one small subapical tooth (Fig. 214).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5A2BBFFF3ED32FFEA4FE0C.taxon	description	Description. ♀ (Fig. 213). Apterous; body length 4.2 mm. Head brown, except mandible, clypeus, malar space and anterior third of frons testaceous; antenna testaceous; mesosoma black, except lateral regions of pronotum testaceous, mesoscutum mostly yellow, metanotum testaceous, metapectal-propodeal disc and propodeal declivity brown; metasoma brown; proleg testaceous, except club of femur brown and brown spots on coxa, trochanter and tibia; mesoleg testaceous, except brown spot on coxa and club and distal extremity of femur; metaleg testaceous, except brown spot on coxa and distal extremity of femur. Antenna clavate; antennomeres in following proportions: 9: 7: 24: 17: 14: 10: 8: 8: 7: 9. Head (Fig. 213 D) excavated, dull, granulate, with no ADOs; frontal line complete; occipital carina absent; temple distinct; POL = 2; OL = 2; OOL = 9. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum dull, granulate, with two lateral pointed apophyses. Mesoscutellum shiny, unsculptured. Metanotum dull, slightly rugose, hollow behind mesoscutellum (Fig. 213 B). Metapectal-propodeal disc dull, granulate and with many longitudinal striae; propodeal declivity granulate and strongly transversely striate. Mesopleuron partly unsculptured and partly transversely striate, with lateral pointed apophysis (Fig. 213 C). Metapleuron transversely striate. Meso-metapleural suture step shaped, distinct and com- plete. Protarsomeres in following proportions: 20: 4: 6: 25: 36. Enlarged claw (Fig. 214) with one small subapical tooth, 1 seta and 6 peg-like setae. Protarsomere 5 (Fig. 214) with inner side proximally slightly serrate, with two rows of 4 (proximal) + 24 lamellae; apex with at least 12 lamellae. Tibial spurs 1 / 0 / 1.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE5A2BBFFF3ED32FFEA4FE0C.taxon	materials_examined	♂. Unknown Material examined. Type: ♀ holotype, DR 18001: DEMOCRATIC REPUBLIC OF THE CONGO: Ituri, Djugu Territory, Mongbwalu mining camp, abandoned shamba behind old fences, 01 ° 56 ’ 39.32 ’’ N 30 ° 02 ’ 39.46 ’’ E, 75 Moericke traps, 20 – 27. III. 2015, A. Gumovsky leg. (SIZK).. Hosts. Unknown. Distribution. Democratic Republic of the Congo. Etymology. The species is named after the collector, Dr. Alex Gumovsky (Schmalausen Institute of Zoology, Kiev, Ukraine).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	description	(Figs 215 B, 216, 225 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; temple without sharp carina; pronotum crossed by strong transverse furrow; mesoscutum less than twice as long as broad, without lateral pointed apophyses (Fig. 225 A); meso-metapleural suture distinct and complete; mesopleuron with lateral pointed prominence; metapectalpropodeal disc transversely striate, without strong median furrow; protarsomere 1 slightly shorter than 4; enlarged claw (Fig. 215 B) with one small subapical tooth and proximal bristles.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	description	Description. ♀ (Figs 216, 225 A). Apterous; body length 3.2 – 3.9 mm. Head brown, except mandible, clypeus, malar space, anterior surface of frons and ventral side testaceous; antenna testaceous, except antennomeres 7 – 10 darkened; prothorax brown; rest of mesosoma black; metasoma brown; legs brown-yellow. Antenna clavate; anten- nomeres in following proportions: 8: 4: 14: 5: 4: 4: 4: 4: 4: 5. Head (Fig. 216 C) excavated, shiny, unsculptured, except occiput and temple slightly granulate; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 6. Palpal formula 6 / 3. Pronotum crossed by strong transverse furrow, shiny, unsculptured, except lateral regions sculp- tured by longitudinal keels. Mesoscutum shiny, with numerous longitudinal keels, without lateral pointed apophyses (Figs 216 A, 225 A). Mesoscutellum sloping anteriorly. Metanotum sloping anteriorly, transversely striate, exca- vated behind mesoscutellum (Fig. 216 B). Metapectal-propodeal disc and propodeal declivity transversely striate. Mesopleuron transversely striate and with one small lateral pointed apophysis. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres (Fig. 216 D) in following proportions: 10: 2: 4: 12: 19. Enlarged claw (Fig. 215 B) with one small subapical tooth and three proximal bristles. Protarsomere 5 (Fig. 215 B) with inner side not serrate proximally and two rows of 3 (proximal) + 24 lamellae; apex with about nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19464): MADAGASCAR: Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 49.85 ’ S 46 ° 32.17 ’ E, 80 m, 2 – 11. IV. 2004, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 49 (CAS). Paratypes: MADAGASCAR: Mahajanga, Bale de Baly National Park, 12.4 km 337 ° NNW Soalala, 16 ° 00 ’ 36 ’’ S 45 ° 15 ’ 54 ’’ E, 10 m, 26 – 30. XI. 2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6811, 1 ♀ (CAS); Toliara, same locality label as holotype, 20. V – 3. VI. 2003, MA- 02 - 21 - 20, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE582BB0FF3ED03EFBFFF84F.taxon	etymology	Etymology. The species is named after one of the collectors, R. Harin’Hala.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE562BB2FF3ED06FFD5BFD48.taxon	description	(Figs 209 H, I, 215 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE562BB2FF3ED06FFD5BFD48.taxon	description	Description. ♀. Apterous; body length 2.6 – 3.1 mm. Completely yellow-testaceous, except petiole black. Antenna clavate; antennomeres in following proportions: 6: 4: 13: 9.5: 8: 7: 6.5: 6: 6: 8. Head excavated, shiny, unsculptured, with occiput granulate; POL = 1; OL = 1; OOL = 6. In specimen from Uganda, Kawanda, frontal line complete; POL = 1; OL = 1; OOL = 5; greatest breadth of lateral ocelli about as long as OL. Palpal formula of holotype 6 / 3; palpal formula of specimen from Kenya, Embakasi Forest, 5 / 2. Pronotum shiny, slightly granulate, crossed by strong transverse impression. Mesoscutum without lateral pointed apophyses (Fig. 209 H). Metanotum transversely striate, about coplanar to mesoscutellum (Fig. 209 I), with sides rounded. Mesopleuron and metapleuron slightly transversely striate. Metapectal-propodeal disc slender shiny, unsculptured, weakly sloping anteriorly (Fig. 209 I); propodeal declivity slightly transversely striate. Meso-metapleural suture obsolete. Protarsomeres of holotype in following proportions: 13: 2.5: 3: 14: 18. Protarsomeres of specimen from Uganda, Kawanda, in following proportions: 10.5: 1.5: 3: 10: 15. Enlarged claw (Fig. 215 C) with one small subapical tooth and one row of four bristles, or three peg-like setae + one bristle. Protarsomere 5 of holotype (Fig. 215 C) with inner side proximally serrate, with two rows of 2 (proximal) + 14 lamellae (two rows of 1 (proximal) + 19 lamellae in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E); in holotype, apex of protarsomere 5 with approximately 12 lamellae (eight lamellae in specimen from South Africa, 32 ° 24.561 ’ S 25 ° 05.946 ’ E). Protarsomere 5 of specimen from Uganda, Kawanda, with one row of 16 lamellae; apex with about seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE562BB2FF3ED06FFD5BFD48.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, V. 1924, R. E. Turner leg. (NHMUK). Other material: KENYA: Central Prov., Nairobi, Embakasi Forest, 24. II. 1975, W. R. B. Hynd leg., 1 ♀ (NHMUK). SOUTH AFRICA: Eastern Cape, Huntly Glen Farm, 32 ° 24.561 ’ S 25 ° 05.946 ’ E, 1065 m, 10. IV. 2010, sweeping, Great Fish Thicket, HUN 10 - ACA 1 - S 01, S. van Noort leg., 1 ♀ (SAMC). UGANDA: Kawanda, VI. 1943, T. H. C. Taylor leg., 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE562BB2FF3ED06FFD5BFD48.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE562BB2FF3ED06FFD5BFD48.taxon	distribution	Distribution. Kenya, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	description	(Figs 215 D, E, 220 B, C, 224 B, C, 225 B, C, 229 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	description	Description. ♀ (Figs 224 B, C). Apterous; body length 2.0 – 3.5 mm; usually 2.0 – 2.5 mm; 3.5 mm in specimen from South Africa, Blue Mountain Ranch Bed & Breakfast). Head brown-black, except mandible, clypeus and malar space yellow; antenna brown, except antennomeres 1 – 3 yellow; mesosoma and metasoma black; legs yellowbrown, except coxae, clubs of femora and tibiae brownish. Antenna clavate; antennomeres of holotype in following proportions: 6: 4.5: 8.5: 5: 5: 5: 4: 4: 4: 6. Antennomeres of specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, in following proportions: 7: 5: 13: 7: 6: 6: 6: 6: 5: 7. Head (Fig. 229 F) very excavated, shiny, unsculptured, except frons and occiput granulate. Head of specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, with POL = 2; OL = 3; OOL = 7.5. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, shiny, granulate, bare. Mesoscutum dull, with some longitudinal keels. Metanotum transversely striate. Metapectal-propodeal disc dull, granulate, often with slight longitudinal striae; propodeal declivity transversely striate. Meso- and metapleuron transversely striate. Occasionally top of metapectal-propodeal disc with track of median longitudinal furrow. Meso-metapleural suture obsolete; rarely track of meso-metapleural suture slightly visible. Protarsomeres of specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, in following proportions: 13.5: 3: 4: 15: 21. Enlarged claw (Figs 215 D, E) with one small subapical tooth and 5 – 6 peg-like setae. Protarsomere 5 (Fig. 215 D) with inner side proximally serrate and one row of 15 – 19 lamellae; apex with 7 – 9 lamellae (in specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, apex with 20 lamellae (Fig. 215 E )). Tibial spurs 1 / 0 / 1. ♂ (Figs 225 B, C). Fully winged; body length 1.3 – 1.8 mm. Head brown, except mandible testaceous; antenna brown; mesosoma and metasoma brown; legs brown. Antenna filiform; antennomeres in following proportions: 3: 3.5: 4: 4: 4: 4: 3.5: 3.5: 3.5: 4.5; antennomere 3 less than 3 × as long as broad (2.75). Head shiny, setose, slightly granulate; frontal line absent; occipital carina absent; POL = 4; OL = 1.5; OOL = 2.5; greatest breadth of lateral ocelli about as long as OOL; vertex with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas anteriorly surrounded by high carina). Palpal formula 4 / 2. Mesoscutum shiny, setose, slightly granulate. Notauli often absent; in specimens reared in Nigeria, IITA, notauli incomplete and reaching approximately 0.25 – 0.30 × length of mesoscutum; in specimens from Mozambique, Maputo, notauli incomplete and reaching approximately 0.5 × length of mesoscutum. Mesoscutellum and metanotum shiny, setose, unsculptured. Metapectal-propodeal disc shiny, unsculptured, with median longitudinal furrow; in specimens reared in Nigeria, IITA, metapectal-propodeal disc sculptured by irregular striae and unsculptured among striae. Forewing hyaline, without dark transverse bands, only with costal cell distinctly enclosed by pigmented veins (other basal cells enclosed by fading veins). Dorsal process of paramere (Figs 220 B, C) long and narrow. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Kawanda, 7. VII. 1943, T. H. C. Taylor leg. (NHMUK). Paratypes: TANZANIA: Pongwe, 1935, ex egg laid parthenogenetically by following ♀, Kirkpatrick leg., 1 ♂ (NHMUK); same locality label, 1 ♀ (AMNH). Other material: BURKINA FASO: Saria, 1 ♀ (AMNH). CAM- EROON: Maroua, Banks of Mayo Makabay River, 1 ♀ (CIRAD). CAPE VERDE ISLANDS: Cape Verde Is- lands, 8 ♀♀ (AMNH). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Katanga, Lubumbashi, II. 1989, ex Cicadulina? mbila, M. S. Alam leg., Bosque-Perez CIE A 20475, 1 ♀ (NHMUK); Ituri Prov., Mongbwalu AGK camp, 01 ° 56 ’ 32.81 ’’ N 30 ° 02 ’ 25.62 ’’ E, 21 - 28. II. 2015, 75 YPT, behind the former President Mobutu’s Doctor’s house, A. Gumovsky leg., 1 ♀ (SIZK); Oriental Province, Ituri Region, Mongbwalu, AGK Camp, 01 ° 56 ’ 34.12 ’’ N 30 ° 02 ’ 24.85 ’’ E, nursery, just short weeds, 75 Yellow Pan Traps, 21 – 28. II. 2015, A. Gumovsky leg., 1 ♀ (SIZK). GAMBIA: Western Division, Kombo St Mary, Bakau, 1 ♀ (KJHC); Western Division, Kombo St Mary, Jeswang, 1 ♀ (KJHC). GUINEA-BISSAU: Região Bafatá, 5 km W of Bijine, 2. XI. 1992, 1 ♀ (ZIL); Região Gabú, 7 km NE of Gabú, 3. XI. 1992, 1 ♀ (ZIL). MADAGASCAR: Toliara, 12 km NW of Amboasary, Berenty, 1 ♀ (AMNH). MOZAMBIQUE: Maputo Prov., Maputo, 2 ♀♀ (AMNH, USNM); Maputo Prov., 26 km S of Maputo, Umbelúzi, 4 ♀♀ (AMNH); Niassa, Cuamba, M. Olmi reared from Cicadellid, C. 8. XI. 1999, B. 9. XI. 1999, Sf. 27. XI. 1999, 1 ♀ (MOLC); same locality label, C. 8. XI. 1999, B. 10. XI. 1999, Sf. 28. XI. 1999, 1 ♀ (MOLC); same locality label, 8. XI. 1999, sweeping, M. Olmi leg., 1 ♀ (MOLC); Cuamba, Campus of the Catholic University of Mozambique, C. 6. III. 2005, B. 7. III. 2005, Sf. 24. III. 2005, M. Olmi reared from Cicadellid, 1 ♀ (MOLC). NIGERIA: Ibadan, In- ternational Institute of Tropical Agricolture (IITA), IX. 1989, ex Cicadulina triangula, 1 ♀, 1 ♂ (AMNH). OMAN: Salalah, 1 ♀ (PMA). REPUBLIC OF THE CONGO: Pool, Kindamba, Kisanga, 4 ♀♀ (AMNH). SIERRA LEONE: Makeni, 12 ° 03 ’ W 08 ° 53 ’ N, 28. XI. 1993, 1 ♀ (ZIL). SOUTH AFRICA: Eastern Cape, 16 km E of Port Elizabeth, S. George’s Strand, 1 ♀ (AMNH); Free State, Harrismith, 1 ♀ (NHMUK); Free State, Sasolburg, Wonderwater Colliery, 26 ° 45.37 ’ S 27 ° 48.31 ’ E, 1450 m, 23. XI. 1999, D-Vac, IS- 18, entrance garden, translocated rehab., artificial marsh (IV. 1993), game park simulation, dom. Aristida junciformis, M. Stiller leg., 1 ♀ (SANC); Free State, NE Kroonstad, Heuningspruit, 27 ° 26 ’ S 27 ° 22 ’ E, 1400 m, 4. III. 2002, DVac off grass, common: Eragrostis gummuflua (Poaceae), and forbs, M. Stiller & J. du Plessis leg., 1 ♀ (SANC); KwaZulu-Natal, St. Lucia Estuary, 25. I. 1977, D. J. Brothers leg., 1 ♀ (DJBC); KwaZulu-Natal, Northern Drakensberg, 4 km S of Cavern Berg Resort, along the road from R 74 to Cavern Berg Resort, C. 9. IV. 2007, B. 13. IV. 2007, M. Olmi reared, 1 ♀ (MOLC); Western Cape, Swellendam, Blue Mountain Ranch Bed & Breakfast, C. 29. I. 2003, B. 6. II. 2003, Sf. 5. II. 2003, M. Olmi reared from Exitianus taeniaticeps (Kirschbaum), 1 ♀ (MOLC). YEMEN: Sana’a, III. 1991, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Burkina Faso: Cicadulina similis China; in Democratic Republic of the Congo: Cicadulina mbila (Naudé); in Nigeria: Cicadulina storeyi China (= Cicadulina triangula Ruppel); in South Africa: Exitianus taeniaticeps (Kirschbaum); in Tanzania: Cicadulina bipunctata (Melichar) (= Cicadulina zeae China). Natural enemies. Hymenoptera Encyrtidae: in Mozambique: Cheiloneurus gonatopodis R. Perkins. This Encyrtid is one of the most common natural enemies of Dryinids in Mozambique (see also Echthrodelphax tauricus Ponomarenko, Gonatopus amoenus Olmi and Gonatopus mossambicus Olmi). The reared specimens have the following locality labels: Mozambique, Maputo Prov., Maputo, municipal plant nursery, parasitized leafhopper collected on 10. X. 1993; cocoon spun on 11. X. 1993; encyrtids emergence on 27. X. 1993. 3 ♂♂, 4 ♀♀.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	distribution	Distribution. Burkina Faso, Cameroon, Cape Verde Islands, Democratic Republic of the Congo, Gambia, Guinea-Bissau, Madagascar, Mozambique, Nigeria, Oman, Republic of the Congo, Sierra Leone, South Africa, Tanzania, Uganda, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE552BB3FF3ED56FFB19F91C.taxon	discussion	Remarks. The ♂ paratype of G. incognitus was obtained parthenogenetically by Mr. Kirkpatrick from an egg laid by the ♀ paratype in Tanzania; this means that in this species parthenogenesis is arrhenotokous.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE542BB4FF3ED0A3FD09FC90.taxon	description	(Fig. 222 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE542BB4FF3ED0A3FD09FC90.taxon	description	Description. ♀. Apterous; body length 2.5 – 3.4 mm. Head black, except mandible and clypeus yellow; antenna brown, except scape, pedicel and part of flagellomere 1 yellow; mesosoma black, except distal extremity of propo- deal declivity reddish; metasoma black; legs testaceous, except coxae and club of profemur brown or black. Head excavated, shiny, unsculptured, except occiput granulate; POL = 2; OL = 8; OOL = 23. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured, except lateral regions granulate. Mesoscutum with two pointed apophyses laterally. Metanotum transversely striate, excavated behind mesoscutellum; sides of metanotum protruding; lateral protrusions of metanotum rounded. Metapectal-propodeal disc shiny, unsculptured or slightly alutaceous, with deep median longitudinal furrow; propodeal declivity transversely striate. In specimen from Botswana, 19 ° 37 ’ S 23 ° 47 ’ E, metapectal propodeal disc slightly granulate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (13: 15). Enlarged claw (Fig. 222 A) with one small subapical tooth and eight peg-like setae. Protarsomere 5 (Fig. 222 A) with two rows of 4 (proximal) + 21 lamellae; apex with 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE542BB4FF3ED0A3FD09FC90.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 5 – 30. IV. 1923, R. E. Turner leg. (NHMUK). Paratype: NAMIBIA: 30 mi. SE Keetmanshoop, 1 ♀ (AMNH). Other material: BO- TSWANA: Smiti Mopane Woddland, 19 ° 37 ’ S 23 ° 47 ’ E, SE corner of swamps, S of the Moxemi Wildlife Reserve, 4 – 11. IX. 1975, Pitfall traps, A. Russel-Smith leg., 2 ♀♀ (AMG). NAMIBIA: Namib / Naukluft Park, near Gobabeb, Kuiseb River, 1 ♀ (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE542BB4FF3ED0A3FD09FC90.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE542BB4FF3ED0A3FD09FC90.taxon	distribution	Distribution. Botswana, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE532BB4FF3ED527FE27F8EA.taxon	description	(Figs 222 B, 226 A-D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE532BB4FF3ED527FE27F8EA.taxon	description	Description. ♀ (Figs 226 A-D). Apterous; body length 3.5 – 4.0 mm. Head brown-black, except mandible, clypeus, anterior half of frons (with short stripe along orbits) and ventral side whitish; antenna brown, except scape, pedicel and proximal third of flagellomere 1 whitish, antennomere 10 testaceous; mesosoma black, except posterior third of mesoscutum yellow; metasoma brown-testaceous; proleg brown, except part of coxa, trochanter, stalk of femur, part of tibia and part of chela testaceous; mesoleg brown, except part of coxa, trochanter, part of femur and tibia, tarsus testaceous; metaleg brown, except distal extremity of coxa, trochanter, part of femur, tibia and tarsus testaceous. Antenna clavate; antennomeres in following proportions: 9: 6: 21: 12: 9: 9: 7: 6: 5: 9. Head excavated, shiny, unsculptured; frontal line incomplete, absent in anterior third of frons; occipital carina absent; POL = 2.5; OL = 2; OOL = 10. Palpal formula 6 / 3. Pronotum crossed by strong transverse furrow, shiny, unsculptured. Mesoscutum shiny, sculptured by numerous longitudinal striae, laterally with two strong pointed apophyses situated on sides of mesoscutellum (Figs 226 A, C). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum very long, transversely striate and excavated behind mesoscutellum (Fig. 226 B), with sides protruding (protrusions rounded) (Figs 226 A, C). Metapectal-propodeal disc shiny, sculptured by many longitudinal striae, unsculptured among striae; propodeal declivity, meso- and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 17: 3: 6: 20: 31. Protarsomere 3 produced into hook. Enlarged claw (Fig. 222 B) with one small subapical tooth and one row of four small lamellae + one bristle. Protarsomere 5 (Fig. 222 B) with two rows of 1 (proximal) + 20 lamellae; apex with approximately 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE532BB4FF3ED527FE27F8EA.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Kibale National Park, Kanyawara, Makerere University Biological Field Station, 0 ° 33.871 ’ N 30 ° 21.355 ’ E, 1495 m, 4 – 26. VIII. 2008, YPT, primary mid-altitude rainforest, S. van Noort leg., SAM-HYM-A 023422 (SAMC). Paratype: same locality label as holotype, SAM-HYM-A 023422, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE532BB4FF3ED527FE27F8EA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE532BB4FF3ED527FE27F8EA.taxon	distribution	Distribution. Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB5FF3ED6D7FE31FC6D.taxon	description	(Fig. 222 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB5FF3ED6D7FE31FC6D.taxon	description	Description. ♀. Apterous; body length 2.2. Head brown, except mandible, clypeus and anterior region of frons testaceous; antenna testaceous; mesosoma testaceous-reddish, except propleuron and mesopleuron darkened; metasoma brown; legs testaceous, except coxae and club of metafemur brown. Antenna clavate; antennomeres of holotype in following proportions: 6: 4: 5.5: 4: 3: 3: 2.5: 4: 3: 5. Head excavated, shiny, slightly granulate; temple distinct; frontal line incomplete, only present in posterior half of frons; occipital carina absent; POL = 1; OL = 1; OOL = 6.5; greatest breadth of lateral ocelli about as long as OL. Labial palpi 2 – segmented; maxillary palpi of holotype missing. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum less than twice as long as broad, shiny, without lateral pointed apophyses. Mesoscutellum shiny, smooth, circular, weakly sloping anteriorly. Metanotum very short, sloping anteriorly, transversely striate, about coplanar to mesoscutellum. Metapectal-propodeal disc shiny, with median longitudinal furrow, with few median longitudinal striae; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 9: 2: 3: 8: 14. Enlarged claw (Fig. 222 C) with one small subapical tooth, four large lamellae and one bristle. Protarsomere 5 (Fig. 222 C) with one row of 12 lamellae; apex with about eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB5FF3ED6D7FE31FC6D.taxon	materials_examined	Material examined. Type: ♀ holotype: NAMIBIA: 17 km S Karibib, along Road C 32, 8. III. 2006, ♀ holotype attacking Paradorydium spatulatum (Naudé), 24 A, M. Olmi leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB5FF3ED6D7FE31FC6D.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Namibia: Paradorydium spatulatum (Naudé).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB5FF3ED6D7FE31FC6D.taxon	distribution	Distribution. Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	description	(Figs 218, 220 D, 222 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 4 / 2; pronotum crossed by strong transverse furrow; head deeply excavated; temple without sharp carina; mesoscutum very long, approximately 3.0 × as long as broad; mesometapleural suture distinct and complete; protarsomere 1 slightly shorter than 4; enlarged claw (Fig. 222 D) with one small subapical tooth and one row of peg-like setae. ♂ with palpal formula 6 / 3; region of head between lateral ocelli and eyes with ovoidal area anteriorly delimited by strong and high carina; antennomere 3 less than 3 × as long as broad; notauli complete; dorsal process of paramere large and with apex broadened (Fig. 220 D).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	description	Description. ♀ (Fig. 218). Apterous; body length 4.1 mm. Head black, except mandible testaceous; antenna black, except ventral side of antennomere 1 whitish; mesosoma and metasoma black; legs brown. Antenna clavate, without ADOs; antennomeres in following proportions: 10: 6: 15: 11: 8: 7: 6: 6: 5: 7. Head deeply excavated, with frons shiny, unsculptured; temple prominent, without sharp carina, finely punctate, unsculptured among punctures; frontal line complete; occipital carina very shortly present behind lateral ocelli; POL = 0.5; OL = 1; OOL = 8. Palpal formula 4 / 2. Pronotum shiny, finely punctate, unsculptured among punctures, crossed by strong transverse furrow. Mesoscutum slender, about 3 × as long as broad (14: 5), shiny, finely punctate, unsculptured among punctures, without lateral pointed apophyses. Metanotum transversely striate, weakly excavated behind mesoscutellum. Sides of metanotum laterally protruding; protrusions rounded. Metapectal-propodeal disc subspheroidal, shiny, finely punctate, unsculptured among punctures; propodeal declivity strongly transversely striate. Mesopleuron finely transversely striate. Metapleuron strongly transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 3: 5: 16: 24. Enlarged claw (Fig. 222 D) with one small subapical tooth and one row of 6 peg-like setae. Protarsomere 5 (Fig. 222 D) with inner side proximally not serrate and one row of 17 lamellae; apex with approximately 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.6 mm. Head black, except mandible testaceous; mesosoma black; metasoma brown; legs brown, except tarsomeres 1 – 4 and protibia testaceous – brown. Antenna filiform; antennomeres in following proportions: 6: 4: 8: 8: 7: 7: 7: 6.5: 6.5: 8; antennomere 3 less than 3 × as long as broad (8: 3). Head setose, dull, granulate and slightly rugose; frons without frontal line, with shiny and unsculptured central longitudinal stripe from anterior ocellus to clypeus; POL = 9; OL = 3; OOL = 5; greatest breadth of lateral ocelli slightly shorter than OL (2: 3); vertex with shiny, unsculptured, ovoidal area between lateral ocelli and eyes, bordered anteriorly by strong and high carina (as in Fig. 208 G); temple distinct. Palpal formula 5 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli about as long as OL. Mesoscutellum and metanotum shiny, finely punctate, unsculptured among punctures. Metapectal-propodeal disc dull, reticulate rugose, without transverse or longitudinal keels; top with central longitudinal furrow. Forewing hyaline, without dark transverse bands or spots; distal part of 2 r-rs & Rs vein slightly longer than proximal part (13: 11); costal cell surrounded by strongly pigmented; other basal veins diaphanous. Dorsal process of paramere (Fig. 220 D) shorter than volsella and with distal half broadened. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Koeberg Nature Reserve, near dams, 33.637152 ° S 18.414520 ° E, 8. XII. 2014, sweeping, M. Olmi leg., SAM-HYM-P 086344 (SAMC). Paratype: same locality label as holotype, C. 8. XII. 2014, B. 11. XII. 2014, Sf. Summer 2015, reared from a nymph of Strongylodemas breviceps Fennah, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	biology_ecology	Hosts. Fulgoridae, Strongylodematinae: Strongylodemas breviceps Fennah (A. Emeljanov det.). Strongylodematinae belong to the family Fulgoridae (Emeljanov, personal comm.) and not Dictyopharidae, as attributed by other authors.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	etymology	Etymology. The species is named koebergensis after the collection site in the Koeberg Nature Reserve.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE522BB6FF3ED272FCB8F871.taxon	discussion	Remarks. The chela of G. koebergensis has one small subapical tooth on the enlarged claw. Additionally, its antenna lacks ADOs. Normally, species with this combination of characters parasitize Cicadomorpha. On the contrary, G. koebergensis parasitizes Fulgoridae (Fulgoromorpha).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE502849FF3ED0CBFCC4FEE0.taxon	description	(Fig. 222 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE502849FF3ED0CBFCC4FEE0.taxon	description	Description. ♀. Apterous; body length 2.6 – 3.3 mm. Head brown, except mandible, clypeus and short frontal medi- an longitudinal stripe near clypeus testaceous (in specimen from Central African Republic, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, head brown, except mandible, clypeus and anterior third of frons testaceous); antenna testaceous-darkened, except ventral side of antennomere 1 yellow; mesosoma brown, except mesoscutum, metanotum and apex of propodeal declivity testaceous-darkened (in specimen from Central African Republic, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, mesosoma brown-black); petiole black; metasoma brown; legs brown, with tarsi and articulations testaceous-darkened. An- tenna clavate; antennomeres in following proportions: 7: 5: 13: 7: 6: 5.5: 5: 5: 4: 6. Head flat, dull, granulate; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 7; greatest breadth of lateral ocelli about as long as OL. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, with dorsal surface shiny and unsculptured and lateral regions shiny and slightly granulate. Mesoscutum slightly granulate, with two lateral pointed apophyses at border with mesoscutellum. Metanotum granulate, excavated behind mesoscutellum. Metapectal-propodeal disc dull, granulate; propodeal declivity granulate, not transversely striate (only with few transverse striae at posterior extremity). In specimen from Central African Republic, 3 ° 00.27 ’ N 16 ° 11.55 ’’ E, top of metapectal-propodeal disc with track of longitudinal furrow. Mesopleuron and metapleuron granulate, not transversely striate. Meso-metapleu- ral suture distinct and complete. Protarsomeres in following proportions: 12: 2: 5: 16: 24. Enlarged claw (Fig. 222 E) with one small subapical tooth and 3 – 4 proximal bristles. Protarsomere 5 (Fig. 222 E) with two rows of 3 (proximal) + 19 – 22 lamellae; apex with nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE502849FF3ED0CBFCC4FEE0.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Pietermaritzburg, Hilton, 19 – 21. IV. 2005, YPT, V. Kolyada leg. (NMSA). Other material: CENTRAL AFRICAN REPUBLIC: Prefecture Sangha-Mbaéré, Reserve Speciale de forêt dense de Dzanga-Sangha, 12.7 km 326 ° NW Bayanga, 3 ° 00.27 ’ N 16 ° 11.55 ’ E, 420 m, 11 – 12. V. 2001, MT, lowland rainforest, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE502849FF3ED0CBFCC4FEE0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FE502849FF3ED0CBFCC4FEE0.taxon	distribution	Distribution. Central African Republic, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE2849FF3ED7F7FCB8F9F6.taxon	description	(Figs 223 A, 229 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE2849FF3ED7F7FCB8F9F6.taxon	description	Description. ♀ (Fig. 229 A). Apterous; body length 3.7 – 4.5 mm. Head black, except mandible, clypeus and ante- rior region of frons (more along orbits) reddish (in specimen from Tanzania, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, occiput and temple partly ferruginous); antenna brown, except scape and pedicel yellow or testaceous; pronotum black, with reddish nuances; rest of mesosoma and metasoma black (in specimen from Tanzania, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, mesosoma completely ferruginous-testaceous and metasoma testaceous, with brown transverse stripes); legs yellow, except meso- and metacoxa and clubs of pro- and metafemur brownish (in specimen from Tanzania, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, legs completely ferruginous). Head excavated, shiny, bare, smooth, with some short keels near the ocelli; lateral regions of occiput rugose, setose; temple sticking out, reticulate rugose and crossed by longitudinal keel. Antenna clavate; antennomeres of holotype in following proportions: 10: 10: 22: 21: 17: 13: 11: 10: 10: 11; antennomeres of specimen from Tanzania, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, in following proportions: 8: 10: 18: 17: 14: 12: 9: 8: 7: 9. Third antennomere as broad as 2. POL = 1; OL = 2; OOL = 8. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, granulate. Mesoscutum very long, about 3 × as long as broad (15: 5), dull, granulate. Mesoscutellum shiny, smooth, circular. Stalk between pronotum and metanotum (Fig. 229 A) composed of mesoscutum + mesoscutellum (in almost all other Gonatopus species, stalk only composed of mesoscutum), about 4 × as long as broad (20: 5). Metanotum very long, transversely striate, not excavated behind mesoscutellum. Metapectal-propodeal disc dull, with four median longitudinal keels, and, in some specimens, with track of median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 slightly longer than 4 (17: 16). Protarsomeres of specimen from Tanzania, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, in following proportions: 18: 3: 4.5: 17: 26. Enlarged claw (Fig. 223 A) with one small subapical tooth and 5 – 6 lamellae or peg-like setae. Protarsomere 5 (Fig. 223 A) with one row of 17 – 23 lamellae; apex with 11 – 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE2849FF3ED7F7FCB8F9F6.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Bekily, IV. 1933, A. Seyrig leg. (MNHN). Other material: LESOTHO: Mamathes, 10. II. 1951, A. Jacot-Guillarmod leg., 2 ♀♀ (NHMUK, AMNH). MAURI- TANIA: 32 km S Nouakchott, 6. XI. 1993, 1 ♀ (AMCR). TANZANIA: Mkomazi Game Reserve, Inselberg between Mabata and Kamakota Hill, 04 ° 15.60 ’ S 38 ° 26.35 ’ E, sweep, Ficus glumosa & mixed grasses in Acacia / Commiphora bushland, 4. XII. 1995, S. van Noort leg., SAM-HYM-PO 16879, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE2849FF3ED7F7FCB8F9F6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE2849FF3ED7F7FCB8F9F6.taxon	distribution	Distribution. Lesotho, Madagascar, Mauritania, Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE284AFF3ED08AFE1FFD00.taxon	description	(Figs 223 B, 229 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE284AFF3ED08AFE1FFD00.taxon	description	Description. ♀. Apterous; body length 4.4 mm. Head testaceous-reddish; antenna brown, except scape and pedicel testaceous; pronotum and mesoscutum testaceous-reddish, except lateral regions darkened; rest of mesosoma black, except distal extremity of propodeal declivity reddish and some reddish nuances on metanotum; petiole black; metasoma brown-reddish; legs testaceous-reddish. Head very excavated, dull, granulate; occipital carina incomplete, only shortly present on sides of lateral ocelli; temple very long; frontal line hardly visible, apparently complete; POL = 3; OL = 5; OOL = 21. Palpal formula 4 / 2. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum very long and slender (Fig. 229 B), about 3 – 4 × as long as broad, dull, granulate, without lateral pointed apophyses. Mesoscutellum not visible. Metanotum with sides rounded, about coplanar to mesoscutellum. Metapectal-propodeal disc dull, granulate; propodeal declivity granulate, not transversely striate; mesopleuron and metapleuron not transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 slightly shorter than 4 (15: 17). Enlarged claw (Fig. 223 B) with one small subapical tooth and one row of seven peg-like setae. Protarsomere 5 (Fig. 223 B) with inner side proximally serrate, with one row of 17 lamellae; apex with approximately 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE284AFF3ED08AFE1FFD00.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, 1 – 14. V. 1923, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE284AFF3ED08AFE1FFD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAE284AFF3ED08AFE1FFD00.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284AFF3ED497FD22F917.taxon	description	(Figs 223 C, D, 229 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284AFF3ED497FD22F917.taxon	description	Description. ♀. Apterous; body length 1.6 – 3.1 mm. Head brown, except clypeus, mandible and anterior region of frons yellow; antenna yellow, except antennomeres 4 – 10 brown; mesosoma black; metasoma black, with some brown nuances; legs yellow, except clubs of femora brownish and both extremities of meso- and metatibia brownish (legs appear to be banded such as in ♀♀ of group 1). Head (Fig. 229 C) weakly excavated, almost flat, shiny, granulate; POL = 3; OL = 5; OOL = 15. Palpal formula 4 / 2 or 5 / 2. Pronotum crossed by strong transverse impression, shiny, granulate. Mesoscutum dull, strongly granulate, without lateral pointed apophyses. Metanotum dull, granulate, with some transverse striae. Metapectal-propodeal disc dull, completely granulate; propodeal declivity granulate, not transversely striate. Mesopleuron and metapleuron not transversely striate. Meso-metapleural suture obsolete. Enlarged claw (Fig. 223 C) with one small subapical tooth and with or without bristles, or small lamellae, or peg-like setae. Protarsomere 5 of holotype (Fig. 223 C) with inner side proximally not serrate and one row of one long lamella + four very short lamellae, in addition to many bristles; apex with four lamellae, including one very long. In specimen from South Africa, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, Protarsomere 5 with five lamellae and many bristles; apex with five lamellae, including one very long. In specimen from Namibia, Okahandja, protarsomere 5 (Fig. 223 D) with one row of 20 lamellae and group of eight distal lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284AFF3ED497FD22F917.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Central Prov., Olgasalic, IV. 1945, S. Patrizi leg. (MRAC). Other material: NAMIBIA: Okahandja, 19 – 29. XII. 1927, R. E. Turner leg., 1 ♀ (NHMUK). SOUTH AFRICA: Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 29. IX – 30. XII. 2010, MT, Hantam Succulent Karoo, KEL 09 - SUC 1 - M 05, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284AFF3ED497FD22F917.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284AFF3ED497FD22F917.taxon	distribution	Distribution. Kenya, Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	description	(Fig. 223 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	description	Description. ♀. Apterous; body length 6.1 mm. Head black, except anterior region of frons, clypeus and mandible testaceous; antenna brown, except scape, pedicel and proximal third of flagellomere 1 testaceous; mesosoma and metasoma black; legs black, except junctions, tarsi and part of protibia testaceous. Antenna clavate, without ADOs; antennomeres in following proportions: 13: 7: 36: 14: 10: 19: 13: 12: 10: 14. Head excavated, dull, with long sparse setae, with frons granulate and sculptured by many longitudinal keels; temple distinct and strongly granulate and slightly rugose; frontal line complete; occipital carina absent; POL = 2; OL = 2; OOL = 10. Palpal formula 6 / 3. Pronotum dull, crossed by strong transverse impression, strongly granulate and slightly rugose, with long sparse setae. Mesoscutum dull, rugose, granulate and sculptured by strong longitudinal keels, with two lateral pointed apophyses situated on sides of mesoscutellum, with long sparse setae. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum about coplanar to mesoscutellum, sloping anteriorly, granulate and reticulate rugose. Metapectal-propodeal disc dull, with long sparse setae, reticulate rugose and granulate; propodeal declivity strongly transversely striate, granulate and partly reticulate rugose. Mesopleuron dull, strongly transversely striate and granulate. Metapleuron dull, reticulate rugose. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 26: 4: 8: 21: 35. Enlarged claw (Fig. 223 E) without subdistal tooth, with one row of four lamellae and one bristle, with proximal half very broad. Protarsomere 5 (Fig. 223 E) with one row of 10 lamellae; apex with approximately 30 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Niassa, Cuamba, Mozambique Catholic University Campus, 10. IV. 2002, on grass, M. Olmi leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	biology_ecology	Hosts. Unknown. However, the ♀ holotype was seen to attack unidentified planthoppers belonging to Meeno- plidae, Delphacidae and Tropiduchidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAD284BFF3ED0ACFE8EFC04.taxon	discussion	Remarks. The ♀ of this species has no antennal ADOs. Despite this character, it attacks Fulgoromorpha and not Cicadomorpha.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	description	(Figs 223 F, 229 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	description	Description. ♀. Apterous; body length 2.1 – 2.6 mm. Head black, except mandible and borders of clypeus yellow; antenna brown, except ventral side of antennomere 1 yellow; mesosoma black; metasoma brown; legs brown, except tarsi and articulations testaceous. Head excavated, shiny, unsculptured, except vertex and temple slightly granulate; frontal line complete; occipital carina shortly present behind and on sides of the ocellar triangle; POL = 1; OL = 2; OOL = 7; temple very long, slightly shorter than half of eyes (7: 18). Palpal formula 5 / 2. Antenna clavate; antennomeres in following proportions: 5: 4: 11: 7: 6: 5.5: 5: 5.5: 4.5: 7. Pronotum crossed by strong transverse impression, shiny, setose, slightly granulate, with anterior collar partly unsculptured. Mesoscutum dull, granulate, with two strong lateral pointed apophyses (Fig. 229 D). Mesoscutellum very sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 229 E). Metapectal-propodeal disc dull, slightly granulate, with strong median longitudinal furrow; propodeal declivity strongly transversely striate. In paratype, propodeal declivity rugose and with undistinctly visible transverse striae. Mesopleuron slightly granulate, not transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 11: 2: 3: 14: 21. Enlarged claw (Fig. 223 F) with one small subapical tooth and six peg-like setae (in holotype) or five peg-like setae + one bristle (in paratype). Protarsomere 5 (Fig. 223 F) with inner side proximally not serrate and two rows of 5 (proximal) + 21 lamellae (in holotype) or 3 (proximal) + 16 lamellae (in paratype); apex with at least 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Western Cape, West Coast National Park, about 2 km S Dainepos Rd. Junction, 2. II. 2006, M. Olmi leg. (attacking adults of Colistra n. sp.), SAM-HYM-P 024976 (SAMC). Paratypes: NAMIBIA: 51 km S Grünau, along Rd B 1, C. 1. III. 2006, B. 5. III. 2006, Sf. 5. IV. 2006, reared from a ♀ adult of Aconurella compta (Naudé), M. Olmi leg., 1 ♀ (MOLC); 40 km W Keetmanshoop, along Rd B 4, C. 10. III. 2006, B. 14. III. 2006, Sf. 11. IV. 2006, reared from an adult of Aconurella compta (Naudé), M. Olmi leg., 27 T, 1 ♀ (MOLC); 6.5 km W Aus, along Rd B 4, C. 12. III. 2006, B. 14. III. 2006, Sf. 13. IV. 2006, reared from an adult of Aconurella compta (Naudé), M. Olmi leg., 28 A, 1 ♀ (MOLC). About further paratypes, see remarks.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in South Africa: Colistra sp.; in Namibia: Aconurella compta (Naudé). About Agallia cuneata Cogan (= Austroagallia cuneata (Cogan )), considered by mistake host of G. maritimus by Olmi (2007 b) and Guglielmino & Olmi (2007), see remarks.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	distribution	Distribution. Namibia, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAC284CFF3ED59BFEB1FDC7.taxon	discussion	Remarks. The following three ♀ paratypes were attributed by mistake to G. maritimus: (Guglielmino & Olmi 2007; Olmi 2007 b): NAMIBIA: 27.6 km S Omaruru, along Rd C 33, C. 8. III. 2006, B. 12. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 A, 1 ♀ paratype (MOLC); 27.6 Km S Omaruru, along Rd C 33, C. 8. III. 2006, B. 13. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 B, 1 ♀ paratype (SMWN); 27.6 km S Omaruru, along Rd C 33, C. 8. III. 2006, B. 11. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 C, 1 ♀ paratype (MOLC). The above three specimens are now attributed to G. okahandjae Olmi (see Material examined for this species).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284CFF3ED4DFFD9BFA65.taxon	description	(Figs 223 G, 228 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284CFF3ED4DFFD9BFA65.taxon	description	Description. ♀ (Fig. 228 A). Apterous; body length 3.1 – 3.7 mm. Holotype and paratype ferruginous, except peti- ole black, metasoma mostly brown, antennomeres 6 – 10 darkened, legs partly darkened; in specimen from South Africa, 22 ° 27 ’ S 31 ° 17 ’ E, head and mesosoma ferruginous-brown. Antenna clavate; antennomeres in following proportions: 11: 6: 11: 6: 6: 6: 4.5: 4.5: 4.5: 7.5. Head flat, dull, granulate; frontal line absent (only track); occipital carina almost completely absent (shortly present on sides of lateral ocelli, not reaching eyes); occiput dull, granulate, setose; POL = 3.5; OL = 3.5; OOL = 9; OPL = 0.5; TL = 4. Palpal formula 6 / 3. Pronotum, crossed by strong transverse impression, with anterior collar and disc dull, granulate. Mesoscutum slightly longer than broad (11: 10), shiny, sculptured by longitudinal striae, laterally without pointed apophyses. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum short, longitudinally striate and excavated behind mesoscutellum. Metapectal-propodeal disc dull, granulate; propodeal declivity, mesopleura and metapleura transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 3: 5: 16: 24. Enlarged claw (Fig. 223 G) with one small subapical tooth and one row of four peg-like setae, in addition to one bristle. Protarsomere 5 (Fig. 223 G) with two rows of 3 (proximal) + 20 lamellae; apex with approximately 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284CFF3ED4DFFD9BFA65.taxon	materials_examined	Material examined. Types: ♀ holotype: BOTSWANA: Serowe, MT, VIII. 1989, P. Forchhammer leg. (CAS). Paratype: same locality label as holotype, VI. 1988, 1 ♀ (MOLC). Other material: SOUTH AFRICA: Limpopo, Krüger National Park, Pafuri, 22 ° 27 ’ S 31 ° 17 ’ E, 2 – 13. II. 1980, MT, L. E. O. Braack leg., 1 ♀ (DJBC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284CFF3ED4DFFD9BFA65.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284CFF3ED4DFFD9BFA65.taxon	distribution	Distribution. Botswana, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284DFF3ED07AFE14FD00.taxon	description	(Figs 226 E, 229 G, 233 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284DFF3ED07AFE14FD00.taxon	description	Description. ♀ (Fig. 226 E). Apterous; body length 3.0 mm. Head brown, except mandible, malar space and anterior fourth of frons testaceous; clypeus testaceous, except central brown spot; antenna testaceous, except antennomere 8 darkened and antennomeres 9 – 10 brown; mesosoma brown-testaceous, except posterior extremity of propodeal declivity testaceous; mesoscutum slightly testaceous; petiole black; metasoma brown; legs brown, except tarsi and trochanters testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 14: 8: 7: 6: 6: 5: 4: 7. Head weakly excavated, shiny, unsculptured; frontal line incomplete (shortly present in front of anterior ocellus); occipital carina absent; POL = 2; OL = 3; OOL = 7. Palpal formula 6 / 3. Pronotum, crossed by strong transverse impression, shiny, unsculptured. Mesoscutum slightly longer than broad, shiny, sculptured by few longitudinal striae, laterally with two pointed apophyses (Fig. 226 E). Mesoscutellum shiny, smooth, flat. Length of mesoscutum + mesoscutellum less than twice as long as breadth of mesoscutum (12: 7). Metanotum short, unsculptured, with sides protruding (protrusions rounded) (Fig. 226 E), about coplanar to mesoscutellum (Fig. 229 G). Metapectal-propodeal disc shiny, unsculptured, with slight median furrow; propodeal declivity with few transverse striae near posterior extremity. Mesopleuron and metapleuron shiny, smooth, not transversely striate. Meso-metapleural obsolete (short track present near metanotum). Protarsomeres in following proportions: 12: 2.5: 4: 16: 24.5. Enlarged claw (Fig. 233 A) with one small subapical tooth and one row of seven peg-like setae, in addition to one bristle. Protarsomere 5 (Fig. 233 A) with two rows of 4 (proximal) + 23 lamellae; apex with approximately 12 lamellae. tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284DFF3ED07AFE14FD00.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Masoala National Park, 39.4 km 150 ° SSE Maroantsetra, 15 ° 42.6 ’ S 49 ° 58.2 ’ E, 200 m, canopy moss and leaf litter, rainforest, 28. XI – 3. XII. 2001, Fisher et al. leg., BLF (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284DFF3ED07AFE14FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAB284DFF3ED07AFE14FD00.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAA284EFF3ED497FAB4FEE0.taxon	description	(Fig. 233 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAA284EFF3ED497FAB4FEE0.taxon	description	Description. ♀. Apterous; body length 2.2 – 2.7 mm. Head black, except mandible and clypeus yellow; occasion- ally anterior region of frons testaceous; antenna brown; mesosoma black; metasoma black; legs black, except tarsi brown. Head flat, shiny, bare, granulate; frontal line complete; POL = 2.5; OL = 1.5; OOL = 7. Palpal formula 3 / 2 or 4 / 2. Pronotum shiny, slightly granulate, crossed by strong transverse impression. Mesoscutum dull, granulate, with some longitudinal keels, without lateral pointed apophyses. Metanotum dull, slightly transversely striate, with sides protruding; lateral protrusions of metanotum rounded. Metapectal-propodeal disc shiny, granulate; propodeal declivity, mesopleuron and metapleuron not or slightly transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 approximately twice as long as 4 (13: 6). Enlarged claw (Fig. 233 B) with one small subapical tooth and one row of 3 – 5 peg-like setae. Protarsomere 5 (Fig. 233 B) with inner side proximally not serrate, with two rows of 13 – 15 usually small lamellae; apex with approximately 4 – 9 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAA284EFF3ED497FAB4FEE0.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Esira, IX. 1940, A. Seyrig leg. (MNHN). Paratypes: MADAGASCAR: Toliara, Bekily, V. 1939, A. Seyrig leg., 1 ♀ (MNHN), Toliara, Behara, IX. 1940, A. Seyrig leg., 1 ♀ (MRAC). Other material: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Rutshuru, 1 ♀ (MRAC). ETHIOPIA: Shoa Province, 95 km W Addis Ababa, Ambo, near Plant Protection Research Institute, 2400 – 2500 m, 27. V. 1990, 22. VII. 1990, A. Emel’yanov leg., 2 ♀♀ (ZISP). KENYA: Central Prov., Gatamayu For- est, 00 ° 58.68 ’ S 36 ° 41.62 ’ E, near fishing camp, 20 – 30. I. 1999, MT, R. Copeland leg., 1 ♀ (NMK); Central Prov., Nairobi, ICIPE Research Station, 01 ° 17 ’ S 36 ° 49 ’ E, 24. V. 2001, sweeping tall grass, M. Stiller leg., 2 ♀♀ (SANC). MADAGASCAR: Toamasina, Rogez, 1 ♀ (MRAC); Toliara, Bekily, V. 1939, A. Seyrig leg., 1 ♀ (MRAC); Toliara, 12 km NW Amboasary, Berenty, 1 ♀ (AMNH); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 01.26 ’ S 46 ° 18.33 ’ E, 35 m, 7 – 17. X. 2004, MT, spiny forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02 - 22 A- 28, 1 ♀ (CAS). RWANDA: Kisenyi, 2 ♀♀ (AMNH, MRAC). SOUTH AFRICA: Eastern Cape, Somerset East, X. 1930, R. E. Turner leg., 16 ♀♀ (NHMUK); Eastern Cape, Katberg, 15 – 30. I. 1933, R. E. Turner leg., 1 ♀ (NHMUK); Gauteng, Pretoria, National Botanical Gardens, 1 ♀ (AMNH); KwaZulu-Natal, Eshowe, 1 – 22. IV. 1926, VI. 1926, R. E. Turner leg., 2 ♀♀ (NHMUK); KwaZulu-Natal, Eshowe, near Ntumeni N. Res., 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 26 – 28. X. 2007, YPT, Kolyada & Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Louwsberg, iGwala Gwala, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 2 – 3. VI. 2005, YPT, Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Karkloof, 29 ° 19.1 ’ S 30 ° 15.5 ’ E, 1325 m, 22. I – 6. II. 2007, YPT, Kolyada & Mostovski leg., 3 ♀♀ (NMSA); KwaZulu-Natal, Cathedral Peak Nature Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 3 – 15. XII. 2005, MT, M. Mostovski leg., 1 ♀ (NMSA); Limpopo, Klaserie Nature Reserve, 1 ♀ (PMA); Limpopo, Thabazimbi and vicinity, 24 ° 35 ’ S 27 ° 22 ’ E, 2. IV. 2000, swept off Hyparrhenia sp. (Poaceae), M. Stiller leg., 1 ♀ (SANC); same locality label, 2. IV. 2000, swept off grass, open and under tree canopy, ungrazed / unburnt, 1 ♀ (SANC); Western Cape, 25 km N Prince Albert, Kat River Crossing, 33 ° 05 ’ 39 ’’ S 21 ° 49 ’ 59 ’’ E, 500 m, MT, damp wsh, 3 – 24. X. 2004, M. E. Irwin, F. D. Parker & M. Hauser leg., 1 ♀ (UCFC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAA284EFF3ED497FAB4FEE0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDAA284EFF3ED497FAB4FEE0.taxon	distribution	Distribution. Democratic Republic of the Congo, Ethiopia, Kenya, Madagascar, Rwanda, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	description	(Figs 221, 228 B, 233 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with body small (1.9 mm), totally yellow-testaceous, except petiole black (Fig. 228 B); palpal formula 5 / 3; pronotum crossed by strong transverse furrow; mesoscutum less than 3 × as long as broad, with two lateral pointed prominences; meso-metapleural suture obsolete; protarsomere 1 as long as 4; enlarged claw (Fig. 233 C) with one small subapical tooth; protarsomere 5 (Fig. 233 C) with two rows of lamellae situated in distal half.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	description	Description. ♀ (Figs 221, 228 B). Apterous; body length 1.9 mm. Totally yellow-testaceous, except petiole black and antenna brown, except scape and pedicel testaceous and flagellomeres 7 – 8 whitish. Antenna clavate; antennomeres in following proportions: 8: 3: 7: 4: 3: 3: 4: 3: 2: 4. Head (Fig. 221 A, D) slightly excavated, shiny, unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 4.5. Palpal formula 5 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, unsculptured, laterally with two small pointed apophyses (Fig. 228 B). Mesoscutellum shiny, unsculptured, almost flat. Metanotum short, unsculp- tured, about coplanar to mesoscutellum (Figs 221 B). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron shiny, unsculptured. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 8: 2: 3: 8: 13. Enlarged claw (Fig. 233 C) with one small subapical tooth and two peg-like setae. Protarsomere 5 (Fig. 233 C) with two rows of 3 + 11 lamellae situated in distal half; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	materials_examined	Material examined. Type: ♀ holotype (CASTYPE 19466): MADAGASCAR: Toliara, Lake Ranobe, 23 ° 02.21 ’ S 43 ° 36.42 ’ E, 30 m, 25 – 28. IV. 2003, sifted litter (leaf mold, rotten wood), spiny forest thicket, MGF 064, Frontier Wilderness Project (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA9284FFF3ED7F7FAC2FDB8.taxon	etymology	Etymology. The name of the species derives from the Latin adjective minutus, meaning very small.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA72841FF3ED3E1FE5DFE98.taxon	description	(Figs 230 A, 233 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA72841FF3ED3E1FE5DFE98.taxon	description	Description. ♀. Apterous; body length 3.6 mm. Completely yellow-testaceous, except petiole black. Antenna distally thickened; antennomeres in following proportions: 7: 5: 15: 10: 7: 7: 6: 6: 5: 8. Head excavated, shiny, slightly granulate, except large frontal unsculptured area in front of anterior ocellus; frontal line complete; occipital carina almost completely absent (short track present behind ocellar triangle); occiput setose; POL = 1.5; OL = 1.5; OOL = 7. Palpal formula 5 / 2. Pronotum setose, crossed by strong transverse impression, with anterior collar and disc shiny, unsculptured. Mesoscutum shiny, unsculptured, setose, laterally without pointed apophyses. Mesoscutellum shiny, smooth, flat. Metanotum short, transversely striate and about coplanar to mesoscutellum (Fig. 230 A). Metapectalpropodeal disc shiny, setose, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 15: 2.5: 5: 14: 23. Enlarged claw (Fig. 233 D) with one small subapical tooth and one row of three peg-like setae, in addition to three bristles. Protarsomere 5 (Fig. 233 D) with two rows of 4 (medial) + 16 lamellae; apex with approximately nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA72841FF3ED3E1FE5DFE98.taxon	materials_examined	Material examined. Type: ♀ holotype: YEMEN: Ja’ar, 15. I. 2001, N. 5152, A. van Harten leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA72841FF3ED3E1FE5DFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA72841FF3ED3E1FE5DFE98.taxon	distribution	Distribution. Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	description	(Figs 230 B, 233 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	description	Description. ♀. Apterous; body length 3.1 mm. Completely ferruginous, except petiole black. Antenna clavate. Head (Fig. 230 B) almost flat, shiny, unsculptured or slightly granulate; temple very prominent; POL = 4; OL = 6; OOL = 19. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, with some longitudinal keels, without lateral pointed apophyses. Metanotum shiny, transversely striate, about coplanar to mesoscutellum. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron not transversely striate. Metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 slightly shorter than 4 (10: 11). Enlarged claw (Fig. 233 E) with one small subapical tooth and six lamellae, in addition to one bristle. Protarsomere 5 (Fig. 233 E) with inner side proximally serrate and one row of approximately 16 lamellae; apex with approximately 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, V. 1928, R. E. Turner leg. (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA62842FF3ED041FAA8FE28.taxon	discussion	Remarks. Based on the presence of the track of the transverse furrow on the pronotum, G. yemenensis (group 6) can be included in group 7, where it is similar to Gonatopus mosseli Olmi 1984. However, the subapical tooth of the enlarged claw of G. yemenensis is farther from the apex (Fig. 203 F) than it is in G. mosseli (Fig. 233 E).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA52844FF3ED0ACFE1FFE28.taxon	description	(Fig. 234 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA52844FF3ED0ACFE1FFE28.taxon	description	Description. ♀. Apterous; body length 2.5 mm. Head black, except clypeus, mandible and anterior third of frons testaceous; antenna brown, except antennomere 10 whitish and scape, pedicel and proximal third of flagellomere 1 testaceous; mesosoma black, except part of mesoscutum testaceous. metasoma black; legs testaceous, except coxae and clubs of femora partly brown-black (in paratype, metalegs completely testaceous, except stalks of femora brown-black). Head excavated, shiny, bare, unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1.5; OOL = 7. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Me- soscutum shiny, with some longitudinal keels, without lateral pointed apophyses. Metanotum transversely striate. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleu- ron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 11: 2: 3: 12: 19. Enlarged claw (Fig. 234 A) with one small subapical tooth and 7 – 8 peg-like setae. Protarsomere 5 (Fig. 234 A) with inner side proximally not serrate and one row of 11 – 13 lamellae situated in distal half; apex with 13 – 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA52844FF3ED0ACFE1FFE28.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: KwaZulu-Natal, Port Edward, along road from R 61 to Umtamvuna Nature Reserve, C. 13. IV. 2007, B. 17. IV. 2007, M. Olmi reared from an adult of Maiestas angustisecta (Linnavuori), SAM-HYM-P 025112 (SAMC). Paratype: SOUTH AFRICA: Free State, Royal Natal National Park, 11 km E of Park Gate along Road from R 74 to Park, grass along the road, C. 9 April 2007, B. 12. IV. 2007, M. Olmi reared from an adult of Maiestas angustisecta (Linnavuori), 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA52844FF3ED0ACFE1FFE28.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in South Africa: Maiestas angustisecta (Linnavuori).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA52844FF3ED0ACFE1FFE28.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	description	(Figs 227, 230 C, D, 234 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; temple without sharp carina; mesosoma black; pro- notum crossed by strong transverse furrow; mesoscutum less than twice as long as broad, laterally with two strong pointed apophyses; mesopleuron without lateral pointed prominence; metanotum deeply excavated behind mesos- cutellum; meso-metapleural suture distinct and complete; metapectal-propodeal disc without strong median furrow; protarsomere 1 as long as 4; enlarged claw (Fig. 234 B) long, straight and slender, with one small subapical tooth and one row of peg-like setae; subapical tooth of enlarged claw situated very near apex (Fig. 234 B).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	description	Description. ♀ (Fig. 227). Apterous; body length 2.9 – 4.4 mm. Head black, except mandible, clypeus, malar space and two short stripes along orbits testaceous; antenna brown-testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae partly brown and clubs of femora brown. In specimens from Democratic Republic of the Congo, 04 ° 15 ’ S 17 ° 10 ’ E, head brown, except mandible, clypeus, anterior half of frons testaceous; antenna brown-testaceous, except scape, pedicel and part of flagellomere 1 whitish; mesosoma black, except mesoscutum mostly testaceous; metasoma brown; legs brown, except coxae, trochanters and femora partly whitish; tarsi testaceous. Antenna clavate; antennomeres in following proportions: 7: 6: 12: 8: 5: 5: 4: 4: 4: 6. Head excavated, shiny, with frons unsculptured; frontal line complete; occipital carina absent; POL = 1; OL = 1; OOL = 8; greatest breadth of lateral ocelli as long as POL. Palpal formula 6 / 3. Pronotum crossed by strong transverse furrow, shiny, unsculptured. Mesoscutum shiny, unsculptured, laterally with two strong pointed apophyses situated at border of mesoscutellum (Fig. 230 C). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum rugose, deeply excavated behind mesoscutellum (Fig. 230 D). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity sculptured by numerous transverse striae. Mesopleuron partly unsculptured and partly slightly transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 14: 2: 3: 14: 18. Enlarged claw (Fig. 234 B) with one small subapical tooth and one row of 6 – 9 peg-like setae + one bristle. Protarsomere 5 (Fig. 234 B) with two rows of 1 – 2 (long and proximal) + 15 – 18 lamellae; apex with 6 – 8 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Northern Cape, West Driefontein farm, Succulent Karoo 1 Site, 13.1 km 106 ° ESE Nieuwoudtville, 31 ° 22.178 ’ S 19 ° 15.064 ’ E, 16. X. 2000, sweep, Succulent Karoo on shales, S. van Noort leg., SAM-HYM-A 022692 (SAMC). Paratypes: DEMOCRATIC REPUBLIC OF THE CON- GO: Haut-Uele, Wamba Territory, Kikongo Mission, 04 ° 15 ’ S 17 ° 10 ’ E, 6. IV. 2006, forest pantrap, S. L. Heydon & S. E. Stevenson leg., 1 ♀ (MOLC); same locality label, 30. III. 2006, forest pantrap, 1 ♀ (DEUCD). SOUTH AFRICA: Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 29. IX – 30. XII. 2010, YPT, Hantam Succulent Karoo, KEL 09 - SUC 1 - Y 07, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	distribution	Distribution. Democratic Republic of the Congo, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA32845FF3ED44FFBBDFC68.taxon	etymology	Etymology. The species is named nigropictus, because of its mostly black colour.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA22847FF3ED20FFA95F8CA.taxon	description	(Figs 220 E, 230 E – G, 232 A, B, 234 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA22847FF3ED20FFA95F8CA.taxon	description	Description. ♀ (Figs 232 A, B). Apterous; body length 3.1 – 3.6 mm. Head totally reddish, or brown-black, except mandible, clypeus and anterior region of frons (more along orbits) yellow; antenna brown, except scape, pedicel and part of flagellomere 1 yellow; mesosoma brown-black or reddish; metasoma brown-black; legs testaceous, except coxae and club of profemur brown. Head almost flat (Fig. 230 E), dull, granulate; occipital carina incomplete, only present behind and on sides of lateral ocelli, laterally not reaching half-way of OOL; POL = 7; OL = 7; OOL = 21. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum with (Fig. 232 A) or without (Fig. 230 F) two pointed apophyses laterally. Metanotum short, transversely striate, excavated behind mesoscutellum (Fig. 230 G). Metapectal-propodeal disc shiny, unsculptured, with deep median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 as long as 4. Enlarged claw (Fig. 234 C) with one small subapical tooth and 6 – 7 peg-like setae. Protarsomere 5 (Fig. 234 C) with inner side proximally not serrate and 1 – 2 rows of 1 – 4 (proximal) + 11 – 19 lamellae; apex with at least seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.6 – 2.1 mm. Head black; antenna brown; mesosoma and metasoma black; legs brown, except stalks of femora, protibia and tarsi whitish (except brown last tarsomere). Antenna filiform; antennomeres in following proportions: 4: 3.5: 6: 6: 5: 5.5: 5: 5: 5: 7.5; antennomere 3 less than 3 × as long as broad (6: 2.5). Head dull, granulate and slightly rugose; frontal line absent; occipital carina absent; POL = 5.5; OL = 2; OOL = 4; greatest breadth of lateral ocelli slightly longer than OL (2.5: 2); area between lateral ocelli and eyes flat and rugose. Palpal formula 5 / 2. Mesoscutum dull, granulate. Notauli complete, posteriorly separated; minimum distance between notauli as long as greatest breadth of lateral ocelli. Mesoscutellum slightly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc shiny, unsculptured, without furrows or keels. Forewing hyaline, without dark transverse bands. Dorsal process of paramere (Fig. 220 E) long and slender. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA22847FF3ED20FFA95F8CA.taxon	materials_examined	Material examined. Type: ♀ holotype: NAMIBIA: Okahandja, 19 – 29. XII. 1927, R. E. Turner leg. (NHMUK). Other material: CAPE VERDE ISLANDS: Santiago Island, São Jorge dos Orgãos, 1 ♀, 1 ♂ (AMNH). MAURI- TANIA: Nouakchott, 1 ♀ (AMCR). MOZAMBIQUE: Maputo Prov., Maputo, E. Mondlane University, Faculty of Agriculture campus, 1985, on lettuce, reared from Agalliopsis sp., 1 ♀ (AMNH); Maputo Prov., 26 km S of Ma- puto, Umbelúzi, 1 ♀ (AMNH). NAMIBIA: 27.6 km S Omaruru, along Rd C 33, C. 8. III. 2006, B. 12. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 A, designated by mistake as ♀ paratype of G. maritimus Olmi, 1 ♀ (MOLC); same locality label, C. 8. III. 2006, B. 13. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 B, designated by mistake as ♀ paratype of G. maritimus Olmi, 1 ♀ (MOLC); same locality label, C. 8. III. 2006, B. 11. III. 2006, Sf. 12. IV. 2006, reared from a nymph of Austroagallia cuneata (Cogan), M. Olmi leg., 23 C, designated by mistake as ♀ paratype of G. maritimus Olmi, 1 ♀ (MOLC). NORTH SUDAN (= REPUBLIC OF THE SUDAN): 39 km W Al Qadarif (= Gedaref), 19 – 29. IX. 1987, L. J. Pinto leg., 1 ♀ (USNM). SOUTH AFRICA: Northern Cape, Kimberley, 1 ♀ (MCZ); Northern Cape, Hopetown, 29 ° 37 ’ S 24 ° 05 ’ E, X. 1887, SAM-HYM-PO 05918, 2 ♀♀ (SAMC). YEMEN: Wadi Bana, 20. III. 1993, A. van Harten leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA22847FF3ED20FFA95F8CA.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Agalliopsis sp.; in Namibia: Agallia cuneata Cogan (= Austroagallia cuneata (Cogan )) (Olmi 2007 b; Guglielmino & Olmi 2007), attributed by mistake to G. maritimus (Olmi 2007 b; see remarks for G. maritimus).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDA22847FF3ED20FFA95F8CA.taxon	distribution	Distribution. Cape Verde Islands, Mauritania, Mozambique, Namibia, North Sudan, South Africa, Yemen.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED6D7FE55FC25.taxon	description	(Fig. 234 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED6D7FE55FC25.taxon	description	Description. ♀. Apterous; body length 3.1 mm. Head black, except mandible, clypeus and malar space testaceous; antenna black, except scape and pedicel testaceous; mesosoma and metasoma black; legs black, except trochanters, tarsi, stalk of profemur, protibia and proximal half of meso- and metatibia testaceous. Antenna clavate; antennomeres in following proportions: 8: 7: 14: 11: 11: 10: 9: 8: 7: 8; antennomere 3 about twice as broad as 2. Head almost flat, with vertex and frons shiny, unsculptured or slightly alutaceous; temple and occiput slightly granulate; temple very prominent, without sharp carina; frontal line complete; occipital carina absent; POL = 1; OL = 1.5; OOL = 6. Palpal formula 5 / 2. Pronotum shiny, slightly granulate, crossed by strong transverse impression. Mesoscutum narrow, dull, granulate, laterally without pointed apophyses. Metanotum transversely striate, about coplanar to mesoscutellum, without lateral pointed protrusions. Metapectal-propodeal disc shiny, slightly granulate; propodeal declivity transversely striate. Mesopleuron and metapleuron unsculptured or slightly granulate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 13: 2: 3.5: 11: 18. Enlarged claw (Fig. 234 D) with one small subapical tooth and one row of six peg-like setae. Protarsomere 5 (Fig. 234 D) with inner side proximally serrate and one row of approximately 16 lamellae; apex with approximately seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED6D7FE55FC25.taxon	materials_examined	Material examined. Type: ♀ holotype: OMAN: Salalah, 12. X. 1985, J. Huber leg. (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED6D7FE55FC25.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED6D7FE55FC25.taxon	distribution	Distribution. Oman.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED5BAFD61F874.taxon	description	(Fig. 234 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED5BAFD61F874.taxon	description	Description. ♀. Apterous; body length 3.0 – 6.3 mm. Head black, except mandible yellow; antenna black, except ventral side of scape yellow; mesosoma and metasoma black; legs brown-black, except tarsi testaceous. In small specimen (3.0 mm) from South Africa, Die Dam, head black, except mandible, clypeus, malar space and anterior part of frons testaceous; mesosoma black, except mesoscutum and metanotum testaceous. Antenna clavate; antennomeres in following proportions: 15: 8: 27: 15: 12: 10: 9: 6.5: 7: 11. Head excavated, dull, granulate; frontal line complete; occipital carina absent; temple distinct; POL = 3; OL = 1.5; OOL = 10.5. Palpal formula 6 / 3. Pronotum dull, granulate, crossed by strong transverse impression. Mesoscutum dull, granulate, with some longitudinal keels or completely granulate, with two lateral pointed apophyses. Mesoscutellum sloping anteriorly, dull, granulate. Metanotum not transversely striate, with sides rounded, excavated behind mesoscutellum. Metapectal-propodeal disc dull, granulate; propodeal declivity with lateral regions transversely striate and median region granulate and not striate. In specimen from South Africa, Die Dam, propodeal declivity completely transversely striate. Mesopleuron and metapleuron transversely striate. In specimen from Uganda, 01 ° 45 ’ N 31 ° 35 ’ E, mesopleuron and metapleuron granulate and not transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 of holotype slightly longer than 4 (20: 19). Enlarged claw (Fig. 234 F) with one small subapical tooth and two bristles (or two peg-like setae and one bristle). Protarsomere 5 (Fig. 234 F) with two rows of 1 – 3 (proximal) + 20 – 23 lamellae; apex with about 12 – 16 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED5BAFD61F874.taxon	materials_examined	Material examined. Type: ♀ holotype: UGANDA: Kyagur, XI. 1938, T. H. C. Taylor leg. (NHMUK). Oth- er material: CAMEROON: 6 mi. S Kumba, 180 m, 1 ♀ (AMNH). SOUTH AFRICA: Western Cape, Die Dam, 24. II. 2006, M. Olmi reared ex Caffrolix cyclopia (Cogan), 1 ♀ (MOLC). UGANDA: Masindi District, Budongo Forest near Sonso, 01 ° 45 ’ N 31 ° 35 ’ E, VI. 1995, Th. Wagner leg., 1 ♀ (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED5BAFD61F874.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in South Africa: Caffrolix cyclopia (Cogan).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBF2858FF3ED5BAFD61F874.taxon	distribution	Distribution. Cameroon, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE2859FF3ED6D7FDB4FA8D.taxon	description	(Fig. 234 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE2859FF3ED6D7FDB4FA8D.taxon	description	Description. ♀. Apterous; body length 3.0 – 3.4 mm. Head testaceous-reddish; antenna testaceous-reddish, except antennomere 10 brown (in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, antenna testaceous, except antennomeres 9 – 10 brown); mesosoma testaceous reddish; petiole black; metasoma testaceous-reddish, partly brownish; legs testaceous-reddish (in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, legs testaceous, except stalk of metafemur and both extremities of metatibia brown). Specimen from Kenya, 00 ° 25.45 ’ S 37 ° 32.71 ’ E, brownish, except anterior surface of frons, clypeus, mandible, mesoscutum and metanotum testaceous-reddish. Head excavated, shiny. In specimen from Kenya, 00 ° 25.45 ’ S 37 ° 32.71 ’ E, head excavated, dull, granulate, with frontal line complete, POL = 2; OL = 2; OOL = 7. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured, except lateral regions granulate. Mesoscutum shiny, unsculptured, with some longitudinal keels and two lateral pointed apophyses. Metanotum shiny, excavated behind mesoscutellum, rugose. Metapectal-propodeal disc shiny, granulate; propodeal declivity not transversely striate or slightly striate; in specimen from South Africa, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, propodeal declivity granulate, transversely striate near posterior extremity. Mesopleuron and metapleuron not transversely striate or slightly striate. Meso-metapleural suture distinct and complete, broad and step-shaped (mesopleuron and metapleuron situated on different planes). Protarsomeres in following proportions: 13: 2: 4: 16: 22. Enlarged claw (Fig. 234 E) with one small subapical tooth and 4 – 5 peg-like setae. Protarsomere 5 (Fig. 234 E) with inner side proximally not serrate, with two rows of 2 – 4 + 21 – 28 lamellae; apex with 9 – 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE2859FF3ED6D7FDB4FA8D.taxon	materials_examined	Material examined. Type: ♀ holotype: KENYA: Central Prov., Nairobi, IV. 1945, S. Patrizi leg. (MRAC). Other material: KENYA: Eastern Prov., Kirimiri Hill, 00 ° 25.45 ’ S 37 ° 32.71 ’ E, 1745 m, 16 – 29. III. 2005, MT, R. Copeland leg., 1 ♀ (RSC). SOUTH AFRICA: Eastern Cape, Port St. Johns, 1 – 11. VI. 1923, R. E. Turner leg., 1 ♀ (NHMUK); same locality label, 1 – 9. VII. 1923, 1 ♀ (NHMUK); same locality label, 6 – 25. II. 1924, 1 ♀ (AMNH); KwaZulu-Natal, Eshowe, near Ntumeni Nature Reserve, 28 ° 52.08 ’ S 31 ° 22.41 ’ E, 680 m, 23. II – 20. IV. 2007, MT, Kolyada & Mostovski leg., 1 ♀ (NMSA); KwaZulu-Natal, Cathedral Peak Natural Reserve, Rainbow Gorge, 28 ° 57.60 ’ S 29 ° 13.61 ’ E, 1480 m, 22. IX – 17. XI. 2006, MT, M. Mostovski leg., 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE2859FF3ED6D7FDB4FA8D.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE2859FF3ED6D7FDB4FA8D.taxon	distribution	Distribution. Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE285AFF3ED312FE5BFEC4.taxon	description	(Fig. 235 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE285AFF3ED312FE5BFEC4.taxon	description	Description. ♀. Apterous; body length 4.0 mm. Head black, except mandible and clypeus testaceous; antenna black, except scape and pedicel testaceous; mesosoma and metasoma black; legs black, except tarsi, part of procoxa, part of protrochanter and fore articulations testaceous. Antenna clavate; antennomeres in following proportions: 10: 7: 24: 17: 13: 10: 9.5: 9: 8: 11. Head excavated, shiny, strongly sculptured by many longitudinal keels on frons, vertex and part of occiput; frontal line complete; occipital carina absent; POL = 2; OL = 2; OOL = 9. Palpal formula 6 / 3. Pronotum shiny, partly granulate and partly unsculptured, with some transverse keels on lateral regions, crossed by strong transverse impression. Mesoscutum dull, completely reticulate rugose, with two prominent lateral point- ed apophyses. Mesoscutellum short, sloping anteriorly, rugose. Metanotum sloping anteriorly, reticulate rugose. Metapectal-propodeal disc dull, completely reticulate rugose; propodeal declivity with few transverse striae near posterior extremity. Mesopleuron and metapleuron reticulate rugose, not transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 16: 3: 6: 25: 37. Enlarged claw (Fig. 235 A) with one small subapical tooth and nine lamellae in addition to one bristle. Protarsomere 5 (Fig. 235 A) with inner side proximally serrate and two rows of 2 (proximal) + 20 lamellae; apex with approximately 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE285AFF3ED312FE5BFEC4.taxon	materials_examined	Material examined. Type: ♀ holotype: GABON: Estuaire Province, Donguila, 5. I. 1985, on Borreria verticillata, A. Pauly leg. (FSAE).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE285AFF3ED312FE5BFEC4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBE285AFF3ED312FE5BFEC4.taxon	distribution	Distribution. Gabon.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285AFF3ED7DBFD62FBFE.taxon	description	(Figs 230 H, 235 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285AFF3ED7DBFD62FBFE.taxon	description	Description. ♀. Apterous; body length 4.5 – 7.4 mm. Head brown; antenna with antennomeres 1 – 3 reddish; other an- tennomeres missing in only two examined specimens, including holotype; pronotum black, except posterior surface and lateral regions reddish; rest of mesosoma and metasoma black; legs black, except tarsi testaceous. Head (Fig. 230 H) slightly excavated, shiny, granulate, with some longitudinal keels on frons; occiput setose; POL = 5; OL = 7; OOL = 23. Palpal formula 6 / 3. Pronotum shiny, unsculptured, with long setae, with some keels on lateral regions, crossed by strong transverse impression. Mesoscutum shiny, with long setae, with two lateral pointed apophyses. Metapectal-propodeal disc with long setae, sculptured by numerous longitudinal keels; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 24: 4: 8: 23: 35. Enlarged claw (Fig. 235 B) with one small subapical tooth and one row of eight lamellae. Protarsomere 5 (Fig. 235 B) with inner side proximally serrate, with two rows of 17 lamellae (shortest row not extending beyond 0.5 length of protarsomere); apex with about 19 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285AFF3ED7DBFD62FBFE.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Free State, Bothaville, 12. X. 1898, Hans Brauns leg. (MCPM). Other material: ZIMBABWE: Bulawayo, 9. IX. 1923, Rhodesia Museum, 1 ♀ (NHMUK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285AFF3ED7DBFD62FBFE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285AFF3ED7DBFD62FBFE.taxon	distribution	Distribution. South Africa, Zimbabwe.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	description	(Figs 212 B, 217 A, B, 230 I, 231, 235 C, 239 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; head granulate, black, except mandible, clypeus, small area between antennal toruli, malar space and narrow stripe near malar space along orbits testaceous; temple without sharp carina; mesosoma brown-black; pronotum crossed by strong transverse impression; mesoscutum about twice as long as broad, with two lateral pointed apophyses (Fig. 217 A); metanotum about coplanar to mesoscutellum; mesopleuron without lateral pointed prominence; meso-metapleural suture distinct and complete; metapectal-propodeal disc without strong median furrow, unsculptured; protarsomere 1 slightly shorter than 4; enlarged claw (Figs 212 B, 235 C) with one small subapical tooth and one row of peg-like setae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	description	Description. ♀ (Figs 217 A, B, 231). Apterous; body length 3.5 – 3.7 mm. Head black, except mandible, clypeus, small area between antennal toruli, malar space and short narrow stripe along orbits near malar space testaceous; antenna brown, except scape and pedicel brown-testaceous and flagellomere 8 whitish (in paratype flagellomeres 7 and 8 whitish); mesosoma brown-black, except apex of propodeal declivity yellow; petiole black; metasoma brownblack; proleg brown, except part of coxa, trochanter and tarsus testaceous; mesoleg brown, except tarsus testaceous; metaleg brown, except tarsus and part of tibia testaceous (in paratype proleg brown, except part of coxa and trochanter whitish; meso- and metaleg brown, except part of tibiae and tarsi testaceous). Antenna clavate; antennomeres in following proportions: 8: 5: 19: 10: 8: 7: 6: 6: 4: 8. Head (Fig. 231 C) excavated, dull, granulate; frontal line complete; occipital carina shortly present behind and on sides of lateral ocelli; POL = 1; OL = 1.5; OOL = 8; greatest breadth of lateral ocellus about as long as OL. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum shiny, unsculptured, about twice as long as broad (11: 6) (shortest breadth), laterally with two pointed apophyses (Fig. 230 I). Mesoscutellum shiny, flat, unsculptured. Length of mesoscutum + mesoscutel-lum about 3 × as long as breadth of mesoscutum (17: 6). Metanotum shiny, unsculptured, sloping anteriorly, not or weakly excavated behind mesoscutellum (Fig. 239 A), with sides rounded, not protruding. Metapectal-propodeal disc (Fig. 231 A, B) fusiform, shiny, unsculptured; propodeal declivity with transverse striae on distal half. Meso- pleuron and metapleuron unsculptured. Meso-metapleural suture distinct and complete. Protarsomeres (Fig. 231 D) in following proportions: 16: 3: 5: 17: 26. Enlarged claw (Figs 212 B, 235 C) with one small subapical tooth and one row of five-six peg-like setae + one bristle. Protarsomere 5 (Figs 212 B, 235 C) without proximal serrate area, with two rows of 3 – 4 (proximal) + 18 – 19 lamellae; apex with five-seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	materials_examined	Material examined. Types: ♀ holotype (CASTYPE 19467): MADAGASCAR: Fianarantsoa, Ranomafana National Park, Vohiparara, 21 ° 13.67 ’ S 47 ° 22.19 ’ E, at broken bridge, 1110 m, 12 – 19. II. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 08 A- 16 (CAS). Paratype: MADAGASCAR: same locality label as holo- type, 26. II – 4. IV. 2002, MA- 02 - 09 A- 18 (MOLC). DEMOCRATIC REPUBLIC OF THE CONGO: Ituri Prov., Djugu Territory, confluence of Ituri and Kelebo rivers, Uphill “ Camp Kelebo ”, 30. III – 6. IV. 2015, primary / secondary forest, 75 YPTs, A. Gumovsky leg., 1 ♀ (SIZK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	distribution	Distribution. Democratic Republic of the Congo, Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBD285BFF3ED282FA68FD74.taxon	etymology	Etymology. The species is named ranomafanensis after the collection site in the Ranomafana National Park.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBC285DFF3ED120FC70FF7C.taxon	description	(Figs 220 F, 235 D, 239 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBC285DFF3ED120FC70FF7C.taxon	description	Description. ♀. Apterous; body length 2.5 – 3.0 mm. Head testaceous-reddish; antenna brown, except scape, pedicel and part of flagellomere 1 yellow, flagellomeres 7 – 8 and partly 6 whitish; pronotum brown-reddish; mesoscutum yellow, except anterior margin brown; rest of mesosoma black, except distal extremity of propodeal declivity reddish; petiole black; metasoma brown-reddish; legs testaceous. In very dark specimen from Mozambique, Cuamba, head with large black band on vertex; antenna black, except scape and pedicel testaceous and flagellomere 8 whitish; propleuron brown; pronotum black; legs testaceous, except femora darkened. In specimen from Madagascar, antenna brown, except scape and pedicel testaceous and flagellomere 8 whitish. Head (Fig. 239 B) excavated, shiny, unsculptured; temple prominent. POL = 4; OL = 4; OOL = 15. Palpal formula 5 / 2. Pronotum shiny, setose, unsculptured, crossed by strong transverse impression. Mesoscutum shiny, with some longitudinal keels. Metanotum, mesopleuron and metapleuron transversely striate. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 slightly shorter than 4 (10: 11). Enlarged claw (Fig. 235 D) with one small subapical tooth and one row of seven peg-like setae. Protarsomere 5 (Fig. 235 D) with inner side proximally not serrate, with one row of 15 – 19 lamellae; apex with at least 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 1.1 – 1.9 mm. Head black, except mandible testaceous; antenna brown; mesosoma and metasoma black; legs brown, except tarsi, protibia and stalk of profemur testaceous. Antenna setose, filiform; antennomeres of specimen from Mozambique, Cuamba, in following proportions: 4: 4: 5: 4: 4.5: 4.5: 4: 4.5: 4: 6.5; antennomeres of specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, in following proportions: 5: 4: 5: 5: 4.5: 4.5: 4.5: 4: 4: 6. Antennomere 3 less than 3 × as long as broad: 5: 1.7 in specimen from Mozambique, Cuamba; 5: 2.3 in specimen from South Africa, Blue Mountain Ranch Bed & Breakfast. Head dull, granulate; frontal line absent; occipital carina absent; in ♂ specimen from Mozambique, Cuamba, POL = 5; OL = 2; OOL = 4; in ♂ specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, POL = 5; OL = 3; OOL = 4. Vertex of head with two oval smooth and shiny areas situated between lateral ocelli and eyes (areas anteriorly and posteriorly surrounded by carina, anterior carina being much higher. Temple distinct, crossed by sharp carina parallel to posterior margin of eyes. Palpal formula 4 / 2. Mesoscutum dull, granulate; notauli complete, posteriorly separated. In specimen from Mozambique, Cuamba, minimum distance between notauli less than 0.5 × as long as greatest breadth of lateral ocelli (0.5: 1.5). In specimen from South Africa, Blue Mountain Ranch Bed & Breakfast, minimum distance between notauli shorter than greatest breadth of lateral ocelli (1.5: 2.0). Mesoscutellum shiny, weakly granulate. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, completely reticulate rugose; areolae of metapectal-propodeal disc small; propodeal declivity without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands, with only costal cell surrounded by well pigmented veins (other basal cells surrounded by obsolete veins); 2 R 1 cell open; 2 r-rs & Rs vein regularly curved, with distal part longer than proximal part (approximately 9: 7). Dorsal process of paramere (Fig. 220 F) short and slender, with apex pointed. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBC285DFF3ED120FC70FF7C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, Mossel Bay, I. 1922, R. E. Turner leg. (NHMUK). Other material: JUAN DE NOVA ISLAND (FRANCE; in Mozambique Channel): East Point, 17.06182 S, 42.72513 E, 10 m, 12. IV. 2011, ex used bird nest in littoral verge, B. L. Fisher leg., BLF 26478, 1 ♀ (CAS). MOZAMBIQUE: Maputo Prov., Maputo, 1 ♀ (AMNH); Maputo Prov., Maputo race-course, 1 ♀ (AMNH); Niassa, Cuamba, reared from Aconurella aethiopica (Cogan), C. 8. XI. 1999, B. 10. XI. 1999, Sf. 27. XI. 1999, 1 ♀ (MOLC); same locality label, C. 8. XI. 1999, B. 11. XI. 1999, Sf. 2. XII. 1999, 1 ♂ (MOLC); same locality label, C. 8. XI. 1999, B. 12. XI. 1999, Sf. 2. XII. 1999, 1 ♂ (MOLC); same locality label, C. 8. XI. 1999, B. 9. XI. 1999, Sf. 1. XII. 1999, 1 ♂ (MOLC); same locality label, C. 8. XI. 1999, B. 10. XI. 1999, Sf. 1. XII. 1999, 1 ♂ (MOLC); same locality label, reared from Aconurella shaba Ghauri, C. 22. III. 2003, B. 24. III. 2003, Sf. 14. IV. 2003, 3 ♂♂ (2 in MOLC, 1 in SAM); Niassa, Maúa, C. 10. XI. 1999, B. 14. XI. 1999, Sf. 10. XII. 1999, 1 ♂ (MOLC). SOUTH AFRICA: Eastern Cape, Februarie Farm, 40.2 km 267 ° W Kirkwood, 33 ° 33.124 ’ S 25 ° 03.043 ’ E, 10 – 17. II. 2001, Winkler, Valley Bushveld (non-trashed), H. G. Robertson & R. Tourle leg., 1 ♀ (SAMC); Western Cape, near George, N 12 - Road to Montagu Pass junction, reared from unidentified Cicadellid, C. 25. I. 2003, B. 29. I. 2003, Sf. 19. II. 2003, M. Olmi leg., 1 ♀ (SAMC); Western Cape, near Swellendam, Marloth Nature Reserve, C. 29. I. 2003, B. 7. II. 2003, Sf. 27. II. 2003, M. Olmi reared from Recilia lobata (Linnavuori), 1 ♀ (MOLC); Western Cape, Swellendam, Blue Mountain Ranch Bed & Breakfast, C. 29. I. 2003, B. 6. II. 2003, Sf. 25. II. 2003, M. Olmi reared from Aconurella aethiopica (Cogan), 1 ♂ (MOLC), same locality label, 16. II. 2006, M. Olmi reared from Aconurella aethiopica (Cogan), 1 ♂ (MOLC); Western Cape, near Citrusdal, The Baths, C. 22. I. 2003, B. 30. I. 2003, M. Olmi reared from unidentified Cicadellid, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBC285DFF3ED120FC70FF7C.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Aconurella aethiopica (Cogan) and Aconurella shaba Ghauri; in South Africa: Aconurella aethiopica (Cogan); Maiestas lobata (Linnavuori) (= Recilia lobata (Linnavuori )). Distribution. Juan de Nova Island, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBA285EFF3ED0E5FD76FCBC.taxon	description	(Figs 235 E, F, 239 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBA285EFF3ED0E5FD76FCBC.taxon	description	Description. ♀. Apterous; body length 2.2 – 3.0 mm. Yellow-reddish, except head slightly brownish, antennomeres 3 – 10 brownish, petiole black, metasoma black-brown and club of profemur brown. Specimen from Mozambique, Cuamba, completely yellow-testaceous, except petiole black. Antennomeres in following proportions: 4: 4: 11: 6: 4.5: 5: 4.4: 4: 5.5. Head (Fig. 239 C) excavated, shiny, bare, unsculptured. Palpal formula 4 / 2. Pronotum dull, crossed by strong transverse impression. Mesoscutum dull, with some longitudinal keels, without lateral pointed apophy- ses. Metanotum transversely striate. Metapectal-propodeal disc smooth; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Chelae missing in holotype. In specimen from Madagascar, Bereboka, enlarged claw (Fig. 235 E) with one small subapical tooth, two peg-like setae and four bristles; protarsomere 5 (Fig. 235 E) with two rows of 3 + 13 lamellae situated in distal half; apex of protarsomere 5 with at least 13 lamellae. In specimen from Madagascar, Berenty, protarsomere 5 (Fig. 235 F) with one row of 15 lamellae situated in proximal and distal half. In specimen from Mozambique, Cuamba, enlarged claw with five peg-like setae; protarsomere 5 with one row of 22 lamellae situated in proximal and distal half; apex of protarsomere 5 with nine lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBA285EFF3ED0E5FD76FCBC.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, Bekily, III. 1941, A. Seyrig leg. (MNHN). Paratypes: same locality label as holotype, VII. 1937, A. Seyrig leg., 11 ♀♀ (MNHN, MRAC). Other material: MADAGASCAR: Toliara, 60 km NE of Morondava, Bereboka, 4 ♀♀ (2 in NHMUK, 2 in AMNH); Toliara, 12 km NW of Amboasary, Berenty, 4 ♀♀ (AMNH). MOZAMBIQUE: Niassa, Cuamba, Farm of the Catholic University of Mozambique, C. 11. III. 2005, B. 13. III. 2005, Sf. 4. IV. 2005, M. Olmi reared from nymph of an unidentified species of Cicadellidae, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBA285EFF3ED0E5FD76FCBC.taxon	biology_ecology	Hosts. In Mozambique: unidentified species of Cicadellidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDBA285EFF3ED0E5FD76FCBC.taxon	distribution	Distribution. Madagascar, Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	description	(Figs 235 G, 236 A, 237, 239 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; temple without sharp carina; mesosoma black; pronotum crossed by strong transverse furrow; mesoscutum less than twice as long as broad, with two lateral pointed apophyses; mesopleuron without lateral pointed apophyses; metanotum deeply excavated behind mesoscutellum, with sides rounded (metapectal-propodeal disc fusiform); meso-metapleural suture distinct and complete; metapectal-propodeal disc without strong median furrow, only with slight track of median furrow, sculptured by some longitudinal keels; protarsomere 1 slightly shorter than 4; enlarged claw (Figs 235 G, 236 A) with one small subapical tooth, situated very far from apex, and proximal peg-like setae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	description	Description. ♀ (Fig. 237). Apterous; body length 2.5 – 3.0 mm. Head black, except mandible, clypeus and malar space (with stripe along orbits) testaceous; antenna brown, except ventral side of antennomere 1 whitish; mesosoma black; metasoma and legs brown. Paratypes with head black, except mandible, clypeus and anterior surface of frons testaceous; mesosoma black, except part of prothorax, part of mesoscutum and metanotum testaceous-brown. Head excavated, dull, granulate; frontal line complete; occipital carina absent; POL = 1.5; OL = 2; OOL = 8. Palpal formu- la 6 / 3. Pronotum crossed by strong transverse furrow, shiny, slightly granulate (unsculptured in paratypes). Mesos- cutum shiny, with some longitudinal keels, with two lateral pointed apophyses (Fig. 239 D). Mesoscutellum sloping anteriorly. Metanotum excavated behind mesoscutellum (Fig. 239 E), not protruding laterally. Metapectal-propodeal disc fusiform, dull, granulate, sculptured by some longitudinal irregular keels, with slight median longitudinal fur- row; propodeal declivity strongly transversely striate. Paratypes with metapectal-propodeal disc shiny, sculptured by longitudinal striae, unsculptured among striae. Mesopleuron slightly granulate, not transversely striate. Meta- pleuron strongly transversely striate. Meso-metapleural suture distinct and complete, step shaped. Protarsomeres in following proportions: 12: 2.5: 4: 5: 14: 22. Enlarged claw (Figs 235 G, 236 A) with one small subapical tooth and two proximal peg-like setae + 1 – 2 bristles. Protarsomere 5 (Figs 235 G, 236 A) with inner side proximally not serrate and two rows of 3 – 4 (proximal) + 17 – 18 lamellae; apex with about 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Northern Cape, Nieuwoudtville, Hantam National Botanical Garden, 31 ° 24.841 ’ S 19 ° 09.551 ’ E, 797 m, 27. X. 2009 – 9. II. 2010, MT, Nieuwoudtville Dolerite, Koppie Renosterveld, S. van Noort leg., SAM-HYM-A 022243 (SAMC). Paratypes: SOUTH AFRICA: Northern Cape, Swaarweerberg, Vredehoek Farm, 32 ° 26.387 ’ S 20 ° 34.501 ’ E, 1613 m, 28. X. 2009 – 9. II. 2010, MT, Succulent Karoo, S. van Noort leg., SAM-HYM-A 023397, 1 ♀ (SAMC); same locality label, 1 ♀ MOLC); same locality label, 9. II – 31. III. 2010, SAM-HYM-A 022710, 1 ♀ (SAMC); Northern Cape, Sterboom Farm, 32 ° 36.794 ’ S 20 ° 44.660 ’ E, 1599 m, 28. X. 2009 – 9. II. 2010, MT, Succulent Karoo, S. van Noort leg., SAM-HYM-A 022705, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB82851FF3ED002FB47F8C6.taxon	etymology	Etymology. The species is named rugithorax, because of the rugose aspect of its thorax.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB42853FF3ED203FDE4F845.taxon	description	(Figs 236 B, 239 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB42853FF3ED203FDE4F845.taxon	description	Description. ♀. Apterous; body length 5.9 mm. Head black, except mandible, clypeus, malar space and two short stripes along orbits testaceous; antenna testaceous, except antennomere 3, proximal half of 4, 8 – 10 darkened; mesosoma black, except lateral regions of pronotum partly testaceous; metasoma brown; legs testaceous, except clubs of femora slightly darkened. Antenna clavate; antennomeres in following proportions: 12: 6: 26: 17: 15: 11: 10: 9.5: 8.5: 10. Head excavated, shiny, with frons unsculptured; vertex, temple and occiput granulate; frontal line complete; occipital carina absent; POL = 2; OL = 2; OOL = 9; greatest breadth of lateral ocelli about as long as OL. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, with anterior collar and disc shiny, unsculptured; lateral regions of pronotum partly granulate and partly sculptured by longitudinal striae. Mesoscutum shiny, sculptured by few longitudinal striae, laterally with two strong pointed apophyses situated at border of mesoscutellum. Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum very short, transversely striate and deeply excavated behind mesoscutellum (Fig. 239 F). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity sculptured by many transverse striae; mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 20: 3.5: 7: 18: 27. Enlarged claw (Fig. 236 B) with one small subapical tooth and one row of six peg-like setae + one bristle. Protarsomere 5 (Fig. 236 B) with two rows of 7 + 13 lamellae; apex with approximately 21 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB42853FF3ED203FDE4F845.taxon	materials_examined	Material examined. Type: ♀ holotype: BURKINA FASO: Kompienga Prov., 20 km S Pama, 3 – 24. IX. 1988, savana, interception trap, F. Génier, M. Sanborne & F. M. Tou leg. (CNC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB42853FF3ED203FDE4F845.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB42853FF3ED203FDE4F845.taxon	distribution	Distribution. Burkina Faso.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32854FF3ED6D7FE13FC37.taxon	description	(Figs 236 C, 239 G-I)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32854FF3ED6D7FE13FC37.taxon	description	Description. ♀. Apterous; body length 3.1 mm. Completely testaceous-reddish, except petiole black. Antenna cla- vate; antennomeres in following proportions: 10: 4: 13: 7: 6: 6: 5: 5: 5: 7. Head excavated (Fig. 239 G), shiny, slightly granulate, with occiput more strongly granulate; frontal line complete; temple distinct; occipital carina incomplete, shortly present behind and on sides of lateral ocelli; POL = 2; OL = 2; OOL = 9; greatest breadth of lateral ocelli about as long as POL. Palpal formula 5 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured, except lateral regions granulate. Mesoscutum less than twice as long as broad (10: 6), with two pointed apophyses laterally (Fig. 239 H). Metanotum transversely striate, excavated behind mesoscutellum (Fig. 239 I). Sides of metanotum protruding; lateral protrusions of metanotum rounded (Fig. 239 H). Metapectal-propodeal disc shiny, unsculptured, with deep median longitudinal furrow; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete, step shaped. Protarsomeres in following proportions: 14: 3: 5: 17: 28. Enlarged claw (Fig. 236 C) with one small subapical tooth and six small lamellae + one bristle. Protarsomere 5 (Fig. 236 C) with two rows of 5 (proximal) + 26 lamellae; apex with at least 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32854FF3ED6D7FE13FC37.taxon	materials_examined	Material examined. Type: ♀ holotype: IVORY COAST: Bouaké Dept., Pli-Agakro, 10. IV. 1993, savannah, grass below trees, Frank Schierl leg. (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32854FF3ED6D7FE13FC37.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32854FF3ED6D7FE13FC37.taxon	distribution	Distribution. Ivory Coast.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	description	(Figs 236 D, 240 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	description	Description. ♀. Apterous; body length 3.2 – 4.4 mm. Head black, except mandible, clypeus and anterior region of frons testaceous; antenna brown, except antennomeres 5 – 6 testaceous and 9 – 10 testaceous-whitish (in specimen from Madagascar, 14 ° 26.36 ’ S 49 ° 44.36 ’’ E, ventral side of antennomere 1 whitish, rest of antenna testaceous-brown, except antennomeres 6 – 8 brown and 9 – 10 testaceous); mesosoma black; metasoma brown; legs brown. Antennal clavate; antennomeres in following proportions: 10: 6: 19: 9: 8: 8: 7: 7: 6: 10.5; antenna with ADOs on antennomeres 9 and 10. Head excavated, dull, granulate, with tracks of irregular striae; frontal line complete; occipital carina absent; occiput dull, sculptured by numerous transverse striae; POL = 2; OL = 2; OOL = 8. Palpal formula 6 / 3. Pronotum shiny, unsculptured, crossed by strong transverse impression. Mesoscutum about twice as long as broad, dull, with some longitudinal striae, with two pointed apophyses laterally (Fig. 240 A). Mesoscutellum shiny, unsculptured. Metapectal-propodeal disc dull, with track of median longitudinal furrow, sculptured by numerous longitudinal striae (in specimen from Madagascar, 14 ° 26.36 ’ S 49 ° 44.36 ’ E, metapectal-propodeal disc partly unsculptured and partly sculptured by slight striae); propodeal declivity with anterior half sculptured by many longitudinal striae and posterior half transversely striate (in specimen from Madagascar, 14 ° 26.36 ’ S 49 ° 44.36 ’ E, propodeal declivity unsculptured, except posterior half transversely striate and lateral regions of anterior half sculptured by slight transverse striae). Mesopleuron smooth, shiny, unsculptured. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 15: 3: 5: 17.5: 25. Enlarged claw (Fig. 236 D) with one small subapical tooth and one row of 5 – 7 lamellae in addition to 1 – 2 bristles. Protarsomere 5 (Fig. 236 D) with two rows of approximately 20 – 23 lamellae; apex with 10 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Fianarantsoa, 7 km SW of Ranomafana, 1200 m, 23. X. 1988, montane rainforest, Pyrethrin fogging of dead leaves and sticks in bamboo vine tangle, M. Steiner, C. Kremen & R. van Epps leg. (USNM). Other material: MADAGASCAR: Antsiranana, Marojejy National Park, Antranohofa, 26.6 km 31 ° NNE Andapa, 10.7 km 318 ° NW Manantenina, 14 ° 26.36 ’ S 49 ° 44.36 ’ E, 1325 m, 18. XI. 2003, on low vegetation, montane rainforest, B. L. Fisher leg., 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB32855FF3ED5B1FA6BFE98.taxon	discussion	Remarks. In this species the enlarged claw has a small subapical tooth. In spite of this character, the last two an- tennomeres show ADOs. This character combination is uncommon, though the enlarged claw has some lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB22856FF3ED0DFFCD2FCBC.taxon	description	(Fig. 236 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB22856FF3ED0DFFCD2FCBC.taxon	description	Description. ♀. Apterous; body length 4.5 – 6.0 mm. Head black, except mandible and clypeus yellow; occasionally malar space testaceous; antenna black; occasionally antennnomere 1 testaceous; mesosoma and metasoma black; legs black, except trochanters, meso- and metatibia and meso- and metatarsus brownish. Antennomeres in follow- ing proportions: 11: 9: 26: 23: 15: 15: 12: 12: 9: 14. Head dull, excavated; frons bare; occiput setose; frons and vertex with some longitudinal keels. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, smooth, with anterior surface of disc alutaceous and lateral regions and collar setose. Mesoscutum with some longitudinal keels. Metanotum sculptured by longitudinal keels. Metapectal-propodeal disc setose, sculptured by longitudinal keels; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Metasoma and legs very setose. Enlarged claw of holotype (Fig. 236 E) with one small subapical tooth and 10 small lamellae; enlarged claw of specimen from Mozambique, Cuamba, without subapical tooth and with one row of 12 lamellae. In holotype, protarsomere 5 (Fig. 236 E) with one row of 25 lamellae and apex provided with at least 16 lamellae. In specimen from South Africa, Irene, protarsomere 5 with two rows of 2 (proximal) + 18 lamellae. In specimen from Mozambique, Cuamba, protarsomere 5 with two rows of 1 (proximal) + 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB22856FF3ED0DFFCD2FCBC.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: [no locality], A. Seyrig leg. (MNHN). Other material: MOZAMBIQUE: Niassa, Cuamba, Campus of the Faculty of Agriculture of the Catholic University of Mo- zambique, sweeping, 7. VI. 2007, M. Olmi leg., 1 ♀ (MOLC); same locality label, C. 30. V. 2007, B. 10. VI. 2007, Sf. 15. VII. 2007, M. Olmi reared from an adult of Exitianus sp., 1 ♀ (MOLC). SOUTH AFRICA: Gauteng, Irene, 1 ♀ (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB22856FF3ED0DFFCD2FCBC.taxon	biology_ecology	Hosts. Cicadellidae (Guglielmino et al. 2013): in Mozambique: Exitianus sp.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB22856FF3ED0DFFCD2FCBC.taxon	distribution	Distribution. Madagascar, Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB12856FF3ED5C3FD2FF832.taxon	description	(Figs 236 F, 240 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB12856FF3ED5C3FD2FF832.taxon	description	Description. ♀. Apterous; body length 2.1 – 2.4 mm. Head black, except mandible, clypeus and anterior third of frons (including region between antennal toruli) testaceous; ventral side of head black, except anterior half testaceous; antenna completely testaceous (holotype with antennomeres 1 – 3 testaceous; remaining antennomeres missing; in specimen from South Africa, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, antenna brown, except antennomeres 1 – 3 testaceous); mesosoma black; metasoma brown; legs testaceous, except clubs of femora and meso- and metacoxa partly brown. Antenna clavate; antennomeres in following proportions: 7: 5: 5: 4.5: 5: 4: 3.5: 3.5: 3.5: 5. Head excavated, shiny, unsculptured, except temple and occiput granulate; frontal line complete; occipital carina shortly present behind and on sides of lateral ocelli; POL = 1; OL = 1; OOL = 7; greatest breadth of lateral ocelli longer than OL (1.5: 1). Palpal formula 4 / 2. Pronotum bare, crossed by strong transverse impression, shiny, unsculptured, except few longitudinal striae on laterial regions. Mesoscutum dull, sculptured by few longitudinal striae, without lateral pointed apophyses. Mesoscutellum shiny, unsculptured, flat. Metanotum short, rugose and transversely striate, about coplanar to mesoscutellum (Fig. 240 B), laterally with two rounded protrusions. Metapectal-propodeal disc shiny, with few irregular striae, unsculptured among striae, with track of median longitudinal furrow; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 10: 2.5: 3: 12: 19. Enlarged claw (Fig. 236 F) with one small subapical tooth and one row of 5 – 6 peg-like setae, in addition to one bristle. Protarsomere 5 (Fig. 236 F) with one row of 16 – 19 lamellae; apex with approximately 8 – 11 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB12856FF3ED5C3FD2FF832.taxon	materials_examined	Material examined. Type: ♀ holotype: ETHIOPIA: Shewa (= Shoa) Province, 95 km W Addis Abeba, Ambò (= Hagere Hiywot), near Plant Protection Research Institute, 2400 – 2500 m, 19. V. 1990, A. Emelyanov leg. (ZISP). Other material: SOUTH AFRICA: Eastern Cape, Asante Sana Game Reserve, 32 ° 14.990 ’ S 24 ° 55.962 ’ E, 2183 m, 29. X. 2009 – 23. II. 2010, MT, Karoo Escarpment Grassland, S. van Noort leg., 1 ♀ (SAMC). ZAMBIA: Copperbelt Province, 50 km W Chingola, 9 – 11. XII. 2002, J. Halada leg., 1 ♀ (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB12856FF3ED5C3FD2FF832.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB12856FF3ED5C3FD2FF832.taxon	distribution	Distribution. Ethiopia, South Africa, Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	description	(Figs 238 A, B, 240 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	description	Description. ♀. Apterous; body length 3.4 – 4.4 mm. Head brown, except anterior region of frons, clypeus and man- dible yellow; antenna testaceous (in specimen from South Africa, 34 ° 15 ’ S 18 ° 21.5 ’ E, antennomere 10 darkened); pronotum with collar black and disc brown-reddish or testaceous-reddish; mesoscutum yellow, except anterior margin black; rest of mesosoma black; metasoma brown-black; legs testaceous, except part of coxae and part of clubs of femora brown. Antenna clavate; antennomeres in following proportions: 9: 6.5: 19: 15: 12: 10: 7.5: 7: 6: 9. Head excavated, shiny, unsculptured, except occiput and anterior region of frons granulate; temple distinct; frontal line incomplete, not present in anterior third of frons; occipital carina absent; POL = 1; OL = 2; OOL = 8. Palpal formula 6 / 3. Pronotum dull, slightly granulate, crossed by strong transverse impression. Mesoscutum dull, granulate, with two pointed apophyses laterally. In specimen from Kenya, Kora, mesoscutum without lateral pointed apophyses. Metanotum transversely striate, about coplanar to mesoscutellum (Fig. 240 C). Metapectal-propodeal disc shiny, slightly alutaceous; propodeal declivity transversely striate. Mesopleuron and metapleuron granulate or transversely striate. Meso-metapleural suture obsolete. Protarsomere 1 approximately as long as 4. Enlarged claw (Figs 238 A, B) with one large subdistal tooth and one row of 8 – 10 lamellae + 1 seta. Protarsomere 5 (Fig. 238 A) with two rows of 3 – 4 (proximal) + 14 – 16 lamellae; apex with approximately 14 – 18 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Somerset East, 25 – 30. XI. 1930, R. E. Turner leg. (NHMUK). Other material: KENYA: North Eastern Prov., Kora, 1. I. 1984, C. West leg., 1 ♀ (AMNH). SOUTH AFRICA: Western Cape, Cape Peninsula National Park, 34 ° 15 ’ S 18 ° 21.5 ’ E, 100 m, 14 – 31. XII. 1999, fynbos, FIT, S. & J. Peck leg., 99 – 141, 1 ♀ (CNC); Western Cape, Gamkaberg Nature Reserve, 33 ° 44.090 ’ S 21 ° 55.654 ’ E, 997 m, 4. X. 2010 – 25. I. 2011, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., GB 09 - FYN 2 - M 53, 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	distribution	Distribution. Kenya, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED6D7FB8BFB7C.taxon	discussion	Remarks. Gonatopus somerseti was previously attributed to group 4 (former genus Apterodryinus) because the large subapical tooth and the lamellae of the enlarged claw were considered reliable characters for placing species in that group. However, the absence of antennal ADOs and the large distance between the subapical tooth and the apex of the enlarged claw, are convincing evidence that this species belongs in group 7.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED30FFE14F87A.taxon	description	(Figs 238 C, 240 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED30FFE14F87A.taxon	description	Description. ♀. Apterous; body length 3.4 mm. Head brown-black, except mandible, clypeus and anterior region of frons yellow; antenna yellow; mesosoma and metasoma black; legs yellow, except clubs of femora and meso- and metacoxa brown. Head excavated, shiny, unsculptured, except occiput and frons granulate. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured or slightly granulate. Mesoscutum shiny, with some longitudinal keels, without lateral pointed apophyses (Fig. 240 D). Metanotum shiny, transversely striate, sloping anteriorly, about coplanar to mesoscutellum (Fig. 240 E). Metapectal-propodeal disc shiny, unsculptured, with median longitudinal keel connecting metanotum to disc; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 as long as 4. Enlarged claw (Fig. 238 C) with one small subapical tooth and seven peg-like setae. Protarsomere 5 (Fig. 238 C) with inner side proximally serrate and one row of 16 lamellae; apex with eight lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED30FFE14F87A.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo Prov., Tananarive [= Antananarivo] (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED30FFE14F87A.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDB02857FF3ED30FFE14F87A.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2868FF3ED6D7FC42FAA9.taxon	description	(Figs 238 D, 240 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2868FF3ED6D7FC42FAA9.taxon	description	Description. ♀. Apterous; body length 4.1 – 4.4 mm. Head black, except mandible, clypeus and malar space yellow; antenna brown, except antennomeres 4 – 7 yellow; mesosoma black, with reddish nuances on disc and lateral regions of pronotum; metasoma black; legs brown-yellow. In specimen from Ethiopia, Ambo, head black, except mandible and lateral margins of clypeus testaceous; mesosoma totally black, except lateral regions of pronotum partly testaceous. Head excavated, shiny, unsculptured, except occiput and anterior region of frons slightly rugose and granulate; POL = 6; OL = 6; OOL = 25. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured, except lateral regions of collar and disc granulate and sculptured by longitudinal keels. Mesoscutum shiny, with many longitudinal keels, without lateral pointed apophyses. Mesoscutellum shiny, unsculptured. Metanotum shiny, not transversely striate, hollow behind mesoscutellum. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron transversely striate and with lateral pointed protrusion (Fig. 240 F). Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 as long as 4. Enlarged claw (Fig. 238 D) with one small subapical tooth and 2 – 6 peg-like setae or bristles. Protarsomere 5 (Fig. 238 D) with inner side proximally not serrate and one row of 17 lamellae, or two rows of 3 – 4 (proximal) + 18 – 30 lamellae; apex with at least 8 – 20 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2868FF3ED6D7FC42FAA9.taxon	materials_examined	Material examined. Types: ♀ holotype: UGANDA: Eastern Region, Sironko District, Bugusege (= Bugusaga, misspelt in Olmi 1984) [01 ° 07 ’ 20 ” N, 034 ° 15 ’ 55 ” E], XI. 1938, T. H. C. Taylor leg. (NHMUK). Paratypes: KENYA: Eastern Prov., Kibwezi, 30. XII. 1959, E. S. Ross leg., 1 ♀ (CAS). SOUTH AFRICA: Eastern Cape, Port St. Johns, XI. 1923, R. E. Turner leg., 1 ♀ (NHMUK). UGANDA: Central Region, Mubende District, Namutamba, 9. VII. 1940, T. H. C. Taylor leg., 1 ♀ (NHMUK); Eastern Region, Mayuge district, Bugota [00 ° 20 ’ 00 ” N, 033 ° 37 ’ 00 ” E], 1 ♀ (AMNH). Other material: BOTSWANA: Serowe, Farmer’s Brigade, 1 ♀ (USNM). ETHIOPIA: Shoa Province, 95 km W Addis Ababa, Ambo, near Plant Protection Research Institute, 2400 – 2500 m, 14. V. 1990, A. Emel’yanov leg., 1 ♀ (ZISP). SOUTH AFRICA: Limpopo, Naboomspruit, Nylsvley Nature Reserve, 1100 m, 9. IV. 1976, Sa- vanna Ecosystem Research Project, C. S. I. R., Habitat S. pungens, series No 2893, E. Holm, F. Kirsten & C. Scholtz leg., SAM-HYM-PO 21007, 1 ♀ (SAMC); Limpopo, near Waterpoort, Rochdale Farm, 22 ° 53 ’ S 29 ° 42 ’ E, 780 m, 28 – 31. XII. 1999, swept off herbaceous shrubs and grass under tree canopy, M. Stiller leg., 1 ♀ (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2868FF3ED6D7FC42FAA9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2868FF3ED6D7FC42FAA9.taxon	distribution	Distribution. Botswana, Ethiopia, Kenya, South Africa, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2869FF3ED3CEFE1FFD9C.taxon	description	(Fig. 238 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2869FF3ED3CEFE1FFD9C.taxon	description	Description. ♀. Apterous; body length 2.6 – 3.0 mm. Head black, except mandible and part of clypeus testaceous; antenna black, except scape and pedicel testaceous; mesosoma and metasoma black; legs testaceous, except coxae and clubs of femora darkened. Antenna clavate; antennomeres in following proportions: 6: 5: 13: 9: 7: 7: 6.5: 6: 5: 7. Head excavated, with frons unsculptured, and vertex and occiput dull and granulate; temple distinct; frontal line complete; occipital carina absent. Palpal formula 5 / 2. Pronotum slightly granulate, crossed by strong transverse impression. Mesoscutum dull, irregularly sculptured by keels, without lateral points. Metapectal-propodeal disc dull, granulate; propodeal declivity not transversely striate (or occasionally transversely striate). Mesopleuron transversely striate. Metapleuron not transversely striate. Meso-metapleural suture obsolete. Protarsomeres in following proportions: 14: 2: 4: 14: 20. Enlarged claw (Fig. 238 E) with one small subapical tooth and three peg-like setae or four lamellae + two bristles. Protarsomere 5 (Fig. 238 E) with two rows of approximately 15 – 17 lamellae situated in distal half of protarsomere; apex with approximately 9 – 14 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2869FF3ED3CEFE1FFD9C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Gauteng, Pretoria Distr., Soutpan, 25 ° 24 ’ S 28 ° 06 ’ E (SANC). Other material: SOUTH AFRICA: Free State, NE Kroonstad, Heuningspruit, 27 ° 26 ’ S 27 ° 22 ’ E, 1400 m, 4. III. 2002, DVac off grass, common: Eragrostis gummuflua (Poaceae), and forbs, M. Stiller & J. du Plessis leg., 1 ♀ (SANC); Free State, Sasolburg, Mullers Rust 352, 26 ° 24.53 ’ S 27 ° 46.42 ’ E, 1450 m, 23. XI. 1999, D-Vac, IS- 00, natural moist grassland (25 – 30 years since ploughing disturbance), slpe: none, M. Stiller leg., 1 ♀ (SANC); Free State, Sasolburg, Wonderwater Colliery, 26 ° 46.11 ’ S 27 ° 46.59 ’ E, 1450 m, 10. IV. 2000, D-Vac, IS- 08 b, west burm, seeded rehab., M. Stiller leg., 1 ♀ (SANC); Gauteng, Faerie Glen N / R Pretoria, 25 ° 46 ’ S 28 ° 17 ’ E, 1450 m, 18. III. 1998, D-Vac off shallow, rocky W-slope, M. Stiller leg., 1 ♀ (MOLC); KwaZulu-Natal, Vernon Crookes Nature Reserve, 23. XI. 2003, G. B. P. Davies leg., 1 ♀ (NMSA); Mpumalanga, E Greylingstad, near Val Station, Lawreston Farm, 26 ° 28 ’ S 28 ° 54 ’ E, 1600 m, 14. III. 2002, DVac short grass regrowth after mowing, M. Stiller leg., 1 ♀ (SANC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2869FF3ED3CEFE1FFD9C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8F2869FF3ED3CEFE1FFD9C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8E2869FF3ED423FD73F848.taxon	description	(Figs 240 G, H, K)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8E2869FF3ED423FD73F848.taxon	description	Description. ♀. Apterous; body length 2.5 – 4.5 mm. Head black, except mandible, clypeus and anterior region of frons (more along orbits) yellow (in very large specimen (4.5 mm) from Madagascar, 22 ° 18.48 ’ S 45 ° 17.30 ’ E, frons black, mandible and clypeus yellow); antenna yellow-reddish, except antennomeres 6 – 10, or 7 – 10, or 9 – 10 yellow; occasionally antenna totally brown, or testaceous, except antennomere 10 brown; mesosoma and metasoma black; legs yellow-reddish, except coxae and clubs of femora brownish; occasionally legs brown, except trochanters, tibiae and tarsi testaceous. Head excavated, shiny, unsculptured, except occiput and anterior region of frons granulate. Palpal formula 5 / 2 (in scutellaris), or 6 / 3 (in trochantericus). Pronotum crossed by strong transverse impression, shiny, unsculptured, bare. Mesoscutum shiny, with some longitudinal keels, with two pointed apophyses laterally (Fig. 240 G). Metanotum excavated behind mesoscutellum (Fig. 240 H). Metapectal-propodeal disc shiny, unsculptured; propodeal declivity more or less transversely striate. In specimen from South Africa, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, top of metapectal-propodeal disc with track of median longitudinal furrow. Mesopleuron not transversely striate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 shorter than 4 (14: 21). Enlarged claw (Fig. 240 K) with one small subapical tooth and 1 – 9 peg-like setae or bristles. Protarsomere 5 (Fig. 240 K) with inner side proximally not serrate and two rows of 25 – 38 lamellae; apex with 7 – 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8E2869FF3ED423FD73F848.taxon	materials_examined	Material examined. Types: ♀ holotype of E. trochantericus: MADAGASCAR: Toliara, Bekily, IV. 1937, A. Seyrig leg. (MNHN); ♀ holotype of G. scutellaris: MADAGASCAR: Toliara, Bekily, I. 1939, A. Seyrig leg. (MNHN). Paratype of E. trochantericus: same locality label as holotype, V. 1937, 1 ♀ (MRAC). Paratypes of G. scutellaris: same locality label as holotype, V. 1937, 2 ♀♀ (MNHN); same locality label as holotype, VIII. 1938, 1 ♀ (MNHN); same locality label as holotype, X. 1938, 1 ♀ (MRAC); same locality label as holotype, XI. 1938, 1 ♀ (MRAC). Other material: MADAGASCAR: Fianarantsoa, Isalo, Analalava, 1 ♀ (AMNH); Fianarantsoa, Isalo National Park, Sahanafa River, 29.2 km 351 ° N Ranohira, 22 ° 18.48 ’ S 45 ° 17.30 ’ E, 500 m, 10 – 13. II. 2003, MT, gallery forest, BLF 7652, Fisher, Griswold et al. leg., 4 ♀♀ (3 in CAS, 1 in MOLC); Mahajanga, Tsingy de Bemaraha National Park, 2.5 km 62 ° ENE Bekopaka, Ankidrodroa River, 19 ° 07.56 ’ S 44 ° 48.53 ’ E, 100 m, 11 – 15. XI. 2001, sifted litter, tropical dry forest on Tsingy, Fisher, Griswold et al. leg., BLF 4340, 1 ♀ (CAS). SOUTH AFRICA: Northern Cape, Three Sisters, 1 ♀ (SANC); Northern Cape, 36 km N Garies, roadside N 7, 9. VIII. 2008, R. Whar- ton leg., 1 ♀ (TAMU); Western Cape, Matjesfontein, 4. XII. 1931, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Koeberg Nature Reserve, 33 ° 37.622 ’ S 18 ° 24.259 ’ E, 17. IV – 15. V. 1998, MT, West Coast Strandveld, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8E2869FF3ED423FD73F848.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8E2869FF3ED423FD73F848.taxon	distribution	Distribution. Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8D286AFF3ED6D7FE14F8E9.taxon	description	(Figs 232 C, 241 A, 243 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8D286AFF3ED6D7FE14F8E9.taxon	description	Description. ♀ (Figs 232 C, 243 A). Apterous; body length 2.7 – 4.0 mm. Head black, except mandible, clypeus and anterior region of frons (with stripe along orbits) testaceous; antenna testaceous, except antennomeres 6 – 10 brown; occasionally antenna almost completely brown; mesosoma black, except lateral regions of pronotum brown; occasionally pronotum mostly brown or brown-testaceous; metasoma brown; legs brown, except trochanters, stalk of meso- and metafemur, part of metatibia and tarsi testaceous (except brown last tarsomere of meso- and metalegs). Antenna clavate; antennomeres in following proportions: 9: 4: 10: 6: 5: 5: 4: 4: 3.5: 5. Head excavated, shiny, unsculptured, except temple and occiput granulate; frontal line complete; occipital carina absent; POL = 2; OL = 2; OOL = 7.5. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, with anterior collar and disc shiny, unsculptured; lateral regions of pronotum sculptured by many longitudinal striae. Mesoscutum shiny, sculptured by many longitudinal striae, laterally with two strong pointed apophyses situated on sides of mesoscutellum (Fig. 232 C). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum very short, transversely striate and deeply excavated behind mesoscutellum (Fig. 243 A). Metapectal-propodeal disc shiny, sculptured by many transverse striae, with top unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 13: 2: 4: 15: 25. Enlarged claw (Fig. 241 A) with one small subapical tooth and one row of four proximal setae. Protarsomere 5 (Fig. 241 A) with two rows of 3 – 5 (proximal) + 26 – 30 lamellae; apex with approximately 11 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8D286AFF3ED6D7FE14F8E9.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Toliara, 7.0 km 156 ° SSE Lavanono, 25 ° 28.3 ’ S 44 ° 59.3 ’ E, 50 m, 17. II. 2002, beating low vegetation, spiny forest / thicket, Fisher et al. leg. (CAS). Paratypes: MAD- AGASCAR: Toliara, Andohahela National Park, Ambohibory Forest, 1.7 km 61 ° ENE of Tsimelahy, 36.1 km 308 ° NW of Tolagnaro, 24 ° 55.48 ’ S 46 ° 38.44 ’ E, 300 m, 16 – 20. I. 2002, beating low vegetation, tropical dry forest, Fisher, Griswold et al. leg., BLF 4918, 1 ♀ (CAS); Toliara, Mahafaly Plateau, 6.2 km 74 ° ENE of Itampolo, 24 ° 39.13 ’ S 43 ° 59.48 ’ E, 80 m, MT, spiny forest thicket, 21 – 25. II. 2002, Fisher, Griswold et al. leg., BLF 5759, 1 ♀ (MOLC); same locality label, beating low vegetation, in spiny forest thicket, 21 – 25. II. 2002, Fisher, Griswold et al. leg., BLF 5760, 1 ♀ (CAS); Toliara, Andohahela National Park, Manantalinjo Forest, 33.6 km 63 ° ENE Amboasary, 7.6 km 99 ° E Hazofotsy, 24 ° 49.1 ’ S 46 ° 36.36 ’ E, 150 m, 12 – 16. I. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 4811, 1 ♀ (CAS); same locality label, 12 – 16. I. 2002, beating vegetation, spiny forest / thicket, Fisher, Griswold et al. leg., BLF 4814, 2 ♀♀ (CAS); Toliara, Tsimanampetsotsa National Park, Bemanateza Forest, 20.7 km 81 ° E Efoetse, 23 km 131 ° SE Beheloka, 23 ° 59.32 ’ S 43 ° 52.50 ’ E, 90 m, 22 – 26. III. 2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 6257, 1 ♀ (CAS); same locality label, 22 – 26. III. 2002, beating low vegetation, in spiny forest thicket, Fisher, Griswold et al. leg., BLF 6256, 4 ♀♀ (3 in CAS, 1 in MOLC); Toliara, Beroboka Forest, 5.9 km 131 ° SE Ankidranoka, 22 ° 13.59 ’ S 43 ° 21.59 ’ E, 80 m, 12 – 16. III. 2002, pitfall trap, in tropical dry forest, Fisher, Griswold et al. leg., BLF 6068, 1 ♀ (CAS); Toliara, Cap Sainte Marie Special Reserve, 12.3 km 262 ° W Marovato, 25 ° 34.54 ’ S 45 ° 10.06 ’ E, 200 m, 11 – 15. II. 2002, beating low vegetation, spiny forest thicket, Fisher, Griswold et al. leg., BLF 5502, 1 ♀ (CAS); Toliara, Tsinjoriaky Forest, 6.2 km 84 ° E Tsifota, 22 ° 48.08 ’ S 43 ° 25.14 ’ E, 70 m, 6 – 10. III. 2002, beating low vegetation, spiny forest thicket, Fisher, Griswold et al. leg., BLF 5968, 2 ♀♀ (CAS). Other material: MADAGASCAR: Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24 ° 56.21 ’ S 46 ° 37.60 ’ E, 180 m, 22 – 29. VI. 2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 20 - 30, 1 ♀ (CAS); Toliara, along national Road 10 from Betioky to Ejeda at Km 119, 21. II. 1984, M. von Tschirnhaus leg., 1 ♀ (MTC); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 49.85 ’ S 46 ° 32.17 ’ E, 80 m, 30. XI – 7. XII. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 40, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8D286AFF3ED6D7FE14F8E9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8D286AFF3ED6D7FE14F8E9.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8C286CFF3ED3A0FC86FC4C.taxon	description	(Fig. 241 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8C286CFF3ED3A0FC86FC4C.taxon	description	Description. ♀. Apterous; body length 3.1 – 4.4 mm. Head yellow-reddish or brown-reddish; antenna yellow-red- dish; occasionally antennomere 10 brown; mesosoma yellow-reddish or brown-reddish; occasionally mesosoma brown, except mesoscutum and metanotum reddish; petiole black; metasoma brownish; legs yellow-reddish, except meso- and metatibia and both extremities of coxae brown. Head excavated, shiny, unsculptured, except anterior and lateral regions of frons and occiput granulate; POL = 5; OL = 6; OOL = 22. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsculptured. Mesoscutum dull, with two pointed apophyses laterally. Metanotum flat, transversely striate, excavated behind mesoscutellum. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomere 1 slightly shorter than 4 (14: 16). Enlarged claw (Fig. 241 B) with one small subapical tooth and 5 – 6 peg-like setae. Protarsomere 5 (Fig. 241 B) with inner side proximally serrate or not serrate, with two rows of 3 – 5 (proximal) + 17 – 20 lamellae; apex with at least 10 – 17 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8C286CFF3ED3A0FC86FC4C.taxon	materials_examined	Material examined. Types: ♀ holotype: SOUTH AFRICA: Eastern Cape, Port St. Johns, IX. 1923, R. E. Turner leg. (NHMUK). Paratypes: SOUTH AFRICA: same locality label as holotype, VIII. 1923, 1 ♀ (MRAC); same lo- cality label as holotype, XI. 1923, 1 ♀ (MRAC); same locality label, I. 1924, 1 ♀ (MRAC); same locality label as holotype, III. 1924, 2 ♀♀ (NHMUK); same locality label as holotype, 15 – 31. VIII. 1923, 1 ♀ (NHMUK); KwaZulu- Natal, near Durban, Kloof, 1500 ft., VIII. 1926, 1 ♀ (NHMUK). Other material: BOTSWANA: Serowe, Farmer’s Brigade, 1 ♀ (USNM). GABON: Ogoové-Maritime Province, Reserve des Monts Doudou, 25.2 km 304 ° NW Dous- sala, 2 ° 13.63 ’ S 10 ° 23.67 ’ E, 660 m, 15. III. 2000, sweep, coastal lowland rainforest, low canopy in forest, S. van Noort leg., 1 ♀ (SAMC). MADAGASCAR: Toliara, Andohahela National Park, Ambohibory Forest, 1.7 km 61 ° ENE of Tsimelahy, 36.1 km 308 ° NW of Tolagnaro, 24 ° 55.48 ’ S 46 ° 38.44 ’ E, 300 m, 16 – 20. I. 2002, beating low vegetation, tropical dry forest, Fisher, Griswold et al. leg., BLF 4918, 1 ♀ (CAS). SOUTH AFRICA: Eastern Cape, Kat- berg, 4000 ft., 14 – 26. XI. 1932, R. E. Turner leg., 1 ♀ (NHMUK); Eastern Cape, Grahamstown, 1 ♀ (SANC); Western Cape, Worcester, 17 – 31. VIII. 1928, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Cape Town, Milnerton, I. 1926, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Matjesfontein, 19 – 31. XII. 1928, R. E. Turner leg., 1 ♀ (NHMUK); Western Cape, Ceres, I. 1925, II. 1925, R. E. Turner leg., 2 ♀♀ (NHMUK); same locality label, 1 ♀ (AMNH); Western Cape, Cape Town, SE of Table Mt., Newlands Forest, 33 ° 58 ’ S 18 ° 28 ’ E, indigenous forest, 4. IV. 2001, 1 ♀ (CAS); Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, 24. I – 21. II. 1998, YPT, S. van Noort & B. Fisher leg., 1 ♀ (SAMC); Western Cape, Koeberg Nature Reserve, 33 ° 37.62 ’ S 18 ° 24.26 ’ E. 20. III – 19. IV. 1998, YPT, S. van Noort leg., 1 ♀ (SAMC); same locality label, 23. I – 20. II. 1998, YPT, S. van Noort leg., 3 ♀♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8C286CFF3ED3A0FC86FC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8C286CFF3ED3A0FC86FC4C.taxon	distribution	Distribution. Botswana, Gabon, Madagascar, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8B286DFF3ED254FCAFFE54.taxon	description	(Figs 240 I, J, 241 C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8B286DFF3ED254FCAFFE54.taxon	description	Description. ♀. Apterous; body length 4.4 – 5.6 mm. Head black, except mandible and clypeus yellow; antenna black, except ventral side of antennomere 1 yellow; mesosoma and metasoma black; legs reddish, except coxae and profemur black or brown. In specimens from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E and 2.07836 ° S 38.22517 ° E, head black, except mandible, part of clypeus and part of malar space testaceous; antenna testaceous, except antennomeres 1, 9, 10 and proximal half of 3 brown; mesosoma and metasoma black; legs testaceous, except coxae and clubs of femora brown. Head excavated, dull, granulate and with short irregular keels. In specimens from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E and 2.07836 ° S 38.22517 ° E, and South Africa, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, head completely rugose; frontal line complete; POL = 2; OL = 2; OOL = 15. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, dull, granulate. In specimens from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E and 2.07836 ° S 38.22517 ° E, pronotum granulate, with disc completely or partly unsculptured. In specimen from South Africa, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, pronotum with many longitudinal striae on lateral regions of anterior collar and around disc. Mesoscutum granulate, with some longitudinal keels, without lateral pointed apophyses (Fig. 240 I). In specimen from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E, mesoscutum completely sculptured by many longitudinal keels. Metanotum rugose, excavated behind mesoscutellum (Fig. 240 J). Metapectal-propodeal disc dull, granulate; propodeal declivity transversely striate (in specimens from Kenya, 2.07836 ° S 38.22517 ° E, and South Africa, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, metapectal-propodeal disc shiny, punctate, unsculptured among punctures). In specimen from South Africa, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, top of metapectal-propodeal disc with track of median longitudinal furrow. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 20: 4: 6: 23: 35. In specimen from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E, Protarsomeres in following proportions: 22: 5: 8: 29: 48. Enlarged claw (Fig. 241 C) with one small subapical tooth and four peg-like setae or bristles. In specimens from Kenya, 00 ° 38.1 ’ N 37 ° 40.4 ’ E and 2.07836 ° S 38.22517 ° E, enlarged claw with one peg-like seta + one bristle. Protarsomere 5 (Fig. 241 C) with inner side proximally not serrate and two rows of 4 + 23 – 24 lamellae, or two rows of 2 – 3 (proximal) + 18 – 30 lamellae; apex with at least 10 – 13 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8B286DFF3ED254FCAFFE54.taxon	materials_examined	Material examined. Type: ♀ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Lomami, Up- emba National Park (MRAC). Other material: KENYA: Eastern Prov., near Ewaso Ngiro River opposite Archer’s Post, 00 ° 38.1 ’ N 37 ° 40.4 ’ E, 2 –. XII. 2002, W. J. Pulawski leg., 1 ♀ (CAS); Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 24. IV – 10. V. 2015, MT, just inside isolated woodland patch, R. Copeland leg., 1 ♀ (NMK). SOUTH AFRICA: Western Cape, Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 26. I – 9. III. 2013, MT, Afromontane Forest, S. van Noort leg., GVB 12 - FOR 1 - M 15, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8B286DFF3ED254FCAFFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8B286DFF3ED254FCAFFE54.taxon	distribution	Distribution. Democratic Republic of the Congo, Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	description	(Figs 241 D, 242)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	diagnosis	Diagnosis. Apterous ♀ of Gonatopus with palpal formula 6 / 3; head excavated; temple without sharp carina; me- sosoma ferruginous (Fig. 242); pronotum crossed by strong transverse furrow; mesoscutum less than twice as long as broad, without lateral pointed apophyses; mesopleuron without lateral pointed apophysis; metanotum deeply ex- cavated behind mesoscutellum; meso-metapleural suture distinct and complete; metapectal-propodeal disc without strong median furrow, shiny, unsculptured; protarsomere 1 shorter than 4; enlarged claw (Fig. 241 D) with one small subapical tooth situated far from apex and some peg-like setae. Description. ♀ (Fig. 242). Apterous; body length 4.0 mm. Ferruginous, except petiole black and antennomeres 9 – 10 darkened. Antenna clavate; antennomeres in following proportions: 10: 7: 20: 13: 9: 8: 7: 6: 5: 8. Head excavated, shiny, with frons partly unsculptured and partly rugose; temple granulate; frontal line complete; occipital carina almost completely absent (shortly present on sides of lateral ocelli, not reaching eyes); occiput setose, partly dull and granulate, partly shiny and unsculptured; POL = 2; OL = 2; OOL = 10. Palpal formula 6 / 3. Pronotum, crossed by strong transverse impression, shiny, with anterior collar and disc slightly granulate. Mesoscutum less than twice as long as broad, shiny, sculptured by longitudinal keels, laterally without pointed apophyses (Fig. 242 C). Mesoscutellum shiny, smooth, sloping anteriorly. Metanotum short, excavated behind mesoscutellum (Fig. 242 B). Metapectalpropodeal disc shiny, unsculptured; propodeal declivity, mesopleuron and metapleuron transversely striate. Mesometapleural suture distinct and complete. Protarsomeres in following proportions: 17: 3: 6: 22: 32. Enlarged claw (Fig. 241 D) with one small subapical tooth and one row of two peg-like setae + one bristle. Protarsomere 5 (Fig. 241 D) with two rows of 3 (proximal) + 29 lamellae; apex with approximately 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	materials_examined	Material examined. Type: ♀ holotype, NMK / INV / T- 645: KENYA: Eastern Prov., Kasaala area, 2.07836 ° S 38.22517 ° E, 733 m, 24. IV – 10. V. 2015, MT, just inside isolated woodland patch, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD8A286EFF3ED46BFDFEF8EA.taxon	etymology	Etymology. The species is named after Carol Jean Wikstrand, renowned immunologist, teacher and longtime friend of one of the authors (RSC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD88286FFF36D6D7FE24FB89.taxon	description	(Fig. 241 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD88286FFF36D6D7FE24FB89.taxon	description	Description. ♀. Apterous; body length 5.5 mm. Head black, except mandible and narrow stripe along margin of clypeus testaceous; clypeus testaceous, except brown median region; antenna brown, except ventral side of antennomere 1 testaceous; mesosoma and metasoma black; legs testaceous, except coxae, trochanters and clubs of femora brown. Antenna clavate, without ADOs; antennomeres in following proportions: 15: 7: 31: 22: 15: 11: 10: 10: 9: 12. Head excavated, dull, with short sparse setae, granulate; temple distinct; frontal line complete; occipital carina absent; POL = 1.5; OL = 2; OOL = 14; greatest breadth of lateral ocelli about as long as OL. Palpal formula 6 / 3. Pronotum dull, crossed by strong transverse impression, strongly granulate, with short sparse setae. Mesoscutum dull, rugose, granulate and sculptured by strong longitudinal keels, without lateral pointed apophyses. Mesoscutellum dull, granulate, sloping anteriorly. Metanotum excavated behind mesoscutellum, flat, granulate and crossed by strong transverse keel. Metapectal-propodeal disc dull, with short sparse setae, granulate; propodeal declivity granulate and with slight tracks of few transverse striae near apex. Mesopleuron and metapleuron dull, granulate, not transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 25: 5: 7.5: 27: 43. Enlarged claw (Fig. 241 E) with one small subdistal tooth and one row of two peg-like setae + one bristle. Protarsomere 5 (Fig. 241 E) with two rows of 5 (proximal) + 27 lamellae; apex with about 16 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD88286FFF36D6D7FE24FB89.taxon	materials_examined	Material examined. Type: ♀ holotype: ZAMBIA: North-Western Province, NC, 82 km SSW Solwezi, 3. XII. 2004, M. Snizek leg. (OLL).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD88286FFF36D6D7FE24FB89.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD88286FFF36D6D7FE24FB89.taxon	distribution	Distribution. Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD882861FF3ED391FDE2FE54.taxon	description	(Figs 244 A, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD882861FF3ED391FDE2FE54.taxon	description	Description. ♀. Apterous; body length 3.5 – 4.0 mm. Completely brown-reddish. Antenna clavate, with ADOs on antennomeres 6 – 10; antennomeres in following proportions: 8: 6: 22: 12: 10: 8: 7: 6: 6: 9. Head slightly excavated, dull, completely strongly granulate; frontal line absent; occipital carina absent; POL = 2; OL = 1.5; OOL = 9. Palpal formula 4 / 2. Pronotum crossed by strong transverse impression, with anterior collar shiny, unsculptured, and with disc dull and granulate. Mesoscutum dull, granulate, without lateral pointed apophyses. Metanotum rugose, excavated behind mesoscutellum, with sides protruding; lateral protrusions of metanotum rounded. Mesopleuron granulate. Metapleuron transversely striate. Meso-metapleural suture distinct and complete. Metapectal-propodeal disc shiny, unsculptured, with longitudinal median furrow; propodeal declivity transversely striate. Protarsomeres in following proportions: 16: 2.5: 5: 13: 22. Enlarged claw (Fig. 244 A) without subapical tooth, with two proximal bristles, two subdistal lamellae and group of four distal lamellae. Protarsomere 5 (Fig. 244 A) with two rows of 10 lamellae; apex with group of approximately 15 lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.2 mm. Head black, except ventral side, temple, malar space, clypeus and man- dible testaceous-dark; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna filiform, with antennomere 3 about 4.5 × as long as broad (9: 2); antennomeres in following proportions: 5: 5: 9: 7: 6.5: 6.5: 6.5: 6.5: 6: 9. Head dull, granulate and rugose; frontal line absent; occipital carina absent; POL = 5; OL = 2; OOL = 2.5; temple very short. Palpal formula 4 / 2. Mesoscutum shiny, slightly granulate. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of lateral ocelli (2: 3). Mesoscutellum and metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, with deep median longitudinal furrow, with anterior surface mostly smooth. Forewing hyaline, without dark transverse bands; 2 R 1 cell open; 2 r-rs & Rs vein curvilinear, with distal part much longer than proximal part (15: 8). Dorsal process of paramere (Fig. 244 C) reduced to inner membranous band. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD882861FF3ED391FDE2FE54.taxon	materials_examined	Material examined. Type: ♀ holotype: MOZAMBIQUE: Maputo Prov., Maputo, Costa do Sol, Municipal Ag- ricultural Nursery, 10. X. 1992, M. Olmi leg. (AMNH). Paratypes: same locality label as holotype, 2 ♀♀ (AMNH); same locality label as holotype, reared from a parasitized adult of Nisia nervosa (Mots.), C. 11. X. 1992, Sf. 6. XI. 1992, M. Olmi reared, 1 ♀ (AMNH). Other material: same locality label as holotype, reared from a parasitized adult of Nisia nervosa (Mots.), C. 26. XI. 1993, Sf. 20. XII. 1993, M. Olmi reared, 1 ♂ (AMNH); same locality label as ho- lotype, reared from a parasitized adult of Nisia nervosa (Mots.), C. 21. IX. 1996, B. 25. IX. 1996, Sf. 22. X. 1996, M. Olmi reared, 1 ♂ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD882861FF3ED391FDE2FE54.taxon	biology_ecology	Hosts. Meenoplidae (Guglielmino et al. 2013): in Mozambique: Nisia nervosa (Motschulsky).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD882861FF3ED391FDE2FE54.taxon	distribution	Distribution. Mozambique.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD862861FF3ED46BFE29FACB.taxon	description	(Figs 243 B, C, 244 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD862861FF3ED46BFE29FACB.taxon	description	Description. ♀ (Figs 243 B, C). Apterous; body length 2.6 mm. Head brown, except mandible, clypeus and anterior region of frons testaceous; antenna with antennomeres 1 – 4 testaceous, other antennomeres missing in holotype; mesosoma brown, with reddish nuances; metasoma black; legs brown, except trochanters and tarsi testaceous. Antennomeres 1 – 4 in following proportions: 9: 6: 24: 13 (other antennomeres missing in only known specimen). Head slightly excavated, shiny, slightly granulate; frontal line thin and complete; occipital carina absent; POL = 2; OL = 2; OOL = 9. Palpal formula 3 / 2, or 4 / 2. Pronotum crossed by strong transverse impression, shiny, unsculptured, except some slight striae on lateral regions. Mesoscutum dull, with many longitudinal striae. Mesoscutellum shiny, unsculptured. Metanotum transversely striate, excavated behind mesoscutellum, with sides protruding; lateral protrusions of metanotum rounded. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture obsolete. Metapectal-propodeal disc shiny, with strong median furrow, unsculptured; propodeal declivity transversely striate. Enlarged claw (Fig. 244 B) without one subapical tooth and one row of six lamellae, in addition to group of five distal lamellae. Protarsomere 5 (Fig. 244 B) with two rows of 11 lamellae; apex with group of seven lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD862861FF3ED46BFE29FACB.taxon	materials_examined	Material examined. Type: ♀ holotype: SENEGAL: Louga Region, 22 km W of Kébémer, Konkoyo (FSAE).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD862861FF3ED46BFE29FACB.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD862861FF3ED46BFE29FACB.taxon	distribution	Distribution. Senegal.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	description	(Figs 244 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	description	Description. ♀. Fully winged; body length 4.5 – 5.6 mm. Holotype with head black, except mandible and clypeus testaceous; antenna testaceous; propleuron black; pronotum black, except lateral regions reddish; rest of mesosoma black; metasoma black; legs testaceous, except coxae and clubs of femora brown or black. In specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, clypeus testaceous-brown; antenna testaceous, except antennomeres 8 – 10 darkened; legs brown, except protrochanter partly yellowish and coxae partly black. Antenna clavate; in holotype antennomeres in following proportions: 11: 6: 21: 13: 11: 7.5: 6.5: 4.5: 6: 6. ADOs present in antennomeres 6 – 10. In holotype, head excavated, shiny, smooth, slightly alutaceous; occipital carina absent (no tracks behind ocelli); frontal line complete; OL longer than POL. In specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’’ E, head unsculptured; POL = 3; OL = 3; OOL = 12; occipital carina shortly present behind ocellar triangle (lateral ocelli touching occipital carina). Palpal formula 6 / 3. In holotype pronotum shiny, slightly granulate, with anterior third crossed by transverse impression; pronotum with anterior collar and disc, without posterior collar (Fig. 244 D); disc of pronotum slightly humped. Mesoscutum shiny, bare, granulate. In holotype and in specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, notauli incomplete, strongly converging, almost reaching posterior margin of mesoscutum. Minimum distance between notauli shorter than POL (2: 3). Mesoscutellum shiny, unsculptured or slightly granulate. Metanotum very reduced and short, reticulate rugose. Holotype and specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, with metapectal-propodeal disc not reticulate rugose, sculptured by many parallel transverse keels, with two median longitudinal keels and two further longitudinal keels situated on lateral region; propodeal declivity with two longitudinal keels. In holotype forewing with three brown spots. Forewing of specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, with two dark transverse bands. Forewing of holotype with distal part of 2 r-rs & Rs vein about as long as proximal part. In holotype and in specimen from Madagascar, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, hind wing hyaline, not darkened. Protarsomere 1 shorter than 4 (18: 22 in holotype). Enlarged claw (Fig. 244 E) with one large subapical tooth and one row of 13 – 16 lamellae. Protarsomere 5 (Fig. 244 E) with two rows of about 41 lamellae extending continuously to apex. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Behara, IV. 1937, A. Seyrig leg. (MNHN). Other material: MADAGASCAR: Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25 ° 00.40 ’ S 46 ° 18.20 ’ E, elev 85 m, 16 – 27. XII. 2002, M. Irwin, F. Parker & R. Harin’Hala leg., MT, spiny forest, MA- 02 - 22 - 10, 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852862FF3ED6D7FD4EF916.taxon	discussion	Remarks. Gonatopus bisulcatus was previously included in Dryinus because of the presence of a small lateral tibial spur on mesotibia. Apparently this small tibial spur was absent in G. obscurus (Ceballos), so it was treated as a Gonatopodine. However, a further study of all known specimens of both species showed that this small tibial spur is present in all specimens. The presence of a small lateral spur on the mesotibia suggests that both species belong to Dryininae. However, in Dryininae the mesotibial spur is very long and situated in a central position at the extremity of the mesotibia, whereas in these specimens the mesotibial spur is very short and situated laterally. Because of this difference, the above species have been moved to the Gonatopodinae. The apparent spur is considered a modified bristle present at the extremity of the mesotibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	description	(Figs 245 A, 246)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	diagnosis	Diagnosis. Macropterous ♀ of Gonatopus with palpal formula 6 / 3; pronotum crossed by deep transverse furrow; forewing with two dark transverse bands; propodeal declivity with two incomplete longitudinal keels; enlarged claw (Fig. 245 A) with one large subapical tooth and one row of 8 – 14 lamellae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	description	Description. ♀ (Fig. 246). Fully winged; body length 3.2 – 4.1 mm. Head black, except mandible and clypeus testaceous (in paratypes area between antennal toruli also testaceous); antenna brown, except ventral side of antennomere 1 testaceous-whitish (in paratype last two antennomeres testaceous); mesosoma black; metasoma brown; legs brown, except articulations and tarsi testaceous. Head dull, bare, granulate; occipital carina incomplete, only present behind and shortly on sides of lateral ocelli; POL = 3; OL = 3; OOL = 8; OPL = 1; greatest breadth of lateral ocelli shorter than OL (2: 3). Palpal formula 6 / 3. Pronotum shiny, bare, granulate, crossed by strong transverse furrow; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum shiny, bare, very slightly granulate. Notauli complete, posteriorly very close and strongly converging; minimum distance between notauli shorter than POL (1: 3). Mesoscutellum shiny, punctate, unsculptured among punctures. Metanotum very reduced and short, reticulate rugose. Metapectal-propodeal disc reticulate rugose, with tracks of two median longitudinal keels; propodeal declivity with two incomplete longitudinal keels. In paratype from South Africa, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, metapectal-propodeal disc reticulate rugose, without tracks of two median longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (17: 6). Hind wing with distal region dark. Protarsomeres of holotype in following proportions: 18: 3: 6: 15: 27. In paratype protarsomere 1 slightly shorter than 4 (21: 22). Enlarged claw (Fig. 245 A) with one large subapical tooth and one row of 8 – 14 lamellae. Protarsomere 5 (Fig. 245 A) with two rows of 18 – 27 lamellae; apex with at least 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Western Cape, West Coast Fossil Park, 5.5 km 270 ° W Langebaanweg, 32 ° 57.759 ’ S 18 ° 05.519 ’ E, 16 – 23. X. 2002, MT, Rehabilitated slimes dam, S. van Noort leg., SAM-HYM-A 023993 (SAMC). Paratypes: SOUTH AFRICA: Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 29. IX – 30. XII. 2010, MT, Hantam Succulent Karoo, KEL 09 - SUC 1 - M 05, SAM-HYM-A 035670, 1 ♀ (SAMC); Northern Cape, Hantam National Botanical Garden, 31 ° 20.602 ’ S 19 ° 11.695 ’ E, 749 m, 28. IX – 29. XII. 2010, MT, Hantam Succulent Karoo, S. van Noort leg., GL 07 - SUC 1 - M 153, SAM-HYM- A 035761 (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD852864FF3ED0ADFD44FC4C.taxon	etymology	Etymology. The species is named costalis after the collection site in the West Coast Fossil Park (costalis is a Latin adjective meaning “ living on the coast ”).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD832865FF3ED253FE14FE0C.taxon	description	(Fig. 245 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD832865FF3ED253FE14FE0C.taxon	description	Description. ♀. Fully winged; body length 4.5 – 6.9 mm. Holotype with head ferruginous-reddish, except occiput and ocellar region black; antenna testaceous, except antennomere 10 darkened; mesosoma and metasoma black; legs testaceous, except clubs of femora darkened. In specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, head ferruginous, except vertex, occiput and posterior half of ventral side black; legs brown, except part of procoxa and protrochanter testaceous. Antenna clavate; in holotype antennomeres in following proportions: 10: 6: 22: 12: 11: 9: 8: 6: 5: 8. ADOs present in antennomeres 6 – 10. In holotype head dull, bare, granulate; occipital carina incomplete, only present behind and shortly on sides of lateral ocelli (laterally not reaching eyes); ocellar triangle equilateral. In specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, POL = 3; OL = 3; OOL = 11. Palpal formula 6 / 3. In holotype pronotum crossed by strong transverse impression, dull, bare, smooth, alutaceous, with some punctures on disc; disc humped; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, bare, granulate. In holotype and in specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, notauli complete, posteriorly very close and strongly converging; minimum distance between notauli shorter than POL (1: 3). Mesoscutellum of holotype shiny, unsculptured. Mesoscutellum of specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, dull, granulate. Metanotum very reduced and short, reticulate rugose. Holotype and specimen from Madagascar, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, with metapectal-propodeal disc reticulate rugose, with two median subparallel longitudinal keels; propodeal declivity with two longitudinal keels. In holotype forewing with two dark transverse bands. Forewing of holotype with distal part of 2 r-rs & Rs vein much longer than proximal part (31: 10). Hind wing with distal region dark. Protarsomere 1 shorter than 4 (15: 19 in holotype). Enlarged claw (Fig. 245 B) with one large subapical tooth and one row of 10 – 12 lamellae. Protarsomere 5 (Fig. 245 B) with two rows of about 29 lamellae; apex with group of 25 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD832865FF3ED253FE14FE0C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Périnet, XI. 1930, A. Seyrig leg. (MNMS). Other material: MADAGASCAR: Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 4 – 12. III. 2002, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B – 19, 1 ♀ (CAS); Mahajanga, Namoroka National Park, 16.40818 S 45.31437 E, 21 – 31. X. 2016, MT, C. Villemant leg., 1 ♀ (MNHN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD832865FF3ED253FE14FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD832865FF3ED253FE14FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822865FF3ED46BFAFBFA13.taxon	description	Description. ♂. Fully winged; body length 1.8 mm (according to the original description). Holotype very damaged, with head, metasoma, legs and right wings missing (originally they were present). According to the original description, head and antenna black, mesosoma black (but it is presently brown); metasoma black; legs black, except tarsi and tibiae yellow. According to the original description, head with frons and vertex shiny and smooth; frontal line incomplete, only anteriorly present; occiput emarginated and concave; POL longer than OOL; mandible tridentate; clypeus regularly rounded and convex; eyes setose. According to the original description, antennomeres of approximately equal lengths. Palpal ratio unknown. According to the direct personal study of the holotype: mesoscutum shiny, slightly granulate; notauli incomplete, reaching approximately 0.5 × length of mesoscutum; mesoscutellum and metanotum shiny, unsculptured; metapectal-propodeal disc shiny, unsculptured, not reticulate rugose; mesopleuron and metapleuron shiny, unsculptured; forewing hyaline, without dark transverse bands, with only costal cell distinctly visible, with distal part of 2 r-rs & Rs vein approximately as long as proximal part. According to the original description, metasoma laterally flat. Genital armature missing. Tibial spurs unknown. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822865FF3ED46BFAFBFA13.taxon	materials_examined	Material examined. Type: ♂ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: North Kivu, Virunga National Park (= Albert National Park), May-ya-Moto, 950 m, 16. IX. 1934, Miss. G. F. de Witte (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822865FF3ED46BFAFBFA13.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822865FF3ED46BFAFBFA13.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822865FF3ED46BFAFBFA13.taxon	discussion	Remarks. The emarginated occiput and unsculptured metapectal-propodeal disc place this species with Gonatopus. This genus is divided into many groups based on ♀♀. The ♀ of this species is unknown, so that it is impos- sible to place it in a group. It is inevitable that it needs to be included among the incertae sedis species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822866FF3ED3A8FAABFF50.taxon	description	Description (according to Kieffer 1913 c). ♀. Apterous; length 3.5 mm. Colour black, except mandible red (except black teeth); clypeus, margins of mouth, narrow stripe along anterior half of orbits, antennomere 1, tarsi and slender part of tibiae white; petiole and lateral regions of pronotum brown-red. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822866FF3ED3A8FAABFF50.taxon	materials_examined	Type material (not examined). ♀ holotype (only known specimen) originally in SAMC. Labelled “ South Af- rica, Cape Town ”. This specimen was sent for identification to Kieffer by Louis Albert Péringuey, former director of the South African Museum. Presently, it is not deposited in the collections of that museum. It was not found in any of the main world museums (including the Natural History Museum in London, England, and the Natural History Museum of Paris, France, where the Kieffer collection was housed). Probably this specimen has to be considered lost.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822866FF3ED3A8FAABFF50.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822866FF3ED3A8FAABFF50.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD822866FF3ED3A8FAABFF50.taxon	discussion	Remarks. The original description is not sufficient to permit the identification of this species, or to include it in the key to the ♀♀ of the Afrotropical Gonatopus. Therefore we include it among the incertae sedis species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812866FF3ED767FBA5FD4E.taxon	diagnosis	Diagnosis. ♀: micropterous (Figs 247 B, C); antenna with ADOs; palpal formula: 6 / 3; pronotum crossed by strong transverse impression (Fig. 247 C), not produced into lobes directed towards tegulae (Fig. 247 C); enlarged claw with apex pointed, with one large subapical tooth and lamellae (Figs 248 A, B); tibial spurs 1 / 0 / 1. ♂: un- known.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812866FF3ED767FBA5FD4E.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812866FF3ED767FBA5FD4E.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812866FF3ED767FBA5FD4E.taxon	discussion	World species. Two species are known, both present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812867FF3ED280FE1FFE99.taxon	description	(Figs 247 B, C, 248 A)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812867FF3ED280FE1FFE99.taxon	description	Description. ♀ (Figs 247 B, C). Micropterous; body length 4.0 mm. Head reddish, except brown spots on vertex; antenna reddish; mesosoma reddish, except brown spots on pronotum, mesopleuron ad metapleuron; petiole black; metasoma reddish. Antenna very slender and long, with ADOs in antennomeres 6 – 10 (two ADOs in antennomere 10); antennomeres in the following proportions: 13: 7: 30: 28: 27: 1: 11: 9: 8: 12; antennomere 3 approximately eleven × as long as broad. Head dull, granulate, with irregular striae and rugosities; vertex swollen; occipital carina incomplete, only visible behind and shortly on sides of lateral ocelli; temple distinct; POL = 6; OL = 5; OOL = 10; OPL = 1. Palpal formula 6 / 3. Pronotum dull, granulate, crossed by strong transverse impression; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, granulate, approximately circular, as long as collar of pronotum. Notauli complete, posteriorly separated. Mesoscutellum shiny, smooth, about 0.5 × as long as mesoscutum. Metanotum shiny, smooth, about 0.5 × as long as mesoscutellum, not excavated behind mesoscutellum. Forewing reduced, as long as mesoscutum. Hind wing absent. Metapectal-propodeal disc humped (as in apterous species of Gonatopodinae), granulate, dull, with anterior surface granulate and rugose; propodeal declivity transversely striate. Mesopleuron and metapleuron transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 16: 3: 5: 18.5: 28. Protrochanter approximately 4 × as long as broad. Enlarged claw (Fig. 248 A) with one large subdistal tooth and one row of seven lamellae. Protarsomere 5 (Fig. 248 A) with two rows of 20 lamellae; apex with approximately 12 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812867FF3ED280FE1FFE99.taxon	materials_examined	Material examined. Type: ♀ holotype: SOUTH AFRICA: Eastern Cape, Algoa Bay, Hans Brauns leg. (MCPM).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812867FF3ED280FE1FFE99.taxon	description	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD812867FF3ED280FE1FFE99.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD802878FF3ED0D5FE31FD3C.taxon	description	(Fig. 248 B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD802878FF3ED0D5FE31FD3C.taxon	description	Description. ♀. Micropterous; body length 2.7 mm. Head brown, except mandible testaceous and clypeus partly testaceous; mesosoma black, except sides of pronotum ferruginous; metasoma brown; legs brown, except junctions, tarsi, trochanters and part of coxae testaceous. Antennae clavate; antennomeres in following proportions: 7: 5: 14: 8: 7: 4.5: 4.5: 4: 3: 5.5. Head flat, dull, granulate, except central area of frons completely unsculptured and shiny; occiput granulate; frontal line complete; occipital carina incomplete, present behind and shortly on sides of lateral ocelli; temple distinct, very long; POL = 1; OL = 2; OOL = 7; OPL = 0.5. Palpal formula 6 / 3. Pronotum shiny, crossed by strong transverse impression, with many longitudinal striae around disc and on lateral regions. Mesoscutum shiny, sculptured by few longitudinal keels, without lateral pointed apophyses. Mesoscutellum shiny, smooth, sloping an- teriorly. Forewing reduced, very short, much shorter than mesoscutum, approximately as long as one third of mesos- cutellum. Hind wing absent. Metanotum deeply excavated behind mesoscutellum, very short. Metapectal-propodeal disc shiny, strongly sculptured by many longitudinal striae, except median unsculptured stripe; metapectal-propo- deal disc laterally rounded, not protruding; propodeal declivity transversely striate. Mesopleuron dull, granulate. Metapleuron dull, granulate and transversely striate. Meso-metapleural suture distinct and complete. Protarsomeres in following proportions: 12: 3: 5: 13.5: 22. Enlarged claw (Fig. 248 B) with one large subdistal tooth and one row of seven lamellae and one bristle. Protarsomere 5 (Fig. 248 B) with two rows of 14 lamellae; apex with group of ap- proximately 16 lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD802878FF3ED0D5FE31FD3C.taxon	materials_examined	Material examined. Type: ♀ holotype: NAMIBIA: Brandberg Plateau, 21 ° 10.50 ’ S 14 ° 32.50 ’ E, 1960 m, 19 – 21. X. 1998, YPT 7, R. Butlin & J. Altringham leg. (SMWN).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD802878FF3ED0D5FE31FD3C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD802878FF3ED0D5FE31FD3C.taxon	distribution	Distribution. Namibia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9F2879FF3ED156FB75FDDB.taxon	diagnosis	Diagnosis. ♀: apterous (Figs 249 A, B); palpal formula 2 / 1; antenna with ADOs; pronotum not crossed by strong transverse furrow or weakly impressed (Fig. 249 B); enlarged claw with one large subapical tooth and one row of lamellae (Fig. 248 C); tibial spurs 1 / 0 / 1. ♂: fully winged (Figs 249 C, D); occipital carina usually absent, oc- casionally incomplete, then present behind and shortly on sides of lateral ocelli; occiput concave; temple present; palpal formula 2 / 1; tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9F2879FF3ED156FB75FDDB.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9F2879FF3ED156FB75FDDB.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9F2879FF3ED156FB75FDDB.taxon	discussion	World species. Six species are known, one in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9F2879FF3ED156FB75FDDB.taxon	discussion	Remarks. Larvae of Haplogonatopus were described by Hernandez & Bellotti (1984).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	description	(Figs 248 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	description	Description. ♀. Apterous; body length 2.0 – 2.2 mm. Head brown, except anterior region of frons, clypeus and mandible testaceous; antenna testaceous, except antennomeres 4 – 10 darkened (in specimens from South Africa, Rocher Pan Nature Reserve, antenna brown, except scape and pedicel testaceous and flagellomere 8 partly yellowwhitish); mesosoma brown-testaceous (in specimens from South Africa, Rocher Pan Nature Reserve, mesosoma mostly brown, except mesoscutum partly or completely yellow and apex of propodeal declivity testaceous); metasoma brown; in ♀ from South Africa, 29 ° 56.086 ’ S 29 ° 26.721 ’ E, mesoscutellum and metapectal-propodeal disc black, except distal extremity of propodeal declivity testaceous; legs testaceous-brown; in specimens from South Africa, Rocher Pan Nature Reserve, legs mostly brown. Antennae clavate; antennomeres in the following proportions: 5: 4: 6: 3: 3: 3: 3: 3: 3: 5.5. Head excavated, granulate; frontal line complete; occipital carina incomplete, present behind and shortly on sides of lateral ocelli; POL = 1; OL = 1; OOL = 4.5. Palpal formula 2 / 1. Pronotum shiny, unsculptured, not crossed by strong transverse impression. Mesoscutum shiny, unsculptured. Metapectal-propodeal disc shiny, unsculptured; propodeal declivity, meso- and metapleuron transversely striate. Meso-metapleural suture obsolete. Protarsomeres of holotype in following proportions: 10: 1.5: 3: 6: 11. Enlarged claw (Fig. 248 C) with one subapical tooth and one row of four lamellae. Protarsomere 5 (Fig. 248 C) with two rows of seven lamellae; apex with three lamellae. Tibial spurs 1 / 0 / 1. ♂. Fully winged; body length 2.0 – 2.3 mm. Head black, except mandible yellow; antenna brown; mesosoma black; metasoma brown; legs testaceous. Antenna filiform; antennomeres in following proportions: 7: 5: 10: 10: 9: 10: 9: 9: 9: 11. Head shiny, setose, slightly granulate; frontal line absent; occipital carina absent; POL = 8; OL = 3; OOL = 5; greatest breadth of lateral ocelli slightly longer than OL (3.5: 3); temple distinct; occiput concave. Palpal formula 2 / 1. Mesoscutum setose, shiny, slightly granulate. Notauli complete, posteriorly joint. Mesoscutellum and metanotum shiny, slightly punctate. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part. Dorsal process of paramere (Fig. 248 D) long, slender, with apex serrate, not distally broadened. Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	materials_examined	Material examined. Type: ♂ holotype: DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Katanga, Lubumbashi (= Elisabethville), II. 1940, H. J. Brédo leg. (MRAC). Other material: MADAGASCAR: Toliara, 60 km NE of Morondava, Bereboka, 1 ♀ (AMNH). NIGER: Maradi, Tarna goulbin, 22 – 25. IX. 1985, G. J. Steck leg., 3 ♂♂ (2 in TAMU, 1 in AMNH). SOUTH AFRICA: Eastern Cape, Port St. Johns, along the road from first to sec- ond beach, picnic area, C. 17. IV. 2007, B. 18. IV. 2007; C. 17. IV. 2007, B. 19. IV. 2007, M. Olmi reared from nymphs of Falcotoya aglauros (Fennah) and Syndelphax capellana (Jacobi), 2 ♀♀ (MOLC, SAMC), 1 ♂ (MOLC); Kwa- Zulu-Natal, Coleford Nature Reserve, 29 ° 56.086 ’ S 29 ° 26.721 ’ E, 2 – 10. XII. 2001, winkler trap, H. Robertson D. et al. leg., 1 ♀ (SAMC); Western Cape, near Velddrif, Rocher Pan Nature Reserve, M. Olmi reared from Falcotoya sporoboli (Lindberg), C. 31. I. 2003, B. 31. I. 2003, Sf. 16. II. 2003, 1 ♀, 1 ♂ (MOLC); same locality label, C. 31. I. 2003, B. 2. II. 2003, Sf. 16. II. 2003, 2 ♀♀, 3 ♂♂ (MOLC); same locality label, 31. I. 2003, B. 3. II. 2003, Sf. 16. II. 2003, 4 ♀♀, 2 ♂♂ (MOLC); same locality label, 31. I. 2003, B. 4. II. 2003, Sf. 16. II. 2003, 1 ♀, 2 ♂♂ (MOLC); 31. I. 2003, B. 6. II. 2003, Sf. 16. II. 2003, 1 ♀, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	biology_ecology	Hosts. Delphacidae (Guglielmino et al. 2013): in South Africa: Falcotoya aglauros (Fennah), Falcotoya sporoboli (Lindberg); Syndelphax capellana (Jacobi) (= Syndelphax nigricaudata (Muir )).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	distribution	Distribution. Democratic Republic of the Congo, Madagascar, Niger, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287AFF3ED6D7FF12F8B8.taxon	discussion	Remarks. The record from China of Xu & He (1995) is erroneous; it is a specimen of Haplogonatopus apicalis Perkins.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287DFF3ED1DFFEE4FEBE.taxon	diagnosis	Diagnosis. ♀: fully winged (Figs 250 A, B); palpal formula 6 / 3; antenna with ADOs; pronotum crossed by strong transverse furrow (Fig. 250 B); enlarged claw with one subapical tooth and one row of lamellae, or peg-like setae, or bristles (Figs 251 B, D); tibial spurs 1 / 0 / 2. ♂: fully winged (Figs 250 C, D); occiput straight in dorsal view; temple absent (Fig. 250 D); occipital carina usually absent; occasionally incomplete, present behind and shortly on sides of lateral ocelli, laterally not reaching eyes; palpal formula 6 / 3; tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287DFF3ED1DFFEE4FEBE.taxon	distribution	Distribution. Worldwide.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287DFF3ED1DFFEE4FEBE.taxon	biology_ecology	Hosts. Flatidae, Nogodinidae, Ricaniidae (Guglielmino et al. 2013).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287DFF3ED1DFFEE4FEBE.taxon	discussion	World species. 53 species are known, six in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9D287DFF3ED1DFFEE4FEBE.taxon	discussion	Remarks. Larvae of Neodryinus were described by Guglielmino & Bückle (2003).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287DFF3ED7C2FAF7FAEB.taxon	description	(Fig. 248 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287DFF3ED7C2FAF7FAEB.taxon	description	Description. ♀. Fully winged; body length 4.4 – 5.0 mm. Head black, except mandible reddish; antenna totally brown, or with flagellomere 3, distal part of 2 and proximal part of 4 yellow; occasionally antenna brown, except ventral side of scape whitish; mesosoma black, occasionally with lateral regions of pronotum testaceous; metasoma brown-black; legs brown-reddish; occasionally legs brown, except tarsi testaceous (except brown last tarsomere of meso- and metalegs). Antenna clavate; antennomeres in following proportions: 15: 7: 37: 17: 13: 10: 9: 8: 7: 10. Head slightly excavated, with many longitudinal keels on frons, vertex and occiput; occipital carina absent; POL = 3; OL = 3; OOL = 13. Palpal formula 6 / 3. Pronotum crossed by transverse impression, dull, granulate and with many longitudinal striae on collar and disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, as sculptured as pronotum, with many longitudinal keels. Notauli absent. Mesoscutellum as sculptured as mesoscutum. Metanotum short, very reduced. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, with or without two longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (13: 10). Protrochanter approximately 4 × as long as broad. Protarsomere 1 longer than 4 (20: 10). Enlarged claw (Fig. 248 E) with one subapical tooth and 4 – 11 peg-like setae or bristles. Protarsomere 5 (Fig. 248 E) with two rows of 9 – 21 lamellae; apex with at least 20 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287DFF3ED7C2FAF7FAEB.taxon	materials_examined	Material examined. Type: ♀ holotype (by mistake the holotype was labelled Neodryinus berlandi n. sp.): MADAGASCAR: [no locality], 1919, De Gaulle collection (MNHN). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 6. V. 2001, sweep, lowland rainforest, marsh clearing, CAR 01 - S 70, S. van Noort leg., 1 ♀ (SAMC). DEMOCRATIC REPUBLIC OF THE CONGO: Haut-Katanga, Lubumbashi (= Elisabethville), 30. III. 1939, 1 ♀ (ISNB). ZAMBIA: Lusaka, 14 – 17. I. 1980, MT, R. A. beaver leg., 1 ♀ (AMNH).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287DFF3ED7C2FAF7FAEB.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287DFF3ED7C2FAF7FAEB.taxon	distribution	Distribution. Central African Republic, Democratic Republic of the Congo, Madagascar, Zambia.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	description	(Figs 251 A, 252)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	diagnosis	Diagnosis. Macropterous ♀ of Neodryinus with mesoscutum unsculptured and forewing with two dark transverse bands.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	description	Description. ♀ (Fig. 252). Fully winged; body length 6.5 – 6.7 mm. Completely testaceous-ferruginous (Fig. 252 A, B), except ocelli partly black, one black spot on mesopleuron and black petiole. Antennae clavate; antennomeres in following proportions: 12: 7: 37: 30: 23: 13: 10: 8: 7: 9; ADOs in antennomeres 6 – 10. Head (Fig. 252 C) shiny, very slightly granulate; frontal line incomplete, only present in posterior half of frons; occipital carina incomplete, only present behind and on sides of lateral ocelli, laterally not reaching eyes; lateral ocelli touching occipital carina; temple slightly prominent; POL = 2.5; OL = 5; OOL = 11; greatest breadth of lateral ocelli shorter than OL (2: 5). Palpal formula 6 / 3. Pronotum crossed by transverse impression, with disc humped; anterior collar and disc shiny, very slightly granulate, slightly rugose on lateral regions of disc; pronotal tubercle absent. Mesoscutum shiny, unsculptured. Notauli (Fig. 252 A) complete, posteriorly separated; minimum distance between notauli shorter than OL (3: 5). Mesoscutellum shiny, unsculptured. Metanotum short and rugose. Metapectal-propodeal disc reticulate rugose, with two complete subparallel longitudinal keels; propodeal declivity reticulate rugose, with two subparallel longitudinal keels. Forewing (Fig. 252 D) with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (27: 13). Protarsomeres in following proportions: 28: 4: 8: 31: 42. Protarsomere 3 produced into hook. Enlarged claw (Fig. 251 A) with one subapical tooth and one row of 12 lamellae. Protarsomere 5 (Fig. 251 A) with two rows of 18 + 24 lamellae; apex with at least 20 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	materials_examined	Material examined. Types: ♀ holotype (CASTYPE 19470): MADAGASCAR: Antsiranana, Antasahabe For- est, 11.4 km 275 ° W Daraina, 13 ° 12.42 ’ S 49 ° 33.24 ’ E, 550 m, 12. XII. 2003, YPT, tropical dry forest, BLF 10115, B. L. Fisher leg. (CAS). Paratype: MADAGASCAR: Antsiranana, Analamerana Reserve, 16.7 km 123 ° Anivorano-Nord, 12 ° 48.17 ’ S 49 ° 22.26 ’ E, 225 m, 3. XII. 2004, YPT, tropical dry forest, B. L. Fisher leg., BLF 11277, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD9A287EFF3ED3F0FED8FDE3.taxon	etymology	Etymology. The species is named bimaculatus, in reference to the two black spots on the mesopleura (one per mesopleuron).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD99287FFF3ED085FE14FBDC.taxon	description	(Figs 251 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD99287FFF3ED085FE14FBDC.taxon	description	Description. ♀. Fully winged; body length 5.6 – 7.8 mm. Head black, except mandible, clypeus and anterior region of frons reddish (in specimen from Madagascar, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, head black, except mandible testaceous); antenna reddish, except antennomeres 7 – 10 darkened (in specimen from Madagascar, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, an- tenna brown, except antennomere 5, distal extremity of 4 and ventral side of 1 – 2 whitish); mesosoma black, except lateral regions and posterior margin of pronotum partly reddish and mesopleuron partly reddish (in specimen from Madagascar, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, mesosoma black, except lateral margins of pronotum ferruginous); metasoma brown-reddish or brown; legs reddish (in specimen from Madagascar, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, legs brown, except tarsi ferruginous and part of procoxa and protrochanter testaceous). Antenna filiform; antennomeres in following proportions: 17: 8: 42: 22: 18: 11: 8: 8: 6: 10. Head (Fig. 251 C) dull, slightly excavated (almost flat), granulate; frons and vertex completely sculptured by longitudinal keels; occipital carina incomplete, only present behind and on sides of lateral ocelli; occiput strongly sculptured by keels or reticulate rugose; temple very prominent (Fig. 251 C). In specimen from Madagascar, 16 ° 19.16 ' S 46 ° 48.80 ' E, POL = 3; OL = 2; OOL = 12; OPL = 1; TL = 7; greatest breadth of lateral ocelli as long as OL. Palpal formula 6 / 3. Pronotum crossed by transverse impression, unsculptured, or partly slightly granulate and with many longitudinal keels; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, completely reticulate rugose and with irregular longitudinal keels. Notauli absent. Mesoscutellum dull, completely reticulate rugose. Metanotum very reduced and short, reticulate rugose. Metapectal-propodeal disc completely reticulate rugose; propodeal declivity reticulate rugose, with two incomplete longitudinal keels. Forewing with two dark transverse bands (proximal band divided into two parts); distal part of 2 r-rs & Rs vein much longer than proximal part (18: 10). Protrochanter approximately six × as long as broad (30: 5). In holotype protarsomere 1 longer than 4 (30: 18). Protarsomeres of specimen from Madagascar, 16 ° 19.16 ' S 46 ° 48.80 ' E, in following proportions: 25: 3: 8: 16: 29. Enlarged claw (Fig. 251 B) with one subapical tooth and one row of six lamellae (in holotype, lamellae missing (Fig. 721 in Olmi 1984 )). Protarsomere 5 (Fig. 251 B) with one row of 14 – 15 lamellae; apex with at least 15 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD99287FFF3ED085FE14FBDC.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Rogez (= Andekaleka), X. 1930, A. Sey- rig leg. (MNMS). Other material: MADAGASCAR: Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16 ° 19.16 ’ S 46 ° 48.80 ’ E, 43 m, 17 – 24. VIII. 2003, MT, deciduous forest, R. Harin’Hala leg., MA- 25 - 11, 1 ♀ (CAS); no locality, 1 ♀ (MRAC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD99287FFF3ED085FE14FBDC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD99287FFF3ED085FE14FBDC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	description	(Figs 251 G, 253)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	diagnosis	Diagnosis. Macropterous ♀ of Neodryinus with occipital carina present behind and on sides of lateral ocelli; mesosoma mostly black; mesoscutum reticulate rugose; hind wing with distal third darkened (Fig. 253 A); enlarged claw with one row of long and slender lamellae (Fig. 251 G).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	description	Description. ♀. Fully winged (Figs 253 A, B); body length 6.2 – 4.4 mm (holotype 5.8 mm). Head black, except mandible, clypeus and area between antennal toruli ferruginous; antenna testaceous-dark, except antennomeres 3, 4 and part of 5 brown; mesosoma black, except lateral regions of pronotum testaceous; metasoma brown; proleg brown, except coxae and trochanters partly whitish. Antenna clavate; antennomeres in following proportions: 15: 6: 31: 16: 12: 10: 8: 7: 7: 9; antennal ADOs present in antennomeres 6 – 10. Head (Figs 253 A, C) dull, slightly excavated, reticulate rugose and granulate; frontal line complete, slender in anterior third of frons; occipital carina incomplete, only present behind and on sides of lateral ocelli (laterally reaching about 0.7 length of OOL); POL = 1; OL = 1; OOL = 13; lateral ocelli almost touching occipital carina; greatest breadth of lateral ocelli longer than POL (3: 1); temple absent. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, shiny, unsetose, sculptured by many longitudinal and parallel striae on anterior collar, lateral regions and disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum (Fig. 253 A) dull, unsetose, reticulate rugose. Notauli absent. Mesoscutellum dull, reticulate rugose, except posterior margin unsculptured. Metanotum dull, short, transverse, rugose. Metapectal-propodeal disc dull, reticulate rugose, sculptured by numerous parallel and longitudinal keels (with many transverse keels among longitudinal keels); propodeal declivity with two complete longitudinal keels; median and lateral areas of propodeal declivity rugose. Mesopleuron dull and rugose. Metapleuron sculptured by many transverse keels. Forewing (Figs 253 A, B) with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (27: 16). Hind wing with distal third darkened. Protarsomeres in following proportions: 28: 4: 9: 21: 35. Enlarged claw (Fig. 251 G) with one subapical tooth and one row of 15 long and slender lamellae + one long seta. Protarsomere 5 (Fig. 251 G) with two rows of 8 + 11 (in holotype) or 8 + 9 lamellae; apex with at least 14 – 20 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	materials_examined	Material examined. Types: ♀ holotype, DR 18002: DEMOCRATIC REPUBLIC OF THE CONGO: Ituri Prov., Djugu Territory, confluence of Ituri and Kelebo rivers, Uphill “ Camp Kelebo ”, 30. III – 6. IV. 2015, primary / secondary forest, 75 YPTs, A. Gumovsky leg., 1 ♀ (SIZK). Paratypes: same locality label, 3 ♀♀ (2 in SIZK, 1 in MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD982870FF3ED2E3FBB1FE70.taxon	etymology	Etymology. The species is named after the type locality, Uphill “ Camp Kelebo ”.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD972871FF3ED075FE32FDE4.taxon	description	(Figs 251 D, E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD972871FF3ED075FE32FDE4.taxon	description	Description. ♀. Fully winged; body length 4.5 mm. Head testaceous-reddish, except ocellar triangle brown; an- tenna testaceous-reddish; mesosoma testaceous-reddish, except mesopleuron, metapectal-propodeal disc, propodeal declivity (except testaceous apex) and sutures around mesoscutum, mesoscutellum and metanotum black; petiole black; metasoma testaceous-reddish; legs testaceous-reddish. Antenna clavate; antennomeres in following proportions: 14: 7: 32: 20: 17: 10: 8: 7: 6: 9; ADOs in antennomeres 6 – 10. Head (Fig. 251 E) shiny, reticulate rugose; frontal line incomplete, only present in front of anterior ocellus and reaching approximately 0.5 × length of frons; occipital carina incomplete, only present on sides of lateral ocelli; lateral ocelli touching occipital carina; temple slightly promi- nent (Fig. 251 E); POL = 3; OL = 2.5; OOL = 12. Pronotum crossed by transverse impression, with disc humped; anterior collar and disc slightly granulate and rugose, with many striae around disc; pronotal tubercles absent. Mesoscutum shiny, with median region reticulate rugose and lateral regions unsculptured. Notauli absent. Mesoscutellum rugose. Metanotum short and rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two complete longitudinal keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein longer than proximal part (18: 8). Protarsomeres in following proportions: 22: 3: 10: 20: 33. Protarsomere 3 produced into hook. Enlarged claw (Fig. 251 D) with one subapical tooth and one row of seven bristles. Protarsomere 5 (Fig. 251 D) with two rows of 27 lamellae; apex with at least 20 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD972871FF3ED075FE32FDE4.taxon	materials_examined	Material examined. Type: ♀ holotype: TANZANIA: Kilimanjaro Region, near Moshi, Kware, 27. XII. 1951 – 13. I. 1952, D. O. Africa Expedition (SMNS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD972871FF3ED075FE32FDE4.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD972871FF3ED075FE32FDE4.taxon	distribution	Distribution. Tanzania.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD962871FF3ED4FBFC91F8E9.taxon	description	(Fig. 251 F)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD962871FF3ED4FBFC91F8E9.taxon	description	Description. ♀. Fully winged; body length 4.7 – 5.0 mm. Head black, except mandible testaceous (in specimen from Uganda, 0 ° 34.390 ’ N 30 ° 21.658 ’ E, clypeus brown); antenna brown (in specimen from Uganda, 0 ° 34.390 ’ N 30 ° 21.658 ’ E, ventral side of scape and pedicel and part of flagellomere 1 testaceous); mesosoma black, except lateral regions of pronotum testaceous; metasoma brown; proleg black, except protarsomeres 3 – 5 and enlarged claw testaceous; meso- and metalegs black, except tarsi testaceous (except brown last tarsomere). In specimen from Uganda, 0 ° 34.390 ’ N 30 ° 21.658 ’ E, extremities of metafemur testaceous. Antenna clavate; antennomeres in following proportions: 10: 6: 29: 17: 12: 11: 9: 8: 7: 9.5; antennal ADOs present in antennomeres 6 – 10. Head dull, slightly excavated, reticulate rugose and granulate; frontal line incomplete, absent in anterior third of frons; occipital carina incomplete, only present behind and on sides of lateral ocelli (laterally reaching about 0.5 length of OOL); POL = 2; OL = 2.5; OOL = 10; lateral ocelli touching occipital carina; temple absent. Palpal formula 6 / 3. Pronotum crossed by strong transverse impression, dull, setose, granulate and with tracks of many longitudinal and parallel striae on anterior collar, lateral regions and disc; posterior edges of pronotum rounded, not produced into lobes. Mesoscutum dull, setose, granulate and reticulate rugose. Notauli absent. Mesoscutellum dull, granulate and reticulate rugose. Metanotum dull, short, transverse, rugose. Metapectal-propodeal disc dull, reticulate rugose, sculptured by numerous parallel and longitudinal keels (with many transverse keels among longitudinal keels); propodeal declivity with two complete longitudinal keels; median and lateral areas of propodeal declivity rugose. Mesopleuron dull and rugose. Metapleuron sculptured by many transverse keels. Forewing with two dark transverse bands; distal part of 2 r-rs & Rs vein much longer than proximal part (18: 9). Hind wing hyaline. Protarsomeres in following proportions: 25: 3: 7: 14: 24. Enlarged claw (Fig. 251 F) with one subapical tooth and one row of 9 – 12 narrow lamellae. Protarsomere 5 (Fig. 251 F) with two rows of 9 – 13 lamellae; apex with at least 14 lamellae. Tibial spurs 1 / 0 / 2. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD962871FF3ED4FBFC91F8E9.taxon	materials_examined	Material examined. Type: ♀ holotype: CAMEROON: Messabengondo, 1 – 15. III. 2003, Marc Tussac leg. (MNHN). Other material: CENTRAL AFRICAN REPUBLIC: Sangha-Mbaéré Pref., Dzanga-Ndoki National Park, Mabéa Bai, 21.4 km 53 ° NE Bayanga, 3 ° 02.01 ’ N 16 ° 24.57 ’ E, 510 m, 4. V. 2001, sweep, lowland rainforest, marsh clearing, CAR 01 - S 24, S. van Noort leg., 1 ♀ (SAMC). UGANDA: Kibale National Park, Kanyawara, Mak- erere University Biological Field Station, 0 ° 34.390 ’ N 30 ° 21.658 ’ E, 1587 m, 4 – 26. VIII. 2008, YPT, primary midaltitude rainforest, near stream, S. van Noort leg., 1 ♀ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD962871FF3ED4FBFC91F8E9.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD962871FF3ED4FBFC91F8E9.taxon	distribution	Distribution. Cameroon, Central African Republic, Uganda.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD952874FF3ED6D4FA7DF877.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD952874FF3ED6D4FA7DF877.taxon	discussion	World species. 13 species are known, 11 in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD952874FF3ED6D4FA7DF877.taxon	discussion	World genera. Seven genera are known, four in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD952874FF3ED6D4FA7DF877.taxon	discussion	Remarks. The following three genera have not been recorded from the Afrotropical region: Bocchopsis Olmi 1991 (Australian), Apodryinus Olmi 1984 (Neotropical) and Peckius Olmi & Virla 2014 (Neotropical).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922875FF3ED6D7FEECFBDE.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922875FF3ED6D7FEECFBDE.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922877FF3ED2E5FE14FAE0.taxon	description	(Figs 254 A, B, 255 A, 256 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922877FF3ED2E5FE14FAE0.taxon	description	Description. ♀ (Figs 254 A, B, 255 A). Apterous; body length 1.9 – 2.2 mm. Head black, except clypeus and mandible testaceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum and apex of propodeal declivity testaceous; metasoma brown, except distal third testaceous; legs testaceous, except metacoxa almost completely brown. Antenna clavate, slender; antennomeres of holotype in following proportions: 10: 4: 5: 6: 6: 6: 6: 6: 6: 10; antennomeres longer than broad; length / breadth ratio of antennomere 9 in lateral view: 6: 3; antennomere 2 longer than broad; ADOs present in antennomeres 3 – 10; antennal toruli contiguous with lateral margins of clypeus. Head spheroidal in dorsal view (Figs 254 B, 255 A), slightly longer than broad (28: 23), smooth, shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head convex (Fig. 255 A); eyes small, much shorter than head (7: 28 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, incomplete on lateral regions; frontal line absent; mandible with four irregular teeth; clypeus with ventral margin rounded; temple very long, less than twice as long as eye (12: 7) (Fig. 254 B); subocular sulcus absent (short slight track present). Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma almost completely fused, with transverse suture separating mesosoma in anterior and posterior part (suture laterally formed by meso-metapleural suture); anterior part of mesosoma composed of pronotum + mesothorax; posterior part of mesosoma composed of metapectal-propodeal disc and propodeal declivity; pronotum + mesothorax completely shiny and unsculptured. Mesothoracic spiracles present. In pronotum + mesothorax, no dorsal sutures visible (laterally two incomplete sutures present, first one separating pronotum from mesopleuron and second one corresponding to epicnemial sulcus); suture between pronotum + mesothorax and metapectal-propodeal disc with two small pointed structures (metathoracic spiracles) situated on sides of dorsal part (Fig. 254 A); posterior part of mesosoma dorsally divided into two regions, first one dorsal and composed of metanotum + metapectal-propodeal disc, second one posterior and inclined corresponding to propodeal declivity; propodeal declivity completely surrounded by keel separating this area from metapectal-propodeal disc and metapleuron (keel laterally with two small pointed apophyses per side (Fig. 254 A); metapectal-propodeal disc laterally with two prominent propodeal spiracles (Fig. 254 A). Metapectal-propodeal disc composed of completely fused sclerites, not showing any dorsal or lateral suture, completely shiny, with few short setae, unsculptured, except few lateral rugosities situated near meso- and metacoxa and two very short distinct lateral longitudinal sutures leaving from lateral pointed apophyses of metapectal-propodeal disc and directed towards propodeal spiracles, not reaching them (Fig. 256 A). Propodeal declivity unsculptured, except few dorsal transverse striae situated near apex. Protarsomeres in following proportions: 11: 3: 2.5: 7: 9. Protarsomere 3 produced into hook. Enlarged claw of holotype (Fig. 256 B) with four small subapical and medial teeth and one row of six slender lamellae. Protarsomere 5 of holotype (Fig. 256 B) with two rows of 4 + 9 lamellae; apex with five lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922877FF3ED2E5FE14FAE0.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Antsiranana, Ampasindava, Ambilanivy Forest, 3.9 km 181 ° S of Ambaliha, 13 ° 47.55 ’ S 48 ° 9.42 ’ E, 600 m, 4 – 9. III. 2001, sifted litter in rainforest, Fisher, Griswold et al. leg., BLF 3252 (CAS). Paratype: same locality label as holotype, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922877FF3ED2E5FE14FAE0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD922877FF3ED2E5FE14FAE0.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD902808FF3ED3F7FE14FBDC.taxon	description	(Figs 255 B, 256 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD902808FF3ED3F7FE14FBDC.taxon	description	Description. ♀ (Figs 255 B, C). Apterous; body length 2.0 mm. Head black, except clypeus and mandible tes- taceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum and apex of propodeal declivity testaceous; metasoma brown, except distal third testaceous; legs testaceous, except metacoxa almost completely brown. Antenna clavate, slender; antennomeres in following proportions: 8: 3.5: 4: 5: 5.5: 5.5: 5.5: 5.5: 5: 10; antennomeres longer than broad; length / breadth ratio of antennomere 9 in lateral view: 5: 3; antennomere 2 longer than broad; ADOs present in antennomeres 3 – 10; antennal toruli contiguous with lateral margins of clypeus. Head spheroidal in dorsal view, slightly broader than long (20: 17), shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head convex; eye small, much shorter than head (5: 17 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, incomplete on lateral regions; frontal line absent; clypeus humped, with ventral margin rounded; temple very long, less than twice as long as eye (7: 5); subocular sulcus absent (short slight track present). Palpal formula 4 / 3. Propleuron apparently mobile and articu- lated, hidden under apparent pronotum. Remaining regions of mesosoma almost completely fused, with transverse suture separating mesosoma in anterior and posterior part (laterally suture separating two above parts formed by meso-metapleural suture); anterior part of mesosoma composed of pronotum + mesothorax; posterior part of mesosoma composed of metapectal-propodeal disc + propodeal declivity; pronotum + mesothorax completely shiny and unsculptured. Mesothoracic spiracles present. In pronotum + mesothorax no dorsal sutures visible (laterally two incomplete sutures present, first one separating pronotum from mesopleuron and second one corresponding to epicnemial sulcus); suture between pronotum + mesothorax and metapectal-propodeal disc with two small pointed structures (metathoracic spiracles) situated on sides of dorsal part (Fig. 255 B); posterior part of mesosoma dorsally divided into two regions, first one dorsal and composed of metanotum + metapectal-propodeal disc, second one posterior and inclined corresponding to propodeal declivity; propodeal declivity completely surrounded by keel separating this area from metapectal-propodeal disc and metapleuron (keel laterally with two small pointed structures per side (Fig. 255 B); metapectal-propodeal disc laterally with two prominent propodeal spiracles (Fig. 255 B). Metapectal-propodeal disc composed of completely fused sclerites, not showing any dorsal or lateral suture, completely smooth, shiny, with few short setae, unsculptured, except few lateral rugosities situated near meso- and metacoxa and two long distinct lateral longitudinal sutures joining lateral pointed structures of metapectal-propodeal disc to three spiracles (mesothoracic, metathoracic, propodeal) and separating metapectal-propodeal disc + metanotum from metapleura and mesonotum from mesopleura (Figs 255 C, 256 C). Propodeal declivity unsculptured, except few dorsal transverse striae situated near apex. Protarsomeres in following proportions: 12: 3: 2: 6.5: 9. Protarsomere 3 produced into hook. Enlarged claw (Fig. 256 D) with two small subapical and medial teeth and one row of eight slender lamellae. Protarsomere 5 (Fig. 256 D) with two rows of 2 + 8 lamellae; apex with five lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD902808FF3ED3F7FE14FBDC.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Fianarantsoa, Atsirakamblaty Forest, 7.6 km 285 ° WNW Itremo, 20 ° 35.36 ’ S 46 ° 33.48 ’ E, 1550 m, 22 – 26. I. 2003, sifted litter, montane rainforest, Fisher, Griswold et al. leg., BLF 7151 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD902808FF3ED3F7FE14FBDC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FD902808FF3ED3F7FE14FBDC.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2808FF3ED2E3FC18F912.taxon	type_taxon	Type species. Gondwanadryinus fisheri Olmi 2007, by monotypy and original designation.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2808FF3ED2E3FC18F912.taxon	diagnosis	Diagnosis. ♀: apterous (Figs 255 D, E): antenna geniculate, with ADOs; occipital carina complete; ocelli absent; propleuron and pronotum mobile and articulated; mesoscutum very short; mesoscutellum, metanotum and metapectal-propodeal disc fused; sutures separating mesoscutellum, metanotum and metapectal-propodeal disc completely obliterated; sutures delimiting propleura, pronotum and mesoscutum distinctly visible; meso-metapleural suture absent; pronotum not crossed by transverse impression (Fig. 255 E); enlarged claw (Fig. 256 E) without subapical teeth; protarsomere 5 (Fig. 256 E) with rows of lamellae; tibial spurs 1 / 0 / 1. ♂: unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2808FF3ED2E3FC18F912.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2808FF3ED2E3FC18F912.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2808FF3ED2E3FC18F912.taxon	discussion	World species. One species is known, present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2809FF3ED0A6FE14FC20.taxon	description	(Figs 255 D, E, 256 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2809FF3ED0A6FE14FC20.taxon	description	Description. ♀ (Figs 255 D, E). Apterous; body length 1.3 mm. Completely testaceous, except petiole and reduced eye black. Antenna clavate, thickened, with ADOs in antennomeres 3 – 10 (two in antennomere 10); antennomeres in following proportions: 6: 2: 1: 2: 2: 2: 2: 2: 2: 3; antennomeres 1 and 10 longer than broad; antennomeres 2 – 9 as long as broad, or broader than long; antennal toruli contiguous with lateral margins of clypeus. Head trapezoidal in dorsal view (Fig. 255 D), broader than long (13: 10), shiny, covered with short and fine setae, slightly granulate; vertex of head flat; frons slightly convex (Fig. 255 E); eye very small, reduced to small group of ommatidia (Figs 255 D, E); ocelli absent; occipital carina complete; frontal line absent; clypeus with ventral margin straight; temple very long (Fig. 255 E). Propleuron mobile and articulated. Pronotum as sculptured as head, with posterior margin curved and with dorsal surface flat. Mesoscutum very short, slightly visible between two posterior angles of pronotum. Remaining part of mesosoma with no sutures separating mesoscutellum, metanotum and metapectal-propodeal disc (sutures also absent laterally); meso-metapleural suture obsolete; mesosoma as sculptured as head; fused part of mesosoma cylindric and divided into two regions (anterior flat dorsal region (corresponding to mesoscutellum, metanotum and metapectal-propodeal disc) and posterior inclined region (corresponding to propodeal declivity )); in anterior dorsal region, no sutures visible; epicnemium not visible (holotype very small, so that is not easy to distinguish an epicnemium); metapectal-propodeal disc as sculptured as head; propodeal declivity anteriorly delimited by strong keel separating it from anterior dorsal region, separated from lateral regions of mesosoma by numerous short transverse keels, almost completely unsculptured, except few transverse striae near apex. Protarsomeres in following proportions: 3: 1: 1.5: 2: 6. Protarsomere 3 produced into hook. Enlarged claw (Fig. 256 E) without subapical teeth, with two proximal bristles. Protarsomere 5 (Fig. 256 E) with two rows of 1 (long) + 8 (short) lamellae; apex with three lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2809FF3ED0A6FE14FC20.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Vohibasia Forest, 59 km N Sakaraha, 22 ° 28 ’ S 44 ° 51 ’ E, 780 m, 13. I. 1996, B. L. Fisher leg., 1310 (DEUCD).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2809FF3ED0A6FE14FC20.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEF2809FF3ED0A6FE14FC20.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEE280AFF3ED5B7FA7DFBE5.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEE280AFF3ED5B7FA7DFBE5.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEE280AFF3ED5B7FA7DFBE5.taxon	discussion	World species. Seven species are known, all in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEE280AFF3ED5B7FA7DFBE5.taxon	discussion	Remarks. Based on the presence of an incomplete suture that is only laterally visible between the pronotum and mesothorax in ♀♀, the genus Madecadryinus is very similar to the Neotropical genus Apodryinus Olmi 1984 (in contrast the Australian genus Bocchopsis Olmi 1991 has the suture between pronotum and mesothorax complete and visible not only laterally, but also dorsally). The main differences between Apodryinus and Madecadryinus are in respect of the meso-metapleural suture (completely absent in Apodryinus, laterally partly present in Madecadryinus), the occipital carina (absent in Apodryinus, almost complete in Madecadryinus), the shape of the head (spheroidal in Apodryinus, cuboidal in Madecadryinus), the eye (larger in Apodryinus, smaller in Madecadryinus), the palpal formula (4 / 3 in Apodryinus, 2 / 2 in Madecadryinus), the shape of the antennae (slenderer in Apodryinus, more thickened in Madecadryinus), the ADOs (present in antennomeres 3 – 10 in Apodryinus, in 4 – 10 in Madecadryinus), the enlarged claw (with lamellae and one subapical tooth in Apodryinus, with bristles or very slender lamellae and with small subapical and medial teeth in Madecadryinus). Based on the absence of an inner slender process of the parameres, the ♂ of Madecadryinus is more similar to the ♂♂ of Apodryinus than those of Bocchopsis.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	description	(Figs 257 A, 258 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	description	Description. ♀ (Fig. 257 A). Apterous; body length 1.4 – 2.2 mm. Head black, except clypeus and mandible testa- ceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum and distal extremity of propodeal declivity testaceous; metasoma brown, except distal half testaceous; legs brown, except tarsi and meso- and metatibia testaceous. In two paratypes from Madagascar, 18 ° 13.31 ’ S 47 ° 17.13 ’ E and 18 ° 28.24 ’ S 47 ° 57.36 ’ E, legs testaceous, except metacoxa partly brown. Antenna (Fig. 257 A) clavate, thickened; antennomeres of holotype in following proportions: 10: 3.5: 1: 3: 3.5: 3.5: 3.5: 3.5: 3.5: 5; antennomeres 1 and 10 longer than broad; antennomeres 3 – 9 longer than broad (length / breadth ratio of antennomere 9 in lateral view: 3.5: 3; antennomere 6 longer than broad: 3.5: 2); antennomere 2 longer than broad (3.5: 2); ADOs present in antennomeres 4 – 10; antennal toruli contiguous with lateral margins of clypeus. Head rectangular in dorsal view (Fig. 257 A), slightly longer than broad (20: 19), shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head flat; frons very convex (Fig. 258 A); eye (Fig. 258 A) very small, much shorter than head (3: 20 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, incomplete on lateral regions; frontal line absent; clypeus with ventral margin rounded; temple very long, about 3 × as long as eye (10: 3); subocular sulcus absent. Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma completely fused with no sutures separating fused regions; laterally two sutures visible, suture between apparent pronotum and mesothorax and meso-metapleural suture (sutures complete, their upper third or fourth being thiner than rest of sutures). In two paratypes from Madagascar, 18 ° 13.31 ’ S 47 ° 17.13 ’ E and 18 ° 28.24 ’ S 47 ° 57.36 ’ E, and in five paratypes from Madagascar, 13 ° 07.06 ’ S 49 ° 13.48 ’ E, lateral suture between apparent pronotum and mesothorax incomplete, upper third being absent. Mesosoma shiny, covered with fine and short setae, finely punctate, unsculptured among punctures; mesosoma cylindric and divided into two regions (anterior flat dorsal region (corresponding to pronotum, mesonotum, metanotum and metapectal-propodeal disc) and posterior inclined region (corresponding to propodeal declivity )); in anterior dorsal region no sutures dorsally visible, except suture separating dorsal region from lateral regions (suture incomplete, being absent in anterior third of dorsal region (probably corresponding to pronotum )) (Fig. 257 A); propodeal declivity completely surrounded by strong keel separating it from metapectal-propodeal disc and from lateral regions of mesosoma; propodeal declivity finely punctate, unsculptured among punctures, with few transverse striae near apex. Protarsomeres in following proportions: 8: 2: 2: 7: 9. Protarsomere 3 produced into hook. Enlarged claw (Fig. 258 B) without subapical teeth (with tracks of three small teeth), with eight very slender lamellae and many setae on outer side. Protarsomere 5 (Fig. 258 B) with two rows of 4 + 10 lamellae; apex with four lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	materials_examined	Material examined. Types: ♀ holotype: MADAGASCAR: Antsiranana, Binara Forest, 9.1 km 233 ° SW Da- raina, 13 ° 15.48 ’ S 49 ° 36.12 ’ E, 650 – 800 m, 3. XII. 2003, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher leg., code BLF 9656, (CAS). Paratypes: MADAGASCAR: Antananarivo, Ambohitantely Special Reserve, Ambohitantely Forest, 20.9 km 72 ° NE of Ankazobe, 18 ° 13.31 ’ S 47 ° 17.13 ’ E, 1410 m, 17 – 22. IV. 2001, sifted litter in montane rainforest, Fisher, Griswold et al. leg., BLF 3694, 1 ♀ (CAS); Antananarivo, 3 km 41 ° NE of Andranomay, 11.5 km 147 ° SSE of Anjozorobe, 18 ° 28.24 ’ S 47 ° 57.36 ’ E, 1300 m, 5 – 13. XII. 2000, montane rainforest, sifted litter, Fisher, Griswold et al. leg., BLF 2378, 1 ♀ (MOLC); Antsiranana, Andavakoera Forest, 21.4 km 75 ° ENE Ambilobe, 4.6 km 356 ° N Betsiaka, 13 ° 07.06 ’ S 49 ° 13.48 ’ E, 425 m, 15. XII. 2003, sifted litter (leaf mold, rotten wood), rainforest, B. L. Fisher leg., BLF 10260, 5 ♀♀ (4 in CAS, 1 in MOLC). Other material: MADAGASCAR: Antsiranana, Binara Forest, 7.5 km 230 ° SW Daraina, 13 ° 15.18 ’ S 49 ° 37.00 ’ E, 375 m, 1. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9556, 1 ♀ (CAS); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 15. II – 6. III. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 03, M. E. Irwin & R. Harin’Hala leg., 1 ♀ (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDED280BFF3ED2FBFD35FBF8.taxon	discussion	Remarks. The two paratypes of M. humicolus from Antananarivo Province. are very similar to the holotype, though they have legs that are differently coloured.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEC280CFF3ED29FFE14FDB8.taxon	description	(Figs 257 B, 258 C, D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEC280CFF3ED29FFE14FDB8.taxon	description	Description. ♀ (Fig. 257 B). Apterous; body length 1.4 mm. Head black, except clypeus and mandible testaceous; antenna testaceous; mesosoma black, except anterior margin of apparent pronotum testaceous; metasoma brown, except apex testaceous; legs testaceous. Antenna (Fig. 257 B) clavate, thickened; antennomeres of holotype in following proportions: 8: 3: 1.5: 3: 3: 3: 3: 3.5: 3.5: 6; antennomeres 1 and 10 longer than broad; antennomeres 3 – 9 as broad as long (length / breadth ratio of antennomere 9 in lateral view: 3.5: 3.5; antennomere 6 as long as broad (3: 3 )); antennomere 2 approximately as long as broad; ADOs present in antennomeres 4 – 10; antennal toruli contiguous with lateral margins of clypeus. Head rectangular in dorsal view, less broad than long (18: 22), dull, covered with short and fine setae, granulate; vertex of head slightly convex; frons very convex; eye (Fig. 258 C) very small, much shorter than head (4: 22 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, but incomplete on lateral regions of head; frontal line absent; mandible with three teeth; clypeus with ventral margin rounded; temple very long, almost 3 × as long as eye (10: 4); subocular sulcus absent. Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma completely fused with no sutures separating fused regions; laterally two incomplete sutures visible (suture between apparent pronotum and mesothorax and meso-metapleural suture; sutures incomplete and only partly visible on lateral regions of mesosoma); mesosoma smooth, covered with fine and short setae; mesosoma (Fig. 257 B) cylindric and divided into two regions (anterior flat dorsal dull and granulate region (corresponding to pronotum, mesonotum, metanotum and metapectal-propodeal disc) and posterior inclined unsculptured region (corresponding to propodeal declivity and showing few transverse striae near apex); in anterior dorsal region no sutures visible, except suture separating dorsal region from lateral regions (suture incomplete, being absent in anterior third of dorsal region probably corresponding to pronotum); propodeal declivity completely surrounded by strong keel separating it from metapectal-propodeal disc and from lateral regions of mesosoma. Protarsomeres in following proportions: 8: 2: 1: 6: 7. Protarsomere 3 produced into hook. Enlarged claw (Fig. 258 D) with one subapical tooth and six bristles. Protarsomere 5 (Fig. 258 D) with two rows of 3 (long) + 9 lamellae; apex with four lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEC280CFF3ED29FFE14FDB8.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toliara, Analavelona Classified Forest, 29.2 km 343 ° NNW Mahaboboka, 22 ° 40.30 ’ S 44 ° 11.24 ’ E, 1100 m, 18 – 22. II. 2003, sifted litter, in montane rainforest, Fish- er, Griswold et al. leg., BLF 7820 (CAS).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEC280CFF3ED29FFE14FDB8.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEC280CFF3ED29FFE14FDB8.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEA280EFF3ED31FFE14FC4C.taxon	description	(Figs 257 C, 258 E, G)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEA280EFF3ED31FFE14FC4C.taxon	description	Description. ♀ (Fig. 257 C). Apterous; body length 1.2 – 2.1 mm. Head black, except clypeus and mandible testa- ceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum and distal extremity of propodeal declivity testaceous; metasoma brown, except distal extremity testaceous; legs testaceous, except club of profemur brown, metacoxa partly brown and meso- and metafemur darkened. Antenna (Fig. 257 C) clavate, thickened; antennomeres in following proportions: 9: 2.5: 1.5: 3: 3.5: 3.5: 3: 3: 3: 7; antennomeres 1 and 10 longer than broad; antennomeres 3 – 9 broader than long or as long as broad (length / breadth ratio of antennomere 9 in lateral view: 3: 3.5; antennomere 6 as broad as long: 3.5: 3.5); antennomere 2 as long as broad; ADOs present in antennomeres 4 – 10; antennal toruli contiguous with lateral margins of clypeus. Head rectangular in dorsal view (Fig. 257 C), broader than long (21.5: 19), shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head flat; frons very convex (Fig. 258 E); eye (Fig. 258 E) small, much shorter than head (4: 19 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, but incomplete on lateral regions of head; frontal line absent; clypeus with ventral margin rounded; temple very long, almost 3 × as long as eye (11: 4); subocular sulcus absent. Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma completely fused with no sutures separating fused regions; laterally in- complete sutures visible (suture between apparent pronotum and mesothorax and meso-metapleural suture); above two sutures incomplete and partly visible on lateral regions of mesosoma. Mesosoma (Fig. 257 C) shiny, covered with fine and short setae, finely punctate, unsculptured among punctures; mesosoma cylindric and divided into two regions (anterior flat dorsal region (corresponding to pronotum, mesonotum, metanotum and metapectal-propodeal disc) and posterior inclined region (corresponding to propodeal declivity )); in anterior dorsal region no sutures visible, except suture separating dorsal region from lateral regions (suture incomplete, being absent in anterior third of dorsal region (probably corresponding to pronotum )) (Fig. 257 C); propodeal declivity completely surrounded by strong keel separating it from metapectal-propodeal disc and from lateral regions of mesosoma; anterior dorsal region and posterior inclined region completely unsculptured, except few transverse striae near apex of posterior region. Protarsomeres in following proportions: 8: 2: 2: 6: 9. Protarsomere 3 produced into hook. Enlarged claw (Fig. 258 G) with three small teeth, 10 bristles and many setae on outer side. Protarsomere 5 (Fig. 258 G) with two rows of 3 + 10 lamellae; apex with four lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEA280EFF3ED31FFE14FC4C.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antsiranana, Marojejy National Park, 25.7 km 32 ° NNE Andapa, 10.3 km 314 ° NW Manantenina, 14 ° 26.42 ’ S 49 ° 44.30 ’ E, 1575 m, 21. XI. 2003, sifted litter (leaf mold, rotten wood), montane rainforest, B. L. Fisher leg., BLF 9242 (CAS). Paratypes: MADAGASCAR: Toamasina, E of Maroantsetra, Antongil Bay, Ambodivoangy, lavage de terre, J. Vadon leg., IV. 1959, 1 ♀ (MRAC); same local- ity label, VIII. 1959, 1 ♀ (MRAC). Other material: Toamasina, Ambatovaky Special Reserve, Sandrangato River, 16 ° 49.03 ’ S 49 ° 17.42 ’ E, 360 m, 25 – 27. II. 2010, sifted litter rainforest, B. L. Fisher et al. leg., BLF 24750, 1 ♀ (CAS); Toamasina, Akirindro Mt., 7.6 km 341 ° Ambinanitelo, 15 ° 17.18 ’ S 49 ° 32.54 ’ E, 600 m, 17 – 21. III. 2003, sifted litter / wood, Fisher, Griswold et al. leg., BLF 8250, 1 ♀ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEA280EFF3ED31FFE14FC4C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDEA280EFF3ED31FFE14FC4C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	description	(Figs 258 F, H, 259)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	diagnosis	Diagnosis. ♂ of Madecadryinus with POL about 3 × as long as OOL (Fig. 258 F) and mesoscutum granulate.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	description	Description. ♂ (Figs 259). Fully winged; body length 1.1 – 1.5 mm. Head brown, except mandible and clyp- eus testaceous; antenna brown-testaceous; mesosoma brown, occasionally black; metasoma brown-testaceous; legs yellow. Antenna filiform, setose, very slender, about as long as body; antennomeres in following proportions: 5: 3: 7: 8: 8: 8: 8: 8: 7: 8. Head (Fig. 259 C) convex, shiny, unsculpured; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; ocelli large (Figs 258 F, 259 C); POL = 3; OL = 1; OOL = 1; minimum distance from eyes to occipital carina as long as OL; TL = 4; greatest breadth of lateral ocelli longer than OL (2: 1); lateral ocelli touching occipital carina; temple very long, shorter than eye (4: 8); clypeus without short median longitudinal keel. Palpal formula 6 / 3. Mesoscutum (Fig. 259 A) dull, completely slightly granulate. Notauli complete, posteriorly separated; minimum distance between notauli less than twice as long as breadth of lateral ocelli (3: 2). Mesoscutellum and metanotum shiny, unsculptured. Epicnemium exposed. Mesopleuron and metapleuron shiny, unsculptured. Metapectal-propodeal disc and propodeal declivity reticulate rugose, with some areolae as large as tegulae. Forewing (Fig. 259 D) hyaline without slight dark transverse bands; distal part of 2 r-rs & Rs vein shorter than proximal part (8: 12); 2 R 1 cell open. Hind wing hyaline, without dark transverse band. Paramere (Fig. 258 H) without inner apical and pointed expansion; basivolsella with apex parallel to distivolsella (Fig. 258 H). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19469): MADAGASCAR: Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 19 – 26. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 17 (CAS). Paratypes: MADAGASCAR: same locality label as holotype, 4 – 16. V. 2003, MA- 02 - 09 C- 60, 1 ♂ (CAS); same locality label as holotype, 12 – 23. IV. 2003, MA- 02 - 09 C- 58, 1 ♂ (MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21 ° 13.57 ’ S 47 ° 22.19 ’ E, 1110 m, 26. II – 4. IV. 2002, MT, high altitude rainforest, R. Harin’Hala leg., MA- 02 - 09 A- 18, 1 ♂ (MOLC); same locality label, 26 – 31. III. 2002, MA- 02 - 09 A- 22, 3 ♂♂ (2 in CAS, 1 in MOLC); Fianarantsoa, Manombo Special Reserve camp site, 32 km SSE of Farafangana, 23 ° 01.31 ’ S 47 ° 43.20 ’ E, 36 m, 27. VII – 10. VIII. 2005, MT, lowland rainforest, M. Irwin, R. Harin’Hala leg., MA- 28 - 24, 2 ♂♂ (CAS, MOLC); same locality label, 24. IV – 10. V. 2005, MA- 28 - 18, 1 ♂ (CAS); same locality label, 6 – - 24. VI. 2005, MA- 28 - 21, 1 ♂ (CAS); same locality label, 30. X – 12. XI. 2004, MA- 28 - 03, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE9280FFF3ED253FA68FE0C.taxon	etymology	Etymology. The species is named ranomafanensis after the collection site in the Ranomafana National Park.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE82802FF3ED045FE14FD00.taxon	description	(Figs 257 D, E, 260 A – D, 261 A – C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE82802FF3ED045FE14FD00.taxon	description	Description. ♀ (Figs 257 D, E, 260 A, B). Apterous; body length 1.6 – 2.0 mm. Head black, except clypeus and mandible testaceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum testaceous; apparent mesothorax black, except small region near mesocoxa testaceous; metasoma brown, except distal half testaceous; legs testaceous. Antenna (Figs 257 D, E, 260 A, B) clavate, thickened; antennomeres of holotype in following proportions: 7: 2: 2: 2.5: 2.5: 2.5: 2.5: 2.5: 2.5: 7; antennomeres 1 and 10 longer than broad; antennomeres 3 – 9 broader than long (length / breadth ratio of antennomere 9 in lateral view: 3.5: 4.5; in dorsal view: 2: 4; antennomere 6 broader than long: 3.5: 2.5); antennomere 2 approximately as long as broad; ADOs present in antennomeres 4 – 10; antennal toruli contiguous with lateral margins of clypeus. Head rectangular in dorsal view (Figs 257 D, 260 A), broader than long (23: 18), shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head flat; frons very convex (Figs 257 E, 260 B); eye very small, much shorter than head (3: 17 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, incomplete on lateral regions of head; frontal line absent; mandible with three teeth; clypeus with ventral margin rounded; temple very long, almost 4 × as long as eye (11: 3) (Fig. 260 A); subocular sulcus absent. Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma (Figs 257 D, E, 260 A, B) completely fused with no sutures separating fused regions; laterally two incomplete sutures visible (suture between apparent pronotum and mesothorax and meso-metapleural suture); above two sutures incomplete and only partly visible on lateral regions of mesosoma (upper third of meso-metapleural suture absent). Mesosoma (Figs 257 D, E, 260 A, B) shiny, covered with fine and short setae, finely punctate, unsculptured among punctures; mesosoma cylindric and divided into two regions (anterior flat dorsal region (corresponding to pronotum, mesonotum, metanotum and metapectal-propodeal disc) and posterior inclined region (corresponding to propodeal declivity )); in anterior dorsal region, no sutures dorsally visible, except suture (Fig. 261 A) separating dorsal region from lateral regions (suture incomplete, being absent in anterior third of dorsal region (probably corresponding to pronotum) and from propodeal ocelli and lateral corners of keel surrounding posterior inclined region of mesosoma); posterior inclined region completely surrounded by strong keel separating it from anterior dorsal region and from lateral regions of mesosoma; posterior inclined region completely unsculptured, except few transverse striae near apex of posterior region. Protarsomeres in following proportions: 7: 2: 2: 4.5: 7.5. Protarsomere 3 produced into hook. Enlarged claw of holotype (Fig. 261 B) without subapical teeth (tracks of small teeth visible), with three proximal bristles and many setae on outer side; enlarged claw of paratype from Madagascar, 18 ° 10.17 ’ S 47 ° 16.55 ’ E, with long row of 11 bristles and three small teeth. Protarsomere 5 of holotype (Fig. 261 B) with two rows of 2 + 9 lamellae (in paratype from Madagascar, 18 ° 10.17 ’ S 47 ° 16.55 ’ E, protarsomere 5 with two rows of 3 + 10 lamellae). Apex of protarsomere 5 with five lamellae. Tibial spurs 1 / 0 / 1. ♂ (Figs 260 C, D, 261 C, 262 A). Fully winged; body length 1.6 mm. Head black, except mandible testaceous (with teeth dark) and clypeus testaceous-dark; antenna testaceous-darkened; mesosoma black; metasoma brown; legs yellow, except metacoxa and club of metafemur brown. Antenna filiform, very slender and long, much longer than body (Fig. 260 C); antennomeres in following proportions: 5.5: 4.5: 9: 11: 10: 10.5: 10: 10: 10: 10. Head convex (Fig. 260 D), shiny, unsculpured; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; ocelli small (Figs 260 C, 262 A); POL = 4.5; OL = 2.5; OOL = 3.5; minimum distance from eyes to occipital carina shorter than OL (2: 2.5); TL = 4.5; lateral ocelli touching occipital carina; temples very long, shorter than eyes (4.5: 7); clypeus with short median longitudinal keel; mandible with four irregular teeth. Palpal formula 6 / 3. Mesoscutum dull, completely granulate. Notauli complete, posteriorly separated; minimum distance between notauli twice as long as greatest breadth of lateral ocelli (4: 2). Mesoscutellum and metanotum shiny, unsculptured. Epicnemium exposed. Metapectal-propodeal disc and propo-deal declivity reticulate rugose, with areolae at least partly as large as tegulae. Forewing with slight dark transverse band; distal part of 2 r-rs & Rs vein longer than proximal part (15: 10); 2 R 1 cell open. Hind wing hyaline, without dark transverse band. Paramere (Fig. 261 C) without inner apical and pointed expansion; basivolsella with apex parallel to distivolsella (Fig. 261 C). Tibial spurs 1 / 1 / 2.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE82802FF3ED045FE14FD00.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, F. C. Sandranantitra, 18 ° 02.9 ’ S 49 ° 05.5 ’ E, 450 m, sifted litter (leaf mold, rotten wood), rainforest, 18 – 21. I. 1999, H. J. Ratsirarson leg. 101 (CAS). Paratype: MADAGASCAR: Toamasina, F. C. Sandranantitra, 18 ° 02.9 ’ S 49 ° 05.5 ’ E, 450 m, sifted litter (leaf mold, rotten wood), rainforest, 18 – 21. I. 1999, H. J. Ratsirarson leg., 101, 1 ♂ (CAS). Other material: MADAGASCAR: Fianarantsoa, Manombo Special Reserve camp site, 33 km SSE of Farafangana, 23 ° 01.31 ’ S 47 ° 43.20 ’ E, 36 m, 24. VI – 10. VII. 2005, MT, lowland rainforest, M. Irwin, R. Harin’Hala leg., MA- 28 - 22, 1 ♂ (MOLC); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km 332 ° Toamasina, 17 ° 55.26 ’ S 49 ° 11.59 ’ E, 390 m, 28. XI. 2005, sifted litter, rainforest, Brian L. Fisher leg., BLF 13125, 2 ♀♀ (CAS, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE82802FF3ED045FE14FD00.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE82802FF3ED045FE14FD00.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE52803FF3ED151FE14FA18.taxon	description	(Figs 261 D, E, G, 262 B, C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE52803FF3ED151FE14FA18.taxon	description	Description. ♀ (Figs 262 B, C). Apterous; body length 1.7 – 2.1 mm. Head black, except clypeus and mandible testaceous; antenna testaceous; mesosoma black, except lateral margin of apparent pronotum and distal extremity of propodeal declivity testaceous; metasoma brown, except distal half testaceous; legs testaceous. Antenna (Figs 262 B, C) clavate, thickened; antennomeres of holotype in following proportions: 10: 3.5: 2: 2.5: 2.5: 2.5: 2.5: 2: 2: 6; antennomeres 1 and 10 longer than broad; antennomeres 3 – 9 broader than long (length / breadth ratio of antennomere 9 in lateral view: 3.5: 2; antennomere 6 broader than long: 3: 2.5); antennomere 2 slightly longer than broad; ADOs present in antennomeres 4 – 10; antennal toruli contiguous with lateral margins of clypeus. Head rectangular in dorsal view, broader than long (25: 20), shiny, covered with short and fine setae, finely punctate, unsculptured among punctures; vertex of head flat; frons very convex (Figs 261 D, 262 C); eye very small, much shorter than head (2: 20 in dorsal view); ocelli absent; occipital carina distinct, complete on dorsal region of head, incomplete on lateral regions of head; frontal line absent; clypeus with ventral margin rounded; temple very long, almost six × as long as eyes (13: 2) (Fig. 261 D); subocular sulcus absent. Palpal formula 2 / 2. Propleuron apparently mobile and articulated, hidden under apparent pronotum. Remaining regions of mesosoma completely fused with no sutures separating fused regions; laterally two incomplete sutures visible (suture between apparent pronotum and mesothorax and meso-metapleural suture); above two sutures incomplete and only partly visible on lateral regions of mesosoma. Mesosoma (Figs 262 B, C) shiny, covered with fine and short setae, finely punctate, unsculptured among punctures; mesosoma cylindric and divided into two regions (anterior flat dorsal region (corresponding to pronotum, mesonotum, metanotum and metapectal-propodeal declivity) and posterior inclined region (corresponding to propodeal declivity); in anterior dorsal region no sutures dorsally visible, except suture separating dorsal region from lateral regions (suture incomplete, being absent in anterior third of dorsal region (probably corresponding to pronotum )) (Fig. 261 E); posterior inclined region completely surrounded by strong keel separating it from anterior dorsal region and from lateral regions of mesosoma; anterior dorsal regions and posterior inclined region completely unsculptured, except few transverse striae near apex of posterior region. Protarsomeres in following proportions: 7: 2: 2: 5: 9. Protarsomere 3 produced into hook. Enlarged claw (Fig. 261 G) without subapical teeth (only with tracks of three small teeth), with 11 bristles and many setae on outer side. Protarsomere 5 (Fig. 261 G) with two rows of 3 + 10 lamellae; apex with five lamellae. Tibial spurs 1 / 0 / 1. ♂. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE52803FF3ED151FE14FA18.taxon	materials_examined	Material examined. Type: ♀ holotype: MADAGASCAR: Antananarivo, Ambohitantely Special Reserve, Ambohitantely Forest, 24.1 km 59 ° NE of Ankazobe, 18 ° 10.17 ’ S 47 ° 16.55 ’ E, 1620 m, 17 – 22. IV. 2001, in botanical garden, sifted litter in montane rainforest, Fisher, Griswold et al. leg., BLF 3720 (CAS). Paratypes: MADA- GASCAR: same locality label as holotype, 2 ♀♀ (CAS, MOLC); Toamasina, Analamay, 18 ° 48.22 ’ S 48 ° 20.13 ’ E, 1068 m, 21. III. 2004, sifted litter (leaf mold, rotten wood), montane rainforest, Malagasy ant team, BLF 10502, 2 ♀♀ (CAS). Other material: MADAGASCAR: Fianarantsoa, Vevembe Forest, 66.6 km 293 ° WNW Farafangana, 22 ° 47.28 ’ S 47 ° 10.55 ’ E, 600 m, 23. IV. 2006, maxi Winkler litter extraction, rainforest transitioning to montane forest, BLF 14120, B. L. Fisher et al. leg., 2 ♀♀ (CAS, MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE52803FF3ED151FE14FA18.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE52803FF3ED151FE14FA18.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE42804FF3ED3BFFE14FA60.taxon	description	(Figs 261 F, 262 D)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE42804FF3ED3BFFE14FA60.taxon	description	Description. ♂ (Fig. 262 D). Fully winged; body length 1.4 – 2.2 mm. Head black, except mandible testaceous and clypeus testaceous-dark; antenna testaceous-darkened; mesosoma black; metasoma brown; legs yellow-testaceous. In paratypes, mesosoma brown-black. In specimen from Madagascar, 17 ° 41.36 ’ S 49 ° 27.37 ’ E, MG- 37 B- 18, mesosoma totally testaceous-dark and metasoma yellow-testaceous. Antenna filiform, very slender and long, much longer than body; antennomeres in following proportions: 6: 5: 10: 12: 10: 12: 12: 11: 10: 10. Head (Fig. 262 D) convex, shiny, unsculpured; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; ocelli large (Fig. 262 D); holotype with POL = 4; OOL = 3; paratype with POL = 4; OL = 1.5; OOL = 2; minimum distance from eye to occipital carina about as long as OL; TL = 4; greatest breadth of lateral ocelli longer than OOL (3: 2); lateral ocelli touching occipital carina; temple very long, shorter than eye (4: 9); mandible with four irregular teeth. Palpal formula 6 / 3. Mesoscutum, mesoscutellum and metanotum shiny, unsculptured (in specimen from Madagascar, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, mesoscutum finely punctate, unsculptured among punctures); notauli complete, posteriorly separated; minimum distance between notauli less than twice as long as greatest breadth of lateral ocelli (5: 3). Epicnemium exposed. Metapectal-propodeal disc and propodeal declivity reticulate rugose, with areolae at least partly as large as tegulae. Forewing very slightly broadly darkened, with darker spot around 2 r-rs & Rs vein; distal part of 2 r-rs & Rs vein slightly longer than proximal part (12: 11); 2 R 1 cell open. Hind wing very slightly darkened. Paramere (Fig. 261 F) without inner apical and pointed expansion; basivolsella with apex parallel to distivolsella (Fig. 261 F). Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE42804FF3ED3BFFE14FA60.taxon	materials_examined	Material examined. Types: ♂ holotype: MADAGASCAR: Toamasina, Akirindro Mt., 7.6 km 341 ° Ambi- nanitelo, 15 ° 17.18 ’ S 49 ° 32.54 ’ E, 600 m, 17 – 21. III. 2003, MT, rainforest, Fisher, Griswold et al. leg., BLF 8253 (CAS). Paratypes: MADAGASCAR: Toamasina, same locality label as holotype, 1 ♂ (CAS); Toamasina, Andasibe National Park, 18 ° 55.58 ’ S 48 ° 24.47 ’ E, botanic garden near entrance to park, 1025 m, 1 – 5. IX. 2001, MT, tropical forest, R. Harin’Hala leg., MA- 01 - 08 B- 11, 2 ♂♂ (CAS, MOLC). Other material: MADAGASCAR: Antsiranana, Binara Forest, 7.5 km 230 ° SW Daraina, 13 ° 15.18 ’ S 49 ° 37.00 ’ E, 375 m, 1. XII. 2003, MT, tropical dry forest, B. L. Fisher leg., BLF 9557, 1 ♂ (CAS); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Man- tella, 14 ° 26.29 ’ S 49 ° 46.44 ’ E, 490 m, 26. IX – 4. X. 2005, MT, low altitude rainforest, M. Irwin, R. Harin’Hala leg., MA- 31 - 32, 1 ♂ (MOLC); Antsiranana, Bekaraoka Forest, 6.8 km 60 ° ENE Daraina, 13 ° 00.00 ’ S 49 ° 42.36 ’ E, 150 m, 7. XII. 2003, tropical dry forest, general collecting night spiders, B. L. Fisher leg., BLF 9876, 1 ♂ (CAS); Antsiranana, Montagne d’Ambre National Park, 12 ° 30.52 ’ S 49 ° 10.53 ’ E, 900 m, 29. I. 2001, netted by hand, M. E. Irwin leg., MA- 01 - 01 A- 05, 2 ♂♂ (CAS, MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31.13 ’ S 49 ° 10.45 ’ E, 1125 m, 29. I – 11. II. 2001, MT, R. Harin’Hala leg., MA- 01 - 01 D- 03, 1 ♂ (CAS); same locality label, 11. II – 4. III. 2001, MA- 01 - 01 D- 04, 1 ♂ (MOLC); Antsiranana, Montagne d’Ambre National Park, 12 ° 31 ’ S 49 ° 11 ’ E, 975 m, 4 – 10. III. 2001, MT, M. E. Irwin, E. I. Schlinger & R. Harin’Hala leg., MA- 01 - 01 B- 06, 3 ♂♂ (2 in CAS, 1 in MOLC); Antsiranana, Sakalava Beach, 12 ° 15.46 ’ S 49 ° 23.51 ’ E, 10 m, 15. II – 6. III. 2001, dwarf littoral forest, MT, across sandy trail, MA- 01 - 04 B- 03, M. E. Irwin & R. Harin’Hala leg., 2 ♂♂ (CAS, MOLC); Fianarantsoa, Ranomafana National Park, Belle Vue at Talatakely, 21 ° 15.99 ’ S 47 ° 25.21 ’ E, 1020 m, 4 – 12. II. 2002, MT, secondary tropical forest, R. Harin’Hala leg., MA- 02 - 09 C- 15, 1 ♂ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21 ° 15.05 ’ S 47 ° 24.43 ’ E, 1130 m, 18 – 27. II. 2003, 9 – 20. III. 2003, MT, mixed tropical forest, R. Harin’Hala leg., MA- 02 - 09 B- 53, 2 ♂♂ (MOLC); Toamasina, Mananara-Nord National Park, 7.1 km 261 ° Antanambe, 16 ° 27.18 ’ S 49 ° 47.15 ’ E, 225 m, 16. XI. 2005, YPT, rainforest, B. L. Fisher et al. leg., BLF 12705, 1 ♂ (CAS); Toamasina, Mobot site, Analalava, 7 km SW of Foulpointe, 17 ° 41.36 ’ S 49 ° 27.37 ’’ E, 60 ft., 11 – 18. I. 2008, MT, on sand in low altitude dense humid forest, MG- 37 B- 18, 1 ♂ (MOLC); same locality label, 21 – 28. IX. 2007, MG- 37 B- 02, 1 ♂ (CAS); same locality label, 25. IV – 2. V. 2008, MG- 37 B- 33, 1 ♂ (CAS); same locality label, 9 – 16. XI. 2007, MG- 37 B- 09, 1 ♂ (MOLC); Toamasina, Ankerana, 18 ° 24.23 ’ S 48 ° 48.09 ’ E, 1108 m, 19 – 24. I. 2012, MT, montane forest, B. L. Fisher et al. leg., BLF 27860, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE42804FF3ED3BFFE14FA60.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE42804FF3ED3BFFE14FA60.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE32805FF3ED077FEC5FEE0.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE32805FF3ED077FEC5FEE0.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE32805FF3ED077FEC5FEE0.taxon	discussion	World species. One species is known, present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE32805FF3ED077FEC5FEE0.taxon	discussion	Remarks. The ♂ of Vannoortia shares the absence of an inner slender process of the parameres and complete notauli with Madecadryinus, but these two genera can be easily separated by the shape of the head (occiput very excavated and minimum distance between eyes and occipital carina much shorter than OOL in Madecadryinus; occiput less excavated and minimum distance between eyes and occipital carina approximately as long as OOL in Vannoortia).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE22806FF3ED7F7FE1FFCAB.taxon	description	(Figs 263 A, B, 264 A, B)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE22806FF3ED7F7FE1FFCAB.taxon	description	♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE22806FF3ED7F7FE1FFCAB.taxon	materials_examined	Material examined. Types: ♂ holotype: SOUTH AFRICA: Western Cape, Walker Bay Nature Reserve, 34 ° 27.41 ’ S 19 ° 21.39 ’ E, IX – XII. 1997, MT, S. van Noort leg., SAM-HYM-P 024845 (SAMC). Paratypes: same locality label as holotype, SAM-HYM-P 024845, 6 ♂♂ (SAMC); same locality label as holotype, 1 ♂ (MOLC). Other material: SOUTH AFRICA: Western Cape, Fynbos Estate, Dragonridge, 33 ° 34.092 ’ S 18 ° 47.680 ’ E, 635 m, 2. IV – 30. VII. 2010, YPT, Boland Granite Fynbos, S. van Noort leg., 1 ♂ (SAMC); Western Cape, Kogelberg Nature Reserve, 34 ° 16.481 ’ S 19 ° 01.033 ’ E, 16. II – 16. III. 1999, MT, Mesic Mountain Fynbos, last burnt c. 1978, S. van Noort leg., 3 ♂♂ (2 in SAMC, 1 in MOLC); same locality label, 16. I – 16. II. 1999, 1 ♂ (SAMC); same locality label, 16. III – 16. IV. 1999, 90 ♂♂ (86 in SAMC, 4 in MOLC); same locality label, 16. IV – 16. V. 1999, 1 ♂ (SAMC); Western Cape, Western Cape, Banghoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 14. IV – 2. VI. 2014, MT, Mesic Mountain Fynbos, BH 12 - FYN 3 - M 18, S. van Noort leg., 2 ♂♂ (SAMC); same locality label, 6 – 28. VI. 2013, Burnt Mesic Mountain Fynbos, BH 12 - FYN 3 - M 10, 5 ♂♂ (3 in SAMC, 2 in MOLC); same locality label, 10. XII. 2012 – 24. I. 2013, BH 12 - FYN 3 - M 05, 42 ♂♂ (39 in SAMC, 3 in MOLC); same locality label, 16. V – 6. VI. 2013, BH 12 - FYN 3 - M 09, 12 ♂♂ (SAMC); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 59.030 ’ S 20 ° 49.128 ’ E, 340 m, 13. V – 21. VI. 2012, MT, over stream, Afromontane Forest, S. van Noort leg., GVB 12 - FOR 1 - M 10, 5 ♂♂ (3 in SAMC, 2 in MOLC); same locality label, 18. V – 30. VI. 2013, GVB 12 - FOR 1 - M 18, 2 ♂♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE22806FF3ED7F7FE1FFCAB.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE22806FF3ED7F7FE1FFCAB.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02807FF3ED552FBEEFBD6.taxon	diagnosis	Diagnosis. ♀: unknown. ♂: see the above diagnosis of the subfamily.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02807FF3ED552FBEEFBD6.taxon	distribution	Distribution. Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02807FF3ED552FBEEFBD6.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02807FF3ED552FBEEFBD6.taxon	discussion	World species. Four species are known, present in the Afrotropical region.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	description	(Figs 264 C, 265)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	diagnosis	Diagnosis. ♂ of Apoaphelopus with POL about as long as OOL; notauli incomplete, reaching about 0.5 × length of mesoscutum.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	description	Description. ♂ (Fig. 265). Fully winged; length 0.7 – 1.1 mm. Head brown-testaceous, except mandible tes- taceous; antenna testaceous; mesosoma and metasoma testaceous; legs yellow-testaceous. Antenna filiform, very slender and long, shorter than body; antennomeres in following proportions: 5: 2.5: 2.5: 2.5: 2.5: 2: 2.5: 2.5: 2.5: 3.5. Head (Figs 265 A, C, E) convex, subspheroidal, dull, granulate, finely setose; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; POL = 2; OL = 1; OOL = 2; TL = 1; posterior ocelli touching occipital carina; temple very long, shorter than eye (1: 3.5). Mesoscutum (Figs 265 A, C), mesoscutellum and metanotum shiny, completely unsculptured. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Epicnemium exposed. Metapectal-propodeal disc and propodeal declivity shiny, almost completely unsculptured, slightly rugose on sides of metapectal-propodeal disc. Mesopleuron and metapleuron shiny, unsculptured. Forewing (Fig. 265 F) without pterostigma, with costal vein very thick; costal cell absent; 2 r-rs & Rs vein present and not reaching margin of forewing; other veins of forewing absent, veins M and Cu being marked by dark stripes. Hind wing with costal vein absent, with two dark longitudinal median stripes. Basivolsella (Fig. 264 C) with distal process parallel to distivolsella. Palpi absent. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	materials_examined	Material examined. Types: ♂ holotype (CASTYPE 19447): MADAGASCAR: Mahajanga, Namoroka National Park, 16.9 km 317 ° NW Vilanandro, 16 ° 24.4 ’ S 45 ° 18.6 ’ E, 100 m, 12 – 16. XI. 2002, MT, tropical dry forest, Fisher et al. leg., BLF 6581 (CAS). Paratypes: MADAGASCAR: Antsiranana, Manongarivo Special Reserve, 17.3 km 218 ° SW Antanambao, 14 ° 01.3 ’ S 48 ° 25.1 ’ E, 1580 m, 27. X. 1998, B. L. Fisher # 1970, sifted litter (leaf mold, rotten wood), rainforest, 1 ♂ (MOLC); Mahajanga, Beanka Forest Reserve, 50.7 km E Maintirano, 17 ° 52.49 ’ S 44 ° 28.08 ’ E, 140 m, 28 – 31. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 23001, 1 ♂ (MOLC); Mahajanga, Beanka Forest Reserve, 50.2 km E Maintirano, 18 ° 01.35 ’ S 44 ° 03.02 ’ E, 250 m, 19 – 26. X. 2009, tropical dry forest on tsingy, MT, B. L. Fisher et al. leg., BLF 22787, 1 ♂ (CAS); Mahajanga, Bemarivo Special Reserve, 23.8 km 223 ° SW Besalampy, 16 ° 55.30 ’ S 44 ° 22.06 ’ E, 19 – 23. XI. 2002, 30 m, sifted litter (leaf mold, rotten wood), in tropical dry forest, Fisher, Griswold et al. leg., BLF 6693, 1 ♂ (CAS); Mahajanga, Namoroka National Park, 21 – 31. X. 2016, MT, C. Villemant leg., 4 ♂♂ (3 in MNHN, 1 in MOLC); Toliara, Andohahela National Park, Ihazofotsy Parcelle III, 24 ° 50.09 ’ S 46 ° 29.21 ’ E, 80 m, 28. IV – 6. V. 2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02 - 21 - 17, 1 ♂ (CAS); Toliara, Makay Mts., 21 ° 13.11 ’ S, 45 ° 19.26 ’ E, 500 m, 24. XI – 12. XII. 2010, MT, gallery forest, sandy soil, B. L. Fisher et al. leg., BLF 25357, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	distribution	Distribution. Madagascar.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	etymology	Etymology. The species is named after Brian L. Fisher, one of the collectors.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDE02819FF3ED03DFBFEFC91.taxon	discussion	Remarks. Apoaphelopus fisheri is the smallest known species of Dryinidae.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFE2819FF3ED526FE1FF816.taxon	description	(Figs 264 D, 266 A-C)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFE2819FF3ED526FE1FF816.taxon	description	Description. ♂ (Figs 266 A-C). Fully winged; length 0.8 – 1.1 mm. Head brown, except mandible and clypeus testa- ceous; antenna testaceous-darkened, except scape and pedicel yellow-testaceous; mesosoma brown; metasoma and legs yellow-testaceous; petiole black. Antenna filiform, very slender and long, shorter than body (Figs 266 A-C); antennomeres in following proportions: 6: 3: 3: 3: 3: 3.5: 3: 3.5: 4: 5. Head convex, shiny, unsculptured; occiput very concave, unsculptured; frontal line absent; occipital carina complete; POL = 2; OL = 1; OOL = 3; minimum distance from eye to occipital carina about as long as POL; TL = 3; posterior ocelli touching occipital carina; temple very long, shorter than eye (3: 4); mandible with one tooth. Mesoscutum, mesoscutellum and metanotum shiny, completely unsculptured. Notauli absent. Epicnemium exposed. Metapectal-propodeal disc and propodeal declivity dull, reticulate rugose. Mesopleuron and metapleuron shiny, unsculptured. Forewing without pterostigma, with costal cell absent, with costal vein very thick; 2 r-rs & Rs vein present and not reaching margin of forewing; other veins of forewing absent, veins M and Cu being marked by dark stripes. Hind wing with costal vein present and with two dark longitudinal median stripes. Basivolsella (Fig. 264 D) with distal process parallel to distivolsella. Palpi absent. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFE2819FF3ED526FE1FF816.taxon	materials_examined	Material examined. Type: ♂ holotype: SOUTH AFRICA: KwaZulu-Natal, Louwsberg, Gwala Gwala, 27 ° 34 ’ S 31 ° 17.9 ’ E, 1090 m, 2 – 3. VI. 2005, YPT, M. Mostovski leg. (NMSA). Other material: SOUTH AFRICA: KwaZulu- Natal, Vernon Crookes Nature Reserve, 250 m, 30 ° 17.4 ’ S 30 ° 36.9 ’ E, 24. I – 10. V. 2006, MT, M. Mostovski leg., 9 ♂♂ (6 in NMSA, 3 in MOLC); Northern Cape, 30 km N of Calvinia, Kelkiewyn Farm, 31 ° 12.923 ’ S 19 ° 40.812 ’ E, 762 m, 29. IX – 30. XII. 2010, MT, Hantam Succulent Karoo, KEL 09 - SUC 1 - M 05, 5 ♂♂ (3 in SAMC, 2 in MOLC); same lo- cality label, YPT, KEL 09 - SUC 1 - Y 08, 1 ♂ (SAMC); same locality label, YPT, KEL 09 - SUC 1 - Y 07, 10 ♂♂ (SAMC); Western Cape, Grootvadersbosch Nature Reserve, 33 ° 58.888 ’ S 20 ° 48.885 ’ E, 454 m, 26. X. 2010 – 26. I. 2011, YPT, South Langeberg Sandstone Fynbos, S. van Noort leg., GVB 10 - FYN 1 - Y 09, 1 ♂ (SAMC); same locality label, MT, GVB 10 - FYN 1 - M 08, 1 ♂ (SAMC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFE2819FF3ED526FE1FF816.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFE2819FF3ED526FE1FF816.taxon	distribution	Distribution. South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFD281BFF3ED333FD7EFE54.taxon	description	(Fig. 264 E)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFD281BFF3ED333FD7EFE54.taxon	description	Description. ♂. Fully winged; length 1.1 – 1.3 mm. Head brown-testaceous, except mandible testaceous; antenna testaceous; mesosoma and metasoma testaceous-darkened; legs yellow-testaceous. Antenna filiform, very slender and long, shorter than body; antennomeres in following proportions: 7: 3: 2.5: 3: 3.5: 4: 3.5: 3.5: 3: 5. Head convex, subspheroidal, dull, granulate, finely setose; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; POL = 3; OL = 1.5; OOL = 2; minimum distance from eye to occipital carina about as long as OL; TL = 2.5; posterior ocelli touching occipital carina; temple very long, shorter than eye (2.5: 5); mandible with one tooth. Mesoscutum, mesoscutellum and metanotum shiny, completely unsculptured. Notauli absent. Epicnemium exposed. Metapectal-propodeal disc and propodeal declivity shiny, completely unsculptured. Meso- and metapleuron shiny, unsculptured. In specimens from South Africa, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, meso- and metapleuron rugose. Forewing without pterostigma, without costal cell, with costal vein very thick; 2 r-rs & Rs vein present and not reaching margin of forewing; other veins of forewing absent, veins M and Cu being marked by dark stripes. Hind wing with costal vein absent, with two dark longitudinal median stripes. Basivolsella (Fig. 264 E) with distal process parallel to distivolsella. Palpi absent. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFD281BFF3ED333FD7EFE54.taxon	materials_examined	Material examined. Type: ♂ holotype: MOZAMBIQUE: Niassa, Cuamba District, Cuamba, Farm of the Catholic University of Mozambique, 600 m, 7. IV – 2. VI. 2006, MT, M. Olmi leg. (MOLC). Other material: MO- ZAMBIQUE: same locality label as holotype, 4 – 18. IX. 2007, 1 ♂ (MOLC). SOUTH AFRICA: Western Cape, Bang- hoek Valley, Dwarsriviershoek Farm, 33 ° 56.232 ’ S 18 ° 57.711 ’ E, 410 m, 3 – 26. IV. 2013, MT, Burnt Mesic Mountain Fynbos, BH 12 - FYN 3 - M 07, S. van Noort leg., 1 ♂ (SAMC); same locality label, 10. XII. 2012 – 24. I. 2013, BH 12 - FYN 3 - M 05, 1 ♂ (MOLC).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFD281BFF3ED333FD7EFE54.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFD281BFF3ED333FD7EFE54.taxon	distribution	Distribution. Mozambique, South Africa.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	description	(Figs 264 F, 266 D, 267)	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	diagnosis	Diagnosis. ♂ of Apoaphelopus with POL about twice as long as OOL; notauli incomplete, reaching about 0.5 × length of mesoscutum.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	description	Description. ♂ (Figs 266 D, 267). Fully winged; length 0.9 mm. Head testaceous-dark, except mandible tes- taceous; antenna whitish; mesosoma and metasoma testaceous; legs yellow-testaceous. Antenna filiform, shorter than body; antennomeres in following proportions: 7: 3: 2: 2.5: 2.5: 3: 2.5: 2: 2: 5. Head convex, subspheroidal, dull, granulate, finely setose; occiput very concave, unsculptured; frontal line absent; central area of frons with slight short median longitudinal furrow; occipital carina complete; POL = 3; OL = 1.5; OOL = 1.5; TL = 1; posterior ocelli touching occipital carina; temple shorter than eye (1: 5). Mesoscutum, mesoscutellum and metanotum shiny, completely unsculptured. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Epicnemium exposed. Metapectal-propodeal disc and propodeal declivity shiny, almost completely unsculptured, slightly rugose on sides of metapectal-propodeal disc. Mesopleuron and metapleuron shiny, unsculptured. Forewing without pterostigma, with costal vein very thick; costal cell absent; 2 r-rs & Rs vein present and not reaching margin of forewing; other veins of forewing absent, veins M and Cu being marked by dark stripes. Hind wing with costal vein absent, with two dark longitudinal median stripes. Basivolsella (Fig. 264 F) with distal process parallel to distivolsella. Palpi absent. Tibial spurs 1 / 1 / 2. ♀. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	materials_examined	Material examined. Type: ♂ holotype, NMK / INV / T- 638: KENYA: Coast Prov., Diani Beach area, 4.27559 ° S 39.59337 ° E, 10 m, MT, shrubland off Diani Beach Rd., 8 – 22. I. 2014, R. Copeland leg. (NMK).	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	biology_ecology	Hosts. Unknown.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	distribution	Distribution. Kenya.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
03E987E2FDFB281CFF3ED6D7FE9FFBDC.taxon	etymology	Etymology. The species is named in honour of the great naturalist, explorer and writer, Alfred Russel Wallace, co-founder, together with Charles Darwin, of modern evolutionary theory, and himself a collector of many rare and undescribed species.	en	Olmi, Massimo, Copeland, Robert S., Noort, Simon Van (2019): Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea). Zootaxa 4630 (1): 1-619, DOI: 10.11646/zootaxa.4630.1.1
