taxonID	type	description	language	source
03E9D425FFC3FF8EFECFF289FEDB76C3.taxon	materials_examined	Type species: Omophron variegatum Olivier, 1811	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC3FF8EFECFF129FD0B7643.taxon	materials_examined	Type species: Carabus limbatus Fabricius, 1776 = Scolytus Fabricius, 1790: 221 [HM] (type sp. Carabus limbatus Fabricius, 1777) (preoccupied by Scolytus Geoffroy, 1762)	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC3FF8FFECFF309FCB271AE.taxon	description	Description. Body length 8.12 – 9.01 mm; width 5.24 – 5.38 mm (holotype 8.14 mm and 5.28 mm, respectively). Habitus (Fig. 1). Colour: Head, pronotum, elytra, legs, and antennae yellowish brown. Patches of pronotum and elytral patterns brown. Mandibles yellowish brown, their apical part dark brown, almost black. Ventral side excluding epipleura yellowish brown; epipleural sides of elytra, pronotum, and prosternum lighter. Head fairly flat, coarsely punctured laterally and posteriorly, sparsely punctured around clypeus and on the middle of frons. In the middle of head punctures smaller and more shallow. With one supraorbital setiferous puncture on each side of head near the eye. Subocular ridge well developed, curved upwards and slightly rounded. Clypeus wrinkled, bisetose. Labrum distinctly emarginated, with jagged sides and six setae near the front margin. Anterior angles of labrum acute, expressly curved upwards. First segment of antennae unisetose at apex, 5 – 11 segments covered with thin setae, 1 – 4 segments bare, only with some longer setae near apex, HW: 2.64 – 2.72 mm. Pronotum moderately convex, base bisinuated on each side, sides slightly rounded, with edge expanding toward the anterior margin. Anterior angles nearly acute. Apex and base of pronotum coarsely punctured, disc finely punctured. Median impression well developed. The front part of proepisternum smooth, posterior part with coarse puncture, strongly inclined towards the mesepisternum. Prosternum with gentle and irregular puncture. PL: 2.32 – 2.39 mm, PW: 4.02 – 4.15 mm. Elytra moderately convex, oval. Elytra with 15 full striae. 1 – 5 interstriae flat on the base, striae composed of large punctures only. 6 – 13 interstriae convex, striae deep. At the apical two thirds of elytra 3 rd stria joining 4 th and 7 th joining 8 th, 1 stria remains evanescent in the middle of elytra. 12 th stria joining 13 th on the base of elytron. Metasternum and metepimeron gently punctured. Sides of sterna slightly wrinkled. EL: 5.35 – 5.51 mm, EW: 5.24 – 5.38 mm.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC3FF8FFECFF309FCB271AE.taxon	discussion	Comments. Until now O. (P.) alluaudi was considered to be a synonym of O. (P.) vittulatum, however, when comparing the holotypes of both taxa, sufficient morphological differences have been observed to regard them as two valid species. In comparison with O. (P.) vittulatum, O. (P.) alluaudi is larger, its dark elytral pattern clearly bordered, and the impression on base of pronotum more pronounced. Sides of pronotum in the case of O. (P.) alluaudi are moderately rounded, but in the case of O. (P.) vittulatum are almost straight. There are no specific differences observed among the species of genus Omophron (Phrator) in the structure of aedeagus.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC3FF8FFECFF309FCB271AE.taxon	distribution	Distribution. Western to Central Tropical Africa. Distribution map (Fig. 5).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC3FF8FFECFF309FCB271AE.taxon	materials_examined	Material examined. Bimu (2 female, 1 male, Glauning leg.); Angola: Lunda Norte Province, Dundo, 08. - 09.1948 (1 male, de Baross Machado leg.) (MRAC); Lunda Norte Province, Riv. Tshlkapa, 50 km S. O. de Dundo, 07.1948 (1 male, 1 female, de Baross Machado leg.) (MRAC); Cameroon: Edea, Bsse-Sanaga, Vadon! (1 female, 1 male) (MNHN); Edea (1, Le Moult leg.) (ETHZ); Edea, (Base Senegal) (1 female, Vadon leg.) (MNHN), Younde (1 male) (MRAC); Benue b. Lau, 17.04.1905 (1 female, Glauning leg.); Benue (2 female, 1 male, Glauning leg.) (ZMHB); Central African Republic: Kemoperfecture, Sibut, (1, Le Moult leg.) (ETHZ); Democratic Republic of Congo: Congo: (1) (ETHZ); Congo (1 female) (MNHN); Kasai, Kondue (1 female, Luja leg.) (holotype) (ISNB); “ Bata ”, 05.1924 (1 female, Donchrind) (ETHZ); “ Lambarinia ”, 1924 (2 males, Haug) (ETHZ); “ Lambarinia ”, 1896 (2 females, 1 male, Haug leg.) (MNHN); Kasai, Djokopunda, 10.08.1921 (1 female, Schouteden leg.) (ETHZ); Kasai, Djokopunda, 10.08.1921 (18 males, 20 females, Schouteden leg.) (MRAC); Kasai, Ilebo, 14 - 15.07.1925 (1 male, Prince Leopold leg.) (MRAC); Kasai-Occidental Province, Bena Makima, 08.08.1921 (3 males, 3 females, Ghesquiere leg.) (MRAC); Sud-Ubangi Province, Libenge, 04.01.1948 (25 males, 18 females, Cremer & Neuman leg.); 06.01.1948 (5 males, 8 females, Cremer & Neuman leg.); Sud-Ubangi Province, Libenge, Mawuya, 06.01.1948 (1 male, Cremer & Neuman leg.) (ISNB); Sud-Ubangi Province, Libenge, Ubangi river, 04.01.1948 (29 males, 33 females, Cremer & Neuman leg.) (ISNB); Sud-Ubangi Province, Libenge, Zambi, 06.01.1948 (2 males, 23 females, Cremer & Neuman leg.) (ISNB); Sud-Ubangi Province, Libenge, Zambi, 06.01.1948 (7 males, 9 females, Cremer & Neuman leg.) (MRAC); Mai-Ndombe Province, Inongo, Lake Mai-Ndombe, 11. - 24.06.1925 (1 male, Prince Leopold leg.) (ISNB); Sankuru Province, Sankuru river, 09.1924 (19 males, 23 females, Ghesquiere leg.) (MRAC); Orientale Province, Kungulu; 06.1923 (1 male, 2 females, Ghesquiere leg.) (MRAC); Lomami river, 08.1924 (1 female, Ghesquiere leg.) (MRAC); De Kwamoutha, Port Irangui, 05.1930 (1 male, 1 female, Vanderyst leg.) (MRAC); Basongo, 10.09.1930 (1 male, De Vitte leg.) (MRAC); Basongo, River Kasai, 06.1929 (1 male, 1 female, Overlaet leg.) (MRAC); Basongo, 10.09.1930 (1 female, Witte leg.) (MNHN); “ Env. Lac Leipold II ”, 11. - 24.06.1925 (1 female, Prince Leopold leg.); Haut-Zaire, Bambesa, Alipago, 02.1935 (9 males, 4 females, Leroy leg.) (MRAC); Haut-Zaire, Equater, Bokuma, 1938 (1 male, Hulstaert leg.) (MRAC); Equater, Bokuma, 07.1952 (1 male, Lootens leg.) (MRAC); 1953 (2 males, 2 females, Lootens leg.); Tshuapa, Bokuma, 1953 (2 males, Lootens leg.); 03.1954 (2 males, 2 females, Lootens leg.) (MRAC); Kimpangu river, Fulezi, 850 m, 06.07.1949 (6 males, 3 females, Leleup leg.) (MRAC); Lomami, 08.1924 (1 female, Ghesquiere leg.) (MNHN); Dutch Zambezi (1 male) (MNHN); Fort Archambault, Mission Chari-Tchad, 1904 (1 female, Decorse leg.) (MNHN); Fort Archambault, Moyen Chari, Bongoul, Mission Chari-Tchad (Ba Kare) (MNHN), 04.1904 (2 male, Decorse leg.) (MNHN); Fort Archambault, Baha Sara, Mission Chari-Tchad, 04.1904 (1 male, Decorse leg.) (MNHN); Congo Belge, Rives du Congo, Mission du Bourg de Bozas, 1903 (1 female) (MNHN); Gabon: Ogowe (1 male) (MNHN); Ogooue, Lambarene, 1898 (5 females, 9 males, Haug leg.) (MNHN), Bas. Ogooue, Lambarene, 1901 (1 male, Haug leg.) (MNHN); Ogooue, Lambarene, 1901 (1 female, 1 male, Haug leg.) (MNHN); Congo, Libreville, 1899 (1 female, 3 male, Boucher leg.) (MNHN); Mali: Macina, Dioura, OICMA, 20.09.1969 (1 female, Farrow leg.) (BMNH); Macina, Tilembeya, Niger Bank, OICMA, 05.06.1969 (2 males, 1 female, Farrow leg.) (BMNH); Kara, Flood plain, OICMA, 29.06.1969 (5 males, 6 females, Farrow leg.) (BMNH); Nigeria: Adamawa State, Yola, Benue river, 12. - 20.04.03 (1 male, Schultze leg.) (ETHZ); Benue, 16.05.1903 (1 male) (ZMHB); Tarbata State, Sardauna, Sunkuru, 09.1924 (1 male, Ghesquiere) (ETHZ); Senegal: Clermont (1, Nodier leg.) (ETHZ); Haut Senegal (1 female) Comp. Type Basilewsky (MRAC); Haut Senegal (1 female) (MRAC); Senegal (1 female) (MNHN).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC2FF8BFCE6F5F4FD9775EE.taxon	description	Description: Body length 6.7 – 7.11 mm; width 4.29 – 4.43 mm (holotype 7.09 mm and 4.41 mm, respectively). Habitus (Fig. 2). Colour: Head, pronotum, and elytra brown. Legs and antennae amber yellow. Patches of pronotum and elytral patterns phthalo green, with metalic lustre. Mandibles near base amber yellow, their apical part dark brown, almost black. Ventral side excluding epipleura and sternites brown; epipleura and sternites amber yellow. Head fairly flat, sparsely punctured, gently wrinkled on vertex and frons, more wrinkled near eyes. Subocular ridge well developed, curved upwards and slightly rounded. Head with one supraorbital setiferous puncture on each side of head near the eye. Frons with gentle but broad impression toward clypeus. Clypeus convex, frontal part smooth, near labrum wrinkled, bisetose. Labrum with moderately rounded sides and six setae near the front margin. First segment of antennae unisetose at apex, 5 – 11 segments covered with thin setae, 1 – 4 segments bare, only with some longer setae near apex, HW: 2.21 – 2.35 mm. Pronotum moderately convex, base bisinuated, edge weakly expressed. On the base lateral edge curved towards the exterior, but near the head curved towards the interior. Anterior angles nearly acute. Base, disc and apex of pronotum wrinkled, coarsely punctured, sides almost smooth. Median impression well developed. Green pattern on pronotum weakly expressed, tripartite, without clear borders. Prosternum almost smooth, with some coarse punctures around coxa. Front part of proepisternum smooth, posterior part gently wrinkled, coarsely punctured, strongly inclined towards the mesepisternum. PL: 1.76 – 1.83 mm, PW: 3.32 – 3.39 mm. Elytra moderately convex, oval, with weakly expressed edge. Elytra with 15 striae, all striae evanescent towards apex. Striae well developed but shallow, extremely slightly punctured on the disc and smooth on the base. At the apical two thirds of elytra 3 rd stria joining 4 th. Intervals smooth. Elytral pattern divided into three irregular bands. In 6 th and 10 th intervals apical band of elytral pattern may converge with median band. Metasternum and metepimeron almost smooth. Sides of sterna with weak wrinkles. Last three sternites bisetose. EL: 4.10 – 4.30 mm, EW: 4.29 – 4.43 mm.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC2FF8BFCE6F5F4FD9775EE.taxon	discussion	Comments. O. (P.) barsevskisi n. sp. clearly differs from other Omophron (Phrator) species. The shape of the new species resembles the species of subgenus Omophron, although peculiar mandibles as well as metafemurs and mesofemurs strongly projecting beyond elytral borders undeniably indicate the association of this species to subgenus Omophron (Phrator). The new species can be easily distinguished from other species of the Omophron (Phrator) ‘ vittulatum’ group by evenly rounded elytra, smooth elytral striae, and a different elytral pattern. The only male sample known up to now has been damaged, therefore aedeagus is not described in this article.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC2FF8BFCE6F5F4FD9775EE.taxon	etymology	Etymology. This species is named after my colleague and supervisor of my Ph. D. thesis Prof. Dr. biol. Arvīds Barševskis (Institute of Systematic Biology, Daugavpils University, Daugavpils, Latvia).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC2FF8BFCE6F5F4FD9775EE.taxon	distribution	Distribution. Up to now known only from three locations in Cameroon and one location in Nigeria. Distribution map (Fig. 6).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC2FF8BFCE6F5F4FD9775EE.taxon	materials_examined	Material examined. Holotype: Cameroon: Edea (Bsse-Sanaga) (1 female) (MRAC). Paratypes: Cameroon: Younde (2 females) (MRAC); Vadon (1 male) (MRAC); Nigeria: Taraba State, Lau, Benue river, 14.04 (1 female).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC6FF8BFF03F1D4FB7470EE.taxon	description	Description. Holotype body length 11.9 mm; width 7.5 mm. Habitus (Fig. 3). Colour: Head and pronotum orange brown. Elytra amber yellow. Legs and antennae yellowish brown. Patches of pronotum and elytral patterns umber, almost black. Mandibles yellowish brown, their apical part umber, almost black. Ventral side yellowish brown; epipleural sides of elytra, pronotum, and prosternum lighter. Head fairly flat, rarely and finely punctured. With one supraorbital setiferous puncture on each side of head near the eye. Subocular ridge well developed, curved upwards and slightly rounded. Clypeus wrinkled, bisetose. Labrum distinctly emarginated, with moderately jagged sides and six setae near the front margin. Neck smooth. First segment of antennae unisetose at apex, 5 – 11 segments covered with thin setae, 1 – 4 segments bare, only with some longer setae near apex. Last segment of maxillary palps longer than penultimate segment, with longitudinal impression. HW of holotype: 3.8 mm. Pronotum moderately convex, base bisinuated, sides almost straight, gently rounded only towards anterior angles. Sides with well developed edge. Anterior angles nearly acute. Apex and base of pronotum coarsely punctured, disc moderately wrinkled without puncture. Basal impression weakly expressed. Median impression well developed. Pattern on pronotum tripartite, brown, almost reduced. Front part of proepisternum smooth, posterior part coarsely punctured, strongly inclined towards the mesepisternum. Prosternum weakly punctured. PL of holotype: 3.30 mm, PW of holotype: 5.70 mm. Elytra moderately convex, longitudinally oval. Sides slightly concave in two thirds of elytra. Elytra with 15 striae reaching from base to apex. Striae well developed, deep, punctures clearly marked. Near apex, striae become shallower. Intervals expressly convex, smooth, narrower at sides. Elytral pattern consists of rambling dark brown spots, which may be divided in three bands. The median band of elytral pattern converges with apical and basal bands in 6 th and 10 th intervals. Metasternum and metepimeron gently punctured. Sterna smooth, last three sterna with two setae. EL of holotype: 7.65 mm, EW of holotype: 7.50 mm.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC6FF8BFF03F1D4FB7470EE.taxon	discussion	Comments. Morphologically, this species resembles O. (P.) vittulatum, although O. (P.) schoutedeni is at least 1.5 times bigger. O. (P.) schoutedeni can be distinguished by its orange brown pronotum, darker elytral pattern, and less expressed puncture on the base of pronotum. Significant differences in the build of aedeagus compared to other species of subgenus Omophron (Phrator) were not observed in this species.	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
03E9D425FFC6FF8BFF03F1D4FB7470EE.taxon	distribution	Distribution. Known only from one type location (Djokopunda, Kasai Province, Democratic Republic of Congo). Distribution map (Fig. 7).	en	Valainis, Uldis (2011): Revision of the Omophron (Phrator) ‘ vittulatum’ species group (Coleoptera: Carabidae). Baltic Journal of Coleopterology 11 (2): 117-126, DOI: 10.5281/zenodo.13204272
