identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E9E50844597C6AFE54FCD7FAF04ECD.text	03E9E50844597C6AFE54FCD7FAF04ECD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valgina Mulsant 1842	<div><p>Subtribe Valgina Mulsant, 1842 (Bouchard et al. 2011)</p><p>Checklist of Chinese Hybovalgus</p><p>Hybovalgus bioculatus Kolbe, 1904</p><p>calvus sp. nov.</p><p>fraternus Moser, 1915</p><p>= Excisivalgus csikii Endrödi, 1952</p><p>fulvosquamosus Moser, 1904</p><p>sexdentatus Arrow, 1944</p><p>jordansi (Endrödi, 1952)</p><p>thibetanus (Nonfried, 1891)</p><p>= versicolor Moser, 1908</p><p>= laticollis Arrow, 1944</p><p>= Valgus pubicollis Pic, 1928</p><p>= Valgus savioi (Pic, 1928)</p><p>= yunnanus Moser, 1906</p><p>tonkinensis Moser, 1904</p><p>= Excisivalgus klapperichi Endrödi, 1952</p><p>Key to males</p><p>1. Pronotal carinae present as rounded elevations or barely noticeable ................................................ 2</p><p>– Pronotal carinae sharp (sometimes only slightly sharp).....................................................................4</p><p>2. Pronotal posterior margin slightly protruding backwards, apex rounded ......................................... 3</p><p>– Pronotal posterior margin simply rounded ............................................... H. bioculatus Kolbe, 1904</p><p>3. Pronotal carinae parallel, barely noticeable or nearly obsolete, short or long (at most ⅓ pronotal length). Meso- and metatibiae each without central tooth on posterior sides ........ H. calvus sp. nov.</p><p>– Pronotal carinae arched outwardly, ending well after middle of disc. Meso- and metatibiae each with a prominent central tooth on posterior side ............................................ H. tonkinensis Moser, 1904</p><p>4. Protibia with 6 external teeth: teeth 1–3 long and sharp, teeth 4–6 short and slightly sharp ................ ................................................................................................................ H. sexdentatus Arrow, 1944</p><p>– Protibia with 5 external teeth: teeth 1–3 long and sharp, teeth 4–5 short and slightly sharp ............ 5</p><p>5. Posterior margin of pronotum simply rounded; pronotal lateral margin not arched toward inside on posterior half ..................................................................................................................................... 6</p><p>– Posterior margin of pronotum at center more or less prominent; pronotal lateral margins noticeably arched on posterior toward inside in the second half ........................................................................ 7</p><p>6. Pronotum suboval (Fig. 7A), lateral small carina obsolete. Anal sternites covered with recumbent, testaceous scales, sparse on disc and dense laterally ................................. H. fraternus Moser, 1915</p><p>– Pronotum subtrapezoidal (Fig. 9A), lateral small carina present, posteriorly arching outwards. Anal sternites completely covered with thick, recumbent, testaceous scales ............................................... ........................................................................................................... H. thibetanus (Nonfried, 1891)</p><p>7. Pronotal posterior margin at center slightly prominent, with apex rounded (Fig. 8K); pronotal lateral small carina obsolete ....................................................................... H. fulvosquamosus Moser, 1904</p><p>– Pronotal posterior margin at center prominent, V-shaped, apex rounded on top only (Fig. 8G); pronotal lateral small carina present, posteriorly arching outwards ....... H. jordansi (Endrödi, 1952)</p><p>Key to females (females of H. fulvosquamosus and H. jordansi are unknown)</p><p>1. Pronotal carinae present as rounded elevations or noticeable and short ........................................... 2</p><p>– Pronotal carinae sharp (sometimes only slightly sharp) .................................................................... 3</p><p>2. Pronotal posterior margin at center excavated in shape of an enlarged “E”. Metatibia without central tooth at center of posterior margin ........................................................... H. bioculatus Kolbe, 1904</p><p>– Pronotal posterior margin at center prominent, V-shaped. Metatibia with a barely noticeable tooth at center of posterior margin ....................................................................................... H. calvus sp. nov.</p><p>3. Protibia with 5 external teeth ............................................................................................................ 4</p><p>– Protibia with 6 external teeth ................................................................. H. sexdentatus Arrow, 1944</p><p>4. Pronotal posterior margin at center deeply excavated as an enlarged “E” (Fig. 11E) .................. .......................................................................................................... H. tonkinensis Moser, 1904</p><p>– Pronotal posterior margin at center slightly excavated as an enlarged “C” (Fig. 9G) ......... 5</p><p>5. Elytra near scutellum deeply depressed and with prominent ridges. Pygidium without cretaceous maculae ............................................................................................. H. thibetanus (Nonfried, 1891)</p><p>– Elytra near scutellum depressed but without ridges. Pygidium on sides covered with cretaceous maculae ...................................................................................................... H. fraternus Moser, 1915</p></div>	https://treatment.plazi.org/id/03E9E50844597C6AFE54FCD7FAF04ECD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E508445E7C64FD06F9D9FD034AE6.text	03E9E508445E7C64FD06F9D9FD034AE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus Kolbe 1904	<div><p>Hybovalgus Kolbe, 1904</p><p>Hybovalgus Kolbe, 1904: 55 .</p><p>Excisivalgus Endrödi, 1952: 62, syn. nov.</p><p>Type species</p><p>Hybovalgus bioculatus Kolbe, 1904 by monotypy.</p><p>Diagnosis</p><p>In the years after 1904 some entomologists tried to better define Hybovalgus (Arrow 1944; Paulian 1961; Krikken 1978; Miyake 1985), but could not find even one morphological synapomorphy. Krikken (1984) and Miyake (1985) wrote that the shape of the parameters was peculiar to Hybovalgus . Miyake (1985) distinguished Hybovalgus from Dasyvalgus as follows: “Parameres more or less prolonged, not confluent each other, with a excision on each inner edge, Hybovalgus . Parameres vestigial, sometimes confluent each other, Dasyvalgus ”. The paramere shape of each genus is different, and easily distinguishable, although the meaning given to the word “vestigial” by Miyake is not clear.</p><p>After studying many specimens (including all the species of Hybovalgus in Taiwan), we conclude that the males of Hybovalgus can be distinguished from those of Dasyvalgus by a series of non-exclusive characters and by the shape of the parameres (Fig. 3 A–F). The special shape of Hybovalgus parameres is distinctive compared with the variable parameres of males of Dasyvalgus species. This paramere shape, that we define as “ Hybovalgus shape”, within the Chinese Valgina is shared only with Dasyvalgoides denticulatus Endrödi, 1952, in a monotypic genus. The relationship between these two genera must still be investigated, but the latter species is readily distinguishable from any Hybovalgus by the protibia with six teeth; and from H. sexdentatus by its flattened body with inconspicuous scales tufts.</p><p>The series of non-exclusive characters that separate Hybovalgus from the other Oriental Valgina genera are:</p><p>- Protibial external teeth 1–3 long and sharp in males, where tooth 2 is sometime a bit shorter than 1 and 3; the same for females but often they are worn by digging (see remarks on page 7 about differences between the two sexes) and in some cases tooth 2 is slightly shorter than 1 or 3 (Fig. 1).</p><p>- Both sexes have 5–6 protibial teeth.</p><p>- Suctorial brush long in males, short in females (Ricchiardi &amp; Li 2015). See remarks below about differences between the two sexes.</p><p>- Anterior margin of clypeus noticeably sinuate in both sexes, a bit less in females of H. thibetanus .</p><p>- Pronotal lateral margins more or less crenate in both sexes.</p><p>- Pronotal lateral margin depression always present in both sexes.</p><p>- Pronotal posterior margin of most females (but not all) more or less excavated.</p><p>- Parameres of Hybovalgus shape (Fig. 3).</p><p>- Protibia with five teeth on the external margin, in one case six ( H. sexdentatus).</p><p>- Pygidium of males always narrowed from the middle to the apex; apex rounded in caudal view, not prominent in lateral view. Pygidium of females often (but not always) modified at apex (see Remarks under species accounts below).</p><p>- Pygidium of males with a ventral small, flat or slightly hollowed surface with posterior margin rounded or pointed at center. In females this area is bigger and deeply hollowed.</p><p>- Pygidium of females without pygidial stylus.</p><p>- Propygidial scale tufts of females darker than those of males.</p><p>Some species have protibial teeth 1–2 and/or 3–4 closer to one another than to the others. This character is not consistent among all the specimens of Hybovalgus as it varies within the same species.</p><p>Another character that we decided not to use for the separation of species is the presence or absence of cretaceous maculae on the propygidium or pygidium of males. In some species those maculae are really difficult to see because they are covered with thick scales or by the built-up body grease.</p><p>Identification of females</p><p>Generally, it is easy to separate females from males because the pronotal posterior margin is more or less deeply excavated ( H. bioculatus, H. fraternus, H. thibetanus, H. tonkinensis) and/or the apex of the pygidium is modified into a pointed projection ( H. fraternus, H. sexdentatus, H. thibetanus, H. tonkinensis), a rounded projection ( H. bioculatus), or the apex is simply rounded ( H. calvus sp. nov.).</p><p>Often, females are larger than males. All the females we studied bear black propygidial scale tufts, even when their male counterparts have testaceous propygidial scale tufts.</p><p>Females of two species, H. fulvosquamosus and H. jordansi, are unknown.</p><p>Variability of paramere shape</p><p>After analysis of the males of a many Valgina belonging to hundreds of species, we concluded that the study of the shape of the parameres is a reliable way to define species. There are, however, a couple of notable exceptions. Jameson &amp; Swoboda (2005), revising the New World Valgina, wrote that: “However, one species of New World valgine, Valgus seticollis Beauvois, 1807, has extremely variable male genitalia although there is an overall similarity in pattern” (Jameson &amp; Swoboda 2005: figs 31–35); “Other characters that we examined are consistent within this species, including form of the tarsomeres, pygidial spine, pronotum, sternites, and spiracles. Lacking other morphological characters to separate the genitalic forms, we conservatively group all the forms as one species. A thorough study of the genitalic forms of V. seticollis would be an excellent topic for a molecular analysis and could reveal interesting evolutionary patterns”.</p><p>Kolbe (1897) described a new Valgina genus, Homovalgus, to include Valgus seticollis . Jameson &amp; Swoboda (2005) clarified the nomenclature of the species and placed it under Valgus . It is our opinion that because the shape of the parameres is similar to that of Hybovalgus / Dasyvalgoides, and the lack of a pygidial stylus in females, this species must be separated from Valgus and that its valid name is Homovalgus seticollis . The relationships of this species to the Oriental Valgina could add appeal to further study.</p><p>In the Oriental Region, another species, Hybovalgus fraternus, is a similar case to that of Homovalgus seticollis . Hybovalgus fraternus also exhibits limited variation in morphology and great variation in paramere shape. This is why we decided to conservatively avoid splitting this species merely on the basis of variation in paramere shape.</p><p>Description of aedeagus</p><p>Figure 3 shows several examples of the aedeagus of Chinese Hybovalgus species along with some nomenclature. The outline of the aedeagus is distinctive. The phallobase is always V-shaped ventroapically (vs) and with a sclerotized upper anterior part (spb) notched, flat or pointed, sometime variable within the same species. The parameres are prominent, with the apex extended and with different shapes, with or without a sub-apical external or internal tooth; basal lobe more or less developed and in some cases variable in shape within a species. In lateral view the parameres shows a more or less deep, basomedial notch (bmn).</p><p>Distribution</p><p>Hybovalgus species are mostly distributed in south-central continental China, with most species extending to northern Laos, northern Vietnam, Hainan Island (5–6 species) and Taiwan. In Taiwan there are six species: Hybovalgus kamiyai (Sawada, 1941); H. major Sawada, 1939; H. matsudai Miyake, 1985; H. obsoletosquamosus Miyake, 1985; H. takasagoensis (Sawada, 1941) and H. thoracicus Moser 1911 . These six Taiwanese species are considered endemic, but their relationships with continental Hybovalgus must still be analyzed.</p><p>Figure 4 shows the approximate distribution of Hybovalgus species. In the west, the genus inhabits Yunnan and Sichuan (there are no records so far for Myanmar). In the east some species reach Shandong, Jangsu and Zhejiang. In the north there are extensions into the southestern part of Palaearctic China (sensu Chen et al. 2008). We found no records of Hybovalgus from Henan and Anhui provinces or Chongqing Municipality, but this may be due to a lack of collecting.</p><p>Summary of Chinese species distribution</p><p>Hybovalgus bioculatus Kolbe, 1904: Laos, Vietnam, China (Hainan, Guangxi).</p><p>Hybovalgus calvus sp. nov.: Vietnam, China (Hainan).</p><p>Hybovalgus fraternus Moser, 1915: Vietnam, Laos, China (Fujian, Zhejiang, Sichuan, Guangxi, Jangxi, Hunan, Guizhou).</p><p>Hybovalgus fulvosquamosus Moser, 1904: Laos, Vietnam, China (Hainan, Hunan, Guangdong).</p><p>Hybovalgus jordansi (Endrödi, 1952): China (Fujian).</p><p>Hybovalgus sexdentatus Arrow, 1944: Vietnam, Laos, China (Guangxi, Yunnan, Sichuan, Hainan, Shandong?).</p><p>Hybovalgus thibetanus (Nonfried,1891): China (Yunnan, Sichuan, Jangsu, Shaanxi, Jiangxi, Hubei, Gansu, Zhejiang).</p><p>Hybovalgus tonkinensis Moser, 1904: Vietnam, China (Fujian, Zhejiang, Jiangxi, Guangxi, Hunan, Guizhou, Hainan, Gansu, Sichuan).</p></div>	https://treatment.plazi.org/id/03E9E508445E7C64FD06F9D9FD034AE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844507C62FDC3FDDCFD4C4EEA.text	03E9E50844507C62FDC3FDDCFD4C4EEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus bioculatus Kolbe 1904	<div><p>Hybovalgus bioculatus Kolbe, 1904</p><p>Fig. 5</p><p>Hybovalgus bioculatus Kolbe, 1904: 56 .</p><p>Hybovalgus bioculatus – Schenkling 1922: 51. — Ma 1995: 196, fig. 168, misidentification, it is Hybovalgus thibetanus .</p><p>Material examined</p><p>VIETNAM: 7 ♂♂ (MNHUB), 1 ♂, 1 ♀ (ERC), Montes Mauson, 2000–3000’, Apr.– May, H. Fruhstorfer leg. (same locality and data of type series but these specimens are not labelled as such); 1 ♂, Vinh Phuc Prov., Tamdao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.61139&amp;materialsCitation.latitude=21.483612" title="Search Plazi for locations around (long 103.61139/lat 21.483612)">Tam Dao N.P.</a>, about 21°29′01″ N, 103°36′41″ E, May 1997 (ERC); 1 ♂, Ht. Tonkin, Lao Kay, Lào Cai, 22°20′14″ N, 104°08′54″ E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.14833&amp;materialsCitation.latitude=22.337223" title="Search Plazi for locations around (long 104.14833/lat 22.337223)">Bourgoin Coll. B.M.</a> 1938-252 (NHM); 2 ♂♂, Chapa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.87278&amp;materialsCitation.latitude=22.323057" title="Search Plazi for locations around (long 103.87278/lat 22.323057)">Sa Pa</a>, 22°19′23″ N, 103°52′22″ E, May–Jul. 1918, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.87278&amp;materialsCitation.latitude=22.323057" title="Search Plazi for locations around (long 103.87278/lat 22.323057)">Bourgoin Coll. B.M.</a> 1938-252 (NHM); 2 ♂♂, Tonkin, Chapa, 1 Jun. 1916, R.Vitalis de Salvaza leg., Bourgoin Coll. B.M. 1938-252 (NHM).</p><p>LAOS: 3 ♂♂, 1 ♀, Huaphan Prov, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.01667&amp;materialsCitation.latitude=20.2" title="Search Plazi for locations around (long 104.01667/lat 20.2)">Ban Saleui</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.01667&amp;materialsCitation.latitude=20.2" title="Search Plazi for locations around (long 104.01667/lat 20.2)">Phou Pan Mt.</a>, about 20°12′ N, 104°01′ E, 1300–1900⁄m, 11 Apr.–15 May 2012, C.C. Holzschuh leg. (ZFMK); 1 ♂, Huaphan Prov., Mt. Phu Phan, 2060 m, Apr.– May 2012, local collectors (ERC).</p><p>CHINA: 3 ♂♂, Hainan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.51667&amp;materialsCitation.latitude=18.774721" title="Search Plazi for locations around (long 109.51667/lat 18.774721)">Wuzhishan</a>, about 18°46′29″ N, 109°30′60″ E, 1 Jun. 2009 (ERC); 2 ♂♂, Guangxi, Mt. Dayaoshan, 24°08′11″ N, 110°11′19″ E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.188614&amp;materialsCitation.latitude=24.136389" title="Search Plazi for locations around (long 110.188614/lat 24.136389)">Linxiang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.188614&amp;materialsCitation.latitude=24.136389" title="Search Plazi for locations around (long 110.188614/lat 24.136389)">Jinxiu</a>, 700 m, 25 Apr.–5 May 2014, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.188614&amp;materialsCitation.latitude=24.136389" title="Search Plazi for locations around (long 110.188614/lat 24.136389)">Huang Yuanyong and Li Dong</a> leg. (ERC); 1 ♂, Guangxi, Mt. Dayaoshan, Pingzhao, Jinxiu, 1200 m, 15–25 May 2014, Zhao Jinsheng leg. (ERC).</p><p>Remarks</p><p>Kolbe (1904: 57) wrote that the locality where Fruhstorfer collected the type series was at 2–3000 m, but this is incorrect, as the original labels bears the mention “ 2-3000’ ” (’ = feet). In fact, the highest point of Montes Mauson is actually 1541 m (5249’). Many, or all of the specimens of H. bioculatus preserved at MNHUB and ERC most probably belong to the H. bioculatus type series, but they were not labelled as such by Kolbe who wrote “Eine Anzahl Exemplare”. Scanning the literature dealing with H. bioculatus, we concluded that the female of this species was never described and probably Kolbe himself failed to pair the two sexes. This is clear because one of the specimens (ERC) is a female (described below), and Kolbe wrote on one small label “keine Forceps” without recognizing it as female.</p><p>Description of a female from Montes Mauson (ERC; type locality of the species)</p><p>MEASUREMENTS. Length 8.0 mm; width 5.1 mm. Body colour mostly castaneous.</p><p>HEAD. Black, slightly shiny, covered with large, round punctures each with a bristle-like, erect, testaceous scale. Clypeus black, with anterior margin castaneous, anterior margin sinuate, suctorial brush shorter than that of male.</p><p>PRONOTUM. Black, nearly glabrous, trapezoidal, slightly crenate at sides, laterally depressed at center of lateral margins; anterior angle acute, posterior angle obtuse, apical margin rounded with a small ridge inclined at approximately 45°; posterior margin with central part deeply excavated as an enlarged “E” (Fig. 5F); carinae rounded, parallel, ending around middle of disk; central small carina not present; lateral carinae obsolete; with 2 testaceous, barely noticeable, central carina scale tufts; with 2 more or less noticeable, small, scales tufts at posterior angle.</p><p>SCUTELLUM. Black, long, apically rounded, nearly glabrous.</p><p>ELYTRA. Castaneous, slightly shiny, mostly glabrous, with shallow, black juxtascutellar depression, the anterior half of which is surrounded by a prominent ridge; humeral humbones prominent, with a small, testaceous scale tuft; anteapical umbones slightly prominent, with a prominent testaceous scale tuft directed posteriorly. Part of disk black, with thick, small, recumbent, barely noticeable scales.</p><p>PROPYGIDIUM. Castaneous, slightly shiny, nearly glabrous, with scattered, small, recumbent black scales; posterior margin curved inward at middle, with 2 prominent, black propygidial scale tufts. Cones at posterior margin short and blunt.</p><p>PYGIDIUM. Black, wider than long, slightly shiny, nearly glabrous, covered with scattered, small, recumbent, black scales; narrowing to apex, where there is a prominent rounded projection, showy in lateral view. Ventrally with a hollowed surface, glabrous and posteriourly semicircular. Apex with a prominent, testaceous scale tuft.</p><p>PROTIBIA. Black, with 5 external teeth: 1–3 long, 4–5 smaller, all rounded at apex. Teeth 4–5 closer to one another than to 1–3.</p><p>MESO- AND METAFEMUR. Castaneous with a small, testaceous scale tuft at centre of posterior margin.</p><p>METATIBIA. Castaneous, centrally enlarged, without a tooth at center of posterior margin. First metatarsomere long, twice length of second.</p><p>ABDOMEN. Castaneous, covered with thick, recumbent, testaceous scales. Anal sternites longer than other visible sternites, with same scales but centrally glabrous.</p><p>Species variability</p><p>Males</p><p>Length 7.7–8.5 mm; width 4.5–5.1 mm. The colour of the body is stable. Along posterior border of pygidium, in the places where Valgina often have scale tufts, H. bioculatus sometimes shows a glabrous surface, while in other specimens there are small, thick, erect scales whose colour varies from testaceous, to dirty testaceous to blackish. Lastly, in Chinese specimens the two posterior margin propygidial scale tufts and humeral humbone scale tufts are dirty testaceous instead of testaceous. Two of the three specimens from Guangxi have the basomedial notches of the parameres closed.</p><p>Females</p><p>The only two known females do not show any significant variation.</p><p>Flight period</p><p>The adults of this species were collected from mid-April to early June at an elevation of 700–1900 m.</p><p>Distribution</p><p>Hybovalgus bioculatus was described from northern Vietnam. New records reported here came from northern Laos and China (Hainan, Guangxi).</p></div>	https://treatment.plazi.org/id/03E9E50844507C62FDC3FDDCFD4C4EEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844567C61FDF7F9D3FD7A4AFC.text	03E9E50844567C61FDF7F9D3FD7A4AFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus calvus Ricchiardi 2017	<div><p>Hybovalgus calvus sp. nov.</p><p>urn:lsid:zoobank.org:act: 2AE1E317-E510-4210-B843-973C1985BF3E</p><p>Fig. 6</p><p>Diagnosis</p><p>Hybovalgus calvus sp. nov. can easily be separated from all other Chinese Hybovalgus (except from H. jordansi) by the V-shaped protrusion at the posterior margin of the pronotum (Fig. 6A, D, J). The female pygidium apex lacks a protrusion. The parameres are different from those of H. bioculatus in the C-shaped apex (Fig. 6H). Females of Hybovalgus calvus sp. nov. can be distinguished from those of H. jordansi by the prominent testaceous scale tufts at the posterior margin of the pronotum; by the fringe of small, testaceous scales at the pygidial apex, instead of prominent, double-pointed testaceous scale tufts. The apex of the parameres of both species (Figs 6H, 8I) are inwardly C-shaped, but in H. jordansi they are thinner and tapered.</p><p>Etymology</p><p>The name comes from the Latin word “ calvus ”, meaning glabrous, as the integument of this species appears at first glance.</p><p>Type material</p><p>Holotype</p><p>VIETNAM: ♂, Tonkin, Montes Mauson, Apr.–May, 2000–3000’, H. Fruhstorfer leg. (MNHUB).</p><p>Paratypes</p><p>VIETNAM: 1 ♂ (ERC), 1 ♀ (MNHUB), same data as for HT.</p><p>CHINA: 1 ♂, Hainan, Wuzhishan, 1 Jun. 2009 (ERC).</p><p>Description</p><p>Holotype</p><p>MEASUREMENTS. Length 8.2 mm; width 5.1 mm.</p><p>BODY. Colour mostly black, with propygidium and pygidium dark castaneous. HEAD. Black, slightly shiny, covered with large round punctures; frons and vertex covered with small, recumbent, testaceous scales that do not form scale tufts. Clypeus black, anteriorly sinuate, suctorial brush long.</p><p>PRONOTUM. Black, nearly glabrous, trapezoidal, slightly crenate at sides, laterally depressed at center of lateral margins; anterior angle acute, posterior angle right-angled, with apex rounded; posterior margin with central part showily produced backwards, V-shaped, with apex rounded. Carinae rounded, parallel, ending around middle of disk; central small carina not present; lateral carinae not present; with 2 testaceous barely noticeable central carina scale tufts; without any other scale tufts. Covered with small, decumbent, scattered, testaceous scales.</p><p>SCUTELLUM. Black, long, apically rounded, covered with small, decumbent, thick, testaceous scales.</p><p>ELYTRA. Black, slightly shiny, mostly glabrous, with barely noticeable juxtascutellar depression. Humbones slightly prominent, without scale tufts; anteo-central part of disk covered with hardly noticeable small, thick, decumbent black scales.</p><p>PROPYGIDIUM. Dark castaneous, covered with thick, decumbent, small, coffee bean shaped, testaceous scales that form two small propygidium scale tufts; centre of posterior margin depressed; cones short and blunt.</p><p>PYGIDIUM. Dark castaneous, wider than long, slightly depressed at sides in ventral half. Covered with thick, decumbent, small, coffee grain shaped, testaceous scales which form a fringe of long scales at apex. Apex rounded, without projection. Under apex, in ventral view, there is a hollowed, semicircular, glabrous, horizontal surface.</p><p>PROTIBIA. Black, with 5 external teeth: 1–3, long, 4–5 smaller, all rounded at apex. Teeth 3–4 closer to one another than 1–3; tooth 5 rather spaced apart from 4 (Fig. 6B).</p><p>MESO- AND METAFEMUR. Black, with a fringe of small, testaceous scales at centre of posterior margin which do not form a small scale tuft.</p><p>METATIBIA. Black, centrally enlarged at posterior margin, without a tooth at centre. First metatarsomere long 1.8 times the length of second.</p><p>ABDOMEN. Dark castaneous, covered with thick, recumbent, testaceous scales, scattered at centre of 1–3 visible sternites. Anal sternites, with same scales but centrally glabrous.</p><p>Paratype ♀ (differences from HT)</p><p>MEASUREMENTS. Length 9.5 mm; width 5.7 mm. Body dark castaneous.</p><p>The female is quite similar to the male, not showing great differences in the shape of the posterior margins of the pronotum or pygidium apex. The first is pointed at the apex, the latter is regularly rounded. The propygidium is much wider than long, shaped as in the male but with a small depression close to each superior corner.</p><p>Type series variability</p><p>Males: length 8.0– 8.4 mm; width 4.7–5.3 mmm. A PT ♂ has the propygidium, pygidium and abdomen black.</p><p>Flight period</p><p>The Vietnamese types were collected from May to early June at an elevation of 700–1500 m.</p><p>Distribution</p><p>Present in northern Vietnam and China (Hainan).</p></div>	https://treatment.plazi.org/id/03E9E50844567C61FDF7F9D3FD7A4AFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844557C7FFDC9FDC1FCEF4E7A.text	03E9E50844557C7FFDC9FDC1FCEF4E7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus fraternus Moser 1915	<div><p>Hybovalgus fraternus Moser, 1915</p><p>Fig. 7</p><p>Hybovalgus fraternus Moser, 1915: 603 .</p><p>Excisivalgus csikii Endrödi, 1952: 63, syn. nov.</p><p>Hybovalgus fraternus – Schenkling 1922: 51. — Krajčík 2011: 92, fig. 48.</p><p>Excisivalgus csikii – Krajčík 2011: 91.</p><p>Type material</p><p>Lectotype (here designated)</p><p>VIETNAM: ♂, Montes Mauson, Lang Son Pr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Mt Mau Son</a>, 21°50′56″ N, 106°58′00″ E, 2–3000 feet, Apr. – May, H. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Fruhstorfer</a> leg. (MNHUB).</p><p>Paralectotype (here designated)</p><p>VIETNAM: 1 ♂, same data as the LT (MNHUB).</p><p>Other type material examined</p><p>Holotype of Excisivalgus csikii Endrödi, 1952</p><p>CHINA: ♀, Fujian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (Guadun), 27°40′ N, 117°40′ E, 2300 m, 20 Apr. 1938, J. Klapperich leg. (ZFMK).</p><p>Other material examined</p><p>VIETNAM: 2 ♂♂ (MNHUB), 1 ♂ (ERC), Montes Mauson, Lang Son Pr., Mt Mau Son, 21°50′56″ N, 106°58′00″ E, 2–3000 feet, Apr. – May, H. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Fruhstorfer</a> leg.</p><p>LAOS: 2 ♂♂ (ZFMK), 1 ♂ (ERC), Hua Phan Prov., Ban Saleui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.01667&amp;materialsCitation.latitude=20.2" title="Search Plazi for locations around (long 104.01667/lat 20.2)">Phou Pan Mt.</a>, 20°12′ N, 104°01′ E, 1300–1900 m, 11 Apr.–15 May 2012, C. Holzschuh leg.; 4 ♂♂ (ZFMK), 1 ♂ (ERC), same data but 1–31 May 2011.</p><p>CHINA: 65 ♂♂, Fujian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 24–28 Apr. 1946, Tschung-Seng leg. (MHNG); 26 ♂♂, same data but 21 Mar. 1946 (MHNG); 28 ♂♂, Fujian, Kuatun, 28 Apr. 1946, Tschung-Seng leg. (ERC); 12 ♂♂, Fujian, Kuatun, 24 Apr. 1946, Tschung-Seng leg. (ERC); 26 ♂♂ (MHNG), 5 ♂♂ (ERC), Fujian, Kuatun, 21 Mar. 1946, Tschung-Seng leg.; 1 ♂ (MNHUB), 8 ♂♂ (HNHM), 1 ♂ (ERC), Fujian, Kuatun, 18 Apr. 1946, Tschung-Sen leg.; 2 ♂♂, Fujian, Kuatun, 26 Apr. 1946, Tschung-Sen leg. (HNHM); 2 ♂♂, Fujian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 2300 m, 25 May 1938, J. Klapperich leg. (HNHM); 1 ♂, same data but 6 Apr. 1938 (HNHM); 1 ♂, same data but 20 May 1938 (HNHM); 1 ♂, same data but 3 Apr. 1938 (HNHM); 1 ♂, same data but 19 Apr. 1938 (HNHM); 1 ♂, same data but 4 Apr. 1938 (ERC); 1 ♂, Fujian, Fukien <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 18 Apr. 1946, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Tschung-Sen</a> leg. (NHM 1952 -443); 1 ♂, Fujian, Fukien <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′ N, 117°40′ E, 26 Apr. 1946, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Tschung-Sen</a> leg. (NHM 1952 -443); 6 ♂♂, Zhejiang, Changlongshan, Anji, 800–900 m, 28 Apr. 2015, Xi Huangshun leg. (ERC); 3 ♂♂, Jangxi, Jinggang Shan – Ciping, 2–14 Jun. 1994, E. Jendek and O. Sausa leg. (ERC); 1 ♂, Hunan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.933334&amp;materialsCitation.latitude=26.066668" title="Search Plazi for locations around (long 113.933334/lat 26.066668)">Guidong</a> env., 26°04′ N, 113°56′ E, 23–31 May 1994, E. Jendek and O. Sausa leg. (ERC); 3 ♂♂, 1 ♀, Guizhou, 60 km North of Kaili, Shibing-Yuntai Shan, 21–26 May 1995, E. Jendek leg. (ERC); 6 ♂♂, Guangxi, Mt. Dayaoshan, Linxiang, Jingxiu, 700 m, 25 Apr.–5 May 2014, Huang Yuanyong and Li Dong leg. (ERC); 1 ♂, Guangxi, Mt. Dayaoshan, Pingban, Jingxiu, 1200 m, 5 May 2014, local collectors (ERC); 1 ♂, Sichuan, Mt. Quingcheng Shan, 26 May 1990, Jima Nara leg. (ERC).</p><p>Remarks</p><p>The analysis of many males from northern Vietnam, northern Laos and China shows that, despite the great variability in paramere shape (particularly the size of the basomedial notch and basal lobe, but many males show intermediate dimensions), there is otherwise little morphological variation within males: the black posterior corner and carina posterior scale tufts are sometimes reduced or barely noticeable. The size of males varies from 5.4 mm (width 3.5 mm) to 6.8 mm (width 4.3 mm). Sometimes the elytra scale pattern is reduced (Fig. 7A), but it is unclear if this is due to wear or real pattern variability.</p><p>One female (ERC) from Guizhou was collected at the same locality and date as three males and its general aspect and all the morphology allows it to be assigned to H. fraternus . This female can be separated from the male by the posterior margin of the pronotum slightly excavated as an enlarged ‘C’, the apex of the pygidium prominently pointed.</p><p>Many males of H. fraternus were collected by J. Klapperich in April 1938 near Kuatun (Guadun, Fujian, 29°37′46″ N, 117°40′01″ E). At least one of these males (HNHM) was collected on the same day (20 April) and at the same locality as the type series of Excisivalgus csikii Endrödi, 1952 . The morphology of the HT of E. klapperichi csikii being identical to the Guizhou H. fraternus female, we concluded that Excisivalgus csikii Endrödi, 1952 is a junior synonym (♀) of Hybovalgus fraternus Moser, 1915 .</p><p>The two females are larger than the males. The female from Fujian (HT of E. csikii) is 7.3 mm long and 4.2 mm wide, while the one from Guizhou (ERC) is 6.9 mm long and 4.5 mm wide.</p><p>Flight period</p><p>Adults of H. fraternus were collected from the first days of April to the beginning of June, at an elevation of 700–2300 m.</p><p>Distribution</p><p>This species was described from northern Vietnam. Records from Laos and China (Fujian, Zhejiang, Sichuan, Guangxi, Hunan, Guizhou, and Jiangxi) are new.</p></div>	https://treatment.plazi.org/id/03E9E50844557C7FFDC9FDC1FCEF4E7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E508444B7C7CFD9FFA41FEA44E79.text	03E9E508444B7C7CFD9FFA41FEA44E79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus fulvosquamosus Moser 1904	<div><p>Hybovalgus fulvosquamosus Moser, 1904</p><p>Fig. 8 K–N</p><p>Hybovalgus fulvosquamosus Moser, 1904: 272 .</p><p>Hybovalgus fulvosquamosus – Schenkling 1922: 51.</p><p>Type material</p><p>Lectotype (here designated)</p><p>VIETNAM: ♂, Montes Mauson, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Lang Son Pr</a>., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Mt Mau Son</a>, 21°50′56″ N, 106°58′00″ E, 2–3000 feet, Apr.–May, H. Fruhstorfer leg. (MNHUB).</p><p>Paralectotypes (here designated)</p><p>VIETNAM: 4 ♂♂, same data as LT (MNHUB).</p><p>Other material examined</p><p>VIETNAM: 1 ♂, Tonkin, Tam Dao, May 1997 (ERC); 1 ♂, same data but May 1993 (ERC). LAOS: 3 ♂♂ (ZFMK), 1 ♂ (ERC), Ban Saleui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.01667&amp;materialsCitation.latitude=20.2" title="Search Plazi for locations around (long 104.01667/lat 20.2)">Phou Pan Mts</a>, 20°12′ N, 104°01′ E, 1300–1900 m, 7 Apr.–25 May 2010, C. Holzschuh leg.; 1 ♂, same data but 11 Apr.–15 May 2012 (ZFMK).</p><p>CHINA: 25 ♂♂, Hainan, Sanya County, Mt. Janfeng, 1–12 Jul. 2004 (ERC); 2 ♂♂, Hunan, Yizhang, Mangshan, 1270–1340 m, 14–17 Jul. 2008, Zhu Xiaoyu and Yang Zhuo leg., IOZ(E) 1945060/63 (IZAS); 2 ♂♂, Hunan, Yizhang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.93222&amp;materialsCitation.latitude=24.956667" title="Search Plazi for locations around (long 112.93222/lat 24.956667)">Mangshan</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.93222&amp;materialsCitation.latitude=24.956667" title="Search Plazi for locations around (long 112.93222/lat 24.956667)">Guizizhai</a>, 24°57′24″ N, 112°55′56″ E, 1270 m, 14–15 Jul. 2008, Zhu Xiaoyu and Yang Zhuo leg., IOZ(E) 1945061/64 (IZAS); 3 ♂♂, Hunan, Yizhang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.88722&amp;materialsCitation.latitude=24.98611" title="Search Plazi for locations around (long 112.88722/lat 24.98611)">Mangshan</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.88722&amp;materialsCitation.latitude=24.98611" title="Search Plazi for locations around (long 112.88722/lat 24.98611)">Forest Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.88722&amp;materialsCitation.latitude=24.98611" title="Search Plazi for locations around (long 112.88722/lat 24.98611)">Mt. Tiantaishan</a>, 24°59′10″ N, 112°53′14″ E, 1572 m, 15 Jul. 2008, Zhu Xiaoyu leg. (IZAS); 1 ♂, border of Hunan, Mangshan and Guangdong Nanling, 17 Jul. 2008, Liang Hongbin leg., IOZ(E) 1945062 (IZAS); 1 ♂, Guangdong, Shixing, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.21833&amp;materialsCitation.latitude=24.737778" title="Search Plazi for locations around (long 114.21833/lat 24.737778)">Chebaling Nature Reserve</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.21833&amp;materialsCitation.latitude=24.737778" title="Search Plazi for locations around (long 114.21833/lat 24.737778)">Xianrendong</a>,(24°44′16″ N, 114°13′6″ E, 508 m, 23 Jul. 2008, Zhu Xiaoyu leg. (IZAS) .</p><p>Remarks</p><p>The female is unknown.</p><p>Species variability</p><p>Species of variable size: length 6.7–8.3 mm; width 3.9–4.5 mm.</p><p>The specimens from Hainan have the pronotum, elytron and legs darker, sometimes nearly black. One specimen from Laos has the scales on the propygidium and pygidium darker. In general all scale tufts are small to barely noticeable.</p><p>Flight period</p><p>Hybovalgus fulvosquamosus adults appear in Vietnam and Laos between April and June. Instead, all Chinese specimens were collected in July. Collected between 500 and 1500 m.</p><p>Distribution</p><p>Described from northern Vietnam. New records are from Laos and China (Guangdong, Hunan and Hainan).</p></div>	https://treatment.plazi.org/id/03E9E508444B7C7CFD9FFA41FEA44E79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844487C7DFE73FABDFBDF48DB.text	03E9E50844487C7DFE73FABDFBDF48DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus jordansi (Endrodi 1952) Ricchiardi 2017	<div><p>Hybovalgus jordansi (Endrödi, 1952), comb. nov.</p><p>Fig. 8 H–J</p><p>Dasyvalgus jordansi Endrödi, 1952: 65 .</p><p>Dasyvalgus jordansi – Krajčík 2011 (gives incorrect year of description: Endrödi, 1953): 89.</p><p>Type material</p><p>Paratypes</p><p>CHINA: 2 ♂♂, Fujian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 8 Apr. 1946, Tschung-Sen leg. (HNHM); 2 ♂♂, Fujian, Kuatun (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 2300 m, 8 Apr. 1938, J. Klapperich leg. (HNHM); 2 ♂♂, Fujian, same data but 1938 (HNHM).</p><p>Other material examined</p><p>CHINA: 3 ♂♂, Fujian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Kuatun</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′N, 117°40′E, 21 Mar. 1946, Tschung-Sen leg. (ERC); 4 ♂♂, same data but 28 Apr. 1946 (ERC); 1 ♂, same data but 24 Apr. 1946 (ERC); 1 ♂, same data but 29 Apr. 1946 (ERC).</p><p>Remarks</p><p>This species was described by Endrödi (1952) as Dasyvalgus jordansi Endrödi, 1952 . Having its parameres clearly of the same shape as in Hybovalgus (Fig. 8 I–J), we include this species in the latter genus (new combination).</p><p>The female is unknown.</p><p>Species variability</p><p>Hybovalgus jordansi shows little morphological variation, but varies in size (length 5.6–6.5 mm; width 3.3–3.9 mm). In some specimens the posterior half of the lateral margin of the pronotum narrows in a straight line from the middle to the posterior corner; in other specimens it narrows in the same way, but noticeably arched inwards. The latter is a morphological character used by Miyake (1985) as a main character in his key to Taiwanese Hybovalgus species, but it seems like an unstable character to separate the continental species of the genus.</p><p>Flight period</p><p>Adults of H. jordansi were collected from the end of March to the end of April.</p><p>Distribution</p><p>This species seems to be endemic to the northern mountains of Fujian (China).</p></div>	https://treatment.plazi.org/id/03E9E50844487C7DFE73FABDFBDF48DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844497C7BFDBAFBDDFA864C33.text	03E9E50844497C7BFDBAFBDDFA864C33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus sexdentatus Arrow 1944	<div><p>Hybovalgus sexdentatus Arrow, 1944</p><p>Fig. 8 A–F</p><p>Hybovalgus sexdentatus Arrow, 1944: 238 .</p><p>Hybovalgus sexdentatus – Ma 1995: 197, fig. 169.</p><p>Type material</p><p>Lectotype</p><p>LAOS: ♂, Ban Naban, 15 Mar. 1915, R. Vitalis de Salvaza leg. (NHM).</p><p>Paralectotypes</p><p>LAOS: 1 ♂, Ban Naban, 18°54′23″ N, 102°54′12″ E, 15 Mar. 1915, R. Vitalis de Salvaza leg. (NHM); 1 ♀, Xieng Khouang (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.5&amp;materialsCitation.latitude=19.5" title="Search Plazi for locations around (long 103.5/lat 19.5)">Khoueng Xieng Khouang</a>), 19°30′00″ N, 103°30′00″ E, 20 May 1915, R. Vitalis de Salvaza leg. (NHM).</p><p>Other material examined</p><p>LAOS: 41 ♂♂ (ZFMK), 2 ♂♂ (ERC), Ban Saleui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.01667&amp;materialsCitation.latitude=20.2" title="Search Plazi for locations around (long 104.01667/lat 20.2)">Phou Pan Mts</a>, 20°12′ N, 104°01′ E, 1300–1900 m, 11 Apr.–15 May 2012, C. Holzschuh leg.; 45 ♂♂ (ZFMK), 10 ♂♂, 1 ♀ (ERC), same data but 1–31 May 2001; 1 ♂, Ban Saleui, Phu Pan Mts, Apr.–May 2012, local collectors (ERC).</p><p>VIETNAM: 1 ♂ (NMER), 1 ♂ (ERC), Cao Bang Prov., vic. Vin Den, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.88139&amp;materialsCitation.latitude=22.564722" title="Search Plazi for locations around (long 105.88139/lat 22.564722)">Nui Piac Oac Nt. Res.</a> 22°33′53″ N, 105°52′53″ E, 900–1300 m, 6–10 May 2012, A. Weigel leg.</p><p>CHINA: 1 ♂, Shandong, Kiautschau ( Jaozhou Bay) (ERC); 29 ♂♂, Guangxi, Mt. Dayaoshan, Linxiang, Jinxiu, 700 m, 25 Apr.–5 May 2014, HuangYuanyong &amp; Lidong leg. (ERC); 1 ♂, Guangxi, same data but May 2015, 1200 m, local collectors (ERC); 1 ♂, Guangxi, Jinxiu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.12&amp;materialsCitation.latitude=24.157778" title="Search Plazi for locations around (long 110.12/lat 24.157778)">Huawang Shanzhuang</a>, 24°9′28″ N, 110°7′12″ E, 600 m, 20 May 1999, Yang Xingke leg., IOZ(E) 902246 (IZAS); 1 ♂, Guangxi, Daxin Xialei Town, 22°54′49″ N, 106°45′50″ E, 680 m, 31 Mar. 1998, Li Wenzhu leg., IOZ(E) 902246 (IZAS); 1 ♂, Hainan, Bawangling, 19°7′43″ N, 109°6′14″ E, 8 Apr. 1984, Gu Maobing leg., IOZ(E) 902438 (IZAS); 1 ♂, Sichuan, Dukou, Pingdi, 7 Jun. 1981, Zhang Baolin leg., IOZ(E) 902243 (IZAS); 3 ♂♂, Yunnan, Menglong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.57&amp;materialsCitation.latitude=22.058887" title="Search Plazi for locations around (long 100.57/lat 22.058887)">Banna</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.57&amp;materialsCitation.latitude=22.058887" title="Search Plazi for locations around (long 100.57/lat 22.058887)">Mengsong</a>, 22°3′32″ N, 100°34′12″ E, 160 m, 24–25 May 1958, Wang Shuyong, Zheng Leyi and Pu Fuj leg., IOZ(E) 902255-57 (IZAS).</p><p>Remarks</p><p>According to Arrow (1944), the type series comprised three specimens, all from Laos. In his work Arrow seemed not to realize that one of the types was a female, as he described only the male. Paulian (1961), applying the International Code of Zoological Nomenclature, amended the name of this species from H sex-dentatus (Arrow, 1944) to H. sexdentatus, but Bacchus (1974) erroneously misspelled the name of the species again and designated the type series of three specimens, LT, one PLT male and one PLT female, but did not describe the latter, which remained undescribed until now. In this work we restate that the hyphen must be eliminated as the correct name is H. sexdentatus, and give the differences between sexes.</p><p>Identification of female (from the ♀ at ERC)</p><p>The size of females is more or less the same as that of the biggest males and the shape of the posterior margin of the pronotum is simply rounded, without protrusions or excavations. The protibia external margin has six teeth, as in the male.</p><p>At first glance the female can be distinguished by the pygidium shape: its apex is protruding and pointed (Fig. 8D).</p><p>Species variability</p><p>The length of males varies from 4.7 to 7.5 mm; the width from 3.0 to 4.1 mm. The only female known so far is 7.4 mm long and 3.5 mm wide.</p><p>The study of 148 specimens leads to the conclusion that the species is morphologically slightly variable. In males the only variations observed are:</p><p>- Propygidial scales and scale tufts testaceous or whitish.</p><p>- The scales on the elytron are sometimes black and testaceous, sometimes black and whitish.</p><p>- The legs are black or castaneous.</p><p>- On occasion, protibia teeth 3–4 and 5–6 are closer to one another than to 1–3.</p><p>Flight period</p><p>Specimens were collected between mid-March and the first week of June at an elevation comprised between 160 and 1900 m.</p><p>Distribution</p><p>Hybovalgus sexdentatus was described from Laos and recorded by Ma (1995) from Yunnan. New records reported here are from northern Vietnam and China (Guangxi, Hainan, Sichuan). One specimen is labelled as from Shandong (Jaozhou Bay), but this record is doutbful without further finds.</p></div>	https://treatment.plazi.org/id/03E9E50844497C7BFDBAFBDDFA864C33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E508444F7C74FDA8F88AFE36498F.text	03E9E508444F7C74FDA8F88AFE36498F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus thibetanus (Nonfried 1891)	<div><p>Hybovalgus thibetanus (Nonfried, 1891)</p><p>Fig. 9</p><p>New synonyms: Hybovalgus laticollis Arrow, 1944; Hybobalgus yunnanus Moser, 1906; Valgus savioi Pic, 1928; Dasyvalgus pubicollis Pic, 1928, Hybovalgus versicolor Moser, 1908 .</p><p>Valgus thibetanus Nonfried, 1891: 372 .</p><p>Hybovalgus yunnanus Moser, 1906: 403 . syn. nov.</p><p>Hybovalgus versicolor Moser, 1908: 259 . syn. nov.</p><p>Dasyvalgus pubicollis Pic, 1928b: 4 . syn. nov.</p><p>Valgus savioi Pic, 1928a: 7 . syn. nov.</p><p>Hybovalgus laticollis Arrow, 1944: 238 . syn. nov.</p><p>Dasyvalgus thibetanus Krajčík, 2011: 92, fig. 46.</p><p>Valgus thibetanus – Schenkling 1922: 43.</p><p>Hybovalgus yunnanus – Schenkling 1922: 51. — Krajčík 2011: 93.</p><p>Hybovalgus versicolor – Schenkling 1922: 51. — Krajčík 2011: 93.</p><p>Dasyvalgus pubicollis – Krajčík 2011: 90.</p><p>Valgus savioi – Krajčík 2011: 96.</p><p>Hybovalgus laticollis – Krajčík 2011: 92.</p><p>non Hybovalgus bioculatus – Ma 1995: 196, fig. 168. Misidentification.</p><p>Type material</p><p>Note: all types here designated have a red type label. Alla data reported are copied from specimen labels.</p><p>Lectotype (here designated)</p><p>CHINA: ♂, Yunnan, (Thibet) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.89833&amp;materialsCitation.latitude=20.00639" title="Search Plazi for locations around (long 98.89833/lat 20.00639)">Tsekou</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.89833&amp;materialsCitation.latitude=20.00639" title="Search Plazi for locations around (long 98.89833/lat 20.00639)">Cigu</a>), 20°00′23″ N, 98°53′54″ E, R. P. Dubernard leg. (MNHUB).</p><p>Paralectotype (here designated)</p><p>CHINA: 1 ♂, same data as LT (MNHUB).</p><p>Other type material examined</p><p>Lectotype of Hybovalgus laticollis Arrow, 1944</p><p>CHINA: ♂, Shandong, Wei-Hai-Wei, 37°30′53″ N, 122°07′10″ E, on oak, 25 Apr. 1899, T.B. Fletcher leg. (NHM).</p><p>Syntype of Hybovalgus yunnanus Moser, 1906</p><p>CHINA: ♂, Yunnan (SDEI).</p><p>Type of Valgus savioi Pic, 1928</p><p>CHINA: ♂, China, 23 Apr. 1918 (MNHN).</p><p>Lectoype of Dasyvalgus pubicollis Pic, 1928</p><p>CHINA: ♂, Thibet (MNHN).</p><p>Lectotype of Hybovalgus versicolor Moser, 1908 (here designated)</p><p>CHINA: ♀, Yunnan (MNHUB).</p><p>Other material examined</p><p>CHINA: 12 ♂♂, Yunnan, 9 Sep. 1919, Simeon Ten leg. (ZMUC); 2 ♂♂, Yunnan, Thibet <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.89833&amp;materialsCitation.latitude=20.00639" title="Search Plazi for locations around (long 98.89833/lat 20.00639)">Tse-Kou</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.89833&amp;materialsCitation.latitude=20.00639" title="Search Plazi for locations around (long 98.89833/lat 20.00639)">Cigu</a>), 20°00′23″ N, 98°53′54″ E, R. P. Dubernard leg. (ERC); 7 ♂♂, Yunnan, 100 km W of Kummig, Diaolin Nat. Res. (on Castanea flowers), 13–14 Jun. 1995, E. Jendek leg. (ERC); 8 ♂♂, Yunnan, 100 km W of Kummig, Diaolin Nat. Res., 22 May–2 Jun. 1993, E. Jendek and O. Sausa leg. (ERC); 3 ♂♂, Yunnan (ERC); 1 ♂, Yunnan, Tse Kou, 28°02′N, 98°54′E, 2000 m, 7–8 Jun. 2006, Jifi Klir leg. (ERC); 1 ♂, Shaanxi, Foping env., 120 km SW of Xian, 7 Jun. 1992, M. Bok leg. (SNMS); 1 ♀, Shaanxi, Chin-Lins Mts ( Quing Ling Mts), Apr.–May 1904 (USNM); 1 ♀, Shaanxi, Chin-Lins Mts ( Quing Ling Mts) (ERC); 1 ♂ (ERC), 2 ♂♂ (MHNG), Sichuan, Wassuland, BZK Sankiangkou, Yengshaping, 1600 m, 15 May 1935, H. Becker leg.; 1 ♂, Sichuan, Shanglian, 24–26 May 2005, E. Kucera leg. (ERC); 2 ♂♂, Sichuan, same data but 24–29 May 2002 (ERC); 1 ♀, Sichuan, Luding Co, Moxi env., 22 May–10 Jun. 1993, Beneš leg. (ERC); 1 ♂ (MNHUB), 1 ♂ (ERC), Jangxi, Kiukiang ( Jujiang), Jul. 1887, A.E. Pratt leg.; 1 ♂, Hubei, Ichang ( Ychang), 6–8000 ft, May (MNHUB); 1 ♂, Gansu, South Gansu, Venxian env., 18–25 Jun. 1995, Beneš leg. (ERC); 1 ♂, Gansu, Kansou mer., Hoei Sein (ERC); 7 ♂♂, 5♀♀ (AMNH), 1 ♂, 2 ♀♀ (ERC), Yunnan, “ Thibet ”; 1 ♂, Jiangsu, Nanking, 29 Oct 1923 (USNM); 1 ♂, Shaanxi, Wanrong, 35°25′18″ N, 110°50′40″ E, 27 Mar. 1995, Li Qingsen leg. (IZAS); 2 ♂♂, Jiangsu, Chemo, 11–12 May 1935, O. Piel coll., IOZ(E) 902241-42 (IZAS); 4 ♂♂, 1 ♀, Jiangsu, Chemo, 23–26 Apr. 1918, IOZ(E) 902238-40, IOZ(E) 902473 (IZAS); 1 ♂, Zhejiang, Mokanshan, IOZ(E) 902251 (IZAS); 1 ♂, Zhejiang, Suichang suburb, 24 Apr. 1988, Dong Sansong leg., IOZ(E) 902440 (IZAS); 1 ♂, Zhejiang, Chusan, 23 Apr. 1935, IOZ(E) 902253 (IZAS); 1 ♀, Hubei, Xingshan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.497215&amp;materialsCitation.latitude=31.32889" title="Search Plazi for locations around (long 110.497215/lat 31.32889)">Longmen River</a>, 31°19′44″ N, 110°29′50″ E, 1300 m, 10 May 1994, Li Wenzhu leg., 31572 Inst. Zool. Acad. Sin. (IZAS); 4 ♂♂, Sichuan, Luding, Detuo to Moxi, 1300 m, 20 May 2009, Yang Ganyan leg. (IZAS); 1 ♂, Sichuan, Dujiangyan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.71527&amp;materialsCitation.latitude=31.053888" title="Search Plazi for locations around (long 103.71527/lat 31.053888)">Huaxi</a>, 31°3′14″ N, 103°42′55″ E, 721 m, 12 May 2009, Yang Ganyan leg. (IZAS); 1 ♂, 4 ♀♀, Sichuan, Luding, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.132225&amp;materialsCitation.latitude=29.648611" title="Search Plazi for locations around (long 102.132225/lat 29.648611)">Moxi</a>, 29°38′55″ N, 102°7′56″ E, 1500 m, 17–18 Jun. 1983, Wang Shuyong and Chen Yuanqing leg., IOZ(E) 902156-57/235 (IZAS); 1 ♀, Sichuan, Wanxian, Wang’erbao, 30°40′7″ N, 108°45′30″ E, 1200 m, 28 May 1994, Zhang Youwei leg., 36965 Inst. Zool. Acad. Sin., IOZ(E) 902466 (IZAS); 1 ♂, Yunnan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.98167&amp;materialsCitation.latitude=22.791111" title="Search Plazi for locations around (long 100.98167/lat 22.791111)">Simao</a>, 22°47′28″ N, 100°58′54″ E, IOZ(E) 902236 (IZAS); 1 ♂ ( H. bioculatus sensu Ma Wenzhen, 1992) Yunnan, Gongshan County, Bingzhongluo, 28°0′55″ N, 98°37′58″ E, 1790 m, Sino-Ameica Exped., 6 Jul. 2000, Liang H.B. N28.01 E98.61 (IZAS); 1 ♀, Yunnan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.6725&amp;materialsCitation.latitude=27.746944" title="Search Plazi for locations around (long 98.6725/lat 27.746944)">Gongshan</a>, 27°44′49″ N, 98°40′21″ E, 1560 m, 15 Jun. 1983, IOZ(E) 902249 (IZAS); 1 ♂, Yunnan, Diqing, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.094444&amp;materialsCitation.latitude=27.367222" title="Search Plazi for locations around (long 99.094444/lat 27.367222)">Weixi</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.094444&amp;materialsCitation.latitude=27.367222" title="Search Plazi for locations around (long 99.094444/lat 27.367222)">Baijixun</a>, 27°22′2″ N, 99°5′40″ E, 1900 m, 3 Jul. 1979, Yan Xiangqun leg. (IZAS); 1 ♂, Yunnan, Lushui, 25°49′45″ N, 98°51′52″ E, 2150 m, 11 Jun. 1981, Wang Shuyong leg., IOZ(E) 902244 (IZAS); 1 ♂, Yunnan, Dali, May 1981, IOZ(E) 902469 (IZAS); 1 ♂, Yunnan, Heqing, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.34417&amp;materialsCitation.latitude=26.42472" title="Search Plazi for locations around (long 100.34417/lat 26.42472)">Liuhe</a>, 26°25′29″ N, 100°20′39″ E, 2000 m, 1 Jun. 1980, IOZ(E) 902252 (IZAS); 1 ♂, Yunnan, Kaiyuan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.42555&amp;materialsCitation.latitude=23.743889" title="Search Plazi for locations around (long 103.42555/lat 23.743889)">Mazheshao</a>, 23°44′38″ N, 103°25′32″ E, 1800 m, 26 Apr. 1970, Wang Zhengjun etc. leg., IOZ(E) 902250 (IZAS); 1 ♂, Yunnan, Fugong, 1196 m, 18 May 1983, IOZ(E) 902468 (IZAS).</p><p>Remarks</p><p>In a long period of time when valgine specimens rarely reached European museums from China, specimens of H. thibetanus were described several times under different names. Study of the types of several Chinese species, i.e., Hybovalgus yunnanus Moser, 1906; Valgus pubicollis Pic, 1928; Dasyvalgus savioi Pic, 1928 and Hybovalgus laticollis Arrow, 1944, showed us that these species are morphologically close to one another and their aedeagi are nearly identical. Even Pic described this species twice under two different names in the same year (Pic 1928a, 1928b). Finally, when Arrow (1944) described H. laticollis, due to World War II, types deposited in other European museums were inaccessible for study and he had to work on the basis of descriptions only, and it is known that Pic’s descriptions were too concise.</p><p>Until now the female of H. thibetanus was unknown. Krajčík (2011), listing H. versicolor Moser, 1908 within the Chinese Valginae (sensu Krikken 1984), doubtfully proposed that this species is the female of, and thus a junior synonym of, H. thibetanus . The study of the LT (Moser 1908 did not specify how many specimens he had for study) of H. versicolor and morphological comparison with H. thibetanus confirm that the two specimens are male and female of the same species, whose correct name is H. thibetanus (new junior synonym). The synonymy is reinforced by the study of eleven other females collected in the same locality and at the same day as the males (see list of material examined).</p><p>Species variability</p><p>Males</p><p>In most specimens the legs are black, but in a few they are castaneous. The same is true for the propygidium and pygidium. The scales on the propygidium and pygidium vary in colour, from light testaceous to dark testaceous (see Fig. 9 A–D). Sometimes the last visible sternites and the anal sternite are glabrous instead of covered with thick scales. The size of males is variable: length from 5.7 to 7.6 mm; width from 3.5 to 4.2 mm.</p><p>Females</p><p>The colour of the integuments and scales varies in the same way as in males. The same with size: length from to 5.8 to 8.3 mm; width from 3.3 to 4.0 mm.</p><p>Flight period</p><p>Specimes were collected between the end of March and early June. Some specimens are labelled as having been collected in September and October (see list of studied material), but these records must be confirmed by other finds. Specimens have been collected from 700 to 2000 m.</p><p>Distribution</p><p>Hybovalgus thibetanus is endemic to continental China and inhabits most of the central provinces: Yunnan, Sichuan, Jiangsu, Shaanxi, Jiangxi, Hubei, Gansu and Zhejiang. The absence from the other central provinces of China (Guizhou, Hunan, Anhui and Henan) is probably due to a lack of collecting. Hybovalgus thibetanus seems to be absent from the southern Chinese Provinces; we have no records from Guangxi, Guangdong, Hainan or Fujian.</p><p>The name of this species, “ thibetanus ”, is from ancient collecting when the northernmost parts of Yunnan were considered to be part of Tibet. To date, there are no specimens of this species known to us collected in Myanmar, Arunachal Pradesh or the southern Xizang valleys, but the presence of H. thibetanus there is possible (see Fig. 10, which shows the distribution of H. thibetanus in Yunnan’s valleys close to Myanmar's borders).</p></div>	https://treatment.plazi.org/id/03E9E508444F7C74FDA8F88AFE36498F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
03E9E50844407C73FDBCFD36FEC14900.text	03E9E50844407C73FDBCFD36FEC14900.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hybovalgus tonkinensis Moser 1904	<div><p>Hybovalgus tonkinensis Moser, 1904</p><p>Fig. 11</p><p>Hybovalgus tonkinensis Moser, 1904: 272 .</p><p>Excisivalgus klapperichi Endrödi, 1952: 63 syn. nov.</p><p>Hybovalgus tonkinensis – Schenkling 1922: 51. — Krajčík 2011: 93, fig. 47.</p><p>Excisivalgus klapperichi – Krajčík 2011: 91, fig. 54.</p><p>non Hybovalgus thoracius Ma 1995: 196 . Misidentification.</p><p>Type material</p><p>Lectotype (here designated)</p><p>VIETNAM: ♂, Tonkin, Montes Mauson (Lang Son Pr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.96667&amp;materialsCitation.latitude=21.84889" title="Search Plazi for locations around (long 106.96667/lat 21.84889)">Mt Mau Son</a>) 21°50′56″ N, 106°58′00″ E, Apr– May, 2000 – 3000’, H. Fruhstorfer leg. (MNHUB).</p><p>Paralectotype (here designated)</p><p>VIETNAM: 1 ♂, same data as LT (MNHUB).</p><p>Other type material examined</p><p>Excisivalgus klapperichi Endrödi, 1952</p><p>Holotype</p><p>CHINA: ♀, Fujian, Kuatun (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′ N, 117°40′ E, 2300 m, 21 Apr. 1938, J. Klapperich leg. (ZFMK).</p><p>Paratypes</p><p>CHINA: 1 ♀, same data as HT but 24 Mar. 1938 (HNHM); 1 ♀, same data but 8 Apr. 1938 (ZFMK); 4 ♀♀, same data but 28 Mar. 1938 (ZFMK); 1 ♀, same data but 28 Apr. 1938 (ZFMK); 1 ♀, same data but 24 Apr. 1938 (ZFMK); 1 ♀, Fujian, Kuatun, (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Guadun</a>), 27°40′ N, 117°40′ E, 28 Apr. 1946, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.666664&amp;materialsCitation.latitude=27.666666" title="Search Plazi for locations around (long 117.666664/lat 27.666666)">Tschung Sen</a> leg. (SNMS).</p><p>Other material examined</p><p>CHINA: 1 ♂ (HNHM), 1 ♂ (ERC), Fujian, Kuatun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.64166&amp;materialsCitation.latitude=27.733612" title="Search Plazi for locations around (long 117.64166/lat 27.733612)">Guadun</a>, 27°44′01″ N, 117°38′30″ E, 2300 m, 3 Apr. 1938, L. Klapperich leg.; 4 ♂♂, 2 ♀♀, Fujian, Kuatun, 28 Apr. 1946, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.64166&amp;materialsCitation.latitude=27.733612" title="Search Plazi for locations around (long 117.64166/lat 27.733612)">Tchung Sen</a> leg. (ERC); 1 ♂, Fujian, Kuatun, 21 Mar. 1946, Tchung Sen leg. (ERC); 2 ♂♂, Fujian, Kuatun, 24 Apr. 1946, Tchung Sen leg. (ERC); 1 ♂, Fujian, Kuatun, 22 Apr. 1946, Tchung Sen leg. (MNHUB); 1 ♂, Fujian, same data but 18 Apr. 1946 (HNHM); 1 ♂, Fujian, Kuatun, 27.40 N, 117.40 O, 2300 m, 20 Apr. 1938, L. Klapperich leg. (HNHM); 2 ♂♂, Fujian, same data but 8 Apr. 1938 (HNHM); 1 ♂, Fujian, same data but 5Apr. 1938 (HNHM); 1 ♂, Fujian, same data but 27 Apr. 1938 (HNHM); 4 ♂♂, Zhejiang, Mt. Changlongshan, Anji, 800–900 m, 28 Apr. 2015, Xi Huangshun leg. (ERC); 2 ♂♂, Hunan, Hunan SE, Guidong env., 26–31 May 1994, E. Jendek and O. Sausa leg. (ERC); 6 ♂♂, Guizhou, 20 km NW of Jangkow, Fanjing Shan, 27 May–3 Jun. 1995, E. Jendek leg. (ERC); 1 ♂, Hainan, Wuzhishan, 1 Jun. 2009 (ERC); 4 ♂♂, China, Jangxi, Jinggang Shan, Ciping, 2–14 Jun. 1994, E. Jendek and O. Sousa leg. (ERC); 1 ♂, Guangxi, Mt. Dayaoshan, Jinxin, Pingzhao, 1200 m, May 2015 (ERC); 1 ♂, Guangxi, same data but 15–25 May 2014, Zanjingshen leg. (ERC); 12 ♂♂, Tibet Province (AMNH); 1 ♂, Zhejiang, Mt. Tianmushan, Laodian 30°20′448″ N, 119°26′44″ E, 9 May 1998, Yu Zhiyong leg. (IZAS); 1 ♂, Zhejiang, Tiemushan, 13 May 1937, O. Piel Coll., IOZ(E) 902263 (IZAS); 1 ♂, Zhejiang, Tiemushan, 10 May 1937, O. Piel Coll., IOZ(E) 902263 (IZAS); 1 ♂, Zhejiang, E. Tiemushan, 30°19′2″ N, 119°31′60″ E, 12 Jun. 1936, IOZ(E) 902269 (IZAS); 1 ♂, 1 ♀, Fujian, Mt. Wuyishan, 22 Apr. 1982, Zhang Baolin leg., IOZ(E) 1945182 (IZAS); 1 ♀, Fujian, Chong’an, Xingcun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.66873&amp;materialsCitation.latitude=27.728067" title="Search Plazi for locations around (long 117.66873/lat 27.728067)">Shilichang</a>, 27°43′41.0376″ N, 117°40′7.446″ E, 840 m, 25 May 1960, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.66873&amp;materialsCitation.latitude=27.728067" title="Search Plazi for locations around (long 117.66873/lat 27.728067)">Jiang Shengqiao</a> leg., IOZ(E) 902233 (IZAS); 1 ♀, Fujian, Chongan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.68778&amp;materialsCitation.latitude=27.75361" title="Search Plazi for locations around (long 117.68778/lat 27.75361)">Xingcun</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.68778&amp;materialsCitation.latitude=27.75361" title="Search Plazi for locations around (long 117.68778/lat 27.75361)">Sangang</a>, 27°45′13″ N, 117°41′16″ E, 720 m, 14 May 1960, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.68778&amp;materialsCitation.latitude=27.75361" title="Search Plazi for locations around (long 117.68778/lat 27.75361)">Pu Fuji</a> leg., IOZ(E) 902227 (IZAS); 1 ♂, Fujian, Jiangyang, road from Huangkeng to Guilin, 290–320 m, 12 Apr. 1960, Jiang Shengqiao leg., IOZ(E) 902259 (IZAS); 1 ♀, Fujian, Chongan, Tongmuguan, Guanping, 27°48′15″ N, 117°43″ E, 800– 900 m, 6 Jun. 1960, Jiang Shengqiao leg., IOZ(E) 902228 (IZAS); 1 ♀, Fujian, Chongan, Tongmuguan, Guanping 27°48′15″ N, 117°43″ E, 800–900 m, 6 Jun. 1960, Jiang Shengqiao leg., IOZ(E) 902229 (IZAS); 1 ♀, Fujian, Jianyang, road from Huangkeng to Guilin, 270 m, 11 Apr. 1960, Ma Chenglin leg., IOZ(E) 902231 (IZAS); 1 ♂, Fujian, Jianyang, Huangkeng to Guilin, 290–320 m, 14 Apr. 1960, Jiang Shengqiao leg., IOZ(E) 902260 (IZAS); 1 ♂, Fujian, Jianyang, road from Huangkeng to Guilin, 290–320 m, 12 Apr. 1960, Jiang Shengqiao leg., IOZ(E) 902258 (IZAS); 1 ♀, Fujian, Jianyang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Huangkeng</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Aotou</a>, 27°40′21″ N, 117°39′20″ E, 800–1050 m, 26 Apr. 1960, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Zhang Yiran</a> leg., IOZ(E) 902232 (IZAS); 1 ♀, Fujian, Jianyang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Huangkeng</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Aotou</a>, 27°40′21″ N, 117°39′20″ E, 950 m, 23 Apr. 1960, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.655556&amp;materialsCitation.latitude=27.672499" title="Search Plazi for locations around (long 117.655556/lat 27.672499)">Zuoyong</a> leg., IOZ(E) 902264 (IZAS); 1 ♂, Fujian, Chongan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.67528&amp;materialsCitation.latitude=27.734446" title="Search Plazi for locations around (long 117.67528/lat 27.734446)">Xingcun</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.67528&amp;materialsCitation.latitude=27.734446" title="Search Plazi for locations around (long 117.67528/lat 27.734446)">Qiliqiao</a>, 27°44′4″ N, 117°40′31″ E, 840 m, 1 Jun. 1960, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.67528&amp;materialsCitation.latitude=27.734446" title="Search Plazi for locations around (long 117.67528/lat 27.734446)">Zuo Yong</a> leg., IOZ(E) 902261 (IZAS); 1 ♀, Guizhou, Suiyang, Kuankuoshui Nature Reserve, 27°57′51″ N, 107°11′32″ E, 9 Jun. 2010, Wang Zhiliang leg. (IZAS).</p><p>VIETNAM: 1 ♂, Tamdao, 17–21 May 1990, P. Pacholatko leg. (ERC); 1 ♂, Tamdao, 12–24 May 1990, P. Pacholatko leg. (ERC).</p><p>Remarks</p><p>Endrödi (1952) described Excisivalgus klapperichi Endrödi, 1952 on the basis of 57 females from Kuatun (Guadun, Fujian) as a new species belonging to the new genus Excisivalgus Endrödi, 1952 . He did not try to pair this species (and a second one, Excisivalgus csikii Endrödi, 1952; see section Remarks above under H. fraternus) to the many males of Hybovalgus collected from the same locality and named the new genus after the wide excision of the posterior margin of the pronotum. Actually, we found that E. klapperichi is the female of H. tonkinensis, as evidenced by the capture of both sexes in the same place and time. An additional proof is the presence in the IZAS collection of a pair in copula (see the material examined list and Fig. 2).</p><p>Identification of female</p><p>The female (formerly Excisivalgus klapperichi Endrödi, 1952) can be easily separated from males by the wide and deep excision of the pronotal posterior margin; by the modified apex of the pygidium and by the black propygidial scale tufts. Furthermore, the ventral side of females is glabrous (Fig. 11H).</p><p>Species variability</p><p>Male</p><p>Males are variable, even in size (length 6.9–9.2 mm, width 3.8–5.2 mm). The pygidium of males has a double pointed testaceous scale tuft at the apex, rarely reduced to a fringe of scales. The small scales that cover the propygidium and pygidium vary from testaceous to whitish. The abdomen varies from nearly glabrous to covered with whitish scales at the sides. The posterior half of the pronotal lateral margin varies from slightly indented to slightly arched outwardly. The interteguments of some specimens from Zhejiang are castaneous instead of black, with the pronotum darkened at the anterior and posterior margins; one specimen from the same Chinese province shows black pronotal posterior margin scale tufts (as, in general, is the case in females) instead of testaceous. The parameres do not vary much in shape.</p><p>Female</p><p>Females show variable scale patterns on the elytra (Fig. 11 E–I). Furthermore, the colour of the small scales covering the propygidium and pygidium varies from dirty testaceous (sometimes somewhat orange) to black. Less variable in size than males, generally constantly greater, but not attaining the size of the bigger males (female length 8.2–8.6 mm, width 4.9–5.2 mm).</p><p>Flight period</p><p>Adults were captured from early April to mid-June, at elevations from 300 to 1600 m.</p><p>Distribution</p><p>Described from northern Vietnam. New records are from China (Fujian, Zhejiang, Jangxi, Guangxi, Hunan, Guizhou, Hainan, Gansu and Sichuan). New finds are necessary to better define the distribution of H. tonkinensis in Sichuan and Gansu. The absence of reports from Yunnan is probably due to a lack of research.</p></div>	https://treatment.plazi.org/id/03E9E50844407C73FDBCFD36FEC14900	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ricchiardi, Enrico	Ricchiardi, Enrico (2017): Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae). European Journal of Taxonomy 340 (340): 1-32, DOI: 10.5852/ejt.2017.340
