taxonID	type	description	language	source
03E902352767FF858A49E80DFC0BF9C7.taxon	diagnosis	Diagnosis Adults with asymmetrical valves with LV overlapping RV. LV antero-ventrally with a beak-like expansion of the valve margin; anterior calcified inner lamella with two incomplete inner lists, forming a sulcus and extending in the lip-like structure. RV with short antero-ventral inner list, and antero-ventrally either with a small, transparent ‘ lapel’ (see below) or with the valve forming a bulbous expansion. A 1 and A 2 with medium-long natatory setae; expopodite of A 2 with two long and one short seta. Mdpalp with alpha-seta long and smooth, beta-seta stout and hirsute in distal half, gamma-seta relatively slender and hirsute in distal half. T 2 (walking leg) with penultimate segment divided. T 3 a cleaning leg. Caudal ramus and attachment slender. Hemipenis consisting of at least two asymmetrical penal sheets.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF858A49E80DFC0BF9C7.taxon	discussion	Genus allocated: Bennelongia De Deckker & McKenzie, 1981 Remarks Bennelongia was previously allocated to the subfamily Cypridinae Baird, 1845 within the family Cyprididae Baird, 1845, but since the regional revision of this subfamily by Martens (1990, 1992, 2007) it is clear that Bennelongia lacks the additional post-labyrinthal coils of the spermiduct in the hemipenis that are characteristic of Cypridinae s. s. In addition, the remarkable morphology of the anterior part of the valves sets this genus aside from all other extant genera in the Cyprididae. Whether Bennelongiinae subfam. nov. will eventually need to be transferred to the Cyprideidae Martin, 1940 will, hopefully, be resolved by the time our revision of Bennelongia is completed. Meanwhile the new subfamily is lodged in the Cyprididae. Some general aspects of the morphology of Alboa De Deckker, 1981 indicate that it might be close to Bennelongia and thus might also be a candidate to be included in Bennelongiinae subfam. nov., but initial molecular screening shows that Alboa is closer to Eucypris (Vavra, 1891) Daday, 1900 and Heterocypris Claus, 1832 than to Bennelongia (results not shown).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	diagnosis	Diagnosis (modified after De Deckker & McKenzie 1981) Adults with asymmetrical valves, especially anteriorly. LV overlapping RV anteriorly, ventrally and posteriorly, sometimes also dorsally. LV antero-ventrally with a beak-like expansion of the valve margin; anterior calcified inner lamella with two incomplete inner lists, a ventro-proximal one and a dorso-distal one, both extending to about the middle of the valve and slightly or significantly overlapping each other; ventro-proximal list protruding and forming a sulcus running along the proximal inner list and extending in the lip-like structure. RV with short antero-ventral inner list, and antero-ventrally either with a small, transparent flange (here termed ‘ lapel’) or with valve forming a bulbous expansion; RV set with tubercles along most of the anterior, ventral and posterior valve margin. Juveniles with completely different valve morphology, with symmetrical valves, without beak-like expansion on LV or lapel on RV and with external surface either pitted or reticulated or set with wartlike tubercles in most lineages in the genus. A 1 and A 2 with medium-long natatory setae. Md-palp with alpha-seta long and smooth, beta-seta stout and hirsute in distal half, gamma-seta relatively slender and hirsute in distal half. Mx 1 - palp with distal segment rectangular. T 1 with prehensile palps in males strongly asymmetrical, right palp with broad terminal segment, left palp with sickle shaped distal segment. T 2 with seta d 1> d 1, sometimes twice as long, penultimate segment of endopod (segment 3) divided. T 3 a cleaning leg. Caudal ramus and attachment slender. Hemipenis consisting of at least two, asymmetrical penal sheets.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	discussion	Remarks The extent of the difference in morphology between juveniles and adults in most lineages of Bennelongia is unusual in non-marine Ostracoda. The difference of the extent of this dimorphism between the lineages, and its relevance, will be discussed elsewhere in a separate paper. Differential diagnosis Bennelongia is immediately distinguishable from all other cypridid genera by the peculiar morphology of the anterior parts of the valves, especially of the LV (see above).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	type_taxon	Type species Bennelongia harpago De Deckker & McKenzie, 1981 (Queensland, Australia).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	discussion	Other species allocated (only area of type locality given) See Table 2.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	distribution	Distribution The genus is most likely endemic to Australia and New Zealand and can be considered as one of the more typical ostracod groups of the Australasian region. De Deckker (1981 a) suggested that Strandesia flavescens Klie, 1932 and Strandesia feuerborni Klie, 1932, both from Indonesia (Sumatra, Java), might also belong in this genus, but Savatenalinton & Martens (2010) and Martens & Savatenalinton (2011) retained both species in their original genus, Strandesia Stuhlmann, 1888.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352767FF878986ECA0FD27F9D4.taxon	description	General valve morphology (Figure 4) In order to allow accurate descriptions of species in this genus, it is necessary to unequivocally establish the homology of the different marginal structures in both valves, as these have undergone remarkable evolutionary changes (Figure 4 A). In the LV, the valve margin and two inner lists are of importance. The valve margin shows an anteroventral beak-like expansion. The anterior calcified inner lamella carries two inner lists, a proximal ventral and a distal dorsal one. Together, these lists line a sulcus, a depression in the calcified inner lamella which further expands in the beak-like expansion. The proximal inner list can be significantly elevated (Figure 4 B). In the RV, an outer list, the valve margin, a selvage and two inner lists form a complex structure. The outer list is invariably modified, completely or partially. In B. cygnus sp. nov., the middle part of the outer list is still a list, the posterior part is modified into a row of tubercles and the anterior part in a row of tubercles and a lapel (Figure 4 C). In most (all?) species of the B. australis group (see below), the entire outer list is modified into a row of tubercles and the antero-ventral lapel (Figure 4 D). In the species of the B. pinpi lineage, the entire outer list is modified into a row of tubercles, which stops where in other species the lapel is formed and where in these species the bulbous expansion occurs (Figure 4 E). What looks to be the anterior valve margin in the RV is actually a selvage, which can be modified in places, with swollen or flange-like parts near the lapel. The actual anterior valve margin is strongly reduced, but can in most species still be detected as a row of small tubercles set with setae (Figure 4 C-E). The calcified inner lamella also has two incomplete inner lists, here the ventral one is distal and the dorsal one more proximal. Internal to the ventral inner list, the calcified inner lamella forms a shallow sulcus, which matches the deeper one in the LV. The posterior selvage is more easily recognizable as such. Most Bennelongia species have some form of valve ornamentation, either a pitted surface or set with tubercles and / or with long setae. This is species-dependent (though seemingly with similar patterns within a lineage) and can vary with water-chemistry. But even species with almost smooth valve surfaces (e. g. the B. cygnus lineage, see below) invariably have a field of external tubercles along the anterior margin on the RV (see Figure 4 F, G - not to be confused with the marginal row of tubercles in the RV, which is mostly internally visible). This field can be narrower or wider, according to species, but is always present. Its function remains unknown.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352765FF878A06EC8DFEE0F85E.taxon	discussion	Remarks This lineage so far consists of two species, here described as new: B. cygnus sp. nov. and B. frumenta sp. nov. They are characterized by a relatively small carapace (L = 1.6 mm) with triangular shape in lateral view, a RV with a pointed lapel and by a hemipenis of the B. australis- type, i. e. with ms with oblique, nearly straight distal margin and a boot-shaped LS, a sickle shape Lpp and a Rpp with elongated second segment.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	description	(Figs 5 - 9)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	etymology	Etymology The type locality of this new species is situated in the Swan Valley near Perth. We thus name this species after the (black) swan, Cygnus in Latin.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	diagnosis	Diagnosis Valves triangular in lateral view, weakly pitted, relatively narrow in dorsal view and with rostrum weakly developed. LV with beak weakly developed. RV with lapel long and narrow, ventrally tear-like, almost pointed. Hemipenis with ls extending beyond ms, ls distally rounded and with bluntly pointed apex. ms dorsally with additional lobe-like expansion. Lpp with distal segment sickle-shaped, but relatively short. Rpp with sub-trapezoidal distal segment.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1330; H = 933 - 954. LV: L = 1330 - 1410; H = 968 - 969. Female: RV: L = 1450 - 1600; H = 1000 - 1070. LV: L = 1550 - 1750; H = 1030 - 1130. Cp: L = 1690 - 1790; W = 1120 - 1130; H = 1100.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	distribution	Type locality Ellen Brook Nature Reserve, Swan Valley, near Perth (WA); approximate coordinates: 31 º 44 ’ 00 ” S 116 º 01 ’ 00 ” E. Material used for the present description was collected on 25 Sep. 1991, 3 Oct. 1997 and 2 Oct. 1998, all by SH.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	materials_examined	Type material Holotype Male (WAM. C 49370), with soft parts dissected in a sealed slide and valves stored dry in a micropalaeontological slide. Allotype Female (WAM. C 49371), with soft parts dissected in a sealed slide, and valves stored dry in a micropalaeontological slide. Paratypes Five males dissected and stored as the holotype (WAM. C 49372 - 49375, OC. 3310); RV + LV of one female (OC. 3311). Several in toto specimens in EtOH (voucher specimens WAM. C 49376). Other material investigated Perth Airport unnamed swamp, collected by SH on 12 Sep. 1995, approximate coordinates: 31 º 56 ’ 00 ” S 115 º 58 ’ 00 ” E Goonaping Swamp, collected by Adrian Pinder and Jane McRae on 3 Oct. 1997, approximate coordinates: 32 º 27 ’ 46 ” S 119 º 25 ’ 1 ” E. (illustrated specimens from this locality: RV + LV of a female (WAM. C 49377); two female carapaces (WAM. C 49378) all from OST 11 C). Christopher Brook, collected by Adrian Pinder and Jane McRae on 28 Oct. 1997, approximate coordinates: 32 º 10 ’ 12 ” S 116 º 47 ’ 39 ” E. (illustrated specimens from this locality: RV + LV of a female (WAM. C 49379) from OST 11 A). Cobertup Swamp, collected by Andrew Storey and SH on 19 Oct. 1996, approximate coordinates: 34 º 27 ’ 00 ” S 116 º 49 ’ 00 ” E. One Tree Hill, collected by SH and Adrian Pinder on 11 Aug. 1999, approximate coordinates: 29 º 35 ’ 19 ” S 115 º 26 ’ 31 ” E.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	diagnosis	Differential diagnosis Bennelongia cygnus sp. nov. defines the B. cygnus lineage by its triangular shape, the simple type of hemipensis and the pointed lapel. It can be distinguished from the other species in this lineage, B. frumenta sp. nov. (see below), by the absence of a cavity in the selvage near the lapel, the presence of a dorsal lobe on the ms of the hemipenes, the short second segment of the Lpp and the broad base of the second segment of the Rpp.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	description	Additional descriptions Male valves (Figure 5 C, D) slightly smaller and more highly arched than female valves (Figure 5 A, B, H), otherwise very similar in appearance. Both valves triangular, with greatest height situated in the middle of the valves, dorsal margins equally sloping to both anterior and posterior margins, ventral margin almost straight. Valves weakly pitted and set with few, very short setae. LV (Figure 5 A, C) with posterior calcified lamella narrow, inner list running along valve margin and creating a narrow sulcus; the latter continuing towards the anterior side and widening up in between both inner lists (see diagnosis of genus); antero-ventral beak weakly developed. RV (Figure 5 B, D, H) of similar shape as LV, smaller and slightly less high; posterior and ventral margin set with tubercles, anterior calcified lamella with short inner list; lapel relatively long, rather ventrally situated and tear-shaped at its ventral edge (Figure 5 I, J); posterior side with long inner list (reaching almost up to dorsal margin) and with selvage clearly inwardly displaced. Width of carapace in dorsal (Figure 5 E) and ventral (Figure 5 F) views more than half the length, greatest width situated in the middle, LV overlapping RV on all sides, especially anteriorly and posteriorly (Figure 5), anterior rostrum very weakly built, to almost absent. A 1 (Figure 6 E) with all segments relatively short and narrow, natatory setae long, chaetotaxy as typical of the family. A 2 (Figure 6 A, C) with 5 natatory setae extending beyond tips of end claws, basic chaetotaxy and sexual dimorphism in chaetotaxy of penultimate segment as typical of the family: in female with claws G 1 - G 3 and z 1 - 3 setae; in males with G 1 a short claw, G 2 a large claw and G 3 a seta of intermediate length; z 1 and z 2 large claws, z 3 a long seta. Md coxa (Figure 7 A) relatively slender, without special features. Mandibular palp (Figure 7 B-D, G) with chaetotaxy as typical of the family, endclaws and gamma seta unusually slender. Mx 1 (Figure 7 E, F) with second palp segment rectangular, c. twice as long as basal width, 3 claws of this segment relatively slender. Third endite with smooth ‘ zahnborsten’. Sideways directed bristles on first endite of unequal length. T 1 (Figure 8 C-E) with endite bearing 16, mostly hirsute, setae of unequal length. Females with endopod a palp bearing 3 unequal apical setae. Males with endepod developed in asymmetrical prehensile palps. Rpp (Figures 8 D, 9 C) with basal segment stout and only slightly longer than the largest width, subapically with two unequal sensory organs; second segment trapezioidal, with blunt dorsal and pointed ventral edge. Lpp (Figures 8 C, 9 D) with first segment more slender, almost twice as long as wide, distal segment sickle-shaped but relatively short (L = <half L of first segment). T 2 (Figure 6 B) a walking leg with seta d 1> d 2. T 3 (Figure 6 D) a cleaning limb. CR (Figure 9 E) and its attachment (Figure 9 F) slender. Hemipenes (Figures 8 A, B; 9 A, B) almost symmetrical, with ls slender, with rounded dorsal margin and bluntly pointed distal edge; ms consisting of two sub-lobes, one rectangular, the second one elongated and ventrally directed, with rounded edge.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352763FF9E8A0EEB4AFDAAFE5B.taxon	discussion	Remarks One male specimen (WAM. C 49375 – Figure 9 A, B) had aberrant morphology of hemipenes and Rpp. The hemipenes were asymmetrical, with one being typical of the species, the other bearing an additional thumb-like expansion of the ls. The same specimen also had an additional distal thumb-like expansion of the second segment of the Rpp. Rather than considering this a different species, we decided that it is most likely a teratological specimen.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	description	(Figs 10 - 11)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	etymology	Etymology Bennelongia frumenta sp. nov. is typical of the south-western side of the Western Australian wheatbelt. Wheat = frumenta in Latin.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	diagnosis	Diagnosis Valves triangular in lateral view, weakly pitted, width more than half the length in dorsal / ventral view and with rostrum well-developed. LV with beak weakly developed. RV with lapel triangular, ventrally pointed; selvage near lapel with a cavity, visible with SEM, but especially with transparent light. Hemipenis with ls slightly extending beyond ms, ls distally rounded and with bluntly pointed apex; ms ventrally and dorsally without lobe-like expansion. Rpp with trapezoidal distal segment; with nearly straight distal margin and very narrow base. Lpp with distal segment sickle-shaped, elongated, almost as long as first segment.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1310; H = 907. LV: L = 1430; H = 933. Cp: L = 1320 - 1330; H = 841; W = 810 - 826. Female: RV: L = 1320 - 1390; H = 851 - 905. LB: L = 1460 - 1500; H = 962. Cp: L = 1400 - 1520; H = 878; W = 901 - 907.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	distribution	Type locality Kodjinup Melaleuca Swamp, 6 km N of Lake Muir in the Cranbrook shire (WA), collected by Adrian Pinder and Jane McRae on 2 Oct. 1998, approximate coordinates; 34 º 23 ’ 45 ” S 116 º 39 ’ 1 ” E (OSTR 11 A, SPS 105).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	materials_examined	Type material Holotype Female (WAM. C 49380), with soft parts dissected in a sealed slide and valves stored dry in a micropalaeontological slide. Allotype Male (WAM. C 49381), with soft parts dissected in a sealed slide, valves kept in EtOH (decalcified). Paratypes LV + RV of a female (OC. 3312); one female carapace (WAM. C 49383); three male carapaces (WAM. C 49382 A-C); soft parts of one male (WAM. C 49384 – valves lost). Several in toto specimens in EtOH (voucher specimens WAM. C 49385). Other material investigated Wetland south east of Kodjinup Swamp, collected by Andrew Storey and SH on 21 Oct. 1997, approximate coordinates: 34 º 23 ’ 00 ” S 116 º 40 ’ 00 ” E. Specimens from this locality: a dissected male (WAM. C 49386); LV + RV of a female (WAM. C 49387). Job’s Sump, collected by Jane McRae and Mick Smith on 10 Oct. 1997, approximate coordinates: 32 º 21 ’ 15 ” S 117 º 39 ’ 27 ” E (SPS 060). West Kulunilup Swamp, collected by Andrew Storey and SH on 22 Oct. 1997, approximate coordinates: 34 º 20 ’ 00 ” S 116 º 47 ’ 00 ” E. Specimens from this locality: several voucher specimens in EtOH (OSTR 13 E), slide # 38, a dissected male (OC. 3313) and a female CP (WAM. C 49388). Unnamed claypan, Pingrup, collected by Adrian Pinder on 13 Sep. 2007, approximate coordinates: 33 º 26 ’ 49 ” S 118 º 30 ’ 41 ” E. Specimens from this locality: several voucher specimens in EtOH (OSTR 13 G, ABP 051); LV + RV of a female (OC. 3314). Lake Wheatfield, collected by David Cale on 26 Oct. 2005, approximate coordinates: 33 º 48 ’ 46 ” S 121 º 55 ’ 38 ” E (SPM 005 B).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	diagnosis	Differential diagnosis Bennelongia frumenta sp. nov. belongs to the B. cygnus lineage because of its triangular shape, the simple type of hemipenis and the pointed lapel. It can be distinguished from the other species in this lineage, B. cygnus sp. nov. (see above), by the presence of a cavity in the selvage near the lapel, the absence of a dorsal lobe on the ms of the hemipenes, the elongated second segment of the Lpp and the narrow base of the second segment of the Rpp.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235277CFF9B8A03E81DFB84FA6F.taxon	description	Additional description Valves triangular (Figure 10 A-D), with greatest height situated in the middle; LV overlapping RV on all sides (Figure 10 K), dorsally only in the first half of the carapace, ventral margin almost straight; width of carapace in dorsal and ventral views (Figure 10 E-H) more than half the length, greatest width situated in the middle, anterior rostrum strong; carapace weakly pitted and set with few, very short setae. Male valves slightly smaller and more highly arched than female valves, otherwise very similar in appearance. LV (Figure 10 A, C) with posterior calcified inner lamella narrow, inner list running along valve margin and creating a narrow sulcus; this sulcus continuing towards the anterior side and widening up in between both anterior inner lists; antero-ventral beak weakly developed. RV (Figure 10 B, D) of similar shape as LV, smaller and slightly less high; posterior and ventral margin set with tubercles, anterior calcified inner lamella with short inner list, the latter forming a ‘ cavity’ in and on the selvage near the lapel; lapel relatively long, rather ventrally situated and ventrally pointed (Figure 10 I, J); inner margin of posterior calcified inner lamella with long inner list (reaching almost up to dorsal margin) and with selvage submarginal. Most appendages as typical of the genus and without special features. Rpp (Figure 11 B) with basal segment elongated, c. 1.5 x the central width; subapically with two unequal sensory organs; second segment trapezoidal, with blunt dorsal and pointed ventral edge, distal margin nearly straight, base of segment very narrow. Lpp (Figure 11 C, D) with first segment more slender, more than twice as long as wide, subapically with an elongated outgrowth, subapically with a short sensory organ; distal segment sickle-shaped and elongated, longer than dorsal margin of first segment. Walking leg (Figure 11 E) stout and hirsute. Hemipenes (Figure 11 A) symmetrical; LS with rounded dorsal margin and bluntly pointed distal edge; MS with oblique but straight distal margin, ventrally with a broadly rounded lobe, ventrally without additional lobe.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352779FF9A8A3DEC02FC3DFC1B.taxon	discussion	Remarks Brady (1886) described Chlamydotheca australis based on empty valves collected by Prof R. Tate from Penola (South Australia). The description was incomplete and, as pointed out by De Deckker (1981 a), most likely also erroneous: although the LV (Brady 1886, plate 9 fig. 7) is undoubtedly of a species of Bennelongia, the RV is not. De Deckker (loc. cit.) investigated the type material of Chlamydotheca australis in the British Musuem (presently the Natural History Museum, London) and found that the slide contained the illustrated Bennelongia LV as well as a second LV, most likely belonging to a species of Heterocypris. It is likely that this is the valve illustrated by Brady (1886, plate 9 fig. 8). As it is the RV that shows most specific features of Bennelongia species, the true identity of Bennelongia australis will be impossible to establish based on the type material only. New material from Penola will be necessary and topotypes must be established to determine what species is the true B. australis. Below, we describe 5 sibling species in this lineage, which can be identified based on shape of the valves, form of the lapel on the RV, shape of the hemipenis and of the prehensile palps. The identity of four of these species was confirmed by molecular screening (see above). De Deckker (1981 a) extensively redescribed what he thought to be B. australis, based on material from Western Australia (mainly from pools near Leonora and Cunderdin). Based on the illustrated lapels, it is clear that this redescription is based on at least two different species within the B. australis lineage, neither of which are present in our collections. Given the regional specificity of the species within this lineage, it is unlikely that the true B. australis, described from South Australia is amongst the species used by De Deckker (1981 a). Davies & Christidis (1997) also illustrated a specimen of what they labelled B. australis, collected from lakes in and around Perth. We have collected material from two of the same lakes and describe this species below as B. gwelupensis sp. nov. The species figured by Davies & Christidis (1997) as Bennelongia sp. was described above as B. cygnus sp. nov. Karanovic (2008) illustrated specimens from Pilbara as B. australis s. l. These specimens belong to B. strellyensis sp. nov. in the B. pinpi lineage (see below). The B. australis lineage is characterized by relatively large carapaces, LV with intermediately sized beaks and RV with lapel. Most species described here also have rounded dorsal margins and have pronounced anterior (and sometimes posterior) rostrum in dorsal view.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	description	(Figs 12, 13, 14 A, B)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	etymology	Etymology The species is named after its type locality, Coondiner Pool, Pilbara.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, and weaker posterior rostrum. LV with pronounced anteroventral beak and rounded dorsal margin. Lapel on RV tear-shaped, dorsally sloping towards valve margin. Hemipenis with MS with straight margin, ventrally widely produced as a rounded lobe; ls in one hemipene rounded, and distally bluntly pointed, in second hemipenis distally with thumb-like process; tips of ls and ms well-separated, ls extending clearly beyond ms. Rpp with distal segment triangular, rather broad. Lpp with proximal segment bearing rectangular apical outgrowth; distal segment sickleshaped, tapering and rather elongated.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1640, H = 971. LV: L = 1690, H = 911. Cp: L = 1690 - 1710; W = 911 - 913. Female: RV: L = 1750, H = 1040. LV: L = 1860, H = 1120. Cp: L = 1810 - 1890; W = 1020 - 1080.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	distribution	Type locality Coondiner Pool, Pilbara, WA (sample KIES 14); approximate coordinates: 22 º 43 ’ 26 ” S 119 º 39 ’ 23 ” E. All material used for the present description collected on 24 Apr. 2006 by the authors.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	materials_examined	Type material Holotype Male (WAM. C 49389), with soft parts dissected in a sealed slide and valves stored dry in a micropalaeontological slide. Allotype Female (WAM. C 49390), with soft parts dissected in a sealed slide and valves stored dry in a micropalaeontological slide. Paratypes One male dissected and stored as the holotype (OC. 3315); three male carapaces (WAM. C 49391 A, WAM. C 49392), two female carapaces (WAM. C 49391 B-C). 2 RV + 1 LV of a female (OC. 3317 A-C). Several in toto specimens in EtOH (WAM. C 49393). Other material investigated Ethel Creek Clay pan, Pilbara, approximate coordinates: 22 º 49 ’ 32 ” S 120 º 15 ’ 32 ” E. Collected by the authors on 24 Apr. 2006.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	diagnosis	Differential diagnosis The species belongs to the B. australis group because of the generally large size (L> 1500 µm), the presence of a lapel on the RV and of a strong anterior rostrum in dorsal view. It can be distinguished from the other species in this lineage by the rounded dorsal margin of the LV, the tear-shaped lapel, the broad second segment of the Rpp, the fact that tips of ls and ms of the hemipenes are well-separated from each other and that ls extends well beyond the ms. The shape of the lapel somewhat resembles that of the second species redescribed by De Deckker (1981 a) as B. australis (from a pool close to Cunderdin), yet these latter specimens have much more highly arched valves, while also the edge of the beak in the LV is less pointed than in B. coondinerensis sp. nov.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352778FF978A5EEE56FE76FDDD.taxon	description	Additional description Valves in lateral view with rounded dorsal margin (Figure 12 A-D), LV overlapping RV on all sides (Figure 12 K, L), greatest height anterior to the middle; in dorsal and ventral views (Figure 12 E-H) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side weakly pointed, LV dorsally ridge-like; external surface weakly pitted and set with setae of intermediate length. LV in inner view (Figure 12 A, C) with rounded dorsal margin, greatest height situated in the middle; antero-ventral beak-like expansion rather large; posterior part of ventral margin markedly sloping in dorsal direction. RV (Figure 12 D) in inner view with greatest height situated in front of the middle, dorsal margin almost straight for about the middle third; posterior selvage clearly inwardly displaced in the posterior half of the valve, posterior inner list merging with posterior selvage at about halfway the length of the latter; lapel tear-shaped (Figure 12 I, J), dorsally sloping towards the valve margin, ventrally abruptly curving towards it; antero-ventral inner list running to about halfway the lapel; selvage at height of lapel expanded and slightly striate. Most appendages as typical of the genus and without special features. Rpp (Figure 13 A) with first segment c. 1.5 times as long as wide, subapically with one long but slender, and one short sensory organ; second palp segment triangular, rather broad, with almost straight distal margin; apically with one small sensory organ. Lpp (Figure 13 B) with first segment elongated, more than twice as long as central width, subapically with one stout sensory organ, apically with a rectangular outgrowth, bearing one small sensory organ; second palp segment sickle-shaped and relatively elongated (L> ½ L of first segment). T 2 (Figure 13 C) a hirsute walking leg, with seta d 1> seta d 2. Hemipenes (Figure 14 A, B) with tips of ls and ms well separated from one another, ls reaching well beyond tip of ms, distal part of ms produced into an elongated lobe; distal part of ls bluntly pointed with distal margin rounded in one hemipenis, ls almost rectangular with distal thumb-like process in the other.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	description	(Figs 14 C-F, 15)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	etymology	Etymology Named after the type locality, Lake Gwelup, Perth.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, and weaker posterior rostrum. LV with pronounced anteroventral beak and rounded dorsal margin. Lapel on RV almost evenly rounded, dorsally sloping towards valve margin. Hemipenis with MS with straight margin, ventrally widely produced as a bluntly pointed lobe; ls in both hemipenes evenly rounded, distally with blunt tip; tips of ls and ms well-separated, ls and ms nearly equally long. Rpp with distal segment triangular, relatively elongated. Lpp with proximal segment bearing rectangular apical outgrowth with rounded tip; distal segment sickle-shaped, elongated, as long as first segment, and with almost parallel margins, except for distal part.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	description	Measurements (all measurements in µm) Female: RV: L = 1710; H = 1040. LV: L = 1830; H = 1110. Cp: L = 1610 - 1820; H = 1080; W = 860 - 916.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	distribution	Type locality Lake Gwelup, Perth (WA); approximate coordinates: 31 º 52 ’ 37 ” S 115 º 47 ’ 18 ” E. Material used for the present description collected on 23 Dec. 2008 and on 15 July 2010 by the authors.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	materials_examined	Type material Holotype A female (WAM. C 49394), with soft parts dissected in glycerine in a sealed slide, valves stored dry in a micropalaeontological slide. Allotype As the population collected at the type locality is fully parthenogenetic, no males occurred (at least at the time of collection) and no allotype can be designated. Paratypes Four female carapaces (WAM. C 49395 A-D) and several in toto specimens in EtOH (WAM. C 49396). Other material investigated Big Carine Swamp, Perth, collected by the authors on 15 July 2010, approximate coordinates: 31 º 51 ’ 08 ” S 115 º 47 ’ 04 ” E. Several pools between Bunbury and Perth, collected by the authors on 04 July 2010 and 31 July 2010. The only male found thus far was in a dense population in a large ditch next to a parking lot beside Forrest Highway, just northeast of the Greenlands Road intersection, 32 º 38 ’ 29 ” S 115 º 48 ’ 34 ” E (sample MR / 10). Specimens from this locality: a male (WAM. C 49397) dissected and stored as the holotype; 3 female carapaces (WAM. C 49398 A-C). All other pools, as well as Lake Gwelup and Big Carine Swamp, held only parthenogenetic populations. Small unnamed streamlet, inflowing in Leschenault Estuary, collected by the authors on 04 July 2010 and 31.7.2010 This is the most southern locality thus far of this species; approximate coordinates: 33 º 19 ’ 10.3 ” S 115 º 41 ’ 17.3 ” E (sample MR / 09). Specimens from this locality: RV + LV of one female and a Cp of a female (OC. 3316 A-B);	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	diagnosis	Differential diagnosis This species belongs to the B. australis group because of the generally large size (L> 1500 µm), the presence of a lapel on the RV and of a strong anterior rostrum in dorsal view. It can be distinguished from the other species in this lineage by the rounded dorsal margin of the LV, the rounded lapel, the rather narrow second segment of the Rpp, the fact that tips of ls and ms of the hemipenes are wellseparated from each other and that ls and ms are almost equally long.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	description	Additional description Valves in lateral view with rounded dorsal margin (Figure 15 A-B), LV overlapping RV on all sides (Figure 15 C, H), greatest height anterior to the middle; in dorsal and ventral view (Figure 15 E, G) with greatest width in the middle of the carapace; anterior rostrum well-developed (Figure 15 F), posterior side bluntly pointed, LV dorsally ridge-like; external surface weakly pitted and almost devoid of setae. LV (Figure 15 A) in inner view with rounded dorsal margin, greatest height situated anterior to the middle; antero-ventral beak-like expansion rather large; ventral margin almost straight over most of its length. RV (Figure 15 B) in inner view with greatest height situated well in front of the middle, dorsal margin almost straight for about the middle third; posterior selvage submarginal, posterior inner list merging with posterior selvage at about halfway the length of the latter; lapel (Figure 15 G, I, J) rounded, dorsally sloping towards the valve margin, ventrally gently curving towards it; antero-ventral inner list running to about halfway the lapel; selvage at height of lapel expanded, not striate. Most appendages as typical of the genus and without special features. Rpp (Figure 14 C) with first segment c. 1.5 times as long as wide, subapically with one long but slender, and one short sensory organ; second palp segment triangular, rather elongate, with sinuous distal margin; apically with one small sensory organ. Lpp (Figure 14 E) with first segment elongated, more than twice as long as central width, subapically with one long but slender sensory organ, slightly swollen in the middle; apically with a short and distally rounded outgrowth, bearing one very small sensory organ; second palp segment nearly straight, sickleshaped and relatively elongated (L = L of first segment); distal part of this segment straight and with nearly parallel margins. Hemipenes (Figure 14 D, F) symmetrical, with tips of ls and ms well separated from one another, ls and ms almost equally long, distal part of ms produced into an elongated lobe; distal part of ls bluntly pointed with distal margin rounded.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352773FF908A2CEB4AFB86F85E.taxon	discussion	Remarks Davies & Christidis (1997) mention the presence of B. australis in 11 Perth wetlands. We have been able to identify two of these populations (Lake Gwelup, Big Carina swamp) as B. gwelupensis sp. nov., but most likely all of the populations of these 11 wetlands belong to this species.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	description	(Figs 16, 17 A-C)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	etymology	Etymology The species has a broad lapel on the RV. Broad = lata in Latin.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, posteriorly bluntly pointed. LV with relatively weak anteroventral beak and perfectly rounded dorsal margin. Lapel on RV broad, running parallel to selvage. Hemipenis with ms with straight margin, ventrally widely produced as a broad lobe; ls in both hemipenes boot shaped, distally with blunt tip; tips of ls and ms close together, ls slightly longer than ms. Rpp with distal segment triangular, broad. Lpp with proximal segment without apical outgrowth; distal segment sickleshaped, elongated, as long as first segment, and with almost parallel margins, except for distal part.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1660 - 1700; H = 930 - 962. LV: L = 1770 - 1810; H = 995 - 1030. Cp: L = 1670 - 1800; H = 881 - 995. Female: Cp: L = 2000 - 2040; H = 1130; W = 1120 - 1170.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	distribution	Type locality Yandoo Billabong, Boolathana Station, Gascoyne, WA (sample SIEK 3); approximate coordinates 24 º 38 ’ 25 ” S 113 º 55 ’ 20 ” E. All material was collected on 7 Apr. 2006 by the authors.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	materials_examined	Type material Holotype Male (WAM. C 49399), with soft parts dissected in a sealed slide, valves stored dry in a micropalaeontological slide. Allotype LV and RV of a female (WAM. C 49400) stored dry in a micropalaeontological slide. Paratypes Two males dissected and stored as the holotype (WAM. C 49401, OC. 3318); two males and one female (WAM. C 49402 A-C) carapaces in a micropalaeontological slide. Several juvenile specimens: one cp of a male (A- 1) (WAM. C 49403); 2 Cp and 1 LV female (A- 1) and 1 Cp female (A- 3) (WAM. C 49404 A-D). Several specimens in EtOH (WAM. C 49405). Other material investigated Tirigie Claypan, Boolathana Station, Gascoyne, WA (sample SIEK 4), approximate coordinates: 24 º 38 ’ 29 ” S 113 º 59 ’ 44 ” E. All material was collected on 7 Apr. 2006 by the authors.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	diagnosis	Differential diagnosis The species belongs to the B. australis group because of the generally large size (L> 1500 µm), the presence of a lapel on the RV and of a strong anterior rostrum in dorsal view. It can be distinguished from the other species in this lineage by the very rounded dorsal margin of both valves, the elongated lapel running parallel to the selvage, the very broad second segment of the Rpp, the fact that tips of ls and ms of the hemipenes are hardly separated from each other and that the ls is slightly longer than the ms.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352770FFAD8AE3EB4AFE55FA10.taxon	description	Additional description Valves in lateral view (Figure 16 A-D) with rounded dorsal margin, LV overlapping RV on all sides (Figure 16 K), greatest height in the middle; in dorsal and ventral views (Figure 16 E-H) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side bluntly pointed, LV dorsally less ridge-like than in the other species of this lineage; external surface weakly pitted and sparsely set with short setae. LV (Figure 16 A, C) in inner view with rounded dorsal margin, greatest height situated in the middle; antero-ventral beak-like expansion moderate; ventral margin almost straight over most of its length. RV (Figure 16 B, D) in inner view with greatest height situated slightly in front of the middle, dorsal margin rounded; posterior selvage submarginal, posterior inner list merging with posterior selvage; remnant of valve margin visible between lapel and valve margin; antero-ventral inner list running to about halfway the lapel; selvage at height of lapel expanded, not striate, lapel (Figure 16 I, J) broad and long, see specific name. Most appendages as typical of the genus and without special features. Rpp (Figure 17 C) with first segment c. 1.5 times as long as wide, subapically with one long but slender, and one short sensory organ; second palp segment triangular, broad, with straight distal margin; apically with one small sensory organ. Lpp (Figure 17 A) with first segment elongated, more than twice as long as central width, subapically with one long, but slender sensory organ, slightly striate in the middle; apically without distinct outgrowth, but with one small sensory organ; second palp segment nearly straight, sickle-shaped and relatively elongated (L = c. L of first segment); distal part of this segment straight and with nearly parallel margins. Hemipenes (Figure 17 B) asymmetrical, with tips of ls and ms situated closely together, ls slightly longer than ms, distal part of ms produced into a blunt lobe; distal part of ls boot-shaped, bluntly pointed with distal margin rounded in one hemipenis, with thumb-like expansion in the other.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	description	(Figs 17 D-H, 18)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	etymology	Etymology The species is named after a small outback town, Cue, near its type locality.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, posteriorly bluntly pointed; in right lateral view, LV overlapping RV on all sides, but with distinct overlap on the dorsal side. LV with relatively rounded anteroventral beak and almost straight dorsal margin. Lapel on RV broad, running parallel to selvage. Hemipenis with ms with straight and broad distal margin, ventrally produced as a long lobe, dorsally also protruding main peniferum as a rounded lobe; ls asymmetrical, in both hemipenes with rounded distal margin and distally with blunt tip, but in one hemipenis without and in the other hemipenis with additional (and unusually elongated) thumb-like expansion; tips of ls and ms well-separated, ls significantly longer than ventral lobe of ms. Rpp with distal segment triangular, relatively elongated and with narrow base. Lpp with proximal segment with small, triangular apical outgrowth; distal segment sickle-shaped, elongated, as long as first segment, and with almost parallel margins, except for distal part.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1810; H = 1040. LV: L = 1930; H = 1130. Cp: L = 1820 - 1960. H = 1090. W = 1040 - 1080. Female: Cp: L = 2170 - 2230; H = 1360; H = 1180 - 1300.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	distribution	Type locality Pool along Great Northern Highway, near Cue, WA, approximate coordinates: 27 º 15 ’ 20 ” S 117 º 58 ’ 58 ” E. All specimens collected by SH on 6 Sep. 2006.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	materials_examined	Type material Holotype Male (WAM. C 49406), with soft parts dissected in a sealed slide, RV stored dry in a micropalaeontological slide (LV lost). Allotype A carapace of a female (WAM. C 49407) stored dry in a micropalaeontological slide. Paratypes Three males (WAM. C 49408 - 49409, OC. 3319) dissected and stored as the holotype; two male carapaces (WAM. C 49410). Several specimens in EtOH (WAM. C 49411). Other material investigated The species is thus far known from its type locality only.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	diagnosis	Differential diagnosis The species belongs to the B. australis group because of the generally large size (L> 1500 µm – it is the largest species here described), the presence of a lapel on the RV and of a strong anterior rostrum in dorsal view. It can be distinguished from the other species in this lineage by the considerable dorsal overlap of the RV by the LV, the straight dorsal margin of the LV, the elongated lapel running parallel to the selvage, the rather narrow second segment of the Rpp, the fact that tip of ls and ms of the hemipenes are well-separated from each other and that the ls extends much longer than the ventral lobe of the ms. Bennelongia cuensis sp. nov. is similar to the previously described species, B. lata sp. nov., especially in the shape of the lapel on the RV, but can be easily distinguished by the shape of the carapace in right lateral view (rounded with limited dorsal overlap in B. lata sp. nov., sub quadrate with considerable dorsal and ventral overlap in B. cuensis sp. nov.) and especially by the shape of the ms on the hemipenis (ventrally bluntly pointed and short and without dorsal lobe in B. lata sp. nov., with elongated and narrow lobes on both dorsal and ventral sides in B. cuensis sp. nov.)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274FFFA88A08EFB0FD5DFC59.taxon	description	Additional description Valves (Figure 18 A, B) in lateral view with straight dorsal margin, LV overlapping RV on all sides (Figure 18 G, H), but with considerable dorsal overlap, greatest height situated in the middle; in dorsal and ventral views (Figure 18 C-F) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side bluntly pointed, LV dorsally ridge-like; external surface weakly pitted and set with short setae. LV (Figure 18 A) in inner view with rather straight dorsal margin, greatest height situated in the middle; antero-ventral beak-like expansion moderate and rounded; ventral margin slightly sinuous in the middle. RV (Figure 18 B) in inner view with greatest height situated slightly in front of the middle, dorsal margin rounded; posterior selvage sub marginal, posterior inner list merging with posterior selvage; remnant of valve margin visible between lapel and valve margin; antero-ventral inner list running to about halfway the lapel; selvage at height of lapel expanded, striate; lapel (Figure 18 I-K) broad, running parallel to selvage, slightly striate. Most appendages as typical of the genus and without special features. Rpp (Figure 17 D) with first segment c 1.5 times as long as wide, subapically with one long but slender, and one short sensory organ; second palp segment elongate, with narrow base, apically with one small sensory organ. Lpp (Figure 17 G, H) with first segment elongated, more than twice as long as central width, subapically with one long, stout sensory organ; apically with small, rounded outgrowth, a small sensory organ at the base of this lobe; second palp segment nearly straight, sickle-shaped and relatively elongated, as long as first segment; distal part of this segment straight and with nearly parallel margins. Hemipenes (Figure 17 E, F) asymmetrical, with tips of ls and ms well-separated, ls extending well beyond ms, ventral part of ms produced into an elongated, pointed lobe, dorsally also extending beyond peniferum with a rounded lobe; distal part of ls bluntly pointed with distal margin rounded in one hemipenis, with elongated thumb-like expansion in the other.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	description	(Figs 19, 20)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	etymology	Etymology The species is named after it type locality, Bidgelang Pool on Callytharra Springs Station in the Gascoyne, WA.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, posteriorly bluntly pointed; highly arched in right lateral view, LV overlapping RV on all sides. LV with relatively rounded anteroventral beak, rounded dorsal margin and large postero-ventral flange. RV in lateral view highly arched, with highest point of dorsal margin at about 1 / 3 of the total length. Lapel on RV broad, dorsally slightly sloping towards selvage. Hemipenis with ms with straight distal margin, ventrally produced as a pointed lobe, dorsally slightly protruding main peniferum as a bluntly pointed lobe; ls asymmetrical, in both hemipenes with straight distal margin and distally with blunt tip, but in one hemipenis with additional elongated thumb-like expansion; tips of ls and ms almost touching, ls very slightly longer than ventral lobe of ms; hallway perniferum with additional small internal lobe. Rpp with distal segment elongated and with narrow base. Lpp with proximal segment with broad, rectangular apical outgrowth; distal segment sickle-shaped, elongated, as long as first segment, and with almost parallel margins, except for distal part.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1700 - 1780; H = 952 - 1030. LV: L = 1890; H = 1040 - 1110. Cp: L = 1920; W: 1030. Female: RV: L = 1950; H = 1170. LV: L = 2130; H = 1260. Cp: L = 2030 - 2090; H = 1230; W = 1130 - 1210.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	distribution	Type locality Bidgelang Pool, Callytharra Springs Station, Gascoyne (WA), approximate coordinates: 24 º 34 ’ 32 ” S 115 º 36 ’ 01 ” E. All material collected by SH on 25 Aug. 1994 (OSTR 14 D).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	materials_examined	Type material Holotype Male (WAM. C 49412), with soft parts dissected in a sealed slide, valves stored dry in a micropalaeontological slide. Allotype LV and RV of a female (WAM. C 49413) stored dry in a micropalaeontological slide. Paratypes Three males dissected and stored as the holotype (WAM. C 49414, WAM. C 49417, OC. 3320); one male (WAM. C 49415 A) and three females (WAM. C 49415 B-D) carapaces stored dry in a micropalaeontological slide, RV + LV of one male (WAM. C 49416). Several specimens in EtOH (WAM. C 49418). Other material investigated The species is known from its type locality only.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	diagnosis	Differential diagnosis The species belongs to the B. australis group because of the generally large size (L> 1500 µm), the presence of a lapel on the RV and of a strong anterior rostrum in dorsal view. It can be distinguished from the other species in this lineage by the highly arched aspect of carapace and valves, the strongly produced postero-ventral flange on the LV (unique character for this species in the B. australis lineage), the broad and elongated lapel on the RV, dorsally sloping towards the selvage, the narrow second segment of the Rpp, and the fact that the tips of ls and ms of the hemipenes are almost touching each other.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E90235274AFFA48A5CEE1BFC82FECB.taxon	description	Additional description Valves in lateral view (Figure 19 A-D) with middle part of dorsal margin almost straight, LV overlapping RV on all sides (Figure 19 K), greatest height situated posteriorly from the middle; in dorsal and ventral views (Figure 19 E-H) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side bluntly pointed, LV dorsally ridge-like; external surface weakly pitted and set with short setae. LV (Figure 19 A, C) in inner view with slightly rounded dorsal margin, greatest height situated in the middle; antero-ventral beak-like expansion moderate and rounded; ventral margin slightly sinuous in the middle; postero-ventral flange strongly produced. RV (Figure 19 B, D) in inner view with greatest height situated at about 1 / 3 of total length, dorsal margin rounded; posterior selvage not sub-marginal, well inwardly displaced, posterior inner list merging with posterior selvage below half of the height of the valve; remnant of valve margin visible between lapel and valve margin; antero-ventral inner list running to about 1 / 4 of length of the lapel; selvage at height of lapel expanded, striate; lapel broad, dorsally slightly sloping towards selvage, slightly striate (Figure 19 I, J). Most appendages as typical of the genus and without special features. Rpp (Figure 20 A, E) with first segment c. 1.5 times as long as wide, subapically with one long and basally stout, and one very short sensory organ; second palp segment elongate and with narrow base, with straight distal margin; apically with one small sensory organ. Lpp (Figure 20 C, D) with first segment elongated, length c. 1.5 x central width as central width, subapically with one long, stout sensory organ; apically with broad, rectangular outgrowth, a small sensory organ at the base of this lobe; second palp segment nearly straight, sickle-shaped and relatively elongated, as long as first segment; distal part of this segment straight and with nearly parallel margins. Hemipenes (Figure 20 B) asymmetrical, with tips of ls and ms almost touching, ls and ms almost equally long; ventral part of ms produced into a bluntly pointed lobe, dorsally also extending beyond peniferum with a rounded lobe; ls almost rectangular, distal margin almost straight in both hemipenes, ventrally bluntly pointed in one hemipenis, with elongated thumb-like expansion in the other.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA48A10EBAFFBC6FCCD.taxon	discussion	Remarks Bennelongia pinpi De Deckker, 1981 was described from Pine Tree Creek Lagoon in Queensland and is characterized by a very pronounced antero-ventral beak-like expansion in the LV, more so than in any other species of Bennelongia, and also by the absence of an antero-ventral lapel on the RV and by the presence of an antero-ventral expansion of the RV. The following two species are allocated to this species group based on the latter two features, as the beak on the LV is not particularly large in either of them. An additional character might be the heavily sclerotized dorsal margins of the A 1 in this species group (Figure 24 F).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	description	(Figs 21, 22)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	etymology	Etymology Named after the type locality, Strelley Station, Pilbara (WA).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, posteriorly bluntly pointed; rounded in right lateral view, LV overlapping RV on all sides, external valve surface centrally almost completely smooth and devoid of setae. LV with pronounced anteroventral beak, and highly arched, rounded dorsal margin. RV in lateral view rather elongated, with highest point of dorsal margin at about 1 / 3 of the total length; lapel on RV absent, but valves building a triangular bulbous expansion in the same location; anterior selvage with flange-like expansion and with additional inner submarginal list. Hemipenis symmetrical, heavily sclerotized. ms forming a three-dimensional structure, embracing ls along three sides (proximal, ventral, distal). ls a large, slightly curved lobe, with almost parallel sides, distally bluntly pointed in ventral direction. About halfway the hemipenal body, one additional lobe present. Rpp with distal segment short and rounded, with blunt tip bearing a sensory organ. Lpp with distal segment sickle-shaped, rounded and short.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1710 - 1900; H = 1090 - 1130. LV: L = 790 - 2010; H = 1080 - 1220. Cp: L = 1860 - 1880; H = 1120; W = 1140 - 1150. Female: RV: L = 1870; H = 1130. LV: L = 1980; H = 1200. Cp: L = 1880 - 2000; H = 1170; W: 1160 - 1230.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	distribution	Type locality Unnamed roadside pool on Strelley Station, Pilbara, WA (KIES 3), approximate coordinates: 20 º 31 ’ 24 ” S 119 º 03 ’ 40 ” E. All material collected on 22 Apr. 2006 by the authors.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	materials_examined	Type material Holotype Male (WAM. C 49419), with soft parts dissected in a sealed slide, valves stored dry in a micropalaeontological slide. Allotype LV and RV of a female (WAM. C 49420) stored dry in a micropalaeontological slide. Paratypes Three males (WAM. C 49421, 49422, OC. 3321) dissected and stored as the holotype; three females (WAM. C 49423 A-C) and two males (WAM. C 49424 A, B) carapaces stored dry in a micropalaeontological slide; RV + LV of a male (WAM. C 49425). Several juvenile specimens: RV + LV of males (A- 1) (WAM. C 49426); 3 Cp of females (A- 1) and 3 Cp of females (A- 3) (WAM. C 49427 AF). Several specimens in EtOH (WAM. C 49428). Other material investigated Cooliarin Pool, Pilbara (WA), collected by Adrian Pinder on 10 Sep. 2004, approximate coordinates: 20 º 30 ’ 20 ” S 118 º 37 ’ 20 ” E (PSW 033). Specimens from this locality: RV + LV of two females (WAM. C 49429, 49330). Errawallana Spring, Pilbara (WA), collected by Adrian Pinder on 10 Sep. 2004, approximate coordinates: 21 º 37 ’ 48 ” S 117 º 46 ’ 14 ” E (PSW 055). Specimens from this locality: a male, dissected and stored as the holotype (WAM. C 49431).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	diagnosis	Differential diagnosis Bennelongia strellyensis sp. nov. belongs to the B. pinpi lineage, and therefore differs from all other species in the genus not belonging to this lineage by a combination of the following characters: large species, with well-developed anterior rostrum and bluntly pointed posterior end in dorsal view; RV without antero-ventral lapel, but with bulbous expansion of the valve there; antero-ventral selvage with an additional submarginal inner list. LV with antero-ventral beak well-developed. Hemipenis stout and well-sclerotized; ms a ventral, 3 - dimensional structure enveloping the ls. Rpp with short and rounded second segment. Lpp with second segment sickle-shaped and short. Within this lineage, B. strellyensis sp. nov. differs from B. pinpi by the length of the antero-ventral beak in the LV (longer in B. pinpi) and by the shape of the valves in lateral view (more highly arched in B. pinpi). The differences between B. strellyensis sp. nov. and B. kimberleyensis sp. nov. will be outlined below.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	description	Additional description Valves in lateral view (Figures 21 A-D) with rounded dorsal margin, LV overlapping RV on all sides (Figures 21 K, L), greatest height in the middle; in dorsal and ventral views (Figures 21 E-H) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side bluntly pointed, LV. Dorsally only anteriorly ridge-like; external valve surface centrally almost completely smooth and devoid of setae. LV (Figure 21 A, C) in inner view with rounded dorsal margin, greatest height situated in front of the middle; antero-ventral beak-like expansion large. RV (Figure 21 B, D) in inner view with greatest height situated well in front of the middle, dorsal margin rounded; posterior selvage submarginal; lapel on RV absent, but valves building a triangular bulbous expansion in the same location (Figure 21 I, J); anterior selvage with flange-like expansion and with additional inner submarginal list dorsally merging with selvage. Most appendages as typical of the genus and without special features. Rpp (Figures 22 B, 24 D) with first segment c. twice as long as central width, subapically with one longer (but less so than in species of the previous lineage), and one short sensory organ; second palp segment triangular, broad, without clear ventro-apical corner and rounded distal margin; apically with one small sensory organ. Lpp (Figure 22 C, D) with first segment elongated, more than 2.5 x as long as central width, subapically with one large and stout sensory organ, apically with rounded outgrowth, without sensory organ; second palp segment sickle-shaped short, and asymmetrically curved over c. 180 º, distally with a small sensory organ. Hemipenes (Figure 22 A) symmetrical, ls protruding well beyond ms, distally bluntly pointed towards the ventral side, proximally with nearly parallel sides; ms actually consisting of three lobes: distal part of ms (ms 1) produced into a three-dimensional lobe, embracing the ls on proximal and distal sides (this not visible in drawing, which is made of a flattened hemipenis in a slide) and with a small additional dorsal lobe; ms 2 with distal margin nearly straight, ms 3 straight and incomplete. (Remark: ms 2 in some populations weakly developed and almost invisible – see below).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352746FFA08A36E9A6FC66FDCF.taxon	discussion	Remarks The present species closely resembles B. kimberleyensis sp. nov. described below, as was the case of the 5 sibling species in the previous B. australis lineage. The specific status of both new taxa was nevertheless confirmed by molecular methods (see results above), where all the Pilbara and all the Kimberley specimens clustered together in two distinct groups.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	description	(Figs 23, 24)	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	etymology	Etymology The species is named after the area in which it occurs, namely the Kimberley in the northern part of Western Australia.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	diagnosis	Diagnosis Cp with pronounced anterior rostrum, posteriorly bluntly pointed; rounded in right lateral view, LV overlapping RV on all sides, external valve surface centrally almost completely smooth and devoid of setae. LV with pronounced anteroventral beak, and highly arched, rounded dorsal margin. RV in lateral view rather elongated, with highest point of dorsal margin at about 1 / 3 of the total length; lapel on RV absent, but valves building a triangular bulbous expansion in the same location, this expansion very pronounced; anterior selvage with flange-like expansion and with additional inner submarginal list. Hemipenis symmetrical, heavily sclerotized. ms forming a three-dimensional structure, embracing ls along three sides (proximal, ventral, distal). ls a large, slightly curved lobe, with almost parallel sides, distally bluntly pointed in ventral direction. About halfway the hemipenal body, one additional lobe present. Rpp with distal segment short and triangular, with a distinct dorso-apical corner, and with blunt tip bearing a sensory organ. Lpp with distal segment sickle-shaped, very curved, and short.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	description	Measurements (all measurements in µm) Male: RV: L = 1998; H = 1159. LV: L = 2093; H = 1230. Cp: L = 2150 - 2177; H = 1289; W = 1298. Female: RV: L = 2068; H = 1243. Cp: L = 2139 - 2230; H = 1300; W = 1223 - 1268.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	materials_examined	Type locality Taylors Lagoon, Broome, collected by the authors on 21 July 2011, approximate coordinates: 17 º 49 ’ 184 ” S 122 º 41 ’ 936 ” E (KIMB- 01). Type material Holotype Male (WAM. C 49432), with soft parts dissected in a sealed slide, valves stored dry in a micropalaeontological slide. Allotype LV and RV of a female (WAM. C 49433) stored dry in a micropalaeontological slide. Paratypes Two males (WAM. C 49434, OC. 3322) dissected and stored as the holotype; three female (WAM. C 49435) and three male (WAM. C 49436) carapaces stored dry in micropalaeontological slides; RV + LV of a male (WAM. C 49437) stored dry in micropalaeontological slides. Several specimens in EtOH (WAM. C 49438). Other material investigated (all from Kimberley) Parry Lagoons, collected by Andrew Storey on 21 Sep. 2000, approximate coordinates: 15 º 34 ’ 00 ” S 128 º 17 ’ 00 ” E (sample NS- 028). Specimens from this locality: two males, dissected and stored as the holotype (OC. 3323; WAM. C 49441); RV + LV of a male (WAM. C 49440); soft parts of a male (WAM. C 49442); RV + LV of a female (WAM. C 49439). Small dam beside natural wetland along Great Northern Highway, collected by the authors on 21 July 2011, approximate coordinates: 17 º 45 ’ 05 ” S 122 º 56 ’ 14 ” E (KIMB- 04). Roadside pool E of Fitzroy River, Willare, collected by the authors on 21 July 2011, approximate coordinates: 17 º 44 ’ 19 ” S 123 º 38 ’ 30 ” E (KIMB- 05). Roadside pool containing cane grass, E of Fitzroy River along Great Northern Highway, collected by the authors on 21 July 2011, approximate coordinates: 17 º 44 ’ 22 ” S 123 º 37 ’ 54 ” E (KIMB- 06). Myall Bore, E pool, collected by the authors on 22 July 2011, approximate coordinates: 17 º 20 ’ 53 ” S 123 º 39 ’ 59 ” E (KIMB- 10). Pool N side of Gibb Rd, collected by the authors on 22 July 2011, approximate coordinates: 17 º 21 ’ 09 ” S 123 º 44 ’ 11 ” E (KIMB- 11). (Identification of KIMB 04,06,10 & 11 was confirmed using barcoding).	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	diagnosis	Differential diagnosis The present species is slightly larger than B. strellyensis sp. nov. (up to 2.2 mm in Kimberley, up to 2.0 mm in Pilbara), and has a slightly larger antero-ventral beak in the LV, while also the antero-ventral bulbous expansion in the RV is slightly larger. The second segment of the Rpp in B. kimberleyensis sp. nov. is more triangular, i. e. with a clear dorsoapical corner. The second segment of the Lpp is less curved, but this also depends partially on the position of this segment relative to the first segment (the second segment can hinge). The hemipenis has a broader ls, while in some Kimberley populations, lobe ms 2 is hardly visible, but still present (visible at least on the ventral side at high magnification). Additional description Valves in lateral view (Figure 23 A-D) with rounded dorsal margin, LV overlapping RV on all sides (Figure 23 K, L), greatest height in the middle; in dorsal and ventral views (Figure 21 E-H) with greatest width in the middle of the carapace; anterior rostrum well-developed, posterior side bluntly pointed, LV dorsally only anteriorly ridge-like; external valve surface centrally almost completely smooth and devoid of setae. LV (Figure 23 A, C) in inner view with rounded dorsal margin, greatest height situated in front of the middle; antero-ventral beak-like expansion large. RV (Figure 23 B, D) in inner view with greatest height situated well in front of the middle, dorsal margin rounded; posterior selvage submarginal; lapel on RV absent, but valves building a triangular bulbous expansion in the same location (Figure 23 I, J); anterior selvage with flange-like expansion and with additional inner submarginal list dorsally merging with selvage. Most appendages as typical of the genus and without special features. Rpp (Figure 24 A) with first segment c. twice as long as central width, subapically with one longer (but less so than in species of the previous lineage), and one short sensory organ; second palp segment triangular, broad, with clear ventro-apical corner and almost straight distal margin; apically with one small sensory organ. Lpp (Figure 24 B) with first segment elongated, more than 2.5 x as long as central width, subapically with one large and stout sensory organ, apically with rounded outgrowth, without sensory organ; second palp segment sickle-shaped short, and asymmetrically curved over c. 180 º, distally with a small sensory organ. T 2 (Figure 24 E) a hirsute walking leg. Hemipenes (Figure 24 C) symmetrical, ls protruding well beyond ms, distally bluntly pointed towards the ventral side, proximally with nearly parallel sides; ms actually consisting of three lobes: distal part of ms (ms 1) produced into a three-dimensional lobe, embracing the ls on proximal and distal sides (this not visible in drawing, which is made of a flattened hemipenis in a slide) and with a small additional dorsal lobe; ms 2 with distal margin nearly straight, ms 3 straight and incomplete.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
03E902352742FFBC8A5EE8A5FD5DFBA4.taxon	biology_ecology	Ecology and distribution This species occurred in a third of all our localities (pools, clay pans) sampled during 2010 in the southwestern Kimberley and was also found in the north-east near the mouth of the Ord River. The species appears to be common through the Kimberley and occurs in fresh, clear or slightly turbid water. No further environmental data are available.	en	Martens, Koen, Halse, Stuart, Schön, Isa (2012): Nine new species of Bennelongia De Deckker & McKenzie, 1981 (Crustacea, Ostracoda) from Western Australia, with the description of a new subfamily. European Journal of Taxonomy 8: 1-56, DOI: 10.5852/ejt.2012.8
