taxonID	type	description	language	source
03F6F264FFECFF90B828FA42FD2F2781.taxon	biology_ecology	Biology Authors: Pape, Robert B., Thomas, Donald B., and Aalbu, Rolf L. Source: The Coleopterists Bulletin, 61 (4): 519 - 540 Published By: The Coleopterists Society URL: https: // doi. org / 10.1649 / 0010 - 065 X (2007) 61 [519: AROTGE] 2.0. CO; 2 BioOne Complete (complete. BioOne. org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www. bioone. org / terms-of-use. Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. A REVISION OF THE GENUS ESCHATOMOXYS BLAISDELL (TENEBRIONIDAE: PIMELIINAE: EDROTINI) WITH NOTES ON THE BIOLOGY ROBERT B. PAPE Department of Entomology University of Arizona Tucson, AZ 85721, USA DONALD B. THOMAS USDA-ARS Subtropical Agriculture Research Center 2413 E. Hwy 83 Weslaco, TX 78596, USA AND ROLF L. AALBU Department of Entomology California Academy of Sciences Golden Gate Park San Francisco, CA 94118, USA	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE6FF9DB970FEF1FEF524CD.taxon	description	Head. Prognathous, slightly deflexed, dorso-ventrally compressed; subquadrate in dorsal view, cranium somewhat shorter (1.5 – 1.9 mm) than width (1.8 – 2.1 mm), greatest width across epistoma. Lateral epistomal margins rounded and converging toward eyes. Median epistomal lobe produced, arcuate, margin serrulate. Epistomal sutures vague to obsolescent. Supraorbital carina weak and short, beginning at middle of lateral epistomal lobe and fading above middle of eye; cuticle behind eye smooth. Cranial surface set with shallow punctures on dorsum separated from one another by about three to five diameters of a puncture, becoming denser to contiguous on sides of head at base, vague to obsolescent on venter. Eyes, oval, not bulging, anteriorly emarginated to accommodate epistomal canthus; facets in ten horizontal rows, submedian row longest with 8 facets. Posterior margin of eye separated from cervix by distance equal to about one and one-half diameter of eye. Labrum retractile, apex broadly emarginated, edge bristling with golden setae. Mandibles thick, robust, left side dominant (overlapping right), apex bifid; superior margin with stout angular cusp, somewhat larger on right mandible than left. Maxillary palpus approximately equal in length to first two antennomeres, basal segment longest (apical segment more elongate, slender in male, subsecuriform in female). Mentum hexagonal, surface coarsely punctate. Submentum with large ovoid fossa at middle. Gula indistinct; tentorial pits obsolete. Antennae filiform, antennomeres slender except basal segment broadened apically and last segment spindle-shaped, acuminate; third segment longest; fourth to seventh subequal; eighth to tenth successively shorter until penultimate segment, which is about twice as long as wide; last segment about 1 K times length of penultimate; last two and apical half of third to last clothed in short golden pubescence. Thorax. Pronotum cylindrical, shorter than head, wider (1.9 – 2.4 mm) than long (1.4 – 1.7 mm), widest at middle; surface of dorsum with very shallow, widely spaced punctures on anterior and middle of disc, becoming larger and denser posterolaterally; large and scattered ventrally. Distinct, complete lateral carinate bead on side of thorax, ending anteriorly in acute anterolateral angle. Base of pronotum constricted with thick, marginal bead demarcating posterior edge. Elytra inflated, embracing abdominal venter; surface glabrous, with widely spaced punctures tending to run in rows, strongest in humeral area. Epipleura widest basally, about as wide as metepisternum, gradually narrowing to about third abdominal segment, parallel thereafter, apex about half width at base. Femora long, clavate; tibiae and tarsi long, slender. Abdomen. Ovoid in form, widest at middle (3.6 – 4.2 mm), length 5.3 – 6.4 mm; base slightly wider than apposing pronotal base. Surface of abdominal sternites smooth, impunctate. Length of first abdominal sternite from metacoxa to posterior margin slightly shorter or subequal to length of second abdominal sternite; third about two-thirds length of second; fourth about half length of third; fifth about twice length of fourth at midline. Males with elongate, ovoid, pubescent fovea at middle of first abdominal sternite. Genitalia as in E. pholeter.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE6FF9DB970FEF1FEF524CD.taxon	materials_examined	Holotype. Male, CALIF: Riverside Co. Painted Canyon, IX- 13 - 78 / 1 - 7 - 1979, Fred G. Andrews, (b) Ethylene Glycol Pit trap in Desert Wash. Paratypes: 1 male, 2 females, same data as type; 2 females, CALIF. Riverside Co. Painted Canyon, V- 18 - 78 / IX- 14 - 1978 Antifreeze Pit trap, Fred G. Andrews; 2 females, CALIF. Riverside Co. Painted Canyon, V- 18 - 78 /! X- 13 - 1979, F. Andrews & K. Cooper (b) Ethylene Glycol Pit Trap; 1 female, CALIF. Riverside Co. Painted Canyon IV- 15 - 1979, Fred G. Andrews & M. S. Wasbauer colls.; 1 female, CALIF. Riverside Co. Painted Canyon, el. 500 9 Sand Wash-Larrea-Palo Verde, IX- 13 - 78 to I- 7 - 79, 530 Ethylene glycol can trap, R. Aalbu collr.; 1 female, Painted Canyon, Cal. April 8, 25, Timberlake coll. (b) Eschatomoxys wagneri det. Blaisd.?; 3 males, 1 female, CALIF. Riverside Co., Box Cyn, 10 mi. E. Mecca, VIII- 9 / 10 - 86 pit trap, Rolf L. Aalbu Coll.; 1 female, CALIF. Imperial Co. 3.9 mi. N. Walters Camp, IV- 27 - 70 to VII- 19 - 1978, Hardy & Andrews.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE6FF9DB970FEF1FEF524CD.taxon	etymology	Etymology. The specific epithet honors Fred G. Andrews, the collector of the holotype and many of the paratype specimens.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE6FF9DB970FEF1FEF524CD.taxon	diagnosis	Diagnosis. Similar to E. wagneri but with a complete lateral bead on the side of the pronotum.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE5FF9BB922FF31FD2020F1.taxon	description	(Fig. 7) Form elongate, fiddle-shaped, apterous. Color uniformly castaneous to piceous, shining. Males: length, 8.5 – 10.9 mm; width 4.0 – 4.8 mm. Females: length, 9.0 – 11.1 mm; width, 3.9 – 5.1 mm. Head. Prognathous, slightly deflexed, dorso-ventrally compressed; subquadrate in dorsal view, cranium slightly wider (1.7 – 2.4 mm) than long (1.4 – 2.0 mm), greatest width across epistoma. Lateral epistomal margins rounded and converging toward eyes. Median epistomal lobe produced, angularly arcuate, margin denticulate. Epistomal sutures obsolescent. Supraorbital carina prominent anteriorly, beginning at middle of lateral epistomal lobe and fading above middle of eye. Cranial surface dense with shallow punctures on dorsum, each separated from one another by about one diameter of a puncture on disc, becoming denser to contiguous on sides of head at base, degenerating to wrinkles on venter. Eyes small, oval, not bulging, anteriorly indented to accommodate epistomal canthus; facets in nine horizontal rows, median rows longest with 7 – 8 facets. Posterior margin of eye separated from cervix by 528 distance equal to about twice the diameter of eye. Labrum retractile, apex broadly emarginated, emargination bristling with marginal setae. Mandibles thick, robust, left side dominant (overlapping right), apex bifid; superior margin with stout angular cusp, somewhat larger on right mandible than left. Maxillary palpus approximately equal in length to first two antennomeres, basal segment longest (apical segment more elongate, slender in male, subsecuriform in female). Mentum hexagonal, surface coarsely punctate. Submentum with large ovoid fossa at middle. Gula weak to obsolescent, surface strigose. Tentorial pits weak to obsolete. Antennae filiform, antennomeres slender except basal segment broadened apically and last segment spindle-shaped, acuminate. Third segment longest; fourth to seventh subequal; eighth to tenth successively shorter until penultimate segment, which is about twice as long as wide; last segment about 1 K length of penultimate. Last two and apical third of the third to last segment set with short pale pubescence imparting a contrasting golden-yellow color to end of antennae. Thorax. Pronotum cylindrical, shorter than head, wider (2.0 – 2.6 mm) than long (1.5 – 1.9 mm), widest at middle; surface set with shallow punctures dorsally, the latter becoming less defined and more widely spaced laterally, inconspicuous ventrally. Complete lateral margin present. Anterolateral angle produced forward as an acute, angular cusp. Base of pronotum constricted with thick, marginal bead demarcating posterior edge. Femora long, clavate; tibiae and tarsi long, slender. Elytra inflated, embracing abdominal venter; surface glabrous, obscurely punctate at base. Epipleura widest basally, about as wide as metepisternum, gradually narrowing to about third abdominal segment, parallel thereafter, apex about half width at base. Abdomen. Ovoid in form, widest at middle (3.9 – 5.1 mm), length 5.9 – 7.6 mm, base slightly wider than apposing pronotal base. Surface of abdominal sternites smooth, impunctate. Length of first abdominal sternite from metacoxa to posterior margin subequal to length of second or slightly shorter; third about twothirds length of second; fourth about half length of third; fifth about twice length of fourth at midline. Males with elongate, ovoid, pubescent fovea at middle of first abdominal sternite. Genitalia as in E. pholeter.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE5FF9BB922FF31FD2020F1.taxon	materials_examined	Holotype. Male, MEXICO: Baja Calif. Isla Calaveras, V- 27 - 99, Oatmeal / Peanut butter pitfall trap, F. Sanchez-Piñero col. Paratypes: 2 males, 4 females, labeled same data as holotype; 2 males, 9 females, MEXICO, Baja Calif., Isla Calavera, VII- 17 - 97, F. Sanchez-Piñero col. 1 female, MEXICO, Baja Calif. Isla Mitlan, VII- 18 - 97, F. Sanchez-Piñero col.; 1 male, MEXICO, Baja Calif. Isla Mitlan PCPT VI- 25 / 27 - 1999 A. Boulton # D 00750 - 1502; 1 male, 1 female, MEXICO, Baja Calif., Isla Bota, VI- 23 - 98, F. Sanchez-Piñero col.; 1 male, MEXICO, Baja Calif. Isla Smith PGPT V- 13 - 14 / VII- 8 - 99 Area H: arroyo near sand area F. Piñero # D 00750 - 1502; 1 male, MEXICO, Baja Calif. Isla Ventana VII- 17 - 97, F. Sanchez-Piñero col.; 3 females, MEX: Gulf of Baja Calif. Pond Is. V- 26 - 1962, R. & A. Ryckman, C. Christianson collectors.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE5FF9BB922FF31FD2020F1.taxon	etymology	Etymology. The specific epithet is a noun in apposition and honors our colleague Francisco ‘‘ Paco’ ’ Sanchez-Piñero, who collected most of the specimens.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE5FF9BB922FF31FD2020F1.taxon	diagnosis	Diagnosis. The acutely produced anterior angles of the pronotum and the complete lateral bead on the thorax distinguish this species.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFEBFF98B95FF8EAFC9B212A.taxon	description	(Figs. 1, 5, 8 A, B) Form elongate, fiddle-shaped (Fig. 1), apterous. Color uniformly castaneous to piceous, shining (in life with a bluish, waxy bloom). Males: length, 9.3 – 11.3 mm; width, 4.1 – 5.1 mm. Females: length, 10.1 – 11.7 mm; width, 4.3 – 5.1 mm. Head. Prognathous, slightly deflexed, dorso-ventrally compressed; subquadrate in dorsal view, cranium about equal length (1.8 – 2.2 mm) and width (1.9 – 2.3 mm), greatest width across epistoma. Lateral epistomal margins rounded, converging toward eyes. Median epistomal lobe produced, arcuate, margin serrulate, weakly, broadly emarginated on each side at epistomal suture. Epistomal sutures obsolescent. Supraorbital carina feeble, short, beginning at middle of lateral epistomal lobe, fading above middle of eye; cuticle behind eye smooth. Cranial surface set with shallow punctures on dorsum separated from one another by about two diameters of a puncture, becoming denser to contiguous on sides of head at base, degenerating to wrinkles on venter. Eyes small, oval, anteriorly emarginated to accommodate epistomal canthus; facets in nine horizontal rows, median row longest with eight facets. Posterior margin of eye separated from cervix by distance equal to two and one-half to three times diameter of eye. Labrum retractile, apex broadly emarginated, emargination bristling with marginal setae. Mandibles thick, robust, left side dominant (overlapping right), apex bifid; superior margin with stout angular cusp, somewhat larger on right mandible than left. Maxillary palpus approximately equal in length to first two antennomeres, basal segment longest (apical segment more elongate, slender in 526 male, subsecuriform in female). Mentum hexagonal, surface coarsely punctate. Submentum with large ovoid fossa at middle. Gula rectangular, surface shagreened. Tentorial pits distinct, each connected to cervix by thin sulcus demarcating gula. Antennae filiform, antennomeres slender except basal segment broadened apically and last segment pyriform, acuminate. Third segment longest; fourth to seventh subequal; eighth, ninth and tenth successively shorter until penultimate segment, which is about twice as long as wide; last segment spindle shaped, 1 K times length of penultimate. Thorax. Pronotum cylindrical, shorter than head, wider (2.1 – 2.5 mm) than long (1.5 – 1.9 mm), widest at middle; surface set with shallow punctures dorsally, the latter becoming less defined and more widely spaced laterally, inconspicuous ventrally. Weak lateral margin present in apical fourth behind anterolateral angle, which is obtuse. Base of pronotum constricted with thick, marginal bead demarcating posterior edge. Femora long, clavate; tibiae. tarsi long, slender. Elytra inflated, embracing abdominal venter; surface glabrous, obscurely punctate at base; base slightly wider than apposing pronotal base. Epipleura widest basally, about as wide as metepisternum, gradually narrowing to about third abdominal segment, parallel thereafter, apex about half width at base. Abdomen. Ovoid in form, widest at middle (4.1 – 5.1 mm), longer than wide (6.0 – 7.7 mm). Surface of abdominal sternites smooth, impunctate. Length of first abdominal sternite from metacoxa to posterior margin about three-quarters length of second; third about two-thirds length of second; fourth about half width of third; fifth about 1 K length of fourth. Males with elongate, ovoid, pubescent fovea at middle of first abdominal sternite. Genitalia. Male aedeagus 3.2 mm long (Fig. 8 A); tegmen elongate, spatulate, cleft at apex. Median lobe (penis) extrusible, narrow, elongate, a simple flattened rod acute at apex, its ventral surface narrowly sulcate. Female ovipositor 2.2 mm long (Fig. 8 B); sclerotized dorsally, open ventrally. Paraproct slightly longer and narrower than coxite; bacula (valvifers) prominent, thick, flattened, sclerotized rods; basicoxites wider and shorter than terminal, triangular gonocoxites; apex of each gonocoxite with digitoid lobe on each side, bearing a small seta-like stylus. Vulva terminating in a pair of digitoid lobes, each ental to and subequal in size to terminal lobe of gonocoxite.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFEBFF98B95FF8EAFC9B212A.taxon	materials_examined	Holotype. Male. ARIZONA: Mohave Co., Grand Canyon N. P., Bat Cave, 29 - III- 96, R. B. Pape coll. Allotype. Female. Labeled same data as holotype. Paratypes: Three males, one female, labeled same data as holotype; Four males, six females: AZ. Mohave Co., Bat Cave, Gr. Canyon, GRCA Cave SCI- 0043, 26 - X- 01, R. B. Pape (deposited CAS and DB Thomas collection). One female: AZ. Mohave Co., Eldel Cave, twilight, IX- 23 - 05, J. Judson Wynne col. Three females: ARIZONA: Mohave Co., Grand Canyon N. P., Rampart Cave, teneb on 7,200 yr old sloth shit.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFEBFF98B95FF8EAFC9B212A.taxon	etymology	Etymology. The specific epithet pholeter (Greek) meaning, one who lurks in a hole.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFEBFF98B95FF8EAFC9B212A.taxon	diagnosis	Diagnosis. The new species averages larger in length than the other U. S. species. The smaller eyes differentiate this species from E. wagneri. It differs from E. tanneri and E. wagneri in the configuration of the gula. Presumably the longer head requires strengthening of the tentorial endoskeleton such that the tentorial pits are pronounced and the gula well demarcated by a narrow sulcus on each side, its surface contrastingly smooth compared to the rugose surface lateral to the sulci; whereas in E. tanneri and E. wagneri, with proportionately shorter heads, the gula is obsolete with neither tentorial pits or sulci prominent, and the rugosity of the venter is irregularly continuous towards the midline. The presence of this species in three separate but nearby caves suggests that this species is a troglophile and likely occurs in crevices rich in organic matter such as rodent nests in this surrounding area (see discussion of biology below).	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE0FF9CB94CFA8BFD1427B9.taxon	description	(Fig. 6) Form elongate, fiddle-shaped, apterous. Color uniformly castaneous to piceous, shining. Length: 10.1 – 10.7 mm, width: 4.6 – 4.8 mm. Head. Prognathous, slightly deflexed, dorso-ventrally compressed; subquadrate in dorsal view, cranium somewhat wider (2.0 – 2.3 mm) than long (1.7 – 2.0 mm), greatest width across epistoma. Lateral epistomal margins rounded and converging toward eyes. Median epistomal lobe produced, arcuate, margin feebly serrulate. Epistomal sutures obsolescent. Supraorbital carina feeble and short, beginning at middle of lateral epistomal lobe and fading above middle of eye; oval area of cuticle immediately behind eye impunctate. Cranial surface set with shallow punctures on dorsum separated from one another by about two diameters of a puncture, becoming denser to contiguous on sides of head at base; coarsest punctures on cranium posterior to mandibular insertion. Eyes small, oval, not bulging, anteriorly indented to accommodate epistomal canthus; facets in eight horizontal rows, median row longest with seven facets. Posterior margin of eye separated from cervix by distance equal to twice diameter of eye. Labrum retractile, apex broadly emarginated, emargination bristling with marginal setae. Mandibles thick, robust, left side dominant (overlapping right), apex bifid; superior margin with stout angular cusp, somewhat larger on right mandible than left. Maxillary palpus approximately equal in length to first two antennomeres, basal segment longest (apical segment more elongate, slender in male, subsecuriform in female). Mentum roundly hexagonal, surface rugosely punctate. Submentum with large ovoid fossa at middle. Gula ill-defined; tentorial pits obsolescent. Antennae filiform, antennomeres slender except basal segment broadened apically and last segment spindle-shaped, acuminate; third segment longest; fourth to seventh subequal; eighth to tenth successively shorter until penultimate segment, which is about twice as long as wide; last segment about 1 K times length of penultimate. Last two segments, and apical third of third to last segment, clothed in short golden setae. Thorax. Pronotum cylindrical, shorter than head, wider (2.5 – 2.6 mm) than long (1.7 – 1.9 mm), widest at middle; surface set with shallow punctures anterodorsally, becoming larger and more closely set postero-laterally, deeper but more widely spaced ventrally. Weak lateral margin present from acute antero-lateral angle to about mid-thorax. Base of pronotum constricted with thick, marginal bead demarcating posterior edge. Elytra inflated, embracing abdominal venter; surface glabrous, obscurely punctate at base. Epipleura widest basally, about as wide as metepisternum, gradually narrowing to about third abdominal segment, parallel thereafter, apex about half width at base. Femora long, clavate; tibiae and tarsi long, slender. Abdomen. Ovoid in form, widest at middle (4.6 – 4.8 mm), length: 6.7 – 6.8 mm, base slightly wider than apposing pronotal base. Sternal surface smooth, glabrous. Length of second sternite subequal to length of first sternite from coxae to posterior margin; third sternite about two-thirds length of second; fourth about half length of third; fifth about twice length of fourth at midline. Males with elongate, ovoid, pubescent fovea at middle of first abdominal sternite. Genitalia as described for E. pholeter.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE0FF9CB94CFA8BFD1427B9.taxon	materials_examined	Holotype. Female; MEXICO: Baja Calif. Isla Cabeza PGP V- 15 / VII- 5 - 99. Area B. SW slope near light. F. Piñero # D 00750 - 1502. Paratypes: 1 Male: MEXICO: Baja Calif. Isla San Lorenzo Norte, VII- 26 - 2001, Carrion trap, F. Sanchez-Piñero col. 1 Female: MEXICO: Baja Calif. San Lorenzo Is. 4 - vii- 1996, F. S. Piniero. [OSU]. 1 Female: MEXICO, B. Cal. Isla San Esteban, IV- 3 - 1953, Col. By J. P. Figg Hobiya. [CAS] 2 Females, 1 male: MEXICO: Baja Calif. Bahia de Los Angeles, Vermilion Sea Field Station, VII- 25 - 2005 F. Sánchez Piñero & D. Talley cols.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE0FF9CB94CFA8BFD1427B9.taxon	etymology	Etymology. The species is named to honor Mike Rose, a dedicated, enthusiastic scientist, who among others, lost his life in the tragic sinking of their research vessel near Isla Cabeza in March 2000.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE0FF9CB94CFA8BFD1427B9.taxon	diagnosis	Diagnosis. The anterior angles of the pronotum are not produced as in E. paco, and the lateral bead of the pronotum is incomplete.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE1FF9FB933FE28FF252484.taxon	diagnosis	Diagnosis. Form elongate, fiddle-shaped, apterous, with long legs and antennae. Color uniformly castaneous to piceous, shining. Head subquadrate in form, length about equal to width; dorso-ventrally compressed. Eyes medium size, posterior margin of eye separated from cervix by at least two diameters of eye. Although Sorenson & Stones (1959) cited differences in the proportions of 532 the antennal segments between this species and E. wagneri we did not find these slight differences to be consistent. Length of body: 9.3 – 10.0 mm; width: 4.0 – 4.1 mm. Type locality: Padre Creek, Kane County, Utah. Holotype and paratypes at BYU were examined. Additional Material Examined: 1 male, UTAH: San Juan Co., 1 mi. N, 17 mi. W. Bluff, 5600 9 1 X- 13 - 1985, D. Giuliani [CDFA]. 1 female, UTAH: Garfield Co., Calf Ck Cmpg N of Escalante, 2 – 3 Aug 1984, Nelson Baumann [BYUC]. 1 female, UTAH: Garfield Co. Calf Creek Cmpg, 1 August 1985, R. W. Baumann [BYUC]. 1 female, ARIZONA, Marble Canyon, 4 Sept 1950, D. R. Estes Collector.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE1FF9CB9F5F85FFEC8203F.taxon	description	(Figs. 2 – 4)	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE1FF9CB9F5F85FFEC8203F.taxon	diagnosis	Diagnosis. Form elongate, fiddle-shaped, apterous, with long legs and long antennae. Color uniformly castaneous to piceous. Head subrectangular in form, wider than long. Eyes large, posterior margin of eye separated from anterior margin of pronotum by less than two diameters of the eye. Tentorial pits obsolete. Thorax orbicular, wider than long; greatest width slightly before middle. Lateral pronotal margin effaced at middle, anterolateral pronotal angle obtuse, feebly produced. Length of body: 8.4 – 9.6 mm; width: 3.6 – 4.2 mm.	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
03F6F264FFE1FF9CB9F5F85FFEC8203F.taxon	materials_examined	Type Locality. Death Valley, Inyo County, California (holotype, CAS, examined). Material Examined. 1 male, 1 female: Furnace Creek, Death Valley, Calif. 2 - 17 - 54, FB Turner Coll. [CAS] 1 male: CALIF. S. Bern. Co. near Barstow, 3 - VI- 1975, J. & F. Enik, under stones & debris [CAS]. 1 male, 1 female: Mine, 160 ft. level, Laguna Dam, Calif. May 20, 1940, Glen M. Kohls Coll. (b) wagneri det. by F. E. Blaisdell [CAS]. 1 male: California, Inyo Co. Funeral Mountains, Pyramid Peak, 17 January 1962, C. Jorgensen [BYUC]. 1 male: USA, California, San Bernardino Co. Saratoga Springs, 10 June 1959, Ernest R. Tinkham. (b) Eschatomoxys sp. Det. J. Doyen. 1 female: Salt Creek, Death Valley, Calif. Mar. 27 – Ap. 3, 1949. (b) Eschatomoxys wagneri Blaisd., det. J. Doyen 64 [CAS]. 1 female: Moencopie, Ariz. Moencopie Wash, Coconino Co., VII- 24 - 52. M. Cazier, R. Schrammel coll. [AMNH].	en	López-Guerrero, Irma (2007): Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits. The Coleopterists Bulletin 61 (4): 519-540, DOI: 10.1649/0010, URL: http://dx.doi.org/10.5281/zenodo.5326738
