taxonID	type	description	language	source
03F787804302FFFAF0A2FD54FA43CB61.taxon	description	(Figs 1 B, 2, 3; Tabs III-V)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	materials_examined	Material examined Syntypes. – MNHN-IC- 1932 - 0108 from Madagascar (1 male and 2 females). Others. – Fifteen males and five females collected from Seychelles and Madagascar with a size range of 29.3 - 64 mm SL. MNHN-IC- 2007 - 0204, 2 specimens, Grande Anse, Mahé, Seychelles Islands, 20 Nov. 2004, Accouche et al. coll.; tags M- 0204 A & M- 0204 B. MNHN-IC- 2020 - 0043, Silhouette, Anse Mondon, Seychelles Islands, 20 Nov. 2004, Accouche et al. coll.; tag 13970. MNHN-IC- 2007 - 0200, 2 specimens (tags A, B) of 3, Mahé, Seychelles Islands, 22 Nov. 2004, Accouche et al. coll. MNHN-IC- 2020 - 0049, Andrepona, Madagascar, 04 Jul. 08, Diamsoi coll.; tag 7546. MNHN-IC- 2020 - 0050, same data as 2020 - 0049, tag 7547. MNHN-IC- 2020 - 0051, same data as 2020 - 0049, tag 7548. MNHN-IC- 2020 - 0052, Ambanizana, Madagascar, May 2010, Diamsoi coll.; tag 13015. MNHN-IC- 2020 - 0053, Farambana, Madagascar, July 2008, Diamsoi coll.; tag 13007. MNHN-IC- 2020 - 0054, Sahana, Madagascar, 07 Jul. 2008; Diamsoi coll.; tag 12947. MNHN-IC- 2020 - 0055, same data as 2020 - 0053; tag 13001. MNHN-IC- 1962 - 0172, 3 specimens, Madagascar, Lampahana, Kiener coll. MNHN-IC- 2020 - 0056, Fond B’Offay, Praslin, Seychelles Islands, 22 Feb. 2020, Keith, Mennesson, Henriette et al., coll.; tag 12728. MNHN-IC- 2020 - 0057, same data as 2020 - 0056, tag 12730. MNHN-IC- 2020 - 0058, Grand St Louis River, Mahé, Seychelles Islands, 15 Feb. 2020, Keith, Mennesson, Henriette et al., coll.; tag 12593.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	diagnosis	Diagnosis The species is distinguished by: second and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.4 ’ pattern), row os not connected with row oi at ventro-posterior margin of operculum (‘ open’ pattern); 15 pectoral fin rays; no teeth on vomer; scales in zigzag series 10 - 12, and 18 - 24 in transverse forward series.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	description	Description Scale and ray counts in Eleotris pellegrini and related species are given in tables III-IV, morphometrics in table V. The body is elongated but stocky. The body depth at anus is 17 - 24 (% SL) and the caudal peduncle depth is 10 - 17 (% SL). Predorsal length 42 - 49 (% SL) and preanal length 62 - 70 (% SL). Size: up to 64 mm SL. Head (32 - 37 % SL) slightly depressed, the snout is pointed. Eyes high on head. The mouth is large, as jaw length is 9 - 14 % SL, and oblique, the posterior margin of upper jaw reaches the vertical through the half of the eye. Upper and lower jaws with multiple rows of small inwardly curved teeth, larger on outer row; a few caniniform teeth in posterior position. Dorsal fins mostly VI-I, 8 - 9; D 1 separate from and same height as D 2; spines not elongated. Anal fin I, 8 - 9 and directly opposite to second dorsal fin. Pelvic fins separate, I, 5. Pectoral fins 15. Caudal fin rounded with 15 branched rays. Cycloid scales on top of head, nape, cheek, operculum, pectoral fin base, prepelvic region, and abdomen. Ctenoid scales covering flanks. No lateral line canals. 48 - 56 scales in lateral series, 28 - 37 in predorsal series, 14 - 16 in transverse back series, 18 - 24 in transverse forward series and 10 - 12 in zigzag series. A downward pointing spine at the posterior margin of the preoperculum is covered with skin. Gill opening is moderately broad, extending to below the preoperculum. Tubular anterior nares overhanging half upper lip, posterior nares open pits. Cephalic lateralis: Adults with five transverse suborbital free neuromast rows of which second and fourth suborbital free neuromast rows on cheek extend ventrally past horizontal row d (‘ 2.4 ’ pattern); row os not connected with row oi at ventro-posterior margin of operculum (‘ open pattern’) (Fig. 1 B). Urogenital papilla in females rounded, elongated and tapered in males. Colour in life (Fig. 3) Male and female similar. Body marbled from light to dark brown, more beige on the back and striped and spotty on the flanks, alternating dark brown and beige. Three horizontal dark brown to reddish stripes between eyes and preoperculum and three vertical white ones below the eye. First dorsal fin translucent with two large horizontal dark spotted stripes, one at fin base, one in the middle of the fin. Second dorsal and anal fins with 5 - 6 brownish wavy spotted horizontal stripes. Pelvic fins base yellowish. Caudal fin brown, with two brown spots anteriorly, several brownish vertical spotted wavy rows. Pectoral fins translucent. Colour in preservation Male and female similar. Body marbled brown, lighter on the back and striped on the flanks. Three horizontal dark brown stripes between eyes and preoperculum. First dorsal fin with two large horizontal dark stripes, one at fin base, one in the middle of the fin. Second dorsal and anal fins with 5 - 6 brownish wavy spotted horizontal stripes. Caudal fin brown, with two brown spots anteriorly, several brownish vertical wavy spotted stripes. Pectoral fins greyish.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	diagnosis	Comparison Eleotris pellegrini differs from the other species sequenced that occur in the area studied by having a high % of divergence in COI gene (9.4 - 15.2 %). Within the pattern ‘ 2.4 ’ group, E. pellegrini differs from E. acanthopoma in having 15 pectoral fin rays versus 16 - 17 and 28 - 34 scales in predorsal series versus 30 - 42. It differs from E. vomerodentata by the absence of teeth in the vomer versus presence. It differs from E. sahanaensis by a greater head length (31 - 37 versus 30 - 32 % SL) and more scales in zigzag series (10 - 12 versus 10).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	distribution	Distribution Currently known only from Madagascar and the Seychelles Islands. It was found in Seychelles in sympatry with E. acanthopoma and in Madagascar with E. sahanaensis (this paper).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804302FFFAF0A2FD54FA43CB61.taxon	biology_ecology	Ecology The species is supposed to be mainly amphidromous (Mennesson and Keith, 2017). It lives in the lower part of the river. It lives in muddy to clear rivers with sandy to gravel bottom between 5 to 10 m in altitude. It is carnivorous.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	description	(Figs 1 B, 2, 4; Tabs III-V)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	materials_examined	Material examined Holotype. – MNHN-IC- 1984 - 0803 from Madagascar (male).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	diagnosis	Diagnosis The species is distinguished by: second and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.4 ’ pattern), row os not connected with row oi at ventro-posterior margin of operculum (‘ open’ pattern); 15 pectoral fin rays and the presence of teeth in the vomer.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	description	Description Scale and ray counts in Eleotris vomerodentata and related species are given in tables III-IV, morphometrics in table V. The body is elongated. The body depth at anus is 21 (% SL) and the caudal peduncle depth is 13 (% SL). (% SL). Comparison Within the pattern ‘ 2.4 ’ Eleotris vomerodentata differs from all other species by the presence of teeth in the vomer versus absence.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	distribution	Distribution Currently known only from Madagascar.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	biology_ecology	Ecology Unknown.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	description	Description of the new species	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430DFFF8F0C1FC1EFB9BC941.taxon	materials_examined	Comparative material Adults with five transverse suborbital free neuromast rows of which second and fourth suborbital free neuromast rows on cheek extend ventrally past horizontal row d (‘ 2.4 ’ pattern); row os not connected with row oi at ventro-posterior margin of operculum (‘ open pattern’). Eleotris acanthopoma: MNHN-IC- 2006 - 0617, 3 specimens, Mohéli, Mdjawaché, Comoros Islands, 31 Oct. 2005, Keith et al. coll. MNHN-IC- 2020 - 0029, Mayotte, Kwale, Comoros Islands, 22 Apr. 2009, Feutry coll.; tag 12426. MNHN-IC- 2020 - 0030, Comoros Islands, Mayotte, Apr. 2009, Feutry coll.; tag 12417. MNHN-IC- 2020 - 0031, Mayotte, Ouroveni, Comoros Islands, 06 Sep. 2007, Marquet coll.; tag 13951. MNHN-IC- 2020 - 0032, same data as 2020 - 0031, 06 Sep. 2007, Marquet coll.; tag 13952. MNHN-IC- 2020 - 0033, same data as 2020 - 0031; tag 13953. MNHN-IC- 2020 - 0034, same data as 2020 - 0031; tag 13954. MNHN-IC- 2020 - 0041, Mayotte, Ouroveni, Comoros Islands, 06 Sep. 2007, Marquet coll.; tag 11817. MNHN-IC- 2020 - 0035, Mayotte, Coconi, Comoros Islands, 06 Aug. 19, Valade et al. coll.; tag 14519. MNHN-IC- 2020 - 0036, same data as 2020 - 0035, tag 14520. MNHN-IC- 2020 - 0037, same data as 2020 - 0035, tag 14521. MNHN-IC- 2020 - 0038, Comoros Islands, Mayotte, Ouroveni, 06 Aug. 2019, Valade et al. coll.; tag 14562. MNHN-IC- 2020 - 0039, Comoros Islands, Mayotte, Longoni, 07 Aug. 2019, Valade et al. coll.; tag 14563. MNHN-IC- 2020 - 0040, same data as 2020 - 0039, tag 14564. MNHN-IC- 2007 - 0185, 2 specimens, Mahé, Seychelles Islands, 24 Nov. 2004, Accouche et al. coll. MNHN-IC- 2007 - 0199, Mahé, Anse aux poules bleues riv., Seychelles Islands, 14 Oct. 2003, Keith et al. coll. MNHN-IC- 2007 - 0205, Praslin, Fond B’Offay riv., Seychelles Islands, 10 Oct. 2003, Keith et al. coll.; tag 14509. MNHN-IC- 2020 - 0042, Praslin, Nouvelle découverte, Seychelles Islands, 09 Oct. 2003, Keith et al. coll.; tag 13749. MNHN-IC- 2007 - 0200, 1 (tag C) of 3, Mahé, Seychelles Islands, 22 Nov. 2004, Accouche et al. coll. Eleotris vomerodentata Maugé, 1984: holotype from Madagascar (MNHN-IC- 1984 - 0803). Eleotris pellegrini Maugé, 1984: syntypes from Madagascar (MNHN-IC- 1932 - 0108) and others (see above, E. pellegrini redescription).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	description	(Figs 1 B, 2, 5; Tabs III-V)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	materials_examined	Material examined One male and one juvenile collected from Madagascar with a size range of 26 - 44 mm SL. Holotype. – MNHN-IC- 2020 - 0078, male (44 mm SL), Sahana River, Madagascar; July 2008; Diamsoi coll.; tag 13003. Paratype. – MNHN-IC- 2020 - 0077, juv. (26 mm SL), Andrepona River, Madagascar; 4 Jul. 2008; Diamsoi coll.; tag 7278.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	diagnosis	Diagnosis The species is distinguished by: second and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.4 ’ pattern), row os not connected with row oi at ventro-posterior margin of operculum (‘ open’ pattern); 15 pectoral fin rays; no teeth on vomer; 54 - 55 scales in lateral series and 29 - 30 scales in predorsal series.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	description	Description Scale counts in Eleotris sahanaensis sp. nov. and related species are given in table III, ray counts in table IV and morphometrics in table V. Below, the holotype counts are given first followed in brackets, if different, by the paratype counts. The body is elongated. The body depth at anus is 22 (16 % SL) and the caudal peduncle depth is 12 (11 % SL). Predorsal length 47 (46 % SL) and preanal length 63 (61 % SL). The head 32 (30 % SL) is broad and depressed, the snout is pointed. Eyes high on head. The mouth is quite large, as jaw length 11 (9 % SL), and oblique, the posterior margin of upper jaw reaches the vertical through the two thirds of the eye. Upper and lower jaws with multiple rows of small inwardly curved teeth. Dorsal fins VI-I, 8 - 9; D 1 separate from and smaller than D 2; spines not elongated. Anal fin I, 8 - 9 and directly opposite to second dorsal fin. Pelvic fins separate, I, 5. Pectoral fins 15. Caudal fin pointed with 15 branched rays. Cycloid scales on top of head, nape, cheek, operculum, pectoral fin base, prepelvic region, and abdomen. Ctenoid scales covering flanks. No lateral line canals. 55 (54) scales in lateral series, 29 (30) in predorsal series, 15 (14) in transverse back series, 19 (20) in transverse forward series and 10 in zigzag series. A downward pointing spine at the posterior margin of the preoperculum is covered with skin. Gill opening is moderately broad, extending to below the preoperculum. Tubular anterior nares overhanging half upper lip, posterior nares open pits. Cephalic lateralis: Adults with five transverse suborbital free neuromast rows of which second and fourth suborbital free neuromast rows on cheek extend ventrally past horizontal row d (‘ 2.4 ’ pattern); row os not connected with row oi at ventro-posterior margin of operculum (‘ open pattern’) (Fig. 1 B). Urogenital papilla elongated and tapered in male. Colour in life Unknown. Colour in preservation (Fig. 5) Head and preoperculum greyish, finely spotted and with 3 - 4 oblique brown lines from the eye to the edge of operculum. Back brown, flanks light brown with numerous dots more or less aligned from the operculum to the hypural. A big blackish area at the anterior part of the flanks, below the medial part. Abdomen and gular region whitish to yellowish and finely spotted. A black dot at the base of the pectoral fins. Anal and second dorsal fins with 3 - 4 dark horizontal bands alternating with 3 small white ones. Caudal fin greyish, with 2 - 3 small spots anteriorly. Pelvic fins whitish. Comparison Eleotris sahanaensis differs from the other species sequenced that occur in the area studied by having a high % of divergence in COI gene [79 bp alignment] (3.8 - 10.1 %). Within the pattern ‘ 2.4 ’, E. sahanaensis differs from E. acanthopoma in having 15 pectoral fin rays versus 16 - 17, 10 scales in zigzag series versus 11 - 14, and 29 - 30 scales in predorsal series versus 30 - 42. It differs from E. vomerodentata by the absence of teeth in the vomer versus presence. It differs from E. pellegrini by a smaller head length (30 - 32 versus 31 - 37 % SL) and fewer scales in zigzag series (10 versus 10 - 12).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	distribution	Distribution Currently known only from Madagascar. It was found in sympatry with E. pellegrini and E. diamsoi (this paper).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	biology_ecology	Ecology Unknown.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430FFFFFF03AFE3EFA91CA61.taxon	etymology	Etymology The new species is named after the Sahana River, type locality, where many species of Eleotris were found.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	description	(Figs 1 D, 2, 6; Tabs VI-VIII)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	materials_examined	Material examined Syntypes: BMNH 1864.11. 15.127 (1), 1865.3.18.26 - 27 (2), 1867.3.9.521 (1) from Mozambique.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	diagnosis	Diagnosis The species is distinguished by: second, third and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.3.4 ’ pattern), row os connected with row oi at ventro-posterior margin of operculum (‘ closed’ pattern); 17 - 18 pectoral fin rays; 50 - 56 scales in lateral series, and 12 - 13 scales in zigzag series.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	description	Description Scale and ray counts in Eleotris soaresi and related species are given in tables VI-VII and morphomeristics in table VIII. The body is elongated. The body depth at anus is 20 - 21 (% SL) and the caudal peduncle depth is 15 (% SL). Predorsal length 47 (% SL) and preanal length 65 - 69 (% SL). Size: up to 100 mm SL. The head (33 - 37 % SL) is broad and depressed, the snout is pointed. Eyes high on head. The mouth is large, as the jaw length is 11 - 13 % SL, and oblique, the posterior margin of upper jaw reaches the vertical through the two thirds of the eye. Upper and lower jaws with multiple rows of small inwardly curved teeth. Dorsal fins VI-I, 8 - 9; D 1 separate from and same height as D 2; spines not elongated. Anal fin I, 8 - 9 and directly opposite to second dorsal fin. Pelvic fins separate, I, 5. Pectoral fins 17 - 18. Caudal fin rounded with 15 branched rays. Cycloid scales on top of head, nape, cheek, operculum, pectoral fin base, prepelvic region, and abdomen. Ctenoid scales covering flanks. No lateral line canals. 50 - 56 scales in lateral series; 39 - 43 in predorsal series, 15 - 18 in transverse back series, 22 in transverse forward series and 12 - 13 in zigzag series. A downward pointing spine at the posterior margin of the preoperculum is covered with skin. Gill opening is moderately broad, extending to below the preoperculum. Cephalic lateralis: Adults with five transverse suborbital free neuromast rows of which second, third and fourth suborbital free neuromast rows on cheek extend ventrally past horizontal row d (‘ 2.3.4 ’ pattern); row os connected with row oi at ventro-posterior margin of operculum (‘ closed pattern’) (Fig. 1 C). Urogenital papilla elongated and tapered in males. Colour in life (Fig. 6) According to Playfair (in Playfair and Günther, 1867) ‘ body dark brown, dorsal and anal punctulated with black’. Colour in preservation Head, preoperculum, body, abdomen and fins dark brown. Gular and isthmus beige or dark brown depending on the specimen.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	diagnosis	Comparison E. soaresi differs from the other species of Indian Ocean with a ‘ 2.3.4 ’ pattern in having fewer scales in lateral series (50 - 56 versus 59 - 64) and in zigzag series (12 - 13 versus 13 - 15). Moreover, it differs from E. valadei n. sp., this paper, in having fewer scales in transverse forward series (22 - 24 versus 27 - 30) and predorsal series (39 - 43 versus 45 - 50), and from E. diamsoi in having 17 - 18 pectoral fin rays versus 16.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	distribution	Distribution Currently known only from Mozambique.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F787804308FFFDF0ABFD7EFC0BCA41.taxon	biology_ecology	Ecology Unknown.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	description	(Figs 1 D, 2, 7; Tabs VI-VIII)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	materials_examined	Material examined One male and one female collected from Madagascar with a size range of 61 - 64.6 mm SL. Holotype. – MNHN-IC- 2020 - 0068, male (64.6 mm SL), Sahana River, Madagascar; 7 July 2008; Diamsoi coll.; tag 13004. Paratype. – MNHN-IC- 2020 - 0067, female (61 mm SL), Ankazofotsy River, Madagascar; May 2010; Diamsoi coll.; tag 13016.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	diagnosis	Diagnosis The species is distinguished by: second, third and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.3.4 ’ pattern) (Fig. 1 D); row os connected with row oi at ventro-posterior margin of operculum (‘ closed’ pattern); 16 pectoral fin rays; 63 scales in lateral series; a small body depth at anal and second dorsal fins origin (15 - 18 % SL).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	description	Description Scale counts in Eleotris diamsoi sp. nov. and related species are given in table VI, ray counts in table VII and morphometrics in table VIII. Below, the holotype counts are given first followed in brackets, if different, by the paratype counts. The body is elongated. The body depth at anus is 15 (18 % SL) and the caudal peduncle depth is 12 (14 % SL). Predorsal length 44 (47 % SL) and preanal length 63 (66 % SL). The head 32 (33 % SL) is broad and depressed, the snout is pointed. Eyes high on head. The mouth is quite large, as jaw length is 10 (11 % SL), and oblique, the posterior margin of upper jaw reaches the vertical through the two thirds of the eye. Upper and lower jaws with multiple rows of small inwardly curved teeth, larger on outer row; a few caniniform teeth in anterior position on lower jaw. Dorsal fins VI-I, 8; D 1 separate from and smaller than D 2; spines not elongated. Anal fin I, 8 and directly opposite to second dorsal fin. Pelvic fins separate, I, 5. Pectoral fins 16. Caudal fin pointed with 15 branched rays. Cycloid scales on top of head, nape, cheek, operculum, pectoral fin base, prepelvic region, and abdomen. Ctenoid scales covering flanks. No lateral line canals. 63 scales in lateral series, 36 (38) in predorsal series, 18 in transverse back series, 17 (23) in transverse forward series and 14 in zigzag series. A downward pointing spine at the posterior margin of the preoperculum is covered with skin. Gill opening is moderately broad, extending to below the preoperculum. Tubular anterior nares overhanging one third of upper lip, posterior nares open pits. Cephalic lateralis: Adults with five transverse suborbital free neuromast rows of which second, third and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.3.4 ’ pattern); row os connected with row oi at ventro-posterior margin of operculum (‘ closed’ pattern) (Fig. 1 D). Urogenital papilla in females rounded, elongated and tapered in males. Colour in life Unknown. Colour in preservation (Fig. 7) Male and female similar. Head and preoperculum beige. Back and flanks brownish. A big brown dot at the base of the pectoral fins. Abdomen and gular region yellowish. Anal and first and second dorsal fins spotted with spots aligned on rays. Caudal fin brownish, with several yellow spots. Pelvic and pectoral fins whitish.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	diagnosis	Comparison Eleotris diamsoi differs from the other species sequenced that occur in the area studied by having a high % of divergence in COI gene (9.9 - 14.9 %). Eleotris diamsoi n. sp. differs from the other species of the Indian Ocean with a ‘ 2.3.4 ’ pattern by several characters. It differs from E. soaresi in having 16 pectoral fin rays versus 17 - 18, more scales in lateral series (63 versus 50 - 56) and in zigzag series (14 versus 12 - 13), and a smaller body depth at anal fin origin (15 - 18 versus 20 - 21). It differs from E. valadei, this paper, in having 16 pectoral fin rays versus 18, fewer scales in predorsal series in males (36 - 38 versus 45 - 50), fewer scales in transverse forward series (17 - 23 versus 27 - 30) and a smaller body depth at anal fin origin (15 - 18 versus 19 - 24).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	distribution	Distribution Currently known only from Madagascar. It was found in sympatry with E. pellegrini and E. sahanaensis (this paper).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	biology_ecology	Ecology Unknown.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430AFFFCF05AFB5FFCBFCC4A.taxon	etymology	Etymology The new species is named after the MNHN ‘ Diamsoi’ program which enabled to collect the specimens in Madagascar.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	description	(Figs 1 D, 2, 8; Tabs VI-VIII)	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	materials_examined	Material examined Five males and two females collected from Mayotte, Praslin (Seychelles), Reunion Island and Madagascar, with a size range of 45 - 99.3 mm SL. Holotype. – MNHN-IC- 2020 - 0063, male (57 mm SL), Ouroveni River, Mayotte; 06 Aug. 2019; Valade coll.; tag 14561. Paratypes. – MNHN-IC- 2020 - 0060, male (60.6 mm SL), Ambodiforaha River, Madagascar; Jul. 2008; diamsoi coll.; tag 12950. MNHN-IC- 2020 - 0062, male (45 mm SL), Ouroveni River, Mayotte; 20 Apr. 2009; Feutry coll.; tag 11822. MNHN-IC- 2020 - 0059, male (46 mm SL), Mtchangachehi, Mayotte; 12 Dec. 2003; tag 12418. MNHN-IC- 2020 - 0061, female (48 mm SL), Ouroveni river, Mayotte; date 06 Aug. 2007; Marquet coll.; tag 11818. MNHN-IC- 2020 - 0064, female (79.3 mm SL), Reunion Island, Dec. 2007, Zimmermann et al. coll.; tag 12409. MNHN-IC- 2020 - 0065, male (99.3 mm SL), Nouvelle-Découverte River, Praslin, Seychelles; 21 Feb. 2020; Keith, Mennesson, Henriette et al. coll.; tag 12584. Non types. – MNHN-IC- 1966 - 0959, two males (105 - 110 mm), Anjanambo, Tamatave Province, Madagascar, Dec. 1962, Kiener & Therezien coll.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	diagnosis	Diagnosis The species is distinguished by: second, third and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.3.4 ’ pattern), row os connected with row oi at ventro-posterior margin of operculum (‘ closed’ pattern); 18 pectoral fin rays; 59 - 64 scales in lateral series and 27 - 30 scales in transverse forward series.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	description	Description Scale counts in Eleotris valadei sp. nov. and related species are given in table VI, ray counts in table VII and morphometrics in table VIII. Below, the holotype counts are given first followed in brackets, if different, by the paratype counts. The body is elongated. The body depth at anus is 21 (19 - 24 % SL) and the caudal peduncle depth is 14 (13 - 17 % SL). Predorsal length 45 (42 - 49 % SL) and preanal length 66 (67 - 70 % SL). The head 33 (33 - 39 % SL) is broad and depressed, the snout is pointed. Eyes high on head. The mouth is large, as the jaw length is 12 (11 - 15 % SL), and oblique, the posterior margin of upper jaw reaches the vertical through the two thirds of the eye. Upper and lower jaws with multiple rows of small inwardly curved teeth; larger on outer row on upper jaw and larger on inner jaw on lower jaw; a few small caniniform teeth in posterior position. Dorsal fins VI-I, 8 - 9; D 1 separate from and slightly higher than D 2; spines not elongated. Anal fin I, 8 - 9 and directly opposite to second dorsal fin. Pelvic fins separate, I, 5. Pectoral fins 18. Caudal fin pointed with 15 branched rays. Cycloid scales on top of head, nape, cheek, operculum, pectoral fin base, prepelvic region, and abdomen. Ctenoid scales covering flanks. No lateral line canals. Scales in lateral series 63 (59 - 64), in predorsal series 50 (45 - 49). Scales in transverse back series 21 (17 - 20), in transverse forward series 30 (27 - 30) and 13 (14 - 15) in zigzag series. A downward pointing spine at the posterior margin of the preoperculum is covered with skin. Gill opening is moderately broad, extending to below the preoperculum. Tubular anterior nares overhanging upper lip, posterior nares open pits. Cephalic lateralis: Adults with five transverse suborbital free neuromast rows of which second, third and fourth suborbital free neuromast rows on cheek extending ventrally past horizontal row d (‘ 2.3.4 ’ pattern); row os connected with row oi at ventro-posterior margin of operculum (‘ closed’ pattern) (Fig. 1 D). Urogenital papilla in females rounded, elongated and tapered in males. Colour in life (Fig. 8) Male and female similar. Top of head, top of the eye and back beige with brown freckles. Lateral part of head and body dark brown. Abdomen and gular region dark brown. Lips with dark brown with evenly spaced yellow spots. First dorsal fin with 2 - 3 large horizontal dark bands alternating with 2 white bands; rays with yellow to orange spotted wavy rows alternating with black. Second dorsal, anal and pelvic fins with yellow to orange spotted wavy rows alternating with black. Caudal fin dark brown, with in the superior part, at anterior position, a small beige and black striped area. Pectoral fins translucent grey, pectoral base dark brown. Colour in preservation Male and female similar. Head and body dark brown. Abdomen and gular region dark brown. First dorsal fin with 2 - 3 large horizontal dark bands alternating with 2 greyish bands; rays with greyish spots alternating with black wavy spotted rows. Second dorsal, anal and pelvic fins with greyish spots alternating with black wavy spotted rows. Caudal fin dark brown, with in the superior part, at anterior position, a small beige and black striped area. Pectoral fins greyish, pectoral base dark brown.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	diagnosis	Comparison Eleotris valadei differs from the other species sequenced that occur in the area studied by having a high % of divergence in COI gene (6.8 - 14.6 %). Eleotris valadei n. sp. differs from the other species of the Indian Ocean with a ‘ 2.3.4 ’ pattern by several characters. It differs from E. soaresi in having more scales in lateral series (59 - 64 versus 50 - 56), more scales in transverse forward series (27 - 30 versus 22 - 24), in predorsal series (45 - 50 versus 39 - 43) and in zigzag series (13 - 15 versus 12 - 13). It differs from E. diamsoi n. sp., this paper, in having 18 pectoral fin rays versus 16, more scales in transverse forward series (27 - 30 versus 17 - 23), in predorsal series (45 - 50 versus 36 - 38), and a higher body depth at dorsal fin origin (19 - 24 versus 15 - 18).	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	distribution	Distribution Currently known from Madagascar, Reunion Island, Mayotte and the Seychelles islands. It was found in Seychelles in sympatry with E. pellegrini and E. acanthopoma.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	biology_ecology	Ecology The species is supposed to be amphidromous. It is found in the lower part of the river, near the estuary. It lives in muddy to clear rivers with sandy to gravel bottom between 0 to 5 meters in altitude. It is carnivorous.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
03F78780430BFFE2F059FB37FE90CE7B.taxon	etymology	Etymology The name of the species honours Pierre Valade, who collected several specimens of this new species in Reunion Island and Mayotte.	en	Mennesson, Marion I., Keith, Philippe (2020): Eleotris (Teleostei: Eleotridae) of the Indian Ocean: an overview with the description of three new species. Cybium 44 (3): 185-203, DOI: 10.26028/cybium/2020-443-002, URL: http://dx.doi.org/10.26028/cybium/2020-443-002
