taxonID	type	description	language	source
03F787E6FF82992E91917311FE49FD88.taxon	description	When the second author (Hardy) first saw this Venezuelan specimen in the Henry Howden collection, he could not readily place it to genus. When the first author (Ratcliffe) received the specimen from Hardy for examination in mid- 2003, he was immediately taken with its gestalt similarity with the genus Coscinocephalus Prell that occurs in Arizona and Mexico. Unfortunately, there is only one specimen, but our examination revealed such unique character states that were indicative of a new, undescribed genus and species that we describe it here. We utilize the Phylogenetic Species Concept as outlined by Wheeler and Platnick (2000). This concept defines species as the smallest aggregation of sexual populations diagnosable by a unique combination of character states. Whether this species occurs rarely in nature remains unknown, but the area from which it was collected is readily accessible if someone wanted to make a concerted effort to find additional specimens. During a visit by one of us (Ratcliffe) to the extensive collections of the Universidad Central de Venezuela in Maracay in 1999, similar specimens were not found. With the new genus described here, the tribe Pentodontini now contains 26 genera and a few more than a hundred species in the New World. Endrödi (1969, 1985) comprehensively reviewed and defined the Pentodontini, and this tribal concept was used by Ratcliffe (2002) in the latest generic treatment of the North American dynastines. Endrödi’s definition did not address monophyly, and we remain concerned that the Pentodontini (as currently defined) may not be monophyletic. In particular, the head and pronotal armature as well as the form of the meso- and metatibiae used to characterize the tribe are not consistently expressed. Until such time as the monophyly of the Pentodontini is resolved, we place our new genus in the Pentodontini because it possesses the following characters: a weak tubercle on the frons, a strong frontoclypeal carina, tridentate foretibia, and a truncate metatibia with a fringe of short, spine-like setae on the apex. The last generic keys to adults of the New World Pentodontini were by Ratcliffe (1981) and Endrödi (1985). Morón and Ratcliffe (1997) transferred the genus Coscinocephalus from the Cyclocephalini to the Pentodontini, but it has never been incorporated into a comprehensive generic key. In addition, the genus Oryctomorphus Guérin-Meneville has bounced back and forth between the Rutelinae (i. e., Machatschke 1972) and Dynastinae (i. e., Endrödi 1985). It is in the key by Ratcliffe (1981), but molecular data based on 28 S and 18 S DNA gene regions suggest that Oryctomorphus is a member of the subfamily Rutelinae (A. Smith and D. Hawks, unpubl. data), and so it is not included in the present key. We provide a revised key to the adults of all the New World genera of Pentodontini. Morón and Ratcliffe (1997) provided a key to nine genera of larval Pentodontini.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	type_taxon	Type Species. Collagenus dasysternus Ratcliffe and Hardy, new species, here designated.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	description	Description. Scarabaeidae, Dynastinae, Pentodontini. Form: Suboval, pronotum widest at middle, elytra widest past middle. Length of only known specimen 20.1 mm. Color: Light and dark reddish brown. Head: Frons at center with weakly elevated, flattened boss. Frontoclypeal carina strong, elevated completely from side to side. Clypeus with apical rim swollen, strongly reflexed; venter of clypeus greatly swollen and plump. Mandibles small, subtriangular, completely hidden beneath clypeus. Mentum strongly swollen and protuberant ventrally. Pronotum: Surface completely, moderately punctate. Elytra: Surface with 5 distinctly furrowed striae (2 double rows plus 1 sutural stria). Pygidium: Surface completely, moderately punctate, strongly convex. Legs: Legs with moderately dense, long, yellowish brown setae. Foretibia tridentate; apical tooth slightly removed, larger, curved; apex obliquely truncate. Apices of meso- and metatibiae each with fringe of stout bristles. Tarsi and claws slender. Venter: Thoracic sternites with dense, long, yellowish brown setae. Prosternal process a small, rounded protuberance. Abdominal sternites shiny, glabrous.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	diagnosis	Diagnosis. Collagenus can be easily distinguished from similar genera because of the unique form of the greatly thickened rim on the broadly parabolic clypeal apex, ventrally protuberant mentum, strong frontoclypeal carina, absence of tubercles or a fovea on the pronotum, long tarsi, densely setose thoracic sternites, small rounded prosternal process, and the crenulate apex of the posterior tibiae.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	etymology	Etymology. One of the most striking features of this new genus is the greatly swollen apical rim and venter of the clypeus that gives the impression of swollen ‘‘ lips. ’’ In some circles today, it is cosmetically fashionable to enlarge human lips by injecting them with collagen, an insoluble fibrous protein. Collagen is derived from the Greek kolla, meaning glue. Collagenus is a play on combining ‘‘ collagen’ ’ (in reference to its use to enlarge lips) and ‘‘ genus’ ’ (in reference to this new genus), hence (loosely) the big-lipped genus. The name Collagenus is considered masculine. Collagenus dasysternus Ratcliffe and Hardy, new species (Figs. 1 – 4)	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	materials_examined	Type Material. Holotype labeled ‘‘ Ven: Bolivar, 10 km N Corocito, 18. VI- 3. VII. 87, S & J Peck, FIT, R. Caura rainforest. ’’ Holotype deposited at the Canadian Museum of Nature (Ottawa, Canada).	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	description	Description of Holotype. Male (Fig. 1). Length from apex of clypeus to apex of elytra 20.1 mm; width at humerus 9.6 mm. Color of dorsum and legs dark reddish brown, color of pygidium and venter reddish brown. Head: Frons with surface densely, coarsely rugopunctate; center with small, transverse, feebly elevated boss. Frontoclypeal carina strong, elevated, complete from side to side, weakly arcuate towards posterior. Clypeus with surface densely, coarsely rugopunctate; apex broadly parabolic, with strongly thickened anterior rim, rim strongly reflexed (Fig. 2); clypeus greatly swollen ventrally. Mandibles small, subtriangular, slender, completely hidden beneath clypeus. Mentum enlarged and strongly protuberant ventrally. Interocular width equals 5.0 transverse eye diameters. Anterior edge of eye canthus with dense, long, reddish brown setae. Antenna with 10 segments, club subequal in length to segments 2 – 7. Maxillary palpus with last segment longitudinally depressed and roughened on lateral edge, length 3 times longer than preceding (third) segment, third segment half as long as second segment. Pronotum: Surface entirely punctate; punctures moderate in density, size small (becoming slightly larger in anterior and posterior angles). All margins with strong bead. Sides evenly arcuate. Elytra: Surface between suture and humerus with 1 furrowed sutural stria and 2 pair of furrowed discal striae. Intervals slightly convex, those between striae 1 – 2 and 4 – 5 with irregular row of large punctures. Sides laterad of humerus with 4 distinct rows and 2 indistinct rows of punctures. Propygidium: Surface finely alutaceous with moderately dense, small punctures. Pygidium: Surface entirely punctate; punctures moderate in density and size, becoming slightly smaller at apex. In lateral view, surface strongly, evenly convex. Legs: Trochanter and femora with dense, long, yellowish brown setae. Tibiae with moderately dense, long, stout, yellowish brown setae, those on anterior femora mostly on venter. Foretibia tridentate, teeth becoming progressively larger from base to apex; apical tooth slightly removed from others, long, curved; apex of foretibia obliquely truncate. Meso- and metatibiae each with strong, obliquely transverse carina at about middle on lateral edge; apices expanded, each with fringe of long, stout bristles. Tarsi slender, subequal in length to their respective tibiae, each tarsomere at apex with 5 – 10 long, stout, reddish brown setae. Claws slender, equal in size. Vente r: Thoracic sternites clothed with dense, long, yellowish brown setae. Prosternal process a very small, rounded protuberance. Abdominal sternites shining, glabrous except for about 20 small setae in a transverse row on penultimate segment. Last sternite with apex weakly bisinuate. Parameres: Figs. 3 – 4.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	etymology	Etymology. From the Greek roots dasy, meaning shaggy or hairy, and sternon, meaning chest or sternum. So named in reference to the densely setose thoracic sternites, hence dasysternus.	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
03F787E6FF8399289008725CFC2CFF31.taxon	distribution	Distribution. Known only from a single specimen collected in a flight intercept trap just north of Corocito, which is south of Ciudad Piar on the west side of Embalse de Guri in Bolivar state, Venezuela. Nothing is known of its biology. Revised Key to the Genera of New World Pentodontini	en	Ratcliffe, Brett C., Hardy, Martin (2005): Collagenus dasysternus, A New Genus And Species Of Dynastinae From Eastern Venezuela With A Key To The New World Genera Of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 59 (1): 143-150, DOI: 10.1649/0010-065x(2005)059[0143:cdanga]2.0.co;2
