taxonID	type	description	language	source
03F7F326302DFFA0DDCB0E8905C55E6B.taxon	description	Figs 1 – 3	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F326302DFFA0DDCB0E8905C55E6B.taxon	diagnosis	Diagnosis. This species can be distinguished from M. azureus, which is occasionally collected sympatrically with M. rivularis n. sp., by shape of the male segment IX that is strongly extended posteroventrad. Among species of the M. azureus Species Group, this character state is similar to that of M. schmidi Morse & Yang 2002 from Sri Lanka, but distinguishable from the latter by the long sigmoid spine of segment X. The female of this species is characterized by the manta-ray-shaped plate of the spermathecal sclerite. Adult. Forewing length: male 6.4 – 7.6 mm (mean = 7.2 mm, n = 10); female 6.8 – 7.6 mm (mean = 7.3 mm, n = 9). Vertex, thorax, and wings in ethanol brownish black. General morphology typical for the genus. Relative eye size smaller than that in M. azureus for both male and female (Fig. 1). Male genitalia (Figs 2 A – 2 E). Segment IX 1 / 5 as long dorsally as tall laterally, 1 1 / 5 as long ventrally as tall laterally; in ventral view apicoventral process of sternum 3 / 4 as long as rest of sternum, with stout base and slender V-shaped lateral arms. Preanal appendages very long and slender. Tergum X antisymmetrical, produced into two sinuate spines: one spine sigmoid in dorsal aspect, long, extending far beyond apical process of sternum IX; other spine weakly sinuate and shorter than former. Inferior appendages each with dorsomesal lobe oriented vertically, somewhat longer than mid-width in lateral view, its anterior and posterior margins subparallel; posterior margin with three caudal projections: upper caudal projection with acute apex; middle caudal projection narrow and often longest among three projections, with acute apex; lower caudal projection with round apex. Phallus without evident paramere spines or phallicata, with ventromedian spine directed anteroventrad at one third distance from apex and pair of sharp, triangular flanges subapically, apicoventral lip of phallobase constricted subapically in ventral aspect. Variation (Figs 3 A 1 – 3 F 3): The shapes of tergum X and inferior appendages are variable geographically. The shorter process of tergum X is about half the length of the longer process in individuals from central Honshû (Figs 3 F 1 – 3 F 3), but the former is prolonged in those from northern Honshû (Figs 3 C 1 – 3 E 1) and almost reaches the tip of the latter in those from Hokkaidô (Figs 3 A 1 – 3 B 2). The middle projection of each inferior appendage tends to be more strongly developed in individuals from Hokkaidô and northern Honshû (Figs 3 A 1 – 3 E 1) than those from central Honshû (Figs 3 F 1, 3 F 2) with an exception (Fig. 3 F 3). Female genitalia (Figs 2 F – 2 I). Segment IX short in lateral view. Segment X shorter than preanal appendages, narrowly incised ventromedially in dorsal view. Preanal appendages slender, straight, and setose. Lamellae very long, constricted basally, shape in lateral aspect variable geographically; dorsal margin inflated; apicodorsal corner protruded outward; outer surface slightly concave. Gonopod plates triangular in ventral aspect with pair of posterolateral processes short, rounded and apex extending far beyond apex of segment X. Spermathecal sclerite with pigmented manta-ray-shaped plate, anterior and posterior margins extended anterad and posterad mesally and acute. Variation: Lamellae in lateral aspect are each subelliptical and rounded apically in individuals from the type locality Kakida-gawa, central Honshû (Fig. 2 F), but more nearly parallel-sided, gradually heightened caudally, and obliquely truncate apically in those from Hokkaidô (Fig. 2 I).	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F326302DFFA0DDCB0E8905C55E6B.taxon	materials_examined	Holotype: ³, Kakida-gawa, Shimizu-chô, Shizuoka, Honshû, Japan, 35.1031 ° N, 138.9028 ° E, alt. 13 m, 27. iv. 2002, TN. Paratypes: 3 ³, same data as holotype; 5 ³, type locality, 22 – 23. xi. 2002, NK, 2 ³ (pinned), type locality, 5. iv. 2006, TN. Other specimens examined. HOKKAIDÔ: Nemuro: 1 ♀, Shibetsu-chô, Shibetsu-shitsugen, 12. viii. 1996, K. Kuribayashi, 1 ³ 1 ♀, same location, 21. viii. 2013, M. Nakatani; 1 ♀, same location, 2 – 3. viii. 2013, M. Nakatani; 1 ³ 1 ♀, same location, 3. ix. 2013, M. Nakatani. Kushiro: 1 ³, Kushiro-shi, Akan-chô, Ibeshibetsu-gawa R., 6. viii. 1990, NK; 1 ³, same location, 6. viii. 1990, NK; 3 ³ 1 ♀, Shibecha-chô, Gojikkoku, Shirarutoroetoro-gawa R., 16. vii. 2009, TI; 2 ³ 7 ♀, same location, 28. vii. 2012, TI; 1 ³ 2 ♀, Shibecha-chô, Kayanuma, Shirarutoroetoro-gawa R., 16. vii. 2009, TI; Kamikawa: 1 ³, Horokanai-chô, small tributary of Shumarinai-gawa R., 10. viii. 1999, TI & A. Ohkawa; 1 ³, Horokanai-chô, small tributary of Shumarinai-gawa R., 7. vii. 2007, NK. Ishikari: 1 ³, Chitose-shi, Bibi, Bibi-gawa R., 5 – 26. ix. 1993, TI; 1 ³ 1 ♀, same location, 4. viii. 2007, TI; 1 ³ 3 ♀, Eniwa-shi, Izari-gawa R., Eniwa-ôhashi, 11. vii. 1999, TI; 1 ³, same location, 22. vii. 1999, TI; 1 ³, same location, 17. vii. 2015, TI; 1 ♀, Sapporo-shi, Hitsujigaoka, 24 – 31. vii. 2009, K. Konishi; 1 ³, Sapporo-shi, Nopporo-shinrin-kôen, Osawa-guchi, 17. vi. 2002, M. Sakurai. Iburi: 3 ³ 5 ♀, Tomakomai-shi, Misawa, Bibi-gawa R., 29. vii. 1989, NK; 5 ³ 1 ♀, same location, 17. vii. 1990, NK; 1 ♀, same location, 16. viii. 1990, NK; 4 ♀, same location, 18. viii. 1991, TI; 2 ³, same location, 12. ix. 1993, TI; 1 ³, same location, 2. viii. 1998, NK; 1 ³, same location, 20. viii. 2007, TI & A. Ohkawa; 5 ³ 8 ♀, same location, 31. vii. 2009, NK; 2 ³, Tomakomai-shi, Misawa, small stream, 1. viii. 1992, NK; 1 ³, same location, 15. viii. 1992, NK; 5 ³ 2 ♀, Tomakomai-shi, Uenae, Bibi-gawa R., 22. vii. 2001, TI & A. Ohkawa; 1 ³, same location, 25. vii. 2001, TI; 1 ³, same location, 29. vii. 2001, TI; 1 ³, same location, 30. vii. 2001, TI; 3 ³ 2 ♀, same location, 23. viii. 2007, TI; 2 ³ 3 ♀, same location, 16. vii. 2008, TI. HONSHÛ: Aomori: 1 ³, Nishimeya-mura, Anmon-gawa R., alt. 240 m, 16. ix. 2010, TI; 1 ³, Takko-machi, Natsuzaka, Kumahara-gawa R., 18. viii. 1996, Suzuki. Yamagata: 3 ³, Nishikawa-chô, Shizu, Buna-no-izumi, 11. ix. 2003, TI. Shizuoka: 1 ♀, type locality, 11. iii. 1984, reared and emerged on 7. iv. 1984, TN; 3 ³ 1 ♀, same data as holotype; 1 ³ 1 ♀, type locality, 27. iv. 2002, TN; 1 ♀, type locality, 31. viii. 2002, TN; 2 ³ 1 ♀, type locality, 14. xi. 2002, TN; 6 ³ 1 ♀, type locality, 22 – 23. xi. 2002, NK.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F326302DFFA0DDCB0E8905C55E6B.taxon	etymology	Etymology. The specific epithet (Latin adjective, rivularis = of a brook or small stream) refers to the habitat of this species.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F326302DFFA0DDCB0E8905C55E6B.taxon	distribution	Distribution. Japan (Hokkaidô, eastern Honshû). Habitat. Adults are often found beside slowly flowing streams. They are sometimes collected with M. azureus sympatrically, but the habitat preference of the new species is narrower than that of the latter.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263021FFA2DDCB089803FA5C52.taxon	description	Figs 1, 4 A – 4 I Mystacides sp. (af. superatus): Morita 2011, 217.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263021FFA2DDCB089803FA5C52.taxon	diagnosis	Diagnosis. Among species of the M. azureus Species Group, the male of this species shares an unusually large caudal projection of the each inferior appendage with that of M. superatus from China, but can be distinguished from it by lack of the lower projection of the appendage. Female of this species is similar to that of M. azureus but can be distinguished from it by the shape of the lamellae; in lateral aspect, each is parallel sided or slightly tapered in the apical half in the present species but broadest with a dorsal expansion at midpoint in M. azureus. It also somewhat resembles that of M. rivularis n. sp. but can be distinguished from it by the subtriangular plate of the spermathecal sclerite. Adult. Forewing length: male 6.1 – 8.1 mm (mean = 7.5 mm, n = 10); female 6.8 – 8.5 mm (mean = 7.5 mm, n = 8). Vertex, thorax, and wings in ethanol brownish black. General morphology typical for the genus. Relative eye size smaller than that in M. azureus for both male and female (Fig. 1). Male genitalia (Figs 4 A – 4 E). Segment IX 1 / 8 as long dorsally as tall laterally, 1 1 / 4 times as long ventrally as tall laterally; in ventral view apicoventral process of sternum 2 / 3 as long as rest of sternum, with stout base and slender V-shaped lateral arms. Preanal appendages very long and slender. Tergum X antisymmetrical, produced into two spines; longer spine bent inward at one-third from base in dorsal aspect, extending beyond apical process of sternum IX; other spine half as long as longer one, nearly straight in dorsal aspect with acute apex. Inferior appendages each clavate in lateral view with slender base 1 / 3 as thick as broad apex; upper caudal projection directed dorsad, as long as wide; middle caudal projection unusually large, curved dorsomesad, tapering to acute apex; lower caudal projection lacking. Phallus without evident paramere spines or phallicata, with ventromedian spine directed anteroventrad at one third distance from apex and pair of sharp, triangular flanges subapically, apicoventral lip of phallobase constricted subapically in ventral aspect. Female genitalia (Figs 4 F – 4 H). Segment IX short. Segment X shorter than preanal appendages, narrowly incised ventromedially in dorsal view. Preanal appendages slender, straight, and setose. Lamellae very long; in lateral aspect each with apical half twice as broad as basal half, somewhat curved downward, parallel sided or slightly tapered in apical half, with round apex; dorsal margin inflated; outer surface slightly concave. Gonopod plates triangular in ventral aspect with short, rounded pair of posterolateral processes; apex extending far beyond apex of segment X. Spermathecal sclerite rhomboid, rounded anteriorly.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263021FFA2DDCB089803FA5C52.taxon	materials_examined	Holotype: ³, Honshû, Mie, Shima-shi, Isobe-chô, Natsukusa, 34.359 ° N, 136.766 ° E, alt. 20 m, 5. vii. 2008, H. Morita. Paratypes: 1 ³, same data as holotype; 7 ³, Honshû, Hyôgo, Asago-shi, Wadayama, Kudawa, Ishibe-jinjya, 3. xi. 2006, K. Inazu & TI. Other specimens examined. HONSHÛ: Fukui: 5 ³ 9 ♀, Tsuruga-shi, Ikenokôchi-shitsugen, 1. x. 2013, TI; 3 ³ 11 ♀, same location, 30. v. 2015, TI; 8 ³ 7 ♀, same location, 24. vi. 2015, TI; 5 ³ 4 ♀, same location, 21. vii. 2015, TI; 4 ♀, same location, 11. vi. 2016, TI. Hyôgo: 3 ³, Asago-shi, Wadayama, Kudawa, Ishibe-jinjya, 19. v. 2009, R. B. Kuranishi.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263021FFA2DDCB089803FA5C52.taxon	etymology	Etymology. Named moritai after Mr. H. Morita, who first provided us the examined specimens of this species, including the holotype.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263021FFA2DDCB089803FA5C52.taxon	distribution	Distribution. Japan (western Honshû). Habitat. Adults of this species have been collected at three sites: beside a small stream with low gradient, from a stream flowing through a small marsh, and from a pond recharged by spring water. Adults of M. azureus also have been collected at all these sites sympatrically.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263023FFA3DDCB0A88047C5BD7.taxon	description	Figs. 1, 5 A – 5 M, 6 A 1 – 6 E 6	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
03F7F3263023FFA3DDCB0A88047C5BD7.taxon	discussion	Remarks. The males exhibit great variation in the shape of tergum X even among individuals collected at the same time and place. The tergum X is produced into two asymmetrical spines posteriorly: In most populations, one spine is shorter and sinuous in the distal half and the other spine is curved and longer but variable in shape and length. The latter spine is slightly longer than the former and curved mesad in its apical half or bent mesad at midlength in more than half of the examined specimens (short type; Figs 6 A 1, 6 A 2, 6 B 1 – 6 B 4, 6 C 1, 6 C 2, 6 E 1 – 6 E 3), which type well agrees with European individuals (Kumanski 1988; Mosely 1939). However, in some specimens, the latter is far longer than the former and curved sigmoidally in dorsal aspect (long twist type; Figs 6 A 5, 6 C 5, 6 C 6, 6 E 5, 6 E 6). In addition, a few specimens have the longer spine slightly curved outward subapically, representing an intermediate between the two types (Fig. 6 A 3, 6 A 4, 6 C 3, 6 C 4, 6 E 4). In most specimens, the longer spine is crossing beneath the shorter. Frequencies of the long twist type vary geographically. Only a few or no long-twist type individuals were found in specimens collected in Hokkaidô and Kyûshû, but over 20 % of individuals exhibit this type in specimens collected in Honshû and Shikoku (Table 1). The population in lakes in Ômachi, Nagano, and Honshû, however, exhibit a different type of shape of tergum X from the other populations (Figs 5 F – 5 J): The usually short sinuate spine is usually longer than the curved one (reversal type; Figs 6 D 1, 6 D 2), although some specimens have the curved spine extending beyond the sinuate spine (Figs 6 D 3, 6 D 4), resembling the intermediate type. Specimens having more extended spines (Fig. 6 D 5) almost agree with those of the long twist type. The degree of the posteroventral excisions of the inferior appendages in lateral aspect is also variable (Figs 5 A, 5 F, 6). The excision of specimens collected in Honshû (Figs 6 C – 6 E) tends to be deeper than that in Hokkaidô (Figs 6 A, 6 B), but variation of this character within each population is rather small. The reversal type specimens of the Ômachi population closely resemble illustrations of Mystacides elongatus from China by Yang & Morse (2000). In addition, our female specimens of this species show no evident variation in genital morphology and are very similar to illustrations those of M. elongatus by them. These facts suggest that M. elongatus may be included in the variation of M. azureus. Morphological and molecular studies of M. elongatus specimens from China would be needed to clarify the relation of M. elongatus and M. azureus. HOKKAIDO: Kushiro: 57 ³ 2 ³ (L) 3 ³ (I) 13 ♀, Kushiro-shi, Akan-chô, Akan-ko, 14. ix. 1999, TI & N. Minakawa; 6 ³ 10 ♀, same location, 7. viii. 2007, TI; 6 ³ 5 ♀, Kushiro-shi, Akan-chô, Ibeshibetsu-gawa R., 19. ix. 1989, NK; 1 ³ 1 ³ (L) 4 ♀, Kushiro-shi, Akan-chô, Panketô, 21. ix. 1996, TI; 1 ³, same location, 14. ix. 1999, N. Minakawa; 6 ³ 2 ♀, same location, 3 – 4. x. 2006, TI; 1 ³ 2 ♀, same location, 8. viii. 2007, TI; 7 ♀, Shibecha-chô, L. Shirarutoro-ko, 9. viii. 2005, TI; 1 ³, same location, 16. viii. 2007, TI; 1 ³, same location, 25. vii. 2008, TI. Sôya: 16 ³ 8 ♀, Sarufutsumura, Asajino, 31. vii. 2007, NK. Kamikawa: 1 ♀, Tôma-chô, Nakadai, 15. vii. 2007, NK. Sorachi: 1 ♀, Yuni-chô, Kawabata, small stream, 15 – 30. viii. 2007, NK. Ishikari: 1 ♀, Chitose-shi, Bibi, Bibi-gawa R., 29. viii. 1991, TI; 1 ♀, same location, 5 – 26. ix. 1993, TI; 4 ♀, Chitose-shi, Neshikoshi, Chitose-gawa R., 4. ix. 1997, NK; 1 ³, Chitose-shi, Izumisawa, Mamachi-gawa R., 6. viii. 2003, NK; 1 ♀, Chitose-shi, Okotan, Okotanpe-gawa R., 26. ix. 1998, NK; 4 ³ 1 ♀, Chitose-shi, L. Shirarutoro-ko, 29. ix. 1996, TI & A. Ohkawa; 3 ♀, same location, 29. ix. 1996, A. Ohkawa; 1 ³ 5 ♀, same location, 28. viii. 1997, TI & A. Ohkawa; 5 ³ 1 ♀, same location, 25. ix. 1997, NK; 3 ³ 13 ♀, same location, 5. viii- 7. x. 2005, NK; 6 ³ 5 ♀, same location, 4 – 10. viii. 2006, NK; 2 ³ 1 ♀, Chitose-shi, small stream beside L. Shikotsu-ko, 1. x. 1993, NK; 4 ♀, same location, 21. VII- 6. x. 1996, Y. Nagayasu; 1 ♀, Sapporo-shi, Misumai, Toyohiragawa R., 27. vii. 1992, NK. Iburi: 1 ♀, Atsuma-chô, Naganuma, Koi-numa, 13. vii. 2006, NK; 1 ♀, Tomakomai-shi, Misawa, Bibi-gawa R., 18. vi. 1990, NK; 2 ³, same location, 2. viii. 1998, NK; 6 ♀, same location, 5. vii. 2009, NK; 2 ³ 3 ♀, Tomakomai-shi, L. Utonai-ko, 1 – 6. vii. 1998, NK; 1 ³ 1 ♀, same location, 21. vi. 2003, NK; 9 ³, same location, 22. vii. 2004, TI. Oshima: 4 ³ 6 ♀, Nanae-chô, Konuma, 23. vi. 2003, NK; 10 ³ 12 ♀, Nanae-chô, Ônuma, 23. vi. 2003, NK. HONSHU: Aomori: 1 ³ (I), Fujisaki-machi, Shirako, Iwaki-gawa R., 2. viii. 1996, Suzuki; 1 ³ 1 ³ (L) 2 ♀, Hirosaki-shi, Akudo, Iwaki-gawa R., 20. viii. 1996, Suzuki. Akita: 5 ³, Kosaka-machi, Towadako-namariyama, L. Towada-ko, 8. viii. 1999, H. Kato. Fukushima: 1 ³ 1 ³ (L) 3 ♀, Shôwa-mura, Yanohara-kôgen, 19. viii. 2008, N. Katsuma. Ibaraki: 4 ³ 2 ³ (L) 3 ♀, Kasama-shi, Minamikoizumi, 24. vi. 2007, N. Katsuma. Gumma: 1 ³ (L) 1 ³ (I) 2 ♀, Fujioka-shi, Samba-gawa R., 29. v. 1991, S. Ishiwata. Tôkyô: 1 ³ (L), Fussa-shi, Nagatabashi, 9. vi. 1985, S. Sasaki. Kanagawa: 1 ³, Hadano-shi, L. Shinsei-ko, 16. xii. 1979, TN; 2 ³ (I) 15 ♀, Sagamihara-shi, Fujino-chô, Magino, 9. vi. 1988, TN; 2 ³ 1 ³ (L) 5 ♀, Yamakita-machi, Shiraishi-zawa, 5 – 6. vii. 1984, TN; 1 ³ (L), same location, 19. vii. 1982, TN; 10 ³ 4 ³ (L) 7 ♀, Zushi-shi, Sakurayama, Morito-gawa R., 20. vii. 2009, TN. Fukui: 1 ³, Tsuruga-shi, Ikenokôchi-shitsugen, 21. vii. 2015, TI; 1 ³ 1 ³ (L) 13 ♀, same location, 11. vi. 2016. Nagano: 1 ³ 1 ♀, Koumi-chô, Matsubara, L. Chô-ko, 30. v. 1997, TI; 3 ♀, Maruko-machi, Uchimura-kawa, 21. vi. 1997, K. Tojo; 1 ³ (R), Ômachi-shi, L. Aoki-ko, 28. viii. 2008, N. Katsuma; 1 ³ (L) 2 ³ (R), same location, 23. ix. 2009, NK; 1 ³ (L) 2 ³ (I) 9 ³ (R) 17 ♀, Ômachi-shi, L. Nakatsuna-ko, 27. ix. 1990, NK; 3 ³ (L) 2 ³ (I) 14 ³ (R) 19 ♀, same location, 23. ix. 2009, NK. Gifu: 1 ³ (L), Higashishirakawa-mura, Oppara, 27. v. 1988, TN. Shizuoka: 2 ³ 3 ³ (L) 1 ³ (I) 1 ♀, Shimoda-shi, Tateno, 12. iv. 2009, S. Inaba. Mie: 4 ³ 2 ³ (L) 2 ³ (I) 2 ♀, Shima-shi, Isobe-chô, Natsukusa, 5. vii. 2008, H. Morita. Hyôgo: 2 ³ 1 ³ (I), Asago-shi, Wadayama, Kudawa, Ishibe-jinjya, 3. xi. 2006, K. Inazu. & TI. Shimane: 1 ³ 3 ♀, Gôtsu-shi, Sakuraechô, Kawagoe, 2. v. 1999, SN; 2 ³, Oochi-chô, Shiki, 10. viii. 1999, SN; 1 ³ 1 ♀, Kawamoto-machi, Inbara, 12. v. 1999, SN; 4 ³ 1 ♀, Masuda-shi, Mukaiyokotachô, 7 - 8. v. 2000, SN; 1 ³ 4 ♀, same location, 29. vii. 2000, SN; 3 ³, Masuda-shi, Yasudomichô, 18. vii. 2000, SN; 2 ³ 8 ♀, Masuda-shi, Musochô, 6. v. 2001, SN; 2 ³ 1 ³ (L), Masuda-shi, Musochô, Takatsu-gawa R., 27. vii. 2000, SN. Hiroshima: 1 ³ (I), Akitakata-shi, Yachiyo-chô, Haji, Haji-dam, 11. vii. 2000, SN; 1 ³ 2 ♀, Akitakata-shi, Yoshida-chô, Yoshida, 21. iv. 1999, SN; 1 ³ (L) 1 ³ (I), Higashihiroshima-shi, Takaya-chô, Nakashima, 13. vii. 1999, SN; 1 ³, Hiroshima-shi, Asakita-ku, Kabechô, Imaida, 24. iv. 1999, SN; 3 ³ 1 ³ (L) 2 ♀, Kôzan-chô, Uzuto-gawa R., Hattabara-dam, 5. viii. 1998, SN; 6 ³ 1 ³ (L) 1 ♀, same location, 25. vii. 2000, SN; 1 ³ (I), Miyoshi-shi, Minamihatajikimachi, 1. viii. 1999, SN; 2 ³, same location, 6. x. 1999, SN; 1 ³ 2 ♀, Ôtake-shi, Fukase, Kose-gawa R., 8. v. 2001, SN; 1 ³, same location, 23. vii. 2001, SN; 1 ³ (L) 4 ♀, Ôtake-shi, Bouroku, Kose-gawa R., 7. v. 2001, SN; 5 ³ 4 ♀, Ôtake-shi, Kuritanicho, Yasaka-dam, 28. vii. 2000, SN; 1 ³, Sera-chô, Io, Hattabara-dam, 26. vii. 2000, SN; 2 ³ 1 ³ (L), Yuki-chô, Shimominauchi, 25. iv. 1999, SN; 1 ³ (L), same location, 15. v. 2003, TN. Yamaguchi: 1 ³ 2 ♀, Iwakuni-shi, Miwachô, One-gawa, Yasaka-dam, 25. ix. 2000, SN. SHIKOKU: Tokushima: 1 ³ 1 ³ (L), Kitô-son, Takanose-kyô, 18. vii. 1998, I. Yamashita. Kagawa: 2 ³ 1 ³ (I) 1 ♀, Takamatsu-shi, Shionoe, 14. ix. 2006, NK. Kôchi: 11 ³ 2 ³ (L) 6 ♀, Kami-shi, Monobechô, Befu, 2. viii. 2003, I. Yamashita; 2 ³ (L), Muroto-shi, Sakihama, Sakihama-gawa R., 29. iv. 2004, M. Takai; 1 ³ (L) 1 ♀, Nankoku-shi, Nakanogawa-rindô, 29. v. 2004, M. Takai; 1 ³ (L), Ochi-chô, Tokoroyama, Ichigaya, 21. vii. 2004, K. Nio; 1 ³ (L), Sukumo-shi, Itchûbara, 28. iv. 2001, M. Takai; 1 ³, same location, 30. iv. 2004, M. Takai; 1 ³, Tosashimizu-shi, Mochiishi, Kitayamahigashi, 21. vii. 2004, K. Nio. KYÛSHÛ: Fukuoka: 1 ³ 1 ♀, Chikushino-shi, Yoshiki, 30. iv. 1986, N. Gyotoku; 1 ³ 1 ♀, same location, 9. x. 1986, N. Gyotoku; 2 ³, Ukiha-shi, Yoshiimachi, Sakurai, Chikuko-gawa R., 8. iv. 1997, TN. Saga: 4 ³ 9 ♀, Takeo-shi, 22 – 23. iv. 2009, T. Shimizu; 7 ³ 1 ♀, Ureshino-shi, 4. vi. 2009, T. Shimizu. TSUSHIMA: 5 ³ 2 ³ (L) 2 ♀, Tsushima-shi, Kamiagata-chô, Sago, upper reaches of Sago-gawa R., 22. vii. 2009, R. B. Kuranishi.	en	Kuhara, Naotoshi, Nozaki, Takao, Zhang, Ao, Zhou, Xin (2023): DNA barcoding facilitates discovery and description of two new species of the Mystacides azureus Species Group (Trichoptera: Leptoceridae) in Japan. Zootaxa 5306 (2): 215-231, DOI: 10.11646/zootaxa.5306.2.3, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.3
