taxonID	type	description	language	source
03F5672C4E23C14BFE90F5F8A69EBC00.taxon	diagnosis	Diagnosis — Body length, 407 – 500 µm; width, 200 – 286 µm. Integument of notogaster punctuated and irregularly alveolate with seven pairs of round elevations bearing dorsal seta (e); epimerata granulate; other surface punctulate. Rostral tip rounded. Notogaster with 13 pairs of T-shaped setae, 2 pairs of simple setae (p 1 and p 2) and 1 pair of virtual setae f 1. Sensilli consisting of penicillate swollen head and smooth thin stem. Lateral margin of pedotectum 1 bearing more than 20 ciliary’s processes. Semicircular suture (na) of ventral region short, not reaching the level of insertion of aggenital seta ag 1 or lyrifissure ih. Genito-anal setal formula: 9 - 2 - 2 - 3; all setae biramous. Subcapitulum diarthric, subcapitular setae: 1 - 2 - 1; setae h biramous. Epimeral setal formula: 4 - 2 - 3 - 4; all setae simple. Monodactylous. Material examined — Holotype (Adult female) (NSMT-Ac 13611) from sand, and soil at point A; 9 paratypes (Adult female) (NSMT-Ac 13635 to 13640) from humus and soil of the garden under no-tillage manner of Nagasato at Asagiri-cho, Kumamoto Prefecture, Fujikawa T. leg, on March 3, 2008, September 20, 2009, April 11, 2011, October 15, 2011, November 6, 2011, December 16, 2011; 1 paratype (Nymph) (NSMT-Ac 13641): same data as paratypes of adults but on December 16, 2011; 7 paratypes (Adult female) for the scanning electron microscopy: same data as paratypes of adults but on November 5, 2009 and March 3, 2012. Etymology — After the numerous ciliary’s processes of pedotectum. Measurements and body appearance — Body length (14 exs.): 407 (mean 450) 500 µm; width (11 exs.): 200 (229) 286 µm. Body colour light brown, Integument of prodorsum, ventral plate, subcapitural plate, genital plates, anal plates and legs punctulate; epimerata granulate; notogaster punctulate and irregularly alveolate with seven pairs of round elevations bearing one or three dorsal setae. Prodorsum — Rostral tip widely rounded bearing smooth spiniform setae ro (ca. 33 µm) inserting far anterior on prodorsum and close to each other (Fig. 4 A). Setae le (155 µm) and in (281 µm) T-shaped with dilated basal portion and long flagelliform distal portion (Figs. 3 B, C). Setae le bearing a few acute projections, inserting on round elevations. Bothridia barrel-shaped, opening dorso-laterally. Sensilli (ss) (62 µm) consisting of penicillate swollen head and smooth thin stem. Setae ex 1 (6 µm) and ex 2 (3 µm) short, smooth, inserting at the basal portion of bothridium. Lateral margin of pedotectum 1 bearing more than 20 ciliary’s processes (7 – 12 µm). Posterobothridial condyles small acute (Fig. 3 A). Relative lengths and distances: in> le> ss> ro> ex; (in - in: 66 µm)> (le - le: 21 µm)> (ro - ro: 14 µm). Notogaster — Anterior margin straight, shorter than the width of the middle part of notogaster .. Notogaster with 13 pairs of T-shaped setae (241 µm), 2 pairs of simple setae (p 1 and p 2) (73 µm), 1 pair of virtual setae f 1 and seven pairs of round elevations; six elevations bearing each seta of cp, d 1 - 2, e 1 - 2 and f 2; the last pair of posterior elevations bearing three setae h 1 - 3. Virtual seta f 1 present posterior to elevation bearing seta e 1. Posterior projection of notogaster with truncate margin, bearing two pairs of setae p 1 and p 2 at corners. Lyrifissures long, remarkable; ia (20 µm) and ip (9 µm) aligned along lateral margin of notogaster, posterior to c 3 and h 3, respectively; im (9 µm) longitudinally anterior to e 2; ih and ips situated ventrally (Fig. 4 E); ih (12 µm) aligned transversely posterior to ag 1; ips (16 µm) obliquely posterior to ag 2. Ventral region — Genital aperture (64 µm) anterior roughly triangle, posterior parabola in shape; anal aperture (84 µm) elliptical. Semicircular suture (na) on ventral plate between genital and anal apertures, short, not reaching the level of insertion of aggenital seta ag 1 or lyrifissure ih. Genito-anal setal formula: 9 - 2 - 2 - 3; all setae smooth, biramous (Fig. 3 D). Genital (36 – 48 µm) and anal (17 µm) setae inserting at the inner margin of each plate (Figs. 4 D- E). Setae ag 1 inserting at level of insertion of g 7; ag 2 (91 µm) aligned in latero-posteriorly to genital aperture. All adanal setae (64 µm) aligned in adanal. Lyrifissures ian (19 µm) longitudinally anterior to an 2; iad (14 µm) inverse apoanal, located near anterior margin of anal aperture. Subcapitulum diarthric, subcapitular setae: 1 - 2 - 1; setae h (41 µm) smooth, biramous; a (16 µm), m 1 - 2 (11 µm) smooth, simple (Fig. 4 C). Epimeral setal formula: 4 - 2 - 3 - 4; all setae (11 - 65 µm) smooth, simple spiniform (Fig. 5 A); 3 b the longest, 1 c the shortest. Sternal ridge, apodemata 1 - 4 and sj. distinct. Setae cha (8 µm) thick basally; chb (26 µm) long bacilliform (Fig. 5 D). Trägårdh’s organ (31 µm) taeniform. Legs — Monodactylous; claw (22 to 27 µm) smooth (Fig. 4 B). Setal formula of legs: I (1 - 6 - 5 - 6 - 26), II (1 - 8 - 5 - 6 - 23), III (5 - 3 - 3 - 4 - 21), IV (1 - 3 - 3 - 4 - 19). Solenidiotaxy: I (1 - 2 - 2), II (1 - 1 - 3), III (1 - 1 - 0), IV (1 - 1 - 0). Measurements (µm) of segments: Holotype: I ([41 - 46] - [84 - 80] - [54 - 52] - [39 - 41] - [93 - 86]), II ([54 - 48] - [70 - 71] - [39 - 45] - [36 - 36] - [75 - 80]), III ([71 -?] - [61 - 59] - [32 - 27] - [36 - 36] - [61 - 57]), IV ([71 -?] - [71 - 63] - [38 - 30] - [41 - 41] - [71 - 80]). Average value according to paratypes: I ([45 - 45] - [79 - 83] - [49 - 54] - [46 - 45] - [85 - 92]), II ([39 - 41] - [64 - 67] - [44 - 46] - [40 - 37] - [77 - 87]), III ([60 - 62] - [56 - 64] - [33 - 31] - [31 - 36] - [66 - 62]), IV ([62 - 60] - [72 - 67] - [37 - 37] - [38 - 45] - [75 - 75]). Legs I, II and III bearing solenidia of all tibiae and genua contiguous to dorsal seta (Figs. 3 E, 5 B). On tarsus I (Fig. 3 E), famulus ε (5 µm) spiniform situated lateral to seta tc’ (19 µm), posterior to solenidion ω 2; ω 2 (20 µm) thin bacilliform, lateral to seta it’ (25 µm); ω 1 (25 µm) thick bacilliform, narrow basally, posterior to seta ft’ (26 µm); laminate dorsal seta (29 µm) large, smooth, with longitudinally running nerve-like line. Tarsus II bearing short clavate ω 1 (10 µm) and ω 2 (8 µm) posterior, and long thin bacilliform ω 3 (13 µm) at the basal portion of claw (Fig. 5 B). On tibia I, solenidia ’ 1 (12 µm) and ’ 2 (12 µm) contiguously anterior to smooth entire seta d (14 µm); lateral setae (31 µm) broad, large bearing minute barbs. On genu I, σ (21 µm) thin bacilliform, contiguous to smooth entire seta d (28 µm) situated on a small apophysis; lateral setae (41 µm) minutely barbed, long, broad taeniform with truncate, serrate anterior margin. Juvenile instar (Figs. 6 and 7) A single tritonymph: length, 443 µm; width, 179 µm. Body surface wrinkled, light whitish- yellow coloured without any ornamentation. Rostrum widely rounded bearing simple, smooth rostral setae (21 µm) laterally. Setae le (83 µm) and in (152 µm) T-shaped originating from apophysis (Fig. 6 A). Sensillus lost. Lateral margin of pedotectum 1 bearing 8 ciliary’s processes (5 - 16 µm) (Fig. 6 C). Posterobothridial condyles absent. Relative distances: (ro-ro) = (le-le) = (in-in). Gastronotum covered by ten pairs of large round elevations bearing T-shaped seta (e) (238 µm); eighteen pairs of notogastral setae recognized (Fig. 6 B). Semicircular suture between genital and anal apertures absent. Distance (70 µm) between genital (40 µm) and anal (71 µm) apertures almost as long as the length of anal aperture. Genito-anal setal formula: 7 - 2 - 2 - 3; an 2 (10 µm) short, smooth, thin, simple setiform; other setae lost. Seta h (40 µm) and ag (47 µm) smooth, biramous (Figs. 7 A-B). Epimeral setal formula: 4 - 2 - 3 - 4; all setae (6 - 46 µm) smooth, simple spiniform; the longest 2 A, the shortest 2 b. Setal formula of legs: I (1 - 6 - 5 - 5 - 21), II (1 - 8 - 5 - 4 - 20), III (3 - 3 - 3 - 4 - 18), IV (1 - 3 - 3 - 4 - 16). Solenidiotaxy: I (1 - 2 - 2), II (1 - 1 - 2), III (1 - 1 - 0), IV (1 - 1 - 0). Measurements (µm) of segments: I ([30 - 32] - [57 - 59] - [45 - 36] - [36 - 39] - [68 - 59]), II ([36 - 36] - [54 - 63] - [39 - 32] - [32 - 30] - [52 - 50]), III ([36 - 39] - [45 - 36] - [18 - 23] - [21 - 26] - [41 - 50]), IV ([50 - 41] - [36 - 54] - [21 - 23] - [27 - 26] - [59 - 59]). Remarks — The new species differs from all the species of the genus Masthermannia Berlese, 1913 by having (1) short semicircular suture between genital and anal apertures, not reaching the level of lyrifissure ih or insertion of aggenital seta ag 1 and (2) biramous genital, aggenital, anal and posterior subcapitular setae. The new species is distinguished from M. nematophora (Grandjean, 1954) by number of the ciliary’s processes of lateral margin of pedotectum 1, shape of solenidia on tarsus I of leg, and length of solenidion on genu I of leg, from the original description of M. hirsuta (Hartman, 1949) by the shape of ornaments on notogastral surface and shape of notogastral posterior region, and from M. hirsuta sensu Aoki (1980) by shape of notogastral posterior region, shape of genital setae, and number of setae on the last pair of median round elevations on notogaster.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E2AC157FEAAF69EA248BD47.taxon	diagnosis	Diagnosis — Body length 807 µm; width 386 µm. Prodorsal surface punctulate and irregularly alveolate. Notogastral surface medially and laterally tuberculate; alveolate between medial and lateral portions. Rostral incision short. Sensilli long, rod-like bearing sparsely spins. Notogaster anteriorly broadly rounded, posteriorly roundish angular, with a pair of longitudinal ridges extending from c - series line to insertion of seta f 1. Of all notogastral setae, f 1 the longest. Solenidia ’ 1 and σ longer than each dorsal seta. Genito-anal setal formula: 9 - 0 - 2 - 3. Subcapitulum diarthric, subcapitular setae: 1 - 2 - 1. Epimeral setal formula: 11 [9] - 5 - 6 - 5. Heterotridactylous. Material examined — Holotype (Adult female) (NSMT-Ac 13634) from sand, soil and humus at point B. Etymology — After longitudinal ridges on notogaster divergent posteriorly Measurements and body appearance — Body length 807 µm; width 386 µm. Body colour light brown. Prodorsal surface punctulate and irregularly alveolate. Notogastral surface medial and lateral portions of notogastral surface tuberculate; surface between medial and lateral portions alveolate. Prodorsum — Rostral incision short (Fig. 10 B). Transversal ridge distinct between insertions of setae ro, and between those of setae le. Setae ro (21 – 23 µm), le (45 µm) and in (50 µm) thick, short rod-like, closely barbed. Sensilli (179 µm) long, rod-like bearing sparsely spins (Fig. 10 A). Bothridia cup-shaped, opening dorso-laterally. Setae ex (30 µm) barbed setiform inserting at the basal portion of bothridium. Lateral prosomatic mark (m) distinct anterolaterally to bothridium. Notogaster — Anterior margin broadly rounded; posterior margin roundish angular; a pair of longitudinal ridges extending from c - series line to insertion of seta f 1, not joining posteriorly. Of all 16 notogastral setae, the longest f 1 (89 µm), the shortest c 2 = c 3 (39 µm). Lyrifissures ia aligned obliquely posterior to c 3; im longitudinally, between d 3 and e 2; ip perpendicular, postero-laterally to gla; ih and ip s situated ventrally (Fig. 11 A). Relative distances: (f 1 - f 1: 571 µm)> (e 1 - e 1: 104 µm)> (d 2 - d 2: 89 µm)> (h 1 - h 1: 75 µm)> (p 1 - p 1: 68 µm)> (c 1 - c 1: 54 µm) = (d 1 - d 1). Ventral region — Genito-anal setal formula: 9 - 0 - 2 - 3; all setae spiniform; anal setae sparsely barbed, other setae smooth (Fig. 11 B). Genital (35 µm) and anal (24 µm) setae inserting at the inner margin of each plate. All adanal setae (51 µm) aligned in adanal. Lyrifissures ani and adi aligned almost longitudinally at the level setae ad 3. Subcapitulum diarthric, subcapitular setae: 1 - 2 - 1; setae h (26 µm), A (40 µm), m 1 (20 µm), m 2 (14 µm) smooth spiniform. Epimeral setal formula: 11 [9] - 5 - 6 - 5; all setae (14 µm) thin, short, smooth setiform. Epimeral borders distinct. Legs — Heterotridactylous; claw (40 µm) sparsely, minutely dentate (Fig. 11 F). Setal formula: I (1 - 9 - 5 - 6 - 26), II (1 - 9 - 5 - 5 - 26), III (4 - 6 - 5 - 5 - 22), IV (1 - 6 - 5 - 5 - 20). Solenidiotaxy: I (1 - 2 - 3), II (1 - 1 - 1), III (1 - 1 - 0), IV (1 - 1 - 0). Measurements (µm) of segments: [Rightleft] I ([71 - 61] - [164 - 164] - [89 - 89] - [79 - 82] - [148 - 154]), II ([71 - 71] - [136 - 136] - [71 - 75] - [61 - 68] - [125 - 125]), III ([71 - 71] - [100 - 100] - [64 - 61] - [61 - 57] - [129 - 139]), IV ([107 - 107] - [114 - 125] - [68 - 71] - [71 - 68] - [182 - 182]). All legs bearing one solenidion of all tibiae and genua contiguous to dorsal seta (Figs. 11 D-E). On tarsus I (Fig. 11 C), famulus ε (4 µm) spiniform situated antero-laterally to ω 1 (35 µm), posterior to solenidia ω 2; ω 2 (16 µm), postero-laterally to ω 3 (14 µm); ω 2, ω 3 thin bacilliform; ω 1 thick bacilliform, broad basally, situated anterior to seta ft ’ (67 µm). Solenidia ’ 1 and σ longer than each dorsal seta. On tibia I, solenidion ’ 1 (39 µm) and ’ 2 (16 µm) setiform, situated on every apophysis; ’ 1 contiguous to smooth spiniform seta d (26 µm), inserting anterior to d. On genu I, σ (21 µm) spiniform, contiguous to smooth cone-like seta d (10 µm) situated on a small apophysis. Remarks — The new species is similar in shape of prodorsal and notogastral setae, and notogaster with one pair of longitudinal ridges to Nothrus discifer Hammer, 1961 and T. gracilis Hammer, 1961. However, the new species differs from those species in having the longest setae not h 2 or p 1, but f 1 of all notogastral setae, and longitudinal ridges not joining posteriorly.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E37C154FE90F7BAA66CBD88.taxon	diagnosis	Diagnosis — Average body length 587 µm; width 348 µm. Prodorsal surface granulate; notogastral surface postulate of hexagonal pattern. Of all notogastral setae, the longest h 1. Solenidia ’ 1 and σ shorter than each dorsal seta. Genito-anal setal formula: 7 - 0 - 1 - 3. Subcapitulum diarthric, subcapitular setae: 1 - 1 - 1. Epimeral setal formula: 3 - 1 - 3 - 3. Homotridactyle. Material examined — Holotype (Adult female) (NSMT-Ac 13626) from sand, soil and humus at point C; 2 paratypes (NSMT-Ac 13625 and 13627): same data as holotype. Etymology — After the shape of notogastral posterior region. Measurements and body appearance — Body length (26 exs.) 464 (587) 629 µm; width (27 exs.): 307 (348) 407 µm. Body colour light yellowish brown. Prodorsal surface granulate; notogastral surface postulate with hexagonal pattern. All specimens female. Percentage of gravid females 89 % in total, having 1 to 4 eggs. Prodorsum — Triangular in dorsal view (Fig. 12 A). Rostrum rounded. Setae ro (80 µm) thin, barbed setiform inserting almost mid-distance between rostral anterior margin and insertion of lamellar setae. Setae le (71 µm) and in (95 µm) thick, long, densely barbed bacilliform. Sensilli (61 µm) with spiculate fusiform head and thin stem (Fig. 12 B). Bothridia cup-shaped, opening dorsolaterally. Setae ex (4 µm) smooth minute cone-like, inserting at the basal portion of bothridium. Relative distances: (in-in: 100 µm)> (le-in: 70 µm)> (le-le: 50 µm)> (ro-ro: 43 µm)> (ro-le: 32 µm). Notogaster — Notogaster anteriorly straight, posterior angular. Of all 16 notogastral setae including virtual f 1; the longest h 1 (77 µm), the shortest c 2 (18 µm). Setae p of series inserting ventrally (Fig. 13 A). Lyrifissures ia aligned longitudinally posterior to c 3; im obliquely between e 1 and gla; ip perpendicular to notogastral outline, lateral to f 2; ih obliquely, anterior to cp; ip s ventrally situated at the level of the insertion of setae ad 3. Relative distances: (d 2 - d 2: 177 µm)> (e 1 - e 1: 143 µm)> (e 1 - h 3: 136 µm)> (h 3 - h 3: 121 µm)> (e 1 - f 1: 107 µm)> (h 1 - h 3: 102 µm)> (d 1 - d 1: 98 µm)> (f 1 - f 1: 95 µm)> (h 1 - h 1: 93 µm)> (d 1 - e 1: 88 µm)> (p 1 - p 1: 79 µm)> (c 1 - c 1: 73 µm)> (c 1 - d 1: 54 µm). Ventral region — Genito-anal setal formula: 7 - 0 - 1 - 3; all setae thin setiform; genital setae (32 µm) densely barbed; anal (29 µm) and adanal setae (30 – 36 µm) sparsely barbed (Fig. 13 A). All adanal setae aligned in adanal. Lyrifissures ian and iad aligned obliquely at the level of anal anterior margin. Subcapitulum stenarthric, subcapitular setae: 1 - 1 - 1; setae h (13 µm), A (32 µm) sparsely roughened; m (3 µm) smooth, minute cone-like (Fig. 13 E). Epimeral setal formula: 3 - 1 - 3 - 3; all setae (16 – 32 µm) thin, short, smooth setiform; 1 c the longest; 1 A = 2 A the shortest. Epimeral borders distinct. Cheliceral setae cha (42 µm) long, barbed setiform; chb (17 µm) short, smooth, spiniform. Trägårdh’s organ (29 µm) thin. Legs — Homotridactylous; claws (43 µm) dentate dorsally (Fig. 13 B). Setal formula: I (1 - 5 - 5 - 5 - 19), II (1 - 5 - 5 - 5 - 17), III (2 - 4 - 3 - 4 - 12), IV (1 - 2 - 3 - 3 - 10). Solenidiotaxy: I (1 - 2 - 3), II (1 - 1 - 2), III (1 - 1 - 0), IV (0 - 1 - 0). Measurements (µm) of segments according to holotype: [Right-left] I ([50 -?] - [86 -?] - [41 -?] - [27 -?] - [66 -?]), II ([? -?] - [73 -?] - [41 -?] - [27 -?] - [63 -?]), Acarologia 53 (1): 41 – 76 (2013) III ([46 -?] - [63 -?] - [36 - 36] - [27 - 29] - [63 - 61), IV ([54 - 50] - [68 - 63] - [43 - 41] - [38 - 39] - [84 - 80]). Measurements (average value: µm) of segments according to depressed paratypes: [Right-left] I ([47 - 47] - [79 - 73] - [46 - 47] - [31 - 31] - [54 - 59]), II ([47 - 39] - [70 - 71] - [34 - 43] - [29 - 36] - [58 - 54]), III ([57 - 59] - [59 - 55] - [36 - 30] - [29 - 30] - [50 - 54]), IV ([54 - 54] - [63 - 63] - [36 - 36] - [32 - 38] - [71 - 84]). All legs bearing one solenidia of all tibiae and genua contiguous to dorsal seta (Figs. 13 C-D). On tarsus I, famulus ε (14 µm) obtuse situated laterally to ω 1; ω 1 (25 µm) thick bacilliform; ω 2 (25 µm) thin setiform inserting near the base of claws; ω 3 (14 µm) situated lateral to famulus and posterior to ω 2. Solenidia ’ 1 and σ shorter than each dorsal seta. On tibia I, solenidion ’ 1 (31 µm) setiform and ’ 2 (6 µm) spiniform, situated on either side of seta d (55 µm) on common apophysis (Fig. 13 C). On genu I, σ (12 µm) bacilliform, contiguous to barbed ensiform seta d (21 µm) situated on a small apophysis (Fig. 13 D). Remarks — The new species is similar in shape of notogaster, insertion of rostral setae, namely, almost mid-distance between rostral anterior margin and insertion of lamellar setae, and length of notogastral setae to Trhypochthoius tectorum (Berlese, 1896), T. septentrionalis Fujikawa, 1995 and T. fujinitaensis Fujikawa, 2000. The chelicerae of the new species are similar in appearance to those of T. semovitussi Szywilewska, 2004 and T. sphagnicola Weigmann, 1997. However, the new species differs from its conginers in having smaller body size, solenidion ω 1 located between famulus and seta ft ’ on tarsus I, setiform solenidion ω 2 on tarsus I, and sparsely barbed bacilliform dorsal seta on tibia I.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E34C153FC07F019A2B3BC5E.taxon	description	Tectocepheus elegans Ohkubo, 1981, Annot. Zool. Japon., 54 (1), pp. 42 - 52, figs. 1 - 3; Fujikawa et al., 1993, J. Acarol. Soc. Jpn., 2 (Suppl. 1), p. 57; Subías, 2004, Graellsia, 60, p. 154; Nakamura et al., 2006, Mem. Fac. Agr., Ehime Univ., p. 41. Diagnosis — Rostrum with fence-like sclerotic ridge. Sensilli consisting clavate verrucose head and thin, smooth stem. Notogaster with four pairs of large hollows medially. Material examined — One female (NSMT-Ac 13632): from point B; holotype slide NSMT-Ac 9204 bearing the label " Tectocepheus elegans OHOKUBO, 1980 " in the National Museum of Nature and Science, Tokyo. Measurements — Body length 314 µm; width 214 µm. Body colour light brown. Supplementary description — Notogastral surface covered with cerotegument of irregularly granulate. Measurements (µm) of segments of legs: [Right-left]: I ([29 -? - [23 - 23] - [18 -?] - [25 -?]), II ([36 -?] - [50 -?] - [18 - 18] - [34 - 25] - [29 - 26]), III ([29 - 30] - [36 - 36] - [16 - 18] - [34 -?] - [29 -?]), IV ([48 -?] - [39 - 39] - [18 - 21] - [39 - 40] - [36 - 36]). Distribution — Oriental region. Remaks — The present specimen differs from the holotype in shape of granules of cerotegument on notogastral surface.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E33C153FF55F6CCA750B828.taxon	diagnosis	Diagnosis — Rostral anterior margin without incision. Lamellar cuspis without dent. Setae in short. Sensilli with verrucose globular head. Bothridia with deep incision, without swelling or projection. Depressions and dorsosejugal scissure absent. Humeral region with small triangular projection. Ten pairs of notogastral setae. Genito-anal setal formula: 6 - 1 - 2 - 3. Lyrifissures iad located along, near anterior margin of anal aperture. Epimeral setal formula: 3 - 1 - 3 - 3. Trochantera III and IV bearing carina without sharply pointed apex. Monodactylous. Material examined — One female (NSMT-Ac 13633): from point B; 11 exs. and 2 parts of body, slides Nos. 596 - 604 bearing the label " Tegeocranus velatus " in the Michael collection. Measurements — Body length 336 µm; width 229 µm. Body colour light brown. Supplementary description — Morphological variation in shape of lamellar cuspis, sensillus, and situation of adanal lyrifissure iad, type B, B, and A used in figs 6 and 7 by Fujikawa (1999), respectively, that is, the present specimen has narrow cuspis, elongate sensillus, and enclosed angle of the adanae lyrifissure pair iad, 170 º. Genital plates bearing g 1 and g 2 inserting at the same level. Measurements (µm) of segments of legs: [Right-left]: I ([? - 16] - [66 - 50] - [24 - 23] - [38 - 34] - [39 - 38]), II ([? - 31] - [71 - 57] - [20 - 27] - [34 - 36] - [31 - 31]), III ([43 - 43] - [43 - 46] - [18 - 19] - [37 - 32] - [32 - 31]), IV ([50 - 53] - [50 - 54] - [21 - 23] - [46 - 43] - [37 - 36]). Distribution — Cosmopolitan. Remarks — Three specimens from Matsushimacho belonging to the genus Tectocepheus have a body surface ornamented " as with a veil " (Michael, 1880), and the ornamentation has densely, dark, large granules (Fig. 15 A). Lyrifissure im of the present specimen aligned transversely anterior to opisthonotal gland gla.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E33C15FFC43F256A186BD1B.taxon	diagnosis	Diagnosis — Average body length 282 µm; width 171 µm. Rostral anterior margin without incision, broadly rounded with rostral trowel. Lamellar cuspis with dents bilaterally to lamellar seta, not extending to level of rostral anterior margin. Setae in roughened rod-like. Sensilli with spinose, clavate head and smooth, thin stem. Bothridia with deep F – Solenidial region on right genu III; G – Chelicera (B, F: Holotype NSMT-Ac 13630; C, C, E, G: Paratype NSMT-Ac 13631). incision, without swelling or projection. Depressions and dorsosejugal scissure absent. Humeral region with small triangular projection. Ten pairs of notogastral setae. Genito-anal setal formula: 6 - 1 - 2 - 3. Lyrifissures iad located along, near anterior margin of anal aperture. Epimeral setal formula: 3 - 1 - 3 - 3. Trochantera III and IV bearing carina with sharply pointed apex. Monodactylous. Material examined — Holotype (Female) (NSMT-Ac 13630) from point B; 1 paratype (NSMT- AC 13631): same data as holotype. Etymology — After long, sharply pointed apex on trochanter III. Measurements and body appearance — Body length 279 – 286 µm; width 164 – 179 µm. Body colour light brown. Whole body surface covered with cerotegument; cerotegument irregularly dark granulate. Prodorsum — Rostral anterior margin without incision, broadly rounded with rostral trowel (Fig. 15 C). Rostral setae (16 µm) consisting of unilaterally spinose head and thick, smooth stem, inserting dorsally under lamellar cuspis. Lamellar cuspis with dents bilaterally to lamellar seta, not extending to level of rostral anterior margin; outer dent longer than inner dent. Inner sides of cuspides straight. Setae le (33 µm) setiform minutely roughened, strongly curved inside. Setae in (7 µm) minutely roughened rod-like. Sensilli (46 µm) composed of thin, roughened stem and globular head bearing dark coloured spines. Setae ex (4 µm) smooth, short bacilliform. Bothridia opening antero-laterally, with deep incision, without swelling or projection. Notogaster — Depressions and dorsosejugal scissure absent. Dorsophragmatic apophysis (hy) conspicuously long. Humeral region with small triangular projection (Fig. 15 B). Ten pairs of notogastral setae short (4 – 7 µm) setiform, roughened throughout length. Lyrifissures ia aligned longitudinally or obliquely, lateral to setae c; im obliquely, lateral to gla; ih, ip and psi perpendicular to notogastral outline. Relative distances: (h 3 - h 3: 120 µm)> (h 2 - h 2: 95 µm)> (lp - lp: 88 µm)> (h 1 - h 1: 43 µm)> (lm - lm: 34 µm)> (p 1 - p 1: 18 µm). Ventral region — Genital (31 µm) and anal (55 µm) apertures almost pentagonal in shape; distance (23 µm) between them appreciably shorter than half length of anal aperture. Genito-anal setal formula: 6 - 1 - 2 - 3; all setae smooth setiform (Fig. 16). Setae g 1 and g 2 (9 µm) not inserting at the same level at anterior margin of plates. Setae ag (8 µm) inserting latero-posterior to genital aperture. Setae ad 1 and ad 2 (10 µm) aligned in postanal position, ad 3 (7 µm) in adanal; ad 3 inserting almost at mid-distance between an 1 and an 2 (7 µm). Lyrifissures iad located along, near anterior margin of anal aperture with enclosed angle 120 - 140 º. Sternal ridge indistinct. Epimeral borders 1 - 4, sj distinct. Epimeral setal formula: 3 - 1 - 3 - 3; setae smooth, short setiform (3 – 7 µm). Subcapitulum diarthric, subcapitular setae 3 pairs, A (14 µm), m (14 µm), and h (11 µm); setae thin smooth setiform. Cheliceral setae cha (23 µm), chb (16 µm) thin, barbed setiform. Trägårdh’s organ short (14 µm), with a blunt apex (Fig. 15 G). Legs — Monodactylous; claw (21 µm) sparsely dentate dorsally. Setal formula: I (1 - 5 - 3 - 4 - 18), II (1 - 5 - 3 - 4 - 14), III (2 - 4 - 1 - 3 - 14), IV (1 - 2 - 2 - 3 - 10). Measurements (µm) of right segments according to depressed paratype: I (25 - 40 - 16 - 31 - 35), II (21 - 46 - 14 - 25 - 28), III (50 - 36 - 14 - 26 - 23), IV (51 - 43 - 21 -? -?). Trochantera III – IV and femora III-IV bearing carina; trochantera III with long, sharply pointed apex (21 µm) bending to rostral side (Fig. 16). On tarsus I, famulus ε (9 µm) consisting of a fine tip and expanded basal portion, situated antero-laterally contiguous to ω 2; ω 2 (31 µm) terminating in fine tip; ω 1 (21 µm) bacilliform situated posterior to ω 2; ft ’ (8 µm), inserting posterior contiguous to ω 1 (Fig. 15 D). Solenidion ’ 2 (15 µm) originating from a small apophysis on the tip of tibia I; ’ 1 (65 µm) situated at the base of apophysis. On genu I, solenidion σ (18 µm) terminating in a fine tip, longer than seta d (8 µm) (Fig. 15 E). All solenidia on tarsus II, tibiae II-IV and genua II-III short rod-like (σ III: 7 µm), shorter than each dorsal seta (d III: 11 µm) (Fig. 15 F). Remarks — The new species is similar to Tectocepheus minor Berlese, 1903 in shape of dentate cuspis and small enclosed angle of lyrifissure pair iad, however, the former is different from the latter in shape of bothridial ventral extension, long sharply pointed carina on trochantera III, arrangement of genital setae, and solenidia of legs (Laumann et al., 2007; Nübel-Reidelbach, 1994). The new species is similar in shape of long sharply pointed carina on trochantera III, and shape, length and situation of solenidia, famulus and seta ft ’ on tarsus I to T. kumayaensis Nakamura et al., 2010. However, the new species differs from the latter in having a round rostrum, dentate cuspides, lyrifissures im located laterally to opisthonotal gland gla and genital setae aligned almost in a line. The new species is distinguished from its congeners by shape of rostral region, cuspides, dorsosejugal region with long dorsophragmatic apophysis, humeral region, carina on trochantera III and IV, and solenidia of legs.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E3BC158FF56F13AA7CBBC8C.taxon	diagnosis	Diagnosis — Body length 464 µm; width 343 µm. Rostral tip rounded, weakly extended bending to ventral side. Prolamellae developed. Sensilli consisting of sparsely spinose, fusiform head and smooth, thin stem. Four pairs of sacculi, ten pairs of notogastral setae, five pairs of lyrifissures present on notogaster. Porose areas Ah and dosophragmata hy distinct. Material examined — Holotype (Female) (NSMT-Ac 13629); from litter and humus at point C. Etymology — After prolamella. Measurements and body appearance — Body length 464 µm; width 343 µm. Body colour light yellowish brown. Whole integument smooth. Prodorsum — Rostral tip rounded, weakly extended bending to ventral side (Fig. 20 B). Rostral setae (ro) (69 µm) inserting at lateral margins of rostrum (Fig. 20 A). Lamellae convergent, extending from bothridia to about two third-way along length of prodorsum without cuspis nor translamella; setae le (84 µm) arising at the ending of lamellae; prolamella (14 µm) distinct between insertions of le and ro. Setae ro, le extending anterior of rostrum for distance equal to half of their length. Transverse ridge absent in rostral and lamellar region. Setae in (118 µm) extending for a short distance in front of anterior rostral margin. Setae ro, le, in, ex thin setiform, sparsely, minutely barbed throughout the length, terminating in fine tips. Sensilli (91 µm) consisting of sparsely spinose fusiform head and smooth thin stem. Bothridia opening dorsal-laterally; setae ex (41 µm) inserting laterally. Porose areae Ah large, posterior to bothridia. Relative distances, (le-in: 66 µm)> (le-le) = (in-in) (63 µm)> (ro-ro: 59 µm)> (anterior margin of straighten rostrum- le) (54 µm)> (anterior margin of straighten rostrum- ro) (45 µm). Notogaster — Length as long as width; broadly triarched anteriorly, semicircular posteriorly. Dorsophragmata distinct. Ten pairs of notogastral setae present; setae thin smooth, minute; p 1 the longest (11 µm), other setae 9 µm. Four pairs of opening minute pores of sacculi present; Sa located longitudinally laterally to la, antero-laterally to lm; S 1 obliquely lateral to gla, antero-laterally to h 3; S 2 obliquely laterally to h 1; S 3 longitudinally between p 1 and p 2. Lyrifissures ia aligned obliquely anterolaterally to c; im transversely or obliquely anterolaterally to gla; ih perpendicular to notogastral outline; ip and ips situated ventrally. A number of light spots arranged peripherally on notogaster. Ventral region — Genital (59 µm in length) and anal (91 µm in length) apertures with distance (107 µm) between them, roughly circular and square, respectively (Fig. 21 A). Genito-anal setal formula 4 - 1 - 2 - 3; all setae thin, smooth, setiform except for genital setae thin, smooth, and setiform: genital setae (21 µm) sparsely minutely barbed (Fig. 20 C). Setae g 1 inserting at anterior margin of plates; setae g 4 at posterior margins. Setae ag (17 µm) inserting latero-posteriorly to genital aperture, near one third-distance between genital and anal apertures. Setae an 1 and an 2 (21 µm) inserting far from posterior and anterior margins of plates, respectively. Adanal setae (12 µm) ad 1 aligned in postanal position; ad 2 latero-posteriorly to the aperture; ad 3 in preanal. Lyrifissures iad located longitudinally at the level of mid-distance between an 2 and anterior anal margin. Posterial anal locking-pieces indistinct. All epimeral borders distinct. Epimeral setal formula: 3 - 1 - 3 - 3; setae thin, smooth and setiform except for 3 c; 3 c the longest (35 µm) bearing a few barbs; 1 A the shortest (16 µm) (Fig. 21 C). Subcapitulum diarthric, subcapitular setae 3 pairs: a (21 µm) unilatelally sparsely minutely barbed; m (15 µm) and h (31 µm) minutely barbed throughout the length. Legs — Heterotridactylous; claws (39 µm) minutely dentate dorsally. Setal formula: I (1 - 5 - 3 - 4 - 19), II (1 - 5 - 3 - 4 - 16), III (2 - 3 - 1 - 3 - 14), IV (1 - 2 - 2 - 2 - 10). Measurements (µm) of segments of legs: [Rightleft]: I ([38 - 39] - [86 - 84] - [25 - 32] - [63 - 48] - [63 - 63]), II ([36 - 32] - [86 - 80] - [18 - 21] - [48 - 41] - [54 - 52]), III ([? - 48] - [59 - 59] - [21 - 21] - [54 - 52] - [56 - 59]), IV ([? - 50] - [61 - 59] - [36 - 36] - [61 - 68] - [64 - 71]). Femora II-IV and trochantera III-IV bearing a small rounded carina. On tarsus I, famulus ε (6 µm) spiniform, situated contiguously posterior to solenidion ω 2; ω 2 (54 µm) setiform posterior to ω 1; solenidion ω 1 (25 µm) bacilliform. Seta ft ’ (30 µm) unilaterally sparsely barbed setiform, inserting posterior to ε (Fig. 21 E). Solenidion ’ 1 (107 µm) on tibia I situated contiguous to ’ 2 (45 µm) originating from a small apophysis (Fig. 21 D). Solenidion σ (63 µm) originating from a small apophysis (Fig. 21 B) Remarks — The new species differs from members of the genus Scheloribates (Scheloribates) Berlese, 1908 by having weakly protruding and bending rostral tip, fusiform sensilli, prolamella, long interlamellar setae extending in front of rostral anterior margin, and famulus contiguous, posterior to setiform solenidion ω 2 and anterior to seta ft ’ on tarsi I. The new species resembles in the shape of rostral tip, shape of sensilli, presence of prolamella and arrangement of famulus and solenidia such as found in S. (S.) azumaensis Enami et al., 1996, however, the former is different from the latter in shape of rostral extension, length of interlamellar setae, and distance of famulus and ω 2. The new species is similar to S. (S.) shigeruus Fujikawa, 2011 in shape of prolamella and length of interlamellar setae, however, the former is different from the latter in shape of tip of rostrum, shape of sensilli, and arrangement of solenidia and famulus on tarsi I.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
03F5672C4E38C165FC2BF74FA78CBAAD.taxon	diagnosis	Diagnosis — Body length 321 µm; width 250 µm. Pteromorphae lineate; other integument of body smooth. Rostrum broadly rounded with rostral trowel. Setae ro, le and in short. Sensilli fusiform, composed barbed head and thin, smooth stem. Ten pairs of short notogastral setae, four pairs of porose areas, five pairs of lyrifissures present. Movable pteromorphs bearing short furrow, c and ia. Pteromorphae notch distinct. Dorsophragmatic apophysis (hy) small circular in shape. Genito-anal setal formula: 6 - 1 - 2 - 3. Epimeral setal formula: 3 - 0 - 3 - 3. Heterotridactyl Subcapitulum diarthric, subcapitular setae. Material examined — Holotype (Female) (NSMT-Ac 13628) from litter and humus at the point C. Etymology — After rostral trowel. Measurements and body appearance — Body length 321 µm; width 250 µm. Body colour light brown. Integument on pteromorphae lineate; other body surface punctulate. Prodorsum — Rostral tip broadly rounded with rostral trowel (Fig. 22 A). Setae ro (21 µm) thin, short, smooth setiform, inserting on lateral margins of rostrum, not reaching rostral tip. Lamellar and sublamellar lines near each other actuate, parallel. Setae le (39 µm) short, setiform, minutely barbed through the length, inserting between lines L and L, extending for a short distance in front of rostral tip. Setae in (12 µm) short, thin, smooth setiform. Bothridia opening dorso-laterally. Sensilli (ss) (66 µm) composed of thin, long, smooth stem and expanded fusiform head bearing sparsely spicules. Setae ex (4 µm) short, smooth, setiform. Notogaster — Ten pairs of short notogastral setae (4 µm), six pairs of porose areas, five pairs of lyrifissures present. Anterior notogastral margin absent. Movable pteromorphs bearing short furrow, c and ia; pteromorphal notch distinct. Dorsophragmatic apophysis (hy) small circle. Porose areas (major axis, minor axis: µm) A l (9,8) located outer lateral to insertion of in; Ad (9,6) posterior to A l; Aa (9,6) outer lateral to lm; A 1 (6,5) latero-posteriorly to lp or im; A 2 (7,6) latero-posteriorly to A 1 or gla; A 3 (9,6) posterior to h 1. Distance between A 1 and A 2 about one-third as long as that between A 2 and A 3. Lyrifissures ia on pteromorphae aligned almost parallel to the joint line between pteromorpha and notogaster; im transversely latero-posteriorly to lp; ih, ip perpendicular to notogastral outline; ip between p 1 and p 2;? ips between p 2 and p 3. Opening gla situated posterior to h 3. Relative distances: (lp - lp: 38 µm)> (lm - lm: 34 µm)> (h 1 - h 1: 29 µm)> (p 1 - p 1: 14 µm). Ventral region — Genital (45 µmin length) and anal (62 µmin length) apertures almost trapezoid length of anal aperture slightly shorter than distance (66 µm) between genital and anal apertures (Figs. 23 A-B). Genito-anal setal formula: 6 - 1 - 2 - 3; all setae short, thin, smooth setiform. Setae g 1 - g 3 (9 µm) aligned at the anterior margin of the plates; other setae longitudinally in a row at the medial portion of the plates. Setae ag (7 µm) inserting posterior-lateral to genital aperture. Anal setae (5 µm) inserting far from anterior margin of anal plates. Setae ad 1 and ad 2 (4 µm) aligned in postanal position; ad 3 in adanal, at the level of almost mid-distance along the anal aperture. Lyrifissures iad located longitudinally, antero-laterally to ad 3. Epimeral borders short. Epimeral setal formula: 3 - 0 - 3 - 3; setae short, thin, smooth setiform; 1 b (4 µm) the shortest, 1 c (24 µm) the longest (Fig. 23 C). Subcapitulum bearing 3 pairs of setae; setae short, setiform; a (0.7 µm), m (11 µm) sparsely, minutely barbed; h (14 µm) roughened. Legs — Heterotridactyl; claws (23 µm) minutely dentate. Setal formula: I (1 - 5 - 3 - 4 - 19), II (1 - 4 - 3 - 4 - 16), III (2 - 3 - 1 - 3 - 15), IV (1 - 2 - 2 - 3 - 12). Measurements (µm) of segments of left legs: I (25 - 46 - 25 - 39 - 55), II (31 - 75 - 21 - 25 - 36), III (49 - 45 - 16 - 36 - 39), IV (54 - 43 - 17 - 43 - 46). On tarsus I (Fig. 22 B), famulus ε (6 µm) obtuse, situated posterior to solenidia ω 1; ω 1 (24 µm) bacilliform; ω 2 (29 µm) setiform, situated latero-posterior to ω 1; ft ’ (31 µm) smooth, setiform, inserting just lateral to ω 1. On tibia I, solenidia ’ 1 (86 µm), ’ 2 (23 µm) setiform situated at anterior portion of the segment; ’ 2 originating from a small apophysis; dorsal seta (34 µm) smooth setiform, inserting contiguous to ’ 1. On genu I, σ (69 µm) longer than seta d (11 µm). Remarks — The new species is very similar to Trichogalumna chimaera Ohkubo, 1984 in ornamentation of the pteromorphal integument, length of rostral and interlamellar setae, shape of sensilli and insertion of genital-anal setae. However, the former is different from the latter in having a round rostrum with rostral trowel, and the body surface is punctulate except for pteromorphae.	en	Nakamura, K., Nakamura, Y. - N., Fujikawa, T. (2013): Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011. Acarologia 53 (1): 41-76, DOI: 10.1051/acarologia/20132081, URL: http://dx.doi.org/10.1051/acarologia/20132081
