identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F287B9FFDE1B1EFF10FA09094E20E2.text	03F287B9FFDE1B1EFF10FA09094E20E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) okinawanus Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) okinawanus sp. n.</p><p>(Figs 1, 9, 17, 25)</p><p>Type material. Holotype: JAPAN (Okinawa Pref.): ♂, two labels: " JAPAN, OKINAWA Pref. / OKINAWA Is. FIT / OKINAWASHI, 6-9 IX 2003 / CHIBANAGUSUKU-ATO / S. &amp; K. Arai leg." [white, printed]; " MICROSCYD- MUS (s. str.) / okinawanus m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (NSMT).</p><p>Diagnosis. Body length ~ 0.9 mm; posteromedian portion of vertex with punctures larger and denser than those on surrounding areas; bristles on tempora and vertex short and sparse, barely discernible; protibia strongly thickened in apical region, with apical comb of ten robust spines that extend from apical onto mesal margin of tibia, the most proximal spine longer and of different shape than the penultimate one; aedeagus with a pair of subapical arcuate structures connected at middle and with strongly elongate lateral components flanking a long tubular structure; parameres nearly straight.</p><p>Description. Body of male (Fig. 1) slender, slightly flattened, pigmentation brown, vestiture light brown; BL 0.88 mm.</p><p>Head (Fig. 9) broadest at eyes, HL 0.18 mm, HW 0.20 mm; tempora weakly arcuate, slightly longer than eyes; posterior margin of vertex straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex covered with small and shallow but dense and distinct punctures, which on posteromesal portion of vertex are distinctly larger than those on surrounding areas; setae sparse, short and suberect, bristles on tempora and vertex short and sparse; vertex with a pair of long posterolateral setae. Antennae slender, AnL 0.35 mm, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest at middle, PL 0.23 mm, PW 0.23 mm; anterior and posterior margins weakly arcuate, sides strongly rounded, anterior and posterior pronotal corners indistinct, obtuse-angled; pronotal base with a well-defined, short transverse groove, additionally with one tiny pit near each hind corner. Punctures on disc fine and inconspicuous, smaller and shallower than those on head; setae sparse, short, suberect; lateral bristles short and sparse.</p><p>Elytra together oval, broadest slightly in front of middle, EL 0.48 mm, EW 0.33 mm, EI 1.46; humeral calli distinct; elytral base with large and deep basal impression; narrow adsutural region in anterior half of each elytron slightly impressed; apices of elytra separately rounded. Punctures about as indistinct as those on pronotum; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 17) strongly thickened in apical region, broadest at apex, with apical comb composed of ten robust teeth that occupy apical margin and extend onto mesal margin, the most basal spine strongly differs in length and shape from the penultimate one.</p><p>Aedeagus (Fig. 25) relatively stout, AeL 0.14 mm, endophallus with long median tubular component flanked by a pair of shorter elongate and nearly symmetrical sclerites in submedian region of median lobe, in subapical region the tubular component connected to a pair of long, arcuate, transverse sclerites; parameres nearly straight, each with one apical seta.</p><p>Female. Unknown.</p><p>Distribution. Japan, Okinawa-jima.</p><p>Etymology. Locotypical, after Okinawa Island and Prefecture.</p><p>Remarks. Among Japanese species, M. okinawanus is most similar to M. nishimeanus sp. n. described below. They share the protibiae strongly and abruptly thickened in distal region (either not thickened or thickened gradually, not abruptly, in all remaining Far Eastern Microscydmus); they also have a similar general body form, relatively indistinct punctures on the pronotum and elytra, but distinct on the vertex, short and sparse bristles on the tempora and vertex, and similar endophallic structures. The apical protibial comb in M. okinawanus is composed of 10 robust teeth that occupy not only the apical margin, but extend onto the mesal margin, and the most basal spine or tooth of the comb strongly differs in length and shape from the penultimate one. In M. nishimeanus, the comb is composed of only six teeth that do not extend onto the mesal tibial margin, and the most basal tooth is inconspicuous, of the same shape and length as the penultimate one. Microscydmus okinawanus is also distinctly smaller than M. nishimeanus .</p></div>	https://treatment.plazi.org/id/03F287B9FFDE1B1EFF10FA09094E20E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFDD1B1FFF10FA090D10214A.text	03F287B9FFDD1B1FFF10FA090D10214A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) oharaensis Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) oharaensis sp. n.</p><p>(Figs 2, 10, 18, 26)</p><p>Type material. Holotype: JAPAN (Okinawa Pref.): ♂, two labels: " JAPAN, OKINAWA Pref. / IRIOMOTEJIMA / ÔHARA env. / 19 IX 2003, subtr. forest / rotten wood, P. Jałoszyński " [white, printed]; " MICROSCYDMUS (s. str.) / oharaensis m. / det. P. Jałoszyński, '15 / HOLOTYPUS " [red, printed] (NSMT).</p><p>Diagnosis. Body length 0.7 mm; vertex evenly punctate; head with long, well-visible bristles, especially long on posterior margin of vertex; protibia in male slightly tapered in apical region, lacking apical comb; aedeagus with a pair of subapical round structures; parameres nearly straight.</p><p>Description. Body of male (Fig. 2) slender, slightly flattened, pigmentation brown, with head and elytra slightly darker than pronotum, vestiture light brown; BL 0.70 mm.</p><p>Head (Fig. 10) broadest at eyes, HL 0.13 mm, HW 0.16 mm; tempora weakly arcuate, slightly shorter than eyes; posterior margin of vertex slightly arcuate, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex covered with small, shallow but dense punctures with diffuse margins; setae sparse, short and suberect; bristles on tempora, and especially on posterior margin of vertex, long and relatively dense, directed posterad. Antennae slender, AnL 0.30 mm, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest distinctly in front of middle, PL 0.18 mm, PW 0.20 mm; anterior margin confluent with sides, anterior corners not marked, sides strongly rounded in anterior third and weakly so in posterior half, posteriorly strongly convergent toward base, posterior margin nearly straight; posterior corners well-defined, obtuse-angled; pronotal base with a well-defined, short transverse groove, additionally with one tiny pit near each hind corner. Punctures on disc fine and inconspicuous, similar to those on head; setae sparse, short, suberect; lateral bristles long and dense, especially in posterior half of pronotum.</p><p>Elytra together oval, broadest slightly in front of middle, EL 0.40 mm, EW 0.25 mm, EI 1.60; humeral calli distinct; elytral base with large and deep basal impression; adsutural region on each elytron not impressed; apices of elytra separately rounded. Punctures fine and inconspicuous; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 18) slender, slightly tapered in apical region, lacking apical comb.</p><p>Aedeagus (Fig. 26) moderately slender, AeL 0.13 mm, endophallus lightly sclerotized, with two round subapical structures; parameres nearly straight, slightly broadened distad, each with one apical seta.</p><p>Female. Unknown.</p><p>Distribution. Japan, Yaeyama Archipelago of the Ryukyus, Iriomote-jima.</p><p>Etymology. Locotypical, after Ôhara, a town on Iriomote Is.</p><p>Remarks. Among Japanese species, M. oharaensis externally resembles most M. yusuharanus sp. n. described below. They share a similar body form and length, and they both lack the apical protibial comb in the male. The protibial apex in the male of M. oharaensis has its apical margin transverse in relation to the long axis of tibia, and the apical spurs are inserted at the apex. In M. yusuharanus, the protibial apex forms a short, tapered lobe with its apical margin oblique in relation to the long axis of tibia, and the spurs are inserted at its base, far from the tibial apex. The tempora in M. oharaensis are only slightly shorter than eyes; in M. yusuharanus they are yet shorter. The aedeagi of these species clearly differ, both in endophallic structures, and in the relative length of parameres.</p><p>The individual that is fixed as a holotype was sifted from a hollow in a large rotten log of a deciduous tree.</p></div>	https://treatment.plazi.org/id/03F287B9FFDD1B1FFF10FA090D10214A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFDC1B19FF10FB710DFC2646.text	03F287B9FFDC1B19FF10FB710DFC2646.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) nishimeanus Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) nishimeanus sp. n.</p><p>(Figs 3, 11, 19, 27)</p><p>Type material. Holotype: JAPAN (Okinawa Pref.): ♂, three labels: " Mt. Nishimedake (No. 1) / (epiphytic fern: litter) / Kunigami-son, Okinawa Is. / [ Ryukyus: JAPAN] / 20.iv.2001, S. Karasawa " [white, printed]; " Epiphytic Fern Survey / Scydmaenidae / Microscydmus sp. 1 / P. Jałoszyński det.,2005" [printed, creamy], " MICROSCYDMUS (s. str.) / nishimeanus m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (NSMT) . Paratypes (11 exx.): 8 ♀♀ and 3 exx. of unknown sex (undissected), same data as for holotype, except for yellow PARATYPUS labels (cPJ, NSMT) .</p><p>Diagnosis. Body length 0.90–0.98 mm; posteromedian portion of vertex with punctures larger and denser than those on surrounding areas; bristles on tempora and vertex short and sparse, barely discernible; protibia strongly thickened in apical region, with apical comb of six curved teeth that occupy only apical margin, proximal teeth of the same length and shape; aedeagus with a pair of short, transverse subapical structures connected at middle, and with strongly elongated, asymmetrical lateral components flanking a long tubular structure; parameres nearly straight.</p><p>Description. Body of male (Fig. 3) slender, slightly flattened, pigmentation dark brown, vestiture light brown; BL 0.95 mm.</p><p>Head (Fig. 11) broadest at eyes, HL 0.20 mm, HW 0.23 mm; tempora weakly arcuate, about as long as eyes; posterior margin of vertex weakly arcuate, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex covered with small and shallow but dense and distinct punctures, which on posteromesal portion of vertex are slightly larger than those on surrounding areas; setae sparse, short and suberect, bristles on tempora and vertex barely discernible; vertex with a pair of long posterolateral setae. Antennae slender, AnL 0.38 mm, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest at middle, PL 0.25 mm, PW 0.28 mm; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded, except for posterior 1/5, where margins are distinctly sinuate; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove, additionally with one tiny pit near each hind corner. Punctures on disc fine and inconspicuous, smaller and shallower than those on head; setae sparse, short, suberect; lateral bristles short and sparse.</p><p>Elytra together oval, broadest slightly in front of middle, EL 0.50 mm, EW 0.38 mm, EI 1.33; humeral calli distinct; elytral base with large and deep basal impression that extends posterad as a broad, shallow and diffuse groove; narrow adsutural region in anterior half of each elytron slightly impressed; apices of elytra separately rounded. Punctures as indistinct as those on pronotum; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 19) strongly thickened in apical region, with apical comb of six curved teeth that occupy only apical margin, proximal teeth of the same length and shape.</p><p>Aedeagus (Fig. 27) moderately slender, AeL 0.13 mm, endophallus with long median tubular component flanked by a pair of shorter elongate and clearly asymmetrical sclerites in submedian region of median lobe, in subapical region the tubular component connected to a pair of short, oval transverse sclerites; parameres nearly straight, each presumably with one apical seta (seta broken off, but its socket visible).</p><p>Female. Externally similar to male, also with thickened protibiae, but lacking apical combs, most specimens lighter brown than male, punctures on vertex and frons variable, from similar to those in male to more evenly distributed and less conspicuous; BL 0.90–0.98 mm (mean 0.94 mm); HL 0.19–0.20 mm (mean 0.20 mm), HW 0.20–0.23 mm (mean 0.22 mm), AnL 0.33–0.35 mm (mean 0.34 mm); PL 0.24–0.25 mm (mean 0.25 mm), PW 0.25–0.28 mm (mean 0.27 mm); EL 0.48–0.53 mm (mean 0.50 mm), EW 0.33–0.35 mm (mean 0.34 mm), EI 1.41–1.50.</p><p>Distribution. Japan, Okinawa-jima.</p><p>Etymology. Locotypical, after Mt. Nishime (= Nishime-dake).</p><p>Remarks. This species is most similar to M. okinawanus; differences were discussed in remarks for the latter.</p></div>	https://treatment.plazi.org/id/03F287B9FFDC1B19FF10FB710DFC2646	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFDA1B1AFF10FC750B8B26FE.text	03F287B9FFDA1B1AFF10FC750B8B26FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) kochiensis Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) kochiensis sp. n.</p><p>(Figs 4, 12, 20, 28)</p><p>Type material. Holotype: JAPAN (Kôchi Pref.): ♂, two labels: "( Kôchi: JAPAN) / Kuroson / Nishitosa-mura / 23. V. 1999 / M. Sakai leg." [white, printed]; " MICROSCYDMUS (s. str.) / kochiensis m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (EUMJ) . Paratypes (3 exx.): 2 ♀♀, same data as for holotype, except for 21. VI. 1999 and 22-23. V. 1999, both Tatsuya Kan leg.; 1 ♀, " Kôchi: JAPAN / Ohdaba / Kuroson / 18. x. 1997 / M. Sakai (litter)" [white, printed]; all paratypes with yellow PARATYPUS labels (cPJ, EUMJ) .</p><p>Diagnosis. Body length ~ 0.8 mm; entire frons and vertex with distinct, deep, well-defined and very dense punctures; bristles on tempora and vertex short but well-visible; protibia not thickened in apical region, with apical comb of four teeth that occupy distomesal margin; aedeagus with conspicuous ovoid vesicular structure in sub-basal region of median lobe; parameres strongly recurved.</p><p>Description. Body of male (Fig. 4) slender, distinctly flattened, pigmentation moderately dark brown, vestiture light brown; BL 0.81 mm.</p><p>Head (Fig. 12) broadest at eyes, HL 0.15 mm, HW 0.18 mm; tempora weakly arcuate, about as long as eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small but distinct, well-marked and conspicuously dense punctures; setae sparse, short and suberect, bristles on tempora and vertex short but well-visible; long posterolateral setae on vertex not found. Antennae slen- der, AnL 0.30 mm, antennomeres I and II strongly elongate, III–X each distinctly transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest in front of middle, PL 0.21 mm, PW 0.23 mm; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove deepened at each end, lateral pit near each hind corner present, but barely discernible, very small and obscured by dense punctures. Punctures on disc similar as those on head, conspicuously dense; setae sparse, short, suberect; lateral bristles dense and relatively long, well visible.</p><p>Elytra together oval, broadest distinctly in front of middle, EL 0.45 mm, EW 0.30 mm, EI 1.50; humeral calli distinct; elytral base with large and deep basal impression; narrow adsutural region in anterior half of each elytron indistinctly impressed; apices of elytra separately rounded. Punctures dense but superficial and very small and diffuse, much less distinct than those on head and pronotum; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 20) not thickened in apical region, with apical comb of four teeth that occupy distomesal margin.</p><p>Aedeagus (Fig. 28) moderately slender, AeL 0.13 mm, endophallus with long median tubular component gradually broadened in its proximal portion to form an ovoid vesicle; parameres strongly sinuate, each with one apical seta.</p><p>Female. Externally similar to male, also with thickened protibiae, but lacking apical combs; BL 0.83–0.85 mm (mean 0.83 mm); HL 0.15 mm, HW 0.19–0.20 mm (mean 0.20 mm), AnL 0.30–0.33 mm (mean 0.32 mm); PL 0.20–0.23 mm (mean 0.22 mm), PW 0.23–0.25 mm (mean 0.24 mm); EL 0.45–0.48 mm (mean 0.47 mm), EW 0.33 mm, EI 1.38–1.46.</p><p>Distribution. Japan, southern Shikoku.</p><p>Etymology. Locotypical, after Kôchi Prefecture.</p><p>Remarks. No other Far Eastern species have so distinctly and densely punctate both the head and pronotum. The male protibial comb composed of only four teeth is similarly small and occupies a similar area on the tibial apex as those in some European species (revised and illustrated by Orousset (2017)), but the aedeagi of the latter do not have the large, sub-basal vesicular structure in their endophalli, and their parameres are nearly straight or weakly curved, whereas M. kochiensis has conspicuously recurved, sinuate parameres. The shape of parameres and the sub-basal vesicular endophallic structure are similar in M. kochiensis and M. elytratus; both species have been collected in the same area. However, despite similarities in the aedeagi, their body length is conspicuously different ( M. kochiensis is much smaller than M. elytratus), and male secondary sexual characters differ profoundly. In M. elytratus the elytra bear a posteromedian patch of modified setae (absent in M. kochiensis); and the protibial comb in M. elytratus is composed of six teeth, vs. only four teeth in M. kochiensis .</p></div>	https://treatment.plazi.org/id/03F287B9FFDA1B1AFF10FC750B8B26FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFD91B1BFF10FC2D093D278E.text	03F287B9FFD91B1BFF10FC2D093D278E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) elytratus Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) elytratus sp. n.</p><p>(Figs 5, 13, 21, 29)</p><p>Type material. Holotype: JAPAN (Kôchi Pref.): ♂, two labels: "(Kôchi: JAPAN) / Kuroson / Nishitosa-Village / 22-23. V. 1999 / Tatsuya Kan leg." [white, printed]; " MICROSCYDMUS (s. str.) / elytratus m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (EUMJ) . Paratypes (3 exx.): 3 ♀♀, same data as for holotype, except for 21. VI . 1999; all paratypes with yellow PARATYPUS labels (cPJ, EUMJ) .</p><p>Diagnosis. Body length ~ 1.1 mm; entire frons and vertex with distinct but shallow and diffuse dense punctures; bristles on tempora and vertex moderately long and dense, well-visible; protibia gradually thickened distad, with apical comb of six teeth that occupy distomesal margin; elytra between middle and posterior third with an oval circumsutural patch of long and dense setae directed toward suture; aedeagus with conspicuous spherical vesicular structure in sub-basal region of median lobe; parameres strongly recurved.</p><p>Description. Body of male (Fig. 5) moderately slender, slightly flattened, pigmentation moderately dark brown, vestiture light brown; BL 1.10–1.13 mm (mean 1.11 mm).</p><p>Head (Fig. 13) broadest at eyes, HL 0.23 mm, HW 0.25 mm; tempora weakly arcuate, slightly longer than eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small and dense but shallow and diffuse punctures; setae sparse, short and suberect, bristles on tempora and vertex relatively short but well-visible; vertex with a pair of long posterolateral setae. Antennae slender, AnL 0.49–0.50 mm (mean 0.49 mm), antennomeres I and II strongly elongate, III indistinctly transverse, IV–VI each about as long as broad, VII–X each transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest slightly in front of middle, PL 0.26–0.28 mm (mean 0.27 mm), PW 0.33 mm; anterior and posterior margins weakly arcuate; anterior corners not marked; sides strongly rounded in anterior half, in posterior half slightly sinuate; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove deepened at each end, lateral pit near each hind corner present, but barely discernible, very small and obscured by dense punctures. Punctures on disc conspicuously dense, slightly larger than those on head, but very shallow and with diffuse margins; setae sparse, short, suberect; lateral bristles dense and relatively long, well visible.</p><p>Elytra together oval, broadest distinctly in front of middle, EL 0.60–0.63 mm (mean 0.62 mm), EW 0.43–0.45 mm (mean 0.44 mm), EI 1.39–1.41; humeral calli distinct; elytral base with large and deep basal impression; adsutural area not impressed in anterior half; apices of elytra separately rounded; small oval circumsutural region between middle and posterior half slightly impressed and surrounded by dense, long setae directed toward suture. Surface of elytra finely shagreened, individual punctures not discernible; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 21) gradually thickened distad, with apical comb of six teeth that occupy distomesal margin.</p><p>Aedeagus (Fig. 29) moderately slender, AeL 0.18 mm, endophallus with long median tubular component abruptly broadened in its proximal portion to form a spherical vesicle; parameres strongly sinuate, each with one apical seta.</p><p>Female. Externally similar to male, with weakly thickened protibiae lacking apical combs, and elytra lacking posteromesal patch of modified setae; BL 1.10 mm; HL 0.23 mm, HW 0.25 mm, AnL 0.40 mm; PL 0.28 mm, PW 0.30 mm; EL 0.60 mm, EW 0.43 mm, EI 1.41.</p><p>Distribution. Japan, southern Shikoku.</p><p>Etymology. The adjective elytratus refers to the modified elytra in males.</p><p>Remarks. This is a giant among Japanese species of Microscydmus, the only one with adults distinctly exceeding 1 mm in length. The elytral patch of modified setae is unique among Palaearctic species of this genus. Similarities and differences in relation to M. kochiensis, a species with the most similar aedeagus, were discussed in remarks for the latter species.</p></div>	https://treatment.plazi.org/id/03F287B9FFD91B1BFF10FC2D093D278E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFD81B14FF10F8DB092C202B.text	03F287B9FFD81B14FF10F8DB092C202B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) tibialis Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) tibialis sp. n.</p><p>(Figs 6, 14, 22, 30)</p><p>Type material. Holotype: JAPAN (Kôchi Pref.): ♂, two labels: "( Kôchi: JAPAN) / Kuroson / Nishitosa-mura / 21. VI. 1999 / M. Sakai leg." [white, printed]; " MICROSCYDMUS (s. str.) / tibialis m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (EUMJ).</p><p>Diagnosis. Body length ~ 0.7 mm; frons and vertex with very fine, inconspicuous punctures; bristles on tempora and vertex short and sparse; protibia not thickened distad, with apical portion tapered, apical comb of nine teeth occupies apical tibial margin, teeth directed toward base of tibia; aedeagus with symmetrical, lightly sclerotized endophallus that contains two pairs of small dark droplet-like structures in sub-basal region; parameres nearly straight.</p><p>Description. Body of male (Fig. 6) slender, slightly flattened, pigmentation moderately dark brown, vestiture light brown; BL 0.69 mm.</p><p>Head (Fig. 14) broadest at eyes, HL 0.11 mm, HW 0.15 mm; tempora weakly arcuate, about as long as eyes; posterior margin of vertex straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with fine, inconspicuus punctures; setae sparse, short and suberect, bristles on tempora and vertex distinct; long posterolateral setae on vertex not found. Antennae slender, AnL 0.29 mm, antennomeres I and II strongly elongate, III–X each transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest distinctly in front of middle, PL 0.20 mm, PW 0.20 mm; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded; posterior pronotal corners distinct, obtuse-angled; posterior margin nearly straight; pronotal base with an indistinct, short and shallow transverse groove, lateral pits not discernible. Punctures on disc fine, inconspicuous; setae sparse, short, suberect; lateral bristles sparse and moderately long.</p><p>Elytra together oval, broadest slightly in front of middle, EL 0.38 mm, EW 0.25 mm, EI 1.50; humeral calli distinct; elytral base with large and deep basal impression; adsutural area indistinctly impressed in anterior half; apices of elytra separately rounded. Punctures on elytra as fine as those on head and pronotum; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 22) not thickened distad, with apical portion tapered, apical comb of nine teeth occupies apical tibial margin, teeth directed toward base of tibia.</p><p>Aedeagus (Fig. 30) slender, AeL 0.13 mm, endophallus lightly sclerotized, with an elongate cup-shaped median component containing two pairs of darkly sclerotized, small droplet-shaped structures; parameres nearly straight, each with one apical seta.</p><p>Female. Unknown.</p><p>Distribution. Japan, southern Shikoku.</p><p>Etymology. The adjective tibialis refers to the unusually shaped protibial modification in males.</p><p>Remarks. This species has the smallest adults of all Japanese Microscydmus; only M. oharaensis has a comparable body length. The aedeagus of M. tibialis has uniquely shaped endophallic structures, with four small sclerotized 'spots' between base and middle; also the protibial comb composed of nine teeth directed toward base of tibia is unusual.</p></div>	https://treatment.plazi.org/id/03F287B9FFD81B14FF10F8DB092C202B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFD71B16FF10FA5009E524D2.text	03F287B9FFD71B16FF10FA5009E524D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) yusuharanus Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) yusuharanus sp. n.</p><p>(Figs 7, 15, 23, 31)</p><p>Type material. Holotype: JAPAN (Kôchi Pref.): ♂, two labels: "[ Kôchi: JAPAN] / Kami-oriwatari / Yusuharacho / 14. vi. 1997 / M. Sakai (½ * {= Japanese kuchiki, rotten wood})" [white, printed]; " MICROSCYDMUS (s. str.) / yusuharanus m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (EUMJ).</p><p>Diagnosis. Body length ~ 0.75 mm; frons and vertex with fine and shallow but distinct, dense punctures; bristles on tempora and vertex well-visible; protibia not thickened distad, with apical portion forming a slightly protruding, short subrectangular lobe with its apical margin oblique in relation to the long axis of tibia, apical comb absent; aedeagus with a subapical group of dense, needle-like sclerites; parameres slightly recurved.</p><p>Description. Body of male (Fig. 7) moderately slender, slightly flattened, pigmentation moderately dark brown with indistinctly darker head, vestiture light brown; BL 0.74 mm.</p><p>Head (Fig. 15) broadest at eyes, HL 0.13 mm, HW 0.18 mm; tempora weakly arcuate, much shorter than eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex, except for distinctly flattened frontal area between and slightly behind supraantennal tubercles; supraantennal tubercles weakly elevated but welldefined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small, shallow but relatively distinct and dense punctures; setae sparse, short and suberect, bristles on tempora and vertex well-visible; long posterolateral setae on vertex present (only the left seta visible in the studied specimen, the other one presumably broken off). Antennae slender, AnL 0.30 mm, antennomeres I and II strongly elongate, III–X each strongly transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest distinctly in front of middle, PL 0.19 mm, PW 0.23 mm; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded in anterior half, weakly so in posterior half; posterior pronotal corners distinct, obtuse-angled; posterior margin weakly arcuate; pronotal base with distinct, relatively long transverse groove, lateral pits indiscernible. Punctures on disc fine, inconspicuous; setae sparse, short, suberect; lateral bristles sparse and moderately long.</p><p>Elytra together oval, broadest distinctly in front of middle, EL 0.43 mm, EW 0.33 mm, EI 1.31; humeral calli distinct; elytral base with large and deep basal impression; adsutural area indistinctly impressed in anterior half; apices of elytra separately rounded. Punctures on elytra dense but small, very shallow and with diffuse margins, inconspicuous; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 23) not thickened distad, with apical portion forming a slightly protruding, short subrectangular lobe with its apical margin oblique in relation to the long axis of tibia, apical comb absent.</p><p>Aedeagus (Fig. 31) moderately slender, AeL 0.10 mm, endophallus lightly sclerotized, in subapical region with a large group of dense, needle-like sclerites directed distomesad; parameres slightly sinuate, each with one apical seta.</p><p>Female. Unknown.</p><p>Distribution. Japan, southern Shikoku.</p><p>Etymology. Locotypical, after the village of Yusuhara.</p><p>Remarks. This species is most similar to M. oharaensis; similarities and differences were discussed in remarks for the latter species.</p></div>	https://treatment.plazi.org/id/03F287B9FFD71B16FF10FA5009E524D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
03F287B9FFD51B17FF10FDF90D2725AA.text	03F287B9FFD51B17FF10FDF90D2725AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microscydmus (Microscydmus) omogonus Jałoszyński 2019	<div><p>Microscydmus (Microscydmus) omogonus sp. n.</p><p>(Figs 8, 16, 24, 32)</p><p>Type material. Holotype: JAPAN (Ehime Pref.): ♂, two labels: "(SHIKOKU) / Omogo-kei / Ehime Pref. / 26. V. 1969 / M. Sakai leg." [white, printed]; " MICROSCYDMUS (s. str.) / omogonus m. / P. Jałoszyński, 2019 / HOLOTYPUS " [red, printed] (EUMJ).</p><p>Diagnosis. Body length ~ 0.8 mm; frons and vertex with fine and shallow but distinct, dense punctures; bristles on tempora and vertex well-visible; protibia not thickened distad, unmodified, with apical margin transverse in relation to the long axis of tibia; aedeagus with asymmetrical tubular endophallic structure in submedian region; parameres weakly curved.</p><p>Description. Body of male (Fig. 8) moderately slender, slightly flattened, pigmentation moderately dark brown, vestiture light brown; BL 0.83 mm.</p><p>Head (Fig. 16) broadest at eyes, HL 0.13 mm, HW 0.18 mm; tempora weakly arcuate, distinctly shorter than eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex, except for distinctly flattened frontal area between and slightly behind supraantennal tubercles; supraantennal tubercles weakly elevated but welldefined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small, shallow, moderately distinct, dense punctures; setae sparse, short and suberect, bristles on tempora and vertex distinct; long posterolateral setae on vertex not found.Antennae slender, AnL 0.36 mm, antennomeres I and II stron- gly elongate, III–X each strongly transverse, XI about as long as broad.</p><p>Pronotum in dorsal view nearly circular, broadest distinctly in front of middle, PL 0.23 mm, PW 0.23 mm; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded in anterior half, weakly so in posterior half; posterior pronotal corners distinct, obtuse-angled; posterior margin weakly arcuate; pronotal base with distinct, relatively long transverse groove, lateral pits very small, barely discernible. Punctures on disc fine, inconspicuous; setae sparse, short, suberect; lateral bristles sparse and moderately long.</p><p>Elytra together oval, broadest distinctly in front of middle, EL 0.48 mm, EW 0.33 mm, EI 1.46; humeral calli distinct; elytral base with large and deep basal impression; adsutural area indistinctly impressed in anterior half; apices of elytra separately rounded. Punctures on elytra more distinct than those on pronotum, very dense, but very shallow and with diffuse margins; setae short, sparse and nearly recumbent.</p><p>Protibia (Fig. 24) not thickened distad, unmodified, with apical margin transverse in relation to the long axis of tibia.</p><p>Aedeagus (Fig. 32) slender, AeL 0.13 mm, endophallus with asymmetrical long median tubular component funnel-like broadened in its proximal portion; parameres curved in proximal half, nearly straight in distal half, each with one apical seta.</p><p>Female. Unknown.</p><p>Distribution. Japan, northern Shikoku.</p><p>Etymology. Locotypical, after Omogo Gorge (= Omogo-kei).</p><p>Remarks. Microscydmus omogonus has the aedeagus most similar to that of its Far Eastern congener, M. pusio Kurbatov, 2006 that occurs on the Kunashir Is. (a disputed territory of the southern Kuril Islands, until 1945 belonging to Japan, currently to Russia). The endophallus in these species is clearly asymmetrical and contains a recurved tubular component broad proximally and narrowing distad. In M. pusio this structure is longer than half length of the median lobe, nearly parallel-sided in the proximal third, and gradually narrowing; in M. omogonus this structure is as long as about 1/3 of median lobe, and its proximal portion forms a funnel-like, inversely subconical and abrupt broadening. Among Japanese species, the unmodified protibia makes M. omogonus similar to M. oharaensis, but aedeagi of these species clearly differ, and M. omogonus is distinctly larger than its Ryukyuan relative.</p></div>	https://treatment.plazi.org/id/03F287B9FFD51B17FF10FDF90D2725AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2019): Unexpected diversity of Microscydmus Saulcy & Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4679 (3): 539-552, DOI: 10.11646/zootaxa.4679.3.7
