identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F287C3FFADFFD5FF55FF31E4CCFDF1.text	03F287C3FFADFFD5FF55FF31E4CCFDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homalotrichus yana Pérez & Rodríguez & Asenjo 2018	<div><p>Homalotrichus yana Pérez, Rodríguez &amp; Asenjo, sp. nov.</p><p>(Figs. 5A–I)</p><p>Type material (3♂♂, 3♀♀). Holotype: PERU: ♂, labeled “ PERU: CU[Department of Cuzco], Canchis, / Llutuyo, 14°3'5.04''S, 71°/ 7'28.33''W, 4456 m [altitude in relation to sea level], / 15.i [January].2018, bofedal [high Andean wetlands], L. Pérez / leg.”; “ HOLOTYPE [red label] / Homalotrichus yana / Pérez, Rodríguez &amp; Asenjo / Desig. Pérez et al., 2018” (MUSM) . Paratypes (5): 1♂, labeled “ PERU: CU, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.8322&amp;materialsCitation.latitude=-13.246201" title="Search Plazi for locations around (long -71.8322/lat -13.246201)">Calca</a>, / 13°14'46.32''S, / 71°49'55.92''W, 4070 m [altitude in relation to sea level], / 12.xii [December].2016, under stone / in high Andean wetlands, / L. Pérez leg.” (MUSM) . 1♂, labeled: “ PERU: JU[Department of Junin], Laguna / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.99972&amp;materialsCitation.latitude=-10.871944" title="Search Plazi for locations around (long -75.99972/lat -10.871944)">Yanacocha</a>, 10°52'19"S, / 75°59'59"W, 4130 m [altitude in relation to sea level],/ 17.iv [April].2012, J. Grados &amp; I. / Galindo leg.”(FMNH) . 3 ♀♀, labeled “ PERU: CU[Department of Cuzco], Canchis, / Llutuyo, 14°3'5.04''S, 71°/ 7'28.33''W, 4456 m [altitude in relation to sea level], / 15.i [January].2018, bofedal [high Andean wetlands], L. Pérez / leg.” (MUSM), all paratypes labeled “ PARATYPE [yellow label] / Homalotrichus yana / Pérez, Rodríguez &amp; Asenjo / Desig. Pérez et al., 2018”.</p><p>Diagnosis. H. yana sp. nov. can be differentiated from other species of Homalotrichus by the combination of the following characters: head, antennae, pronotum and abdomen black and elytra dark brown (Fig. 5A–B), distinctive shape of head, pronotum and elytra (Fig. 5C), pronotal impressions kidney-shaped, tergum VII with a triangular process near posterior margin more or less lamellar on males (Fig. 5A–B, D) and the internal sclerites of aedeagus as in Figs. 5G–I.</p><p>Description. Holotype male (BL: 3.48; BW: 1.05).</p><p>Head, antennae, pronotum, and abdomen black, elytra dark brown. Mesocoxae, mesotrochanter, femur and tibiae dark brown, first to fifth tarsomeres brown. Mouthparts light brown, labial palps, maxillary palps and mandibles dark brown (Fig. 5A–B).</p><p>Head hexagonal, wider than long (HW: 0.67; HL: 0.48) with anterior margin straight, head width about 11.8× eye width. Neck narrower than head (NW: 0.48). Cephalic disk (Fig. 5A) with micro-punctation spread heterogeneously forming smooth areas. One macrosetae near anterior edge of eye, one macrosetae near posterior edge of eye and one macrosetae near posterior margin of head (Fig. 5C). Eyes medium sized, longer than wide (YL: 0.15 YW: 0.06) located laterally in middle. Antennae longer than length of head and pronotum together, antennomere 1 the longest, slightly incrassate apically, 2–3 elongate and strongly incrassate apically, 4 slightly longer than wide, antennomere 5 subquadrate, from 5 on broader than 2–4 and with distinct basal ridge, antennomere 10 wider than length (width:length = 0.13:0.11), terminal antennomere about 1.4× longer than wide.</p><p>Pronotum (Figs. 5A–C) wider than long (PW: 0.84; PL: 0.64), anterior and posterior margin straight, lateral margin slightly convex and narrowing to base. Pronotal lateral margin irregular. Pronotal disc with setigerous punctures separated irregularly. Two pronotal impressions kidney-shaped near posterior margin. Four macrosetae near each lateral margin and one small setae on each posterior corner, four macrosetae near anterior margin in middle and two macrosetae near posterior margin in middle (Fig. 5C). Scutellum subtriangular with rounded apex and with micro-punctations (Fig. 5A, C).</p><p>Elytra (Figs. 5A–C) longer than wide (EL: 1.01; EW: 1.05) with apex wider than base. Elytron with five well defined lines of punctures and scattered punctures near outer lateral margin. Elytron with line of six setae near inner lateral margin, line of five setae near outer lateral margin, line of six setae very close to outer lateral margin.</p><p>Legs. Procoxae conical narrowly separated by prosternal process. Mesocoxae contiguous, elongated and oblique separated by mesosternal process. Metacoxae transverse, with a conical region that articulates with trochanter. Protrochanter, mesotrochanter and metatrochanter triangular. Profemora, mesofemora and metafemora long, mesofemora tapering slightly towards its extreme. Protibiae with 4 spines and 2 setae arranged in one defined row in external-lateral margin, mesotibiae and metatibiae with spines. Tarsi 5-segmented, tarsomere 1–4 small as long as wide (0.03; 0.04). Tarsomere 5 longer than wide (0.17; 0.04).</p><p>Abdomen margined (Fig. 5A). Terga IV–VII (morphological segments) with subbasal carinae complete. Segments III-VI with two laterosternites per segment, VII with one laterosternite (Fig. 5D). Tergum VII with a triangular process more or less lamellar in midline, near posterior margin (Fig. 5D). Tergum VIII (Fig. 5E) with posterior margin emarginated, trapezoidal shape with three rows of setae arranged horizontally. Row 1 and 3 with four macrosetae each, situated in middle and posterior region respectively. Row 2 with two macrosetae, one macroseta on both lateral margins. Sternite VIII (Fig. 5F) with a lobe on middle of posterior margin, 3 rows of setae arranged horizontally. Row 1 and 3 with four macrosetae situated in middle and posterior region respectively. Row 2 with four macrosetae, two macrosetae on both lateral margins. Tergum IX visible in ventral view.</p><p>Aedeagus, symmetrical, median lobe bulbous at base, narrow in its middle part. Parameres thin, curved, not exceed the length of aedeagus. In ventral view (Fig. 5G), parameres fused basally. In lateral view (Fig. 5H), parameres with acute projection near base. In dorsal view (Fig. 5I), base of median lobe with an oval membranous area. Internal sclerites as in Figs. 5G–I.</p><p>Female. Similar to male, except tergum VII without a triangular process that is more or less lamellar in midline.</p><p>Habitat. According to the collection labels, the specimens were collected in pitfall traps and under a stone, in high Andean wetlands.</p><p>Distribution. Known from Llutuyo and Calca in Cuzco and Laguna Yanacocha in Junin from 4070 m to 4456 m (Fig. 7).</p><p>Etymology. The specific name “ yana ” means black in Quechua and it refers to the species’ body color. This is a noun in apposition.</p><p>Remarks. The number of spines on the protibiae varies between 4–5 in the paratypes.</p></div>	https://treatment.plazi.org/id/03F287C3FFADFFD5FF55FF31E4CCFDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pérez, Luis;Rodríguez, Maryzender;Asenjo, Angélico	Pérez, Luis, Rodríguez, Maryzender, Asenjo, Angélico (2018): Two new species of Homalotrichus Solier, 1849 (Coleoptera: Staphylinidae) from Peru. Zootaxa 4459 (2): 376-386, DOI: 10.11646/zootaxa.4459.2.10
03F287C3FFAFFFDAFF55FD0EE4DEFDAC.text	03F287C3FFAFFFDAFF55FD0EE4DEFDAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homalotrichus shapra Pérez & Rodríguez & Asenjo 2018	<div><p>Homalotrichus shapra Pérez, Rodríguez &amp; Asenjo, sp. nov.</p><p>(Figs. 6A–J)</p><p>Type material (3♂♂, 3♀♀). Holotype: PERU: ♂, labeled “ PERU: CA[Department of Cajamarca], Rio Chonta, / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.57904&amp;materialsCitation.latitude=-7.008275" title="Search Plazi for locations around (long -78.57904/lat -7.008275)">Cerro Negro</a>, 7°0'29.79"S, / 78°34'44.55"W, 3656 m [altitude in relation to sea level], / 29-30.xi [November].2010. F. Chávez / leg.”; “ HOLOTYPE [red label] / Homalotrichus shapra / Pérez, Rodríguez &amp; Asenjo / Desig. / Pérez et al., 2018” (MUSM) . Paratypes (5): 2♂♂, 1♀, labeled “ PERU: CA[Department of Cajamarca], Caserio / Chugurmayo, 6°53'38"S, / 78°19'4.9"W, 3626 m [altitude in relation to sea level], / 20.xi [<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.31802&amp;materialsCitation.latitude=-6.893889" title="Search Plazi for locations around (long -78.31802/lat -6.893889)">November</a>].2012” (1♂ and 1♀ MUSM; 1 ♂ FMNH) . 2♀♀, labeled “ PERU: CA[Department of Cajamarca], Rio Chonta, / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.57904&amp;materialsCitation.latitude=-7.008275" title="Search Plazi for locations around (long -78.57904/lat -7.008275)">Cerro Negro</a>, 7°0'29.79"S, / 78°34'44.55"W, 3656 m [altitude in relation to sea level], / 29-30.xi [November].2010. F. Chávez / leg.” (MUSM); all paratypes labeled “ PARATYPE [yellow label] / Homalotrichus shapra / Pérez, Rodríguez &amp; Asenjo / Desig. Pérez et al., 2018”.</p><p>Diagnosis. H. shapra sp. nov. is easily differentiated from other species of Homalotrichus by the combination of the following characters: head width about 8× eye width (Fig. 6A,C), pronotum convex, lateral margin strongly convex and narrowing to base and pronotal impressions oval-shaped (Fig. 6A,C). The male is easily differentiated from males of other species by not having a marked sexual dimorphism: absence of triangular process on tergum VII (Fig. 6A–B) and the absence of a huge curved horn on tergite VIII.</p><p>Description. Holotype male (BL: 3.80; BW: 1.08).</p><p>Head, antennae, pronotum, and abdomen black, elytra dark brown. Mouthparts light brown, labial palps, maxillary palps dark brown. Coxae and trochanter dark brown. First to fifth tarsomeres brown (Figs. 6A–B).</p><p>Head hexagonal, wider than long (HW: 0.67; HL: 0.46) with anterior margin straight, head width about 8× eye width. Neck narrower than head (NW: 0.49). Cephalic disk (Fig. 6A) with micro-punctation spread heterogeneously forming smooth areas. One macroseta near anterior edge of eye, one macroseta near posterior edge of eye and one macroseta near posterior margin of head (Fig. 6C). Eyes large and prominent, longer than wide (YL: 0.184; YW: 0.08) located laterally in middle part. Antennae longer than length of head and pronotum together, antennomere 1 the longest, slightly incrassate apically, 2–3 elongate and strongly incrassate apically, 4 slightly longer than wide, antennomere 5 subquadrate, from 5 on broader than 2–4 and with distinct basal ridge, antennomere 10 wider than length (width:length = 0.14:0.13), terminal antennomere about 1.4× longer than wide.</p><p>Pronotum convex (Figs. 6A–C) wider than long (PW: 0.85; PL: 0.64), margin anterior slightly convex and posterior margin straight, lateral margin irregular, strongly convex and narrowing to base. Pronotal disc with setigerous punctures spread irregularly. Two pronotal impressions oval-shaped near posterior margin. Three macrosetae near each lateral margin and two setae near each posterior corner. Two pairs of macroseta near anterior margin. Two macrosetae near posterior margin. Scutellum subtriangular with rounded apex and with micropunctation (Figs. 6A,C).</p><p>Elytra (Figs. 6A–C) longer than wide (EL: 1.15; EW: 1.11) with apex wider than base. Elytron with five well defined lines of punctures and scattered punctures near outer lateral margin. First longitudinal line with five setae near inner lateral margin, line of seven setae near outer lateral margin, line of six setae very close outer lateral margin.</p><p>Legs. Procoxae conical narrowly separated by prosternal process. Mesocoxae contiguous, elongated and oblique, separated by mesosternal process. Metacoxae transversal, with conical region that articulates with trochanter. Protrochanter triangular, mesotrochanter triangular, and metatrochanter triangular. Profemora, mesofemora and metafemora long, mesofemora tapering slightly towards its extreme. Left protibiae with a defined row of four spines and two setae in external-lateral margin, right protibiae with well defined row of five spines and two setae in external-lateral margin, mesotibiae and metatibiae with spines. Tarsi 5-segmented, tarsomere 1–4 small, as long as wide (0.03:0.04). Tarsomere 5 longer than wide (0.20; 0.04).</p><p>Abdomen margined (Fig. 6A). Terga III-VII (morphological segments) with subbasal carinae complete. Segments III-VI with two laterosternites per segment, segment VII with only one laterosternite per segment. Tergum VII without a triangular process that is more or less lamellar at midline, near posterior margin. Tergum VIII emarginate, trapezoidal with 16 macrosetae, 6 arranged near each lateral margin and 4 macrosetae in middle forming a rectangle (Fig. 6D). Sternite VIII (Fig. 6E) with membranous lobe on posterior margin, 6 macrosetae in each lateral-posterior third. Tergum IX visible in ventral view, many setae in distal part (dorsal view). Sternite IX an elongate plate.</p><p>Aedeagus symmetrical, with median lobe bulbous at base, narrow in middle part. Parameres thin, curved, not exceed length of aedeagus. In ventral view (Fig. 6F), parameres fused basally. In lateral view (Fig.6G), parameres with acute projection near base. In dorsal view (Fig. 6H), base of median lobe with an oval membranous area. Internal sclerites as in Figs. 6F–H.</p><p>Female similar to male, except sternite IX (Fig. 6J) divided perpendicularly and forming two plates with many setae in distal part and absence of acute projection in tergum IX (Fig. 6I)</p><p>Habitat. Unknown.</p><p>Distribution. Known from Chugurmayo and Cerro Negro localities, Cajamarca, northern region of Peru, at 3626 m and 3656 m respectively (Fig. 7).</p><p>Etymology. The specific name “ shapra ” means hair in Quechua. It refers to the large number of setae on sternite VIII. This is a noun in apposition.</p><p>Remarks. The number of spines on the protibiae varies between 4 – 5 in the paratypes.</p></div>	https://treatment.plazi.org/id/03F287C3FFAFFFDAFF55FD0EE4DEFDAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pérez, Luis;Rodríguez, Maryzender;Asenjo, Angélico	Pérez, Luis, Rodríguez, Maryzender, Asenjo, Angélico (2018): Two new species of Homalotrichus Solier, 1849 (Coleoptera: Staphylinidae) from Peru. Zootaxa 4459 (2): 376-386, DOI: 10.11646/zootaxa.4459.2.10
03F287C3FFA0FFDAFF55FD70E441FC82.text	03F287C3FFA0FFDAFF55FD70E441FC82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homalotrichus nahuelbutensis Coiffait & Saiz 1968	<div><p>Homalotrichus nahuelbutensis Coiffait &amp; Sáiz, 1968</p><p>CHILE: VIII Región, Hualpen, 10. xii. 1971, T. Cekalovic leg. (1♀ VMDC).</p></div>	https://treatment.plazi.org/id/03F287C3FFA0FFDAFF55FD70E441FC82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pérez, Luis;Rodríguez, Maryzender;Asenjo, Angélico	Pérez, Luis, Rodríguez, Maryzender, Asenjo, Angélico (2018): Two new species of Homalotrichus Solier, 1849 (Coleoptera: Staphylinidae) from Peru. Zootaxa 4459 (2): 376-386, DOI: 10.11646/zootaxa.4459.2.10
03F287C3FFA0FFDAFF55FCE4E3FCFC56.text	03F287C3FFA0FFDAFF55FCE4E3FCFC56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homalotrichus striatus Solier 1849	<div><p>Homalotrichus striatus Solier, 1849</p><p>CHILE: V Región, 20 km E. Algarrobo, prov. San Antonio, 4.i.1992, G. Gonzalez leg (1♀ VMDC). VI Región, Tanumé, 20 km N. Pichilemu, 18–26.x.1993, G. Arriagada leg. (1♀ VMDC). VIII Región, Chillán Viejo, Fundo Las Cruces, 18–20.xii.2000, pitfall trap, G. Arriagada leg. (1♀ VMDC).</p></div>	https://treatment.plazi.org/id/03F287C3FFA0FFDAFF55FCE4E3FCFC56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pérez, Luis;Rodríguez, Maryzender;Asenjo, Angélico	Pérez, Luis, Rodríguez, Maryzender, Asenjo, Angélico (2018): Two new species of Homalotrichus Solier, 1849 (Coleoptera: Staphylinidae) from Peru. Zootaxa 4459 (2): 376-386, DOI: 10.11646/zootaxa.4459.2.10
03F287C3FFA0FFDAFF55FC50E57DFBA2.text	03F287C3FFA0FFDAFF55FC50E57DFBA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Homalotrichus substriatus Kraatz 1859	<div><p>Homalotrichus substriatus Kraatz, 1859</p><p>CHILE: V Región, 20 km E. Algarrobo, prov. San Antonio, 4.i.1992, G. Gonzalez leg. (1♀, 1♂ VMDC).</p></div>	https://treatment.plazi.org/id/03F287C3FFA0FFDAFF55FC50E57DFBA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pérez, Luis;Rodríguez, Maryzender;Asenjo, Angélico	Pérez, Luis, Rodríguez, Maryzender, Asenjo, Angélico (2018): Two new species of Homalotrichus Solier, 1849 (Coleoptera: Staphylinidae) from Peru. Zootaxa 4459 (2): 376-386, DOI: 10.11646/zootaxa.4459.2.10
