taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488920/files/figure.png	https://doi.org/10.5281/zenodo.4488920	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488928/files/figure.png	https://doi.org/10.5281/zenodo.4488928	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488924/files/figure.png	https://doi.org/10.5281/zenodo.4488924	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488930/files/figure.png	https://doi.org/10.5281/zenodo.4488930	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488932/files/figure.png	https://doi.org/10.5281/zenodo.4488932	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488934/files/figure.png	https://doi.org/10.5281/zenodo.4488934	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488938/files/figure.png	https://doi.org/10.5281/zenodo.4488938	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488940/files/figure.png	https://doi.org/10.5281/zenodo.4488940	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A2BFFB9FE9B04EE3655BF1C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488942/files/figure.png	https://doi.org/10.5281/zenodo.4488942	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A29FFB7FC0A01963680B69C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488924/files/figure.png	https://doi.org/10.5281/zenodo.4488924	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A29FFB7FC0A01963680B69C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488928/files/figure.png	https://doi.org/10.5281/zenodo.4488928	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A29FFB7FC0A01963680B69C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488930/files/figure.png	https://doi.org/10.5281/zenodo.4488930	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A29FFB7FC0A01963680B69C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB7FC0D019630E2B2A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488920/files/figure.png	https://doi.org/10.5281/zenodo.4488920	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB7FC0D019630E2B2A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488932/files/figure.png	https://doi.org/10.5281/zenodo.4488932	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB7FC0D019630E2B2A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB7FF4200C334EFB0F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488930/files/figure.png	https://doi.org/10.5281/zenodo.4488930	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	Fig. 4. Male genitalia of Thysanarthria species. A–E – T. atriceps (Régimbart, 1903) from Zambia: 20 km W of Gwanda, coll. NMPC (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. atriceps from Republic of South Africa: 8 km NEE of Stanford, coll. NMPC (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–O – T. brittoni Balfour-Browne, 1951, holotype (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – detail of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB7FF4200C334EFB0F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB5FC4504E13540B4DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488934/files/figure.png	https://doi.org/10.5281/zenodo.4488934	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A27FFB5FC4504E13540B4DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB5FF7102D63009B6BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488932/files/figure.png	https://doi.org/10.5281/zenodo.4488932	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB5FF7102D63009B6BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB5FC2400F6317DB1B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488932/files/figure.png	https://doi.org/10.5281/zenodo.4488932	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	Fig. 5. Male genitalia of Thysanarthria species, holotypes.A–E – T.brincki Hebauer, 2001 (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view); F–J – T. cardamona sp. nov. (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view); K–M – T. bengalensis Hebauer, 2001 (K – photo of the aedeagus before re-mounting; L – whole aedeagus in ventral view; M – whole aedeagus in lateral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB5FC2400F6317DB1B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB3FC0C07CA34DDB2E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488934/files/figure.png	https://doi.org/10.5281/zenodo.4488934	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A25FFB3FC0C07CA34DDB2E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A23FFB0FF6C072130C0B1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488924/files/figure.png	https://doi.org/10.5281/zenodo.4488924	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A23FFB0FF6C072130C0B1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488928/files/figure.png	https://doi.org/10.5281/zenodo.4488928	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A23FFB0FF6C072130C0B1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488938/files/figure.png	https://doi.org/10.5281/zenodo.4488938	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A23FFB0FF6C072130C0B1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A20FFB1FC5A07C035C9B19F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488940/files/figure.png	https://doi.org/10.5281/zenodo.4488940	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A20FFB1FC5A07C035C9B19F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A21FFB1FF5C079737B0B3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488920/files/figure.png	https://doi.org/10.5281/zenodo.4488920	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A21FFB1FF5C079737B0B3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488940/files/figure.png	https://doi.org/10.5281/zenodo.4488940	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A21FFB1FF5C079737B0B3DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A21FFAFFC0D05D6350EB41C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488940/files/figure.png	https://doi.org/10.5281/zenodo.4488940	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A21FFAFFC0D05D6350EB41C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3FFFAFFEA902163141B5FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488920/files/figure.png	https://doi.org/10.5281/zenodo.4488920	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3FFFAFFEA902163141B5FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488944/files/figure.png	https://doi.org/10.5281/zenodo.4488944	Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view).	Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3FFFAFFEA902163141B5FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3FFFACFC39020E3696B192.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488942/files/figure.png	https://doi.org/10.5281/zenodo.4488942	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3FFFACFC39020E3696B192.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3CFFADFF6307EC31ACB65C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488920/files/figure.png	https://doi.org/10.5281/zenodo.4488920	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3CFFADFF6307EC31ACB65C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488942/files/figure.png	https://doi.org/10.5281/zenodo.4488942	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 9. Male genitalia of Thysanarthria species, holotypes. A–E – T. saurahana sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. siamensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3CFFADFF6307EC31ACB65C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3DFFABFC3300A53552B25C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488944/files/figure.png	https://doi.org/10.5281/zenodo.4488944	Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view).	Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3DFFABFC3300A53552B25C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3DFFABFC3300A53552B25C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488938/files/figure.png	https://doi.org/10.5281/zenodo.4488938	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3DFFABFC3300A53552B25C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488940/files/figure.png	https://doi.org/10.5281/zenodo.4488940	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFABFEAC045637D0B35C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488934/files/figure.png	https://doi.org/10.5281/zenodo.4488934	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	Fig. 6. Male genitalia of Thysanarthria species, holotypes (A–I, K–O) and paratype (J). A–E – T.bifida sp. nov. (A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. trifida sp. nov. (F–H – whole aedeagus of the holotype in dorsal, lateral and ventral view; I – details of parameres of the holotype; J – same of the paratype). K–O – T. ceylonensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFABFEAC045637D0B35C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFA8FB9805F03024BF1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488946/files/figure.png	https://doi.org/10.5281/zenodo.4488946	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFA8FB9805F03024BF1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4549144/files/figure.png	https://doi.org/10.5281/zenodo.4549144	Fig. 12. Chaetarthria sp. from Saudi Arabia. A–C – aedeagus (A – dorsal, B – lateral, C – ventral). D–E – habitus (D – dorsal, E – lateral). F – details of parameres and apical part of the median lobe in ventral view. G – male tergite VIII; H – male sternite VIII; I – male sternite IX; J–K – elytral setae, SEM micrographs (J – general view; K – detail).	Fig. 12. Chaetarthria sp. from Saudi Arabia. A–C – aedeagus (A – dorsal, B – lateral, C – ventral). D–E – habitus (D – dorsal, E – lateral). F – details of parameres and apical part of the median lobe in ventral view. G – male tergite VIII; H – male sternite VIII; I – male sternite IX; J–K – elytral setae, SEM micrographs (J – general view; K – detail).	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFA8FB9805F03024BF1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488948/files/figure.png	https://doi.org/10.5281/zenodo.4488948	Figs 12A–C, F–I with Figs 2N–S). External characters support this assignment: dorsal setae are simple and cut-off apically (compare Fig. 12K with Fig. 2d), elytra lack longitudinal striae except for sutural stria, and pronotum and elytra are yellowish in color. The specimens above seem to stand close to the Argentinian species C. argentina Miller, 1974 and C. hermani Miller, 1974 of the C. atra group defined by MILLER (1974). When compared with the genitalia drawings and descriptions provided by MILLER (1974) it seems that the specimens examined here represent an undescribed species. We are however leaving it undescribed, as it is likely introduced and a more detailed comparison with the American species would be necessary to diagnose the species properly. The presence of the species which is clearly an element of Neotropical fauna in Saudi Arabia is very unexpected. The first author discussed the problem with both collectors (J. Bezděk and D. Král) and with the person who	Figs 12A–C, F–I with Figs 2N–S). External characters support this assignment: dorsal setae are simple and cut-off apically (compare Fig. 12K with Fig. 2d), elytra lack longitudinal striae except for sutural stria, and pronotum and elytra are yellowish in color. The specimens above seem to stand close to the Argentinian species C. argentina Miller, 1974 and C. hermani Miller, 1974 of the C. atra group defined by MILLER (1974). When compared with the genitalia drawings and descriptions provided by MILLER (1974) it seems that the specimens examined here represent an undescribed species. We are however leaving it undescribed, as it is likely introduced and a more detailed comparison with the American species would be necessary to diagnose the species properly. The presence of the species which is clearly an element of Neotropical fauna in Saudi Arabia is very unexpected. The first author discussed the problem with both collectors (J. Bezděk and D. Král) and with the person who	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
03F387C57A3BFFA8FB9805F03024BF1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4488924/files/figure.png	https://doi.org/10.5281/zenodo.4488924	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale.	2019-06-20	Fikáček, Martin;Liu, Hsing-Che		Zenodo	biologists	Fikáček, Martin;Liu, Hsing-Che			
