identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F0879A1633FFAAFEA1FD542429FCCA.text	03F0879A1633FFAAFEA1FD542429FCCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes Bates 1873	<div><p>Amphimenes Bates, 1873</p> <p>Bates, 1873: 322; Jedlička, 1963: 366; Habu, 1964: 472; 1967: 113; 1982: 90; Fedorenko, 2010: 18; 2014: 310.</p> <p>Type-species: Amphimenes piceolus Bates, 1873 (by monotypy).</p> <p>REDESCRIPTION. Unnecessary here, except for some features not mentioned before, including the following leg characters: (1) profemur with a ventrobasal tubercle, (2) mesotrochanter ventrally (posteriorly) with a very fine longitudinal ridge, (3) mesotibia minutely tuberculate along apical 1/2–2/3 inner margin; and (4) a pointed tubercle at middle of its wider, mesal, part of the metacoxa. For details see below.</p> <p>COMMENTS. Fedorenko [2010] arranged all species into five species groups: the rugulipennis -group, the piceolus -group, the medius -group, the rufipes -group, and the planicollis -group; the former two groups are arboricolous and the remainder include soil-dwelling species. The planicollis - group was then [Fedorenko, 2014] subdivided into the nominal subgroup and the brunneus -subgroup, with the rufipes - group being considered as a subgroup of the planicollis - group. All these groupings are treated as separate species groups below, except that the subgenus Amphinemes subg.n. is introduced for the rugulipennis -group.</p> <p>Out of the generic characters mentioned above, the profemoral tubercle is more or less distinct in A. medius (the medius -group), A. femoralis sp.n. (the brunneus -group), three of four species of Amphinemes subg.n., and A. (Amblops) marginicollis sp.n.</p> <p>The mesotrochanteral ridge adjoins or almost adjoins apical trochanteral seta in almost all Amphimenes species for which males are known. It is longest yet slightly tuberculate apically in A. reflexicollis while more or less shortened and/or modified into a minute preapical tubercle to denticle in most other congeners, being combined with vestigial or totally reduced seta in all species but two. These have no ridge/ tubercle, but the seta is well-developed (A. planipennis) or barely shortened (A. femoralis sp.n.). This may suggest that the seta was gradually reduced as the ridge developed in course of morphological evolution. Accordingly, the pattern observed in A. planipennis and A. femoralis sp.n. could be recognized as plesiomorphous for the genus, and a minute tubercle or nearly indistinct ridge — since the two characters are combined with vestigial or missing seta — as secondary reductions.</p> <p>The mesotibial tubercles are minute and range between species in number, from mostly four (A. bicoloripes sp.n., A. femoralis sp.n., A. konplongensis sp.n., A. marginicollis sp.n., A. medius, A. micros, A. planicollis, A. reflexicollis) or five (A.acutipennis sp.n., A. basipunctatus sp.n., A. planipennis, A. similis sp.n.) through three (A. gracilis, A. maculatus, A. ruficollis sp.n.) to one (A. piceus, A. subcostatus sp.n.) or none (A. giganteus).</p> <p>Most species have the umbilical seta series (USS) consisting of 15 setae/setigerous pores (US) that runs on elytral interval 9; these are arranged into two groups, anterior, or posthumeral (nine setae included), and posterior, or postmedian (six setae). The USS groundplan seems to include these groups more or less widely separated, with US 6, US 8 and US 11 approximating to or touching stria 9 (the piceolus - group). The members of the subgenus Amblops are similar, except for setigerous pores being large, barely smaller than the width of interval 9. The subgenus Amphinemes subg.n. is farther advanced due to the anterior and posterior pore groups more widely gaped in its members, with US 6, US 8 and US 11 adjoining stria 9.</p> <p>In most soil-dwelling species, USS apparently evolved toward a despecialized condition defined by US becoming continuous or almost so, less oscillating across interval 9, and additional US appeared. Most of these species have retained 15 US, whereas the rest have US increased to either 17–18 (A. basipunctatus sp.n., A. reflexicollis) or 16–19 (A. konplongensis sp.n.), or 20–21 (A. acutipennis sp.n.) in number. But A. giganteus which is closely related to A. acutipennis sp.n. has retained nearly groundplan USS pattern.</p> </div>	https://treatment.plazi.org/id/03F0879A1633FFAAFEA1FD542429FCCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A1635FFA1FC32FACE209FFE2C.text	03F0879A1635FFA1FC32FACE209FFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblops piceus Andrewes 1931	<div><p>The piceus species group.</p> <p>2.1. Amphimenes (Amblops) piceus</p> <p>(Andrewes, 1931), comb.n.</p> <p>Figs 1, 27–28, 32.</p> <p>Andrewes, 1931: 521 (Amblops; Dehra Dun, North India).</p> <p>MATERIAL. Two syntypes (BMNH): ♂ with four labels: ‘Dehra Dun, Jharipani 5,222 ft. ’, ‘Bought from Staudinger &amp; Bang- Haas, 1930’, ‘Type’ [red], ‘ Amblops piceus, Type Andr.[hw], H.E.Andrewes det.’; ♂ mounted on back, with same labels, except for ‘Cotype’ (white circle margined with green) substituted for the fourth label. The former syntype is here designated as lectotype. Also, digital image of a syntype at https://carabidae.org.</p> <p>REDESCRIPTION. Unnecessary, except for the following additions: Body (Fig. 1). Pronotum broadest 0.29 times from apex, with median part of base distinctly produced and 2.67 times as wide as lateral lobes; PW/PL 1.48–1.53 (1.51, n=3), PW/HW 1.20–1.24 (1.23), PB/PA 1.37–1.40. EL/EW 1.40–1.47 (1.43), EW/PW 1.55–1.60 (1.58); D1/EL 0.24, D2/EL 0.63–0.66 (0.65), D3/EL 0.92–0.94 (0.93). Mesotibia with a minute preapical fissure instead of tubercles. Tarsomere 5 with one pair of long ventral setae in apical half.</p> <p>Aedeagus (Figs 27–28, 32): median lobe in ventral view barely wider a third from base than just distal to basal bulb; apex rounded and fairly small.</p> <p>DISTRIBUTION. Known from a few localities in the environs of Dehra Dun, northern India.</p> <p>HABITATS AND HABITS. Andrewes [1931] reported that most of the syntypes were taken under bark, including eleven under bark of Pinus longifolia.</p> <p>2.2. Amphimenes (Amblops) subcostatus</p> <p>Fedorenko, sp.n.</p> <p>Figs 2, 29–30, 34.</p> <p>MATERIAL. Holotype ♀ (ZMMU) labelled: ‘ Vietnam, Dak Lak Province, Chu Yang Sin Nat [io]n[al]. Park, 12°25 ' 25 '' N / 108°21 ' 53 '' E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.36472&amp;materialsCitation.latitude=12.423612" title="Search Plazi for locations around (long 108.36472/lat 12.423612)">Krong Kmar</a> riv[er]., upper flow, h= 970 m, 15– 30.V.2014, D. Fedorenko leg.’. Paratype ♂ (SIEE):‘ Laos, Prov [ince]. Vientiane, env. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.433334&amp;materialsCitation.latitude=18.92" title="Search Plazi for locations around (long 102.433334/lat 18.92)">Van Vieng</a>, 18°55 ' 12 '' N / 102°26 ' E, h=~ 230 m, at light, 7–9.XI.2015, leg. I.Melnik’.</p> <p>DESCRIPTION. Body (Fig. 2) macropterous, small, BL 5.2–5.4 mm, shiny black. Mouthparts, basal four antennomeres, legs, including coxae, as well as sternum between coxae and in front of metacoxae, reddish-yellow; tibiae and tarsi indistinctly infuscated while mandibles, hypomera, reflexed margins of both pronotum and elytra barely more so; metaventrite and abdomen reddish-brown; antennomeres 5– 11 dark brown. Head and pronotum with isodiametric microsculpture, more superficial on vertex and neck, nearly granulate before, granulate along pronotal apex and base; elytra with superficial yet distinct microsculpture consisting of very transverse yet large meshes.</p> <p>Head: eyes large and hemispherical; genae nearly indistinct. Posterior supra-ocular seta on a level with posterior margin of eye. Antennae short, not quite reaching basal angles of pronotum.</p> <p>Pronotum quadrate, slightly transverse, PW/PL 1.49– 1.54, PW/HW 1.24–1.26, wider at base than at apex, PB/PA 1.39–1.41, broadest a third from apex. Sides rounded in apical two thirds, indistinctly angulate at and straight on each side of anterolateral seta, gently sinuate in front of basal angles, which are obtuse and rounded at tips. Base with median part convex, distinctly produced beyond and twice as wide as lateral lobes; these slightly oblique forward and rounded at basal angles; basal bead obsolete in the middle. Apex evenly concave and very finely beaded; apical angles blunt and lightly projecting forward. Disc very convex, finely and densely rugulose. Lateral margin explanate and moderately reflexed, narrow in apical half, slightly wider basad. Median line entire and deep. Both transverse impressions, basal and apical, shallow. Basal foveae small, deep and rounded. Paramedian foveae very small and nearly imperceptible.</p> <p>Elytra nearly square, EL/EW 1.45–1.51, EW/PW 1.46– 1.48, broadest at middle, nearly parallel-sided in middle third, with humeri and posterolateral angles widely rounded. Apical truncature oblique and barely sinuate; extreme apices blunt. Striae deep and finely punctate; intervals convex, more so before apex. D1/EL 0.21–0.23 (0.22), D2/EL 0.60–0.64 (0.62), D3/EL 0.92–0.94 (0.93).</p> <p>Metepisternum long.</p> <p>Tarsomere 5 with one pair of ventral setae close to apex. In male, profemur and mesotrochanter without tubercle, mesotibia with one preapical tubercle at inner margin.</p> <p>Aedeagus (Figs 28–29, 33): as in A. piceus, except for median lobe subtriangular (vs. subequally wide) in ventral view owing two a large rounded swell on left side, with apical lamella distinctly wider and a little upturned.</p> <p>DIAGNOSIS. Very similar to A. piceus, being distinctive in having the elytra slightly longer and more convex, with striae deep, inner intervals convex and outer intervals very so. Aedeagus is differently shaped as well.</p> <p>DISTRIBUTION. Southern Vietnam and Laos.</p> <p>NAME. Refers to the convex to very convex elytral intervals.</p> <p>HABITATS AND HABITS. The female specimen was taken under exfoliated bark scales of a living tree in a broad-leaved monsoon forest.</p> <p>2.3. Amphimenes (Amblops) marginicollis Fedorenko, sp.n.</p> <p>Figs 3, 25–26, 36.</p> <p>MATERIAL. Holotype ♂ (ZMMU) and paratype ♂ (SIEE) labelled: ‘N-Vietnam, Nghe An Prov., Que Phong Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.79639&amp;materialsCitation.latitude=19.755278" title="Search Plazi for locations around (long 104.79639/lat 19.755278)">Pu Hoat National Park</a>, 19°45 ' 19 '' N / 104°47 ' 47 '' E, h= 840 m 15–27.V.2019, leg. D.Fedorenko’.</p> <p>DESCRIPTION. As for the previous species, except as follows: Body (Fig. 3) larger, BL 5.9–6.4 mm. Mouthparts (labrum excluded), antennae, legs, reflexed margins of both pronotum and elytra reddish yellow. Underside red within prothorax, along middle of meso- and metaventrites, with abdomen reddish medially, otherwise dark brown.</p> <p>Antennae reaching basal angles of pronotum.</p> <p>Pronotum large and transverse, PW/PL 1.57–1.61, PW/ HW 1.40–1.48, half wider at base than at apex, PB/PA 1.47– 1.50, broadest two thirds from apex. Sides rounded in apical three fourths, with basal basal angles nearly right and blunt. Median part of base half wider than lateral lobes. Explanate lateral margin wide in apical half and very wide in basal half. Median line entire and deep. Basal transverse impression deep, apical one shallow medially, obliterate laterally. Basal foveae small, deep and rounded. Paramedian foveae missing.</p> <p>Elytra a bit more rounded on sides, EL/EW 1.44–1.48, EW/PW 1.35–1.36. Intervals convex in basal third, along sides and close to apex, otherwise inner four or five nearly flat. D1/EL 0.21–0.25 (0.23), seta d1 closer to apex, D2/EL 0.68–0.71 (0.70), D3/EL 0.93–0.95 (0.93). USS anterior (10 US) and posterior (6 US) groups setae slightly separated to almost continuous, consisting of 9+1+6 setae.</p> <p>Tarsomere 5 with two pairs of ventral setae. In male, profemur with a distinct, small and blunt tubercle; mesotibia with four preapical tubercles at inner margin.</p> <p>Aedeagus (Figs 25–26, 36) similar to that of A. absensacidus, except chiefly for median lobe strongly and abruptly desclerotized, i.e., membranous medioventrally.</p> <p>DIAGNOSIS. Similar to A. absensacidus, except that pronotal sides are sinuate (vs. convex) in front of almost straight (vs. obtuse) basal angles, and aedeagus distinctive.</p> <p>DISTRIBUTION. Northern Vietnam.</p> <p>NAME. Refers to the pale and widely explanate lateral margins of the pronotum.</p> <p>HABITATS AND HABITS. Both specimens were taken at night, running on trunks of smaller living trees in a broad-leaved monsoon forest.</p></div> 	https://treatment.plazi.org/id/03F0879A1635FFA1FC32FACE209FFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A1638FFA1FEB0FE2E2598FD57.text	03F0879A1638FFA1FEB0FE2E2598FD57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes ruficollis Fedorenko 2019	<div><p>The ruficollis species group.</p> <p>This group is introduced here for the only species distinctive in having the body smaller, clypeus and labrum pale, elytral pattern variegated, elytra with striae subpunctate and only two non-foveate discal setigerous pores, d2 and d3.</p> <p>2.4. Amphimenes (Amblops) ruficollis</p> <p>Fedorenko, sp.n.</p> <p>Fig. 4.</p> <p>MATERIAL. Holotype ♂ (ZMMU): ‘S Vietnam, Lam Dong Prov [ince]., Bi Doup-Nui Ba Nat [ure]. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.67889&amp;materialsCitation.latitude=12.178889" title="Search Plazi for locations around (long 108.67889/lat 12.178889)">Res</a> [erve]., env[irons]. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.67889&amp;materialsCitation.latitude=12.178889" title="Search Plazi for locations around (long 108.67889/lat 12.178889)">Long Lanh</a>, h= 1400 m, 12°10 ' 44 '' N, 108°40 ' 44 '' E, window-trap h= 25 m, 6–10.V.2009, leg. A.Polilov’.</p> <p>DESCRIPTION. Body (Fig. 4) macropterous and very small, BL 4.5 mm. Elytra, mesosternum and metapleura black, base and apex of elytra brownish, head dark brown in posterior half, reddish before, propleura and abdomen brown laterally, ventral side otherwise reddish yellow; mandibles red; pronotum and narrow lateral margin of elytra reddish-yellow; palps, antennae, legs and widely explanate lateral margin of pronotum (including hypomera), yellow. Dorsum shiny, head and pronotum slightly dull from rather coarse isodiametric, nearly granulate, microsculpture; sides and apex of pronotal disc with granulate microsculpture becoming slightly transverse on each side of median line behind the middle; elytra with superficial yet distinct microsculpture consisting of slightly to moderately transverse meshes, with admixture of isodiametric meshes here and there. Elytral intervals smooth.</p> <p>Head: eyes slightly flattened; genae very short. Posterior supra-ocular seta on a level with posterior margin of eye. Antennae short, surpassing pronotal base by apical half of segment 11.</p> <p>Pronotum transverse, PW/PL 1.59, PW/HW 1.55, evenly rounded on sides, slightly less so in basal third, broadest two fifths from apex. Base broader than apex, PB/PA 1.39, with median part very slightly projecting caudad; lateral lobes rather strongly rounded close to basal angles; these almost rounded off. Apex deeply sinuate; apical angles porrect, subrectangular, with rounded tips. Disc moderately convex, with very fine transverse wrinkles. Lateral margin explanate and moderately reflexed, becoming increasingly wide from very narrow apically to very wide basally. Median line moderately deep, obliterate behind basal transverse impression, which is well defined and very obtusely angulate forward. Basal foveae indistinct, merged into rather wide and smoothly concave lateral grooves. Paramedian foveae very small and nearly indistinct.</p> <p>Elytra nearly square, slightly oval, EL/EW 1.44, EW/PW 1.38, evenly rounded on sides, broadest at middle, with humeri widely rounded yet distinct; posterolateral angles highly obtuse and widely rounded; apices blunt. Apical truncature oblique, indistinctly sinuate. Lateral reflexed margin very narrow just medially, otherwise reduced to a finest bead. Striae rather shallow, crenulate; intervals flat. Interval 3 with two setae, D2/EL 0.56, D3/EL 0.94. USS continuous, consisting of 15 US.</p> <p>Metepisternum 1.45 times as long as wide.</p> <p>Tarsomere 5 of pro-, meso- and metatarsi with one or 1– 2, or two pairs of ventral setae, respectively. In male, profemur with no tubercle, mesotrochanter with a minute tubercle; mesotibia with three preapical tubercles along inner margin.</p> <p>Aedeagus (lost through mounting): median lobe long, narrow, subequally wide in apical three fourths, almost straight in lateral view, with a well defined apical orifice in almost left position; apex moderately long, tapered, rounded apically, not upturned.</p> <p>DIAGNOSIS. As for the group.</p> <p>DISTRIBUTION. Known from the type locality only.</p> <p>NAME. Refers to the colour of the pronotum.</p> <p>HABITATS AND HABITS. The only specimen has been collected by a window-trap at 25 m above ground level in a lowland monsoon broad-leaved forest.</p></div> 	https://treatment.plazi.org/id/03F0879A1638FFA1FEB0FE2E2598FD57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A1638FFA7FC4CFBD023DCFDF0.text	03F0879A1638FFA7FC4CFBD023DCFDF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes piceolus Bates 1873	<div><p>The piceolus species group.</p> <p>The group includes six arboricolous species, five macropterous and one, A. asahinai from Taiwan, brachypterous; the latter may form the subgroup or group of itself. Four species included, three from Vietnam (A. bidoupensis, A. gracilis, A. montanus) and A. ryukyuensis from southern Japan and Taiwan are very similar to one another and thence hard to discriminate. Smaller details of aedeagus, including those of internal sac, are useful for the purpose. Among them, the following four need to be briefly described (Figs. 39–50): (1) dc, preapical dorsal carina; (2) dcl, shorter left dorsolateral carina, diverging from or from near the base of dc; (3) dorsal groove between dc and dcl; and (4) ppl, left preapical protuberance just outside the base of dcl. Dcl does mark the external margin of the dorsal groove, therefore it looks like carina only when the aedeagus is viewed laterally; the deeper dorsal groove (and/or the more prominent ppl) the more distinct dcl.</p> <p>3.1. Amphimenes (s. str.) bidoupensis</p> <p>Fedorenko, 2010</p> <p>Figs 39, 42–44.</p> <p>Fedorenko, 2010: 23, 27 (Lam Dong Province, southern Vietnam).</p> <p>MATERIAL. Type series; additional material includes many specimens, ♂♂ and ♀♀ (SIEE, D. Fedorenko leg.), from Central Vietnam: Gia Lai, ~ 50 km N of An Khe, Kon Chu Rang Nature Reserve, 14°312 N 108°32 ' E, h= 1000–1040 m, 24.V–2.VI. 2016; Kon Tum, Kon Plong Distr., 14°43 ' N, 108°19 ' 10 '' E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.31944&amp;materialsCitation.latitude=14.716666" title="Search Plazi for locations around (long 108.31944/lat 14.716666)">Dak Khe River</a>, h= 1030 m, 8–23.IV.2015; same locality, except for 14°45 ' N 108°18 ' 10 '' E, env.ngock Boc 1 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.30278&amp;materialsCitation.latitude=14.75" title="Search Plazi for locations around (long 108.30278/lat 14.75)">Mt.</a>, h= 1200–1300 m, 8–23.IV.2015; Quang N am, Nam Gian District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.37889&amp;materialsCitation.latitude=15.571111" title="Search Plazi for locations around (long 107.37889/lat 15.571111)">Song Thanh National Park</a>, 15°34 ' 16 '' N 107°22 ' 44 '' E, h= 1030 m, 23.IV–11.V.2019; same data except 15°33 ' 48 '' N 107°23 ' 22 '' E, h= 1050 m. Aedeagus examined in 13 males.</p> <p>DIAGNOSIS. Elytral intervals flat behind middle of disc. Body (BL 6.7–8 mm) and thence aedeagus (Figs 39, 42–44) barely larger than in the other two species from Vietnam in general. Median lobe in lateral view subequally wide between basal curve and apical orifice, with ppl indistinct, dc fine, low, not quite reaching apical orifice; dcl diverging from dc, straight and rather short, about half as long as distance between base of dc and apical orifice, sometimes vestigial or no. Everted and inflated internal sac (Figs 42–44) with apical bulb (a) apical in position; right lateral bulb (lr) large, curved, directed basad and contiguous to fairly large dorsomedial bulb (dm); left lateral bulb (ll) large.</p> <p>DISTRIBUTION. The records listed are within the species range that extends from Quang Binh Province in Central Vietnam to Lam Dong Province in southern Vietnam.</p> <p>3.2. Amphimenes (s. str.) gracilis</p> <p>Fedorenko, 2010</p> <p>Figs 40, 45–47.</p> <p>Fedorenko, 2010: 23, 30 (env. Thai Nguen, northern Vietnam).</p> <p>MATERIAL. Type series. Additional material (SIEE; D. Fedorenko leg.): Northern Vietnam: ♀, N ghe An, Que Phong Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.79639&amp;materialsCitation.latitude=19.755278" title="Search Plazi for locations around (long 104.79639/lat 19.755278)">Pu Hoat National Park</a>, 19°45 ' 19 '' N 104°47 ' 47 '' E, h= 840 m, 15– 27.V.2019; 3♂♂, 2♀♀, same data, except 19°46 ' 35 '' N 104°48 ' 13 '' E, h= 1350 m; ♀, Cao Bang, 40 km W of Cao Bang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.88528&amp;materialsCitation.latitude=22.600834" title="Search Plazi for locations around (long 105.88528/lat 22.600834)">Phia Oac — Phia Den Nat. Park</a>, Phia Oac Mt, 22°36 ' 03 '' N 105°53 ' 07 '' E, h= 1300 m, 3–11.X.2018. Aedeagus examined in five males.</p> <p>DIAGNOSIS. Elytral intervals convex (type specimens) to nearly flat behind middle of disc (the other specimens examined). Aedeagus (Figs 40, 45–47) smaller following a bit smaller body, BL 6.2–7.6 mm. Median lobe in lateral view more robust, distinctly narrower basally than preapically, broadened toward apical orifice in third fourth, with apical orifice more transverse; dc conspicuous, high and reaching apical orifice; dcl starting from near it, longer and more or less sinuate, with a small ppl at base; dorsal groove deep.</p> <p>41</p> <p>Everted and inflated internal sac (Figs 45–47) with a on right side, lr small, dm very small and separate from lr; an additional, small, basal dorsomedial bulb (dmb) present.</p> <p>DISTRIBUTION. Northern Vietnam (Thai N guen, Cao Bang, N ghe An).</p> <p>HABITATS AND HABITS. Under bark scales of live trees (N ghe An) and under bark of a dead tree (Cao Bang) in rainforests.</p> <p>3.3. Amphimenes (s. str.) montanus Fedorenko, 2010</p> <p>Figs 41, 48–50.</p> <p>Fedorenko, 2010: 23, 30 (env. Sa Pa, northern Vietnam).</p> <p>MATERIAL. Type specimens and additional material as follows: 20♂♂, 14♀♀ (SIEE, D. Fedorenko leg.), Northern Vietnam: Lao Cai, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat National Park, 22°37 ' –37.6 ' N 103°37.32–38.3 ' E, h= 1600–1900 m, 16–21.X.2018; ♂, same data, except for 22°36 ' 31 '' N 103°37 ' 23 '' E, h= 2000–2100 m, 4–14.VI.2019; ♂, same data except 22°36 ' 02 '' N 103°37 ' 13 '' E, h= 2200 m; 5♂♂, ♀, same data except 22°37 ' 36 '' N 103°37 ' 32 '' E, h= 1850 m. Aedeagus examined in seven males.</p> <p>DIAGNOSIS. Aedeagus (Figs 41, 48–50) similar to that of A. bidoupensis, except that dc and dcl are well-developed, higher and longer, dc reaching apical orifice, dorsal groove deeper, and small ppl present. Everted and inflated internal sac (Figs 48–50) with a on right side; lr large and long, much longer than wide at base; dm vesicular and medium-sized.</p> <p>DISTRIBUTION.This species is only known from Hoang Lien mountain ridge in northern Vietnam. The locality listed above is the second exact locality, next to Mt. Phang Xi Pang (= Fansipan).</p> <p>HABITATS AND HABITS. Common under bark of standing dead trees in cloudy forests.</p> <p>COMMENTS. Based on the material listed, the following additions and corrections to the species description are made: BL 6.2–7.6 mm. PW/PL 1.30–1.45 (1.37, n=5), 1.49– 1.61 (1.54), PB/PA 1.43–1.53 (1.48), EL/EW 1.43–1.50 (1.48), EW/PW 1.42–1.55 (1.49), D1/EL 0.20–0.27 (0.23, n=5×2), D2/EL 0.67–0.76 (0.72). Besides, the specimens listed share entirely pale legs, whereas the type specimens has the legs with femora infuscated. This, firstly, suggests the type specimens may only represent a local population or a high-montane form of the species and, secondly, reduces differences of the species from A. bidoupensis and A. gracilis to the following ones: the pronotum barely larger, PW/HW 1.49–1.61 (vs. 1.34–1.50), with sides not or barely converging towards the base, the base broader relative to the apex, PB/PA 1.43–1.53 (vs. 1.31–1.43), and the reflexed lateral margin being basally wider than in the related species; the elytra are less square due to due to their narrower bases and both humeri and sides more rounded.</p> <p>3.4. Amphimenes (s. str.) ryukyuensis Habu, 1964</p> <p>Habu, 1964: 474, 475 (Ishigaki Is., Ryukius, Japan); 1967: 114, 116; 1982: 91; Fedorenko, 2010: 23. — carinacaulis Hunting et Yang, 2019: 10, 20 (Taiwan), syn.n.</p> <p>MATERIAL. No specimen, except for digital images of the holotype of A. ryukyuensis at NIAES, Institute for Agro-Environmental Sciences, Tsukuba, Japan (http://www.naro.affrc.go.jp).</p> <p>DISTRIBUTION. Southern Japan: Satsunans (Amamioshima Is. and Tokunoshima Is.), Ryukius (Iriomote Is. and Ishigaki Is.).</p> <p>COMMENTS. The above synonymy is based on the fact that A. carinacaulis has been described and, surprisingly, compared with the species of Amphimenes from Taiwan only, instead of with similar and closely related species outside the island. Of these, A. ryukyuensis is widespread in southern Japan, including Iriomote Is. and Ishigaki Is., these being much closer to Taiwan than to Satsunan Islands Amamioshima and Tokunoshima as other known localities of the species. I see no difference between the two species in body appearance and the shape of aedeagus. Accordingly I consider the two names as conspecific until evidence to the contrary.</p> </div>	https://treatment.plazi.org/id/03F0879A1638FFA7FC4CFBD023DCFDF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A163EFFA5FEA6FDE42403FC67.text	03F0879A163EFFA5FEA6FDE42403FC67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes medius Fedorenko 2010	<div><p>The medius species group.</p> <p>The group includes five apterous soil-dwelling species, two from southern Vietnam, two from Central Vietnam, and one from northern Vietnam. Elytral seta d1 inserted close to base is their main distinctive feature.</p> <p>3.5. Amphimenes (s. str.) acutipennis</p> <p>Fedorenko, sp.n.</p> <p>Figs 6, 11–12, 31.</p> <p>MATERIAL. Holotype ♂ (ZMMU): ‘ Vietnam, Kon Tum Province, Kon Plong Distr [ict]., 14°45 ' N / 108°18210 '' E, env. ngock Boc I Mt., h= 1200–1300 m, 8–23.IV.2015, D. Fedorenko leg.’. Paratypes (ZISP, ZMMU, SIEE): 6♂♂, 3♀♀, same data; ♂, ♀, same data except 14°44 ' 20 '' N / 108°19 ' E, h= 1100–1200 m; 2♂♂, 3♀♀, same data, except for 14°45 ' N 108°17 ' 51 '' E, h= 1300–1400 m, 8–10.VI.2016; ♂, ♀, ‘ Vietnam, Gia Lai Province,~ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.33&amp;materialsCitation.latitude=14.219166" title="Search Plazi for locations around (long 108.33/lat 14.219166)">40 km ENE of Pleiku</a>, 14°13 ' 09 '' N / 108°19 ' 48 '' E, Kon Ka Kinh Nat [io]n[al]. Park, h= 1470 m, 21– 30.V.2017, D. Fedorenko leg.’. Aedeagus examined in three males.</p> <p>DESCRIPTION. Body apterous, large and slender (Fig. 6), BL 10.1–11.9 mm. Black; mouthparts, including labrum, anterior half to two thirds of clypeus, antennae, tarsi and trochanters red; lateral margins of elytra and extreme lateral margins of pronotum barely translucent with reddish-brown (more distinctly translucent under spotlight). Sometimes antennomeres 3 and 4 infuscated. Microsculpture isodiametric, distinct yet slightly superficial on head, very coarse, granulate-aciculate over pronotum. Elytra with transverse, mostly moderately transverse, microsculpture becoming isodiametric on explanate lateral margins. As a result, head and elytra moderately shiny, much shinier than dull pronotum. Elytral cross-striated sculpture dense and conspicuous in basal half, becoming increasingly weak and sparse apicad until nearly indistinct before apex.</p> <p>Head: eyes slightly flattened, twice as long as genae, these smoothly extended into neck. Posterior supra-ocular seta clearly behind posterior margin of eye (about 1/6 eye length distant from eye). Antennae very long, surpassing base of pronotum by about apical 4.5 antennomeres.</p> <p>Pronotum subcordate, broadest at anterolateral seta a fourth from apex, only a fifth wider than long, PW/PL 1.18–1.23 (mean 1.20, n=5), PW/HW 1.45–1.48 (1.47). Base a third broader than apex, PB/PA 1.28–1.37 (1.34), straight medially, with lateral lobes evenly convex caudad, half as wide as and slightly produced beyond median part. Basal angles highly obtuse yet mostly distinct. Apex deeply sinuate; apical angles porrect, slightly acute and sharp. Sides evenly rounded in apical two thirds, gently sinuate behind. Disc slightly convex, with fine and dense transverse wrinkles. Lateral margins rather strongly reflexed, narrow anteriorly, moderately wide in basal half; lateral grooves fairly wide and more or less flat at bottom. Median line entire and moderately deep; basal transverse impression well defined, angulate or convex forward. Basal foveae deep, round, merged into reflexed lateral margin and thence fairly wide; each as a very shallow yet fairly wide groove reaching middle or just apical margin. Paramedian foveae as small and very shallow to indistinct pits or short longitudinal impressions a third from apex and equidistant from median line and lateral groove.</p> <p>Elytra elliptic, EL/EW 1.41–1.61 (1.53), EW/PW 1.40– 1.52 (1.44), oblique from mesothoracic peduncle, with humeri rounded off, evenly rounded on sides, broadest at middle. Apical truncature oblique, deeply sinuate, more so close to acute and pointed posterolateral angles. Apices rectangular combined, with extreme tips slightly separate and sharp to blunt. Lateral margin very widely explanate behind the level of seta d1 and flat in horizontal plane. Striae deep; intervals convex. Interval 3 with three setae, D1/EL 0.16–0.18 (0.17, n=5×2), D2/EL 0.63–0.72 (0.67), D3/EL 0.91–0.94 (0.93).</p> <p>Metepisternum 1.03–1.04 times as long as wide.</p> <p>Tarsomere 5 with two pairs of ventral setae. Male profemur without tubercle, mesotrochanter with a small tubercle; mesotibia 5-tuberculate along inner margin.</p> <p>Aedeagus (Figs 11–12, 31): as in A. giganteus, except that median lobe in lateral view is much broader medially than just behind basal bulb (vs. subequally wide), with apex in ventral view being slightly more curved to the left.</p> <p>DIAGNOSIS. The largest species within the genus. It is similar to A. giganteus, except that that is smaller, with elytral base convex forward and outer angles obtuse and apically rounded, etc.</p> <p>DISTRIBUTION. Known from two fairly close localities in central Vietnam (Gia Lai, Kon Tum).</p> <p>NAME. Refers to acute posterolateral angles of the elytra.</p> <p>HABITATS AND HABITS. This species occurs on the underside of logs and smaller tree fragments on the ground in monsoon forests at 1100–1470 m altitude. It lives syntopically with A. konplongensis sp.n. and/or A. basipunctatus sp.n.</p> <p>3.6. Amphimenes (s. str.) basipunctatus</p> <p>Fedorenko, sp.n.</p> <p>Figs 7, 17–18, 33.</p> <p>MATERIAL. Holotype ♂ (ZMMU): ‘ Vietnam, Kon Tum Prov [ince]., Chu Mom Ray National Park, 14°30 ' N / 107°42 ' 30 '' E, ngock <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.70833&amp;materialsCitation.latitude=14.5" title="Search Plazi for locations around (long 107.70833/lat 14.5)">To Lum mt.</a>, h= 1100–1350 m 28.III–4.IV.2015, D. Fedorenko leg.’. Paratypes (ZISP, ZMMU, SIEE), 7♂♂, 6♀♀, same data; ♀ (SIEE), Vietnam, Kon Tum Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.2975&amp;materialsCitation.latitude=14.75" title="Search Plazi for locations around (long 108.2975/lat 14.75)">Kon Plong Distr.</a>, 14°45 ' N / 108°17 ' 51 '' E, env. ngock Boc 1 Mt., h= 1300–1400 m, 8–10.VI.2016, D. Fedorenko leg.; 2♂♂, 2♀♀, ‘ Vietnam, Gia Lai Province, ~ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.33&amp;materialsCitation.latitude=14.219166" title="Search Plazi for locations around (long 108.33/lat 14.219166)">40 km ENE of Pleiku</a>, 14°13 ' 09 '' N / 108°19 ' 48 '' E, Kon Ka Kinh Nat [io]n[al]. Park, h= 1470 m, 21–30.V.2017, D. Fedorenko leg.’; ♂, ♀, same data except 14°13 ' 40 '' N / 108°18 ' 35 '' E, h= 1300–1380 m; ♂, ‘ Vietnam, Quang Nam Prov., Nam Gian Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.38333&amp;materialsCitation.latitude=15.547501" title="Search Plazi for locations around (long 107.38333/lat 15.547501)">Song Thanh Natn. Park</a>, 15°32 ' 51 '' N / 107°23 ' E, h= 1230 m, 23.IV–11.V.2019, leg. D.Fedorenko’; 3♂♂, ♀, same data, except for either 15°34 ' 07 '' N / 107°23 ' 13 '' E, h= 1010 m; or 15°32–33 ' N, 103°22–23 ' E, h= 1100– 1230 m; or 15°33 ' 48 '' N / 107°23 ' 22 '' E, h= 1050 m; or 15°33 ' 19 '' N / 107°23 ' 29 '' E, h= 1070 m. Aedeagus examined in eight males.</p> <p>DESCRIPTION. As for A. acutipennis sp.n. except as follows: Body (Fig. 7) medium-sized, BL 6.7–8.7 mm. Black yet barely paler than A. acutipennis; antennae entirely pale, tibiae and tarsi red or pale brown, or tibiae slightly darker than tarsi while distinctly paler than femora. Lateral margin of elytra more clearly translucent with reddish-yellow. Microsculpture coarse, isodiametric on head, granulate-aciculate on pronotum, much more superficial on elytra, consisting of very transverse meshes mixed with wider transverse meshes here and there; explanate lateral margin of elytra with isodiametric microsculpture. Head and pronotum subequally dull, elytra moderately shiny.</p> <p>Antennae surpassing pronotal base by apical 3–3.5 antennomeres.</p> <p>Pronotum cordate, much narrower relative to elytra, PW/ PL 1.16–1.21 (mean 1.18, n=5), PW/HW 1.42–1.50 (1.47). Base straight or indistinctly convex posteriad, on a level with gently rounded lateral lobes or barely produced, PB/PA 1.33–1.43 (1.38). Basal angles slightly to very obtuse. Basal foveae rather small and mostly not extended apicad. Paramedian foveae usually indistinct, situated midway between median line and narrow lateral grooves. Reflexed lateral margin slightly to moderately reflexed, narrow from apical angle to anterolateral seta, slightly and increasingly wider toward basal angles.</p> <p>Elytra: EL/EW 1.41–1.47 (1.44), EW/PW 1.55–1.69 (1.59); base very narrow yet distinct; posterolateral angles very obtuse and rounded; apices obtuse and pointed combined. Apical truncature barely sinuate, more deeply near apex. Reflexed lateral margin slightly concave dorsally, wide medially, much narrower before and behind. Intervals convex, inner ones slightly convex to nearly flat. Interval 3 with three setae; d1 very close to base, D1/EL 0.09–0.12 (0.11, n=5×2), D2/EL 0.60–0.69 (0.66), D3/EL 0.89–0.93 (0.92).</p> <p>Metepisternum short, ES3L/ES3W 0.85-1.0 (0.92, n=3).</p> <p>In male, profemur without tubercle, mesotrochanter with a small tubercle, mesotibia with four tubercles along inner margin.</p> <p>Aedeagus (Figs 17–18, 33): as in the previous species, except for apical lamella slenderer and nearly straight in ventral view.</p> <p>DIAGNOSIS. This species is transitional from A. medius and A. tonkinensis sp.n. to A. giganteus and A. acutipennis sp.n. in size. The three species of this group, that share similar elytral chetome, A. basipunctatus sp.n., A. giganteus, and A. acutipennis sp.n., have distinctive elytral apices each. The apices are either contiguous and sharp, with indistinct sinuation outside, in A. basipunctatus sp.n. or slightly separate at sutural angle, with the apical truncature either deeply and evenly concave (A. giganteus) or very deeply concave just behind outer angle (A. acutipennis sp.n.). For other differences see the key.</p> <p>VARIATION. Specimens from the northern population (Quang N am) may be barely larger than those from southern ones, BL being 7.3–8.7 or 6.7–8.1 mm, respectively.</p> <p>DISTRIBUTION. Central Vietnam (Gia Lai, Kon Tum).</p> <p>NAME. Refers to the anterior discal seta (d1) inserted closer to elytral base than in the congeners.</p> <p>HABITATS AND HABITS. As for the previous species.</p> <p>3.7. Amphimenes (s. str.) tonkinensis</p> <p>Fedorenko, sp.n.</p> <p>Fig. 8.</p> <p>MATERIAL. Holotype ♀ (ZMMU): ‘N-Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat N [ational]P[ark], 22°36 ' 02 '' N / 103°37 ' 13 '' E, h= 2200 m, 4–14.VI.2019, leg. D.Fedorenko’.</p> <p>DESCRIPTION. Body apterous, small (Fig. 8), BL 6 mm, shiny black, with pronotum dull. Clypeus, except for brownish base, labrum, mouthparts, antennae, legs, gula, prosternum, hypomera, meso- and metaventrite, and abdominal sternite III between metacoxae red; abdomen widely reddish along middle, underside otherwise rather dark brown; reflexed lateral margins of elytra translucent with red, those of pronotum translucent with reddish-brown. Microsculpture isodiametric on head; pronotal microsculpture very coarse, granulate-aciculate in basal half and along sides, otherwise consisting of isodiametric or barely transverse meshes. Elytra with meshed microsculpture superficial, isodiametric or barely transverse. Elytral cross-striated sculpture vestigial, hardly traceable along middle of elytral intervals basally, nearly indistinct behind.</p> <p>Head narrow, with eyes rather flat and small, as long as genae which almost smoothly extend into neck. Posterior supra-ocular seta distant far from posterior margin of eye. Antennae fairly short, surpassing base of pronotum by apical antennomere.</p> <p>Pronotum narrow, subcordate, nearly square, broadest at anterolateral seta slightly less than a third (0.30) from apex, slightly wider than long, PW/PL 1.16, PW/HW 1.39. Base a fourth broader than apex, PB/PA 1.24, barely concave between lateral lobes; these convex caudad and more rounded laterally. Basal angles very obtuse and blunt. Apex deeply sinuate; apical angles porrect and acute. Sides evenly round- ed in apical three fourths, gently sinuate behind. Disc flattened. Lateral margins together with basal angles and laterobasal lobes strongly reflexed, slightly wider before than behind. Median line entire and moderately deep; basal transverse impression short and deep. Basal foveae short and deep, each extended apicad into a very shallow impression that disappears medially. Paramedian foveae indistinct.</p> <p>Elytra elliptic, EL/EW 1.45, EW/PW 1.71, broadest medially, with common base narrow, humeri round yet traceable, and sides evenly rounded up to rounded posterolateral angles. Apical truncature oblique, slightly sinuate. Apices very slightly obtuse combined and nearly contiguous. Lateral margin rather widely explanate in inner three fifths. Striae deep, intervals slightly convex, almost flat on disc. Interval 3 with three setae, D1/EL 0.13, D2/EL 0.37/0.40, D3/EL 0.90/ 0.91 (0.93).</p> <p>Metepisternum short, about as long as wide.</p> <p>Tarsomere 5 with one pair of ventral setae, preapical.</p> <p>DIAGNOSIS. Within the medius -group, this species is easily recognizable by the combination of the body small, with narrow forebody and pale appendages, nearly isodiametric elytral microsculpture, three elytral discal setae present, with seta d1 inserted before middle.</p> <p>DISTRIBUTION. Known from the type locality only, that is on the Hoang Lien Mt. Ridge, Lao Cai, northern Vietnam.</p> <p>NAME. Toponymic; refers to the former historical name of northern Vietnam.</p> <p>HABITATS AND HABITS. The only female specimen was hand collected under a small tree fragment on the ground in a cloudy forest. The species appears to live syntopically with A. reflexicollis.</p> <p>COMMENTS. Two elytral characters, nearly isodiametric microsculpture and seta d1 inserted distinctly before middle, are not characteristic of the other members of the medius -group and the species range is distinctive. Accordingly this species may require to be placed in the separate group of itself.</p> </div>	https://treatment.plazi.org/id/03F0879A163EFFA5FEA6FDE42403FC67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A163CFFBBFC58FC742587FDF7.text	03F0879A163CFFBBFC58FC742587FDF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes planicollis Fedorenko 2010	<div><p>The planicollis species group.</p> <p>The group includes four apterous soil-dwelling species, three from northern Vietnam and one from Central Vietnam. The group is defined by the combination of the elytral seta d1 missing and seta d2 situated behind middle.</p> <p>3.8. Amphimenes (s. str.) konplongensis</p> <p>Fedorenko, sp.n.</p> <p>Figs 9, 21–22, 38.</p> <p>MATERIAL. Holotype ♂ (ZMMU) labelled: ‘ Vietnam, Kon Tum Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.30278&amp;materialsCitation.latitude=14.75" title="Search Plazi for locations around (long 108.30278/lat 14.75)">Kon Plong Distr</a> [ict]., 14°45 ' N / 108°18 ' 10 '' E, env. ngock Boc I Mt., h = 1200–1300 m, 8–23.IV.2015, D. Fedorenko leg.’. Paratypes (SIEE, ZISP, ZMMU): 6♂♂, 3♀♀, same data; ♂, ♀, same data, except for 14°44 ' 20 '' N / 108°19 ' E, h = 1100–1200 m; 5♂♂, 2♀♀, same data, except for 14°45 ' N / 108°17 ' 51 '' E, h= 1300–1400 m, 8–10.VI.2016; ♀, ‘ Vietnam, Quang Nam Prov., Nam Gian Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.38333&amp;materialsCitation.latitude=15.547501" title="Search Plazi for locations around (long 107.38333/lat 15.547501)">Song Thanh Natn. Park</a>, 15°32 ' 51 '' N / 107°23 ' E, h= 1230 m, 23.IV– 11.V.2019, leg. D.Fedorenko’. Aedeagus examined in three males.</p> <p>DESCRIPTION. Body (Fig. 9) apterous, medium-sized for the genus, BL 6–7.1 mm, and black. Mouthparts, including labrum, antennae, trochanters, tibiae, tarsi and femoral apices reddish-yellow; femora otherwise dark brown to brownblack; lateral margins of elytra and narrow lateral margins of pronotum barely to distinctly translucent with red. Microsculpture coarse, isodiametric on head, granulate at middle of pronotum, granulate-aciculate around; elytral microsculpture much more superficial yet distinct, consisting of moderately to very transverse meshes becoming isodiametric ones on explanate lateral margin. Head and pronotum equally dull, slightly shining, elytra shinier. Elytral intervals moderately densely cross-striated, less so behind.</p> <p>Head: eyes slightly flattened; genae about a third as long as eyes, mostly very smoothly extended into neck. Posterior supra-ocular seta just behind posterior margin of eye. Antennae long, surpassing pronotal base by about apical three antennomeres.</p> <p>Pronotum cordate, broadest slightly less than two fifths from apex, a third to two fifths wider than long, PW/PL 1.34–1.40 (mean 1.37, n=5), PW/HW 1.47–1.60 (1.51). Base two fifths wider than apex, PB/PA 1.40–1.44 (1.42), with median part on a level with and a third wider than lateral lobes, all slightly rounded or the lobes more rounded toward basal angles. Basal angles slightly to highly obtuse. Apex deeply sinuate; apical angles porrect, slightly acute, and blunt. Sides evenly rounded in apical four fifths, mostly distinctly sinuate behind and barely diverging basad. Disc moderately convex, very finely transversely rugulose. Lateral margins moderately explanate-reflexed, narrow apically and gradually broadening to rather wide at basal angles; lateral groove narrow. Median line moderately deep and mostly obsolete behind well defined basal transverse impression. Basal foveae round and moderately deep, not extended forward. Paramedian foveae usually wide yet very shallow to indistinct.</p> <p>Elytra broadly oval, EL/EW 1.39–1.48 (1.44), EW/PW 1.45–1.56 (1.49), at base as wide as pronotal base, evenly rounded on sides, broadest at middle, with humeri distinct yet widely rounded and posterolateral angles very obtuse and rounded; apices contiguous, subrectangular to obtuse combined, sharp or slightly blunted. Apical truncature oblique, slightly sinuate near apex. Striae deep; intervals moderately convex, outer ones slightly convex to nearly flat. Interval 3 with two setae, D2/EL 0.58–0.63 (0.60, n=5×2), D3/EL 0.92–0.94 (0.93).</p> <p>Metepisternum short, ES3L/ES3W 0.84-0.90 (0.87, n=3).</p> <p>Tarsomere 5 with one pair of ventral setae, metatarsomere often with additional vestigial pair of setae proximal to normal setae. In male, profemur without tubercle, mesotrochanter with a small tubercle; mesotibia with three or four minute tubercles along inner margin.</p> <p>Aedeagus median lobe cylindrical (Figs 21–22, 38).</p> <p>DIAGNOSIS. For details see the key.</p> <p>DISTRIBUTION. Known from the type locality only.</p> <p>NAME. Given after the provenance of the species, Kon Plong District, Kon Tum, Vietnam.</p> <p>HABITATS AND HABITS. As for A. acutipennis sp.n.</p> <p>3.9. Amphimenes (s. str.) bicoloripes Fedorenko, sp.n.</p> <p>Figs 23–24, 27.</p> <p>MATERIAL. Holotype ♂ (ZMMU) labelled:‘N-Vietnam, Nghe An Prov., Que Phong Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.80361&amp;materialsCitation.latitude=19.776388" title="Search Plazi for locations around (long 104.80361/lat 19.776388)">Pu Hoat National Park</a>, 19°46 ' 35 '' N / 104°48 ' 13 '' E, h= 1350 m 15–27.V.2019, leg. D.Fedorenko’. Paratypes (SIEE) 8♂♂, 12♀♀, same data; 5♂♂, 4♀♀, same data, except 19°45 ' N / 104°46 ' 38 '' E, h= 1220 m; ♀, same data, except 19°46 ' 13 '' N / 104°47 ' 08 '' E, h= 1030 m. Aedeagus examined in eight males.</p> <p>DESCRIPTION. As for A. konplongensis, except for small differences that concern body proportions, elytral microsculpture, and shape of aedeagus. Body of nearly the same facies as in A. konplongensis (see Fig. 9), BL 5.8–7.5 mm. elytral microsculpture consisting of dense transverse lines to extremely narrow and almost indistinct transverse meshes. Pronotum slightly narrower, PW/PL 1.24–1.35 (mean 1.29, n=7), PW/HW 1.44–1.54 (1.48), PB/PA 1.34–1.41 (1.36). Elytra broadly oval, EL/EW 1.41–1.50 (1.45), EW/PW 1.49– 1.60 (1.57). D2/EL 0.66–0.72 (0.69, n=6×2), 0.56/ 0.59 in one specimen measured, D3/EL 0.91–0.94 (0.93).</p> <p>Tarsomere 5 with two pairs of ventral setae; mesotibia with four tubercles along inner margin.</p> <p>Aedeagus (Figs 23–24, 27) with median lobe strongly twisted; left margin angulate at middle.</p> <p>DIAGNOSIS. For details see the key.</p> <p>DISTRIBUTION. Known from the type locality only.</p> <p>NAME. Refers to the bicoloured legs, pale, with femora infuscated.</p> <p>HABITATS AND HABITS. Under various tree fragments on the ground in monsoon forests, mostly above 1200 m elevation.</p> <p>3.10. Amphimenes (s. str.) reflexicollis</p> <p>Fedorenko, 2010</p> <p>Fedorenko, 2010: 39 (Tam Dao, northern Vietnam).</p> <p>MATERIAL. Northern Vietnam (SIEE; D. Fedorenko leg.): 4♂♂, 3♀♀, Cao Bang, 40 km W of Cao Bang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.88528&amp;materialsCitation.latitude=22.600834" title="Search Plazi for locations around (long 105.88528/lat 22.600834)">Phia Oac — Phia Den Nat. Park</a>, Phia Oac Mt, 22°36 ' 03 '' N 105°53 ' 07 '' E, h= 1300 m, 3– 11.X.2018; ♂ (SIEE), same data, except for 22°36 ' 25 '' N 105°52 ' 08 '' E, h= 1650–1700 m; 5♂♂, 4♀♀, same data, except for 22°36 ' 30 '' N 105°52 ' 20 '' E, h= 1600–1650 m; 9♂♂, 3♀♀, Lao Cai, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Nat. Park, 22°372– 37.6 ' N 103°37.32–38.3 ' E, h= 1600–1900 m, 16–21.X.2018; 5♂♂, 5♀♀, same data, except for 22°36 ' 31 '' N 103°37 ' 23 '' E, h= 2000–2100 m, 4–14.VI.2019; 10♂♂, 5♀♀, same data except 22°37 ' 36 '' N 103°37 ' 32 '' E, h= 1850 m. Aedeagus examined in seven males.</p> <p>REDESCRIPTION. Based on material listed, the following additions are only necessary: BL 7.6–9.3 mm. PW/PL 1.31–1.42 (1.35, n=10), PW/HW 1.50–1.68 (1.59), PB/PA 1.41–1.52 (1.47), EL/EW 1.43–1.51 (1.48), EW/PW 1.45– 1.56 (1.51), D2/EL 0.61–0.71 (0.67, n=9×2).</p> <p>DISTRIBUTION. The species range covers montane areas of northern Vietnam from Lao Cai Province in the west to Vinh Phuc (type locality) and Cao Bang provinces in the east.</p> <p>HABITATS AND HABITS. This species occurs under smaller tree fragments on the ground. Individual adults were taken under bark of standing dead trees up to 1–2 m above ground.</p> <p>COMMENTS. There are some subtle differences between two populations of the species, western (Lao Cai) and eastern (Cao Bang). The adults from the former have the pronotum barely larger, PW/HW 1.54–1.68 (1.61) vs. 1.50– 1.61 (1.56), EW/PW 1.45–1.52 (1.48) vs. 1.49–1.56 (1.54), with base wider, PB/PA 1.47–1.52 (1.49) vs. 1.41–1.49 (1.45); explanate and reflexed lateral margin barely wider, sides indistinctly sinuate to subconvex (vs. subsinuate to straight), and basal angles subrectangular to obtuse (vs. nearly right with narrowly rounded apices); and USS continuous and consisting of 16–18 US (vs. 15–17).</p></div> 	https://treatment.plazi.org/id/03F0879A163CFFBBFC58FC742587FDF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A1622FFBAFC43FDE420FDFACA.text	03F0879A1622FFBAFC43FDE420FDFACA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes brunneus Kirschenhofer 1999	<div><p>The brunneus species group.</p> <p>The group includes five apterous soil-dwelling species from northern Vietnam; they all share elytral seta d1 missing and seta d2 situated before middle, combined with elytral cross-striated sculpture strongly reduced to nearly indistinct in most species. Three species of the group are known based on female holotypes only.</p> <p>3.11. Amphimenes (s. str.) femoralis Fedorenko, sp.n.</p> <p>Figs 10, 19–20, 35.</p> <p>MATERIAL. Holotype ♀ (ZMMU) and paratypes ♂, 4♀♀ (SIEE), labelled: ‘N[orthern]- Vietnam, 40 km W of Cao Bang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.86889&amp;materialsCitation.latitude=22.606945" title="Search Plazi for locations around (long 105.86889/lat 22.606945)">Phia Oac Mt.</a>, 22°36 ' 25 '' N / 105°52 ' 08 '' E, h~ 1650–1700 m, deciduous forest, 3–11.X.2018, leg. D.Fedorenko’.</p> <p>DESCRIPTION. Body (Fig. 10) apterous, small, BL 6.2– 6.7 mm, and shiny black. Head and pronotum slightly dull, sometimes indistinctly paler; clypeus entirely or in apical half, labrum, mouthparts, antennae, and legs reddish-yellow; femora infuscated except apically. Pronotal lateral margin barely translucent with brown red, reflexed lateral margins of elytra narrowly red, becoming reddish brown just inside. Microsculpture coarse, isodiametric on head and at middle of pronotum, nearly granulate around; elytral microsculpture very superficial, consisting of very transverse meshes that become isodiametric ones on explanate lateral margin. Elytral cross-striated sculpture vestigial, hardly traceable in basal 1/2–1/3, slightly more distinct close to base.</p> <p>Head with slightly flattened eyes; genae about half as long as eyes. Posterior supra-ocular seta about 2/5 eye length distant from posterior margin of eye. Antennae surpassing pronotal base by about apical two antennomeres.</p> <p>Pronotum cordate, broadest slightly more than a third from apex, less than a third wider than long, PW/PL 1.28– 1.31 (1.29, n=6), and half wider than head, PW/HW 1.48– 1.55 (1.51). Base a third wider than apex, PB/PA 1.29–1.39 (1.34), with median part about as wide as lateral lobes; these evenly rounded and slightly projecting, more rounded towards highly obtuse basal angles. Apex very deeply sinuate; apical angles porrect, slightly acute and blunt. Sides rounded, more so toward apical angles, distinctly sinuate before basal ones. Disc convex, explanate lateral margins strongly reflexed, very narrow before anterolateral seta, moderately and increasingly wide behind. Median line deep, reaching apex and not quite reaching base. Basal foveae rather small, round, deeper just in front of meeting point of basal median part and lateral lobes. Basal transverse impression deep. Paramedian foveae small, moderately deep to indistinct, situated about midway between median line and lateral margin 2/5 from apex.</p> <p>Elytra broadly oval, a third longer than wide, EL/EW 1.31–1.41 (1.34), and two thirds wider than pronotum, EW/ PW 1.59–1.68 (1.66), broadest at middle, with evenly round- ed sides. Humeri distinct yet more or less widely rounded, posterolateral angles very obtuse and blunt. Apical truncature oblique and rather transverse, barely sinuate; apices mostly very obtuse combined and blunt. Striae deep; intervals convex basally and in apical fourth, slightly less convex to nearly flat in between. Interval 3 bisetose, D2/EL 0.42–0.55 (0.49, n=6×2), posterior discal seta close to apex.</p> <p>Metepisternum short: ES3L/ES3W 0.86–0.89 (n=2).</p> <p>Tarsomere 5 with one pair of ventral setae. In male, profemur a third from base with a large ventral tubercle; mesotibia 4-tuberculate along apical 2/5 inner margin.</p> <p>Aedeagus median lobe (Figs 19–20, 35) slightly twisted and dilated before apical orifice because a conspicuous left ventrolateral bulge is present.</p> <p>DIAGNOSIS. This species belongs to the brunneus - group that includes totally five species, being very similar to A. brunneus and A. kabakovi. The latter two species share unicoloured pale legs, being distinctive also in having either a weaker microsculpture on the head and pronotum (A. kabakovi) or the body smaller, with elytral apices more pointed combined, subrectangular (A. brunneus) vs. very obtuse. Differences between the three species in body proportions are subtle, yet not overlapped (Table).</p> <p>DISTRIBUTION. Known from the type locality only.</p> <p>NAME. Refers to the femora infuscated.</p> <p>HABITATS AND HABITS. An apterous soil-dwelling species occurring under smaller tree fragments on the ground and sharing these habitats with the sympatric species A. reflexicollis.</p> <p>COMMENTS. Relationships between A. brunneus, A. kabakovi and A. femoralis sp.n. are obscured by their great similarity and the males unknown for the former two. More material is required to clarify whether they are species or subspecies, or conspecific.</p> </div>	https://treatment.plazi.org/id/03F0879A1622FFBAFC43FDE420FDFACA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
03F0879A1623FFBAFEB3FA8125D2FE0A.text	03F0879A1623FFBAFEB3FA8125D2FE0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphimenes brunneus Kirschenhofer 1999	<div><p>The brunneus species group.</p> <p>This group includes two species, one apterous soil-dwelling from southern Vietnam, the other macropterous arboricolous and widespread. Both share the body small-sized, with elytral seta d1 missing, seta d2 situated at or just before middle, and the antennae short.</p> <p>3.12. Amphimenes (s. str.) micros Fedorenko, 2014</p> <p>Fedorenko, 2014: 309 (Chu Yang Sin NP, southern Vietnam).</p> <p>MATERIAL. Central Vietnam: 2♂♂, ♀ (SIEE; D. Fedorenko leg.), Gia Lai, ~ 50 km N of An Khe, Kon Chu Rang <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.53333&amp;materialsCitation.latitude=14.516666" title="Search Plazi for locations around (long 108.53333/lat 14.516666)">Nature Reserve</a>, 14°31 ' N 108°32 ' E, h= 1000–1040 m, 24.V–2.VI.2016; ♀ (SIEE), Kon Tum, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.30278&amp;materialsCitation.latitude=14.75" title="Search Plazi for locations around (long 108.30278/lat 14.75)">Kon Plong Distr.</a>, 14°45 ' N 108°18 ' 10 '' E, env. ngock Boc 1 Mt., h= 1300–1400 m, 8–10.VI.2016; ♂, Quang N am, Nam Gian Distr., Song Thanh National Park, 15°32–33 ' N 103°22–23 ' E, h= 1100–1230 m, 23.IV–11.V.2019.</p> <p>REDESCRIPTION. Unnecessary except the following corrections: BL 4.3–5.2 mm. PW/PL 1.25–1.40 (1.37, n=6), PW/HW 1.34–1.47 (1.43), PB/PA 1.21–1.35 (1.26), EL/EW 1.35–1.41 (1.39), EW/PW 1.49–1.59 (1.55), D2/EL 0.39– 0.46 (0.42, n=6×2), D3/EL 0.91–0.95 (0.92).</p> <p>DISTRIBUTION. This species was previously known from Dak Lak Province only. The listed records extend the species range farther northwards.</p> <p>Acknowledgements. I am very indebted to Dr. Beulah Garner (BMNH) and Dr. Kirill Makarov (MPSU) for loan of material under their care, and to Dr. Alexei Polilov and Mr. Igor Melnik (both from Moscow) for donating specimens. This study was funded by the Presidium of the Russian academy of sciences, Program No.41 “Biodiversity of natural systems and biological resources of Russiaâ”.</p></div> 	https://treatment.plazi.org/id/03F0879A1623FFBAFEB3FA8125D2FE0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fedorenko, D. N.	Fedorenko, D. N. (2019): A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233-250, DOI: 10.15298/rusentj.28.3.02, URL: http://dx.doi.org/10.15298/rusentj.28.3.02
