taxonID	type	description	language	source
03F1878DDC56D82BF9F4C403CF3CF827.taxon	diagnosis	Diagnosis. Wiles 1997 a: 202; Di Sabatino et al. 2010: 180.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D828F9F4C3EDCE49FF08.taxon	diagnosis	Diagnosis. Di Sabatino et al. 2010: 180.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D82EF9F4C23BCDEEFF24.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province: stream near Kampung Kleublouw between Jayapura and Sentani, 28. iii. 2010, 2 ˚ 35.249 S 140 ˚ 35.187 E, 82 m a. s. l. Paratypes: 1 / 0 (mounted), River Pos 7, Sentani, 9. iii. 2010, 2 ˚ 33.740 S 140 ˚ 30.784 E, 103 m a. s. l.; 1 / 0 (mounted), River Kamp Walker near Uncen, Abepura, 31. iii. 2010, 2 ˚ 34.202 S 140 ˚ 38.886 E, 144 m a. s. l. Further records. Indonesia, New Guinea, Papua Province, River Yabawi, upstream of Harapan, 28. iii. 2010, 2 ˚ 34.216 S 140 ˚ 33.723 E, 120 m a. s. l. 0 / 2 (0 / 1 mounted).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D82EF9F4C23BCDEEFF24.taxon	diagnosis	Diagnosis. Idiosoma dimensions small (L / W 700 – 744 / 447 – 494 in males); idiosoma elongated (dorsal shield L / W 1.6 – 1.7); capitular bay rectangular at its proximal end; distal margins of P- 3 without denticles, P- 4 with well developed tubercles near the insertion of the ventral hairs; Cx- 4 with slightly accentuated posterior suture line; postgenital area large; excretory pore close to the line of primary sclerotization, Vgl- 2 posterior and well distanced from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D82EF9F4C23BCDEEFF24.taxon	description	Description. Male (holotype from stream near Kampung Kleublouw, in parentheses paratype from River Pos 7, in square brackets paratype from River Kamp Walker): Idiosoma (ventral view: Fig. 2 B) L 700 (744) [744], W 447 (450) [494]; dorsal shield (Fig. 2 A) L 597 (631) [650], W 356 (375) [391], L / W ratio 1.68 (1.68) [1.66]; dorsal plate 569 (606) [588]; shoulder plate L 163 (175) [167], W 50 (52) [53], L / W ratio 3.26 (3.37) [3.15]; frontal plate L 108 (117) [113], W 44 (44) [44], L / W ratio 2.46 (2.66) [2.57]; shoulder / frontal plate L ratio 1.5 (1.5) [1.48]; capitular bay L 138 (151) [149], W 66 (67) [78], L / W ratio 2.1 (2.25) [1.9]; Cx- 1 total L 234 (250) [253], Cx- 1 medial L 95 (99) [103], Cx- 2 + 3 medial 56 (63) [63]; ratio Cx- 1 L / Cx- 2 + 3 medial L 4.2 (4.0) [4.0]; Cx- 1 medial L / Cx- 2 + 3 medial L 1.7 (1.6) [1.6]; genital field L / W 135 (137) [136] / 106 (106) [113], L / W ratio 1.27 (1.29) [1.2], ejaculatory complex (Fig. 2 D) L 140 (145); distance genital field – excretory pore 154 (172) [167], genital field – caudal idiosoma margin 269 (283) [300]; capitulum (Fig. 2 C) ventral L 166 (185) [182]; chelicera L 220 (235) [226]; palp (Figs. 2 E – F) total L 206 (210) [206], dL: P- 1, 25 (25) [26]; P- 2, 60 (59) [60]; P- 3, 40 (43) [40]; P- 4, 50 (52) [52]; P- 5, 31 (31) [28]; % L: P- 1, 12.1 (11.9) [12.6]; P- 2, 29.1 (28.1) [29.1]; P- 3, 19.4 (20.5) [19.4]; P- 4, 24.3 (24.8) [25.2]; P- 5, 15.1 (14.8) [13.6]; L P- 2 / P- 4 ratio, 1.2 (1.14) [1.15]; L I-L- 4 - 6 (Fig. 2 G): 105 (108) [111], 115 (114) [117], 112 (119) [115]. Additionally we give the measurements for one specimen from River Yabawi which we suspect represents the female sex of Monatractides novaeguineae nov. sp. Female: Idiosoma (ventral view: Fig. 4 A) L 736, W 464; dorsal shield L 631, W 366, L / W ratio 1.72; dorsal plate L 606; shoulder plate L 164, W 55, L / W ratio 3.0; frontal plate L 106, W 46, L / W ratio 2.3; shoulder / frontal plate L ratio 1.55; capitular bay L 145, W 69, L / W ratio 2.2; Cx- 1 total L 231, Cx- 1 medial L 87, Cx- 2 + 3 medial 45; ratio Cx- 1 L / Cx- 2 + 3 medial L 5.1; Cx- 1 medial L / Cx- 2 + 3 medial L 1.9; genital field L / W 147 / 125, L / W ratio 1.18; distance genital field – excretory pore 181, genital field – caudal idiosoma margin 311; capitulum ventral L 175; chelicera total L 236; palp total L 212, dL and % L (in parentheses): P- 1, 24 (11.3); P- 2, 62 (29.3); P- 3, 42 (19.8); P- 4, 52 (24.5); P- 5, 32 (15.1); P- 2 / P- 4 ratio 1.19; palp segments as in male; L I-L- 5 - 6: 118, 114.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D82EF9F4C23BCDEEFF24.taxon	discussion	Discussion. This species belongs to the M. luteus (K. Viets, 1935) species-complex, characterized by the presence of anteriorly broad and shorter Cx- 1, a relatively wide capitular bay, a relatively short medial suture line of Cx- 2 + 3 in both sexes and I-L- 6 not greatly expanded (Pešiċ & Smit 2010). Due to the considerably elongated idiosoma (e. g., dorsal shield L / W 1.6 – 1.7) and the shape of the capitular bay which is rectangular at its proximal end, Monatractides novaeguineae sp. nov. resembles M. papuensis sp. nov. (see below), but differs in its minor idiosoma and gnathosoma dimensions (e. g., in males: Idiosoma L / W <800 / 500 in M. novaeguineae, L / W> 900 / 600 in M. papuensis, genital field L / W <140 / 120 in M. novaeguineae, L / W> 180 / 130 in M. papuensis, palp total L <215 in M. novaeguineae,> 250 in M. papuensis). Further differences are found in a shorter ejaculatory complex in the male with a relatively longer proximal chamber, distal margins of P- 3 without denticles and a generally slightly accentuated posterior suture line of Cx- 4. In the females, the additional differences are found in the shape of genital field with rounded (in M. papuensis angled) anterolateral margins. Apart from the diagnostic features, there are ecological differences between these two species: Monatractides novaeguineae sp. nov. was found only in the lowlands, while M. papuensis sp. nov. seems to be restricted to higher elevations.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC55D82EF9F4C23BCDEEFF24.taxon	etymology	Etymology. The species is named after the country where it was collected. Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province: Bion River, upstream, Pass Valley, 15. iii. 2010, 3 ˚ 51.294 S 139 ˚ 05.733 E, 1962 m a. s. l. Paratypes: 1 / 5 (1 / 1 mounted), same data as holotype; 0 / 1 / 0, Bion River, 15. iii. 2010, 3 ˚ 51.513 S 139 ˚ 05.570 E, 2007 m a. s. l.; 1 / 3 (0 / 1 mounted), same data as holotype, 02. iii. 2010, 3 ˚ 51.267 S 139 ˚ 05.789 E, 1962 m a. s. l. Further records: Indonesia, New Guinea, Papua Province, unnamed creek crossing road to Pass Valley, 13. iii. 2010, 3 ˚ 52.849 S 139 ˚ 04.194 E, 2253 m a. s. l. 0 / 1.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	diagnosis	Diagnosis. Idiosoma dimensions large (L> 900 in male,> 1000 in female); idiosoma elongated (dorsal shield L / W 1.6 – 1.7); capitular bay rectangular at its proximal end; distal margins of P- 3 bearing at least one denticle, P- 4 with well developed tubercles near the insertion of the ventral hairs; Cx- 4 with well accentuated posterior suture line; postgenital area large; excretory pore close to the line of primary sclerotization, Vgl- 2 posterior and well away from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	description	Description. Male (holotype, in parentheses paratype): Idiosoma (ventral view: Fig. 3 B) L 931 (969), W 600 (625); dorsal shield (Fig. 3 A) L 809 (825), W 478 (475), L / W ratio 1.69 (1.74); dorsal plate 769 (794); shoulder plate L 205 (200), W 74 (72), L / W ratio 2.77 (2.77); frontal plate L 144 (153), W 66 (61), L / W ratio 2.18 (2.5); shoulder / frontal plate L ratio 1.42 (1.31); capitular bay L 166 (184), W 94, L / W ratio 1.8; Cx- 1 total L 297 (322), Cx- 1 medial L 129 (134), Cx- 2 + 3 medial 61 (52); ratio Cx- 1 L / Cx- 2 + 3 medial L 4.9 (6.2); Cx- 1 medial L / Cx- 2 + 3 medial L 2.1 (2.6); genital field L / W 184 (181) / 139 (141), L / W ratio 1.32 (1.28); ejaculatory complex (Fig. 3 F) L 197 (219); distance genital field – excretory pore 191 (197), genital field – caudal idiosoma margin 381 (398); capitulum (Fig. 3 C) ventral L 214 (215); chelicera total L 262 (266); palp (Fig. 3 D – E): total L 258 (260), dL: P- 1, 32 (29); P- 2, 73 (75); P- 3, 50 (49); P- 4, 69 (72); P- 5, 34 (35); % L: P- 1, 12.4 (8.1); P- 2, 28.3 (28.9); P- 3, 19.4 (18.9); P- 4, 26.7 (27.7); P- 5, 13.2 (13.5); P- 2 / P- 4 ratio, 1.06 (1.06); L I-L- 4 - 6 (Fig. 3 G): 129 (140), 147 (154), 143 (151). Female (from Bion River, upstream, n = 2): Idiosoma (ventral view: Fig. 4 B) L 1013 – 1059, W 619; dorsal shield L 881 – 894, W 519 – 531, L / W ratio 1.68 – 1.7; dorsal plate 846 – 856; shoulder plate L 209 – 212, W 72 – 77, L / W ratio 2.7 – 2.9; frontal plate L 153, W 62 – 63, L / W ratio 2.43 – 2.47; shoulder / frontal plate L ratio 1.37 – 1.39; capitular bay L 178, W 84, L / W ratio 2.1; Cx- 1 total L 322 – 341, Cx- 1 medial L 144 – 147, Cx- 2 + 3 medial 33 – 37; ratio Cx- 1 L / Cx- 2 + 3 medial L 9.2 – 9.8; Cx- 1 medial L / Cx- 2 + 3 medial L 4.0 – 4.4; genital field L / W 203 – 209 / 169 – 172, L / W ratio 1.18 – 1.24; distance genital field – excretory pore 230, genital field – caudal idiosoma margin 444 – 453; capitulum ventral L 223 – 245; chelicera total L 275 – 284; palp: total L 267 – 288, dL and % L (in parentheses): P- 1, 31 – 36 (11.6 – 12.5); P- 2, 78 – 85 (29.2 – 29.5); P- 3, 57 – 57 (19.8 – 21.3); P- 4, 66 – 74 (24.7 – 25.7); P- 5, 35 – 36 (12.5 – 13.1); L P- 2 / P- 4 ratio, 1.15 – 1.18; L I-L- 4 - 6: 135 – 142, 150 – 158, 149 – 154.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	etymology	Etymology. The species is named after the country where it was collected.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	discussion	Discussion. See discussion section under the preceeding species.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82EF9F4C255CDEEF9BB.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province, Wao & Pao streams near confluence, Walmak, Nipsan district, 18. iii. 2010, 4 ˚ 07.097 S 139 ˚ 38.520 E, 1739 m a. s. l.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	diagnosis	Diagnosis (Female unknown). Idiosoma L> 900; idiosoma elongated (L / W ratio 1.6); capitular bay rounded at its proximal end; distal margins of P- 2 and - 3 without denticles, P- 4 with well visible denticle near the insertion of the ventral hairs; genital field elongated (L / W ratio 1.6); postgenital area large; excretory pore on the line of primary sclerotization, Vgl- 2 posterior and well away from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	description	Description. Male: Idiosoma (ventral view: Fig. 5 B) L 1013, W 675; dorsal shield (Fig. 5 A) L 894, W 556, L / W ratio 1.6; dorsal plate 831; shoulder plate L 247, W 94, L / W ratio 2.6; frontal plate L 166, W 75, L / W ratio 2.2; shoulder / frontal plate L ratio 1.49; capitular bay L 198, W 99, L / W ratio 2.0; Cx- 1 total L 344, Cx- 1 medial L 147, Cx- 2 + 3 medial 76; ratio Cx- 1 L / Cx- 2 + 3 medial L 4.5; Cx- 1 medial L / Cx- 2 + 3 medial L 1.9; genital field L / W 204 / 128, L / W ratio 1.6; ejaculatory complex conventional in shape, L 216; distance genital field – excretory pore 226, genital field – caudal idiosoma margin 375; capitulum (Fig. 6 C) ventral L 243; chelicera total L 295; palp (Figs. 6 A – B) total L 289, dL and % L (in parentheses): P- 1, 34 (11.8); P- 2, 85 (29.4); P- 3, 55 (19.0); P- 4, 77 (26.6); P- 5, 38 (13.2); L P- 2 / P- 4 ratio, 1.1; L I-L- 4 - 6 (Fig. 6 D): 151, 166, 162. Female: Unknown.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	etymology	Etymology. The species is named after the district where it was collected.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	discussion	Discussion. Due to the elongated idiosoma (e. g., dorsal shield L / W> 1.5) and the shape of the capitular bay which is more rounded at its proximal end, Monatractides nipsanicus sp. nov. resembles M. kootoma Cook, 1986, known from Queensland (Australia, Cook 1986). Monatractides nipsanicus sp. nov. differs from the latter species in its major idiosoma and gnathosoma dimensions (e. g., M. kootoma dorsal shield L / W 608 / 391, ventral shield L / W 714 / 425, genital field L / W 140 / 129, palp total L 189; data taken from Cook 1986) and considerably elongated genital field (L / W ratio 1.6 vs. 1.1. in M. kootoma).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC53D82CF9F4C5E2CDEEF80B.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province: Bion River, upstream, Pass Valley, 15. iii. 2010, 3 ˚ 51.84 S 139 ˚ 05.733 E, 1962 m a. s. l. Paratypes: 0 / 1 (mounted), same data as holotype; 1 / 1, ibid., 15. iii. 2010, 3 ˚ 51.513 S 139 ˚ 05.570 E, 2007 m a. s. l.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	diagnosis	Diagnosis. Idiosoma L> 1000 in males,> 1100 in females; relatively wide capitular bay (L / W ratio 2.1 – 2.3); P- 4 with well visible denticle near the insertion of the ventral hairs; Cx- 4 strongly extended posterior to the genital flaps; postgenital area large; excretory pore slightly away from the line of primary sclerotization, Vgl- 2 posterior and well separated from the excretory pore. Morphology. Male (holotype, in parentheses some mesaurements of the non-dissected paratype specimen): Idiosoma (ventral view: Fig. 7 B) L 1044 (1135), W 794 (753); dorsal shield (Fig. 7 A) L 928, W 650, L / W ratio 1.43; dorsal plate L 877; shoulder plate L 254, W 108, L / W ratio 2.35; frontal plate L 194, W 88, L / W ratio 2.2; shoulder / frontal plate L ratio 1.31; capitular bay L 206, W 91, L / W ratio 2.3; Cx- 1 total L 341, Cx- 1 medial L 134, Cx- 2 + 3 medial 84; ratio Cx- 1 L / Cx- 2 + 3 medial L 4.1; Cx- 1 medial L / Cx- 2 + 3 medial L 1.6; genital field L / W 208 / 149, L / W ratio 1.4, ejaculatory complex L 197; distance genital field – excretory pore 234, genital field – caudal idiosoma margin 394; capitulum (Fig. 8 C) ventral L 240; chelicera L 302; palp (Fig. 8 A) total L 289, L and % L (in parentheses): P- 1, 34 (11.8); P- 2, 86 (30.0); P- 3, 53 (18.3); P- 4, 79 (17.3); P- 5, 37 (12.8); L P- 2 / P- 4 ratio, 1.09; L I- L- 4 - 6 (Fig. 8 D): 143, 159, 160.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	description	Female (paratype, in parentheses some mesaurements of the non-dissected paratype specimen): Idiosoma (ventral view: Fig. 9) L 1288 (1313), W 913 (875); dorsal shield L 1128, W 777, L / W ratio 1.45; dorsal plate L 1056; shoulder plate L 309, W 134, L / W ratio 2.31; frontal plate L 225, W 116, L / W ratio 1.9; shoulder / frontal plate L ratio 1.37; capitular bay L 247, W 116, L / W ratio 2.1; Cx- 1 total L 425, Cx- 1 medial L 178, Cx- 2 + 3 medial 58; ratio Cx- 1 L / Cx- 2 + 3 medial L 7.3; Cx- 1 medial L / Cx- 2 + 3 medial L 3.1; genital field L / W 272 / 225, L / W ratio 1.2; distance genital field – excretory pore 309, genital field – caudal idiosoma margin 503; capitulum ventral L 288; chelicera total L 361; palp (Fig. 7 B) total L 346, L and % L (in parentheses): P- 1, 40 (11.6); P- 2, 104 (30.1); P- 3, 64 (18.5); P- 4, 95 (27.5); P- 5, 43 (12.4); L P- 2 / P- 4 ratio, 1.1; L I-L- 4 - 6: 172, 197, 186.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	etymology	Etymology. The species is named after the river where it was collected.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	discussion	Remarks. This species belongs to the M. luteus (K. Viets, 1935) species-complex (see above). Due the larger dimensions of idiosoma and gnathosoma, and in having broad frontal platelets, Monatractides bionus sp. nov. resembles M. landbergi (Lundblad, 1941) and M. roseus (Lundblad, 1941), both known from Java, Indonesia (Lundblad 1956, 1971); the latter species has been tentatively reported from Malaysia (Pešiċ & Smit 2010). Monatractides landbergi differs from the latter species in slightly slender idiosoma, and in having less broad frontal platelets (see Lundblad 1956). Due to these features, M. bionus sp. nov. is more similar to M. landbergi, but can be distinguished from the latter species (in parentheses, data taken from original descriptions, see: Lundblad 1956, 1971) in its larger idiosoma and gnathosoma dimensions (e. g., dorsal shield L / W 670 / 492 in male, L 690 in female; ventral shield L / W 770 / 542 in males, L 800 in female; genital field L / W 167 / 125 in male, 186 / 157 in female; palp total L 218 in male), proportionally wider capitular bay, Cx- 4 strongly extended posterior to the genital flaps and Vgl- 2 more separated from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC50D823F9F4C6E2CDEEFE2C.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	materials_examined	Type series. Holotype, female, dissected and slide mounted, Indonesia, New Guinea, Papua Province, River Pos 7, Sentani, 9. iii. 2010, 2 ˚ 33.740 S 140 ˚ 30.784 E, 103 m asl.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	diagnosis	Diagnosis (Male unknown). Idiosoma elongated (dorsal shield L / W 1.7); the area of primary sclerotization of the dorsal and ventral plate (= dorsal and ventral plate sensu Wiles 1997 b) extending almost to the posterior margin of the dorsal or ventral shield respectively; ventral margin of P- 2 and - 3 with straight stout seta, P- 4 with well developed denticles near the insertion of the ventral hairs; genital field pentagonal; excretory pore incorporated into primary sclerotization of the ventral shield.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	description	Description. Female: Idiosoma (ventral view: Fig. 10 B) L 666, W 444; dorsal shield (Fig. 10 A) L 584, W 344, L / W ratio 1.7; dorsal plate 556; shoulder plate L 178, W 57, L / W ratio 3.1; frontal plate L 106, W 44, L / W ratio 2.4; shoulder / frontal plate L ratio 1.68; capitular bay L 147, W 78, L / W ratio 1.9; Cx- 1 total L 250, Cx- 1 medial L 103, Cx- 2 + 3 medial 25; ratio Cx- 1 L / Cx- 2 + 3 medial L 10.0; Cx- 1 medial L / Cx- 2 + 3 medial L 4.1; genital field L / W 143 / 128, L / W ratio 1.12; distance genital field – excretory pore 137, genital field – caudal idiosoma margin 247; capitulum (Fig. 10 F) ventral L 185; chelicera L 249; palp (Figs. 10 D – E) total L 209, dL and % L (in parentheses): P- 1 29 (13.9), P- 2 59 (28.2), P- 3 38 (18.2), P- 4 55 (26.3), P- 5 28 (13.4); L P- 2 / P- 4 ratio, 1.07; straight stout seta on ventral P- 2 and - 3; L I-L- 4 - 6 (Fig. 10 C): 78, 95, 103. Male: unknown.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	etymology	Etymology. The species is named after the area where it was collected.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	discussion	Discussion. Due to the similar shape of the palp, narrow frontal platelets and shoulder dorsal plates which are not swollen anteriorly, the new species from New Guinea resembles M. neoapratima (Wiles, 1991), known from Malaysia and Thailand (Wiles 1991, Pešiċ & Smit 2009). Monatractides sentanicus sp. nov. can be easily distinguished from the latter species by the following features: the area of primary sclerotization extending almost to the posterior margin of the dorsal or ventral shield respectively, excretory pore incorporated into primary sclerotization of the ventral shield and the genital field which is pentagonal in shape.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5ED821F9F4C579CDEEFEE0.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	materials_examined	Type series. Holotype, female, dissected and slide mounted; Indonesia, New Guinea, Papua Province, River Kamp Walker near Uncen, Abepura, 31. iii. 2010, 2 ˚ 34.202 S 140 ˚ 38.886 E, 144 m a. s. l.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	diagnosis	Diagnosis (Male unknown). Frontal platelets broad (L / W 1.7); the area of primary sclerotization extending almost to the posterior margin of the dorsal or ventral shield respectively; ventral margin of P- 2 and - 3 with slender, hair-like seta, distal margins of P- 3 with denticles; genital field pentagonal; excretory pore incorporated into primary sclerotization of the ventral shield.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	description	Description. Female: Idiosoma (ventral view: Fig. 11 B) L 709, W 484; dorsal shield (Fig. 11 A) L 616, W 422, L / W ratio 1.46; dorsal plate L 559; shoulder plate L 206, W 72, L / W ratio 2.9; frontal plate L 112, W 66, L / W ratio 1.7; shoulder / frontal plate L ratio 1.84; capitular bay L 167, W 69, L / W ratio 2.4; Cx- 1 total L 272, Cx- 1 medial L 103, Cx- 2 + 3 medial 24; ratio Cx- 1 L / Cx- 2 + 3 medial L 11.3; Cx- 1 medial L / Cx- 2 + 3 medial L 4.3; genital field L / W 162 / 141, L / W ratio 1.15; distance genital field – excretory pore 156, genital field – caudal idiosoma margin 252; capitulum (Fig. 11 D) ventral L 177; chelicera total L 229; palp (Fig. 11 C) total L 196, dL and % L (in parentheses): P- 1, 28 (14.3); P- 2, 57 (29.1); P- 3, 34 (17.4); P- 4, 50 (25.5); P- 5, 27 (13.8); L P- 2 / P- 4 ratio, 1.14; L I-L- 5 - 6 (Fig. 11 E): 99, 100. Male: unknown.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	etymology	Etymology. The species is named after the area where it was collected.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	discussion	Discussion. Due to the area of primary sclerotization extending almost to the posterior margin of the dorsal or ventral shield respectively, and the excretory pore incorporated into primary sclerotization of the ventral shield, M. abepurus sp. nov. is close to M. sentanicus sp. nov. It can easily be distinguished from the latter species in the presence of a slender, hair-like seta, not straight stout seta on the ventral margin of P- 2 and P- 3, and in having noticeably broader frontal platelets.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC5BD824F9F4C5F2CDEEFE6C.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province, unnamed creek crossing road to Pass Valley, 13. iii. 2010, 3 ˚ 52.849 S 139 ˚ 04.194 E, 2253 m a. s. l. Paratypes: 1 / 4 (0 / 1 mounted), same data as holotype.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	diagnosis	Diagnosis. The shoulder and frontal platelets of each side are fused to each other to form a pair of elongated platelets; the area of primary sclerotization of the dorsal plate with four dorsoglandularia; capitular bay rounded at its proximal end; P- 4 with well developed tubercles near the insertion of the ventral hairs; postgenital area large; excretory pore on the line of primary sclerotization, Vgl- 2 posterior and well away from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	description	Description. Male: Idiosoma (ventral view: Fig. 12 B) L 938, W 777; dorsal shield (Fig. 12 A) L 822, W 672, L / W ratio 1.22; dorsal plate 769; the shoulder and frontal platelets of each side are fused to each other to form a pair of elongated platelets, L 378, W 109, L / W ratio 3.5; capitular bay L 205, W 82, L / W ratio 2.5; Cx- 1 total L 335, Cx- 1 medial L 130, Cx- 2 + 3 medial 31; ratio Cx- 1 L / Cx- 2 + 3 medial L 10.8; Cx- 1 medial L / Cx- 2 + 3 medial L 4.2; genital field L / W 209 / 156, L / W ratio 1.34; ejaculatory complex (Fig. 13 F) L 225; distance genital field – excretory pore 163, genital field – caudal idiosoma margin 350; capitulum (Fig. 13 E) ventral L 212; chelicera total L 269; palp (Figs. 13 B – C) total L 257, L and % L (in parentheses): P- 1, 27 (10.5); P- 2, 76 (30.0); P- 3, 51 (19.8); P- 4, 71 (27.6); P- 5, 32 (12.5); L P- 2 / P- 4 ratio, 1.07; L I-L- 4 - 6 (Fig. 13 D): 115, 141, 142. Female: Idiosoma (ventral view: Fig. 13 A) L 1022, W 906; dorsal shield L 881, W 756, L / W ratio 1.17; dorsal plate 831; the shoulder and frontal platelets of each side are fused to each other to form a pair of elongated platelets, L 372, W 113, L / W ratio 3.3; capitular bay L 214, W 83, L / W ratio 2.58; Cx- 1 total L 333, Cx- 1 medial L 119, Cx- 2 + 3 medial 18; ratio Cx- 1 L / Cx- 2 + 3 medial L 18.5; Cx- 1 medial L / Cx- 2 + 3 medial L 6.6; genital field L / W 231 / 195, L / W ratio 1.19; distance genital field – excretory pore 184, genital field – caudal idiosoma margin 428; capitulum ventral L 225; chelicera total L 289; palp total L 278, dL and % L (in parentheses): P- 1, 32 (11.5); P- 2, 83 (30.0); P- 3, 52 (18.7); P- 4, 77 (27.7); P- 5, 34 (12.2); P- 2 / P- 4 ratio, 1.08; L I-L- 4 - 6: 125, 146, 150.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	etymology	Etymology. The species is named after the Sahul Shelf.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	discussion	Discussion. Monatractides sahuli sp. nov. is distinguished from all other New Guinean species of the genus by the shoulder and frontal platelets of each side being fused to each other. This character is also found in several other species of Monatractides: M. acutiscutatus (Viets, 1913) from Cameroon (Viets 1913), M. hesperia (Lundblad, 1941) from Colombia (Lundblad 1941) and M. veracruzensis (Cook, 1980) from Mexico (Cook 1980). K. Viets (1931) proposed the subgenus ‘ Rusetriella ’ for these species, which has been rejected by Wiles (1997 b) based on phylogenetic analysis. Monatractides sahuli sp. nov. can be easily distinguished by the shape of the capitular bay which is more rounded at its proximal end and the presence of four dorsoglandularia in the area of primary sclerotization of the dorsal plate vs. the more pointed, V-shaped capitular bay and the area of primary sclerotization of the dorsal plate with two dorsoglandularia in the three aforementioned species.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC59D83AF9F4C29CCDEEFCB8.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province: Bion River, upstream, Pass Valley, 15. iii. 2010, 3 ˚ 51.84 S 139 ˚ 05.733 E, 1962 m asl. Paratypes: 1 / 1 (0 / 1 mounted), same data as holotype; 2 / 2 (1 / 1 mounted), same data as holotype, 15. iii. 2010, 3 ˚ 51.513 S 139 ˚ 05.570 E, 2007 m a. s. l ..; 3 / 1, same data as holotype, 02. iii. 2010, 3 ˚ 51.267 S 139 ˚ 05.789 E, 1962 m a. s. l. Further records: Indonesia, New Guinea, Papua Province: Wao & Pao streams near confluence Walmak, Nipsan district, 18. iii. 2010, 4 ˚ 07.097 S 139 ˚ 38.520 E, 1739 m a. s. l. 1 / 0 (mounted); Yango stream, Walmak, Nipsan district, 19. iii. 2010, 4 ˚ 06.830 S 139 ˚ 38.587 E, 1757 m a. s. l. 2 / 2 (1 / 0 mounted).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	diagnosis	Diagnosis. Idiosoma L> 1100 in males,> 1200 in females; area of primary sclerotization of the dorsal plate with four dorsoglandularia; capitular bay narrow (L / W ratio ca. 3.0); distal margins of P- 2 and - 3 without denticles, P- 4 with small denticle near the insertion of the ventral hairs, P- 4 ventral hairs inserted more distally, P- 4 L / H 2.5 – 2.6; postgenital area large; excretory pore away from the line of primary sclerotization, Vgl- 2 posterior and well separated from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	description	Description. Male (holotype, in parentheses paratype from Bion River, in square brackets specimen from Wao & Pao streams): Idiosoma: (ventral view: Fig. 14 B) L 1163 (1219) [1100], W 806 (900) [744]; dorsal shield (Fig. 14 A) L 906 (996) [878], W 700 (781) [644], L / W ratio 1.29 (1.28) [1.36]; dorsal plate 860 (934) [825]; shoulder plate L 259 (278) [228], W 97 (109) [97], L / W ratio 2.7 (2.6) [2.4]; frontal plate L 166 (209) [178], W 71 (92) [81], L / W ratio 2.3 (2.3) [2.2]; capitular bay L 258 (256) [238], W 82 (81) [80], L / W ratio 3.15 (3.16) [3.0]; Cx- 1 total L 416 (433) [392], Cx- 1 medial L 157 (175) [153], Cx- 2 + 3 medial 80 (70) [68]; ratio Cx- 1 L / Cx- 2 + 3 medial L 5.2 (6.2) [5.8]; Cx- 1 medial L / Cx- 2 + 3 medial L 2.0 (2.5) [2.25]; genital field L / W 238 (261) [219] / 169 (184) [159], L / W ratio 1.4 (1.4) [1.38], ejaculatory complex (Fig. 15 B) L 266 (292) [225]; distance genital field – excretory pore 216 (259) [206], genital field – caudal idiosoma margin 409 (440) [409]; capitulum (Fig. 15 D) ventral L 272 (275) [247]; chelicera total L 313 (314) [303]; palp (Fig. 15 A) total L 292 (316) [294], dL: P- 1, 39 (40) [39]; P- 2, 87 (92) [85]; P- 3, 66 (65) [60]; P- 4, 74 (86) [79]; P- 5, 26 (33) [31]; % L: P- 1, 13.4 (12.7) [13.3]; P- 2, 29.8 (29.1) [28.9]; P- 3, 22.6 (20.6) [20.4]; P- 4, 25.3 (27.2) [26.9]; P- 5, 8.9 (10.4) [10.5]; L P- 2 / P- 4 ratio, 1.18 (1.07) [1.08]; L I-L- 4 - 6 (Fig. 15 C): 151 (169) [160], 162 (178) [157], 145 (159) [142]. Female (from Bion River, upstream, n = 2): Idiosoma (ventral view: Fig. 16 A) L 1263 – 1356, W 988 – 1000; dorsal shield L 1081 – 1109, W 850 – 856, L / W ratio 1.27 – 1.3; dorsal plate L 1013 – 1044; shoulder plate L 256 – 272, W 109 – 122, L / W ratio 2.1 – 2.5; frontal plate L 203 – 213, W 97 – 100, L / W ratio 2.03 – 2.26; shoulder / frontal plate L ratio 1.24 – 1.26; capitular bay L 266 – 279, W 86 – 91, L / W ratio 3.1; Cx- 1 total L 431 – 466, Cx- 1 medial L 164 – 184, Cx- 2 + 3 medial 37 – 38; ratio Cx- 1 L / Cx- 2 + 3 medial L 11.3 – 12.3; Cx- 1 medial L / Cx- 2 + 3 medial L 4.3 – 5.0; genital field L / W 269 – 278 / 218 – 222, L / W ratio 1.2 – 1.28; distance genital field – excretory pore 248 – 306, genital field – caudal idiosoma margin 497 – 552; capitulum ventral L 291 – 294; chelicera L 328 – 335; palp (Fig. 16 B) total L 330 – 331, dL and % L (in parentheses): P- 1, 43 – 47 (13.0 – 14.2); P- 2, 96 (29.0 – 29.1); P- 3, 68 – 70 (20.6 – 21.1); P- 4, 86 – 89 (26.0 – 27.0); P- 5, 32 – 34 (9.7 – 10.3); P- 2 / P- 4 ratio, 1.08 – 1.12; L I-L- 4 - 6: 174 – 175, 182 – 188, 162 – 169.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	discussion	Discussion. Monatractides alticolus sp. nov. and M. humilis sp. nov. (see below) belong to M. macroporus (Viets, 1935) species-complex (see Pešiċ & Smit 2009). Due to the less narrow capitular bay (L / W 3.0), larger idiosoma and gnathosoma dimensions and the presence of four dorsoglandularia in the area of primary sclerotization of the dorsal plate, the specimens from New Guinea are close to M. major (Viets, 1935), a species described originally from Java as a form of M. macrognathus (Viets 1935). Lundblad (1971) considered M. macrognathus and M. macrognathus major, synonymous with M. macroporus, but Wiles (1991) raised M. macrognathus major in ranking to a full species. Both species of the M. macroporus (Viets, 1935) species-complex from New Guinea, M. alticolus sp. nov. and M. humilis sp. nov. (see below) differ from M. major (and the other members of the macroporus - complex from Java, see Viets 1935) in Vgl- 2 shifted far away from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	etymology	Etymology. Named for its occurrence at higher elevations.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC47D839F9F4C0E0CDEEFCBC.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	materials_examined	Type series. Holotype, male, dissected and slide mounted; Indonesia, New Guinea, Papua Province: River Pos 7, Sentani, 9. iii. 2010, 2 ˚ 33.740 S 140 ˚ 30.784 E, 103 m a. s. l .. Paratypes: 1 / 1 (mounted), same data as holotype; 8 / 5 (2 / 2 mounted), River Yabawi, upstream of Harapan, 28. iii. 2010, 2 ˚ 34.216 S 140 ˚ 33.723 E, 120 m a. s. l.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	diagnosis	Diagnosis. Idiosoma L <900 in males, <1000 in females; area of primary sclerotization of the dorsal plate with four dorsoglandularia; capitular bay narrow (L / W ratio ca. 3.0); distal margins of P- 2 and - 3 without denticles, P- 4 with well well visible denticle near the insertion of the ventral hairs, P- 4 L / H 2.2 – 2.3; postgenital area large; excretory pore away from the line of primary sclerotization, Vgl- 2 posterior and well separated from the excretory pore.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	description	Description. Male (holotype, in parentheses paratype from River Pos 7, in square brackets paratype from River Yabawi): Idiosoma (ventral view: Fig. 17 B) L 788 (813) [878], W 543 (537) [576]; dorsal shield (Fig. 17 A) L 625 (697) [681], W 481 (488) [500], L / W ratio 1.3 (1.43) [1.36]; dorsal plate 597 [644]; shoulder plate L 153 (184) [168], W 69 (75) [72], L / W ratio 2.2 (2.5) [2.3]; frontal plate L 113 (128) [131], W 47 (59) [61], L / W ratio 2.4 (2.2) [2.15]; shoulder / frontal plate L ratio 1.35 (1.44) [1.28]; capitular bay L 169 (178) [194], W 59 (67) [66], L / W ratio 2.9 (2.7) [2.9]; Cx- 1 total L 284 (297) [325], Cx- 1 medial L 114 (119) [131], Cx- 2 + 3 medial 76 (81) [74]; ratio Cx- 1 L / Cx- 2 + 3 medial L 3.7 (3.7) [4.39]; Cx- 1 medial L / Cx- 2 + 3 medial L 1.5 (1.47) [1.77]; genital field L / W 160 (172) [179] / 113 (118) [126], L / W ratio 1.4 (1.46) [1.42], ejaculatory complex (Fig. 17 D) L 189 (188); distance genital field – excretory pore 153 (158) [170], genital field – caudal idiosoma margin 259 (253) [281]; capitulum (Fig. 17 G) ventral L 188 (202) [213]; chelicera total L 222 (234) [250]; palp (Fig. 17 G) total L 207 (212) [229], dL: P- 1, 28 (29) [33]; P- 2, 59 (62) [68]; P- 3, 39 (42) [42]; P- 4, 58 (55) [60]; P- 5, 23 (24) [26]; % L: P- 1, 13.5 (13.7) [14.4]; P- 2, 28.5 (29.2) [29.7]; P- 3, 18.8 (19.8) [18.3]; P- 4, 28.0 (26.0) [26.2]; P- 5, 11.1 (11.3) [11.4]; P- 2 / P- 4 ratio, 1.02 (1.13) [1.14]; L I-L- 4 - 6 (Fig. 17 C): 114 (119) [128], 114 (125) [130], 115 (116) [126]. Female (from River Pos 7, in parentheses paratype from River Yabawi): Idiosoma (ventral view: Fig. 17 H) L 925 (975), W 625 (669); dorsal shield L 744 (794), W 550 (563), L / W ratio 1.35 (1.4); dorsal plate 706 (759); shoulder plate L 197 (191), W 78 (78), L / W ratio 2.5 (2.45); frontal plate L 124 (141), W 59 (59), L / W ratio 2.1 (2.39); shoulder / frontal plate L ratio 1.59 (1.36); capitular bay L 194 (200), W 72 (72), L / W ratio 2.7 (2.8); Cx- 1 total L 319 (325), Cx- 1 medial L 125 (125), Cx- 2 + 3 medial 59 (68); ratio Cx- 1 L / Cx- 2 + 3 medial L 5.4 (4.8); Cx- 1 medial L / Cx- 2 + 3 medial L 2.1 (1.8); genital field L / W 188 (194) / 154 (158), L / W ratio 1.22 (1.23); distance genital field – excretory pore 200 (197), genital field – caudal idiosoma margin 350 (367); capitulum ventral L 225 (222); chelicera total L 257 (259); palp total L 240 (245), dL: P- 1, 33 (34); P- 2, 69 (71); P- 3, 45 (46); P- 4, 65 (66); P- 5, 28 (28); % L: P- 1, 13.8 (13.9); P- 2, 28.8 (29.0); P- 3, 18.8 (18.8); P- 4, 27.1 (26.9); P- 5, 11.7 (11.4); L P- 2 / P- 4 ratio, 1.06 (1.08); L I-L- 4 - 6: 129 (132), 125 (132), 119 (127).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	etymology	Etymology. Named for its occurrence at lower elevations.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	discussion	Discussion. See discussion section under the preceeding species.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC44D83CF9F4C0EDCDEEFE94.taxon	distribution	Distribution. New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC41D83CF9F4C2C5CE58FB93.taxon	materials_examined	Material examined. Holotype male, Torrenticola papillata, River Rumei, Waigeo, Indonesia, 17. x. 1993, leg. M. Argeloo (slide INDON 31, ZMAN). Morphology. Male (in parentheses measurements taken from the original description): Idiosoma (ventral view: Fig. 18 B) L 766, W 691; dorsal shield (Fig. 18 A) L 706, W 509, L / W ratio 1.39; dorsal plate 644 (689); shoulder plate L 209 (204 - 213), W 97, L / W ratio 2.2; frontal plate L 138 - 141 (132), W 75 - 78, L / W ratio 1.8 - 1.84; shoulder / frontal plate L ratio 1.48; capitular bay L 128; Cx- 1 total L 233 (233), Cx- 1 medial L 103, Cx- 2 + 3 medial 55 (60); ratio Cx- 1 L / Cx- 2 + 3 medial L 4.2; Cx- 1 medial L / Cx- 2 + 3 medial L 1.87; genital field L / W 148 (155) / 120 (126), L / W ratio 1.23; ejaculatory complex conventional in shape, L 178; distance genital field – excretory pore 176, genital field – caudal idiosoma margin 325; capitulum ventral L 181; chelicera total L 229; palp (Fig. 18 C) total L 224, dL: P- 1, 29 (29); P- 2, 66 (67); P- 3, 42 (43); P- 4, 61 (58); P- 5, 26 (26); L P- 2 / P- 4 ratio, 1.08; P- 4 with well developed setal tubercles; postgenital area large; excretory pore on the line of primary sclerotization, Vgl- 2 posterior and well away from the excretory pore; L I- 5 - 6 (Fig. 18 D): 135 (127), 136 (108).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC41D83CF9F4C2C5CE58FB93.taxon	discussion	Remarks. Due to the broad frontal platelets (L / W 1.8), shoulder dorsal plates which are swollen anteriorly, and P- 4 with large setal tubercles, Monatractides papillatus can be easily distinguished from the other members of this genus from New Guinea.	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
03F1878DDC41D83CF9F4C2C5CE58FB93.taxon	distribution	Distribution. Waigeo (Indonesia).	en	Pešić, Vladimir, Smit, Harry (2011): Water mites of the genus Monatractides Viets (Acari: Hydrachnidia, Torrenticolidae) from New Guinea, with descriptions of nine new species. Zootaxa 2779: 39-62, DOI: 10.5281/zenodo.207512
