identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F187A07C04CD3127B79AE8FDEBFCB6.text	03F187A07C04CD3127B79AE8FDEBFCB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta Mickel 1936	<div><p>Genus Lomachaeta Mickel, 1936</p><p>Type species: L. hicksi Mickel, 1936</p></div>	https://treatment.plazi.org/id/03F187A07C04CD3127B79AE8FDEBFCB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3127B79B3DFC16FA85.text	03F187A07C04CD3127B79B3DFC16FA85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Smicromutilla Mickel 1964	<div><p>Smicromutilla Mickel, 1964 .</p><p>Type species: S. powelli Mickel, 1964 . Synonymized by Williams &amp; Pitts (2009).</p><p>Diagnosis. FEMALE. Females of Lomachaeta can be recognized by the following combination of characters: the body setae are sparse, nowhere obscuring the sculpture; the mesosoma has coarse sculpture and lacks a scutellar scale; T1 is slender, with its shape either sub-sessile or sub-disciform; the tergal fringes are not composed of distinctly plumose setae; and T6 lacks a defined pygidium.</p><p>MALE. Males of Lomachaeta can be recognized by the following combination of characters: the body setae are sparse, nowhere obscuring the sculpture; the axilla is armed with a posteriorly directed tooth; T1 is slender, with its shape either sub-sessile or disciform; the tergal fringes are not composed of distinctly plumose setae; and the digitus is sessile with the usually short pad-like cuspis.</p><p>Distribution. Widespread in the New World, from Canada to Argentina.</p></div>	https://treatment.plazi.org/id/03F187A07C04CD3127B79B3DFC16FA85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3027B79D2AFA7EFA3C.text	03F187A07C04CD3027B79D2AFA7EFA3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta (Females) Mickel 1936	<div><p>Key to species of Lomachaeta (Females)</p><p>1. Mesosoma and T2 disc with matching thickened posteriorly-directed subparallel appressed silvery to golden setae (Figs 1–3; North America)...................................................................................... 2</p><p>- Mesosoma and T2 not having matching appressed posteriorly-directed silvery golden setae; dorsum usually without any appressed thickened setae, but if thick setae present on mesonotum, then they are not predominantly subparallel and the T2 disc has erect sparse setae only (e.g. Figs 9, 12, 21, 33; North or South America).................................. 3</p><p>2(1). Mesosoma elongate, thoracic dorsum nearly as long as wide; head wider (~1.2×) than mesosoma (Figs 2, 3, 6, 7; Texas)........................................................................ L. vacamuerta Williams &amp; Pitts, 2009</p><p>- Mesosoma more compact, thoracic dorsum shorter (0.85×) than wide; head as wide as mesosoma (Figs 1, 5; Arizona, Nevada, Utah).............................................................. L. argenta Pitts &amp; Manley, 2004</p><p>3(1). Areolations on frons and vertex broad, most transverse intervals obliterated, leaving variously defined apparent longitudinal carinae; lateral pronotal and propodeal faces with areolations having intervals more clearly defined, at least one complete areolation on mesopleuron and lateral propodeal face; body variegated dark and light brown or primarily dark-brown (Figs 9, 10, 15, 16; Central and South America only)................................... L. hyphantria Pitts &amp; Manley, 2004</p><p>- Areolations on frons and vertex variable, either complete or with obliterated edges not oriented to leave longitudinal carinae; lateral pronotal and propodeal faces often with areolations largely obliterated; body often predominantly orange-brown (North, Central, or South America)...................................................................... 4</p><p>4(3). South American species; mesosomal areolations dense, intervals narrow and even, not scabrous (Figs 12–14)........... 5</p><p>- North and Central American species; mesosomal sculpture variable, areolations often sparser with intervals partially obliterated, sometimes appearing scabrous (e.g. Figs 22, 23, 33, 34)................................................. 7</p><p>5(4). Mesosoma, metasoma, and legs black, head orange-brown; entire body except T2 disc with appressed setae thickened, white, contrasting with integument (Figs 12, 18; São Paulo, Brazil)............................. L. meloi Williams, sp. nov.</p><p>- Head and mesosoma reddish brown, often darkened laterally, metasoma blackish, always darker than head; body setae predominantly brown.................................................................................... 6</p><p>6(5). Propodeal spiracle prominently dentiform; scape as long as pedicel and F1–3 combined (Figs 11, 17; Buenos Aires, Argentina).............................................................................. L. ibarrai Casal, 1969</p><p>- Propodeal spiracle lowly rounded, not dentiform; scape shorter than pedicel and F1–3 combined (Figs 13, 14, 19, 20; Argentina, Brazil, and Paraguay)............................................................. L. vianai Casal, 1969</p><p>7(4). Mandible with baso-ventral subhyaline lamella, mandible narrowed beyond lamella (Fig. 65); pronotal and propodeal spiracles swollen and apparently tuberculate (Figs 22, 23); T1 shape sub-disciform, often with dorsal and anterior faces set apart by angle or incomplete carina; T2 fringe usually composed of thick silvery setae (Figs 21, 23–26); mesosomal dorsum often with dense areolations, at least anteriorly (Figs 21, 24); frequently encountered in trap nests......................... 8</p><p>- Mandible without basal lamella, if weak lamella apparently present, it does not interrupt ventral mandible contour (Fig. 66); pronotal and propodeal spiracles usually not so swollen (e.g. Figs 33–36); T1 shape sub-sessile, dorsal and anterior faces evenly rounded together; T2 fringe usually composed of sparse slender setae (e.g. Figs 33–35); mesosomal dorsum never with areolations dense; not known from trap nests......................................................... 10</p><p>8(7). Head and mesosoma reddish brown, metasomal segments 2–6 uniform blackish; mesosomal dorsum with tight subuniform areolations; T6 with dense thickened long setae that are black basally and whitish apically (Figs 21, 27; subtropical Mexico to Costa Rica)........................................................... L. chionothrix Pitts &amp; Manley, 2004</p><p>- Variably colored, usually variegapted light and dark brown, but if metasoma predominantly blackish, then head and mesosoma also dark; mesosomal dorsum usually with fewer areolations that are often broader posteriorly; T6 lacking long thick bi-colored setae (widespread in North America)............................................................ 9</p><p>9(8). Body unicolorous pale orange-brown, at most with T6 largely blackened and/or legs and antennae at most darkened apically; T3–6 setae mostly blackish (Figs 22, 23, 28, 29; southwestern USA and northwestern Mexico).............................................................................................. L. cirrhomeris Pitts &amp; Manley, 2004</p><p>- Body generally darker, variegated with orange, brown, and blackish, legs and antennae variably blackened; T3–6 with interspersed silvery and blackish setae (Figs 24–26, 30–32; widespread in North America)............ L. hicksi Mickel, 1936</p><p>10(7). Mesosoma elongate, thoracic length subequal to width; mesosomal areolations deep and distinct, with all intervals distinct; frons mostly with complete areolations (Figs 4, 8; Madrean Archipelago in Arizona)........... L. osita Williams, sp. nov.</p><p>- Mesosoma not so elongate, thoracic length less than 0.9 × width; mesosomal areolations with margins less distinct and often partly obliterated; frons generally with obscure punctures and shallow rounded intervals (e.g. Figs 33–36)............. 11</p><p>11(10). T2 sculpture mostly smooth between punctures (microreticulations not visible at 30 × magnification, obscure apparent reticulations sometimes visible at&gt; 80 × magnification); dorsal bristles of mesonotum and T2 blackish, many bristles nearly as long as scape (Figs 35, 39; Central Valley and Coast Ranges, California).................... L. powelli (Mickel), 1964</p><p>- T2 sculpture generally microreticulate between punctures (visible at 30 × magnification); if microreticulations obscure, then dorsal bristles of mesonotum and T2 mostly brown or yellowish, generally much shorter than scape (arid and semi-arid habitats South or East of Sierra Nevada and related ranges)...................................................... 12</p><p>12(11). Antennae and forelegs entirely blackish, middle and hind legs mostly black (Figs 33, 37; Baja California Sur).................................................................................... L. hedera Williams &amp; Pitts, 2009</p><p>- Antennae and legs largely orange-brown, at least scape and fore coxa lighter brown (Figs 34, 36, 38, 40).............. 11</p><p>13(12). Body orange-brown, at most with T6 blackened (Figs 34, 38; Mojave and Sonoran Deserts in California and Arizona).......................................................................... L. beadugrimi (Pitts &amp; Manley, 2004)</p><p>- Femora and T2–5 variably darkened brown, in part clearly darker than orange-brown mesosoma (Figs 36, 40; Colorado Plateau and Sonoran Desert in Arizona).............................................. L. warneri Williams, sp. nov.</p></div>	https://treatment.plazi.org/id/03F187A07C04CD3027B79D2AFA7EFA3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C05CD3727B79DB7FA79FA11.text	03F187A07C05CD3727B79DB7FA79FA11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta (Males) Mickel 1936	<div><p>Key to species of Lomachaeta (Males)</p><p>1. Ventral margin of mandible armed with strong tooth basally................................................... 2</p><p>- Ventral margin of mandible lacking tooth basally............................................................ 4</p><p>2(1). Femora and tegulae orange brown, contrasting with entirely black mesosoma and metasoma (Fig. 42; southwestern USA and northwestern Mexico)..................................................... L. cirrhomeris Pitts &amp; Manley, 2004</p><p>- Femora and tegula usually blackish, mesosoma blackish, metasoma often partly reddened; if femora orange, then metasoma also predominantly orange (Figs 44–45)................................................................... 3</p><p>3(2). Penis valve with one basal tooth (see Pitts &amp; Manley 2004, Fig. 16–18); body color variable, entirely black or with metasoma cuticle variably orange to red (Figs 43–46; widespread in Nearctic)............................ L. hicksi Mickel, 1936</p><p>- Penis valve with two basal teeth (see Pitts &amp; Manley 2004, Fig. 14); body entirely black (Fig. 42; Nayarit, Mexico South to Costa Rica)............................................................. L. chionothrix Pitts &amp; Manley, 2004</p><p>4(1). Metasoma largely red to orange (e.g. Figs 47–52; North America only).......................................... 5</p><p>- Body entirely black except tegula, head, or legs sometimes orange or lighter brown (Figs 53–64; North or South America). 9</p><p>5(4). Paramere broadly flattened throughout, rounded apically (Arizona, California, and Nevada; Fig. 47, also see Pitts &amp; Manley 2004, Fig. 23).......................................................... L. beadugrimi (Pitts &amp; Manley 2004)</p><p>- Paramere acuminate apically, at most laterally compressed (e.g. Pitts &amp; Manley 2004, Figs 14–22).................... 6</p><p>6(5). Wing venation reduced, cells contained in basal half of forewing (Figs 48, 49); paramere lacking row of elongate setae ventrally (e.g. Fig. 67).................................................................................... 7</p><p>- Wing venation normal, cells extending beyond basal half of forewing (e.g. Fig. 52); paramere with somewhat uniform row of long setae ventrally (see Manley &amp; Pitts 2004; Fig. 19, 20)................................................... 8</p><p>7(1). Head and mesosoma orange, concolorous with T2 (Fig. 49); paramere laterally compressed, moderately setose (Fig. 49, 50; Riverside County, California)................................................. L. calamondin Williams, sp. nov.</p><p>- Head and mesosoma black (Fig. 48); paramere not so compressed, basically subcylindrical, sparsely setose (Central Valley and Coast Ranges, California)....................................................... L. powelli (Mickel), 1964</p><p>8(7). Pronotum and mesoscutum with IS more than twice PD; metasoma blackish with orange color restricted to T2 and T3 (Fig. 51; Sonoran Desert in Arizona and California, USA and Oaxaca, Mexico)............ L. ptilohyalus Pitts &amp; Manley, 2004</p><p>- Pronotum and mesoscutum with IS less than twice PD; metasoma mostly orange (Fig. 52; Great Basin to Chihuahuan Desert in southwestern Nearctic Region)............................................ L. crocopinna Pitts &amp; Manley, 2004</p><p>9(7). Apical fringe of T2 with row of convergent thick bristles (e.g. Figs 53, 54, 57; North and South America).............. 10</p><p>- Apical fringe of T2 with simple setae only (e.g. Figs 58, 60, 63; North America).................................. 13</p><p>10(9). Vertex with separated punctures, IS=PD; paramere having long setae on external surface medially (see Williams &amp; Pitts 20 0 9, Fig. 15; Sonora, Mexico)..................................................... L. theresa Williams &amp; Pitts, 2009</p><p>- Frons and vertex coarsely punctate or areolate; paramere lacking long setae except sometimes at apex (South America)... 11</p><p>11(10). T1 shape nodose, anterior face convex (Fig. 56); apical third of paramere distinctly downcurved (Fig. 71; Colombia).................................................................................... L. juanita Williams, sp. nov.</p><p>- T1 shape disciform, anterior face flat or concave (Figs 53, 55); paramere basically straight or evenly downcurving throughout its length (e.g. Manley &amp; Pitts 2004, Fig. 21).............................................................. 12</p><p>12(11). Gena weakly carinate, apex of paramere with weak tuft of long setae (see Williams &amp; Pitts 2009, Fig. 7; Argentina, Brazil, and Paraguay)....................................................................... L. vianai Casal, 1969</p><p>- Genal carina distinct; apex of paramere lacking setal tuft (see Manley &amp; Pitts 2004, Fig. 21; Northern South America)............................................................................. L. hyphantria Pitts &amp; Manley, 2004</p><p>13(9). Paramere broadly flattened, rounded apically (Fig. 69; also see Williams &amp; Pitts 2009, Fig. 13)..................... 14</p><p>- Paramere acuminate apically, at most laterally compressed (e.g. see Williams &amp; Pitts 2009, Fig. 1)................... 15</p><p>14(13). Cuspis and paramere with all setae shorter than paramere width (see Williams &amp; Pitts 2009, Fig. 13; Sonoran Desert in California)................................................................ L. snellingella Williams &amp; Pitts, 2009</p><p>- Cuspis and paramere with setae much longer than paramere width (Figs 59, 69; Sonora, Mexico)................................................................................................... L. eironeia Williams, sp. nov.</p><p>15(13). Paramere lacking long setae, all setae shorter than paramere width (see Williams &amp; Pitts 2009, Fig. 9; Arizona, USA and Sonora, Mexico).................................................... L. polemomechana Williams &amp; Pitts, 2009</p><p>- Paramere with at least some ventral or apical setae longer than mid-paramere width............................... 16</p><p>16(15). Entire paramere with continuous ventral setal row (see Williams &amp; Pitts 2009, Fig. 3; Mohave and Sonoran Deserts in California and Nevada)............................................................... L. ilex Williams &amp; Pitts, 2009</p><p>- Long ventral setae of paramere absent from basal half....................................................... 17</p><p>17(16). Paramere narrow throughout, long cylindrical, apically convergent, and evenly curving ventrally throughout length (Fig. 63, also see Williams &amp; Pitts 2009, Fig. 5; Arizona, USA and Sonora, Mexico)........... L. litosisyra Williams &amp; Pitts, 2009</p><p>- Paramere evenly tapering toward apex, somewhat laterally compressed, subparallel, and virtually straight to apex or scarcely downcurving (e.g. Figs 62, 64)......................................................................... 18</p><p>18(17). Long setae of paramere restricted to tuft in apical fifth of free length (see Williams &amp; Pitts 2009, Fig. 17; Texas, New Mexico, and Arizona, USA and Sonora, Mexico)..................................... L. vacamuerta Williams &amp; Pitts, 2009</p><p>- Long setae of paramere scattered through apical half of free length (see Williams &amp; Pitts 2009, Fig. 1; Baja California Sur, Mexico).................................................................. L. hedera Williams &amp; Pitts, 2009</p></div>	https://treatment.plazi.org/id/03F187A07C05CD3727B79DB7FA79FA11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.text	03F187A07C02CD3627B79E5BFD66FD0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta argenta Pitts & Manley 2004	<div><p>Lomachaeta argenta Pitts &amp; Manley, 2004</p><p>(Figs 1, 5)</p><p>Lomachaeta argenta Pitts &amp; Manley, 2004: 5 . Holotype, ♀, USA, Utah, Washington Co. (EMUS). Pitts &amp; Manley (2004): Hosts.</p><p>Diagnosis. FEMALE. This species can be recognized by its setal distribution and body shape: the mesosoma and T2 have matching subparallel subappressed long silver to golden setae and the dorsal thoracic length is 0.85 × its width. The following characters may also aid in diagnosis: the head width is subequal to the mesosomal width; the baso-ventral mandibular margin lacks a hyaline lamella; the pronotal spiracle is only weakly swollen; the lateral mesonotal tooth is distinct; the propodeal spiracle is slightly more sharply swollen than the pronotal spiracle; the mesonotal dorsum has about 12 areolations between the pronotal spiracles; the T1 shape is sub-disciform; T2 has dense punctures and mostly smooth intervals; the fringes of T2–3 are composed of dense setae thicker than the erect setae surrounding them; and the S6 lateral carina is small and sharply triangular. Body length 3–3.5 mm.</p><p>MALE. Unknown.</p><p>Material examined. USA: Arizona: Yavapai County, 11 mi W Jerome, Rearing 7598, F. D. Parker (2♀, EMUS) ; Coconino County, 7 mi S. Sedona, Reared, F. D. Parker (2♀, EMUS); Mohave County, Oatman 3 mi S, Reared, F. D. Parker (1♀, EMUS) ; Nevada, Esmeralda County, 1.5 mi. S, 4 mi. W Lida Summit, 7400’, 28.III– 1.X.1982, D. Giuliani, ethylene glycol pit trap (1♀, CSCA) ; Nye County, Ash Meadows, NWR [Natural Wildlife Reserve], Non-dune salt soil site, S. Devils hole rd. BR, 11s0563221 4029908, 11.V.2009, N. Boehme (1♀, EMUS) ; Utah, Washington County, 3 mi W Hurricane, reared, F. D. Parker (1♀, paratype, EMUS, Figs 1, 5) .</p><p>Distribution. Known from the Great Basin, Mojave Desert, and the Arizona Mountains forests in Arizona, Nevada, and Utah, USA.</p><p>Remarks. This species, formerly recognized from one Great Basin locality in southern Utah, is newly recorded from Arizona and Nevada in two additional ecoregions. The newly identified specimens from Arizona were collected from trap nests that also included L. crocopinna, L. cirrhomeris, and L. hicksi . Since the females of L. hicksi and L. cirrhomeris are already known, L. crocopinna could be the unknown male of L. argenta . These specimens, however, are far removed from the type locality of L. crocopinna, so we prefer to wait for more evidence before formalizing this synonymy.</p></div>	https://treatment.plazi.org/id/03F187A07C02CD3627B79E5BFD66FD0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.text	03F187A07C03CD3527B79AA0FDE0FCB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta beadugrimi (Pitts & Manley 2004)	<div><p>Lomachaeta beadugrimi (Pitts &amp; Manley, 2004)</p><p>(Figs 34, 38, 47, 66)</p><p>Smicromutilla beadugrimi Pitts &amp; Manley, 2004: 20 . Holotype, ♂, USA, California, San Bernardino Co. (EMUS). Williams &amp; Pitts (2009): Combination under Lomachaeta .</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. beadugrimi: the body is entirely black, except T2–3 are largely orange; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the forewing has veins encompassing the basal 0.7 × wing; the T1 shape is sub-sessile; the T2 fringe is composed of simple setae; and the paramere is dorsoventrally flattened, apically rounded, and has short setae only. Body length 3.5–5 mm.</p><p>FEMALE. This species can be recognized by the following combination of characters: the body is predominantly orange, except the antennae, legs, and T6 are largely dark-brown; the mandible lacks a distinct ventral lamella; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the mesosoma is compact, with the thoracic dorsal length 0.8 × its width; the T2 disc is sparsely punctate, clothed with erect blackish brachyplumose setae, with the intervals microreticulate; the T2 fringe is sparse and simple; and the S6 lateral carina is distinctly triangular. Body length 2.5–3 mm.</p><p>Description. FEMALE. (hitherto unrecognized, based on female from Kern County). Body length 2.5–3 mm. Coloration. Body reddish-orange except F2–10 dark brown; femora, tibiae, and tarsi dark brown apically; and T6 dark brown. Tibial spurs white. Erect dorsal brachyplumose setae on frons, vertex, mesonotum, T2 disc, and T6 black-brown; on vertex, pronotum, propodeum, T1, and T2 base white; and on T3–5 interspersed brown and white. Mesonotum with few scattered subappressed simple white setae; fringes of T2–5 composed of sparse subappressed simple white setae. Ventral and pleural setae white, except S6 setae black-brown. Head. Head width subequal to pronotal width. Frons, vertex and gena areolate, some vertex intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe without dorsal carina. Length of F1 subequal to pedicel; F2 1.1 × pedicel length. Mesosoma. Mesosoma compact; dorsal thoracic length 0.8 × width. Humeral carina angulate dorsally, weak ventrally, not reaching epaulet. Mesosomal dorsum areolate; most intervals clearly defined, not raised into tubercles; areolations evenly distributed; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle distinct, small. Lateral propodeal face impunctate. Metasoma. T1 shape sub-sessile, subareolate. T2 slightly longer than wide. Disc of T2 with sparse punctures, intervals microreticulate. T3–5 and S2–5 with separated punctures, intervals microreticulate. T6 convex, incomplete lateral carina triangular.</p><p>Material examined. USA: Arizona: Maricopa County, Gila River, 10 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.758835&amp;materialsCitation.latitude=33.22167" title="Search Plazi for locations around (long -112.758835/lat 33.22167)">Arlington</a>, malaise on sand beach, 200 m, 33°13.3’N 112°45.53’W, 25.V–03.VI.2010, M. E. Irwin (1♂, EMUS) ; Maricopa County, Gila River, 8 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.771164&amp;materialsCitation.latitude=33.223667" title="Search Plazi for locations around (long -112.771164/lat 33.223667)">Arlington</a>, malaise in opening nr. tamerisk, 200 m, 33°13.42’N 112°46.27’W, 7–13.VI.2010, M. E. Irwin (1♂, EMUS) ; California: Kern County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.9223&amp;materialsCitation.latitude=35.2872" title="Search Plazi for locations around (long -117.9223/lat 35.2872)">Munsey Road</a>, 5.8 km E Neuralia Road, 35.2872 o N 117.9223 o W, 590m, 24– 25.VI.2017, K. A. &amp; E. E. Williams (1♀, CSCA, Figs 34, 38) ; San Bernardino County: 6 mi E Phelan, Baldy Mesa, 21.IV–27.IV.1981, pan traps, J. T. Hubert (1♂, CSCA, Fig. 47) ; 10 mi. NW Barstow, 2200’, 4.VI.1971, R. R. &amp; R. A. Snelling (1♀, EMUS) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.433334&amp;materialsCitation.latitude=34.65" title="Search Plazi for locations around (long -117.433334/lat 34.65)">4 mi. NW Adelanto</a>, 2900 ft., 34°39’N 117°26’W, 22.V.1993, G. C. Snelling (71♀ 45♂, EMUS, Fig. 66) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.433334&amp;materialsCitation.latitude=34.65" title="Search Plazi for locations around (long -117.433334/lat 34.65)">4 mi. NW Adelanto</a>, 2900 ft., 34°39’N 117°26’W, 11.VI.1993, G. C. Snelling, on Chamaesyce sp. (6♀, EMUS) ; 4 mi. NW Adelanto, 880 ft., 34°65’N 117°43’W, 02.VII.1995, R. R. &amp; G. C. Snelling (11♀, EMUS) ; Los Angeles County, 3.2 km S. Pearblossom, 1067m, 15.VIII.1977, R. R. &amp; G. C. Snelling (1♀, EMUS) .</p><p>Distribution. Hot deserts in Arizona, California, and Nevada.</p><p>Remarks. This species was originally placed in the genus Smicromutilla because it lacked thickened bristles on the T2 fringe. It is closely related to L. snellingella, which may eventually be recognized as a darkened southern variety of L. beadugrimi .</p><p>This sex association is based on a series of 45 males and 88 females collected near Adelanto, California by G.C. Snelling in the early 1990’s. These wasps were apparently feeding together on nectar from flowers of ground covering spurges in the subgenus Euphorbia (Chamaesyce) S.F. Gray.</p><p>The female from Kern County was collected in a sandy wash where workers of the ant genus Forelius Emery were foraging. The mutillid superficially resembled these ants, differing by slightly larger size and a not easily describable difference in gait. It is not clear whether Lomachaeta are purposeful co-mimics with ants or this similarity is purely coincidental.</p></div>	https://treatment.plazi.org/id/03F187A07C03CD3527B79AA0FDE0FCB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.text	03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta calamondin Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta calamondin Williams, sp. nov.</p><p>(Figs 49, 50, 67, 68)</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique coloration, wherein the entire body is orange, except the darkened apical tergites. The following characters are also useful for diagnosis: the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins restricted to the basal half of wing; the T1 shape is sub-disciform; the T2 disc is has separated punctures; the T2 fringe is composed of simple setae; and the paramere is laterally compressed and moderately setose.</p><p>Description. MALE. Body length 4 mm. Coloration (Figs 36, 49). Entire body pale orange-brown, except frons, scape, tibiae, and T3 largely brown; flagellum, tarsi, and T4–7 dark brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, mesoscutum, and T3–7 with interspersed erect silvery and brownish brachyplumose setae. Head. Rounded posteriorly, with coarse punctures, sparser around ocelli. Mandible tridentate apically, unarmed ventrally. Clypeus punctate, anterior margin smoothly rounded. Genal carina weak. Ocelli small; ocellocular distance 3 × length of lateral ocellus, interocellar distance subequal to lateral ocellar diameter. F1 1.0 × pedicel length; F2 1.2 × pedicel length. Mesosoma. Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins sparsely setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron smooth. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.5 × wing. Metasoma. T1 shape sub-disciform, without carina separating anterior and dorsal faces. T2 with separated punctures, intervals smooth; T2 fringe setae simple. S2 with coarse oblique punctures. T3–6 punctures sparse. T7 smooth. Hypopygium punctate, sharp bidentate posteriorly. Genitalia (Fig. 67, 68). Paramere laterally compressed, straight, subacuminate apically, with short setae along inner and outer surfaces, setae longer and denser apicolaterally, few long setae ventrally in basal half. Cuspis with posteroventral setae subequal to cuspis length. Penis valve unidentate apically.</p><p>FEMALE. Unknown.</p><p>Material examined. Holotype, ♂, USA, California, Riverside County, Riverside, 14.VIII.1969, J.C. Hall, malaise trap (EMUS, Figs 49, 50, 67, 68).</p><p>Distribution. Known only from the type locality in southern California.</p><p>Etymology. Named after the calamondin, a small orange-like fruit frequently grown in southern California, in reference to the small orange-ish color of this southern Californian species. Treat as a noun in apposition.</p><p>Remarks. Based on the reduced wing venation, simple parameres, and distribution in California, this is likely the sister species to L. powelli . In the western USA, this is apparently the only diurnal male mutillid with the head and mesosoma pale orange-brown (Fig. 49), although many nocturnal species can be superficially similar.</p></div>	https://treatment.plazi.org/id/03F187A07C00CD3B27B79B3CFAD8FB4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.text	03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta chionothrix Pitts & Manley 2004	<div><p>Lomachaeta chionothrix Pitts &amp; Manley, 2004</p><p>(Figs 21, 27, 41)</p><p>Lomachaeta chionothrix Pitts &amp; Manley, 2004: 6 . Holotype, ♂, Guatemala, Zacapa, Rio Hondo (CNCI).</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. chionothrix: the body is entirely black; the mandible has a ventral tooth basally; the forewing has its veins encompassing the basal 0.7 of the wing; the mesoscutum is densely coarsely punctate; the T1 shape is disciform; the head and T2 disc have separated punctures; the T2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks elongate setae. Body length 3.5–5 mm.</p><p>FEMALE. As in males, this species is similar to the variable L. hicksi . Lomachaeta chionothrix has tighter areolations on the head and mesosoma, with 13 areolations between the pronotal spiracles; and has T6 basally with dense long setae that are black basally and white apically. The following characters may also be useful for diagnosis: the baso-ventral mandibular margin has a weak hyaline lamella; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the pronotal and propodeal spiracles are swollen and tuberculate; the lateral mesonotal tooth is undefined; the lateral propodeal face is impunctate; the mesosomal dorsum lacks subparallel thickened setae; the T1 shape is sub-disciform; T2 has sparse punctures, mostly erect brachyplumose setae, and mostly smooth intervals; the T2 fringe is composed of dense thick white setae; and the S6 lateral carina is shallow.</p><p>Description. FEMALE. (hitherto unrecognized, based on female from Jalisco). Body length 4.5 mm. Coloration. Body orange-brown except flagellum, mesosomal dorsum, legs, and T1 variegated with brown and metasomal segments 2–6 dark brown to black. Tibial spurs white. Erect dorsal brachyplumose setae on mesonotum, T2 disc, and T3–6 black-brown; remaining setae whitish. Fringe of T2 composed of dense subappressed thick white setae, fringes of T3–4 with sparse thick white setae. Head. Head width 1.1 × pronotal width. Frons, vertex and gena tightly areolate, all areolations complete. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with weak ventral lamella basally. Antennal scrobe without dorsal carina. Length of F1 1.4 × pedicel; F2 length subequal to pedicel length. Mesosoma. the Mesosoma is compact, thoracic dorsal length 0.85 × width. Humeral carina distinct, rounded dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, some raised to form apparent tubercles; intervals slightly tighter anteriorly than posteriorly; dorsally with 13 areolations between pronotal spiracles. Pronotal and propodeal spiracles swollen, tuberculiform; lateral mesonotal tubercle undefined. Lateral propodeal face with a few nearly-complete areolations. Metasoma. T1 shape sub-disciform, punctate. T2 as long as wide. Disc of T2 with sparse oblique punctures, intervals mostly smooth. T3–5 and S2–5 with separated punctures, intervals smooth. T6 convex. Incomplete lateral S6 carina shallow but distinct.</p><p>Material examined. COSTA RICA, Guanacaste: EJN 14 km S Cañas, 14.III.1990, F. D. Parker (1♂, CSCA, Fig. 41); P.N. Barra Honda, 100m: III.1995 (1♂, MIUP); IV.1995 (1♂, MIUP), M. Reyes . HONDURAS, Fco. Morazan, San Antonio de Oriente, El Zamorano, Malaise trap in coffee plantation, 25.II–6.III.1990, R. Cave (1♂, MIUP) . MEXICO, Jalisco, Chamela, 10.VII.1989, J. G. Rozen (1♀, MIUP, Figs 21, 27) . NICARAGUA, Granada, Volcan Mombacho, Santa Ana: 15.IV.1998 (1♂, MIUP); 15.V.1998 (1♂, MIUP), malaise trap, J. M. Maes .</p><p>Distribution. Mexico (Nayarit and Jalisco) South to Costa Rica. First records for Honduras and Nicaragua.</p><p>Remarks. This species is closely related to the widespread L. hicksi, differing in males by mesosomal punctation and genitalic differences only. Until now, this species is the only Lomachaeta known from Central America and although 10 species are known from Mexico, only L. chionothrix has been found in Jalisco.</p><p>Only two females have been observed from the northern Neotropical region: one from Costa Rica and one from Jalisco, Mexico. These females differ in significant structural features, especially the head and mesonotum punctation. The female from Jalisco was collected in Chamela, where L. chionothrix males have been recorded (Williams &amp; Pitts 2009). This female is similar to L. hicksi, differing mainly by its tighter mesosomal areolations. Lomachaeta chionothrix males also have tighter mesosomal punctation than L. hicksi . Morphological similarity to L. hicksi and co-occurrence with L. chionothrix males in Chamela supports the association of this female with L. chionothrix . The female from Costa Rica cannot be differentiated from the newly associated L. hyphantria females and is discussed below.</p><p>Lomachaeta cirrhomeris Pitts &amp; Manley, 2004</p><p>(Figs 22, 23, 28, 29, 42)</p><p>Lomachaeta cirrhomeris Pitts &amp; Manley, 2004: 7 . Holotype, ♂, USA, Arizona, Santa Cruz Co. (CNCI). Pitts &amp; Manley (2004): Hosts.</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique coloration, wherein the body is largely black, except the orange tegula and femora. The following characters are also useful for diagnosis: the head and mesosoma have coarse separated punctures; the mandible has a ventral tooth basally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is disciform; the T2 disc has separated punctures; the T2 fringe is composed of thickened bristles; and the paramere is straight and lacks elongate setae. Body length 4.5–6.5 mm.</p><p>FEMALE. This species can be recognized by the body color, which is entirely orange, including the legs and antennae, except T6 is blackish; and T3–5 are clothed mostly with black erect setae except for the dense thick silver setae of the T2 and sometimes T3 fringe. The following characters will also aid in diagnosis: the baso-ventral mandibular margin has a hyaline lamella; the head width is 1.1 × the mesosomal width; thoracic dorsal length 0.9 × its width; the pronotal spiracle is usually swollen and nearly tuberculate; the lateral mesonotal tooth is usually distinct; the propodeal spiracle is sharply swollen and tuberculate; the lateral propodeal face is impunctate; the mesosomal dorsum has about 10 areolations between the pronotal spiracles, many areolations have the intervals either obliterated or raised to tuberculate corners; the T1 shape is sub-sessile; T2 has sparse punctures and mostly smooth intervals; and the S6 lateral carina is low sub-triangular. Body length 2.5–4 mm.</p><p>Material examined. USA: Arizona: Maricopa County: Gila River, 10 km S. Arlington, malaise on sand beach, 200 m, 33°13.3’N 112°45.53’W, 25.V–03.VI.2010, M. E. Irwin (1♂, EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.771164&amp;materialsCitation.latitude=33.223667" title="Search Plazi for locations around (long -112.771164/lat 33.223667)">Gila River</a>, 8 km S. Arlington, malaise in opening nr. tamerisk, 200 m, 33°13.42’N 112°46.27’W, 7–13.VI.2010, M. E. Irwin (1♂, EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.771164&amp;materialsCitation.latitude=33.223667" title="Search Plazi for locations around (long -112.771164/lat 33.223667)">Pima County</a>, Tucson, 9.I.1939, J. Knull (1♀, EMUS, Figs 23, 29); additional data unavailable (1♂, FSCA, Fig. 42) ; California, Riverside County, San Timoteo Canyon, Malaise trap, 14.IX.1972, M. Wasbauer, A. Hardy (1♀, CSCA, Figs 22, 28) .</p><p>Distribution. Widespread in the western Nearctic, including Arizona, California, Nevada, New Mexico, Texas, and Utah (USA) and Baja California and Baja California Sur (Mexico).</p><p>Remarks. This species was placed in the L. hicksi species-group (Williams &amp; Pitts 2009) with L. hicksi and L. chionothrix . Males in this species-group can be easily recognized because of the distinct ventral mandibular tooth. Females are more difficult to define, as the main diagnostic trait of males, the ventral mandibular tooth, is reduced to a subtle lamella in females. Additional traits for L. hicksi species-group females (couplet 7 below), are more distinct in L. hicksi and L. chionothrix than in L. cirrhomeris . If one were to mis-step at couplet seven, females of L. cirrhomeris would key out to L. beadugrimi, which can be recognized by having the T2 intervals microreticulate (mostly smooth in L. cirrhomeris) and T3–5 with interspersed silver and blackish erect setae (T3–5 setae mostly black in L. cirrhomeris).</p></div>	https://treatment.plazi.org/id/03F187A07C0ECD3927B79D6DFE4BFD6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.text	03F187A07C0CCD3827B79B43FAA9FF72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta crocopinna Pitts & Manley 2004	<div><p>Lomachaeta crocopinna Pitts &amp; Manley, 2004</p><p>(Fig. 52)</p><p>Lomachaeta crocopinna Pitts &amp; Manley, 2004: 10 . Holotype, ♂, USA, Texas, Val Verde Co. (UMSP). Pitts &amp; Manley (2004): Host.</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. crocopinna: the body is entirely black, except the metasoma is almost entirely orange; the mandible lacks a ventral tooth basally; the forewing has its veins encompassing the basal 0.65 × of the wing; the head, mesoscutum, and T2 disc have separated punctures; the T1 shape is sub-sessile; the T2 fringe is composed of slightly thickened setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 3–5 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. MEXICO, Sonora: Alamos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.92284&amp;materialsCitation.latitude=27.026167" title="Search Plazi for locations around (long -106.92284/lat 27.026167)">Rancho Acosta</a>, malaise in dry wash, tropical deciduous forest, 395 m, 27°01.57’N 106°55.37’W, 26.V–6.VI.2007, M. E. Irwin (1♂, EMUS) ; 3 km NE Alamos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.9&amp;materialsCitation.latitude=27.016666" title="Search Plazi for locations around (long -108.9/lat 27.016666)">La Huerta Orchard</a>, 27°01’N 108°54’W, 19–24.[Month unknown].2008, M. E. Irwin &amp; O. Figuero (2 ♂, EMUS) ; USA, Arizona: Yavapal County, 11 mi W. Jerome, Reared, F. D. Parker (2♂, EMUS); Yavapal County, Clarksdale, Reared, F. D. Parker (1♂, EMUS) ; Coconino County, 7 mi S. Sedona, Reared, F. D. Parker (3♂, EMUS); Gila County, 2 mi S. Pine, F. D. Parker (1♂, EMUS); Pima County, 550m, Silver Reef Wash, 4 km E. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.90784&amp;materialsCitation.latitude=32.651" title="Search Plazi for locations around (long -111.90784/lat 32.651)">Vaiva Vo</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.90784&amp;materialsCitation.latitude=32.651" title="Search Plazi for locations around (long -111.90784/lat 32.651)">Cockelburr</a>), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.90784&amp;materialsCitation.latitude=32.651" title="Search Plazi for locations around (long -111.90784/lat 32.651)">Tat Momoli Mts.</a>, malaise in wash, 32°39.06’N 111°54.47’W, 07–14.V.2006, M. E. Irwin (5♂, EMUS) ; Maricopa County, Gila River, 10 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.758835&amp;materialsCitation.latitude=33.22167" title="Search Plazi for locations around (long -112.758835/lat 33.22167)">Arlington</a>, malaise on sand flat, 200m, 33°13.30’N 112°45.53’W, 07–13.VI.2010, M. E. Irwin (1♂, EMUS) ; 4 km E. Vaiva Vo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.90784&amp;materialsCitation.latitude=32.651" title="Search Plazi for locations around (long -111.90784/lat 32.651)">Silver Reef Wash</a>, 32°39.06’N 111°54.47’W, 550m, 1–7.V.2006, M. E. Irwin, malaise trap (1♂, EMUS) ; Kitt Peak rd., Coyote Mts., 5 km S. jct. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.56316&amp;materialsCitation.latitude=31.988667" title="Search Plazi for locations around (long -111.56316/lat 31.988667)">Hwy.</a> 36 and 286, malaise in wash, 1300 m, 31°59.32’N 111°33.79’W, 2–12.V.2006, M. E. Irwin (1♂, EMUS) ; New Mexico: Socorro, NWR [Natural Wildlife Resort] LTER [Long Term Ecological Reserve] site 1, 26.V.1992 (1♂, EMUS) ; Texas, Culberson County, Choza Springs, Guadalupe Mountains, 1610 m, 8.IV.1996 (1♂, EMUS, Fig. 52) ; Utah: Garfield County, Alvey Wash, 7 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.63&amp;materialsCitation.latitude=37.708332" title="Search Plazi for locations around (long -111.63/lat 37.708332)">Escalante</a>, pan trap in dry wash, 1990 m, 37°42.5’N 111°37.8’W, 24– 25.V.2002, F. D. Parker &amp; M. E. Irwin (2♂, EMUS) ; Garfield County, Alvey Wash, 7 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.63&amp;materialsCitation.latitude=37.708332" title="Search Plazi for locations around (long -111.63/lat 37.708332)">Escalante</a>, malaise in dry wash, 1990m, 37°42.5’N 111°37.8’W, 24–25.V.2002, M. E. Irwin &amp; F. D. Parker (1♂, EMUS) .</p><p>Distribution. Known from the Arizona Mountains forests, Colorado Plateau, Great Basin, and Chihuahuan Desert in Arizona, Nevada, New Mexico, and Texas (USA) and Sonora (Mexico).</p><p>Remarks. This is the apparent sister species to L. ptilohyalus; the two species share similar coloration and genitalic morphology. The distribution of L. crocopinna spans multiple ecoregions in the western USA and parallels that of Dilophotopsis concolor (Cresson) . Wilson and Pitts (2008) described the distribution of D. concolor and its sister species, D. paron (Cameron) . Dilophotopsis paron occurs in the Mojave and Sonoran deserts, as does L. ptilohyalus . The parallel distributions of these sister-species pairs should be studied more closely, as they could shed light on historic events that impacted the biogeography of numerous velvet ants.</p></div>	https://treatment.plazi.org/id/03F187A07C0CCD3827B79B43FAA9FF72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.text	03F187A07C0DCD3827B79975FDC7FA39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta eironeia Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta eironeia Williams, sp. nov.</p><p>(Figs 59, 69, 70)</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the cuspis and lateral paramere margins have numerous elongate (~5 × paramere width) setae. The following characters are also useful for diagnosis: the body is entirely blackish; the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-disciform; the T2 disc is densely coarsely punctate; the T2 fringe is composed of simple setae.</p><p>Description. MALE. Body length 4 mm. Coloration (Fig. 40). Entire body black, except mandible, tegula, and legs largely brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, and mesoscutum with interspersed erect silvery and brownish brachyplumose setae. Head. Rounded posteriorly, with coarse punctures, coarser on face. Mandible tridentate apically, unarmed ventrally. Gena not carinate. Ocelli small; ocellocular distance 6 × length of lateral ocellus, interocellar distance 2 × lateral ocellar diameter. F1 1.0 × pedicel length; F2 1.6 × pedicel length. Mesosoma. Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron glabrous. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.7 × wing. Metasoma. T1 shape sub-disciform, with defined anterior and dorsal faces. T2 with coarse dense punctures, intervals smooth; T2–4 fringe setae slightly thicker than disc setae, but not forming bristles. S2 with coarse separated punctures. T3–6 punctate. T7 smooth. Hypopygium punctate, sharply bidentate posteriorly. Genitalia (Figs 69, 70). Paramere dorsoventrally flattened, obliquely downcurving to apex, sub-truncate apically, with long (~5 × paramere width) ventral setae around inner and lateral margins, setae longer and denser apically. Cuspis with posteroventral setae ~5 × cuspis length. Penis valve unidentate apically.</p><p>FEMALE. Unknown.</p><p>Material examined. Holotype, ♂, MEXICO, Sonora, km 23, Hwy 16, 5.5 km ENE [Los] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.844&amp;materialsCitation.latitude=27.7285" title="Search Plazi for locations around (long -109.844/lat 27.7285)">Hornos</a>, 23– 26.VII.2008, 27 o 43.71’N 109 o 50.64’W, 40m, M.E. Irwin &amp; R.L. Wescott, malaise in dense riparian forest (EMUS, Figs 59, 69, 70).</p><p>Distribution. Known only from the type locality in Sonora, Mexico</p><p>Etymology. From the Greek, eironeia, irony, in reference to the fact that this specimen was recognized as a new species by KAW only a few months after Williams &amp; Pitts 2009 was published.</p><p>Remarks. The genitalia of this species are truly bizarre (Fig. 59), even for a genus with such variable parameres (Williams &amp; Pitts 2009).</p></div>	https://treatment.plazi.org/id/03F187A07C0DCD3827B79975FDC7FA39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.text	03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta hedera Williams & Pitts 2009	<div><p>Lomachaeta hedera Williams &amp; Pitts, 2009</p><p>(Figs 33, 37, 62)</p><p>Lomachaeta hedera Williams &amp; Pitts, 2009: 231 . Holotype, ♂, Mexico, Baja California Sur, Arroyo San Gregorio (CSCA).</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is virtually straight, laterally subcompressed, and long setae throughout apical half of free paramere length ventrally. The following characters are also useful for diagnosis: the body is entirely blackish; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the head and T2 disc have separated punctures; and the T2 fringe is composed of simple setae. Body length 4–5 mm.</p><p>FEMALE. This species can be recognized by the body color, which is mostly orange, except the legs, antennae and T6 are entirely blackish. Additional useful diagnostic features include: the mandible lacks a ventral lamella; the mesosoma is compact, with the thoracic dorsal length 0.8 × its width; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the T2 disc is sparsely punctate, clothed with erect blackish brachyplumose setae, and the intervals mostly micropunctate; the T2 fringe is sparse and simple; and the S6 lateral carina is distinctly triangular.</p><p>Description. FEMALE (hitherto unrecognized, based on female from Arroyo San Gregario). Body length 2.5– 3 mm. Coloration. Body reddish-orange except antenna, legs (including coxae), and T6 blackish, mid and hind legs (including coxae) sometimes partly orange. Tibial spurs white. Erect dorsal brachyplumose setae on frons, anterior vertex, mesonotum, T2–5 disc, and T6 black-brown; on posterior vertex, pronotum, propodeum, T1, and T2 base yellowish. Mesonotum with few scattered subappressed simple white setae; fringes of T2–5 composed of sparse subappressed simple whitish setae. Ventral and pleural setae white, except S6 setae black-brown. Head. Head width subequal to pronotal width. Frons, vertex and gena areolate, edges of many frons and vertex areolations obliterated, areolation edges on frons and vertex forming parallel longitudinal apparent striae. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe lacking dorsal carina. Length of F1 subequal to pedicel; F2 1.2 × pedicel length. Mesosoma. Mesosoma compact, dorsal thoracic length 0.8 × width. Humeral carina weak, not reaching epaulet. Mesosomal dorsum areolate; many intervals obliterated; areolations evenly distributed; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle small, but distinct. Lateral propodeal face impunctate. Metasoma. T1 shape sub-sessile, punctate. T2 as wide as long, laterally slightly swollen subapically. Disc of T2 with sparse incomplete apparent areolations, intervals microreticulate. T3–5 and S2–5 with separated punctures, intervals microreticulate. T6 convex. Incomplete lateral S6 carina distinct triangular.</p><p>Material examined. Paratypes, 2♂, MEXICO, Baja California Sur, Arroyo San Gregorio, 13 air km WNW La Purissima, 24–26.Apr.1983, M.S. Wasbauer coll. (CSCA, Fig. 62). Other material. MEXICO: Baja California Sur, same data as above (4♀, CSCA, Figs 33, 37) .</p><p>Distribution. Baja California Sur in Mexico.</p><p>Remarks. The females here associated with L. hedera have identical label data to two of the male paratypes (Williams &amp; Pitts 2009). Only one other Lomachaeta species has been found in Baja California Sur, L. cirrhomeris . Unlike L. cirrhomeris, L. hedera females have black legs and antennae and a completely unmodified ventral mandible margin. These females are structurally similar to L. beadugrimi and L. warneri sp. nov., especially in their microreticulate T2 intervals.</p></div>	https://treatment.plazi.org/id/03F187A07C0DCD3F27B79DAFFDF2FBDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.text	03F187A07C0ACD2227B79C93FDA4FEC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta hicksi Mickel 1936	<div><p>Lomachaeta hicksi Mickel, 1936</p><p>(Figs 24–26, 30–32, 43–46, 65)</p><p>Lomachaeta hicksi Mickel, 1936: 289 . Holotype, ♂, USA, California, Los Angeles Co. (UMSP). Pitts &amp; Manley (2004): Hosts.</p><p>Lomachaeta coloradensis Mickel, 1936: 292 . Holotype, ♂, USA, Colorado, Boulder Co. (UMSP). Pitts &amp; Manley (2004): Synonymy.</p><p>Lomachaeta punctinota Mickel, 1936: 293 . Holotype, ♂, USA, Illinois, Jackson Co. (INHS). Pitts &amp; Manley (2004): Synonymy.</p><p>Lomachaeta minutula Mickel, 1936: 294 . Holotype, ♀, USA, Tennessee, Shelby Co. (USNM). Pitts &amp; Manley: Synonymy.</p><p>Lomachaeta variegata Mickel, 1940: 128 . Holotype, ♂, USA, California, Riverdale Co. (CASC). Pitts &amp; Manley (2004): Synonymy.</p><p>Lomachaeta formosula Mickel, 1940: 130 . Holotype, ♂, Mexico, Coahuila, Puerta de la Goriona (UMSP). Pitts &amp; Manley: Synonymy.</p><p>Lomachaeta mickeli Manley &amp; Deyrup, 1987: 132 . Holotype, ♂, USA, Florida, Highlands Co. (UMSP). Pitts &amp; Manley (2004): Synonymy.</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. hicksi: the body is black, though T2–3 are often largely red-brown; the mandible has a ventral tooth basally; the mesoscutum is variably punctate, but never contiguously punctate throughout; the forewing has its veins encompassing the basal 0.8 × of the wing; the T1 shape is disciform; the head and T2 disc have separated punctures; the T2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks elongate setae. Body length 3–6 mm.</p><p>FEMALE. This is a hugely variable species in color, setation, and punctation. The following characters are consistent to all L. hicksi: the baso-ventral mandibular margin has a hyaline lamella; the pronotal spiracle is swollen, almost tuberculate; the lateral mesonotal tooth is usually distinct; the propodeal spiracle is sharply swollen and tuberculate; the lateral propodeal face is impunctate; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the mesosomal dorsum is areolate without subparallel thickened setae, many intervals are either obliterated or raised to tuberculate corners; the T1 shape is sub-disciform, often with the dorsal and anterior faces separated by a weak carina; T2 has sparse punctures, mostly erect brachyplumose setae, and mostly smooth intervals; and the S6 lateral carina is low angular. Body length 2.5–5 mm.</p><p>Material examined. USA: Arizona, Coconino County, 3 mi S Sedona, reared, F. D. Parker (1♀, EMUS, Figs 25, 31) ; California: Kern County, Short Canyon, 35.7093 o N 117.9112 o W, 1200m, 21–29.V.2008, M.E. Irwin, malaise trap (1♂, EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50667&amp;materialsCitation.latitude=32.803333" title="Search Plazi for locations around (long -116.50667/lat 32.803333)">Los Angeles County</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50667&amp;materialsCitation.latitude=32.803333" title="Search Plazi for locations around (long -116.50667/lat 32.803333)">Placentia County</a> Park, Walker Ranch, 34.38 o N 118.44 o W, 8– 17.IX.1998, B.V. Brown, malaise trap (1♂, EMUS, Fig. 45); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50667&amp;materialsCitation.latitude=32.803333" title="Search Plazi for locations around (long -116.50667/lat 32.803333)">Riverside County</a>: Palms Springs, Araby Wash, 33.750 o N 116.5319 o W, 538m, F.D. Parker and T.D. McIntyre, malaise trap: 14–21.IV.2015 (1♂, EMUS, Fig. 44); 21–28.IV.2015 (1♂, EMUS); 21–24.IV.2016 (1♂, CSCA); 17–21.V.2016 (1♂, CSCA); Riverside, 30.VI.1969, J.C. Hall, malaise trap (1♂, EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50667&amp;materialsCitation.latitude=32.803333" title="Search Plazi for locations around (long -116.50667/lat 32.803333)">San Diego County</a>, 2 km SE Pine Valley near Laguna Summit, 32 ° 48.2’N 116°30.4’W, 1280m, 31.V–5.VI.2002, M.E. Irwin and F.D. Parker, malaise trap in sandy ravine (2♂, CSCA EMUS) ; Florida, Alachua County, additional data unavailable (1♂ 1♀, FSCA, Figs 24, 30, 43), Utah, Cache County, Logan Canyon #3, reared, F. D. Parker (1♀, EMUS, Figs 26, 32) .</p><p>Distribution. This is the most widespread Nearctic mutillid species, ranging from the Atlantic to Pacific Oceans, North to Massachussetts and South to Florida (USA) and Puebla (Mexico).</p><p>Remarks. As one of the World’s most widely distributed mutillid species, L. hicksi has the longest synonym list in this genus, belying its tremendous morphological variation. Although fewer females than males have been available to study, they also reveal high variation in color and punctation (Figs 24–26). The mesosoma and metasoma each vary from nearly entirely pale orange to nearly black. The mesonotal areolations vary from broadly to tightly areolate, though when the mesonotal areolations are tightly spaced, there are fewer than 12 areolations between the pronotal spiracles and the propodeal areolations are much broader.</p><p>The males mentioned above in the Material Examined section are the first known to have orange femora, a trait that was apparently unique to males of L. cirrhomeris . Males of L. cirrhomeris, however, always have cuticle of the metasoma entirely black and contrasting with their orange femora (Fig. 42), while these L. hicksi from California have the metasoma predominantly orange and concolorous with the femora (Fig. 44, 45). Pitts &amp; Manley (2004) suggested that L. cirrhomeris may eventually prove to be a synonymous color variant of L. hicksi . These males lend additional credence to that hypothesis.</p></div>	https://treatment.plazi.org/id/03F187A07C0ACD2227B79C93FDA4FEC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.text	03F187A07C17CD2127B799EBFC92FAF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta hyphantria Pitts & Manley 2004	<div><p>Lomachaeta hyphantria Pitts &amp; Manley, 2004</p><p>(Figs 9, 10, 15, 16, 53, 54)</p><p>Lomachaeta hyphantria Pitts &amp; Manley, 2004: 11 . Holotype, #m, Bolivia, Dep. Beni, Rio Itenez (AMNH). Lomachaeta garm Williams &amp; Pitts, 2007: 299 . Holotype, #m, Colombia, Bolivar, PNN Gorgona La Suiris (IAvH). Williams &amp; Pitts (2009): Synonymy.</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. hyphantria: the head and mesosoma are variegated orange and dark brown; the metasoma is black, except the pale yellow T1 apex; the mandible lacks a ventral tooth basally; the head and pronotum are areolate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is disciform; the T2 disc has small punctures; the T2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks an apical setal tuft. Body length 3–6 mm.</p><p>FEMALE. This species can be recognized by the combination of a broadly areolate mesosomal dorsum and well-defined areolate lateral propodeal face. The areolations of the head are broad and have many transverse intervals obliterated, leaving a few apparent subparallel rugae. Additional useful diagnostic features include: the mandible lacks a ventral lamella; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the mesosoma lacks appressed subparallel pale setae; the T2 disc is moderately punctate, clothed with erect blackish brachyplumose setae, and the intervals mostly smooth; the T2 fringe is sparse and simple; and the S6 lateral carina is weak.</p><p>Description. FEMALE (hitherto unrecognized, based on female from Venezuela). Body length 2.5–3.5 mm. Coloration. Head, mesosoma, legs, and T1 variegated orange brown and dark brown; remainder of metasoma blackish. Tibial spurs white. Erect dorsal brachyplumose setae yellowish to black-brown, blackish setae often whitish at bases. Head, mesonotum, and T3–5 with scattered subappressed simple usually white setae; fringes of T2–5 composed of sparse subappressed simple white setae. Ventral and pleural setae predominantly whitish. Head. Head width 1.1 × pronotal width. Frons, vertex and gena broadly areolate; frons and vertex many transverse intervals obliterated, leaving subparallel subrugae. Clypeus without distinct teeth or ridges. Genal carina distinct to hypostomal carina, angulate at junction. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe with dorsal carina. Length of F1 1.2 × pedicel; F2 length subequal to pedicel length. Mesosoma. Dorsal thoracic length 0.85 × width. Humeral carina distinct to epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 8 areolations between pronotal spiracles. Pronotal spiracles low rounded; propodeal spiracle tuberculiform; lateral mesonotal tubercle weakly defined. Lateral propodeal face areolate, with multiple complete areolations. Metasoma. T1 shape sub-disciform, punctate. T2 slightly longer than wide. Disc of T2 with dense oblique punctures, intervals smooth. T3–5 and S2–5 with separated punctures, intervals smooth. T6 convex. Incomplete lateral S6 carina weak.</p><p>Material examined. BOLIVIA, Beni, Rio Itenez about 4 km above Costa Marques ( Brazil), 12–18.IX.1964, Bouseman and Lussenhop (1♂, MIUP) ; BRAZIL: Roraima, Caracaraí, Parque Nacional de Viruá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.966667&amp;materialsCitation.latitude=1.4666667" title="Search Plazi for locations around (long -60.966667/lat 1.4666667)">Vicinal</a> na estrada perdida, 01°28’N 60°58’W, 24.XII.2015 – 07.I.2016, J.A. Rafael e equipe, malaise (1♂, INPA) ; Alto Alegre, Estação Ecológica de Maracá, 03°21’59’’N 61°26’04’’W, 01–15.IV.2016, R. Boldrini &amp; J.A. Rafael, malaise (1♂, 1♀, INPA); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.115276&amp;materialsCitation.latitude=-2.5891666" title="Search Plazi for locations around (long -60.115276/lat -2.5891666)">Amazonas</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.115276&amp;materialsCitation.latitude=-2.5891666" title="Search Plazi for locations around (long -60.115276/lat -2.5891666)">Manaus</a>, ZF2, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.115276&amp;materialsCitation.latitude=-2.5891666" title="Search Plazi for locations around (long -60.115276/lat -2.5891666)">Km</a> 14, 02°35’21’’S 60°06’55’’W, 3–17.VIII.2016, J.A. Rafael &amp; F.F. Xavier-Filho, malaise em igarapé perto da torre (1♀, INPA, Figs 9, 15) ; 15–31.XII.2016, J.A. Rafael &amp; F.F. Xavier- Filho, malaise a 18m na torre (1♂, INPA, Fig. 53) ; Reserva Ducke, 30.IX–9.X.2005, A. Aguiar, Malaise (1♂, INPA) ; 2– 4.X. 20005, A. Aguiar, yellow-pan trap (1♀, INPA) ; Baixio Leste / Oeste, 14.II–6.III.2007, Freitas, G. &amp; Feitosa, M, Malaise suspense no dossel (1♂, INPA) ; EMBRAPA, Guaraná convencional borda, 02°53’42.18’’S 59°59’10.58’’W, 23.XI.2012, K. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.98627&amp;materialsCitation.latitude=-2.8950498" title="Search Plazi for locations around (long -59.98627/lat -2.8950498)">Schoeninger</a>, malaise (1♀, INPA) ; Tefé, 03°19’45’’S 64°41’13’’W, 20.IX– 5.X.2016, J.A. Oliveira, D.M.M. Mendes &amp; J.A. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.68694&amp;materialsCitation.latitude=-3.3291667" title="Search Plazi for locations around (long -64.68694/lat -3.3291667)">Rafael</a>, malaise (1♂) ; 1–5.XI.2016, J.A. Oliveira, D.M.M. Mendes &amp; J.A. Rafael, malaise (1♂, INPA) ; 10–23.X.2016, J.A. Oliveira, D.M.M. Mendes &amp; J.A. Rafael, malaise (1♂) ; Rondônia, Itapuã do Oeste, FLONA do Jamari, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.082027&amp;materialsCitation.latitude=-9.194278" title="Search Plazi for locations around (long -63.082027/lat -9.194278)">Trilha Pedra Grande</a>, 09°11’39.4’’S 63°04’55.3’’W, 6.X.2014, J.A. Rafael, F.F. Xavier-Filho, R.M. Vieira &amp; R.H. Aquino, malaise (2♀, INPA) ; Rio Guapore, opp. Mouth Rio Baures ( Bolivia), 26.IX.1964, Bouseman and Lussenhop (1♂, MIUP) . COLOMBIA: Magdalena, Parque Nacional Natural Tayrona, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03333&amp;materialsCitation.latitude=11.333333" title="Search Plazi for locations around (long -74.03333/lat 11.333333)">Zaino</a>, 11°20’N 74°2’W, 50m, 29.V–14.VI. 200, Malaise, R. Henriquez, M. 241 (1♂, IAvH) ; Meta: Parque Nacional Natural Sierra de la Macarena, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Caño Curia</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Sendero Cachicamos</a>, 3°21’N 73°56’W, 460m, 10.XI–21.XII.2002, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Malaise</a>, M. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Duarte</a>, M.2985 (1♂, IavH); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Parque Nacional Natural Tinigua</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.8&amp;materialsCitation.latitude=2.1833334" title="Search Plazi for locations around (long -73.8/lat 2.1833334)">Caño Nevera</a>, 2°11’N 73°48’W, 390m, 23.XII.2002 – 7.I.2003, Malaise, C. Sanchez, M.3476 (2♂, IAvH, Fig. 54) . COSTA RICA, Puntarenas, Est. Queb. Bonita, Res. Biol. Carara, 50m, L-N-194500, 469850, IV.1993, J. C. Saborío (1♀, INBio) . PERU, Madre de Dios, Reserva El Manu, Estac. Biol. Pakitza, 1–6.III.1992, Malaise trap, R. Cambra (1♂, MIUP) . TRINIDAD, W. I., Arima Valley, 800–1200 ft, 10–22.II.1964, Rozen and Wygodzinsky (1♂, MIUP) . VENEZUELA: Aragua, Rancho Grande, 1100m, IV.1987, Bordón (1♀, MIUP, Figs 10, 16) .</p><p>Distribution. This species is widespread in northern South America, including Bolivia, Brazil, Colombia, Ecuador, Peru, Trinidad, and Venezuela. First records for Peru and Trinidad.</p><p>Remarks. Females herein associated with L. hyphantria have been repeatedly and consistently collected in the same traps as males of L. hyphantria in different areas of the Brazilian Amazon (Roraima and Amazonas states). These females and the males of L. hyphantria are the only species of this genus widely distributed throughout the Brazilian Amazon region to be found so far. Additional females were also found in regions where L. hyphantria is the only recognized male species, such as Venezuela.</p><p>Both males and females of this species show conspicuous variations related to where they occur. This is especially evident when comparing females collected in the Brazilian Amazon with ones from Venezuela wherein females from the first are much darker and have the variegated pattern overall less evident in comparison with Venezuelan specimens. Males from the Brazilian Amazon are also overall darker than males from further north in South America, in accordance to what was mentioned by Williams &amp; Pitts (2009), even specimens from areas near the Brazilian-Venezuelan border (Roraima state). The main differences that can be added to the observations already made by Williams &amp; Pitts (2009) is that specimens from Rondônia, Amazonas and Roraima states in Brazil have the legs and antennae almost completely black, the thick bristles of T2 are black, mandibles are darker and the integument of the head is almost completely reddish-brown, except around the ocelli where it is black.</p><p>The second northern Neotropical female specimen, mentioned above in the Remarks for L. chionothrix, cannot be readily separated from South American L. hyphantria specimens. Given that no males of L. hyphantria have been found in Central America, and that females are more difficult to differentiate than males, the distribution of L. hyphantria in Central America should be considered as tentative.</p></div>	https://treatment.plazi.org/id/03F187A07C17CD2127B799EBFC92FAF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.text	03F187A07C14CD2727B79DFBFCEAFA69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta ibarrai Casal 1969	<div><p>Lomachaeta ibarrai Casal, 1969</p><p>(Figs 11, 17)</p><p>Lomachaeta ibarrai Casal, 1969: 33 . Holotype, ♀, Argentina, Buenos Aires, Burzaco (AMNH).</p><p>Diagnosis. FEMALE. The following characters are diagnostic: the head and mesosoma are reddish-brown, the metasoma is blackish; the pronotal spiracle is scarcely swollen; the propodeal spiracle is sharply dentiform; the head is 1.1 × wider than the mesosoma; the lateral propodeal face has a few complete areolations; the mesosoma is elongate, with the thoracic dorsal length slightly less than its width; the mesosomal dorsum is tightly areolate without subparallel thickened setae; the T1 shape is sub-disciform, often with dorsal and anterior faces separated by a weak carina; T2 has distinct separated punctures, numerous subappressed black setae, and mostly smooth intervals; and the S6 lateral carina is low angular. Body length 5 mm.</p><p>MALE. Unknown.</p><p>Material examined. Type material. Holotype of Lomachaeta ibarrai, ♀, ARGENTINA, Buenos Aires, Burzaco, XII.1947, F.C. Sud (AMNH, Figs 11, 17). Other material. ARGENTINA: Buenos Aires, Glew, I.1975, Carpintero (1 ♀, MIUP).</p><p>Distribution. Known only from Buenos Aires Province, Argentina.</p><p>Remarks. Based on the tight mesosomal sculpture, this species forms an apparent group with L. vianai and L. meloi, sp. nov., which are the only females known from southern South America. To this point, only one of those three species, L. vianai, has been associated with a male. The apparently disparate ranges of these three females, coupled with the morphological differences listed in the key below, make treating them as separate species the most reasonable hypothesis for now. Association of males with L. ibarrai and L. meloi or molecular sequence data might disprove or support the current taxonomic arrangement.</p></div>	https://treatment.plazi.org/id/03F187A07C14CD2727B79DFBFCEAFA69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.text	03F187A07C12CD2627B79E7EFDBAFEBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta ilex Williams & Pitts 2009	<div><p>Lomachaeta ilex Williams &amp; Pitts, 2009</p><p>(Fig. 61)</p><p>Lomachaeta ilex Williams &amp; Pitts, 2009: 233 . Holotype, ♂, USA, Nevada, South of Kaolin Wash (EMUS).</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. ilex: the body is largely darkbrown to black, the tegulae are often orange; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is subsessile; the T2 fringe is composed of simple setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 4–5 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. USA, Nevada, Clark County, 4.5 mi SW of Boulder, 17.IX.1997, Andrus, Griswold, and Messinger (1♂, paratype, CSCA, Fig. 61) .</p><p>Distribution. Mojave and western Sonoran Deserts in California and Nevada.</p><p>Remarks. This could be the sister species to L. hedera based on structural similarities in the mandible, T2 fringe, and genitalia. It could be the opposite sex of L. warneri, given that they have overlapping distribution and are both apparently related to L. hedera .</p></div>	https://treatment.plazi.org/id/03F187A07C12CD2627B79E7EFDBAFEBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.text	03F187A07C13CD2627B79933FD87F979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta juanita Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta juanita Williams, sp. nov.</p><p>(Figs 56, 71, 72)</p><p>Diagnosis. MALE. This species can be recognized by the following combination of characters: the body is black, except the obscure pale yellow T1 apex; the mandible lacks a ventral tooth basally; the gena is weakly carinate; the head and pronotum are areolate; the forewing has veins encompassing the basal 0.7 × wing; the T1 shape is disciform; the T2 disc has dense small punctures; the T2 fringe is composed of thick bristles; and the paramere is virtually straight except its downcurved apical third with a weak apical setal tuft and a few long setae along inner face medially. Body length 4.2 mm.</p><p>Description. MALE. Body length 4.2 mm. Coloration. Entire body black, except mandible, tegula, and trochanter partly orange-brown and T1 apex obscurely pale yellow. Tibial spurs white. Wings pale brown, lighter basally, veins brown. Body setae whitish, except tergal fringes with pale yellow or blackish bristles and apical tergites with more or less extensive blackish setae. Head. Rounded posteriorly. Frons areolate-rugose, vertex punctate, gena areolate. Mandible tridentate apically, unarmed ventrally. Gena with weak ventral carina. Ocelli small; ocellocular distance 5 × length of lateral ocellus, interocellar distance 1.8 × lateral ocellar diameter. F1 1.1 × pedicel length; F2 1.7 × pedicel length. Mesosoma. Pronotum and mesoscutum areolate. Pronotum anterior and dorsal faces separated by complete carina. Tegula smooth with few punctures, margin setigerously punctate. Mesopleuron coarse areolate. Metapleuron smooth. Scutellum slightly convex, areolate. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face smooth anteriorly with areolations posteriorly. Forewing with veins encompassing basal 0.65 × wing; marginal cell as measured on costa 1.8 × stigma. Metasoma. T1 shape nodose, with obscurely defined anterior and dorsal faces, anterior face convex. T2 with coarse separated punctures, intervals smooth; T2–4 fringes composed of thick pale yellow convergent bristles. S2 punctures coarser and sparser than T2 punctures. T3–6 punctate. T7 rugo-punctate. Hypopygium punctate, sharply bidentate posteriorly. Genitalia (Figs 71, 72). Paramere laterally subcompressed, straight in basal two thirds, apical third downcurved, acuminate apically, with three long (~2 × paramere width) setae along inner margin medially and small clump of setae at inner apex. Cuspis with posteroventral setae sparse and ~1.5 × cuspis length. Penis valve unidentate apically.</p><p>FEMALE. Unknown.</p><p>Material examined. Holotype, ♂, COLOMBIA, Magdalena, Parque Nacional Natural Tayrona, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03333&amp;materialsCitation.latitude=11.333333" title="Search Plazi for locations around (long -74.03333/lat 11.333333)">Neguanje</a>, 11°20’N 74°2’W, 10m, 9–17.IX.2001, Malaise, R. Henriquez, M. 2137 (IAvH, Figs 56, 71, 72).</p><p>Distribution. Known only from the type locality in Magdalena, Colombia.</p><p>Etymology. KAW is happy to name this species in honor of his friend and colleague Juanita Rodriquez- Arrieta, in honor of her contributions to wasp systematics and in gratitude for her help with finding and borrowing the type specimen from the IAvH in 2011. Treat as a noun in apposition.</p><p>Remarks. This is the third South American species based on males to be discovered. It shares its range with L. hyphantria, which has even been found in the same National Park (PNN Tayrona) but can be immediately differentiated from that species by the weak genal carina (strong in L. hyphantria) and the short marginal cell (~3 × stigma in L. hyphantria). It therefore seems closely related to L. vianai, but the nodose T1 shape and downcurved paramere set it apart as a distinct species.</p></div>	https://treatment.plazi.org/id/03F187A07C13CD2627B79933FD87F979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.text	03F187A07C13CD2527B79F6EFCB1FD23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta litosisyra Williams & Pitts 2009	<div><p>Lomachaeta litosisyra Williams &amp; Pitts, 2009</p><p>(Fig. 63)</p><p>Lomachaeta litosisyra Williams &amp; Pitts, 2009: 234 . Holotype, ♂, USA, Arizona, Santa Cruz Co. (EMUS).</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is elongate, gradually downcurving, cylindrical, and having an apical tuft of long setae. The following characters are also useful for diagnosis: the body is entirely blackish, except the sometimes reddish tegulae; the head has separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the T2 disc has coarse punctures; and the T2 fringe is composed of simple setae. Body length 4–6 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. USA: Arizona: Pima County, Vail Mountain Creek Ranch, 18–25.IV.2006, 1100 m, malaise trap, M. E. Irwin (1♂, paratype, CSCA, Fig. 63) .</p><p>Distribution. Sonoran Desert in southern Arizona, USA and northern Sonora, Mexico.</p><p>Remarks. The elongate, down-curving, cylindrical paramere shape is unique in Lomachaeta . This morphology is superficially similar to that of L. vacamuerta, except that species has shorter, straighter parameres. The length and curve, however, are similar to that of L. beadugrimi and L. snellingella, except those species have dorsoventrally flattened parameres without an apical setal tuft.</p></div>	https://treatment.plazi.org/id/03F187A07C13CD2527B79F6EFCB1FD23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.text	03F187A07C10CD2527B79A8BFE56F979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta meloi Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta meloi Williams, sp. nov.</p><p>(Figs 12, 18)</p><p>Diagnosis. FEMALE. This species can be recognized by the coloration and setation: the entire body is mostly blackish except the head is orange-red; the body is clothed with distinct subappressed white setae. Additional useful diagnostic features include: the head is broader than the mesosoma; the mesosoma is elongate, with the thoracic dorsal length subequal to its width; the pronotal and propodeal spiracles are scarcely swollen; the T1 shape is sub-disciform; and T2 is densely punctate.</p><p>Description. FEMALE. Body length 2.8 mm. Coloration. Body blackish except head and legs orange-red. Tibial spurs white. Body clothed predominantly with distinct subappressed white setae. Clypeus and lateral margins of metasoma with many erect setae black-brown. Fringes of T2–5 composed of appressed white setae thickened, contrasting with integument. Head. Head width 1.4 × pronotal width. Frons, vertex and gena areolate, vertex with some intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Length of F1 0.9 × pedicel length; F2 0.7 × pedicel length. Mesosoma. Mesosoma elongate; dorsal thoracic length subequal to width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles scarcely swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T1 shape sub-disciform, punctate. T2 slightly longer than wide. Disc of T2 densely punctate, intervals obscurely microreticulate. T3–5 and S2–5 with separated punctures, intervals microreticulate. T6 convex. Incomplete lateral S6 carina shallow but distinct.</p><p>MALE. Unknown.</p><p>Material examined. Holotype, ♀, BRAZIL, São Paulo, Ribeirão Preto, Campus da USP, 13.X.1999, G.A.R. Melo (DZUP, Figs 12, 18).</p><p>Distribution. São Paulo State in Brazil.</p><p>Etymology. Named in honor of Gabriel A. R. Melo, who collected the type specimen and has contributed to many other projects on Mutillidae and other Hymenoptera .</p><p>Remarks. The short bristle-like setae covering this insect are unique in Lomachaeta . No males have yet been found in Southern Brazil.</p></div>	https://treatment.plazi.org/id/03F187A07C10CD2527B79A8BFE56F979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.text	03F187A07C10CD2427B79F6EFB66FB4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta osita Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta osita Williams, sp. nov.</p><p>(Figs 4, 8)</p><p>Diagnosis. FEMALE. This species can be recognized by the following combination of characters: the head and mesosoma are orange-brown and the legs and metasoma are lighter yellow-brown with T2 having a brown transverse subapical cuticular band; the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the erect dorsal brachyplumose setae of the whole body are interspersed dark brown and yellow-brown; the mesosomal dorsum is elongate, coarsely areolate, and lacks appressed subparallel pale setae; the T2 disc is sparsely punctate with the intervals obscurely microreticulate; the T2 fringe is composed of simple silvery setae; and the S6 lateral carina is shallow, but distinct.</p><p>Description. FEMALE. Body length 3.1 mm. Coloration. Head and mesosoma orange-brown, legs and metasoma lighter yellow-brown; except F2–10 brown, femoral and tibial apices obscurely brown; T2 with transverse subapical brown band; and T6 dark brown apically. Tibial spurs white. Erect dorsal brachyplumose setae interspersed pale yellow and darker brown; except T6 with setae predominantly brown. Mesonotum with few scattered subappressed simple white setae; fringes of T2–5 composed of subappressed pale golden or silvery setae, those of T2 silvery, not thicker than erect T2 disc setae. Ventral and pleural setae whitish, except S6 setae largely brown. Head. Head width 1.2 × pronotal width. Frons areolate; vertex areolate anteriorly, smooth posteriorly; gena areolate dorsally, smooth ventrally. Clypeus without distinct teeth or ridges. Genal carina continuous to mandible base, subinterrupted medially by areolation margins. Mandible oblique, tapering, bidentate apically, with barely perceptible ventral lamella that does not interrupt ventral mandibular contour. Antennal scrobe without dorsal carina. Lengths of F1 and F2 each subequal to pedicel length. Mesosoma. Mesosoma elongate; thoracic dorsum length 0.92 × width. Humeral carina distinct, angulate dorsally, continuous to epaulet. Mesosomal dorsum areolate; intervals forming uniform carinae, appearing darker than surrounding cuticle; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles scarcely swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T1 shape sub-disciform, mostly smooth. T2 as wide as long. Disc of T2 with separated punctures, intervals microreticulate. T3–5 and S2–5 with separated weak punctures, intervals microreticulate. T6 convex. Incomplete lateral S6 carina shallow but distinct.</p><p>MALE. Unknown.</p><p>Material examined. Holotype, ♀, USA, Arizona, Santa Cruz County, 2 mi. S Sonoita, malaise, 31 o 38’N 111 o 39’W, 5100’ [~ 1650 m], 22–30.VIII.2010, E. E. Grissell (EMUS, Figs 4, 8).</p><p>Distribution. Madrean Archipelago in Arizona.</p><p>Etymology. From the Spanish osita “little bear”, based on colloquial reference to these small brown wasps as teddy bears by some authors.</p><p>Remarks. Based on mesosomal shape and overall habitus, this species is similar to the newly associated female of L. vacamuerta, although it lacks the appressed subparallel setae on T2. This similarity, coupled with distribution in southeastern Arizona suggests that this could be the female of L. litosisyra .</p></div>	https://treatment.plazi.org/id/03F187A07C10CD2427B79F6EFB66FB4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.text	03F187A07C11CD2427B79D1DFE94F8C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta polemomechana Williams & Pitts 2009	<div><p>Lomachaeta polemomechana Williams &amp; Pitts, 2009</p><p>(Fig. 60)</p><p>Lomachaeta polemomechana Williams &amp; Pitts, 2009: 236 . Holotype, ♂, Mexico, Sonora, 30km Agua Prieta (EMUS).</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. polemomechana: the body is almost entirely black; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the T2 fringe is composed of simple setae; and the paramere is virtually straight, subcylindrical, and having evenly distributed short setae. Body length 4–6 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. USA: Arizona: Santa Cruz County, Ruby Mountain, 20 km SSE Arivaca, 3–7.V.2004, M. E. Irwin and F. D. Parker (1♂, EMUS, Fig. 60) .</p><p>Distribution. Southern Arizona, USA and northern Sonora, Mexico.</p><p>Remarks. This species has only been found in transition zones between the Madrean Sky Islands and Chihuahuan Desert.</p></div>	https://treatment.plazi.org/id/03F187A07C11CD2427B79D1DFE94F8C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.text	03F187A07C1CCD2827B798CBFDCBFD9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta powelli (Mickel 1964)	<div><p>Lomachaeta powelli (Mickel, 1964)</p><p>(Figs 35, 39, 48)</p><p>Smicromutilla powelli Mickel, 1964: 108 . Holotype, ♂, USA, California, San Luis Obispo Co. (CISC). Williams &amp; Pitts (2009): Combination under Lomachaeta .</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for males of L. powelli: the body is entirely black, except T2–3 are largely orange; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the forewing venation is restricted to the basal half of the wing; the T1 shape is subsessile; the T2 fringe is composed of simple setae; and the paramere is virtually straight, subcylindrical, and lacking tufts or rows of long setae. Body length: 3–6 mm.</p><p>FEMALE. This species can be recognized by the numerous elongate blackish dorsal bristles throughout the body. The following characters will also aid in diagnosis: the baso-ventral mandibular margin lacks a hyaline lamella; the head width is 1.2 × the mesosomal width; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the pronotal spiracle is weakly swollen; the lateral mesonotal tooth is indistinct; the propodeal spiracle is weakly swollen; the lateral propodeal face is impunctate; the mesosomal dorsum has about 10 areolations between the pronotal spiracles, many intervals are partially obliterated; the T1 shape is sub-sessile; T2 has sparse punctures and smooth intervals; and the S6 lateral carina is low rounded. Body length 2–3 mm.</p><p>Material examined. USA: California: Fresno Co., Panoche Road, S 31 T15 R12 MO, 14.VI.1982, R. F. Gill &amp; N. J. Smith (2♀, CSCA); Monterey County, Arroyo Seco Camp, 15.VI.1957, R.C. Bechtel (1♀, EMUS, Figs 35, 39); San Diego County, San Diego, Harkins Collection (1♀, EMUS); San Luis Obispo County, Pozo, 30.IV.1962, J. Powell (1♂, paratype, CISC, Fig. 48) .</p><p>Distribution. Known only from the Central Valley and Coast Range areas of California, USA.</p><p>Remarks. Both sexes of this species were collected in nesting aggregations of Diodontus occidentalis Fox, a ground-nesting pemphredonine wasp (Mickel 1964). Where known, the other North American species parasitize twig nesters (Pitts &amp; Manley 2004).</p></div>	https://treatment.plazi.org/id/03F187A07C1CCD2827B798CBFDCBFD9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.text	03F187A07C1DCD2827B79AD3FE6FFACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta ptilohyalus Pitts & Manley 2004	<div><p>Lomachaeta ptilohyalus Pitts &amp; Manley, 2004</p><p>(Fig. 51)</p><p>Lomachaeta ptilohyalus Pitts &amp; Manley, 2004: 12 . Holotype, ♂, Mexico, Oaxaca, 10 m North of Huajuapan de Leon (CNCI). Pitts &amp; Manley (2004): Host.</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. ptilohyalus: the body is entirely black, except T2–3 are largely orange; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the mesoscutum has sparse punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the T2 fringe is composed of simple setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 4–6 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. USA: Arizona: Maricopa County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.758835&amp;materialsCitation.latitude=33.22167" title="Search Plazi for locations around (long -112.758835/lat 33.22167)">Gila River</a>, 10 km S. Arlington, malaise on sand beach, 200 m, 33°13.3’N 112°45.53’W, 25.V–03.VI.2010, M. E. Irwin (1♂, CSCA, Fig. 51) .</p><p>Distribution. This species has an apparently disjunct distribution in the Mojave and Sonoran Deserts (Arizona and California, (USA) and Balsas Dry Forests (Oaxaca, Mexico).</p><p>Remarks. The holotype from Oaxaca, Mexico is separated from the other known specimens of L. ptilohyalus in hot deserts of the USA by over 2300 km. Further collections in Mexico may reveal other populations, or molecular data comparisons between Oaxacan and hot desert specimens may reveal that they are not conspecific. Either way, this is a compelling link between disjunct arid habitats of southern Pacific Mexico and the North American hot deserts.</p></div>	https://treatment.plazi.org/id/03F187A07C1DCD2827B79AD3FE6FFACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.text	03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta snellingella Williams & Pitts 2009	<div><p>Lomachaeta snellingella Williams &amp; Pitts, 2009</p><p>(Fig. 58)</p><p>Lomachaeta snellingella Williams &amp; Pitts, 2009: 237 . Holotype, ♂, USA, California, San Diego Co. (UCDC).</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. snellingella: the body is entirely blackish; the mandible lacks a ventral tooth basally; the head and T2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-sessile; the T2 fringe is composed of simple setae; and the paramere is dorsoventrally flattened, apically rounded, and having short setae only. Body length 3–5 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. USA, California, San Diego County, Borrego, 30.IV.1957, F.X. Williams (1♂, paratype, CASC, Fig. 58) .</p><p>Distribution. Western Sonoran Desert in California, USA.</p><p>Remarks. Lomachaeta snellingella is clearly a close relative of L. beadugrimi, given the nearly identical paramere shape. If females are found that key out to L. beadugrimi but occur in Imperial or San Diego Counties, those would be good candidates for the female of L. snellingella . If no recognizable differences were found between these females, that might provide evidence for synonymizing these taxa.</p></div>	https://treatment.plazi.org/id/03F187A07C1DCD2F27B79DE3FBC4FF72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.text	03F187A07C1ACD2E27B7997BFDC6FE76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta theresa Williams & Pitts 2009	<div><p>Lomachaeta theresa Williams &amp; Pitts, 2009</p><p>(Fig. 57)</p><p>Lomachaeta theresa Williams &amp; Pitts, 2009: 238 . Holotype, ♂, Mexico, Sonora, 42 km ENE Alamos (EMUS).</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. theresa: the body is entirely blackish; the mandible lacks a ventral tooth basally; the gena is ecarinate; the head and T2 disc have separated punctures; the forewing has its veins encompassing the basal 0.65 × of the wing; the T1 shape is sub-sessile; the T2 fringe is composed of thickened bristles; and the paramere is subcylindrical, virtually straight, and lacks long tufts of setae. Body length 5–6 mm.</p><p>FEMALE. Unknown.</p><p>Material examined. MEXICO: Sonora: 43 km E Alamos, Rancho San Pablo, 1–5.VI.2007, M.E. Irwin (1♂, EMUS, Fig. 57) .</p><p>Distribution. Known only from Sonora, Mexico.</p><p>Remarks. Among males of Lomachaeta, some of the most easily interpreted features for sorting the taxa are: 1) the presence or absence or a ventral mandibular tooth, 2) the presence or absence of thick bristles on the T2 fringe, and 3) distribution in either North or South America. Lomachaeta theresa defies the functionality of some of these traits for establishing species-groups. Lomachaeta chionothrix, L. cirrhomeris, and L. hicksi have T2 bristles like L. theresa, but have a ventral mandibular tooth, which L. theresa lacks. The remaining North American species lack a ventral mandibular tooth, like L. theresa, but also lack the T2 bristles. The South American species, L. hyphantria and L. vianai, have the combination of unarmed mandible and T2 bristles, but they differ from L. theresa in having the gena carinate.</p></div>	https://treatment.plazi.org/id/03F187A07C1ACD2E27B7997BFDC6FE76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.text	03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta vacamuerta Williams & Pitts 2009	<div><p>Lomachaeta vacamuerta Williams &amp; Pitts, 2009</p><p>(Figs 2, 3, 6, 7, 64)</p><p>Lomachaeta vacamuerta Williams &amp; Pitts, 2009: 239 . Holotype, ♂, USA, New Mexico, Chaves Co. (EMUS).</p><p>Lomachaeta vacamuerte Williams &amp; Pitts, 2009: 242 . lapsus calami.</p><p>Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is straight, subcylindrical, and having an apical tuft of long setae. The following characters are also useful for diagnosis: the body is entirely blackish, except the sometimes reddish tegulae; the mandible is unarmed ventrally; the gena is ecarinate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is subsessile; the head and T2 disc have separated punctures; and the T2 fringe is composed of simple setae. Body length 5–6 mm.</p><p>FEMALE. This species can be recognized by the setae and mesosoma shape: the mesosomal dorsum and T2 disc have matching thickened posteriorly-directed subparallel appressed pale golden setae and the mesosoma is elongate, with the thoracic dorsum nearly as long as wide. Additional useful diagnostic features include: the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the head is clearly broader than the mesosoma; the T2 fringe is sparse and simple; and the S6 lateral carina is raised, rounded.</p><p>Description. FEMALE (hitherto unrecognized, based on female from Kennedy County). Body length 3 mm. Coloration. Head, mesosoma, antenna, legs, and T6 predominantly dark brown; metasoma and bases of leg segments and antenna orange. Tibial spurs white. Body setae whitish yellow, except frons, mesonotum, and T6 with many erect setae darkened, at least apically. Mesonotum and T2 disc with long thick posteriorly-directed subparallel setae; fringes of T2–3 composed of dense thick subappressed simple white setae; fringes of T4–5 setae sparse, simple. Head. Head width 1.2 × pronotal width. Frons, vertex and gena areolate, vertex with some intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with shallow lamella baso-ventrally that does not interrupt the ventral mandible contour. Antennal scrobe without dorsal carina. Lengths of F1 and F2 each subequal to pedicel length. Mesosoma. Mesosoma elongate; dorsal thoracic length 0.95 × width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T1 shape sub-disciform, punctate. T2 slightly longer than wide. Disc of T2 with dense oblique punctures, intervals smooth. T3–5 and S2–5 with separated punctures, intervals obscure microreticulate. T6 convex. Incomplete lateral S6 carina rounded, raised.</p><p>Material examined. USA, Arizona, Pima County, 550m, Silver Reef Wash, 4 km E. Vaiva Vo, Tat Momoli Mountains, malaise, 1–7.V.2006, M. E. Irwin (1♂, EMUS, Fig. 64) ; Texas: Kennedy County, 2.5 mi. S Sarita, 18–20.X.2002, B. Raber &amp; E. Riley, pit-fall in sand (1♀, TAMU, Figs 3, 7); LaSalle County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-99.383&amp;materialsCitation.latitude=28.293" title="Search Plazi for locations around (long -99.383/lat 28.293)">Chaparral Wildlife Management Area</a>, 13 mi. SW Cotulla, 28.293°N 99.383°W, 19.V.2006, J.S. Wilson and K.A. Williams (2♀, CSCA EMUS, Figs 2, 6) .</p><p>Distribution. Widespread in the western Nearctic, including Arizona, California, New Mexico and Texas (USA) and Sonora (Mexico).</p><p>Remarks. This widespread species has variably colored tegulae in males. In Arizona and California, this species has orange tegulae, resembling the sympatric L. cirrhomeris (Fig. 42), L. ilex (Fig. 61), and L. litosisyra (Fig. 63); farther East, the males are uniform blackish.</p><p>The putative sex association presented here is based on a process of elimination. Lomachaeta hicksi and L. vacamuerta are the only Lomachaeta species known from southern Texas and the female of L. hicksi is distinct (in nearly all species-level characters) from the females described above.</p><p>The female from Kennedy County (Fig. 3) has different coloration than the two females from LaSalle County (Fig. 2). It is the only Lomachaeta we have seen with a dark head and mesosoma that contrast with an orange metasoma, a pattern rarely seen in mutillids, except among Dasymutilla Ashmead and Pseudomethoca Ashmead from the Texan mimicry ring (Wilson et al., 2015). If natural selection has favored the similarities of this species to the Texan mimicry ring, various hypotheses could explain its limited mimetic fidelity. These include phylogenetic constraints of the genus Lomachaeta, the relaxed selection hypothesis based on its small size (Penney et al., 2012), or the community diversity hypothesis based on presence of multiple potential models (Wilson et al., 2013).</p></div>	https://treatment.plazi.org/id/03F187A07C1BCD2D27B79A4AFAB0FE52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.text	03F187A07C18CD2D27B79A1BFB09F95F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta vianai Casal 1969	<div><p>Lomachaeta vianai Casal, 1969</p><p>(Figs 13, 14, 19, 20, 55)</p><p>Lomachaeta vianai Casal, 1969: 34 . Holotype, ♀, Argentina, Misiones, Dep. Concep. (AMNH, examined). Lomachaeta megomicron Williams &amp; Pitts, 2009: 235 . Holotype, ♂, Argentina, Salta, 8km N La Viña (EMUS, examined), syn. nov.</p><p>Diagnosis. MALE. The following combination of characters is diagnostic for L. vianai: the body is black, except the pale yellow T1 apex; the mandible lacks a ventral tooth basally; the gena is weakly carinate; the head and pronotum are areolate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is disciform; the T2 disc has dense small punctures; the T2 fringe is composed of thick bristles; and the paramere is virtually straight and has an weak apical setal tuft. Body length 4–6 mm.</p><p>FEMALE. The following characters are diagnostic: the head and mesosoma are dark reddish, the metasoma is blackish; the pronotal and propodeal spiracles are scarcely swollen; the head is 1.2 × wider than the mesosoma; the lateral propodeal face has a few complete areolations; the mesosoma is elongate, with the thoracic dorsum nearly as long as wide; the mesosomal dorsum is tightly areolate without subparallel thickened setae; the T1 shape is subdisciform; T2 has dense oblique punctures, mostly subappressed pale yellow setae, and mostly smooth intervals; and the S6 lateral carina is low angular. Body length 2.5–4.5 mm.</p><p>Material examined. Type material. Holotype of Lomachaeta vianai, ♀, ARGENTINA, Misiones, Dep. Concepcion, Santa María, X.1956, M.J. Viana (AMNH, Figs 13, 19) ; Holotype of Lomachaeta megamicron, ♂, ARGENTINA, Salta, 8 km N La Viña, 26.X–13.XI.2003, M.E. Irwin &amp; F.D. Parker coll. (EMUS). Other material. ARGENTINA: Salta: 8 km N La Viña, 26.X–13.XI.2003, M.E. Irwin &amp; F.D. Parker (1♂, EMUS, Fig. 55); Chaco Occidental, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Los Colorados</a>, 100 km NE Joaquin V. Gonzales, 14.III.1992, B. Bestelmeyer (1♀, MIUP); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Pocitos</a>, XI.1978, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Fritz</a> (1♂, MIUP). BRAZIL, Goiás, Parque Nacional Chapada dos Veadeiros, A.P. Aguiar and B.F. dos Santos, Yellow Pan Trap: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Ponto</a> 1, 13–15.IX.2005 (1♂, UFES); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Ponto</a> 3, 19-21.IX.2005 (2♂, UFES); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Ponto</a> 6, 9–11.IX.2005 (1♂, UFES); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Ponto</a> 10, 9–11.IX.2005 (2♂, UFES); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Ponto</a> 15, IX.2005 (1♂, UFES). PARAGUAY: Boquerón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Mariscal Estigarribia</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.77186&amp;materialsCitation.latitude=-22.69014" title="Search Plazi for locations around (long -59.77186/lat -22.69014)">Escuela Agricola</a>, 174m, 21 o 59’45”S 60 o 38’31”W, 5–7.X.2003, B. Garcette, Malaise (1♂, MIUP); Presidente Hayes, Los Pioneros, 22.69014 o S 59.77186 o W, 9–11.X.2003, M. LePonce, Pit Fall Trap (1♀, MIUP, Figs 14, 20) .</p><p>Distribution. Argentina (Catamarca, Misiones, and Salta Provinces), Brazil (Goiás), and Paraguay (Boquerón and Presidente Hayes Departments).</p><p>Remarks. Previously known from the unique type only, this species is now recognized as a widely distributed species in South America. Although this species is widespread, it is known from only seven localities. The sex association is based on a male and female collected in similar habitats (Chaco Boreal) in Paraguay.</p></div>	https://treatment.plazi.org/id/03F187A07C18CD2D27B79A1BFB09F95F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.text	03F187A07C18CD1227B79F10FC40FC5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lomachaeta warneri Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019	<div><p>Lomachaeta warneri Williams, sp. nov.</p><p>(Figs 36, 40)</p><p>Diagnosis. FEMALE. This species can be recognized by the following combination of characters: the body is predominantly orange-brown, except the antennae, legs, and T3–6 are somewhat darker brown; the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the erect dorsal brachyplumose setae of the whole body are mostly yellow-brown; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the T2 disc is sparsely punctate with the intervals partly smooth and partly microreticulate; the T2 fringe is somewhat dense with slightly thickened silvery setae; and the S6 lateral carina is shallow, but distinct.</p><p>Description. FEMALE. Body length 2.5–3 mm. Coloration. Body reddish-orange except F2–10 dark brown; femora, tibiae, and tarsi largely brown; T2 apex and T3–5 brown; and T6 dark brown. Tibial spurs white. Erect dorsal brachyplumose setae entirely white to pale brown, many setae lighter at base; except T6 with setae predominantly brown. Mesonotum with few scattered subappressed simple white setae; fringes of T2–5 composed of subappressed white setae, those of T2 silvery, thicker than surrounding erect setae. Ventral and pleural setae white, except S6 setae largely brown. Head. Head width 1.1 × pronotal width. Frons, vertex and gena areolate, vertex with many intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with barely perceptible ventral lamella that does not interrupt ventral mandibular contour. Antennal scrobe without dorsal carina. Lengths of F1 and F2 each subequal to pedicel length. Mesosoma. Mesosoma compact; dorsal thoracic length 0.85 × width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; areolations clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T1 shape sub-sessile, mostly smooth. T2 slightly longer than wide. Disc of T2 with sparse punctures, intervals smooth. T3–5 and S2–5 with separated punctures, intervals shagreened. T6 convex. Incomplete lateral S6 carina shallow but distinct.</p><p>MALE. Unknown.</p><p>Material examined. Holotype, ♀, USA, Arizona, Coconino County, Hwy. 89A, 4.9 mi. [7.8 km] ESE of jct. 389, 36 o 55’20”N 112 o 15’10”W, 24.IV–25.V.2014, W.B. Warner, barrier pitfall with black cups (CSCA, Figs 36, 40) . Paratypes: USA: Arizona, Coconino County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.25278&amp;materialsCitation.latitude=36.922222" title="Search Plazi for locations around (long -112.25278/lat 36.922222)">Hwy.</a> 89A, 4.9 mi. [7.8 km] ESE of jct. 389, 36 o 55’20”N 112 o 15’10”W, 29.VI–26.VII.2014, W.B. Warner, barrier pitfall with black cups (2♀: ASUT, FSCA) ; Utah, Washington County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.3259&amp;materialsCitation.latitude=37.2649" title="Search Plazi for locations around (long -113.3259/lat 37.2649)">Mills Rd.</a>, 1.6 mi. [2.6 km] S I-15 exit 27; 37.2649 o N 113.3259 o W, 15.IV–3.VII.2015, barrier pitfall with black cups (1♀, CSCA) . Other Material. USA: Arizona, Pinal County, 4 mi. [6.4 km] W Stanfield, 12.IV.1963, G.I. Stage (1♀, EMUS) .</p><p>Distribution. Colorado Plateau in Arizona and Utah.</p><p>Etymology. Named in honor of William B. Warner, the collector of the holotype and many other important mutillid specimens.</p><p>Remarks. This female apparently belongs to the L. crocopinna species-group (Williams &amp; Pitts 2009) in having the ventral mandibular lamella obliterated and the pronotal and propodeal spiracles lowly produced. Within this apparent group, the thickened T2 fringe setae are unique. The specimen from Pinal County Arizona has the cuticle somewhat darker, the dorsal setae longer and darker, and the intervals of T2 with the microreticulations more distinct. Although these differences are not great enough to justify describing it as a distinct species at this point, they are enough to justify removing it from the paratype series.</p></div>	https://treatment.plazi.org/id/03F187A07C18CD1227B79F10FC40FC5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto A.;Bartholomay, Pedro R.;Luz, David R.;Quintero, Diomedes;Pitts, James P.	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
