taxonID	type	description	language	source
03F187A07C04CD3127B79AE8FDEBFCB6.taxon	materials_examined	Type species: L. hicksi Mickel, 1936	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3127B79B3DFC16FA85.taxon	materials_examined	Type species: S. powelli Mickel, 1964. Synonymized by Williams & Pitts (2009).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3127B79B3DFC16FA85.taxon	diagnosis	Diagnosis. FEMALE. Females of Lomachaeta can be recognized by the following combination of characters: the body setae are sparse, nowhere obscuring the sculpture; the mesosoma has coarse sculpture and lacks a scutellar scale; T 1 is slender, with its shape either sub-sessile or sub-disciform; the tergal fringes are not composed of distinctly plumose setae; and T 6 lacks a defined pygidium.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3127B79B3DFC16FA85.taxon	description	MALE. Males of Lomachaeta can be recognized by the following combination of characters: the body setae are sparse, nowhere obscuring the sculpture; the axilla is armed with a posteriorly directed tooth; T 1 is slender, with its shape either sub-sessile or disciform; the tergal fringes are not composed of distinctly plumose setae; and the digitus is sessile with the usually short pad-like cuspis.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C04CD3127B79B3DFC16FA85.taxon	distribution	Distribution. Widespread in the New World, from Canada to Argentina.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	description	(Figs 1, 5)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	diagnosis	Diagnosis. FEMALE. This species can be recognized by its setal distribution and body shape: the mesosoma and T 2 have matching subparallel subappressed long silver to golden setae and the dorsal thoracic length is 0.85 × its width. The following characters may also aid in diagnosis: the head width is subequal to the mesosomal width; the baso-ventral mandibular margin lacks a hyaline lamella; the pronotal spiracle is only weakly swollen; the lateral mesonotal tooth is distinct; the propodeal spiracle is slightly more sharply swollen than the pronotal spiracle; the mesonotal dorsum has about 12 areolations between the pronotal spiracles; the T 1 shape is sub-disciform; T 2 has dense punctures and mostly smooth intervals; the fringes of T 2 – 3 are composed of dense setae thicker than the erect setae surrounding them; and the S 6 lateral carina is small and sharply triangular. Body length 3 – 3.5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	description	MALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	materials_examined	Material examined. USA: Arizona: Yavapai County, 11 mi W Jerome, Rearing 7598, F. D. Parker (2 ♀, EMUS); Coconino County, 7 mi S. Sedona, Reared, F. D. Parker (2 ♀, EMUS); Mohave County, Oatman 3 mi S, Reared, F. D. Parker (1 ♀, EMUS); Nevada, Esmeralda County, 1.5 mi. S, 4 mi. W Lida Summit, 7400 ’, 28. III – 1. X. 1982, D. Giuliani, ethylene glycol pit trap (1 ♀, CSCA); Nye County, Ash Meadows, NWR [Natural Wildlife Reserve], Non-dune salt soil site, S. Devils hole rd. BR, 11 s 0563221 4029908, 11. V. 2009, N. Boehme (1 ♀, EMUS); Utah, Washington County, 3 mi W Hurricane, reared, F. D. Parker (1 ♀, paratype, EMUS, Figs 1, 5).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	distribution	Distribution. Known from the Great Basin, Mojave Desert, and the Arizona Mountains forests in Arizona, Nevada, and Utah, USA.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C02CD3627B79E5BFD66FD0E.taxon	discussion	Remarks. This species, formerly recognized from one Great Basin locality in southern Utah, is newly recorded from Arizona and Nevada in two additional ecoregions. The newly identified specimens from Arizona were collected from trap nests that also included L. crocopinna, L. cirrhomeris, and L. hicksi. Since the females of L. hicksi and L. cirrhomeris are already known, L. crocopinna could be the unknown male of L. argenta. These specimens, however, are far removed from the type locality of L. crocopinna, so we prefer to wait for more evidence before formalizing this synonymy.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	description	(Figs 34, 38, 47, 66)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. beadugrimi: the body is entirely black, except T 2 – 3 are largely orange; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the forewing has veins encompassing the basal 0.7 × wing; the T 1 shape is sub-sessile; the T 2 fringe is composed of simple setae; and the paramere is dorsoventrally flattened, apically rounded, and has short setae only. Body length 3.5 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	description	FEMALE. This species can be recognized by the following combination of characters: the body is predominantly orange, except the antennae, legs, and T 6 are largely dark-brown; the mandible lacks a distinct ventral lamella; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the mesosoma is compact, with the thoracic dorsal length 0.8 × its width; the T 2 disc is sparsely punctate, clothed with erect blackish brachyplumose setae, with the intervals microreticulate; the T 2 fringe is sparse and simple; and the S 6 lateral carina is distinctly triangular. Body length 2.5 – 3 mm. Description. FEMALE. (hitherto unrecognized, based on female from Kern County). Body length 2.5 – 3 mm. Coloration. Body reddish-orange except F 2 – 10 dark brown; femora, tibiae, and tarsi dark brown apically; and T 6 dark brown. Tibial spurs white. Erect dorsal brachyplumose setae on frons, vertex, mesonotum, T 2 disc, and T 6 black-brown; on vertex, pronotum, propodeum, T 1, and T 2 base white; and on T 3 – 5 interspersed brown and white. Mesonotum with few scattered subappressed simple white setae; fringes of T 2 – 5 composed of sparse subappressed simple white setae. Ventral and pleural setae white, except S 6 setae black-brown. Head. Head width subequal to pronotal width. Frons, vertex and gena areolate, some vertex intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe without dorsal carina. Length of F 1 subequal to pedicel; F 2 1.1 × pedicel length. Mesosoma. Mesosoma compact; dorsal thoracic length 0.8 × width. Humeral carina angulate dorsally, weak ventrally, not reaching epaulet. Mesosomal dorsum areolate; most intervals clearly defined, not raised into tubercles; areolations evenly distributed; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle distinct, small. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-sessile, subareolate. T 2 slightly longer than wide. Disc of T 2 with sparse punctures, intervals microreticulate. T 3 – 5 and S 2 – 5 with separated punctures, intervals microreticulate. T 6 convex, incomplete lateral carina triangular.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	materials_examined	Material examined. USA: Arizona: Maricopa County, Gila River, 10 km S. Arlington, malaise on sand beach, 200 m, 33 ° 13.3 ’ N 112 ° 45.53 ’ W, 25. V – 03. VI. 2010, M. E. Irwin (1 ♂, EMUS); Maricopa County, Gila River, 8 km S. Arlington, malaise in opening nr. tamerisk, 200 m, 33 ° 13.42 ’ N 112 ° 46.27 ’ W, 7 – 13. VI. 2010, M. E. Irwin (1 ♂, EMUS); California: Kern County, Munsey Road, 5.8 km E Neuralia Road, 35.2872 o N 117.9223 o W, 590 m, 24 – 25. VI. 2017, K. A. & E. E. Williams (1 ♀, CSCA, Figs 34, 38); San Bernardino County: 6 mi E Phelan, Baldy Mesa, 21. IV – 27. IV. 1981, pan traps, J. T. Hubert (1 ♂, CSCA, Fig. 47); 10 mi. NW Barstow, 2200 ’, 4. VI. 1971, R. R. & R. A. Snelling (1 ♀, EMUS); 4 mi. NW Adelanto, 2900 ft., 34 ° 39 ’ N 117 ° 26 ’ W, 22. V. 1993, G. C. Snelling (71 ♀ 45 ♂, EMUS, Fig. 66); 4 mi. NW Adelanto, 2900 ft., 34 ° 39 ’ N 117 ° 26 ’ W, 11. VI. 1993, G. C. Snelling, on Chamaesyce sp. (6 ♀, EMUS); 4 mi. NW Adelanto, 880 ft., 34 ° 65 ’ N 117 ° 43 ’ W, 02. VII. 1995, R. R. & G. C. Snelling (11 ♀, EMUS); Los Angeles County, 3.2 km S. Pearblossom, 1067 m, 15. VIII. 1977, R. R. & G. C. Snelling (1 ♀, EMUS).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	distribution	Distribution. Hot deserts in Arizona, California, and Nevada.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C03CD3527B79AA0FDE0FCB1.taxon	discussion	Remarks. This species was originally placed in the genus Smicromutilla because it lacked thickened bristles on the T 2 fringe. It is closely related to L. snellingella, which may eventually be recognized as a darkened southern variety of L. beadugrimi. This sex association is based on a series of 45 males and 88 females collected near Adelanto, California by G. C. Snelling in the early 1990 ’ s. These wasps were apparently feeding together on nectar from flowers of ground covering spurges in the subgenus Euphorbia (Chamaesyce) S. F. Gray. The female from Kern County was collected in a sandy wash where workers of the ant genus Forelius Emery were foraging. The mutillid superficially resembled these ants, differing by slightly larger size and a not easily describable difference in gait. It is not clear whether Lomachaeta are purposeful co-mimics with ants or this similarity is purely coincidental.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	description	(Figs 49, 50, 67, 68)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique coloration, wherein the entire body is orange, except the darkened apical tergites. The following characters are also useful for diagnosis: the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins restricted to the basal half of wing; the T 1 shape is sub-disciform; the T 2 disc is has separated punctures; the T 2 fringe is composed of simple setae; and the paramere is laterally compressed and moderately setose.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	description	Description. MALE. Body length 4 mm. Coloration (Figs 36, 49). Entire body pale orange-brown, except frons, scape, tibiae, and T 3 largely brown; flagellum, tarsi, and T 4 – 7 dark brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, mesoscutum, and T 3 – 7 with interspersed erect silvery and brownish brachyplumose setae. Head. Rounded posteriorly, with coarse punctures, sparser around ocelli. Mandible tridentate apically, unarmed ventrally. Clypeus punctate, anterior margin smoothly rounded. Genal carina weak. Ocelli small; ocellocular distance 3 × length of lateral ocellus, interocellar distance subequal to lateral ocellar diameter. F 1 1.0 × pedicel length; F 2 1.2 × pedicel length. Mesosoma. Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins sparsely setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron smooth. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.5 × wing. Metasoma. T 1 shape sub-disciform, without carina separating anterior and dorsal faces. T 2 with separated punctures, intervals smooth; T 2 fringe setae simple. S 2 with coarse oblique punctures. T 3 – 6 punctures sparse. T 7 smooth. Hypopygium punctate, sharp bidentate posteriorly. Genitalia (Fig. 67, 68). Paramere laterally compressed, straight, subacuminate apically, with short setae along inner and outer surfaces, setae longer and denser apicolaterally, few long setae ventrally in basal half. Cuspis with posteroventral setae subequal to cuspis length. Penis valve unidentate apically. FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	materials_examined	Material examined. Holotype, ♂, USA, California, Riverside County, Riverside, 14. VIII. 1969, J. C. Hall, malaise trap (EMUS, Figs 49, 50, 67, 68).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	distribution	Distribution. Known only from the type locality in southern California.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	etymology	Etymology. Named after the calamondin, a small orange-like fruit frequently grown in southern California, in reference to the small orange-ish color of this southern Californian species. Treat as a noun in apposition.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C00CD3B27B79B3CFAD8FB4A.taxon	discussion	Remarks. Based on the reduced wing venation, simple parameres, and distribution in California, this is likely the sister species to L. powelli. In the western USA, this is apparently the only diurnal male mutillid with the head and mesosoma pale orange-brown (Fig. 49), although many nocturnal species can be superficially similar.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	description	(Figs 21, 27, 41)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. chionothrix: the body is entirely black; the mandible has a ventral tooth basally; the forewing has its veins encompassing the basal 0.7 of the wing; the mesoscutum is densely coarsely punctate; the T 1 shape is disciform; the head and T 2 disc have separated punctures; the T 2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks elongate setae. Body length 3.5 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	description	FEMALE. As in males, this species is similar to the variable L. hicksi. Lomachaeta chionothrix has tighter areolations on the head and mesosoma, with 13 areolations between the pronotal spiracles; and has T 6 basally with dense long setae that are black basally and white apically. The following characters may also be useful for diagnosis: the baso-ventral mandibular margin has a weak hyaline lamella; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the pronotal and propodeal spiracles are swollen and tuberculate; the lateral mesonotal tooth is undefined; the lateral propodeal face is impunctate; the mesosomal dorsum lacks subparallel thickened setae; the T 1 shape is sub-disciform; T 2 has sparse punctures, mostly erect brachyplumose setae, and mostly smooth intervals; the T 2 fringe is composed of dense thick white setae; and the S 6 lateral carina is shallow. Description. FEMALE. (hitherto unrecognized, based on female from Jalisco). Body length 4.5 mm. Coloration. Body orange-brown except flagellum, mesosomal dorsum, legs, and T 1 variegated with brown and metasomal segments 2 – 6 dark brown to black. Tibial spurs white. Erect dorsal brachyplumose setae on mesonotum, T 2 disc, and T 3 – 6 black-brown; remaining setae whitish. Fringe of T 2 composed of dense subappressed thick white setae, fringes of T 3 – 4 with sparse thick white setae. Head. Head width 1.1 × pronotal width. Frons, vertex and gena tightly areolate, all areolations complete. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with weak ventral lamella basally. Antennal scrobe without dorsal carina. Length of F 1 1.4 × pedicel; F 2 length subequal to pedicel length. Mesosoma. the Mesosoma is compact, thoracic dorsal length 0.85 × width. Humeral carina distinct, rounded dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, some raised to form apparent tubercles; intervals slightly tighter anteriorly than posteriorly; dorsally with 13 areolations between pronotal spiracles. Pronotal and propodeal spiracles swollen, tuberculiform; lateral mesonotal tubercle undefined. Lateral propodeal face with a few nearly-complete areolations. Metasoma. T 1 shape sub-disciform, punctate. T 2 as long as wide. Disc of T 2 with sparse oblique punctures, intervals mostly smooth. T 3 – 5 and S 2 – 5 with separated punctures, intervals smooth. T 6 convex. Incomplete lateral S 6 carina shallow but distinct.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	materials_examined	Material examined. COSTA RICA, Guanacaste: EJN 14 km S Cañas, 14. III. 1990, F. D. Parker (1 ♂, CSCA, Fig. 41); P. N. Barra Honda, 100 m: III. 1995 (1 ♂, MIUP); IV. 1995 (1 ♂, MIUP), M. Reyes. HONDURAS, Fco. Morazan, San Antonio de Oriente, El Zamorano, Malaise trap in coffee plantation, 25. II – 6. III. 1990, R. Cave (1 ♂, MIUP). MEXICO, Jalisco, Chamela, 10. VII. 1989, J. G. Rozen (1 ♀, MIUP, Figs 21, 27). NICARAGUA, Granada, Volcan Mombacho, Santa Ana: 15. IV. 1998 (1 ♂, MIUP); 15. V. 1998 (1 ♂, MIUP), malaise trap, J. M. Maes.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	distribution	Distribution. Mexico (Nayarit and Jalisco) South to Costa Rica. First records for Honduras and Nicaragua.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	discussion	Remarks. This species is closely related to the widespread L. hicksi, differing in males by mesosomal punctation and genitalic differences only. Until now, this species is the only Lomachaeta known from Central America and although 10 species are known from Mexico, only L. chionothrix has been found in Jalisco. Only two females have been observed from the northern Neotropical region: one from Costa Rica and one from Jalisco, Mexico. These females differ in significant structural features, especially the head and mesonotum punctation. The female from Jalisco was collected in Chamela, where L. chionothrix males have been recorded (Williams & Pitts 2009). This female is similar to L. hicksi, differing mainly by its tighter mesosomal areolations. Lomachaeta chionothrix males also have tighter mesosomal punctation than L. hicksi. Morphological similarity to L. hicksi and co-occurrence with L. chionothrix males in Chamela supports the association of this female with L. chionothrix. The female from Costa Rica cannot be differentiated from the newly associated L. hyphantria females and is discussed below.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique coloration, wherein the body is largely black, except the orange tegula and femora. The following characters are also useful for diagnosis: the head and mesosoma have coarse separated punctures; the mandible has a ventral tooth basally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is disciform; the T 2 disc has separated punctures; the T 2 fringe is composed of thickened bristles; and the paramere is straight and lacks elongate setae. Body length 4.5 – 6.5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	description	FEMALE. This species can be recognized by the body color, which is entirely orange, including the legs and antennae, except T 6 is blackish; and T 3 – 5 are clothed mostly with black erect setae except for the dense thick silver setae of the T 2 and sometimes T 3 fringe. The following characters will also aid in diagnosis: the baso-ventral mandibular margin has a hyaline lamella; the head width is 1.1 × the mesosomal width; thoracic dorsal length 0.9 × its width; the pronotal spiracle is usually swollen and nearly tuberculate; the lateral mesonotal tooth is usually distinct; the propodeal spiracle is sharply swollen and tuberculate; the lateral propodeal face is impunctate; the mesosomal dorsum has about 10 areolations between the pronotal spiracles, many areolations have the intervals either obliterated or raised to tuberculate corners; the T 1 shape is sub-sessile; T 2 has sparse punctures and mostly smooth intervals; and the S 6 lateral carina is low sub-triangular. Body length 2.5 – 4 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	materials_examined	Material examined. USA: Arizona: Maricopa County: Gila River, 10 km S. Arlington, malaise on sand beach, 200 m, 33 ° 13.3 ’ N 112 ° 45.53 ’ W, 25. V – 03. VI. 2010, M. E. Irwin (1 ♂, EMUS); Gila River, 8 km S. Arlington, malaise in opening nr. tamerisk, 200 m, 33 ° 13.42 ’ N 112 ° 46.27 ’ W, 7 – 13. VI. 2010, M. E. Irwin (1 ♂, EMUS); Pima County, Tucson, 9. I. 1939, J. Knull (1 ♀, EMUS, Figs 23, 29); additional data unavailable (1 ♂, FSCA, Fig. 42); California, Riverside County, San Timoteo Canyon, Malaise trap, 14. IX. 1972, M. Wasbauer, A. Hardy (1 ♀, CSCA, Figs 22, 28).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	distribution	Distribution. Widespread in the western Nearctic, including Arizona, California, Nevada, New Mexico, Texas, and Utah (USA) and Baja California and Baja California Sur (Mexico).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ECD3927B79D6DFE4BFD6A.taxon	discussion	Remarks. This species was placed in the L. hicksi species-group (Williams & Pitts 2009) with L. hicksi and L. chionothrix. Males in this species-group can be easily recognized because of the distinct ventral mandibular tooth. Females are more difficult to define, as the main diagnostic trait of males, the ventral mandibular tooth, is reduced to a subtle lamella in females. Additional traits for L. hicksi species-group females (couplet 7 below), are more distinct in L. hicksi and L. chionothrix than in L. cirrhomeris. If one were to mis-step at couplet seven, females of L. cirrhomeris would key out to L. beadugrimi, which can be recognized by having the T 2 intervals microreticulate (mostly smooth in L. cirrhomeris) and T 3 – 5 with interspersed silver and blackish erect setae (T 3 – 5 setae mostly black in L. cirrhomeris).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	description	(Fig. 52)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. crocopinna: the body is entirely black, except the metasoma is almost entirely orange; the mandible lacks a ventral tooth basally; the forewing has its veins encompassing the basal 0.65 × of the wing; the head, mesoscutum, and T 2 disc have separated punctures; the T 1 shape is sub-sessile; the T 2 fringe is composed of slightly thickened setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 3 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	materials_examined	Material examined. MEXICO, Sonora: Alamos, Rancho Acosta, malaise in dry wash, tropical deciduous forest, 395 m, 27 ° 01.57 ’ N 106 ° 55.37 ’ W, 26. V – 6. VI. 2007, M. E. Irwin (1 ♂, EMUS); 3 km NE Alamos, La Huerta Orchard, 27 ° 01 ’ N 108 ° 54 ’ W, 19 – 24. [Month unknown]. 2008, M. E. Irwin & O. Figuero (2 ♂, EMUS); USA, Arizona: Yavapal County, 11 mi W. Jerome, Reared, F. D. Parker (2 ♂, EMUS); Yavapal County, Clarksdale, Reared, F. D. Parker (1 ♂, EMUS); Coconino County, 7 mi S. Sedona, Reared, F. D. Parker (3 ♂, EMUS); Gila County, 2 mi S. Pine, F. D. Parker (1 ♂, EMUS); Pima County, 550 m, Silver Reef Wash, 4 km E. Vaiva Vo (Cockelburr), Tat Momoli Mts., malaise in wash, 32 ° 39.06 ’ N 111 ° 54.47 ’ W, 07 – 14. V. 2006, M. E. Irwin (5 ♂, EMUS); Maricopa County, Gila River, 10 km S. Arlington, malaise on sand flat, 200 m, 33 ° 13.30 ’ N 112 ° 45.53 ’ W, 07 – 13. VI. 2010, M. E. Irwin (1 ♂, EMUS); 4 km E. Vaiva Vo, Silver Reef Wash, 32 ° 39.06 ’ N 111 ° 54.47 ’ W, 550 m, 1 – 7. V. 2006, M. E. Irwin, malaise trap (1 ♂, EMUS); Kitt Peak rd., Coyote Mts., 5 km S. jct. Hwy. 36 and 286, malaise in wash, 1300 m, 31 ° 59.32 ’ N 111 ° 33.79 ’ W, 2 – 12. V. 2006, M. E. Irwin (1 ♂, EMUS); New Mexico: Socorro, NWR [Natural Wildlife Resort] LTER [Long Term Ecological Reserve] site 1, 26. V. 1992 (1 ♂, EMUS); Texas, Culberson County, Choza Springs, Guadalupe Mountains, 1610 m, 8. IV. 1996 (1 ♂, EMUS, Fig. 52); Utah: Garfield County, Alvey Wash, 7 km S. Escalante, pan trap in dry wash, 1990 m, 37 ° 42.5 ’ N 111 ° 37.8 ’ W, 24 – 25. V. 2002, F. D. Parker & M. E. Irwin (2 ♂, EMUS); Garfield County, Alvey Wash, 7 km S. Escalante, malaise in dry wash, 1990 m, 37 ° 42.5 ’ N 111 ° 37.8 ’ W, 24 – 25. V. 2002, M. E. Irwin & F. D. Parker (1 ♂, EMUS).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	distribution	Distribution. Known from the Arizona Mountains forests, Colorado Plateau, Great Basin, and Chihuahuan Desert in Arizona, Nevada, New Mexico, and Texas (USA) and Sonora (Mexico).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0CCD3827B79B43FAA9FF72.taxon	discussion	Remarks. This is the apparent sister species to L. ptilohyalus; the two species share similar coloration and genitalic morphology. The distribution of L. crocopinna spans multiple ecoregions in the western USA and parallels that of Dilophotopsis concolor (Cresson). Wilson and Pitts (2008) described the distribution of D. concolor and its sister species, D. paron (Cameron). Dilophotopsis paron occurs in the Mojave and Sonoran deserts, as does L. ptilohyalus. The parallel distributions of these sister-species pairs should be studied more closely, as they could shed light on historic events that impacted the biogeography of numerous velvet ants.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	description	(Figs 59, 69, 70)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the cuspis and lateral paramere margins have numerous elongate (~ 5 × paramere width) setae. The following characters are also useful for diagnosis: the body is entirely blackish; the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-disciform; the T 2 disc is densely coarsely punctate; the T 2 fringe is composed of simple setae.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	description	Description. MALE. Body length 4 mm. Coloration (Fig. 40). Entire body black, except mandible, tegula, and legs largely brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, and mesoscutum with interspersed erect silvery and brownish brachyplumose setae. Head. Rounded posteriorly, with coarse punctures, coarser on face. Mandible tridentate apically, unarmed ventrally. Gena not carinate. Ocelli small; ocellocular distance 6 × length of lateral ocellus, interocellar distance 2 × lateral ocellar diameter. F 1 1.0 × pedicel length; F 2 1.6 × pedicel length. Mesosoma. Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron glabrous. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.7 × wing. Metasoma. T 1 shape sub-disciform, with defined anterior and dorsal faces. T 2 with coarse dense punctures, intervals smooth; T 2 – 4 fringe setae slightly thicker than disc setae, but not forming bristles. S 2 with coarse separated punctures. T 3 – 6 punctate. T 7 smooth. Hypopygium punctate, sharply bidentate posteriorly. Genitalia (Figs 69, 70). Paramere dorsoventrally flattened, obliquely downcurving to apex, sub-truncate apically, with long (~ 5 × paramere width) ventral setae around inner and lateral margins, setae longer and denser apically. Cuspis with posteroventral setae ~ 5 × cuspis length. Penis valve unidentate apically. FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	materials_examined	Material examined. Holotype, ♂, MEXICO, Sonora, km 23, Hwy 16, 5.5 km ENE [Los] Hornos, 23 – 26. VII. 2008, 27 o 43.71 ’ N 109 o 50.64 ’ W, 40 m, M. E. Irwin & R. L. Wescott, malaise in dense riparian forest (EMUS, Figs 59, 69, 70).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	distribution	Distribution. Known only from the type locality in Sonora, Mexico	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	etymology	Etymology. From the Greek, eironeia, irony, in reference to the fact that this specimen was recognized as a new species by KAW only a few months after Williams & Pitts 2009 was published.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3827B79975FDC7FA39.taxon	discussion	Remarks. The genitalia of this species are truly bizarre (Fig. 59), even for a genus with such variable parameres (Williams & Pitts 2009).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	description	(Figs 33, 37, 62)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is virtually straight, laterally subcompressed, and long setae throughout apical half of free paramere length ventrally. The following characters are also useful for diagnosis: the body is entirely blackish; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-sessile; the head and T 2 disc have separated punctures; and the T 2 fringe is composed of simple setae. Body length 4 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	description	FEMALE. This species can be recognized by the body color, which is mostly orange, except the legs, antennae and T 6 are entirely blackish. Additional useful diagnostic features include: the mandible lacks a ventral lamella; the mesosoma is compact, with the thoracic dorsal length 0.8 × its width; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the T 2 disc is sparsely punctate, clothed with erect blackish brachyplumose setae, and the intervals mostly micropunctate; the T 2 fringe is sparse and simple; and the S 6 lateral carina is distinctly triangular. Description. FEMALE (hitherto unrecognized, based on female from Arroyo San Gregario). Body length 2.5 – 3 mm. Coloration. Body reddish-orange except antenna, legs (including coxae), and T 6 blackish, mid and hind legs (including coxae) sometimes partly orange. Tibial spurs white. Erect dorsal brachyplumose setae on frons, anterior vertex, mesonotum, T 2 – 5 disc, and T 6 black-brown; on posterior vertex, pronotum, propodeum, T 1, and T 2 base yellowish. Mesonotum with few scattered subappressed simple white setae; fringes of T 2 – 5 composed of sparse subappressed simple whitish setae. Ventral and pleural setae white, except S 6 setae black-brown. Head. Head width subequal to pronotal width. Frons, vertex and gena areolate, edges of many frons and vertex areolations obliterated, areolation edges on frons and vertex forming parallel longitudinal apparent striae. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe lacking dorsal carina. Length of F 1 subequal to pedicel; F 2 1.2 × pedicel length. Mesosoma. Mesosoma compact, dorsal thoracic length 0.8 × width. Humeral carina weak, not reaching epaulet. Mesosomal dorsum areolate; many intervals obliterated; areolations evenly distributed; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle small, but distinct. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-sessile, punctate. T 2 as wide as long, laterally slightly swollen subapically. Disc of T 2 with sparse incomplete apparent areolations, intervals microreticulate. T 3 – 5 and S 2 – 5 with separated punctures, intervals microreticulate. T 6 convex. Incomplete lateral S 6 carina distinct triangular.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	materials_examined	Material examined. Paratypes, 2 ♂, MEXICO, Baja California Sur, Arroyo San Gregorio, 13 air km WNW La Purissima, 24 – 26. Apr. 1983, M. S. Wasbauer coll. (CSCA, Fig. 62). Other material. MEXICO: Baja California Sur, same data as above (4 ♀, CSCA, Figs 33, 37).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	distribution	Distribution. Baja California Sur in Mexico.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0DCD3F27B79DAFFDF2FBDB.taxon	discussion	Remarks. The females here associated with L. hedera have identical label data to two of the male paratypes (Williams & Pitts 2009). Only one other Lomachaeta species has been found in Baja California Sur, L. cirrhomeris. Unlike L. cirrhomeris, L. hedera females have black legs and antennae and a completely unmodified ventral mandible margin. These females are structurally similar to L. beadugrimi and L. warneri sp. nov., especially in their microreticulate T 2 intervals.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	description	(Figs 24 – 26, 30 – 32, 43 – 46, 65)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. hicksi: the body is black, though T 2 – 3 are often largely red-brown; the mandible has a ventral tooth basally; the mesoscutum is variably punctate, but never contiguously punctate throughout; the forewing has its veins encompassing the basal 0.8 × of the wing; the T 1 shape is disciform; the head and T 2 disc have separated punctures; the T 2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks elongate setae. Body length 3 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	description	FEMALE. This is a hugely variable species in color, setation, and punctation. The following characters are consistent to all L. hicksi: the baso-ventral mandibular margin has a hyaline lamella; the pronotal spiracle is swollen, almost tuberculate; the lateral mesonotal tooth is usually distinct; the propodeal spiracle is sharply swollen and tuberculate; the lateral propodeal face is impunctate; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the mesosomal dorsum is areolate without subparallel thickened setae, many intervals are either obliterated or raised to tuberculate corners; the T 1 shape is sub-disciform, often with the dorsal and anterior faces separated by a weak carina; T 2 has sparse punctures, mostly erect brachyplumose setae, and mostly smooth intervals; and the S 6 lateral carina is low angular. Body length 2.5 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	materials_examined	Material examined. USA: Arizona, Coconino County, 3 mi S Sedona, reared, F. D. Parker (1 ♀, EMUS, Figs 25, 31); California: Kern County, Short Canyon, 35.7093 o N 117.9112 o W, 1200 m, 21 – 29. V. 2008, M. E. Irwin, malaise trap (1 ♂, EMUS); Los Angeles County, Placentia County Park, Walker Ranch, 34.38 o N 118.44 o W, 8 – 17. IX. 1998, B. V. Brown, malaise trap (1 ♂, EMUS, Fig. 45); Riverside County: Palms Springs, Araby Wash, 33.750 o N 116.5319 o W, 538 m, F. D. Parker and T. D. McIntyre, malaise trap: 14 – 21. IV. 2015 (1 ♂, EMUS, Fig. 44); 21 – 28. IV. 2015 (1 ♂, EMUS); 21 – 24. IV. 2016 (1 ♂, CSCA); 17 – 21. V. 2016 (1 ♂, CSCA); Riverside, 30. VI. 1969, J. C. Hall, malaise trap (1 ♂, EMUS); San Diego County, 2 km SE Pine Valley near Laguna Summit, 32 ° 48.2 ’ N 116 ° 30.4 ’ W, 1280 m, 31. V – 5. VI. 2002, M. E. Irwin and F. D. Parker, malaise trap in sandy ravine (2 ♂, CSCA EMUS); Florida, Alachua County, additional data unavailable (1 ♂ 1 ♀, FSCA, Figs 24, 30, 43), Utah, Cache County, Logan Canyon # 3, reared, F. D. Parker (1 ♀, EMUS, Figs 26, 32).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	distribution	Distribution. This is the most widespread Nearctic mutillid species, ranging from the Atlantic to Pacific Oceans, North to Massachussetts and South to Florida (USA) and Puebla (Mexico).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C0ACD2227B79C93FDA4FEC3.taxon	discussion	Remarks. As one of the World’s most widely distributed mutillid species, L. hicksi has the longest synonym list in this genus, belying its tremendous morphological variation. Although fewer females than males have been available to study, they also reveal high variation in color and punctation (Figs 24 – 26). The mesosoma and metasoma each vary from nearly entirely pale orange to nearly black. The mesonotal areolations vary from broadly to tightly areolate, though when the mesonotal areolations are tightly spaced, there are fewer than 12 areolations between the pronotal spiracles and the propodeal areolations are much broader. The males mentioned above in the Material Examined section are the first known to have orange femora, a trait that was apparently unique to males of L. cirrhomeris. Males of L. cirrhomeris, however, always have cuticle of the metasoma entirely black and contrasting with their orange femora (Fig. 42), while these L. hicksi from California have the metasoma predominantly orange and concolorous with the femora (Fig. 44, 45). Pitts & Manley (2004) suggested that L. cirrhomeris may eventually prove to be a synonymous color variant of L. hicksi. These males lend additional credence to that hypothesis.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	description	(Figs 9, 10, 15, 16, 53, 54)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. hyphantria: the head and mesosoma are variegated orange and dark brown; the metasoma is black, except the pale yellow T 1 apex; the mandible lacks a ventral tooth basally; the head and pronotum are areolate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is disciform; the T 2 disc has small punctures; the T 2 fringe is composed of thick bristles; and the paramere is virtually straight and lacks an apical setal tuft. Body length 3 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	description	FEMALE. This species can be recognized by the combination of a broadly areolate mesosomal dorsum and well-defined areolate lateral propodeal face. The areolations of the head are broad and have many transverse intervals obliterated, leaving a few apparent subparallel rugae. Additional useful diagnostic features include: the mandible lacks a ventral lamella; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the mesosoma lacks appressed subparallel pale setae; the T 2 disc is moderately punctate, clothed with erect blackish brachyplumose setae, and the intervals mostly smooth; the T 2 fringe is sparse and simple; and the S 6 lateral carina is weak. Description. FEMALE (hitherto unrecognized, based on female from Venezuela). Body length 2.5 – 3.5 mm. Coloration. Head, mesosoma, legs, and T 1 variegated orange brown and dark brown; remainder of metasoma blackish. Tibial spurs white. Erect dorsal brachyplumose setae yellowish to black-brown, blackish setae often whitish at bases. Head, mesonotum, and T 3 – 5 with scattered subappressed simple usually white setae; fringes of T 2 – 5 composed of sparse subappressed simple white setae. Ventral and pleural setae predominantly whitish. Head. Head width 1.1 × pronotal width. Frons, vertex and gena broadly areolate; frons and vertex many transverse intervals obliterated, leaving subparallel subrugae. Clypeus without distinct teeth or ridges. Genal carina distinct to hypostomal carina, angulate at junction. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe with dorsal carina. Length of F 1 1.2 × pedicel; F 2 length subequal to pedicel length. Mesosoma. Dorsal thoracic length 0.85 × width. Humeral carina distinct to epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 8 areolations between pronotal spiracles. Pronotal spiracles low rounded; propodeal spiracle tuberculiform; lateral mesonotal tubercle weakly defined. Lateral propodeal face areolate, with multiple complete areolations. Metasoma. T 1 shape sub-disciform, punctate. T 2 slightly longer than wide. Disc of T 2 with dense oblique punctures, intervals smooth. T 3 – 5 and S 2 – 5 with separated punctures, intervals smooth. T 6 convex. Incomplete lateral S 6 carina weak.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	materials_examined	Material examined. BOLIVIA, Beni, Rio Itenez about 4 km above Costa Marques (Brazil), 12 – 18. IX. 1964, Bouseman and Lussenhop (1 ♂, MIUP); BRAZIL: Roraima, Caracaraí, Parque Nacional de Viruá, Vicinal na estrada perdida, 01 ° 28 ’ N 60 ° 58 ’ W, 24. XII. 2015 – 07. I. 2016, J. A. Rafael e equipe, malaise (1 ♂, INPA); Alto Alegre, Estação Ecológica de Maracá, 03 ° 21 ’ 59 ’’ N 61 ° 26 ’ 04 ’’ W, 01 – 15. IV. 2016, R. Boldrini & J. A. Rafael, malaise (1 ♂, 1 ♀, INPA); Amazonas, Manaus, ZF 2, Km 14, 02 ° 35 ’ 21 ’’ S 60 ° 06 ’ 55 ’’ W, 3 – 17. VIII. 2016, J. A. Rafael & F. F. Xavier-Filho, malaise em igarapé perto da torre (1 ♀, INPA, Figs 9, 15); 15 – 31. XII. 2016, J. A. Rafael & F. F. Xavier- Filho, malaise a 18 m na torre (1 ♂, INPA, Fig. 53); Reserva Ducke, 30. IX – 9. X. 2005, A. Aguiar, Malaise (1 ♂, INPA); 2 – 4. X. 20005, A. Aguiar, yellow-pan trap (1 ♀, INPA); Baixio Leste / Oeste, 14. II – 6. III. 2007, Freitas, G. & Feitosa, M, Malaise suspense no dossel (1 ♂, INPA); EMBRAPA, Guaraná convencional borda, 02 ° 53 ’ 42.18 ’’ S 59 ° 59 ’ 10.58 ’’ W, 23. XI. 2012, K. Schoeninger, malaise (1 ♀, INPA); Tefé, 03 ° 19 ’ 45 ’’ S 64 ° 41 ’ 13 ’’ W, 20. IX – 5. X. 2016, J. A. Oliveira, D. M. M. Mendes & J. A. Rafael, malaise (1 ♂); 1 – 5. XI. 2016, J. A. Oliveira, D. M. M. Mendes & J. A. Rafael, malaise (1 ♂, INPA); 10 – 23. X. 2016, J. A. Oliveira, D. M. M. Mendes & J. A. Rafael, malaise (1 ♂); Rondônia, Itapuã do Oeste, FLONA do Jamari, Trilha Pedra Grande, 09 ° 11 ’ 39.4 ’’ S 63 ° 04 ’ 55.3 ’’ W, 6. X. 2014, J. A. Rafael, F. F. Xavier-Filho, R. M. Vieira & R. H. Aquino, malaise (2 ♀, INPA); Rio Guapore, opp. Mouth Rio Baures (Bolivia), 26. IX. 1964, Bouseman and Lussenhop (1 ♂, MIUP). COLOMBIA: Magdalena, Parque Nacional Natural Tayrona, Zaino, 11 ° 20 ’ N 74 ° 2 ’ W, 50 m, 29. V – 14. VI. 200, Malaise, R. Henriquez, M. 241 (1 ♂, IAvH); Meta: Parque Nacional Natural Sierra de la Macarena, Caño Curia, Sendero Cachicamos, 3 ° 21 ’ N 73 ° 56 ’ W, 460 m, 10. XI – 21. XII. 2002, Malaise, M. Duarte, M. 2985 (1 ♂, IavH); Parque Nacional Natural Tinigua, Caño Nevera, 2 ° 11 ’ N 73 ° 48 ’ W, 390 m, 23. XII. 2002 – 7. I. 2003, Malaise, C. Sanchez, M. 3476 (2 ♂, IAvH, Fig. 54). COSTA RICA, Puntarenas, Est. Queb. Bonita, Res. Biol. Carara, 50 m, L-N- 194500, 469850, IV. 1993, J. C. Saborío (1 ♀, INBio). PERU, Madre de Dios, Reserva El Manu, Estac. Biol. Pakitza, 1 – 6. III. 1992, Malaise trap, R. Cambra (1 ♂, MIUP). TRINIDAD, W. I., Arima Valley, 800 – 1200 ft, 10 – 22. II. 1964, Rozen and Wygodzinsky (1 ♂, MIUP). VENEZUELA: Aragua, Rancho Grande, 1100 m, IV. 1987, Bordón (1 ♀, MIUP, Figs 10, 16).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	distribution	Distribution. This species is widespread in northern South America, including Bolivia, Brazil, Colombia, Ecuador, Peru, Trinidad, and Venezuela. First records for Peru and Trinidad.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C17CD2127B799EBFC92FAF3.taxon	discussion	Remarks. Females herein associated with L. hyphantria have been repeatedly and consistently collected in the same traps as males of L. hyphantria in different areas of the Brazilian Amazon (Roraima and Amazonas states). These females and the males of L. hyphantria are the only species of this genus widely distributed throughout the Brazilian Amazon region to be found so far. Additional females were also found in regions where L. hyphantria is the only recognized male species, such as Venezuela. Both males and females of this species show conspicuous variations related to where they occur. This is especially evident when comparing females collected in the Brazilian Amazon with ones from Venezuela wherein females from the first are much darker and have the variegated pattern overall less evident in comparison with Venezuelan specimens. Males from the Brazilian Amazon are also overall darker than males from further north in South America, in accordance to what was mentioned by Williams & Pitts (2009), even specimens from areas near the Brazilian-Venezuelan border (Roraima state). The main differences that can be added to the observations already made by Williams & Pitts (2009) is that specimens from Rondônia, Amazonas and Roraima states in Brazil have the legs and antennae almost completely black, the thick bristles of T 2 are black, mandibles are darker and the integument of the head is almost completely reddish-brown, except around the ocelli where it is black. The second northern Neotropical female specimen, mentioned above in the Remarks for L. chionothrix, cannot be readily separated from South American L. hyphantria specimens. Given that no males of L. hyphantria have been found in Central America, and that females are more difficult to differentiate than males, the distribution of L. hyphantria in Central America should be considered as tentative.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	description	(Figs 11, 17)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	diagnosis	Diagnosis. FEMALE. The following characters are diagnostic: the head and mesosoma are reddish-brown, the metasoma is blackish; the pronotal spiracle is scarcely swollen; the propodeal spiracle is sharply dentiform; the head is 1.1 × wider than the mesosoma; the lateral propodeal face has a few complete areolations; the mesosoma is elongate, with the thoracic dorsal length slightly less than its width; the mesosomal dorsum is tightly areolate without subparallel thickened setae; the T 1 shape is sub-disciform, often with dorsal and anterior faces separated by a weak carina; T 2 has distinct separated punctures, numerous subappressed black setae, and mostly smooth intervals; and the S 6 lateral carina is low angular. Body length 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	description	MALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	materials_examined	Material examined. Type material. Holotype of Lomachaeta ibarrai, ♀, ARGENTINA, Buenos Aires, Burzaco, XII. 1947, F. C. Sud (AMNH, Figs 11, 17). Other material. ARGENTINA: Buenos Aires, Glew, I. 1975, Carpintero (1 ♀, MIUP).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	distribution	Distribution. Known only from Buenos Aires Province, Argentina.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C14CD2727B79DFBFCEAFA69.taxon	discussion	Remarks. Based on the tight mesosomal sculpture, this species forms an apparent group with L. vianai and L. meloi, sp. nov., which are the only females known from southern South America. To this point, only one of those three species, L. vianai, has been associated with a male. The apparently disparate ranges of these three females, coupled with the morphological differences listed in the key below, make treating them as separate species the most reasonable hypothesis for now. Association of males with L. ibarrai and L. meloi or molecular sequence data might disprove or support the current taxonomic arrangement.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	description	(Fig. 61)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. ilex: the body is largely darkbrown to black, the tegulae are often orange; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is subsessile; the T 2 fringe is composed of simple setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 4 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	materials_examined	Material examined. USA, Nevada, Clark County, 4.5 mi SW of Boulder, 17. IX. 1997, Andrus, Griswold, and Messinger (1 ♂, paratype, CSCA, Fig. 61).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	distribution	Distribution. Mojave and western Sonoran Deserts in California and Nevada.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C12CD2627B79E7EFDBAFEBA.taxon	discussion	Remarks. This could be the sister species to L. hedera based on structural similarities in the mandible, T 2 fringe, and genitalia. It could be the opposite sex of L. warneri, given that they have overlapping distribution and are both apparently related to L. hedera.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	description	(Figs 56, 71, 72)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	diagnosis	Diagnosis. MALE. This species can be recognized by the following combination of characters: the body is black, except the obscure pale yellow T 1 apex; the mandible lacks a ventral tooth basally; the gena is weakly carinate; the head and pronotum are areolate; the forewing has veins encompassing the basal 0.7 × wing; the T 1 shape is disciform; the T 2 disc has dense small punctures; the T 2 fringe is composed of thick bristles; and the paramere is virtually straight except its downcurved apical third with a weak apical setal tuft and a few long setae along inner face medially. Body length 4.2 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	description	Description. MALE. Body length 4.2 mm. Coloration. Entire body black, except mandible, tegula, and trochanter partly orange-brown and T 1 apex obscurely pale yellow. Tibial spurs white. Wings pale brown, lighter basally, veins brown. Body setae whitish, except tergal fringes with pale yellow or blackish bristles and apical tergites with more or less extensive blackish setae. Head. Rounded posteriorly. Frons areolate-rugose, vertex punctate, gena areolate. Mandible tridentate apically, unarmed ventrally. Gena with weak ventral carina. Ocelli small; ocellocular distance 5 × length of lateral ocellus, interocellar distance 1.8 × lateral ocellar diameter. F 1 1.1 × pedicel length; F 2 1.7 × pedicel length. Mesosoma. Pronotum and mesoscutum areolate. Pronotum anterior and dorsal faces separated by complete carina. Tegula smooth with few punctures, margin setigerously punctate. Mesopleuron coarse areolate. Metapleuron smooth. Scutellum slightly convex, areolate. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face smooth anteriorly with areolations posteriorly. Forewing with veins encompassing basal 0.65 × wing; marginal cell as measured on costa 1.8 × stigma. Metasoma. T 1 shape nodose, with obscurely defined anterior and dorsal faces, anterior face convex. T 2 with coarse separated punctures, intervals smooth; T 2 – 4 fringes composed of thick pale yellow convergent bristles. S 2 punctures coarser and sparser than T 2 punctures. T 3 – 6 punctate. T 7 rugo-punctate. Hypopygium punctate, sharply bidentate posteriorly. Genitalia (Figs 71, 72). Paramere laterally subcompressed, straight in basal two thirds, apical third downcurved, acuminate apically, with three long (~ 2 × paramere width) setae along inner margin medially and small clump of setae at inner apex. Cuspis with posteroventral setae sparse and ~ 1.5 × cuspis length. Penis valve unidentate apically. FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	materials_examined	Material examined. Holotype, ♂, COLOMBIA, Magdalena, Parque Nacional Natural Tayrona, Neguanje, 11 ° 20 ’ N 74 ° 2 ’ W, 10 m, 9 – 17. IX. 2001, Malaise, R. Henriquez, M. 2137 (IAvH, Figs 56, 71, 72).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	distribution	Distribution. Known only from the type locality in Magdalena, Colombia.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	etymology	Etymology. KAW is happy to name this species in honor of his friend and colleague Juanita Rodriquez- Arrieta, in honor of her contributions to wasp systematics and in gratitude for her help with finding and borrowing the type specimen from the IAvH in 2011. Treat as a noun in apposition.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2627B79933FD87F979.taxon	discussion	Remarks. This is the third South American species based on males to be discovered. It shares its range with L. hyphantria, which has even been found in the same National Park (PNN Tayrona) but can be immediately differentiated from that species by the weak genal carina (strong in L. hyphantria) and the short marginal cell (~ 3 × stigma in L. hyphantria). It therefore seems closely related to L. vianai, but the nodose T 1 shape and downcurved paramere set it apart as a distinct species.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	description	(Fig. 63)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is elongate, gradually downcurving, cylindrical, and having an apical tuft of long setae. The following characters are also useful for diagnosis: the body is entirely blackish, except the sometimes reddish tegulae; the head has separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-sessile; the T 2 disc has coarse punctures; and the T 2 fringe is composed of simple setae. Body length 4 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	materials_examined	Material examined. USA: Arizona: Pima County, Vail Mountain Creek Ranch, 18 – 25. IV. 2006, 1100 m, malaise trap, M. E. Irwin (1 ♂, paratype, CSCA, Fig. 63).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	distribution	Distribution. Sonoran Desert in southern Arizona, USA and northern Sonora, Mexico.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C13CD2527B79F6EFCB1FD23.taxon	discussion	Remarks. The elongate, down-curving, cylindrical paramere shape is unique in Lomachaeta. This morphology is superficially similar to that of L. vacamuerta, except that species has shorter, straighter parameres. The length and curve, however, are similar to that of L. beadugrimi and L. snellingella, except those species have dorsoventrally flattened parameres without an apical setal tuft.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	description	(Figs 12, 18)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	diagnosis	Diagnosis. FEMALE. This species can be recognized by the coloration and setation: the entire body is mostly blackish except the head is orange-red; the body is clothed with distinct subappressed white setae. Additional useful diagnostic features include: the head is broader than the mesosoma; the mesosoma is elongate, with the thoracic dorsal length subequal to its width; the pronotal and propodeal spiracles are scarcely swollen; the T 1 shape is sub-disciform; and T 2 is densely punctate.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	description	Description. FEMALE. Body length 2.8 mm. Coloration. Body blackish except head and legs orange-red. Tibial spurs white. Body clothed predominantly with distinct subappressed white setae. Clypeus and lateral margins of metasoma with many erect setae black-brown. Fringes of T 2 – 5 composed of appressed white setae thickened, contrasting with integument. Head. Head width 1.4 × pronotal width. Frons, vertex and gena areolate, vertex with some intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Length of F 1 0.9 × pedicel length; F 2 0.7 × pedicel length. Mesosoma. Mesosoma elongate; dorsal thoracic length subequal to width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles scarcely swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-disciform, punctate. T 2 slightly longer than wide. Disc of T 2 densely punctate, intervals obscurely microreticulate. T 3 – 5 and S 2 – 5 with separated punctures, intervals microreticulate. T 6 convex. Incomplete lateral S 6 carina shallow but distinct. MALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	materials_examined	Material examined. Holotype, ♀, BRAZIL, São Paulo, Ribeirão Preto, Campus da USP, 13. X. 1999, G. A. R. Melo (DZUP, Figs 12, 18).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	distribution	Distribution. São Paulo State in Brazil.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	etymology	Etymology. Named in honor of Gabriel A. R. Melo, who collected the type specimen and has contributed to many other projects on Mutillidae and other Hymenoptera.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2527B79A8BFE56F979.taxon	discussion	Remarks. The short bristle-like setae covering this insect are unique in Lomachaeta. No males have yet been found in Southern Brazil.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	description	(Figs 4, 8)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	diagnosis	Diagnosis. FEMALE. This species can be recognized by the following combination of characters: the head and mesosoma are orange-brown and the legs and metasoma are lighter yellow-brown with T 2 having a brown transverse subapical cuticular band; the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the erect dorsal brachyplumose setae of the whole body are interspersed dark brown and yellow-brown; the mesosomal dorsum is elongate, coarsely areolate, and lacks appressed subparallel pale setae; the T 2 disc is sparsely punctate with the intervals obscurely microreticulate; the T 2 fringe is composed of simple silvery setae; and the S 6 lateral carina is shallow, but distinct.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	description	Description. FEMALE. Body length 3.1 mm. Coloration. Head and mesosoma orange-brown, legs and metasoma lighter yellow-brown; except F 2 – 10 brown, femoral and tibial apices obscurely brown; T 2 with transverse subapical brown band; and T 6 dark brown apically. Tibial spurs white. Erect dorsal brachyplumose setae interspersed pale yellow and darker brown; except T 6 with setae predominantly brown. Mesonotum with few scattered subappressed simple white setae; fringes of T 2 – 5 composed of subappressed pale golden or silvery setae, those of T 2 silvery, not thicker than erect T 2 disc setae. Ventral and pleural setae whitish, except S 6 setae largely brown. Head. Head width 1.2 × pronotal width. Frons areolate; vertex areolate anteriorly, smooth posteriorly; gena areolate dorsally, smooth ventrally. Clypeus without distinct teeth or ridges. Genal carina continuous to mandible base, subinterrupted medially by areolation margins. Mandible oblique, tapering, bidentate apically, with barely perceptible ventral lamella that does not interrupt ventral mandibular contour. Antennal scrobe without dorsal carina. Lengths of F 1 and F 2 each subequal to pedicel length. Mesosoma. Mesosoma elongate; thoracic dorsum length 0.92 × width. Humeral carina distinct, angulate dorsally, continuous to epaulet. Mesosomal dorsum areolate; intervals forming uniform carinae, appearing darker than surrounding cuticle; dorsally with 10 areolations between pronotal spiracles. Pronotal and propodeal spiracles scarcely swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-disciform, mostly smooth. T 2 as wide as long. Disc of T 2 with separated punctures, intervals microreticulate. T 3 – 5 and S 2 – 5 with separated weak punctures, intervals microreticulate. T 6 convex. Incomplete lateral S 6 carina shallow but distinct. MALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	materials_examined	Material examined. Holotype, ♀, USA, Arizona, Santa Cruz County, 2 mi. S Sonoita, malaise, 31 o 38 ’ N 111 o 39 ’ W, 5100 ’ [~ 1650 m], 22 – 30. VIII. 2010, E. E. Grissell (EMUS, Figs 4, 8).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	distribution	Distribution. Madrean Archipelago in Arizona.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	etymology	Etymology. From the Spanish osita “ little bear ”, based on colloquial reference to these small brown wasps as teddy bears by some authors.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C10CD2427B79F6EFB66FB4B.taxon	discussion	Remarks. Based on mesosomal shape and overall habitus, this species is similar to the newly associated female of L. vacamuerta, although it lacks the appressed subparallel setae on T 2. This similarity, coupled with distribution in southeastern Arizona suggests that this could be the female of L. litosisyra.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	description	(Fig. 60)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. polemomechana: the body is almost entirely black; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-sessile; the T 2 fringe is composed of simple setae; and the paramere is virtually straight, subcylindrical, and having evenly distributed short setae. Body length 4 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	materials_examined	Material examined. USA: Arizona: Santa Cruz County, Ruby Mountain, 20 km SSE Arivaca, 3 – 7. V. 2004, M. E. Irwin and F. D. Parker (1 ♂, EMUS, Fig. 60).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	distribution	Distribution. Southern Arizona, USA and northern Sonora, Mexico.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C11CD2427B79D1DFE94F8C9.taxon	discussion	Remarks. This species has only been found in transition zones between the Madrean Sky Islands and Chihuahuan Desert.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	description	(Figs 35, 39, 48)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for males of L. powelli: the body is entirely black, except T 2 – 3 are largely orange; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the forewing venation is restricted to the basal half of the wing; the T 1 shape is subsessile; the T 2 fringe is composed of simple setae; and the paramere is virtually straight, subcylindrical, and lacking tufts or rows of long setae. Body length: 3 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	description	FEMALE. This species can be recognized by the numerous elongate blackish dorsal bristles throughout the body. The following characters will also aid in diagnosis: the baso-ventral mandibular margin lacks a hyaline lamella; the head width is 1.2 × the mesosomal width; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the pronotal spiracle is weakly swollen; the lateral mesonotal tooth is indistinct; the propodeal spiracle is weakly swollen; the lateral propodeal face is impunctate; the mesosomal dorsum has about 10 areolations between the pronotal spiracles, many intervals are partially obliterated; the T 1 shape is sub-sessile; T 2 has sparse punctures and smooth intervals; and the S 6 lateral carina is low rounded. Body length 2 – 3 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	materials_examined	Material examined. USA: California: Fresno Co., Panoche Road, S 31 T 15 R 12 MO, 14. VI. 1982, R. F. Gill & N. J. Smith (2 ♀, CSCA); Monterey County, Arroyo Seco Camp, 15. VI. 1957, R. C. Bechtel (1 ♀, EMUS, Figs 35, 39); San Diego County, San Diego, Harkins Collection (1 ♀, EMUS); San Luis Obispo County, Pozo, 30. IV. 1962, J. Powell (1 ♂, paratype, CISC, Fig. 48).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	distribution	Distribution. Known only from the Central Valley and Coast Range areas of California, USA.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1CCD2827B798CBFDCBFD9A.taxon	discussion	Remarks. Both sexes of this species were collected in nesting aggregations of Diodontus occidentalis Fox, a ground-nesting pemphredonine wasp (Mickel 1964). Where known, the other North American species parasitize twig nesters (Pitts & Manley 2004).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	description	(Fig. 51)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. ptilohyalus: the body is entirely black, except T 2 – 3 are largely orange; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the mesoscutum has sparse punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-sessile; the T 2 fringe is composed of simple setae; and the paramere is virtually straight, laterally subcompressed, and having long setae ventrally throughout its length. Body length 4 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	materials_examined	Material examined. USA: Arizona: Maricopa County, Gila River, 10 km S. Arlington, malaise on sand beach, 200 m, 33 ° 13.3 ’ N 112 ° 45.53 ’ W, 25. V – 03. VI. 2010, M. E. Irwin (1 ♂, CSCA, Fig. 51).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	distribution	Distribution. This species has an apparently disjunct distribution in the Mojave and Sonoran Deserts (Arizona and California, (USA) and Balsas Dry Forests (Oaxaca, Mexico).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2827B79AD3FE6FFACB.taxon	discussion	Remarks. The holotype from Oaxaca, Mexico is separated from the other known specimens of L. ptilohyalus in hot deserts of the USA by over 2300 km. Further collections in Mexico may reveal other populations, or molecular data comparisons between Oaxacan and hot desert specimens may reveal that they are not conspecific. Either way, this is a compelling link between disjunct arid habitats of southern Pacific Mexico and the North American hot deserts.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	description	(Fig. 58)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. snellingella: the body is entirely blackish; the mandible lacks a ventral tooth basally; the head and T 2 disc have separated punctures; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is sub-sessile; the T 2 fringe is composed of simple setae; and the paramere is dorsoventrally flattened, apically rounded, and having short setae only. Body length 3 – 5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	materials_examined	Material examined. USA, California, San Diego County, Borrego, 30. IV. 1957, F. X. Williams (1 ♂, paratype, CASC, Fig. 58).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	distribution	Distribution. Western Sonoran Desert in California, USA.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1DCD2F27B79DE3FBC4FF72.taxon	discussion	Remarks. Lomachaeta snellingella is clearly a close relative of L. beadugrimi, given the nearly identical paramere shape. If females are found that key out to L. beadugrimi but occur in Imperial or San Diego Counties, those would be good candidates for the female of L. snellingella. If no recognizable differences were found between these females, that might provide evidence for synonymizing these taxa.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	description	(Fig. 57)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. theresa: the body is entirely blackish; the mandible lacks a ventral tooth basally; the gena is ecarinate; the head and T 2 disc have separated punctures; the forewing has its veins encompassing the basal 0.65 × of the wing; the T 1 shape is sub-sessile; the T 2 fringe is composed of thickened bristles; and the paramere is subcylindrical, virtually straight, and lacks long tufts of setae. Body length 5 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	description	FEMALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	materials_examined	Material examined. MEXICO: Sonora: 43 km E Alamos, Rancho San Pablo, 1 – 5. VI. 2007, M. E. Irwin (1 ♂, EMUS, Fig. 57).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	distribution	Distribution. Known only from Sonora, Mexico.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1ACD2E27B7997BFDC6FE76.taxon	discussion	Remarks. Among males of Lomachaeta, some of the most easily interpreted features for sorting the taxa are: 1) the presence or absence or a ventral mandibular tooth, 2) the presence or absence of thick bristles on the T 2 fringe, and 3) distribution in either North or South America. Lomachaeta theresa defies the functionality of some of these traits for establishing species-groups. Lomachaeta chionothrix, L. cirrhomeris, and L. hicksi have T 2 bristles like L. theresa, but have a ventral mandibular tooth, which L. theresa lacks. The remaining North American species lack a ventral mandibular tooth, like L. theresa, but also lack the T 2 bristles. The South American species, L. hyphantria and L. vianai, have the combination of unarmed mandible and T 2 bristles, but they differ from L. theresa in having the gena carinate.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	description	(Figs 2, 3, 6, 7, 64)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	diagnosis	Diagnosis. MALE. This species can be immediately recognized by the unique genitalia, wherein the paramere is straight, subcylindrical, and having an apical tuft of long setae. The following characters are also useful for diagnosis: the body is entirely blackish, except the sometimes reddish tegulae; the mandible is unarmed ventrally; the gena is ecarinate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is subsessile; the head and T 2 disc have separated punctures; and the T 2 fringe is composed of simple setae. Body length 5 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	description	FEMALE. This species can be recognized by the setae and mesosoma shape: the mesosomal dorsum and T 2 disc have matching thickened posteriorly-directed subparallel appressed pale golden setae and the mesosoma is elongate, with the thoracic dorsum nearly as long as wide. Additional useful diagnostic features include: the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the head is clearly broader than the mesosoma; the T 2 fringe is sparse and simple; and the S 6 lateral carina is raised, rounded. Description. FEMALE (hitherto unrecognized, based on female from Kennedy County). Body length 3 mm. Coloration. Head, mesosoma, antenna, legs, and T 6 predominantly dark brown; metasoma and bases of leg segments and antenna orange. Tibial spurs white. Body setae whitish yellow, except frons, mesonotum, and T 6 with many erect setae darkened, at least apically. Mesonotum and T 2 disc with long thick posteriorly-directed subparallel setae; fringes of T 2 – 3 composed of dense thick subappressed simple white setae; fringes of T 4 – 5 setae sparse, simple. Head. Head width 1.2 × pronotal width. Frons, vertex and gena areolate, vertex with some intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with shallow lamella baso-ventrally that does not interrupt the ventral mandible contour. Antennal scrobe without dorsal carina. Lengths of F 1 and F 2 each subequal to pedicel length. Mesosoma. Mesosoma elongate; dorsal thoracic length 0.95 × width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; intervals clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-disciform, punctate. T 2 slightly longer than wide. Disc of T 2 with dense oblique punctures, intervals smooth. T 3 – 5 and S 2 – 5 with separated punctures, intervals obscure microreticulate. T 6 convex. Incomplete lateral S 6 carina rounded, raised.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	materials_examined	Material examined. USA, Arizona, Pima County, 550 m, Silver Reef Wash, 4 km E. Vaiva Vo, Tat Momoli Mountains, malaise, 1 – 7. V. 2006, M. E. Irwin (1 ♂, EMUS, Fig. 64); Texas: Kennedy County, 2.5 mi. S Sarita, 18 – 20. X. 2002, B. Raber & E. Riley, pit-fall in sand (1 ♀, TAMU, Figs 3, 7); LaSalle County, Chaparral Wildlife Management Area, 13 mi. SW Cotulla, 28.293 ° N 99.383 ° W, 19. V. 2006, J. S. Wilson and K. A. Williams (2 ♀, CSCA EMUS, Figs 2, 6).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	distribution	Distribution. Widespread in the western Nearctic, including Arizona, California, New Mexico and Texas (USA) and Sonora (Mexico).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C1BCD2D27B79A4AFAB0FE52.taxon	discussion	Remarks. This widespread species has variably colored tegulae in males. In Arizona and California, this species has orange tegulae, resembling the sympatric L. cirrhomeris (Fig. 42), L. ilex (Fig. 61), and L. litosisyra (Fig. 63); farther East, the males are uniform blackish. The putative sex association presented here is based on a process of elimination. Lomachaeta hicksi and L. vacamuerta are the only Lomachaeta species known from southern Texas and the female of L. hicksi is distinct (in nearly all species-level characters) from the females described above. The female from Kennedy County (Fig. 3) has different coloration than the two females from LaSalle County (Fig. 2). It is the only Lomachaeta we have seen with a dark head and mesosoma that contrast with an orange metasoma, a pattern rarely seen in mutillids, except among Dasymutilla Ashmead and Pseudomethoca Ashmead from the Texan mimicry ring (Wilson et al., 2015). If natural selection has favored the similarities of this species to the Texan mimicry ring, various hypotheses could explain its limited mimetic fidelity. These include phylogenetic constraints of the genus Lomachaeta, the relaxed selection hypothesis based on its small size (Penney et al., 2012), or the community diversity hypothesis based on presence of multiple potential models (Wilson et al., 2013).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	description	(Figs 13, 14, 19, 20, 55)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	diagnosis	Diagnosis. MALE. The following combination of characters is diagnostic for L. vianai: the body is black, except the pale yellow T 1 apex; the mandible lacks a ventral tooth basally; the gena is weakly carinate; the head and pronotum are areolate; the forewing has its veins encompassing the basal 0.7 × of the wing; the T 1 shape is disciform; the T 2 disc has dense small punctures; the T 2 fringe is composed of thick bristles; and the paramere is virtually straight and has an weak apical setal tuft. Body length 4 – 6 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	description	FEMALE. The following characters are diagnostic: the head and mesosoma are dark reddish, the metasoma is blackish; the pronotal and propodeal spiracles are scarcely swollen; the head is 1.2 × wider than the mesosoma; the lateral propodeal face has a few complete areolations; the mesosoma is elongate, with the thoracic dorsum nearly as long as wide; the mesosomal dorsum is tightly areolate without subparallel thickened setae; the T 1 shape is subdisciform; T 2 has dense oblique punctures, mostly subappressed pale yellow setae, and mostly smooth intervals; and the S 6 lateral carina is low angular. Body length 2.5 – 4.5 mm.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	materials_examined	Material examined. Type material. Holotype of Lomachaeta vianai, ♀, ARGENTINA, Misiones, Dep. Concepcion, Santa María, X. 1956, M. J. Viana (AMNH, Figs 13, 19); Holotype of Lomachaeta megamicron, ♂, ARGENTINA, Salta, 8 km N La Viña, 26. X – 13. XI. 2003, M. E. Irwin & F. D. Parker coll. (EMUS). Other material. ARGENTINA: Salta: 8 km N La Viña, 26. X – 13. XI. 2003, M. E. Irwin & F. D. Parker (1 ♂, EMUS, Fig. 55); Chaco Occidental, Los Colorados, 100 km NE Joaquin V. Gonzales, 14. III. 1992, B. Bestelmeyer (1 ♀, MIUP); Pocitos, XI. 1978, Fritz (1 ♂, MIUP). BRAZIL, Goiás, Parque Nacional Chapada dos Veadeiros, A. P. Aguiar and B. F. dos Santos, Yellow Pan Trap: Ponto 1, 13 – 15. IX. 2005 (1 ♂, UFES); Ponto 3, 19 - 21. IX. 2005 (2 ♂, UFES); Ponto 6, 9 – 11. IX. 2005 (1 ♂, UFES); Ponto 10, 9 – 11. IX. 2005 (2 ♂, UFES); Ponto 15, IX. 2005 (1 ♂, UFES). PARAGUAY: Boquerón, Mariscal Estigarribia, Escuela Agricola, 174 m, 21 o 59 ’ 45 ” S 60 o 38 ’ 31 ” W, 5 – 7. X. 2003, B. Garcette, Malaise (1 ♂, MIUP); Presidente Hayes, Los Pioneros, 22.69014 o S 59.77186 o W, 9 – 11. X. 2003, M. LePonce, Pit Fall Trap (1 ♀, MIUP, Figs 14, 20).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	distribution	Distribution. Argentina (Catamarca, Misiones, and Salta Provinces), Brazil (Goiás), and Paraguay (Boquerón and Presidente Hayes Departments).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD2D27B79A1BFB09F95F.taxon	discussion	Remarks. Previously known from the unique type only, this species is now recognized as a widely distributed species in South America. Although this species is widespread, it is known from only seven localities. The sex association is based on a male and female collected in similar habitats (Chaco Boreal) in Paraguay.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	description	(Figs 36, 40)	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	diagnosis	Diagnosis. FEMALE. This species can be recognized by the following combination of characters: the body is predominantly orange-brown, except the antennae, legs, and T 3 – 6 are somewhat darker brown; the mandible has a barely perceptible ventral lamella that does not interrupt the ventral mandibular contour; the erect dorsal brachyplumose setae of the whole body are mostly yellow-brown; the mesosoma is coarsely areolate and lacks appressed subparallel pale setae; the mesosoma is compact, with the thoracic dorsal length 0.85 × its width; the T 2 disc is sparsely punctate with the intervals partly smooth and partly microreticulate; the T 2 fringe is somewhat dense with slightly thickened silvery setae; and the S 6 lateral carina is shallow, but distinct.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	description	Description. FEMALE. Body length 2.5 – 3 mm. Coloration. Body reddish-orange except F 2 – 10 dark brown; femora, tibiae, and tarsi largely brown; T 2 apex and T 3 – 5 brown; and T 6 dark brown. Tibial spurs white. Erect dorsal brachyplumose setae entirely white to pale brown, many setae lighter at base; except T 6 with setae predominantly brown. Mesonotum with few scattered subappressed simple white setae; fringes of T 2 – 5 composed of subappressed white setae, those of T 2 silvery, thicker than surrounding erect setae. Ventral and pleural setae white, except S 6 setae largely brown. Head. Head width 1.1 × pronotal width. Frons, vertex and gena areolate, vertex with many intervals obliterated. Clypeus without distinct teeth or ridges. Genal carina extending anteriorly nearly to hypostomal carina. Mandible oblique, tapering, bidentate apically, with barely perceptible ventral lamella that does not interrupt ventral mandibular contour. Antennal scrobe without dorsal carina. Lengths of F 1 and F 2 each subequal to pedicel length. Mesosoma. Mesosoma compact; dorsal thoracic length 0.85 × width. Humeral carina distinct, angulate dorsally, not reaching epaulet. Mesosomal dorsum areolate; areolations clearly defined, not raised into tubercles; areolations slightly tighter anteriorly than posteriorly; dorsally with 12 areolations between pronotal spiracles. Pronotal and propodeal spiracles weakly swollen; lateral mesonotal tubercle weakly defined. Lateral propodeal face impunctate. Metasoma. T 1 shape sub-sessile, mostly smooth. T 2 slightly longer than wide. Disc of T 2 with sparse punctures, intervals smooth. T 3 – 5 and S 2 – 5 with separated punctures, intervals shagreened. T 6 convex. Incomplete lateral S 6 carina shallow but distinct. MALE. Unknown.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	materials_examined	Material examined. Holotype, ♀, USA, Arizona, Coconino County, Hwy. 89 A, 4.9 mi. [7.8 km] ESE of jct. 389, 36 o 55 ’ 20 ” N 112 o 15 ’ 10 ” W, 24. IV – 25. V. 2014, W. B. Warner, barrier pitfall with black cups (CSCA, Figs 36, 40). Paratypes: USA: Arizona, Coconino County, Hwy. 89 A, 4.9 mi. [7.8 km] ESE of jct. 389, 36 o 55 ’ 20 ” N 112 o 15 ’ 10 ” W, 29. VI – 26. VII. 2014, W. B. Warner, barrier pitfall with black cups (2 ♀: ASUT, FSCA); Utah, Washington County, Mills Rd., 1.6 mi. [2.6 km] S I- 15 exit 27; 37.2649 o N 113.3259 o W, 15. IV – 3. VII. 2015, barrier pitfall with black cups (1 ♀, CSCA). Other Material. USA: Arizona, Pinal County, 4 mi. [6.4 km] W Stanfield, 12. IV. 1963, G. I. Stage (1 ♀, EMUS).	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	distribution	Distribution. Colorado Plateau in Arizona and Utah.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	etymology	Etymology. Named in honor of William B. Warner, the collector of the holotype and many other important mutillid specimens.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
03F187A07C18CD1227B79F10FC40FC5E.taxon	discussion	Remarks. This female apparently belongs to the L. crocopinna species-group (Williams & Pitts 2009) in having the ventral mandibular lamella obliterated and the pronotal and propodeal spiracles lowly produced. Within this apparent group, the thickened T 2 fringe setae are unique. The specimen from Pinal County Arizona has the cuticle somewhat darker, the dorsal setae longer and darker, and the intervals of T 2 with the microreticulations more distinct. Although these differences are not great enough to justify describing it as a distinct species at this point, they are enough to justify removing it from the paratype series.	en	Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes, Pitts, James P. (2019): Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations. Zootaxa 4564 (1): 101-136, DOI: 10.11646/zootaxa.4564.1.4
